455 lines
53 KiB
XML
455 lines
53 KiB
XML
<document id="4BCE42A6DF10850680EC661B7D4BB191" ID-DOI="10.1093/zoolinnean/zlae138" ID-ISSN="0024-4082" ID-ZooBank="681F645-F70D-4E1F-BF7F-D0251528BD51" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1734033805634" checkinUser="plazi" docAuthor="Lopes, Matheus Vieira, Pérez, Thierry & Klautau, Michelle" docDate="2024" docId="038F87F9FFF5FF99FEC5788B1D3FF817" docLanguage="en" docName="zlae138.pdf" docOrigin="Zoological Journal of the Linnean Society (zlae 138) 202 (3)" docSource="https://doi.org/10.1093/zoolinnean/zlae138" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Borojevia moana Lopes, Pérez & Klautau, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="FFB6FF81FFF3FF9EFFF379501853FFD2" masterDocTitle="Cave-dwelling calcareous sponges (Porifera: Calcarea) from the Marquesas Islands, French Polynesia" masterLastPageNumber="29" masterPageNumber="1" pageNumber="7" updateTime="1734544474156" updateUser="guilherme">
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<mods:titleInfo id="DC6EFB055AEE04E1D4DF1C38E5692598">
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<mods:title id="95CC9663DB0BDD7F5619BC3E83B346E8">Cave-dwelling calcareous sponges (Porifera: Calcarea) from the Marquesas Islands, French Polynesia</mods:title>
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<mods:name id="083D1DC139699698978D8853C349E011" type="personal">
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<mods:namePart id="ED5E9FB62792DD5A75E190BA8368DAD6">Lopes, Matheus Vieira</mods:namePart>
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<mods:affiliation id="D1206787B5BAA94A5E7A0CD36E47BDA0">Federal University of Rio de Janeiro, Biology Institute, Zoology Department, TaxoN Laboratory, Avenue Carlos Chagas Filho 791, CEP 21941 - 599, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:namePart id="01489644DF26C49523123B0297C76DD1">Pérez, Thierry</mods:namePart>
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<mods:affiliation id="246EE8BB49F9F3A9AC6563980B973348">Institut Méditerranéen de Biodiversité et d’Ecologie marine et continentale, CNRS, Aix-Marseille Université, IRD, Université d’Avignon, Station Marine d’Endoume, Chemin de la Batterie des Lions, 13007 Marseille, France</mods:affiliation>
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</mods:name>
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<mods:name id="BC5F6DE157968E978B19E6AB369FBF34" type="personal">
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<mods:roleTerm id="4BE40717C0BF760AFDAAF9FA4A5AA1E8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5932068EC650BA3C332F9C37C588C748">Klautau, Michelle</mods:namePart>
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<mods:affiliation id="22B104999722BE4AC87EF9CD772DD576">Federal University of Rio de Janeiro, Biology Institute, Zoology Department, TaxoN Laboratory, Avenue Carlos Chagas Filho 791, CEP 21941 - 599, Rio de Janeiro, Brazil</mods:affiliation>
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<mods:nameIdentifier id="9008A32A62881F6BA4F2804A10D8D7C0" type="email">mklautau@biologia.ufrj.br</mods:nameIdentifier>
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<mods:title id="969A69D2292C2E87C4F3BEBEBA6D2FAE">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="33AE9A61B0B5F870DDFD0DFFE27533A1">
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<mods:date id="82CF6B12120F1B267C603C12BF826ADF">2024</mods:date>
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<mods:title id="79A261830DAC8A5CF79732D0169FC10A">zlae 138</mods:title>
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<mods:number id="7F862422982A3768F764C4E47CD828A8">2024-11-06</mods:number>
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<mods:number id="9C897009848B2AA6AEBC63F0704D3A41">202</mods:number>
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<mods:number id="949E9CF3E7A5202335BF0BBC7E6D4B94">3</mods:number>
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<mods:url id="BB8691E12DACB59C69151654F723E001">https://doi.org/10.1093/zoolinnean/zlae138</mods:url>
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<mods:classification id="ACA8DA164C1322D06ED9FB046550865E">journal article</mods:classification>
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<mods:identifier id="25C641F5FAF7FDB10D2F564469513F1D" type="DOI">10.1093/zoolinnean/zlae138</mods:identifier>
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<treatment id="038F87F9FFF5FF99FEC5788B1D3FF817" LSID="urn:lsid:plazi:treatment:038F87F9FFF5FF99FEC5788B1D3FF817" httpUri="http://treatment.plazi.org/id/038F87F9FFF5FF99FEC5788B1D3FF817" lastPageId="7" lastPageNumber="7" pageId="6" pageNumber="7">
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<subSubSection id="C33C6564FFF5FF98FEC5788B1A07FE27" box="[310,596,475,502]" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="8B9936EFFFF5FF98FEC5788B1A07FE27" blockId="6.[310,596,475,502]" box="[310,596,475,502]" pageId="6" pageNumber="7">
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<heading id="D0D18183FFF5FF98FEC5788B1A07FE27" bold="true" box="[310,596,475,502]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
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<emphasis id="B952EAFDFFF5FF98FEC5788B1A07FE27" bold="true" box="[310,596,475,502]" pageId="6" pageNumber="7">
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||
<taxonomicName id="4C264D6CFFF5FF98FEC5788B19A4FE24" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[310,503,475,502]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
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<emphasis id="B952EAFDFFF5FF98FEC5788B19A4FE24" bold="true" box="[310,503,475,502]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A2615786FFF5FF98FE0F788B1A07FE27" box="[508,596,475,501]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33C6564FFF5FF98FE907B491A54FD43" pageId="6" pageNumber="7" type="description">
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||
<paragraph id="8B9936EFFFF5FF98FE907B491A74FDE1" blockId="6.[355,551,537,564]" box="[355,551,537,564]" pageId="6" pageNumber="7">
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||
<emphasis id="B952EAFDFFF5FF98FE907B491A74FDE1" bold="true" box="[355,551,537,564]" pageId="6" pageNumber="7">
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||
(
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<figureCitation id="131D2A6AFFF5FF98FE9D7B4A19F8FDE6" box="[366,427,538,564]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Figs 5</figureCitation>
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,
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<figureCitation id="131D2A6AFFF5FF98FE467B4A1990FDE6" box="[437,451,538,564]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">6</figureCitation>
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; Table 2)
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</emphasis>
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||
</paragraph>
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<paragraph id="8B9936EFFFF5FF98FF727B0A1A54FD43" blockId="6.[129,777,602,657]" pageId="6" pageNumber="7">
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<emphasis id="B952EAFDFFF5FF98FF727B0A1939FDA0" box="[129,362,602,626]" italics="true" pageId="6" pageNumber="7">Zoobank registration:</emphasis>
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<uri id="FFB73AEDFFF5FF98FE7B7B0A1A50FD43" pageId="6" pageNumber="7">
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||
urn:lsid:zoobank.org:act:
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||
<uuid id="FF800C3AFFF5FF98FD797B0A1A50FD43" pageId="6" pageNumber="7">7F25DF3B-A045-47C7-8EDC-ED25B5EFC60A</uuid>
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</uri>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6564FFF5FF98FF727BE41A1BFD18" box="[129,584,691,715]" pageId="6" pageNumber="7" type="type_taxon">
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||
<paragraph id="8B9936EFFFF5FF98FF727BE41A1BFD18" blockId="6.[129,584,691,715]" box="[129,584,691,715]" pageId="6" pageNumber="7">
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||
<emphasis id="B952EAFDFFF5FF98FF727BE418AFFD19" box="[129,252,692,715]" italics="true" pageId="6" pageNumber="7">
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<typeStatus id="549D884DFFF5FF98FF727BE418E2FD19" box="[129,177,692,715]" pageId="6" pageNumber="7">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C264D6CFFF5FF98FEF27BE31A11FD19" authority="Haeckel, 1872" authorityName="Haeckel" authorityYear="1872" box="[257,578,691,715]" class="Calcarea" family="Leucascidae" genus="Ascaltis" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="cerebrum">
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<emphasis id="B952EAFDFFF5FF98FEF27BE319F8FD19" box="[257,427,691,715]" italics="true" pageId="6" pageNumber="7">Ascaltis cerebrum</emphasis>
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||
<bibRefCitation id="EFB74B1EFFF5FF98FE437BE31A11FD19" author="Haeckel E" box="[432,578,691,715]" pageId="6" pageNumber="7" refId="ref16300" refString="Haeckel E. Die Kalkschwamme. Eine Monographie in zwei Banden Text und einem Atlas mit 60 Tafeln Abbildungen. Vols 1 - 3. Berlin: Reimer, 1872." type="book" year="1872">Haeckel, 1872</bibRefCitation>
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||
</taxonomicName>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33C6564FFF5FF98FF727BBD1945FC50" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph id="8B9936EFFFF5FF98FF727BBD1945FC50" blockId="6.[128,778,748,898]" pageId="6" pageNumber="7">
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<emphasis id="B952EAFDFFF5FF98FF727BBD18B4FCD6" box="[129,231,749,772]" italics="true" pageId="6" pageNumber="7">Diagnosis:</emphasis>
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White
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<taxonomicName id="4C264D6CFFF5FF98FEB17BBD19CDFCD6" box="[322,414,749,772]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="genus">
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<emphasis id="B952EAFDFFF5FF98FEB17BBD19CDFCD6" box="[322,414,749,772]" italics="true" pageId="6" pageNumber="7">Borojevia</emphasis>
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</taxonomicName>
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, cormus composed of regular and tightly anastomosed tubes. Multiple water-collecting tubes present. Skeleton composed of tripods, triactines, and tetractines. Apical actine of the tetractines either smooth or with five to nine rows of spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6564FFF5FF98FF727AF3193CFC28" pageId="6" pageNumber="7" type="etymology">
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||
<paragraph id="8B9936EFFFF5FF98FF727AF3193CFC28" blockId="6.[129,778,931,1018]" pageId="6" pageNumber="7">
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<emphasis id="B952EAFDFFF5FF98FF727AF318A3FC69" box="[129,240,931,955]" italics="true" pageId="6" pageNumber="7">Etymology:</emphasis>
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From ‘Moana’ (Ocean in Maori), for the oceanographic expedition Pakaihi i te Moana, which collected the sponges published here
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C6564FFF5FF98FF727D4C1A36FB18" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="8B9936EFFFF5FF98FF727D4C197EFB43" blockId="6.[129,778,1051,1169]" pageId="6" pageNumber="7">
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<emphasis id="B952EAFDFFF5FF98FF727D4C195CFBE1" box="[129,271,1051,1075]" italics="true" pageId="6" pageNumber="7">Type material:</emphasis>
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||
<materialsCitation id="3B4E3CB2FFF5FF98FEEF7D4B197EFB43" collectingDate="2012-01-25" collectorName="T. Perez" country="French Polynesia" elevation="8" latitude="-8.676666" location="Motu Iti Island" longLatPrecision="129" longitude="-140.62" pageId="6" pageNumber="7" specimenCode="UFRJPOR 7524" specimenCount="1" stateProvince="Marquesas Islands" typeStatus="holotype">
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<typeStatus id="549D884DFFF5FF98FEEF7D4B19D0FBE1" box="[284,387,1051,1075]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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||
:
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<collectingCountry id="F331767FFFF5FF98FE7E7D4B1A6DFBE1" box="[397,574,1051,1075]" name="French Polynesia" pageId="6" pageNumber="7">French Polynesia</collectingCountry>
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||
,
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<collectingRegion id="49E2F80DFFF5FF98FDBB7D4B1B54FBE1" box="[584,775,1051,1075]" country="French Polynesia" name="Iles Marquises" pageId="6" pageNumber="7">Marquesas Islands</collectingRegion>
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||
,
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||
<location id="8EF96034FFF5FF98FF727D6B1967FB80" LSID="urn:lsid:plazi:treatment:038F87F9FFF5FF99FEC5788B1D3FF817:8EF96034FFF5FF98FF727D6B1967FB80" box="[129,308,1082,1106]" country="French Polynesia" latitude="-8.676666" longLatPrecision="129" longitude="-140.62" name="Motu Iti Island" pageId="6" pageNumber="7" stateProvince="Marquesas Islands">Motu Iti Island</location>
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||
:
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||
<emphasis id="B952EAFDFFF5FF98FEB47D6B19A5FB80" box="[327,502,1082,1106]" italics="true" pageId="6" pageNumber="7">
|
||
<specimenCode id="DB809E94FFF5FF98FEB47D6B19A5FB80" box="[327,502,1082,1106]" pageId="6" pageNumber="7">UFRJPOR 7524</specimenCode>
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||
</emphasis>
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||
, lava tunnel (
|
||
<geoCoordinate id="EE125028FFF5FF98FD6F7D6A1B56FB80" box="[668,773,1081,1106]" degrees="8" direction="south" minutes="40.6" orientation="latitude" pageId="6" pageNumber="7" precision="92" value="-8.676666">
|
||
8°40.6
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||
<emphasis id="B952EAFDFFF5FF98FD127D691AB4FB80" box="[737,743,1081,1106]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||
S
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||
</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE125028FFF5FF98FF727D0A1956FBA0" box="[129,261,1112,1138]" degrees="140" direction="west" minutes="37.2" orientation="longitude" pageId="6" pageNumber="7" precision="92" value="-140.62">
|
||
140°37.2
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||
<emphasis id="B952EAFDFFF5FF98FF2F7D0818B1FBA3" box="[220,226,1112,1137]" italics="true" pageId="6" pageNumber="7">ʹ</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
),
|
||
<elevation id="000BD1DCFFF5FF98FEE47D0A19D1FBA0" box="[279,386,1114,1138]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="m" value="8.0">
|
||
<quantity id="4CDE9B0AFFF5FF98FEE47D0A196CFBA3" box="[279,319,1114,1138]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="7" unit="m" value="8.0">8 m</quantity>
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||
depth
|
||
</elevation>
|
||
,
|
||
<date id="FF98102FFFF5FF98FE7F7D0919BEFBA3" box="[396,493,1113,1137]" pageId="6" pageNumber="7" value="2012-01-25">
|
||
<collectingDate id="EFDCE9C7FFF5FF98FE7F7D0919BEFBA3" box="[396,493,1113,1137]" pageId="6" pageNumber="7" value="2012-01-25">25.i.2012</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26D35339FFF5FF98FDDB7D0A1A2FFBA0" box="[552,636,1114,1138]" pageId="6" pageNumber="7">T. Pérez</collectorName>
|
||
, field number MQ27-GR-TP2.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FF727DE21A36FB18" blockId="6.[129,613,1202,1226]" box="[129,613,1202,1226]" pageId="6" pageNumber="7">
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||
<emphasis id="B952EAFDFFF5FF98FF727DE21898FB18" box="[129,203,1202,1226]" italics="true" pageId="6" pageNumber="7">Colour:</emphasis>
|
||
White alive and in ethanol (
|
||
<figureCitation id="131D2A6AFFF5FF98FE027DE21A07FB18" box="[497,596,1202,1226]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5A, B</figureCitation>
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).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6564FFF5FF98FF727DBC1BC2FED4" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="8B9936EFFFF5FF98FF727DBC18B5F9CC" blockId="6.[128,779,1260,1566]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FF727DBC18A4FAD1" box="[129,247,1260,1283]" italics="true" pageId="6" pageNumber="7">Description:</emphasis>
|
||
Sponge massive, with cormus formed by regular and tightly anastomosed tubes (
|
||
<figureCitation id="131D2A6AFFF5FF98FE147C5B1A0BFAF1" box="[487,600,1291,1315]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5A–C</figureCitation>
|
||
). Outer surface smooth and, in some regions, folded. Various water-collecting tubes emerge from conical projections on the surface (
|
||
<figureCitation id="131D2A6AFFF5FF98FD427C1A1AAAFAB3" box="[689,761,1353,1378]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5B</figureCitation>
|
||
). On the thicker parts of the cormus, there is a space lined by the walls of internal tubes, giving the appearance of an atrial cavity, but lacking membrane around it (
|
||
<figureCitation id="131D2A6AFFF5FF98FE167CF81A62FA6D" box="[485,561,1447,1472]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5C</figureCitation>
|
||
). Aquiferous system asconoid. Skeleton typical of the genus, with rare tripods present on the surface of the external tubes, triactines, and tetractines (
|
||
<figureCitation id="131D2A6AFFF5FF98FF787F561885F9CF" box="[139,214,1541,1566]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FEA27F131A6BF98C" blockId="6.[128,778,1603,1760]" box="[337,568,1603,1630]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FEA27F131A6BF98C" box="[337,568,1603,1630]" italics="true" pageId="6" pageNumber="7">
|
||
Spicules (
|
||
<figureCitation id="131D2A6AFFF5FF98FE587F14198EF98F" box="[427,477,1603,1629]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">Fig.6</figureCitation>
|
||
; Table 2)
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FF727F3A18A1F932" blockId="6.[128,778,1603,1760]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FF727F3A1880F950" box="[129,211,1642,1666]" italics="true" pageId="6" pageNumber="7">Tripods:</emphasis>
|
||
Regular to subregular. Actines conical, stout, and undulated, with blunt to sharp tips. They resemble large triactines and are very rare (
|
||
<figureCitation id="131D2A6AFFF5FF98FEFC7FF8190BF912" box="[271,344,1704,1728]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">Fig. 6A</figureCitation>
|
||
). Size (
|
||
<emphasis id="B952EAFDFFF5FF98FE5A7FF91A1DF912" box="[425,590,1704,1728]" italics="true" pageId="6" pageNumber="7">UFRJPOR 7524</emphasis>
|
||
): 63.4 (±5.5)/8.4 (±1.1) µm.
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FF727E5218A0F88A" blockId="6.[128,777,1793,1880]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FF727E5218B4F8CB" box="[129,231,1794,1817]" italics="true" pageId="6" pageNumber="7">Triactines:</emphasis>
|
||
Regular. Actines slightly conical to conical, with blunt to sharp tips (
|
||
<figureCitation id="131D2A6AFFF5FF98FEE37E70190BF8EA" box="[272,344,1824,1848]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">Fig. 6B</figureCitation>
|
||
). Size (
|
||
<emphasis id="B952EAFDFFF5FF98FE5B7E711A1DF8EA" box="[424,590,1824,1848]" italics="true" pageId="6" pageNumber="7">UFRJPOR 7524</emphasis>
|
||
): 56.6 (±6.3)/6.8 (±1.0) µm.
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FF727E2A1BC2FED4" blockId="6.[129,778,1913,1968]" lastBlockId="6.[825,1474,144,262]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FF727E2A18A2F843" box="[129,241,1914,1937]" italics="true" pageId="6" pageNumber="7">Tetractines:</emphasis>
|
||
Regular. Actines slightly conical to conical, with blunt to sharp tips (
|
||
<figureCitation id="131D2A6AFFF5FF98FEA77EC81991F862" box="[340,450,1944,1968]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">Fig. 6C, D</figureCitation>
|
||
). Apical actine slightly conical, smooth, or with well-developed spines organized in five to nine rows (
|
||
<figureCitation id="131D2A6AFFF5FF98FC8879FF1B94FF15" box="[891,967,175,199]" captionStart="Figure 6" captionStartId="11.[115,180,1306,1330]" captionTargetBox="[148,1425,144,1279]" captionTargetId="figure-335@11.[148,1425,144,1279]" captionTargetPageId="11" captionText="Figure 6.Spicules of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, tripod. B, triactine.C, tetractine. D, two types of apical actines, one smooth (arrow head) and the other with spines (arrow)." pageId="6" pageNumber="7">Fig. 6D</figureCitation>
|
||
). It is straight and sharp. Size (
|
||
<emphasis id="B952EAFDFFF5FF98FAF979E01DE3FF15" box="[1290,1456,175,199]" italics="true" pageId="6" pageNumber="7">UFRJPOR 7524</emphasis>
|
||
): 56.9 (±5.1)/6.4 (±1.1) µm (basal); 53.1 (±9.2)/4.9 (±0.7) µm (apical).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6564FFF5FF98FCCA78771BCEFE8D" pageId="6" pageNumber="7" type="biology_ecology">
|
||
<paragraph id="8B9936EFFFF5FF98FCCA78771BCEFE8D" blockId="6.[825,1474,295,351]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA78771BDFFEED" box="[825,908,295,319]" italics="true" pageId="6" pageNumber="7">Ecology:</emphasis>
|
||
Large patch of the specimen was encrusting a boulder (
|
||
<figureCitation id="131D2A6AFFF5FF98FCB778171BDEFE8C" box="[836,909,326,351]" captionStart="Figure 5" captionStartId="10.[129,194,1277,1301]" captionTargetBox="[129,1473,144,1249]" captionTargetId="figure-297@10.[129,1473,144,1249]" captionTargetPageId="10" captionText="Figure 5. Morphology of the holotype of Borojevia moana sp. nov. (UFRJPOR 7524). A, specimens in situ encrusting the surface of a rock (no scale available). B, detail of the anastomosis of the holotype. C, section of a thicker part of the body evidencing the spaces between anastomosed tubes (*). D, tangential section of the body surface." pageId="6" pageNumber="7">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6564FFF5FF98FCCA78D01C2AFE65" pageId="6" pageNumber="7" type="distribution">
|
||
<paragraph id="8B9936EFFFF5FF98FCCA78D01C2AFE65" blockId="6.[825,1476,384,439]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA78D01C6AFE4A" box="[825,1081,384,408]" italics="true" pageId="6" pageNumber="7">Geographical distribution:</emphasis>
|
||
<materialsCitation id="3B4E3CB2FFF5FF98FBBA78D01C26FE65" location="Ecoregion" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Iles Marquises">
|
||
Currently known only from the
|
||
<typeStatus id="549D884DFFF5FF98FA6578D01D97FE4A" box="[1430,1476,384,408]" pageId="6" pageNumber="7">type</typeStatus>
|
||
locality.
|
||
<location id="8EF96034FFF5FF98FC7C78CF1BAFFE65" LSID="urn:lsid:plazi:treatment:038F87F9FFF5FF99FEC5788B1D3FF817:8EF96034FFF5FF98FC7C78CF1BAFFE65" box="[911,1020,415,439]" name="Ecoregion" pageId="6" pageNumber="7" stateProvince="Iles Marquises">Ecoregion</location>
|
||
:
|
||
<collectingRegion id="49E2F80DFFF5FF98FBF778F01C26FE65" box="[1028,1141,416,439]" country="French Polynesia" name="Iles Marquises" pageId="6" pageNumber="7">Marquesas</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33C6564FFF5FF99FCCA78891D3FF817" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
|
||
<paragraph id="8B9936EFFFF5FF98FCCA78891DD5FBD7" blockId="6.[825,1476,473,1969]" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA78891BC4FE23" box="[825,919,473,497]" italics="true" pageId="6" pageNumber="7">Remarks:</emphasis>
|
||
Considering the currently accepted
|
||
<taxonomicName id="4C264D6CFFF5FF98FAC278891B32FDC2" pageId="6" pageNumber="7">
|
||
<emphasis id="B952EAFDFFF5FF98FAC278891DDEFE22" box="[1329,1421,473,496]" italics="true" pageId="6" pageNumber="7">Borojevia</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, those with a cormus formed by regular and tightly anastomosed tubes, and skeleton composed of tripods, triactines and tetractines are:
|
||
<taxonomicName id="4C264D6CFFF5FF98FC2B7B671D09FD9D" authority="(Klautau et al., 1994)" baseAuthorityName="Klautau" baseAuthorityYear="1994" box="[984,1370,567,591]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="aspina">
|
||
<emphasis id="B952EAFDFFF5FF98FC2B7B671C29FD9C" box="[984,1146,567,590]" italics="true" pageId="6" pageNumber="7">Borojevia aspina</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FB797B671D1CFD9D" author="Klautau M & Sole-Cava AM & Borojevic R" box="[1162,1359,567,591]" pageId="6" pageNumber="7" pagination="367 - 75" refId="ref16850" refString="Klautau M, Sole-Cava AM, Borojevic R. Biochemical systematics of sibling sympatric species of Clathrina (Porifera: Calcarea). Biochemical Systematics and Ecology 1994; 22: 367 - 75. https: // doi. org / 10.1016 / 0305 - 1978 (94) 90027 - 2" type="journal article" year="1994">
|
||
Klautau
|
||
<emphasis id="B952EAFDFFF5FF98FB137B671D59FD9D" box="[1248,1290,567,591]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
., 1994
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D6CFFF5FF98FA957B671CC9FDBC" authority="(Sole-Cava et al., 1991)" baseAuthorityName="Sole-Cava" baseAuthorityYear="1991" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="brasiliensis">
|
||
<emphasis id="B952EAFDFFF5FF98FA957B671BF1FDBC" italics="true" pageId="6" pageNumber="7">Borojevia brasiliensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FC407B061CDCFDBC" author="Sole-Cava AM & Klautau M & Boury-Esnault N" box="[947,1167,598,622]" pageId="6" pageNumber="7" pagination="381 - 6" refId="ref18319" refString="Sole-Cava AM, Klautau M, Boury-Esnault N et al. Genetic evidence for cryptic speciation in allopatric populations of two cosmopolitan species of the calcareous sponge genus Clathrina. Marine Biology 1991; 111: 381 - 6." type="journal article" year="1991">
|
||
Solé-Cava
|
||
<emphasis id="B952EAFDFFF5FF98FBD37B071C19FDBC" box="[1056,1098,598,622]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
., 1991
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D6CFFF5FF98FB557B071BC3FD5F" authority="Klautau et al., 2016" authorityName="Klautau" authorityYear="2016" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="croatica">
|
||
<emphasis id="B952EAFDFFF5FF98FB557B071D07FDBC" box="[1190,1364,599,622]" italics="true" pageId="6" pageNumber="7">Borojevia croatica</emphasis>
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FAA87B061BC3FD5F" author="Klautau M & Imesek M & Azevedo F" pageId="6" pageNumber="7" pagination="1 - 52" refId="ref16956" refString="Klautau M, Imesek M, Azevedo F et al. Adriatic calcarean sponges (Porifera, Calcarea), with the description of six new species and a richness analysis. European Journal of Taxonomy 2016; 178: 1 - 52." type="journal article" year="2016">
|
||
Klautau
|
||
<emphasis id="B952EAFDFFF5FF98FA427B071B18FD5F" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
., 2016
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C264D6CFFF5FF98FC6F7B261D7CFD5F" authority="Fontana et al., 2018" authorityName="Fontana" authorityYear="2018" box="[924,1327,629,653]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="crystallina">
|
||
<emphasis id="B952EAFDFFF5FF98FC6F7B261C32FD5F" box="[924,1121,629,653]" italics="true" pageId="6" pageNumber="7">Borojevia crystallina</emphasis>
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FB947B261D7CFD5F" author="Fontana T & Condor-Lujan B & Azevedo F" box="[1127,1327,629,653]" pageId="6" pageNumber="7" pagination="331 - 69" refId="ref16026" refString="Fontana T, Condor-Lujan B, Azevedo F et al. Diversity and distribution patterns of calcareous sponges (subclass Calcinea) from Martinique. Zootaxa 2018; 4410: 331 - 69. https: // doi. org / 10.11646 / zootaxa. 4410.2.5" type="journal article" year="2018">
|
||
Fontana
|
||
<emphasis id="B952EAFDFFF5FF98FB327B261CB9FD5F" box="[1217,1258,629,653]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
|
||
., 2018
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B952EAFDFFF5FF98FA947B261C19FD7E" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C264D6CFFF5FF98FA947B261BCAFD7E" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="trispinata">Borojevia trispinata</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C264D6CFFF5FF98FC567BC51C19FD7E" box="[933,1098,661,684]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="aspina">Borojevia aspina</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C264D6CFFF5FF98FB717BC51D16FD7E" box="[1154,1349,661,684]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="trispinata">
|
||
<emphasis id="B952EAFDFFF5FF98FB717BC51D16FD7E" box="[1154,1349,661,684]" italics="true" pageId="6" pageNumber="7">Borojevia trispinata</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from our new species mainly by the spine pattern on the apical actines of tetractines. While the apical actine of
|
||
<taxonomicName id="4C264D6CFFF5FF98FCCA7BA31B8DFCD8" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[825,990,755,778]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA7BA31B8DFCD8" box="[825,990,755,778]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FC167BA31C61FCD8" box="[997,1074,755,778]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
varies from smooth to spiny, with long spines displayed in many rows, the apical actine of
|
||
<taxonomicName id="4C264D6CFFF5FF98FA957A431B2AFC9B" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="aspina">
|
||
<emphasis id="B952EAFDFFF5FF98FA957A431B2AFC9B" italics="true" pageId="6" pageNumber="7">Borojevia aspina</emphasis>
|
||
</taxonomicName>
|
||
is smooth or the spines are vestigial.
|
||
<taxonomicName id="4C264D6CFFF5FF98FAF07A621D97FC9B" box="[1283,1476,818,841]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="trispinata">
|
||
<emphasis id="B952EAFDFFF5FF98FAF07A621D97FC9B" box="[1283,1476,818,841]" italics="true" pageId="6" pageNumber="7">Borojevia trispinata</emphasis>
|
||
</taxonomicName>
|
||
has a single row with three spines only. The size and abundance of the tripods also differentiates
|
||
<taxonomicName id="4C264D6CFFF5FF98FB787A211D61FC5A" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[1163,1330,881,904]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FB787A211D61FC5A" box="[1163,1330,881,904]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FAC47A201DD6FC55" box="[1335,1413,880,903]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
[63.4 (±5.5)/8.4 (±1.1) µm; with rare tripods] from
|
||
<taxonomicName id="4C264D6CFFF5FF98FAE77AC01D92FC75" box="[1300,1473,912,935]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="croatica">
|
||
<emphasis id="B952EAFDFFF5FF98FAE77AC01D92FC75" box="[1300,1473,912,935]" italics="true" pageId="6" pageNumber="7">Borojevia croatica</emphasis>
|
||
</taxonomicName>
|
||
[102.6 (±10.0)/11.9 (±1.5) µm; with common tripods (
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FA817AFF1BE3FC34" author="Klautau M & Imesek M & Azevedo F" pageId="6" pageNumber="7" pagination="1 - 52" refId="ref16956" refString="Klautau M, Imesek M, Azevedo F et al. Adriatic calcarean sponges (Porifera, Calcarea), with the description of six new species and a richness analysis. European Journal of Taxonomy 2016; 178: 1 - 52." type="journal article" year="2016">
|
||
Klautau
|
||
<emphasis id="B952EAFDFFF5FF98FCCA7A9F1B23FC34" box="[825,880,974,998]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
)], and
|
||
<taxonomicName id="4C264D6CFFF5FF98FBFF7A9F1C8AFC34" box="[1036,1241,974,998]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="crystallina">
|
||
<emphasis id="B952EAFDFFF5FF98FBFF7A9F1C8AFC34" box="[1036,1241,974,998]" italics="true" pageId="6" pageNumber="7">Borojevia crystallina</emphasis>
|
||
</taxonomicName>
|
||
[169.2 (±15.5)/21.9 (±2.5) µm; with numerous tripods (
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FB597ABE1D38FBD7" author="Fontana T & Condor-Lujan B & Azevedo F" box="[1194,1387,1005,1029]" pageId="6" pageNumber="7" pagination="331 - 69" refId="ref16026" refString="Fontana T, Condor-Lujan B, Azevedo F et al. Diversity and distribution patterns of calcareous sponges (subclass Calcinea) from Martinique. Zootaxa 2018; 4410: 331 - 69. https: // doi. org / 10.11646 / zootaxa. 4410.2.5" type="journal article" year="2018">
|
||
Fontana
|
||
<emphasis id="B952EAFDFFF5FF98FAF77ABE1D61FBD7" box="[1284,1330,1005,1029]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
)].
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF98FCA67D5D1C44F927" blockId="6.[825,1476,473,1969]" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C264D6CFFF5FF98FCA67D5D1BADFBF6" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[853,1022,1037,1060]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FCA67D5D1BADFBF6" box="[853,1022,1037,1060]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FBFB7D5D1C0AFBF6" box="[1032,1113,1037,1060]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is morphologically different from
|
||
<taxonomicName id="4C264D6CFFF5FF98FCCA7D7D1BA1FB96" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[825,1010,1068,1092]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA7D7D1BA1FB96" box="[825,1010,1068,1092]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
by the encrusting growth form of the new species, spines in rows of five to nine, and frequently shorter spicules (Table 2), while
|
||
<taxonomicName id="4C264D6CFFF5FF98FBA87D3B1D4BFB51" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[1115,1304,1131,1155]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FBA87D3B1D4BFB51" box="[1115,1304,1131,1155]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
is tubular, has spines in rows of three, and has larger spicules. Nevertheless, in our molecular analysis (C-LSU),
|
||
<taxonomicName id="4C264D6CFFF5FF98FB4D7DFA1D3BFB13" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[1214,1384,1194,1217]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FB4D7DFA1D3BFB13" box="[1214,1384,1194,1217]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FA817DFA1D90FB13" box="[1394,1475,1194,1217]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
grouped, with low bootstrap (<60%), with
|
||
<taxonomicName id="4C264D6CFFF5FF98FAF97D991D91FB33" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[1290,1474,1225,1249]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FAF97D991D91FB33" box="[1290,1474,1225,1249]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F331767FFFF5FF98FC857DB81BB3FAD2" box="[886,992,1256,1280]" name="Seychelles" pageId="6" pageNumber="7">Seychelles</collectingCountry>
|
||
(ZMAPOR 12435). In the original description, authors mentioned morphological variation among the specimens of
|
||
<taxonomicName id="4C264D6CFFF5FF98FC2B7C771CC5FAED" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[984,1174,1319,1343]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FC2B7C771CC5FAED" box="[984,1174,1319,1343]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F331767FFFF5FF98FB147C771D17FAED" box="[1255,1348,1319,1343]" name="Maldives" pageId="6" pageNumber="7">Maldives</collectingCountry>
|
||
(
|
||
<typeStatus id="549D884DFFF5FF98FAA97C771DEDFAED" box="[1370,1470,1319,1343]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
:
|
||
<emphasis id="B952EAFDFFF5FF98FCCA7C161C52FA8C" box="[825,1025,1350,1374]" italics="true" pageId="6" pageNumber="7">RMNHPOR 10158</emphasis>
|
||
) and
|
||
<collectingCountry id="F331767FFFF5FF98FBB37C161CFFFA8C" box="[1088,1196,1350,1374]" name="Seychelles" pageId="6" pageNumber="7">Seychelles</collectingCountry>
|
||
, but they considered these differences as intraspecific variability (
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FB307C361B3EFA4F" author="Van Soest RWM & De Voogd NJ" pageId="6" pageNumber="7" pagination="1 - 160" refId="ref18644" refString="Van Soest RWM, De Voogd NJ. Calcareous sponges of the Western Indian ocean and red sea. Zootaxa 2018; 4426: 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest and De Voogd 2018</bibRefCitation>
|
||
). These differences are mainly in colour (the specimens of
|
||
<taxonomicName id="4C264D6CFFF5FF98FCC97CF51BA3FA6E" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[826,1008,1444,1468]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FCC97CF51BA3FA6E" box="[826,1008,1444,1468]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F331767FFFF5FF98FBC37CF41CDEFA6E" box="[1072,1165,1444,1468]" name="Maldives" pageId="6" pageNumber="7">Maldives</collectingCountry>
|
||
are white to cream, while the Seychelles’ specimen is yellow) and spicules (sagittal triactines and tetractines are present in the Maldives’ specimens and absent in the Seychelles’ specimen). Moreover, we noticed that the spines of the apical actines are in the middle of the actine in the
|
||
<typeStatus id="549D884DFFF5FF98FCC97F111BC5F98B" box="[826,918,1601,1625]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C264D6CFFF5FF98FC407F111C34F98B" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[947,1127,1601,1625]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FC407F111C34F98B" box="[947,1127,1601,1625]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
, while in the Seychelles’ specimen they are only on the distal portion of the actine (
|
||
<bibRefCitation id="EFB74B1EFFF5FF98FAC27F301C59F945" author="Van Soest RWM & De Voogd NJ" pageId="6" pageNumber="7" pagination="1 - 160" refId="ref18644" refString="Van Soest RWM, De Voogd NJ. Calcareous sponges of the Western Indian ocean and red sea. Zootaxa 2018; 4426: 1 - 160. https: // doi. org / 10.11646 / zootaxa. 4426.1.1" type="journal article" year="2018">Van Soest and De Voogd 2018: 30</bibRefCitation>
|
||
, 32). For that reason, the Seychelles’ specimen most likely represents a different species from
|
||
<taxonomicName id="4C264D6CFFF5FF98FA957FCF1B2FF904" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FA957FCF1B2FF904" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FC757FEE1B85F907" box="[902,982,1726,1749]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C264D6CFFF5FF98FBFD7FEF1C94F904" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[1038,1223,1726,1750]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FBFD7FEF1C94F904" box="[1038,1223,1726,1750]" italics="true" pageId="6" pageNumber="7">Borojevia tubulata</emphasis>
|
||
</taxonomicName>
|
||
. Hence, we assign it as
|
||
<taxonomicName id="4C264D6CFFF5FF98FCCA7F8E1C42F927" ID-CoL="MKQ7" authorityName="Van Soest and De Voogd" authorityYear="2018" box="[825,1041,1757,1781]" class="Calcarea" family="Clathrinidae" genus="Borojevia" isUncertain="true" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="tubulata">
|
||
<emphasis id="B952EAFDFFF5FF98FCCA7F8E1BC6F927" box="[825,917,1758,1781]" italics="true" pageId="6" pageNumber="7">Borojevia</emphasis>
|
||
aff.
|
||
<emphasis id="B952EAFDFFF5FF98FC4D7F8D1C42F927" box="[958,1041,1757,1781]" italics="true" pageId="6" pageNumber="7">tubulata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9936EFFFF5FF99FCA67FAD1D3FF817" blockId="6.[825,1476,473,1969]" lastBlockId="7.[810,1459,1840,1989]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
The most similar species to
|
||
<taxonomicName id="4C264D6CFFF5FF98FB8C7FAD1D74F8C6" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[1151,1319,1789,1812]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FB8C7FAD1D74F8C6" box="[1151,1319,1789,1812]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FAC37FAD1DD3F8C6" box="[1328,1408,1789,1812]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the Brazilian species
|
||
<taxonomicName id="4C264D6CFFF5FF98FC177E4D1CFDF8E6" ID-CoL="MKPW" box="[996,1198,1820,1844]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="brasiliensis">
|
||
<emphasis id="B952EAFDFFF5FF98FC177E4D1CFDF8E6" box="[996,1198,1820,1844]" italics="true" pageId="6" pageNumber="7">Borojevia brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
. However, the spines in the apical actine of the tetractines of
|
||
<taxonomicName id="4C264D6CFFF5FF98FB6A7E6C1D35F881" ID-CoL="MKPW" box="[1177,1382,1851,1875]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="brasiliensis">
|
||
<emphasis id="B952EAFDFFF5FF98FB6A7E6C1D35F881" box="[1177,1382,1851,1875]" italics="true" pageId="6" pageNumber="7">Borojevia brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
are very short, while in
|
||
<taxonomicName id="4C264D6CFFF5FF98FC217E0B1C24F8A0" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[978,1143,1883,1906]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF5FF98FC217E0B1C24F8A0" box="[978,1143,1883,1906]" italics="true" pageId="6" pageNumber="7">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF5FF98FB8F7E0B1C9BF8A0" box="[1148,1224,1883,1906]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
they are well developed. Besides, the actines of the triactines and tetractines of
|
||
<taxonomicName id="4C264D6CFFF5FF98FA947E2B1BF1F863" ID-CoL="MKPW" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="6" pageNumber="7" phylum="Porifera" rank="species" species="brasiliensis">
|
||
<emphasis id="B952EAFDFFF5FF98FA947E2B1BF1F863" italics="true" pageId="6" pageNumber="7">Borojevia brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
are conical with blunt tips, while in the new species they are slightly conical and blunt to sharp. Overall, spicules of
|
||
<taxonomicName id="4C264D6CFFF4FF99FF807E00196EF8B5" ID-CoL="MKPW" box="[115,317,1871,1895]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="7" pageNumber="8" phylum="Porifera" rank="species" species="brasiliensis">
|
||
<emphasis id="B952EAFDFFF4FF99FF807E00196EF8B5" box="[115,317,1871,1895]" italics="true" pageId="7" pageNumber="8">Borojevia brasiliensis</emphasis>
|
||
</taxonomicName>
|
||
are larger and thicker [
|
||
<typeStatus id="549D884DFFF4FF99FDEC7E1F1AD3F8B5" box="[543,640,1871,1895]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
of
|
||
<emphasis id="B952EAFDFFF4FF99FD6D7E0018A6F854" italics="true" pageId="7" pageNumber="8">
|
||
<taxonomicName id="4C264D6CFFF4FF99FD6D7E00188FF854" ID-CoL="MKPW" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="7" pageNumber="8" phylum="Porifera" rank="species" species="brasiliensis">Borojevia brasiliensis</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
tripods: 81.0 (±8.2)/11.0 (±1.7) µm; triactines: 78.2 (±10.6)/10.8 (±1.5) µm; tetractines: 75.3 (±10.0)/10.4 (±1.3) µm (basal); 36.4 (±9.4)/8.0 (±2.2) µm (apical) (
|
||
<bibRefCitation id="EFB74B1EFFF4FF99FD597EFD1BA4F89A" author="Klautau M & Valentine C" box="[682,1015,1840,1989]" pageId="7" pageNumber="8" pagination="1 - 62" refId="ref16803" refString="Klautau M, Valentine C. Revision of the genus Clathrina (Porifera, Calcarea). Zoological Journal of the Linnean Society 2003; 139: 1 - 62. https: // doi. org / 10.1046 / j. 0024 - 4082.2003.00063. x" type="journal article" year="2003">Klautau and Valentine 2003</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C264D6CFFF4FF99FBE27E601CEAF89A" authorityName="Lopes & Pérez & Klautau" authorityYear="2024" box="[1041,1209,1840,1864]" class="Calcarea" family="Clathrinidae" genus="Borojevia" kingdom="Animalia" order="Clathrinida" pageId="7" pageNumber="8" phylum="Porifera" rank="species" species="moana" status="sp. nov.">
|
||
<emphasis id="B952EAFDFFF4FF99FBE27E601CEAF89A" box="[1041,1209,1840,1864]" italics="true" pageId="7" pageNumber="8">Borojevia moana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2615786FFF4FF99FB317E601D40F89A" box="[1218,1299,1840,1864]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
—tripods: 63.4 (±5.5)/8.4 (±1.1) µm; triactines: 56.6 (±6.3)/6.8 (±1.0) µm; tetractines: 56.9 (±5.1)/6.4 (±1.1) µm (basal); 53.1 (±9.2)/4.9 (±0.7) µm (apical)]. Moreover, our molecular results clearly show that they are different evolutive lineages (
|
||
<figureCitation id="131D2A6AFFF4FF99FAFB7EFD1D17F817" box="[1288,1348,1965,1989]" captionStart="Figure 2" captionStartId="7.[113,178,1740,1764]" captionTargetBox="[115,1456,145,1710]" captionTargetId="figure-96@7.[114,1458,144,1712]" captionTargetPageId="7" captionText="Figure 2. Maximum likelihood phylogenetic tree of the C-LSU of Calcinea. Sequences generated in the present work are highlighted in bold. Bootstrap values are shown at the nodes (<means value below 60%). Tree rooted with Neoernsta, Clathrina, and Janusya." pageId="7" pageNumber="8">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="131D2A6AFFF4FF99FABD7EFD1D08F817" box="[1358,1371,1965,1989]" captionStart="Figure 3" captionStartId="8.[127,192,1644,1668]" captionTargetBox="[161,1441,144,1616]" captionTargetId="figure-106@8.[161,1441,144,1616]" captionTargetPageId="8" captionText="Figure 3. Maximum likelihood phylogenetic tree of the ITS of Calcinea.Sequences generated in the present work are highlighted in bold. Bootstrap values are shown at the nodes (<means value below 60%). Tree rooted with Neoernsta, Clathrina, and Janusya." pageId="7" pageNumber="8">3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |