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<document id="27F269F016E647FE0589CA4DD4481735" ID-DOI="10.11646/phytotaxa.375.4.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13728284" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725680695066" checkinUser="felipe" docAuthor="Tagane, Shuichiro, Dang, Van-Son, Souladeth, Phetlasy, Nagamasu, Hidetoshi, Toyama, Hironori, Naiki, Akiyo, Fuse, Kengo, Tran, Hop, Yang, Ceng-Jui, Prajaksood, Amornrat &amp; Yahara, Tetsukazu" docDate="2018" docId="039D87D0FF908D14FF45C940DF00F8C7" docLanguage="en" docName="phytotaxa.375.4.1.pdf" docOrigin="Phytotaxa 375 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.375.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Syzygium phoukhaokhouayense Soulad., Tagane &amp; Yahara. A 2018, sp. nov." docType="treatment" docVersion="2" lastPageNumber="253" masterDocId="FFA4FFA8FF968D12FFCDC80ADB71FFCF" masterDocTitle="Five new species of Syzygium (Myrtaceae) from Indochina and Thailand" masterLastPageNumber="260" masterPageNumber="247" pageNumber="253" updateTime="1725692588566" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="95405916D019B28D4053139A2A4264AE">Five new species of Syzygium (Myrtaceae) from Indochina and Thailand</mods:title>
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<mods:namePart id="7EC218632907BC1CEA22F6F439E17ED2">Tagane, Shuichiro</mods:namePart>
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<mods:namePart id="6811C945D8218477F84457F118D5C141">Dang, Van-Son</mods:namePart>
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<mods:namePart id="8381912FE197A8D945EEDC98482ED95D">Souladeth, Phetlasy</mods:namePart>
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<mods:affiliation id="995D2E39D6FE91A9C343CCBEBFFF128A">The Kyoto University Museum, Kyoto University, Yoshida Honmachi, Sakyo-ku, Kyoto, 606 - 8501, Japan.</mods:affiliation>
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<mods:namePart id="51C2AA5ABFA4FDD6D986B5DDBB8ED586">Toyama, Hironori</mods:namePart>
<mods:affiliation id="8A614B2E9543B359349164A6C92C733D">Center for Asian Conservation Ecology, Kyushu University, 744 Motooka, Fukuoka, 819 - 0395, Japan. &amp; Iriomote Station, Tropical Biosphere Research Center, University of the Ryukyus, 870 Uehara, Taketomi-cho, Yaeyama-gun, Okinawa, 907 - 1541 Japan.</mods:affiliation>
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<mods:namePart id="0207EA43BAFABBE2B0F6159CE953F3C8">Fuse, Kengo</mods:namePart>
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<mods:namePart id="51C8B69BD77AB64D1FF8B6DBC8DB5886">Yang, Ceng-Jui</mods:namePart>
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<mods:namePart id="86550288884D26A0133AE5ED7968B88E">Prajaksood, Amornrat</mods:namePart>
<mods:affiliation id="F8E3C89CFFFC6E9B66BE4970CEB0222A">Department of Biology, Khon Kaen University, Khon Kaen 40002, Thailand.</mods:affiliation>
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<mods:namePart id="71A960AFF7B662B078746C72BC7CA89C">Yahara, Tetsukazu</mods:namePart>
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<mods:date id="DEBA20898C6885F42CBF68845C88CDE7">2018</mods:date>
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<emphasis id="B940EAD4FF908D14FF45C940DA8DFEAA" bold="true" box="[136,508,330,357]" italics="true" pageId="6" pageNumber="253">Syzygium phoukhaokhouayense</emphasis>
Soulad., Tagane &amp; Yahara
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,
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</emphasis>
(
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).
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TYPE:—
<collectingCountry id="F3237656FF908D14FF38C965DA31FE47" box="[245,320,367,392]" name="Laos" pageId="6" pageNumber="253">LAOS</collectingCountry>
.
<collectingRegion id="49F0F824FF908D14FE84C965D950FE47" box="[329,545,367,392]" country="Laos" name="Viangchan" pageId="6" pageNumber="253">Vientiane Province</collectingRegion>
, Thoulakhom district, Ban Pa Paek, Phou Khao Khouay National Protected Area,
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,
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,
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elev.,
<date id="FF8A1006FF908D14FD5FC998D81DFE64" box="[658,876,402,427]" pageId="6" pageNumber="253" value="2017-12-26">26 December 2017</date>
,
<emphasis id="B940EAD4FF908D14FCBAC998D9ECFE01" italics="true" pageId="6" pageNumber="253">Yahara T., Tagane S., Souladeth P., Nagamasu H., Naiki A., Chayer S. &amp; Kongxaisavath D. L1827</emphasis>
[fl.] (
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FOF!,
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BKF, KAG, KYO!, NHL!, P).
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<taxonomicName id="4C344D45FF908D14FF70C9D3D951FE3D" ID-CoL="546LQ" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[189,544,473,498]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FF70C9D3D951FE3D" box="[189,544,473,498]" italics="true" pageId="6" pageNumber="253">Syzygium phoukhaokhouayense</emphasis>
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is similar to
<taxonomicName id="4C344D45FF908D14FD7AC9D3DE0EFE3D" ID-CoL="546VZ" authority="(Miquel 1855: 431) Merrill &amp; Perry (1938: 109)" authorityName="Merrill &amp; Perry" authorityPageNumber="109" authorityYear="1938" baseAuthorityName="Miquel" baseAuthorityPageNumber="431" baseAuthorityYear="1855" box="[695,1407,473,498]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="syzygioides">
<emphasis id="B940EAD4FF908D14FD7AC9D3D825FE3D" box="[695,852,473,498]" italics="true" pageId="6" pageNumber="253">S. syzygioides</emphasis>
(
<bibRefCitation id="EFA54B37FF908D14FCAEC9D3DF40FE3D" author="Miquel, F. A. W." box="[867,1073,473,498]" pageId="6" pageNumber="253" refId="ref7958" refString="Miquel, F. A. W. (1855) Flora van Nederlandsch Indie. Vol. 1. Fleischer, Leipzig, 1116 pp. https: // doi. org / 10.5962 / bhl. title. 93" type="book" year="1855">Miquel 1855: 431</bibRefCitation>
)
<bibRefCitation id="EFA54B37FF908D14FB8CC9D3DE0EFE3D" author="Merrill, E. D. &amp; Perry, L. M." box="[1089,1407,473,498]" pageId="6" pageNumber="253" pagination="99 - 116" refId="ref7876" refString="Merrill, E. D. &amp; Perry, L. M. (1938) On the Indo-Chinese species of Syzygium Gaertner. Journal of Arnold Arboretum 19: 99 - 116." type="journal article" year="1938">Merrill &amp; Perry (1938: 109)</bibRefCitation>
</taxonomicName>
but differs in having coriaceous leaves (vs. chartaceous to subcoriaceous in
<taxonomicName id="4C344D45FF908D14FC66C9F6DF36FDDA" ID-CoL="546VZ" authorityName="Merrill &amp; Perry" authorityYear="1938" baseAuthorityName="Miquel" baseAuthorityYear="1855" box="[939,1095,508,533]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="syzygioides">
<emphasis id="B940EAD4FF908D14FC66C9F6DF36FDDA" box="[939,1095,508,533]" italics="true" pageId="6" pageNumber="253">S. syzygioides</emphasis>
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), longer petiole (
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long vs.
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long), larger and reddish-purple hypanthium (
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long vs. ca.
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long, greenish), longer styles (ca.
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long vs. ca.
<quantity id="4CCC9B23FF908D14FE47CA48DA91FD94" box="[394,480,578,603]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="6" pageNumber="253" unit="mm" value="8.3">8.3 mm</quantity>
long) and fewer ovules per locule in ovary (45 per locule in
<taxonomicName id="4C344D45FF908D14FB53CA48DEDAFD94" ID-CoL="546LQ" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[1182,1451,578,603]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FB53CA48DEDAFD94" box="[1182,1451,578,603]" italics="true" pageId="6" pageNumber="253">S. phoukhaokhouayense</emphasis>
</taxonomicName>
vs. 1014 per locule in
<taxonomicName id="4C344D45FF908D14FE40CA6FD958FDB1" ID-CoL="546VZ" authorityName="Merrill &amp; Perry" authorityYear="1938" baseAuthorityName="Miquel" baseAuthorityYear="1855" box="[397,553,613,638]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="syzygioides">
<emphasis id="B940EAD4FF908D14FE40CA6FD958FDB1" box="[397,553,613,638]" italics="true" pageId="6" pageNumber="253">S. syzygioides</emphasis>
</taxonomicName>
). Also, it is apparently similar to
<taxonomicName id="4C344D45FF908D14FC53CA6FDA3CFD6D" ID-CoL="7BBBB" authority="(De Candolle 1828: 287) Merrill &amp; Perry (1938: 109)" authorityName="Merrill &amp; Perry" authorityPageNumber="109" authorityYear="1938" baseAuthorityName="De Candolle" baseAuthorityPageNumber="287" baseAuthorityYear="1828" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="lineatum">
<emphasis id="B940EAD4FF908D14FC53CA6FDF6DFDB1" box="[926,1052,613,638]" italics="true" pageId="6" pageNumber="253">S. lineatum</emphasis>
(
<bibRefCitation id="EFA54B37FF908D14FBE7CA6FDE40FDB1" author="De Candolle, A." box="[1066,1329,613,638]" pageId="6" pageNumber="253" refId="ref7307" refString="De Candolle, A. P (1828) Prodromus Systematis Naturalis Regni Vegetabilis. Vol. 3. Treuttel &amp; Wurts, Paris, 745 pp." type="book" year="1828">De Candolle 1828: 287</bibRefCitation>
)
<bibRefCitation id="EFA54B37FF908D14FA8DCA6FDA3CFD6D" author="Merrill, E. D. &amp; Perry, L. M." pageId="6" pageNumber="253" pagination="99 - 116" refId="ref7876" refString="Merrill, E. D. &amp; Perry, L. M. (1938) On the Indo-Chinese species of Syzygium Gaertner. Journal of Arnold Arboretum 19: 99 - 116." type="journal article" year="1938">Merrill &amp; Perry (1938: 109)</bibRefCitation>
</taxonomicName>
but easily distinguished by having more secondary veins (2832 pairs in
<taxonomicName id="4C344D45FF908D14FBB4CA83DEF4FD6D" ID-CoL="546LQ" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[1145,1413,649,674]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FBB4CA83DEF4FD6D" box="[1145,1413,649,674]" italics="true" pageId="6" pageNumber="253">S. phoukhaokhouayense</emphasis>
</taxonomicName>
vs. 1620 pairs in
<taxonomicName id="4C344D45FF908D14FEE2CAA6DADCFD0A" ID-CoL="7BBBB" authorityName="Merrill &amp; Perry" authorityYear="1938" baseAuthorityName="De Candolle" baseAuthorityYear="1828" box="[303,429,684,709]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="lineatum">
<emphasis id="B940EAD4FF908D14FEE2CAA6DADCFD0A" box="[303,429,684,709]" italics="true" pageId="6" pageNumber="253">S. lineatum</emphasis>
</taxonomicName>
) and single intramarginal veins (vs. 2).
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CAC5DA2EFB7D" blockId="6.[136,1452,367,1800]" pageId="6" pageNumber="253">
Tree,
<quantity id="4CCC9B23FF908D14FECCCAC5DA5FFD27" box="[257,302,719,744]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="253" unit="m" value="6.0">6 m</quantity>
tall. Young twigs terete, reddish brown, glabrous,
<quantity id="4CCC9B23FF908D14FCB6CAC5D881FD27" box="[891,1008,719,744]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="6" pageNumber="253" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">22.5 mm</quantity>
in diam., old twigs grayish to reddish brown, internode (1)
<quantity id="4CCC9B23FF908D14FEB5CAF8DA97FCC4" box="[376,486,754,779]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.6" metricValueMin="2.0" pageId="6" pageNumber="253" unit="cm" value="2.8" valueMax="3.6" valueMin="2.0">23.6 cm</quantity>
long. Leaves opposite, blade elliptic to elliptic-oblong, 4.28.9 ×
<quantity id="4CCC9B23FF908D14FB01CAF8DE3DFCC4" box="[1228,1356,754,779]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="6" pageNumber="253" unit="cm" value="2.5" valueMax="3.2" valueMin="1.8">1.83.2 cm</quantity>
, 2.33.1 times longer than wide, coriaceous, reddish brown dark grayish brown adaxially, light yellowish brown abaxially, glabrous on both surfaces, apex acuminate, acumen to
<quantity id="4CCC9B23FF908D14FD33CB33D846FC9D" box="[766,823,825,850]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="253" unit="cm" value="1.0">1 cm</quantity>
long, base cuneate to attenuate, margin entire, slightly recurved when dry, midrib sunken abaxially, prominent and distinct adaxially, secondary veins 2832 pairs,
<quantity id="4CCC9B23FF908D14FA95CB56DBC5FC57" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="6" pageNumber="253" unit="mm" value="2.15" valueMax="2.8" valueMin="1.5">1.52.8 mm</quantity>
apart, slightly prominent on both surfaces, intersecondary veins distinct, intramarginal veins 1, ca.
<quantity id="4CCC9B23FF908D14FADACB75DE1FFC57" box="[1303,1390,895,920]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="253" unit="mm" value="0.8">0.8 mm</quantity>
from margin; petiole
<quantity id="4CCC9B23FF908D14FEFACBA8DAC2FC74" box="[311,435,930,955]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="6" pageNumber="253" unit="cm" value="0.95" valueMax="1.2" valueMin="0.7">0.71.2 cm</quantity>
long, concave adaxially, rounded abaxially, blackish, glabrous. Inflorescences terminal and axillary near the top of branch, paniculate,
<quantity id="4CCC9B23FF908D14FDB9CBCFD983FC11" box="[628,754,965,990]" metricMagnitude="-2" metricUnit="m" metricValue="3.15" metricValueMax="3.8" metricValueMin="2.5" pageId="6" pageNumber="253" unit="cm" value="3.15" valueMax="3.8" valueMin="2.5">2.53.8 cm</quantity>
long, 840 flowered, peduncle up to
<quantity id="4CCC9B23FF908D14FB6DCBCFDF9FFC11" box="[1184,1262,965,990]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="253" unit="cm" value="2.7">2.7 cm</quantity>
long, rachis and lateral branches compressed or slightly angled; bracteoles triangular, ca.
<quantity id="4CCC9B23FF908D14FC03CBE3DF56FBCD" box="[974,1063,1001,1026]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="253" unit="mm" value="0.9">0.9 mm</quantity>
long, glabrous, caducous. Flower buds ca.
<quantity id="4CCC9B23FF908D14FF27CC06DA31FBEA" box="[234,320,1036,1061]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="253" unit="mm" value="2.1">2.1 mm</quantity>
in diam., whitish to purplish (
<emphasis id="B940EAD4FF908D14FD5FCC06D9ACFBEA" box="[658,733,1036,1061]" italics="true" pageId="6" pageNumber="253">in vivo</emphasis>
). Hypanthium funnel-shaped or campanulate,
<quantity id="4CCC9B23FF908D14FB2BCC06DE1DFBEA" box="[1254,1388,1036,1061]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="6" pageNumber="253" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.14.5 mm</quantity>
long, blackish when dry, glabrous, pseudostipe ca.
<quantity id="4CCC9B23FF908D14FD41CC25D9BEFB87" box="[652,719,1071,1096]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="253" unit="mm" value="2.0">2 mm</quantity>
long. Sepals 4, depressed semi-circular, 0.5 ×
<quantity id="4CCC9B23FF908D14FB2BCC25DE4EFB87" box="[1254,1343,1071,1096]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="253" unit="mm" value="1.6">1.6 mm</quantity>
, petals 4, orbicular, ca.
<quantity id="4CCC9B23FF908D14FEE9CC58DAE9FBA4" box="[292,408,1106,1131]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="3.0" metricValueMin="2.4" pageId="6" pageNumber="253" unit="mm" value="2.7" valueMax="3.0" valueMin="2.4">2.43 mm</quantity>
in diam., membranous, gland dot lacking or 1 per petal. Stamens 3032,
<quantity id="4CCC9B23FF908D14FB2FCC58DE1AFBA4" box="[1250,1387,1106,1131]" metricMagnitude="-3" metricUnit="m" metricValue="6.6499999999999995" metricValueMax="8.2" metricValueMin="5.1" pageId="6" pageNumber="253" unit="mm" value="6.6499999999999995" valueMax="8.2" valueMin="5.1">5.18.2 mm</quantity>
long, anthers ca.
<quantity id="4CCC9B23FF908D14FEC8CC7FDA2BFB41" box="[261,346,1141,1166]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="253" unit="mm" value="0.4">0.4 mm</quantity>
long, filaments white (
<emphasis id="B940EAD4FF908D14FD96CC7FD9D7FB41" box="[603,678,1141,1166]" italics="true" pageId="6" pageNumber="253">in vivo</emphasis>
). Style ca.
<quantity id="4CCC9B23FF908D14FCEFCC7FD809FB41" box="[802,888,1141,1166]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="6" pageNumber="253" unit="mm" value="8.3">8.3 mm</quantity>
long. Ovary 2-locular, ovules 45 per locule. Fruit and seeds not seen.
</paragraph>
</subSubSection>
<subSubSection id="C32E654DFF908D14FF70CCB6DF91FAF1" pageId="6" pageNumber="253" type="distribution">
<paragraph id="8B8B36C6FF908D14FF70CCB6DFBCFB1A" blockId="6.[136,1452,367,1800]" box="[189,1229,1212,1237]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CCB6DA04FB1A" bold="true" box="[189,373,1212,1237]" pageId="6" pageNumber="253">Distribution:—</emphasis>
<collectingCountry id="F3237656FF908D14FEB8CCB6DADAFB1A" box="[373,427,1212,1237]" name="Laos" pageId="6" pageNumber="253">Laos</collectingCountry>
(so far only known from Phou Khao Khouay National Protected Area).
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CCD5D808FB37" blockId="6.[136,1452,367,1800]" box="[189,889,1247,1272]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CCD5DAA9FB37" bold="true" box="[189,472,1247,1272]" pageId="6" pageNumber="253">Habitat and Ecology:—</emphasis>
Open pine forest, at
<quantity id="4CCC9B23FF908D14FD71CCD5D872FB37" box="[700,771,1247,1272]" metricMagnitude="2" metricUnit="m" metricValue="9.05" pageId="6" pageNumber="253" unit="m" value="905.0">905 m</quantity>
elevation.
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CD08D8E4FAD4" blockId="6.[136,1452,367,1800]" box="[189,917,1282,1307]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CD08DA2EFAD4" bold="true" box="[189,351,1282,1307]" pageId="6" pageNumber="253">Phenology:—</emphasis>
Flowering specimens were collected in December.
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CD2FDF91FAF1" blockId="6.[136,1452,367,1800]" box="[189,1248,1317,1342]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CD2FDAFEFAF1" bold="true" box="[189,399,1317,1342]" pageId="6" pageNumber="253">DNA barcodes:—</emphasis>
LC381851 (
<emphasis id="B940EAD4FF908D14FDD5CD2FD93CFAF1" box="[536,589,1317,1342]" italics="true" pageId="6" pageNumber="253">rbcL</emphasis>
), LC381854 (
<emphasis id="B940EAD4FF908D14FD21CD2FD858FAF1" box="[748,809,1317,1342]" italics="true" pageId="6" pageNumber="253">matK</emphasis>
); sequenced from
<emphasis id="B940EAD4FF908D14FC37CD2FDFA8FAF1" box="[1018,1241,1317,1342]" italics="true" pageId="6" pageNumber="253">Tagane et al. L1827</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C32E654DFF908D14FF70CD43DF00F8C7" pageId="6" pageNumber="253" type="etymology">
<paragraph id="8B8B36C6FF908D14FF70CD43DA44FA4A" blockId="6.[136,1452,367,1800]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CD43DA15FAAD" bold="true" box="[189,356,1353,1378]" pageId="6" pageNumber="253">Etymology:—</emphasis>
The species epithet refers to the geographical location of the find, Phou Khao Khouay National Protected Area.
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CD85D9E7FA67" blockId="6.[136,1452,367,1800]" box="[189,662,1418,1448]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CD85D971FA67" bold="true" box="[189,512,1418,1448]" pageId="6" pageNumber="253">Vernacular name:—ຫວາພ</emphasis>
(Wa Phou).
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CDB8D8DBF9B4" blockId="6.[136,1452,367,1800]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CDB8D9E6FA04" bold="true" box="[189,663,1458,1483]" pageId="6" pageNumber="253">Preliminary conservation assessment:—</emphasis>
Critically Endangered (CR).At present,
<taxonomicName id="4C344D45FF908D14FB81CDB8DEDAFA04" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[1100,1451,1458,1483]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FB81CDB8DEDAFA04" box="[1100,1451,1458,1483]" italics="true" pageId="6" pageNumber="253">Syzygium phoukhaokhouayense</emphasis>
</taxonomicName>
is known from only a single individual in the Phou Khao Khouay National Protected Area, at
<quantity id="4CCC9B23FF908D14FB71CDDFDE75FA21" box="[1212,1284,1493,1518]" metricMagnitude="2" metricUnit="m" metricValue="9.05" pageId="6" pageNumber="253" unit="m" value="905.0">905 m</quantity>
elev. The area is close to the
<collectingRegion id="49F0F824FF908D14FEE6CDF3DAE7F9DD" box="[299,406,1529,1554]" country="Laos" name="Viangchan" pageId="6" pageNumber="253">Vientiane</collectingRegion>
Capital, and has been relatively intensively surveyed by many botanists (e.g.
<bibRefCitation id="EFA54B37FF908D14FACBCDF3DBB2F9FA" author="Newman, M. &amp; Thomas, P. &amp; Lanorsavah, S. &amp; Ketphanh, S. K. &amp; Svengsuka, B. &amp; Lamxay, V." pageId="6" pageNumber="253" pagination="222 - 251" refId="ref8242" refString="Newman, M., Thomas, P., Lanorsavah, S., Ketphanh, S. K., Svengsuka, B. &amp; Lamxay, V. (2007) New records of angiosperms and pteridophytes in the flora of Laos. Edinburgh Journal of Botany 64 (2): 222 - 251. https: // doi. org / 10.1017 / S 0960428607000923" type="journal article" year="2007">
Newman
<emphasis id="B940EAD4FF908D14FABECDF3DEDDF9DD" box="[1395,1452,1529,1554]" italics="true" pageId="6" pageNumber="253">et al.</emphasis>
2007
</bibRefCitation>
,
<bibRefCitation id="EFA54B37FF908D14FF00CE16D963F9FA" author="Souladeth, P. &amp; Meesawat, A." box="[205,530,1564,1589]" pageId="6" pageNumber="253" pagination="927 - 931" refId="ref8794" refString="Souladeth, P. &amp; Meesawat, A. (2012) Myrtaceae in Phou Khao Khouay National Protected Area, Lao PDR. Proceedings: 1 st ASEAN Plus Three Graduate Research Congress 2012: 927 - 931." type="journal article" year="2012">Souladeth &amp; Meesawat 2012</bibRefCitation>
,
<bibRefCitation id="EFA54B37FF908D14FDD1CE16D9AEF9FA" author="Lucas, C. &amp; Nanthavong, K. &amp; Millet, J." box="[540,735,1564,1589]" pageId="6" pageNumber="253" pagination="410 - 420" refId="ref7816" refString="Lucas, C., Nanthavong, K. &amp; Millet, J. (2013) Environmental and human influnce on forest composition, structure and diversity in Laos. Journal of Tropical Forest Science 25 (3): 410 - 420." type="journal article" year="2013">
Lucas
<emphasis id="B940EAD4FF908D14FDA8CE16D9EDF9FA" box="[613,668,1564,1589]" italics="true" pageId="6" pageNumber="253">et al.</emphasis>
2013
</bibRefCitation>
). However, we could not find any specimen of this species in the major herbaria (BKF, FOF, HN, KAG, NHL, P, RUPP, TNS, VNM), indicating that this is a rare species. This situation is qualified as CR according to IUCN Red List criteria D (
<bibRefCitation id="EFA54B37FF908D14FCD9CE68D8EEF9B4" author="IUCN" box="[788,927,1634,1659]" pageId="6" pageNumber="253" refId="ref7560" refString="IUCN (2012) IUCN Red List Categories and Criteria. Version 3.1. Second edition. IUCN, Switzerland and Cambridge, Gland, 32 pp." type="book" year="2012">IUCN 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8B8B36C6FF908D14FF70CE8FDF00F8C7" blockId="6.[136,1452,367,1800]" pageId="6" pageNumber="253">
<emphasis id="B940EAD4FF908D14FF70CE8FDA6AF951" bold="true" box="[189,283,1669,1694]" pageId="6" pageNumber="253">Note:—</emphasis>
The
<emphasis id="B940EAD4FF908D14FE80CE8FDAFBF951" box="[333,394,1669,1694]" italics="true" pageId="6" pageNumber="253">matK</emphasis>
sequence of
<taxonomicName id="4C344D45FF908D14FDD1CE8FD858F951" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[540,809,1669,1694]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FDD1CE8FD858F951" box="[540,809,1669,1694]" italics="true" pageId="6" pageNumber="253">S. phoukhaokhouayense</emphasis>
</taxonomicName>
is identical with
<taxonomicName id="4C344D45FF908D14FC24CE8FDFF4F951" authorityName="Merrill &amp; Perry" authorityYear="1938" baseAuthorityName="Miquel" baseAuthorityYear="1855" box="[1001,1157,1669,1694]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="syzygioides">
<emphasis id="B940EAD4FF908D14FC24CE8FDFF4F951" box="[1001,1157,1669,1694]" italics="true" pageId="6" pageNumber="253">S. syzygioides</emphasis>
</taxonomicName>
(783/783 bp for GenBank accession no. AB924771 and AB925281, 780 bp for AB924710 and AB924734, 768/768 bp for AB924947). However,
<taxonomicName id="4C344D45FF908D14FF45CEC6DAE5F92A" authorityName="Soulad., Tagane &amp; Yahara. A" authorityYear="2018" box="[136,404,1740,1765]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="phoukhaokhouayense">
<emphasis id="B940EAD4FF908D14FF45CEC6DAE5F92A" box="[136,404,1740,1765]" italics="true" pageId="6" pageNumber="253">S. phoukhaokhouayense</emphasis>
</taxonomicName>
is easily distinguished from
<taxonomicName id="4C344D45FF908D14FD1CCEC6D81DF92A" authorityName="Merrill &amp; Perry" authorityYear="1938" baseAuthorityName="Miquel" baseAuthorityYear="1855" box="[721,876,1740,1765]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="6" pageNumber="253" phylum="Tracheophyta" rank="species" species="syzygioides">
<emphasis id="B940EAD4FF908D14FD1CCEC6D81DF92A" box="[721,876,1740,1765]" italics="true" pageId="6" pageNumber="253">S. syzygioides</emphasis>
</taxonomicName>
by the diagnostic characters mentioned above, such as the differences in leaf thickness, length of petiole and style, and colour of hypanthium.
</paragraph>
</subSubSection>
</treatment>
</document>