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<document ID-DOI="10.5733/afin.054.0213" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918661" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1683583268648" checkinUser="felipe" docAuthor="Niekerk, P. van &amp; Dippenaar-Schoeman, A. S." docDate="2013" docId="03D8D254FFFAA93D21A6FC5BFC83FDDD" docLanguage="en" docName="AfrInvertebr.54.2.447-476.pdf" docOrigin="African Invertebrates 54 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.054.0213" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Heriaeus madagascar Niekerk &amp; Dippenaar-Schoeman 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="465" masterDocId="FFE1AA2CFFEBA92F2055FF99FFE4FFE6" masterDocTitle="A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region" masterLastPageNumber="447" masterPageNumber="447" pageNumber="464" updateTime="1683701492856" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the crab spider genus Heriaeus Simon, 1875 (Araneae: Thomisidae) in the Afrotropical Region</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Niekerk, P. van</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dippenaar-Schoeman, A. S.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-11-14</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>54</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>447</mods:start>
<mods:end>447</mods:end>
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<mods:location>
<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.054.0213</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.054.0213</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7918661</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7917985" ID-Zenodo-Dep="7917985" LSID="urn:lsid:plazi:treatment:03D8D254FFFAA93D21A6FC5BFC83FDDD" httpUri="http://treatment.plazi.org/id/03D8D254FFFAA93D21A6FC5BFC83FDDD" lastPageId="18" lastPageNumber="465" pageId="17" pageNumber="464">
<subSubSection box="[499,824,962,989]" pageId="17" pageNumber="464" type="nomenclature">
<paragraph blockId="17.[181,1142,962,1677]" box="[499,824,962,989]" pageId="17" pageNumber="464">
<heading box="[499,824,962,989]" centered="true" fontSize="11" level="2" pageId="17" pageNumber="464" reason="2">
<taxonomicName ID-CoL="3KYVB" authority="Niekerk &amp; Dippenaar-Schoeman, 2013" authorityName="Niekerk &amp; Dippenaar-Schoeman" authorityYear="2013" box="[499,758,962,988]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="464" phylum="Arthropoda" rank="species" species="madagascar" status="sp. n.">
<emphasis bold="true" box="[499,758,962,988]" pageId="17" pageNumber="464">Heriaeus madagascar</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[765,824,963,989]" pageId="17" pageNumber="464" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="465" pageId="17" pageNumber="464" type="description">
<paragraph blockId="17.[181,1142,962,1677]" box="[564,759,1002,1029]" pageId="17" pageNumber="464">
<figureCitation box="[564,634,1002,1028]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="17" pageNumber="464">Figs 8</figureCitation>
,
<figureCitation box="[647,719,1002,1029]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="17" pageNumber="464">35, 36</figureCitation>
,
<figureCitation box="[731,759,1002,1029]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="17" pageNumber="464">59</figureCitation>
</paragraph>
<paragraph blockId="17.[181,1142,962,1677]" box="[181,1080,1042,1069]" pageId="17" pageNumber="464">
Etymology: The species name is a noun in apposition derived from
<collectingCountry box="[940,1074,1042,1068]" name="Madagascar" pageId="17" pageNumber="464">Madagascar</collectingCountry>
.
</paragraph>
<paragraph blockId="17.[181,1142,962,1677]" pageId="17" pageNumber="464">
Diagnosis: The species is recognized by a combination of long spiniform setae with acute tips on the abdomen, a single broad central stripe on the carapace, short strong setae on tibiae and metatarsi I and II (
<figureCitation box="[605,673,1146,1172]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="17" pageNumber="464">Fig. 8</figureCitation>
), and only four setae on the anterior edge of the clypeus. The male palp is distinguished by the angular shape of the cymbium, which is extended into a blunt extension laterally, the shape of the embolus, a broad flat ITA and a ventral tooth-like lobe on the RTA (
<figureCitation box="[696,826,1242,1269]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="17" pageNumber="464">Figs 35, 36</figureCitation>
). Female unknown.
</paragraph>
<paragraph blockId="17.[181,1142,962,1677]" box="[181,321,1282,1308]" pageId="17" pageNumber="464">Description:</paragraph>
<paragraph blockId="17.[181,1142,962,1677]" box="[181,370,1322,1348]" pageId="17" pageNumber="464">
<emphasis box="[181,239,1322,1348]" italics="true" pageId="17" pageNumber="464">Male</emphasis>
(
<typeStatus box="[254,359,1322,1348]" pageId="17" pageNumber="464">holotype</typeStatus>
).
</paragraph>
<paragraph blockId="17.[181,1142,962,1677]" lastBlockId="18.[144,1104,203,571]" lastPageId="18" lastPageNumber="465" pageId="17" pageNumber="464">
<emphasis box="[181,226,1363,1389]" italics="true" pageId="17" pageNumber="464">Size</emphasis>
: TL 3.02, CL 1.56, CW 1.41.
<emphasis box="[551,630,1362,1388]" italics="true" pageId="17" pageNumber="464">Colour</emphasis>
: Carapace with a broad brown stripe centrally, darker brown posteriorly; setae brown; chelicerae yellow-brown with brown markings; clypeus brown; eye region yellow-brown; eye tubercles grey.Abdomen creamy with dark mottling; cream with brown spots ventrally; setae brown. Palp with all segments yellow-brown. Legs yellow-brown, with brown marks on femora, tibiae and metatarsi I and II.
<emphasis box="[181,291,1523,1549]" italics="true" pageId="17" pageNumber="464">Carapace</emphasis>
:
<emphasis box="[307,516,1523,1548]" italics="true" pageId="17" pageNumber="464">Eye measurements</emphasis>
: AMEAME 0.19, ALEAME 0.11, PMEPME 0.11, PLEPME 0.29, AMEPME 0.15, MOQL AMEPME 0.22, MOQAW AMEAME 0.25, MOQPW PMEPME 0.21.
<emphasis box="[560,767,1587,1612]" italics="true" pageId="17" pageNumber="464">Leg measurements</emphasis>
: leg I Fe 2.20, Pat 0.97, Tib 1.50, Mt 1.38, Ta 0.76, total 6.81; leg II Fe 2.15, Pat 0.89, Tib 1.41, Mt 1.38, Ta 0.70, total 6.53; leg III Fe 1.13, Pat 0.58, Tib 1.13, Mt 0.76, Ta 0.44, total 4.04; leg IV Fe 0.96, Pat 0.53, Tib 0.78, Mt 0.81, Ta 0.45, total 3.53.
<emphasis box="[669,759,204,230]" italics="true" pageId="18" pageNumber="465">Clypeus</emphasis>
: Four setae on anterior clypeal edge.
<emphasis box="[208,314,236,262]" italics="true" pageId="18" pageNumber="465">Abdomen</emphasis>
: Oval, truncated anteriorly, slightly pointed posteriorly (
<figureCitation box="[934,999,236,262]" captionStart="Figs 19" captionStartId="3.[181,220,1090,1112]" captionTargetBox="[187,1132,204,1065]" captionTargetId="figure-241@3.[187,1132,204,1065]" captionTargetPageId="3" captionText="Figs 19. Heriaeus spp., habitus:(1) Heriaeus sp. long spiniform abdominal setae; (2) H.allenjonesi antoni sp. n. female; (3) H.antoni sp. n. female; (4) H.copricola sp. n. male; (5) H. crassispinus Lawrence, 1942 male; (6) H. foordi sp. n. male; (7) H. latifrons Lessert, 1919 female; (8) H.madagascar sp. n. male; (9) H. muizenberg sp. n. female." figureDoi="http://doi.org/10.5281/zenodo.7918663" httpUri="https://zenodo.org/record/7918663/files/figure.png" pageId="18" pageNumber="465">Fig. 8</figureCitation>
), bearing scattered long spiniform setae with acute tips.
<emphasis box="[663,716,268,293]" italics="true" pageId="18" pageNumber="465">Legs</emphasis>
: Tibiae and metatarsi I and II with short strong setae.
<emphasis box="[345,456,300,326]" italics="true" pageId="18" pageNumber="465">Male palp</emphasis>
: Cymbium with blunt extension laterally, forming a narrow ridge for tip of embolus to fit into, embolus long, winding once around tegulum, with sharp angle distally, tip straight and extending dorsally onto cymbium; ITA broad and flat; RTA with long distal tooth and ventral lobe below; VTA knob­like (
<figureCitation box="[950,1081,395,422]" captionStart="Figs 3036" captionStartId="15.[181,220,1193,1215]" captionTargetBox="[189,1133,212,1162]" captionTargetId="figure-164@15.[181,1141,204,1174]" captionTargetPageId="15" captionText="Figs 3036.(3033) Heriaeus foordi sp. n.: (30) male palp, ventral view; (31) male palp, lateral view; (32) epigyne, ventral view; (33) epigyne, dorsal view; (34) Heriaeus latifrons Lessert, 1919, epigyne, ventral view; (35, 36) Heriaeus madagascar sp. n.: (35) male palp, ventral view; (36) male palp, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7918671" httpUri="https://zenodo.org/record/7918671/files/figure.png" pageId="18" pageNumber="465">Figs 35, 36</figureCitation>
).
</paragraph>
<paragraph blockId="18.[144,1104,203,571]" box="[144,1068,434,456]" pageId="18" pageNumber="465">
<materialsCitation box="[144,1068,434,456]" collectingDate="1970-02" collectionCode="MRAC" collectorName="A. Lambillon" latitude="-15.73" longLatPrecision="770" longitude="49.47" pageId="18" pageNumber="465" specimenCode="MRAC 142.650" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[144,232,434,456]" pageId="18" pageNumber="465">Holotype</typeStatus>
:
<specimenCount box="[240,257,434,455]" pageId="18" pageNumber="465" type="male"></specimenCount>
<taxonomicName authority=": Beanana" authorityName="Beanana" box="[263,514,434,455]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="465" phylum="Arthropoda" rank="species" species="madagascar">MADAGASCAR: Beanana</taxonomicName>
(
<geoCoordinate box="[526,600,434,456]" degrees="15.73" direction="south" orientation="latitude" pageId="18" pageNumber="465" precision="555" value="-15.73">15.73°S</geoCoordinate>
<geoCoordinate box="[605,683,434,455]" degrees="49.47" direction="east" orientation="longitude" pageId="18" pageNumber="465" precision="555" value="49.47">49.47°E</geoCoordinate>
),
<date box="[698,764,434,456]" pageId="18" pageNumber="465" value="1970-02">
<collectingDate box="[698,764,434,456]" pageId="18" pageNumber="465" value="1970-02">ii.1970</collectingDate>
</date>
,
<collectorName box="[772,894,434,456]" pageId="18" pageNumber="465">A. Lambillon</collectorName>
(
<specimenCode box="[906,1058,434,456]" collectionCode="MRAC" country="Belgium" name="Musée Royal de lAfrique Centrale" pageId="18" pageNumber="465" type="Museum">MRAC 142.650</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="18.[144,1104,203,571]" box="[144,1091,468,491]" pageId="18" pageNumber="465">
<materialsCitation box="[144,1087,468,491]" collectingDate="1972-11-10" collectionCode="MRAC" collectorName="A. Dhondt" latitude="-18.15" longLatPrecision="765" longitude="49.42" pageId="18" pageNumber="465" specimenCode="MRAC 142.296" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus box="[144,227,469,490]" pageId="18" pageNumber="465">Paratype</typeStatus>
:
<specimenCount box="[235,252,469,490]" pageId="18" pageNumber="465" type="male"></specimenCount>
<taxonomicName authority=": Tananarive" authorityName="Tananarive" box="[258,530,469,491]" class="Arachnida" family="Thomisidae" genus="Heriaeus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="465" phylum="Arthropoda" rank="species" species="madagascar">MADAGASCAR: Tananarive</taxonomicName>
(
<geoCoordinate box="[543,617,468,490]" degrees="18.15" direction="south" orientation="latitude" pageId="18" pageNumber="465" precision="555" value="-18.15">18.15°S</geoCoordinate>
<geoCoordinate box="[622,700,469,490]" degrees="49.42" direction="east" orientation="longitude" pageId="18" pageNumber="465" precision="555" value="49.42">49.42°E</geoCoordinate>
),
<date box="[715,814,469,491]" pageId="18" pageNumber="465" value="1972-11-10">
<collectingDate box="[715,814,469,491]" pageId="18" pageNumber="465" value="1972-11-10">10.xi.1972</collectingDate>
</date>
,
<collectorName box="[822,918,469,491]" pageId="18" pageNumber="465">A. Dhondt</collectorName>
(
<specimenCode box="[929,1081,468,490]" collectionCode="MRAC" country="Belgium" name="Musée Royal de lAfrique Centrale" pageId="18" pageNumber="465" type="Museum">MRAC 142.296</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[144,1104,203,571]" box="[144,541,505,532]" pageId="18" pageNumber="465">
Distribution:
<collectingCountry box="[294,429,505,531]" name="Madagascar" pageId="18" pageNumber="465">Madagascar</collectingCountry>
(
<figureCitation box="[444,529,505,532]" captionStart="Figs 5759" captionStartId="27.[347,386,1654,1676]" captionTargetBox="[181,1141,609,1633]" captionTargetId="figure-176@27.[132,1190,602,1636]" captionTargetPageId="27" captionText="Figs 5759. Distribution of Heriaeus spp. in the Afrotropical Region." figureDoi="http://doi.org/10.5281/zenodo.7918679" httpUri="https://zenodo.org/record/7918679/files/figure.png" pageId="18" pageNumber="465">Fig. 59</figureCitation>
).
</paragraph>
<paragraph blockId="18.[144,1104,203,571]" box="[144,871,545,571]" pageId="18" pageNumber="465">Natural history: Males collected during February and November.</paragraph>
</subSubSection>
</treatment>
</document>