194 lines
36 KiB
XML
194 lines
36 KiB
XML
<document id="733AED3F7D4EAA03A7D0206C2BF17FED" ID-CLB-Dataset="45970" ID-DOI="10.11646/zootaxa.3790.1.6" ID-GBIF-Dataset="bc52b393-d1b1-43c3-9491-d3b9d47ef9b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="226715" ID-ZooBank="87AB27EC-DC05-48F3-8AB7-5C317B275AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460410436114" checkinUser="plazi" docAuthor="Sueyoshi, Masahiro" docDate="2014" docId="49168794FFE7564C37ADF8CE215A371D" docLanguage="en" docName="zt03790p164.pdf" docOrigin="Zootaxa 3790 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoempheria biceltisuta Sueyoshi, 2014, sp. n." docType="treatment" docVersion="9" lastPageNumber="147" masterDocId="B52FFFECFFE15644373AFFCC222B360A" masterDocTitle="Taxonomy of fungus gnats allied to Neoempheria ferruginea (Brunetti, 1912) (Diptera: Mycetophilidae), with descriptions of 11 new species from Japan and adjacent areas" masterLastPageNumber="164" masterPageNumber="139" pageNumber="145" updateTime="1698300419586" updateUser="ExternalLinkService">
|
||
<mods:mods id="3383BE4E4A80124D669858A6A86FA97A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="C77E75F0FA21EBBA9F8ADCF0F575137C">
|
||
<mods:title id="28BBAB607F7C5B3CA80E7F153CA9763E">Taxonomy of fungus gnats allied to Neoempheria ferruginea (Brunetti, 1912) (Diptera: Mycetophilidae), with descriptions of 11 new species from Japan and adjacent areas</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0AC9A144FE1E1EE31DEE2EFB0C5353E4" type="personal">
|
||
<mods:role id="36E718F47BB8574580A0BB9E4B36D2DE">
|
||
<mods:roleTerm id="0A0EF702790840EFEFAC9141E84351C4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="62344DEA7B331C0D0EB844C33C2672B9">Sueyoshi, Masahiro</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="48F06BF792406298E4EB08400A0922F6">text</mods:typeOfResource>
|
||
<mods:relatedItem id="92BE59A6149B57BFCAABB983AC02712A" type="host">
|
||
<mods:titleInfo id="7B9ACE281AC9ED681DDF16E83F06DBBA">
|
||
<mods:title id="02B4FFC6C1B8F6E08242CBA991601E27">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="686BDE46DD52A22CE87370CD2E006B09">
|
||
<mods:date id="BAD1359890473B9373C6541A1E226E82">2014</mods:date>
|
||
<mods:detail id="CE76EF65312651469C05D2D5E21D6B05" type="volume">
|
||
<mods:number id="5A242A520047674295AB5A28B16840D1">3790</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2DD825FD75168DE2203BAB70490F3693" type="issue">
|
||
<mods:number id="A62E131754BF57A157CCD6E86F39E7F5">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8EE3AF245757DD130BE57B4FD976D518" unit="page">
|
||
<mods:start id="A05933B2DBF4B81D6B3B3D9452E3AC48">139</mods:start>
|
||
<mods:end id="A51689190F5421D488FA17A4D67A8D57">164</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="1DBBDD20F412A8B05DA311E1D1063475">journal article</mods:classification>
|
||
<mods:identifier id="EAF1B608EE7246E66AA536E1E5923078" type="CLB-Dataset">45970</mods:identifier>
|
||
<mods:identifier id="1E345ADCD152FB131542156D5E5A06E5" type="DOI">10.11646/zootaxa.3790.1.6</mods:identifier>
|
||
<mods:identifier id="9399F218ECCA679B2266F100EB836E2B" type="GBIF-Dataset">bc52b393-d1b1-43c3-9491-d3b9d47ef9b0</mods:identifier>
|
||
<mods:identifier id="5FD907EC4A8467587A74D51E702A94E1" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="6F5045BE16D6F88081471C4C6593C2FC" type="Zenodo-Dep">226715</mods:identifier>
|
||
<mods:identifier id="EBB904B2312058E6DBD29F91CB7D804C" type="ZooBank">87AB27EC-DC05-48F3-8AB7-5C317B275AF5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="49168794FFE7564C37ADF8CE215A371D" ID-DOI="http://doi.org/10.5281/zenodo.6138786" ID-GBIF-Taxon="119423262" ID-Zenodo-Dep="6138786" LSID="urn:lsid:plazi:treatment:49168794FFE7564C37ADF8CE215A371D" httpUri="http://treatment.plazi.org/id/49168794FFE7564C37ADF8CE215A371D" lastPageId="8" lastPageNumber="147" pageId="6" pageNumber="145">
|
||
<subSubSection id="89A56509FFE7564237ADF8CE23A2316E" pageId="6" pageNumber="145" type="nomenclature">
|
||
<paragraph id="C1003682FFE7564237ADF8CE23DD3116" blockId="6.[151,712,1794,1892]" box="[151,502,1794,1820]" pageId="6" pageNumber="145">
|
||
<heading id="9A4881EEFFE7564237ADF8CE23DD3116" bold="true" box="[151,502,1794,1820]" fontSize="10" level="2" pageId="6" pageNumber="145" reason="2">
|
||
<emphasis id="F3CBEA90FFE7564237ADF8CE23DD3116" bold="true" box="[151,502,1794,1820]" pageId="6" pageNumber="145">
|
||
<taxonomicName id="06BF4D01FFE7564237ADF8CE23843111" ID-CoL="9HYXD" box="[151,431,1794,1819]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="biceltisuta" status="sp. nov.">
|
||
<emphasis id="F3CBEA90FFE7564237ADF8CE23843111" bold="true" box="[151,431,1794,1819]" italics="true" pageId="6" pageNumber="145">Neoempheria biceltisuta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8F857EBFFE75642368CF8CF23DD3116" box="[438,502,1795,1820]" pageId="6" pageNumber="145" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="C1003682FFE7564237ADF8E423A2316E" blockId="6.[151,712,1794,1892]" pageId="6" pageNumber="145">
|
||
[Japanese name: futanomi-nagamado-kinoko-bae] (
|
||
<figureCitation id="59842A07FFE7564237A5F88722C6316E" box="[159,237,1867,1892]" captionStart="FIGURE 2" captionStartId="3.[151,250,1812,1834]" captionTargetBox="[151,1436,345,1790]" captionTargetId="figure@3.[151,1436,345,1791]" captionTargetPageId="3" captionText="FIGURE 2. Thoraxes of Neoempheria in dorsal view. a, N. biceltisuta sp. n. (holotype). b, N. bifurcata sp. n. (holotype). c, N. bisecuriata sp. n. (holotype). d, N. brevispathulata sp. n. (holotype). e, N. carinata sp. n. (holotype). f, N. cuneata sp. n. (holotype). g, N. denticulata sp. n. (holotype). h, N. dilatata sp. n. (holotype). i, N. forficulata sp. n. (holotype). j, N. latisternata sp. n. (holotype). k, N. muticata sp. n. (holotype). l, N. sakhalinensis Zaitzev, 2001 (Ne. 2042)." httpUri="https://zenodo.org/record/226716/files/figure.png" pageId="6" pageNumber="145">Figs. 2</figureCitation>
|
||
a, 3a, 5, 9a–e)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A56509FFE7564337ADF8412623319B" lastPageId="7" lastPageNumber="146" pageId="6" pageNumber="145" type="description">
|
||
<paragraph id="C1003682FFE7564337ADF841277737FF" blockId="6.[151,1436,1933,2033]" lastBlockId="7.[151,1437,151,501]" lastPageId="7" lastPageNumber="146" pageId="6" pageNumber="145">
|
||
<emphasis id="F3CBEA90FFE7564237ADF841230A31AC" bold="true" box="[151,289,1933,1958]" pageId="6" pageNumber="145">Description</emphasis>
|
||
. Body length:
|
||
<quantity id="06479B67FFE7564236F7F842207D31AC" box="[461,598,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="6" pageNumber="145" unit="mm" value="4.1" valueMax="4.4" valueMin="3.8">3.8–4.4 mm</quantity>
|
||
(n= 2) in male,
|
||
<quantity id="06479B67FFE75642342FF842214631AC" box="[789,877,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="145" unit="mm" value="4.2">4.2 mm</quantity>
|
||
(n= 1) in female. Wing length:
|
||
<quantity id="06479B67FFE7564233E4F842271D31AC" box="[1246,1334,1934,1958]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="145" unit="mm" value="4.4">4.4 mm</quantity>
|
||
(4.0–
|
||
<quantity id="06479B67FFE756423243F84122EE31C0" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="145" unit="mm" value="4.6">4.6 mm</quantity>
|
||
, n = 3) in male,
|
||
<quantity id="06479B67FFE756423642F87E23E531C0" box="[376,462,1970,1995]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="6" pageNumber="145" unit="mm" value="5.3">5.3 mm</quantity>
|
||
(n= 1) in female.
|
||
<emphasis id="F3CBEA90FFE7564235AFF87F20E631C0" box="[661,717,1971,1994]" italics="true" pageId="6" pageNumber="145">Wing</emphasis>
|
||
(
|
||
<figureCitation id="59842A07FFE7564235E7F87E210B31C0" box="[733,800,1970,1995]" captionStart="FIGURE 3" captionStartId="4.[151,250,1644,1666]" captionTargetBox="[151,1436,193,1620]" captionTargetId="figure@4.[151,1436,193,1622]" captionTargetPageId="4" captionText="FIGURE 3. Right wings of Neoempheria in dorsal view. a, N. biceltisuta sp. n. (holotype). b, N. bifurcata sp. n. (holotype). c, N. bisecuriata sp. n. (holotype). d, N. brevispathulata sp. n. (holotype). e, N. carinata sp. n. (holotype). f, N. cuneata sp. n. (holotype). g, N. denticulata sp. n. (holotype). h, N. dilatata sp. n. (holotype). i, N. forficulata sp. n. (holotype). j, N. latisternata sp. n. (holotype). k, N. muticata sp. n. (holotype). l, N. sakhalinensis Zaitzev, 2001 (Ne. 2042)." httpUri="https://zenodo.org/record/226717/files/figure.png" pageId="6" pageNumber="145">Fig. 3</figureCitation>
|
||
a): vein sc-r ending basal 1/3 of anterior margin of cell r1. Vein Rs as long as distance between basal end of vein Rs and apical end of vein sc-r.
|
||
<emphasis id="F3CBEA90FFE756423346F814269D31FB" box="[1148,1206,2008,2033]" italics="true" pageId="6" pageNumber="145">Male</emphasis>
|
||
: genitalia yellow in ground color, gonocoxal projection black, apex of gonocoxal lobe and surrounding parts brown to dark brown, basal portion of aedeagus yellow (
|
||
<figureCitation id="59842A07FFE65643352FFF70207336DF" box="[533,600,188,213]" captionStart="FIGURE 5" captionStartId="6.[151,250,1636,1658]" captionTargetBox="[151,1436,591,1614]" captionTargetId="figure@6.[151,1436,591,1614]" captionTargetPageId="6" captionText="FIGURE 5. Distribution map of Asian Neoempheria. Neoempheria biceltisuta sp. n. (filled star), N. brevispathulata sp. n. (filled up-side-down triangle), N. denticulata sp. n. (filled diamond), N. dilatata sp. n. (filled circle), N. ferruginea (Brunetti, 1912) (filled square), and N. latisternata sp. n. (filled triangle.)" httpUri="https://zenodo.org/record/226719/files/figure.png" pageId="7" pageNumber="146">Fig. 5</figureCitation>
|
||
a). Sternite 9 (
|
||
<figureCitation id="59842A07FFE6564335CFFF70211236DE" box="[757,825,188,213]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a: S9) narrow, less than 1/10 as wide as long, visible as seam between gonocoxites, without sternal projection. Gonocoxite 9 (
|
||
<figureCitation id="59842A07FFE656433490FF1321C636F2" box="[938,1005,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a, b: gc) with gonocoxal lobe (
|
||
<figureCitation id="59842A07FFE65643327FFF1327A136F2" box="[1349,1418,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a: gl) truncate at apex. Gonostylus (
|
||
<figureCitation id="59842A07FFE656433530FEC820653716" box="[522,590,260,285]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a: gs) slender, apical 1/4 less than 1.5 times as wide as base. Aedeagus (
|
||
<figureCitation id="59842A07FFE656433255FEC8228D374A" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a, b) dilated on apical half, without projections. Sclerotized part of aedeagus (
|
||
<figureCitation id="59842A07FFE65643331BFEEB264C374A" box="[1057,1127,295,320]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a: sa) tapered to apex, with slender, angled lateral extension.
|
||
<emphasis id="F3CBEA90FFE656433529FE80204C376F" box="[531,615,332,357]" italics="true" pageId="7" pageNumber="146">Female</emphasis>
|
||
: S7 with uniform setae on posterior margin, posterior margin with deep medial indentation. S8 triangular in ventral view. Gonocoxite 8 (
|
||
<figureCitation id="59842A07FFE6564334B5FEA321FC3782" box="[911,983,367,392]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
c: gc8) small, recognizable as bump at sublateral portion in posterior margin of S8. Gonapophysis 8 (
|
||
<figureCitation id="59842A07FFE65643346EFE5821B237A6" box="[852,921,404,429]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
c: gp8) dilated in apical half, with acute apex, dentate on dorsal margin. Gonocoxite 9 (
|
||
<figureCitation id="59842A07FFE656433552FE7B208737DA" box="[616,684,439,464]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
d, e: gc9) yellow, without projections. Gonapophysis 9 (
|
||
<figureCitation id="59842A07FFE65643321DFE7B274737DA" box="[1319,1388,439,464]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
d, e: gp9) yellow, with 2 indistinct ventral sublateral ridges (
|
||
<figureCitation id="59842A07FFE656433439FE10216C37FE" box="[771,839,476,501]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
e: vr), gradually tapered to apex with gonopore.
|
||
</paragraph>
|
||
<caption id="95C0660AFFE6564337ADF91927283100" httpUri="https://zenodo.org/record/226720/files/figure.png" pageId="7" pageNumber="146" targetBox="[212,1380,555,1735]" targetPageId="7">
|
||
<paragraph id="C1003682FFE6564337ADF91927283100" blockId="7.[151,1436,1749,1802]" pageId="7" pageNumber="146">
|
||
<emphasis id="F3CBEA90FFE6564337ADF919212330E1" bold="true" box="[151,776,1749,1771]" pageId="7" pageNumber="146">
|
||
FIGURE 6. Distribution map of Japanese
|
||
<emphasis id="F3CBEA90FFE65643354FF919212330E1" box="[629,776,1749,1771]" italics="true" pageId="7" pageNumber="146">
|
||
<taxonomicName id="06BF4D01FFE65643354FF919212F30E1" box="[629,772,1749,1771]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="genus">Neoempheria</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</emphasis>
|
||
<taxonomicName id="06BF4D01FFE656433429F919262A30E1" box="[787,1025,1749,1771]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="bifurcata" status="sp. nov.">Neoempheria bifurcata</taxonomicName>
|
||
<taxonomicNameLabel id="E8F857EBFFE656433331F919266330E1" box="[1035,1096,1749,1771]" pageId="7" pageNumber="146" rank="species">
|
||
<emphasis id="F3CBEA90FFE656433331F919266330E1" bold="true" box="[1035,1096,1749,1771]" pageId="7" pageNumber="146">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(filled circle: locality of indoor facility; double circle: locality in field),
|
||
<taxonomicName id="06BF4D01FFE656433516F93920B13100" box="[556,666,1781,1802]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="cuneata" status="sp. nov.">
|
||
<emphasis id="F3CBEA90FFE656433516F93920163100" box="[556,573,1781,1802]" italics="true" pageId="7" pageNumber="146">N</emphasis>
|
||
.
|
||
<emphasis id="F3CBEA90FFE656433570F93920B13100" box="[586,666,1781,1802]" italics="true" pageId="7" pageNumber="146">cuneata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8F857EBFFE65643359AF93820F13100" box="[672,730,1780,1802]" pageId="7" pageNumber="146" rank="species">
|
||
<emphasis id="F3CBEA90FFE65643359AF93820F13100" bold="true" box="[672,730,1780,1802]" pageId="7" pageNumber="146">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(filled square), and
|
||
<taxonomicName id="06BF4D01FFE65643349CF93926363100" box="[934,1053,1781,1802]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="muticata" status="sp. nov.">
|
||
<emphasis id="F3CBEA90FFE65643349CF939219C3100" box="[934,951,1781,1802]" italics="true" pageId="7" pageNumber="146">N</emphasis>
|
||
.
|
||
<emphasis id="F3CBEA90FFE6564334FEF93926363100" box="[964,1053,1781,1802]" italics="true" pageId="7" pageNumber="146">muticata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8F857EBFFE656433318F93826703100" box="[1058,1115,1780,1802]" pageId="7" pageNumber="146" rank="species">
|
||
<emphasis id="F3CBEA90FFE656433318F93826703100" bold="true" box="[1058,1115,1780,1802]" pageId="7" pageNumber="146">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(filled triangle).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C1003682FFE6564337FDF8FD2623319B" blockId="7.[151,1437,1841,2010]" pageId="7" pageNumber="146">
|
||
<emphasis id="F3CBEA90FFE6564337FDF8FD23E73140" bold="true" box="[199,460,1841,1866]" pageId="7" pageNumber="146">Specimens examined.</emphasis>
|
||
<typeStatus id="1E048820FFE6564336E2F8FD206F3140" box="[472,580,1841,1866]" pageId="7" pageNumber="146" type="holotype">Holotype</typeStatus>
|
||
. Male, “
|
||
<collectingCountry id="B9A87612FFE656433596F8FD213E3140" box="[684,789,1841,1866]" name="Thailand" pageId="7" pageNumber="146">Thailand</collectingCountry>
|
||
, Phuket/ Marine Biol. Center/
|
||
<date id="B5011042FFE6564333B3F8FD27003140" box="[1161,1323,1841,1866]" pageId="7" pageNumber="146" value="1987-11-13" valueMax="1987-11-16" valueMin="1987-11-13">13.-16.xi.1987</date>
|
||
/ Mogens Andersen leg.”, green disk label, “Ne.7004” (ZMCU).
|
||
<typeStatus id="1E048820FFE656433439F89A215F3164" box="[771,884,1878,1902]" pageId="7" pageNumber="146" type="paratype">Paratypes</typeStatus>
|
||
: same data as
|
||
<typeStatus id="1E048820FFE65643332DF89926523164" box="[1047,1145,1877,1902]" pageId="7" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="D7B9FD0BFFE6564333B1F89A26863164" box="[1163,1197,1878,1902]" pageId="7" pageNumber="146" type="generic">1♂</specimenCount>
|
||
, 1♀, Ne.7005, 7006; ZMCU); same data as
|
||
<typeStatus id="1E048820FFE6564336AEF8B523DD3198" box="[404,502,1913,1938]" pageId="7" pageNumber="146" type="holotype">holotype</typeStatus>
|
||
except,
|
||
<date id="B5011042FFE65643356EF8B520DC3198" box="[596,759,1913,1938]" pageId="7" pageNumber="146" value="1987-11-17" valueMax="1987-11-21" valueMin="1987-11-17">17–21.xi.1987</date>
|
||
(
|
||
<specimenCount id="D7B9FD0BFFE65643343DF8B52107319B" box="[775,812,1913,1937]" pageId="7" pageNumber="146" type="generic">1♂</specimenCount>
|
||
, Ne.7007; ZMCU).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A56509FFE6564337FDF851208231D3" pageId="7" pageNumber="146" type="etymology">
|
||
<paragraph id="C1003682FFE6564337FDF851208231D3" blockId="7.[151,1437,1841,2010]" pageId="7" pageNumber="146">
|
||
<emphasis id="F3CBEA90FFE6564337FDF851236D31BC" bold="true" box="[199,326,1949,1974]" pageId="7" pageNumber="146">Etymology</emphasis>
|
||
. The specific epithet is derived from Latin and refers to the broad, truncate, chisel-like (
|
||
<emphasis id="F3CBEA90FFE656433274F85127B931BC" box="[1358,1426,1949,1974]" italics="true" pageId="7" pageNumber="146">celtis-</emphasis>
|
||
) apex of the two (
|
||
<emphasis id="F3CBEA90FFE65643366EF80D235831D0" box="[340,371,1985,2010]" italics="true" pageId="7" pageNumber="146">bi-</emphasis>
|
||
) gonocoxal lobes (
|
||
<figureCitation id="59842A07FFE656433570F80D20A631D3" box="[586,653,1985,2010]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="7" pageNumber="146">Fig. 9</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A56509FFE9564C37FDFF5B20BF36BA" box="[199,660,151,176]" pageId="8" pageNumber="147" type="distribution">
|
||
<paragraph id="C1003682FFE9564C37FDFF5B20BF36BA" blockId="8.[151,1437,151,279]" box="[199,660,151,176]" pageId="8" pageNumber="147">
|
||
<emphasis id="F3CBEA90FFE9564C37FDFF5B234B36BA" bold="true" box="[199,352,151,176]" pageId="8" pageNumber="147">Distribution.</emphasis>
|
||
<collectingCountry id="B9A87612FFE9564C365DFF5B23E036BA" box="[359,459,151,176]" name="Thailand" pageId="8" pageNumber="147">Thailand</collectingCountry>
|
||
(Phuket) (
|
||
<figureCitation id="59842A07FFE9564C357BFF5B20AD36BA" box="[577,646,151,176]" captionStart="FIGURE 5" captionStartId="6.[151,250,1636,1658]" captionTargetBox="[151,1436,591,1614]" captionTargetId="figure@6.[151,1436,591,1614]" captionTargetPageId="6" captionText="FIGURE 5. Distribution map of Asian Neoempheria. Neoempheria biceltisuta sp. n. (filled star), N. brevispathulata sp. n. (filled up-side-down triangle), N. denticulata sp. n. (filled diamond), N. dilatata sp. n. (filled circle), N. ferruginea (Brunetti, 1912) (filled square), and N. latisternata sp. n. (filled triangle.)" httpUri="https://zenodo.org/record/226719/files/figure.png" pageId="8" pageNumber="147">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89A56509FFE9564C37FDFF70215A371D" pageId="8" pageNumber="147" type="discussion">
|
||
<paragraph id="C1003682FFE9564C37FDFF70215A371D" blockId="8.[151,1437,151,279]" pageId="8" pageNumber="147">
|
||
<emphasis id="F3CBEA90FFE9564C37FDFF70231036DF" bold="true" box="[199,315,188,213]" pageId="8" pageNumber="147">Remarks.</emphasis>
|
||
This species is distinguished from similar species by the broad, truncate apex of the male gonocoxal lobe (
|
||
<figureCitation id="59842A07FFE9564C37E6FF11230E36FC" box="[220,293,221,246]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="8" pageNumber="147">Fig. 9</figureCitation>
|
||
a), and in the female by the dentate dorsal margin of the gonapophysis 8 (
|
||
<figureCitation id="59842A07FFE9564C3393FF1126D936FC" box="[1193,1266,221,246]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="8" pageNumber="147">Fig. 9</figureCitation>
|
||
c), and yellow gonapophysis 9 with indistinct ventral sublateral ridges (
|
||
<figureCitation id="59842A07FFE9564C342BFF32217E371D" box="[785,853,254,279]" captionStart="FIGURE 9" captionStartId="12.[151,250,916,938]" captionTargetBox="[169,1416,205,885]" captionTargetId="figure@12.[163,1425,195,895]" captionTargetPageId="12" captionText="FIGURE 9. Neoempheria biceltisuta sp. n. a, b, male (holotype). c – e, female (paratype, Ne. 7006). a, 9 th abdominal segment and aedeagus in ventral view. b, gonocoxite, gonostylus, and aedeagus in left lateral view. c, 8 th abdominal sternite and projections in left lateral view. d, e, 9 th abdominal segment in left lateral view (d) and posterior view (e). Scale bars: a, b = 0.50 mm; c – e = 0.25 mm. Abbreviations: gc, male gonocoxite; gl, gonocoxal lobe of male 9 th gonocoxite; gp, gonocoxal projection; gs, gonostylus; lb, labia; ma, membranous part of aedeagus; sa, sclerotized part of aedeagus; vr, ventral sublateral ridge of female gonapophysis 9; S 8, sternite 8; S 9, sternite 9; T 9, tergite 9; gc 8, female gonocoxite 8; gc 9, female gonocoxite 9; gp 8, female gonapophysis 8; gp 9, female gonapophysis 9." httpUri="https://zenodo.org/record/226723/files/figure.png" pageId="8" pageNumber="147">Fig. 9</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |