treatments-xml/data/E0/BD/74/E0BD74DF82145003A5B61763858FD1E1.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1024.60473" ID-GBIF-Dataset="b5167639-dcd0-4edc-b502-e8863632d628" ID-PMC="PMC7985131" ID-Pensoft-Pub="1313-2970-1024-117" ID-Pensoft-UUID="34BB99A9758355CBB3170F765063BCF2" ID-PubMed="33776523" ID-ZooBank="1E36885A6C0349BCA22F982A9CEA710" ModsDocID="1313-2970-1024-117" checkinTime="1615893247286" checkinUser="pensoft" docAuthor="Ramirez-Reyes, Tonatiuh, Barraza-Soltero, Ilse K., Nolasco-Luna, Jose Rafael, Flores-Villela, Oscar &amp; Escobedo-Galvan, Armando H." docDate="2021" docId="E0BD74DF82145003A5B61763858FD1E1" docLanguage="en" docName="ZooKeys 1024: 117-136" docOrigin="ZooKeys 1024" docPubDate="2021-03-15" docSource="http://dx.doi.org/10.3897/zookeys.1024.60473" docTitle="Phyllodactylus cleofasensis Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván 2021, sp. nov." docType="treatment" docUuid="0FCA70B9-5FF9-4AE4-B422-7210A3F1ED4F" docUuidSource="ZooBank" docVersion="6" id="34BB99A9758355CBB3170F765063BCF2" lastPageNumber="117" masterDocId="34BB99A9758355CBB3170F765063BCF2" masterDocTitle="A new species of leaf-toed gecko (Phyllodactylidae, Phyllodactylus) from Maria Cleofas Island, Nayarit, Mexico" masterLastPageNumber="136" masterPageNumber="117" pageNumber="117" updateTime="1668149897153" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of leaf-toed gecko (Phyllodactylidae, Phyllodactylus) from Maria Cleofas Island, Nayarit, Mexico</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ramirez-Reyes, Tonatiuh</mods:namePart>
<mods:affiliation>Posgrado en Ciencias Biologicas, Unidad de Posgrado, Edificio D, primer piso, Universidad Nacional Autonoma de Mexico, Ciudad Universitaria 3000, Coyoacan, Ciudad de Mexico, Mexico &amp; Museo de Zoologia, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, Circuito Exterior de CU, Ciudad Universitaria, 04510, Ciudad de Mexico, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barraza-Soltero, Ilse K.</mods:namePart>
<mods:affiliation>Centro Universitario de la Costa, Universidad de Guadalajara, Av. Universidad 203, 48280, Puerto Vallarta, Jalisco, Mexico &amp; Laboratorio de Ecologia del Comportamiento, Centro de Investigaciones Biologicas del Noroeste, S. C., Av. Instituto Politecnico Nacional 195, Col. Playa Palo de Santa Rita Sur, Apdo. Postal 128; La Paz, BCS 23096, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nolasco-Luna, Jose Rafael</mods:namePart>
<mods:affiliation>Centro Universitario de la Costa, Universidad de Guadalajara, Av. Universidad 203, 48280, Puerto Vallarta, Jalisco, Mexico &amp; Universidad Autonoma de Nayarit, Ciudad de la Cultura &quot; Amado Nervo &quot; s / n, 63155, Tepic, Nayarit, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Flores-Villela, Oscar</mods:namePart>
<mods:affiliation>Museo de Zoologia, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, Circuito Exterior de CU, Ciudad Universitaria, 04510, Ciudad de Mexico, Mexico</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Escobedo-Galvan, Armando H.</mods:namePart>
<mods:affiliation>Centro Universitario de la Costa, Universidad de Guadalajara, Av. Universidad 203, 48280, Puerto Vallarta, Jalisco, Mexico</mods:affiliation>
<mods:nameIdentifier type="email">elchorvis@gmail.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-15</mods:number>
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<mods:detail type="volume">
<mods:number>1024</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1024.60473</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1024.60473</mods:identifier>
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<treatment ID-GBIF-Taxon="180583900" LSID="urn:lsid:zoobank.org:act:0FCA70B9-5FF9-4AE4-B422-7210A3F1ED4F" httpUri="http://treatment.plazi.org/id/E0BD74DF82145003A5B61763858FD1E1" lastPageNumber="117" pageId="0" pageNumber="117">
<subSubSection pageId="0" pageNumber="117" type="nomenclature">
<paragraph pageId="0" pageNumber="117">
<taxonomicName LSID="http://zoobank.org/0FCA70B9-5FF9-4AE4-B422-7210A3F1ED4F" authority="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván, 2021" authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus cleofasensis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="cleofasensis" status="sp. nov.">Phyllodactylus cleofasensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="117">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Dorsal view of Phyllodactylus cleofasensis sp. nov., in life. (Photo by AHE-G)." figureDoi="10.3897/zookeys.1024.60473.figure7" httpUri="https://binary.pensoft.net/fig/520452" pageId="0" pageNumber="117">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="reference_group">
<paragraph pageId="0" pageNumber="117">
<taxonomicName authority="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván, 2021" authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus tuberculosus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="tuberculosus" status="sp. nov.">Phyllodactylus tuberculosus</taxonomicName>
(in part) Wiegmann 1835 (
<bibRefCitation DOI="https://doi.org/10.3996/nafa.14.0004" author="Stejneger, L" journalOrPublisher="North American Fauna" pageId="0" pageNumber="117" pagination="63 - 71" refId="B37" refString="Stejneger, L, 1899. Reptiles of the Tres Marias and Isabel islands. North American Fauna 14: 63 - 71, DOI: https://doi.org/10.3996/nafa.14.0004" title="Reptiles of the Tres Marias and Isabel islands." url="https://doi.org/10.3996/nafa.14.0004" volume="14" year="1899">Stejneger 1899</bibRefCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authority="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván, 2021" authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus lanei" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="lanei" status="sp. nov.">Phyllodactylus lanei</taxonomicName>
(in part) 
<bibRefCitation author="Smith, HM" journalOrPublisher="University Kansas Science Bulletin" pageId="0" pageNumber="117" pagination="119 - 155" refId="B36" refString="Smith, HM, 1935. Miscellaneous notes on Mexican lizards. University Kansas Science Bulletin 22: 119 - 155" title="Miscellaneous notes on Mexican lizards." volume="22" year="1935">Smith 1935</bibRefCitation>
(
<bibRefCitation author="Zweifel, RG" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="117" pagination="77 - 128" refId="B43" refString="Zweifel, RG, 1960. Results of the Puritan-American Museum of Natural History expedition to western Mexico. 9. Herpetology of the Tres Marias islands. American Museum Novitates 119: 77 - 128" title="Results of the Puritan-American Museum of Natural History expedition to western Mexico. 9. Herpetology of the Tres Marias islands." volume="119" year="1960">Zweifel 1960</bibRefCitation>
)
</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authority="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván, 2021" authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Insecta" family="Phyllodactylidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="P. tuberculosus subsp. saxatilis" order="Squamata" pageId="0" pageNumber="117" phylum="Nematoda" rank="subSpecies" species="tuberculosus" status="sp. nov." subSpecies="saxatilis">P. tuberculosus saxatilis</taxonomicName>
(in part) 
<bibRefCitation author="Dixon, JR" journalOrPublisher="New Mexico State University Science Bulletin" pageId="0" pageNumber="117" pagination="1 - 145" refId="B13" refString="Dixon, JR, 1964. The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America. New Mexico State University Science Bulletin 64 (1): 1 - 145" title="The systematics and distribution of lizards of the genus Phyllodactylus in North and Central America." volume="64" year="1964">Dixon 1964</bibRefCitation>
(
<bibRefCitation author="McDiarmid, RW" journalOrPublisher="Natural History Museum of Los Angeles County Contributions in Science" pageId="0" pageNumber="117" pagination="1 - 17" refId="B25" refString="McDiarmid, RW, Copp, JF, Breedlove, DE, 1976. Notes on the herpetofauna of western Mexico: new records from Sinaloa and the Tres Marias Islands. Natural History Museum of Los Angeles County Contributions in Science 275: 1 - 17" title="Notes on the herpetofauna of western Mexico: new records from Sinaloa and the Tres Marias Islands." volume="275" year="1976">McDiarmid et al. 1976</bibRefCitation>
; 
<bibRefCitation author="Casas-Andreu, G" journalOrPublisher="Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico Serie Zoologia" pageId="0" pageNumber="117" pagination="95 - 112" refId="B7" refString="Casas-Andreu, G, 1992. Anfibios y reptiles de las Islas Marias y otras islas adyacentes a la costa de Nayarit, Mexico. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico Serie Zoologia 63 (1): 95 - 112" title="Anfibios y reptiles de las Islas Marias y otras islas adyacentes a la costa de Nayarit, Mexico." volume="63" year="1992">Casas-Andreu 1992</bibRefCitation>
; 
<bibRefCitation author="Woolrich-Pina, GA" journalOrPublisher="Mesoamerican Herpetology" pageId="0" pageNumber="117" pagination="375 - 448" refId="B42" refString="Woolrich-Pina, GA, Ramirez-Silva, JP, Loc-Barragan, J, Ponce Campos, P, Mata-Silva, V, Johnson, JD, Garcia-Padilla, E, Wilson, LD, 2016. The herpetofauna of Nayarit, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3: 375 - 448" title="The herpetofauna of Nayarit, Mexico: composition, distribution, and conservation status." volume="3" year="2016">
<normalizedToken originalValue="Woolrich-Piña">Woolrich-Pina</normalizedToken>
et al. 2016
</bibRefCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="common name">
<paragraph pageId="0" pageNumber="117">Common name.</paragraph>
<paragraph pageId="0" pageNumber="117">
<normalizedToken originalValue="María">Maria</normalizedToken>
Cleofas leaf-toed Gecko, Salamanquesa de la 
<collectingCountry name="Iceland">Isla</collectingCountry>
<normalizedToken originalValue="María">Maria</normalizedToken>
Cleofas.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="type species">
<paragraph pageId="0" pageNumber="117">
<typeStatus>Type</typeStatus>
species.
</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">
<emphasis italics="true" pageId="0" pageNumber="117">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3063617302" collectingDate="2018-05-24" collectingDateMax="2018-05-25" collectingDateMin="2018-05-24" collectorName="Ilse K. Barraza Soltero &amp; Armando H. Escobedo Galvan. Paratypes. All" latitude="21.3095" location="Maria Cleofas Island" longLatPrecision="7" longitude="-106.234" municipality="Maria Cleofas Island" specimenCode="MZFC-HE 35623, MZFC-HE 35618-35622" specimenCount="1" typeStatus="holotype">
Adult female (
<specimenCode collectionCode="MZFC-HE">MZFC-HE 35623</specimenCode>
) collected on 
<location LSID="urn:lsid:plazi:treatment:E0BD74DF82145003A5B61763858FD1E1:B4A9A3B38C22727D5776D9E75303EA40" latitude="21.3095" longLatPrecision="7" longitude="-106.234" municipality="Maria Cleofas Island" name="Maria Cleofas Island">
<normalizedToken originalValue="María">Maria</normalizedToken>
Cleofas Island
</location>
(
<geoCoordinate degrees="21.3095" direction="north" orientation="latitude" precision="5" value="21.3095">21.3095°N</geoCoordinate>
, 
<geoCoordinate degrees="106.2340" direction="west" orientation="longitude" precision="5" value="-106.234">106.2340°W</geoCoordinate>
, WGS84, 64 m elev.) on 
<date value="2018-05-24" valueMax="2018-05-25" valueMin="2018-05-24">
<collectingDate value="2018-05-24" valueMax="2018-05-25" valueMin="2018-05-24">24-25 May 2018</collectingDate>
</date>
by 
<collectorName>Ilse K. Barraza Soltero</collectorName>
and 
<collectorName>
Armando H. Escobedo 
<normalizedToken originalValue="Galván">Galvan</normalizedToken>
. 
<emphasis bold="true" pageId="0" pageNumber="117">
<emphasis italics="true" pageId="0" pageNumber="117">
<typeStatus>Paratypes</typeStatus>
</emphasis>
.
</emphasis>
All
</collectorName>
collected from the type locality, 
<collectingMunicipality>
<normalizedToken originalValue="María">Maria</normalizedToken>
Cleofas Island
</collectingMunicipality>
(Six adult specimens and two juveniles). 
<specimenCode collectionCode="MZFC-HE">MZFC-HE 35618-35622</specimenCode>
and
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3063617301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Ilse K. Barraza Soltero &amp; Armando H. Escobedo Galvan. Paratypes. All" isTailEnumeration="true" latitude="21.3095" location="Maria Cleofas Island" longLatPrecision="7" longitude="-106.234" municipality="Maria Cleofas Island" specimenCode="MZFC-HE 35624-35626" specimenCount="1" typeStatus="holotype">
<specimenCode collectionCode="MZFC-HE">MZFC-HE 35624-35626</specimenCode>
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/zookeys.1024.60473.figure7" httpUri="https://binary.pensoft.net/fig/520452" pageId="0" pageNumber="117" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 7.</emphasis>
Dorsal view of 
<taxonomicName authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus cleofasensis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus cleofasensis</emphasis>
</taxonomicName>
sp. nov., in life. (Photo by AHE-G).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="diagnosis">
<paragraph pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus cleofasensis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus cleofasensis</emphasis>
</taxonomicName>
is a species of medium to large body size. Snout-vent length of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
measured during fieldwork ranged from 44.8 to 77.0 mm (mean 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.95" unit="mm" value="59.5">59.5 mm</quantity>
). Concerning body size, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
differs (in mean size) from 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. magnus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="magnus">
<emphasis italics="true" pageId="0" pageNumber="117">P. magnus</emphasis>
</taxonomicName>
(73.03), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. nolascoensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="nolascoensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. nolascoensis</emphasis>
</taxonomicName>
(56.29), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. partidus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="partidus">
<emphasis italics="true" pageId="0" pageNumber="117">P. partidus</emphasis>
</taxonomicName>
(55.12) and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. saxatilis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="saxatilis">
<emphasis italics="true" pageId="0" pageNumber="117">P. saxatilis</emphasis>
</taxonomicName>
(65.44). 
<taxonomicName authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus cleofasensis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus cleofasensis</emphasis>
</taxonomicName>
has a white venter, variable dorsal coloration, and a greater number of paravertebral dorsal tubercles (mean 47.89). Three meristic characters mainly differentiate 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
from the rest of the species of the 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus saxatilis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="saxatilis">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus saxatilis</emphasis>
</taxonomicName>
clade (
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. saxatilis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="saxatilis">
<emphasis italics="true" pageId="0" pageNumber="117">P. saxatilis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. nolascoensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="nolascoensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. nolascoensis</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. partidus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="partidus">
<emphasis italics="true" pageId="0" pageNumber="117">P. partidus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. homolepidurus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="homolepidurus">
<emphasis italics="true" pageId="0" pageNumber="117">P. homolepidurus</emphasis>
</taxonomicName>
and 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus</emphasis>
</taxonomicName>
sp.) (Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Mean (and standard deviation) of morphometric characters (in millimeters) for the studied species." httpUri="http://table.plazi.org/id/77CED25F4DFB443B29ABA99018B292A1" pageId="0" pageNumber="117" tableUuid="77CED25F4DFB443B29ABA99018B292A1">3</tableCitation>
): paravertebral dorsal tubercles from head to tail (mean 47.89); number of scales across the snout, starting from the 3rd labial scale (mean 26.5); and number of longitudinal ventral scales from an imaginary line of the forelimbs to the cloacal opening (mean 61). 
<taxonomicName authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus cleofasensis" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus cleofasensis</emphasis>
</taxonomicName>
has the highest values of the mentioned characters, while the rest of the studied species presented lower counts compared to other 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus</emphasis>
</taxonomicName>
species in 
<collectingCountry name="Mexico">Mexico</collectingCountry>
(Tables 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean and (standard deviation) of meristic characters examined for species studied. In bold we show the set of diagnostic meristic characters of P. cleofasensis sp. nov." httpUri="http://table.plazi.org/id/8DA7767F59C61BEA0AC24964AC6A0247" pageId="0" pageNumber="117" tableUuid="8DA7767F59C61BEA0AC24964AC6A0247">2</tableCitation>
and 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Mean (and standard deviation) of morphometric characters (in millimeters) for the studied species." httpUri="http://table.plazi.org/id/77CED25F4DFB443B29ABA99018B292A1" pageId="0" pageNumber="117" tableUuid="77CED25F4DFB443B29ABA99018B292A1">3</tableCitation>
). The new species has the second highest mean number of dorsal tubercles after 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. delcampi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="delcampi">
<emphasis italics="true" pageId="0" pageNumber="117">P. delcampi</emphasis>
</taxonomicName>
(63.4). Other 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus</emphasis>
</taxonomicName>
species have fewer mean dorsal tubercles (
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. paucituberculatus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="paucituberculatus">
<emphasis italics="true" pageId="0" pageNumber="117">P. paucituberculatus</emphasis>
</taxonomicName>
, 28.7; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. duellmani" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="duellmani">
<emphasis italics="true" pageId="0" pageNumber="117">P. duellmani</emphasis>
</taxonomicName>
, 37.1; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. bordai" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="bordai">
<emphasis italics="true" pageId="0" pageNumber="117">P. bordai</emphasis>
</taxonomicName>
, 32.9; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. davisi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="davisi">
<emphasis italics="true" pageId="0" pageNumber="117">P. davisi</emphasis>
</taxonomicName>
, 41.6; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. muralis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="muralis">
<emphasis italics="true" pageId="0" pageNumber="117">P. muralis</emphasis>
</taxonomicName>
, 33.1; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. homolepidurus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="homolepidurus">
<emphasis italics="true" pageId="0" pageNumber="117">P. homolepidurus</emphasis>
</taxonomicName>
, 35.9; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. xanti" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="xanti">
<emphasis italics="true" pageId="0" pageNumber="117">P. xanti</emphasis>
</taxonomicName>
, 37; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lanei" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lanei">
<emphasis italics="true" pageId="0" pageNumber="117">P. lanei</emphasis>
</taxonomicName>
, 32.4; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. papenfussi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="papenfussi">
<emphasis italics="true" pageId="0" pageNumber="117">P. papenfussi</emphasis>
</taxonomicName>
, 33.2; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. isabelae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="isabelae">
<emphasis italics="true" pageId="0" pageNumber="117">P. isabelae</emphasis>
</taxonomicName>
, 32.3; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lupitae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lupitae">
<emphasis italics="true" pageId="0" pageNumber="117">P. lupitae</emphasis>
</taxonomicName>
, 28.8; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. rupinus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="rupinus">
<emphasis italics="true" pageId="0" pageNumber="117">P. rupinus</emphasis>
</taxonomicName>
, 28; 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. benedettii" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="benedettii">
<emphasis italics="true" pageId="0" pageNumber="117">P. benedettii</emphasis>
</taxonomicName>
, 28.8; and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. kropotkini" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="kropotkini">
<emphasis italics="true" pageId="0" pageNumber="117">P. kropotkini</emphasis>
</taxonomicName>
, 28.4). Regarding the number of scales crossing the snout, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
has a very similar number of scales to 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. muralis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="muralis">
<emphasis italics="true" pageId="0" pageNumber="117">P. muralis</emphasis>
</taxonomicName>
(26.1) and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lupitae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lupitae">
<emphasis italics="true" pageId="0" pageNumber="117">P. lupitae</emphasis>
</taxonomicName>
(25.5); lower counts occur in 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. unctus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="unctus">
<emphasis italics="true" pageId="0" pageNumber="117">P. unctus</emphasis>
</taxonomicName>
(21.7), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. paucituberculatus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="paucituberculatus">
<emphasis italics="true" pageId="0" pageNumber="117">P. paucituberculatus</emphasis>
</taxonomicName>
(19.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. duellmani" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="duellmani">
<emphasis italics="true" pageId="0" pageNumber="117">P. duellmani</emphasis>
</taxonomicName>
(19.9), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. delcampi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="delcampi">
<emphasis italics="true" pageId="0" pageNumber="117">P. delcampi</emphasis>
</taxonomicName>
(21), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. bordai" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="bordai">
<emphasis italics="true" pageId="0" pageNumber="117">P. bordai</emphasis>
</taxonomicName>
(19.4), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. davisi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="davisi">
<emphasis italics="true" pageId="0" pageNumber="117">P. davisi</emphasis>
</taxonomicName>
(23.7), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. muralis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="muralis">
<emphasis italics="true" pageId="0" pageNumber="117">P. muralis</emphasis>
</taxonomicName>
(26.1), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. homolepidurus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="homolepidurus">
<emphasis italics="true" pageId="0" pageNumber="117">P. homolepidurus</emphasis>
</taxonomicName>
(22.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. xanti" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="xanti">
<emphasis italics="true" pageId="0" pageNumber="117">P. xanti</emphasis>
</taxonomicName>
(16.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lanei" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lanei">
<emphasis italics="true" pageId="0" pageNumber="117">P. lanei</emphasis>
</taxonomicName>
(21.6), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. papenfussi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="papenfussi">
<emphasis italics="true" pageId="0" pageNumber="117">P. papenfussi</emphasis>
</taxonomicName>
(17.8), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. isabelae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="isabelae">
<emphasis italics="true" pageId="0" pageNumber="117">P. isabelae</emphasis>
</taxonomicName>
(21.1), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. rupinus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="rupinus">
<emphasis italics="true" pageId="0" pageNumber="117">P. rupinus</emphasis>
</taxonomicName>
(20.6), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. benedettii" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="benedettii">
<emphasis italics="true" pageId="0" pageNumber="117">P. benedettii</emphasis>
</taxonomicName>
(22), and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. kropotkini" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="kropotkini">
<emphasis italics="true" pageId="0" pageNumber="117">P. kropotkini</emphasis>
</taxonomicName>
(20.2). Concerning ventral scales, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
has a mean of 61, similar to three species of the 
<taxonomicName authorityName="Ramírez-Reyes &amp; Barraza-Soltero &amp; Nolasco-Luna &amp; Flores-Villela &amp; Escobedo-Galván" authorityYear="2021" class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus lanei" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="lanei">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus lanei</emphasis>
</taxonomicName>
clade (or clade I), namely, 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lupitae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lupitae">
<emphasis italics="true" pageId="0" pageNumber="117">P. lupitae</emphasis>
</taxonomicName>
(61.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. rupinus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="rupinus">
<emphasis italics="true" pageId="0" pageNumber="117">P. rupinus</emphasis>
</taxonomicName>
(62) and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. lanei" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="lanei">
<emphasis italics="true" pageId="0" pageNumber="117">P. lanei</emphasis>
</taxonomicName>
(62.86). The rest of the 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus</emphasis>
</taxonomicName>
in 
<collectingCountry name="Mexico">Mexico</collectingCountry>
have less than 61 scales: 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. kropotkini" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="kropotkini">
<emphasis italics="true" pageId="0" pageNumber="117">P. kropotkini</emphasis>
</taxonomicName>
(60), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. benedettii" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="benedettii">
<emphasis italics="true" pageId="0" pageNumber="117">P. benedettii</emphasis>
</taxonomicName>
(59.71), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. xanti" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="xanti">
<emphasis italics="true" pageId="0" pageNumber="117">P. xanti</emphasis>
</taxonomicName>
(58.75), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. angelensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="angelensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. angelensis</emphasis>
</taxonomicName>
(56.75), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. davisi" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="davisi">
<emphasis italics="true" pageId="0" pageNumber="117">P. davisi</emphasis>
</taxonomicName>
(55.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. magnus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="magnus">
<emphasis italics="true" pageId="0" pageNumber="117">P. magnus</emphasis>
</taxonomicName>
(55.3), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. partidus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="partidus">
<emphasis italics="true" pageId="0" pageNumber="117">P. partidus</emphasis>
</taxonomicName>
(55.2), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. saxatilis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="saxatilis">
<emphasis italics="true" pageId="0" pageNumber="117">P. saxatilis</emphasis>
</taxonomicName>
(55), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. santacruzensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="santacruzensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. santacruzensis</emphasis>
</taxonomicName>
(54), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. muralis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="muralis">
<emphasis italics="true" pageId="0" pageNumber="117">P. muralis</emphasis>
</taxonomicName>
(53.9), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. isabelae" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="isabelae">
<emphasis italics="true" pageId="0" pageNumber="117">P. isabelae</emphasis>
</taxonomicName>
(53.85), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. nolascoensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="nolascoensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. nolascoensis</emphasis>
</taxonomicName>
(52.75), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. homolepidurus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="homolepidurus">
<emphasis italics="true" pageId="0" pageNumber="117">P. homolepidurus</emphasis>
</taxonomicName>
(52.5), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. bordai" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="bordai">
<emphasis italics="true" pageId="0" pageNumber="117">P. bordai</emphasis>
</taxonomicName>
(51.91), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. bugastrolepis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="bugastrolepis">
<emphasis italics="true" pageId="0" pageNumber="117">P. bugastrolepis</emphasis>
</taxonomicName>
(51.57), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. duellmani" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="duellmani">
<emphasis italics="true" pageId="0" pageNumber="117">P. duellmani</emphasis>
</taxonomicName>
(48.75), 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. paucituberculatus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="paucituberculatus">
<emphasis italics="true" pageId="0" pageNumber="117">P. paucituberculatus</emphasis>
</taxonomicName>
(48.66), and 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. unctus" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="unctus">
<emphasis italics="true" pageId="0" pageNumber="117">P. unctus</emphasis>
</taxonomicName>
(48.5).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="description">
<paragraph pageId="0" pageNumber="117">
Description of the 
<typeStatus>holotype</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="117">
Adult female with SVL 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="7.117" unit="mm" value="71.17">71.17 mm</quantity>
, robust body, head not flattened, neck slightly differentiated from head. Head width 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.44" unit="mm" value="14.4">14.4 mm</quantity>
and snout length 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.2109999999999999" unit="mm" value="12.11">12.11 mm</quantity>
. Rostral scale is flat (no grooves or stretch marks) and in contact with the two internasal scales. Nostril in contact mostly with the rostral scale and marginally with the first labial scale on both sides; supranasal scales in contact with 12 scales crossing from right to left side; 22 interorbital scales counted from middle of eye, interorbital distance 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="9.75" unit="mm" value="9.75">9.75 mm</quantity>
. 24 scales crossing snout between contralateral second labial scales 27 between third labial scales. Number of loreal scales 14 on right side and 15 on left side; equal number of supralabial scales (14), labial scales (8), and infralabial scales (7) on each side. Auricular opening oval (
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.48" unit="mm" value="2.48">2.48 mm</quantity>
) smaller than the ocular opening, occipital scales similar in size and shape to interorbitals (not greatly differentiated in size and shape). Mental scale slightly wider than long, forming a 
<normalizedToken originalValue="“V”">
&quot; 
<collectionCode country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" type="Museum">V</collectionCode>
&quot;
</normalizedToken>
but not with pronounced angles; eight postmental scales in contact with first infralabials on both sides (right and left). Body with granular and circular scales interspersed with tubercles of different sizes; the specimen presents a fragmented tail. Internarinal distance 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.67" unit="mm" value="2.67">2.67 mm</quantity>
and axilla-groin length 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.619" unit="mm" value="26.19">26.19 mm</quantity>
. 14 rows of dorsal tubercles, 20 axilla-groin tubercles on right side and 19 on left side. Presents 29 transversal ventral scales with first ventral scale differentiated from lateral scales (which are small and circular). Ventral scales differentiated in size and shape from lateral and gular scales. Scales imbricate on extremities (anterior and posterior), as well as on dorsal region of the tail. No femoral or precloacal pores. Digital lamella formulae: right posterior (9-11-15-14-13), left posterior (8-10-11-12-13), right anterior (8-11-12-13-10), left anterior (8-10-12-13-11); fourth finger of extremities longer that others (
<quantity metricMagnitude="-3" metricUnit="m" metricValue="5.55" unit="mm" value="5.55">5.55 mm</quantity>
manus, 
<quantity metricMagnitude="-3" metricUnit="m" metricValue="7.49" unit="mm" value="7.49">7.49 mm</quantity>
pes); digital toepads longer than wide on all fingers.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="etymology">
<paragraph pageId="0" pageNumber="117">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="117">
Specific epithet is taken from the 
<typeStatus>type</typeStatus>
locality 
<normalizedToken originalValue="María">Maria</normalizedToken>
Cleofas Island, with the Latin suffix - 
<emphasis italics="true" pageId="0" pageNumber="117">ensis</emphasis>
meaning, &quot;originating from.&quot; Specific epithet is masculine, in agreement with the gender of 
<taxonomicName class="Reptilia" family="Phyllodactylidae" genus="Phyllodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phyllodactylus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Phyllodactylus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="variation">
<paragraph pageId="0" pageNumber="117">Variation.</paragraph>
<paragraph pageId="0" pageNumber="117">
All meristic and morphometric characters are presented with mean values and standard deviation in Tables 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean and (standard deviation) of meristic characters examined for species studied. In bold we show the set of diagnostic meristic characters of P. cleofasensis sp. nov." httpUri="http://table.plazi.org/id/8DA7767F59C61BEA0AC24964AC6A0247" pageId="0" pageNumber="117" tableUuid="8DA7767F59C61BEA0AC24964AC6A0247">2</tableCitation>
and 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Mean (and standard deviation) of morphometric characters (in millimeters) for the studied species." httpUri="http://table.plazi.org/id/77CED25F4DFB443B29ABA99018B292A1" pageId="0" pageNumber="117" tableUuid="77CED25F4DFB443B29ABA99018B292A1">3</tableCitation>
, respectively.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="natural history">
<paragraph pageId="0" pageNumber="117">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="117">
Individuals of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
were observed active during night surveys under single rocks, abandoned anthropogenic structures, or in some cases on the trunk of 
<taxonomicName class="Magnoliopsida" family="Picrodendraceae" genus="Piranhea" higherTaxonomySource="GBIF,IPNI" kingdom="Plantae" lsidName="Piranhea mexicana" order="Malpighiales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis italics="true" pageId="0" pageNumber="117">Piranhea mexicana</emphasis>
</taxonomicName>
. Some were observed on the ground while they moved between rocks. In some parts of the island, they can be seen in abandoned man-made structures, which are shelters for species such as geckos, anoles, iguanids, and bats. Although predation of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
has not been reported, we suggest that 
<taxonomicName authorityName="Barbour &amp; Amaral" authorityYear="1926" class="Reptilia" family="Colubridae" genus="Oxybelis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Oxybelis microphthalmus" order="Squamata" pageId="0" pageNumber="117" phylum="Chordata" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="117">Oxybelis microphthalmus</emphasis>
</taxonomicName>
(for taxonomic status see 
<bibRefCitation DOI="https://doi.org/10.1007/s13127-020-00461-0" author="Jadin, RC" journalOrPublisher="Organisms Diversity &amp; Evolution" pageId="0" pageNumber="117" pagination="723 - 746" refId="B19" refString="Jadin, RC, Blair, C, Orlofske, SA, Jowers, MM, Rivas, GA, Vitt, LJ, Ray, JM, Smith, EN, Murphy, JC, 2020. Not withering on the evolutionary vine: systematic revision of the Brown Vine Snake (Reptilia: Squamata: Oxybelis) from its northern distribution. Organisms Diversity &amp; Evolution 20: 723 - 746, DOI: https://doi.org/10.1007/s13127-020-00461-0" title="Not withering on the evolutionary vine: systematic revision of the Brown Vine Snake (Reptilia: Squamata: Oxybelis) from its northern distribution." url="https://doi.org/10.1007/s13127-020-00461-0" volume="20" year="2020">Jadin et al. 2020</bibRefCitation>
) could be a potential predator given that we captured two 
<taxonomicName lsidName="O. microphthalmus" pageId="0" pageNumber="117" rank="species" species="microphthalmus">
<emphasis italics="true" pageId="0" pageNumber="117">O. microphthalmus</emphasis>
</taxonomicName>
at the same site of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
during nocturnal surveys. During our visits (
<date value="2017-05" valueMax="2017-08-31" valueMin="2017-05-00">May to August 2017</date>
and 
<date value="2018-04" valueMax="2018-08-31" valueMin="2018-04-00">April to August 2018</date>
), we did not observe reproductive activity or females with a shelled egg in one of their oviducts. Eleven distinct prey items were found in 36 stomach contents. It has been mentioned that the diets of lizards in island environments contain a high percentage of vegetation matter due to the lack of food or scarcity of prey belonging to the Class 
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="" pageId="0" pageNumber="117" phylum="Nematoda" rank="class">Insecta</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1016/S0169-5347(03)00004-1" author="Olesen, JM" journalOrPublisher="Trends in Ecology and Evolution" pageId="0" pageNumber="117" pagination="177 - 181" refId="B28" refString="Olesen, JM, Valido, A, 2003. Lizards as pollinators and seed dispersers: an island phenomenon. Trends in Ecology and Evolution 18 (4): 177 - 181, DOI: https://doi.org/10.1016/S0169-5347(03)00004-1" title="Lizards as pollinators and seed dispersers: an island phenomenon." url="https://doi.org/10.1016/S0169-5347(03)00004-1" volume="18" year="2003">Olesen and Valido 2003</bibRefCitation>
). However, in this work, a great variety of arthropods was reported in the stomach contents of geckos (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Prey items identified from stomach contents of P. cleofasensis A Pycnoselus surinamensis B crickets of the family Rhaphidophoridae C beetles of the genus Telabis D Centruroides elegans insularis E Euphoria germinata F and G beetles of the family Elateridae H spider of the family Caponiidae I some individuals of lepidopterans." figureDoi="10.3897/zookeys.1024.60473.figure8" httpUri="https://binary.pensoft.net/fig/520453" pageId="0" pageNumber="117">8</figureCitation>
). Prey items of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
were mostly composed of orthopterans (Family 
<taxonomicName class="Insecta" family="Rhaphidophoridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
) and coleopterans (Family 
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Passalidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="117" rank="family">Passalidae</taxonomicName>
); they also consumed plant matter, arachnids, lepidopterans, scorpions (
<taxonomicName authorityName="Pocock" authorityYear="1902" class="Insecta" family="Buthidae" genus="Centruroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centruroides elegans subsp. insularis" order="Blattodea" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="insularis">
<emphasis italics="true" pageId="0" pageNumber="117">Centruroides elegans insularis</emphasis>
</taxonomicName>
), and cockroaches (
<taxonomicName class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Blattodea" pageId="0" pageNumber="117" rank="order">Blattodea</taxonomicName>
: 
<taxonomicName genus="Pycnoselus" lsidName="Pycnoselus surinamensis" pageId="0" pageNumber="117" rank="species" species="surinamensis">
<emphasis italics="true" pageId="0" pageNumber="117">Pycnoselus surinamensis</emphasis>
</taxonomicName>
) (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Prey items identified from stomach contents of P. cleofasensis A Pycnoselus surinamensis B crickets of the family Rhaphidophoridae C beetles of the genus Telabis D Centruroides elegans insularis E Euphoria germinata F and G beetles of the family Elateridae H spider of the family Caponiidae I some individuals of lepidopterans." figureDoi="10.3897/zookeys.1024.60473.figure8" httpUri="https://binary.pensoft.net/fig/520453" pageId="0" pageNumber="117">8</figureCitation>
). Additionally, during fieldwork in 
<date value="2018-05">May 2018</date>
, we captured two individuals with remains of shed skin in their stomach contents (
<bibRefCitation author="Barraza-Soltero, IK" journalOrPublisher="Biharean Biologist" pageId="0" pageNumber="117" pagination="33 - 35" refId="B3" refString="Barraza-Soltero, IK, Escobedo-Galvan, AH, 2020. Keratophagy in two lizards from Isla Maria Cleofas, Mexico. Biharean Biologist 14 (1): 33 - 35" title="Keratophagy in two lizards from Isla Maria Cleofas, Mexico." volume="14" year="2020">
Barraza-Soltero and 
<normalizedToken originalValue="Escobedo-Galván">Escobedo-Galvan</normalizedToken>
2020
</bibRefCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1024.60473.figure8" httpUri="https://binary.pensoft.net/fig/520453" pageId="0" pageNumber="117" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 8.</emphasis>
Prey items identified from stomach contents of 
<taxonomicName class="Insecta" family="Phyllodactylidae" kingdom="Animalia" lsidName="P. cleofasensis" order="Blattodea" pageId="0" pageNumber="117" phylum="Nematoda" rank="species" species="cleofasensis">
<emphasis italics="true" pageId="0" pageNumber="117">P. cleofasensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">A</emphasis>
<taxonomicName genus="Pycnoselus" lsidName="Pycnoselus surinamensis" pageId="0" pageNumber="117" rank="species" species="surinamensis">
<emphasis italics="true" pageId="0" pageNumber="117">Pycnoselus surinamensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">B</emphasis>
crickets of the family 
<taxonomicName class="Insecta" family="Rhaphidophoridae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Orthoptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="family">Rhaphidophoridae</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">C</emphasis>
beetles of the genus 
<taxonomicName authorityName="Casey" authorityYear="1890" class="Insecta" family="Tenebrionidae" genus="Telabis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Telabis" order="Coleoptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Telabis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">D</emphasis>
<taxonomicName authorityName="Pocock" authorityYear="1902" class="Insecta" family="Buthidae" genus="Centruroides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Centruroides elegans subsp. insularis" order="Blattodea" pageId="0" pageNumber="117" phylum="Arthropoda" rank="subSpecies" species="elegans" subSpecies="insularis">
<emphasis italics="true" pageId="0" pageNumber="117">Centruroides elegans insularis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">E</emphasis>
<taxonomicName class="Insecta" family="Cetoniidae" genus="Euphoria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Euphoria germinata" order="Coleoptera" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="germinata">
<emphasis italics="true" pageId="0" pageNumber="117">Euphoria germinata</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">F</emphasis>
and 
<emphasis bold="true" pageId="0" pageNumber="117">G</emphasis>
beetles of the family 
<taxonomicName authorityName="Leach" authorityYear="1815" class="Insecta" family="Elateridae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="117" rank="family">Elateridae</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">H</emphasis>
spider of the family 
<taxonomicName authorityName="Simon" authorityYear="1890" class="Arachnida" family="Caponiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="family">Caponiidae</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="117">I</emphasis>
some individuals of lepidopterans.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>