334 lines
24 KiB
XML
334 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit & Jochum, Adrienne" docDate="2023" docId="92466D68DA315792829018A068703ACE" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila babel Pall-Gergely & Vermeulen 2023, sp. nov." docType="treatment" docUuid="405BA4E4-A738-488E-8061-4EDEA99667E3" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunyadi, Andras</mods:namePart>
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<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
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<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grego, Jozef</mods:namePart>
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<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reischuetz, Alexander</mods:namePart>
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<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
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<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Oerstan, Aydin</mods:namePart>
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<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fekete, Judit</mods:namePart>
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<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jochum, Adrienne</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
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<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-13</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1147</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>177</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1147-1</mods:identifier>
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<mods:identifier type="ZooBank">9BB9881B0076473D8E53155D37CA1F50</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FF2B6B317B505F9EA0E1000BDCD16CE7</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:405BA4E4-A738-488E-8061-4EDEA99667E3" httpUri="http://treatment.plazi.org/id/92466D68DA315792829018A068703ACE" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="https://zoobank.org/405BA4E4-A738-488E-8061-4EDEA99667E3" authority="Páll-Gergely & Vermeulen" authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila babel" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="babel" status="sp. nov.">
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Angustopila babel
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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& Vermeulen
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Angustopila babel Pall-Gergely & Vermeulen, sp. nov. (holotype, RMNH 347765). Apertural (A), ventral (B), lateral (C) and apical (D) sides of the shell; microstructure of the protoconch (E), ventral (F) and frontal (G) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure28" httpUri="https://binary.pensoft.net/fig/808837" pageId="0" pageNumber="1">Figs 28</figureCitation>
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<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Variability of Angustopila fabella Pall-Gergely & Hunyadi, 2015 in the Halong Bay A JJV 6217 B WMVT. 0338 and Angustopila babel Pall-Gergely & Vermeulen, sp. nov. C JJV 16590 D JJV 17634, Specimen 1 E JJV 17634, Specimen 2." figureDoi="10.3897/zookeys.1147.93824.figure29" httpUri="https://binary.pensoft.net/fig/808838" pageId="0" pageNumber="1">, 29C-E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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: Vietnam • 1 empty shell (H: 1.07 mm, D: 0.97 mm); Quang Ninh Province, Halong Bay area, unnamed island in Dau Moi Temple area (locality code: WMVT.0328);
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<geoCoordinate degrees="20" direction="north" minutes="55.69" orientation="latitude" precision="9" value="20.928167">20°55.69'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="09.40" orientation="longitude" precision="9" value="107.15667">107°09.40'E</geoCoordinate>
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; 13 Sep. 2003; W.J.M. Maassen leg.; RMNH 347765.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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: Vietnam • 12 shells; same data as for holotype; RMNH 347766 • 491 shells; Quang Ninh Province, Halong Bay, Cap La Cave, deposit of soil fallen in through roof in pristine cave, vegetation outside cave tall and woody;
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<geoCoordinate degrees="20" direction="north" minutes="51.79" orientation="latitude" precision="9" value="20.863167">20°51.79'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="13.54" orientation="longitude" precision="9" value="107.22567">107°13.54'E</geoCoordinate>
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; 7 Mar. 2018; J.J. Vermeulen & K. Anker leg.; JJV 17632 • 10 shells; same data as for preceding; coll. HA • 20 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, drift material washed together over sinkhole in cave;
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<geoCoordinate degrees="20" direction="north" minutes="50.83" orientation="latitude" precision="9" value="20.847166">20°50.83'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="05.67" orientation="longitude" precision="9" value="107.0945">107°05.67'E</geoCoordinate>
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; J.J. Vermeulen & A.J. Whitten leg.; 2 Oct. 1998; JJV 6220 • 12 shells; Quang Ninh Province, Halong Bay area, Dao Bo Hon, Song Sot Cave, guano enriched sediments in cave;
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<geoCoordinate degrees="20" direction="north" minutes="50.83" orientation="latitude" precision="9" value="20.847166">20°50.83'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="05.67" orientation="longitude" precision="9" value="107.0945">107°05.67'E</geoCoordinate>
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; 2 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6221 • 1 shell; Haiphong Province, Cat Ba Island, Cave Qua Vang, around cave entrance, rocky limestone slope with low, somewhat mature forest;
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<geoCoordinate degrees="20" direction="north" minutes="48.64" orientation="latitude" precision="9" value="20.810667">20°48.64'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="04.64" orientation="longitude" precision="9" value="107.07733">107°04.64'E</geoCoordinate>
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; 100 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 17657 (ex JJV 16612) • 20 shells; Haiphong Province, Cat Ba Archipelago, Dao Dau Be, rocks and small soil deposits bordering beach;
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<geoCoordinate degrees="20" direction="north" minutes="45.68" orientation="latitude" precision="9" value="20.761333">20°45.68'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="07.47" orientation="longitude" precision="9" value="107.1245">107°07.47'E</geoCoordinate>
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; 10 m a.s.l.; 8 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16591 • 73 shells; Haiphong Province, Cat Ba Island, Cave Qua Vang, inside cave, large, ecologically intact active cave with speleothems;
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<geoCoordinate degrees="20" direction="north" minutes="48.64" orientation="latitude" precision="9" value="20.810667">20°48.64'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="04.64" orientation="longitude" precision="9" value="107.07733">107°04.64'E</geoCoordinate>
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; 60 m a.s.l.; 6 Jun. 2017; J.J. Vermeulen & K. Anker leg.; JJV 16590 • 1 shell; Quang Ninh Province, Halong Bay area, May Den Island, Tam Cung Cave, densely-vegetated limestone hill, near cave; approx. GPS coordinates:
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<geoCoordinate degrees="20" direction="north" minutes="52.33" orientation="latitude" precision="9" value="20.872168">20°52.33'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="07.09" orientation="longitude" precision="9" value="107.118164">107°07.09'E</geoCoordinate>
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; 3 Oct. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6248 • 2 shells; Quang Ninh Province, Halong-Campha area, 2.5 km SW Quang Hanh, foot of limestone cliff with degraded regrowth, near gardens;
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<geoCoordinate degrees="20" direction="north" minutes="59.750" orientation="latitude" precision="1" value="20.995832">20°59.750'N</geoCoordinate>
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,
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<geoCoordinate degrees="107" direction="east" minutes="12.750" orientation="longitude" precision="1" value="107.2125">107°12.750'E</geoCoordinate>
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; 29 Sep. 1998; J.J. Vermeulen & A.J. Whitten leg.; JJV 6218.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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A medium-sized to large
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<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species with a conical shell, regularly growing whorls, a strong parietal fold reaching the peristome and a narrow umbilicus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell medium-sized to large-sized for the genus, higher than wide, very rarely slightly wider than high; conical, body whorl widest in standard apertural view; protoconch consists of 1.25 whorls with weak spiral striae preceding the first teleoconch whorl; teleoconch with very fine irregularly spaced, weak radial growth lines crossed with equally strong, rather regularly and sparsely-spaced spiral striae (ca. 12 on body whorl from apertural view); on both ventral and dorsal surfaces of body whorl spiral lines dominant or some radial lines are of comparable strength; sculpture overall weaker than in most other
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<taxonomicName authorityName="Jochum, Slapnik & Pall-Gergely" authorityYear="2014" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
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</taxonomicName>
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species; whorls ca. 3.25-4, rounded or very slightly shouldered; aperture oblique to shell axis from lateral view; umbilicus narrow; aperture ovate-oblong, parietal callus straight; sinulus very wide, weakly isolated due to the rather low parietal tooth; peristome expanded, not reflected; parietal callus only very slightly detached from penultimate whorl, not smeared onto it; parietal tooth moderately elevated, of normal length, reaches parietal callus (in some, probably subadult shells it does not), perpendicular to parietal wall.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
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<paragraph pageId="0" pageNumber="1">
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H = 0.94-1.17, D = 0.92-1.16, H/D*100 = 89.0-121.7 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 18), RUD = 22.1-25.2 (
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<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
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= 8).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shells of
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<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
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<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
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</taxonomicName>
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lots in the Halong Bay area differ from
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<taxonomicName lsidName="A. babel" pageId="0" pageNumber="1" rank="species" species="babel">
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<emphasis italics="true" pageId="0" pageNumber="1">A. babel</emphasis>
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</taxonomicName>
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sp. nov. by having a concave-conical shell (instead of the conical shape of
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<taxonomicName lsidName="A. babel" pageId="0" pageNumber="1" rank="species" species="babel">
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<emphasis italics="true" pageId="0" pageNumber="1">A. babel</emphasis>
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</taxonomicName>
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sp. nov.), a wider umbilicus and a more deeply set parietal tooth.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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From
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<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
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<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
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</taxonomicName>
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populations living outside of the Halong Bay area,
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<taxonomicName lsidName="A. babel" pageId="0" pageNumber="1" rank="species" species="babel">
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<emphasis italics="true" pageId="0" pageNumber="1">A. babel</emphasis>
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</taxonomicName>
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sp. nov. can be distinguished by the following traits:
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<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila babel" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="babel">
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<emphasis italics="true" pageId="0" pageNumber="1">Angustopila babel</emphasis>
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</taxonomicName>
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sp. nov. is usually larger, has a less pointed (rather blunt) apex, and a comparatively larger aperture, a narrower umbilicus, and the parietal tooth reaches the parietal callus. Some
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<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
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<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
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</taxonomicName>
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populations are, however, strikingly similar in shape. For example, sample 2020/58, which is smaller and has a more deeply situated parietal tooth, and sample 2020/32, which is also smaller, and has a more pointed apex and a smaller aperture. See also under
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<taxonomicName lsidName="A. fratermajor" pageId="0" pageNumber="1" rank="species" species="fratermajor">
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<emphasis italics="true" pageId="0" pageNumber="1">A. fratermajor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
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|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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|
<paragraph pageId="0" pageNumber="1">Named after the conical shape, resembling the Tower of Babel as depicted by Pieter Breughel the Elder (ca. 1563). To be used as a noun in apposition.</paragraph>
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</subSubSection>
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|
<subSubSection pageId="0" pageNumber="1" type="distribution">
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|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Known from several populations in the Halong Bay area only (Fig.
|
|
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of Angustopila apiostoma Pall-Gergely & Vermeulen, sp. nov. and Angustopila babel Pall-Gergely & Vermeulen, sp. nov." figureDoi="10.3897/zookeys.1147.93824.figure11" httpUri="https://binary.pensoft.net/fig/808820" pageId="0" pageNumber="1">11</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
Conical or concave-conical shells with a single parietal tooth belong to two distinct morphological groups in the Halong Bay Area. Both types would otherwise fit the morphological continuum of the widespread and variable
|
|
<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila fabella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fabella">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila fabella</emphasis>
|
|
</taxonomicName>
|
|
. However, since the two types are clearly distinct within such a small area, only one of them is identified
|
|
<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
|
|
</taxonomicName>
|
|
, and the other one must be a distinct species. There are three populations in the Halong Bay area with concave-conical shells bearing a single parietal tooth. Individuals from these populations are identified here as
|
|
<taxonomicName lsidName="A. fabella" pageId="0" pageNumber="1" rank="species" species="fabella">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">A. fabella</emphasis>
|
|
</taxonomicName>
|
|
. Conversely, the other populations of morphospecies that are conical, have a blunt apex, and possess a single parietal tooth are described as
|
|
<taxonomicName lsidName="A. babel" pageId="0" pageNumber="1" rank="species" species="babel">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">A. babel</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</paragraph>
|
|
<caption doi="10.3897/zookeys.1147.93824.figure28" httpUri="https://binary.pensoft.net/fig/808837" pageId="0" pageNumber="1" start="Figure 28" startId="F28">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 28.</emphasis>
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila babel" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="babel">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila babel</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov. (holotype, RMNH 347765). Apertural (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
), ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
), lateral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
) and apical (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
) sides of the shell; microstructure of the protoconch (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
), ventral (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
|
) and frontal (
|
|
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
|
|
) surface of the body whorl.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/zookeys.1147.93824.figure29" httpUri="https://binary.pensoft.net/fig/808838" pageId="0" pageNumber="1" start="Figure 29" startId="F29">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">Figure 29.</emphasis>
|
|
Variability of
|
|
<taxonomicName authorityName="Pall-Gergely & Hunyadi" authorityYear="2015" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila fabella" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="fabella">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila fabella</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Hunyadi, 2015 in the Halong Bay
|
|
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
|
JJV 6217
|
|
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
|
WMVT.0338 and
|
|
<taxonomicName authorityName="Pall-Gergely & Vermeulen" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila babel" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="babel">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila babel</emphasis>
|
|
</taxonomicName>
|
|
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
|
|
& Vermeulen, sp. nov.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
|
JJV 16590
|
|
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
|
JJV 17634, Specimen 1
|
|
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
|
JJV 17634, Specimen 2.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |