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<mods:title id="95E147CADD9C42B304FB70585BB3D366">A revision and addition to Zopheridae (Coleoptera: Tenebrionoidea) in Baltic amber: possible connections between modern Holarctic distributions and Eocene amber forests</mods:title>
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<mods:namePart id="181D87A6D9B8325C74B85008017B137F">Alekseev, Vitalii</mods:namePart>
<mods:affiliation id="CB70947EF65EDF7DF5B216463015D73E">Immanuel Kant Baltic Federal University, Nevskogo 14, Kaliningrad 236016, Russia. &amp; Royal Saskatchewan Museum, 2445 Albert St., Regina, SK, S 4 P 4 W 7, Canada. &amp; Biology Department, University of Regina, Regina, SK, S 4 S 0 A 2, Canada; &amp; Department of Ecology &amp; Evolutionary Biology, University of Kansas, Lawrence, Kansas, 66045, USA;</mods:affiliation>
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<mods:namePart id="A56474B326EE56CCD82FFA21E8F93074">Bukejs, Andris</mods:namePart>
<mods:affiliation id="10F461C528D1E02197EDC3D6349C4ADC">Institute of Life Sciences and Technologies, Daugavpils University, Vienības 13, Daugavpils LV- 5401, Latvia.</mods:affiliation>
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<paragraph id="8BE26E2EFFE9664AFF506AD6BA8356B6" blockId="9.[151,456,151,178]" box="[151,456,151,178]" pageId="9" pageNumber="210">
<heading id="D0AAD942FFE9664AFF506AD6BA8356B6" bold="true" box="[151,456,151,178]" fontSize="11" level="1" pageId="9" pageNumber="210" reason="1">
<emphasis id="B929B23CFFE9664AFF506AD6BA8356B6" bold="true" box="[151,456,151,178]" pageId="9" pageNumber="210">
Genus
<taxonomicName id="4C5D15ADFFE9664AFF296AD6BA8356B6" ID-CoL="63JML" authority="Dajoz, 1984" authorityName="Dajoz" authorityYear="1984" box="[238,456,151,178]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE9664AFF296AD6BA6556B5" bold="true" box="[238,302,151,178]" italics="true" pageId="9" pageNumber="210">Paha</emphasis>
<bibRefCitation id="EFCC13DFFFE9664AFEF26AD6BA8356B6" author="Dajoz, R." box="[309,456,151,177]" pageId="9" pageNumber="210" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz, 1984</bibRefCitation>
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<paragraph id="8BE26E2EFFE9664AFF506AA1B9E256FE" blockId="9.[151,681,223,250]" box="[151,681,223,250]" pageId="9" pageNumber="210">
<typeStatus id="54E6D08CFFE9664AFF506AA1BB8456FD" box="[151,207,224,250]" pageId="9" pageNumber="210">Type</typeStatus>
species:
<taxonomicName id="4C5D15ADFFE9664AFEF26A9EB9E256FE" authority="Dajoz, 1984" authorityName="Dajoz" authorityYear="1984" box="[309,681,223,249]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="species" species="guadalupensis">
<emphasis id="B929B23CFFE9664AFEF26A9EB95156FE" box="[309,538,223,249]" italics="true" pageId="9" pageNumber="210">Paha guadalupensis</emphasis>
<bibRefCitation id="EFCC13DFFFE9664AFDE76A9EB9E256FE" author="Dajoz, R." box="[544,681,223,249]" pageId="9" pageNumber="210" pagination="145 - 157" refId="ref25423" refString="Dajoz, R. (1984) Note sur quatre genres de Coleopteres Colydiidae: Nematidium Erichson, Monoedus Horn, Lobogestoria Reitter et Paha n. gen. Bulletin Mensuel de la Societe Linneenne de Lyon, 53 (4), 145 - 157. https: // doi. org / 10.3406 / linly. 1984.10633" type="journal article" year="1984">Dajoz, 1984</bibRefCitation>
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<paragraph id="8BE26E2EFFE9664AFF506B66B9F15466" blockId="9.[151,1437,295,610]" pageId="9" pageNumber="210">
<emphasis id="B929B23CFFE9664AFF506B66BAFA5746" bold="true" box="[151,433,295,321]" pageId="9" pageNumber="210">Taxonomic assignment.</emphasis>
The studied inclusion is placed in the tribe
<taxonomicName id="4C5D15ADFFE9664AFC686B66BF695746" authorityName="Erichson" authorityYear="1845" box="[943,1058,295,321]" class="Insecta" family="Zopheridae" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="tribe" tribe="Synchitini">Synchitini</taxonomicName>
within the subfamily
<taxonomicName id="4C5D15ADFFE9664AFAE66B66BED75746" authorityName="Erichson" authorityYear="1842" box="[1313,1436,295,321]" class="Insecta" family="Colydiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="family">Colydiinae</taxonomicName>
based on the combination of the following characters: (1) dorsally concealed antennal insertions; (2) antennae clubbed, lacking scale-like setae; (3) procoxal cavities open posteriorly; (4) all tarsi tetramerous, not dilated; (5) protibiae without apical spurs; and (6) metacoxae narrowly separated by triangular process of abdominal ventrite 1. The new extinct species belongs to the genus
<taxonomicName id="4C5D15ADFFE9664AFD166BF6B84757D6" authorityName="Dajoz" authorityYear="1984" box="[721,780,439,465]" class="Insecta" family="Zopheridae" genus="Paha" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B929B23CFFE9664AFD166BF6B84757D6" box="[721,780,439,465]" italics="true" pageId="9" pageNumber="210">Paha</emphasis>
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based on its: 10-segmented antennae with 1-segmented, rounded club; antennal groove ventrad eye absent; antennomere 3 not twice as long as antennomere 4; scutellary striole absent; dorsum lacking obvious pubescense (setation short and fine, visible under high magnification only); pronotal disc with elevated central area limited by two parallel longitudinal carinae; odd elytral intervals carinate; and elytral striae with rows of elongate granules.
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