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<document id="431CBBBAAEA56F312349CB7243F41A1C" ID-DOI="10.5281/zenodo.204823" ID-GBIF-Dataset="848522c8-578f-4575-abfb-7c465d110cd7" ID-ISSN="1175-5326" ID-Zenodo-Dep="204823" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460240152533" checkinUser="plazi" docAuthor="Błażewicz-Paszkowycz, Magdalena, Bamber, Roger N &amp; Cunha, Marina R" docDate="2011" docId="023B130FFF91F735B1C031A2FC9BFE1A" docLanguage="en" docName="zt02769p053.pdf" docOrigin="Zootaxa 2769" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pseudotanais (Pseudotanais) tympanobaculum Błażewicz-Paszkowycz, Bamber &amp; Cunha, 2011, sp. nov." docType="treatment" docVersion="10" lastPageNumber="48" masterDocId="FE026B77FFBBF71AB1573264FF90FF9D" masterDocTitle="New tanaidomorph Tanaidacea (Crustacea: Peracarida) from submarine mud-volcanoes in the Gulf of Cadiz (North-east Atlantic)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="43" updateTime="1698240523580" updateUser="plazi">
<mods:mods id="B77EE82AFF52F45CDB78C440A9C9520E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="40CAA12EB9A507F80BF19CAF5BE6E9EA">New tanaidomorph Tanaidacea (Crustacea: Peracarida) from submarine mud-volcanoes in the Gulf of Cadiz (North-east Atlantic)</mods:title>
</mods:titleInfo>
<mods:name id="201EE443F07555A451E745E97CF12352" type="personal">
<mods:role id="55EF3619DC13E00D41F286750B1C06D9">
<mods:roleTerm id="3C1ADBABB02045302A86D468C326F8D6">Author</mods:roleTerm>
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<mods:namePart id="35065A37F7CFD60C52FAC921169A1F35">Błażewicz-Paszkowycz, Magdalena</mods:namePart>
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<mods:name id="181B8CA74FA43440FA6D53EC93F80EC6" type="personal">
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<mods:namePart id="AE2E87410749F7078EFFA90A544D3A5E">Bamber, Roger N</mods:namePart>
</mods:name>
<mods:name id="BBD87493F79E300597A3830B2EFD0CD3" type="personal">
<mods:role id="06A2EE936449FFB1FC8D913D1535B3CF">
<mods:roleTerm id="02BA236D02315CC8A09CCCF51E09947A">Author</mods:roleTerm>
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<mods:namePart id="2465755550B92FEA389769E5AAB14AC2">Cunha, Marina R</mods:namePart>
</mods:name>
<mods:typeOfResource id="67951CA6CDAA8A99E4D6DFEE34BED1D5">text</mods:typeOfResource>
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<mods:title id="AC46228080C8B2243E18A432C562EF26">Zootaxa</mods:title>
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<mods:part id="0DA566038E12C91F7817BD03528C0925">
<mods:date id="F6F1835D8D7A89AD995466211CB85617">2011</mods:date>
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<mods:number id="653A162A4F08093CBD8826E5716E3B78">2769</mods:number>
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<mods:classification id="1CD4CA6A01E2816C43A063DC35DFA793">journal article</mods:classification>
<mods:identifier id="930BD82C2C34706EAF1F8DA1CA30202E" type="DOI">10.5281/zenodo.204823</mods:identifier>
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<mods:identifier id="A72B26502F8187460FBFFD50F94886FF" type="Zenodo-Dep">204823</mods:identifier>
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<treatment id="023B130FFF91F735B1C031A2FC9BFE1A" ID-DOI="http://doi.org/10.5281/zenodo.6189446" ID-GBIF-Taxon="119389580" ID-Zenodo-Dep="6189446" LSID="urn:lsid:plazi:treatment:023B130FFF91F735B1C031A2FC9BFE1A" httpUri="http://treatment.plazi.org/id/023B130FFF91F735B1C031A2FC9BFE1A" lastPageId="47" lastPageNumber="48" pageId="42" pageNumber="43">
<subSubSection id="C288F192FF91F730B1C031A2FE84FB9C" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8A2DA219FF91F730B1C031A2FCA1FC7D" blockId="42.[151,817,966,1025]" box="[151,817,966,992]" pageId="42" pageNumber="43">
<heading id="D1651575FF91F730B1C031A2FCA1FC7D" bold="true" box="[151,817,966,992]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<emphasis id="B8E67E0BFF91F730B1C031A2FCA1FC7D" bold="true" box="[151,817,966,992]" pageId="42" pageNumber="43">
<taxonomicName id="4D92D99AFF91F730B1C031A2FD5CFC7D" ID-CoL="4PCRQ" box="[151,716,966,992]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="tympanobaculum" status="sp. nov." subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF91F730B1C031A2FD5CFC7D" bold="true" box="[151,716,966,992]" italics="true" pageId="42" pageNumber="43">Pseudotanais (Pseudotanais) tympanobaculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D5C370FF91F730B38331A2FCA1FC7D" box="[724,817,966,992]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8A2DA219FF91F730B1C0318DFE84FB9C" blockId="42.[151,817,966,1025]" box="[151,276,1001,1025]" pageId="42" pageNumber="43">
<figureCitation id="12A9BE9CFF91F730B1C0318DFE84FB9C" box="[151,276,1001,1025]" captionStart-0="FIGURE 26" captionStart-1="FIGURE 27" captionStart-2="FIGURE 28" captionStartId-0="43.[151,250,1861,1884]" captionStartId-1="44.[151,250,1953,1976]" captionStartId-2="45.[151,250,1979,2002]" captionTargetBox-0="[224,1377,310,1815]" captionTargetBox-1="[261,1331,181,1919]" captionTargetBox-2="[291,1316,194,1945]" captionTargetId-0="figure@43.[195,1392,301,1833]" captionTargetId-1="figure@44.[239,1353,169,1949]" captionTargetId-2="figure@45.[261,1326,151,1956]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionTargetPageId-2="45" captionText-0="FIGURE 26. Pseudotanais (P.) tympanobaculum sp. nov., A, Ƥ holotype, dorsal; B, Ƥ holotype, lateral. Scale line = 1 mm." captionText-1="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." captionText-2="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri-0="https://zenodo.org/record/204848/files/figure.png" httpUri-1="https://zenodo.org/record/204849/files/figure.png" httpUri-2="https://zenodo.org/record/204850/files/figure.png" pageId="42" pageNumber="43">Figs 2628</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C288F192FF91F730B1C03648FE55F9D9" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8A2DA219FF91F730B1C03648FAAEFB11" blockId="42.[151,1437,1067,2036]" pageId="42" pageNumber="43">
<emphasis id="B8E67E0BFF91F730B1C03648FEC7FBD9" bold="true" box="[151,343,1067,1092]" pageId="42" pageNumber="43">
<typeStatus id="55291CBBFF91F730B1C03648FF42FBD9" box="[151,210,1068,1092]" pageId="42" pageNumber="43">Type</typeStatus>
material.
</emphasis>
1Ƥ,
<typeStatus id="55291CBBFF91F730B0E53648FD8AFBD9" box="[434,538,1068,1092]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
(
<collectionCode id="EC833ADCFF91F730B36D3648FD02FBD9" box="[570,658,1068,1092]" pageId="42" pageNumber="43">BMNH</collectionCode>
.2010.422), 12ƤƤ (2 with empty brood-pouches),
<typeStatus id="55291CBBFF91F730B4663649FA0CFBD8" box="[1329,1436,1069,1093]" pageId="42" pageNumber="43" type="paratype">paratypes</typeStatus>
(
<collectionCode id="EC833ADCFF91F730B1C9362BFF66FBFA" box="[158,246,1103,1127]" pageId="42" pageNumber="43">BMNH</collectionCode>
.2010.423-432), 2 further females, dissected, Station CA 546, Captain Arutyunov mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B1C03610FEB6FB11" box="[151,294,1140,1164]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.661533">35º39.692N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B07A3610FE53FB11" box="[301,451,1140,1164]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-7.3341">07º20.046W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B0993610FDB3FB11" box="[462,547,1140,1164]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="42" pageNumber="43" unit="m" value="1345.0">1345 m</quantity>
depth, mud-breccia and gas-hydrate, TV-grab, 0 6.08.2004, coll.
<collectionCode id="EC833ADCFF91F730B5AF3610FAAAFB11" box="[1272,1338,1140,1164]" httpUri="http://grbio.org/cool/sdrz-mymz" name="TUBITAK Marmara Research Center Culture Collection" pageId="42" pageNumber="43">MRC</collectionCode>
.
</paragraph>
<paragraph id="8A2DA219FF91F730B19236F3FE55F9D9" blockId="42.[151,1437,1067,2036]" pageId="42" pageNumber="43">
<emphasis id="B8E67E0BFF91F730B19236F3FEEBFB2D" bold="true" box="[197,379,1175,1200]" pageId="42" pageNumber="43">Other records.</emphasis>
<specimenCount id="9C946990FF91F730B0DF36F3FD8CFB32" box="[392,540,1175,1199]" pageId="42" pageNumber="43" type="generic">2 specimens</specimenCount>
, Stn CA 393, Captain Arutyunov mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B53236F3FB64FB32" box="[1125,1268,1175,1199]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.662334">35º39.740N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B45536F3FA07FB32" box="[1282,1431,1175,1199]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-7.3323665">07º19.942W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B1C036D8FF7CFB49" box="[151,236,1212,1236]" metricMagnitude="3" metricUnit="m" metricValue="1.327" pageId="42" pageNumber="43" unit="m" value="1327.0">1327 m</quantity>
depth, mud-breccia, TV-grab,
<date id="FE2C84D9FF91F730B30B36D8FD4EFB49" box="[604,734,1212,1236]" pageId="42" pageNumber="43" value="2002-09-07">09.07.2002</date>
;
<specimenCount id="9C946990FF91F730B3B836D8FCE7FB49" box="[751,887,1212,1236]" pageId="42" pageNumber="43" type="generic">1 specimen</specimenCount>
, Stn Ki 528, Kidd mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B45A36D8FA0CFB49" box="[1293,1436,1212,1236]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.421734">35º25.304N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B1C036BBFEBDFB6A" box="[151,301,1247,1271]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-6.732867">06º43.972W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B06D36BBFE11FB65" box="[314,385,1247,1272]" metricMagnitude="2" metricUnit="m" metricValue="4.89" pageId="42" pageNumber="43" unit="m" value="489.0">489 m</quantity>
depth, mud-breccia and H2S, TV-grab,
<date id="FE2C84D9FF91F730B21B36BBFC5EFB6A" box="[844,974,1247,1271]" pageId="42" pageNumber="43" value="2004-03-08">03.08.2004</date>
;
<specimenCount id="9C946990FF91F730B28C36BBFBCFFB6A" box="[987,1119,1247,1271]" pageId="42" pageNumber="43" type="generic">1 specimen</specimenCount>
, Stn CA 543, Captain Arutyunov mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B0D83763FD8EFA82" box="[399,542,1287,1311]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.66147">35º39.688N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B3713763FD2CFA82" box="[550,700,1287,1311]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-7.333017">07º19.981W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B39F3763FC8DFA82" box="[712,797,1287,1311]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="42" pageNumber="43" unit="m" value="1345.0">1345 m</quantity>
depth, mud-breccia and H2S, Gravity corer,
<date id="FE2C84D9FF91F730B4413763FA08FA82" box="[1302,1432,1287,1311]" pageId="42" pageNumber="43" value="2004-06-08">06.08.2004</date>
;
<specimenCount id="9C946990FF91F730B1C03749FEA6FAD8" box="[151,310,1325,1349]" pageId="42" pageNumber="43" type="generic">26 specimens</specimenCount>
, Stn CA 547, Captain Arutyunov mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B2263749FC6FFAD8" box="[881,1023,1325,1349]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.661682">35º39.701N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B55C3749FB31FAD8" box="[1035,1185,1325,1349]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-7.33395">07º20.037W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B5E73749FA95FAD8" box="[1200,1285,1325,1349]" metricMagnitude="3" metricUnit="m" metricValue="1.344" pageId="42" pageNumber="43" unit="m" value="1344.0">1344 m</quantity>
depth, mudbreccia and gas-hydrate, TV-grab,
<date id="FE2C84D9FF91F730B3613735FD28FAF4" box="[566,696,1361,1385]" pageId="42" pageNumber="43" value="2004-06-08">06.08.2004</date>
;
<specimenCount id="9C946990FF91F730B39D3735FCCDFAF4" box="[714,861,1361,1385]" pageId="42" pageNumber="43" type="generic">2 specimens</specimenCount>
, Stn CA 548, Captain Arutyunov mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B1C03711FEB6FA10" box="[151,294,1397,1421]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.6618">35º39.708N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B0783711FE55FA10" box="[303,453,1397,1421]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-7.3339">07º20.034W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B0843711FDB8FA10" box="[467,552,1397,1421]" metricMagnitude="3" metricUnit="m" metricValue="1.345" pageId="42" pageNumber="43" unit="m" value="1345.0">1345 m</quantity>
depth, mud-breccia and H2S, Kasten-corer,
<date id="FE2C84D9FF91F730B5743711FB35FA10" box="[1059,1189,1397,1421]" pageId="42" pageNumber="43" value="2004-06-08">06.08.2004</date>
;
<specimenCount id="9C946990FF91F730B5E53711FAD4FA10" box="[1202,1348,1397,1421]" pageId="42" pageNumber="43" type="generic">21 specimen</specimenCount>
, Stn Ki 559, Kidd mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B0E237F8FDD4FA29" box="[437,580,1436,1460]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.41295">35º24.777N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B31B37F8FD72FA29" box="[588,738,1436,1460]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-6.7297">06º43.782W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B3B837F8FCA6FA29" box="[751,822,1436,1460]" metricMagnitude="2" metricUnit="m" metricValue="5.52" pageId="42" pageNumber="43" unit="m" value="552.0">552 m</quantity>
depth, mud, Boxcorer,
<date id="FE2C84D9FF91F730B51A37F8FB5FFA29" box="[1101,1231,1436,1460]" pageId="42" pageNumber="43" value="2004-08-08">08.08.2004</date>
;
<specimenCount id="9C946990FF91F730B58B37F8FAFAFA29" box="[1244,1386,1436,1460]" pageId="42" pageNumber="43" type="generic">2 specimens</specimenCount>
, Stn Ki 561, Kidd mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B08D37A4FDFAFA45" box="[474,618,1472,1496]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.4267">35º25.602N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B32437A4FC99FA45" box="[627,777,1472,1496]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-6.7349834">06º44.099W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B24137A4FCCDFA45" box="[790,861,1472,1496]" metricMagnitude="2" metricUnit="m" metricValue="5.26" pageId="42" pageNumber="43" unit="m" value="526.0">526 m</quantity>
depth, mud, Boxcorer,
<date id="FE2C84D9FF91F730B53B37A4FB7EFA45" box="[1132,1262,1472,1496]" pageId="42" pageNumber="43" value="2004-08-08">08.08.2004</date>
;
<specimenCount id="9C946990FF91F730B5AC37A4FA08FA45" box="[1275,1432,1472,1496]" pageId="42" pageNumber="43" type="generic">21 specimens</specimenCount>
, Stn Mer 576, Mercator mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B36A3780FD5DFA61" box="[573,717,1508,1532]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.294285">35º17.657N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B3843780FCF9FA61" box="[723,873,1508,1532]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-6.65215">06º39.129W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B2243780FC2AFA61" box="[883,954,1508,1532]" metricMagnitude="2" metricUnit="m" metricValue="4.279999999999999" pageId="42" pageNumber="43" unit="m" value="428.0">428 m</quantity>
depth, mud, Boxcorer,
<date id="FE2C84D9FF91F730B5973780FAD2FA61" box="[1216,1346,1508,1532]" pageId="42" pageNumber="43" value="2005-07-26">26.07.2005</date>
;
<specimenCount id="9C946990FF91F730B41B3780FF4DF9BD" pageId="42" pageNumber="43" type="generic">2 specimens</specimenCount>
, Stn Bom 597, Bonjardim mud-volcano,
<geoCoordinate id="EFA6C4DEFF91F730B3E2346CFCD4F9BD" box="[693,836,1544,1568]" direction="north" orientation="latitude" pageId="42" pageNumber="43" precision="1" value="35.459385">35º27.563N</geoCoordinate>
<geoCoordinate id="EFA6C4DEFF91F730B21A346CFC73F9BD" box="[845,995,1544,1568]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="1" value="-9.0005">09º00.030W</geoCoordinate>
,
<quantity id="4D6A0FFCFF91F730B2A7346CFBD5F9BD" box="[1008,1093,1544,1568]" metricMagnitude="3" metricUnit="m" metricValue="3.061" pageId="42" pageNumber="43" unit="m" value="3061.0">3061 m</quantity>
depth, mud-breccia, TV-grab,
<date id="FE2C84D9FF91F730B1C03448FE89F9D9" box="[151,281,1580,1604]" pageId="42" pageNumber="43" value="2005-02-08">02.08.2005</date>
; all coll.
<collectionCode id="EC833ADCFF91F730B0293448FE50F9D9" box="[382,448,1580,1604]" httpUri="http://grbio.org/cool/sdrz-mymz" name="TUBITAK Marmara Research Center Culture Collection" pageId="42" pageNumber="43">MRC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C288F192FF91F734B1923434FEE5FA91" lastPageId="46" lastPageNumber="47" pageId="42" pageNumber="43" type="description">
<paragraph id="8A2DA219FF91F730B1923434FDE2F8F9" blockId="42.[151,1437,1067,2036]" pageId="42" pageNumber="43">
<emphasis id="B8E67E0BFF91F730B1923434FEC9F9F4" bold="true" box="[197,345,1616,1641]" pageId="42" pageNumber="43">Description:</emphasis>
body (
<figureCitation id="12A9BE9CFF91F730B0FE3434FE6CF9F5" box="[425,508,1616,1640]" captionStart="FIGURE 26" captionStartId="43.[151,250,1861,1884]" captionTargetBox="[224,1377,310,1815]" captionTargetId="figure@43.[195,1392,301,1833]" captionTargetPageId="43" captionText="FIGURE 26. Pseudotanais (P.) tympanobaculum sp. nov., A, Ƥ holotype, dorsal; B, Ƥ holotype, lateral. Scale line = 1 mm." httpUri="https://zenodo.org/record/204848/files/figure.png" pageId="42" pageNumber="43">Fig. 26</figureCitation>
A, B) compact,
<typeStatus id="55291CBBFF91F730B3E53434FC84F9F5" box="[690,788,1616,1640]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
<quantity id="4D6A0FFCFF91F730B24B3434FCE2F9F5" box="[796,882,1616,1640]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="42" pageNumber="43" unit="mm" value="2.4">2.4 mm</quantity>
long, four times as long as wide. Cephalothorax trapezoidal, two-thirds as long as wide, as long as pereonites 2 and 3 together, single setae in anterolateral corners, rostrum eyelobes and eyes absent. Pereonites naked; pereonite 1 shortest by far, pereonite 2 2.5 times as long as pereonite 1, pereonite 3 three times as long as pereonite 1, pereonites 4 and 6 subequal, four times as long as pereonite 1, pereonite 5 longest, five times as long as pereonite 1 (all pereonites respectively 7.7, 3.4, 3.2, 2.1, 1.7 and 1.9 times as wide as long). Pleonites becoming narrower posteriorly, each pleonite about 6.5 times as wide as long and bearing pleopods. Pleotelson semicircular but extended into blunt posterior apex, 2.5 times as long as each pleonite, nearly twice as wide as long, naked.
</paragraph>
<paragraph id="8A2DA219FF91F730B1923514FBBDF869" blockId="42.[151,1437,1067,2036]" pageId="42" pageNumber="43">
Antennule (
<figureCitation id="12A9BE9CFF91F730B01E3514FE0AF815" box="[329,410,1904,1928]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="42" pageNumber="43">Fig. 27</figureCitation>
A) proximal article 4.5 times as long as wide, much longer than distal two articles together, with simple mesial seta, one longer and one shorter distal simple setae and two penicillate distal setae; second article 2.5 times as long as wide, 0.4 times as long as first article; third article nearly half as long as second article, distally with three simple, two bifurcate and one penicillate setae and one aesthetasc.
</paragraph>
<paragraph id="8A2DA219FF90F731B19232FCFADBFF6A" blockId="43.[151,1436,151,247]" pageId="43" pageNumber="44">
Antenna (
<figureCitation id="12A9BE9CFF90F731B06332F3FE16FF32" box="[308,390,151,175]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="43" pageNumber="44">Fig. 27</figureCitation>
B) proximal article compact, naked; second article almost as wide as long, with small dorsodistal spine; third just longer than second, naked; fourth article longest, six times as long as wide, with four distal setae; fifth article 0.4 times as long as fourth, with one distal seta; sixth article minute with five distal setae.
</paragraph>
<caption id="DEEDF291FF90F731B1C03521FA11F8C1" box="[151,1409,1861,1884]" httpUri="https://zenodo.org/record/204848/files/figure.png" pageId="43" pageNumber="44" targetBox="[224,1377,310,1815]" targetPageId="43">
<paragraph id="8A2DA219FF90F731B1C03521FA11F8C1" blockId="43.[151,1409,1861,1884]" box="[151,1409,1861,1884]" pageId="43" pageNumber="44">
<emphasis id="B8E67E0BFF90F731B1C03521FE8FF8C6" bold="true" box="[151,287,1861,1884]" pageId="43" pageNumber="44">FIGURE 26.</emphasis>
<taxonomicName id="4D92D99AFF90F731B0723522FD1CF8C1" box="[293,652,1862,1884]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="tympanobaculum" status="sp. nov." subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF90F731B0723522FD1CF8C1" box="[293,652,1862,1884]" italics="true" pageId="43" pageNumber="44">Pseudotanais (P.) tympanobaculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D5C370FF90F731B3C43522FD73F8C6" box="[659,739,1862,1883]" pageId="43" pageNumber="44" rank="species">
<emphasis id="B8E67E0BFF90F731B3C43522FD73F8C6" bold="true" box="[659,739,1862,1883]" pageId="43" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
, A, Ƥ holotype, dorsal; B, Ƥ holotype, lateral. Scale line = 1 mm.
</paragraph>
</caption>
<caption id="DEEDF291FF97F736B1C035C5FF68F869" httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="44" pageNumber="45" targetBox="[261,1331,181,1919]" targetPageId="44">
<paragraph id="8A2DA219FF97F736B1C035C5FF68F869" blockId="44.[151,1436,1953,2036]" pageId="44" pageNumber="45">
<emphasis id="B8E67E0BFF97F736B1C035C5FEB0F82B" bold="true" box="[151,288,1953,1976]" pageId="44" pageNumber="45">FIGURE 27.</emphasis>
<taxonomicName id="4D92D99AFF97F736B07135C5FD1EF82A" box="[294,654,1953,1975]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="tympanobaculum" status="sp. nov." subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF97F736B07135C5FD1EF82A" box="[294,654,1953,1975]" italics="true" pageId="44" pageNumber="45">Pseudotanais (P.) tympanobaculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D5C370FF97F736B3C235C6FD76F82A" box="[661,742,1954,1975]" pageId="44" pageNumber="45" rank="species">
<emphasis id="B8E67E0BFF97F736B3C235C6FD76F82A" bold="true" box="[661,742,1954,1975]" pageId="44" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
, Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm.
</paragraph>
</caption>
<caption id="DEEDF291FF96F737B1C035DFFD49F86D" httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="45" pageNumber="46" targetBox="[291,1316,194,1945]" targetPageId="45">
<paragraph id="8A2DA219FF96F737B1C035DFFD49F86D" blockId="45.[151,1436,1979,2032]" pageId="45" pageNumber="46">
<emphasis id="B8E67E0BFF96F737B1C035DFFEB2F84C" bold="true" box="[151,290,1979,2002]" pageId="45" pageNumber="46">FIGURE 28.</emphasis>
<taxonomicName id="4D92D99AFF96F737B07C35D8FD07F84F" box="[299,663,1980,2002]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="tympanobaculum" status="sp. nov." subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF96F737B07C35D8FD07F84F" box="[299,663,1980,2002]" italics="true" pageId="45" pageNumber="46">Pseudotanais (P.) tympanobaculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D5C370FF96F737B3F735D8FD63F84C" box="[672,755,1980,2001]" pageId="45" pageNumber="46" rank="species">
<emphasis id="B8E67E0BFF96F737B3F735D8FD63F84C" bold="true" box="[672,755,1980,2001]" pageId="45" pageNumber="46">sp. nov.</emphasis>
</taxonomicNameLabel>
, A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8A2DA219FF95F734B19232F3FA1AFE1A" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Labrum (
<figureCitation id="12A9BE9CFF95F734B07B32F3FEEDFF32" box="[300,381,151,175]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
C) rounded, naked. Left mandible (
<figureCitation id="12A9BE9CFF95F734B25432F3FCC4FF32" box="[771,852,151,175]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
D) with wide, crenulate lacinia mobilis, right mandible (
<figureCitation id="12A9BE9CFF95F734B19E32D8FE89FF49" box="[201,281,188,212]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
E) with crenulated upper margin; pars molaris of both mandibles acuminate with two or three fine subdistal denticles. Labium (
<figureCitation id="12A9BE9CFF95F734B0EE32BBFD9CFF6A" box="[441,524,223,247]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
H) simple, bilobed, naked. Maxillule (
<figureCitation id="12A9BE9CFF95F734B29432BBFB85FF6A" box="[963,1045,223,247]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
F) with nine distal spines and two fine setae, palp (
<figureCitation id="12A9BE9CFF95F734B0023360FE36FE86" box="[341,422,258,284]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
Fʹ) with two distal setae. Maxilla (
<figureCitation id="12A9BE9CFF95F734B2633360FC17FE81" box="[820,903,260,284]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
G) simple, ovoid, naked. Maxilliped (
<figureCitation id="12A9BE9CFF95F734B4603360FA1AFE81" box="[1335,1418,260,284]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">
Fig.
<date id="FE2C84D9FF95F734B43A3360FA1AFE81" box="[1389,1418,260,284]" pageId="46" pageNumber="47" value="1927-01">27</date>
</figureCitation>
I) first palp article naked, second article with one outer and two inner finely setulose setae, third article with two inner setae, fourth article with three inner, two distal and one outer subdistal setae; basis fused, naked; endites almost completely fused, distally with single seta and slight oval tubercle. Epignath (
<figureCitation id="12A9BE9CFF95F734B2AB330BFBDDFE1A" box="[1020,1101,367,391]" captionStart="FIGURE 27" captionStartId="44.[151,250,1953,1976]" captionTargetBox="[261,1331,181,1919]" captionTargetId="figure@44.[239,1353,169,1949]" captionTargetPageId="44" captionText="FIGURE 27. Pseudotanais (P.) tympanobaculum sp. nov., Ƥ paratype. A, antennule; B, antenna; C, labrum; D, left mandible; E, right mandible; F, maxillule; Fˏ, maxillule palp; G, maxilla; H, I, maxilliped; J, epignath. Scale lines: A-B, = 0.1 mm; C-I = 0.01 mm." httpUri="https://zenodo.org/record/204849/files/figure.png" pageId="46" pageNumber="47">Fig. 27</figureCitation>
J) elongate, distally pointed.
</paragraph>
<paragraph id="8A2DA219FF95F734B19233F0FDCAFDA1" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Cheliped (
<figureCitation id="12A9BE9CFF95F734B06D33F0FE1CFE31" box="[314,396,404,428]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
A) with rounded, naked basis 1.4 times as long as wide; merus subtriangular, naked; carpus 1.6 times as long as wide, with single mid-dorsal and midventral setae; propodus 1.7 times as long as wide, with comb-row of one longer and eight shorter setae near base of dactylus; fixed finger 0.75 times as long as propodus, with one ventral seta and three setae adjacent to cutting edge; dactylus with small proximal seta, cutting edge with central fine spinule. Chela not forcipate.
</paragraph>
<paragraph id="8A2DA219FF95F734B1923023FBA8FC1D" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Pereopod 1 (
<figureCitation id="12A9BE9CFF95F734B0043023FE34FDC2" box="[339,420,583,607]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
B) coxa naked; basis slender, 5.5 times as long as wide with one mesial seta; ischium compact, with small ventral seta; merus 0.24 times as long as basis and with small ventrodistal seta; carpus as long as merus, naked; propodus nearly as long as carpus and merus together, with one dorsodistal seta; dactylus with distal seta and distinct slender unguis twice as long as dactylus, both together 1.45 times as long as propodus. Pereopod 2 (
<figureCitation id="12A9BE9CFF95F734B1C830B3FF63FD72" box="[159,243,727,751]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
C), coxa similar to but stouter than pereopod 1, basis 5.2 times as long as wide, with one ventroproximal seta; merus 0.35 times as long as basis and with slender, blunt ventrodistal spine; carpus as long as merus, with dorsodistal seta and blade-like ventrodistal spine; propodus 1.5 times as long as carpus, with ventrodistal spine; dactylus with slightly longer unguis together two-thirds as long as propodus. Pereopod 3 (
<figureCitation id="12A9BE9CFF95F734B5183120FB31FCC1" box="[1103,1185,836,860]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
D) similar to but more compact than pereopod 2, merus ventrodistally with two spines and one small seta.
</paragraph>
<paragraph id="8A2DA219FF95F734B19231E8FCA7FBC5" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Pereopod 4 (
<figureCitation id="12A9BE9CFF95F734B00531E8FE32FC39" box="[338,418,908,932]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
E) basis stout, 2.7 times as long as wide; ischium with two ventral setae; merus 0.4 times as long as carpus and with one ventrodistal spine; carpus with one simple and one blade-like spines, and two small ventrodistal setae; propodus 0.8 times as long as carpus, with two ventrodistal spines and dorsodistal spine longer than dactylus plus unguis; dactylus and unguis short, 0.6 times as long as propodus. Pereopod 5 (
<figureCitation id="12A9BE9CFF95F734B583319CFAB4FB8D" box="[1236,1316,1016,1040]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
F) as pereopod 4, but additional ventrodistal spine on carpus, single ventrodistal spine but additional dorsodistal seta on propodus. Pereopod 6 (
<figureCitation id="12A9BE9CFF95F734B0CC3624FE7CFBC5" box="[411,492,1088,1112]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
G) as pereopod 5, but stouter.
</paragraph>
<paragraph id="8A2DA219FF95F734B1923600FD5DFB3D" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Pleopods (
<figureCitation id="12A9BE9CFF95F734B06C3600FE1CFBE1" box="[315,396,1124,1148]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">Fig. 28</figureCitation>
H) all alike, with naked basis, slender endopod shorter than exopod and with four distal plumose setae, exopod with five distal plumose setae.
</paragraph>
<paragraph id="8A2DA219FF95F734B19236C8FEC6FB75" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Uropods (
<figureCitation id="12A9BE9CFF95F734B06136C8FE18FB59" box="[310,392,1196,1220]" captionStart="FIGURE 28" captionStartId="45.[151,250,1979,2002]" captionTargetBox="[291,1316,194,1945]" captionTargetId="figure@45.[261,1326,151,1956]" captionTargetPageId="45" captionText="FIGURE 28. Pseudotanais (P.) tympanobaculum sp. nov., A, cheliped, with detail of chela (A); B to G, pereopods 1 to 6, respectively; H, pleopod; I, uropod. Scale line = 0.1 mm." httpUri="https://zenodo.org/record/204850/files/figure.png" pageId="46" pageNumber="47">
Fig.
<date id="FE2C84D9FF95F734B03C36C8FE18FB59" box="[363,392,1196,1220]" pageId="46" pageNumber="47" value="1928-01">28</date>
</figureCitation>
I) biramous, both exopod and endopod of two segments, setose as figured; exopod almost as long as endopod.
</paragraph>
<paragraph id="8A2DA219FF95F734B1923690FEE5FA91" blockId="46.[151,1437,151,1976]" box="[197,373,1268,1292]" pageId="46" pageNumber="47">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C288F192FF95F734B192377CFEECFAE5" pageId="46" pageNumber="47" type="etymology">
<paragraph id="8A2DA219FF95F734B192377CFEECFAE5" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
<emphasis id="B8E67E0BFF95F734B192377CFEDCFAAD" bold="true" box="[197,332,1304,1328]" pageId="46" pageNumber="47">Etymology.</emphasis>
from the Latin
<emphasis id="B8E67E0BFF95F734B0AC377DFDE0FAAD" box="[507,624,1305,1328]" italics="true" pageId="46" pageNumber="47">tympanum</emphasis>
a drum, and
<emphasis id="B8E67E0BFF95F734B240377CFCE7FAAD" box="[791,887,1304,1328]" italics="true" pageId="46" pageNumber="47">baculum</emphasis>
a stick, thus drum-stick (noun in apposition), a term for the stout back legs of edible phasianid birds, the present species being characterized by stout bases on the posterior pereopods.
</paragraph>
</subSubSection>
<subSubSection id="C288F192FF95F735B19237E0FC9BFE1A" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8A2DA219FF95F734B19237E0FB25F97D" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
<emphasis id="B8E67E0BFF95F734B19237E0FEA9FA01" bold="true" box="[197,313,1412,1436]" pageId="46" pageNumber="47">Remarks.</emphasis>
The
<taxonomicName id="4D92D99AFF95F734B02237E0FDB3FA01" box="[373,547,1412,1436]" class="Malacostraca" family="Pseudotanaidae" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="family">Pseudotanaidae</taxonomicName>
of the northeast Atlantic was comprehensively analyzed by
<bibRefCitation id="EE03DFE8FF95F734B58137E0FF73FA5D" author="Bird" pageId="46" pageNumber="51" refString="Bird, G. J. &amp; Holdich, D. M. (1989) Tanaidacea (Crustacea) of the north-east Atlantic: the subfamily Pseudotanainae (Pseudotanaidae) and the family Nototanaidae. Zoological Journal of the Linnean Society, 97, 233 - 298." type="journal article" year="1989">Bird and Holdich (1989)</bibRefCitation>
, who included a key to the species then known. With its blade-like carpal spines on pereopods 2 to 6 (characteristic of the nominate subgenus
<taxonomicName id="4D92D99AFF95F734B37F37A8FD2FFA79" box="[552,703,1484,1508]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B8E67E0BFF95F734B37F37A8FD2FFA79" box="[552,703,1484,1508]" italics="true" pageId="46" pageNumber="47">Pseudotanais</emphasis>
</taxonomicName>
), distally-bifurcate setae on the antennule, one ventral seta on the non-forcipate chela, short inferodistal spines on the propodus of pereopods 2 and 3, and subequal uropod rami, of the species in that key
<taxonomicName id="4D92D99AFF95F734B0F73471FDEDF9B1" box="[416,637,1556,1580]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="tympanobaculum" status="sp. nov.">
<emphasis id="B8E67E0BFF95F734B0F73471FDEDF9B1" box="[416,637,1556,1580]" italics="true" pageId="46" pageNumber="47">P. tympanobaculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3D5C370FF95F734B3D23470FD71F9B1" box="[645,737,1556,1580]" pageId="46" pageNumber="47" rank="species">
<emphasis id="B8E67E0BFF95F734B3D23470FD71F9B1" bold="true" box="[645,737,1556,1580]" pageId="46" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
is closest to
<taxonomicName id="4D92D99AFF95F734B22C3471FABCF9B1" authority="Bird &amp; Holdich, 1989" authorityName="Bird &amp; Holdich" authorityYear="1989" box="[891,1324,1556,1580]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spicatus" subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF95F734B22C3471FBB4F9B1" box="[891,1060,1556,1580]" italics="true" pageId="46" pageNumber="47">P. (P.) spicatus</emphasis>
<bibRefCitation id="EE03DFE8FF95F734B57A3470FABCF9B1" author="Bird" box="[1069,1324,1556,1580]" pageId="46" pageNumber="51" refString="Bird, G. J. &amp; Holdich, D. M. (1989) Tanaidacea (Crustacea) of the north-east Atlantic: the subfamily Pseudotanainae (Pseudotanaidae) and the family Nototanaidae. Zoological Journal of the Linnean Society, 97, 233 - 298." type="journal article" year="1989">Bird &amp; Holdich, 1989</bibRefCitation>
</taxonomicName>
, recorded from the Rockall Trough, Porcupine Seabight and Abyssal Plain and Bay of Biscay at depths between 2227 and
<quantity id="4D6A0FFCFF95F734B1C03438FF7FF9E9" box="[151,239,1628,1652]" metricMagnitude="3" metricUnit="m" metricValue="4.829" pageId="46" pageNumber="47" unit="m" value="4829.0">4829 m</quantity>
. However, unlike the present species,
<taxonomicName id="4D92D99AFF95F734B3CF3439FCACF9E9" box="[664,828,1628,1652]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="spicatus" subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF95F734B3CF3439FCACF9E9" box="[664,828,1628,1652]" italics="true" pageId="46" pageNumber="47">P. (P.) spicatus</emphasis>
</taxonomicName>
has a small dorsodistal spine on the third article of the antennule, more inner setae on the second and third maxilliped palp articles, more complex and dendritic denticulation on the mandibular pars molaris, a one-segmented uropod endopod, many more setae on the pleopod rami, and, on the posterior pereopods, much more slender bases and a stouter, fused dactylus and unguis.
</paragraph>
<paragraph id="8A2DA219FF95F734B1903488FC58F825" blockId="46.[151,1437,151,1976]" pageId="46" pageNumber="47">
Of the two similar species described since 1989 (non-forcipate members of the nominate subgenus, with subequal uropod rami),
<emphasis id="B8E67E0BFF95F734B0D43574FDD6F8B5" box="[387,582,1808,1832]" italics="true" pageId="46" pageNumber="47">
<taxonomicName id="4D92D99AFF95F734B0D43574FD8AF8B5" box="[387,538,1808,1832]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Pseudotanais</taxonomicName>
(P)
</emphasis>
<taxonomicName id="4D92D99AFF95F734B3183574FCD4F8B5" authority="Bird, 1999" authorityName="Bird" authorityYear="1999" box="[591,836,1808,1832]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="baresnauti">
<emphasis id="B8E67E0BFF95F734B3183574FD53F8B5" box="[591,707,1808,1832]" italics="true" pageId="46" pageNumber="47">baresnauti</emphasis>
<bibRefCitation id="EE03DFE8FF95F734B39B3574FCD4F8B5" author="Bird" box="[716,836,1808,1832]" pageId="46" pageNumber="50" refString="Bird, G. J. (1999) A new species of Pseudotanais (Crustacea, Tanaidacea) from cold seeps in the deep Caribbean, collected by the French submersible Nautile. Zoosystema, 21 (3), 445 - 451." type="journal article" year="1999">Bird, 1999</bibRefCitation>
</taxonomicName>
(q.v.) (from a Caribbean cold seep), which shares a naked merus with
<taxonomicName id="4D92D99AFF95F734B0303551FDD4F8D1" box="[359,580,1844,1868]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="tympanobaculum">
<emphasis id="B8E67E0BFF95F734B0303551FDD4F8D1" box="[359,580,1844,1868]" italics="true" pageId="46" pageNumber="47">P. tympanobaculum</emphasis>
</taxonomicName>
, has subequal anterior pereonites, a simpler pars molaris, a small dorsodistal spine on the third article of the antennule, and more slender posterior pereopod bases;
<taxonomicName id="4D92D99AFF95F734B52D353DFF5FF809" authority="McLelland, 2007" authorityName="McLelland" authorityYear="2007" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="nipponicus">
<emphasis id="B8E67E0BFF95F734B52D353DFA80F8ED" box="[1146,1296,1881,1904]" italics="true" pageId="46" pageNumber="47">P. nipponicus</emphasis>
<bibRefCitation id="EE03DFE8FF95F734B44D353CFF5FF809" author="McLelland" pageId="46" pageNumber="52" refString="McLelland, J. C. (2007) Family Pseudotanaidae Sieg, 1976. In: Larsen K &amp; Shimomura M. (Eds) 2007. Tanaidacea (Crustacea: Peracarida) from Japan III. The deep trenches; the Kurile-Kamchatka Trench and Japan Trench. Zootaxa, 1599, 87 - 99." type="journal article" year="2007">McLelland, 2007</bibRefCitation>
</taxonomicName>
(q.v.) (from the Kurile-Kamchatka Trench) has more, or more complex setation on the maxilliped (palps, bases and endites) and maxillule, and particularly on the pereopod bases.
</paragraph>
<paragraph id="8A2DA219FF94F735B19032F3FC3AFF6A" blockId="47.[151,1436,151,391]" pageId="47" pageNumber="48">
Indeed, other than the
<collectingCountry id="F285E289FF94F735B09E32F3FDA1FF32" box="[457,561,151,175]" name="Antarctica" pageId="47" pageNumber="48">Antarctic</collectingCountry>
species
<taxonomicName id="4D92D99AFF94F735B3C332FCFBEFFF32" authority="Sieg, 1977" authorityName="Sieg" authorityYear="1977" box="[660,1151,151,176]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="nordenskioldi" subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF94F735B3C332FCFC6DFF2D" box="[660,1021,152,176]" italics="true" pageId="47" pageNumber="48">Pseudotanais (P.) nordenskioldi</emphasis>
<bibRefCitation id="EE03DFE8FF94F735B55132F3FBEFFF32" author="Sieg" box="[1030,1151,151,175]" pageId="47" pageNumber="52" refString="Sieg, J. (1977) Taxonomische Monographie der familie Pseudotanaidae (Crustacea, Tanaidacea). Mitteilungen aus dem Zoologischen Museum in Berlin, 53, 1 - 109." type="journal article" year="1977">Sieg, 1977</bibRefCitation>
</taxonomicName>
, only
<taxonomicName id="4D92D99AFF94F735B59032FDFE1FFF49" authority="Bird &amp; Holdich, 1989" authorityName="Bird &amp; Holdich" authorityYear="1989" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="denticulatus" subGenus="Pseudotanais">
<emphasis id="B8E67E0BFF94F735B59032FDFA0CFF2D" box="[1223,1436,152,176]" italics="true" pageId="47" pageNumber="48">P. (P.) denticulatus</emphasis>
<bibRefCitation id="EE03DFE8FF94F735B1C032D8FE1FFF49" author="Bird" box="[151,399,188,212]" pageId="47" pageNumber="51" refString="Bird, G. J. &amp; Holdich, D. M. (1989) Tanaidacea (Crustacea) of the north-east Atlantic: the subfamily Pseudotanainae (Pseudotanaidae) and the family Nototanaidae. Zoological Journal of the Linnean Society, 97, 233 - 298." type="journal article" year="1989">Bird &amp; Holdich, 1989</bibRefCitation>
</taxonomicName>
approaches
<taxonomicName id="4D92D99AFF94F735B34B32D9FD69FF49" box="[540,761,188,212]" class="Malacostraca" family="Pseudotanaidae" genus="Pseudotanais" kingdom="Animalia" order="Tanaidacea" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="tympanobaculum">
<emphasis id="B8E67E0BFF94F735B34B32D9FD69FF49" box="[540,761,188,212]" italics="true" pageId="47" pageNumber="48">P. tympanobaculum</emphasis>
</taxonomicName>
in having wide bases to the posterior pereopods, but in that species the bases are still 3.3 to 4 times as long as their greatest width.
</paragraph>
<paragraph id="8A2DA219FF94F735B1903360FC9BFE1A" blockId="47.[151,1436,151,391]" pageId="47" pageNumber="48">
The new species was collected from the Captain Arutyunov mud-volcano (type-locality), in the deep-water field within the Portuguese margin, at depths from
<quantity id="4D6A0FFCFF94F735B3853343FC15FEA2" box="[722,901,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" metricValueMax="1.345" metricValueMin="1.327" pageId="47" pageNumber="48" unit="m" value="1336.0" valueMax="1345.0" valueMin="1327.0">
<date id="FE2C84D9FF94F735B3853343FCF6FEA2" box="[722,870,295,319]" pageId="47" pageNumber="48" value="1327" valueMax="1345">1327 to 1345</date>
m
</quantity>
, from the Kidd and Mercator mud-volcanoes in the El Arraiche field on the Moroccan margin, at
<quantity id="4D6A0FFCFF94F735B3E83328FCC4FEF9" box="[703,852,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.9" metricValueMax="5.52" metricValueMin="4.279999999999999" pageId="47" pageNumber="48" unit="m" value="490.0" valueMax="552.0" valueMin="428.0">428 to 552 m</quantity>
depth, and from the Bomjardim mud-volcano in the western Moroccan field at
<quantity id="4D6A0FFCFF94F735B094330BFD8AFE1A" box="[451,538,367,391]" metricMagnitude="3" metricUnit="m" metricValue="3.061" pageId="47" pageNumber="48" unit="m" value="3061.0">3061 m</quantity>
, on muddy-substrata.
</paragraph>
</subSubSection>
</treatment>
</document>