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<mods:title id="CE8D74A2A1073A00825B01F21490E653">A new coral reef mantis shrimp from the northwestern Indian Ocean, Gonodactylellus celosinus sp. nov. (Crustacea: Stomatopoda: Gonodactylidae)</mods:title>
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<heading id="D0FE4B22D43FFFB8E8FF37166F45F975" bold="true" box="[151,567,1662,1689]" fontSize="11" level="1" pageId="1" pageNumber="454" reason="1">
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<taxonomicName id="4C0987CDD43FFFB8E8FF37166CA0F975" authorityName="Ahyong" authorityYear="2024" box="[151,466,1662,1689]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="454" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43FFFB8E8FF37166CA0F975" bold="true" box="[151,466,1662,1689]" italics="true" pageId="1" pageNumber="454">Gonodactylellus celosinus</emphasis>
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<taxonomicNameLabel id="A24E9D27D43FFFB8E9B237176F45F975" box="[474,567,1663,1689]" pageId="1" pageNumber="454" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="1332E0CBD43FFFB8E8F737CB6D90F951" box="[159,226,1699,1725]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="1" pageNumber="454">Fig. 1</figureCitation>
, 
<figureCitation id="1332E0CBD43FFFB8E89837CB6D8DF951" box="[240,255,1699,1725]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="1" pageNumber="454">2</figureCitation>
)
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<paragraph id="8BB6FC4ED43FFFB8E8FF378F6EB3F8D1" blockId="1.[151,1436,1767,1975]" pageId="1" pageNumber="454">
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<taxonomicName id="4C0987CDD43FFFB8E8FF378F6C08F913" ID-CoL="4Q8P" authority="demani." authorityName="Demani." box="[151,378,1767,1791]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B97D205CD43FFFB8E8FF378F6C04F913" box="[151,374,1767,1791]" italics="true" pageId="1" pageNumber="454">Gonodactylus demani</emphasis>
.
</taxonomicName>
— 
<treatmentCitation id="0AA8DA5FD43FFFB8E9FC37806F36F913" author="Kemp, S." box="[404,580,1767,1791]" page="164" pageId="1" pageNumber="454" year="1913">
<bibRefCitation id="EF9881BFD43FFFB8E9FC37806F36F913" author="Kemp, S." box="[404,580,1767,1791]" pageId="1" pageNumber="454" pagination="1 - 217" refId="ref6528" refString="Kemp, S. (1913) An account of the Crustacea Stomatopoda of the Indo-Pacific region, based on the collection in the Indian Museum. Memoirs of the Indian Museum, 4, 1 - 217, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 12631" type="journal article" year="1913">Kemp 1913: 164</bibRefCitation>
</treatmentCitation>
, figs. 108, 110 [part, 
<collectingRegion id="49CD32ACD43FFFB8EB49378F6E05F913" box="[801,887,1767,1791]" country="Egypt" name="Al Bahr al Ahmar" pageId="1" pageNumber="454">Red Sea</collectingRegion>
and Persian Gulf specimens].— 
<treatmentCitation id="0AA8DA5FD43FFFB8ECD7378F68EEF913" author="Tattersall, W. M." box="[1215,1436,1767,1791]" page="359" pageId="1" pageNumber="454" year="1921">
<bibRefCitation id="EF9881BFD43FFFB8ECD7378F68EEF913" author="Tattersall, W. M." box="[1215,1436,1767,1791]" pageId="1" pageNumber="454" pagination="345 - 398" refId="ref7536" refString="Tattersall, W. M. (1921) Report on the Stomatopoda and Macrurous Decapoda Collected by Mr. Cyril Crossland in the Sudanese Red Sea. Journal of the Linnean Society, London, Zoology, 34, 345 - 398, pls. 27 + 28. https: // doi. org / 10.1111 / j. 1096 - 3642.1921. tb 01838. x" type="journal article" year="1921">Tattersall 1921: 359–</bibRefCitation>
</treatmentCitation>
360.— 
<bibRefCitation id="EF9881BFD43FFFB8E963366E6F54F8F2" author="Kemp, S. &amp; Chopra, B." box="[267,550,1798,1822]" pageId="1" pageNumber="454" pagination="297 - 311" refId="ref6587" refString="Kemp, S. &amp; Chopra, B. (1921) Notes on Stomatopoda. Records of the Indian Museum, 22, 297 - 311. https: // doi. org / 10.5962 / bhl. part. 1475" type="journal article" year="1921">Kemp &amp; Chopra 1921: 297</bibRefCitation>
, 309–310 [part, 
<collectingRegion id="49CD32ACD43FFFB8EAA6366E6E51F8F2" box="[718,803,1798,1822]" country="Egypt" name="Al Bahr al Ahmar" pageId="1" pageNumber="454">Red Sea</collectingRegion>
and Persian Gulf specimens].— 
<bibRefCitation id="EF9881BFD43FFFB8EC02366E686CF8F2" author="Ramadan, M. M." box="[1130,1310,1798,1822]" pageId="1" pageNumber="454" pagination="1 - 48" refId="ref7349" refString="Ramadan, M. M. (1936) Report on a collection of Stomatopoda and Decapoda from Ghardaqa, Red Sea. The Egyptian University, Bulletin of the Faculty of Science, 6, 1 - 48, pls. 41 - 42." type="journal article" year="1936">Ramadan 1936: 8</bibRefCitation>
.— 
<bibRefCitation id="EF9881BFD43FFFB8ED55366E68EEF8F2" author="Makarov, R. R." box="[1341,1436,1798,1822]" pageId="1" pageNumber="454" pagination="141 - 155" refId="ref6749" refString="Makarov, R. R. (1971) Fauna and distribution of Red Sea Stomatopoda. In: Kovalevsky, A. O. (Ed.), Benthos of the shelf of the Red Sea. Academia nauk SSSR, Kiev, pp. 141 - 155." type="book chapter" year="1971">Makarov</bibRefCitation>
1971: 154.— 
<bibRefCitation id="EF9881BFD43FFFB8E923364C6F43F8D1" author="Basson, P. W. &amp; Burchard, J. E., Jr. &amp; Hardy, J. T. &amp; Price, A. R. G." box="[331,561,1828,1853]" pageId="1" pageNumber="454" refId="ref5938" refString="Basson, P. W., Burchard, J. E., Jr., Hardy, J. T. &amp; Price, A. R. G. (1977) Biotopes of the western Arabian Gulf. Marine life and environments of Saudi Arabia. Aramco, Dharhan, 284 pp." type="book" year="1977">
Basson 
<emphasis id="B97D205CD43FFFB8E9F4364D6CBBF8D0" box="[412,457,1828,1852]" italics="true" pageId="1" pageNumber="454">et al</emphasis>
. 1977: 47
</bibRefCitation>
, 73, 227, 234, 237.— 
<bibRefCitation id="EF9881BFD43FFFB8EB63364C6ECEF8D1" author="Hughes, R. N." box="[779,956,1828,1853]" pageId="1" pageNumber="454" pagination="77 - 96" refId="ref6355" refString="Hughes, R. N. (1977) The biota of reef-flats and limestone cliffs near Jeddah, Saudi Arabia. Journal of Natural History, 11, 77 - 96. https: // doi. org / 10.1080 / 00222937700770051" type="journal article" year="1977">Hughes 1977: 90</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB6FC4ED43FFFB8E8FF362B6E74F8B7" blockId="1.[151,1436,1767,1975]" box="[151,774,1859,1883]" pageId="1" pageNumber="454">
<taxonomicName id="4C0987CDD43FFFB8E8FF362B6C0CF8B7" authority="Demani." authorityName="Demani." box="[151,382,1859,1883]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B97D205CD43FFFB8E8FF362B6C08F8B7" box="[151,378,1859,1883]" italics="true" pageId="1" pageNumber="454">Gonodactylus Demani</emphasis>
.
</taxonomicName>
— 
<bibRefCitation id="EF9881BFD43FFFB8E9F0362B6F45F8B7" author="Nobili, G." box="[408,567,1859,1883]" pageId="1" pageNumber="454" pagination="1 - 12" refId="ref7145" refString="Nobili, G. (1905) Crostacei di Zanzibar. Bollettino dei Musei di Zoologia ed Anatomia comparata della Regia, Universita di Torino, 20 (506), 1 - 12. https: // doi. org / 10.5962 / bhl. part. 9304" type="journal article" year="1905">Nobili 1905: 11</bibRefCitation>
.— 
<bibRefCitation id="EF9881BFD43FFFB8EA3D362B6F9AF8B7" author="Parisi, B." box="[597,744,1859,1883]" pageId="1" pageNumber="454" pagination="1 - 7" refId="ref7297" refString="Parisi, B. (1940) Gli stomatopodi raccolti dal Prof. L. Sanzo nella campagna idrografica nel Mar Rosso della R. N. Ammiraglio Magnaghi 1923 - 1924. Memoria 16. a della Campagna. Reale Comitato Talassografico Italiano, Memoria 275, 1 - 7." type="journal article" year="1940">Parisi 1940: 4</bibRefCitation>
, 6.
</paragraph>
<paragraph id="8BB6FC4ED43FFFB8E8FF360A6C76F874" blockId="1.[151,1436,1767,1975]" pageId="1" pageNumber="454">
<taxonomicName id="4C0987CDD43FFFB8E8FF360A6CFCF896" authority="De Mani." authorityName="De Mani." box="[151,398,1890,1914]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B97D205CD43FFFB8E8FF360A6CFBF895" box="[151,393,1890,1914]" italics="true" pageId="1" pageNumber="454">Gonodactylus De Mani</emphasis>
.
</taxonomicName>
— 
<bibRefCitation id="EF9881BFD43FFFB8E9C0360A6F1AF896" author="Nobili, G." box="[424,616,1890,1914]" pageId="1" pageNumber="454" pagination="1 - 347" refId="ref7194" refString="Nobili, G. (1906 a) Faune carcinologique de la Mer Rouge. Decapodes et Stomatopodes. Annales des Sciences Naturelles, Series 9, 4, 1 - 347, pls. 1 - 11. https: // doi. org / 10.5962 / bhl. title. 10635" type="journal article" year="1906">Nobili 1906a: 330</bibRefCitation>
.— 
<bibRefCitation id="EF9881BFD43FFFB8EAEE360A6E11F896" author="Maccagno, T." box="[646,867,1890,1914]" pageId="1" pageNumber="454" pagination="171 - 186" refId="ref6705" refString="Maccagno, T. (1936) Spedizione del Barone Raimondo Franchetti in Doncalia (1928 - 29). Crostacei di Assab. Decapodi Stomatopodi Anfipodi. Annali del Museo civico di Storia naturale di Genova, 59, 171 - 186." type="journal article" year="1936">Maccagno 1936: 179</bibRefCitation>
.— 
<bibRefCitation id="EF9881BFD43FFFB8EBE9360A6930F896" author="Dollfus, R. P." box="[897,1090,1890,1914]" pageId="1" pageNumber="454" pagination="185 - 236" refId="ref6033" refString="Dollfus, R. P. (1938) Mission Robert Ph. Dollfus en Egypte, X. Stomatopoda (II): Catalogue synonymique des especes jusqu'a present recoltees dans la Mer Rouge, y compris la partie sud du Canal de Suez et le Golfe d'Aden. Memoires de l'Institut d'Egypte, 37, 185 - 236." type="journal article" year="1938">Dollfus 1938: 213</bibRefCitation>
[part, 
<collectingRegion id="49CD32ACD43FFFB8ECE3360A6991F896" box="[1163,1251,1890,1914]" country="Egypt" name="Al Bahr al Ahmar" pageId="1" pageNumber="454">Red Sea</collectingRegion>
and Persian Gulf only].
</paragraph>
<paragraph id="8BB6FC4ED43FFFB8E8FF36F76F2CF85B" blockId="1.[151,1436,1767,1975]" box="[151,606,1951,1975]" pageId="1" pageNumber="454">
<taxonomicName id="4C0987CDD43FFFB8E8FF36F76CF0F85B" authority="de Mani." authorityName="de Mani." box="[151,386,1951,1975]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="1" pageNumber="454" phylum="Arthropoda" rank="genus">
<emphasis id="B97D205CD43FFFB8E8FF36F76C0FF85B" box="[151,381,1951,1975]" italics="true" pageId="1" pageNumber="454">Gonodactylus de Mani</emphasis>
.
</taxonomicName>
— 
<bibRefCitation id="EF9881BFD43FFFB8E9F436F76F28F85B" author="Nobili, G." box="[412,602,1951,1975]" pageId="1" pageNumber="454" pagination="13 - 159" refId="ref7247" refString="Nobili, G. (1906 b) Mission J. Bonnier et Ch. Perez (Golfe Persique, 1901). Crustaces Decapodes et Stomatopodes. Bulletin Scientifique de la France et de la Belgique, 40, 13 - 159, pls. 2 - 5." type="journal article" year="1906">Nobili 1906b: 158</bibRefCitation>
.
</paragraph>
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<paragraph id="8BB6FC4ED43CFFBBE8FF36766DB4F805" blockId="2.[151,1437,1822,2025]" pageId="2" pageNumber="455">
<emphasis id="B97D205CD43CFFBBE8FF36766C66F8DA" bold="true" box="[151,276,1822,1846]" pageId="2" pageNumber="455">FIGURE 1.</emphasis>
<taxonomicName id="4C0987CDD43CFFBBE97336766F55F8DA" authorityName="Ahyong" authorityYear="2024" box="[283,551,1822,1846]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="2" pageNumber="455" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43CFFBBE97336766F55F8DA" box="[283,551,1822,1846]" italics="true" pageId="2" pageNumber="455">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43CFFBBEA4636766FF2F8DA" bold="true" box="[558,640,1822,1846]" pageId="2" pageNumber="455">
<taxonomicNameLabel id="A24E9D27D43CFFBBEA4636766FF2F8DA" box="[558,640,1822,1846]" pageId="2" pageNumber="455" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm.
</paragraph>
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<taxonomicName id="4C0987CDD43DFFBAE8FF31FF6C0CFF43" authorityName="Henderson" authorityYear="1893" box="[151,382,151,175]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43DFFBAE8FF31FF6C0CFF43" box="[151,382,151,175]" italics="true" pageId="3" pageNumber="456">Gonodactylus demanii</emphasis>
</taxonomicName>
.—Holthuis 1941: 282, fig. 8a [part, 
<collectingRegion id="49CD32ACD43DFFBAEA9631FF6E26FF43" box="[766,852,151,175]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
specimens only].— 
<treatmentCitation id="0AA8DA5FD43DFFBAEC4831FF69B2FF43" author="Stella, E." box="[1056,1216,150,175]" page="69" pageId="3" pageNumber="456" year="1955">
<bibRefCitation id="EF9881BFD43DFFBAEC4831FF69B2FF43" author="Stella, E." box="[1056,1216,150,175]" pageId="3" pageNumber="456" pagination="51 - 70" refId="ref7499" refString="Stella, E. (1955) Crostacei decapodi e stomatopodi, pt. III: Spedizione subacquea Italiana nel Mar Rosso, Ricerche zoologiche. Rivista di Biologia coloniale, 13, 51 - 70." type="journal article" year="1955">Stella 1955: 69</bibRefCitation>
</treatmentCitation>
.— 
<treatmentCitation id="0AA8DA5FD43DFFBAECB631FF68EEFF43" author="Dollfus, R. P." box="[1246,1436,150,175]" page="242" pageId="3" pageNumber="456" year="1959">
<bibRefCitation id="EF9881BFD43DFFBAECB631FF68EEFF43" author="Dollfus, R. P." box="[1246,1436,150,175]" pageId="3" pageNumber="456" pagination="241 - 245" refId="ref6094" refString="Dollfus, R. P. (1959) Mission Robert Ph. Dollfus en Egypte, XXX. Stomatopoda (III). Mission Robert Ph. Dollfus en Egypte (Decembre 1927 - Mars 1929). S. S. Al Sayad. Resultats scientifiques, 3 e partie (XXIII - XXXIV), 241 - 245." type="book chapter" year="1959">Dollfus 1959: 242</bibRefCitation>
</treatmentCitation>
[part; discussion].— 
<treatmentCitation id="0AA8DA5FD43DFFBAE9FE31DD6F08FF21" author="Holthuis, L. B." box="[406,634,181,205]" page="32" pageId="3" pageNumber="456" year="1967">
<bibRefCitation id="EF9881BFD43DFFBAE9FE31DD6F08FF21" author="Holthuis, L. B." box="[406,634,181,205]" pageId="3" pageNumber="456" pagination="1 - 45" refId="ref6299" refString="Holthuis, L. B. (1967) The stomatopod Crustacea collected by the 1962 and 1965 Israel South Red Sea Expeditions. The Second Israel South Red Sea Expedition, 1965, Report No. 1. Israel Journal of Zoology, 16, 1 - 45, figs. 1 - 7." type="journal article" year="1967">Holthuis 1967: 32–34</bibRefCitation>
</treatmentCitation>
, 41.— 
<treatmentCitation id="0AA8DA5FD43DFFBAEAD331DD6EEAFF21" author="Manning, R. B." box="[699,920,181,205]" page="8" pageId="3" pageNumber="456" year="1967">
<bibRefCitation id="EF9881BFD43DFFBAEAD331DD6EEAFF21" author="Manning, R. B." box="[699,920,181,205]" pageId="3" pageNumber="456" pagination="1 - 27" refId="ref6800" refString="Manning, R. B. (1967) Notes on the demanii section of genus Gonodactylus Berthold with descriptions of three new species (Crustacea: Stomatopoda). Proceedings of the United States National Museum, 123, 1 - 27. https: // doi. org / 10.5479 / si. 00963801.123 - 3618.1" type="journal article" year="1967">Manning 1967: 8–10</bibRefCitation>
</treatmentCitation>
, fig. 3 [part, except Gulf of Mannar specimen].— 
<treatmentCitation id="0AA8DA5FD43DFFBAE8AF31BC6CD6FF00" author="Shanbhogue, S. L." box="[199,420,212,236]" page="35" pageId="3" pageNumber="456" year="1969">
<bibRefCitation id="EF9881BFD43DFFBAE8AF31BC6CD6FF00" author="Shanbhogue, S. L." box="[199,420,212,236]" pageId="3" pageNumber="456" pagination="33 - 36" refId="ref7395" refString="Shanbhogue, S. L. (1969) Catalogue of stomatopods in reference collections of the Central Marine Fisheries Research Institute. Bulletin of the Central Marine Fisheries Research Institute, 9, 33 - 36." type="journal article" year="1969">Shanbhogue 1969: 35</bibRefCitation>
</treatmentCitation>
[part, Gulf of Kutch record only].— 
<treatmentCitation id="0AA8DA5FD43DFFBAEB7031BA6E8FFF00" author="Fishelson, L." box="[792,1021,210,236]" page="119" pageId="3" pageNumber="456" year="1971">
<bibRefCitation id="EF9881BFD43DFFBAEB7031BA6E8FFF00" author="Fishelson, L." box="[792,1021,210,236]" pageId="3" pageNumber="456" pagination="113 - 133" refId="ref6156" refString="Fishelson, L. (1971) Ecology and distribution of the benthic fauna in the shallow waters of the Red Sea. Marine Biology, 10, 113 - 133. https: // doi. org / 10.1007 / BF 00354828" type="journal article" year="1971">Fishelson 1971: 119</bibRefCitation>
</treatmentCitation>
, 128.— 
<treatmentCitation id="0AA8DA5FD43DFFBAEC3E31BC68E5FF00" author="Manning, R. B. &amp; Lewinsohn, C." box="[1110,1431,212,236]" page="6" pageId="3" pageNumber="456" year="1986">
<bibRefCitation id="EF9881BFD43DFFBAEC3E31BC68E5FF00" author="Manning, R. B. &amp; Lewinsohn, C." box="[1110,1431,212,236]" pageId="3" pageNumber="456" pagination="1 - 19" refId="ref7049" refString="Manning, R. B. &amp; Lewinsohn, C. (1986) Notes on some stomatopod Crustacea from the Sinai Peninsula, Red Sea. Smithsonian Contributions to Zoology, 433, i - iv + 1 - 19. https: // doi. org / 10.5479 / si. 00810282.433" type="journal article" year="1986">Manning &amp; Lewinsohn 1986: 6</bibRefCitation>
</treatmentCitation>
, 7, 15, 16.— 
<treatmentCitation id="0AA8DA5FD43DFFBAE956319B6F42FEE6" author="Shanbhogue, S. L." box="[318,560,242,267]" page="519" pageId="3" pageNumber="456" year="1986">
<bibRefCitation id="EF9881BFD43DFFBAE956319B6F42FEE6" author="Shanbhogue, S. L." box="[318,560,242,267]" pageId="3" pageNumber="456" pagination="515 - 567" refId="ref7433" refString="Shanbhogue, S. L. (1986) Studies on stomatopod Crustacea from the seas around India. In: James, P. S. B. R. (Ed.), Recent Advances in Marine Biology: Dr Jones' 70 th birthday commemoration volume. Today &amp; Tomorrow's Printers &amp; Publishers, New Delhi, pp. 515 - 567." type="book chapter" year="1986">Shanbhogue 1986: 519</bibRefCitation>
</treatmentCitation>
, 521 [part, Gulf of Kutch record only].— 
<bibRefCitation id="EF9881BFD43DFFBAEBB0319B6906FEE7" author="Manning, R. B." box="[984,1140,243,267]" pageId="3" pageNumber="456" pagination="113 - 119" refId="ref6900" refString="Manning, R. B. (1989) Zoogeographical relationships of stomatopod Crustacea from the northern Arabian Sea. In: Thompson, M. - S. &amp; Tirmizi, N. (Eds.), Marine Science of the Arabian Sea, Proceedings of an International Conference. American Institute of Biological Sciences, Washington, D. C., pp. 113 - 119." type="book chapter" year="1989">Manning 1989</bibRefCitation>
: tab. 1.— 
<treatmentCitation id="0AA8DA5FD43DFFBAECBB319B68EAFEE7" author="Manning, R. B." box="[1235,1432,243,267]" page="98" pageId="3" pageNumber="456" year="1990">
<bibRefCitation id="EF9881BFD43DFFBAECBB319B68EAFEE7" author="Manning, R. B." box="[1235,1432,243,267]" pageId="3" pageNumber="456" pagination="93 - 108" refId="ref6971" refString="Manning, R. B. (1990) Stomatopod Crustacea from the Persian Gulf, with the description of a new Manningia. Steenstrupia, 16, 93 - 108." type="journal article" year="1990">Manning 1990: 98</bibRefCitation>
</treatmentCitation>
, 104.— 
<treatmentCitation id="0AA8DA5FD43DFFBAE96130796C8DFEC5" author="Tirmizi, N. M. &amp; Kazmi, Q. B. &amp; Manning, R. B." box="[265,511,273,297]" page="161" pageId="3" pageNumber="456" year="1994">
<bibRefCitation id="EF9881BFD43DFFBAE96130796C8DFEC5" author="Tirmizi, N. M. &amp; Kazmi, Q. B. &amp; Manning, R. B." box="[265,511,273,297]" pageId="3" pageNumber="456" pagination="125 - 169" refId="ref7605" refString="Tirmizi, N. M., Kazmi, Q. B. &amp; Manning, R. B. (1994) An illustrated key to the Malacostraca (Crustacea) of the northern Arabian Sea. Part II: Stomatopoda. Pakistan Journal of Marine Science, 3, 125 - 169." type="journal article" year="1994">
Tirmizi 
<emphasis id="B97D205CD43DFFBAE936307A6CF9FEC5" box="[350,395,273,297]" italics="true" pageId="3" pageNumber="456">et al</emphasis>
. 1994: 161
</bibRefCitation>
</treatmentCitation>
, fig. 35.— 
<treatmentCitation id="0AA8DA5FD43DFFBAEA0230796E0AFEC5" author="Joydas, T. V. &amp; Borja, A." box="[618,888,273,297]" page="234" pageId="3" pageNumber="456" year="2019">
<bibRefCitation id="EF9881BFD43DFFBAEA0230796E0AFEC5" author="Joydas, T. V. &amp; Borja, A." box="[618,888,273,297]" pageId="3" pageNumber="456" pagination="222 - 253" refId="ref6445" refString="Joydas, T. V. &amp; Borja, A. (2019) Benthic ecosystems (soft and hard substrata). In: Al-Abdulkader, K. A., Loughland, R. A. &amp; Qurban, M. A. (Eds.), Ecosystems and biodiversity of the Arabian Gulf: fifty years of scientific research. Saudi Aramco &amp; King Fahd University of Petroleum &amp; Minerals, Dhahran, pp. 222 - 253." type="book chapter" year="2019">Joydas &amp; Borja 2019: 234</bibRefCitation>
</treatmentCitation>
, 239.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB6FC4ED43DFFBAE8FF30586ED3FEA4" blockId="3.[151,1437,150,451]" box="[151,929,304,328]" pageId="3" pageNumber="456">
<treatmentCitationGroup id="AB19DB60D43DFFBAE8FF30586ED3FEA4" box="[151,929,304,328]" pageId="3" pageNumber="456">
<emphasis id="B97D205CD43DFFBAE8FF30586C90FEA4" box="[151,482,304,328]" italics="true" pageId="3" pageNumber="456">
<taxonomicName id="4C0987CDD43DFFBAE8FF30586CB7FEA4" box="[151,453,304,328]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="subSpecies" species="demani" subSpecies="demani">Gonodactylus demani demani</taxonomicName>
.—
</emphasis>
<treatmentCitation id="0AA8DA5FD43DFFBAE98B30586F04FEA4" author="Ingle, R. W." box="[483,630,304,328]" page="27" pageId="3" pageNumber="456" year="1963">
<bibRefCitation id="EF9881BFD43DFFBAE98B30586F04FEA4" author="Ingle, R. W." box="[483,630,304,328]" pageId="3" pageNumber="456" pagination="1 - 69" refId="ref6398" refString="Ingle, R. W. (1963) Contributions to the knowledge of the Red Sea, No. 26. Crustacea Stomatopoda from the Red Sea and Gulf of Aden. Sea Fisheries Research Station Bulletin, Haifa, 33, 1 - 69." type="journal article" year="1963">Ingle 1963: 27</bibRefCitation>
</treatmentCitation>
[part, 
<collectingRegion id="49CD32ACD43DFFBAEAD130586E7FFEA4" box="[697,781,304,328]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
records only].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB6FC4ED43DFFBAE8FF302769E2FE8B" blockId="3.[151,1437,150,451]" box="[151,1168,335,359]" pageId="3" pageNumber="456">
<treatmentCitationGroup id="AB19DB60D43DFFBAE8FF302769E2FE8B" box="[151,1168,335,359]" pageId="3" pageNumber="456">
<treatmentCitation id="0AA8DA5FD43DFFBAE8FF30276F24FE8B" author="Manning, R. B." box="[151,598,335,359]" page="4" pageId="3" pageNumber="456" year="1967">
<taxonomicName id="4C0987CDD43DFFBAE8FF30276F24FE8B" authority="Manning, 1967: 4" authorityName="Manning" authorityPageNumber="4" authorityYear="1967" box="[151,598,335,359]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="hendersoni">
<emphasis id="B97D205CD43DFFBAE8FF30276CE9FE8B" box="[151,411,335,359]" italics="true" pageId="3" pageNumber="456">Gonodactylus hendersoni</emphasis>
<bibRefCitation id="EF9881BFD43DFFBAE9C930276F24FE8B" author="Manning, R. B." box="[417,598,335,359]" pageId="3" pageNumber="456" pagination="1 - 27" refId="ref6800" refString="Manning, R. B. (1967) Notes on the demanii section of genus Gonodactylus Berthold with descriptions of three new species (Crustacea: Stomatopoda). Proceedings of the United States National Museum, 123, 1 - 27. https: // doi. org / 10.5479 / si. 00963801.123 - 3618.1" type="journal article" year="1967">Manning, 1967: 4</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[part, 
<typeStatus id="54B242ECD43DFFBAEAF130276F80FE8B" box="[665,754,335,359]" pageId="3" pageNumber="456" type="holotype">holotype</typeStatus>
only; 
<typeStatus id="54B242ECD43DFFBAEB5A30276EE1FE8B" box="[818,915,335,359]" pageId="3" pageNumber="456" type="paratype">paratypes</typeStatus>
represent other species].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB6FC4ED43DFFBAE8FF300568EFFE69" blockId="3.[151,1437,150,451]" box="[151,1437,365,389]" pageId="3" pageNumber="456">
<treatmentCitationGroup id="AB19DB60D43DFFBAE8FF300568EFFE69" box="[151,1437,365,389]" pageId="3" pageNumber="456">
<taxonomicName id="4C0987CDD43DFFBAE8FF30056C8FFE69" box="[151,509,365,389]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="variety" species="demani" variety="spinosus">
<emphasis id="B97D205CD43DFFBAE8FF30056C07FE69" box="[151,373,365,389]" italics="true" pageId="3" pageNumber="456">Gonodactylus demani</emphasis>
var. 
<emphasis id="B97D205CD43DFFBAE9CF30066C8FFE69" box="[423,509,366,389]" italics="true" pageId="3" pageNumber="456">spinosus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43DFFBAE99530066F53FE69" box="[509,545,366,389]" italics="true" pageId="3" pageNumber="456">?—</emphasis>
<treatmentCitation id="0AA8DA5FD43DFFBAEA4930066E47FE69" author="Kemp, S. &amp; Chopra, B." box="[545,821,365,389]" page="311" pageId="3" pageNumber="456" year="1921">
<bibRefCitation id="EF9881BFD43DFFBAEA4930066E47FE69" author="Kemp, S. &amp; Chopra, B." box="[545,821,365,389]" pageId="3" pageNumber="456" pagination="297 - 311" refId="ref6587" refString="Kemp, S. &amp; Chopra, B. (1921) Notes on Stomatopoda. Records of the Indian Museum, 22, 297 - 311. https: // doi. org / 10.5962 / bhl. part. 1475" type="journal article" year="1921">Kemp &amp; Chopra 1921: 311</bibRefCitation>
</treatmentCitation>
[Pearl Banks, Persian Gulf; not 
<taxonomicName id="4C0987CDD43DFFBAEC15300668E1FE69" authority="Bigelow, 1893" authorityName="Bigelow" authorityYear="1893" box="[1149,1427,365,389]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B97D205CD43DFFBAEC1530066981FE69" box="[1149,1267,366,389]" italics="true" pageId="3" pageNumber="456">G. spinosus</emphasis>
<bibRefCitation id="EF9881BFD43DFFBAEC92300568E1FE69" author="Bigelow, R. P." box="[1274,1427,365,389]" pageId="3" pageNumber="456" pagination="100 - 102" refId="ref5995" refString="Bigelow, R. P. (1893) Preliminary notes on the Stomatopoda of the Albatross collections and on other specimens in the National Museum. Johns Hopkins University Circulars, 12, 100 - 102." type="journal article" year="1893">Bigelow, 1893</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB6FC4ED43DFFBAE8FF30E46CBBFE2F" blockId="3.[151,1437,150,451]" pageId="3" pageNumber="456">
<treatmentCitationGroup id="AB19DB60D43DFFBAE8FF30E46CBBFE2F" pageId="3" pageNumber="456">
<taxonomicName id="4C0987CDD43DFFBAE8FF30E46CE7FE48" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[151,405,396,420]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43DFFBAE8FF30E46CE7FE48" box="[151,405,396,420]" italics="true" pageId="3" pageNumber="456">Gonodactylellus demanii</emphasis>
</taxonomicName>
.— 
<treatmentCitation id="0AA8DA5FD43DFFBAE9DA30E46FD0FE48" author="Trivedi, J. N. &amp; Ahyong, S. T. &amp; Vachhrajani, K. D. &amp; Kumar, A. B." box="[434,674,396,420]" page="224" pageId="3" pageNumber="456" year="2020">
<bibRefCitation id="EF9881BFD43DFFBAE9DA30E46FD0FE48" author="Trivedi, J. N. &amp; Ahyong, S. T. &amp; Vachhrajani, K. D. &amp; Kumar, A. B." box="[434,674,396,420]" pageId="3" pageNumber="456" pagination="221 - 238" refId="ref7710" refString="Trivedi, J. N., Ahyong, S. T., Vachhrajani, K. D. &amp; Kumar, A. B. (2020) An annotated checklist of the mantis shrimps of India (Crustacea: Stomatopoda). Zootaxa, 4768 (2), 221 - 238. https: // doi. org / 10.11646 / zootaxa. 4768.2.4" type="journal article" year="2020">
Trivedi 
<emphasis id="B97D205CD43DFFBAEA6C30E56F42FE48" box="[516,560,396,420]" italics="true" pageId="3" pageNumber="456">et al</emphasis>
. 2020: 224
</bibRefCitation>
</treatmentCitation>
[part, 
<collectingRegion id="49CD32ACD43DFFBAEA8C30E46E42FE48" box="[740,816,396,420]" country="India" name="Gujarat" pageId="3" pageNumber="456">Gujarat</collectingRegion>
and 
<collectingRegion id="49CD32ACD43DFFBAEB0830E46EADFE48" box="[864,991,396,420]" country="India" name="Maharashtra" pageId="3" pageNumber="456">Maharashtra</collectingRegion>
records only].— 
<treatmentCitation id="0AA8DA5FD43DFFBAECED30E468FDFE48" author="Vadher, P. &amp; Kardani, H. &amp; Beleem, I." box="[1157,1423,396,420]" page="21034" pageId="3" pageNumber="456" year="2022">
<bibRefCitation id="EF9881BFD43DFFBAECED30E4683AFE48" author="Vadher, P. &amp; Kardani, H. &amp; Beleem, I." box="[1157,1352,396,420]" pageId="3" pageNumber="456" pagination="21032 - 21042" refId="ref7778" refString="Vadher, P., Kardani, H. &amp; Beleem, I. (2022) Diversity and distribution of mantis shrimps (Arthropoda: Crustacea: Stomatopoda) in the Gulf of Kachchh, Gujarat, India. Journal of Threatened Taxa, 14, 21032 - 21042. https: // doi. org / 10.11609 / jott. 7471.14.5.21032 - 21042" type="journal article" year="2022">
Vadher 
<emphasis id="B97D205CD43DFFBAECB030E56877FE48" box="[1240,1285,396,420]" italics="true" pageId="3" pageNumber="456">et al</emphasis>
. 2022
</bibRefCitation>
: 21034
</treatmentCitation>
– 21036, image 1, tab. 1, 2.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF76ACC6D43DFFBAE8FF345D6F6DFA54" ID-DOI="http://doi.org/10.5281/zenodo.12780239" ID-Zenodo-Dep="12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="3" pageNumber="456" startId="3.[151,250,1333,1357]" targetBox="[176,1410,481,1302]" targetPageId="3" targetType="figure">
<paragraph id="8BB6FC4ED43DFFBAE8FF345D6F6DFA54" blockId="3.[151,1437,1333,1465]" pageId="3" pageNumber="456">
<emphasis id="B97D205CD43DFFBAE8FF345D6C61FAA1" bold="true" box="[151,275,1333,1357]" pageId="3" pageNumber="456">FIGURE 2.</emphasis>
<taxonomicName id="4C0987CDD43DFFBAE972345D6F57FAA1" authorityName="Ahyong" authorityYear="2024" box="[282,549,1333,1357]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="3" pageNumber="456" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43DFFBAE972345D6F57FAA1" box="[282,549,1333,1357]" italics="true" pageId="3" pageNumber="456">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43DFFBAEA44345D6F0FFAA1" bold="true" box="[556,637,1333,1357]" pageId="3" pageNumber="456">
<taxonomicNameLabel id="A24E9D27D43DFFBAEA44345D6F0FFAA1" box="[556,637,1333,1357]" pageId="3" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C313AFC5D43DFFBDE8FF37636C26FCB1" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456" type="materials_examined">
<paragraph id="8BB6FC4ED43DFFBAE8FF376369B9F95A" blockId="3.[151,1437,1547,2042]" pageId="3" pageNumber="456">
<emphasis id="B97D205CD43DFFBAE8FF37636C4CF9C9" bold="true" box="[151,318,1547,1573]" pageId="3" pageNumber="456">Type material</emphasis>
. 
<materialsCitation id="3B61F613D43DFFBAE924376368EBF9A5" collectingDate="2013-06-26" collectionCode="SMF, RSS" collectorName="A. Brosing &amp; Brosing, A &amp; B. Werding" elevation="5" latitude="28.014" location="Alkhuraybah" longLatPrecision="1" longitude="35.163086" pageId="3" pageNumber="456" specimenCode="SMF 62255, RSS1-2013, 35" specimenCount="1" stateProvince="Al Bahr al Ahmar" typeStatus="holotype">
<typeStatus id="54B242ECD43DFFBAE92437636CCBF9C9" box="[332,441,1547,1573]" pageId="3" pageNumber="456" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="DBAF5435D43DFFBAE9AA37636F38F9CA" box="[450,586,1547,1574]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="456">SMF 62255</specimenCode>
, male (TL 
<quantity id="4CF151ABD43DFFBAEAAC37636E65F9C9" box="[708,791,1547,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="456" unit="mm" value="27.0">27 mm</quantity>
), 
<location id="8ED6AA95D43DFFBAEB4037636ECFF9C9" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAEB4037636ECFF9C9" box="[808,957,1547,1573]" latitude="28.014" longLatPrecision="1" longitude="35.163086" name="Alkhuraybah" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Alkhuraybah</location>
, 
<collectingRegion id="49CD32ACD43DFFBAEBA037636955F9C9" box="[968,1063,1547,1573]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEC5A37636993F9CA" box="[1074,1249,1547,1574]" degrees="28" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="456" precision="1" seconds="50.40" value="28.014">28°00’50.40”N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEC84376368EAF9CA" box="[1260,1432,1547,1574]" degrees="35" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="456" precision="1" seconds="47.10" value="35.163086">35°09’47.10”E</geoCoordinate>
, 
<quantity id="4CF151ABD43DFFBAE8FF37476DACF9A6" box="[151,222,1583,1610]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">
<elevation id="00241B7DD43DFFBAE8FF37476DACF9A6" box="[151,222,1583,1610]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, sandy-bottom &amp; coral rocks, 
<specimenCode id="DBAF5435D43DFFBAEA2137476FBEF9A6" box="[585,716,1583,1610]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2013</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAEABD37476F80F9A6" box="[725,754,1583,1610]" collectionCode="RSS" pageId="3" pageNumber="456">35</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAEB5537476ECDF9A5" box="[829,959,1583,1609]" pageId="3" pageNumber="456">A. Brösing</collectorName>
, A. Alhaj &amp; 
<collectorName id="26FC9998D43DFFBAEC3737586995F9A5" box="[1119,1255,1583,1610]" pageId="3" pageNumber="456">B. Werding</collectorName>
, 
<date id="FFB7DA8ED43DFFBAEC9E374768EBF9A5" box="[1270,1433,1583,1610]" pageId="3" pageNumber="456" value="2013-06-26">
<collectingDate id="EFF32366D43DFFBAEC9E374768EBF9A5" box="[1270,1433,1583,1610]" pageId="3" pageNumber="456" value="2013-06-26">26 June 2013</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B61F613D43DFFBAE8FF373C6CECF982" box="[151,414,1619,1646]" collectionCode="SMF" pageId="3" pageNumber="456" specimenCode="SMF 62256" specimenCount="1" stateProvince="Al Bahr al Ahmar" typeStatus="paratype">
<typeStatus id="54B242ECD43DFFBAE8FF373C6C7AF982" box="[151,264,1620,1646]" pageId="3" pageNumber="456" type="paratype">Paratypes</typeStatus>
: 
<specimenCode id="DBAF5435D43DFFBAE97D373B6CECF982" box="[277,414,1619,1646]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="456">SMF 62256</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAE9C3373B6F90F97D" elevation="5" latitude="28.014" location="Alkhuraybah" longLatPrecision="1" longitude="35.163086" pageId="3" pageNumber="456" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Al Bahr al Ahmar" typeStatus="paratype">
<specimenCount id="9D0F37C7D43DFFBAE9C3373B6C8AF981" box="[427,504,1619,1645]" count="1" pageId="3" pageNumber="456" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEA5C373B6FFBF981" box="[564,649,1619,1645]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="456" unit="mm" value="36.0">36 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAEAF6373B6E79F981" box="[670,779,1619,1645]" count="2" pageId="3" pageNumber="456" type="female">2 females</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEB2F373B6EB4F981" box="[839,966,1619,1645]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="3" pageNumber="456" unit="mm" value="22.0" valueMax="24.0" valueMin="20.0">20–24 mm</quantity>
), 
<location id="8ED6AA95D43DFFBAEBB1373B691CF981" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAEBB1373B691CF981" box="[985,1134,1619,1645]" latitude="28.014" longLatPrecision="1" longitude="35.163086" name="Alkhuraybah" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Alkhuraybah</location>
, 
<collectingRegion id="49CD32ACD43DFFBAEC13373B69AEF981" box="[1147,1244,1619,1645]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEC81373B68EAF982" box="[1257,1432,1619,1646]" degrees="28" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="456" precision="1" seconds="50.40" value="28.014">28°00’50.40”N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAE8FF371F6C31F97E" box="[151,323,1655,1682]" degrees="35" direction="east" minutes="09" orientation="longitude" pageId="3" pageNumber="456" precision="1" seconds="47.10" value="35.163086">35°09’47.10”E</geoCoordinate>
, 
<quantity id="4CF151ABD43DFFBAE938371F6CE1F97E" box="[336,403,1655,1682]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">
<elevation id="00241B7DD43DFFBAE938371F6CE1F97E" box="[336,403,1655,1682]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, sandy-bottom &amp; coral rocks
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAEA87371F6C7FF959" collectingDate="2013-06-26" collectionCode="RSS" collectorName="A. Brosing &amp; A. Alhaj &amp; B. Werding" pageId="3" pageNumber="456" specimenCode="RSS1-2013, 35" specimenCount="1" stateProvince="Al Bahr al Ahmar" typeStatus="paratype">
<specimenCode id="DBAF5435D43DFFBAEA87371F6E00F97E" box="[751,882,1655,1682]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2013</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAEB13371F6EEAF97E" box="[891,920,1655,1682]" collectionCode="RSS" pageId="3" pageNumber="456">35</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAEBB5371F692EF97D" box="[989,1116,1655,1681]" pageId="3" pageNumber="456">A. Brösing</collectorName>
, 
<collectorName id="26FC9998D43DFFBAEC00371F69BAF97D" box="[1128,1224,1655,1681]" pageId="3" pageNumber="456">A. Alhaj</collectorName>
&amp; 
<collectorName id="26FC9998D43DFFBAEC8737106806F97D" box="[1263,1396,1655,1682]" pageId="3" pageNumber="456">B. Werding</collectorName>
, 
<date id="FFB7DA8ED43DFFBAEDE8371F6C7FF959" pageId="3" pageNumber="456" value="2013-06-26">
<collectingDate id="EFF32366D43DFFBAEDE8371F6C7FF959" pageId="3" pageNumber="456" value="2013-06-26">26 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43DFFBAE97037F36CBBF95A" box="[280,457,1691,1718]" collectionCode="KAUMM" pageId="3" pageNumber="456" specimenCode="KAUMM 1059" specimenCount="1" stateProvince="Al Bahr al Ahmar" typeStatus="paratype">
<specimenCode id="DBAF5435D43DFFBAE97037F36CBBF95A" box="[280,457,1691,1718]" collectionCode="KAUMM" pageId="3" pageNumber="456">KAUMM 1059</specimenCode>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAE9BC37F369B4F95A" box="[468,1222,1691,1718]" collectionCode="RSS" pageId="3" pageNumber="456" specimenCode="RSS1-2013, 35" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Al Bahr al Ahmar" typeStatus="paratype">
<specimenCount id="9D0F37C7D43DFFBAE9BC37F36F6DF959" box="[468,543,1691,1717]" count="1" pageId="3" pageNumber="456" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEA3037F36FD9F959" box="[600,683,1691,1717]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="456" unit="mm" value="27.0">27 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAEAD637F36E58F959" box="[702,810,1691,1717]" count="3" pageId="3" pageNumber="456" type="female">3 females</specimenCount>
(
<quantity id="4CF151ABD43DFFBAEB5037F36EC5F959" box="[824,951,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="5.0" metricValueMin="2.3" pageId="3" pageNumber="456" unit="mm" value="36.5" valueMax="50.0" valueMin="23.0">23–50 mm</quantity>
), same (
<specimenCode id="DBAF5435D43DFFBAEC7B37F369EBF95A" box="[1043,1177,1691,1718]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2013</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAECC937F369CCF95A" box="[1185,1214,1691,1718]" collectionCode="RSS" pageId="3" pageNumber="456">35</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB6FC4ED43DFFBDE8AF37D7692BFD69" blockId="3.[151,1437,1547,2042]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="457" pageId="3" pageNumber="456">
<emphasis id="B97D205CD43DFFBAE8AF37D76C86F935" bold="true" box="[199,500,1727,1753]" pageId="3" pageNumber="456">Other material examined</emphasis>
. 
<materialsCitation id="3B61F613D43DFFBAEA6A37A86907F935" box="[514,1141,1727,1754]" collectionCode="SMF" location="Al Wajh" pageId="3" pageNumber="456" specimenCode="SMF 62257" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahr al Ahmar">
<collectingRegion id="49CD32ACD43DFFBAEA6A37A86F08F935" box="[514,634,1727,1754]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">RED SEA</collectingRegion>
: 
<specimenCode id="DBAF5435D43DFFBAEAEC37D76E7EF936" box="[644,780,1727,1754]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="456">SMF 62257</specimenCode>
, 
<specimenCount id="9D0F37C7D43DFFBAEB7F37D76E0AF935" box="[791,888,1727,1753]" count="1" pageId="3" pageNumber="456" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEBD837D76971F935" box="[944,1027,1727,1753]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="456" unit="mm" value="17.0">17 mm</quantity>
), 
<location id="8ED6AA95D43DFFBAEC7C37D76907F935" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAEC7C37D76907F935" box="[1044,1141,1727,1753]" name="Al Wajh" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Al Wajh</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAEC1737D769A8F911" collectingDate="2013-06-11" collectionCode="RSS" collectorName="A. A. J. Kumar" latitude="26.058277" location="Red Sea" longLatPrecision="1" longitude="36.594723" pageId="3" pageNumber="456" specimenCode="RSS1-2013, 1" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<collectingRegion id="49CD32ACD43DFFBAEC1737D769ACF935" box="[1151,1246,1727,1753]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEC8137D768EAF935" box="[1257,1432,1727,1753]" degrees="26" direction="north" minutes="03" orientation="latitude" pageId="3" pageNumber="456" precision="1" seconds="29.80" value="26.058277">26°03’29.80”N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAE8FF378B6C31F912" box="[151,323,1763,1790]" degrees="36" direction="east" minutes="35" orientation="longitude" pageId="3" pageNumber="456" precision="1" seconds="41.00" value="36.594723">36°35’41.00”E</geoCoordinate>
, intertidal reef flat, dead corals, 
<specimenCode id="DBAF5435D43DFFBAEADC378B6E4FF911" box="[692,829,1763,1789]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2013</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAEB2D378B6E3CF911" box="[837,846,1763,1789]" collectionCode="RSS" pageId="3" pageNumber="456">1</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAEBFB378B6946F912" box="[915,1076,1763,1790]" pageId="3" pageNumber="456">A.A.J. Kumar</collectorName>
, 
<date id="FFB7DA8ED43DFFBAEC28378B69A8F911" box="[1088,1242,1763,1790]" pageId="3" pageNumber="456" value="2013-06-11">
<collectingDate id="EFF32366D43DFFBAEC28378B69A8F911" box="[1088,1242,1763,1790]" pageId="3" pageNumber="456" value="2013-06-11">11 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43DFFBAEC8F378B6C5EF8A9" collectionCode="KAUMM" location="Alkhuraybah" pageId="3" pageNumber="456" specimenCode="KAUMM 1060" specimenCount="10" specimenCount-female="4" specimenCount-juvenile="5" specimenCount-male="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43DFFBAEC8F378B68EAF911" box="[1255,1432,1763,1789]" collectionCode="KAUMM" pageId="3" pageNumber="456">KAUMM 1060</specimenCode>
, 
<specimenCount id="9D0F37C7D43DFFBAE8FF366F6D91F8CD" box="[151,227,1799,1825]" count="1" pageId="3" pageNumber="456" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAE975366F6C03F8CD" box="[285,369,1799,1825]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="456" unit="mm" value="31.0">31 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAE9EC366F6C86F8CD" box="[388,500,1799,1825]" count="4" pageId="3" pageNumber="456" type="juvenile">4 juvenile</specimenCount>
males (TL 
<quantity id="4CF151ABD43DFFBAEA1F366F6F87F8CD" box="[631,757,1799,1826]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="3" pageNumber="456" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAEB60366F6E07F8CD" box="[776,885,1799,1825]" count="4" pageId="3" pageNumber="456" type="female">4 females</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEBC7366F695FF8CD" box="[943,1069,1799,1825]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.1" metricValueMin="2.3" pageId="3" pageNumber="456" unit="mm" value="27.0" valueMax="31.0" valueMin="23.0">23–31 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAEC28366F69C2F8CD" box="[1088,1200,1799,1825]" count="1" pageId="3" pageNumber="456" type="juvenile">1 juvenile</specimenCount>
female (TL 
<quantity id="4CF151ABD43DFFBAED56366F68E2F8CD" box="[1342,1424,1799,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="456" unit="mm" value="11.0">11 mm</quantity>
), 
<location id="8ED6AA95D43DFFBAE8FF36436C5EF8A9" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAE8FF36436C5EF8A9" box="[151,300,1835,1861]" name="Alkhuraybah" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Alkhuraybah</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAE95E36436F70F885" collectingDate="2013-06-23" collectionCode="RSS" collectorName="A. Alhaj" elevation="5" latitude="28.107668" location="Red Sea" longLatPrecision="1" longitude="34.99089" pageId="3" pageNumber="456" specimenCode="RSS1-2013, 37" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<collectingRegion id="49CD32ACD43DFFBAE95E36436CE1F8A9" box="[310,403,1835,1861]" country="Egypt" name="Al Bahr al Ahmar" pageId="3" pageNumber="456">Red Sea</collectingRegion>
, 
<geoCoordinate id="EE3D9A89D43DFFBAE9F536436F3EF8A9" box="[413,588,1835,1861]" degrees="28" direction="north" minutes="06" orientation="latitude" pageId="3" pageNumber="456" precision="1" seconds="27.60" value="28.107668">28°06’27.60”N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEA3E36436E70F8AA" box="[598,770,1835,1862]" degrees="34" direction="east" minutes="59" orientation="longitude" pageId="3" pageNumber="456" precision="1" seconds="27.20" value="34.99089">34°59’27.20”E</geoCoordinate>
, 
<quantity id="4CF151ABD43DFFBAEB6436436E3EF8AA" box="[780,844,1835,1862]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">
<elevation id="00241B7DD43DFFBAEB6436436E3EF8AA" box="[780,844,1835,1862]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="456" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, sandy bottom with coral rocks, digging, 
<specimenCode id="DBAF5435D43DFFBAED7A364368E7F8A9" box="[1298,1429,1835,1861]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2013</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAE8FF36276DC7F885" box="[151,181,1871,1897]" collectionCode="RSS" pageId="3" pageNumber="456">37</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAE89136276C2EF885" box="[249,348,1871,1897]" pageId="3" pageNumber="456">A. Alhaj</collectorName>
, 
<date id="FFB7DA8ED43DFFBAE90036276F70F885" box="[360,514,1871,1898]" pageId="3" pageNumber="456" value="2013-06-23">
<collectingDate id="EFF32366D43DFFBAE90036276F70F885" box="[360,514,1871,1898]" pageId="3" pageNumber="456" value="2013-06-23">23 June 2013</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43DFFBAEA673627692BF885" box="[527,1113,1871,1897]" collectionCode="KAUMM" collectorName="Al Lith" pageId="3" pageNumber="456" specimenCode="KAUMM 1061" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43DFFBAEA6736276FCDF885" box="[527,703,1871,1897]" collectionCode="KAUMM" pageId="3" pageNumber="456">KAUMM 1061</specimenCode>
, juvenile male (TL 
<quantity id="4CF151ABD43DFFBAEBF736276E80F885" box="[927,1010,1871,1897]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="456" unit="mm" value="13.0">13 mm</quantity>
), 
<collectorName id="26FC9998D43DFFBAEC6D3627692BF885" box="[1029,1113,1871,1897]" pageId="3" pageNumber="456">Al Lith</collectorName>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAEC0E36276F94F861" collectingDate="2012-03-06" collectionCode="RSS" latitude="20.2002" longLatPrecision="1" longitude="40.1184" pageId="3" pageNumber="456" specimenCode="RSS1-2012, 58" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43DFFBAEC0E36276988F885" box="[1126,1274,1871,1897]" degrees="20" direction="north" minutes="12.012" orientation="latitude" pageId="3" pageNumber="456" precision="1" value="20.2002">20°12.012’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAED6F362768EAF885" box="[1287,1432,1871,1897]" degrees="40" direction="east" minutes="07.104" orientation="longitude" pageId="3" pageNumber="456" precision="1" value="40.1184">40°07.104’E</geoCoordinate>
, 
<specimenCode id="DBAF5435D43DFFBAE8FF361B6C69F862" box="[151,283,1907,1934]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAE94B361B6C32F862" box="[291,320,1907,1934]" collectionCode="RSS" pageId="3" pageNumber="456">58</specimenCode>
, reeftop, coral rubble, 
<date id="FFB7DA8ED43DFFBAEA2B361B6F94F861" box="[579,742,1907,1933]" pageId="3" pageNumber="456" value="2012-03-06">
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</date>
</materialsCitation>
; 
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<specimenCode id="DBAF5435D43DFFBAEA9B361B6E0EF862" box="[755,892,1907,1934]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="456">SMF 62258</specimenCode>
, 
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(TL 
<quantity id="4CF151ABD43DFFBAEC73361B69EBF861" box="[1051,1177,1907,1933]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.7" metricValueMin="1.9" pageId="3" pageNumber="456" unit="mm" value="23.0" valueMax="27.0" valueMin="19.0">19–27 mm</quantity>
), 
<specimenCount id="9D0F37C7D43DFFBAECC6361B6869F861" box="[1198,1307,1907,1934]" count="5" pageId="3" pageNumber="456" type="female">5 females</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAED3E361B6DB5F85D" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="3.6" metricValueMin="1.6" pageId="3" pageNumber="456" unit="mm" value="26.0" valueMax="36.0" valueMin="16.0">16–36 mm</quantity>
), 
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, 
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</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBAEA5C36FF6CBBF839" collectingDate="2012-02-26" collectionCode="RSS" collectorName="A. Brosing" latitude="16.734867" longLatPrecision="1" longitude="41.74465" pageId="3" pageNumber="456" specimenCode="RSS1-2012, 36" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43DFFBAEA5C36FF6FBBF85D" box="[564,713,1943,1969]" degrees="16" direction="north" minutes="44.092" orientation="latitude" pageId="3" pageNumber="456" precision="1" value="16.734867">16°44.092’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEABE36FF6E14F85D" box="[726,870,1943,1969]" degrees="41" direction="east" minutes="44.679" orientation="longitude" pageId="3" pageNumber="456" precision="1" value="41.74465">41°44.679’E</geoCoordinate>
, 0.5–1.0 m, under stones, 
<specimenCode id="DBAF5435D43DFFBAECFB36FF6865F85D" box="[1171,1303,1943,1969]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43DFFBAED7736FF684EF85D" box="[1311,1340,1943,1969]" collectionCode="RSS" pageId="3" pageNumber="456">36</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAEDE936FF6D80F839" pageId="3" pageNumber="456">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43DFFBAE89636D36CBBF839" box="[254,457,1979,2005]" pageId="3" pageNumber="456" value="2012-02-26">
<collectingDate id="EFF32366D43DFFBAE89636D36CBBF839" box="[254,457,1979,2005]" pageId="3" pageNumber="456" value="2012-02-26">26 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43DFFBAE9BB36D36C44F815" collectingDate="2011-04-26" collectionCode="SMF, RSS" collectorName="A. Brosing" location="Jeddah" pageId="3" pageNumber="456" specimenCode="SMF 62259, RSS1-2011" specimenCount="2" specimenCount-female="2" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43DFFBAE9BB36D36F29F83A" box="[467,603,1979,2006]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="456">SMF 62259</specimenCode>
, 
<specimenCount id="9D0F37C7D43DFFBAEA0E36D36FA0F839" box="[614,722,1979,2005]" count="2" pageId="3" pageNumber="456" type="female">2 females</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEB6236D36EF5F839" box="[778,903,1979,2005]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="3" pageNumber="456" unit="mm" value="23.0" valueMax="24.0" valueMin="22.0">22–24 mm</quantity>
), north of 
<location id="8ED6AA95D43DFFBAEB9236D36939F839" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAEB9236D36939F839" box="[1018,1099,1979,2005]" name="Jeddah" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Jeddah</location>
, 
<specimenCode id="DBAF5435D43DFFBAEC3E36D369ABF839" box="[1110,1241,1979,2005]" collectionCode="RSS" pageId="3" pageNumber="456">RSS1-2011</specimenCode>
, coll. 
<collectorName id="26FC9998D43DFFBAED7236D368EAF839" box="[1306,1432,1979,2005]" pageId="3" pageNumber="456">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43DFFBAE8FF36B76C44F815" box="[151,310,2015,2041]" pageId="3" pageNumber="456" value="2011-04-26">
<collectingDate id="EFF32366D43DFFBAE8FF36B76C44F815" box="[151,310,2015,2041]" pageId="3" pageNumber="456" value="2011-04-26">26 April 2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43DFFBAE92A36B76EEEF815" box="[322,924,2015,2042]" collectionCode="KAUMM" location="Farasan Islands" pageId="3" pageNumber="456" specimenCode="KAUMM 1062" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43DFFBAE92A36B76C81F815" box="[322,499,2015,2041]" collectionCode="KAUMM" pageId="3" pageNumber="456">KAUMM 1062</specimenCode>
, 
<specimenCount id="9D0F37C7D43DFFBAE99736B76F38F815" box="[511,586,2015,2041]" count="1" pageId="3" pageNumber="456" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43DFFBAEAEC36B76FAAF815" box="[644,728,2015,2041]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="456" unit="mm" value="29.0">29 mm</quantity>
), 
<location id="8ED6AA95D43DFFBAEA8336886EEEF815" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43DFFBAEA8336886EEEF815" box="[747,924,2015,2042]" name="Farasan Islands" pageId="3" pageNumber="456" stateProvince="Al Bahr al Ahmar">Farasan Islands</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43DFFBDEBC036B7696EFF5D" collectingDate="2012-02-24" collectionCode="RSS" collectorName="A. Brosing" lastPageId="4" lastPageNumber="457" latitude="16.755249" longLatPrecision="1" longitude="42.061565" pageId="3" pageNumber="456" specimenCode="RSS1-2012, 27" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43DFFBAEBC036B7694EF816" box="[936,1084,2015,2042]" degrees="16" direction="north" minutes="45.315" orientation="latitude" pageId="3" pageNumber="456" precision="1" value="16.755249">16°45.315’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43DFFBAEC2036B769ABF815" box="[1096,1241,2015,2041]" degrees="42" direction="east" minutes="03.694" orientation="longitude" pageId="3" pageNumber="456" precision="1" value="42.061565">42°03.694’E</geoCoordinate>
, low tide in front of mangroves, hand dredge, 
<specimenCode id="DBAF5435D43AFFBDE9B131FF6F2FFF5D" box="[473,605,151,177]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDEA0D31FF6FF0FF5D" box="[613,642,151,177]" collectionCode="RSS" pageId="4" pageNumber="457">27</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDEAAD31FF6E31FF5D" box="[709,835,151,177]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEB3831FF696EFF5D" box="[848,1052,151,177]" pageId="4" pageNumber="457" value="2012-02-24">
<collectingDate id="EFF32366D43AFFBDEB3831FF696EFF5D" box="[848,1052,151,177]" pageId="4" pageNumber="457" value="2012-02-24">24 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDEC4031FF6F6BFF39" collectionCode="KAUMM" location="Farasan Islands" municipality="Sasu Island" pageId="4" pageNumber="457" specimenCode="KAUMM 1063" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDEC4031FF69ABFF5D" box="[1064,1241,151,177]" collectionCode="KAUMM" pageId="4" pageNumber="457">KAUMM 1063</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEC8C31FF6834FF5D" box="[1252,1350,151,177]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEDE831FF6DB5FF39" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="457" unit="mm" value="20.0">20 mm</quantity>
), 
<collectingMunicipality id="6BD26634D43AFFBDE8B231D36C2FFF39" box="[218,349,187,213]" pageId="4" pageNumber="457">Sasu Island</collectingMunicipality>
, 
<location id="8ED6AA95D43AFFBDE90131D36F6BFF39" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDE90131D36F6BFF39" box="[361,537,187,213]" municipality="Sasu Island" name="Farasan Islands" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Farasan Islands</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDEA4C31D26F50FF15" collectingDate="2012-02-21" collectionCode="RSS" collectorName="A. Brosing" latitude="16.870184" longLatPrecision="1" longitude="41.592415" pageId="4" pageNumber="457" specimenCode="RSS1-2012, 18" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDEA4C31D26FC5FF39" box="[548,695,186,213]" degrees="16" direction="north" minutes="52.211" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="16.870184">16°52.211’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDEAAB31D26E27FF39" box="[707,853,186,213]" degrees="41" direction="east" minutes="35.545" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="41.592415">41°35.545’E</geoCoordinate>
, 0.5–2.0 m, bench with coral heads, 
<specimenCode id="DBAF5435D43AFFBDEC8731D36807FF39" box="[1263,1397,187,213]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDED1531D368EAFF39" box="[1405,1432,187,213]" collectionCode="RSS" pageId="4" pageNumber="457">18</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDE8A631B76C39FF15" box="[206,331,223,249]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDE93F31B76F50FF15" box="[343,546,223,249]" pageId="4" pageNumber="457" value="2012-02-21">
<collectingDate id="EFF32366D43AFFBDE93F31B76F50FF15" box="[343,546,223,249]" pageId="4" pageNumber="457" value="2012-02-21">21 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDEA4531B76929FF15" box="[557,1115,223,249]" collectionCode="SMF" location="Farasan Islands" pageId="4" pageNumber="457" specimenCode="SMF 62260" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDEA4531B76FC7FF15" box="[557,693,223,249]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="457">SMF 62260</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEAA931B76E7EFF15" box="[705,780,223,249]" count="1" pageId="4" pageNumber="457" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEB2D31B76EEAFF15" box="[837,920,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="457" unit="mm" value="23.0">23 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDEBC331B76929FF15" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEBC331B76929FF15" box="[939,1115,223,249]" name="Farasan Islands" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Farasan Islands</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDEC0F31B769FEFEF1" collectingDate="2012-02-23" collectionCode="RSS" collectorName="A. Brosing" latitude="16.718834" longLatPrecision="1" longitude="42.065033" pageId="4" pageNumber="457" specimenCode="RSS1-2012, 10" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDEC0F31B76989FF15" box="[1127,1275,223,249]" degrees="16" direction="north" minutes="43.130" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="16.718834">16°43.130’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDED6F31B768EAFF15" box="[1287,1432,223,249]" degrees="42" direction="east" minutes="03.902" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="42.065033">42°03.902’E</geoCoordinate>
, 0.5–1.0 m, beach with dead corals, 
<specimenCode id="DBAF5435D43AFFBDEA40306B6FDCFEF1" box="[552,686,259,285]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDEADF306B6FA1FEF1" box="[695,723,259,285]" collectionCode="RSS" pageId="4" pageNumber="457">10</specimenCode>
(2), coll. 
<collectorName id="26FC9998D43AFFBDEB5D306B6EC1FEF1" box="[821,947,259,285]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEBA8306B69FEFEF1" box="[960,1164,259,285]" pageId="4" pageNumber="457" value="2012-02-23">
<collectingDate id="EFF32366D43AFFBDEBA8306B69FEFEF1" box="[960,1164,259,285]" pageId="4" pageNumber="457" value="2012-02-23">23 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDECF0306B6CB8FEAD" collectionCode="SMF" location="Al Lith" pageId="4" pageNumber="457" specimenCode="SMF 62261" specimenCount="1" specimenCount-juvenile="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDECF0306B6853FEF1" box="[1176,1313,259,285]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="457">SMF 62261</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDED44306B68EEFEF1" box="[1324,1436,259,285]" count="1" pageId="4" pageNumber="457" type="juvenile">1 juvenile</specimenCount>
male (TL 
<quantity id="4CF151ABD43AFFBDE963304F6C12FEAD" box="[267,352,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="457" unit="mm" value="12.0">12 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDE91B304F6CB8FEAD" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDE91B304F6CB8FEAD" box="[371,458,295,321]" name="Al Lith" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Al Lith</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDE9B0304F68E5FEAD" box="[472,1431,294,321]" collectingDate="2011-03-29" collectionCode="RSS" collectorName="A. Brosing" latitude="20.266716" longLatPrecision="1" longitude="39.990715" pageId="4" pageNumber="457" specimenCode="RSS1-2011, 01" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDE9B0304F6F1EFEAD" box="[472,620,295,321]" degrees="20" direction="north" minutes="16.003" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="20.266716">20°16.003’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDEA12304E6E79FEAD" box="[634,779,294,321]" degrees="39" direction="east" minutes="59.443" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="39.990715">39°59.443’E</geoCoordinate>
, beach, 
<specimenCode id="DBAF5435D43AFFBDEB02304F6E9CFEAD" box="[874,1006,295,321]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2011</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDEB9E304F6960FEAD" box="[1014,1042,295,321]" collectionCode="RSS" pageId="4" pageNumber="457">01</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDEC30304F69AAFEAD" box="[1112,1240,295,321]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEC8E304F68E5FEAD" box="[1254,1431,295,321]" pageId="4" pageNumber="457" value="2011-03-29">
<collectingDate id="EFF32366D43AFFBDEC8E304F68E5FEAD" box="[1254,1431,295,321]" pageId="4" pageNumber="457" value="2011-03-29">29 March 2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDE8FF30236EB1FE89" box="[151,963,331,357]" collectionCode="KAUMM" location="Farasan Islands" municipality="Khotib Harbour" pageId="4" pageNumber="457" specimenCode="KAUMM 1064" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDE8FF30236C38FE89" box="[151,330,331,357]" collectionCode="KAUMM" pageId="4" pageNumber="457">KAUMM 1064</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDE93F30236CD7FE89" box="[343,421,331,357]" count="1" pageId="4" pageNumber="457" type="male">1 male</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDE98A30236F4AFE89" box="[482,568,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="457" unit="mm" value="20.0">20 mm</quantity>
), 
<collectingMunicipality id="6BD26634D43AFFBDEA2530236E71FE89" box="[589,771,331,357]" pageId="4" pageNumber="457">Khotib Harbour</collectingMunicipality>
, 
<location id="8ED6AA95D43AFFBDEB7930236EB1FE89" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEB7930236EB1FE89" box="[785,963,331,357]" municipality="Khotib Harbour" name="Farasan Islands" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Farasan Islands</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDEBB930226CE7FE65" collectingDate=" 21 February 2012" collectionCode="RSS" latitude="16.9152" longLatPrecision="1" longitude="41.842968" pageId="4" pageNumber="457" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDEBB930226917FE89" box="[977,1125,330,357]" degrees="16" direction="north" minutes="54.912" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="16.9152">16°54.912’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDEC1B30226876FE89" box="[1139,1284,330,357]" degrees="41" direction="east" minutes="50.578" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="41.842968">41°50.578’E</geoCoordinate>
, 
<collectionCode id="ED18648BD43AFFBDED7A30236834FE89" box="[1298,1350,331,357]" pageId="4" pageNumber="457">RSS</collectionCode>
1-2012- 14, 
<date id="FFB7DA8ED43AFFBDE8AC30076CE7FE65" box="[196,405,367,393]" pageId="4" pageNumber="457" value="2012-02-21">
<collectingDate id="EFF32366D43AFFBDE8AC30076CE7FE65" box="[196,405,367,393]" isEnumeration="true" pageId="4" pageNumber="457">21 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDE9CB30076987FE65" box="[419,1269,367,393]" collectionCode="SMF" location="Farasan Islands" municipality="Aboshok Island" pageId="4" pageNumber="457" specimenCode="SMF 62262" specimenCount="3" specimenCount-female="3" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDE9CB30076F5CFE65" box="[419,558,367,393]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="457">SMF 62262</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEA5430076FD9FE65" box="[572,683,367,393]" count="3" pageId="4" pageNumber="457" type="female">3 females</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEA8130076E1BFE65" box="[745,873,367,393]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="3.0" metricValueMin="1.9" pageId="4" pageNumber="457" unit="mm" value="24.5" valueMax="30.0" valueMin="19.0">19–30 mm</quantity>
), 
<collectingMunicipality id="6BD26634D43AFFBDEB1630076946FE65" box="[894,1076,367,393]" pageId="4" pageNumber="457">Aboshok Island</collectingMunicipality>
, 
<location id="8ED6AA95D43AFFBDEC2A30076987FE65" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEC2A30076987FE65" box="[1090,1269,367,393]" municipality="Aboshok Island" name="Farasan Islands" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Farasan Islands</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDED6B30066C33FE3D" collectingDate="2012-02-27" collectionCode="V" collectorName="A. Brosing &amp; Spiridonov" elevation="9" latitude="16.986" longLatPrecision="1" longitude="41.79093" pageId="4" pageNumber="457" specimenCode="RSS1-2012, 38" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDED6B300668EAFE65" box="[1283,1432,366,393]" degrees="16" direction="north" minutes="59.160" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="16.986">16°59.160’N</geoCoordinate>
, 
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, 
<quantity id="4CF151ABD43AFFBDE95D30FB6CDBFE41" box="[309,425,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="4" pageNumber="457" unit="m" value="0.85" valueMax="0.9" valueMin="0.8">
<elevation id="00241B7DD43AFFBDE95D30FB6CDBFE41" box="[309,425,403,429]" metricMagnitude="-1" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="4" pageNumber="457" unit="m" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 m</elevation>
</quantity>
, limestone bench with brown algae, 
<specimenCode id="DBAF5435D43AFFBDEB2130FB6EBFFE41" box="[841,973,403,429]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDEBBD30FB6E80FE41" box="[981,1010,403,429]" collectionCode="RSS" pageId="4" pageNumber="457">38</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDEC5E30FB69C0FE41" box="[1078,1202,403,429]" pageId="4" pageNumber="457">A. Brösing</collectorName>
&amp; 
<collectionCode id="ED18648BD43AFFBDECBF30FB6999FE41" box="[1239,1259,403,429]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="457" type="Museum">V</collectionCode>
. 
<collectorName id="26FC9998D43AFFBDEC9F30FB6807FE41" box="[1271,1397,403,429]" pageId="4" pageNumber="457">Spiridonov</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEDE830FB6C33FE3D" pageId="4" pageNumber="457" value="2012-02-27">
<collectingDate id="EFF32366D43AFFBDEDE830FB6C33FE3D" pageId="4" pageNumber="457" value="2012-02-27">27 February 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDE92630DF6FCDFE19" collectingDate="2012-03-04" collectionCode="TL" collectorName="A. Brosing" elevation="5" location="Al Lith" longitude="39.9916" pageId="4" pageNumber="457" specimenCode="SMF 62263, RSS1-2012, 49" specimenCount="1" specimenCount-male="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDE92630DF6CAAFE3D" box="[334,472,439,465]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="457">SMF 62263</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDE98D30DF6F40FE3D" box="[485,562,439,465]" count="1" pageId="4" pageNumber="457" type="male">1 male</specimenCount>
(
<collectionCode id="ED18648BD43AFFBDEA2B30DF6F14FE3D" box="[579,614,439,465]" pageId="4" pageNumber="457">TL</collectionCode>
<quantity id="4CF151ABD43AFFBDEA0730DF6FB6FE3D" box="[623,708,439,465]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="457" unit="mm" value="19.0">19 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDEABF30DF6E5FFE3D" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEABF30DF6E5FFE3D" box="[727,813,439,465]" longitude="39.9916" name="Al Lith" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Al Lith</location>
, 
<geoCoordinate id="EE3D9A89D43AFFBDEB5330DE6ECDFE3D" box="[827,959,438,465]" pageId="4" pageNumber="457">20°15.922’,</geoCoordinate>
<geoCoordinate id="EE3D9A89D43AFFBDEBA030DE692BFE3D" box="[968,1113,438,465]" degrees="39" direction="east" minutes="59.496" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="39.9916">39°59.496’E</geoCoordinate>
, 
<quantity id="4CF151ABD43AFFBDEC0F30DE69D9FE3D" box="[1127,1195,438,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="457" unit="m" value="0.5">
<elevation id="00241B7DD43AFFBDEC0F30DE69D9FE3D" box="[1127,1195,438,465]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="457" unit="m" value="0.5">0.5 m</elevation>
</quantity>
, beach, hand dredge, 
<specimenCode id="DBAF5435D43AFFBDE8FF30B36C69FE19" box="[151,283,475,501]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDE94A30B36C32FE19" box="[290,320,475,501]" collectionCode="RSS" pageId="4" pageNumber="457">49</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDE9E130B36F7BFE19" box="[393,521,475,501]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEA7030B36FCDFE19" box="[536,703,475,501]" pageId="4" pageNumber="457" value="2012-03-04">
<collectingDate id="EFF32366D43AFFBDEA7030B36FCDFE19" box="[536,703,475,501]" pageId="4" pageNumber="457" value="2012-03-04">4 March 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDEAA530B36987FE19" box="[717,1269,474,501]" collectionCode="KAUMM" location="Al Lith" pageId="4" pageNumber="457" specimenCode="KAUMM 1065" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDEAA530B36EF3FE19" box="[717,897,474,501]" collectionCode="KAUMM" pageId="4" pageNumber="457">KAUMM 1065</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEBE730B36E81FE19" box="[911,1011,475,501]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEC5B30B369FBFE19" box="[1075,1161,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="457" unit="mm" value="20.0">20 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDECF530B36987FE19" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDECF530B36987FE19" box="[1181,1269,475,501]" name="Al Lith" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Al Lith</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDED6B30B26C8CFDD1" collectingDate="2012-03-06" collectionCode="RSS" collectorName="A. Grishchenko" elevation="3" latitude="20.25385" longLatPrecision="1" longitude="39.9892" pageId="4" pageNumber="457" specimenCode="RSS1-2012, 53" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDED6B30B268E5FE19" box="[1283,1431,474,501]" degrees="20" direction="north" minutes="15.231" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="20.25385">20°15.231’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDE8FF30966C5AFDF5" box="[151,296,510,537]" degrees="39" direction="east" minutes="59.352" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="39.9892">39°59.352’E</geoCoordinate>
, 
<quantity id="4CF151ABD43AFFBDE95D30976CDBFDF5" box="[309,425,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="457" unit="m" value="0.3" valueMax="0.4" valueMin="0.2">
<elevation id="00241B7DD43AFFBDE95D30976CDBFDF5" box="[309,425,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="457" unit="m" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 m</elevation>
</quantity>
, 
<collectingMethod id="52488459D43AFFBDE9DE30976FA8FDF5" box="[438,730,511,537]" pageId="4" pageNumber="457">hand dredge and hand net</collectingMethod>
in debris of seagrass and algae, reef flat, 
<specimenCode id="DBAF5435D43AFFBDECDE30966848FDF5" box="[1206,1338,510,537]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDED2A3096682DFDF5" box="[1346,1375,510,537]" collectionCode="RSS" pageId="4" pageNumber="457">53</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDE8FF334B6C3FFDD1" box="[151,333,547,573]" pageId="4" pageNumber="457">A. Grishchenko</collectorName>
, 
<date id="FFB7DA8ED43AFFBDE932334B6C8CFDD1" box="[346,510,547,573]" pageId="4" pageNumber="457" value="2012-03-06">
<collectingDate id="EFF32366D43AFFBDE932334B6C8CFDD1" box="[346,510,547,573]" pageId="4" pageNumber="457" value="2012-03-06">6 March 2012</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDEA62334B6C66FD8D" collectingDate="2011-04-17" collectionCode="KAUMM, RSS" collectorName="A. Brosing" location="Jeddah" pageId="4" pageNumber="457" specimenCode="KAUMM 1066, RSS1-2011" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDEA62334B6FCEFDD1" box="[522,700,547,573]" collectionCode="KAUMM" pageId="4" pageNumber="457">KAUMM 1066</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEAA0334B6E59FDD1" box="[712,811,547,573]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEB0F334B6EC9FDD1" box="[871,955,547,573]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="4" pageNumber="457" unit="mm" value="24.0">24 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDEBB8334B6953FDD1" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEBB8334B6953FDD1" box="[976,1057,547,573]" name="Jeddah" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Jeddah</location>
, 
<specimenCode id="DBAF5435D43AFFBDEC45334B69DDFDD1" box="[1069,1199,547,573]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2011</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDEC9C334B6801FDD1" box="[1268,1395,547,573]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEDE8334B6C66FD8D" pageId="4" pageNumber="457" value="2011-04-17">
<collectingDate id="EFF32366D43AFFBDEDE8334B6C66FD8D" pageId="4" pageNumber="457" value="2011-04-17">17 April 2011</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B61F613D43AFFBDE948332F6E74FD8D" box="[288,774,583,609]" collectionCode="SMF" location="Al Lith" pageId="4" pageNumber="457" specimenCode="SMF 62264" specimenCount="1" specimenCount-female="1" stateProvince="Al Bahr al Ahmar">
<specimenCode id="DBAF5435D43AFFBDE948332F6CDAFD8D" box="[288,424,583,609]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="4" pageNumber="457">SMF 62264</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDE9DB332F6F67FD8D" box="[435,533,583,609]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEA26332F6FD2FD8D" box="[590,672,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="457" unit="mm" value="11.0">11 mm</quantity>
), 
<location id="8ED6AA95D43AFFBDEADA332F6E74FD8D" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEADA332F6E74FD8D" box="[690,774,583,609]" name="Al Lith" pageId="4" pageNumber="457" stateProvince="Al Bahr al Ahmar">Al Lith</location>
</materialsCitation>
, 
<materialsCitation id="3B61F613D43AFFBDEB7A332F6924FD69" collectingDate="2012-03-06" collectionCode="V" collectorName="A. Brosing &amp; S. Trankner &amp; Spiridonov &amp; A. Mal" latitude="20.236534" longLatPrecision="1" longitude="40.06235" pageId="4" pageNumber="457" specimenCode="RSS1-2012, 57" specimenCount="1" stateProvince="Al Bahr al Ahmar">
<geoCoordinate id="EE3D9A89D43AFFBDEB7A332F6ED5FD8D" box="[786,935,583,609]" degrees="20" direction="north" minutes="14.192" orientation="latitude" pageId="4" pageNumber="457" precision="1" value="20.236534">20°14.192’N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDEBDA332F6931FD8D" box="[946,1091,583,609]" degrees="40" direction="east" minutes="03.741" orientation="longitude" pageId="4" pageNumber="457" precision="1" value="40.06235">40°03.741’E</geoCoordinate>
, mangroves, algae &amp; seagrass, 
<specimenCode id="DBAF5435D43AFFBDE8FF33026C69FD69" box="[151,283,618,645]" collectionCode="RSS" pageId="4" pageNumber="457">RSS1-2012</specimenCode>
- 
<specimenCode id="DBAF5435D43AFFBDE94B33026C32FD69" box="[291,320,618,645]" collectionCode="RSS" pageId="4" pageNumber="457">57</specimenCode>
, coll. 
<collectorName id="26FC9998D43AFFBDE9EA33036C8DFD69" box="[386,511,619,645]" pageId="4" pageNumber="457">A. Brösing</collectorName>
, 
<collectorName id="26FC9998D43AFFBDEA6333036FE2FD69" box="[523,656,619,645]" pageId="4" pageNumber="457">S. Tränkner</collectorName>
, 
<collectionCode id="ED18648BD43AFFBDEAF233036FDCFD69" box="[666,686,619,645]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="457" type="Museum">V</collectionCode>
. 
<collectorName id="26FC9998D43AFFBDEAD133036E47FD69" box="[697,821,619,645]" pageId="4" pageNumber="457">Spiridonov</collectorName>
&amp; 
<collectorName id="26FC9998D43AFFBDEB3F33036ED8FD69" box="[855,938,619,645]" pageId="4" pageNumber="457">A. Mal</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEBDD33036924FD69" box="[949,1110,619,645]" pageId="4" pageNumber="457" value="2012-03-06">
<collectingDate id="EFF32366D43AFFBDEBDD33036924FD69" box="[949,1110,619,645]" pageId="4" pageNumber="457" value="2012-03-06">6 March 2012</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF33E76DA5FD1D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">
<materialsCitation id="3B61F613D43AFFBDE8AF33E76DA1FD1D" collectingDate="2012-09-26" collectionCode="GULF, OF, ADEN, FLMNH" collectorName="G. Paulay" country="Djibouti" elevation="1" latitude="11.6012" location="Sheraton Hotel" longLatPrecision="54" longitude="43.159" pageId="4" pageNumber="457" specimenCode="UF32746" specimenCount="1" specimenCount-female="1">
<collectionCode id="ED18648BD43AFFBDE8AF33E76CE1FD45" box="[199,403,655,681]" pageId="4" pageNumber="457">GULF OF ADEN</collectionCode>
: 
<collectionCode id="ED18648BD43AFFBDE9F633E76F72FD45" box="[414,512,655,681]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="4" pageNumber="457" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBAF5435D43AFFBDEA6F33E76F06FD45" box="[519,628,655,681]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="4" pageNumber="457" type="Museum">UF32746</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEA1733E76F92FD45" box="[639,736,655,681]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEB7133E76E1FFD45" box="[793,877,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="457" unit="mm" value="26.0">26 mm</quantity>
), 
<collectingCountry id="F31EBCDED43AFFBDEBE833E76E92FD45" box="[896,992,655,681]" name="Djibouti" pageId="4" pageNumber="457">Djibouti</collectingCountry>
, in front of 
<location id="8ED6AA95D43AFFBDEC0D33E76862FD45" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEC0D33E76862FD45" box="[1125,1296,655,681]" country="Djibouti" latitude="11.6012" longLatPrecision="54" longitude="43.159" name="Sheraton Hotel" pageId="4" pageNumber="457">Sheraton Hotel</location>
, 
<geoCoordinate id="EE3D9A89D43AFFBDED7433E768EAFD45" box="[1308,1432,655,681]" degrees="11.6012" direction="north" orientation="latitude" pageId="4" pageNumber="457" precision="5" value="11.6012">11.6012°N</geoCoordinate>
, 
<geoCoordinate id="EE3D9A89D43AFFBDE8FF33DA6C76FD21" box="[151,260,690,717]" degrees="43.159" direction="east" orientation="longitude" pageId="4" pageNumber="457" precision="55" value="43.159">43.159°E</geoCoordinate>
, 
<quantity id="4CF151ABD43AFFBDE97A33DB6C2DFD21" box="[274,351,691,717]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="4" pageNumber="457" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">
<elevation id="00241B7DD43AFFBDE97A33DB6C2DFD21" box="[274,351,691,717]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="4" pageNumber="457" unit="m" value="1.0" valueMax="2.0" valueMin="0.0">0–2 m</elevation>
</quantity>
, fringing sand flat with some scuzz, BDJRS-0018, DJRS-001, coll. 
<collectorName id="26FC9998D43AFFBDEC1433DB699DFD21" box="[1148,1263,691,717]" pageId="4" pageNumber="457">G. Paulay</collectorName>
, 
<date id="FFB7DA8ED43AFFBDEC9633DB6DA1FD1D" pageId="4" pageNumber="457" value="2012-09-26">
<collectingDate id="EFF32366D43AFFBDEC9633DB6DA1FD1D" pageId="4" pageNumber="457" value="2012-09-26">26 September 2012</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF33936C26FCB1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">
<materialsCitation id="3B61F613D43AFFBDE8AF33936C3DFCB1" collectingDate="1963-11-27" collectionCode="USNM" collectorName="International Indian Ocean Expedition Sta. &amp; L. P. Woods" country="Pakistan" elevation="1" location="Astola Island" municipality="Karachi" pageId="4" pageNumber="457" specimenCode="USNM 120473" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F31EBCDED43AFFBDE8AF33936C23FCF9" box="[199,337,763,789]" name="Pakistan" pageId="4" pageNumber="457">PAKISTAN</collectingCountry>
: 
<specimenCode id="DBAF5435D43AFFBDE93433936F79FCF9" box="[348,523,763,789]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="457" type="Museum">USNM 120473</specimenCode>
, 
<specimenCount id="9D0F37C7D43AFFBDEA7E33936F0AFCF9" box="[534,632,763,789]" count="1" pageId="4" pageNumber="457" type="female">1 female</specimenCount>
(TL 
<quantity id="4CF151ABD43AFFBDEAD933936E76FCF9" box="[689,772,763,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="457" unit="mm" value="22.0">22 mm</quantity>
), western end 
<location id="8ED6AA95D43AFFBDEBCC33936949FCF9" LSID="urn:lsid:plazi:treatment:03A04D58D43FFFBEE8FF37166F75FD45:8ED6AA95D43AFFBDEBCC33936949FCF9" box="[932,1083,763,789]" country="Pakistan" municipality="Karachi" name="Astola Island" pageId="4" pageNumber="457">Astola Island</location>
, ca. 
<quantity id="4CF151ABD43AFFBDEC06339369AEFCF9" box="[1134,1244,763,789]" metricMagnitude="5" metricUnit="m" metricValue="2.84853888" pageId="4" pageNumber="457" unit="mi" value="177.0">177 miles</quantity>
west of 
<collectingMunicipality id="6BD26634D43AFFBDED55339368EBFCF9" box="[1341,1433,763,789]" pageId="4" pageNumber="457">Karachi</collectingMunicipality>
, 
<quantity id="4CF151ABD43AFFBDE8FF32776D84FCD5" box="[151,246,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.2" metricValueMax="2.4" metricValueMin="0.0" pageId="4" pageNumber="457" unit="m" value="1.2" valueMax="2.4" valueMin="0.0">
<elevation id="00241B7DD43AFFBDE8FF32776D84FCD5" box="[151,246,799,825]" metricMagnitude="0" metricUnit="m" metricValue="1.2" metricValueMax="2.4" metricValueMin="0.0" pageId="4" pageNumber="457" unit="m" value="1.2" valueMax="2.4" valueMin="0.0">0–2.4 m</elevation>
</quantity>
, rocks, sand, scattered coral, 
<collectorName id="26FC9998D43AFFBDEA2B3277695AFCD5" box="[579,1064,799,825]" pageId="4" pageNumber="457">International Indian Ocean Expedition Sta.</collectorName>
LW-l, coll. 
<collectorName id="26FC9998D43AFFBDECD832776842FCD5" box="[1200,1328,799,825]" pageId="4" pageNumber="457">L.P. Woods</collectorName>
<emphasis id="B97D205CD43AFFBDED5032776818FCD5" box="[1336,1386,799,825]" italics="true" pageId="4" pageNumber="457">et al</emphasis>
., 
<date id="FFB7DA8ED43AFFBDEDE832776C3DFCB1" pageId="4" pageNumber="457" value="1963-11-27">
<collectingDate id="EFF32366D43AFFBDEDE832776C3DFCB1" pageId="4" pageNumber="457" value="1963-11-27">27 November 1963</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D43AFFBDE8AF320E6978FAE1" pageId="4" pageNumber="457" type="diagnosis">
<paragraph id="8BB6FC4ED43AFFBDE8AF320E6978FAE1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">
<emphasis id="B97D205CD43AFFBDE8AF320E6C48FC6C" bold="true" box="[199,314,870,896]" pageId="4" pageNumber="457">Diagnosis</emphasis>
. Ocular scales rounded, separate. Rostral plate basal portion with straight anterior margins, transverse to sloping posteriorly; anterolateral angles rounded. Raptorial claw dactylus with shallow notch on outer proximal margin. Mandibular palp 3-segmented. Telson intermediate teeth distinct, apices sharp, extending posteriorly well beyond apices of intermediate denticles; lateral teeth projecting well off margin of telson by wide notch (in specimens&gt; TL 
<quantity id="4CF151ABD43AFFBDE93F329F6CA4FBFD" box="[343,470,1015,1041]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="4" pageNumber="457" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
). Telson median and accessory median carinae together with group of usually 5 or 6 posterior spines; anterior submedian carina with 1–3 (usually 2) spines; submedian tooth with proximal cluster of 1–3 (usually 2 or 3) spines proximally; intermediate tooth with 1 dorsal spine preceded by 1–3 (usually 2) spines on anterior intermediate carina; ventral surface of submedian and intermediate teeth flat, without longitudinal carina along midline. Uropodal protopod with low obtuse swelling behind dorsal carina. Uropodal exopod proximal article inner margin smooth, glabrous; distal margin with ventral spine; exopod distal article with outer margin setose, inner margin smooth, glabrous. Uropodal endopod about 3 times as long as wide; distal half of outer margin setose, inner margin smooth, glabrous except for 3–11 (usually 4 or 5) proximal setae.
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D43AFFBCE8AF347E6C84FE3D" lastPageId="5" lastPageNumber="458" pageId="4" pageNumber="457" type="description">
<paragraph id="8BB6FC4ED43AFFBDE8AF347E6EF4FAB9" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">
<emphasis id="B97D205CD43AFFBDE8AF347E6C23FADC" bold="true" box="[199,337,1302,1328]" pageId="4" pageNumber="457">Description</emphasis>
. Eyes elongate; cornea subconical. Ocular scales rounded, separate, bases transverse. Antennular peduncle length 0.61–0.83CL. Antennal scale length 0.45–0.50CL.
</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF34376FDCFA2D" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">Rostral plate longer than wide; basal portion with straight anterior margins, transverse to sloping posteriorly; anterolateral angles rounded; lateral margins divergent anteriorly; median spine longer than base, laterally compressed, with obtusely angular ventral keel.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF34A3690EF9E5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">Raptorial claw dactylus with shallow notch on outer proximal margin. Propodus proximal movable spine slender; opposable margin sparsely pectinate proximally. Carpus dorsal margin unarmed.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF377B6ED8F9C1" blockId="4.[151,1437,151,2013]" box="[199,938,1554,1581]" pageId="4" pageNumber="457">Mandibular palp 3-segmented. Maxillipeds 1–5 each with epipod.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF375F6FA5F999" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">Thoracic somite 6 lateral process subequal to that of thoracic somite 7; lower margins subtruncate. TS8 anterolateral margin rounded; sternal keel obsolete.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF37176934F975" blockId="4.[151,1437,151,2013]" box="[199,1094,1663,1689]" pageId="4" pageNumber="457">Pereopods 1–3 endopod slender, linear, 2-segmented; proximal article unarmed.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF37CB6942F951" blockId="4.[151,1437,151,2013]" box="[199,1072,1699,1725]" pageId="4" pageNumber="457">Male pleopod 1 endopod with well-developed lateral lobe on posterior endite.</paragraph>
<paragraph id="8BB6FC4ED43AFFBDE8AF37AF6C6DF8C5" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="457">Abdominal somites 1–5 posterolateral angles unarmed. AWCLI 792–875. Abdominal somite 6 with posterior spine of submedian, intermediate and lateral carinae distinct in juveniles and females, often obscure or obsolete in adult males.</paragraph>
<paragraph id="8BB6FC4ED43AFFBCE8AF365B6837FF15" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="458" pageId="4" pageNumber="457">
Telson wider than long; with 7–12 (usually 8 or 9) spiniform submedian denticles; intermediate teeth distinct, apices sharp, extending posteriorly well beyond apices of intermediate denticles; lateral teeth demarcated by wide notch in specimens&gt; TL 
<quantity id="4CF151ABD43AFFBDE9C436136F5AF879" box="[428,552,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="4" pageNumber="457" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
), apex angular, projecting well off margin of telson. Telson median carina ovate, usually obscuring accessory median carina, together with a group of usually 5 or 6 posterior spines (1–3 [usually 1 or 2] spines on median, 0–3 [usually 2] spines on each accessory median); anterior submedian carina with 1–3 (usually 2) spines in longitudinal row; submedian tooth with proximal cluster of 1–3 (usually 2 or 3) spines proximally; intermediate tooth with 1 dorsal spine preceded by 1–3 (usually 2) spines on anterior intermediate carina; knob absent; ventral surface flat, without longitudinal carina along midline of submedian and intermediate teeth.
</paragraph>
<paragraph id="8BB6FC4ED43BFFBCE8AF306B6C84FE3D" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="458">Uropodal protopod terminal spines with outer slightly longer than inner; upper proximal surface with low obtuse swelling behind dorsal carina. Uropodal exopod proximal segment outer margin with 9–15 (usually 11) movable spines, distalmost slightly exceeding apex of distal segment; inner margin smooth, glabrous; distal margin with ventral spine; exopod distal segment with outer margin setose, inner margin smooth, glabrous. Uropodal endopod length 2.81–3.16 times width; distal half of outer margin setose, inner margin smooth, glabrous except for 3–11 (usually 4 or 5) proximal setae.</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D43BFFBCE8AF30B36FFBFDD1" pageId="5" pageNumber="458" type="etymology">
<paragraph id="8BB6FC4ED43BFFBCE8AF30B36FFBFDD1" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="458">
<emphasis id="B97D205CD43BFFBCE8AF30B36C3AFE19" bold="true" box="[199,328,475,501]" pageId="5" pageNumber="458">Etymology</emphasis>
. The specific epithet is derived from the Latin, 
<emphasis id="B97D205CD43BFFBCEBE130B36ECAFE19" box="[905,952,475,501]" italics="true" pageId="5" pageNumber="458">celo</emphasis>
(hide, conceal), and 
<emphasis id="B97D205CD43BFFBCECDD30B3699DFE18" box="[1205,1263,475,500]" italics="true" pageId="5" pageNumber="458">sinus</emphasis>
(bay, gulf), in recognition that the new species has been “hiding in plain sight” in the bays and gulfs of the Arabian Sea for more than a century; used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D43BFFBCE8AF332F6874FD45" pageId="5" pageNumber="458" type="description">
<paragraph id="8BB6FC4ED43BFFBCE8AF332F6874FD45" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="458">
<emphasis id="B97D205CD43BFFBCE8AF332F6C04FD8D" bold="true" box="[199,374,583,609]" pageId="5" pageNumber="458">Measurements</emphasis>
. Male (n = 17) TL 
<quantity id="4CF151ABD43BFFBCEA0C332F6F91FD8D" box="[612,739,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="3.6" metricValueMin="1.2" pageId="5" pageNumber="458" unit="mm" value="24.0" valueMax="36.0" valueMin="12.0">12–36 mm</quantity>
, female (n= 27) TL 
<quantity id="4CF151ABD43BFFBCEBBE332F6927FD8D" box="[982,1109,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="3.6" metricValueMin="1.1" pageId="5" pageNumber="458" unit="mm" value="23.5" valueMax="36.0" valueMin="11.0">11–36 mm</quantity>
. Other measurements of the 
<typeStatus id="54B242ECD43BFFBCE8FF33036D8FFD69" box="[151,253,619,645]" pageId="5" pageNumber="458" type="holotype">holotype</typeStatus>
: CL 
<quantity id="4CF151ABD43BFFBCE95C33026CE9FD69" box="[308,411,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.44" pageId="5" pageNumber="458" unit="mm" value="5.44">5.44 mm</quantity>
, antennular peduncle length 
<quantity id="4CF151ABD43BFFBCEA8933026E3AFD69" box="[737,840,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.59" pageId="5" pageNumber="458" unit="mm" value="3.59">3.59 mm</quantity>
, antennal scale length 
<quantity id="4CF151ABD43BFFBCEC22330369C3FD69" box="[1098,1201,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="5" pageNumber="458" unit="mm" value="2.69">2.69 mm</quantity>
, abdominal somite 5 width 
<quantity id="4CF151ABD43BFFBCE8B633E66C37FD45" box="[222,325,654,681]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="458" unit="mm" value="4.5">4.50 mm</quantity>
. 
<bibRefCitation id="EF9881BFD43BFFBCE93833E76F70FD45" author="Holthuis, L. B." box="[336,514,655,681]" pageId="5" pageNumber="458" pagination="1 - 45" refId="ref6299" refString="Holthuis, L. B. (1967) The stomatopod Crustacea collected by the 1962 and 1965 Israel South Red Sea Expeditions. The Second Israel South Red Sea Expedition, 1965, Report No. 1. Israel Journal of Zoology, 16, 1 - 45, figs. 1 - 7." type="journal article" year="1967">Holthuis (1967)</bibRefCitation>
recorded specimens from the 
<collectingRegion id="49CD32ACD43BFFBCEB3C33E76EC2FD45" box="[852,944,655,681]" country="Egypt" name="Al Bahr al Ahmar" pageId="5" pageNumber="458">Red Sea</collectingRegion>
to TL 
<quantity id="4CF151ABD43BFFBCEB9433E76939FD45" box="[1020,1099,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="458" unit="mm" value="40.0">40 mm</quantity>
(as 
<taxonomicName id="4C0987CDD43BFFBCEC1133E76987FD45" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1145,1269,655,681]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEC1133E76987FD45" box="[1145,1269,655,681]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D43BFFBFE8AF33DB699BFEF1" lastPageId="6" lastPageNumber="459" pageId="5" pageNumber="458" type="discussion">
<paragraph id="8BB6FC4ED43BFFBCE8AF33DB6F17F951" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="458">
<emphasis id="B97D205CD43BFFBCE8AF33DB6C46FD21" bold="true" box="[199,308,691,717]" pageId="5" pageNumber="458">Remarks</emphasis>
. 
<taxonomicName id="4C0987CDD43BFFBCE92C33DB6F1FFD21" authorityName="Ahyong" authorityYear="2024" box="[324,621,691,717]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCE92C33DB6F1FFD21" box="[324,621,691,717]" italics="true" pageId="5" pageNumber="458">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEA1F33DB6FA1FD21" bold="true" box="[631,723,691,717]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEA1F33DB6FA1FD21" box="[631,723,691,717]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has long been mistaken for 
<taxonomicName id="4C0987CDD43BFFBCEC7433DB69E9FD21" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1052,1179,691,717]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEC7433DB69E9FD21" box="[1052,1179,691,717]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
, probably because the latter species has never been adequately characterized or figured. To facilitate recognition of 
<taxonomicName id="4C0987CDD43BFFBCECBC33BF6821FD1D" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1236,1363,727,753]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCECBC33BF6821FD1D" box="[1236,1363,727,753]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D43BFFBCED3633BF6D92FCF9" pageId="5" pageNumber="458" sensu="stricto">sensu stricto</taxonomicNameLabel>
, figures are provided of material from the Gulf of Mannar (
<figureCitation id="1332E0CBD43BFFBCEB1233936ECDFCF9" box="[890,959,763,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1856]" captionTargetBox="[210,1375,315,1805]" captionTargetId="figure-95@6.[209,1378,309,1809]" captionTargetPageId="6" captionText="FIGURE 3. Gonodactylellus demanii (Henderson, 1893), Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780241" httpUri="https://zenodo.org/record/12780241/files/figure.png" pageId="5" pageNumber="458">Fig. 3</figureCitation>
). Within the 
<emphasis id="B97D205CD43BFFBCEC38339369C9FCF9" box="[1104,1211,763,789]" italics="true" pageId="5" pageNumber="458">molyneux</emphasis>
group, 
<taxonomicName id="4C0987CDD43BFFBCED7A339368EEFCF9" authorityName="Ahyong" authorityYear="2024" box="[1298,1436,763,789]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCED7A339368EEFCF9" box="[1298,1436,763,789]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCE8FF32776D86FCD5" bold="true" box="[151,244,799,825]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCE8FF32776D86FCD5" box="[151,244,799,825]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most closely resembles 
<taxonomicName id="4C0987CDD43BFFBCEA7A32776FE0FCD5" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[530,658,799,825]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEA7A32776FE0FCD5" box="[530,658,799,825]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
(
<typeStatus id="54B242ECD43BFFBCEACC32776FA4FCD5" box="[676,726,799,825]" pageId="5" pageNumber="458">type</typeStatus>
locality: Rameswaram, Gulf of Mannar, 
<collectingCountry id="F31EBCDED43BFFBCECDE32776983FCD5" box="[1206,1265,799,825]" name="India" pageId="5" pageNumber="458">India</collectingCountry>
) and 
<taxonomicName id="4C0987CDD43BFFBCED5D32776C49FCB1" authority="Ahyong, 2012" authorityName="Ahyong" authorityYear="2012" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCED5D327768EEFCD5" box="[1333,1436,799,825]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
<bibRefCitation id="EF9881BFD43BFFBCE8FF322B6C49FCB1" author="Ahyong, S. T." box="[151,315,835,861]" pageId="5" pageNumber="458" pagination="1 - 111" refId="ref5754" refString="Ahyong, S. T. (2012) The marine fauna of New Zealand: mantis shrimps (Crustacea: Stomatopoda). NIWA Biodiversity Memoir, 125, 1 - 111." type="journal article" year="2012">Ahyong, 2012</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54B242ECD43BFFBCE925322B6C0DFCB1" box="[333,383,835,861]" pageId="5" pageNumber="458">type</typeStatus>
locality: 
<collectingCountry id="F31EBCDED43BFFBCE998322B6FFEFCB1" box="[496,652,835,861]" name="New Zealand" pageId="5" pageNumber="458">New Zealand</collectingCountry>
) in sharing the combination of a slender uropodal endopod (length about 3× width) with several proximal setae on an otherwise glabrous inner margin, and with the ventral surface of the submedian primary teeth of the telson either non-carinate (
<taxonomicName id="4C0987CDD43BFFBCEBEE32E36960FC49" authorityName="Ahyong" authorityYear="2024" box="[902,1042,907,933]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCEBEE32E36960FC49" box="[902,1042,907,933]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEC7332E36905FC49" bold="true" box="[1051,1143,907,933]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEC7332E36905FC49" box="[1051,1143,907,933]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C0987CDD43BFFBCECEF32E3699FFC49" authorityName="Ahyong" authorityYear="2012" box="[1159,1261,907,933]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCECEF32E3699FFC49" box="[1159,1261,907,933]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
) or unicarinate (
<taxonomicName id="4C0987CDD43BFFBCE8C932C76C6DFC25" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[161,287,943,969]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCE8C932C76C6DFC25" box="[161,287,943,969]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
), rather than bicarinate (
<taxonomicName id="4C0987CDD43BFFBCEA5332C76FADFC25" baseAuthorityName="Manning" baseAuthorityYear="1968" box="[571,735,943,969]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="bicarinatus">
<emphasis id="B97D205CD43BFFBCEA5332C76FADFC25" box="[571,735,943,969]" italics="true" pageId="5" pageNumber="458">G. bicarinatus</emphasis>
</taxonomicName>
). 
<taxonomicName id="4C0987CDD43BFFBCEA9032C76952FC25" authorityName="Ahyong" authorityYear="2024" box="[760,1056,943,969]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCEA9032C76952FC25" box="[760,1056,943,969]" italics="true" pageId="5" pageNumber="458">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEC4132C769F7FC25" bold="true" box="[1065,1157,943,969]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEC4132C769F7FC25" box="[1065,1157,943,969]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C0987CDD43BFFBCED7632C768EEFC25" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1310,1436,943,969]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCED7632C768EEFC25" box="[1310,1436,943,969]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D43BFFBCE8FF32BB6C56FC01" box="[151,292,979,1005]" pageId="5" pageNumber="458" sensu="stricto">sensu stricto</taxonomicNameLabel>
in features of the rostral plate with the anterior margin sloping posteriorly to almost transverse with rounded anterolateral corners (
<figureCitation id="1332E0CBD43BFFBCE993329F6F11FBFD" box="[507,611,1015,1041]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Figs. 1A</figureCitation>
, 
<figureCitation id="1332E0CBD43BFFBCEA1A329F6FF3FBFD" box="[626,641,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="5" pageNumber="458">2</figureCitation>
) (versus anterior margin concave and inclined anteriorly with acute anterolateral corners; 
<figureCitation id="1332E0CBD43BFFBCE9E735736C9BFBD9" box="[399,489,1051,1077]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1856]" captionTargetBox="[210,1375,315,1805]" captionTargetId="figure-95@6.[209,1378,309,1809]" captionTargetPageId="6" captionText="FIGURE 3. Gonodactylellus demanii (Henderson, 1893), Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780241" httpUri="https://zenodo.org/record/12780241/files/figure.png" pageId="5" pageNumber="458">Fig. 3A</figureCitation>
), ocular scales distinctly separate rather than medially fused and separated only by shallow concavity (
<figureCitation id="1332E0CBD43BFFBCE91B35576C8AFBB5" box="[371,504,1087,1113]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Figs. 1A, C</figureCitation>
, 
<figureCitation id="1332E0CBD43BFFBCEA6C35576F66FBB5" box="[516,532,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="5" pageNumber="458">2</figureCitation>
, 
<figureCitation id="1332E0CBD43BFFBCEA4935576FFDFBB5" box="[545,655,1087,1113]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1856]" captionTargetBox="[210,1375,315,1805]" captionTargetId="figure-95@6.[209,1378,309,1809]" captionTargetPageId="6" captionText="FIGURE 3. Gonodactylellus demanii (Henderson, 1893), Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780241" httpUri="https://zenodo.org/record/12780241/files/figure.png" pageId="5" pageNumber="458">3A, C, M</figureCitation>
), presence of a notch on the outer proximal margin of the dactylus of the raptorial claw (
<figureCitation id="1332E0CBD43BFFBCE90F350B6CCBFB91" box="[359,441,1123,1149]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Fig. 1E</figureCitation>
) (absent in 
<taxonomicName id="4C0987CDD43BFFBCEA52350B6FC7FB91" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[570,693,1123,1149]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEA52350B6FC7FB91" box="[570,693,1123,1149]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
; 
<figureCitation id="1332E0CBD43BFFBCEAAB350B6E65FB91" box="[707,791,1123,1149]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1856]" captionTargetBox="[210,1375,315,1805]" captionTargetId="figure-95@6.[209,1378,309,1809]" captionTargetPageId="6" captionText="FIGURE 3. Gonodactylellus demanii (Henderson, 1893), Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780241" httpUri="https://zenodo.org/record/12780241/files/figure.png" pageId="5" pageNumber="458">Fig. 3E</figureCitation>
), and absence of a carina along the ventral midline of each submedian telson tooth (present in 
<taxonomicName id="4C0987CDD43BFFBCEA7235EF6FE7FB4D" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[538,661,1159,1185]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEA7235EF6FE7FB4D" box="[538,661,1159,1185]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
; 
<figureCitation id="1332E0CBD43BFFBCEACB35EF6F8AFB4D" box="[675,760,1159,1185]" captionStart="FIGURE 3" captionStartId="6.[151,250,1832,1856]" captionTargetBox="[210,1375,315,1805]" captionTargetId="figure-95@6.[209,1378,309,1809]" captionTargetPageId="6" captionText="FIGURE 3. Gonodactylellus demanii (Henderson, 1893), Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780241" httpUri="https://zenodo.org/record/12780241/files/figure.png" pageId="5" pageNumber="458">Fig. 3L</figureCitation>
). Whereas the ventral surface of each submedian telson tooth carries a prominent longitudinal carina along the midline in 
<taxonomicName id="4C0987CDD43BFFBCEB5335C36ECAFB29" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[827,952,1195,1221]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEB5335C36ECAFB29" box="[827,952,1195,1221]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
, the ventral surface of the submedian teeth in 
<taxonomicName id="4C0987CDD43BFFBCE8DC35A76C4FFB05" authorityName="Ahyong" authorityYear="2024" box="[180,317,1231,1257]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCE8DC35A76C4FFB05" box="[180,317,1231,1257]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCE92C35A76CEFFB05" bold="true" box="[324,413,1231,1257]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCE92C35A76CEFFB05" box="[324,413,1231,1257]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is essentially smooth and flat, with the mesial margin forming a blunt, narrow step adjacent to the submedian denticles (
<figureCitation id="1332E0CBD43BFFBCE9B1359B6F46FAE1" box="[473,564,1267,1293]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Fig. 1K</figureCitation>
). Note that 
<bibRefCitation id="EF9881BFD43BFFBCEAD6359B6E04FAE1" author="Manning, R. B." box="[702,886,1267,1293]" pageId="5" pageNumber="458" pagination="1 - 27" refId="ref6800" refString="Manning, R. B. (1967) Notes on the demanii section of genus Gonodactylus Berthold with descriptions of three new species (Crustacea: Stomatopoda). Proceedings of the United States National Museum, 123, 1 - 27. https: // doi. org / 10.5479 / si. 00963801.123 - 3618.1" type="journal article" year="1967">Manning (1967)</bibRefCitation>
diagnosed 
<taxonomicName id="4C0987CDD43BFFBCEB92359B690BFAE1" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1018,1145,1267,1293]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCEB92359B690BFAE1" box="[1018,1145,1267,1293]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
as lacking the prominent ventral carina of the submedian telson teeth (at most a low mesial ridge), but re-examination of the 
<typeStatus id="54B242ECD43BFFBCECB4347F687EFADD" box="[1244,1292,1303,1329]" pageId="5" pageNumber="458">type</typeStatus>
material and other Indian material showed the carinae to be present. Although 
<taxonomicName id="4C0987CDD43BFFBCEB1034536971FAB9" authorityName="Ahyong" authorityYear="2024" box="[888,1027,1339,1365]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCEB1034536971FAB9" box="[888,1027,1339,1365]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEC6334536917FAB9" bold="true" box="[1035,1125,1339,1365]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEC6334536917FAB9" box="[1035,1125,1339,1365]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has long been mistaken for 
<taxonomicName id="4C0987CDD43BFFBCE8FF34376C66FA95" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[151,276,1375,1401]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCE8FF34376C66FA95" box="[151,276,1375,1401]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
, the new species is morphologically nearest to 
<taxonomicName id="4C0987CDD43BFFBCEB4534376EE3FA95" authorityName="Ahyong" authorityYear="2012" box="[813,913,1375,1401]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCEB4534376EE3FA95" box="[813,913,1375,1401]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
from the southwestern Pacific (
<collectingCountry id="F31EBCDED43BFFBCEC92343768E7FA95" box="[1274,1429,1375,1401]" name="New Zealand" pageId="5" pageNumber="458">New Zealand</collectingCountry>
) in sharing similar rostral plate form and ventral telson ornamentation. 
<taxonomicName id="4C0987CDD43BFFBCEBD534EB6994FA71" authorityName="Ahyong" authorityYear="2024" box="[957,1254,1411,1437]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCEBD534EB6994FA71" box="[957,1254,1411,1437]" italics="true" pageId="5" pageNumber="458">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEC8734EB6839FA71" bold="true" box="[1263,1355,1411,1437]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEC8734EB6839FA71" box="[1263,1355,1411,1437]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C0987CDD43BFFBCE8BC34CF6C44FA2D" authorityName="Ahyong" authorityYear="2012" box="[212,310,1447,1473]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCE8BC34CF6C44FA2D" box="[212,310,1447,1473]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
in having rounded ocular scales (versus truncate in 
<taxonomicName id="4C0987CDD43BFFBCEB1B34CF6EA4FA2D" authorityName="Ahyong" authorityYear="2012" box="[883,982,1447,1473]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCEB1B34CF6EA4FA2D" box="[883,982,1447,1473]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
), and, in specimens exceeding TL 
<quantity id="4CF151ABD43BFFBCED3E34CF6DB7FA09" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="5" pageNumber="458" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
, the orientation of the lateral primary teeth of the telson. In 
<taxonomicName id="4C0987CDD43BFFBCEB3534A36EB2FA09" authorityName="Ahyong" authorityYear="2012" box="[861,960,1483,1509]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCEB3534A36EB2FA09" box="[861,960,1483,1509]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
, the lateral primary teeth are sharply acute and set-off from the margin of the intermediate tooth by a narrow notch (
<bibRefCitation id="EF9881BFD43BFFBCEBB33487690EF9E5" author="Ahyong, S. T." box="[987,1148,1519,1545]" pageId="5" pageNumber="458" pagination="1 - 111" refId="ref5754" refString="Ahyong, S. T. (2012) The marine fauna of New Zealand: mantis shrimps (Crustacea: Stomatopoda). NIWA Biodiversity Memoir, 125, 1 - 111." type="journal article" year="2012">Ahyong 2012</bibRefCitation>
: fig. 9B). In 
<taxonomicName id="4C0987CDD43BFFBCED78348768EEF9E5" authorityName="Ahyong" authorityYear="2024" box="[1296,1436,1519,1545]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCED78348768EEF9E5" box="[1296,1436,1519,1545]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCE8FF377B6D86F9C1" bold="true" box="[151,244,1555,1581]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCE8FF377B6D86F9C1" box="[151,244,1555,1581]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, however, the apex of the lateral tooth is angular but not spiniform and set-off from the margin of the intermediate tooth by a wide notch (
<figureCitation id="1332E0CBD43BFFBCEA4D375F6FD8F9BD" box="[549,682,1591,1617]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Figs. 1H, K</figureCitation>
, 
<figureCitation id="1332E0CBD43BFFBCEADC375F6F8AF9BD" box="[692,760,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="5" pageNumber="458">2A, B</figureCitation>
). Note that the prominence of the lateral primary tooth varies allometrically in 
<taxonomicName id="4C0987CDD43BFFBCE93137336C96F999" authorityName="Ahyong" authorityYear="2024" box="[345,484,1627,1653]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCE93137336C96F999" box="[345,484,1627,1653]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCE98437336F35F999" bold="true" box="[492,583,1627,1653]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCE98437336F35F999" box="[492,583,1627,1653]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="1332E0CBD43BFFBCEA3F37336FA6F999" box="[599,724,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="5" pageNumber="458">Fig. 2C, D</figureCitation>
). In specimens of 
<taxonomicName id="4C0987CDD43BFFBCEBCC37336942F999" authorityName="Ahyong" authorityYear="2024" box="[932,1072,1627,1653]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCEBCC37336942F999" box="[932,1072,1627,1653]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEC50373369E1F999" bold="true" box="[1080,1171,1627,1653]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEC50373369E1F999" box="[1080,1171,1627,1653]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
below about TL 
<quantity id="4CF151ABD43BFFBCED3E37336DB7F975" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="5" pageNumber="458" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
, the lateral primary tooth may approach that of 
<taxonomicName id="4C0987CDD43BFFBCEABC37176E44F975" authorityName="Ahyong" authorityYear="2012" box="[724,822,1663,1689]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="oshea">
<emphasis id="B97D205CD43BFFBCEABC37176E44F975" box="[724,822,1663,1689]" italics="true" pageId="5" pageNumber="458">G. oshea</emphasis>
</taxonomicName>
, so in smaller specimens, the shape of the ocular scales should be used to distinguish the species.
</paragraph>
<paragraph id="8BB6FC4ED43BFFBFE8AF37AF699BFEF1" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1436,151,285]" lastPageId="6" lastPageNumber="459" pageId="5" pageNumber="458">
Sexual dimorphism in 
<taxonomicName id="4C0987CDD43BFFBCE9AE37AF6F23F90D" authorityName="Ahyong" authorityYear="2024" box="[454,593,1735,1761]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD43BFFBCE9AE37AF6F23F90D" box="[454,593,1735,1761]" italics="true" pageId="5" pageNumber="458">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD43BFFBCEA3037AF6FC0F90D" bold="true" box="[600,690,1735,1761]" pageId="5" pageNumber="458">
<taxonomicNameLabel id="A24E9D27D43BFFBCEA3037AF6FC0F90D" box="[600,690,1735,1761]" pageId="5" pageNumber="458" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is evident in the more inflated dorsal telson carinae in males, as is typical of other gonodactylids.As in other members of the 
<emphasis id="B97D205CD43BFFBCEB7937836E0EF8E9" box="[785,892,1771,1797]" italics="true" pageId="5" pageNumber="458">molyneux</emphasis>
group (
<bibRefCitation id="EF9881BFD43BFFBCEBA53783699CF8E9" author="Ahyong, S. T. &amp; Erdmann, M. V." box="[973,1262,1771,1797]" pageId="5" pageNumber="458" pagination="89 - 95" refId="ref5903" refString="Ahyong, S. T. &amp; Erdmann, M. V. (2007) Two new species of Gonodactylellus from Indonesia. Raffles Bulletin of Zoology, 55, 89 - 95." type="journal article" year="2007">Ahyong &amp; Erdmann 2007</bibRefCitation>
; 
<bibRefCitation id="EF9881BFD43BFFBCEC9D378368E2F8E9" author="Ahyong, S. T." box="[1269,1424,1771,1797]" pageId="5" pageNumber="458" pagination="41 - 55" refId="ref5702" refString="Ahyong, S. T. (2008) Stomatopod Crustacea from the Dampier Archipelago, Western Australia. Records of the Western Australian Museum, Supplement 73, 41 - 55. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 73.2007.041 - 055" type="journal article" year="2008">Ahyong 2008</bibRefCitation>
), the degree of dorsal spination of the telson (but also bluntness of the spines), progressively increases with increasing body size (
<figureCitation id="1332E0CBD43BFFBCE979365B6CF9F8A1" box="[273,395,1843,1869]" captionStart="FIGURE 1" captionStartId="2.[151,250,1822,1846]" captionTargetBox="[207,1382,186,1789]" captionTargetId="figure-18@2.[203,1384,179,1796]" captionTargetPageId="2" captionText="FIGURE 1. Gonodactylellus celosinus sp. nov., male holotype, TL 27 mm, Red Sea, SMF 62255; A, anterior cephalothorax; B, rostral plate, right lateral view; C, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somites 5–6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, telson, ventral surface; L, right pleopod 1 endopod, anterior view. Scale: A–K = 2.0 mm, L = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780237" httpUri="https://zenodo.org/record/12780237/files/figure.png" pageId="5" pageNumber="458">Figs. 1H, I</figureCitation>
, 
<figureCitation id="1332E0CBD43BFFBCE9FF365B6CDAF8A1" box="[407,424,1843,1869]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="5" pageNumber="458">2</figureCitation>
). In females and smaller males, the apices of the dorsal telson spines are typically pointed, but in males above about TL 
<quantity id="4CF151ABD43BFFBCE980363E6F16F89D" box="[488,612,1878,1905]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="5" pageNumber="458" unit="mm" value="26.0" valueMax="27.0" valueMin="25.0">25–27 mm</quantity>
, the pointed apices may become obsolete and blunt as figured by 
<bibRefCitation id="EF9881BFD43BFFBCED30363F6DAFF879" author="Kemp, S." pageId="5" pageNumber="458" pagination="1 - 217" refId="ref6528" refString="Kemp, S. (1913) An account of the Crustacea Stomatopoda of the Indo-Pacific region, based on the collection in the Indian Museum. Memoirs of the Indian Museum, 4, 1 - 217, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 12631" type="journal article" year="1913">Kemp (1913</bibRefCitation>
: pl. 9 fig. 109; as 
<taxonomicName id="4C0987CDD43BFFBCE9C136136F55F879" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[425,551,1915,1941]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD43BFFBCE9C136136F55F879" box="[425,551,1915,1941]" italics="true" pageId="5" pageNumber="458">G. demanii</emphasis>
</taxonomicName>
). Similarly, the apices of the submedian and intermediate carina of abdominal somite 6 become smaller or even obsolete with increasing body size. As in other stomatopods, the relative length of the antennular peduncle decreases with increasing size, being about 0.8CL in the smallest specimens to about 0.6–0.7CL in adults. The mandibular palp is present in all specimens examined, but is unsegmented in the smallest specimen (juvenile female, TL 
<quantity id="4CF151ABD438FFBFE99831D36F33FF39" box="[496,577,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="459" unit="mm" value="11.0">11 mm</quantity>
). At TL 
<quantity id="4CF151ABD438FFBFEAF131D36E61FF39" box="[665,787,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="6" pageNumber="459" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
, the second article of the mandibular palp is differentiated, and by TL 
<quantity id="4CF151ABD438FFBFE97C31B76C14FF15" box="[276,358,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="459" unit="mm" value="14.0">14 mm</quantity>
, the third palp article is differentiated. The dorsal swelling on the uropodal protopod is low but evident by TL 
<quantity id="4CF151ABD438FFBFE953306A6CFFFEF1" box="[315,397,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="459" unit="mm" value="15.0">15 mm</quantity>
. In males, the modified pleopod 1 endopod is fully developed by TL 
<quantity id="4CF151ABD438FFBFECFB306B6997FEF1" box="[1171,1253,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="6" pageNumber="459" unit="mm" value="19.0">19 mm</quantity>
.
</paragraph>
</subSubSection>
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<paragraph id="8BB6FC4ED438FFBFE8FF36406935F818" blockId="6.[151,1436,1832,2037]" pageId="6" pageNumber="459">
<emphasis id="B97D205CD438FFBFE8FF36406C66F8AD" bold="true" box="[151,276,1832,1857]" pageId="6" pageNumber="459">FIGURE 3.</emphasis>
<taxonomicName id="4C0987CDD438FFBFE97536416F94F8AD" authority="(Henderson, 1893)" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[285,742,1833,1857]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="6" pageNumber="459" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD438FFBFE97536416F6EF8AD" box="[285,540,1833,1857]" italics="true" pageId="6" pageNumber="459">Gonodactylellus demanii</emphasis>
(
<bibRefCitation id="EF9881BFD438FFBFEA4336416FACF8AD" author="Henderson, J. R." box="[555,734,1833,1857]" pageId="6" pageNumber="459" pagination="325 - 458" refId="ref6240" refString="Henderson, J. R. (1893) A contribution to Indian carcinology. Transactions of the Linnean Society of London, Series 2, Zoology, 5, 325 - 458, pls. 336 - 340. https: // doi. org / 10.1111 / j. 1096 - 3642.1893. tb 00653. x" type="journal article" year="1893">Henderson, 1893</bibRefCitation>
)
</taxonomicName>
, Kilakarai, Gulf of Mannar, India, AM P3967: A–L, female, TL 26 mm; M–N, female, TL 18 mm. A, anterior cephalothorax; B, rostral plate, right lateral view; C, M, ocular scales, dorsal view; D, right antennal protopod, lateral view; E, right raptorial claw, lateral view; F, thoracic somites 6–8, right lateral view; G, abdominal somites 4–5 posterolateral angles, right lateral view; H, abdominal somite 6, telson and right uropod, dorsal view (setae omitted); I, abdominal somite 6 and telson, lateral view; J, right uropod, ventral view; K, left uropodal endopod, dorsal view (setae omitted); L, telson, ventral surface; N, rostral plate, dorsal view. Scale = 2.0 mm.
</paragraph>
</caption>
<subSubSection id="C313AFC5D439FFBEE8AF31FF6CEDFE19" pageId="7" pageNumber="460" type="reference_group">
<paragraph id="8BB6FC4ED439FFBEE8AF31FF6817FE41" blockId="7.[151,1436,151,681]" pageId="7" pageNumber="460">
<taxonomicName id="4C0987CDD439FFBEE8AF31FF6C82FF5D" authorityName="Ahyong" authorityYear="2024" box="[199,496,151,177]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD439FFBEE8AF31FF6C82FF5D" box="[199,496,151,177]" italics="true" pageId="7" pageNumber="460">Gonodactylellus celosinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D439FFBEE99131FF6F24FF5D" box="[505,598,151,177]" pageId="7" pageNumber="460" rank="species">
<emphasis id="B97D205CD439FFBEE99131FF6F22FF5D" bold="true" box="[505,592,151,177]" pageId="7" pageNumber="460">sp. nov</emphasis>
.
</taxonomicNameLabel>
apparently occurs only in the northwestern Indian Ocean, from 
<collectingCountry id="F31EBCDED439FFBEED5131FF68EAFF5D" box="[1337,1432,151,177]" name="Somalia" pageId="7" pageNumber="460">Somalia</collectingCountry>
, the Red Sea, the Persian Gulf and the Arabian Sea. The previous record of 
<taxonomicName id="4C0987CDD439FFBEEBB431D3692AFF39" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[988,1112,187,213]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD439FFBEEBB431D3692AFF39" box="[988,1112,187,213]" italics="true" pageId="7" pageNumber="460">G. demanii</emphasis>
</taxonomicName>
from Astola Island, 
<collectingCountry id="F31EBCDED439FFBEED5531D368EEFF39" box="[1341,1436,187,213]" name="Pakistan" pageId="7" pageNumber="460">Pakistan</collectingCountry>
(
<bibRefCitation id="EF9881BFD439FFBEE8F731B76C35FF15" author="Manning, R. B." box="[159,327,223,249]" pageId="7" pageNumber="460" pagination="1 - 27" refId="ref6800" refString="Manning, R. B. (1967) Notes on the demanii section of genus Gonodactylus Berthold with descriptions of three new species (Crustacea: Stomatopoda). Proceedings of the United States National Museum, 123, 1 - 27. https: // doi. org / 10.5479 / si. 00963801.123 - 3618.1" type="journal article" year="1967">Manning 1967</bibRefCitation>
; 
<bibRefCitation id="EF9881BFD439FFBEE93A31B76F1EFF15" author="Tirmizi, N. M. &amp; Manning, R. B." box="[338,620,223,249]" pageId="7" pageNumber="460" pagination="1 - 48" refId="ref7660" refString="Tirmizi, N. M. &amp; Manning, R. B. (1968) Stomatopod Crustacea from West Pakistan. Proceedings of the United States National Museum, 125, 1 - 48. https: // doi. org / 10.5479 / si. 00963801.125 - 3666.1" type="journal article" year="1968">Tirmizi &amp; Manning 1968</bibRefCitation>
) is referable to 
<taxonomicName id="4C0987CDD439FFBEEB7331B76ED6FF15" authorityName="Ahyong" authorityYear="2024" box="[795,932,223,249]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD439FFBEEB7331B76ED6FF15" box="[795,932,223,249]" italics="true" pageId="7" pageNumber="460">G. celosinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D439FFBEEBC331B76974FF15" box="[939,1030,223,249]" pageId="7" pageNumber="460" rank="species">
<emphasis id="B97D205CD439FFBEEBC331B76E8DFF15" bold="true" box="[939,1023,223,249]" pageId="7" pageNumber="460">sp. nov</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation id="1332E0CBD439FFBEEC7D31B76918FF15" box="[1045,1130,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1333,1357]" captionTargetBox="[176,1410,481,1302]" captionTargetId="figure-160@3.[170,1416,475,1309]" captionTargetPageId="3" captionText="FIGURE 2. Gonodactylellus celosinus sp. nov., telson, ocular scales and rostral plate, dorsal view: A, female paratype, TL 24 mm, Red Sea, RSS1-2013-35, SMF 62256; B, female, TL 21 mm, Astola Island, Pakistan, USNM 120473; C, juvenile female, TL 11 mm, Red Sea, RSS1-2013-37, KAUMM 1060; D, juvenile male, TL 14 mm, Red Sea, RSS1-2013-37, KAUMM 1060. Scale: A, B = 2.0 mm; C, D = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12780239" httpUri="https://zenodo.org/record/12780239/files/figure.png" pageId="7" pageNumber="460">Fig. 2B</figureCitation>
) The only previous records of 
<taxonomicName id="4C0987CDD439FFBEE8DF306B6C47FEF1" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[183,309,259,285]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD439FFBEE8DF306B6C47FEF1" box="[183,309,259,285]" italics="true" pageId="7" pageNumber="460">G. demanii</emphasis>
</taxonomicName>
from western 
<collectingCountry id="F31EBCDED439FFBEE9B5306B6F65FEF1" box="[477,535,259,285]" name="India" pageId="7" pageNumber="460">India</collectingCountry>
are from the far north: 
<collectingCountry id="F31EBCDED439FFBEEB40306B6EF5FEF1" box="[808,903,259,285]" name="India" pageId="7" pageNumber="460">Mumbai</collectingCountry>
(as 
<collectingCountry id="F31EBCDED439FFBEEBD1306B6968FEF1" box="[953,1050,259,285]" name="India" pageId="7" pageNumber="460">Bombay</collectingCountry>
) and the Gulf of Kachchh (
<bibRefCitation id="EF9881BFD439FFBEED3E306B6DA0FEAD" author="Kemp, S." pageId="7" pageNumber="460" pagination="1 - 217" refId="ref6528" refString="Kemp, S. (1913) An account of the Crustacea Stomatopoda of the Indo-Pacific region, based on the collection in the Indian Museum. Memoirs of the Indian Museum, 4, 1 - 217, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 12631" type="journal article" year="1913">Kemp 1913</bibRefCitation>
; 
<bibRefCitation id="EF9881BFD439FFBEE8B5304F6CC2FEAD" author="Shanbhogue, S. L." box="[221,432,295,321]" pageId="7" pageNumber="460" pagination="33 - 36" refId="ref7395" refString="Shanbhogue, S. L. (1969) Catalogue of stomatopods in reference collections of the Central Marine Fisheries Research Institute. Bulletin of the Central Marine Fisheries Research Institute, 9, 33 - 36." type="journal article" year="1969">Shanbhogue, 1969</bibRefCitation>
; 
<bibRefCitation id="EF9881BFD439FFBEE9D3304F6FE6FEAD" author="Trivedi, J. N. &amp; Ahyong, S. T. &amp; Vachhrajani, K. D. &amp; Kumar, A. B." box="[443,660,295,321]" pageId="7" pageNumber="460" pagination="221 - 238" refId="ref7710" refString="Trivedi, J. N., Ahyong, S. T., Vachhrajani, K. D. &amp; Kumar, A. B. (2020) An annotated checklist of the mantis shrimps of India (Crustacea: Stomatopoda). Zootaxa, 4768 (2), 221 - 238. https: // doi. org / 10.11646 / zootaxa. 4768.2.4" type="journal article" year="2020">
Trivedi 
<emphasis id="B97D205CD439FFBEEA7B304F6F38FEAD" box="[531,586,295,321]" italics="true" pageId="7" pageNumber="460">et al.</emphasis>
, 2020
</bibRefCitation>
; 
<bibRefCitation id="EF9881BFD439FFBEEAF5304F6E07FEAD" author="Vadher, P. &amp; Kardani, H. &amp; Beleem, I." box="[669,885,295,321]" pageId="7" pageNumber="460" pagination="21032 - 21042" refId="ref7778" refString="Vadher, P., Kardani, H. &amp; Beleem, I. (2022) Diversity and distribution of mantis shrimps (Arthropoda: Crustacea: Stomatopoda) in the Gulf of Kachchh, Gujarat, India. Journal of Threatened Taxa, 14, 21032 - 21042. https: // doi. org / 10.11609 / jott. 7471.14.5.21032 - 21042" type="journal article" year="2022">
Vadher 
<emphasis id="B97D205CD439FFBEEA9B304F6E59FEAD" box="[755,811,295,321]" italics="true" pageId="7" pageNumber="460">et al.</emphasis>
, 2022
</bibRefCitation>
). The Gulf of Kachchh records are referable to 
<taxonomicName id="4C0987CDD439FFBEEDE9304F6D8DFE89" authorityName="Ahyong" authorityYear="2024" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD439FFBEEDE9304F6D8DFE89" italics="true" pageId="7" pageNumber="460">G. celosinus</emphasis>
</taxonomicName>
<emphasis id="B97D205CD439FFBEE96E30236C2DFE89" bold="true" box="[262,351,331,357]" pageId="7" pageNumber="460">
<taxonomicNameLabel id="A24E9D27D439FFBEE96E30236C2DFE89" box="[262,351,331,357]" pageId="7" pageNumber="460" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, as clearly depicted by 
<bibRefCitation id="EF9881BFD439FFBEEA0D30236E32FE89" author="Vadher, P. &amp; Kardani, H. &amp; Beleem, I." box="[613,832,331,357]" pageId="7" pageNumber="460" pagination="21032 - 21042" refId="ref7778" refString="Vadher, P., Kardani, H. &amp; Beleem, I. (2022) Diversity and distribution of mantis shrimps (Arthropoda: Crustacea: Stomatopoda) in the Gulf of Kachchh, Gujarat, India. Journal of Threatened Taxa, 14, 21032 - 21042. https: // doi. org / 10.11609 / jott. 7471.14.5.21032 - 21042" type="journal article" year="2022">
Vadher 
<emphasis id="B97D205CD439FFBEEAD330236F9EFE89" box="[699,748,331,357]" italics="true" pageId="7" pageNumber="460">et al</emphasis>
. (2022
</bibRefCitation>
: image 1a–f). The record from 
<collectingCountry id="F31EBCDED439FFBEECF23023698BFE89" box="[1178,1273,331,357]" name="India" pageId="7" pageNumber="460">Mumbai</collectingCountry>
(
<bibRefCitation id="EF9881BFD439FFBEED6F302368E6FE89" author="Kemp, S." box="[1287,1428,331,357]" pageId="7" pageNumber="460" pagination="1 - 217" refId="ref6528" refString="Kemp, S. (1913) An account of the Crustacea Stomatopoda of the Indo-Pacific region, based on the collection in the Indian Museum. Memoirs of the Indian Museum, 4, 1 - 217, pls. 1 - 10. https: // doi. org / 10.5962 / bhl. title. 12631" type="journal article" year="1913">Kemp, 1913</bibRefCitation>
) requires confirmation but is probably also referable to 
<taxonomicName id="4C0987CDD439FFBEEA9430076EF5FE65" authorityName="Ahyong" authorityYear="2024" box="[764,903,367,393]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="celosinus" status="sp. nov.">
<emphasis id="B97D205CD439FFBEEA9430076EF5FE65" box="[764,903,367,393]" italics="true" pageId="7" pageNumber="460">G. celosinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D439FFBEEBE630076E9BFE65" box="[910,1001,367,393]" pageId="7" pageNumber="460" rank="species">
<emphasis id="B97D205CD439FFBEEBE630076E91FE65" bold="true" box="[910,995,367,393]" pageId="7" pageNumber="460">sp. nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="4C0987CDD439FFBEEB993007687BFE65" baseAuthorityName="Henderson" baseAuthorityYear="1893" box="[1009,1289,367,393]" class="Malacostraca" family="Gonodactylidae" genus="Gonodactylellus" kingdom="Animalia" order="Stomatopoda" pageId="7" pageNumber="460" phylum="Arthropoda" rank="species" species="demanii">
<emphasis id="B97D205CD439FFBEEB993007687BFE65" box="[1009,1289,367,393]" italics="true" pageId="7" pageNumber="460">Gonodactylellus demanii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24E9D27D439FFBEED79300768EEFE65" box="[1297,1436,367,393]" pageId="7" pageNumber="460" sensu="stricto">sensu stricto</taxonomicNameLabel>
is reliably now known only from the Bay of Bengal between southeastern 
<collectingCountry id="F31EBCDED439FFBEEBBF30FB6963FE41" box="[983,1041,403,429]" name="India" pageId="7" pageNumber="460">India</collectingCountry>
(Gulf of Mannar) and 
<collectingCountry id="F31EBCDED439FFBEED7930FB6813FE41" box="[1297,1377,403,429]" name="Myanmar" pageId="7" pageNumber="460">Burma</collectingCountry>
.
</paragraph>
<paragraph id="8BB6FC4ED439FFBEE8AF30DE6CEDFE19" blockId="7.[151,1436,151,681]" pageId="7" pageNumber="460">
<emphasis id="B97D205CD439FFBEE8AF30DE6C51FE3C" bold="true" box="[199,291,438,464]" pageId="7" pageNumber="460">Habitat</emphasis>
. Intertidal and shallow subtidal reefs to 
<quantity id="4CF151ABD439FFBEEA8030DF6E61FE3D" box="[744,787,439,465]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="460" unit="m" value="3.0">3 m</quantity>
depth (
<bibRefCitation id="EF9881BFD439FFBEEB0030DF697FFE3D" author="Holthuis, L. B." box="[872,1037,439,465]" pageId="7" pageNumber="460" pagination="1 - 45" refId="ref6299" refString="Holthuis, L. B. (1967) The stomatopod Crustacea collected by the 1962 and 1965 Israel South Red Sea Expeditions. The Second Israel South Red Sea Expedition, 1965, Report No. 1. Israel Journal of Zoology, 16, 1 - 45, figs. 1 - 7." type="journal article" year="1967">Holthuis 1967</bibRefCitation>
); amongst rubble and in crevices in coral sponge and algae.
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D439FFBEE8AF309668E2FD8D" pageId="7" pageNumber="460" type="description">
<paragraph id="8BB6FC4ED439FFBEE8AF309668E2FD8D" blockId="7.[151,1436,151,681]" pageId="7" pageNumber="460">
<emphasis id="B97D205CD439FFBEE8AF30966CAEFDF4" bold="true" box="[199,476,510,536]" pageId="7" pageNumber="460">Colour in preservative</emphasis>
. Largely faded, with scattered chromatophores on carapace, thorax, abdomen and raptorial claw; with rectangular median patch of chromatophores on thoracic somite 6 and abdominal somite 1, and small median cluster on abdominal somites 3–5; abdominal somite 6 and telson with diffuse olive green mottling.
</paragraph>
</subSubSection>
<subSubSection id="C313AFC5D439FFBEE8AF33026F75FD45" pageId="7" pageNumber="460" type="distribution">
<paragraph id="8BB6FC4ED439FFBEE8AF33026F75FD45" blockId="7.[151,1436,151,681]" pageId="7" pageNumber="460">
<emphasis id="B97D205CD439FFBEE8AF33026C2BFD68" bold="true" box="[199,345,618,644]" pageId="7" pageNumber="460">Distribution</emphasis>
. Northwestern Indian Ocean: 
<collectingCountry id="F31EBCDED439FFBEEAD533036E6EFD69" box="[701,796,619,645]" name="Somalia" pageId="7" pageNumber="460">Somalia</collectingCountry>
, the Red Sea, Persian Gulf and Arabian Sea including 
<collectingCountry id="F31EBCDED439FFBEE8FF33E76D84FD45" box="[151,246,655,681]" name="Pakistan" pageId="7" pageNumber="460">Pakistan</collectingCountry>
and northwestern 
<collectingCountry id="F31EBCDED439FFBEE9AE33E76F71FD45" box="[454,515,655,681]" name="India" pageId="7" pageNumber="460">India</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>