372 lines
50 KiB
XML
372 lines
50 KiB
XML
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<mods:title id="F6DF2BCCB1BA41EF3EDF92BFA3D7A437">Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae): (2) the species-group of Heteroponera relicta (Wheeler), with descriptions of nine new species and observations on morphology, biogeography and phylogeny of the genus</mods:title>
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<mods:namePart id="A04B5DB158623CAC92AABA5D29267155">Taylor, Robert W.</mods:namePart>
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<mods:title id="31024C258A589C1C2214A97492253008">Zootaxa</mods:title>
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<mods:date id="D5AF7120800F04B690B1644E3A11CC7E">2015</mods:date>
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<mods:number id="894715F963C4067DC8F8FC7A2AB0AF9C">3947</mods:number>
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<treatment id="627587EEFFF92A03DEE6FCBC8EABB6EE" ID-DOI="http://doi.org/10.5281/zenodo.6114520" ID-GBIF-Taxon="119609655" ID-Zenodo-Dep="6114520" LSID="urn:lsid:plazi:treatment:627587EEFFF92A03DEE6FCBC8EABB6EE" httpUri="http://treatment.plazi.org/id/627587EEFFF92A03DEE6FCBC8EABB6EE" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156">
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<subSubSection id="A2C66573FFF92A00DEE6FCBC8FCFB3FC" pageId="5" pageNumber="156" type="nomenclature">
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<paragraph id="EA6336F8FFF92A00DEE6FCBC8A13B36B" blockId="5.[151,1398,834,895]" pageId="5" pageNumber="156">
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<heading id="B12B8194FFF92A00DEE6FCBC8A13B36B" bold="true" fontSize="11" level="1" pageId="5" pageNumber="156" reason="1">
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<emphasis id="D8A8EAEAFFF92A00DEE6FCBC8A13B36B" bold="true" pageId="5" pageNumber="156">
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Key to the Australasian species-groups of
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<taxonomicName id="2DDC4D7BFFF92A00DCD3FCBC887DB348" ID-CoL="4WNX" box="[674,842,834,860]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
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<emphasis id="D8A8EAEAFFF92A00DCD3FCBC887DB348" bold="true" box="[674,842,834,860]" italics="true" pageId="5" pageNumber="156">Heteroponera</emphasis>
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</taxonomicName>
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and species of the
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<taxonomicName id="2DDC4D7BFFF92A00DA45FCBC8F92B348" box="[1076,1189,834,860]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis id="D8A8EAEAFFF92A00DA45FCBC8F92B348" bold="true" box="[1076,1189,834,860]" italics="true" pageId="5" pageNumber="156">H. relicta</emphasis>
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</taxonomicName>
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group (workers, and gynes).
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="EA6336F8FFF92A00DEE6FC528FCFB3FC" blockId="5.[151,1436,940,1000]" pageId="5" pageNumber="156">
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The Australasian species groups are discriminated in couplets 1 to 3, thence the species of the
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<taxonomicName id="2DDC4D7BFFF92A00DAC0FC538E2CB3D1" box="[1201,1307,940,965]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis id="D8A8EAEAFFF92A00DAC0FC538E2CB3D1" box="[1201,1307,940,965]" italics="true" pageId="5" pageNumber="156">H. relicta</emphasis>
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</taxonomicName>
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group. The characteristics of all known Australasian
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<taxonomicName id="2DDC4D7BFFF92A00DC14FC2F8836B3FC" box="[613,769,977,1000]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
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<emphasis id="D8A8EAEAFFF92A00DC14FC2F8836B3FC" box="[613,769,977,1000]" italics="true" pageId="5" pageNumber="156">Heteroponera</emphasis>
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</taxonomicName>
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species were considered in this compilation.
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</paragraph>
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</subSubSection>
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<subSubSection id="A2C66573FFF92A03DEE6FBE98EABB6EE" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156" type="key">
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<keyStep id="51282E5DFFF92A00DEE6FBE98EACB503" pageId="5" pageNumber="156">
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<paragraph id="EA6336F8FFF92A00DEE6FBE98EA1B490" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
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<keyLead id="512D95CDFFF92A00DEE6FBE98EA1B490" pageId="5" pageNumber="156">
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1 Petiole armed dorsally with a small erect median spine. Propodeum with a pair of similarly proportioned posterodorsolateral spines or homologous lamellar spine-like processes. Second gastral segment in side view markedly reflexed, so that the gastral apex is directed strongly anteroventrally. Antennal scrobes and enclosing frontal carinae lacking (see
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<bibRefCitation id="8E4D4B09FFF92A00DAD6FBAC8E1DB473" author="Taylor" box="[1191,1322,1106,1127]" pageId="5" pageNumber="156" refString="Taylor, R. W. (2011) Australasian ants of the subfamily Heteroponerinae (Hymenoptera: Formicidae) (1) General introduction and review of the Heteroponera leae (Wheeler) species-group, with descriptions of two new species. Myrmecological News, 15, 117 - 123." type="journal article" year="2011">Taylor (2011)</bibRefCitation>
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for illustrations and taxonomy of the three known species)...........................................
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<taxonomicName id="2DDC4D7BFFF92A00DAB4FB8E8E3CB491" ID-CoL="3LCK6" box="[1221,1291,1135,1157]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="leae">
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<emphasis id="D8A8EAEAFFF92A00DAB4FB8E8E3CB491" box="[1221,1291,1135,1157]" italics="true" pageId="5" pageNumber="156">H. leae</emphasis>
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</taxonomicName>
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species-group
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</keyLead>
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</paragraph>
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<paragraph id="EA6336F8FFF92A00DEE6FB738EACB503" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
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<keyLead id="512D95CDFFF92A00DEE6FB738EACB503" pageId="5" pageNumber="156">
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- Petiole lacking a dorsomedial spine, though sometimes drawn to a tapering obtuse summit. Propodeum lacking slender erect spines, though sometimes with sculpturally jagged posterolateral edges and with very small accompanying spines in one species (known only from the Mt Elliot summit near Townsville). Reflexure of second gastral segment unexceptional. Antennal scrobes in most species bordered dorsally and posteriorly by extended frontal carinae, sometimes strongly developed, (
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<figureCitation id="72E72A7DFFF92A00DB3CFB1B8EA0B4EE" box="[1357,1431,1253,1274]" captionStart="FIGURES 22 – 25" captionStartId="17.[151,264,1955,1977]" captionTargetBox="[202,1384,193,1932]" captionTargetId="figure@17.[202,1385,193,1933]" captionTargetPageId="17" captionText="FIGURES 22 – 25. Heteroponera viviennae sp. n., holotype worker, standard views (HW = 1.11 mm, PW = 0.79 mm, WL = 1.25, GW = 0.80 mm)." httpUri="https://zenodo.org/record/398960/files/figure.png" pageId="5" pageNumber="156">Figs 23</figureCitation>
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,
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<figureCitation id="72E72A7DFFF92A00DE90FAFC8A28B503" box="[225,287,1282,1303]" captionStart="FIGURES 30 – 33" captionStartId="19.[151,264,1951,1973]" captionTargetBox="[198,1388,193,1929]" captionTargetId="figure@19.[198,1388,193,1929]" captionTargetPageId="19" captionText="FIGURES 30 – 33. Heteroponera monteithi sp. n., Gyne (Mt Dryander), standard views (HW = 1.04 mm, PW = 0.77 mm, WL = 1.41 mm, GW = 1.26 mm)." httpUri="https://zenodo.org/record/398962/files/figure.png" pageId="5" pageNumber="156">30, 31</figureCitation>
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), otherwise vestigial or absent (
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<figureCitation id="72E72A7DFFF92A00DC3EFAFC898AB503" box="[591,701,1282,1303]" captionStart="FIGURES 34 – 37" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[179,1406,193,1930]" captionTargetId="figure@21.[179,1407,193,1931]" captionTargetPageId="21" captionText="FIGURES 34 – 37. Heteroponera lioprocta sp. n., holotype worker, standard views (HW = 1.06 mm, PW = 0.79 mm, WL = 1.23 mm, GW = 0.99 mm)." httpUri="https://zenodo.org/record/398963/files/figure.png" pageId="5" pageNumber="156">Figs 34, 35</figureCitation>
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,
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<figureCitation id="72E72A7DFFF92A00DCB6FAFC8833B503" box="[711,772,1282,1303]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="5" pageNumber="156">42, 43</figureCitation>
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)..................................................... 2
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="51282E5DFFF92A00DEE6FAE18EACB656" pageId="5" pageNumber="156">
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<paragraph id="EA6336F8FFF92A00DEE6FAE18EACB598" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
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<keyLead id="512D95CDFFF92A00DEE6FAE18EACB598" pageId="5" pageNumber="156">
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2(1) Size generally smaller (HW less than
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). Second gastral segment in dorsal view not notably narrower than first, or markedly tapered posteriorly. Antennal scrobes lacking, the scrobal areas sculpturally undifferentiated (widespread, all states (including Australian Capital Territory) except Tasmania. Doubtfully represented from the Northern Territory. (The group currently includes the only known
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<collectingCountry id="92CB7668FFF92A00DC61FA8989A7B598" box="[528,656,1399,1420]" name="New Zealand" pageId="5" pageNumber="156">New Zealand</collectingCountry>
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<taxonomicName id="2DDC4D7BFFF92A00DCE7FA86882BB599" ID-CoL="4WNX" box="[662,796,1400,1421]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
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<emphasis id="D8A8EAEAFFF92A00DCE7FA86882BB599" box="[662,796,1400,1421]" italics="true" pageId="5" pageNumber="156">Heteroponera</emphasis>
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</taxonomicName>
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species,
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<taxonomicName id="2DDC4D7BFFF92A00DD02FA8688F8B599" box="[883,975,1399,1421]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="brouni">
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<emphasis id="D8A8EAEAFFF92A00DD02FA8688F8B599" box="[883,975,1399,1421]" italics="true" pageId="5" pageNumber="156">H. brouni</emphasis>
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</taxonomicName>
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)................
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<taxonomicName id="2DDC4D7BFFF92A00DAD6FA868E26B599" ID-CoL="3LCK3" box="[1191,1297,1399,1421]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="imbellis">
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<emphasis id="D8A8EAEAFFF92A00DAD6FA868E26B599" box="[1191,1297,1399,1421]" italics="true" pageId="5" pageNumber="156">H. imbellis</emphasis>
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</taxonomicName>
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species-group
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</keyLead>
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</paragraph>
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<paragraph id="EA6336F8FFF92A00DEE6FA6B8EACB656" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
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<keyLead id="512D95CDFFF92A00DEE6FA6B8EACB656" pageId="5" pageNumber="156">
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- Generally larger species (HW almost always exceeding
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<quantity id="2D249B1DFFF92A00DD7BFA6B880BB5BE" box="[778,828,1429,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="156" unit="mm" value="1.0">1mm</quantity>
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(rarely as low as
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<quantity id="2D249B1DFFF92A00DD80FA6B8F72B5BE" box="[1009,1093,1429,1450]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="5" pageNumber="156" unit="mm" value="0.92">0.92mm</quantity>
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). Second gastral segment in dorsal view notably narrower than first—often tapering posteriorly (
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<figureCitation id="72E72A7DFFF92A00DD5DFA4C88AEB5D3" box="[812,921,1458,1479]" captionStart="FIGURES 10 – 13" captionStartId="11.[151,264,1954,1976]" captionTargetBox="[195,1392,193,1931]" captionTargetId="figure@11.[195,1392,193,1933]" captionTargetPageId="11" captionText="FIGURES 10 – 13. Heteroponera rhodopygea sp. n., holotype worker, standard views (HW = 1.27 mm, PW = 0.91 mm, WL = 1.55, GW = 1.02 mm)." httpUri="https://zenodo.org/record/398957/files/figure.png" pageId="5" pageNumber="156">Figs 12, 13</figureCitation>
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,
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<figureCitation id="72E72A7DFFF92A00DDD2FA4C88D5B5D3" box="[931,994,1458,1479]" captionStart="FIGURES 34 – 37" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[179,1406,193,1930]" captionTargetId="figure@21.[179,1407,193,1931]" captionTargetPageId="21" captionText="FIGURES 34 – 37. Heteroponera lioprocta sp. n., holotype worker, standard views (HW = 1.06 mm, PW = 0.79 mm, WL = 1.23 mm, GW = 0.99 mm)." httpUri="https://zenodo.org/record/398963/files/figure.png" pageId="5" pageNumber="156">36, 37</figureCitation>
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). Antennal scrobes and frontal carinae usually strongly developed (
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<figureCitation id="72E72A7DFFF92A00DFD4FA318ACBB5F0" box="[421,508,1487,1508]" captionStart="FIGURES 2 – 5" captionStartId="8.[151,264,1953,1975]" captionTargetBox="[189,1397,196,1931]" captionTargetId="figure@8.[189,1397,193,1931]" captionTargetPageId="8" captionText="FIGURES 2 – 5. Heteroponera relicta (Wheeler), worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68 mm); Figs 3 – 5 Black Mountain Rd, Kuranda (HW = 1.27 mm, PW = 0.91 mm, GW = 1.05 mm)." httpUri="https://zenodo.org/record/398955/files/figure.png" pageId="5" pageNumber="156">Figs 2, 3</figureCitation>
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); the carinae vestigial in several species, in which the scrobal areas lack an enclosing carina but are differentiated from the frons as relatively finely-sculptured tracts above the eyes (
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<figureCitation id="72E72A7DFFF92A00DA7BFA138F4EB616" box="[1034,1145,1517,1538]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="5" pageNumber="156">Figs 42, 43</figureCitation>
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). A species-group known only from NE coastal Queensland and adjacent highlands south from Cooktown to Cannonvale, between 15°S and
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<geoCoordinate id="8FE8503FFFF92A00DB73F9F08E65B637" box="[1282,1362,1546,1572]" direction="south" orientation="latitude" pageId="5" pageNumber="156" precision="925" value="-20.5">20°30'S</geoCoordinate>
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. (
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<taxonomicName id="2DDC4D7BFFF92A00DBF5F9F08A28B656" ID-CoL="3LCKB" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="relicta">
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<emphasis id="D8A8EAEAFFF92A00DBF5F9F08EA2B637" box="[1412,1429,1550,1571]" italics="true" pageId="5" pageNumber="156">H</emphasis>
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.
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<emphasis id="D8A8EAEAFFF92A00DE90F9D28A28B656" box="[225,287,1580,1602]" italics="true" pageId="5" pageNumber="156">relicta</emphasis>
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</taxonomicName>
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species group)................................................................................. 3
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</keyLead>
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</paragraph>
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</keyStep>
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3(2) Antennal scrobes fully configured—each almost completely enclosed frontally and posteriorly by a distinct extension of the frontal carina and by a similar ventral carina adjacent to the eye. Scrobe partly divided longitudinally by a median dividing carina. Gastral dorsum (usually much-) less coarsely sculptured than mesosomal dorsum (
|
||
<figureCitation id="72E72A7DFFF92A00DA30F97B8F98B68E" box="[1089,1199,1669,1690]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,264,1953,1975]" captionTargetBox="[198,1388,193,1931]" captionTargetId="figure@18.[198,1388,193,1932]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Heteroponera monteithi sp. n., holotype worker, standard views (HW = 0.93 mm, PW = 0.66 mm, WL = 1.12, GW = 0.90 mm)." httpUri="https://zenodo.org/record/398961/files/figure.png" pageId="5" pageNumber="156">Figs 26, 27</figureCitation>
|
||
)(
|
||
<taxonomicName id="2DDC4D7BFFF92A00DACFF97B8E2EB68E" ID-CoL="3LCKB" box="[1214,1305,1668,1690]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis id="D8A8EAEAFFF92A00DACFF97B8E2EB68E" box="[1214,1305,1668,1690]" italics="true" pageId="5" pageNumber="156">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
species complex)............................................................................................... 4.
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFF92A00DEE6F9418EABB738" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
|
||
<keyLead id="512D95CDFFF92A00DEE6F9418EABB738" pageId="5" pageNumber="156">
|
||
- Antennal scrobes vestigial, represented on each side by a shallowly concave tract above the eye differentiated from the frons by relatively light sculpturation; the enclosing frontal carinae barely defined; scrobal dividing carina vestigially represented or lacking. Body tagmata including the gastral dorsum almost uniformly very coarsely foveolate-rugose (
|
||
<figureCitation id="72E72A7DFFF92A00DAB4F9048E01B71B" box="[1221,1334,1786,1807]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="5" pageNumber="156">Figs 42, 43</figureCitation>
|
||
)(
|
||
<taxonomicName id="2DDC4D7BFFF92A00DB35F9048A3BB739" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="ecarinata">
|
||
<emphasis id="D8A8EAEAFFF92A00DB35F9048A3BB739" italics="true" pageId="5" pageNumber="156">H. ecarinata</emphasis>
|
||
</taxonomicName>
|
||
species complex)................................................................................ 10.
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFF92A03DEE6F8CB8EACB12B" lastPageId="6" lastPageNumber="157" pageId="5" pageNumber="156">
|
||
<paragraph id="EA6336F8FFF92A00DEE6F8CB8EACB7C8" blockId="5.[151,1436,1047,2012]" pageId="5" pageNumber="156">
|
||
<keyLead id="512D95CDFFF92A00DEE6F8CB8EACB7C8" pageId="5" pageNumber="156">
|
||
4(3) Vertexal (=occipital) border of head in square full-face view distinctly emarginate, never squarely transverse. Dorsal surfaces of head, mesosoma and node coarsely, sharply punctate-rugose; first gastral tergite with somewhat polished, effaced large punctures, readily interpreted as a somewhat reduced, smoothed-over version of the stronger, more clearly incised dorsal propodeal sculpturing but less defined and well-marked. Declivitous face of propodeum smooth and strongly shining, with no or very little trace of sculpturation; its lateral borders sculpturally defined, but lacking extra sculptural elements such as a minutely ragged or serrate edge or propodeal-spine-like extensions (
|
||
<figureCitation id="72E72A7DFFF92A00DD1FF839889AB7C8" box="[878,941,1991,2012]" captionStart="FIGURES 2 – 5" captionStartId="8.[151,264,1953,1975]" captionTargetBox="[189,1397,196,1931]" captionTargetId="figure@8.[189,1397,193,1931]" captionTargetPageId="8" captionText="FIGURES 2 – 5. Heteroponera relicta (Wheeler), worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68 mm); Figs 3 – 5 Black Mountain Rd, Kuranda (HW = 1.27 mm, PW = 0.91 mm, GW = 1.05 mm)." httpUri="https://zenodo.org/record/398955/files/figure.png" pageId="5" pageNumber="156">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="72E72A7DFFF92A00DDC6F83988E7B7C8" box="[951,976,1991,2012]" captionStart="FIGURES 10 – 13" captionStartId="11.[151,264,1954,1976]" captionTargetBox="[195,1392,193,1931]" captionTargetId="figure@11.[195,1392,193,1933]" captionTargetPageId="11" captionText="FIGURES 10 – 13. Heteroponera rhodopygea sp. n., holotype worker, standard views (HW = 1.27 mm, PW = 0.91 mm, WL = 1.55, GW = 1.02 mm)." httpUri="https://zenodo.org/record/398957/files/figure.png" pageId="5" pageNumber="156">10</figureCitation>
|
||
).................................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FF698EACB12B" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FF698EACB12B" pageId="6" pageNumber="157">- Vertexal border either as above, or more usually slightly emarginate or squarely transverse—the latter condition conclusive here. Sculpturation differing from the alternative: sculpture of first gastral tergite clearly dissimilar to that of pronotal and propodeal dorsa, always (and usually very distinctly) less strongly developed—ranging from smooth and shining with minute piligerous punctures, to sub-opaque, with very effaced scattered medium puncturation. Declivitous face of propodeum often with superficial sculpturation—shagreening or radial, dorsally-directed effaced striae; the lateral borders usually, but not always, defined by a jagged sculptural edge................................................................ 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FEB98EABB219" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FEB98EACB1A0" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FEB98EACB1A0" pageId="6" pageNumber="157">
|
||
5(4) Body tagmata uniformly dark chocolate-brown; mandibles, antennae and legs dark reddish-brown. Size relatively large: worker HW
|
||
<quantity id="2D249B1DFFFA2A03DF2DFE9B8AD6B16E" box="[348,481,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="1.335" metricValueMax="1.47" metricValueMin="1.2" pageId="6" pageNumber="157" unit="mm" value="1.335" valueMax="1.47" valueMin="1.2">1.20–1.47mm</quantity>
|
||
(mean
|
||
<quantity id="2D249B1DFFFA2A03DC5DFE9B8948B16E" box="[556,639,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" pageId="6" pageNumber="157" unit="mm" value="1.36">1.36mm</quantity>
|
||
), with ca 40% of specimens larger (by HW measurement) than the alternative
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DBF5FE9B8A64B183" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="rhodopygea">
|
||
<emphasis id="D8A8EAEAFFFA2A03DBF5FE9B8A64B183" italics="true" pageId="6" pageNumber="157">H. rhodopygea</emphasis>
|
||
</taxonomicName>
|
||
, which has maximum known HW
|
||
<quantity id="2D249B1DFFFA2A03DCD5FE7C89C1B183" box="[676,758,386,407]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="6" pageNumber="157" unit="mm" value="1.39">1.39mm</quantity>
|
||
. Distribution more southerly (known from Endemism Areas 4, 6, 7, 8, 9, 10, 11, 12 and 13)(
|
||
<figureCitation id="72E72A7DFFFA2A03DFDAFE618935B1A0" box="[427,514,415,436]" captionStart="FIGURES 2 – 5" captionStartId="8.[151,264,1953,1975]" captionTargetBox="[189,1397,196,1931]" captionTargetId="figure@8.[189,1397,193,1931]" captionTargetPageId="8" captionText="FIGURES 2 – 5. Heteroponera relicta (Wheeler), worker, standard views: Fig 2, Lake Eacham NP (WL = 1.68 mm); Figs 3 – 5 Black Mountain Rd, Kuranda (HW = 1.27 mm, PW = 0.91 mm, GW = 1.05 mm)." httpUri="https://zenodo.org/record/398955/files/figure.png" pageId="6" pageNumber="157">Figs 2–5</figureCitation>
|
||
)...........................................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DAA9FE5E8EACB1A0" ID-CoL="3LCKB" authority="Wheeler" authorityName="Wheeler" box="[1240,1435,415,437]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis id="D8A8EAEAFFFA2A03DAA9FE5E8E02B1A1" box="[1240,1333,415,437]" italics="true" pageId="6" pageNumber="157">H. relicta</emphasis>
|
||
(Wheeler)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FE438EABB219" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FE438EABB219" pageId="6" pageNumber="157">
|
||
- Bicolored: head, mesosoma and node dark reddish-brown: the gaster contrasting orange brown, matching the mandibles, antennae and legs. Workers smaller on average: HW
|
||
<quantity id="2D249B1DFFFA2A03DCABFE248868B1FB" box="[730,863,474,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.285" metricValueMax="1.39" metricValueMin="1.18" pageId="6" pageNumber="157" unit="mm" value="1.285" valueMax="1.39" valueMin="1.18">1.18–1.39mm</quantity>
|
||
(mean
|
||
<quantity id="2D249B1DFFFA2A03DDD7FE2488CEB1FB" box="[934,1017,474,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.32" pageId="6" pageNumber="157" unit="mm" value="1.32">1.32mm</quantity>
|
||
), HW not known to exceed
|
||
<quantity id="2D249B1DFFFA2A03DB73FE248E63B1FB" box="[1282,1364,474,495]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" pageId="6" pageNumber="157" unit="mm" value="1.39">1.39mm</quantity>
|
||
. Distribution more northerly (Endemism Areas 1, 2 and 3)(
|
||
<figureCitation id="72E72A7DFFFA2A03DCA6FE098871B218" box="[727,838,503,524]" captionStart="FIGURES 10 – 13" captionStartId="11.[151,264,1954,1976]" captionTargetBox="[195,1392,193,1931]" captionTargetId="figure@11.[195,1392,193,1933]" captionTargetPageId="11" captionText="FIGURES 10 – 13. Heteroponera rhodopygea sp. n., holotype worker, standard views (HW = 1.27 mm, PW = 0.91 mm, WL = 1.55, GW = 1.02 mm)." httpUri="https://zenodo.org/record/398957/files/figure.png" pageId="6" pageNumber="157">Figs 10–13</figureCitation>
|
||
)...............................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DAA1FE068E57B219" box="[1232,1376,503,525]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="rhodopygea" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DAA1FE068E57B219" box="[1232,1376,503,525]" italics="true" pageId="6" pageNumber="157">H. rhodopygea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB17FE098EABB219" box="[1382,1436,503,525]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB17FE098EABB219" bold="true" box="[1382,1436,503,525]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FDEB8EACB28B" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FDEB8EABB271" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FDEB8EABB271" pageId="6" pageNumber="157">
|
||
6(4) First gastral tergite shining, but without smooth areas; closely sown with dense fine puncturation, with a few very weak vestiges of small foveate punctures. Propodeal declivity smooth and shining with its lateral borders sculpturally unelaborated (Known only from EA16, in the
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DC6BFDB18955B270" ID-CoL="8XNJJ" box="[538,610,591,612]" class="Insecta" family="Staphylinidae" genus="Paluma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="165" phylum="Arthropoda" rank="genus">Paluma</taxonomicName>
|
||
area)...........................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DADBFDAE8E68B271" box="[1194,1375,591,613]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="darlingtonorum" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DADBFDAE8E68B271" box="[1194,1375,591,613]" italics="true" pageId="6" pageNumber="157">H. darlingtonorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB17FDB18EABB271" box="[1382,1436,591,613]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB17FDB18EABB271" bold="true" box="[1382,1436,591,613]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FD938EACB28B" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FD938EACB28B" pageId="6" pageNumber="157">- First gastral tergite moderately strongly shining, either virtually lacking sculpturation, or smooth, with scattered point-punctures and somewhat obscure, separated, smoothed-over larger foveate punctures. (Including species from most other EAs).. 7</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FD598EACB3B3" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FD598EACB35B" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FD598EACB35B" pageId="6" pageNumber="157">
|
||
7(6) Exposed section of second gastral tergite (abdominal 4) on each side near its base with an unusual, minute, anterolaterallydirected denticle (or “gastral II spur” —see description in text above) which may extend beyond the outline of the gaster in dorsal view (
|
||
<figureCitation id="72E72A7DFFFA2A03DF13FD1C8AE1B2E3" box="[354,470,738,759]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,264,1953,1975]" captionTargetBox="[198,1388,193,1931]" captionTargetId="figure@18.[198,1388,193,1932]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Heteroponera monteithi sp. n., holotype worker, standard views (HW = 0.93 mm, PW = 0.66 mm, WL = 1.12, GW = 0.90 mm)." httpUri="https://zenodo.org/record/398961/files/figure.png" pageId="6" pageNumber="157">Figs 26, 28</figureCitation>
|
||
,
|
||
<figureCitation id="72E72A7DFFFA2A03DF93FD1C8913B2E3" box="[482,548,738,759]" captionStart="FIGURES 34 – 37" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[179,1406,193,1930]" captionTargetId="figure@21.[179,1407,193,1931]" captionTargetPageId="21" captionText="FIGURES 34 – 37. Heteroponera lioprocta sp. n., holotype worker, standard views (HW = 1.06 mm, PW = 0.79 mm, WL = 1.23 mm, GW = 0.99 mm)." httpUri="https://zenodo.org/record/398963/files/figure.png" pageId="6" pageNumber="157">34, 36</figureCitation>
|
||
). First gastral tergite in dorsal view smooth and shining, with scattered minute piligerous point-punctures visible in reflected light, and no trace of larger foveate punctures or other sculpture. Southern species from EAs 17 and 18. (The gastral II spurs may be obscured if the tubulate second gastral exoskeleton is deeply telescoped into the first gastral segment.)................................................................................. 9
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FCA98EACB3B3" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FCA98EACB3B3" pageId="6" pageNumber="157">- Lateral spurs lacking from second gastral tergite. First gastral tergite in dorsal view with relatively strong point punctures and somewhat obscure scattered foveate punctures, or densely microsculptured and subopaque. Northern species from EAs 2, 3 or 4.................................................................................................. 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FC518EABB44C" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FC518EA1B414" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FC518EA1B414" pageId="6" pageNumber="157">
|
||
8(7) Larger species (HW
|
||
<quantity id="2D249B1DFFFA2A03DFD4FC518918B3D0" box="[421,559,943,964]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.31" metricValueMin="1.24" pageId="6" pageNumber="157" unit="mm" value="1.275" valueMax="1.31" valueMin="1.24">1.24–1.31mm</quantity>
|
||
). Mesosomal proportions as in
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DD26FC4E8885B3D1" ID-CoL="3LCKB" box="[855,946,943,965]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="relicta">
|
||
<emphasis id="D8A8EAEAFFFA2A03DD26FC4E8885B3D1" box="[855,946,943,965]" italics="true" pageId="6" pageNumber="157">H. relicta</emphasis>
|
||
</taxonomicName>
|
||
and most other group species (
|
||
<figureCitation id="72E72A7DFFFA2A03DAA8FC518E7EB3D0" box="[1241,1353,943,964]" captionStart-0="FIGURES 18 – 21" captionStart-1="FIGURES 22 – 25" captionStartId-0="15.[151,264,1955,1977]" captionStartId-1="17.[151,264,1955,1977]" captionTargetBox-0="[196,1390,193,1933]" captionTargetBox-1="[202,1384,193,1932]" captionTargetId-0="figure@15.[196,1390,193,1933]" captionTargetId-1="figure@17.[202,1385,193,1933]" captionTargetPageId-0="15" captionTargetPageId-1="17" captionText-0="FIGURES 18 – 21. Heteroponera wilsoni sp. n., holotype worker, standard views (HW = 1.27 mm, PW = 0.86 mm, WL = 1.55 mm, GW = 1.05)." captionText-1="FIGURES 22 – 25. Heteroponera viviennae sp. n., holotype worker, standard views (HW = 1.11 mm, PW = 0.79 mm, WL = 1.25, GW = 0.80 mm)." httpUri-0="https://zenodo.org/record/398959/files/figure.png" httpUri-1="https://zenodo.org/record/398960/files/figure.png" pageId="6" pageNumber="157">Figs 18 -22</figureCitation>
|
||
). Petiole including its summit generally punctate. Self-colored, dull orange-brown; gaster, antennae and legs slightly paler than elsewhere..................................................................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DA86FC148E6EB3EB" box="[1271,1369,1001,1023]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="wilsoni" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DA86FC148E6EB3EB" box="[1271,1369,1001,1023]" italics="true" pageId="6" pageNumber="157">H. wilsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB11FC148EA1B414" box="[1376,1430,1002,1024]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB11FC148EA1B414" bold="true" box="[1376,1430,1002,1024]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FBF98EABB44C" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FBF98EABB44C" pageId="6" pageNumber="157">
|
||
- Smaller species (HW
|
||
<quantity id="2D249B1DFFFA2A03DFC8FBF98975B408" box="[441,578,1031,1052]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" metricValueMax="1.19" metricValueMin="1.07" pageId="6" pageNumber="157" unit="mm" value="1.13" valueMax="1.19" valueMin="1.07">1.07–1.19mm</quantity>
|
||
). Mesosoma relatively short and compact (
|
||
<figureCitation id="72E72A7DFFFA2A03DD9AFBF98F68B408" box="[1003,1119,1031,1052]" captionStart="FIGURES 22 – 25" captionStartId="17.[151,264,1955,1977]" captionTargetBox="[202,1384,193,1932]" captionTargetId="figure@17.[202,1385,193,1933]" captionTargetPageId="17" captionText="FIGURES 22 – 25. Heteroponera viviennae sp. n., holotype worker, standard views (HW = 1.11 mm, PW = 0.79 mm, WL = 1.25, GW = 0.80 mm)." httpUri="https://zenodo.org/record/398960/files/figure.png" pageId="6" pageNumber="157">Figs 22–25</figureCitation>
|
||
). Petiole generally punctate, but with the summit smooth and shining. Bicolored: head, mesosoma and node bright mahogany- brown, gaster, mandibles, antennae and legs bright reddish/orange-brown...................................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DA97FBBC8E68B443" box="[1254,1375,1090,1111]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="viviennae" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DA97FBBC8E68B443" box="[1254,1375,1090,1111]" italics="true" pageId="6" pageNumber="157">H. viviennae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB17FBBC8EABB44C" box="[1382,1436,1090,1112]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB17FBBC8EABB44C" bold="true" box="[1382,1436,1090,1112]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FBA18EA1B574" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FBA18EA1B4D9" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FBA18EA1B4D9" pageId="6" pageNumber="157">
|
||
9(7) Mesosomal dorsum distinctly punctate-rugose; declivitous face of propodeum smooth and shining, with effaced, very fine, obscure vestigial striae radiating from the center of its base (best seen in reflected light). The declivity separated from the sides and dorsum of the sclerite by a minutely ragged sculptural margin, with no trace of lateral bordering carinae or dorsolateral denticles. (
|
||
<figureCitation id="72E72A7DFFFA2A03DF38FB498A81B4D8" box="[329,438,1207,1228]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,264,1953,1975]" captionTargetBox="[198,1388,193,1931]" captionTargetId="figure@18.[198,1388,193,1932]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Heteroponera monteithi sp. n., holotype worker, standard views (HW = 0.93 mm, PW = 0.66 mm, WL = 1.12, GW = 0.90 mm)." httpUri="https://zenodo.org/record/398961/files/figure.png" pageId="6" pageNumber="157">Figs 26–29</figureCitation>
|
||
)(Known only from the Conway Range area (EA18) near Cannonvale)...........
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DA93FB468E6DB4D9" box="[1250,1370,1207,1229]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DA93FB468E6DB4D9" box="[1250,1370,1207,1229]" italics="true" pageId="6" pageNumber="157">H. monteithi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB11FB498EA1B4D9" box="[1376,1430,1207,1229]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB11FB498EA1B4D9" bold="true" box="[1376,1430,1207,1229]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FB2B8EA1B574" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FB2B8EA1B574" pageId="6" pageNumber="157">
|
||
<emphasis id="D8A8EAEAFFFA2A03DEE6FB2B8BA8B4FE" box="[151,159,1237,1258]" italics="true" pageId="6" pageNumber="157">-</emphasis>
|
||
Mesosomal dorsum less-distinctly ornamented, with somewhat effaced sculpture, including vaguely longitudinal elements; declivitous face of propodeum smooth and shining, lacking the fine sculpturing described above, or with only faint vestiges of similar sculpture. The declivity bordered on each side by a fine carina, which is extended dorsally to form a small obtusely triangular denticle somewhat resembling a minute propodeal tooth. (
|
||
<figureCitation id="72E72A7DFFFA2A03DD11FAD388FAB556" box="[864,973,1325,1346]" captionStart="FIGURES 34 – 37" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[179,1406,193,1930]" captionTargetId="figure@21.[179,1407,193,1931]" captionTargetPageId="21" captionText="FIGURES 34 – 37. Heteroponera lioprocta sp. n., holotype worker, standard views (HW = 1.06 mm, PW = 0.79 mm, WL = 1.23 mm, GW = 0.99 mm)." httpUri="https://zenodo.org/record/398963/files/figure.png" pageId="6" pageNumber="157">Figs 34–37</figureCitation>
|
||
)(Known only from sites near the summit of Mt Elliot (EA17) southeast of Townsville).......................................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DA95FAB48E6EB54B" box="[1252,1369,1353,1375]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="lioprocta" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DA95FAB48E6EB54B" box="[1252,1369,1353,1375]" italics="true" pageId="6" pageNumber="157">H. lioprocta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB11FAB48EA1B574" box="[1376,1430,1354,1376]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB11FAB48EA1B574" bold="true" box="[1376,1430,1354,1376]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6FA998EA2B65E" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FA998EA1B5C1" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FA998EA1B5C1" pageId="6" pageNumber="157">
|
||
10(3) Body tagmata generally overall dull medium- to dark-brown; often bicolored, with the gaster dull light reddish-brown. Pronotal epaulets relatively less developed (
|
||
<figureCitation id="72E72A7DFFFA2A03DC3DFA7B898AB58E" box="[588,701,1413,1434]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="6" pageNumber="157">Figs 42–45</figureCitation>
|
||
). Frons very finely longitudinally striate, with several larger additional striae on each side, each of the larger elements about as distinct as the median frontal carina (
|
||
<figureCitation id="72E72A7DFFFA2A03DA6AFA5C8F6AB5A3" box="[1051,1117,1442,1463]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="6" pageNumber="157">Fig 43</figureCitation>
|
||
). Known only from the far northern Endemicity Areas EA1 (Mt Finnigan) and EA2 (Thornton Peak)...............................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DAACFA3E8E6FB5C1" box="[1245,1368,1471,1493]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="trachypyx" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DAACFA3E8E6FB5C1" box="[1245,1368,1471,1493]" italics="true" pageId="6" pageNumber="157">H. trachypyx</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB11FA418EA1B5C1" box="[1376,1430,1471,1493]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB11FA418EA1B5C1" bold="true" box="[1376,1430,1471,1493]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6FA238EA2B65E" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6FA238EA2B65E" pageId="6" pageNumber="157">
|
||
- Body tagmata uniformly very dark blackish-brown, almost black; legs and antennae lighter, medium to dark reddish-brown. Pronotal -epaulets relatively strongly developed (
|
||
<figureCitation id="72E72A7DFFFA2A03DCB9FA04880CB61B" box="[712,827,1530,1551]" captionStart="FIGURES 46 – 49" captionStartId="26.[151,264,1952,1974]" captionTargetBox="[200,1386,193,1931]" captionTargetId="figure@26.[200,1387,193,1931]" captionTargetPageId="26" captionText="FIGURES 46 – 49. Heteroponera ecarinata sp. n., paratype worker, standard views (HW = 1.23 mm, PW = 0.91 mm, WL = 1.55 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398966/files/figure.png" pageId="6" pageNumber="157">Figs 46–49</figureCitation>
|
||
,
|
||
<figureCitation id="72E72A7DFFFA2A03DD36FA0488B0B61B" box="[839,903,1530,1551]" captionStart="FIGURES 50 – 53" captionStartId="27.[151,264,1954,1976]" captionTargetBox="[198,1387,193,1932]" captionTargetId="figure@27.[198,1388,193,1933]" captionTargetPageId="27" captionText="FIGURES 50 – 53. Heteroponera pendergrasti sp. n., holotype worker, standard views (HW = 1.23 mm, PW = 0.86 mm, WL = 1.54 mm, GW = 0.99 mm)." httpUri="https://zenodo.org/record/398967/files/figure.png" pageId="6" pageNumber="157">50–53</figureCitation>
|
||
). Sculpturation of frons otherwise—either moderately coarsely longitudinally striate without intervening fine striae, or much less regularly sculptured, as in
|
||
<figureCitation id="72E72A7DFFFA2A03DAA1F9E98E24B638" box="[1232,1299,1559,1580]" captionStart="FIGURES 42 – 45" captionStartId="24.[151,264,1956,1978]" captionTargetBox="[193,1394,195,1933]" captionTargetId="figure@24.[193,1394,195,1934]" captionTargetPageId="24" captionText="FIGURES 42 – 45. Heteroponera trachypyx sp. n., holotype worker, standard views (HW = 1.18 mm, PW = 0.95 mm, WL = 1.40 mm, GW = 1.04 mm)." httpUri="https://zenodo.org/record/398965/files/figure.png" pageId="6" pageNumber="157">Fig 43</figureCitation>
|
||
. Known from more southerly Endemicity Areas........................................................................11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="51282E5DFFFA2A03DEE6F9AC8EABB6EE" pageId="6" pageNumber="157">
|
||
<paragraph id="EA6336F8FFFA2A03DEE6F9AC8EA1B6B6" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6F9AC8EA1B6B6" pageId="6" pageNumber="157">
|
||
11(10) Petiolar dorsum generally rounded in lateral view, sometimes slightly extended posteroapically, but never as strongly as the alternative. Declivitous face of propodeum with dorsally-directed radiating striae on a shining background, with or without accompanying shagreening. Known from EA8, EA10 & EA12...................................
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DA91F9738E6EB6B6" box="[1248,1369,1677,1698]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="ecarinata" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DA91F9738E6EB6B6" box="[1248,1369,1677,1698]" italics="true" pageId="6" pageNumber="157">H. ecarinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB11F9728EA1B6B6" box="[1376,1430,1676,1698]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB11F9728EA1B6B6" bold="true" box="[1376,1430,1676,1698]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="EA6336F8FFFA2A03DEE6F9548EABB6EE" blockId="6.[151,1437,151,1786]" pageId="6" pageNumber="157">
|
||
<keyLead id="512D95CDFFFA2A03DEE6F9548EABB6EE" pageId="6" pageNumber="157">
|
||
- Petiolar dorsum relatively tall and narrow in lateral view, extended above to form a distinct broadly tooth-like posteroapical process. Declivitous face of propodeum smooth and shining, with at most only vague traces of dorsally-directed radial sculpturation. Known only from EA7 (Lamb Range)............................................. ..
|
||
<taxonomicName id="2DDC4D7BFFFA2A03DAB8F91B8E57B6EE" box="[1225,1376,1764,1786]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="pendergrasti" status="sp. nov.">
|
||
<emphasis id="D8A8EAEAFFFA2A03DAB8F91B8E57B6EE" box="[1225,1376,1764,1786]" italics="true" pageId="6" pageNumber="157">H. pendergrasti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C39B5791FFFA2A03DB17F91A8EABB6EE" box="[1382,1436,1764,1786]" pageId="6" pageNumber="157" rank="species">
|
||
<emphasis id="D8A8EAEAFFFA2A03DB17F91A8EABB6EE" bold="true" box="[1382,1436,1764,1786]" pageId="6" pageNumber="157">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |