270 lines
39 KiB
XML
270 lines
39 KiB
XML
<document id="7E0756DAC80FCD60824BD91EBED03C05" ID-CLB-Dataset="51371" ID-DOI="10.5281/zenodo.156420" ID-GBIF-Dataset="76c096e3-d27c-4abc-a504-a62ae8c7ba92" ID-ISSN="11755326" ID-Zenodo-Dep="156420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459586556453" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="62088784C37CC725E12E75AAFE32FE36" docLanguage="en" docName="zt00142.pdf" docOrigin="Zootaxa 142" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Seeversiella microphthalma Gusarov, sp. n." docType="treatment" docVersion="10" lastPageNumber="86" masterDocId="9E31FFFCC32EC770E0267411FFBAFFC8" masterDocTitle="A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="102" masterPageNumber="1" pageNumber="83" updateTime="1698203169024" updateUser="ExternalLinkService">
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<mods:titleInfo id="6A230BD036F0AC561AF3C50BEDAE9C5D">
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<mods:title id="339DC6DAC4FB7A7B10F7215AA228C707">A revision of the genus Seeversiella Ashe, 1986 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="0A0EA07BAEE9272B828FC720840362D7">Gusarov, Vladimir I.</mods:namePart>
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<mods:title id="FBE4DCE9A9DCBEA3A426B4E83F5E9E1E">Zootaxa</mods:title>
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<mods:part id="92ED02299FBEE67B712A94A64CC37054">
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<mods:date id="70DC9AD41019FDE3F8D040BACE2C97DD">2003</mods:date>
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<mods:identifier id="A6E944E4A7751315C59955C3662F546B" type="CLB-Dataset">51371</mods:identifier>
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<mods:identifier id="D980EF10431AD1E20A67E9039346204F" type="DOI">10.5281/zenodo.156420</mods:identifier>
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<treatment id="62088784C37CC725E12E75AAFE32FE36" ID-DOI="http://doi.org/10.5281/zenodo.6274689" ID-Zenodo-Dep="6274689" LSID="urn:lsid:plazi:treatment:62088784C37CC725E12E75AAFE32FE36" httpUri="http://treatment.plazi.org/id/62088784C37CC725E12E75AAFE32FE36" lastPageId="85" lastPageNumber="86" pageId="82" pageNumber="83">
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<subSubSection id="A2BB6519C37CC722E12E75AAFBBFFE1D" box="[264,1029,443,470]" pageId="82" pageNumber="83" type="key">
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<paragraph id="EA1E3692C37CC722E12E75AAFBBFFE1D" blockId="82.[264,1029,443,470]" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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<heading id="B15681FEC37CC722E12E75AAFBBFFE1D" bold="true" box="[264,1029,443,470]" fontSize="11" level="1" pageId="82" pageNumber="83" reason="1">
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<emphasis id="D8D5EA80C37CC722E12E75AAFBBFFE1D" bold="true" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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<keyStep id="51552E37C37CC722E12E75AAFBBFFE1D" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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<key id="9E3061A3C37CC722E12E75AAFBBFFE1D" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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<keyLead id="515095A7C37CC722E12E75AAFBBFFE1D" box="[264,1029,443,470]" pageId="82" pageNumber="83">
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24.
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<taxonomicName id="2DA14D11C37CC722E11275ADFD4DFE1D" ID-CoL="6YBCX" authority="Gusarov" authorityName="Gusarov" box="[308,759,443,470]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
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<emphasis id="D8D5EA80C37CC722E11275ADFD3FFE1E" bold="true" box="[308,645,444,470]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
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Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="C3E657FBC37CC722E32275AAFCFCFE1D" box="[772,838,443,469]" pageId="82" pageNumber="83" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="729A2A17C37CC722E37375AAFC47FE1D" box="[853,1021,443,469]" captionStart-0="FIGURES 331 334" captionStart-1="FIGURES 335 343" captionStartId-0="82.[264,383,1659,1683]" captionStartId-1="83.[264,383,1660,1684]" captionTargetBox-0="[274,1308,735,1617]" captionTargetBox-1="[283,1301,504,1615]" captionTargetId-0="figure@82.[264,1323,724,1632]" captionTargetId-1="figure@83.[270,1318,489,1633]" captionTargetPageId-0="82" captionTargetPageId-1="83" captionText-0="FIGURES 331 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 332, holotype; 333 334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm." captionText-1="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri-0="https://zenodo.org/record/156478/files/figure.png" httpUri-1="https://zenodo.org/record/156479/files/figure.png" pageId="82" pageNumber="83">Figs. 331343</figureCitation>
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)
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</keyLead>
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</key>
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</keyStep>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A2BB6519C37CC722E12E7617FB42FDB9" pageId="82" pageNumber="83" type="materials_examined">
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<paragraph id="EA1E3692C37CC722E12E7617FC7FFD81" blockId="82.[264,1323,518,625]" pageId="82" pageNumber="83">
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<emphasis id="D8D5EA80C37CC722E12E7617FE04FDE8" bold="true" box="[264,446,518,544]" pageId="82" pageNumber="83">
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<typeStatus id="351A8830C37CC722E12E7617FEF9FDE8" box="[264,323,518,544]" pageId="82" pageNumber="83">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="351A8830C37CC722E1F27616FDFEFDE9" box="[468,580,519,545]" pageId="82" pageNumber="83" type="holotype">Holotype</typeStatus>
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,,
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<emphasis id="D8D5EA80C37CC722E2507617FC78FDE8" bold="true" box="[630,962,518,544]" pageId="82" pageNumber="83">
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<collectingCountry id="92B67602C37CC722E2507617FC8BFDE8" box="[630,817,518,544]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
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: San Jose:
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</emphasis>
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<quantity id="2D599B77C37CC722E3F07616FBA9FDE9" box="[982,1043,519,545]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="82" pageNumber="83" unit="km" value="2.0">2 km</quantity>
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S Villa Mills,
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<quantity id="2D599B77C37CC722E4EE7616FA9CFDE9" box="[1224,1318,519,545]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="82" pageNumber="83" unit="m" value="3000.0">3000 m</quantity>
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, ridgetop oak forest litter (R.Anderson),
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<date id="9E1F1052C37CC722E2FC763EFCEBFD81" box="[730,849,559,585]" pageId="82" pageNumber="83" value="1996-02-10">10.ii.1996</date>
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(
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<collectionCode id="8CB0AE57C37CC722E347763EFC03FD81" box="[865,953,559,585]" pageId="82" pageNumber="83">KSEM</collectionCode>
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).
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</paragraph>
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<paragraph id="EA1E3692C37CC722E11E7646FB42FDB9" blockId="82.[264,1323,518,625]" box="[312,1272,598,625]" pageId="82" pageNumber="83">
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<typeStatus id="351A8830C37CC722E11E7646FE14FDB9" box="[312,430,599,625]" pageId="82" pageNumber="83" type="paratype">Paratypes</typeStatus>
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:
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<emphasis id="D8D5EA80C37CC722E1E67647FCB9FDB8" bold="true" box="[448,771,598,624]" pageId="82" pageNumber="83">
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<collectingCountry id="92B67602C37CC722E1E67647FDC0FDB8" box="[448,634,598,624]" name="Costa Rica" pageId="82" pageNumber="83">COSTA RICA</collectingCountry>
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: San Jose:
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</emphasis>
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8, same data as the
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<typeStatus id="351A8830C37CC722E43B7646FB3EFDB9" box="[1053,1156,599,625]" pageId="82" pageNumber="83" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="8CB0AE57C37CC722E4B27646FB56FDB9" box="[1172,1260,599,625]" pageId="82" pageNumber="83">KSEM</collectionCode>
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).
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</paragraph>
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</subSubSection>
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||
<caption id="BEDE661AC37CC722E12E726AFBEAF91F" httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="82" pageNumber="83" targetBox="[274,1308,735,1617]" targetPageId="82">
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<paragraph id="EA1E3692C37CC722E12E726AFBEAF91F" blockId="82.[264,1323,1659,1751]" pageId="82" pageNumber="83">
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<emphasis id="D8D5EA80C37CC722E12E726AFE5AF95B" bold="true" box="[264,480,1659,1683]" pageId="82" pageNumber="83">FIGURES 331334</emphasis>
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. Abdominal segment 8 of
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<taxonomicName id="2DA14D11C37CC722E329726AFB27F95A" authority="Gusarov" authorityName="Gusarov" box="[783,1181,1659,1682]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
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<emphasis id="D8D5EA80C37CC722E329726AFB80F95A" box="[783,1082,1659,1682]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
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Gusarov
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</taxonomicName>
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,
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<taxonomicNameLabel id="C3E657FBC37CC722E48C726AFB52F95B" box="[1194,1256,1659,1683]" pageId="82" pageNumber="83" rank="species">
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<emphasis id="D8D5EA80C37CC722E48C726AFB52F95B" bold="true" box="[1194,1256,1659,1683]" pageId="82" pageNumber="83">sp. n.</emphasis>
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</taxonomicNameLabel>
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(331 332, holotype; 333334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm.
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</paragraph>
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||
</caption>
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||
<subSubSection id="A2BB6519C37CC724E11E733BFEFFFE1D" lastPageId="84" lastPageNumber="85" pageId="82" pageNumber="83" type="diagnosis">
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||
<paragraph id="EA1E3692C37CC723E11E733BFD37FE66" blockId="82.[264,1323,1834,1901]" lastBlockId="83.[264,1324,284,430]" lastPageId="83" lastPageNumber="84" pageId="82" pageNumber="83">
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<emphasis id="D8D5EA80C37CC722E11E733BFE02F88C" bold="true" box="[312,440,1834,1860]" pageId="82" pageNumber="83">Diagnosis.</emphasis>
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<taxonomicName id="2DA14D11C37CC722E1E8733AFCA0F88C" box="[462,794,1835,1860]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="species" species="microphthalma">
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<emphasis id="D8D5EA80C37CC722E1E8733AFCA0F88C" box="[462,794,1835,1860]" italics="true" pageId="82" pageNumber="83">Seeversiella microphthalma</emphasis>
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</taxonomicName>
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can be distinguished from other species of
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<taxonomicName id="2DA14D11C37CC722E12E7342FE2CF8A4" box="[264,406,1875,1900]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="82" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="D8D5EA80C37CC722E12E7342FE2CF8A4" box="[264,406,1875,1900]" italics="true" pageId="82" pageNumber="83">Seeversiella</emphasis>
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</taxonomicName>
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by having uniformly brown body; small eyes (temples 4.04.3 times as long as eyes); elytra much shorter than pronotum; reduced wings (shorter than elytra); by lacking clear medial emargination at the posterior margin of male tergum 8 (
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<figureCitation id="729A2A17C37DC723E47A7555FB79FE96" box="[1116,1219,324,350]" captionStart="FIGURES 331 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 332, holotype; 333 334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="83" pageNumber="84">Fig. 331</figureCitation>
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); tergum 8 with four pairs of macrosetae; by the distinct shape of aedeagus (
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<figureCitation id="729A2A17C37DC723E41A757DFE8FFE66" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84">Figs. 335340, 342 343</figureCitation>
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) and spermatheca (
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<figureCitation id="729A2A17C37DC723E23F7585FD3AFE66" box="[537,640,404,430]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84">Fig. 341</figureCitation>
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).
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</paragraph>
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<caption id="BEDE661AC37DC723E12E726DFB1FF888" httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="83" pageNumber="84" targetBox="[283,1301,504,1615]" targetPageId="83">
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||
<paragraph id="EA1E3692C37DC723E12E726DFB1FF888" blockId="83.[264,1323,1660,1856]" pageId="83" pageNumber="84">
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||
<emphasis id="D8D5EA80C37DC723E12E726DFE65F95C" bold="true" box="[264,479,1660,1684]" pageId="83" pageNumber="84">FIGURES 335343</emphasis>
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||
. Genitalia of
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<taxonomicName id="2DA14D11C37DC723E25B726CFBB0F95B" authority="Gusarov" authorityName="Gusarov" box="[637,1034,1660,1684]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="microphthalma" status="sp. nov.">
|
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<emphasis id="D8D5EA80C37DC723E25B726CFC12F95C" box="[637,936,1661,1684]" italics="true" pageId="83" pageNumber="84">Seeversiella microphthalma</emphasis>
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Gusarov
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||
</taxonomicName>
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||
,
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<taxonomicNameLabel id="C3E657FBC37DC723E430726DFBE9F95C" box="[1046,1107,1660,1684]" pageId="83" pageNumber="84" rank="species">
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||
<emphasis id="D8D5EA80C37DC723E430726DFBE9F95C" bold="true" box="[1046,1107,1660,1684]" pageId="83" pageNumber="84">sp. n.</emphasis>
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||
</taxonomicNameLabel>
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||
(335340, 342343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342343), 0.1 mm (336337, 339, 341).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EA1E3692C37AC724E11E750DFEFFFE1D" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
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||
<taxonomicName id="2DA14D11C37AC724E11E750DFD3CFEFD" box="[312,646,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
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<emphasis id="D8D5EA80C37AC724E11E750DFD3CFEFD" box="[312,646,284,309]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2DA14D11C37AC724E311750DFC44FEFD" box="[823,1022,284,309]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="impressicollis">
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||
<emphasis id="D8D5EA80C37AC724E311750DFC44FEFD" box="[823,1022,284,309]" italics="true" pageId="84" pageNumber="85">S. impressicollis</emphasis>
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</taxonomicName>
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in having less asperate pronotal punctation; in males pronotum without medial impression, posterior angles of male tergum 3 projecting as spines; lateral diverticula of internal sac without long spiniform sclerites (
|
||
<figureCitation id="729A2A17C37AC724E19F7585FD91FE66" box="[441,555,404,430]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 343</figureCitation>
|
||
; 316, 318); Lshaped spermatheca with larger distal portion (
|
||
<figureCitation id="729A2A17C37AC724E4DD7585FE8FFE1E" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Fig. 341</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2BB6519C37AC725E11E75F5FDC4FE4E" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85" type="description">
|
||
<paragraph id="EA1E3692C37AC724E11E75F5FD4AFDBD" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
<taxonomicName id="2DA14D11C37AC724E11E75F5FD3FFE35" box="[312,645,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
|
||
<emphasis id="D8D5EA80C37AC724E11E75F5FD3FFE35" box="[312,645,484,509]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2DA14D11C37AC724E31275F5FC75FE35" box="[820,975,484,509]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="sulcicollis">
|
||
<emphasis id="D8D5EA80C37AC724E31275F5FC75FE35" box="[820,975,484,509]" italics="true" pageId="84" pageNumber="85">S. sulcicollis</emphasis>
|
||
</taxonomicName>
|
||
in having larger body, less asperate pronotal punctation; in males pronotum without medial furrow, posterior angles of male tergum 3 projecting as spines; lateral diverticula of internal sac without strong denticulate sclerites (
|
||
<figureCitation id="729A2A17C37AC724E226764DFDCBFDBE" box="[512,625,604,630]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 343</figureCitation>
|
||
; 326328).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E7695FC07FCDE" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
<taxonomicName id="2DA14D11C37AC724E11E7695FD3BFD55" box="[312,641,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
|
||
<emphasis id="D8D5EA80C37AC724E11E7695FD3BFD55" box="[312,641,644,669]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2DA14D11C37AC724E3047695FC6EFD55" box="[802,980,644,669]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="geostiboides">
|
||
<emphasis id="D8D5EA80C37AC724E3047695FC6EFD55" box="[802,980,644,669]" italics="true" pageId="84" pageNumber="85">S. geostiboides</emphasis>
|
||
</taxonomicName>
|
||
in having pubescence in the lateral portions of pronotum directed towards the midline and obliquely posteriorly; in wider apex of median lobe (in parameral view) (
|
||
<figureCitation id="729A2A17C37AC724E37F76C5FBBEFD26" box="[857,1028,724,750]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 335336</figureCitation>
|
||
; 348349); and different shape of the distal portion of spermatheca (
|
||
<figureCitation id="729A2A17C37AC724E32476EDFCC9FCDE" box="[770,883,764,790]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 341</figureCitation>
|
||
; 355).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E7735FD62FCAE" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
<taxonomicName id="2DA14D11C37AC724E11E7735FDC5FCF5" box="[312,639,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
|
||
<emphasis id="D8D5EA80C37AC724E11E7735FDC5FCF5" box="[312,639,804,829]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2DA14D11C37AC724E33A7735FC31FCF5" box="[796,907,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="flavida">
|
||
<emphasis id="D8D5EA80C37AC724E33A7735FC31FCF5" box="[796,907,804,829]" italics="true" pageId="84" pageNumber="85">S. flavida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2DA14D11C37AC724E3E37735FB8BFCF5" box="[965,1073,804,829]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="adusta">
|
||
<emphasis id="D8D5EA80C37AC724E3E37735FB8BFCF5" box="[965,1073,804,829]" italics="true" pageId="84" pageNumber="85">S. adusta</emphasis>
|
||
</taxonomicName>
|
||
in having abdominal tergum 8 with four pairs of macrosetae.
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E7765FBC2FC7D" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
<taxonomicName id="2DA14D11C37AC724E11E7765FD3EFC45" box="[312,644,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="microphthalma">
|
||
<emphasis id="D8D5EA80C37AC724E11E7765FD3EFC45" box="[312,644,884,909]" italics="true" pageId="84" pageNumber="85">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="2DA14D11C37AC724E3097765FC60FC45" box="[815,986,884,909]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="paramoana">
|
||
<emphasis id="D8D5EA80C37AC724E3097765FC60FC45" box="[815,986,884,909]" italics="true" pageId="84" pageNumber="85">S. paramoana</emphasis>
|
||
</taxonomicName>
|
||
in having concave but not emarginate apex of median lobe (in lateral view) (
|
||
<figureCitation id="729A2A17C37AC724E374778DFC40FC7E" box="[850,1018,924,950]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="84" pageNumber="85">Figs. 338339</figureCitation>
|
||
; 379380).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E77D2FCCBFC16" blockId="84.[264,1324,284,1870]" box="[312,881,963,990]" pageId="84" pageNumber="85">
|
||
<emphasis id="D8D5EA80C37AC724E11E77D2FE6AFC15" bold="true" box="[312,464,963,989]" pageId="84" pageNumber="85">Description.</emphasis>
|
||
Length
|
||
<quantity id="2D599B77C37AC724E21F77D5FD7EFC16" box="[569,708,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="84" pageNumber="85" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 mm</quantity>
|
||
. Body brown.
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E77FDFE3DFBB6" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">Head surface glossy, with fine and weak isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 12 times their diameter. Temples 4.04.3 times as long as eyes. Antennal article 2 as long as 3, article 4 subquadrate, 510 transverse.</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E709DFD2EFA76" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
Pronotum slightly transverse, 1.2 times as wide as head, width
|
||
<quantity id="2D599B77C37AC724E40D709DFB6CFB6E" box="[1067,1238,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.0" metricValueMin="4.6" pageId="84" pageNumber="85" unit="mm" value="0.48" valueMax="0.5" valueMin="0.46">0.460.50 mm</quantity>
|
||
, length
|
||
<quantity id="2D599B77C37AC724E12E70A5FE0BFB06" box="[264,433,1204,1230]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="4.699999999999999" metricValueMin="4.3" pageId="84" pageNumber="85" unit="mm" value="0.44999999999999996" valueMax="0.47" valueMin="0.43">0.430.47 mm</quantity>
|
||
, width to length ratio 1.1, surface glossy, with fine isodiametric microsculpture, with fine and asperate punctation, distance between punctures equals 12 times their diameter. Elytra wider and much shorter (measured from humeral angle) than pronotum (pronotal length to elytral length ratio 1.4), 1.8 times wider than long, glossy, with poorly visible microsculpture, with fine and asperate punctation, distance between punctures equals 11.5 times their diameter, punctures stronger than on pronotum. Wings reduced to short vestiges, shorter than elytra.
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E71DDFE71F966" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
Abdominal terga glossy, terga 35 with microsculpture consisting of transverse meshes, and with fine punctation, distance between punctures equals 24 times their diameter; tergum 7 with microsculpture consisting of transverse meshes and with fine and sparse punctation, distance between punctures equals 35 times their diameter. Apical margin of tergum 7 without white palisade fringe. Tergum 8 with four pairs of macrosetae (
|
||
<figureCitation id="729A2A17C37AC724E1377285FE04F966" box="[273,446,1684,1710]" captionStart="FIGURES 331 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 332, holotype; 333 334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="84" pageNumber="85">Figs. 331, 333</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37AC724E11E72ADFE56F886" blockId="84.[264,1324,284,1870]" pageId="84" pageNumber="85">
|
||
In the only known male the posterior angles of tergum 3 projecting as sharp spines half as long as tergum (measured medially); tergum
|
||
<quantity id="2D599B77C37AC724E31B72F5FCD1F936" box="[829,875,1764,1790]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="84" pageNumber="85" unit="in" value="7.0">7 in</quantity>
|
||
front of posterior margin with medial carina half as long as the tergum; posterior margin of tergum 8 without clear medial emargination (
|
||
<figureCitation id="729A2A17C37AC724E15E7325FE65F886" box="[376,479,1844,1870]" captionStart="FIGURES 331 334" captionStartId="82.[264,383,1659,1683]" captionTargetBox="[274,1308,735,1617]" captionTargetId="figure@82.[264,1323,724,1632]" captionTargetPageId="82" captionText="FIGURES 331 334. Abdominal segment 8 of Seeversiella microphthalma Gusarov, sp. n. (331 332, holotype; 333 334, paratype from Villa Mills, Costa Rica). 331 – male tergum 8; 332 – male sternum 8; 333 – female tergum 8; 334 – female sternum 8. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156478/files/figure.png" pageId="84" pageNumber="85">Fig. 331</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37BC725E11E750DFC19FE96" blockId="85.[264,1324,284,510]" pageId="85" pageNumber="86">
|
||
Aedeagus as in
|
||
<figureCitation id="729A2A17C37BC725E1D9750DFCA5FEFE" box="[511,799,284,310]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Figs. 335340, 342343</figureCitation>
|
||
. Apex of paramere narrow, proximal seta approximately as long as the other three setae (
|
||
<figureCitation id="729A2A17C37BC725E3097555FC2CFE96" box="[815,918,324,350]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Fig. 337</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37BC725E11E757DFDC4FE4E" blockId="85.[264,1324,284,510]" box="[312,638,364,390]" pageId="85" pageNumber="86">
|
||
Spermatheca as in
|
||
<figureCitation id="729A2A17C37BC725E233757DFDC0FE4E" box="[533,634,364,390]" captionStart="FIGURES 335 343" captionStartId="83.[264,383,1660,1684]" captionTargetBox="[283,1301,504,1615]" captionTargetId="figure@83.[270,1318,489,1633]" captionTargetPageId="83" captionText="FIGURES 335 343. Genitalia of Seeversiella microphthalma Gusarov, sp. n. (335 340, 342 343, holotype; 341, paratype from Villa Mills, Costa Rica). 335 – median lobe, parameral view; 336 – apex of median lobe, parameral view; 337 – apex of left paramere; 338 – median lobe, lateral view; 339 – apex of median lobe, lateral view; 340 – details of internal sac retracted into median lobe, abparameral view; 341 spermatheca; 342 343 – details of internal sac retracted into median lobe, lateral view. Scale bar 0.2 mm (335, 338, 340, 342 343), 0.1 mm (336 337, 339, 341)." httpUri="https://zenodo.org/record/156479/files/figure.png" pageId="85" pageNumber="86">Fig. 341</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A2BB6519C37BC725E11E7582FE32FE36" pageId="85" pageNumber="86" type="distribution">
|
||
<paragraph id="EA1E3692C37BC725E11E7582FBCDFE66" blockId="85.[264,1324,284,510]" box="[312,1143,403,430]" pageId="85" pageNumber="86">
|
||
<emphasis id="D8D5EA80C37BC725E11E7582FE62FE65" bold="true" box="[312,472,403,429]" pageId="85" pageNumber="86">Distribution.</emphasis>
|
||
Known from Cerro de la Muerte,
|
||
<collectingCountry id="92B67602C37BC725E3547585FC49FE66" box="[882,1011,404,430]" name="Costa Rica" pageId="85" pageNumber="86">Costa Rica</collectingCountry>
|
||
(
|
||
<figureCitation id="729A2A17C37BC725E4257585FBD0FE66" box="[1027,1130,404,430]" captionStart="FIGURE 389" captionStartId="99.[264,368,1817,1841]" captionTargetBox="[317,1272,317,1805]" captionTargetId="figure@99.[310,1276,313,1806]" captionTargetPageId="99" captionText="FIGURE 389. Geographical distribution of eleven species of Seeversiella (with very short elytra) in the high mountains of Costa Rica. Isohypses 1000 m, 2000 m and 3000 m are shown. Areas above 2000 m are grey and those above 3000 m are solid black." httpUri="https://zenodo.org/record/156491/files/figure.png" pageId="85" pageNumber="86">Fig. 389</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EA1E3692C37BC725E11E75AAFE32FE36" blockId="85.[264,1324,284,510]" pageId="85" pageNumber="86">
|
||
<emphasis id="D8D5EA80C37BC725E11E75AAFDBCFE1D" bold="true" box="[312,518,443,469]" pageId="85" pageNumber="86">Natural History.</emphasis>
|
||
<taxonomicName id="2DA14D11C37BC725E23075ADFCE7FE1D" box="[534,861,444,469]" class="Insecta" family="Staphylinidae" genus="Seeversiella" kingdom="Animalia" order="Coleoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="microphthalma">
|
||
<emphasis id="D8D5EA80C37BC725E23075ADFCE7FE1D" box="[534,861,444,469]" italics="true" pageId="85" pageNumber="86">Seeversiella microphthalma</emphasis>
|
||
</taxonomicName>
|
||
was collected in forest litter at altitude of
|
||
<quantity id="2D599B77C37BC725E10E75F5FE3EFE36" box="[296,388,484,510]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="85" pageNumber="86" unit="m" value="3000.0">3000 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |