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<mods:title id="4D4E783A6FA49A27C76A192A71E7F3FB">Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species</mods:title>
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<mods:namePart id="5BE21C7EDFDB3B6D10A314BE7440E4BD">Galindo, Lee Ann</mods:namePart>
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<mods:affiliation id="F6B70E3457FC908D71F18A8A2A464D4C">Muséum national dHistoire naturelle, Département Systématique et Evolution, ISyEB Institut (UMR 7205 CNRS / UPMC / MNHN / EPHE), 43, Rue Cuvier, 75231 Paris, France. &amp; urn: lsid: zoobank. org: author: B 84 DC 387 - F 1 A 5 - 4 FE 4 - 80 F 2 - 5 C 93 E 41 CEC 15 &amp; Corresponding author: galindo @ mnhn. fr</mods:affiliation>
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<mods:namePart id="47B8BA4B300A581ED3EB369F1F295966">Kool, Hugo H.</mods:namePart>
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<mods:affiliation id="1D363304C0201B4AF6817AAFEAA9861B">Associate Mollusca Collection, Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands. &amp; Email: hugo. kool @ naturalis. nl &amp; urn: lsid: zoobank. org: author: 5 E 718 E 5 A- 85 C 8 - 404 C- 84 DC- 6 E 53 FD 1 D 61 D 6</mods:affiliation>
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<subSubSection id="08F76522C73978294F2AFC09FC3725BF" box="[598,990,916,942]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="405236A9C73978294F2AFC09FC3725BF" blockId="8.[598,990,916,942]" box="[598,990,916,942]" pageId="8" pageNumber="9">
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Genus
<taxonomicName id="87ED4D2AC73978294FD4FC09FC3725BF" ID-CoL="7PFNQ" authority="Iredale" authorityName="Iredale" authorityYear="1936" box="[680,990,916,942]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294FD4FC09FCD225BF" bold="true" box="[680,827,916,942]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
<bibRefCitation id="247C4B58C73978294E3EFC09FC3725BF" author="Iredale T." box="[834,990,916,942]" pageId="8" pageNumber="9" pagination="267 - 340" refId="ref27495" refString="Iredale T. 1936. Australian molluscan notes. No. 2. Records of the Australian Museum 19: 267 - 340. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale, 1936</bibRefCitation>
</taxonomicName>
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<typeStatus id="9F56880BC73978294DC1FC4BFF1325E1" box="[189,250,982,1008]" pageId="8" pageNumber="9">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="405236A9C73978294DC1FB98FBE6220F" blockId="8.[189,1039,1028,1055]" box="[189,1039,1028,1055]" pageId="8" pageNumber="9">
<treatmentCitationGroup id="60FD1187C73978294DC1FB98FBE6220F" box="[189,1039,1028,1055]" pageId="8" pageNumber="9">
<taxonomicName id="87ED4D2AC73978294DC1FB98FDE1220E" ID-CoL="8YY3D" authority="Gould, 1850" authorityName="Gould" authorityYear="1850" box="[189,520,1028,1055]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="paupera">
<emphasis id="7299EABBC73978294DC1FB98FE87220F" box="[189,366,1029,1054]" italics="true" pageId="8" pageNumber="9">Nassa paupera</emphasis>
<bibRefCitation id="247C4B58C73978294C09FB99FDE1220E" author="Gould A. A." box="[373,520,1028,1055]" pageId="8" pageNumber="9" pagination="169 - 172" refId="ref27168" refString="Gould A. A. 1850. Descriptions with drawings of several new species of shells brought home from the United States Exploring Expedition. Proceedings of the Boston Society of Natural History 3: 151 - 156, 169 - 172." type="journal article" year="1850">Gould, 1850</bibRefCitation>
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by original designation (
<treatmentCitation id="C14C10B8C73978294E51FB99FBEA220F" author="Iredale T." box="[813,1027,1028,1054]" page="322" pageId="8" pageNumber="9" year="1936">
<bibRefCitation id="247C4B58C73978294E51FB99FBEA220F" author="Iredale T." box="[813,1027,1028,1054]" pageId="8" pageNumber="9" pagination="267 - 340" refId="ref27495" refString="Iredale T. 1936. Australian molluscan notes. No. 2. Records of the Australian Museum 19: 267 - 340. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale 1936: 322</bibRefCitation>
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).
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<heading id="1B1A81C5C73978294DC1FBD8FEDF224E" bold="true" box="[189,310,1093,1119]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="7299EABBC73978294DC1FBD8FEDF224E" bold="true" box="[189,310,1093,1119]" pageId="8" pageNumber="9">Diagnosis</emphasis>
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<paragraph id="405236A9C73978294DC1FBE9FD3823FF" blockId="8.[189,1399,1140,1519]" pageId="8" pageNumber="9">
Morphological features of the protoconch are of great importance when recognizing species of
<emphasis id="7299EABBC73978294DC1FB05FEBA22A0" box="[189,339,1176,1201]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="87ED4D2AC73978294DC1FB05FEA622A0" authorityName="Iredale" authorityYear="1936" box="[189,335,1176,1201]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">Reticunassa</taxonomicName>
.
</emphasis>
Several species have a multispiral protoconch, indicating a planktotrophic/planktonic larval development, resulting in a larger range of distribution. Others have a paucispiral protoconch, indicating a short, non-planktotrophic/direct larval development. All species of
<taxonomicName id="87ED4D2AC739782949FAFB42FAFD22E9" authorityName="Iredale" authorityYear="1936" box="[1158,1300,1247,1272]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC739782949FAFB42FAFD22E9" box="[1158,1300,1247,1272]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
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possess an unkeeled protoconch (
<figureCitation id="D8D62A2CC73978294C92FA9CFDDF230A" box="[494,566,1281,1307]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
AB), as do other
<taxonomicName id="87ED4D2AC73978294E64FA9CFC48230A" authorityName="Iredale" authorityYear="1916" box="[792,929,1281,1307]" class="Gastropoda" family="Nassariidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="subFamily" subFamily="Nassariinae">Nassariinae</taxonomicName>
genera in the Atlantic Ocean (
<bibRefCitation id="247C4B58C73978294864FA9CFEAA232E" author="Galindo L. A. &amp; Puillandre N. &amp; Utge J. &amp; Lozouet P. &amp; Bouchet P." pageId="8" pageNumber="9" pagination="337 - 353" refId="ref27105" refString="Galindo L. A., Puillandre N., Utge J., Lozouet P. &amp; Bouchet P. 2016. The phylogeny and systematics of the Nassariidae revisited (Gastropoda, Buccinoidea). Molecular Phylogenetics and Evolution 99: 337 - 353. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.019" type="journal article" year="2016">
Galindo
<emphasis id="7299EABBC73978294DC1FAB8FF1C232F" box="[189,245,1316,1342]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2016
</bibRefCitation>
). However,
<taxonomicName id="87ED4D2AC73978294CA0FAB8FD83232F" authorityName="Iredale" authorityYear="1936" box="[476,618,1317,1342]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294CA0FAB8FD83232F" box="[476,618,1317,1342]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
</taxonomicName>
lives in sympatry with the Indo-Pacific genus
<taxonomicName id="87ED4D2AC739782949D1FAB8FACB232F" authorityName="Dumeril" authorityYear="1805" box="[1197,1314,1317,1342]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus" sensu="stricto">
<emphasis id="7299EABBC739782949D1FAB8FACB232F" box="[1197,1314,1317,1342]" italics="true" pageId="8" pageNumber="9">Nassarius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AA57C0C73978294852FAB8FA9B232E" box="[1326,1394,1317,1343]" pageId="8" pageNumber="9" sensu="stricto">s. str.</taxonomicNameLabel>
, characterized by a keeled protoconch. Protoconchs of
<taxonomicName id="87ED4D2AC73978294E43FAD5FC242370" authorityName="Iredale" authorityYear="1936" box="[831,973,1352,1377]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294E43FAD5FC242370" box="[831,973,1352,1377]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
</taxonomicName>
can be rather broad and flattened; a few are pointed. They might possess microscopic sculpture, such as spiral ridges, axial striae and minute pustules (
<figureCitation id="D8D62A2CC73978294C52FA13FE9A23B9" box="[302,371,1422,1448]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
AB), but frequently a much higher resolution is needed to see details of this sculpture. Due to the obvious importance of protoconch characters, the identification of specimens in this genus lacking a protoconch becomes more difficult.
</paragraph>
<paragraph id="405236A9C73978294DC1F98BFD5720F1" blockId="8.[189,1399,1557,1760]" pageId="8" pageNumber="9">
Species of
<taxonomicName id="87ED4D2AC73978294C3EF98AFE392021" authorityName="Iredale" authorityYear="1936" box="[322,464,1559,1584]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294C3EF98AFE392021" box="[322,464,1559,1584]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
</taxonomicName>
have strongly or weakly developed continuous spiral cords, from 10 to 15 on the body whorl (
<figureCitation id="D8D62A2CC73978294CF6F9A4FE3B2042" box="[394,466,1593,1619]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
DF), crossing over the axial ribs. The space between these spiral cords, the intercordal surface, can be smooth, axially striated or can display a number of (occasionally extremely) fine spiral striae (
<figureCitation id="D8D62A2CC73978294CF9F9E2FE31208B" box="[389,472,1663,1690]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="8" pageNumber="9">Fig. 2F</figureCitation>
). This makes the sculpture of the intercordal surface important for identification purposes within
<taxonomicName id="87ED4D2AC73978294C03F93EFDE420AD" authorityName="Iredale" authorityYear="1936" box="[383,525,1699,1724]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294C03F93EFDE420AD" box="[383,525,1699,1724]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
</taxonomicName>
.
<figureCitation id="D8D62A2CC73978294F60F93FFD8820AC" box="[540,609,1698,1725]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[263,1325,265,1862]" captionTargetId="figure@7.[246,1337,248,1888]" captionTargetPageId="7" captionText="Fig. 1. A. Bayesian tree based on COI sequences of species of Reticunassa. B. Bayesian tree based on 28S sequences of species of Reticunassa. C. Frequency of pairwise distance (K2P model) values among COI sequences. In gray, intra-specific variation. In black, inter-specific variation. PNG = Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.3824546" httpUri="https://zenodo.org/record/3824546/files/figure.png" pageId="8" pageNumber="9">Fig. 1</figureCitation>
AB shows that differences on these intercordal surfaces are useful synapomorphies to separate SC1 from SC2.
</paragraph>
<paragraph id="405236A9C73978294DC1F89AFDE1219A" blockId="8.[189,1399,1799,1931]" pageId="8" pageNumber="9">
Species within
<taxonomicName id="87ED4D2AC73978294C0EF895FDE92130" authorityName="Iredale" authorityYear="1936" box="[370,512,1800,1825]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73978294C0EF895FDE92130" box="[370,512,1800,1825]" italics="true" pageId="8" pageNumber="9">Reticunassa</emphasis>
</taxonomicName>
have an axially-ribbed teleoconch (
<figureCitation id="D8D62A2CC73978294ED6F89AFBEE2130" box="[938,1031,1799,1825]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
). These axial ribs may become weaker or absent on the body whorl towards the base and/or the outer lip. Ribs can be round or flat. The number of ribs differs among species. Most species develop a strong varix near the outer lip. The suture is usually deeply impressed.
</paragraph>
<caption id="14926621C73878284DC1F8BDFE5B21FA" ID-DOI="http://doi.org/10.5281/zenodo.3824548" ID-Zenodo-Dep="3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1824,1850]" targetBox="[233,1354,265,1789]" targetPageId="9">
<paragraph id="405236A9C73878284DC1F8BDFE5B21FA" blockId="9.[189,1399,1824,2027]" pageId="9" pageNumber="10">
<emphasis id="7299EABBC73878284DC1F8BDFEE6212A" bold="true" box="[189,271,1824,1851]" pageId="9" pageNumber="10">Fig. 2.</emphasis>
Characters of
<taxonomicName id="87ED4D2AC73878284CB6F8BCFCEE212B" authority="Iredale, 1936" authorityName="Iredale" authorityYear="1936" box="[458,775,1824,1850]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73878284CB6F8BCFDB1212B" box="[458,600,1825,1850]" italics="true" pageId="9" pageNumber="10">Reticunassa</emphasis>
<bibRefCitation id="247C4B58C73878284F18F8BDFCEE212B" author="Iredale T." box="[612,775,1824,1850]" pageId="9" pageNumber="10" pagination="267 - 340" refId="ref27495" refString="Iredale T. 1936. Australian molluscan notes. No. 2. Records of the Australian Museum 19: 267 - 340. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale, 1936</bibRefCitation>
</taxonomicName>
.
<emphasis id="7299EABBC73878284E64F8BDFCA6212B" bold="true" box="[792,847,1824,1850]" pageId="9" pageNumber="10">AB</emphasis>
. Protoconchs.
<emphasis id="7299EABBC7387828497BF8BDFBF5212B" bold="true" box="[1031,1052,1824,1850]" pageId="9" pageNumber="10">A</emphasis>
.
<taxonomicName id="87ED4D2AC7387828494CF8BCFA9B212A" authority="(Gould, 1850)" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[1072,1394,1824,1851]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paupera">
<emphasis id="7299EABBC7387828494CF8BCFB51212B" box="[1072,1208,1825,1850]" italics="true" pageId="9" pageNumber="10">R. paupera</emphasis>
(
<bibRefCitation id="247C4B58C738782849B0F8BDFA83212A" author="Gould A. A." box="[1228,1386,1824,1851]" pageId="9" pageNumber="10" pagination="169 - 172" refId="ref27168" refString="Gould A. A. 1850. Descriptions with drawings of several new species of shells brought home from the United States Exploring Expedition. Proceedings of the Boston Society of Natural History 3: 151 - 156, 169 - 172." type="journal article" year="1850">Gould, 1850</bibRefCitation>
)
</taxonomicName>
, MNHN IM-2007-31782.
<emphasis id="7299EABBC73878284C8AF8D9FDE3214F" bold="true" box="[502,522,1860,1886]" pageId="9" pageNumber="10">B</emphasis>
.
<taxonomicName id="87ED4D2AC73878284F63F8D8FC31214E" authority="(Kool &amp; Dekker, 2006)" authorityName="" baseAuthorityName="Kool &amp; Dekker" baseAuthorityYear="2006" box="[543,984,1860,1887]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="silvardi">
<emphasis id="7299EABBC73878284F63F8D8FD49214F" box="[543,672,1860,1886]" italics="true" pageId="9" pageNumber="10">R. silvardi</emphasis>
(
<bibRefCitation id="247C4B58C73878284FC9F8D9FC39214E" author="Kool H. H. &amp; Dekker H." box="[693,976,1860,1887]" pageId="9" pageNumber="10" pagination="54 - 75" refId="ref27639" refString="Kool H. H. &amp; Dekker H. 2006. Review of the Nassarius pauper (Gould, 1850) complex (Gastropoda Nassariidae). Part 1, with the description of four new species from the Indo-West Pacific. Visaya 1 (6): 54 - 75." type="journal article" year="2006">Kool &amp; Dekker, 2006</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="7299EABBC73878284E96F8D9FC16214F" bold="true" box="[1002,1023,1860,1886]" pageId="9" pageNumber="10">C</emphasis>
. Suture detail of
<taxonomicName id="87ED4D2AC7387828499AF8D8FA86214F" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[1254,1391,1861,1886]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paupera">
<emphasis id="7299EABBC7387828499AF8D8FA86214F" box="[1254,1391,1861,1886]" italics="true" pageId="9" pageNumber="10">R. paupera</emphasis>
</taxonomicName>
.
<emphasis id="7299EABBC73878284DC1F8FAFF3B2190" bold="true" box="[189,210,1895,1921]" pageId="9" pageNumber="10">D</emphasis>
. Axial striae in
<taxonomicName id="87ED4D2AC73878284CE8F8F5FDE62190" authorityName="" baseAuthorityName="Kool &amp; Dekker" baseAuthorityYear="2006" box="[404,527,1895,1921]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="silvardi">
<emphasis id="7299EABBC73878284CE8F8F5FDE62190" box="[404,527,1895,1921]" italics="true" pageId="9" pageNumber="10">R. silvardi</emphasis>
</taxonomicName>
.
<emphasis id="7299EABBC73878284F63F8FAFDBA2190" bold="true" box="[543,595,1895,1921]" pageId="9" pageNumber="10">EF</emphasis>
. Intercordal spirals.
<emphasis id="7299EABBC73878284E37F8FAFCB62190" bold="true" box="[843,863,1895,1921]" pageId="9" pageNumber="10">E</emphasis>
.
<taxonomicName id="87ED4D2AC73878284E13F8F5FB552193" authority="(Souverbie, 1864)" baseAuthorityName="Souverbie" baseAuthorityYear="1864" box="[879,1212,1895,1922]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="tringa">
<emphasis id="7299EABBC73878284E13F8F5FC302190" box="[879,985,1896,1921]" italics="true" pageId="9" pageNumber="10">R. tringa</emphasis>
(
<bibRefCitation id="247C4B58C73878284E96F8FAFB5A2193" author="Souverbie M. &amp; Montrouzier R. P." box="[1002,1203,1895,1922]" pageId="9" pageNumber="10" pagination="261 - 275" refId="ref28538" refString="Souverbie M. &amp; Montrouzier R. P. 1864. Descriptions d'especes nouvelles de l'Archipel Caledonien. Journal de Conchyliologie 12: 261 - 275." type="journal article" year="1864">Souverbie, 1864</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="7299EABBC738782849B4F8FAFB332190" bold="true" box="[1224,1242,1895,1921]" pageId="9" pageNumber="10">F</emphasis>
.
<emphasis id="7299EABBC73878284997F8F5FA9E2190" box="[1259,1399,1896,1921]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="87ED4D2AC73878284997F8F5FA9A2190" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[1259,1395,1896,1921]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paupera">R. paupera</taxonomicName>
.
</emphasis>
<emphasis id="7299EABBC73878284DC1F817FF1021B5" bold="true" box="[189,249,1930,1956]" pageId="9" pageNumber="10">GH</emphasis>
. Columellar details of
<emphasis id="7299EABBC73878284F6CF816FD7221B5" box="[528,667,1931,1956]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="87ED4D2AC73878284F6CF816FD7E21B5" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[528,663,1931,1956]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paupera">R. paupera</taxonomicName>
.
</emphasis>
<emphasis id="7299EABBC73878284FDEF817FD3921B5" bold="true" box="[674,720,1930,1956]" pageId="9" pageNumber="10">IL</emphasis>
. Type series of
<emphasis id="7299EABBC73878284EF6F816FBFC21B5" box="[906,1045,1931,1956]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="87ED4D2AC73878284EF6F816FBF821B5" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[906,1041,1931,1956]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="paupera">R. paupera</taxonomicName>
.
</emphasis>
<emphasis id="7299EABBC73878284960F817FBAC21B5" bold="true" box="[1052,1093,1930,1956]" pageId="9" pageNumber="10">IJ</emphasis>
. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm.
<emphasis id="7299EABBC73878284F4FF833FD8521D9" bold="true" box="[563,620,1966,1992]" pageId="9" pageNumber="10">KL</emphasis>
. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm.
</paragraph>
</caption>
<paragraph id="405236A9C73B782B4DC1FE96FD07277D" blockId="10.[189,1399,267,364]" pageId="10" pageNumber="11">
The aperture is oval to sub-circular, the inside of the outer lip with a number of denticles or lirae (
<figureCitation id="D8D62A2CC73B782B4DBBFEB3FEE42758" box="[199,269,302,329]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
GH). A columellar callus is present and varies in shape and sculpture. It can be limited to the columella or partly extend over the body whorl.
</paragraph>
<paragraph id="405236A9C73B782B4DC1FE09FDB627E2" blockId="10.[189,1399,403,500]" pageId="10" pageNumber="11">In some species, the color is variable; generally, it is cream-yellowish to orange, or white to dark brown. Banding with darker colors may also be present. Sometimes only some of the spiral cords are darker than the remaining part of the shell.</paragraph>
<paragraph id="405236A9C73B782B4DC1FD81FEAC248E" blockId="10.[189,1399,539,671]" pageId="10" pageNumber="11">
<taxonomicName id="87ED4D2AC73B782B4DC1FD81FEA22424" authorityName="Iredale" authorityYear="1936" box="[189,331,540,565]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4DC1FD81FEA22424" box="[189,331,540,565]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
species are small, usually varying between 5 and
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, but one is up to
<quantity id="87159B4CC73B782B4875FD86FA9E2427" box="[1289,1399,539,566]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="10" pageNumber="11" unit="mm" value="16.5">16.5 mm</quantity>
(
<taxonomicName id="87ED4D2AC73B782B4DBBFDA2FED02449" authority="Galindo &amp; Kool &amp; Dekker, 2017" authorityName="Galindo &amp; Kool &amp; Dekker" authorityYear="2017" box="[199,313,574,600]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="goliath" status="sp. nov.">
<emphasis id="7299EABBC73B782B4DBBFDA2FED02449" box="[199,313,574,600]" italics="true" pageId="10" pageNumber="11">R. goliath</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69AA57C0C73B782B4C42FDA2FE722448" box="[318,411,575,601]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
). Most species live in littoral to sublittoral ecosystems, only empty shells having been found in waters deeper than
<specimenCount id="56EBFD20C73B782B4F76FDFCFDB5246D" box="[522,604,609,636]" pageId="10" pageNumber="11" type="generic">
<quantity id="87159B4CC73B782B4F76FDFCFDB1246D" box="[522,600,609,636]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="11" unit="m" value="260.0">260 m</quantity>
.
</specimenCount>
Their occurrence is in the Indo-Pacific Ocean, from the Miocene to the present.
</paragraph>
</subSubSection>
<subSubSection id="08F76522C73B782B4DC1FD5BFDCE2185" pageId="10" pageNumber="11" type="discussion">
<paragraph id="405236A9C73B782B4DC1FD5BFEC624F1" blockId="10.[189,303,710,736]" box="[189,303,710,736]" pageId="10" pageNumber="11">
<heading id="1B1A81C5C73B782B4DC1FD5BFEC624F1" bold="true" box="[189,303,710,736]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="7299EABBC73B782B4DC1FD5BFEC624F1" bold="true" box="[189,303,710,736]" pageId="10" pageNumber="11">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="405236A9C73B782B4DC1FD69FCEC2353" blockId="10.[189,1399,756,1346]" pageId="10" pageNumber="11">
The genus name
<taxonomicName id="87ED4D2AC73B782B4CF0FD68FDF3251F" authorityName="Iredale" authorityYear="1936" box="[396,538,757,782]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4CF0FD68FDF3251F" box="[396,538,757,782]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
was introduced to group together “species around
<taxonomicName id="87ED4D2AC73B782B49FEFD68FAAC251F" authority="Gould" authorityName="Gould" box="[1154,1349,756,782]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pauperus">
<emphasis id="7299EABBC73B782B49FEFD68FB06251F" box="[1154,1263,757,782]" italics="true" pageId="10" pageNumber="11">pauperus</emphasis>
Gould
</taxonomicName>
” by
<bibRefCitation id="247C4B58C73B782B4DC1FC85FE412523" author="Iredale T." box="[189,424,792,818]" pageId="10" pageNumber="11" pagination="267 - 340" refId="ref27495" refString="Iredale T. 1936. Australian molluscan notes. No. 2. Records of the Australian Museum 19: 267 - 340. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale (1936: 322)</bibRefCitation>
.
<bibRefCitation id="247C4B58C73B782B4CC7FC85FD052522" author="Cernohorsky W. O." box="[443,748,792,819]" pageId="10" pageNumber="11" pagination="1 - 356" refId="ref26672" refString="Cernohorsky W. O. 1984. Systematics of the family Nassariidae (Mollusca: Gastropoda). Bulletin of the Auckland Institute and Museum 14: 1 - 356." type="journal article" year="1984">Cernohorsky (1984: 175)</bibRefCitation>
regarded
<taxonomicName id="87ED4D2AC73B782B4E15FC84FC1E2523" authorityName="Iredale" authorityYear="1936" box="[873,1015,793,818]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4E15FC84FC1E2523" box="[873,1015,793,818]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
as a synonym of the subgenus “
<emphasis id="7299EABBC73B782B4DB6FCA1FEE22544" box="[202,267,828,853]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
Leach in
<bibRefCitation id="247C4B58C73B782B4CF6FCA6FDF92547" author="Gray J. E." box="[394,528,827,854]" pageId="10" pageNumber="11" refId="ref27445" refString="Gray J. E. 1852. Molluscorum Britanniae. Synopsis of the Mollusca of Great Britain. John van Vorst, London." type="book" year="1852">Gray, 1852</bibRefCitation>
”, within the genus
<taxonomicName id="87ED4D2AC73B782B4E7FFCA1FBDC2547" authority="Dumeril, 1805" authorityName="Dumeril" authorityYear="1805" box="[771,1077,827,854]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4E7FFCA1FC912544" box="[771,888,828,853]" italics="true" pageId="10" pageNumber="11">Nassarius</emphasis>
Duméril, 1805
</taxonomicName>
. However,
<emphasis id="7299EABBC73B782B49BCFCA1FAE82544" box="[1216,1281,828,853]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
is not an available name, but an incorrect subsequent spelling of
<emphasis id="7299EABBC73B782B4E04FCC2FC522569" box="[888,955,863,888]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
Leach in Gray, 1847 (
<bibRefCitation id="247C4B58C73B782B49A1FCC3FF04258D" author="Gray J. E." pageId="10" pageNumber="11" pagination="267 - 273" refId="ref27338" refString="Gray J. E. 1847 a. The classification of the British Mollusca by W. E. Leach, M. D. Annals and Magazine of Natural History 20 (133): 267 - 273. http: // dx. doi. org / 10.1080 / 037454809496042" type="journal article" year="1847">Gray 1847a: 269</bibRefCitation>
), since “
<emphasis id="7299EABBC73B782B4C24FC1FFE70258A" box="[344,409,898,923]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
” included the same three species as in the original description of
<emphasis id="7299EABBC73B782B49C0FC1FFB16258A" box="[1212,1279,898,923]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
(during a period in which original descriptions only consisted of introducing new names). Nonetheless, “
<emphasis id="7299EABBC73B782B4854FC38FA8025AF" box="[1320,1385,933,958]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
” and not
<emphasis id="7299EABBC73B782B4C5FFC54FE8F25F3" box="[291,358,969,994]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
was listed by
<bibRefCitation id="247C4B58C73B782B4F66FC55FCA125F2" author="Adams H. &amp; Adams A." box="[538,840,968,995]" pageId="10" pageNumber="11" pagination="129 - 138" refId="ref26296" refString="Adams H. &amp; Adams A. 1853 - 1858. The Genera of Recent Mollusca; Arranged According to their Organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls. [Published in parts: Vol. 1: i-xl (1858), 1 - 256 (1853), 257 - 484 (1854). Vol. 2: 1 - 92 (1854), 93 - 284 (1855), 285 - 412 (1856), 413 - 540 (1857), 541 - 661 (1858). Vol. 3: pl. 1 - 32 (1853), 33 - 96 (1855), 97 - 112 (1856), 113 - 128 (1857), 129 - 138 (1858).]" type="book chapter" year="1853">Adams &amp; Adams (1853)</bibRefCitation>
.
<bibRefCitation id="247C4B58C73B782B4E2BFC55FBC225F3" author="Cossmann M." box="[855,1067,968,994]" pageId="10" pageNumber="11" refId="ref26774" refString="Cossmann M. 1901. Essais de Paleoconchologie Comparee. Fourth edition. The author and Societe d'Editions Scientifiques, Paris." type="book" year="1901">Cossmann (1901)</bibRefCitation>
considered both
<emphasis id="7299EABBC73B782B4982FC54FAA825F3" box="[1278,1345,969,994]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
and “
<emphasis id="7299EABBC73B782B4DB6FC71FEE22214" box="[202,267,1004,1029]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
” as valid names, with “
<emphasis id="7299EABBC73B782B4F3CFC71FD682214" box="[576,641,1004,1029]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
” treated
<emphasis id="7299EABBC73B782B4F85FC71FCD32214" box="[761,826,1004,1029]" italics="true" pageId="10" pageNumber="11">sensu</emphasis>
<bibRefCitation id="247C4B58C73B782B4E39FC76FB902217" author="Adams H. &amp; Adams A." box="[837,1145,1003,1030]" pageId="10" pageNumber="11" pagination="129 - 138" refId="ref26296" refString="Adams H. &amp; Adams A. 1853 - 1858. The Genera of Recent Mollusca; Arranged According to their Organization. London, van Voorst. Vol. 1: xl + 484 pp.; vol. 2: 661 pp.; vol. 3: 138 pls. [Published in parts: Vol. 1: i-xl (1858), 1 - 256 (1853), 257 - 484 (1854). Vol. 2: 1 - 92 (1854), 93 - 284 (1855), 285 - 412 (1856), 413 - 540 (1857), 541 - 661 (1858). Vol. 3: pl. 1 - 32 (1853), 33 - 96 (1855), 97 - 112 (1856), 113 - 128 (1857), 129 - 138 (1858).]" type="book chapter" year="1853">Adams &amp; Adams (1853)</bibRefCitation>
, not Leach
<emphasis id="7299EABBC73B782B486EFC71FAC02214" box="[1298,1321,1004,1029]" italics="true" pageId="10" pageNumber="11">in</emphasis>
<bibRefCitation id="247C4B58C73B782B484AFC76FF132238" author="Gray J. E." pageId="10" pageNumber="11" refId="ref27445" refString="Gray J. E. 1852. Molluscorum Britanniae. Synopsis of the Mollusca of Great Britain. John van Vorst, London." type="book" year="1852">Gray, 1852</bibRefCitation>
, and with “
<typeStatus id="9F56880BC73B782B4CF1FB93FE2B2239" box="[397,450,1038,1064]" pageId="10" pageNumber="11">type</typeStatus>
species”
<taxonomicName id="87ED4D2AC73B782B4F41FB92FC922239" authority="&quot; Muller" authorityName="Muller" box="[573,891,1038,1064]" class="Gastropoda" family="Nassariidae" genus="Nassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="incrassata">
<emphasis id="7299EABBC73B782B4F41FB92FCE22239" box="[573,779,1039,1064]" italics="true" pageId="10" pageNumber="11">Nassa incrassata</emphasis>
“Müller
</taxonomicName>
”.
<bibRefCitation id="247C4B58C73B782B4EE9FB93FB8C2239" author="Woodring W. P." box="[917,1125,1038,1064]" pageId="10" pageNumber="11" pagination="1 - 564" refId="ref28678" refString="Woodring W. P. 1928. Miocene mollusks from Bowden, Jamaica. Part II. Gastropods and discussion of results. Contributions to the geology and paleontology of the West Indies. Carnegie Institution of Washington Publication 385: 1 - 564." type="journal article" year="1928">Woodring (1928)</bibRefCitation>
noted the misspelling and this was reiterated by
<bibRefCitation id="247C4B58C73B782B4C81FBAFFD6C225A" author="Lee H. G." box="[509,645,1073,1100]" pageId="10" pageNumber="11" pagination="3 - 5" refId="ref27806" refString="Lee H. G. 2013. What to do with Hima of authors, not of Leach, 1852? Shell-O-Gram 54 (2): 3 - 5." type="journal article" year="2013">Lee (2013)</bibRefCitation>
. Despite this, the error in the use of “
<emphasis id="7299EABBC73B782B492CFBAFFB78225A" box="[1104,1169,1074,1099]" italics="true" pageId="10" pageNumber="11">Hima</emphasis>
” has persisted. On the other hand,
<emphasis id="7299EABBC73B782B4CFDFBC8FE2D227F" box="[385,452,1109,1134]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
Leach in Gray, 1847 is a junior objective synonym of
<taxonomicName id="87ED4D2AC73B782B490AFBC8FABB227F" authority="Risso, 1826" authorityName="Risso" authorityYear="1826" box="[1142,1362,1108,1134]" class="Gastropoda" family="Nassariidae" genus="Tritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B490AFBC8FB5E227F" box="[1142,1207,1109,1134]" italics="true" pageId="10" pageNumber="11">Tritia</emphasis>
<bibRefCitation id="247C4B58C73B782B49BFFBC9FABB227F" author="Risso A." box="[1219,1362,1108,1134]" pageId="10" pageNumber="11" refId="ref28362" refString="Risso A. 1826. Histoire naturelle des principales Productions de l'Europe meridionale et particulierement de celles des Environs de Nice et des Alpes Maritimes, IV. F. G. Levrault, Paris." type="book" year="1826">Risso, 1826</bibRefCitation>
</taxonomicName>
as they have the same
<typeStatus id="9F56880BC73B782B4CCFFBE5FE0F2283" box="[435,486,1144,1170]" pageId="10" pageNumber="11">type</typeStatus>
species,
<taxonomicName id="87ED4D2AC73B782B4F2BFBE4FBCB2282" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[599,1058,1144,1171]" class="Gastropoda" family="Buccinidae" genus="Buccinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="reticulatum">
<emphasis id="7299EABBC73B782B4F2BFBE4FCB72283" box="[599,862,1144,1170]" italics="true" pageId="10" pageNumber="11">Buccinum reticulatum</emphasis>
Linnaeus, 1758
</taxonomicName>
, designated for both genera by
<bibRefCitation id="247C4B58C73B782B4D9AFB06FE2522A4" author="Gray J. E." box="[230,460,1179,1205]" pageId="10" pageNumber="11" pagination="129 - 219" refId="ref27390" refString="Gray J. E. 1847 b. A list of the genera of recent Mollusca, their synonyma and types. Proceedings of the Zoological Society of London 15: 129 - 219. Available from http: // biodiversitylibrary. org / page / 12862913 [accessed 21 Dec. 2016]." type="journal article" year="1847">Gray (1847b: 139)</bibRefCitation>
. Molecular results of
<bibRefCitation id="247C4B58C73B782B4FA7FB06FC3422A7" author="Galindo L. A. &amp; Puillandre N. &amp; Utge J. &amp; Lozouet P. &amp; Bouchet P." box="[731,989,1179,1206]" pageId="10" pageNumber="11" pagination="337 - 353" refId="ref27105" refString="Galindo L. A., Puillandre N., Utge J., Lozouet P. &amp; Bouchet P. 2016. The phylogeny and systematics of the Nassariidae revisited (Gastropoda, Buccinoidea). Molecular Phylogenetics and Evolution 99: 337 - 353. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.019" type="journal article" year="2016">
Galindo
<emphasis id="7299EABBC73B782B4E39FB01FC9422A4" box="[837,893,1179,1205]" italics="true" pageId="10" pageNumber="11">et al</emphasis>
. (2016)
</bibRefCitation>
indicated that species considered part of the
<taxonomicName id="87ED4D2AC73B782B4C37FB22FDD022C9" box="[331,569,1215,1240]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pauperus">
<emphasis id="7299EABBC73B782B4C37FB22FDD022C9" box="[331,569,1215,1240]" italics="true" pageId="10" pageNumber="11">Nassarius pauperus</emphasis>
</taxonomicName>
complex, including “
<taxonomicName id="87ED4D2AC73B782B4E3AFB22FBF322C9" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[838,1050,1215,1240]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pauper">
<emphasis id="7299EABBC73B782B4E3AFB22FBF322C9" box="[838,1050,1215,1240]" italics="true" pageId="10" pageNumber="11">Nassarius pauper</emphasis>
</taxonomicName>
”, form a clade distant from
<taxonomicName id="87ED4D2AC73B782B4DC1FB7FFE5622EA" authorityName="Linnaeus" authorityYear="1758" box="[189,447,1249,1275]" class="Gastropoda" family="Buccinidae" genus="Buccinum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="reticulatum">
<emphasis id="7299EABBC73B782B4DC1FB7FFE5622EA" box="[189,447,1249,1275]" italics="true" pageId="10" pageNumber="11">Buccinum reticulatum</emphasis>
</taxonomicName>
(accepted as
<taxonomicName id="87ED4D2AC73B782B4F22FB7FFCF122EA" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[606,792,1249,1275]" class="Gastropoda" family="Nassariidae" genus="Tritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="reticulata">
<emphasis id="7299EABBC73B782B4F22FB7FFCF122EA" box="[606,792,1249,1275]" italics="true" pageId="10" pageNumber="11">Tritia reticulata</emphasis>
</taxonomicName>
), the
<typeStatus id="9F56880BC73B782B4E27FB7FFC6722ED" box="[859,910,1250,1276]" pageId="10" pageNumber="11">type</typeStatus>
species of
<taxonomicName id="87ED4D2AC73B782B496CFB7FFBB822EA" authorityName="Risso" authorityYear="1826" box="[1040,1105,1250,1275]" class="Gastropoda" family="Nassariidae" genus="Tritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B496CFB7FFBB822EA" box="[1040,1105,1250,1275]" italics="true" pageId="10" pageNumber="11">Tritia</emphasis>
</taxonomicName>
(=
<emphasis id="7299EABBC73B782B4906FB7FFB5422EA" box="[1146,1213,1250,1275]" italics="true" pageId="10" pageNumber="11">Hinia</emphasis>
), also shown in
<figureCitation id="D8D62A2CC73B782B4DC1FA99FEF4230E" box="[189,285,1284,1311]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[263,1325,265,1862]" captionTargetId="figure@7.[246,1337,248,1888]" captionTargetPageId="7" captionText="Fig. 1. A. Bayesian tree based on COI sequences of species of Reticunassa. B. Bayesian tree based on 28S sequences of species of Reticunassa. C. Frequency of pairwise distance (K2P model) values among COI sequences. In gray, intra-specific variation. In black, inter-specific variation. PNG = Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.3824546" httpUri="https://zenodo.org/record/3824546/files/figure.png" pageId="10" pageNumber="11">Fig. 1A</figureCitation>
. The only available genus group name for this
<taxonomicName id="87ED4D2AC73B782B4E20FA98FC20230F" box="[860,969,1285,1310]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pauperus">
<emphasis id="7299EABBC73B782B4E20FA98FC20230F" box="[860,969,1285,1310]" italics="true" pageId="10" pageNumber="11">pauperus</emphasis>
</taxonomicName>
clade is
<taxonomicName id="87ED4D2AC73B782B4946FA98FA86230F" authority="Iredale, 1936" authorityName="Iredale" authorityYear="1936" box="[1082,1391,1284,1310]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4946FA98FB21230F" box="[1082,1224,1285,1310]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
<bibRefCitation id="247C4B58C73B782B49AEFA99FA86230F" author="Iredale T." box="[1234,1391,1284,1310]" pageId="10" pageNumber="11" pagination="267 - 340" refId="ref27495" refString="Iredale T. 1936. Australian molluscan notes. No. 2. Records of the Australian Museum 19: 267 - 340. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale, 1936</bibRefCitation>
</taxonomicName>
, which is herein reinstated with full generic status.
</paragraph>
<paragraph id="405236A9C73B782B4DC1FAF4FDCE2185" blockId="10.[189,1399,1385,1940]" pageId="10" pageNumber="11">
Other species might be confused as members of the genus
<taxonomicName id="87ED4D2AC73B782B4E0FFAF7FBE82392" authorityName="Iredale" authorityYear="1936" box="[883,1025,1386,1411]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4E0FFAF7FBE82392" box="[883,1025,1386,1411]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
because of their morphological similarities. As it is impossible to treat all the small and ridged species, we focus on the following characters: (1)
<taxonomicName id="87ED4D2AC73B782B4C1BFA2DFE1C23D8" authorityName="Iredale" authorityYear="1936" box="[359,501,1456,1481]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4C1BFA2DFE1C23D8" box="[359,501,1456,1481]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
is an Indo-Pacific group of molluscs, not occurring in Atlantic waters (
<bibRefCitation id="247C4B58C73B782B486AFA2DFED623FF" author="Galindo L. A. &amp; Puillandre N. &amp; Utge J. &amp; Lozouet P. &amp; Bouchet P." pageId="10" pageNumber="11" pagination="337 - 353" refId="ref27105" refString="Galindo L. A., Puillandre N., Utge J., Lozouet P. &amp; Bouchet P. 2016. The phylogeny and systematics of the Nassariidae revisited (Gastropoda, Buccinoidea). Molecular Phylogenetics and Evolution 99: 337 - 353. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.019" type="journal article" year="2016">
Galindo
<emphasis id="7299EABBC73B782B4DC1FA49FF1123FC" box="[189,248,1491,1517]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2016
</bibRefCitation>
); (2) species of
<taxonomicName id="87ED4D2AC73B782B4C8BFA49FD6C23FC" authorityName="Iredale" authorityYear="1936" box="[503,645,1492,1517]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4C8BFA49FD6C23FC" box="[503,645,1492,1517]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
lack keeled protoconchs (all species with this keel belong to the genus
<taxonomicName id="87ED4D2AC73B782B4C76FA6AFE962001" authorityName="Dumeril" authorityYear="1805" box="[266,383,1527,1552]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4C76FA6AFE962001" box="[266,383,1527,1552]" italics="true" pageId="10" pageNumber="11">Nassarius</emphasis>
</taxonomicName>
); (3) fine spiral striae or axial striae on the intercordal surface (
<figureCitation id="D8D62A2CC73B782B490DFA6BFB5E2001" box="[1137,1207,1526,1552]" captionStart="Fig" captionStartId="9.[189,232,1824,1850]" captionTargetBox="[233,1354,265,1789]" captionTargetId="figure@9.[233,1354,265,1789]" captionTargetPageId="9" captionText="Fig. 2. Characters of Reticunassa Iredale, 1936. AB. Protoconchs. A. R. paupera (Gould, 1850), MNHN IM-2007-31782. B. R. silvardi (Kool &amp; Dekker, 2006). C. Suture detail of R. paupera. D. Axial striae in R. silvardi. EF. Intercordal spirals. E. R. tringa (Souverbie, 1864). F. R. paupera. GH. Columellar details of R. paupera. IL. Type series of R. paupera. IJ. Lectotype, USNM 5726, length 9.1 mm, width 4.8 mm. KL. Paralectotype, MCZ 216602, length 7.2 mm, width 3.5 mm. All scale bars = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824548" httpUri="https://zenodo.org/record/3824548/files/figure.png" pageId="10" pageNumber="11">Fig. 2</figureCitation>
DF) may be an indication of at least kinship with
<taxonomicName id="87ED4D2AC73B782B4F28F987FD0B2022" authorityName="Iredale" authorityYear="1936" box="[596,738,1562,1587]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4F28F987FD0B2022" box="[596,738,1562,1587]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
; (4) all species of
<taxonomicName id="87ED4D2AC73B782B4EBDF987FBA62022" authorityName="Iredale" authorityYear="1936" box="[961,1103,1562,1587]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4EBDF987FBA62022" box="[961,1103,1562,1587]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
have a distinct, elongate and slender shape (
<figureCitation id="D8D62A2CC73B782B4CE0F9A1FE012046" box="[412,488,1596,1623]" captionStart="Fig" captionStartId="11.[189,232,1720,1746]" captionTargetBox="[272,1314,265,1685]" captionTargetId="figure@11.[272,1314,265,1685]" captionTargetPageId="11" captionText="Fig. 3. Shells of Reticunassa Iredale, 1936. AC. Possible neotype of R. paupera (Gould, 1850), MNHN IM-2007-31782, length 6.9 mm, width 3.6 mm, Vanuatu, Aore I., 15°36.6 S, 167°10.1 E, 1018 m. DI. R. tringa (Souverbie, 1864). DE. Hologenophore, MNHN IM-2007-31753, length 9.5 mm, width 4.6 mm. FH. Lectotype, MHNBx 2004.TY.89.1, length 9.0 mm, width 5.0 mm. I. Protoconch of the hologenophore. JM. R. visayaensis sp. nov. JL. Holotype, MNHN IM- 2007-31912, length 9.5 mm, width 4.6 mm, Philippines, Panglao I., 9°35.7 N, 123°44.7 E, 03 m. M. Protoconch. NQ. R. poppeorum sp. nov. NP. Holotype, MNHN IM-2007-31779, length 9.0 mm, width 4.5 mm, Vanuatu, Aore I., 15°34.4 S, 167°12.6 E, 23 m. Q. Protoconch. Scale bars: I, M, Q = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824550" httpUri="https://zenodo.org/record/3824550/files/figure.png" pageId="10" pageNumber="11">Figs 3</figureCitation>
,
<figureCitation id="D8D62A2CC73B782B4C89F9A0FDEC2046" box="[501,517,1597,1623]" captionStart="Fig" captionStartId="29.[189,232,1717,1743]" captionTargetBox="[273,1314,263,1679]" captionTargetId="figure@29.[273,1314,263,1679]" captionTargetPageId="29" captionText="Fig. 8. Shells of Reticunassa Iredale, 1936. AD. Reticunassa annabolteae sp. nov. AC. Holotype, MNHN IM-2009-12843, length 11.5 mm, width5.3 mm, S Madagascar,Ambatobe, Bavarama,25°27.9 S, 44°57.6 E, 01 m. D. Protoconch. EH. R. goliath sp. nov. EG. Holotype, MNHN IM-2000- 22724, length 15.5 mm, width 7.9 mm, New Caledonia, Nouméa, leg. D. Massemin, ex coll. H. Kool. H. Protoconch of paratype, MNHN IM-2000-28394. IL. R. intrudens sp. nov. Holotype, MNHN IM-2000-22726, length 7.9 mm, width 4.3 mm, New Caledonia, Touho, 20°44 S, 165°14 E, 4959 m. L. Protoconch. MP. R. thailandensis sp. nov. MO. Holotype, ZMA Moll.4.09.047, length 7.35 mm, width 3.8 mm, Thailand, Gulf of Thailand, 6°40.313 N, 101°43.852 E, 315 m. P. Protoconch. Scale bars: D, H, L, P = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3824560" httpUri="https://zenodo.org/record/3824560/files/figure.png" pageId="10" pageNumber="11">8</figureCitation>
). These criteria exclude from
<taxonomicName id="87ED4D2AC73B782B4E19F9A0FC1A2047" authorityName="Iredale" authorityYear="1936" box="[869,1011,1597,1622]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4E19F9A0FC1A2047" box="[869,1011,1597,1622]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
any Mediterranean species, such as
<taxonomicName id="87ED4D2AC73B782B4D9CF9FDFDBE206B" authority="(Strom, 1768)" baseAuthorityName="Strom" baseAuthorityYear="1768" box="[224,599,1631,1658]" class="Gastropoda" family="Nassariidae" genus="Tritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="incrassata">
<emphasis id="7299EABBC73B782B4D9CF9FDFE4C2068" box="[224,421,1632,1657]" italics="true" pageId="10" pageNumber="11">Tritia incrassata</emphasis>
(
<bibRefCitation id="247C4B58C73B782B4CC4F9FDFDA6206B" author="Strom H." box="[440,591,1631,1658]" pageId="10" pageNumber="11" pagination="313 - 371" refId="ref28565" refString="Strom H. 1768. Beskrivelse over norske Insecter. Det Kongelige Norske Videnskabers Selskabs Skrifter 4: 313 - 371." type="journal article" year="1768">Strøm, 1768</bibRefCitation>
)
</taxonomicName>
(previously
<taxonomicName id="87ED4D2AC73B782B4F8DF9FDFC1E2068" baseAuthorityName="Strøm" baseAuthorityYear="1768" box="[753,1015,1632,1657]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="incrassatus">
<emphasis id="7299EABBC73B782B4F8DF9FDFC1E2068" box="[753,1015,1632,1657]" italics="true" pageId="10" pageNumber="11">Nassarius incrassatus</emphasis>
</taxonomicName>
), West African species, such as
<taxonomicName id="87ED4D2AC73B782B4DC1F919FD49208C" authority="(Von Maltzan, 1884)" baseAuthorityName="Von Maltzan" baseAuthorityYear="1884" box="[189,672,1667,1693]" class="Gastropoda" family="Nassariidae" genus="Tritia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="senegalensis">
<emphasis id="7299EABBC73B782B4DC1F919FE74208C" box="[189,413,1667,1693]" italics="true" pageId="10" pageNumber="11">Tritia senegalensis</emphasis>
(
<bibRefCitation id="247C4B58C73B782B4CD3F91EFD71208C" author="Von Maltzan H." box="[431,664,1667,1693]" pageId="10" pageNumber="11" pagination="65 - 73" refId="ref28656" refString="Von Maltzan H. 1884. Diagnosen neuer Senegambischer Gastropoden. Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft 16: 65 - 73." type="journal article" year="1884">Von Maltzan, 1884</bibRefCitation>
)
</taxonomicName>
(previously
<taxonomicName id="87ED4D2AC73B782B4E47F919FBA7208C" baseAuthorityName="Maltzan" baseAuthorityYear="1884" box="[827,1102,1667,1693]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="senegalensis">
<emphasis id="7299EABBC73B782B4E47F919FBA7208C" box="[827,1102,1667,1693]" italics="true" pageId="10" pageNumber="11">Nassarius senegalensis</emphasis>
</taxonomicName>
) and Caribbean species, such as
<taxonomicName id="87ED4D2AC73B782B4C58F93BFCD820D0" authority="(C. B. Adams, 1845)" baseAuthorityName="C. B. Adams" baseAuthorityYear="1845" box="[292,817,1702,1729]" class="Gastropoda" family="Nassariidae" genus="Phrontis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="candidissima">
<emphasis id="7299EABBC73B782B4C58F93BFDC620D1" box="[292,559,1702,1728]" italics="true" pageId="10" pageNumber="11">Phrontis candidissima</emphasis>
(C.B.
<bibRefCitation id="247C4B58C73B782B4FFAF93BFCC020D0" author="Adams C. B." box="[646,809,1702,1729]" pageId="10" pageNumber="11" pagination="1 - 17" refId="ref26265" refString="Adams C. B. 1845. Specierum novarum conchyliorum, in Jamaica repertorum, synopsis. Proceedings of the Boston Society of Natural History 2: 1 - 17." type="journal article" year="1845">Adams, 1845</bibRefCitation>
)
</taxonomicName>
(previously
<taxonomicName id="87ED4D2AC73B782B4EACF93AFB1120D1" baseAuthorityName="C. B. Adams" baseAuthorityYear="1845" box="[976,1272,1702,1728]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="candidissimus">
<emphasis id="7299EABBC73B782B4EACF93AFB1120D1" box="[976,1272,1702,1728]" italics="true" pageId="10" pageNumber="11">Nassarius candidissimus</emphasis>
</taxonomicName>
) (
<bibRefCitation id="247C4B58C73B782B486AF93BFEAC20F5" author="Galindo L. A. &amp; Puillandre N. &amp; Utge J. &amp; Lozouet P. &amp; Bouchet P." pageId="10" pageNumber="11" pagination="337 - 353" refId="ref27105" refString="Galindo L. A., Puillandre N., Utge J., Lozouet P. &amp; Bouchet P. 2016. The phylogeny and systematics of the Nassariidae revisited (Gastropoda, Buccinoidea). Molecular Phylogenetics and Evolution 99: 337 - 353. http: // dx. doi. org / 10.1016 / j. ympev. 2016.03.019" type="journal article" year="2016">
Galindo
<emphasis id="7299EABBC73B782B4DC1F957FF1220F2" box="[189,251,1737,1763]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
2016
</bibRefCitation>
). Most Indo-Pacific species, such as
<taxonomicName id="87ED4D2AC73B782B4E7AF957FAD520F2" authority="Kool &amp; Galindo, 2014" authorityName="Kool &amp; Galindo" authorityYear="2014" box="[774,1340,1737,1763]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="vanuatuensis">
<emphasis id="7299EABBC73B782B4E7AF957FBF620F2" box="[774,1055,1738,1763]" italics="true" pageId="10" pageNumber="11">Nassarius vanuatuensis</emphasis>
<bibRefCitation id="247C4B58C73B782B4956F954FAD520F2" author="Kool H. H. &amp; Galindo L. A." box="[1066,1340,1737,1763]" pageId="10" pageNumber="11" pagination="147 - 164" refId="ref27751" refString="Kool H. H. &amp; Galindo L. A. 2014. Description and molecular characterization of six new species of Nassarius (Gastropoda, Nassariidae) from the Western Pacific Ocean. American Malacological Bulletin 32: 147 - 164. http: // dx. doi. org / 10.4003 / 006.032.0202" type="journal article" year="2014">Kool &amp; Galindo, 2014</bibRefCitation>
</taxonomicName>
, can be distinguished from
<taxonomicName id="87ED4D2AC73B782B4CB1F970FDB22117" authorityName="Iredale" authorityYear="1936" box="[461,603,1773,1798]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4CB1F970FDB22117" box="[461,603,1773,1798]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
by the protoconch features. Additionally,
<bibRefCitation id="247C4B58C73B782B4925F971FA9E2116" author="Kool H. H. &amp; Galindo L. A." box="[1113,1399,1772,1799]" pageId="10" pageNumber="11" pagination="147 - 164" refId="ref27751" refString="Kool H. H. &amp; Galindo L. A. 2014. Description and molecular characterization of six new species of Nassarius (Gastropoda, Nassariidae) from the Western Pacific Ocean. American Malacological Bulletin 32: 147 - 164. http: // dx. doi. org / 10.4003 / 006.032.0202" type="journal article" year="2014">Kool &amp; Galindo (2014)</bibRefCitation>
demonstrated a 19% genetic distance between
<taxonomicName id="87ED4D2AC73B782B4FA5F88DFC072138" authorityName="Kool &amp; Galindo" authorityYear="2014" box="[729,1006,1808,1833]" class="Gastropoda" family="Nassariidae" genus="Nassarius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="vanuatuensis">
<emphasis id="7299EABBC73B782B4FA5F88DFC072138" box="[729,1006,1808,1833]" italics="true" pageId="10" pageNumber="11">Nassarius vanuatuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="87ED4D2AC73B782B4959F88DFAF22138" baseAuthorityName="Gould" baseAuthorityYear="1850" box="[1061,1307,1808,1833]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="paupera">
<emphasis id="7299EABBC73B782B4959F88DFAF22138" box="[1061,1307,1808,1833]" italics="true" pageId="10" pageNumber="11">Reticunassa paupera</emphasis>
</taxonomicName>
.
<taxonomicName id="87ED4D2AC73B782B4854F88DFE05215F" authority="(Bozzetti, 2006)" baseAuthorityName="Bozzetti" baseAuthorityYear="2006" class="Gastropoda" family="Nassariidae" genus="Naytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="priscardi">
<emphasis id="7299EABBC73B782B4854F88DFECE215C" italics="true" pageId="10" pageNumber="11">Naytia priscardi</emphasis>
(
<bibRefCitation id="247C4B58C73B782B4C4AF8AEFE0D215F" author="Bozzetti L." box="[310,484,1843,1870]" pageId="10" pageNumber="11" pagination="17" refId="ref26589" refString="Bozzetti L. 2006. Nassarius (Hima) priscardi (Gastropoda: Prosobranchia: Nassariidae) nuova specie dal Madagascar Meridionale. Malacologia Mostra Mondiale 53: 17." type="journal article" year="2006">Bozzetti, 2006</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="38FA7639C73B782B4F4EF8AEFD29215C" box="[562,704,1843,1869]" name="Madagascar" pageId="10" pageNumber="11">Madagascar</collectingCountry>
is a member of the sister group of
<taxonomicName id="87ED4D2AC73B782B4928F8A9FB0B215C" authorityName="Iredale" authorityYear="1936" box="[1108,1250,1844,1869]" class="Gastropoda" family="Nassariidae" genus="Reticunassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4928F8A9FB0B215C" box="[1108,1250,1844,1869]" italics="true" pageId="10" pageNumber="11">Reticunassa</emphasis>
</taxonomicName>
and exhibits great morphological similarity to some species of this genus. Molecular synapomorphies indicate that
<taxonomicName id="87ED4D2AC73B782B4DC1F8E7FEA02182" baseAuthorityName="Bozzetti" baseAuthorityYear="2006" box="[189,329,1913,1939]" class="Gastropoda" family="Nassariidae" genus="Naytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="priscardi">
<emphasis id="7299EABBC73B782B4DC1F8E7FEA02182" box="[189,329,1913,1939]" italics="true" pageId="10" pageNumber="11">N. priscardi</emphasis>
</taxonomicName>
belongs to
<taxonomicName id="87ED4D2AC73B782B4CAEF8E7FDC92182" authorityName="Adams &amp; Adams" authorityYear="1853" box="[466,544,1914,1939]" class="Gastropoda" family="Nassariidae" genus="Naytia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7299EABBC73B782B4CAEF8E7FDC92182" box="[466,544,1914,1939]" italics="true" pageId="10" pageNumber="11">Naytia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>