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<document id="E39F70D7C745E3669519CF88F277C7E0" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFD8FFD5219D1604EDF9F8DB" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Protomedetera uncinata Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="321" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="319" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6D0488E19F4359060F29E64A6F861236">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
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<mods:namePart id="DF40B4A90177158E5CB061CCF49BCE7E">Grootaert, Patrick</mods:namePart>
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<mods:namePart id="8F7A5C957DA40B02B6B9E4FB292E6185">Velde,</mods:namePart>
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<mods:role id="4E9028EBB9F49AE5D3E6545CD13227C7">
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<mods:namePart id="56165325B5B197D909357D28082895D6">Isabella Van de</mods:namePart>
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<mods:title id="37FF690C2FF9C04AECDB3DF7ACFB884A">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="7E517B3F3E16E8D353013C95568D2BA1">2024</mods:date>
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<mods:number id="516DE1308823D7203492B3827818CBF7">2024-09-19</mods:number>
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<treatment id="461287F6FFD8FFD5219D1604EDF9F8DB" ID-DOI="http://doi.org/10.5281/zenodo.14683106" ID-Zenodo-Dep="14683106" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB" httpUri="http://treatment.plazi.org/id/461287F6FFD8FFD5219D1604EDF9F8DB" lastPageId="18" lastPageNumber="321" pageId="16" pageNumber="319">
<subSubSection id="86A1656BFFD8FFD7219D1604ED68FA95" box="[946,1363,1378,1403]" pageId="16" pageNumber="319" type="nomenclature">
<paragraph id="CE0436E0FFD8FFD7219D1604ED68FA95" blockId="16.[946,1363,1378,1435]" box="[946,1363,1378,1403]" pageId="16" pageNumber="319">
<heading id="954C818CFFD8FFD7219D1604ED68FA95" bold="true" box="[946,1363,1378,1403]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="319" reason="2">
<emphasis id="FCCFEAF2FFD8FFD7219D1604ED68FA95" bold="true" box="[946,1363,1378,1403]" pageId="16" pageNumber="319">
<taxonomicName id="09BB4D63FFD8FFD7219D1604EC84FA95" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[946,1215,1378,1403]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD8FFD7219D1604EC84FA95" bold="true" box="[946,1215,1378,1403]" italics="true" pageId="16" pageNumber="319">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD726E11605ED68FA95" box="[1230,1363,1379,1403]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7261516E5ECF7FA75" box="[1082,1228,1411,1435]" pageId="16" pageNumber="319" type="description">
<paragraph id="CE0436E0FFD8FFD7261516E5ECF7FA75" blockId="16.[946,1363,1378,1435]" box="[1082,1228,1411,1435]" pageId="16" pageNumber="319">
(
<figureCitation id="56802A65FFD8FFD7266D16E5ECFFFA75" box="[1090,1220,1411,1435]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="16.[838,875,1049,1071]" captionStartId-1="17.[113,149,827,849]" captionStartId-2="17.[813,848,1205,1227]" captionTargetBox-0="[836,1472,189,1034]" captionTargetBox-1="[113,749,189,811]" captionTargetBox-2="[816,1451,194,1187]" captionTargetId-0="figure-264@16.[836,1472,189,1034]" captionTargetId-1="figure-15@17.[113,749,189,811]" captionTargetId-2="figure-17@17.[798,1469,176,1197]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="17" captionText-0="Fig. 16. Protomedetera uncinata, new species, male habitus (ZRCBDP0085280). Photograph by Kristy Chang." captionText-1="Fig. 17. Protomedetera uncinata, new species, female habitus (ZRCBDP008400). Photograph by Kristy Chang." captionText-2="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi-0="http://doi.org/10.5281/zenodo.14683094" figureDoi-1="http://doi.org/10.5281/zenodo.14683098" figureDoi-2="http://doi.org/10.5281/zenodo.14683100" httpUri-0="https://zenodo.org/record/14683094/files/figure.png" httpUri-1="https://zenodo.org/record/14683098/files/figure.png" httpUri-2="https://zenodo.org/record/14683100/files/figure.png" pageId="16" pageNumber="319">Figs. 1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7216C16A5ED97F935" pageId="16" pageNumber="319" type="diagnosis">
<paragraph id="CE0436E0FFD8FFD7216C16A5ED97F935" blockId="16.[834,1476,1475,1755]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C16A5EB81FA35" bold="true" box="[835,954,1475,1499]" pageId="16" pageNumber="319">Diagnosis.</emphasis>
Small species with shiny metallic green head and thorax. Antenna yellow, but pedicel with almost black apex. Pedicel with 3 long marginal bristles covering the spheroid postpedicel. Anterior half uniformly covered with short bristles; 4-serial acrostichals indistinctly separated from dorsocentrals. Only the 2 prescutellar dorsocentral bristles long. All coxae and femora blackish brown, tibiae and tarsi yellow. Tip hypandrium very wide, forked, and dorsally bent. Surstylus lacking squamiform bristles. Cerci small, oval.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7216C1465EBF7F8B5" pageId="16" pageNumber="319" type="etymology">
<paragraph id="CE0436E0FFD8FFD7216C1465EBF7F8B5" blockId="16.[835,1475,1795,1883]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C1465EBFDF8F5" bold="true" box="[835,966,1795,1819]" pageId="16" pageNumber="319">Etymology.</emphasis>
The name
<taxonomicName id="09BB4D63FFD8FFD726661462EC9CF8F5" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1097,1191,1796,1819]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="FCCFEAF2FFD8FFD726661462EC9CF8F5" box="[1097,1191,1796,1819]" italics="true" pageId="16" pageNumber="319">uncinata</emphasis>
</taxonomicName>
(Lat. Feminine uncīnāta: bearing hooks) refers to hook-like bifurcate tip of the hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD5216C14E5E973FA9B" lastPageId="18" lastPageNumber="321" pageId="16" pageNumber="319" type="materials_examined">
<paragraph id="CE0436E0FFD8FFD7216C14E5EB8AF815" blockId="16.[834,1475,1923,2043]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C14E5EC1BF875" bold="true" box="[835,1056,1923,1947]" pageId="16" pageNumber="319">Material examined.</emphasis>
<materialsCitation id="7ED33CBDFFD8FFD7260A14E5EB8AF815" collectingDate="2016-03-16" collectionCode="LKCNHM" collectorName="P. Grootaert &amp; GenBank" country="Singapore" location="Pulau Ubin" pageId="16" pageNumber="319" specimenCode="PU07, ZRCBDP0085280" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFD8FFD7260A14E5ECB3F875" box="[1061,1160,1923,1947]" pageId="16" pageNumber="319" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFD8FFD726E214E5ED07F875" box="[1229,1340,1923,1947]" name="Singapore" pageId="16" pageNumber="319">Singapore</collectingCountry>
,
<location id="CB64603BFFD8FFD7276B14E5ED85F875" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD8FFD7276B14E5ED85F875" box="[1348,1470,1923,1947]" country="Singapore" name="Pulau Ubin" pageId="16" pageNumber="319">Pulau Ubin</location>
, (
<specimenCode id="9E1D9E9BFFD8FFD7216614C5EBB1F855" box="[841,906,1955,1979]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU07</specimenCode>
)
<date id="BA051020FFD8FFD721B314C5EC77F855" box="[924,1100,1955,1979]" pageId="16" pageNumber="319" value="2016-03-16">
<collectingDate id="AA41E9C8FFD8FFD721B314C5EC77F855" box="[924,1100,1955,1979]" pageId="16" pageNumber="319" value="2016-03-16">16 March 2016</collectingDate>
</date>
,
<collectingMethod id="17FA4EF7FFD8FFD7267514C5ECDEF854" box="[1114,1253,1955,1979]" pageId="16" pageNumber="319">Malaise trap</collectingMethod>
(
<specimenCode id="9E1D9E9BFFD8FFD726D814C5EDFAF855" box="[1271,1473,1955,1979]" collectionCode="ZRCBDP" pageId="16" pageNumber="319">ZRCBDP0085280</specimenCode>
in
<collectionCode id="A8AAAE25FFD8FFD7214F14A5EBECF835" box="[864,983,1987,2011]" pageId="16" pageNumber="319">LKCNHM</collectionCode>
; leg.
<collectorName id="634E5336FFD8FFD7263D14A5ECA0F835" box="[1042,1179,1987,2011]" pageId="16" pageNumber="319">P. Grootaert</collectorName>
).
<collectorName id="634E5336FFD8FFD7268214A5ED29F835" box="[1197,1298,1987,2011]" pageId="16" pageNumber="319">GenBank</collectorName>
accession code: PP893312
</materialsCitation>
</paragraph>
<caption id="9AC46668FFD9FFD6225E105DEA6FFC83" ID-DOI="http://doi.org/10.5281/zenodo.14683098" ID-Zenodo-Dep="14683098" httpUri="https://zenodo.org/record/14683098/files/figure.png" pageId="17" pageNumber="320" startId="17.[113,149,827,849]" targetBox="[113,749,189,811]" targetPageId="17" targetType="figure">
<paragraph id="CE0436E0FFD9FFD6225E105DEA6FFC83" blockId="17.[113,748,827,877]" pageId="17" pageNumber="320">
Fig. 17.
<taxonomicName id="09BB4D63FFD9FFD622E5105DE984FCBE" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[202,447,827,848]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="320" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD9FFD622E5105DE984FCBE" box="[202,447,827,848]" italics="true" pageId="17" pageNumber="320">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD9FFD623E0105DEA70FCBF" box="[463,587,827,849]" pageId="17" pageNumber="320" rank="species">new species</taxonomicNameLabel>
, female habitus (ZRCBDP008400). Photograph by Kristy Chang.
</paragraph>
</caption>
<paragraph id="CE0436E0FFD9FFD6225E10C3E912FA13" blockId="17.[113,755,933,1533]" pageId="17" pageNumber="320">
<materialsCitation id="7ED33CBDFFD9FFD6225E10C3E91EFA13" collectingDate="2016-03-16" collectingDateMax="2019-04-02" collectingDateMin="2016-03-16" collectionCode="ZRCBDP, PU" country="Singapore" county="GenBank" location="GenBank" municipality="GenBank" pageId="17" pageNumber="320" specimenCode="ZRCBDP0081888, PU01, ZRCBDP0084331, ZRCBDP0084003, ZRCBDP0085270, PU04, ZRCBDP0085236, ZRCBDP0085235, ZRCBDP0085269, ZRCBDP0085280, PU07, ZRCBDP0083285, PU10, ZRCBDP0257071, PU19, ZRCBDP0256595, ZRCBDP0317777, PU24" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFD9FFD6225E10C3E8D0FC53" bold="true" box="[113,235,933,957]" pageId="17" pageNumber="320">
<typeStatus id="11008842FFD9FFD6225E10C3E8DDFC53" box="[113,230,933,957]" pageId="17" pageNumber="320" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFD9FFD622DA10C3E9B5FC53" box="[245,398,933,957]" name="Singapore" pageId="17" pageNumber="320">SINGAPORE</collectingCountry>
, female,
<specimenCode id="9E1D9E9BFFD9FFD623DA10C3EA86FC53" box="[501,701,933,957]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0081888</specimenCode>
_12_
<specimenCode id="9E1D9E9BFFD9FFD6225E10A2E894FC32" box="[113,175,964,988]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_
<date id="BA051020FFD9FFD6229010A2E97AFC32" box="[191,321,964,988]" pageId="17" pageNumber="320" value="2016-08-10">
<collectingDate id="AA41E9C8FFD9FFD6229010A2E97AFC32" box="[191,321,964,988]" pageId="17" pageNumber="320" value="2016-08-10">10Aug2016</collectingDate>
</date>
;
<location id="CB64603BFFD9FFD6236010A3E98EFC33" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6236010A3E98EFC33" box="[335,437,965,989]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893313; female
<specimenCode id="9E1D9E9BFFD9FFD622E51082E99BFC12" box="[202,416,996,1020]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0084331</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD623811082E9D6FC12" box="[430,493,996,1020]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_
<date id="BA051020FFD9FFD623D31082EA4DFC12" box="[508,630,996,1020]" pageId="17" pageNumber="320" value="2016-06-01">
<collectingDate id="AA41E9C8FFD9FFD623D31082EA4DFC12" box="[508,630,996,1020]" pageId="17" pageNumber="320" value="2016-06-01">01Jun2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD620A81083EACBFC13" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD620A81083EACBFC13" box="[647,752,997,1021]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893314; female
<specimenCode id="9E1D9E9BFFD9FFD6203B1763EAD8FBF3" box="[532,739,1029,1053]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0084003</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E1743E889FBD3" box="[113,178,1061,1085]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_07
<location id="CB64603BFFD9FFD622F21743E937FBD3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD622F21743E937FBD3" box="[221,268,1061,1085]" country="Singapore" county="GenBank" municipality="GenBank" name="Apr" pageId="17" pageNumber="320">Apr</location>
201 (imaged),
<location id="CB64603BFFD9FFD623911743EA11FBD3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD623911743EA11FBD3" box="[446,554,1061,1085]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893315; female
<specimenCode id="9E1D9E9BFFD9FFD623631722EA21FBB2" box="[332,538,1092,1116]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085270</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620071722EA5CFBB2" box="[552,615,1092,1116]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD6205A1722EAD6FBB2" box="[629,749,1092,1116]" pageId="17" pageNumber="320" value="2016-06-08">
<collectingDate id="AA41E9C8FFD9FFD6205A1722EAD6FBB2" box="[629,749,1092,1116]" pageId="17" pageNumber="320" value="2016-06-08">08Jun2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD6225D1703E8D6FB93" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6225D1703E8D6FB93" box="[114,237,1125,1149]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893316; female
<specimenCode id="9E1D9E9BFFD9FFD6225A17E2E953FB72" box="[117,360,1156,1180]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085236</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6235717E2E9F8FB72" box="[376,451,1156,1180]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD623FC17E2EA5DFB72" box="[467,614,1156,1180]" pageId="17" pageNumber="320" value="2016-04-07">
<collectingDate id="AA41E9C8FFD9FFD623FC17E2EA5DFB72" box="[467,614,1156,1180]" pageId="17" pageNumber="320" value="2016-04-07">07Apr2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD6205317E3EACAFB73" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6205317E3EACAFB73" box="[636,753,1157,1181]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893317; female
<specimenCode id="9E1D9E9BFFD9FFD6203917C2EADDFB52" box="[534,742,1188,1212]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085235</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E17A2E884FB32" box="[113,191,1220,1244]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD622FE17A2E950FB32" box="[209,363,1220,1244]" pageId="17" pageNumber="320" value="2016-04-07">
<collectingDate id="AA41E9C8FFD9FFD622FE17A2E950FB32" box="[209,363,1220,1244]" pageId="17" pageNumber="320" value="2016-04-07">07Apr2016</collectingDate>
</date>
;
<location id="CB64603BFFD9FFD623AC17A3E9C5FB33" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD623AC17A3E9C5FB33" box="[387,510,1221,1245]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893317; female
<specimenCode id="9E1D9E9BFFD9FFD6237E1782EA1AFB12" box="[337,545,1252,1276]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085269</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620001782EA55FB12" box="[559,622,1252,1276]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD620521782EAC8FB12" box="[637,755,1252,1276]" pageId="17" pageNumber="320" value="2016-06-01">
<collectingDate id="AA41E9C8FFD9FFD620521782EAC8FB12" box="[637,755,1252,1276]" pageId="17" pageNumber="320" value="2016-06-01">01Jun2016</collectingDate>
</date>
(Ma9036);
<location id="CB64603BFFD9FFD622DC1663E960FAF3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD622DC1663E960FAF3" box="[243,347,1285,1309]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893319; female
<specimenCode id="9E1D9E9BFFD9FFD6225D1642E976FAD2" box="[114,333,1316,1340]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085280</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD623731642E9A4FAD2" box="[348,415,1316,1340]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU07</specimenCode>
_
<date id="BA051020FFD9FFD623821642EA08FAD2" box="[429,563,1316,1340]" pageId="17" pageNumber="320" value="2016-03-16">
<collectingDate id="AA41E9C8FFD9FFD623821642EA08FAD2" box="[429,563,1316,1340]" pageId="17" pageNumber="320" value="2016-03-16">16Mar2016</collectingDate>
</date>
(mitogenome);
<specimenCode id="9E1D9E9BFFD9FFD6225C1622E969FAB2" box="[115,338,1348,1372]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0083285</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6234E1622E998FAB2" box="[353,419,1348,1372]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU10</specimenCode>
,
<location id="CB64603BFFD9FFD6239B1623EA18FAB3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6239B1623EA18FAB3" box="[436,547,1349,1373]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893320;
<specimenCode id="9E1D9E9BFFD9FFD623231602E9CDFA92" box="[268,502,1380,1404]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0257071</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620281602EA77FA92" box="[519,588,1380,1404]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU19</specimenCode>
_
<date id="BA051020FFD9FFD620731602EAD1FA92" box="[604,746,1380,1404]" pageId="17" pageNumber="320" value="2018-04-26">
<collectingDate id="AA41E9C8FFD9FFD620731602EAD1FA92" box="[604,746,1380,1404]" pageId="17" pageNumber="320" value="2018-04-26">26Apr2018</collectingDate>
</date>
;
<collectingMunicipality id="2E60AC9AFFD9FFD6225B16E3E8E1FA73" box="[116,218,1413,1437]" pageId="17" pageNumber="320">GenBank</collectingMunicipality>
accession code: PP893321;
<specimenCode id="9E1D9E9BFFD9FFD6203216E2EADEFA72" box="[541,741,1412,1436]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0256595</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E16C3E894FA53" box="[113,175,1445,1469]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU19</specimenCode>
_
<date id="BA051020FFD9FFD6229216C3E978FA53" box="[189,323,1445,1469]" pageId="17" pageNumber="320" value="2018-05-03">
<collectingDate id="AA41E9C8FFD9FFD6229216C3E978FA53" box="[189,323,1445,1469]" pageId="17" pageNumber="320" value="2018-05-03">03May2018</collectingDate>
</date>
;
<collectingCounty id="27654E6CFFD9FFD6237E16C3E98CFA53" box="[337,439,1445,1469]" pageId="17" pageNumber="320">GenBank</collectingCounty>
accession code: PP893322;
<specimenCode id="9E1D9E9BFFD9FFD6225D16A3E901FA33" box="[114,314,1477,1501]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0317777</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6236716A3E9BEFA33" box="[328,389,1477,1501]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU24</specimenCode>
_
<date id="BA051020FFD9FFD623BD16A3EA37FA33" box="[402,524,1477,1501]" pageId="17" pageNumber="320" value="2019-04-02">
<collectingDate id="AA41E9C8FFD9FFD623BD16A3EA37FA33" box="[402,524,1477,1501]" pageId="17" pageNumber="320" value="2019-04-02">02Apr2019</collectingDate>
</date>
; GenBank accession code: PP893323
</materialsCitation>
.
</paragraph>
<caption id="9AC46668FFD9FFD6210217D3EBF0FAED" ID-DOI="http://doi.org/10.5281/zenodo.14683100" ID-Zenodo-Dep="14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="17" pageNumber="320" startId="17.[813,848,1205,1227]" targetBox="[816,1451,194,1187]" targetPageId="17" targetType="figure">
<paragraph id="CE0436E0FFD9FFD6210217D3EBF0FAED" blockId="17.[813,1453,1205,1284]" pageId="17" pageNumber="320">
Fig. 18.
<taxonomicName id="09BB4D63FFD9FFD621AB17D3EC4FFB24" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[900,1140,1205,1226]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="320" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD9FFD621AB17D3EC4FFB24" box="[900,1140,1205,1226]" italics="true" pageId="17" pageNumber="320">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD9FFD626AC17D3ECC5FB25" box="[1155,1278,1205,1227]" pageId="17" pageNumber="320" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFD9FFD6225E1543E8E1F9B2" blockId="17.[113,752,1572,1628]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E1543E889F9D3" bold="true" box="[113,178,1573,1597]" pageId="17" pageNumber="320">Male.</emphasis>
Length: body
<quantity id="09439B05FFD9FFD6237A1543E982F9D2" box="[341,441,1573,1597]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="17" pageNumber="320" unit="mm" value="1.79">1.79 mm</quantity>
; wing
<quantity id="09439B05FFD9FFD6202B1542EA53F9D2" box="[516,616,1572,1596]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="17" pageNumber="320" unit="mm" value="1.66">1.66 mm</quantity>
; wing width
<quantity id="09439B05FFD9FFD6225E1522E8EDF9B2" box="[113,214,1604,1628]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="320" unit="mm" value="0.55">0.55 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFD9FFD6225E15E3E90FF893" blockId="17.[113,753,1669,1917]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E15E3E88EF973" bold="true" box="[113,181,1669,1693]" pageId="17" pageNumber="320">Head.</emphasis>
Frons and face shiny metallic green. Face triangular, wide below antenna, narrowing down to clypeus, there a little wider than pedicel; indented near middle with a furrow. Clypeus narrow. Hairs and bristles on head brownish. A pair of strong retroclinate ocellar bristles, a pair of strong proclinate vertical bristles (
<figureCitation id="56802A65FFD9FFD623801443EA38F8D2" box="[431,515,1828,1853]" captionStart="Fig" captionStartId="16.[838,875,1049,1071]" captionTargetBox="[836,1472,189,1034]" captionTargetId="figure-264@16.[836,1472,189,1034]" captionTargetPageId="16" captionText="Fig. 16. Protomedetera uncinata, new species, male habitus (ZRCBDP0085280). Photograph by Kristy Chang." figureDoi="http://doi.org/10.5281/zenodo.14683094" httpUri="https://zenodo.org/record/14683094/files/figure.png" pageId="17" pageNumber="320">Fig. 16</figureCitation>
) and a pair of short postvertical bristles pointing inward. Postocular bristles uniseriate, brown.
</paragraph>
<paragraph id="CE0436E0FFD9FFD6225E14C2ED96F9FD" blockId="17.[113,753,1956,2109]" lastBlockId="17.[812,1453,1339,1555]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E14C2E8DDF852" bold="true" box="[113,230,1956,1980]" pageId="17" pageNumber="320">Antennae.</emphasis>
Scape yellow, short, with an interior projection. Pedicel yellow, cup-like, large, with postpedicel sunken in it; in the apical crown of bristles 3 long apical bristles longer than and bending over the postpedicel; below the long ventral apical bristle a second a long finer bristle; with some additional short, scattered bristles, most at the inner side. Postpedicel with apex dark brown, spheroid, with short pubescence and a subapical long arista (shifted toward outside). All bristles on postpedicel brown. Arista 3 times as long as all antennal segments together; black, naked, basal portion of arista short. Palpus brown, small, circular, with a long brown subapical bristle, and a second bristle beneath.
</paragraph>
<paragraph id="CE0436E0FFD9FFD62102155DED90F8BD" blockId="17.[811,1452,1595,1876]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD62102155DEBB3F9BD" bold="true" box="[813,904,1595,1619]" pageId="17" pageNumber="320">Thorax.</emphasis>
Brownish metallic (pollinosity probably absent, not visible in the ethanol conserved specimen). Basal half of mesonotum uniformly covered with short bristles, the rows of 4 serial acrostichals indistinctly separated from dc. Apical half of mesonotum flat before scutellum, lacking bristles in central area. Hairs and bristles on thorax brownish black; 2 long prescutellar dorsocentral bristles, anterior dorsocentrals short followed by 2 long prescutellar dorsocentrals. A pair of very long crossing scutellars with a short bristle at outside.
</paragraph>
<paragraph id="CE0436E0FFD9FFD62102141DED57F83D" blockId="17.[813,1452,1915,2003]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD62102141DEB52F87D" bold="true" box="[813,873,1915,1939]" pageId="17" pageNumber="320">Legs.</emphasis>
Mainly brown but trochanters, tip of all femora, all tibiae and tarsomeres yellow except tarsomere 5 of all legs brownish. Hairs and bristles on legs brownish black.
</paragraph>
<paragraph id="CE0436E0FFD9FFD52102149AEA84FF1B" blockId="17.[813,1451,2043,2099]" lastBlockId="18.[136,775,189,245]" lastPageId="18" lastPageNumber="321" pageId="17" pageNumber="320">Fore legs. Coxa densely covered with short pale bristles, long apical bristles; one bristle in middle longer than the others. Fore femur with short pale ventrals and no distinct posterior preapical bristles. Tibia with short apicals.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7127BEAA2FE3B" blockId="18.[135,776,285,469]" pageId="18" pageNumber="321">Mid legs. Coxa with short anterior bristles and a long pale brownish exterior bristle as long as coxa is long. Femur with a fine preapical anterior bristle. Tibia with a short anterodorsal and a longer posterodorsal near base; a stronger anterodorsal at ⅓ of base and a small anterodorsal in preapical crown; ventral preapical twice as long as width of tibia.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7129BE921FD5B" blockId="18.[136,776,509,693]" pageId="18" pageNumber="321">Hind legs. Coxa with a short brownish exterior bristle near middle. Femur in apical half with a row of short brown anteroventral bristles; preapical anterior bristles indistinct. Hind femur with indistinct preapical bristles. Hind tibia lacking distinct bristles, apart from the short bristles in apical crown.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A711BBE919FCDB" blockId="18.[136,775,733,821]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A711BBE8E8FD1B" bold="true" box="[136,211,733,757]" pageId="18" pageNumber="321">Wings.</emphasis>
Hyaline, tinged yellowish grey. Veins yellowish brown. Squama white, with a single short brown bristle. Haltere white.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7103BE96CFC9B" blockId="18.[136,343,861,885]" box="[136,343,861,885]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7103BE8C5FC9B" bold="true" box="[136,254,861,885]" pageId="18" pageNumber="321">Abdomen.</emphasis>
Brown.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A710FBE90FFB1B" blockId="18.[135,777,925,1269]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A710FBE968FC5B" bold="true" box="[136,339,925,949]" pageId="18" pageNumber="321">Male terminalia.</emphasis>
<figureCitation id="56802A65FFDAFFD5237110FBE98FFC5B" box="[350,436,925,949]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18</figureCitation>
. Foramen situated at base of epandrium. Surstylus brown, wide truncate composed of a single element supposed to be the dorsal surstylus. Apex surstylus rounded with 3 short and 3 long bristles on the ventral side, and 2 strong subapical bristles on the lateral side (
<figureCitation id="56802A65FFDAFFD522EB175BE91DFBBB" box="[196,294,1085,1109]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
). Cercus short oval, with a short and a very long apical bristle (
<figureCitation id="56802A65FFDAFFD52379173BE98EFB9B" box="[342,437,1117,1141]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
), other bristles short (
<figureCitation id="56802A65FFDAFFD520B3173BEAC7FB9B" box="[668,764,1117,1141]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18C</figureCitation>
). Hypandrium bifurcate (
<figureCitation id="56802A65FFDAFFD523BF171BE9C2FB7B" box="[400,505,1149,1173]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18A</figureCitation>
), the fork bent dorsally (
<figureCitation id="56802A65FFDAFFD522BF17FBE8CBFB5B" box="[144,240,1181,1205]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
). Phallus hidden below hypandrium. Apicoventral epandrial lobe lacking, but 3 almost equally long epandrial bristles present.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7167BE973FA9B" blockId="18.[136,775,1309,1397]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7167BE8D9FADB" bold="true" box="[136,226,1309,1333]" pageId="18" pageNumber="321">Female.</emphasis>
Length: body
<quantity id="09439B05FFDAFFD523A9167BEA3EFADB" box="[390,517,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="1.775" metricValueMax="2.0" metricValueMin="1.55" pageId="18" pageNumber="321" unit="mm" value="1.775" valueMax="2.0" valueMin="1.55">1.552 mm</quantity>
; wing
<quantity id="09439B05FFDAFFD5207E167BEADFFADB" box="[593,740,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="2.2" metricValueMin="1.64" pageId="18" pageNumber="321" unit="mm" value="1.92" valueMax="2.2" valueMin="1.64">1.642.2 mm</quantity>
. In most aspects, resembles male as there are no secondary sexual characters.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFDAFFD522A716FBEDF9F8DB" pageId="18" pageNumber="321" type="discussion">
<paragraph id="CE0436E0FFDAFFD522A716FBEA73F9DB" blockId="18.[136,775,1437,1589]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A716FBE936FA5B" bold="true" box="[136,269,1437,1461]" pageId="18" pageNumber="321">Comments.</emphasis>
Variability is noted in the darkening of the postpedicel from brown with a paler base to almost black apex.
<taxonomicName id="09BB4D63FFDAFFD522E716BBE9F0FA1B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[200,459,1501,1525]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFDAFFD522E716BBE9F0FA1B" box="[200,459,1501,1525]" italics="true" pageId="18" pageNumber="321">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFDAFFD523F616B8EA60FA1B" box="[473,603,1501,1525]" pageId="18" pageNumber="321" rank="species">new species</taxonomicNameLabel>
, differs from all other known
<taxonomicName id="09BB4D63FFDAFFD52336169BE982F9FB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[281,441,1533,1557]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD52336169BE982F9FB" box="[281,441,1533,1557]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
by the strong forked tip of the hypandrium. The forks are bent dorsally.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7153BEACAF8BB" blockId="18.[135,775,1629,1877]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7153BE9A5F99B" bold="true" box="[136,414,1629,1653]" pageId="18" pageNumber="321">National distribution.</emphasis>
Thanks to the barcoding,
<specimenCount id="D8BDFD69FFDAFFD520C4153BE8C2F97B" count="13" pageId="18" pageNumber="321" type="generic">13 specimens</specimenCount>
were identified.
<taxonomicName id="09BB4D63FFDAFFD52393151BEAF3F97B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[444,712,1661,1685]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFDAFFD52393151BEAF3F97B" box="[444,712,1661,1685]" italics="true" pageId="18" pageNumber="321">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFDAFFD520F51518E8E3F95B" pageId="18" pageNumber="321" rank="species">new species</taxonomicNameLabel>
, is the most abundant
<taxonomicName id="09BB4D63FFDAFFD523F815FBEAF5F95B" box="[471,718,1693,1717]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD523F815FBEA4CF95B" box="[471,631,1693,1717]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
species
</taxonomicName>
ever found in
<collectingCountry id="B6AC7670FFDAFFD522C015DBE964F93B" box="[239,351,1725,1749]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
; it is exclusive to Pulau Ubin. All the records are from back mangroves, except for a single record from a coastal forest (PU26). This site is very close to a back mangrove, so sample contamination could have occurred during flooding of the area by a storm and a spring tide.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7141BEA43F85B" blockId="18.[136,775,1917,1973]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7141BE9FDF87B" bold="true" box="[136,454,1917,1941]" pageId="18" pageNumber="321">
Other
<taxonomicName id="09BB4D63FFDAFFD522FF141BE9FDF87B" box="[208,454,1917,1941]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFDAFFD522FF141BE955F87B" bold="true" box="[208,366,1917,1941]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
species.
</taxonomicName>
</emphasis>
Two more unidentified species of
<taxonomicName id="09BB4D63FFDAFFD5228814FBE97CF85B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[167,327,1949,1973]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5228814FBE97CF85B" box="[167,327,1949,1973]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
were found on Pulau Ubin.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A714BBED84FEFB" blockId="18.[136,776,2013,2101]" lastBlockId="18.[835,1474,189,277]" pageId="18" pageNumber="321">
1. A female of
<taxonomicName id="09BB4D63FFDAFFD5230014BBE9F4F81B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[303,463,2013,2037]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5230014BBE9F4F81B" box="[303,463,2013,2037]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
with quadriserial acrostichal bristles, 4 long dorsocentrals, and yellow hind metatarsus was found in station PU01 on
<date id="BA051020FFDAFFD523C31B7BEAA7F7DB" box="[492,668,2077,2101]" pageId="18" pageNumber="321" value="2019-03-19">19 March 2019</date>
(labelled sp4). It seems to be different from the other three species reported above. Unfortunately, no barcode is available yet to give an indication if it might belong to another species.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C125BEB82FCFB" blockId="18.[834,1475,317,789]" pageId="18" pageNumber="321">
2. A female, provisionally labelled as sp5 (PU01, ZRCBDP0083912, image on BOS: https://singapore. biodiversity.online/species/A-Arth-Hexa-Diptera-002292) has the proboscis as long as head is high, which is unusually long for
<taxonomicName id="09BB4D63FFDAFFD521B512DBEC02FE3B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[922,1081,445,469]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD521B512DBEC02FE3B" box="[922,1081,445,469]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. It has 2 long dorsocentrals, but since the specimen was severely damaged, some dorsocentrals may have been lost. The basal tarsomere of the hind tarsus is slender and longer than the following tarsomere, which is also unusual for
<taxonomicName id="09BB4D63FFDAFFD5263C115BEC89FDBB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1043,1202,573,597]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5263C115BEC89FDBB" box="[1043,1202,573,597]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. Since the specimen was destroyed during extraction for sequencing, no nomenclatorial actions can be taken. The mitogenome was successful and the species clusters at the outside of the other
<taxonomicName id="09BB4D63FFDAFFD5273411FBED81FD5B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1307,1466,669,693]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5273411FBED81FD5B" box="[1307,1466,669,693]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. Following barcode is available (courtesy of Darren Yeo): ZRCBDP0083912 sp5 PU01, GenBank accession code: PP893324.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C105BEC48FC5B" blockId="18.[835,1476,829,949]" pageId="18" pageNumber="321">
<materialsCitation id="7ED33CBDFFDAFFD5216C105BEC54FC5B" collectingDate="2018-03-22" collectionCode="PU, ZRCBDP, PP" pageId="18" pageNumber="321" specimenCode="PU12, ZRCBDP0281845, PP893325" specimenCount="1">
3. A female belonging to the
<taxonomicName id="09BB4D63FFDAFFD526AC1058ECD2FCBB" box="[1155,1257,829,853]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD526AC1058ECD2FCBB" box="[1155,1257,829,853]" italics="true" pageId="18" pageNumber="321">P. glabra</emphasis>
</taxonomicName>
group was found in a strip of secondary forest in front of Endut Senit Campsite (
<specimenCode id="9E1D9E9BFFDAFFD52163101BEBB4FC7B" box="[844,911,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="18" pageNumber="321">PU12</specimenCode>
,
<specimenCode id="9E1D9E9BFFDAFFD521B0101BEC54FC7B" box="[927,1135,893,917]" collectionCode="ZRCBDP" pageId="18" pageNumber="321">ZRCBDP0281845</specimenCode>
_
<specimenCode id="9E1D9E9BFFDAFFD52651101BEC80FC7B" box="[1150,1211,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="18" pageNumber="321">PU12</specimenCode>
_
<date id="BA051020FFDAFFD526E5101BED70FC7B" box="[1226,1355,893,917]" pageId="18" pageNumber="321" value="2018-03-22">
<collectingDate id="AA41E9C8FFDAFFD526E5101BED70FC7B" box="[1226,1355,893,917]" pageId="18" pageNumber="321" value="2018-03-22">22Mar2018</collectingDate>
</date>
, GenBank accession code:
<specimenCode id="9E1D9E9BFFDAFFD521DA10FBEC53FC5B" box="[1013,1128,925,949]" collectionCode="PP" pageId="18" pageNumber="321">PP893325</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C10B8EDF8FA9B" blockId="18.[834,1475,989,1397]" pageId="18" pageNumber="321">
As already mentioned above,
<taxonomicName id="09BB4D63FFDAFFD5268110B8ED24FC1B" box="[1198,1311,989,1013]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD5268110B8ED24FC1B" box="[1198,1311,989,1013]" italics="true" pageId="18" pageNumber="321">P. glabra</emphasis>
</taxonomicName>
has different morphology from the type species of
<taxonomicName id="09BB4D63FFDAFFD5273C109BEDF9FBFB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1299,1474,1021,1045]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5273C109BEDF9FBFB" box="[1299,1474,1021,1045]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
and should probably be attributed to another genus. The postpedicel does not bear long apical bristles and the position of the male terminalia on the abdomen is different.
<taxonomicName id="09BB4D63FFDAFFD5216B171BECD1FB7B" authority="Tang et al., 2018" authorityName="Tang" authorityYear="2018" box="[836,1258,1149,1174]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD5216B171BEC0AFB7B" box="[836,1073,1149,1173]" italics="true" pageId="18" pageNumber="321">Protomedetera glabra</emphasis>
<bibRefCitation id="AA2A4B11FFDAFFD526181718ECD1FB7B" author="Tang C &amp; Grootaert P &amp; Yang D" box="[1079,1258,1149,1174]" pageId="18" pageNumber="321" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
</taxonomicName>
was found in
<collectingCountry id="B6AC7670FFDAFFD527AE1718EBF0FB5B" name="Papua New Guinea" pageId="18" pageNumber="321">Papua New Guinea</collectingCountry>
(
<typeStatus id="11008842FFDAFFD521F517FBEC00FB5B" box="[986,1083,1181,1205]" pageId="18" pageNumber="321" type="holotype">holotype</typeStatus>
) and in
<collectingCountry id="B6AC7670FFDAFFD526B917FBED3EFB5B" box="[1174,1285,1181,1205]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
, but although the male terminalia of the specimens from
<collectingCountry id="B6AC7670FFDAFFD526DF17D8EDF9FB3B" box="[1264,1474,1213,1237]" name="Papua New Guinea" pageId="18" pageNumber="321">Papua New Guinea</collectingCountry>
and those from
<collectingCountry id="B6AC7670FFDAFFD521D817BBEC5EFB1B" box="[1015,1125,1245,1269]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
are similar, they are distinctly different and hence the taxonomy of this species group needs revision. The species was generally found in anthropogenic habitats, like Clementi Woods (park), Evolution Garden (Botanic Gardens), NUS (university park), and Sime Forest.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C16FBEDF9F8DB" blockId="18.[834,1476,1437,1846]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD5216C16FBED7CFA5B" bold="true" box="[835,1351,1437,1461]" pageId="18" pageNumber="321">
Comments on
<taxonomicName id="09BB4D63FFDAFFD521C116FBECABFA5B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1006,1168,1437,1461]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD521C116FBECABFA5B" bold="true" box="[1006,1168,1437,1461]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
in Pulau Ubin.
</emphasis>
The genus
<taxonomicName id="09BB4D63FFDAFFD5216C16DBEBD8FA3B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[835,995,1469,1493]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5216C16DBEBD8FA3B" box="[835,995,1469,1493]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
is very rare in
<collectingCountry id="B6AC7670FFDAFFD526AC16DBECD4FA3B" box="[1155,1263,1469,1493]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
and although many sites and various biotopes were surveyed during the last twenty years for prolonged periods, only two species were recorded in the past (
<bibRefCitation id="AA2A4B11FFDAFFD5261C157BECC3F9DB" author="Tang C &amp; Grootaert P &amp; Yang D" box="[1075,1272,1565,1589]" pageId="18" pageNumber="321" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
). With six species now, Pulau Ubin seems to be a hot spot for the genus
<taxonomicName id="09BB4D63FFDAFFD5216C153BEBD8F99B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[835,995,1629,1653]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5216C153BEBD8F99B" box="[835,995,1629,1653]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. It is unclear what the reason might be. On Pulau Ubin, all species were found in mangrove areas or very close to mangroves.
<taxonomicName id="09BB4D63FFDAFFD5260A15FBECFEF95B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1061,1221,1693,1717]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5260A15FBECFEF95B" box="[1061,1221,1693,1717]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
is a tree trunk dweller, but its precise biology is unknown. The presence of lots of dead decaying wood present at the surveyed sites might favour the presence of
<taxonomicName id="09BB4D63FFDAFFD5266F159BECE5F8FB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1088,1246,1789,1813]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5266F159BECE5F8FB" box="[1088,1246,1789,1813]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
, where it might feed on wood inhabiting insects like the related genus
<taxonomicName id="09BB4D63FFDAFFD52779147BED87F8DB" authorityName="Fischer von Waldheim" authorityYear="1819" box="[1366,1468,1821,1845]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD52779147BED87F8DB" box="[1366,1468,1821,1845]" italics="true" pageId="18" pageNumber="321">Medetera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>