Add updates up until 2025-01-18 04:52:11

This commit is contained in:
ggserver 2025-01-18 04:54:21 +00:00
parent 75b7e35b8d
commit 4f3467d244
71 changed files with 19903 additions and 0 deletions

View file

@ -0,0 +1,566 @@
<document id="A529306EDF775926FA953E66CAEF348F" ID-DOI="10.26107/RBZ-2024-0035" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682894" ID-ZooBank="227BB34E-7FF6-492E-A3E3-F82B74C714B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171811771" checkinUser="felipe" docAuthor="Kottelat, Maurice, , Tan Heok Hui, Hasan, &amp; Veryl" docDate="2024" docId="038D87D4AD25FFD6E76AFAF6FC02A267" docLanguage="en" docName="RafflesBZool.72.479-487.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Kapuasia falaris Kottelat, , Hasan &amp; Veryl, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="486" masterDocId="FFB4FFACAD24FFD1E660FFDDFFAFAA52" masterDocTitle="Kapuasia falaris, a new species of nemacheilid loach from Borneo (Teleostei: Nemacheilidae)" masterLastPageNumber="487" masterPageNumber="479" pageNumber="480" updateTime="1737175458869" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="129AC694061B2AD2240F3DB1C6CF35B7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6585F57B88D69258AC0FD96D4A6DA3B0">
<mods:title id="68DD995FD50520C7051E103E9A26CA7E">Kapuasia falaris, a new species of nemacheilid loach from Borneo (Teleostei: Nemacheilidae)</mods:title>
</mods:titleInfo>
<mods:name id="08AF3EE6A68E79205004E390F3D4272C" type="personal">
<mods:role id="4F5ED3A85C1F05B5A885325260ECB354">
<mods:roleTerm id="4B8837BD27227956CE41EEFAA49B57BD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="605E6F5A3742880BBE1C1DEF49639D1F">Kottelat, Maurice</mods:namePart>
</mods:name>
<mods:name id="316C51C062A22B8433E5CBBB06DEDE2D" type="personal">
<mods:role id="B22B3901151B77809712F4AE55BAB263">
<mods:roleTerm id="E0E0644F82177AC888CB29C60D12053A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B8974F12DECE98224AC991351A851CA">, Tan Heok Hui</mods:namePart>
</mods:name>
<mods:name id="4CF9AC02E4CA560408DE01DDF4C1C6F7" type="personal">
<mods:role id="275055911E998D61657C07499D8EACA9">
<mods:roleTerm id="40741BE1A05D4551A6694655BD653EB9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AC40D3C0E508A944C6038E3E5C43568C">Hasan,</mods:namePart>
</mods:name>
<mods:name id="9F82B14478B87CAEFA589CCE4050A884" type="personal">
<mods:role id="3E07BA6AFADC78DE39D1CA20D05F0EC7">
<mods:roleTerm id="F547BB55ECC644FC076F21BF56A756FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6C78F63CCB7FE6EB12128149F6264D08">Veryl</mods:namePart>
</mods:name>
<mods:typeOfResource id="A2104735350FD43B756E76F1BA7748DC">text</mods:typeOfResource>
<mods:relatedItem id="06007A79027CDA6AD691C484A70459EF" type="host">
<mods:titleInfo id="3F3A21C6511FEE0929359DD88FDC46EC">
<mods:title id="D2414BFA6F3890A4A632F3BAD39DFCA9">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0DD0201A80C2E5685ED25ED1C335B381">
<mods:date id="79ED834B7AF8BCF1E3DA8803A16E0691">2024</mods:date>
<mods:detail id="84DB9F84C615D8C3AA853E4BA97A7481" type="pubDate">
<mods:number id="50D056F16E36674234CFC41E790A45D3">2024-12-02</mods:number>
</mods:detail>
<mods:detail id="54636E514D6B414085BC40C68F96A0E4" type="volume">
<mods:number id="F70B7ED10258C270158DAC4651E1A813">72</mods:number>
</mods:detail>
<mods:extent id="722D8E1DD8062E4E4CFFFEC325E2DAC0" unit="page">
<mods:start id="068C81BBFCAD73508A0DE672C8A29618">479</mods:start>
<mods:end id="052280D686A4C676D3D1C992F4AD7ED6">487</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="41ADBE1DCFC50185AE7E63F4CD2ED17D">journal article</mods:classification>
<mods:identifier id="4473115AA0E0E67CF387E20746D812A4" type="DOI">10.26107/RBZ-2024-0035</mods:identifier>
<mods:identifier id="F8153ED5801D6B212D77F581F5215D64" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="9F6BC8BF3664F7B1BA0E020C378B99FB" type="Zenodo-Dep">14682894</mods:identifier>
<mods:identifier id="96C3AD29491F13FFE0992A95BFED3B2F" type="ZooBank">227BB34E-7FF6-492E-A3E3-F82B74C714B6</mods:identifier>
</mods:mods>
<treatment id="038D87D4AD25FFD6E76AFAF6FC02A267" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267" httpUri="http://treatment.plazi.org/id/038D87D4AD25FFD6E76AFAF6FC02A267" lastPageId="7" lastPageNumber="486" pageId="1" pageNumber="480">
<subSubSection id="C33E6549AD25FFD0E76AFAF6FDF8AF11" box="[266,599,1323,1347]" pageId="1" pageNumber="480" type="nomenclature">
<paragraph id="8B9B36C2AD25FFD0E76AFAF6FDF8AF11" blockId="1.[266,599,1323,1379]" box="[266,599,1323,1347]" pageId="1" pageNumber="480">
<heading id="D0D381AEAD25FFD0E76AFAF6FDF8AF11" bold="true" box="[266,599,1323,1347]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="480" reason="2">
<emphasis id="B950EAD0AD25FFD0E76AFAF6FDF8AF11" bold="true" box="[266,599,1323,1347]" pageId="1" pageNumber="480">
<taxonomicName id="4C244D41AD25FFD0E76AFAF6FE6DAF11" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[266,450,1323,1347]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris" status="sp. nov.">
<emphasis id="B950EAD0AD25FFD0E76AFAF6FE6DAF11" bold="true" box="[266,450,1323,1347]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26357ABAD25FFD0E7B1FAF6FDF8AF11" box="[465,599,1323,1347]" pageId="1" pageNumber="480" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD25FFD0E715FA96FE43AF31" box="[373,492,1355,1379]" pageId="1" pageNumber="480" type="description">
<paragraph id="8B9B36C2AD25FFD0E715FA96FE43AF31" blockId="1.[266,599,1323,1379]" box="[373,492,1355,1379]" pageId="1" pageNumber="480">
(
<figureCitation id="131F2A47AD25FFD0E71DFA96FE4CAF31" box="[381,483,1355,1379]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD25FFD0E611FA56FE89AD71" pageId="1" pageNumber="480" type="materials_examined">
<paragraph id="8B9B36C2AD25FFD0E611FA56FDCDAC51" blockId="1.[113,752,1419,1540]" pageId="1" pageNumber="480">
<materialsCitation id="3B4C3C9FAD25FFD0E611FA56FDCDAC51" collectingDate="2024-01-03" collectionCode="MZB" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="MZB 17245" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="holotype">
<emphasis id="B950EAD0AD25FFD0E611FA56FF4AAFF1" bold="true" box="[113,229,1419,1443]" pageId="1" pageNumber="480">
<typeStatus id="549F8860AD25FFD0E611FA56FF4FAFF1" box="[113,224,1419,1443]" pageId="1" pageNumber="480" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DB829EB9AD25FFD0E690FA51FED1AFF1" box="[240,382,1419,1443]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="480" type="Museum">MZB 17245</specimenCode>
, male,
<quantity id="4CDC9B27AD25FFD0E7B6FA56FD93AFF1" box="[470,572,1419,1443]" metricMagnitude="-2" metricUnit="m" metricValue="5.1899999999999995" pageId="1" pageNumber="480" unit="mm" value="51.9">51.9 mm</quantity>
SL;
<collectingCountry id="F3337652AD25FFD0E41BFA56FD43AFF1" box="[635,748,1419,1443]" name="Indonesia" pageId="1" pageNumber="480">Indonesia</collectingCountry>
:
<collectingCounty id="62FA4E4EAD25FFD0E611FA71FF65AF91" box="[113,202,1452,1475]" pageId="1" pageNumber="480">Borneo</collectingCounty>
:
<collectingRegion id="49E0F820AD25FFD0E6BAFA76FE69AF91" box="[218,454,1451,1475]" country="Indonesia" name="Kalimantan Selatan" pageId="1" pageNumber="480">Kalimantan Selatan</collectingRegion>
:
<collectingMunicipality id="6BFFACB8AD25FFD0E7B5FA76FDB2AF91" box="[469,541,1451,1475]" pageId="1" pageNumber="480">Barito</collectingMunicipality>
basin, kabupaten
<location id="8EFB6019AD25FFD0E612FA16FF1CAFB1" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E612FA16FF1CAFB1" box="[114,179,1483,1507]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Tapin" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Tapin</location>
, kecamatan
<location id="8EFB6019AD25FFD0E756FA16FEDEAFB1" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E756FA16FEDEAFB1" box="[310,369,1483,1507]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Piani" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Piani</location>
: desa
<location id="8EFB6019AD25FFD0E7D1FA16FD9DAFB6" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E7D1FA16FD9DAFB6" box="[433,562,1483,1508]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Pipitak Jaya" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Pipitak Jaya</location>
[about
<geoCoordinate id="EE105005AD25FFD0E4E3FA17FD5FAFB1" box="[643,752,1482,1507]" degrees="2" direction="south" minutes="57" orientation="latitude" pageId="1" pageNumber="480" precision="15" seconds="13" value="-2.9536111">2°5713″S</geoCoordinate>
<geoCoordinate id="EE105005AD25FFD0E611FA37FEAFAC51" box="[113,256,1514,1539]" degrees="115" direction="east" minutes="19" orientation="longitude" pageId="1" pageNumber="480" precision="15" seconds="38" value="115.327225">115°1938″E</geoCoordinate>
];
<location id="8EFB6019AD25FFD0E774FA31FE08AC51" LSID="urn:lsid:plazi:treatment:038D87D4AD25FFD6E76AFAF6FC02A267:8EFB6019AD25FFD0E774FA31FE08AC51" box="[276,423,1515,1539]" country="Indonesia" county="Borneo" latitude="-2.9536111" longLatPrecision="21" longitude="115.327225" municipality="Barito" name="Noor Aminin" pageId="1" pageNumber="480" stateProvince="Kalimantan Selatan">Noor Aminin</location>
,
<date id="FF9A1002AD25FFD0E7D4FA36FDF1AC51" box="[436,606,1515,1540]" pageId="1" pageNumber="480" value="2024-01-03">
<collectingDate id="EFDEE9EAAD25FFD0E7D4FA36FDF1AC51" box="[436,606,1515,1540]" pageId="1" pageNumber="480" value="2024-01-03">3 January 2024</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9B36C2AD25FFD0E611F9F6FF53ACF1" blockId="1.[113,751,1579,1699]" pageId="1" pageNumber="480">
<materialsCitation id="3B4C3C9FAD25FFD0E611F9F6FDECAC31" collectingDate="2024-01-03" collectionCode="ZRC" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="ZRC 65844, 8" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="paratype">
<emphasis id="B950EAD0AD25FFD0E611F9F6FF45AC11" bold="true" box="[113,234,1579,1603]" pageId="1" pageNumber="480">
<typeStatus id="549F8860AD25FFD0E611F9F6FF4AAC11" box="[113,229,1579,1603]" pageId="1" pageNumber="480" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCode id="DB829EB9AD25FFD0E691F9F6FEDFAC11" box="[241,368,1579,1603]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65844</specimenCode>
,
<specimenCode id="DB829EB9AD25FFD0E71BF9F6FE25AC11" box="[379,394,1579,1603]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">8</specimenCode>
, 43.066.0 mm SL; CMK 29184, 5,
<quantity id="4CDC9B27AD25FFD0E6EBF996FE8AAC31" box="[139,293,1611,1635]" metricMagnitude="-2" metricUnit="m" metricValue="4.99" metricValueMax="5.66" metricValueMin="4.32" pageId="1" pageNumber="480" unit="mm" value="49.9" valueMax="56.6" valueMin="43.2">43.256.6 mm</quantity>
SL; same data as holotype
</materialsCitation>
. —
<materialsCitation id="3B4C3C9FAD25FFD0E40CF996FF56ACF1" collectingDate="2024-03-04" collectionCode="ZRC" country="Indonesia" county="Borneo" latitude="-2.9536111" location="Tapin" longLatPrecision="21" longitude="115.327225" municipality="Barito" pageId="1" pageNumber="480" specimenCode="ZRC 65998, 12" specimenCount="1" stateProvince="Kalimantan Selatan" typeStatus="paratype">
<specimenCode id="DB829EB9AD25FFD0E40CF996FD44AC31" box="[620,747,1611,1635]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65998</specimenCode>
,
<specimenCode id="DB829EB9AD25FFD0E611F9B6FF21ACD1" box="[113,142,1643,1667]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">12</specimenCode>
, 37.0
<quantity id="4CDC9B27AD25FFD0E6B9F9B6FE93ACD1" box="[217,316,1643,1667]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="1" pageNumber="480" unit="mm" value="59.2">59.2 mm</quantity>
SL; same location data as
<typeStatus id="549F8860AD25FFD0E411F9B6FD7CACD1" box="[625,723,1643,1667]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
;
<date id="FF9A1002AD25FFD0E481F9B6FF56ACF1" pageId="1" pageNumber="480" value="2024-03-04">
<collectingDate id="EFDEE9EAAD25FFD0E481F9B6FF56ACF1" pageId="1" pageNumber="480" value="2024-03-04">4 March 2024</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9B36C2AD25FFD0E611F916FE89AD71" blockId="1.[112,751,1739,1827]" pageId="1" pageNumber="480">
<materialsCitation id="3B4C3C9FAD25FFD0E611F916FE8EAD71" collectingDate="2024-02-14" collectionCode="ZRC" pageId="1" pageNumber="480" specimenCode="ZRC 65915, 46" specimenCount="2">
<emphasis id="B950EAD0AD25FFD0E611F916FE93ACB1" bold="true" box="[113,316,1739,1763]" pageId="1" pageNumber="480">
<typeStatus id="549F8860AD25FFD0E611F916FE93ACB1" box="[113,316,1739,1763]" isOtherTypeStatus="true" pageId="1" pageNumber="480">Non-type material</typeStatus>
</emphasis>
.
<specimenCode id="DB829EB9AD25FFD0E728F916FE6BACB1" box="[328,452,1739,1763]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">ZRC 65915</specimenCode>
,
<specimenCode id="DB829EB9AD25FFD0E7ADF916FE45ACB1" box="[461,490,1739,1763]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="480">46</specimenCode>
,
<quantity id="4CDC9B27AD25FFD0E793F916FD26ACB1" box="[499,649,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="5.005" metricValueMax="5.74" metricValueMin="4.27" pageId="1" pageNumber="480" unit="mm" value="50.05" valueMax="57.4" valueMin="42.7">42.757.4 mm</quantity>
SL; same location data as
<typeStatus id="549F8860AD25FFD0E740F936FE2EAD51" box="[288,385,1771,1795]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
;
<date id="FF9A1002AD25FFD0E7ECF936FDE4AD51" box="[396,587,1771,1795]" pageId="1" pageNumber="480" value="2024-02-14">
<collectingDate id="EFDEE9EAAD25FFD0E7ECF936FDE4AD51" box="[396,587,1771,1795]" pageId="1" pageNumber="480" value="2024-02-14">14 February 2024</collectingDate>
</date>
[specimens all dead on arrival]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD25FFD0E611F896FAB4AD71" pageId="1" pageNumber="480" type="diagnosis">
<paragraph id="8B9B36C2AD25FFD0E611F896FAB4AD71" blockId="1.[113,753,1867,2083]" lastBlockId="1.[811,1453,1259,1828]" pageId="1" pageNumber="480">
<emphasis id="B950EAD0AD25FFD0E611F896FF40AD31" bold="true" box="[113,239,1867,1891]" pageId="1" pageNumber="480">Diagnosis.</emphasis>
<taxonomicName id="4C244D41AD25FFD0E69BF891FE6EAD31" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[251,449,1867,1891]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD25FFD0E69BF891FE6EAD31" box="[251,449,1867,1891]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C244D41AD25FFD0E4B6F891FF42ADD1" authorityName="Roberts" authorityYear="1989" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD25FFD0E4B6F891FF42ADD1" italics="true" pageId="1" pageNumber="480">K. maculiceps</emphasis>
</taxonomicName>
by the following combination of characters: colour pattern made of 37 regular black bars, wider than interspaces and extending across back, of constant width and reaching ventral midline (vs. irregular bars or mostly triangular saddles reaching downward about to lateral line, often alternating or connected with blotches in lower half of body); head black, except for white dorsal side of the snout, a bar behind eye and a bar across the nape and along hind margin of opercle (vs. top of head and cheek spotted); in males, presence of a suborbital flap (
<figureCitation id="131F2A47AD25FFD0E2A9FA96FAA0AF36" box="[1225,1295,1355,1380]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="1" pageNumber="480">Fig. 4</figureCitation>
; vs. absence); in males, vertical skin flanges on the dorsal surface of the pectoral fin, between the branched rays (
<figureCitation id="131F2A47AD25FFD0E2B7FA56FAB8AFF1" box="[1239,1303,1419,1443]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="1" pageNumber="480">Fig. 5</figureCitation>
; vs. absence); no large flat pectoral axillary lobe (presence), but instead a cylindrical lobe, with pointed posterior extremity, usually separate from skin of flank by a deep groove, often with a second similar lobe above the groove (
<figureCitation id="131F2A47AD25FFD0E2ABF9D6FAA2AC71" box="[1227,1293,1547,1571]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="1" pageNumber="480">Fig. 6</figureCitation>
). The shape of this lobe is apparently unique among Southeast Asian species of nemacheilids.
<taxonomicName id="4C244D41AD25FFD0E58CF991FB0CAC31" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[1004,1187,1611,1635]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="480" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD25FFD0E58CF991FB0CAC31" box="[1004,1187,1611,1635]" italics="true" pageId="1" pageNumber="480">Kapuasia falaris</emphasis>
</taxonomicName>
is further distinguished from all nemacheilids from Southeast Asia by the shape of the suborbital flap: globulous, at some distance below the eye, and with a deep vertical groove from the flap to the eye; and the strongly upwards curled pectoral fin, appearing with an almost vertical orientation in life (
<figureCitation id="131F2A47AD25FFD0E2A6F936FAA7AD56" box="[1222,1288,1771,1796]" captionStart="Fig" captionStartId="4.[135,170,1513,1535]" captionTargetBox="[135,1473,189,1497]" captionTargetId="figure-7@4.[135,1473,189,1497]" captionTargetPageId="4" captionText="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Fig. 3</figureCitation>
), is apparently unique among Southeast Asian nemacheilids.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD25FFD6E54DF896FD5CAC64" lastPageId="7" lastPageNumber="486" pageId="1" pageNumber="480" type="description">
<paragraph id="8B9B36C2AD25FFD3E54DF896FEE0ACBE" blockId="1.[813,1453,1867,2083]" lastBlockId="2.[135,776,1524,1772]" lastPageId="2" lastPageNumber="481" pageId="1" pageNumber="480">
<emphasis id="B950EAD0AD25FFD0E54DF896FC18AD31" bold="true" box="[813,951,1867,1891]" pageId="1" pageNumber="480">Description.</emphasis>
See
<figureCitation id="131F2A47AD25FFD0E588F896FBE6AD36" box="[1000,1097,1867,1892]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="1" pageNumber="480">Figs. 13</figureCitation>
for general appearance and
<tableCitation id="C6A60379AD25FFD0E30FF896FC95ADD6" captionStart="Table 1" captionStartId="2.[135,190,189,211]" captionText="Table 1. Morphometric data of type material of Kapuasia falaris (n=10). Ranges and means include holotype data." pageId="1" pageNumber="480">Table 1</tableCitation>
for morphometric data of
<typeStatus id="549F8860AD25FFD0E205F8B6FB6DADD1" box="[1125,1218,1899,1923]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D22FD4BAD25FFD0E29DF8B1FAD1ADD6" box="[1277,1406,1900,1924]" count="9" pageId="1" pageNumber="480" type="generic" typeStatus="paratype">9 paratypes</specimenCount>
. An elongate nemacheilid with body depth gradually increasing up to dorsal-fin origin. Behind dorsal fin, body depth almost uniform until caudal-fin base. Dorsal profile continuous between head and body. Head slightly depressed; body slightly compressed anteriorly, gradually more compressed posteriorly. Interorbital area flat. In lateral view, eye protruding over dorsal profile of head. Cheeks not swollen. Snout pointed but not acute. Depth of caudal peduncle 1.31.5 times in its length, depth uniform. No dorsal crest or keel on post-dorsal area. No ventral crest or keel on caudal peduncle. Vertebrae 1820+1314=3234 (modally 19+14=33). Largest recorded size
<quantity id="4CDC9B27AD26FFD3E794F969FDFDAC9E" box="[500,594,1716,1740]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" pageId="2" pageNumber="481" unit="mm" value="57.5">57.5 mm</quantity>
SL in male, 66.0 mm SL in female.
</paragraph>
<caption id="DF5B664AAD26FFD3E6E7FF60FB5DAA81" box="[135,1266,189,211]" pageId="2" pageNumber="481" startId="2.[135,190,189,211]" targetType="table">
<paragraph id="8B9B36C2AD26FFD3E6E7FF60FB5DAA81" blockId="2.[135,1266,189,211]" box="[135,1266,189,211]" pageId="2" pageNumber="481">
Table 1. Morphometric data of type material of
<taxonomicName id="4C244D41AD26FFD3E401FF63FCABAA80" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[609,772,189,211]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="481" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD26FFD3E401FF63FCABAA80" box="[609,772,189,211]" italics="true" pageId="2" pageNumber="481">Kapuasia falaris</emphasis>
</taxonomicName>
(n=10). Ranges and means include holotype data.
</paragraph>
</caption>
<paragraph id="8B9B36C2AD26FFD3E564FED8FAE4AFF4" pageId="2" pageNumber="481">
<table id="F924C462AD26002EE6F6FED8FAF3AFF4" box="[150,1372,261,1446]" gridcols="4" gridrows="13" pageId="2" pageNumber="481">
<tr id="35143480AD26002EE6F6FED8FAF3AB48" box="[150,1372,261,282]" gridrow="0" pageId="2" pageNumber="481" rowspan-0="1">
<th id="76C55DFCAD26002EE564FED8FCCEAB48" box="[772,865,261,282]" gridcol="1" gridrow="0" pageId="2" pageNumber="481">
<emphasis id="B950EAD0AD26FFD3E564FED8FCCEAB48" bold="true" box="[772,865,261,282]" pageId="2" pageNumber="481">Holotype</emphasis>
</th>
<th id="76C55DFCAD26002EE59DFED8FBDAAB48" box="[1021,1141,261,282]" gridcol="2" gridrow="0" pageId="2" pageNumber="481">
<emphasis id="B950EAD0AD26FFD3E278FED8FBF6AB48" bold="true" box="[1048,1113,261,282]" pageId="2" pageNumber="481">Range</emphasis>
</th>
<th id="76C55DFCAD26002EE341FED8FAF3AB48" box="[1313,1372,261,282]" gridcol="3" gridrow="0" pageId="2" pageNumber="481">
<emphasis id="B950EAD0AD26FFD3E341FED8FAF3AB48" bold="true" box="[1313,1372,261,282]" pageId="2" pageNumber="481">Mean</emphasis>
</th>
</tr>
<tr id="35143480AD26002EE6F6FEE6FAF3AB03" box="[150,1372,315,337]" gridrow="1" pageId="2" pageNumber="481" rowspan-3="1">
<th id="76C55DFCAD26002EE6F6FEE6FE43AB03" box="[150,492,315,337]" gridcol="0" gridrow="1" pageId="2" pageNumber="481">Standard length (mm)</th>
<td id="76C55DFCAD26002EE564FEE6FCCEAB03" box="[772,865,315,337]" gridcol="1" gridrow="1" pageId="2" pageNumber="481">51.9</td>
<td id="76C55DFCAD26002EE59DFEE6FBDAAB03" box="[1021,1141,315,337]" gridcol="2" gridrow="1" pageId="2" pageNumber="481">48.666.0</td>
</tr>
<tr id="35143480AD26002EE6F6FEB7FAF3ABD2" box="[150,1372,362,384]" gridrow="2" pageId="2" pageNumber="481" rowspan-3="1">
<th id="76C55DFCAD26002EE6F6FEB7FE43ABD2" box="[150,492,362,384]" gridcol="0" gridrow="2" pageId="2" pageNumber="481">Total length (mm)</th>
<td id="76C55DFCAD26002EE564FEB7FCCEABD2" box="[772,865,362,384]" gridcol="1" gridrow="2" pageId="2" pageNumber="481">64.4</td>
<td id="76C55DFCAD26002EE59DFEB7FBDAABD2" box="[1021,1141,362,384]" gridcol="2" gridrow="2" pageId="2" pageNumber="481">59.783.4</td>
</tr>
<tr id="35143480AD26002EE6F6FE44FAF3A8DF" box="[150,1372,409,653]" gridrow="3" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FE44FE43A8DF" box="[150,492,409,653]" gridcol="0" gridrow="3" pageId="2" pageNumber="481">In percent of standard length Total length Head length (dorsal) Head length (lateral) Predorsal length Prepelvic length Pre-anus length Pre-anal length</th>
<td id="76C55DFCAD26002EE564FE44FCCEA8DF" box="[772,865,409,653]" gridcol="1" gridrow="3" pageId="2" pageNumber="481">124.0 21.9 24.5 53.1 48.4 67.4 74.7</td>
<td id="76C55DFCAD26002EE59DFE44FBDAA8DF" box="[1021,1141,409,653]" gridcol="2" gridrow="3" pageId="2" pageNumber="481">120.5126.5 18.521.9 22.425.4 51.854.8 48.051.5 67.473.2 73.178.1</td>
<td id="76C55DFCAD26002EE341FE44FAF3A8DF" box="[1313,1372,409,653]" gridcol="3" gridrow="3" pageId="2" pageNumber="481">123.0 20.3 23.9 53.3 50.0 69.2 75.4</td>
</tr>
<tr id="35143480AD26002EE6F6FD4AFAF3A8FF" box="[150,1372,663,685]" gridrow="4" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FD4AFE43A8FF" box="[150,492,663,685]" gridcol="0" gridrow="4" pageId="2" pageNumber="481">Head depth</th>
<td id="76C55DFCAD26002EE564FD4AFCCEA8FF" box="[772,865,663,685]" gridcol="1" gridrow="4" pageId="2" pageNumber="481">10.7</td>
<td id="76C55DFCAD26002EE59DFD4AFBDAA8FF" box="[1021,1141,663,685]" gridcol="2" gridrow="4" pageId="2" pageNumber="481">10.011.4</td>
<td id="76C55DFCAD26002EE341FD4AFAF3A8FF" box="[1313,1372,663,685]" gridcol="3" gridrow="4" pageId="2" pageNumber="481">10.7</td>
</tr>
<tr id="35143480AD26002EE6F6FD6AFAF3A97E" box="[150,1372,695,812]" gridrow="5" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FD6AFE43A97E" box="[150,492,695,812]" gridcol="0" gridrow="5" pageId="2" pageNumber="481">Body depth at dorsal-fin origin Depth of caudal peduncle Length of caudal peduncle Head width</th>
<td id="76C55DFCAD26002EE564FD6AFCCEA97E" box="[772,865,695,812]" gridcol="1" gridrow="5" pageId="2" pageNumber="481">13.8 12.0 17.5 14.3</td>
<td id="76C55DFCAD26002EE59DFD6AFBDAA97E" box="[1021,1141,695,812]" gridcol="2" gridrow="5" pageId="2" pageNumber="481">12.615.5 10.612.0 15.317.5 13.515.4</td>
<td id="76C55DFCAD26002EE341FD6AFAF3A97E" box="[1313,1372,695,812]" gridcol="3" gridrow="5" pageId="2" pageNumber="481">13.6 11.3 16.2 14.3</td>
</tr>
<tr id="35143480AD26002EE6F6FCEBFAF3A91E" box="[150,1372,822,844]" gridrow="6" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FCEBFE43A91E" box="[150,492,822,844]" gridcol="0" gridrow="6" pageId="2" pageNumber="481">Body width at dorsal-fin origin</th>
<td id="76C55DFCAD26002EE564FCEBFCCEA91E" box="[772,865,822,844]" gridcol="1" gridrow="6" pageId="2" pageNumber="481">9.8</td>
<td id="76C55DFCAD26002EE59DFCEBFBDAA91E" box="[1021,1141,822,844]" gridcol="2" gridrow="6" pageId="2" pageNumber="481">8.811.9</td>
<td id="76C55DFCAD26002EE341FCEBFAF3A91E" box="[1313,1372,822,844]" gridcol="3" gridrow="6" pageId="2" pageNumber="481">10.2</td>
</tr>
<tr id="35143480AD26002EE6F6FC8BFAF3AE78" box="[150,1372,854,1066]" gridrow="7" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FC8BFE43AE78" box="[150,492,854,1066]" gridcol="0" gridrow="7" pageId="2" pageNumber="481">Snout length Eye diameter Interorbital width Length of dorsal fin Length of upper caudal-fin lobe Length of median caudal-fin rays Length of lower caudal-fin lobe</th>
<td id="76C55DFCAD26002EE564FC8BFCCEAE78" box="[772,865,854,1066]" gridcol="1" gridrow="7" pageId="2" pageNumber="481">8.4 2.9 5.8 16.5 23.4 17.3 23.4</td>
<td id="76C55DFCAD26002EE59DFC8BFBDAAE78" box="[1021,1141,854,1066]" gridcol="2" gridrow="7" pageId="2" pageNumber="481">7.79.8 2.53.3 5.26.2 16.019.9 19.523.6 12.617.9 20.325.4</td>
<td id="76C55DFCAD26002EE341FC8BFAF3AE78" box="[1313,1372,854,1066]" gridcol="3" gridrow="7" pageId="2" pageNumber="481">9.1 3.0 5.9 17.5 22.1 14.9 23.0</td>
</tr>
<tr id="35143480AD26002EE6F6FBE9FAF3AEDB" box="[150,1372,1076,1161]" gridrow="8" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FBE9FE43AEDB" box="[150,492,1076,1161]" gridcol="0" gridrow="8" pageId="2" pageNumber="481">Length of anal fin Length of pelvic fin Length of pectoral fin</th>
<td id="76C55DFCAD26002EE564FBE9FCCEAEDB" box="[772,865,1076,1161]" gridcol="1" gridrow="8" pageId="2" pageNumber="481">16.4 15.9 19.1</td>
<td id="76C55DFCAD26002EE59DFBE9FBDAAEDB" box="[1021,1141,1076,1161]" gridcol="2" gridrow="8" pageId="2" pageNumber="481">16.218.7 15.718.0 16.419.1</td>
<td id="76C55DFCAD26002EE341FBE9FAF3AEDB" box="[1313,1372,1076,1161]" gridcol="3" gridrow="8" pageId="2" pageNumber="481">17.3 16.4 18.3</td>
</tr>
<tr id="35143480AD26002EE6F6FB7FFAF3AF4A" box="[150,1372,1186,1304]" gridrow="9" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FB7FFE43AF4A" box="[150,492,1186,1304]" gridcol="0" gridrow="9" pageId="2" pageNumber="481">In percent of dorsal head length Snout length Eye diameter Interorbital width</th>
<td id="76C55DFCAD26002EE564FB7FFCCEAF4A" box="[772,865,1186,1304]" gridcol="1" gridrow="9" pageId="2" pageNumber="481">38 13 27</td>
<td id="76C55DFCAD26002EE59DFB7FFBDAAF4A" box="[1021,1141,1186,1304]" gridcol="2" gridrow="9" pageId="2" pageNumber="481">3850 1316 2733</td>
<td id="76C55DFCAD26002EE341FB7FFAF3AF4A" box="[1313,1372,1186,1304]" gridcol="3" gridrow="9" pageId="2" pageNumber="481">45 15 29</td>
</tr>
<tr id="35143480AD26002EE6F6FAEDFAF3AF34" box="[150,1372,1328,1382]" gridrow="10" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FAEDFE43AF34" box="[150,492,1328,1382]" gridcol="0" gridrow="10" pageId="2" pageNumber="481">In percent of lateral head length Snout length</th>
<td id="76C55DFCAD26002EE564FAEDFCCEAF34" box="[772,865,1328,1382]" gridcol="1" gridrow="10" pageId="2" pageNumber="481">34</td>
<td id="76C55DFCAD26002EE59DFAEDFBDAAF34" box="[1021,1141,1328,1382]" gridcol="2" gridrow="10" pageId="2" pageNumber="481">3442</td>
<td id="76C55DFCAD26002EE341FAEDFAF3AF34" box="[1313,1372,1328,1382]" gridcol="3" gridrow="10" pageId="2" pageNumber="481">38</td>
</tr>
<tr id="35143480AD26002EE6F6FAADFAF3AFD4" box="[150,1372,1392,1414]" gridrow="11" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FAADFE43AFD4" box="[150,492,1392,1414]" gridcol="0" gridrow="11" pageId="2" pageNumber="481">Eye diameter</th>
<td id="76C55DFCAD26002EE564FAADFCCEAFD4" box="[772,865,1392,1414]" gridcol="1" gridrow="11" pageId="2" pageNumber="481">12</td>
<td id="76C55DFCAD26002EE59DFAADFBDAAFD4" box="[1021,1141,1392,1414]" gridcol="2" gridrow="11" pageId="2" pageNumber="481">1114</td>
<td id="76C55DFCAD26002EE341FAADFAF3AFD4" box="[1313,1372,1392,1414]" gridcol="3" gridrow="11" pageId="2" pageNumber="481">13</td>
</tr>
<tr id="35143480AD26002EE6F6FA4DFAF3AFF4" box="[150,1372,1424,1446]" gridrow="12" pageId="2" pageNumber="481">
<th id="76C55DFCAD26002EE6F6FA4DFE43AFF4" box="[150,492,1424,1446]" gridcol="0" gridrow="12" pageId="2" pageNumber="481">Interorbital width</th>
<td id="76C55DFCAD26002EE564FA4DFCCEAFF4" box="[772,865,1424,1446]" gridcol="1" gridrow="12" pageId="2" pageNumber="481">24</td>
<td id="76C55DFCAD26002EE59DFA4DFBDAAFF4" box="[1021,1141,1424,1446]" gridcol="2" gridrow="12" pageId="2" pageNumber="481">2226</td>
<td id="76C55DFCAD26002EE341FA4DFAF3AFF4" box="[1313,1372,1424,1446]" gridcol="3" gridrow="12" pageId="2" pageNumber="481">25</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9B36C2AD26FFD3E6E8F8C9FB35AD9E" blockId="2.[134,775,1811,2092]" lastBlockId="2.[835,1476,1524,1996]" pageId="2" pageNumber="481">
Dorsal fin with 4 unbranched and 7½ (1), 8½ (12) or 9½ (1*) branched rays; distal margin slightly concave. Usually, second branched ray longest. Pectoral fin curled and slanted, with 1 unbranched and 9 branched rays (including small last ray, usually unbranched), rounded, reaching ⅓ to ⅜ of distance to pelvic-fin base; rays without filamentous extensions. Axillary pectoral lobe present, cylindrical, posteriorly pointed, usually separated from skin of flank by a deep groove; groove often margined above by a second pointed lobe (
<figureCitation id="131F2A47AD26FFD3E43AF7C9FD33A27E" box="[602,668,2068,2092]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="2" pageNumber="481">Fig. 6</figureCitation>
), more or less regularly shaped, more or less distinct. Pelvic fin with 1 unbranched and 6 (2), 7 (11*) or 8 (1) branched rays (including small last ray, usually unbranched); reaching almost to anus (⅔ of distance to anal-fin origin; triangular to rounded; posterior margin straight; origin below base of unbranched dorsal-fin rays 13; axillary lobe present, entirely free, longer than fin base, high, rounded. Anus situated 1.52.5 eye diameter in front of anal-fin origin, distance between anus and anal-fin origin about ½ of distance between anus and pelvic-fin base. Anal fin with 3 unbranched and 5½ branched rays; distal margin slightly concave, first branched ray longest. Caudal fin with 9+8 (11*) or 8+7 (1) branched rays; forked (upper lobe 1.31.6 times longer than median rays), lobes rounded, subequal, lower lobe slightly longer than upper one (1.01.1 times).
</paragraph>
<paragraph id="8B9B36C2AD26FFD5E523F829FE8EACF9" blockId="2.[835,1474,2036,2093]" lastBlockId="4.[136,775,1619,1707]" lastPageId="4" lastPageNumber="483" pageId="2" pageNumber="481">
Especially visible in life, inner pectoral-fin rays curled upwards and slightly forwards, against body, so that fin may appear vertically orientated (
<figureCitation id="131F2A47AD20FFD5E7DAF98EFE55AC39" box="[442,506,1619,1643]" captionStart="Fig" captionStartId="4.[135,170,1513,1535]" captionTargetBox="[135,1473,189,1497]" captionTargetId="figure-7@4.[135,1473,189,1497]" captionTargetPageId="4" captionText="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="4" pageNumber="483">Fig. 3</figureCitation>
). In pelvic fin, innermost rays also curled upwards. No difference between sexes in this character.
</paragraph>
<caption id="DF5B664AAD27FFD2E611F82EFD9BA210" pageId="3" pageNumber="482" startId="3.[113,148,2035,2057]" targetBox="[175,1390,189,2020]" targetPageId="3" targetType="figure">
<paragraph id="8B9B36C2AD27FFD2E611F82EFD9BA210" blockId="3.[113,1451,2035,2115]" pageId="3" pageNumber="482">
Fig. 2.
<taxonomicName id="4C244D41AD27FFD2E6DCF829FECEA25A" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[188,353,2035,2057]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="482" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD27FFD2E6DCF829FECEA25A" box="[188,353,2035,2057]" italics="true" pageId="3" pageNumber="482">Kapuasia falaris</emphasis>
</taxonomicName>
, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL.
</paragraph>
</caption>
<caption id="DF5B664AAD20FFD5E6E7FA34FE75AC49" pageId="4" pageNumber="483" startId="4.[135,170,1513,1535]" targetBox="[135,1473,189,1497]" targetPageId="4" targetType="figure">
<paragraph id="8B9B36C2AD20FFD5E6E7FA34FE75AC49" blockId="4.[135,1474,1513,1563]" pageId="4" pageNumber="483">
Fig. 3.
<taxonomicName id="4C244D41AD20FFD5E6B0FA34FEDBAFAC" authorityName="Kottelat &amp; &amp; Hasan &amp; Veryl" authorityYear="2024" box="[208,372,1513,1534]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="483" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD20FFD5E6B0FA34FEDBAFAC" box="[208,372,1513,1534]" italics="true" pageId="4" pageNumber="483">Kapuasia falaris</emphasis>
</taxonomicName>
, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed).
</paragraph>
</caption>
<paragraph id="8B9B36C2AD20FFD5E6EAF90EFEC3ADF8" blockId="4.[136,777,1747,1963]" pageId="4" pageNumber="483">Body entirely covered by scales, except in a few specimens on belly, from throat to anus. Scales embedded, not externally visible on anterior part of body. Lateral line complete, with approximately 8693 pores (difficult to count with accuracy). Cephalic lateral line system with 6 supraorbital, 4 + 1011 infraorbital, 910 preoperculo-mandibular and 3 supratemporal pores.</paragraph>
<paragraph id="8B9B36C2AD20FFD5E6E8F80EFAF3ADB9" blockId="4.[136,776,2003,2059]" lastBlockId="4.[834,1476,1619,2027]" pageId="4" pageNumber="483">
Anterior nare pierced in front side of a pointed flap-like tube. Posterior nare adjacent to anterior one. Mouth U-shaped, gape about two times wider than long (
<figureCitation id="131F2A47AD20FFD5E29CF98EFAEEAC39" box="[1276,1345,1619,1643]" captionStart="Fig" captionStartId="6.[836,871,518,540]" captionTargetBox="[836,1474,189,503]" captionTargetId="figure-56@6.[836,1474,185,505]" captionTargetPageId="6" captionText="Fig. 7. Kapuasia falaris, CMK 29184, 51.4 mm SL; mouth." pageId="4" pageNumber="483">Fig. 7</figureCitation>
). Lips thin. Upper lip usually with median notch, with a few shallow, thin wrinkles on posterior edge. Processus dentiformis present. Lower lip almost continuous, with a median notch but not forming two median lobes; median part with small irregular ridges, radiating from anterior extremity of postlabial groove, lateral parts smooth. Anterior part of lower lip and space between anterior extremities of postlabial grooves with irregular ridges, continuous with the patch on the lip itself. Tip of lower jaw not exposed when mouth is closed. No median concavity in lower jaw. Inner rostral barbel not reaching corner of mouth; outer one reaching corner of mouth. Maxillary barbel reaching vertical of eye.
</paragraph>
<caption id="DF5B664AAD21FFD4E54DFDBFFC19A8E3" pageId="5" pageNumber="484" startId="5.[813,848,610,632]" targetBox="[814,1452,189,595]" targetPageId="5" targetType="figure">
<paragraph id="8B9B36C2AD21FFD4E54DFDBFFC19A8E3" blockId="5.[813,1452,610,689]" pageId="5" pageNumber="484">
Fig. 5.
<taxonomicName id="4C244D41AD21FFD4E512FDBFFBFFA825" authority=", CMK" authorityName="CMK" box="[882,1104,610,631]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="484" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD21FFD4E512FDBFFBBBA825" box="[882,1044,610,631]" italics="true" pageId="5" pageNumber="484">Kapuasia falaris</emphasis>
,CMK
</taxonomicName>
29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows).
</paragraph>
</caption>
<paragraph id="8B9B36C2AD21FFD4E54DFD35FB7BAE52" blockId="5.[811,1452,744,1024]" pageId="5" pageNumber="484">Branched rays 4 and following: both branches are branched, with narrow membranes between branches. Some individual variation exists in the thickness and branching pattern. In females, pattern roughly similar, but membranes are present between all branches. Branched ray 1: posterior branch may be slightly branched near tip. Branched rays 2 and 3: anterior branch not branched, posterior branch branched near tip. Branched ray 4 and following: both branches are branched, with membranes between all branches.</paragraph>
<caption id="DF5B664AAD21FFD4E611FB50FE31AE8E" pageId="5" pageNumber="484" startId="5.[113,148,1165,1187]" targetBox="[113,751,189,1150]" targetPageId="5" targetType="figure">
<paragraph id="8B9B36C2AD21FFD4E611FB50FE31AE8E" blockId="5.[113,753,1165,1245]" pageId="5" pageNumber="484">
Fig. 4.
<taxonomicName id="4C244D41AD21FFD4E6D8FB53FE3AAEF0" authority=", ZRC" authorityName="ZRC" box="[184,405,1165,1187]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="484" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD21FFD4E6D8FB53FEF6AEF0" box="[184,345,1165,1187]" italics="true" pageId="5" pageNumber="484">Kapuasia falaris</emphasis>
, ZRC
</taxonomicName>
65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow).
</paragraph>
</caption>
<paragraph id="8B9B36C2AD21FFD4E611FAC9FE19AF3E" blockId="5.[113,752,1300,1388]" pageId="5" pageNumber="484">
Stomach elongated, intestine with a bend some distance behind stomach (
<figureCitation id="131F2A47AD21FFD4E759FAE9FE2DAF1E" box="[313,386,1332,1356]" captionStart="Fig" captionStartId="6.[836,872,1329,1351]" captionTargetBox="[1030,1280,581,1315]" captionTargetId="figure-58@6.[989,1325,577,1324]" captionTargetPageId="6" captionText="Fig. 8. Kapuasia falaris, ZRC 65915, 55.0 mm SL; schematic. Scale bar: 1 mm." pageId="5" pageNumber="484">Fig. 8</figureCitation>
). Air bladder without posterior chamber in abdominal cavity.
</paragraph>
<paragraph id="8B9B36C2AD21FFD4E613FA49FD26AC1E" blockId="5.[112,753,1428,1612]" pageId="5" pageNumber="484">
<emphasis id="B950EAD0AD21FFD4E613FA49FEF5AFFE" bold="true" box="[115,346,1428,1452]" pageId="5" pageNumber="484">Sexual dimorphism.</emphasis>
Males with suborbital flap (modified lateral ethmoid;
<figureCitation id="131F2A47AD21FFD4E77EFA69FEC6AF9E" box="[286,361,1460,1484]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="5" pageNumber="484">Fig. 4a</figureCitation>
), in lower position than in most other nemacheilids, globulous, movable, upper outline continued upwards, almost vertically, by a groove reaching orbit. Flap missing in females, but a shallow crease corresponds to position of lower edge of flap in males (
<figureCitation id="131F2A47AD21FFD4E44BF9E9FDD2AC1E" box="[555,637,1588,1612]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="5" pageNumber="484">Fig. 4b</figureCitation>
).
</paragraph>
<paragraph id="8B9B36C2AD21FFD4E611F9A9FE13AD7E" blockId="5.[112,753,1652,1836]" pageId="5" pageNumber="484">
No tubercles on body and fins, in both sexes. In males, on the dorsal surface of pectoral fin, a conspicuous vertical skin flange, translucent, between branched rays 18, most developed between branched rays 16 (
<figureCitation id="131F2A47AD21FFD4E46EF909FDE2ACBE" box="[526,589,1748,1772]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="5" pageNumber="484">Fig. 5</figureCitation>
). These flanges not present in females. Females deeper bodied and apparently reach a larger size than males.
</paragraph>
<paragraph id="8B9B36C2AD21FFD4E611F889FA02AF72" blockId="5.[112,753,1876,2092]" lastBlockId="5.[811,1453,1064,1313]" pageId="5" pageNumber="484">
Unbranched and 1 or 2 anterior branched pectoral-fin rays thicker than others (
<figureCitation id="131F2A47AD21FFD4E733F8A9FE0DADDE" box="[339,418,1908,1932]" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="5" pageNumber="484">Figs. 5</figureCitation>
,
<figureCitation id="131F2A47AD21FFD4E7D2F8A9FE6DADDE" box="[434,450,1908,1932]" captionStart="Fig" captionStartId="7.[114,149,1140,1162]" captionTargetBox="[114,752,189,1112]" captionTargetId="figure-14@7.[114,752,187,1127]" captionTargetPageId="7" captionText="Fig. 9. Kapuasia falaris, pectoral fin in dorsal view; ZRC 65844. a, female, 58.9 mm SL; b, male 52.3 mm SL (reversed)." pageId="5" pageNumber="484">9</figureCitation>
). In males, branched ray 1 branched only once, anterior branch usually thicker than posterior one, without membrane between branches. Branched ray 2: anterior branch thicker than posterior one, unbranched; posterior branch maybe branched at tip; no membranes. Branched rays 3: as ray 2 but branches may be branched.
<emphasis id="B950EAD0AD21FFD4E54DFBF5FC13AE12" bold="true" box="[813,956,1064,1088]" pageId="5" pageNumber="484">Colouration</emphasis>
(
<figureCitation id="131F2A47AD21FFD4E5AEFBF5FB94AE13" box="[974,1083,1064,1089]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[115,150,1182,1204]" captionStartId-1="3.[113,148,2035,2057]" captionStartId-2="4.[135,170,1513,1535]" captionTargetBox-0="[113,1451,189,1167]" captionTargetBox-1="[175,1390,189,2020]" captionTargetBox-2="[135,1473,189,1497]" captionTargetId-0="figure-398@1.[113,1451,189,1167]" captionTargetId-1="figure-14@3.[175,1390,189,2020]" captionTargetId-2="figure-7@4.[135,1473,189,1497]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Kapuasia falaris, holotype, MZB 17245, 51.9 mm SL; Borneo: Barito drainage." captionText-1="Fig. 2. Kapuasia falaris, paratypes, Borneo: Barito drainage. a, ZRC 65844, male, 43.0 mm SL; b, ZRC 65844, male, 48.6 mm SL; c, ZRC 65844, male, 51.5 mm SL; d, CMK 29184, female, 56.6 mm SL; e, CMK 29184, male, 52.3 mm SL; f, ZRC 65844, male, 57.5 mm SL; g, ZRC 65844, female, 66.0 mm SL." captionText-2="Fig. 3. Kapuasia falaris, paratypes, live, Borneo: Barito drainage; ZRC 65844. a, male, 43.0 mm SL (reversed); b, male, 51.5 mm SL; c, female, 66.0 mm SL (reversed)." pageId="5" pageNumber="484">Figs. 13</figureCitation>
). After fixation in formalin and 4 months in ethanol. Head and body background colour yellowish white. Unless otherwise stated, markings black. Head black; snout white on top, between nostrils to extremity, black on sides; white mark or bar on cheek, from immediately behind eye downwards. Posterior part of opercle white, extending across back, over occiput and continuous with contralateral. Outer rostral barbel black, other barbels whitish.
</paragraph>
<paragraph id="8B9B36C2AD21FFD4E54BFA95FAA3AFB2" blockId="5.[811,1452,1352,1504]" pageId="5" pageNumber="484">Body with 37 bars, extending from dorsal midline to ventral midline (faint on belly in front of anal fin), continuous with their contralaterals on back. Bars mostly of regular width and shape, wider than interspaces, sometimes paired or with paler central area, appearing vertically split.</paragraph>
<paragraph id="8B9B36C2AD21FFD4E54FF9D4FB0FADF2" blockId="5.[812,1453,1544,1952]" pageId="5" pageNumber="484">
Pattern at caudal-fin base: a conspicuous black blotch on each lobe of caudal fin, on proximal half, along upper and lower edge, irregular shape, extending for about half of length of lobe, on unbranched rays and adjacent 4 branched rays. Blotch on upper lobe, a spot of more intense black at anterior extremity. In most specimens, anterior extremity of upper blotch more forward than lower one. A faint inner mark in front of lower blotch. Between blotches, rays black, membranes hyaline, sometimes almost forming a bar. In most specimens, a smaller blotch behind both main ones, along dorsal and ventral edges. In most specimens, a narrow margin around part of whole caudal fin. In
<specimenCount id="9D22FD4BAD21FFD4E36FF8B4FA08ADD2" box="[1295,1447,1896,1920]" count="1" pageId="5" pageNumber="484" type="generic">one specimen</specimenCount>
, caudal fin almost entirely hyaline.
</paragraph>
<paragraph id="8B9B36C2AD21FFD4E54DF815FA03A212" blockId="5.[813,1452,1992,2113]" pageId="5" pageNumber="484">Dorsal fin hyaline, with a black blotch at base of unbranched rays and first branched rays (12). A stripe made of melanophores at middle of last unbranched ray, and in middle area of branched rays, on rays near branching points.</paragraph>
<caption id="DF5B664AAD22FFD7E524FDDBFA3DA84E" box="[836,1426,518,540]" pageId="6" pageNumber="485" startId="6.[836,871,518,540]" targetBox="[836,1474,189,503]" targetPageId="6" targetType="figure">
<paragraph id="8B9B36C2AD22FFD7E524FDDBFA3DA84E" blockId="6.[836,1426,518,540]" box="[836,1426,518,540]" pageId="6" pageNumber="485">
Fig. 7.
<taxonomicName id="4C244D41AD22FFD7E5EDFDDAFBDCA849" authority=", CMK" authorityName="CMK" box="[909,1139,518,540]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD22FFD7E5EDFDDAFB81A849" box="[909,1070,518,540]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
, CMK
</taxonomicName>
29184, 51.4 mm SL; mouth.
</paragraph>
</caption>
<caption id="DF5B664AAD22FFD7E524FAECFC42AF31" pageId="6" pageNumber="485" startId="6.[836,872,1329,1351]" targetBox="[1030,1280,581,1315]" targetPageId="6" targetType="figure">
<paragraph id="8B9B36C2AD22FFD7E524FAECFC42AF31" blockId="6.[836,1474,1329,1380]" pageId="6" pageNumber="485">
Fig. 8.
<taxonomicName id="4C244D41AD22FFD7E5EFFAECFBDAAF14" authority=", ZRC" authorityName="ZRC" box="[911,1141,1329,1350]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD22FFD7E5EFFAECFB99AF14" box="[911,1078,1329,1350]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
, ZRC
</taxonomicName>
65915, 55.0 mm SL; schematic. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8B9B36C2AD22FFD7E523FA46FB42AF81" blockId="6.[835,1474,1435,1491]" pageId="6" pageNumber="485">No melanophores on membranes, except between branches immediately distal to branching points.</paragraph>
<paragraph id="8B9B36C2AD22FFD7E523FA26FAC9AC01" blockId="6.[835,1474,1531,1619]" pageId="6" pageNumber="485">Anal fin hyaline, with a row of patches of melanophores on rays near branching points. Pelvic fin hyaline or with a few faint melanophores on rays near branching points.</paragraph>
<paragraph id="8B9B36C2AD22FFD7E523F9A6FC76ACA1" blockId="6.[835,1475,1659,1779]" pageId="6" pageNumber="485">Pectoral fin hyaline. Unbranched and branched rays 12 with a patch of melanophores on dorsal surface of distal ½ to ⅓. On other rays, melanophores along edge of rays and branches.</paragraph>
<paragraph id="8B9B36C2AD22FFD7E523F8C6FA21AD81" blockId="6.[835,1475,1819,2003]" pageId="6" pageNumber="485">In life: Pattern as in preserved specimens. Body cream-white, dorsum slight golden, venter white. Iris of eye flecked with gold specks. Bars on body brown, darker on dorsum. Caudal fin hyaline to light yellow, can be fully red. Upper part of last pale band (between last black bar and blotch at upper caudal base, and top of snout, bright iridescent white.</paragraph>
<caption id="DF5B664AAD22FFD7E6E7F81DFD59A27E" pageId="6" pageNumber="485" startId="6.[135,171,1984,2006]" targetBox="[135,773,189,1969]" targetPageId="6" targetType="figure">
<paragraph id="8B9B36C2AD22FFD7E6E7F81DFD59A27E" blockId="6.[135,774,1984,2093]" pageId="6" pageNumber="485">
Fig. 6.
<taxonomicName id="4C244D41AD22FFD7E6B4F81CFED3AD87" box="[212,380,1984,2006]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="485" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD22FFD7E6B4F81CFED3AD87" box="[212,380,1984,2006]" italics="true" pageId="6" pageNumber="485">Kapuasia falaris</emphasis>
</taxonomicName>
; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL.
</paragraph>
</caption>
<paragraph id="8B9B36C2AD22FFD7E523F826FA6DA260" blockId="6.[835,1474,2043,2099]" pageId="6" pageNumber="485">
<emphasis id="B950EAD0AD22FFD7E523F826FBBFA241" bold="true" box="[835,1040,2043,2067]" pageId="6" pageNumber="485">Notes on biology.</emphasis>
The stomach content of two dissected females (ZRC 65915,
<quantity id="4CDC9B27AD22FFD7E258F7C6FB36A260" box="[1080,1177,2075,2099]" metricMagnitude="-2" metricUnit="m" metricValue="5.92" pageId="6" pageNumber="485" unit="mm" value="59.2">59.2 mm</quantity>
SL, ZRC 65844,
<quantity id="4CDC9B27AD22FFD7E301F7C6FA6DA260" box="[1377,1474,2075,2099]" metricMagnitude="-2" metricUnit="m" metricValue="5.89" pageId="6" pageNumber="485" unit="mm" value="58.9">58.9 mm</quantity>
</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DFF60FB99ABC4" blockId="7.[811,1452,189,406]" pageId="7" pageNumber="486">is continuous, with a narrow shallow median longitudinal groove extending from the mental area to the anterior edge of the lip. The median part of the lower lip has 810 ridges on each side of the mental groove, radiating from the anterior extremity of the postlabial groove, across the whole lip, and also present on median space between the anterior extremities of the postlabial groove.</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DFE63FA1DA847" blockId="7.[813,1458,445,533]" pageId="7" pageNumber="486">
In
<taxonomicName id="4C244D41AD23FFD6E52BFE63FC1DAB87" box="[843,946,445,469]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E52BFE63FC1DAB87" box="[843,946,445,469]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
</taxonomicName>
, the lips are thinner, the groove in the median part of the lower lip is wider (but not forming conspicuous lobes) and the ridges are less obvious and less well organised.
</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DFDE3FAFBA947" blockId="7.[811,1452,573,789]" pageId="7" pageNumber="486">
<taxonomicName id="4C244D41AD23FFD6E54DFDE3FC3DA807" box="[813,914,574,597]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="genus">
<emphasis id="B950EAD0AD23FFD6E54DFDE3FC3DA807" box="[813,914,574,597]" italics="true" pageId="7" pageNumber="486">Kapuasia</emphasis>
</taxonomicName>
was also distinguished from all known nemacheilid loaches by the presence of a large, conspicuous, rounded suprapectoral lobe above the pectoral fin. In
<taxonomicName id="4C244D41AD23FFD6E375FDA3FAD3A8C7" box="[1301,1404,637,661]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E375FDA3FAD3A8C7" box="[1301,1404,637,661]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
</taxonomicName>
, the lobe is cylindrical, posteriorly pointed, usually separated from skin of flank by a deep groove; groove often margined above by a second pointed lobe (
<figureCitation id="131F2A47AD23FFD6E2EAFD00FB64A8A7" box="[1162,1227,733,757]" captionStart="Fig" captionStartId="6.[135,171,1984,2006]" captionTargetBox="[135,773,189,1969]" captionTargetId="figure-8@6.[134,773,189,1969]" captionTargetPageId="6" captionText="Fig. 6. Kapuasia falaris; variability in shape of supra-pectoral lobe(s). a, ZRC 65844, female, 56.1 mm SL; b, CMK 29184, male, 52.3 mm SL; c, ZRC 65844, female, 58.9 mm SL; d, CMK 29184, male, 50.1 mm SL; e, CMK 29184, male, 51.4 mm SL." pageId="7" pageNumber="486">Fig. 6</figureCitation>
), often less regularly shaped and more or less distinct than the first one.
</paragraph>
<caption id="DF5B664AAD23FFD6E612FBA9FD30AEF5" pageId="7" pageNumber="486" startId="7.[114,149,1140,1162]" targetBox="[114,752,189,1112]" targetPageId="7" targetType="figure">
<paragraph id="8B9B36C2AD23FFD6E612FBA9FD30AEF5" blockId="7.[114,753,1140,1191]" pageId="7" pageNumber="486">
Fig. 9.
<taxonomicName id="4C244D41AD23FFD6E6DBFBA8FEF3AEDB" box="[187,348,1140,1162]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E6DBFBA8FEF3AEDB" box="[187,348,1140,1162]" italics="true" pageId="7" pageNumber="486">Kapuasia falaris</emphasis>
</taxonomicName>
, pectoral fin in dorsal view; ZRC 65844. a, female, 58.9 mm SL; b, male 52.3 mm SL (reversed).
</paragraph>
</caption>
<paragraph id="8B9B36C2AD23FFD6E611FB03FD5CAC64" blockId="7.[113,755,1246,1590]" pageId="7" pageNumber="486">
SL) is uninformative because specimens kept in captivity for several days. The ovaries of the
<quantity id="4CDC9B27AD23FFD6E79BFB23FDF4AF44" box="[507,603,1278,1302]" metricMagnitude="-2" metricUnit="m" metricValue="5.89" pageId="7" pageNumber="486" unit="mm" value="58.9">58.9 mm</quantity>
SL specimen contained unripe eggs, the largest ones about
<quantity id="4CDC9B27AD23FFD6E4F7FAC3FD40AF64" box="[663,751,1310,1334]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="486" unit="mm" value="0.5">0.5 mm</quantity>
diameter. We have no information on habitat. The curled and slanted orientation of the pectoral and pelvic fins suggest an adaptation to strong current. In captivity: in a
<quantity id="4CDC9B27AD23FFD6E435FAA3FD39AFC4" box="[597,662,1406,1430]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="486" unit="cm" value="60.0">60 cm</quantity>
(length) tank setup with rocks and stones, individuals remain not visible as they hide amongst the crevices or burrowed into coarse gravel. When food is introduced, the fish will swim out or even forcefully push against the coarse gravel in its foraging activity. They have a preference for a carnivore diet.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD23FFD6E611F983FDA1ACE4" pageId="7" pageNumber="486" type="distribution">
<paragraph id="8B9B36C2AD23FFD6E611F983FDA1ACE4" blockId="7.[112,753,1630,1718]" pageId="7" pageNumber="486">
<emphasis id="B950EAD0AD23FFD6E611F983FEACAC24" bold="true" box="[113,259,1630,1654]" pageId="7" pageNumber="486">Distribution.</emphasis>
<taxonomicName id="4C244D41AD23FFD6E768F982FE17AC24" box="[264,440,1630,1654]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E768F982FE17AC24" box="[264,440,1630,1654]" italics="true" pageId="7" pageNumber="486">Kapuasia falaris</emphasis>
</taxonomicName>
is presently known only from one locality in the Barito drainage, Borneo. It is logically expected to have a wider distribution.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD23FFD6E611F903FD9DAD24" pageId="7" pageNumber="486" type="etymology">
<paragraph id="8B9B36C2AD23FFD6E611F903FD9DAD24" blockId="7.[113,752,1758,1910]" pageId="7" pageNumber="486">
<emphasis id="B950EAD0AD23FFD6E611F903FF5CACA4" bold="true" box="[113,243,1758,1782]" pageId="7" pageNumber="486">Etymology.</emphasis>
From the Greek phalaros (having a patch of white, φάλαρος) and rhis (nose, snout, ῥίς); with a white spot on the nose. Used as a noun in apposition. Falaris or phalaris (φαλαρίς), is the name of the coot,
<taxonomicName id="4C244D41AD23FFD6E439F8E3FD7BAD04" authorityName="Linnaeus" authorityYear="1758" box="[601,724,1854,1878]" class="Aves" family="Rallidae" genus="Fulica" kingdom="Animalia" order="Gruiformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="atra">
<emphasis id="B950EAD0AD23FFD6E439F8E3FD7BAD04" box="[601,724,1854,1878]" italics="true" pageId="7" pageNumber="486">Fulica atra</emphasis>
</taxonomicName>
, a black bird with white beak and forehead.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD23FFD6E611F843FB58AD87" pageId="7" pageNumber="486" type="discussion">
<paragraph id="8B9B36C2AD23FFD6E611F843FB4BAEC7" blockId="7.[113,754,1950,2103]" lastBlockId="7.[812,1454,829,1173]" pageId="7" pageNumber="486">
<emphasis id="B950EAD0AD23FFD6E611F843FF4EADE4" bold="true" box="[113,225,1950,1974]" pageId="7" pageNumber="486">Remarks.</emphasis>
<taxonomicName id="4C244D41AD23FFD6E68BF842FEFDADE4" box="[235,338,1951,1974]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="genus">
<emphasis id="B950EAD0AD23FFD6E68BF842FEFDADE4" box="[235,338,1951,1974]" italics="true" pageId="7" pageNumber="486">Kapuasia</emphasis>
</taxonomicName>
(and
<taxonomicName id="4C244D41AD23FFD6E7F6F842FD9DADE4" authorityName="Roberts" authorityYear="1989" box="[406,562,1950,1974]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E7F6F842FD9DADE4" box="[406,562,1950,1974]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
</taxonomicName>
) is distinguished from all other genera of
<taxonomicName id="4C244D41AD23FFD6E71FF863FDB0AD84" box="[383,543,1982,2006]" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
in the morphology of the mouth. It is strongly arched, U-shaped, the upper lip is swollen and smooth. The lower lip is swollen, with a deep postlabial groove along the lateral parts; its anterior edge
<specimenCount id="9D22FD4BAD23FFD6E54DFCE0FC68A907" box="[813,967,829,853]" count="1" pageId="7" pageNumber="486" type="generic">One specimen</specimenCount>
of
<taxonomicName id="4C244D41AD23FFD6E58FFCE3FB26A907" authorityName="Roberts" authorityYear="1989" box="[1007,1161,829,853]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E58FFCE3FB26A907" box="[1007,1161,829,853]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
</taxonomicName>
had a few small tubercles on the anterior branched pectoral-fin rays, assumed to be a male character. Both sexes are present in the material of
<taxonomicName id="4C244D41AD23FFD6E54DFC43FC3CA9E7" box="[813,915,925,949]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E54DFC43FC3CA9E7" box="[813,915,925,949]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
</taxonomicName>
; no tubercles were present in males. Males of
<taxonomicName id="4C244D41AD23FFD6E3F5FC43FCDCA987" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E3F5FC43FCDCA987" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
</taxonomicName>
are also distinguished by the presence of vertical skin flanges between the branched rays of the pectoral fin (
<figureCitation id="131F2A47AD23FFD6E31FFC00FC93AE47" captionStart="Fig" captionStartId="5.[813,848,610,632]" captionTargetBox="[814,1452,189,595]" captionTargetId="figure-584@5.[813,1452,189,595]" captionTargetPageId="5" captionText="Fig. 5. Kapuasia falaris,CMK 29184, male, 52.3 mm SL; pectoral fin, antero-dorsal view, showing skin flanges between branched rays (arrows)." pageId="7" pageNumber="486">Fig. 5</figureCitation>
); such flanges had not been reported in
<taxonomicName id="4C244D41AD23FFD6E368FC23FA0AAE47" authorityName="Roberts" authorityYear="1989" box="[1288,1445,1021,1045]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E368FC23FA0AAE47" box="[1288,1445,1021,1045]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
</taxonomicName>
, but a shallow ridge of skin between the rays is visible on the photographs of ZRC 56404, of unknown sex, and ZRC 61464, apparently a male, and appears possibly homologous (
<bibRefCitation id="EFB54B33AD23FFD6E553FBA0FB8CAEC7" author="Kottelat M &amp; Tan HH" box="[819,1059,1149,1173]" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M &amp; Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat &amp; Tan, 2024</bibRefCitation>
: figs. 1b, c, 3, 4).
</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DFB60FBCEAC47" blockId="7.[813,1452,1213,1557]" pageId="7" pageNumber="486">
The males
<taxonomicName id="4C244D41AD23FFD6E5CCFB63FBB7AE87" box="[940,1048,1213,1237]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="falaris">
<emphasis id="B950EAD0AD23FFD6E5CCFB63FBB7AE87" box="[940,1048,1213,1237]" italics="true" pageId="7" pageNumber="486">K. falaris</emphasis>
</taxonomicName>
have a conspicuous suborbital flap (modified lateral ethmoid;
<figureCitation id="131F2A47AD23FFD6E232FB00FB3AAEA7" box="[1106,1173,1245,1269]" captionStart="Fig" captionStartId="5.[113,148,1165,1187]" captionTargetBox="[113,751,189,1150]" captionTargetId="figure-14@5.[113,752,189,1150]" captionTargetPageId="5" captionText="Fig. 4. Kapuasia falaris, ZRC 65844, upper part of cheek; a, 57.5 mm SL; male with suborbital flap (arrow); b, 58.9 mm SL; female with suborbital crease (arrow)." pageId="7" pageNumber="486">Fig. 4</figureCitation>
), which is missing in the single specimen of
<taxonomicName id="4C244D41AD23FFD6E266FB23FB0EAF47" authorityName="Roberts" authorityYear="1989" box="[1030,1185,1277,1301]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E266FB23FB0EAF47" box="[1030,1185,1277,1301]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
</taxonomicName>
tentatively identified as male. The flap is globulous, movable, at some distance below the eye, with a deep groove connecting it to the edge of the orbit. In most nemacheilds of Southeast Asia, when a flap is present, it is immediately below the eye and the groove is not present. A somewhat similar condition is reported only in
<taxonomicName id="4C244D41AD23FFD6E58EFA63FABCAF87" box="[1006,1299,1469,1493]" class="Actinopterygii" family="Nemacheilidae" genus="Mustura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="shuangjiangensis">
<emphasis id="B950EAD0AD23FFD6E58EFA63FBE7AF87" box="[1006,1096,1470,1493]" italics="true" pageId="7" pageNumber="486">Mustura</emphasis>
<emphasis id="B950EAD0AD23FFD6E23AFA60FABCAF87" box="[1114,1299,1469,1493]" italics="true" pageId="7" pageNumber="486">shuangjiangensis</emphasis>
</taxonomicName>
, in which the flap is at a distance from the eye, and the groove slanted (see
<bibRefCitation id="EFB54B33AD23FFD6E500FA20FC50AC47" author="Kottelat M" box="[864,1023,1533,1557]" pageId="7" pageNumber="486" pagination="289 - 314" refId="ref4912" refString="Kottelat M (2018) Mustura celata, a new genus and species of loaches from northern Myanmar, and an overview of Physoschistura and related taxa (Teleostei: Nemacheilidae). Ichthyological Exploration of Freshwaters, 28 (4): 289 - 314." type="journal article" year="2018">Kottelat, 2018</bibRefCitation>
: fig. 26).
</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DF9E0FADEACA7" blockId="7.[811,1452,1597,1781]" pageId="7" pageNumber="486">Another character shared by the two species and distinguishing them from all sundaic nemacheilids is the colour pattern of the caudal fin, made of a pair of black blotches in the proximal area of the fin, of which the anterior extremity of the lower one is slightly pushed backwards, the black rays between them, and a bar or row of spots in subdistal position.</paragraph>
<paragraph id="8B9B36C2AD23FFD6E54DF8C0FB58AD87" blockId="7.[811,1453,1821,2005]" pageId="7" pageNumber="486">
The strongly curled pectoral and pevic fins, with innermost rays positioned very close to the body and appearing in life as if the fins were vertical is a feature not observed in any other nemacheilid in Southeast Asia. However, the preserved specimen of
<taxonomicName id="4C244D41AD23FFD6E5D2F843FBE7ADE7" authorityName="Roberts" authorityYear="1989" box="[946,1096,1949,1973]" family="Nemacheilidae" genus="Nemacheilus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E5D2F843FBE7ADE7" box="[946,1096,1949,1973]" italics="true" pageId="7" pageNumber="486">K. maculiceps</emphasis>
</taxonomicName>
figured by
<bibRefCitation id="EFB54B33AD23FFD6E2A0F840FA07ADE7" author="Kottelat M &amp; Tan HH" box="[1216,1448,1949,1973]" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M &amp; Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat &amp; Tan (2024</bibRefCitation>
: fig. 2) shows a possibly similar condition.
</paragraph>
</subSubSection>
<subSubSection id="C33E6549AD23FFD6E54DF820FC02A267" pageId="7" pageNumber="486" type="materials_examined">
<paragraph id="8B9B36C2AD23FFD6E54DF820FC02A267" blockId="7.[813,1452,2045,2101]" pageId="7" pageNumber="486">
<emphasis id="B950EAD0AD23FFD6E54DF820FBA2A247" bold="true" box="[813,1037,2045,2069]" pageId="7" pageNumber="486">Material examined.</emphasis>
<taxonomicName id="4C244D41AD23FFD6E276F823FB52A247" box="[1046,1277,2045,2069]" family="Nemacheilidae" genus="Kapuasia" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="486" phylum="Chordata" rank="species" species="maculiceps">
<emphasis id="B950EAD0AD23FFD6E276F823FB52A247" box="[1046,1277,2045,2069]" italics="true" pageId="7" pageNumber="486">Kapuasia maculiceps</emphasis>
</taxonomicName>
: see
<bibRefCitation id="EFB54B33AD23FFD6E358F820FC06A267" author="Kottelat M &amp; Tan HH" pageId="7" pageNumber="486" pagination="105 - 109" refId="ref5065" refString="Kottelat M &amp; Tan HH (2024) Kapuasia, a genus name for ' Nemacheilus' maculiceps (Teleostei: Nemacheilidae). Raffles Bulletin of Zoology, 72: 105 - 109." type="journal article" year="2024">Kottelat &amp; Tan (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,193 @@
<document id="0A5C3CB7011B86C6B1F0C6EF2AF566EF" ID-DOI="10.26107/RBZ-2024-0024" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683052" ID-ZooBank="F4FB9765-D4A1-48AB-8845-EC63B5A7D794" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171910635" checkinUser="felipe" docAuthor="Hoang, Quang D., , Quoc T. Phan, Vo, &amp; Van M." docDate="2024" docId="03A88780FFDBFF93FEDAD055D29155A6" docLanguage="en" docName="RafflesBZool.72.294-302.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Gelotia zhengi Cao &amp; Li 2016" docType="treatment" docVersion="2" lastPageNumber="302" masterDocId="FF91FFF8FFDCFF9BFFC3D723D3175413" masterDocTitle="First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species" masterLastPageNumber="302" masterPageNumber="294" pageNumber="301" updateTime="1737175710851" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="74778DDFF09D60B3057D6F5193A7BA6A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="160BEACB6A4EC6255D2E03AD8F20BF08">
<mods:title id="1D17BD7F3C891DD2480E069B36C5F77B">First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="A1A7B218F3A814068D8871FBE160ED6F" type="personal">
<mods:role id="34EAACEE3D164EC80207C905A57B3E83">
<mods:roleTerm id="542AB5BFDCFCA4E0A33F64CDB8D964C8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E5FDA159CEF191FDFD059AE331170825">Hoang, Quang D.</mods:namePart>
</mods:name>
<mods:name id="A73431891B2BDAAD4AE3EFF36E952260" type="personal">
<mods:role id="D448D955C32952CF65CD2282A83703B8">
<mods:roleTerm id="6B6451B4E9D981C250A47C30A3B35240">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A3FB15C04A66F7021718217A35D7279">, Quoc T. Phan</mods:namePart>
</mods:name>
<mods:name id="4A5FC85673F63DF31887CF37F7B463C6" type="personal">
<mods:role id="C59016655B035388F1E79E067026FFCA">
<mods:roleTerm id="1B89D174F01CC0F3E0A148871BCCB7C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="491F17A6C7B0409D2F13D094680A5F72">Vo,</mods:namePart>
</mods:name>
<mods:name id="207B96ACCDBDA3023232542EB0585120" type="personal">
<mods:role id="992DA87ABA72BD3E19472C00BFB23ADD">
<mods:roleTerm id="7E21238FD02DF6D326C65473995830FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E6B6E3629A2A63EF0A37ABF81BE93667">Van M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="87975B9A3473F1C566418360B70B475D">text</mods:typeOfResource>
<mods:relatedItem id="2F045F395EFF8ADD0960F777791917A6" type="host">
<mods:titleInfo id="AFB3A2D53EFF414F02DB8D2E2F97215A">
<mods:title id="147767813AD27D5788E49407325F55BB">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="36492A14CE21E6BF981DE49DB6C6BDCD">
<mods:date id="B57C28D78D0840034FD6AB6735319CA3">2024</mods:date>
<mods:detail id="E6180FDDCE66B23151C4122780D6318A" type="pubDate">
<mods:number id="76BD22889EAB18E2D702C491AE9D0C91">2024-08-29</mods:number>
</mods:detail>
<mods:detail id="787AC6F3ADBDB5D6BFA37F1ECD640415" type="volume">
<mods:number id="F5856E54E3DA540F9D211127BF183E48">72</mods:number>
</mods:detail>
<mods:extent id="73BC73852A764A739B717DA837616DE3" unit="page">
<mods:start id="9754AF48BF93FE493786304B01DF557B">294</mods:start>
<mods:end id="992E2FFB1B0E7E2DE58F8EA63723E11C">302</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6BA9B8C406CB8E3A243C87C9F4F4AA32">journal article</mods:classification>
<mods:identifier id="37EB6A730BD90692FDC42677838485DD" type="DOI">10.26107/RBZ-2024-0024</mods:identifier>
<mods:identifier id="55023EA4BFDFB585A29A94B98EA4D22F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F94DCBD7CF790EEF9CE671A53F1A9303" type="Zenodo-Dep">14683052</mods:identifier>
<mods:identifier id="AE0E1F8BDB55D29F174BD47A8DAB135B" type="ZooBank">F4FB9765-D4A1-48AB-8845-EC63B5A7D794</mods:identifier>
</mods:mods>
<treatment id="03A88780FFDBFF93FEDAD055D29155A6" LSID="urn:lsid:plazi:treatment:03A88780FFDBFF93FEDAD055D29155A6" httpUri="http://treatment.plazi.org/id/03A88780FFDBFF93FEDAD055D29155A6" lastPageId="8" lastPageNumber="302" pageId="7" pageNumber="301">
<subSubSection id="C31B651DFFDBFF9CFEDAD055D161539D" box="[281,630,1910,1934]" pageId="7" pageNumber="301" type="nomenclature">
<paragraph id="8BBE3696FFDBFF9CFEDAD055D161539D" blockId="7.[281,630,1910,1966]" box="[281,630,1910,1934]" pageId="7" pageNumber="301">
<heading id="D0F681FAFFDBFF9CFEDAD055D161539D" bold="true" box="[281,630,1910,1934]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="301" reason="2">
<taxonomicName id="4C014D15FFDBFF9CFEDAD055D161539D" ID-CoL="3FK47" authority="Cao &amp; Li, 2016" authorityName="Cao &amp; Li" authorityYear="2016" box="[281,630,1910,1934]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="301" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B975EA84FFDBFF9CFEDAD055D161539D" bold="true" box="[281,630,1910,1934]" pageId="7" pageNumber="301">
<emphasis id="B975EA84FFDBFF9CFEDAD055D2AD539D" bold="true" box="[281,442,1910,1934]" italics="true" pageId="7" pageNumber="301">Gelotia zhengi</emphasis>
<bibRefCitation id="EF904B67FFDBFF9CFE00D055D161539D" author="Cao Q &amp; Li S &amp; Zabka M" box="[451,630,1910,1934]" pageId="7" pageNumber="301" pagination="43 - 104" refId="ref4253" refString="Cao Q, Li S &amp; Zabka M (2016) The jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 630: 43 - 104." type="journal article" year="2016">Cao &amp; Li, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDBFF9CFE4FD0B5D11453BD" box="[396,515,1942,1966]" pageId="7" pageNumber="301" type="description">
<paragraph id="8BBE3696FFDBFF9CFE4FD0B5D11453BD" blockId="7.[281,630,1910,1966]" box="[396,515,1942,1966]" pageId="7" pageNumber="301">
(
<figureCitation id="133A2A13FFDBFF9CFE57D0B5D2EC53BD" box="[404,507,1942,1966]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[113,148,1989,2011]" captionStartId-1="7.[136,171,1324,1346]" captionTargetBox-0="[124,1449,264,1973]" captionTargetBox-1="[136,1474,189,1309]" captionTargetId-0="figure-12@6.[113,1449,264,1973]" captionTargetId-1="figure-43@7.[136,1474,189,1309]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 5. Male of Gelotia zhengi Cao &amp; Li, 2016. A, habitus, dorsal view; B, carapace, anterior view; C, palp, ventral view; D, palp, retrolateral view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.3 mm." captionText-1="Fig. 6. Female of Gelotia zhengi Cao &amp; Li, 2016. A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, B, 1 mm, C, D, 0.2 mm." pageId="7" pageNumber="301">Figs. 56</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDBFF9CFF49D0F6D7FF527C" pageId="7" pageNumber="301" type="reference_group">
<paragraph id="8BBE3696FFDBFF9CFF49D0F6D1575C37" blockId="7.[138,776,2005,2084]" pageId="7" pageNumber="301">
<taxonomicName id="4C014D15FFDBFF9CFF49D0F6D29D53F8" ID-CoL="3FK47" authority="Cao &amp; Li" authorityName="Cao &amp; Li" authorityYear="2016" box="[138,394,2005,2027]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="301" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B975EA84FFDBFF9CFF49D0F6D20A53F9" box="[138,285,2005,2026]" italics="true" pageId="7" pageNumber="301">Gelotia zhengi</emphasis>
Cao &amp; Li
</taxonomicName>
, in
<bibRefCitation id="EF904B67FFDBFF9CFE77D0F6D1C753F9" author="Cao Q &amp; Li S &amp; Zabka M" box="[436,720,2005,2028]" pageId="7" pageNumber="301" pagination="43 - 104" refId="ref4253" refString="Cao Q, Li S &amp; Zabka M (2016) The jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 630: 43 - 104." type="journal article" year="2016">Cao, Li &amp; Żabka, 2016: 78</bibRefCitation>
, figs. 24AD, 25AB (males);
<bibRefCitation id="EF904B67FFDBFF9CFE68D0D1D1905C14" author="Wang C &amp; Li SQ" box="[427,647,2034,2056]" pageId="7" pageNumber="301" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li, 2020: 41</bibRefCitation>
, figs. 9AD, 10AG, 17G, 18G, 19G (male, females).
</paragraph>
<paragraph id="8BBE3696FFDBFF9CFC80D2B5D7FF527C" blockId="7.[834,1475,1430,1647]" pageId="7" pageNumber="301">
<emphasis id="B975EA84FFDBFF9CFC80D2B5D73051BD" bold="true" box="[835,1063,1430,1454]" pageId="7" pageNumber="301">Marterial examined.</emphasis>
<materialsCitation id="3B693CCBFFDBFF9CFBE8D2B5D0A6523D" collectingDate="2021-11-21" collectorName="Q. D. Hoang. Male" country="Vietnam" elevation="121" latitude="11.4063" location="Tan Phu Dist." longLatPrecision="7" longitude="107.437" municipality="Male" pageId="7" pageNumber="301" specimenCount="1" stateProvince="Dong Nai">
<collectingMunicipality id="6BDAACECFFDBFF9CFBE8D2B5D77651BD" box="[1067,1121,1430,1454]" pageId="7" pageNumber="301">Male</collectingMunicipality>
(VNMN-ARA-SAL-117), female (VNMN-ARA-SAL-95),
<collectingCountry id="F3167606FFDBFF9CFBABD295D7FB51DD" box="[1128,1260,1462,1486]" name="Vietnam" pageId="7" pageNumber="301">VIETNAM</collectingCountry>
,
<collectingRegion id="49C5F874FFDBFF9CFB3ED294D6AC51DD" box="[1277,1467,1462,1487]" country="Vietnam" name="Dong Nai" pageId="7" pageNumber="301">Dong Nai Prov.</collectingRegion>
,
<location id="8EDE604DFFDBFF9CFC80D2F4D0F451FD" LSID="urn:lsid:plazi:treatment:03A88780FFDBFF93FEDAD055D29155A6:8EDE604DFFDBFF9CFC80D2F4D0F451FD" box="[835,995,1494,1518]" country="Vietnam" latitude="11.4063" longLatPrecision="7" longitude="107.437" municipality="Male" name="Tan Phu Dist." pageId="7" pageNumber="301" stateProvince="Dong Nai">Tan Phu Dist.</location>
, (
<geoCoordinate id="EE355051FFDBFF9CFC3AD2F5D76451FD" box="[1017,1139,1494,1518]" degrees="11.4063" direction="north" orientation="latitude" pageId="7" pageNumber="301" precision="5" value="11.4063">11.4063°N</geoCoordinate>
,
<geoCoordinate id="EE355051FFDBFF9CFB42D2F5D61151FD" box="[1153,1286,1494,1518]" degrees="107.4370" direction="east" orientation="longitude" pageId="7" pageNumber="301" precision="5" value="107.437">107.4370°E</geoCoordinate>
),
<elevation id="002CD1A5FFDBFF9CFADFD2F5D68F51FD" box="[1308,1432,1494,1518]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="301" unit="m" value="121.0">
<quantity id="4CF99B73FFDBFF9CFADFD2F5D64D51FD" box="[1308,1370,1494,1518]" metricMagnitude="2" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="301" unit="m" value="121.0">121m</quantity>
a.s.l.
</elevation>
,
<date id="FFBF1056FFDBFF9CFA6BD2F5D0EE521D" pageId="7" pageNumber="301" value="2021-11-21">
<collectingDate id="EFFBE9BEFFDBFF9CFA6BD2F5D0EE521D" pageId="7" pageNumber="301" value="2021-11-21">21 November 2021</collectingDate>
</date>
, coll.
<collectorName id="26F45340FFDBFF9CFB82D2D5D606521D" box="[1089,1297,1526,1551]" pageId="7" pageNumber="301">Q.D. Hoang. Male</collectorName>
(VNMN-ARA- SAL-226)
</materialsCitation>
,
<materialsCitation id="3B693CCBFFDBFF9CFC79D135D7FF527C" collectingDate="2022-05-14" collectorName="Q. D. Hoang." country="Vietnam" elevation="893" latitude="12.2686" location="Chu Yang Sin National Park" longLatPrecision="7" longitude="108.2305" municipality="Krong Bong District" pageId="7" pageNumber="301" specimenCount="1" stateProvince="Dak Lak">
<collectingCountry id="F3167606FFDBFF9CFC79D135D720523D" box="[954,1079,1558,1582]" name="Vietnam" pageId="7" pageNumber="301">VIETNAM</collectingCountry>
,
<collectingRegion id="49C5F874FFDBFF9CFB83D135D7CB523D" box="[1088,1244,1558,1582]" country="Vietnam" name="Dak Lak" pageId="7" pageNumber="301">Dak Lak Prov.</collectingRegion>
,
<collectingMunicipality id="6BDAACECFFDBFF9CFB27D134D6A9523D" box="[1252,1470,1558,1583]" pageId="7" pageNumber="301">Krong Bong District</collectingMunicipality>
,
<location id="8EDE604DFFDBFF9CFC81D115D77F525D" LSID="urn:lsid:plazi:treatment:03A88780FFDBFF93FEDAD055D29155A6:8EDE604DFFDBFF9CFC81D115D77F525D" box="[834,1128,1590,1615]" country="Vietnam" latitude="12.2686" longLatPrecision="7" longitude="108.2305" municipality="Krong Bong District" name="Chu Yang Sin National Park" pageId="7" pageNumber="301" stateProvince="Dak Lak">Chu Yang Sin National Park</location>
(
<geoCoordinate id="EE355051FFDBFF9CFBB7D115D7FC525D" box="[1140,1259,1590,1614]" degrees="12.2686" direction="north" orientation="latitude" pageId="7" pageNumber="301" precision="5" value="12.2686">12.2686°N</geoCoordinate>
,
<geoCoordinate id="EE355051FFDBFF9CFB36D115D663525D" box="[1269,1396,1590,1614]" degrees="108.2305" direction="east" orientation="longitude" pageId="7" pageNumber="301" precision="5" value="108.2305">108.2305°E</geoCoordinate>
),
<elevation id="002CD1A5FFDBFF9CFA45D115D061527D" metricMagnitude="2" metricUnit="m" metricValue="8.93" pageId="7" pageNumber="301" unit="m" value="893.0">
<quantity id="4CF99B73FFDBFF9CFA45D115D6D5525D" box="[1414,1474,1590,1614]" metricMagnitude="2" metricUnit="m" metricValue="8.93" pageId="7" pageNumber="301" unit="m" value="893.0">893m</quantity>
a.s.l.
</elevation>
,
<date id="FFBF1056FFDBFF9CFC47D175D70E527D" box="[900,1049,1622,1647]" pageId="7" pageNumber="301" value="2022-05-14">
<collectingDate id="EFFBE9BEFFDBFF9CFC47D175D70E527D" box="[900,1049,1622,1647]" pageId="7" pageNumber="301" value="2022-05-14">14 May 2022</collectingDate>
</date>
, coll.
<collectorName id="26F45340FFDBFF9CFB9ED175D7FF527C" box="[1117,1256,1622,1647]" pageId="7" pageNumber="301">Q.D. Hoang.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDBFF9CFC80D1B5D7C253DD" pageId="7" pageNumber="301" type="description">
<paragraph id="8BBE3696FFDBFF9CFC80D1B5D7C252FD" blockId="7.[834,1474,1686,1774]" pageId="7" pageNumber="301">
<emphasis id="B975EA84FFDBFF9CFC80D1B5D0D952BD" bold="true" box="[835,974,1686,1710]" pageId="7" pageNumber="301">Description.</emphasis>
For both sexes, see
<bibRefCitation id="EF904B67FFDBFF9CFB62D1B5D64752BD" author="Cao Q &amp; Li S &amp; Zabka M" box="[1185,1360,1686,1710]" pageId="7" pageNumber="301" pagination="43 - 104" refId="ref4253" refString="Cao Q, Li S &amp; Zabka M (2016) The jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 630: 43 - 104." type="journal article" year="2016">Cao et al. (2016)</bibRefCitation>
and
<bibRefCitation id="EF904B67FFDBFF9CFA41D1B4D0D552DD" author="Wang C &amp; Li SQ" pageId="7" pageNumber="301" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li (2020)</bibRefCitation>
. Habitus and copulatory organs from Vietnamese specimens as illustrated in
<figureCitation id="133A2A13FFDBFF9CFBAAD1F5D7C552FD" box="[1129,1234,1750,1774]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[113,148,1989,2011]" captionStartId-1="7.[136,171,1324,1346]" captionTargetBox-0="[124,1449,264,1973]" captionTargetBox-1="[136,1474,189,1309]" captionTargetId-0="figure-12@6.[113,1449,264,1973]" captionTargetId-1="figure-43@7.[136,1474,189,1309]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 5. Male of Gelotia zhengi Cao &amp; Li, 2016. A, habitus, dorsal view; B, carapace, anterior view; C, palp, ventral view; D, palp, retrolateral view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.3 mm." captionText-1="Fig. 6. Female of Gelotia zhengi Cao &amp; Li, 2016. A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, B, 1 mm, C, D, 0.2 mm." pageId="7" pageNumber="301">Figs. 56</figureCitation>
.
</paragraph>
<paragraph id="8BBE3696FFDBFF9CFC80D035D7C253DD" blockId="7.[835,1475,1814,1999]" pageId="7" pageNumber="301">
<emphasis id="B975EA84FFDBFF9CFC80D035D0AF533D" bold="true" box="[835,952,1814,1838]" pageId="7" pageNumber="301">Variation.</emphasis>
There are slight differences with the illustration of the female by
<bibRefCitation id="EF904B67FFDBFF9CFC3AD014D7D4535D" author="Wang C &amp; Li SQ" box="[1017,1219,1846,1871]" pageId="7" pageNumber="301" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li (2020)</bibRefCitation>
, as the copulatory ducts are slightly wider and longer than those of the specimens from
<collectingCountry id="F3167606FFDBFF9CFC41D055D0D5539D" box="[898,962,1910,1934]" name="China" pageId="7" pageNumber="301">China</collectingCountry>
(
<figureCitation id="133A2A13FFDBFF9CFC17D055D73C539D" box="[980,1067,1910,1934]" captionStart="Fig" captionStartId="7.[136,171,1324,1346]" captionTargetBox="[136,1474,189,1309]" captionTargetId="figure-43@7.[136,1474,189,1309]" captionTargetPageId="7" captionText="Fig. 6. Female of Gelotia zhengi Cao &amp; Li, 2016. A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, B, 1 mm, C, D, 0.2 mm." pageId="7" pageNumber="301">Fig. 6D</figureCitation>
and fig. 10C in
<bibRefCitation id="EF904B67FFDBFF9CFB29D054D6A1539D" author="Wang C &amp; Li SQ" box="[1258,1462,1910,1935]" pageId="7" pageNumber="301" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li, 2020</bibRefCitation>
). The male is perfectly matched to the illustrations by
<bibRefCitation id="EF904B67FFDBFF9CFA54D0B5D0DF53DD" author="Cao Q &amp; Li S &amp; Zabka M" pageId="7" pageNumber="301" pagination="43 - 104" refId="ref4253" refString="Cao Q, Li S &amp; Zabka M (2016) The jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 630: 43 - 104." type="journal article" year="2016">Cao et al. (2016)</bibRefCitation>
and
<bibRefCitation id="EF904B67FFDBFF9CFBC2D094D7C553DD" author="Wang C &amp; Li SQ" box="[1025,1234,1974,1999]" pageId="7" pageNumber="301" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDBFF93FC80D0D5D29155A6" lastPageId="8" lastPageNumber="302" pageId="7" pageNumber="301" type="distribution">
<paragraph id="8BBE3696FFDBFF9CFC80D0D5D6D65C3D" blockId="7.[835,1473,2038,2094]" pageId="7" pageNumber="301">
<emphasis id="B975EA84FFDBFF9CFC80D0D5D0CF5C1D" bold="true" box="[835,984,2038,2062]" pageId="7" pageNumber="301">Distribution.</emphasis>
<collectingCountry id="F3167606FFDBFF9CFC21D0D5D7345C1D" box="[994,1059,2038,2062]" name="China" pageId="7" pageNumber="301">China</collectingCountry>
(
<collectingRegion id="49C5F874FFDBFF9CFBF6D0D5D7C55C1D" box="[1077,1234,2038,2062]" country="China" name="Yunnan" pageId="7" pageNumber="301">Yunnan Prov.</collectingRegion>
) and
<collectingCountry id="F3167606FFDBFF9CFAD6D0D5D6655C1D" box="[1301,1394,2038,2062]" name="Vietnam" pageId="7" pageNumber="301">Vietnam</collectingCountry>
(
<collectingRegion id="49C5F874FFDBFF9CFA47D0D5D0795C3D" country="Vietnam" name="Dong Nai" pageId="7" pageNumber="301">Dong Nai</collectingRegion>
,
<collectingRegion id="49C5F874FFDBFF9CFCBEDF35D73A5C3D" box="[893,1069,2070,2094]" country="Vietnam" name="Dak Lak" pageId="7" pageNumber="301">Dak Lak Prov.</collectingRegion>
). The species is probably found in
</paragraph>
<paragraph id="8BBE3696FFD4FF93FFB2D79ED29155A6" blockId="8.[113,752,189,437]" pageId="8" pageNumber="302">
<collectingCountry id="F3167606FFD4FF93FFB2D79ED3CC54C6" box="[113,219,189,213]" name="Malaysia" pageId="8" pageNumber="302">Malaysia</collectingCountry>
, as indicated by two Facebook posts (https:// www.facebook.com/photo/?fbid=10158373523431441 &amp;set=a.10150395133571441, Nicky Bay; https://www. facebook.com/photo/?fbid=10219423322027121&amp;set= gm.625020444714442, Paul Ng). The conformation of the RTA observed in the posts closely resembles the illustrations of
<taxonomicName id="4C014D15FFD4FF93FF4DD65ED23C5586" authorityName="Cao &amp; Li" authorityYear="2016" box="[142,299,381,405]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="302" phylum="Arthropoda" rank="species" species="zhengi">
<emphasis id="B975EA84FFD4FF93FF4DD65ED23C5586" box="[142,299,381,405]" italics="true" pageId="8" pageNumber="302">Gelotia zhengi</emphasis>
</taxonomicName>
, but further examination of the specimens is needed to confirm this.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,106 @@
<document id="9EDB6FDD00A2AF3B2924ABB8D5A76793" ID-DOI="10.26107/RBZ-2024-0024" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683052" ID-ZooBank="F4FB9765-D4A1-48AB-8845-EC63B5A7D794" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171910635" checkinUser="felipe" docAuthor="Hoang, Quang D., , Quoc T. Phan, Vo, &amp; Van M." docDate="2024" docId="03A88780FFDDFF9AFC22D035D0C153FD" docLanguage="en" docName="RafflesBZool.72.294-302.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Gelotia Thorell 1890" docType="treatment" docVersion="2" lastPageNumber="295" masterDocId="FF91FFF8FFDCFF9BFFC3D723D3175413" masterDocTitle="First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species" masterLastPageNumber="302" masterPageNumber="294" pageNumber="295" updateTime="1737175710851" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0C982B51AA0454714D3A2CC10F047FF3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F7ACF3CD28934B72D39B375DE0210BDA">
<mods:title id="F7388B35BDF3C16E5F3CA569C968E1E0">First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="835B1DF345B427FC73A37300E1268FD9" type="personal">
<mods:role id="532DE44819F255398CB68DB825C6AD7E">
<mods:roleTerm id="35514BE992241701BBC345F9F03CC19C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="37473E0B18B19786921BEECD27BF5ECC">Hoang, Quang D.</mods:namePart>
</mods:name>
<mods:name id="09F80EA011D5E2C6666803244137EE7F" type="personal">
<mods:role id="195AB7B9C7C07CBCBD3E619061B3053E">
<mods:roleTerm id="FC2BA2637101605E807271F4B0B9C753">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B80C646B2F96EFB7BC3D52BE0465D95D">, Quoc T. Phan</mods:namePart>
</mods:name>
<mods:name id="D58C88E4983E55AECC3F7D8D96829460" type="personal">
<mods:role id="66E7AF57EAE99A543DDFAD645F34B96B">
<mods:roleTerm id="F3880BF6A65C32BC30A032D306CFACE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B19399408A6225C2A8674CB21DE7560C">Vo,</mods:namePart>
</mods:name>
<mods:name id="960B19E76D27F73E18C98E516FD11B32" type="personal">
<mods:role id="8B4B5FF9819A6B906BDED63DD10817EC">
<mods:roleTerm id="8CF32C7BCDEADD71EA32DB08C037961D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F81611B9A765088B4CCDD9E8DDF0777">Van M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="592A2B9192D7A32DF56BA32483CE3927">text</mods:typeOfResource>
<mods:relatedItem id="98B848D82BCE424784687274B2F43F84" type="host">
<mods:titleInfo id="167D7C67A23A8A4886E77393A32FA40C">
<mods:title id="F296DDF2906E909E733B79D51E2FB413">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="C324AE85A87F6931FCB6F37EB3866ADB">
<mods:date id="2C23FBD89B661BB0A338741261835606">2024</mods:date>
<mods:detail id="BA01774C961264D6063D10A8036EFF93" type="pubDate">
<mods:number id="2F3E8BE506B70FDEA6478FEEC339DCDE">2024-08-29</mods:number>
</mods:detail>
<mods:detail id="748CED278E0E2CD072D341B5D1DB7E48" type="volume">
<mods:number id="024D8EC41945A86C5E422C3216C0BF85">72</mods:number>
</mods:detail>
<mods:extent id="E977336122C6D2C934601F4D5C1F04C4" unit="page">
<mods:start id="991686FA012F31F61B6E28A5BB1D2510">294</mods:start>
<mods:end id="1D0D079EFD1596347AE9D528B0007EB6">302</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D9D3D5909F69C59C7E5DA7684BEBD68C">journal article</mods:classification>
<mods:identifier id="B837E1FE4B953D3B0E102BFF6EDE0F5A" type="DOI">10.26107/RBZ-2024-0024</mods:identifier>
<mods:identifier id="4B79D9E5C1BA75FEE24B1FBAD1ED7ACE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="BB4098D79698D3FD498C2D5BEEF581AF" type="Zenodo-Dep">14683052</mods:identifier>
<mods:identifier id="2232E5A10804E6F454EEE31B8038A547" type="ZooBank">F4FB9765-D4A1-48AB-8845-EC63B5A7D794</mods:identifier>
</mods:mods>
<treatment id="03A88780FFDDFF9AFC22D035D0C153FD" ID-Zenodo-Dep="14683056" LSID="urn:lsid:plazi:treatment:03A88780FFDDFF9AFC22D035D0C153FD" httpUri="http://treatment.plazi.org/id/03A88780FFDDFF9AFC22D035D0C153FD" lastPageNumber="295" pageId="1" pageNumber="295">
<subSubSection id="C31B651DFFDDFF9AFC22D035D633533D" box="[993,1316,1814,1838]" pageId="1" pageNumber="295" type="nomenclature">
<paragraph id="8BBE3696FFDDFF9AFC22D035D633533D" blockId="1.[993,1316,1814,1838]" box="[993,1316,1814,1838]" pageId="1" pageNumber="295">
<heading id="D0F681FAFFDDFF9AFC22D035D633533D" bold="true" box="[993,1316,1814,1838]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="295" reason="2">
<emphasis id="B975EA84FFDDFF9AFC22D035D633533D" bold="true" box="[993,1316,1814,1838]" pageId="1" pageNumber="295">
Genus
<taxonomicName id="4C014D15FFDDFF9AFBF1D035D633533D" ID-CoL="4MNX" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[1074,1316,1814,1838]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis id="B975EA84FFDDFF9AFBF1D035D795533D" bold="true" box="[1074,1154,1814,1838]" italics="true" pageId="1" pageNumber="295">Gelotia</emphasis>
<bibRefCitation id="EF904B67FFDDFF9AFB48D035D633533D" author="Thorell T" box="[1163,1316,1814,1838]" pageId="1" pageNumber="295" pagination="132 - 172" refId="ref4521" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDDFF9AFC80D075D639537D" box="[835,1326,1878,1902]" pageId="1" pageNumber="295" type="type_taxon">
<paragraph id="8BBE3696FFDDFF9AFC80D075D639537D" blockId="1.[835,1326,1878,1902]" box="[835,1326,1878,1902]" pageId="1" pageNumber="295">
<emphasis id="B975EA84FFDDFF9AFC80D075D0CC537D" bold="true" box="[835,987,1878,1902]" pageId="1" pageNumber="295">
<typeStatus id="54BA8834FFDDFF9AFC80D075D06A537D" box="[835,893,1878,1902]" pageId="1" pageNumber="295">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C014D15FFDDFF9AFC27D075D63C537D" ID-CoL="3FK42" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[996,1323,1878,1902]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="295" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="B975EA84FFDDFF9AFC27D075D79E537D" box="[996,1161,1878,1902]" italics="true" pageId="1" pageNumber="295">Gelotia frenata</emphasis>
<bibRefCitation id="EF904B67FFDDFF9AFB57D075D63C537D" author="Thorell T" box="[1172,1323,1878,1902]" pageId="1" pageNumber="295" pagination="132 - 172" refId="ref4521" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDDFF9AFC80D0B5D0C153FD" pageId="1" pageNumber="295" type="distribution">
<paragraph id="8BBE3696FFDDFF9AFC80D0B5D0C153FD" blockId="1.[835,1474,1942,2030]" pageId="1" pageNumber="295">
<emphasis id="B975EA84FFDDFF9AFC80D0B5D0C153BD" bold="true" box="[835,982,1942,1966]" pageId="1" pageNumber="295">Distribution.</emphasis>
<collectingCountry id="F3167606FFDDFF9AFC1FD0B5D75C53BD" box="[988,1099,1942,1966]" name="Singapore" pageId="1" pageNumber="295">Singapore</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFB96D0B5D78053BD" box="[1109,1175,1942,1966]" name="China" pageId="1" pageNumber="295">China</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFB62D0B5D61253BD" box="[1185,1285,1942,1966]" name="Malaysia" pageId="1" pageNumber="295">Malaysia</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFAD3D0B5D66D53BD" box="[1296,1402,1942,1966]" name="Indonesia" pageId="1" pageNumber="295">Indonesia</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFA46D0B5D6A953BD" box="[1413,1470,1942,1966]" name="India" pageId="1" pageNumber="295">India</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFC80D095D0A053DD" box="[835,951,1974,1998]" name="Sri Lanka" pageId="1" pageNumber="295">Sri Lanka</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFC05D094D7B053DD" box="[966,1191,1974,1998]" name="Papua New Guinea" pageId="1" pageNumber="295">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F3167606FFDDFF9AFB75D095D63753DD" box="[1206,1312,1974,1998]" name="Australia" pageId="1" pageNumber="295">Australia</collectingCountry>
, and
<collectingCountry id="F3167606FFDDFF9AFAA1D095D6D653DD" box="[1378,1473,1974,1998]" name="Vietnam" pageId="1" pageNumber="295">Vietnam</collectingCountry>
(new record).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,237 @@
<document id="8FF0F58F74B389E316B94C4F138E6997" ID-DOI="10.26107/RBZ-2024-0024" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683052" ID-ZooBank="F4FB9765-D4A1-48AB-8845-EC63B5A7D794" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171910635" checkinUser="felipe" docAuthor="Hoang, Quang D., , Quoc T. Phan, Vo, &amp; Van M." docDate="2024" docId="03A88780FFDEFF98FEE1D284D67653E7" docLanguage="en" docName="RafflesBZool.72.294-302.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Gelotia onoi Hoang, , Vo &amp; M., 2024, new species" docType="treatment" docVersion="2" lastPageNumber="297" masterDocId="FF91FFF8FFDCFF9BFFC3D723D3175413" masterDocTitle="First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species" masterLastPageNumber="302" masterPageNumber="294" pageNumber="296" updateTime="1737175710851" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CD1E75A0FB0693C3276EB522A08AE1AE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="42F3075D53C9636A9F08068E6E1786A0">
<mods:title id="344D7885EE4576684925CBF221A7C897">First record of Gelotia Thorell, 1890 (Araneae: Salticidae) from Vietnam with description of a new species</mods:title>
</mods:titleInfo>
<mods:name id="0461AFC57E063F949337225426CEC385" type="personal">
<mods:role id="B5F39DA6AB7C8B24DA534D619B614FA4">
<mods:roleTerm id="3260CF2C647FF024C644D2473DA4F4C5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E33D52050794F314A42399E0423AB53">Hoang, Quang D.</mods:namePart>
</mods:name>
<mods:name id="3051514ECD6BC678C70CDBBDBBD00246" type="personal">
<mods:role id="D80EA131A40CB352D53165112674356E">
<mods:roleTerm id="7DA6CBD652B5521471D3126998A3FBD7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFF48EF767DF4CDBA331FF5103B15A27">, Quoc T. Phan</mods:namePart>
</mods:name>
<mods:name id="6166258DE65F126F2DBA42D9AD649134" type="personal">
<mods:role id="4D7CA85C16A81E869D7D017B94565543">
<mods:roleTerm id="BF61B73B79BD7503BB320CE7A54AD693">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7959701FCBA671FE7F3C24F372E3C844">Vo,</mods:namePart>
</mods:name>
<mods:name id="EC229720247FB8A1B8546F0316837EAB" type="personal">
<mods:role id="8480390650807B3BE6BF43843E48729C">
<mods:roleTerm id="948B001868978F0A4041036E3154F34E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A581E9318E6ABE9A6C3241E1C0F494DA">Van M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0E2B05616FE595CA4A633048B248F577">text</mods:typeOfResource>
<mods:relatedItem id="611A7A93DFA272BD02B9E594296FC7EA" type="host">
<mods:titleInfo id="A548040560CBFF5DAEADCB900478ADE9">
<mods:title id="52F1C98DFE4C572D29103746E1865149">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="BD9782EA267F058EF3D0F403C0E55748">
<mods:date id="EB9482AF8D8EB8BEAE905274D1143D48">2024</mods:date>
<mods:detail id="DAB1727F4A44FE244C591FDEF7E6EAC1" type="pubDate">
<mods:number id="D89B409C7A129CC011B0C3801956C851">2024-08-29</mods:number>
</mods:detail>
<mods:detail id="E533FD17BA6B2ACC67D2A286F1A67827" type="volume">
<mods:number id="0C0C2B12AED14BE6BCF8D53095DAE02A">72</mods:number>
</mods:detail>
<mods:extent id="419A58E29C2B9D4AFCC92A1152BF12A3" unit="page">
<mods:start id="97E9A814DB39B5FDA76A6E83AFFBC646">294</mods:start>
<mods:end id="8DCE4C888550C8EE30D5F5B7D1E07AE1">302</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D2FAF03D47A267641D5C61FB51FA1C11">journal article</mods:classification>
<mods:identifier id="1A615770BABD9A4D438BC626EB90EFBE" type="DOI">10.26107/RBZ-2024-0024</mods:identifier>
<mods:identifier id="7BC79733F5170940B6C74F3B069FE244" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="7485AEA150BB2E3D2165C8B540F7CE82" type="Zenodo-Dep">14683052</mods:identifier>
<mods:identifier id="CEC159408578A583C00B7812927C343E" type="ZooBank">F4FB9765-D4A1-48AB-8845-EC63B5A7D794</mods:identifier>
</mods:mods>
<treatment id="03A88780FFDEFF98FEE1D284D67653E7" LSID="urn:lsid:plazi:treatment:03A88780FFDEFF98FEE1D284D67653E7" httpUri="http://treatment.plazi.org/id/03A88780FFDEFF98FEE1D284D67653E7" lastPageId="3" lastPageNumber="297" pageId="2" pageNumber="296">
<subSubSection id="C31B651DFFDEFF99FEE1D284D15751AC" box="[290,576,1447,1471]" pageId="2" pageNumber="296" type="nomenclature">
<paragraph id="8BBE3696FFDEFF99FEE1D284D15751AC" blockId="2.[290,576,1447,1503]" box="[290,576,1447,1471]" pageId="2" pageNumber="296">
<heading id="D0F681FAFFDEFF99FEE1D284D15751AC" bold="true" box="[290,576,1447,1471]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="296" reason="2">
<emphasis id="B975EA84FFDEFF99FEE1D284D15751AC" bold="true" box="[290,576,1447,1471]" pageId="2" pageNumber="296">
<taxonomicName id="4C014D15FFDEFF99FEE1D284D2BC51AC" authorityName="Hoang &amp; &amp; Vo &amp; Van M." authorityYear="2024" box="[290,427,1447,1471]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="onoi" status="sp. nov.">
<emphasis id="B975EA84FFDEFF99FEE1D284D2BC51AC" bold="true" box="[290,427,1447,1471]" italics="true" pageId="2" pageNumber="296">Gelotia onoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657FFFFDEFF99FE78D284D15751AC" box="[443,576,1447,1471]" pageId="2" pageNumber="296" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDEFF99FEB6D2E4D2FB51CC" box="[373,492,1479,1503]" pageId="2" pageNumber="296" type="description">
<paragraph id="8BBE3696FFDEFF99FEB6D2E4D2FB51CC" blockId="2.[290,576,1447,1503]" box="[373,492,1479,1503]" pageId="2" pageNumber="296">
(
<figureCitation id="133A2A13FFDEFF99FEBED2E4D2F351CC" box="[381,484,1479,1503]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[114,149,1371,1393]" captionStartId-1="3.[136,171,1554,1576]" captionTargetBox-0="[113,1451,189,1355]" captionTargetBox-1="[136,1474,200,1525]" captionTargetId-0="figure-333@2.[113,1451,189,1355]" captionTargetId-1="figure-7@3.[136,1474,189,1538]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Habitus of Gelotia onoi, new species (female, holotype) in life. A, B, habitus, dorsal view; C, D, habitus, dorso-lateral view." captionText-1="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDEFF99FFB2D124D12F5293" pageId="2" pageNumber="296" type="materials_examined">
<paragraph id="8BBE3696FFDEFF99FFB2D124D12F5293" blockId="2.[113,752,1543,1664]" pageId="2" pageNumber="296">
<emphasis id="B975EA84FFDEFF99FFB2D124D275520C" bold="true" box="[113,354,1543,1567]" pageId="2" pageNumber="296">Material examined.</emphasis>
<materialsCitation id="3B693CCBFFDEFF99FEADD124D12F5293" collectingDate="2024-02-04" collectorName="Q. D. Hoang." country="Vietnam" elevation="140" latitude="12.9462" location="Yok Don National Park" longLatPrecision="7" longitude="107.7294" municipality="Buon Don Dist." pageId="2" pageNumber="296" specimenCount="1" stateProvince="Dak Lak" typeStatus="holotype">
<typeStatus id="54BA8834FFDEFF99FEADD124D2CB520C" box="[366,476,1543,1567]" pageId="2" pageNumber="296" type="holotype">Holotype</typeStatus>
: female (VNMN-ARA- SAL-480),
<collectingCountry id="F3167606FFDEFF99FF32D10BD2665253" box="[241,369,1576,1600]" name="Vietnam" pageId="2" pageNumber="296">VIETNAM</collectingCountry>
,
<collectingRegion id="49C5F874FFDEFF99FEBCD104D13F522C" box="[383,552,1575,1599]" country="Vietnam" name="Dak Lak" pageId="2" pageNumber="296">Dak Lak Prov.</collectingRegion>
,
<collectingMunicipality id="6BDAACECFFDEFF99FDF5D10BD1FB522C" box="[566,748,1575,1599]" pageId="2" pageNumber="296">Buon Don Dist.</collectingMunicipality>
,
<location id="8EDE604DFFDEFF99FFB2D164D293524C" LSID="urn:lsid:plazi:treatment:03A88780FFDEFF98FEE1D284D67653E7:8EDE604DFFDEFF99FFB2D164D293524C" box="[113,388,1607,1631]" country="Vietnam" latitude="12.9462" longLatPrecision="7" longitude="107.7294" municipality="Buon Don Dist." name="Yok Don National Park" pageId="2" pageNumber="296" stateProvince="Dak Lak">Yok Don National Park</location>
,
<geoCoordinate id="EE355051FFDEFF99FE50D164D119524C" box="[403,526,1607,1631]" degrees="12.9462" direction="north" orientation="latitude" pageId="2" pageNumber="296" precision="5" value="12.9462">12.9462°N</geoCoordinate>
,
<geoCoordinate id="EE355051FFDEFF99FDDED164D1B7524C" box="[541,672,1607,1631]" degrees="107.7294" direction="east" orientation="longitude" pageId="2" pageNumber="296" precision="5" value="107.7294">107.7294°E</geoCoordinate>
,
<elevation id="002CD1A5FFDEFF99FD71D164D3B3526C" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="296" unit="m" value="140.0">
<quantity id="4CF99B73FFDEFF99FD71D164D1E7524C" box="[690,752,1607,1631]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="296" unit="m" value="140.0">140m</quantity>
a.s.l.
</elevation>
,
<date id="FFBF1056FFDEFF99FF71D144D27E526C" box="[178,361,1639,1663]" pageId="2" pageNumber="296" value="2024-02-04">
<collectingDate id="EFFBE9BEFFDEFF99FF71D144D27E526C" box="[178,361,1639,1663]" pageId="2" pageNumber="296" value="2024-02-04">4 February 2024</collectingDate>
</date>
, coll.
<collectorName id="26F45340FFDEFF99FE6ED144D12F5293" box="[429,568,1639,1664]" pageId="2" pageNumber="296">Q.D. Hoang.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDEFF99FFB2D184D1CC52EC" pageId="2" pageNumber="296" type="etymology">
<paragraph id="8BBE3696FFDEFF99FFB2D184D1CC52EC" blockId="2.[113,752,1703,1791]" pageId="2" pageNumber="296">
<emphasis id="B975EA84FFDEFF99FFB2D184D3E052AC" bold="true" box="[113,247,1703,1727]" pageId="2" pageNumber="296">Etymology.</emphasis>
This specific epithet is given to honour Dr. Hirotsugu Ono (
<collectingCountry id="F3167606FFDEFF99FEE6D1E4D27052CC" box="[293,359,1735,1759]" name="Japan" pageId="2" pageNumber="296">Japan</collectingCountry>
), who has made great contributions to improving knowledge of the spider fauna in
<collectingCountry id="F3167606FFDEFF99FDBBD1C4D1C052EC" box="[632,727,1767,1791]" name="Vietnam" pageId="2" pageNumber="296">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDEFF99FFB2D004D7695253" pageId="2" pageNumber="296" type="diagnosis">
<paragraph id="8BBE3696FFDEFF99FFB2D004D7695253" blockId="2.[113,755,1831,2111]" lastBlockId="2.[813,1453,1448,1600]" pageId="2" pageNumber="296">
<emphasis id="B975EA84FFDEFF99FFB2D004D3F1532C" bold="true" box="[113,230,1831,1855]" pageId="2" pageNumber="296">Diagnosis.</emphasis>
The female of
<taxonomicName id="4C014D15FFDEFF99FE50D004D10F532C" authorityName="Hoang &amp; &amp; Vo &amp; Van M." authorityYear="2024" box="[403,536,1831,1855]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="onoi" status="sp. nov.">
<emphasis id="B975EA84FFDEFF99FE50D004D10F532C" box="[403,536,1831,1855]" italics="true" pageId="2" pageNumber="296">Gelotia onoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657FFFFDEFF99FDEBD00BD1BD532C" box="[552,682,1831,1855]" pageId="2" pageNumber="296" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="133A2A13FFDEFF99FD78D004D38B534C" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[114,149,1371,1393]" captionStartId-1="3.[136,171,1554,1576]" captionTargetBox-0="[113,1451,189,1355]" captionTargetBox-1="[136,1474,200,1525]" captionTargetId-0="figure-333@2.[113,1451,189,1355]" captionTargetId-1="figure-7@3.[136,1474,189,1538]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Habitus of Gelotia onoi, new species (female, holotype) in life. A, B, habitus, dorsal view; C, D, habitus, dorso-lateral view." captionText-1="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Figs. 12</figureCitation>
) resembles those of the
<typeStatus id="54BA8834FFDEFF99FE5FD06BD2DD5373" box="[412,458,1864,1888]" pageId="2" pageNumber="296">type</typeStatus>
species,
<taxonomicName id="4C014D15FFDEFF99FDE4D06BD3B1536C" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="B975EA84FFDEFF99FDE4D06BD183534C" box="[551,660,1863,1887]" italics="true" pageId="2" pageNumber="296">G. frenata</emphasis>
<bibRefCitation id="EF904B67FFDEFF99FD58D064D3B1536C" author="Thorell T" pageId="2" pageNumber="296" pagination="132 - 172" refId="ref4521" refString="Thorell T (1890) Diagnoses aranearum aliquot novarum in Indo- Malesia inventarum. Annali del Museo Civico di Storia Naturale di Genova, 30: 132 - 172." type="journal article" year="1890">Thorell, 1890</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C014D15FFDEFF99FF1CD04BD2EA536C" authority="Wang &amp; Li, 2020" authorityName="Wang &amp; Li" authorityYear="2020" box="[223,509,1895,1920]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="liuae">
<emphasis id="B975EA84FFDEFF99FF1CD04BD221536C" box="[223,310,1895,1919]" italics="true" pageId="2" pageNumber="296">G. liuae</emphasis>
<bibRefCitation id="EF904B67FFDEFF99FEFED04BD2EA536C" author="Wang C &amp; Li SQ" box="[317,509,1895,1920]" pageId="2" pageNumber="296" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">
Wang &amp; Li,
<quantity id="4CF99B73FFDEFF99FE0BD044D2EA536C" box="[456,509,1895,1919]" metricMagnitude="1" metricUnit="m" metricValue="5.1308" pageId="2" pageNumber="296" unit="in" value="2020.0">2020</quantity>
</bibRefCitation>
</taxonomicName>
in having the epigyne with a fold and rounded window, but differs from two latter species by the following characteristics: the epigynal window located in the near middle in the new species (
<figureCitation id="133A2A13FFDEFF99FD04D0E4D38453EC" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2C</figureCitation>
) while anterior in both
<taxonomicName id="4C014D15FFDEFF99FE54D0CBD11053EC" authorityName="Thorell" authorityYear="1890" box="[407,519,2023,2047]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="B975EA84FFDEFF99FE54D0CBD11053EC" box="[407,519,2023,2047]" italics="true" pageId="2" pageNumber="296">G. frenata</emphasis>
</taxonomicName>
(fig.
<quantity id="4CF99B73FFDEFF99FD81D0C4D17B53EC" box="[578,620,2023,2047]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="296" unit="in" value="5.0">5 in</quantity>
<bibRefCitation id="EF904B67FFDEFF99FDB7D0C4D3BE5C0C" author="Proszynski J" pageId="2" pageNumber="296" pagination="12 - 20" refId="ref4438" refString="Proszynski J (1969) Redescriptions of type-species of genera of Salticidae (Araneida). III - Remarks on the genera Gelotia Thorell, 1890 and Policha Thorell, 1892. Annali del Museo Civico di Storia Naturale Giacomo Doria, 77: 12 - 20." type="journal article" year="1969">Prószyński, 1969</bibRefCitation>
) and
<taxonomicName id="4C014D15FFDEFF99FF2DDF2BD25D5C0C" authorityName="Wang &amp; Li" authorityYear="2020" box="[238,330,2055,2079]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="liuae">
<emphasis id="B975EA84FFDEFF99FF2DDF2BD25D5C0C" box="[238,330,2055,2079]" italics="true" pageId="2" pageNumber="296">G. liuae</emphasis>
</taxonomicName>
(cf.
<figureCitation id="133A2A13FFDEFF99FE47DF24D2CD5C0C" box="[388,474,2055,2079]" captionStart="Fig" captionStartId="5.[136,171,1545,1567]" captionTargetBox="[143,1474,195,1513]" captionTargetId="figure-210@5.[136,1474,189,1529]" captionTargetPageId="5" captionText="Fig. 4. Female of Gelotia liuae Wang &amp; Li, 2020. A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 2 mm, B, 1 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 4C</figureCitation>
and fig. 8A in
<bibRefCitation id="EF904B67FFDEFF99FD53DF2BD3C05C2C" author="Wang C &amp; Li SQ" pageId="2" pageNumber="296" pagination="25 - 57" refId="ref4598" refString="Wang C &amp; Li SQ (2020) On eight species of jumping spiders from Xishuangbanna, Yunnan, China (Araneae, Salticidae). ZooKeys, 909: 25 - 57." type="journal article" year="2020">Wang &amp; Li, 2020</bibRefCitation>
), copulatory duct significantly longer than the length of spermathecae (
<figureCitation id="133A2A13FFDEFF99FBF6D28BD79D51D3" box="[1077,1162,1448,1472]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2D</figureCitation>
) (shorter in other species), spermathecae almost rounded, relatively very small (
<figureCitation id="133A2A13FFDEFF99FA42D2EBD0595213" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2D</figureCitation>
) in comparision with
<taxonomicName id="4C014D15FFDEFF99FBFCD2CBD7BA5213" authorityName="Thorell" authorityYear="1890" box="[1087,1197,1512,1536]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="frenata">
<emphasis id="B975EA84FFDEFF99FBFCD2CBD7BA5213" box="[1087,1197,1512,1536]" italics="true" pageId="2" pageNumber="296">G. frenata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C014D15FFDEFF99FB27D2CBD62D5213" authorityName="Wang &amp; Li" authorityYear="2020" box="[1252,1338,1512,1536]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="species" species="liuae">
<emphasis id="B975EA84FFDEFF99FB27D2CBD62D5213" box="[1252,1338,1512,1536]" italics="true" pageId="2" pageNumber="296">G. liuae</emphasis>
</taxonomicName>
as well as all other members of the genus
<taxonomicName id="4C014D15FFDEFF99FB56D12BD7F15233" authorityName="Thorell" authorityYear="1890" box="[1173,1254,1544,1568]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B975EA84FFDEFF99FB56D12BD7F15233" box="[1173,1254,1544,1568]" italics="true" pageId="2" pageNumber="296">Gelotia</emphasis>
</taxonomicName>
(see comparative illustrations in
<bibRefCitation id="EF904B67FFDEFF99FC12D10BD7645253" author="Metzner H" box="[977,1139,1576,1600]" pageId="2" pageNumber="296" refId="ref4405" refString="Metzner H (2024) Jumping spiders (Arachnida: Araneae: Salticidae) of the world. https: // www. jumping-spiders. com (Accessed 3 April 2024)." type="url" year="2024">Metzner, 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDEFF98FCEED14BD0EC53A7" lastPageId="3" lastPageNumber="297" pageId="2" pageNumber="296" type="description">
<paragraph id="8BBE3696FFDEFF98FCEED14BD61E52A7" blockId="2.[811,1453,1640,2112]" lastBlockId="3.[834,1473,1660,1716]" lastPageId="3" lastPageNumber="297" pageId="2" pageNumber="296">
<emphasis id="B975EA84FFDEFF99FCEED14BD0AF5293" bold="true" box="[813,952,1640,1664]" pageId="2" pageNumber="296">Description.</emphasis>
Female
<typeStatus id="54BA8834FFDEFF99FBD7D14BD7625293" box="[1044,1141,1640,1664]" pageId="2" pageNumber="296" type="holotype">holotype</typeStatus>
. Measurements: Total length 4.63; carapace length 2.31, width 1.72; abdomen length 2.32, width 1.34; clypeus height 0.08. Carapace brown, clothed with dense iridescent setae, lateral sides dark with yellowish belt (
<figureCitation id="133A2A13FFDEFF99FC19D1CBD7495313" box="[986,1118,1768,1792]" captionStart="Fig" captionStartId="2.[114,149,1371,1393]" captionTargetBox="[113,1451,189,1355]" captionTargetId="figure-333@2.[113,1451,189,1355]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Gelotia onoi, new species (female, holotype) in life. A, B, habitus, dorsal view; C, D, habitus, dorso-lateral view." pageId="2" pageNumber="296">Figs. 1AD</figureCitation>
,
<figureCitation id="133A2A13FFDEFF99FBAED1CBD7D15313" box="[1133,1222,1768,1792]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2A</figureCitation>
); fovea longitudinal. Anterior eyes surrounded by yellow-orange orbital setae (
<figureCitation id="133A2A13FFDEFF99FCF6D00BD0985353" box="[821,911,1832,1856]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2B</figureCitation>
). Clypeus dark brown, narrowest medially with some long setae (
<figureCitation id="133A2A13FFDEFF99FC28D06BD7565373" box="[1003,1089,1864,1888]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2B</figureCitation>
). Sternum yellowish brown, with darker margin and sparse long setae. Endites and labium brownish yellow. Chelicerae brownish yellow (
<figureCitation id="133A2A13FFDEFF99FA87D0ABD68A53B3" box="[1348,1437,1928,1952]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2B</figureCitation>
); promargin with three teeth, and five retromarginal teeth. Abdomen brown, covered with dense iridescent setae and with a large transverse stripes in posterior half of dorsum (
<figureCitation id="133A2A13FFDEFF99FAB7D0CBD0655C33" captionStart="Fig" captionStartId="2.[114,149,1371,1393]" captionTargetBox="[113,1451,189,1355]" captionTargetId="figure-333@2.[113,1451,189,1355]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Gelotia onoi, new species (female, holotype) in life. A, B, habitus, dorsal view; C, D, habitus, dorso-lateral view." pageId="2" pageNumber="296">Figs. 1AD</figureCitation>
,
<figureCitation id="133A2A13FFDEFF99FC43DF2BD0CF5C33" box="[896,984,2056,2080]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="2" pageNumber="296">Fig. 2A</figureCitation>
); venter brownish yellow. Lateral sides of abdomen dark brown covered with yellow setae. Spinnerets very short, dark brown. Legs brown, covered with densely pale setae (
<figureCitation id="133A2A13FFDFFF98FEC4D1BFD27752A7" box="[263,352,1692,1716]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="3" pageNumber="297">Fig. 2A</figureCitation>
). Width of eye rows: anterior eye row 1.67; posterior medial eye row 1.53; posterior lateral eye row 1.64. Distance between ALE-PME 0.51; ALE-PLE 1.09. Diameter of eyes: AME 0.56; ALE 0.32; PME 0.18; PLE 0.29. Length of leg segments: I 1.46 + 0.82 + 1.12 + 0.95 + 0.52 (4.87); II 1.36 + 0.75 + 0.92 + 0.89 + 0.49 (4.41); III 1.29 + 0.63 + 0.91 + 0.99 + 0.43 (4.25); IV 1.73 + 0.74 + 1.40 + 1.62 + 0.48 (5.97). Leg formula IVIIIIII. Leg spination: I: Fm d 114; Pt pr 1; Tb v 222ap, pt 11; Mt v 22. II: Fm d 114; Pt pr 1, rt 1; Tb v 122ap, pt 11; Mt v 22. III: Fm d 114; Pt pr 1, rt 1; Tb d 11, pt 011ap, v 102ap, rt 011ap; Mt d 122ap, v 022ap., and IV: Fm d 114; Pt pr 1, rt 1; Tb d 11, pt 11, rt 11; Mt d 222ap, rt 1ap, v 102ap, pr 1ap.
</paragraph>
<caption id="DF7E661EFFDFFF98FF4BD131D0575257" pageId="3" pageNumber="297" startId="3.[136,171,1554,1576]" targetBox="[136,1474,200,1525]" targetPageId="3" targetType="figure">
<paragraph id="8BBE3696FFDFFF98FF4BD131D0575257" blockId="3.[136,1474,1554,1604]" pageId="3" pageNumber="297">
Fig. 2. Female of
<taxonomicName id="4C014D15FFDFFF98FEF9D131D2A45234" authorityName="Hoang &amp; &amp; Vo &amp; Van M." authorityYear="2024" box="[314,435,1554,1575]" class="Arachnida" family="Salticidae" genus="Gelotia" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="297" phylum="Arthropoda" rank="species" species="onoi" status="sp. nov.">
<emphasis id="B975EA84FFDFFF98FEF9D131D2A45234" box="[314,435,1554,1575]" italics="true" pageId="3" pageNumber="297">Gelotia onoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A24657FFFFDFFF98FE03D131D122523B" box="[448,565,1554,1576]" pageId="3" pageNumber="297" rank="species">new species</taxonomicNameLabel>
(holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BBE3696FFDFFF98FC80D1FFD6A75367" blockId="3.[835,1474,1756,1908]" pageId="3" pageNumber="297">
Epigyne (
<figureCitation id="133A2A13FFDFFF98FC68D1FFD73552E7" box="[939,1058,1756,1780]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="3" pageNumber="297">Fig. 2CD</figureCitation>
): longer than wide, the fold of epigyne wider than the window, and the rim of epigynal window almost rounded (
<figureCitation id="133A2A13FFDFFF98FBC0D03FD74A5327" box="[1027,1117,1820,1844]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="3" pageNumber="297">Fig. 2C</figureCitation>
). Copulatory ducts fused, long, widest medially (
<figureCitation id="133A2A13FFDFFF98FBC4D01FD7745347" box="[1031,1123,1852,1876]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="3" pageNumber="297">Fig. 2D</figureCitation>
). Spermathecae fused, rounded and significantly shorter than copulatory ducts (
<figureCitation id="133A2A13FFDFFF98FA8FD07FD6B35367" box="[1356,1444,1884,1908]" captionStart="Fig" captionStartId="3.[136,171,1554,1576]" captionTargetBox="[136,1474,200,1525]" captionTargetId="figure-7@3.[136,1474,189,1538]" captionTargetPageId="3" captionText="Fig. 2. Female of Gelotia onoi, new species (holotype). A, habitus, dorsal view; B, carapace, anterior view; C, epigyne, ventral view; D, epigyne, dorsal view. Scale bars: A, 1 mm, B, 0.5 mm, C, D, 0.2 mm." pageId="3" pageNumber="297">Fig. 2D</figureCitation>
).
</paragraph>
<paragraph id="8BBE3696FFDFFF98FC80D0BFD0EC53A7" blockId="3.[835,1019,1948,1972]" box="[835,1019,1948,1972]" pageId="3" pageNumber="297">Male: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C31B651DFFDFFF98FC80D0F8D67653E7" box="[835,1377,2011,2036]" pageId="3" pageNumber="297" type="distribution">
<paragraph id="8BBE3696FFDFFF98FC80D0F8D67653E7" blockId="3.[835,1377,2011,2036]" box="[835,1377,2011,2036]" pageId="3" pageNumber="297">
<emphasis id="B975EA84FFDFFF98FC80D0F8D0C153E0" bold="true" box="[835,982,2011,2035]" pageId="3" pageNumber="297">Distribution.</emphasis>
Known only from the
<typeStatus id="54BA8834FFDFFF98FB10D0FFD61653E7" box="[1235,1281,2012,2036]" pageId="3" pageNumber="297">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,201 @@
<document id="734CBE1CAA58FC084F3AD9608EA829E1" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF903B231681FDD2FDE7F97F" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris vittulatus Yasunaga 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="416" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="416" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="50BAE99376CAD92F70983173A45A14ED" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="65B1FD5507DEE6F7D0E82CA101722F15">
<mods:title id="7134B714CA8816124D6BBE15719B7357">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="62F1DF314D01704D0F9E199A50BB07A1" type="personal">
<mods:role id="10A2EFE1619ECFA7F58C1843891B34D0">
<mods:roleTerm id="A4F4FB3F240A8AE20A5BB7C49A8FEB14">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="528A94AFAF5F6EC5186DF98676996372">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="C3BBF848DFA6C5E9838853EF0BD3BB75">text</mods:typeOfResource>
<mods:relatedItem id="7F5ADA8F99CC2C14971BC5E786A7F7AD" type="host">
<mods:titleInfo id="50D42D2427F5052B7A340EBC96BA9EAC">
<mods:title id="633B408A72E461AC7989F0C34EA97197">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="AB957B14C205FC345E318CE7DC0F66D6">
<mods:date id="13D6B8473E9CECC46030ADCDCEB76ED4">2024</mods:date>
<mods:detail id="A66DBF6BCEC5F1E7652F7359D73FE2A5" type="pubDate">
<mods:number id="2ED7505C901190C784780F21425E2D9B">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="8DE9BDF2B0848AF54F94CD6BC78087D6" type="volume">
<mods:number id="F69B4CDAD2D92F5D401D558CD7EAF917">72</mods:number>
</mods:detail>
<mods:extent id="B32B464E0BBEAED448C00335A77A3F0B" unit="page">
<mods:start id="2677EBFE023BDD6117E17E1D646D1E7D">371</mods:start>
<mods:end id="8B4416E02B3AFAAD031DB4B2ED9CE782">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8BECE9E7B9E38037F6880086B1100A02">journal article</mods:classification>
<mods:identifier id="AB08FBC8658707113768A9A922D0D81B" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="B4C1508FA730AC70564968F0245AB7E4" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="3B754619133765CDCB823629AD086440" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="07F8919714615AD148B8405F6717A85E" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF903B231681FDD2FDE7F97F" ID-DOI="http://doi.org/10.5281/zenodo.14683048" ID-Zenodo-Dep="14683048" LSID="urn:lsid:plazi:treatment:03B2879FFF903B231681FDD2FDE7F97F" httpUri="http://treatment.plazi.org/id/03B2879FFF903B231681FDD2FDE7F97F" lastPageNumber="416" pageId="45" pageNumber="416">
<subSubSection id="C3016502FF903B231681FDD2FE43FF21" pageId="45" pageNumber="416" type="nomenclature">
<paragraph id="8BA43689FF903B231681FDD2FE43FF21" blockId="45.[180,685,188,277]" pageId="45" pageNumber="416">
<heading id="D0EC81E5FF903B231681FDD2FE43FF21" bold="true" centered="true" fontSize="10" level="2" pageId="45" pageNumber="416" reason="2">
<emphasis id="B96FEA9BFF903B231681FDD2FE43FF21" bold="true" pageId="45" pageNumber="416">
<taxonomicName id="4C1B4D0AFF903B231681FDD2FDC2FF01" authority="(Poppius, 1914)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[180,630,188,213]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="vittulatus" status="comb. nov.">
<emphasis id="B96FEA9BFF903B231681FDD2FE0FFF00" bold="true" box="[180,443,188,212]" italics="true" pageId="45" pageNumber="416">Paralygocoris vittulatus</emphasis>
(
<bibRefCitation id="EF8A4B78FF903B2317FEFDD3FDDBFF01" author="Poppius B" box="[459,623,189,213]" pageId="45" pageNumber="416" pagination="337 - 398" refId="ref24173" refString="Poppius B (1914) Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici, 12: 337 - 398." type="journal article" year="1914">Poppius, 1914</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A25C57E0FF903B23144AFDD3FE43FF21" pageId="45" pageNumber="416" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231733FD93FDEEFEC1" box="[262,602,253,277]" pageId="45" pageNumber="416" type="description">
<paragraph id="8BA43689FF903B231733FD93FDEEFEC1" blockId="45.[180,685,188,277]" box="[262,602,253,277]" pageId="45" pageNumber="416">
(
<figureCitation id="13202A0CFF903B23173BFD93FED2FEC1" box="[270,358,253,277]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="45" pageNumber="416">Figs. 12</figureCitation>
MO, 17AE, 34AI)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644FC52FEAAFE5F" pageId="45" pageNumber="416" type="reference_group">
<paragraph id="8BA43689FF903B231644FC52FE6CFE85" blockId="45.[113,750,315,395]" box="[113,472,315,337]" pageId="45" pageNumber="416">
<treatmentCitationGroup id="AB0B11A7FF903B231644FC52FE6CFE85" box="[113,472,315,337]" pageId="45" pageNumber="416">
<treatmentCitation id="0ABA1098FF903B231644FC52FE60FE85" author="Poppius B" box="[113,468,315,337]" page="361" pageId="45" pageNumber="416" year="1914">
<taxonomicName id="4C1B4D0AFF903B231644FC52FE60FE85" authority="Poppius, 1914: 361" authorityName="Poppius" authorityPageNumber="361" authorityYear="1914" box="[113,468,315,337]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFF903B231644FC52FEBFFE84" box="[113,267,315,337]" italics="true" pageId="45" pageNumber="416">Lygus vittulatus</emphasis>
<bibRefCitation id="EF8A4B78FF903B231726FC55FE60FE85" author="Poppius B" box="[275,468,315,337]" pageId="45" pageNumber="416" pagination="337 - 398" refId="ref24173" refString="Poppius B (1914) Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici, 12: 337 - 398." type="journal article" year="1914">Poppius, 1914: 361</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA43689FF903B231644FC37FEAAFE5F" blockId="45.[113,750,315,395]" pageId="45" pageNumber="416">
<treatmentCitationGroup id="AB0B11A7FF903B231644FC37FEAAFE5F" pageId="45" pageNumber="416">
<taxonomicName id="4C1B4D0AFF903B231644FC37FEE4FEB9" ID-CoL="3WPZ6" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[113,336,344,366]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFF903B231644FC37FEE4FEB9" box="[113,336,344,366]" italics="true" pageId="45" pageNumber="416">Lygocoris vittulatus</emphasis>
</taxonomicName>
:
<treatmentCitation id="0ABA1098FF903B231756FC36FD1BFEB9" author="Schwartz MD &amp; Cherot F" box="[355,687,344,366]" page="10" pageId="45" pageNumber="416" year="2005">
<bibRefCitation id="EF8A4B78FF903B231756FC36FD1BFEB9" author="Schwartz MD &amp; Cherot F" box="[355,687,344,366]" pageId="45" pageNumber="416" pagination="1 - 24" refId="ref24346" refString="Schwartz MD &amp; Cherot F (2005) Miscellanea Miridologica (Insecta: Heteroptera). Zootaxa, 814: 1 - 24." type="journal article" year="2005">Schwartz &amp; Chérot, 2005: 10</bibRefCitation>
</treatmentCitation>
(new combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644FCDDFE7FFD9F" pageId="45" pageNumber="416" type="materials_examined">
<paragraph id="8BA43689FF903B231644FCDDFE7FFD9F" blockId="45.[113,752,435,587]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644FCDDFEEDFE1F" bold="true" box="[113,345,435,459]" pageId="45" pageNumber="416">Material examined.</emphasis>
<materialsCitation id="3B733CD4FF903B231751FCDDFE72FD9F" collectingDate="1999-12-22" collectingDateMax="1999-12-25" collectingDateMin="1999-12-22" collectionCode="AMNH, TYCN, ZRC" collectorName="M. Takai" country="Indonesia" latitude="-5.3166666" location="Lompobattan" longLatPrecision="1305" longitude="119.933334" pageId="45" pageNumber="416" specimenCode="PBI 00378789" specimenCount="10" specimenCount-female="4" specimenCount-male="6" stateProvince="Sulawesi Selatan">
<emphasis id="B96FEA9BFF903B231751FCDDFDB3FE1F" bold="true" box="[356,519,435,459]" pageId="45" pageNumber="416">
<collectingCountry id="F30C7619FF903B231751FCDDFDB7FE1F" box="[356,515,435,459]" name="Indonesia" pageId="45" pageNumber="416">INDONESIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D1DFD00FF903B231427FCDDFDD9FE1F" box="[530,621,435,459]" count="5" pageId="45" pageNumber="416" type="male">5 males</specimenCount>
,
<specimenCount id="9D1DFD00FF903B23144EFCDDFD58FE1F" box="[635,748,435,459]" count="4" pageId="45" pageNumber="416" type="female">4 females</specimenCount>
,
<collectingRegion id="49DFF86BFF903B231644FCBDFE90FE3F" box="[113,292,467,491]" country="Indonesia" name="Sulawesi Selatan" pageId="45" pageNumber="416">South Sulawesi</collectingRegion>
,
<location id="8EC46052FF903B231701FCBDFE7FFE3F" LSID="urn:lsid:plazi:treatment:03B2879FFF903B231681FDD2FDE7F97F:8EC46052FF903B231701FCBDFE7FFE3F" box="[308,459,467,491]" country="Indonesia" latitude="-5.3166666" longLatPrecision="1305" longitude="119.933334" name="Lompobattan" pageId="45" pageNumber="416" stateProvince="Sulawesi Selatan">Lompobattan</location>
,
<locationDeviation id="9A162FABFF903B2317EFFCBDFD5BFE3F" box="[474,751,467,491]" location="Makassar" pageId="45" pageNumber="416">
<quantity id="4CE39B6CFF903B2317EFFCBDFD89FE3F" box="[474,573,467,491]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="45" pageNumber="416" unit="mi" value="30.0">30 miles</quantity>
E of Makassar
</locationDeviation>
(Ujung Pandang),
<geoCoordinate id="EE2F504EFF903B231779FC9CFE2FFDDF" box="[332,411,498,523]" degrees="5" direction="south" minutes="19" orientation="latitude" pageId="45" pageNumber="416" precision="925" value="-5.3166666">5°19S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF903B231799FC9CFDADFDDF" box="[428,537,498,523]" degrees="119" direction="east" minutes="56" orientation="longitude" pageId="45" pageNumber="416" precision="925" value="119.933334">119°56E</geoCoordinate>
,
<collectorName id="26EE535FFF903B23141FFC9AFD2DFDDF" box="[554,665,499,523]" pageId="45" pageNumber="416">M. Takai</collectorName>
,
<date id="FFA51049FF903B23149CFC9DFE95FDFF" pageId="45" pageNumber="416" value="1999-12-22" valueMax="1999-12-25" valueMin="1999-12-22">
<collectingDate id="EFE1E9A1FF903B23149CFC9DFE95FDFF" pageId="45" pageNumber="416" value="1999-12-22" valueMax="1999-12-25" valueMin="1999-12-22">2225 December 1999</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF903B231701FF7DFE3FFDFF" box="[308,395,531,555]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="45" pageNumber="416">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFF903B2317AFFF7DFE53FDFF" box="[410,487,531,555]" pageId="45" pageNumber="416">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFF903B2317C0FF7DFD9FFDFF" box="[501,555,531,555]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="45" pageNumber="416">ZRC</collectionCode>
) (
<specimenCount id="9D1DFD00FF903B23147DFF7DFD27FDFF" box="[584,659,531,555]" count="1" pageId="45" pageNumber="416" type="male">1 male</specimenCount>
bearing USIs,
<collectionCode id="ED0AAE4CFF903B23168AFF5DFEA5FD9F" box="[191,273,563,587]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="45" pageNumber="416">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FF903B23172BFF5DFE74FD9F" box="[286,448,563,587]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="45" pageNumber="416">PBI 00378789</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644FF1DFE1DFCFF" pageId="45" pageNumber="416" type="diagnosis">
<paragraph id="8BA43689FF903B231644FF1DFE1DFCFF" blockId="45.[113,752,627,811]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644FF1DFF51FD5F" bold="true" box="[113,229,627,651]" pageId="45" pageNumber="416">Diagnosis.</emphasis>
This species is recognised readily by its relatively large size and unique colour pattern on dorsum (
<figureCitation id="13202A0CFF903B2314B6FFFDFF33FD1F" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="45" pageNumber="416">Fig. 12M, O</figureCitation>
), in addition to the characters shown in the key. The male and female genitalic structures are herein described for the first time. Detailed description of external characters was provided by
<bibRefCitation id="EF8A4B78FF903B2316C9FE7DFE11FCFF" author="Poppius B" box="[252,421,787,811]" pageId="45" pageNumber="416" pagination="337 - 398" refId="ref24173" refString="Poppius B (1914) Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici, 12: 337 - 398." type="journal article" year="1914">Poppius (1914)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644FE3DFE05FB7F" pageId="45" pageNumber="416" type="description">
<paragraph id="8BA43689FF903B231644FE3DFE6CFBBF" blockId="45.[113,752,851,1131]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644FE3DFF4BFCBF" bold="true" box="[113,255,851,875]" pageId="45" pageNumber="416">Description.</emphasis>
Male genitalia (
<figureCitation id="13202A0CFF903B231789FE3DFDF9FCBF" box="[444,589,851,875]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="45" pageNumber="416">Figs. 17AE</figureCitation>
,
<figureCitation id="13202A0CFF903B23146EFE3DFD1FFCBF" box="[603,683,851,875]" captionStart="Fig" captionStartId="49.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@49.[113,1451,251,1929]" captionTargetPageId="49" captionText="Fig. 34. Scanning electron micrographs of Paralygocoris vittulatus (AI), Sabactiopus schuhi, new species (JL) and S. sauteri (MO). A, head and pronotum, left lateral view; B, scutellum and hemelytron, left lateral view; C, metathoracic scent efferent system; D, metatarsus; E, pygophore, dorsal view; F, vesica; G, sclerotised ring; H, I, posterior wall; J, left lateral habitus; K, anterior body, dorsal view; L, O, thoracic pleura, left lateral view; M, anterior body, left lateral view; N, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683036" httpUri="https://zenodo.org/record/14683036/files/figure.png" pageId="45" pageNumber="416">34E, F</figureCitation>
): Left paramere L-shaped, with apically inflated sensory lobe and sharply pointed apex of hypophysis; hypophysis of right paramere sharply pointed; vesical spicule elongate, stout, L-shaped. Female genitalia (
<figureCitation id="13202A0CFF903B231793FEBDFDA4FC3F" box="[422,528,979,1003]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="45" pageNumber="416">Figs. 17F</figureCitation>
,
<figureCitation id="13202A0CFF903B231428FEBDFDD2FC3F" box="[541,614,979,1003]" captionStart="Fig" captionStartId="49.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@49.[113,1451,251,1929]" captionTargetPageId="49" captionText="Fig. 34. Scanning electron micrographs of Paralygocoris vittulatus (AI), Sabactiopus schuhi, new species (JL) and S. sauteri (MO). A, head and pronotum, left lateral view; B, scutellum and hemelytron, left lateral view; C, metathoracic scent efferent system; D, metatarsus; E, pygophore, dorsal view; F, vesica; G, sclerotised ring; H, I, posterior wall; J, left lateral habitus; K, anterior body, dorsal view; L, O, thoracic pleura, left lateral view; M, anterior body, left lateral view; N, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683036" httpUri="https://zenodo.org/record/14683036/files/figure.png" pageId="45" pageNumber="416">34GI</figureCitation>
): Sclerotised rings elongate ovoid, narrow, relatively contiguous to each other mesally; dorsal structure basally fused with interramal lobes; and both of interramal lobe and dorsal structure spinulate along posterior margin.
</paragraph>
<paragraph id="8BA43689FF903B231644F9FDFE05FB7F" blockId="45.[113,433,1171,1195]" box="[113,433,1171,1195]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644F9FDFEABFB7F" bold="true" box="[113,287,1171,1195]" pageId="45" pageNumber="416">Measurements.</emphasis>
See
<tableCitation id="C6990332FF903B231762F9FDFE1EFB7F" box="[343,426,1171,1195]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="45" pageNumber="416">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644F9BDFEC3FA9F" pageId="45" pageNumber="416" type="biology_ecology">
<paragraph id="8BA43689FF903B231644F9BDFEC3FA9F" blockId="45.[113,752,1235,1355]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644F9BDFF7BFB3F" bold="true" box="[113,207,1235,1259]" pageId="45" pageNumber="416">Biology.</emphasis>
Unknown. According to the collector (Takai M, pers. comm.), more than a few individuals of this mirid were captured by sweep-netting inflorescences of broadleaf trees around cultivated fields.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231644F81DFDE7F97F" pageId="45" pageNumber="416" type="discussion">
<paragraph id="8BA43689FF903B231644F81DFDE7F97F" blockId="45.[113,752,1395,1707]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231644F81DFF54FA5F" bold="true" box="[113,224,1395,1419]" pageId="45" pageNumber="416">Remarks.</emphasis>
This taxon has been known only by the female
<typeStatus id="54A0882BFF903B231644F8FDFF62FA7F" box="[113,214,1427,1451]" pageId="45" pageNumber="416" type="holotype">holotype</typeStatus>
from S.
<collectingRegion id="49DFF86BFF903B231772F8FDFE16FA7F" box="[327,418,1427,1451]" country="Indonesia" name="Sulawesi Utara" pageId="45" pageNumber="416">Celebes</collectingRegion>
[=
<collectingRegion id="49DFF86BFF903B2317E6F8FDFD8AFA7F" box="[467,574,1427,1451]" country="Indonesia" name="Sulawesi Utara" pageId="45" pageNumber="416">Sulawesi</collectingRegion>
] (Bua-Kraeng) (
<bibRefCitation id="EF8A4B78FF903B23164CF8DDFE05FA1F" author="Schwartz MD &amp; Cherot F" box="[121,433,1459,1483]" pageId="45" pageNumber="416" pagination="1 - 24" refId="ref24346" refString="Schwartz MD &amp; Cherot F (2005) Miscellanea Miridologica (Insecta: Heteroptera). Zootaxa, 814: 1 - 24." type="journal article" year="2005">Schwartz &amp; Chérot, 2005</bibRefCitation>
). During sorting a number of mirine specimens collected by Mr. M. Takai (
<collectingRegion id="49DFF86BFF903B231493F8BDFD5BFA3F" box="[678,751,1491,1515]" country="Japan" name="Ehime" pageId="45" pageNumber="416">Ehime</collectingRegion>
University Expedition in 1999), several samples were found to match Poppius species. Although
<bibRefCitation id="EF8A4B78FF903B23142DFB7DFF0EF99F" author="Schwartz MD &amp; Cherot F" pageId="45" pageNumber="416" pagination="1 - 24" refId="ref24346" refString="Schwartz MD &amp; Cherot F (2005) Miscellanea Miridologica (Insecta: Heteroptera). Zootaxa, 814: 1 - 24." type="journal article" year="2005">Schwartz &amp; Chérot (2005)</bibRefCitation>
placed
<taxonomicName id="4C1B4D0AFF903B231722FB5AFE7EF99F" authorityName="Poppius" authorityYear="1914" box="[279,458,1587,1611]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFF903B231722FB5AFE7EF99F" box="[279,458,1587,1611]" italics="true" pageId="45" pageNumber="416">Lygus vittulatus</emphasis>
</taxonomicName>
in
<taxonomicName id="4C1B4D0AFF903B2317C0FB5AFDD7F99F" authorityName="Reuter" authorityYear="1875" box="[501,611,1588,1611]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF903B2317C0FB5AFDD7F99F" box="[501,611,1588,1611]" italics="true" pageId="45" pageNumber="416">Lygocoris</emphasis>
</taxonomicName>
, the present close examination of the surface structures and genitalia recognised it as a member of
<taxonomicName id="4C1B4D0AFF903B231781FB1DFDF9F95F" authorityName="Yasunaga" authorityYear="2024" box="[436,589,1651,1675]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF903B231781FB1DFDF9F95F" box="[436,589,1651,1675]" italics="true" pageId="45" pageNumber="416">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF903B23146EFB1AFD66F958" box="[603,722,1652,1676]" pageId="45" pageNumber="416" rank="genus">new genus</taxonomicNameLabel>
. A new combination is accordingly established.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,355 @@
<document id="C44241013F925ADAC564E691ADE35448" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF903B3D1583FC72FEF0FBC7" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Sabactiopus schuhi Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="422" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="416" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2A0F7401D52A898C70BC11663B89BBC5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5CAEFEF0A72E2BD46F3BBFFE9F291575">
<mods:title id="9185B755E0DF08CB98715AB9562FA125">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="86FB910421880D7FD1565D133FB83E63" type="personal">
<mods:role id="F53227654C339C75C40EEA88399D7819">
<mods:roleTerm id="F31B4175762DFD5C53C5054877F9338D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6057574920F6466B1BE8D7A0B353237D">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="2BB876062570F94F4515FB3DD6FEC443">text</mods:typeOfResource>
<mods:relatedItem id="BD6EBDA5D76D35B21D712BA9CE4E9231" type="host">
<mods:titleInfo id="E32119F1F1FBD3B02D4E5B9D6CB45FA1">
<mods:title id="5B8AC68DCDF2F0CAF44CB71C31E92C63">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="865BA3FF3DC07A2A165EB660D8ACA47C">
<mods:date id="CC6862082A6A074B8358081633CC786A">2024</mods:date>
<mods:detail id="0B7A09BFB735E8550EFFC373DA74155E" type="pubDate">
<mods:number id="B03A803F428BA3C202AD051689F5E01E">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="301930E013E83F1B6D0DD3CBF08BF980" type="volume">
<mods:number id="44212C45E7FF96DD578AEC9CC9D0BFB5">72</mods:number>
</mods:detail>
<mods:extent id="9D18AFB4D562B9E19454DFCFD2E2B3CA" unit="page">
<mods:start id="C862368D354CB4CD8B3FBAAB21662725">371</mods:start>
<mods:end id="A484EDD155677A6B46CA5433D5F418D6">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="1344A001B8A29DF30606352FF79BF70A">journal article</mods:classification>
<mods:identifier id="8EB2F5DC96826492C348FE2C83748CF7" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="E5AC36B7196D4BD572CB204B608A420E" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="AB98897495E882B237AEB811F588F347" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="D66C155795C4FB5070F45B8692E91BD8" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF903B3D1583FC72FEF0FBC7" LSID="urn:lsid:plazi:treatment:03B2879FFF903B3D1583FC72FEF0FBC7" httpUri="http://treatment.plazi.org/id/03B2879FFF903B3D1583FC72FEF0FBC7" lastPageId="51" lastPageNumber="422" pageId="45" pageNumber="416">
<subSubSection id="C3016502FF903B231583FC72FA97FEE1" box="[950,1315,284,309]" pageId="45" pageNumber="416" type="nomenclature">
<paragraph id="8BA43689FF903B231583FC72FA97FEE1" blockId="45.[933,1331,284,341]" box="[950,1315,284,309]" pageId="45" pageNumber="416">
<heading id="D0EC81E5FF903B231583FC72FA97FEE1" bold="true" box="[950,1315,284,309]" centered="true" fontSize="10" level="2" pageId="45" pageNumber="416" reason="2">
<emphasis id="B96FEA9BFF903B231583FC72FA97FEE1" bold="true" box="[950,1315,284,309]" pageId="45" pageNumber="416">
<taxonomicName id="4C1B4D0AFF903B231583FC72FB3AFEE0" authorityName="Yasunaga" authorityYear="2024" box="[950,1166,284,308]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFF903B231583FC72FB3AFEE0" bold="true" box="[950,1166,284,308]" italics="true" pageId="45" pageNumber="416">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF903B2312A8FC73FA97FEE1" box="[1181,1315,285,309]" pageId="45" pageNumber="416" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231590FC53FA87FE81" box="[933,1331,317,341]" pageId="45" pageNumber="416" type="description">
<paragraph id="8BA43689FF903B231590FC53FA87FE81" blockId="45.[933,1331,284,341]" box="[933,1331,317,341]" pageId="45" pageNumber="416">
(
<figureCitation id="13202A0CFF903B231598FC53FC4CFE81" box="[941,1016,317,341]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="45" pageNumber="416">Figs. 7</figureCitation>
GJ, 18AG, 34JL, 35AI)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231518FC13FBF3FD41" pageId="45" pageNumber="416" type="materials_examined">
<paragraph id="8BA43689FF903B231518FC13FBF3FD41" blockId="45.[813,1452,381,661]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231518FC13FC56FE41" bold="true" box="[813,994,381,405]" pageId="45" pageNumber="416">Type material.</emphasis>
<materialsCitation id="3B733CD4FF903B2315DBFC13FA99FE21" collectingDate="2006-08-02" collectionCode="NMTU, AMNH" country="Nepal" county="Mt. Pulchoki" elevation="1650" latitude="27.579168" location="Kathmandu Valley" longLatPrecision="20" longitude="85.37889" municipality="Godawari" pageId="45" pageNumber="416" specimenCode="PBI 378790" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A0882BFF903B2315DBFC13FBE8FE41" box="[1006,1116,381,405]" pageId="45" pageNumber="416" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="B96FEA9BFF903B231280FC13FAA8FE41" bold="true" box="[1205,1308,381,405]" pageId="45" pageNumber="416">
<collectingCountry id="F30C7619FF903B231280FC13FAACFE41" box="[1205,1304,381,405]" name="Nepal" pageId="45" pageNumber="416">NEPAL</collectingCountry>
:
</emphasis>
<location id="8EC46052FF903B23131DFC13FCCCFE61" LSID="urn:lsid:plazi:treatment:03B2879FFF903B3D1583FC72FEF0FBC7:8EC46052FF903B23131DFC13FCCCFE61" country="Nepal" county="Mt. Pulchoki" latitude="27.579168" longLatPrecision="20" longitude="85.37889" municipality="Godawari" name="Kathmandu Valley" pageId="45" pageNumber="416">Kathmandu Valley</location>
:
<location id="8EC46052FF903B2315B6FCF3FC68FE61" LSID="urn:lsid:plazi:treatment:03B2879FFF903B3D1583FC72FEF0FBC7:8EC46052FF903B2315B6FCF3FC68FE61" box="[899,988,413,437]" country="Nepal" county="Mt. Pulchoki" latitude="27.579168" longLatPrecision="20" longitude="85.37889" municipality="Godawari" name="Lalitpur" pageId="45" pageNumber="416">Lalitpur</location>
,
<collectingMunicipality id="6BC0ACF3FF903B2315D2FCF3FBE7FE61" box="[999,1107,413,437]" pageId="45" pageNumber="416">Godawari</collectingMunicipality>
,
<collectingCounty id="62C54E05FF903B23126BFCF3FB5EFE61" box="[1118,1258,413,437]" pageId="45" pageNumber="416">Mt. Pulchoki</collectingCounty>
(
<quantity id="4CE39B6CFF903B2312CCFCF3FA2BFE61" box="[1273,1439,413,437]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="45" pageNumber="416" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<elevation id="0036D1BAFF903B2312CCFCF3FA2BFE61" box="[1273,1439,413,437]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="45" pageNumber="416" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1,5001,800 m</elevation>
</quantity>
),
<geoCoordinate id="EE2F504EFF903B231518FCD5FC1AFE00" box="[813,942,443,468]" degrees="27" direction="north" minutes="34" orientation="latitude" pageId="45" pageNumber="416" precision="15" seconds="45" value="27.579168">27°3445″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF903B23158FFCD5FB8CFE00" box="[954,1080,443,468]" degrees="85" direction="east" minutes="22" orientation="longitude" pageId="45" pageNumber="416" precision="15" seconds="44" value="85.37889">85°2244″E</geoCoordinate>
,
<collectedFrom id="38077574FF903B231270FCD3FA26FE01" box="[1093,1426,445,469]" pageId="45" pageNumber="416">
on
<taxonomicName id="4C1B4D0AFF903B23125DFCD0FA26FE01" authority=", T. Yasunaga" authorityName="T. Yasunaga" box="[1128,1426,445,469]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="45" pageNumber="416" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFF903B23125DFCD0FB4DFE01" box="[1128,1273,445,469]" italics="true" pageId="45" pageNumber="416">Urtica dioica</emphasis>
, T. Yasunaga
</taxonomicName>
</collectedFrom>
,
<date id="FFA51049FF903B2313ABFCD3FC0FFE21" pageId="45" pageNumber="416" value="2006-08-02">
<collectingDate id="EFE1E9A1FF903B2313ABFCD3FC0FFE21" pageId="45" pageNumber="416" value="2006-08-02">2 August 2006</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF903B2315F8FCB3FB94FE21" box="[973,1056,477,501]" pageId="45" pageNumber="416">NMTU</collectionCode>
) (
<collectionCode id="ED0AAE4CFF903B23120DFCB3FB38FE21" box="[1080,1164,477,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="45" pageNumber="416">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FF903B2312AFFCB3FA91FE21" box="[1178,1317,477,501]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="45" pageNumber="416">PBI 378790</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FF903B23130FFCB3FA94FDE0" collectingDate="2006-08-02" collectionCode="TYCN, T" country="Nepal" county="Mt. Pulchoki" elevation="1650" latitude="28.166666" location="Kathmandu Valley" longLatPrecision="1233" longitude="84.1" municipality="Godawari" pageId="45" pageNumber="416" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54A0882BFF903B23130FFCB3FA1CFE21" box="[1338,1448,477,501]" pageId="45" pageNumber="416" type="paratype">Paratypes</typeStatus>
: NEPAL:
<specimenCount id="9D1DFD00FF903B2315BBFC93FC5FFDC1" box="[910,1003,509,533]" count="1" pageId="45" pageNumber="416" type="female">1 female</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFF903B23133DFC93FAE2FDC1" box="[1288,1366,509,533]" pageId="45" pageNumber="416">TYCN</collectionCode>
); Kaski: Pokhara, Begnas Lakeside,
<geoCoordinate id="EE2F504EFF903B23127BFF75FB13FDE0" box="[1102,1191,539,564]" degrees="28" direction="north" minutes="10" orientation="latitude" pageId="45" pageNumber="416" precision="925" value="28.166666">28°10N</geoCoordinate>
<geoCoordinate id="EE2F504EFF903B231299FF75FAB0FDE0" box="[1196,1284,539,564]" degrees="84" direction="east" minutes="06" orientation="longitude" pageId="45" pageNumber="416" precision="925" value="84.1">84°06E</geoCoordinate>
,
<collectionCode id="ED0AAE4CFF903B231338FF73FA94FDE0" box="[1293,1312,541,564]" name="Tavera, Department of Geology and Geophysics" pageId="45" pageNumber="416">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B733CD4FF903B23131CFF73FBF7FD41" collectingDate="2001-08-15" collectingDateMax="2005-11-06" collectingDateMin="2001-08-15" collectionCode="NMTU, DOAT" collectorName="S. Nagashima" country="Nepal" elevation="850" latitude="18.833334" longLatPrecision="1273" longitude="98.9" pageId="45" pageNumber="416" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
Yasunaga,
<date id="FFA51049FF903B2313ABFF73FC63FD81" pageId="45" pageNumber="416" value="2005-11-06">
<collectingDate id="EFE1E9A1FF903B2313ABFF73FC63FD81" pageId="45" pageNumber="416" value="2005-11-06">6 November 2005</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF903B2315D6FF53FB83FD81" box="[995,1079,573,597]" pageId="45" pageNumber="416">NMTU</collectionCode>
). THAILAND:
<specimenCount id="9D1DFD00FF903B2312E9FF53FA91FD81" box="[1244,1317,573,597]" count="1" pageId="45" pageNumber="416" type="male">1 male</specimenCount>
, Doi Suthep, Chiang Khian,
<quantity id="4CE39B6CFF903B2315E5FF33FBA1FDA0" box="[976,1045,605,629]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="45" pageNumber="416" unit="m" value="850.0">
<elevation id="0036D1BAFF903B2315E5FF33FBA1FDA0" box="[976,1045,605,629]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="45" pageNumber="416" unit="m" value="850.0">850 m</elevation>
</quantity>
alt.,
<geoCoordinate id="EE2F504EFF903B23127EFF35FB10FDA0" box="[1099,1188,603,628]" degrees="18" direction="north" minutes="50" orientation="latitude" pageId="45" pageNumber="416" precision="925" value="18.833334">18°50N</geoCoordinate>
<geoCoordinate id="EE2F504EFF903B231299FF35FAB1FDA0" box="[1196,1285,603,628]" degrees="98" direction="east" minutes="54" orientation="longitude" pageId="45" pageNumber="416" precision="925" value="98.9">98°54E</geoCoordinate>
,
<collectorName id="26EE535FFF903B231325FF33FA1CFDA1" box="[1296,1448,605,629]" pageId="45" pageNumber="416">S. Nagashima</collectorName>
,
<date id="FFA51049FF903B231518FF13FC6FFD41" box="[813,987,637,661]" pageId="45" pageNumber="416" value="2001-08-15">
<collectingDate id="EFE1E9A1FF903B231518FF13FC6FFD41" box="[813,987,637,661]" pageId="45" pageNumber="416" value="2001-08-15">15 August 2001</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF903B2315DEFF13FB8FFD41" box="[1003,1083,637,661]" pageId="45" pageNumber="416">DOAT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B231518FFD3FA12FC00" pageId="45" pageNumber="416" type="diagnosis">
<paragraph id="8BA43689FF903B231518FFD3FA12FC00" blockId="45.[813,1452,701,981]" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231518FFD3FC15FD01" bold="true" box="[813,929,701,725]" pageId="45" pageNumber="416">Diagnosis.</emphasis>
Being closely related to
<taxonomicName id="4C1B4D0AFF903B231287FFD3FA3FFD01" authority="(Poppius)" baseAuthorityName="Poppius" box="[1202,1419,701,725]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFF903B231287FFD3FAADFD01" box="[1202,1305,701,725]" italics="true" pageId="45" pageNumber="416">S. sauteri</emphasis>
(Poppius)
</taxonomicName>
,
<taxonomicName id="4C1B4D0AFF903B2313A2FFD3FCC7FD21" authorityName="Yasunaga" authorityYear="2024" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="45" pageNumber="416" phylum="Arthropoda" rank="species" species="schuhi" status="new species">
<emphasis id="B96FEA9BFF903B2313A2FFD3FCC7FD21" italics="true" pageId="45" pageNumber="416">S. schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF903B2315B6FFB3FBB3FD21" box="[899,1031,733,757]" pageId="45" pageNumber="416" rank="species">new species</taxonomicNameLabel>
can be distinguished by smaller size; narrower vertex; wider yellowish subbasal annulation on antennal antennomere II; antennomere III longer than IV; roundly inflated sensory lobe of left paramere; T-shaped apex of right paramere; anchor-shaped apex and roundly inflated sensory lobe of left paramere; narrowed sclerotised rings; and mesally separated interramal lobes, in addition to being associated with a herb from the
<taxonomicName id="4C1B4D0AFF903B2312B8FED3FAB6FC01" box="[1165,1282,957,981]" class="Magnoliopsida" family="Urticaceae" kingdom="Plantae" order="Rosales" pageId="45" pageNumber="416" phylum="Tracheophyta" rank="family">Urticaceae</taxonomicName>
,
<taxonomicName id="4C1B4D0AFF903B23133BFED0FA14FC01" box="[1294,1440,957,981]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="45" pageNumber="416" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFF903B23133BFED0FA14FC01" box="[1294,1440,957,981]" italics="true" pageId="45" pageNumber="416">Urtica dioica</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF903B3D1518FE93FE04FE40" lastPageId="51" lastPageNumber="422" pageId="45" pageNumber="416" type="description">
<paragraph id="8BA43689FF903B3D1518FE93FDEBFE87" blockId="45.[813,1452,1021,2101]" lastBlockId="51.[113,752,187,339]" lastPageId="51" lastPageNumber="422" pageId="45" pageNumber="416">
<emphasis id="B96FEA9BFF903B231518FE93FC03FBC1" bold="true" box="[813,951,1021,1045]" pageId="45" pageNumber="416">Description.</emphasis>
Body ovoid, stout, rather box-like, small-sized; in female body larger and paler in colour (
<figureCitation id="13202A0CFF903B2312C1F973FAD5FBE1" box="[1268,1377,1053,1077]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="45" pageNumber="416">Fig. 7GI</figureCitation>
); basic colouration shiny dark reddish brown; dorsum polished, finely and sparsely punctate, with uniformly distributed, pale, simple, reclining or semi-erect setae on mesoscutum, scutellum and hemelytra. Head shiny pale brown, partly tinged with red, almost glabrous; eyes small; basal transverse carina on vertex with several upright setae, about as thick as pronotal collar (
<figureCitation id="13202A0CFF903B2315E4F993FB86FAC1" box="[977,1074,1277,1301]" captionStart="Fig" captionStartId="49.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@49.[113,1451,251,1929]" captionTargetPageId="49" captionText="Fig. 34. Scanning electron micrographs of Paralygocoris vittulatus (AI), Sabactiopus schuhi, new species (JL) and S. sauteri (MO). A, head and pronotum, left lateral view; B, scutellum and hemelytron, left lateral view; C, metathoracic scent efferent system; D, metatarsus; E, pygophore, dorsal view; F, vesica; G, sclerotised ring; H, I, posterior wall; J, left lateral habitus; K, anterior body, dorsal view; L, O, thoracic pleura, left lateral view; M, anterior body, left lateral view; N, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683036" httpUri="https://zenodo.org/record/14683036/files/figure.png" pageId="45" pageNumber="416">Fig. 34K</figureCitation>
). Antenna dark brown; antennomere I with darkened apex, base and middle; antennomere II with pale ring at basal ⅓, as long as or slightly shorter than basal width of pronotum; base of antennomere III whitish, longer than IV. Labium pale brown, reaching but not exceeding apex of metacoxa; segment IV reddish brown. Pronotum shiny pale or reddish brown, shallowly punctate or rugose, almost glabrous (
<figureCitation id="13202A0CFF903B2315C7F8B3FBE9FA21" box="[1010,1117,1501,1525]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="45" pageNumber="416">Fig. 29K</figureCitation>
); calli sometimes with a pair of dark spots, flat; pleura pale brown; metathoracic scent efferent system greyish brown, with rather flat peritreme (
<figureCitation id="13202A0CFF903B231501FB53FC2DF981" box="[820,921,1597,1621]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="45" pageNumber="416">Fig. 35A</figureCitation>
); scutellum shiny fuscous, with creamy V-shaped margin. Hemelytron shining, pale brown or reddish brown, widely darkened posteriorly, declivous at cuneal fracture; cuneus yellowish brown apically, with dark apex; membrane pale smoky brown. Coxae and legs pale brown, partly tinged with red; meso- and metacoxa, and base of each femur more or less darkened; tibiae with dark spots or annulations at bases of pale brown spines; tarsomere III darkened; meta-tarsomere II as long as III; pretarsal structures as in
<figureCitation id="13202A0CFF903B2313B4FA53FCE8F8A1" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="45" pageNumber="416">Fig. 30C</figureCitation>
. Abdomen fuscous; female sterna VIIIX pale brown. Male genitalia (
<figureCitation id="13202A0CFF903B2315ECFA13FBDCF841" box="[985,1128,1917,1941]" captionStart="Fig" captionStartId="28.[135,170,1939,1961]" captionTargetBox="[135,1473,285,1924]" captionTargetId="figure-7@28.[135,1473,285,1924]" captionTargetPageId="28" captionText="Fig. 18. Male (AD, H) and female (EG, I, J) genitalia of Sabactiopus schuhi, new species (AG) and S. sauteri (HJ). A, Apex of pygophore with parameres, dorsal view; B, apex of left paramere; C, apical part of right paramere; D, H, vesica; E, genital chamber; F, I, posterior wall; G, J, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682992" httpUri="https://zenodo.org/record/14682992/files/figure.png" pageId="45" pageNumber="416">Figs. 18AD</figureCitation>
,
<figureCitation id="13202A0CFF903B231243FA13FB25F841" box="[1142,1169,1917,1941]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="45" pageNumber="416">35</figureCitation>
DF): Left paramere with roundly inflated sensory lobe (
<figureCitation id="13202A0CFF903B2312B7FAF3FB40F861" box="[1154,1268,1949,1973]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="45" pageNumber="416">Figs. 30D</figureCitation>
); right paramere with elongate hypophysis and T-shaped, flat apex (
<figureCitation id="13202A0CFF903B23130BFAD3FA13F801" box="[1342,1447,1981,2005]" captionStart="Fig" captionStartId="28.[135,170,1939,1961]" captionTargetBox="[135,1473,285,1924]" captionTargetId="figure-7@28.[135,1473,285,1924]" captionTargetPageId="28" captionText="Fig. 18. Male (AD, H) and female (EG, I, J) genitalia of Sabactiopus schuhi, new species (AG) and S. sauteri (HJ). A, Apex of pygophore with parameres, dorsal view; B, apex of left paramere; C, apical part of right paramere; D, H, vesica; E, genital chamber; F, I, posterior wall; G, J, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682992" httpUri="https://zenodo.org/record/14682992/files/figure.png" pageId="45" pageNumber="416">Figs. 18C</figureCitation>
,
<figureCitation id="13202A0CFF903B231518FAB3FCE8F821" box="[813,860,2013,2037]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="45" pageNumber="416">35E</figureCitation>
). Vesica widely membranous, with two lobal-sclerites, one of which is deeply notched; secondary gonopore thick-rimmed (
<figureCitation id="13202A0CFF903B2315A7F573FBB5F7E1" box="[914,1025,2077,2101]" captionStart="Fig" captionStartId="28.[135,170,1939,1961]" captionTargetBox="[135,1473,285,1924]" captionTargetId="figure-7@28.[135,1473,285,1924]" captionTargetPageId="28" captionText="Fig. 18. Male (AD, H) and female (EG, I, J) genitalia of Sabactiopus schuhi, new species (AG) and S. sauteri (HJ). A, Apex of pygophore with parameres, dorsal view; B, apex of left paramere; C, apical part of right paramere; D, H, vesica; E, genital chamber; F, I, posterior wall; G, J, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682992" httpUri="https://zenodo.org/record/14682992/files/figure.png" pageId="45" pageNumber="416">Figs. 18D</figureCitation>
,
<figureCitation id="13202A0CFF903B231225F573FB89F7E1" box="[1040,1085,2077,2101]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="45" pageNumber="416">35F</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFF903B23132CF573FA13F7E1" box="[1305,1447,2077,2101]" captionStart="Fig" captionStartId="28.[135,170,1939,1961]" captionTargetBox="[135,1473,285,1924]" captionTargetId="figure-7@28.[135,1473,285,1924]" captionTargetPageId="28" captionText="Fig. 18. Male (AD, H) and female (EG, I, J) genitalia of Sabactiopus schuhi, new species (AG) and S. sauteri (HJ). A, Apex of pygophore with parameres, dorsal view; B, apex of left paramere; C, apical part of right paramere; D, H, vesica; E, genital chamber; F, I, posterior wall; G, J, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682992" httpUri="https://zenodo.org/record/14682992/files/figure.png" pageId="45" pageNumber="416">Figs. 18EG</figureCitation>
,
<figureCitation id="13202A0CFF8E3B3D1644FDD5FF39FF07" box="[113,141,187,211]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="51" pageNumber="422">35</figureCitation>
GI): Genital chamber with small, narrow sclerotised rings that are separated to each other (
<figureCitation id="13202A0CFF8E3B3D1420FDB5FDCFFF27" box="[533,635,219,243]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="51" pageNumber="422">Fig. 35G</figureCitation>
); posterior wall lacking dorsal structure or lateral lobe; interramal lobe comparatively narrow, densely covered with spinules (
<figureCitation id="13202A0CFF8E3B3D14F1FC75FF23FE87" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="51" pageNumber="422">Fig. 35I</figureCitation>
), each lobe separated mesally (
<figureCitation id="13202A0CFF8E3B3D17D8FC55FDE7FE87" box="[493,595,315,339]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="51" pageNumber="422">Fig. 35H</figureCitation>
).
</paragraph>
<caption id="DF646601FF933B2016BDFAF6FBC0F7D0" ID-DOI="http://doi.org/10.5281/zenodo.14683028" ID-Zenodo-Dep="14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="46" pageNumber="417" startId="46.[136,171,1944,1966]" targetBox="[136,1474,251,1929]" targetPageId="46" targetType="figure">
<paragraph id="8BA43689FF933B2016BDFAF6FBC0F7D0" blockId="46.[136,1475,1944,2053]" pageId="46" pageNumber="417">
Fig. 31. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF933B201405FAF6FCAFF879" authorityName="Yasunaga" authorityYear="2024" box="[560,795,1944,1965]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="417" phylum="Arthropoda" rank="species" species="takaii" status="sp. nov.">
<emphasis id="B96FEA9BFF933B201405FAF6FCAFF879" box="[560,795,1944,1965]" italics="true" pageId="46" pageNumber="417">Himalisundacoris takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF933B20151CFAF7FC15F87A" box="[809,929,1944,1966]" pageId="46" pageNumber="417" rank="species">new species</taxonomicNameLabel>
, holotype male (AG) and
<taxonomicName id="4C1B4D0AFF933B201284FAF6FA08F879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Lu &amp; Zheng" baseAuthorityYear="2004" box="[1201,1468,1944,1965]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="417" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFF933B201284FAF6FA08F879" box="[1201,1468,1944,1965]" italics="true" pageId="46" pageNumber="417">Paralygocoris meridionalis</emphasis>
</taxonomicName>
, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I).
</paragraph>
</caption>
<caption id="DF646601FF923B211644FAF6FECFF7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683030" ID-Zenodo-Dep="14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="47" pageNumber="418" startId="47.[113,149,1944,1966]" targetBox="[113,1451,251,1929]" targetPageId="47" targetType="figure">
<paragraph id="8BA43689FF923B211644FAF6FECFF7D1" blockId="47.[113,1452,1944,2053]" pageId="47" pageNumber="418">
Fig. 32. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF923B21141EFAF6FCECF879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Carvalho" baseAuthorityYear="1987" box="[555,856,1944,1965]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="418" phylum="Arthropoda" rank="species" species="alboscutellatus">
<emphasis id="B96FEA9BFF923B21141EFAF6FCECF879" box="[555,856,1944,1965]" italics="true" pageId="47" pageNumber="418">Paralygocoris alboscutellatus</emphasis>
</taxonomicName>
(AF),
<taxonomicName id="4C1B4D0AFF923B21159BFAF6FBD4F879" authorityName="Yasunaga, Schwartz &amp; Chérot" authorityYear="2018" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[942,1120,1944,1965]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="418" phylum="Arthropoda" rank="species" species="disciger">
<emphasis id="B96FEA9BFF923B21159BFAF6FBD4F879" box="[942,1120,1944,1965]" italics="true" pageId="47" pageNumber="418">Prolygus disciger</emphasis>
</taxonomicName>
(GK),
<taxonomicName id="4C1B4D0AFF923B21128CFAF7FA8CF87A" authorityName="Poppius" authorityYear="1914" box="[1209,1336,1945,1966]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="418" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B96FEA9BFF923B21128CFAF7FA8CF87A" box="[1209,1336,1945,1966]" italics="true" pageId="47" pageNumber="418">P. papuanus</emphasis>
</taxonomicName>
(LO) and
<taxonomicName id="4C1B4D0AFF923B211644FADBFECFF81E" authorityName="Yasunaga, Schwartz, &amp; Cherot" authorityYear="2002" box="[113,379,1973,1994]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="418" phylum="Arthropoda" rank="species" species="furvocarinatus">
<emphasis id="B96FEA9BFF923B211644FADBFECFF81E" box="[113,379,1973,1994]" italics="true" pageId="47" pageNumber="418">Apolygopsis furvocarinatus</emphasis>
</taxonomicName>
(E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe
</paragraph>
</caption>
<caption id="DF646601FF8D3B3E16BDFAF6FEE7F7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683032" ID-Zenodo-Dep="14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="48" pageNumber="419" startId="48.[136,172,1944,1966]" targetBox="[136,1474,251,1929]" targetPageId="48" targetType="figure">
<paragraph id="8BA43689FF8D3B3E16BDFAF6FEE7F7D1" blockId="48.[136,1475,1944,2053]" pageId="48" pageNumber="419">
Fig. 33. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF8D3B3E140BFAF6FCAEF879" authorityName="Yasunaga" authorityYear="2024" box="[574,794,1944,1965]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="48" pageNumber="419" phylum="Arthropoda" rank="species" species="balicus" status="sp. nov.">
<emphasis id="B96FEA9BFF8D3B3E140BFAF6FCAEF879" box="[574,794,1944,1965]" italics="true" pageId="48" pageNumber="419">Paralygocoris balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF8D3B3E151CFAF7FC11F87A" box="[809,933,1944,1966]" pageId="48" pageNumber="419" rank="species">new species</taxonomicNameLabel>
. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall.
</paragraph>
</caption>
<caption id="DF646601FF8C3B3F1644FAF6FC52F7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683036" ID-Zenodo-Dep="14683036" httpUri="https://zenodo.org/record/14683036/files/figure.png" pageId="49" pageNumber="420" startId="49.[113,148,1944,1966]" targetBox="[113,1451,251,1929]" targetPageId="49" targetType="figure">
<paragraph id="8BA43689FF8C3B3F1644FAF6FC52F7D1" blockId="49.[113,1451,1944,2053]" pageId="49" pageNumber="420">
Fig. 34. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF8C3B3F143BFAF6FD42F879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[526,758,1944,1965]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="420" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFF8C3B3F143BFAF6FD42F879" box="[526,758,1944,1965]" italics="true" pageId="49" pageNumber="420">Paralygocoris vittulatus</emphasis>
</taxonomicName>
(AI),
<taxonomicName id="4C1B4D0AFF8C3B3F150AFAF6FC4DF879" authorityName="Yasunaga" authorityYear="2024" box="[831,1017,1944,1965]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="420" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFF8C3B3F150AFAF6FC4DF879" box="[831,1017,1944,1965]" italics="true" pageId="49" pageNumber="420">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF8C3B3F1233FAF7FBCDF87A" box="[1030,1145,1944,1966]" pageId="49" pageNumber="420" rank="species">new species</taxonomicNameLabel>
(JL) and
<taxonomicName id="4C1B4D0AFF8C3B3F12D6FAF7FA8AF87A" box="[1251,1342,1945,1966]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="49" pageNumber="420" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFF8C3B3F12D6FAF7FA8AF87A" box="[1251,1342,1945,1966]" italics="true" pageId="49" pageNumber="420">S. sauteri</emphasis>
</taxonomicName>
(MO). A, head and pronotum, left lateral view; B, scutellum and hemelytron, left lateral view; C, metathoracic scent efferent system; D, metatarsus; E, pygophore, dorsal view; F, vesica; G, sclerotised ring; H, I, posterior wall; J, left lateral habitus; K, anterior body, dorsal view; L, O, thoracic pleura, left lateral view; M, anterior body, left lateral view; N, head, dorsal view.
</paragraph>
</caption>
<caption id="DF646601FF8F3B3C16BDFAC9FDFBF822" ID-DOI="http://doi.org/10.5281/zenodo.14683038" ID-Zenodo-Dep="14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="50" pageNumber="421" startId="50.[136,171,1959,1981]" targetBox="[136,1474,265,1943]" targetPageId="50" targetType="figure">
<paragraph id="8BA43689FF8F3B3C16BDFAC9FDFBF822" blockId="50.[136,1475,1958,2038]" pageId="50" pageNumber="421">
Fig. 35. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF8F3B3C1418FAC9FD5DF868" authorityName="Yasunaga" authorityYear="2024" box="[557,745,1959,1980]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="421" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFF8F3B3C1418FAC9FD5DF868" box="[557,745,1959,1980]" italics="true" pageId="50" pageNumber="421">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF8F3B3C14C2FAC9FCD8F869" box="[759,876,1959,1981]" pageId="50" pageNumber="421" rank="species">new species</taxonomicNameLabel>
(AI) and
<taxonomicName id="4C1B4D0AFF8F3B3C15EEFAC9FB8CF868" box="[987,1080,1959,1980]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="50" pageNumber="421" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFF8F3B3C15EEFAC9FB8CF868" box="[987,1080,1959,1980]" italics="true" pageId="50" pageNumber="421">S. sauteri</emphasis>
</taxonomicName>
(JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view.
</paragraph>
</caption>
<paragraph id="8BA43689FF8E3B3D1644FC12FE04FE40" blockId="51.[113,432,379,404]" box="[113,432,379,404]" pageId="51" pageNumber="422">
<emphasis id="B96FEA9BFF8E3B3D1644FC12FEABFE40" bold="true" box="[113,287,380,404]" pageId="51" pageNumber="422">Measurements.</emphasis>
See
<tableCitation id="C6990332FF8E3B3D1763FC15FE1EFE40" box="[342,426,379,404]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="51" pageNumber="422">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF8E3B3D1644FCD2FF6CFDE7" pageId="51" pageNumber="422" type="etymology">
<paragraph id="8BA43689FF8E3B3D1644FCD2FF6CFDE7" blockId="51.[113,750,443,563]" pageId="51" pageNumber="422">
<emphasis id="B96FEA9BFF8E3B3D1644FCD2FF43FE00" bold="true" box="[113,247,444,468]" pageId="51" pageNumber="422">Etymology.</emphasis>
Named in honour of Dr. Randall T. Schuh (Curator Emeritus, AMNH), one of the worlds most prominent heteropterists and my long-time mentor; an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF8E3B3D1644FF35FEBEFD47" pageId="51" pageNumber="422" type="distribution">
<paragraph id="8BA43689FF8E3B3D1644FF35FEBEFD47" blockId="51.[113,751,603,659]" pageId="51" pageNumber="422">
<emphasis id="B96FEA9BFF8E3B3D1644FF35FEB7FDA7" bold="true" box="[113,259,603,627]" pageId="51" pageNumber="422">Distribution.</emphasis>
<collectingCountry id="F30C7619FF8E3B3D173CFF35FEFDFDA7" box="[265,329,603,627]" name="Nepal" pageId="51" pageNumber="422">Nepal</collectingCountry>
(Kathmandu Valley, Kasuki),
<collectingCountry id="F30C7619FF8E3B3D14A5FF35FD5BFDA7" box="[656,751,603,627]" name="Thailand" pageId="51" pageNumber="422">Thailand</collectingCountry>
(
<collectingRegion id="49DFF86BFF8E3B3D164DFF15FF4AFD47" box="[120,254,635,659]" country="Thailand" name="Chiang Mai" pageId="51" pageNumber="422">Chiang Mai</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF8E3B3D1644FFD5FEF0FBC7" pageId="51" pageNumber="422" type="biology_ecology">
<paragraph id="8BA43689FF8E3B3D1644FFD5FEF0FBC7" blockId="51.[112,752,699,1043]" pageId="51" pageNumber="422">
<emphasis id="B96FEA9BFF8E3B3D1644FFD5FF79FD07" bold="true" box="[113,205,699,723]" pageId="51" pageNumber="422">Biology.</emphasis>
In Kathmandu Valley, several adults of this new species were found to inhabit
<taxonomicName id="4C1B4D0AFF8E3B3D1780FFB2FDF1FD27" box="[437,581,731,755]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="51" pageNumber="422" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFF8E3B3D1780FFB2FDF1FD27" box="[437,581,731,755]" italics="true" pageId="51" pageNumber="422">Urtica dioica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C1B4D0AFF8E3B3D1446FFB5FD5DFD27" box="[627,745,731,755]" class="Magnoliopsida" family="Urticaceae" kingdom="Plantae" order="Rosales" pageId="51" pageNumber="422" phylum="Tracheophyta" rank="family">Urticaceae</taxonomicName>
) and to coexist on the herb with
<taxonomicName id="4C1B4D0AFF8E3B3D17F8FF95FD5BFCC7" authorityName="Yasunaga" authorityYear="2024" box="[461,751,763,787]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="422" phylum="Arthropoda" rank="species" species="duwalae">
<emphasis id="B96FEA9BFF8E3B3D17F8FF95FD5BFCC7" box="[461,751,763,787]" italics="true" pageId="51" pageNumber="422">Himalisundacoris duwalae</emphasis>
</taxonomicName>
under deciduous/evergreen mixed broadleaf forest (
<figureCitation id="13202A0CFF8E3B3D14F7FE75FF3FFC80" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="51" pageNumber="422">Fig. 7J</figureCitation>
). On the other hand, the Taiwanese congener,
<taxonomicName id="4C1B4D0AFF8E3B3D14BCFE52FF6FFCA7" authority="(Poppius)" baseAuthorityName="Poppius" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="51" pageNumber="422" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFF8E3B3D14BCFE52FD5BFC87" box="[649,751,828,851]" italics="true" pageId="51" pageNumber="422">S. sauteri</emphasis>
(Poppius)
</taxonomicName>
, has been collected from the inflorescences and flower-buds of broadleaf trees, such as
<taxonomicName id="4C1B4D0AFF8E3B3D1410FE12FF60FC67" authority="Matsum." authorityName="Matsum." class="Magnoliopsida" family="Styracaceae" genus="Styrax" kingdom="Plantae" order="Ericales" pageId="51" pageNumber="422" phylum="Tracheophyta" rank="species" species="formosanus">
<emphasis id="B96FEA9BFF8E3B3D1410FE12FD5BFC47" box="[549,751,891,915]" italics="true" pageId="51" pageNumber="422">Styrax formosanus</emphasis>
Matsum.
</taxonomicName>
(
<taxonomicName id="4C1B4D0AFF8E3B3D16D2FEF2FEC7FC60" box="[231,371,924,948]" class="Magnoliopsida" family="Styracaceae" kingdom="Plantae" order="Ericales" pageId="51" pageNumber="422" phylum="Tracheophyta" rank="family">Styracaceae</taxonomicName>
) (
<figureCitation id="13202A0CFF8E3B3D17BAFEF5FE5FFC60" box="[399,491,923,948]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="51" pageNumber="422">Fig. 7L</figureCitation>
).
<bibRefCitation id="EF8A4B78FF8E3B3D1431FEF5FD5AFC60" author="Cherot F &amp; Gorczyca J &amp; Schwartz MD &amp; Demol T" box="[516,750,923,948]" pageId="51" pageNumber="422" pagination="55 - 139" refId="ref23668" refString="Cherot F, Gorczyca J, Schwartz MD &amp; Demol T (2017) The Bryocorinae, Cylapinae, Deraeocorinae and Mirinae (Insecta: Heteroptera: Miridae) from Baiteta Forest, Papua New Guinea, with a discussion of their feeding habits and a list of species of the country, pp. 55 - 139, 12 plates. In: Telnov D, Barclay MVL &amp; Pauwels OSG (eds.) Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, Vol. III. The Entomological Society of Latvia, Riga, 658 pp + 172 pls." type="book chapter" year="2017">Chérot et al. (2017)</bibRefCitation>
reported that a
<collectingRegion id="49DFF86BFF8E3B3D1714FED2FED0FC07" box="[289,356,956,979]" country="Indonesia" name="Papua Barat" pageId="51" pageNumber="422">Papua</collectingRegion>
New Guinean congener,
<taxonomicName id="4C1B4D0AFF8E3B3D14BCFED2FDE6FC20" authority="Cherot, Gorczyca, Schwartz &amp; Demol, 2017" authorityName="Cherot, Gorczyca, Schwartz &amp; Demol" authorityYear="2017" class="Magnoliopsida" family="Styracaceae" genus="Styrax" kingdom="Plantae" order="Ericales" pageId="51" pageNumber="422" phylum="Tracheophyta" rank="species" species="zhengi">
<emphasis id="B96FEA9BFF8E3B3D14BCFED2FD5BFC07" box="[649,751,955,979]" italics="true" pageId="51" pageNumber="422">S. zhengi</emphasis>
<bibRefCitation id="EF8A4B78FF8E3B3D1645FEB5FDE6FC20" author="Cherot F &amp; Gorczyca J &amp; Schwartz MD &amp; Demol T" box="[112,594,987,1012]" pageId="51" pageNumber="422" pagination="55 - 139" refId="ref23668" refString="Cherot F, Gorczyca J, Schwartz MD &amp; Demol T (2017) The Bryocorinae, Cylapinae, Deraeocorinae and Mirinae (Insecta: Heteroptera: Miridae) from Baiteta Forest, Papua New Guinea, with a discussion of their feeding habits and a list of species of the country, pp. 55 - 139, 12 plates. In: Telnov D, Barclay MVL &amp; Pauwels OSG (eds.) Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, Vol. III. The Entomological Society of Latvia, Riga, 658 pp + 172 pls." type="book chapter" year="2017">Chérot, Gorczyca, Schwartz &amp; Demol, 2017</bibRefCitation>
</taxonomicName>
, was collected by canopy-fogging.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,232 @@
<document id="E03A9165A8915E84FE27D13AB429D0AF" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF943B271575FF93FBA2F83A" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris meridionalis Yasunaga 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="412" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="412" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C3B4BED2B24BA0EA79CA2B516A4A294A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F2EED29C900FEC3D0AB067779F6276C2">
<mods:title id="3D76B59878B11A3200A601212DABAB42">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="4F332C7EC33505AC411671E4DCF30327" type="personal">
<mods:role id="B4C5AAAF76C8F0CD7B775F2F7EDA2CA6">
<mods:roleTerm id="FB38BAFA60A24D3C679C373744302408">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12E26B5CE5A8B077A8353840C5041976">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="8C5B0701E6F3663D0F3C5F962A15A174">text</mods:typeOfResource>
<mods:relatedItem id="5DEED00E198983654D2C47F2439AA59C" type="host">
<mods:titleInfo id="9F0B58E422E693BA873991071EA7072A">
<mods:title id="0E7B92F5A536FD9EEBE70484D79BA962">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="4760A75B274DFE9A20AE4EC8FD62A5F6">
<mods:date id="405F33B86DC5B41B4E99CBFFD66FC1E5">2024</mods:date>
<mods:detail id="0F5E942D6BD5FBFDA7069C893DC1D583" type="pubDate">
<mods:number id="F3B979516BB118A974F1EDC19B55BE95">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="57F327836FA7ECE23273268FB0D79FBF" type="volume">
<mods:number id="D321630D1638538EA565716E906DA4B1">72</mods:number>
</mods:detail>
<mods:extent id="31A93AD67CD7900DB94CEFB118FD3EBA" unit="page">
<mods:start id="341699B3B61F13F6A7F3BC48C046D1CD">371</mods:start>
<mods:end id="020AC48669A7DF3A056DC6464896160B">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="469EB1C71CBB90206F5475E73D15065A">journal article</mods:classification>
<mods:identifier id="8AB5751574A4D086F28100E9B4E010B9" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="A912CFFB7C542E53B5A14D54EA6A612F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="272F5703988D95271A45442DA9E18CAA" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="D52E80BB5EDF8F9051717F8D768D951A" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF943B271575FF93FBA2F83A" ID-DOI="http://doi.org/10.5281/zenodo.14683046" ID-Zenodo-Dep="14683046" LSID="urn:lsid:plazi:treatment:03B2879FFF943B271575FF93FBA2F83A" httpUri="http://treatment.plazi.org/id/03B2879FFF943B271575FF93FBA2F83A" lastPageNumber="412" pageId="41" pageNumber="412">
<subSubSection id="C3016502FF943B271575FF93FB07FCE1" pageId="41" pageNumber="412" type="nomenclature">
<paragraph id="8BA43689FF943B271575FF93FB07FCE1" blockId="41.[832,1432,765,853]" pageId="41" pageNumber="412">
<heading id="D0EC81E5FF943B271575FF93FB07FCE1" bold="true" centered="true" fontSize="10" level="2" pageId="41" pageNumber="412" reason="2">
<emphasis id="B96FEA9BFF943B271575FF93FB07FCE1" bold="true" pageId="41" pageNumber="412">
<taxonomicName id="4C1B4D0AFF943B271575FF93FAE8FCC1" authority="(Lu &amp; Zheng, 2004)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Lu &amp; Zheng" baseAuthorityYear="2004" box="[832,1372,765,789]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="meridionalis" status="comb. nov.">
<emphasis id="B96FEA9BFF943B271575FF93FBDFFCC1" bold="true" box="[832,1131,765,789]" italics="true" pageId="41" pageNumber="412">Paralygocoris meridionalis</emphasis>
(Lu &amp;
<bibRefCitation id="EF8A4B78FF943B2712F0FF93FAE1FCC1" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[1221,1365,765,789]" pageId="41" pageNumber="412" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng, 2004</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF943B27135FFF93FB07FCE1" pageId="41" pageNumber="412" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B2715F4FE53FAA2FC81" box="[961,1302,829,853]" pageId="41" pageNumber="412" type="description">
<paragraph id="8BA43689FF943B2715F4FE53FAA2FC81" blockId="41.[832,1432,765,853]" box="[961,1302,829,853]" pageId="41" pageNumber="412">
(
<figureCitation id="13202A0CFF943B2715FCFE53FB94FC81" box="[969,1056,829,853]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="41" pageNumber="412">Figs. 12</figureCitation>
KL, 17FH, 31HO)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FE12FB38FC7B" pageId="41" pageNumber="412" type="reference_group">
<paragraph id="8BA43689FF943B271518FE12FB38FC7B" blockId="41.[813,1451,892,943]" pageId="41" pageNumber="412">
<treatmentCitationGroup id="AB0B11A7FF943B271518FE12FB38FC7B" pageId="41" pageNumber="412">
<taxonomicName id="4C1B4D0AFF943B271518FE12FB20FC46" ID-CoL="46LWB" authority="Lu &amp; Zheng" authorityName="Lu &amp; Zheng" authorityYear="2004" box="[813,1172,892,914]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFF943B271518FE12FBBFFC45" box="[813,1035,892,913]" italics="true" pageId="41" pageNumber="412">Neolygus meridionalis</emphasis>
Lu &amp; Zheng
</taxonomicName>
, in
<treatmentCitation id="0ABA1098FF943B27128BFE12FA1FFC45" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[1214,1451,892,914]" page="421" pageId="41" pageNumber="412" year="2004">
<bibRefCitation id="EF8A4B78FF943B27128BFE12FA1FFC45" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[1214,1451,892,914]" pageId="41" pageNumber="412" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng et al., 2004: 421</bibRefCitation>
</treatmentCitation>
(
<quantity id="4CE39B6CFF943B27156CFEF7FC2DFC7B" box="[857,921,921,943]" metricMagnitude="1" metricUnit="m" metricValue="1.89992" pageId="41" pageNumber="412" unit="in" value="748.0">748 in</quantity>
English), fig. 141, pl. 6.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FEB9FABEFB7B" pageId="41" pageNumber="412" type="materials_examined">
<paragraph id="8BA43689FF943B271518FEB9FABEFB7B" blockId="41.[813,1452,983,1199]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FEB9FBAAFC3B" bold="true" box="[813,1054,983,1007]" pageId="41" pageNumber="412">Material examined.</emphasis>
<materialsCitation id="3B733CD4FF943B27121FFEB9FC3FFBFB" collectingDate="1986-04-29" collectionCode="TYCN" collectorName="S. Gotoh" country="Malaysia" latitude="3.7166667" location="Fraser's Hill" longLatPrecision="1307" longitude="101.73333" pageId="41" pageNumber="412" specimenCount="1" specimenCount-female="1" stateProvince="Pahang">
<emphasis id="B96FEA9BFF943B27121FFEB9FB7FFC3B" bold="true" box="[1066,1227,983,1007]" pageId="41" pageNumber="412">
<collectingCountry id="F30C7619FF943B27121FFEB9FB73FC3B" box="[1066,1223,983,1007]" name="Malaysia" pageId="41" pageNumber="412">MALAYSIA</collectingCountry>
:
</emphasis>
<specimenCount id="9D1DFD00FF943B2712E2FEB9FAF5FC3B" box="[1239,1345,983,1007]" count="1" pageId="41" pageNumber="412" type="female">1 female</specimenCount>
,
<collectingRegion id="49DFF86BFF943B271364FEB9FA13FC3B" box="[1361,1447,983,1007]" country="Malaysia" name="Pahang" pageId="41" pageNumber="412">Pahang</collectingRegion>
,
<location id="8EC46052FF943B271518FE99FC0FFBDB" LSID="urn:lsid:plazi:treatment:03B2879FFF943B271575FF93FBA2F83A:8EC46052FF943B271518FE99FC0FFBDB" box="[813,955,1015,1039]" country="Malaysia" latitude="3.7166667" longLatPrecision="1307" longitude="101.73333" name="Fraser's Hill" pageId="41" pageNumber="412" stateProvince="Pahang">Frasers Hill</location>
,
<geoCoordinate id="EE2F504EFF943B2715FFFE9BFBA2FBDA" box="[970,1046,1013,1038]" degrees="3" direction="north" minutes="43" orientation="latitude" pageId="41" pageNumber="412" precision="925" value="3.7166667">3°43N</geoCoordinate>
<geoCoordinate id="EE2F504EFF943B271214FE9BFB3CFBDA" box="[1057,1160,1013,1038]" degrees="101" direction="east" minutes="44" orientation="longitude" pageId="41" pageNumber="412" precision="925" value="101.73333">101°44E</geoCoordinate>
,
<collectorName id="26EE535FFF943B2712A2FE99FB4AFBDB" box="[1175,1278,1015,1039]" pageId="41" pageNumber="412">S. Gotoh</collectorName>
,
<date id="FFA51049FF943B271339FE99FA1FFBDB" box="[1292,1451,1015,1039]" pageId="41" pageNumber="412" value="1986-04-29">
<collectingDate id="EFE1E9A1FF943B271339FE99FA1FFBDB" box="[1292,1451,1015,1039]" pageId="41" pageNumber="412" value="1986-04-29">29 April 1986</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF943B271501F979FC37FBFB" box="[820,899,1047,1071]" pageId="41" pageNumber="412">TYCN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B733CD4FF943B2715ADF979FCCFFBBA" collectionCode="T" country="Malaysia" county="Taiping" elevation="1100" latitude="4.8624997" location="Maxwell Hill" longLatPrecision="21" longitude="100.8" municipality="Bukit Larut" pageId="41" pageNumber="412" specimenCount="1" specimenCount-female="1" stateProvince="Perak">
<specimenCount id="9D1DFD00FF943B2715ADF979FC4FFBFB" box="[920,1019,1047,1071]" count="1" pageId="41" pageNumber="412" type="female">1 female</specimenCount>
,
<collectingRegion id="49DFF86BFF943B27123CF979FBFDFBFB" box="[1033,1097,1047,1071]" country="Malaysia" name="Perak" pageId="41" pageNumber="412">Perak</collectingRegion>
,
<collectingCounty id="62C54E05FF943B271262F979FB19FBFB" box="[1111,1197,1047,1071]" pageId="41" pageNumber="412">Taiping</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FF943B27128FF979FA8EFBFA" box="[1210,1338,1047,1071]" pageId="41" pageNumber="412">Bukit Larut</collectingMunicipality>
(
<location id="8EC46052FF943B27137EF979FCEEFB9B" LSID="urn:lsid:plazi:treatment:03B2879FFF943B271575FF93FBA2F83A:8EC46052FF943B27137EF979FCEEFB9B" country="Malaysia" county="Taiping" latitude="4.8624997" longLatPrecision="21" longitude="100.8" municipality="Bukit Larut" name="Maxwell Hill" pageId="41" pageNumber="412" stateProvince="Perak">Maxwell Hill</location>
),
<geoCoordinate id="EE2F504EFF943B27155BF95BFC56FB9A" box="[878,994,1077,1102]" degrees="4" direction="north" minutes="51" orientation="latitude" pageId="41" pageNumber="412" precision="15" seconds="45" value="4.8624997">4°5145″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF943B2715DAF95BFBCEFB9A" box="[1007,1146,1077,1102]" degrees="100" direction="east" minutes="48" orientation="longitude" pageId="41" pageNumber="412" precision="15" seconds="00" value="100.8">100°4800″E</geoCoordinate>
,
<quantity id="4CE39B6CFF943B2712B2F959FB55FB9A" box="[1159,1249,1079,1103]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="41" pageNumber="412" unit="m" value="1100.0">
<elevation id="0036D1BAFF943B2712B2F959FB55FB9A" box="[1159,1249,1079,1103]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="41" pageNumber="412" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
alt.,
<collectingMethod id="525A4E9EFF943B27132CF959FCEEFBBA" pageId="41" pageNumber="412">mercury light trap</collectingMethod>
,
<collectionCode id="ED0AAE4CFF943B27155DF939FCCFFBBA" box="[872,891,1111,1134]" name="Tavera, Department of Geology and Geophysics" pageId="41" pageNumber="412">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B733CD4FF943B2715BFF939FAB7FBBB" box="[906,1283,1111,1135]" collectingDate="1989-07-14" collectionCode="TYCN" country="Malaysia" location="Yasunaga" pageId="41" pageNumber="412" specimenCount="1">
<location id="8EC46052FF943B2715BFF939FC4CFBBB" LSID="urn:lsid:plazi:treatment:03B2879FFF943B271575FF93FBA2F83A:8EC46052FF943B2715BFF939FC4CFBBB" box="[906,1016,1111,1135]" country="Malaysia" name="Yasunaga" pageId="41" pageNumber="412">Yasunaga</location>
,
<date id="FFA51049FF943B271233F939FB2DFBBB" box="[1030,1177,1111,1135]" pageId="41" pageNumber="412" value="1989-07-14">
<collectingDate id="EFE1E9A1FF943B271233F939FB2DFBBB" box="[1030,1177,1111,1135]" pageId="41" pageNumber="412" value="1989-07-14">14 July 1989</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF943B27129EF939FB4FFBBB" box="[1195,1275,1111,1135]" pageId="41" pageNumber="412">TYCN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FF943B271324F939FA18FB5A" collectionCode="UV" country="Thailand" latitude="18.833334" location="Doi Pui" longLatPrecision="1273" longitude="98.88333" pageId="41" pageNumber="412" specimenCount="2" specimenCount-female="2" stateProvince="Chiang Mai">
<emphasis id="B96FEA9BFF943B271324F939FA18FBBB" bold="true" box="[1297,1452,1111,1135]" pageId="41" pageNumber="412">
<collectingCountry id="F30C7619FF943B271324F939FA13FBBB" box="[1297,1447,1111,1135]" name="Thailand" pageId="41" pageNumber="412">THAILAND</collectingCountry>
:
</emphasis>
<specimenCount id="9D1DFD00FF943B271518F919FC29FB5B" box="[813,925,1143,1167]" count="2" pageId="41" pageNumber="412" type="female">2 females</specimenCount>
,
<collectingRegion id="49DFF86BFF943B271599F919FB81FB5B" box="[940,1077,1143,1167]" country="Thailand" name="Chiang Mai" pageId="41" pageNumber="412">Chiang Mai</collectingRegion>
,
<location id="8EC46052FF943B271276F919FB2AFB5B" LSID="urn:lsid:plazi:treatment:03B2879FFF943B271575FF93FBA2F83A:8EC46052FF943B271276F919FB2AFB5B" box="[1091,1182,1143,1167]" country="Thailand" latitude="18.833334" longLatPrecision="1273" longitude="98.88333" name="Doi Pui" pageId="41" pageNumber="412" stateProvince="Chiang Mai">Doi Pui</location>
,
<geoCoordinate id="EE2F504EFF943B271298F91BFABFFB5A" box="[1197,1291,1141,1166]" degrees="18" direction="north" minutes="50" orientation="latitude" pageId="41" pageNumber="412" precision="925" value="18.833334">18°50N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF943B27132FF91BFAC1FB5A" box="[1306,1397,1141,1166]" degrees="98" direction="east" minutes="53" orientation="longitude" pageId="41" pageNumber="412" precision="925" value="98.88333">98°53E</geoCoordinate>
,
<collectionCode id="ED0AAE4CFF943B2713B0F919FA18FB5A" box="[1413,1452,1143,1166]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="41" pageNumber="412">UV</collectionCode>
</materialsCitation>
<materialsCitation id="3B733CD4FF943B271518F9F9FAB3FB7B" box="[813,1287,1175,1199]" collectingDate="2001-05-15" collectionCode="TYCN" collectorName="Sakurai" country="Thailand" pageId="41" pageNumber="412" specimenCount="1">
lighting,
<date id="FFA51049FF943B2715BAF9F9FB90FB7B" box="[911,1060,1175,1199]" pageId="41" pageNumber="412" value="2001-05-15">
<collectingDate id="EFE1E9A1FF943B2715BAF9F9FB90FB7B" box="[911,1060,1175,1199]" pageId="41" pageNumber="412" value="2001-05-15">15 May 2001</collectingDate>
</date>
, S.
<collectorName id="26EE535FFF943B27127AF9F9FB14FB7B" box="[1103,1184,1175,1199]" pageId="41" pageNumber="412">Sakurai</collectorName>
(
<collectionCode id="ED0AAE4CFF943B271285F9F9FB4BFB7B" box="[1200,1279,1175,1199]" pageId="41" pageNumber="412">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518F9B9FC0BFA5B" pageId="41" pageNumber="412" type="diagnosis">
<paragraph id="8BA43689FF943B271518F9B9FC0BFA5B" blockId="41.[813,1452,1239,1423]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518F9B9FC15FB3B" bold="true" box="[813,929,1239,1263]" pageId="41" pageNumber="412">Diagnosis.</emphasis>
This species is recognised readily by its moderate size and unique colour pattern on the dorsum, as mentioned in the key. Further external characters including the male genitalia, along with a habitus image, were provided by
<bibRefCitation id="EF8A4B78FF943B271518F839FBB6FABB" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[813,1026,1367,1391]" pageId="41" pageNumber="412" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng et al. (2004)</bibRefCitation>
. The female genitalia are described for the first time.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518F8D9FBD8F91B" pageId="41" pageNumber="412" type="description">
<paragraph id="8BA43689FF943B271518F8D9FBBFF95B" blockId="41.[813,1452,1463,1679]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518F8D9FC75FA1B" bold="true" box="[813,961,1463,1487]" pageId="41" pageNumber="412">Description.</emphasis>
Female genitalia (
<figureCitation id="13202A0CFF943B271297F8D9FA8DFA1B" box="[1186,1337,1463,1487]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="41" pageNumber="412">Figs. 17FH</figureCitation>
,
<figureCitation id="13202A0CFF943B27137DF8D9FAD7FA1B" box="[1352,1379,1463,1487]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="41" pageNumber="412">31</figureCitation>
LO): Sclerotised rings narrow, ovoid, relatively contiguous to each other mesally (
<figureCitation id="13202A0CFF943B271239F899FBC3F9DB" box="[1036,1143,1527,1551]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="41" pageNumber="412">Figs. 17F</figureCitation>
,
<figureCitation id="13202A0CFF943B2712B0F899FB02F9DB" box="[1157,1206,1527,1551]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="41" pageNumber="412">31N</figureCitation>
); posterior wall rather with rather wide dorsal structure and interramal lobes (
<figureCitation id="13202A0CFF943B2713B5FB79FCE9F99B" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="41" pageNumber="412">Fig. 31L</figureCitation>
); anterior part of interramal sclerite with a field of minute processes; dorsal structure with uniformly distributed spinules (
<figureCitation id="13202A0CFF943B2715A0FB19FC4BF95B" box="[917,1023,1655,1679]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="41" pageNumber="412">Fig. 31M</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FF943B271518FBD9FBD8F91B" blockId="41.[813,1132,1719,1743]" box="[813,1132,1719,1743]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FBD9FC6FF91B" bold="true" box="[813,987,1719,1743]" pageId="41" pageNumber="412">Measurements.</emphasis>
See
<tableCitation id="C6990332FF943B271227FBD9FBD2F91B" box="[1042,1126,1719,1743]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="41" pageNumber="412">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FB99FBA2F83A" pageId="41" pageNumber="412" type="discussion">
<paragraph id="8BA43689FF943B271518FB99FBA2F83A" blockId="41.[813,1452,1783,2031]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FB99FC2FF8DB" bold="true" box="[813,923,1783,1807]" pageId="41" pageNumber="412">Remarks.</emphasis>
Although this taxon was described as a member of
<taxonomicName id="4C1B4D0AFF943B27157BFA79FC07F8FB" authorityName="Knight" authorityYear="1917" box="[846,947,1815,1839]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF943B27157BFA79FC07F8FB" box="[846,947,1815,1839]" italics="true" pageId="41" pageNumber="412">Neolygus</emphasis>
</taxonomicName>
, the dorsal colour pattern, surface structures and form of the male and female genitalia suggest its best placement lies in
<taxonomicName id="4C1B4D0AFF943B2715C5FA39FB3DF8BB" authorityName="Yasunaga" authorityYear="2024" box="[1008,1161,1879,1903]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF943B2715C5FA39FB3DF8BB" box="[1008,1161,1879,1903]" italics="true" pageId="41" pageNumber="412">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF943B2712ACFA39FAA5F8BB" box="[1177,1297,1879,1903]" pageId="41" pageNumber="412" rank="genus">new genus</taxonomicNameLabel>
. This mirid is now known from S.
<collectingCountry id="F30C7619FF943B2715C8FA19FB88F85B" box="[1021,1084,1911,1935]" name="China" pageId="41" pageNumber="412">China</collectingCountry>
(
<collectingRegion id="49DFF86BFF943B27127DFA19FB14F85B" box="[1096,1184,1911,1935]" country="China" name="Yunnan" pageId="41" pageNumber="412">Yunnan</collectingRegion>
),
<collectingCountry id="F30C7619FF943B271285FA19FAB9F85B" box="[1200,1293,1911,1935]" name="Thailand" pageId="41" pageNumber="412">Thailand</collectingCountry>
and Peninsular
<collectingCountry id="F30C7619FF943B271518FAF9FC22F87B" box="[813,918,1943,1967]" name="Malaysia" pageId="41" pageNumber="412">Malaysia</collectingCountry>
. Its thermophilic habitat preference in tropical and subtropical zones also implies that it is not a genuine member of
<taxonomicName id="4C1B4D0AFF943B27159EFAB9FBBAF83B" authorityName="Knight" authorityYear="1917" box="[939,1038,2007,2031]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF943B27159EFAB9FBBAF83B" box="[939,1038,2007,2031]" italics="true" pageId="41" pageNumber="412">Neolygus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,194 @@
<document id="9A63A6D1892DE5C57ED99DE6D0065B75" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF943B2716DBFED3FBB7FD01" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris balicus Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="412" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="412" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A0FF01B7F5D8E96E0FC30337B2D206A3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D0ECBC480795ED4A258107301ACF7335">
<mods:title id="1C45843630F4E56003EA543FFB7CC3E4">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="AAA75AF7DDDD0E4935147506C4B817BF" type="personal">
<mods:role id="01AB352DC8899F6F8CC44FF1E5CB76D9">
<mods:roleTerm id="911DA77F823827ACDE6BD2557ED95F97">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31784D940A4803C812EC41F8EAE78B36">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="AA6CAE454770FEB40690E6141A02E49D">text</mods:typeOfResource>
<mods:relatedItem id="8680FE3E185E938B7AC7FB5768385889" type="host">
<mods:titleInfo id="501A488D79E779BA72B03057BB571AA9">
<mods:title id="D36F32EBF2861BCC84883A8A36B9373A">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="2827C9BC0BF459BE05706DC0C18DCD31">
<mods:date id="14CD7E0124ED7FE5782BA291F1571911">2024</mods:date>
<mods:detail id="D541C5C94E93ABB552CA225FA7B18CED" type="pubDate">
<mods:number id="6CBC9DE139EAF0050264C1D3D7678E3F">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="733F81BF73676A980F50CCE8728D1688" type="volume">
<mods:number id="BD5C4CEA4B8D2503677DE9F5782F5FEF">72</mods:number>
</mods:detail>
<mods:extent id="54C2DF72D5100B401FE5A357759AFB4B" unit="page">
<mods:start id="F6F34D1951FE7A7B712DD98B94862951">371</mods:start>
<mods:end id="B712D55442D7EAFB4EF2F9F77C8514D0">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6EBAE04C0BB00023BC6178AAE3859835">journal article</mods:classification>
<mods:identifier id="612DC36ACCACD9EFEC05B5C7A64F8CBE" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="D36E1E329C71DB744BFA296CA74C3E22" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="6305366ACC783A3BDAE96B4B7EFA8182" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="9483D8D5403E9289857C6D473EDC1AF7" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF943B2716DBFED3FBB7FD01" ID-DOI="http://doi.org/10.5281/zenodo.14683044" ID-Zenodo-Dep="14683044" LSID="urn:lsid:plazi:treatment:03B2879FFF943B2716DBFED3FBB7FD01" httpUri="http://treatment.plazi.org/id/03B2879FFF943B2716DBFED3FBB7FD01" lastPageNumber="412" pageId="41" pageNumber="412">
<subSubSection id="C3016502FF943B2716DBFED3FDC7FC01" box="[238,627,957,981]" pageId="41" pageNumber="412" type="nomenclature">
<paragraph id="8BA43689FF943B2716DBFED3FDC7FC01" blockId="41.[238,627,957,1013]" box="[238,627,957,981]" pageId="41" pageNumber="412">
<heading id="D0EC81E5FF943B2716DBFED3FDC7FC01" bold="true" box="[238,627,957,981]" centered="true" fontSize="10" level="2" pageId="41" pageNumber="412" reason="2">
<emphasis id="B96FEA9BFF943B2716DBFED3FDC7FC01" bold="true" box="[238,627,957,981]" pageId="41" pageNumber="412">
<taxonomicName id="4C1B4D0AFF943B2716DBFED3FE6BFC01" authorityName="Yasunaga" authorityYear="2024" box="[238,479,957,981]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="balicus" status="sp. nov.">
<emphasis id="B96FEA9BFF943B2716DBFED3FE6BFC01" bold="true" box="[238,479,957,981]" italics="true" pageId="41" pageNumber="412">Paralygocoris balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF943B2717DBFED3FDC7FC01" box="[494,627,957,981]" pageId="41" pageNumber="412" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B27170EFEB3FD92FC21" box="[315,550,989,1013]" pageId="41" pageNumber="412" type="description">
<paragraph id="8BA43689FF943B27170EFEB3FD92FC21" blockId="41.[238,627,957,1013]" box="[315,550,989,1013]" pageId="41" pageNumber="412">
(
<figureCitation id="13202A0CFF943B271776FEB3FE28FC21" box="[323,412,989,1013]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="41" pageNumber="412">Figs. 12</figureCitation>
HJ, 16, 33)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271644F973FE80FB61" pageId="41" pageNumber="412" type="materials_examined">
<paragraph id="8BA43689FF943B271644F973FE80FB61" blockId="41.[113,754,1053,1205]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271644F973FE9DFBE1" bold="true" box="[113,297,1053,1077]" pageId="41" pageNumber="412">Type material.</emphasis>
<materialsCitation id="3B733CD4FF943B271700F973FDC8FBA1" collectingDate="1996-03-08" collectingDateMax="1996-03-12" collectingDateMin="1996-03-08" collectionCode="ZRC, AMNH" country="Indonesia" latitude="-8.283334" location="Candikuning" longLatPrecision="1301" longitude="115.166664" municipality="Tabanan" pageId="41" pageNumber="412" specimenCode="PBI 00378788" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<emphasis id="B96FEA9BFF943B271700F973FE10FBE1" bold="true" box="[309,420,1053,1077]" pageId="41" pageNumber="412">
<typeStatus id="54A0882BFF943B271700F973FE10FBE1" box="[309,420,1053,1077]" pageId="41" pageNumber="412" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFF943B271436F973FD19FBE1" bold="true" box="[515,685,1053,1077]" pageId="41" pageNumber="412">
<collectingCountry id="F30C7619FF943B271436F973FD13FBE1" box="[515,679,1053,1077]" name="Indonesia" pageId="41" pageNumber="412">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFF943B27148CF973FD5FFBE1" box="[697,747,1053,1077]" country="Indonesia" name="Bali" pageId="41" pageNumber="412">Bali</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FF943B271644F953FF66FB81" box="[113,210,1085,1109]" pageId="41" pageNumber="412">Tabanan</collectingMunicipality>
,
<location id="8EC46052FF943B2716D5F953FEC4FB81" LSID="urn:lsid:plazi:treatment:03B2879FFF943B2716DBFED3FBB7FD01:8EC46052FF943B2716D5F953FEC4FB81" box="[224,368,1085,1109]" country="Indonesia" latitude="-8.283334" longLatPrecision="1301" longitude="115.166664" municipality="Tabanan" name="Candikuning" pageId="41" pageNumber="412" stateProvince="Bali">Candikuning</location>
,
<geoCoordinate id="EE2F504EFF943B27174AF952FE7CFB81" box="[383,456,1084,1109]" degrees="8" direction="south" minutes="17" orientation="latitude" pageId="41" pageNumber="412" precision="925" value="-8.283334">8°17S</geoCoordinate>
<geoCoordinate id="EE2F504EFF943B2717E6F952FD8FFB81" box="[467,571,1084,1109]" degrees="115" direction="east" minutes="10" orientation="longitude" pageId="41" pageNumber="412" precision="925" value="115.166664">115°10E</geoCoordinate>
, local collector,
<date id="FFA51049FF943B271644F933FE93FBA1" box="[113,295,1117,1141]" pageId="41" pageNumber="412" value="1996-03-08" valueMax="1996-03-12" valueMin="1996-03-08">
<collectingDate id="EFE1E9A1FF943B271644F933FE93FBA1" box="[113,295,1117,1141]" pageId="41" pageNumber="412" value="1996-03-08" valueMax="1996-03-12" valueMin="1996-03-08">812 March 1996</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF943B271706F933FEDDFBA1" box="[307,361,1117,1141]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="41" pageNumber="412">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFF943B271749F933FE79FBA1" box="[380,461,1117,1141]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="41" pageNumber="412">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FF943B2717EEF933FDC1FBA1" box="[475,629,1117,1141]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="41" pageNumber="412">PBI 00378788</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FF943B2714B1F930FE9BFB61" collectingDate="1996-03-08" collectingDateMax="1996-03-12" collectingDateMin="1996-03-08" collectionCode="AMNH, TYCN" country="Indonesia" latitude="-8.283334" location="Candikuning" longLatPrecision="1301" longitude="115.166664" municipality="Tabanan" pageId="41" pageNumber="412" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Bali" typeStatus="paratype">
<typeStatus id="54A0882BFF943B2714B1F930FD59FBA2" box="[644,749,1118,1142]" pageId="41" pageNumber="412" type="paratype">Paratypes</typeStatus>
: INDONESIA:
<specimenCount id="9D1DFD00FF943B271725F913FED0FB41" box="[272,356,1149,1173]" count="3" pageId="41" pageNumber="412" type="male">3 males</specimenCount>
,
<specimenCount id="9D1DFD00FF943B27175BF913FE6CFB41" box="[366,472,1149,1173]" count="5" pageId="41" pageNumber="412" type="female">5 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFF943B27164DF9F3FF7AFB61" box="[120,206,1181,1205]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="41" pageNumber="412">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFF943B2716EEF9F3FE9CFB61" box="[219,296,1181,1205]" pageId="41" pageNumber="412">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271644F9B3FD35FA01" pageId="41" pageNumber="412" type="diagnosis">
<paragraph id="8BA43689FF943B271644F9B3FD35FA01" blockId="41.[113,753,1245,1493]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271644F9B3FF52FB21" bold="true" box="[113,230,1245,1269]" pageId="41" pageNumber="412">Diagnosis.</emphasis>
This new species can be distinguished from other congeners by the characters mentioned in the above key, in addition to the identical male and female genitalic structures, such as short, rather bulbous right paramere with small, blunt-tipped hypophysis (
<figureCitation id="13202A0CFF943B27174BF833FE69FAA1" box="[382,477,1373,1397]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Fig. 16C</figureCitation>
); slender, elongate vesical spicule recurved apically (
<figureCitation id="13202A0CFF943B2717B0F813FE44FA41" box="[389,496,1405,1429]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16D</figureCitation>
,
<figureCitation id="13202A0CFF943B2717CFF813FD8BFA41" box="[506,575,1405,1429]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33IL</figureCitation>
); elongate-ovoid sclerotised rings contiguous to each other mesally (
<figureCitation id="13202A0CFF943B2714B3F8F3FD58FA61" box="[646,748,1437,1461]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16F</figureCitation>
,
<figureCitation id="13202A0CFF943B271644F8D3FF17FA01" box="[113,163,1469,1493]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33K</figureCitation>
); and triangular dorsal structure (
<figureCitation id="13202A0CFF943B27143AF8D3FDC1FA01" box="[527,629,1469,1493]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">Fig. 33N</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271644F893FBD8FDC1" pageId="41" pageNumber="412" type="description">
<paragraph id="8BA43689FF943B271644F893FB42FE01" blockId="41.[113,753,1533,2037]" lastBlockId="41.[813,1452,189,469]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271644F893FF4EF9C1" bold="true" box="[113,250,1533,1557]" pageId="41" pageNumber="412">Description.</emphasis>
Body yellowish or stramineous brown, elongate oval, subparallel-sided; dorsum shining, impunctate, with uniformly distributed, brown, simple setae. Head shiny pale brown, with a pair of brown, symmetrical, C-shaped maculae in front; apices of clypeus and buccula infuscate. Antenna dark brown; female antennomere II with pale basal ⅓ and obscure subbasal ring (
<figureCitation id="13202A0CFF943B2717F3FBD3FD91F901" box="[454,549,1725,1749]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="41" pageNumber="412">Fig. 12J</figureCitation>
); extreme base of antennomere III creamy yellow. Labium yellowish brown, reaching middle of mesocoxa; apical half of segment IV darkened. Thoracic pleura pale brown; mesal stripe on scutellum not reaching apex; metathoracic scent efferent system as in
<figureCitation id="13202A0CFF943B2716C8FA33FEEAF8A1" box="[253,350,1885,1909]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">Fig. 33C</figureCitation>
. Hemelytron shiny pale brown; inner half of clavus fuscous; corium with a pair of dark spots apically; membrane pale smoky brown. All coxae and legs yellowish brown; metafemur with two brown apical rings subapically; each tarsomere III darkened. Abdomen pale brown. Male genitalia (
<figureCitation id="13202A0CFF943B27121BFDD3FB05FF01" box="[1070,1201,189,213]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Fig. 16AE</figureCitation>
,
<figureCitation id="13202A0CFF943B27128AFDD3FABDFF01" box="[1215,1289,189,213]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33GJ</figureCitation>
): Sensory lobe of left paramere weakly produced apically (
<figureCitation id="13202A0CFF943B271321FDB3FA13FF21" box="[1300,1447,221,245]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16A, B</figureCitation>
,
<figureCitation id="13202A0CFF943B271518FD93FCEAFEC1" box="[813,862,253,277]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33H</figureCitation>
); right paramere rather bulbous, with small, blunt-tipped hypophysis (
<figureCitation id="13202A0CFF943B271584FC73FBA5FEE1" box="[945,1041,285,309]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Fig. 16C</figureCitation>
). Vesica with elongate spicule recurved apically (
<figureCitation id="13202A0CFF943B2715A4FC53FC4BFE81" box="[913,1023,317,341]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16D</figureCitation>
,
<figureCitation id="13202A0CFF943B271239FC53FBE5FE81" box="[1036,1105,317,341]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33IL</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFF943B271312FC53FA13FE81" box="[1319,1447,317,341]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Fig. 16FH</figureCitation>
,
<figureCitation id="13202A0CFF943B271518FC33FC34FEA1" box="[813,896,349,373]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33KO</figureCitation>
): sclerotised rings elongate-ovoid, contiguous to each other mesally (
<figureCitation id="13202A0CFF943B2715FDFC13FB9BFE41" box="[968,1071,381,405]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16F</figureCitation>
,
<figureCitation id="13202A0CFF943B27120DFC13FBDEFE41" box="[1080,1130,381,405]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33K</figureCitation>
); posterior wall with and small interramal lobes (
<figureCitation id="13202A0CFF943B2715D6FCF3FBFEFE61" box="[995,1098,413,437]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="41" pageNumber="412">Figs. 16E</figureCitation>
,
<figureCitation id="13202A0CFF943B271261FCF3FB32FE61" box="[1108,1158,413,437]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">33N</figureCitation>
); dorsal structure triangular, densely covered with spinules (
<figureCitation id="13202A0CFF943B2712B1FCD3FB5DFE01" box="[1156,1257,445,469]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="41" pageNumber="412">Fig. 33O</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FF943B271518FC93FBD8FDC1" blockId="41.[813,1132,509,533]" box="[813,1132,509,533]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FC93FC6FFDC1" bold="true" box="[813,987,509,533]" pageId="41" pageNumber="412">Measurements.</emphasis>
See
<tableCitation id="C6990332FF943B271227FC93FBD2FDC1" box="[1042,1126,509,533]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="41" pageNumber="412">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FF53FA18FD81" box="[813,1452,573,598]" pageId="41" pageNumber="412" type="etymology">
<paragraph id="8BA43689FF943B271518FF53FA18FD81" blockId="41.[813,1452,573,598]" box="[813,1452,573,598]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FF53FC1BFD81" bold="true" box="[813,943,573,597]" pageId="41" pageNumber="412">Etymology.</emphasis>
Named for the
<typeStatus id="54A0882BFF943B271262FF50FB31FD82" box="[1111,1157,574,598]" pageId="41" pageNumber="412">type</typeStatus>
locality,
<collectingRegion id="49DFF86BFF943B2712DCFF53FAADFD81" box="[1257,1305,573,597]" country="Indonesia" name="Bali" pageId="41" pageNumber="412">Bali</collectingRegion>
; an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FF13FB73FD41" box="[813,1223,637,661]" pageId="41" pageNumber="412" type="distribution">
<paragraph id="8BA43689FF943B271518FF13FB73FD41" blockId="41.[813,1223,637,661]" box="[813,1223,637,661]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FF13FC0BFD41" bold="true" box="[813,959,637,661]" pageId="41" pageNumber="412">Distribution.</emphasis>
<collectingCountry id="F30C7619FF943B2715F2FF13FB9BFD41" box="[967,1071,637,661]" name="Indonesia" pageId="41" pageNumber="412">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFF943B27120AFF13FBD9FD41" box="[1087,1133,637,661]" country="Indonesia" name="Bali" pageId="41" pageNumber="412">Bali</collectingRegion>
Island).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF943B271518FFD3FBB7FD01" box="[813,1027,701,725]" pageId="41" pageNumber="412" type="biology_ecology">
<paragraph id="8BA43689FF943B271518FFD3FBB7FD01" blockId="41.[813,1027,701,725]" box="[813,1027,701,725]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271518FFD3FC3EFD01" bold="true" box="[813,906,701,725]" pageId="41" pageNumber="412">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,252 @@
<document id="D4E12CC05B891992F2B7621999C22B03" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF983B271573FB2AFEE4FC41" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris alboscutellatus Yasunaga 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="412" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="408" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1CE81B37B4FBD3DCF16E690CFA1F594E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2855439D5751EFE0AC4F1662D196AD03">
<mods:title id="7C75C30C865BC5E091000FFF68A88520">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="37A91D4168372AC8F8A53B79155F3BFB" type="personal">
<mods:role id="71EC6AA2B14DFD6CC6C88BDC531B8685">
<mods:roleTerm id="4D148970D9A62CE143171BEB94A6DC2F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3FDC814C46C32E221FAF36359344ECAF">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="BE032EA65E4EC73468D5F6AFBAC5DD14">text</mods:typeOfResource>
<mods:relatedItem id="9F0BB9D3E739CE6F0E3F439E97CD01D7" type="host">
<mods:titleInfo id="8C5453C4782CFCBAD175F60EDAC70FE0">
<mods:title id="4EB430A4F35239C5A7C82C23974D4AB2">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0AC1BEEB38ECD99B09F6E9EBE4E60A24">
<mods:date id="5A68A7924B391B1C3E7F5FA2EB720E6E">2024</mods:date>
<mods:detail id="B9873A7A1796D72E6734666EB29E7D65" type="pubDate">
<mods:number id="EAAE8848F9E4B8B9FD8F070FC139102E">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="D036912AD89B85C174D62817B64BECC1" type="volume">
<mods:number id="B5519E5C6920FA31EC20D3918B3AE30B">72</mods:number>
</mods:detail>
<mods:extent id="29C31256C52D752435AEDE341B674989" unit="page">
<mods:start id="6269BA03FB7A41BFFC4C1C8EE30D819A">371</mods:start>
<mods:end id="87C0EF60AEED94F77A8ABCB9A4503FF4">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7B0CAC6B28F671C90B13CD92C6BD11F6">journal article</mods:classification>
<mods:identifier id="61D1033F7CEF85BCD47C767BD2AE3CC0" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="B6C04EBC7B9AB7BDF636E5215F55FF4C" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="B651D52D3F1293AEE79580D6276174EB" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="BC9B84B508D2F5292D0E856ABD79D1EF" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF983B271573FB2AFEE4FC41" LSID="urn:lsid:plazi:treatment:03B2879FFF983B271573FB2AFEE4FC41" httpUri="http://treatment.plazi.org/id/03B2879FFF983B271573FB2AFEE4FC41" lastPageId="41" lastPageNumber="412" pageId="37" pageNumber="408">
<subSubSection id="C3016502FF983B2B1573FB2AFB07F9A8" pageId="37" pageNumber="408" type="nomenclature">
<paragraph id="8BA43689FF983B2B1573FB2AFB07F9A8" blockId="37.[838,1426,1604,1693]" pageId="37" pageNumber="408">
<heading id="D0EC81E5FF983B2B1573FB2AFB07F9A8" bold="true" centered="true" fontSize="10" level="2" pageId="37" pageNumber="408" reason="2">
<emphasis id="B96FEA9BFF983B2B1573FB2AFB07F9A8" bold="true" pageId="37" pageNumber="408">
<taxonomicName id="4C1B4D0AFF983B2B1573FB2AFAE3F988" authority="(Carvalho, 1987)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Carvalho" baseAuthorityYear="1987" box="[838,1367,1604,1628]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="alboscutellatus" status="comb. nov.">
<emphasis id="B96FEA9BFF983B2B1573FB2AFB3FF988" bold="true" box="[838,1163,1604,1628]" italics="true" pageId="37" pageNumber="408">Paralygocoris alboscutellatus</emphasis>
(
<bibRefCitation id="EF8A4B78FF983B2B12AEFB2AFAFBF988" author="Carvalho JCM" box="[1179,1359,1604,1628]" pageId="37" pageNumber="408" pagination="137 - 153" refId="ref23627" refString="Carvalho JCM (1987) Prolygus n. gen. with descriptions of new species and redescription of known ones from Papua New Guinea (Hemiptera, Miridae). Revista Brasileira de Biologia, 47: 137 - 153." type="journal article" year="1987">Carvalho, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1351FB2BFB07F9A8" pageId="37" pageNumber="408" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1210FBEBFB07F949" box="[1061,1203,1669,1693]" pageId="37" pageNumber="408" type="description">
<paragraph id="8BA43689FF983B2B1210FBEBFB07F949" blockId="37.[838,1426,1604,1693]" box="[1061,1203,1669,1693]" pageId="37" pageNumber="408">
(
<figureCitation id="13202A0CFF983B2B1218FBEBFB18F949" box="[1069,1196,1669,1693]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="37" pageNumber="408">Fig. 32AF</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518FBADFB03F921" pageId="37" pageNumber="408" type="reference_group">
<paragraph id="8BA43689FF983B2B1518FBADFB03F921" blockId="37.[813,1452,1731,1782]" pageId="37" pageNumber="408">
<treatmentCitationGroup id="AB0B11A7FF983B2B1518FBADFB03F921" pageId="37" pageNumber="408">
<treatmentCitation id="0ABA1098FF983B2B1518FBADFB51F90C" author="Carvalho JCM" box="[813,1253,1731,1753]" page="140" pageId="37" pageNumber="408" year="1987">
<taxonomicName id="4C1B4D0AFF983B2B1518FBADFB51F90C" ID-CoL="4MRZ6" authority="Carvalho, 1987: 140" authorityName="Carvalho" authorityPageNumber="140" authorityYear="1987" box="[813,1253,1731,1753]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="alboscutellatus">
<emphasis id="B96FEA9BFF983B2B1518FBADFBAEF90C" box="[813,1050,1731,1752]" italics="true" pageId="37" pageNumber="408">Prolygus alboscutellatus</emphasis>
<bibRefCitation id="EF8A4B78FF983B2B122AFBADFB51F90C" author="Carvalho JCM" box="[1055,1253,1731,1753]" pageId="37" pageNumber="408" pagination="137 - 153" refId="ref23627" refString="Carvalho JCM (1987) Prolygus n. gen. with descriptions of new species and redescription of known ones from Papua New Guinea (Hemiptera, Miridae). Revista Brasileira de Biologia, 47: 137 - 153." type="journal article" year="1987">Carvalho, 1987: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 58;
<treatmentCitation id="0ABA1098FF983B2B1378FBAAFC41F922" ID-DOI="http://doi.org/10.5281/zenodo.5061953" ID-Zenodo-Dep="5061953" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" httpUri="http://treatment.plazi.org/id/039D4113FF996902BAF5FF49FB6D7D28" page="359" pageId="37" pageNumber="408" year="2018">
<bibRefCitation id="EF8A4B78FF983B2B1378FBAAFC41F922" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" pageId="37" pageNumber="408" pagination="357 - 387" refId="ref24824" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2018) Review of the plant bug genus Prolygus and related mirine taxa from eastern Asia (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae, 58: 357 - 387." type="journal article" year="2018">Yasunaga et al., 2018: 359</bibRefCitation>
</treatmentCitation>
, figs. 43, 44 (note).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518FA70FBBFF842" pageId="37" pageNumber="408" type="materials_examined">
<paragraph id="8BA43689FF983B2B1518FA70FBBFF842" blockId="37.[813,1452,1822,1942]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1518FA70FBB9F8E2" bold="true" box="[813,1037,1822,1846]" pageId="37" pageNumber="408">Material examined.</emphasis>
<materialsCitation id="3B733CD4FF983B2B1221FA70FCE0F8A2" collectionCode="UV" country="Indonesia" elevation="1400" latitude="3.1833334" location="North of Toba Lake" longLatPrecision="1307" longitude="98.5" municipality="Brastagi" pageId="37" pageNumber="408" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara">
<emphasis id="B96FEA9BFF983B2B1221FA70FB01F8E2" bold="true" box="[1044,1205,1822,1846]" pageId="37" pageNumber="408">
<collectingCountry id="F30C7619FF983B2B1221FA70FB1BF8E2" box="[1044,1199,1822,1846]" name="Indonesia" pageId="37" pageNumber="408">INDONESIA</collectingCountry>
:
</emphasis>
N.
<collectingRegion id="49DFF86BFF983B2B12EBFA70FA8FF8E2" box="[1246,1339,1822,1846]" country="Indonesia" name="Sumatera Utara" pageId="37" pageNumber="408">Sumatra</collectingRegion>
:
<specimenCount id="9D1DFD00FF983B2B1372FA70FA13F8E2" box="[1351,1447,1822,1846]" count="1" pageId="37" pageNumber="408" type="female">1 female</specimenCount>
,
<collectingMunicipality id="6BC0ACF3FF983B2B1518FA50FC3DF882" box="[813,905,1854,1878]" pageId="37" pageNumber="408">Brastagi</collectingMunicipality>
,
<location id="8EC46052FF983B2B15A7FA50FBDCF882" LSID="urn:lsid:plazi:treatment:03B2879FFF983B271573FB2AFEE4FC41:8EC46052FF983B2B15A7FA50FBDCF882" box="[914,1128,1854,1878]" country="Indonesia" latitude="3.1833334" longLatPrecision="1307" longitude="98.5" municipality="Brastagi" name="North of Toba Lake" pageId="37" pageNumber="408" stateProvince="Sumatera Utara">North of Toba Lake</location>
,
<geoCoordinate id="EE2F504EFF983B2B1247FA53FB09F882" box="[1138,1213,1853,1878]" degrees="3" direction="north" minutes="11" orientation="latitude" pageId="37" pageNumber="408" precision="925" value="3.1833334">3°11N</geoCoordinate>
<geoCoordinate id="EE2F504EFF983B2B12F1FA53FAA8F882" box="[1220,1308,1853,1878]" degrees="98" direction="east" minutes="30" orientation="longitude" pageId="37" pageNumber="408" precision="925" value="98.5">98°30E</geoCoordinate>
,
<quantity id="4CE39B6CFF983B2B1313FA50FACAF882" box="[1318,1406,1854,1878]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="408" unit="m" value="1400.0">
<elevation id="0036D1BAFF983B2B1313FA50FACAF882" box="[1318,1406,1854,1878]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="37" pageNumber="408" unit="m" value="1400.0">1,400 m</elevation>
</quantity>
alt.,
<collectionCode id="ED0AAE4CFF983B2B1518FA31FCE0F8A2" box="[813,852,1887,1910]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="37" pageNumber="408">UV</collectionCode>
</materialsCitation>
light trap,
<collectionCode id="ED0AAE4CFF983B2B15F3FA31FC6DF8A2" box="[966,985,1887,1910]" name="Tavera, Department of Geology and Geophysics" pageId="37" pageNumber="408">T</collectionCode>
.
<materialsCitation id="3B733CD4FF983B2B15D6FA31FBB3F842" collectingDate="1989-12-08" collectionCode="AMNH, TYCN" country="Indonesia" location="Yasunaga" pageId="37" pageNumber="408" specimenCode="PBI 00380625" specimenCount="1" stateProvince="Sumatra">
<location id="8EC46052FF983B2B15D6FA31FBFBF8A3" LSID="urn:lsid:plazi:treatment:03B2879FFF983B271573FB2AFEE4FC41:8EC46052FF983B2B15D6FA31FBFBF8A3" box="[995,1103,1887,1911]" country="Indonesia" name="Yasunaga" pageId="37" pageNumber="408" stateProvince="Sumatra">Yasunaga</location>
,
<date id="FFA51049FF983B2B126DFA30FAA0F8A2" box="[1112,1300,1886,1910]" pageId="37" pageNumber="408" value="1989-12-08">
<collectingDate id="EFE1E9A1FF983B2B126DFA30FAA0F8A2" box="[1112,1300,1886,1910]" pageId="37" pageNumber="408" value="1989-12-08">8 December 1989</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF983B2B1314FA30FAC0F8A2" box="[1313,1396,1886,1910]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="37" pageNumber="408">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FF983B2B13B7FA30FC2DF842" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="37" pageNumber="408">PBI 00380625</specimenCode>
) (
<collectionCode id="ED0AAE4CFF983B2B1585FA10FC4BF842" box="[944,1023,1918,1942]" pageId="37" pageNumber="408">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518FAD0FC80F7C2" pageId="37" pageNumber="408" type="distribution">
<paragraph id="8BA43689FF983B2B1518FAD0FBA7F822" blockId="37.[813,1451,1982,2070]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1518FAD0FC71F802" bold="true" box="[813,965,1982,2006]" pageId="37" pageNumber="408">Distribution.</emphasis>
<collectingCountry id="F30C7619FF983B2B15E5FAD0FB88F802" box="[976,1084,1982,2006]" name="Indonesia" pageId="37" pageNumber="408">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFF983B2B127AFAD0FB05F802" box="[1103,1201,1982,2006]" country="Indonesia" name="Sumatera Utara" pageId="37" pageNumber="408">Sumatra</collectingRegion>
),
<collectingCountry id="F30C7619FF983B2B12FDFAD1FA13F802" box="[1224,1447,1982,2006]" name="Papua New Guinea" pageId="37" pageNumber="408">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F30C7619FF983B2B1518FAB0FC11F822" box="[813,933,2014,2038]" name="Philippines" pageId="37" pageNumber="408">Philippines</collectingCountry>
(Negros).
</paragraph>
<paragraph id="8BA43689FF983B2B1518FA91FC80F7C2" blockId="37.[813,1451,1982,2070]" box="[813,820,2047,2070]" pageId="37" pageNumber="408">.</paragraph>
</subSubSection>
<caption id="DF646601FF9B3B2816BDFAF6FCD3F7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683012" ID-Zenodo-Dep="14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="38" pageNumber="409" startId="38.[136,171,1944,1966]" targetBox="[136,1474,251,1929]" targetPageId="38" targetType="figure">
<paragraph id="8BA43689FF9B3B2816BDFAF6FCD3F7D1" blockId="38.[136,1475,1944,2053]" pageId="38" pageNumber="409">
Fig. 25. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF9B3B28141EFAF6FCF9F879" authorityName="Yasunaga" authorityYear="2024" box="[555,845,1944,1965]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="38" pageNumber="409" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFF9B3B28141EFAF6FCF9F879" box="[555,845,1944,1965]" italics="true" pageId="38" pageNumber="409">Euphorbialygus rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9B3B28156EFAF7FC66F87A" box="[859,978,1944,1966]" pageId="38" pageNumber="409" rank="species">new species</taxonomicNameLabel>
. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall.
</paragraph>
</caption>
<caption id="DF646601FF9A3B291644FAF6FAD2F7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683014" ID-Zenodo-Dep="14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="39" pageNumber="410" startId="39.[113,148,1944,1966]" targetBox="[113,1451,251,1929]" targetPageId="39" targetType="figure">
<paragraph id="8BA43689FF9A3B291644FAF6FAD2F7D1" blockId="39.[113,1452,1944,2053]" pageId="39" pageNumber="410">
Fig. 26. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF9A3B29143AFAF6FCA7F879" authorityName="Yasunaga" authorityYear="2024" box="[527,787,1944,1965]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="410" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFF9A3B29143AFAF6FCA7F879" box="[527,787,1944,1965]" italics="true" pageId="39" pageNumber="410">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9A3B291515FAF7FC27F87A" box="[800,915,1944,1966]" pageId="39" pageNumber="410" rank="species">new species</taxonomicNameLabel>
(AI) and
<taxonomicName id="4C1B4D0AFF9A3B2915CAFAF7FBECF87A" authorityName="Yasunaga" authorityYear="2024" box="[1023,1112,1945,1966]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="410" phylum="Arthropoda" rank="species" species="omiae" status="new species">
<emphasis id="B96FEA9BFF9A3B2915CAFAF7FBECF87A" box="[1023,1112,1945,1966]" italics="true" pageId="39" pageNumber="410">H. omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9A3B291250FAF7FB6CF87A" box="[1125,1240,1944,1966]" pageId="39" pageNumber="410" rank="species">new species</taxonomicNameLabel>
(JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view.
</paragraph>
</caption>
<caption id="DF646601FF953B2616BDFAC9FBA9F822" ID-DOI="http://doi.org/10.5281/zenodo.14683018" ID-Zenodo-Dep="14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="40" pageNumber="411" startId="40.[136,171,1959,1981]" targetBox="[136,1474,265,1943]" targetPageId="40" targetType="figure">
<paragraph id="8BA43689FF953B2616BDFAC9FBA9F822" blockId="40.[136,1475,1959,2038]" pageId="40" pageNumber="411">
Fig. 27. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF953B261402FAC9FCF4F868" authorityName="Yasunaga" authorityYear="2024" box="[567,832,1959,1980]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="411" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFF953B261402FAC9FCF4F868" box="[567,832,1959,1980]" italics="true" pageId="40" pageNumber="411">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF953B26157AFAC9FC72F869" box="[847,966,1959,1981]" pageId="40" pageNumber="411" rank="species">new species</taxonomicNameLabel>
(AI) and
<taxonomicName id="4C1B4D0AFF953B26120FFAC9FB21F868" authorityName="Yasunaga" authorityYear="2024" box="[1082,1173,1959,1980]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="40" pageNumber="411" phylum="Arthropoda" rank="species" species="omiae" status="new species">
<emphasis id="B96FEA9BFF953B26120FFAC9FB21F868" box="[1082,1173,1959,1980]" italics="true" pageId="40" pageNumber="411">H. omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF953B261290FAC9FAA8F869" box="[1189,1308,1959,1981]" pageId="40" pageNumber="411" rank="species">new species</taxonomicNameLabel>
(JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring.
</paragraph>
</caption>
<subSubSection id="C3016502FF943B271644FDD3FEE4FC41" pageId="41" pageNumber="412" type="discussion">
<paragraph id="8BA43689FF943B271644FDD3FEE4FC41" blockId="41.[113,753,189,917]" pageId="41" pageNumber="412">
<emphasis id="B96FEA9BFF943B271644FDD3FF50FF01" bold="true" box="[113,228,189,213]" pageId="41" pageNumber="412">Remarks.</emphasis>
<bibRefCitation id="EF8A4B78FF943B2716C5FDD0FE4EFF01" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" box="[240,506,189,214]" pageId="41" pageNumber="412" pagination="357 - 387" refId="ref24824" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2018) Review of the plant bug genus Prolygus and related mirine taxa from eastern Asia (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae, 58: 357 - 387." type="journal article" year="2018">Yasunaga et al. (2018)</bibRefCitation>
retained the original generic placement of this taxon in
<taxonomicName id="4C1B4D0AFF943B2717C7FDB3FDE6FF21" authorityName="Carvalho" authorityYear="1987" box="[498,594,221,245]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF943B2717C7FDB3FDE6FF21" box="[498,594,221,245]" italics="true" pageId="41" pageNumber="412">Prolygus</emphasis>
</taxonomicName>
(
<typeStatus id="54A0882BFF943B271456FDB3FD26FF21" box="[611,658,221,245]" pageId="41" pageNumber="412">type</typeStatus>
species:
<taxonomicName id="4C1B4D0AFF943B271644FD90FE62FEC1" authority="Poppius, 1914" authorityName="Poppius" authorityYear="1914" box="[113,470,253,277]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B96FEA9BFF943B271644FD90FE9CFEC1" box="[113,296,254,277]" italics="true" pageId="41" pageNumber="412">Lygus papuanus</emphasis>
<bibRefCitation id="EF8A4B78FF943B271706FD93FE62FEC1" author="Poppius B" box="[307,470,253,277]" pageId="41" pageNumber="412" pagination="337 - 398" refId="ref24173" refString="Poppius B (1914) Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici, 12: 337 - 398." type="journal article" year="1914">Poppius, 1914</bibRefCitation>
</taxonomicName>
,
<figureCitation id="13202A0CFF943B2717D3FD93FDDAFEC1" box="[486,622,253,277]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32LO</figureCitation>
). However, the following morphological characters support its proper placement in
<taxonomicName id="4C1B4D0AFF943B27173AFC53FE1AFE81" authorityName="Yasunaga" authorityYear="2024" box="[271,430,317,341]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF943B27173AFC53FE1AFE81" box="[271,430,317,341]" italics="true" pageId="41" pageNumber="412">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF943B2717F5FC50FD8AFE82" box="[448,574,318,342]" pageId="41" pageNumber="412" rank="genus">new genus</taxonomicNameLabel>
: clavus lacking rows of lines along claval vein (
<figureCitation id="13202A0CFF943B2717E2FC33FDE9FEA1" box="[471,605,349,373]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32A, B</figureCitation>
); vesica with a single spicule that is coiled subapically and similar in basic colour pattern to
<taxonomicName id="4C1B4D0AFF943B2717BDFCF0FD88FE61" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Lu &amp; Zheng" baseAuthorityYear="2004" box="[392,572,413,437]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFF943B2717BDFCF0FD88FE61" box="[392,572,413,437]" italics="true" pageId="41" pageNumber="412">P. meridionalis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF8A4B78FF943B2714B4FCF3FF1EFE01" author="Carvalho JCM" pageId="41" pageNumber="412" pagination="137 - 153" refId="ref23627" refString="Carvalho JCM (1987) Prolygus n. gen. with descriptions of new species and redescription of known ones from Papua New Guinea (Hemiptera, Miridae). Revista Brasileira de Biologia, 47: 137 - 153." type="journal article" year="1987">Carvalho, 1987</bibRefCitation>
, p. 140, figs. 58); female genital chamber narrow (
<figureCitation id="13202A0CFF943B27164CFCB3FF55FE21" box="[121,225,477,501]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32D</figureCitation>
); and posterior wall with small interramal lobe and triangular, mesally continuous lateral lobe (
<figureCitation id="13202A0CFF943B2714BCFC93FF37FDE1" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32E, F</figureCitation>
).
<taxonomicName id="4C1B4D0AFF943B2716ADFF73FF4CFDE1" authorityName="Carvalho" authorityYear="1987" box="[152,248,541,565]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF943B2716ADFF73FF4CFDE1" box="[152,248,541,565]" italics="true" pageId="41" pageNumber="412">Prolygus</emphasis>
</taxonomicName>
congeners share the different character status, such as clavus at least basally with two or three rows of keeled lines parallel to claval vein (
<figureCitation id="13202A0CFF943B2717C9FF33FDEBFDA1" box="[508,607,605,629]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32L</figureCitation>
); vesica with 24 spicule or lobal sclerites (
<figureCitation id="13202A0CFF943B2717F6FF13FDFDFD41" box="[451,585,637,661]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32J, M</figureCitation>
); wider genital chamber (
<figureCitation id="13202A0CFF943B2716D7FFF3FEF9FD61" box="[226,333,669,693]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32N</figureCitation>
); and posterior wall with developed, wide interramal lobe and paired lateral lobes (
<figureCitation id="13202A0CFF943B27146AFFD3FD57FD01" box="[607,739,701,725]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="41" pageNumber="412">Fig. 32K, O</figureCitation>
). Based on these evidences,
<taxonomicName id="4C1B4D0AFF943B2717BCFFB3FD24FD21" authorityName="Carvalho" authorityYear="1987" box="[393,656,733,757]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="alboscutellatus">
<emphasis id="B96FEA9BFF943B2717BCFFB3FD24FD21" box="[393,656,733,757]" italics="true" pageId="41" pageNumber="412">Prolygus alboscutellatus</emphasis>
</taxonomicName>
is herein transferred to
<taxonomicName id="4C1B4D0AFF943B271737FF93FE2DFCC1" authorityName="Yasunaga" authorityYear="2024" box="[258,409,765,789]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF943B271737FF93FE2DFCC1" box="[258,409,765,789]" italics="true" pageId="41" pageNumber="412">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF943B271790FF90FDACFCC2" box="[421,536,766,790]" pageId="41" pageNumber="412" rank="genus">new genus</taxonomicNameLabel>
. Both
<taxonomicName id="4C1B4D0AFF943B27146CFF93FEA4FCE1" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Carvalho" baseAuthorityYear="1987" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="alboscutellatus">
<emphasis id="B96FEA9BFF943B27146CFF93FEA4FCE1" italics="true" pageId="41" pageNumber="412">Paralygocoris alboscutellatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFF943B271775FE70FE55FCE1" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Lu &amp; Zheng" baseAuthorityYear="2004" box="[320,481,797,821]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="41" pageNumber="412" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFF943B271775FE70FE55FCE1" box="[320,481,797,821]" italics="true" pageId="41" pageNumber="412">P. meridionalis</emphasis>
</taxonomicName>
are considered Sundaland species. However, they appear to be allopatric with each other (the former in Oriental-Wallacea and the latter in continental Eurasia).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,132 @@
<document id="34882E5BDA1B193443CDE28D734E06CA" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF983B2B1518FEB0FA1EF9C8" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris Yasunaga, 2024, new genus" docType="key" docVersion="2" lastPageNumber="408" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="408" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="9D3142F6A3F0BCD1AB71029F6B07A629" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C1FE4209A11E805F968B6FD05DE5526D">
<mods:title id="4A46D74868E46DE186B5644B52E9CA77">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="A7B0E37B3D1307525C9D27EFDEC0FEAC" type="personal">
<mods:role id="6309426604A555A3A198F0B03F1EF49B">
<mods:roleTerm id="1BBF6BCB14A7F52FC7DB32CAF59BADA7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0954B5EC9840F74E6AC352D7DD864157">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="9FCAC6ED08E41D03F6D5992BE5699E6E">text</mods:typeOfResource>
<mods:relatedItem id="4F4A687F71089F3BAC571027323DDA41" type="host">
<mods:titleInfo id="569D1F9F65003109A2B6C9487780B1FD">
<mods:title id="4740A073B6D128CE4E26DF39BCB0E5CC">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="290F7479EB1DC5540C166D279DDC6219">
<mods:date id="22E2EF925E7D542566709E12524BBF4B">2024</mods:date>
<mods:detail id="2C9DDEACDBE496C410CDE8EE69747CC2" type="pubDate">
<mods:number id="967E31071DE224881E0188C03BB97FFD">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="20B53B5D55EFAF368FB0420F81A6CD1A" type="volume">
<mods:number id="F58E1A6E5DFF00A42CD1915861B42222">72</mods:number>
</mods:detail>
<mods:extent id="17B828ABAA103FFF7F4A705A89B31ADA" unit="page">
<mods:start id="BD73710C68858D82F413FA609673C706">371</mods:start>
<mods:end id="A0AD874F815F4BA95C8359F721502948">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="836034204DBA7D467AB8949648A2E920">journal article</mods:classification>
<mods:identifier id="06ABF1ECF22577F1D63DFB3F6D4E9EAD" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="7A814D663FA9CAB5D164411A2CF0832A" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="7AF1DAFAE09DA10DDAE87B7D5554FD56" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="074C98B45FA81A77A2DD9904AB397540" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF983B2B1518FEB0FA1EF9C8" ID-DOI="http://doi.org/10.5281/zenodo.14683042" ID-Zenodo-Dep="14683042" LSID="urn:lsid:plazi:treatment:03B2879FFF983B2B1518FEB0FA1EF9C8" httpUri="http://treatment.plazi.org/id/03B2879FFF983B2B1518FEB0FA1EF9C8" lastPageNumber="408" pageId="37" pageNumber="408">
<subSubSection id="C3016502FF983B2B1518FEB0FAACFC22" box="[813,1304,990,1014]" pageId="37" pageNumber="408" type="nomenclature">
<paragraph id="8BA43689FF983B2B1518FEB0FAACFC22" blockId="37.[813,1304,990,1014]" box="[813,1304,990,1014]" pageId="37" pageNumber="408">
<heading id="D0EC81E5FF983B2B1518FEB0FAACFC22" bold="true" box="[813,1304,990,1014]" centered="true" fontSize="10" level="2" pageId="37" pageNumber="408" reason="2">
<emphasis id="B96FEA9BFF983B2B1518FEB0FAACFC22" bold="true" box="[813,1304,990,1014]" pageId="37" pageNumber="408">
Key to species of
<taxonomicName id="4C1B4D0AFF983B2B15CDFEB0FB25FC22" authorityName="Yasunaga" authorityYear="2024" box="[1016,1169,990,1014]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF983B2B15CDFEB0FB25FC22" bold="true" box="[1016,1169,990,1014]" italics="true" pageId="37" pageNumber="408">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1295FEB0FAACFC22" box="[1184,1304,990,1014]" pageId="37" pageNumber="408" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518F973FA1EF9C8" pageId="37" pageNumber="408" type="key">
<keyStep id="30EF2E2CFF983B2B1518F973FA1EFB5C" pageId="37" pageNumber="408">
<paragraph id="8BA43689FF983B2B1518F973FA1EFB9B" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F973FA1EFB9B" pageId="37" pageNumber="408">1. Vertex pale, without significant dark pattern; labium exceeding apex of mesocoxa, reaching middle or apex of metacoxal ...2</keyLead>
</paragraph>
<paragraph id="8BA43689FF983B2B1518F939FA1EFB5C" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F939FA1EFB5C" pageId="37" pageNumber="408"> Vertex with a pair of dark, symmetrical striae; labium shorter, not reaching apex of mesocoxa ..............................................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFF983B2B1518F9FEFA1EFA87" pageId="37" pageNumber="408">
<paragraph id="8BA43689FF983B2B1518F9FEFA1EFB28" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F9FEFA1EFB28" pageId="37" pageNumber="408">
2. Body larger, more than
<quantity id="4CE39B6CFF983B2B1274F9FEFB39FB72" box="[1089,1165,1168,1190]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="37" pageNumber="408" unit="mm" value="4.9">4.9 mm</quantity>
in total length; antennomere II fuscous; scutellum distinctly darkened mesally; currently considered endemic to
<collectingRegion id="49DFF86BFF983B2B1203F9A4FB39FB34" box="[1078,1165,1226,1248]" country="Indonesia" name="Sulawesi Utara" pageId="37" pageNumber="408">Sulawesi</collectingRegion>
............................................... ......................
<taxonomicName id="4C1B4D0AFF983B2B15E9F989FB43FB28" ID-CoL="3WPZ6" authority="(Poppius, 1914)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[988,1271,1254,1276]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="vittulatus" status="new combination">
<emphasis id="B96FEA9BFF983B2B15E9F989FBE5FB2F" box="[988,1105,1254,1276]" italics="true" pageId="37" pageNumber="408">P. vittulatus</emphasis>
(
<bibRefCitation id="EF8A4B78FF983B2B1255F988FB5BFB28" author="Poppius B" box="[1120,1263,1254,1276]" pageId="37" pageNumber="408" pagination="337 - 398" refId="ref24173" refString="Poppius B (1914) Zur Kenntnis der Indo-Australischen Lygus - Arten. Annales Historico-Naturales Musei Nationalis Hungarici, 12: 337 - 398." type="journal article" year="1914">Poppius, 1914</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1337F989FA1EFB28" box="[1282,1450,1254,1276]" pageId="37" pageNumber="408" rank="species">new combination</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FF983B2B1518F86AFA1EFA87" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F86AFA1EFA87" pageId="37" pageNumber="408">
Body shorter than
<quantity id="4CE39B6CFF983B2B1222F86DFBDDFACD" box="[1047,1129,1283,1305]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="37" pageNumber="408" unit="mm" value="4.5">4.5 mm</quantity>
; antennomere II and scutellum creamy yellow;
<collectingCountry id="F30C7619FF983B2B15C4F84FFB05FAE2" box="[1009,1201,1312,1334]" name="Papua New Guinea" pageId="37" pageNumber="408">Papua New Guinea</collectingCountry>
,
<collectingCountry id="F30C7619FF983B2B1289F84EFA9FFAE2" box="[1212,1323,1312,1334]" name="Philippines" pageId="37" pageNumber="408">Philippines</collectingCountry>
, Sumatra ..... ...........
<taxonomicName id="4C1B4D0AFF983B2B15ACF850FB42FA86" ID-CoL="4MRZ6" authority="(Carvalho, 1987)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Carvalho" baseAuthorityYear="1987" box="[921,1270,1341,1363]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="alboscutellatus" status="new combination">
<emphasis id="B96FEA9BFF983B2B15ACF850FBF2FA86" box="[921,1094,1341,1363]" italics="true" pageId="37" pageNumber="408">P. alboscutellatus</emphasis>
(
<bibRefCitation id="EF8A4B78FF983B2B1260F853FB5BFA86" author="Carvalho JCM" box="[1109,1263,1341,1363]" pageId="37" pageNumber="408" pagination="137 - 153" refId="ref23627" refString="Carvalho JCM (1987) Prolygus n. gen. with descriptions of new species and redescription of known ones from Papua New Guinea (Hemiptera, Miridae). Revista Brasileira de Biologia, 47: 137 - 153." type="journal article" year="1987">Carvalho, 1987</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1337F853FA1EFA87" box="[1282,1450,1341,1363]" pageId="37" pageNumber="408" rank="species">new combination</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFF983B2B1518F834FA1EF9C8" pageId="37" pageNumber="408">
<paragraph id="8BA43689FF983B2B1518F834FA1EFA12" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F834FA1EFA12" pageId="37" pageNumber="408">
3. Antennomere II almost uniformly fuscous; clavus entirely darkened; apical part of corium with dark V-shaped macula; S.
<collectingCountry id="F30C7619FF983B2B155BF8FDFC1EFA7D" box="[878,938,1427,1449]" name="China" pageId="37" pageNumber="408">China</collectingCountry>
, Indochina, W.
<collectingCountry id="F30C7619FF983B2B1272F8FDFB2BFA7D" box="[1095,1183,1427,1449]" name="Malaysia" pageId="37" pageNumber="408">Malaysia</collectingCountry>
............................................ .........
<taxonomicName id="4C1B4D0AFF983B2B15BAF8DFFB43FA11" ID-CoL="46LWB" authority="(Lu &amp; Zheng, 2004)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Lu &amp; Zheng" baseAuthorityYear="2004" box="[911,1271,1456,1478]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="meridionalis" status="new combination">
<emphasis id="B96FEA9BFF983B2B15BAF8DFFB90FA11" box="[911,1060,1456,1478]" italics="true" pageId="37" pageNumber="408">P. meridionalis</emphasis>
(Lu &amp;
<bibRefCitation id="EF8A4B78FF983B2B1244F8DEFB44FA11" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[1137,1264,1456,1478]" pageId="37" pageNumber="408" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng, 2004</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1337F8DFFA1EFA12" box="[1282,1450,1456,1478]" pageId="37" pageNumber="408" rank="species">new combination</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FF983B2B1518F8A3FA1EF9C8" blockId="37.[813,1452,1053,1564]" pageId="37" pageNumber="408">
<keyLead id="30EA95BCFF983B2B1518F8A3FA1EF9C8" pageId="37" pageNumber="408">
Basal part of antennomere II pale; inner part of clavus darkened; apical part of corium with a brown ovoid spot;
<collectingRegion id="49DFF86BFF983B2B1312F884FAFBF9D4" box="[1319,1359,1514,1536]" country="Indonesia" name="Bali" pageId="37" pageNumber="408">Bali</collectingRegion>
.............. .............................................................
<taxonomicName id="4C1B4D0AFF983B2B12F0FB69FA93F9CF" authorityName="Yasunaga" authorityYear="2024" box="[1221,1319,1542,1564]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="balicus" status="new species">
<emphasis id="B96FEA9BFF983B2B12F0FB69FA93F9CF" box="[1221,1319,1542,1564]" italics="true" pageId="37" pageNumber="408">P. balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1300FB69FA1EF9C8" box="[1333,1450,1542,1564]" pageId="37" pageNumber="408" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,233 @@
<document id="9A3825BB936B45C532C2C7BB9BDCC702" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF9C3B2B15E9FAF3FC08FC62" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Paralygocoris Yasunaga, 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="408" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="404" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2C8E4ED3F93B2BF9BF7E36E402C279A9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="732311850E08FDDB0D48D245E554FEC1">
<mods:title id="40F01D12867F6DCA948AAB9583A5D919">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="4DBF0E0AD4ADCE2BFDD7A4F444A42AAB" type="personal">
<mods:role id="A1F31832F08DE6957A1E50EDE4A8E86B">
<mods:roleTerm id="8017037BB9C0AA1036ABB849EBDF5708">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB04741857339B2FCE583BCE051E588C">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="36ADB3E2FAE4B31E9C9CAA53A2CD5F0E">text</mods:typeOfResource>
<mods:relatedItem id="88CFF99D591770FF02698DC49CE46CED" type="host">
<mods:titleInfo id="CE1878DAF38D4F0EFFEEA3FC183B6744">
<mods:title id="AF905B7512A33B7901C1729C14A02385">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="034923F4E76FDAF2630CE733746B0572">
<mods:date id="4A9DDE46DB324B4BF1A4FF6A2D97B6E3">2024</mods:date>
<mods:detail id="0F92F2AF81BCD4B1B4448706998B7F11" type="pubDate">
<mods:number id="FA1C64D3FD9F785E4E3F13EE86130346">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="0556C729E63F93C49F6D8A93BF4BC743" type="volume">
<mods:number id="73F2315DE927DF95AA279F59476297AD">72</mods:number>
</mods:detail>
<mods:extent id="0273165E713BEA315C86FBE75684254C" unit="page">
<mods:start id="5CEAED189795CD499553E97E8A5F7BFB">371</mods:start>
<mods:end id="0D5E3D20833346D4A045A3BC61EEAC8D">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EF04C6D225CE8D987085C1633F9EF50D">journal article</mods:classification>
<mods:identifier id="B1F32CA3F57131E9139C731AB0F0EF0A" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="2205B9AFDBAA1B69322EB233A81FCFE8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="B2FB9FB414723CD4F738DBE3A981936A" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="4B324D172023E73D61B99F99AB9AFCD7" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF9C3B2B15E9FAF3FC08FC62" LSID="urn:lsid:plazi:treatment:03B2879FFF9C3B2B15E9FAF3FC08FC62" httpUri="http://treatment.plazi.org/id/03B2879FFF9C3B2B15E9FAF3FC08FC62" lastPageId="37" lastPageNumber="408" pageId="33" pageNumber="404">
<subSubSection id="C3016502FF9C3B2F15E9FAF3FB48F862" box="[988,1276,1949,1974]" pageId="33" pageNumber="404" type="nomenclature">
<paragraph id="8BA43689FF9C3B2F15E9FAF3FB48F862" blockId="33.[988,1276,1949,1974]" box="[988,1276,1949,1974]" pageId="33" pageNumber="404">
<heading id="D0EC81E5FF9C3B2F15E9FAF3FB48F862" bold="true" box="[988,1276,1949,1974]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="404" reason="2">
<emphasis id="B96FEA9BFF9C3B2F15E9FAF3FB48F862" bold="true" box="[988,1276,1949,1974]" pageId="33" pageNumber="404">
<taxonomicName id="4C1B4D0AFF9C3B2F15E9FAF3FBC1F861" authorityName="Yasunaga" authorityYear="2024" box="[988,1141,1949,1973]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF9C3B2F15E9FAF3FBC1F861" bold="true" box="[988,1141,1949,1973]" italics="true" pageId="33" pageNumber="404">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F12B1FAF0FB48F862" box="[1156,1276,1950,1974]" pageId="33" pageNumber="404" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FAB0FAE0F822" box="[813,1364,2013,2038]" pageId="33" pageNumber="404" type="type_taxon">
<paragraph id="8BA43689FF9C3B2F1518FAB0FAE0F822" blockId="33.[813,1364,2013,2038]" box="[813,1364,2013,2038]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FAB0FC71F821" bold="true" box="[813,965,2013,2038]" pageId="33" pageNumber="404">
<typeStatus id="54A0882BFF9C3B2F1518FAB0FCD3F822" box="[813,871,2014,2038]" pageId="33" pageNumber="404">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1B4D0AFF9C3B2F15F8FAB0FB0FF822" authorityName="Yasunaga" authorityYear="2024" box="[973,1211,2014,2038]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="species" species="balicus" status="sp. nov.">
<emphasis id="B96FEA9BFF9C3B2F15F8FAB0FB0FF822" box="[973,1211,2014,2038]" italics="true" pageId="33" pageNumber="404">Paralygocoris balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F12FEFAB0FAE4F822" box="[1227,1360,2014,2038]" pageId="33" pageNumber="404" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<caption id="DF646601FF9F3B2C16B2FAF6FC7BF7D0" ID-DOI="http://doi.org/10.5281/zenodo.14683004" ID-Zenodo-Dep="14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="34" pageNumber="405" startId="34.[135,170,1944,1966]" targetBox="[135,1473,251,1928]" targetPageId="34" targetType="figure">
<paragraph id="8BA43689FF9F3B2C16B2FAF6FC7BF7D0" blockId="34.[135,1474,1944,2052]" pageId="34" pageNumber="405">
Fig. 22. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF9F3B2C1413FAF6FCABF879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[550,799,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="405" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFF9F3B2C1413FAF6FCABF879" box="[550,799,1944,1965]" italics="true" pageId="34" pageNumber="405">Chelyolygus fuhoshoensis</emphasis>
</taxonomicName>
(AD),
<taxonomicName id="4C1B4D0AFF9F3B2C1544FAF6FC52F87A" authorityName="Yasunaga" authorityYear="2024" box="[881,998,1944,1966]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="405" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFF9F3B2C1544FAF6FC52F87A" box="[881,998,1944,1966]" italics="true" pageId="34" pageNumber="405">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9F3B2C15C7FAF6FBD2F87A" box="[1010,1126,1944,1966]" pageId="34" pageNumber="405" rank="species">new species</taxonomicNameLabel>
(FI),
<taxonomicName id="4C1B4D0AFF9F3B2C129EFAF6FAF5F879" authorityName="Yasunaga" authorityYear="2024" box="[1195,1345,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="405" phylum="Arthropoda" rank="species" species="virgulatellus" status="new species">
<emphasis id="B96FEA9BFF9F3B2C129EFAF6FAF5F879" box="[1195,1345,1944,1965]" italics="true" pageId="34" pageNumber="405">C. virgulatellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9F3B2C137BFAF6FA75F87A" box="[1358,1473,1944,1966]" pageId="34" pageNumber="405" rank="species">new species</taxonomicNameLabel>
(JO) and
<taxonomicName id="4C1B4D0AFF9F3B2C16DAFADBFE4DF81E" authorityName="Yasunaga, Schwartz, &amp; Cherot" authorityYear="2002" box="[239,505,1973,1994]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="405" phylum="Arthropoda" rank="species" species="furvocarinatus">
<emphasis id="B96FEA9BFF9F3B2C16DAFADBFE4DF81E" box="[239,505,1973,1994]" italics="true" pageId="34" pageNumber="405">Apolygopsis furvocarinatus</emphasis>
</taxonomicName>
(E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe.
</paragraph>
</caption>
<caption id="DF646601FF9E3B2D1644FAF6FC69F7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683006" ID-Zenodo-Dep="14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="35" pageNumber="406" startId="35.[113,148,1944,1966]" targetBox="[113,1451,251,1929]" targetPageId="35" targetType="figure">
<paragraph id="8BA43689FF9E3B2D1644FAF6FC69F7D1" blockId="35.[113,1452,1944,2053]" pageId="35" pageNumber="406">
Fig. 23. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF9E3B2D1411FAF6FCA3F879" authorityName="Yasunaga" authorityYear="2024" box="[548,791,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="406" phylum="Arthropoda" rank="species" species="wolskiorum" status="sp. nov.">
<emphasis id="B96FEA9BFF9E3B2D1411FAF6FCA3F879" box="[548,791,1944,1965]" italics="true" pageId="35" pageNumber="406">Chelyolygus wolskiorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9E3B2D1513FAF7FC15F87A" box="[806,929,1944,1966]" pageId="35" pageNumber="406" rank="species">new species</taxonomicNameLabel>
. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall.
</paragraph>
</caption>
<caption id="DF646601FF993B2A16B2FAF6FBBEF7D1" ID-DOI="http://doi.org/10.5281/zenodo.14683010" ID-Zenodo-Dep="14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="36" pageNumber="407" startId="36.[135,170,1944,1966]" targetBox="[135,1473,251,1929]" targetPageId="36" targetType="figure">
<paragraph id="8BA43689FF993B2A16B2FAF6FBBEF7D1" blockId="36.[135,1474,1944,2053]" pageId="36" pageNumber="407">
Fig. 24. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF993B2A1402FAF6FC81F879" authorityName="Yasunaga" authorityYear="2024" box="[567,821,1944,1965]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="36" pageNumber="407" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFF993B2A1402FAF6FC81F879" box="[567,821,1944,1965]" italics="true" pageId="36" pageNumber="407">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF993B2A1570FAF7FC0BF87A" box="[837,959,1944,1966]" pageId="36" pageNumber="407" rank="species">new species</taxonomicNameLabel>
. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall.
</paragraph>
</caption>
<subSubSection id="C3016502FF983B2B1644FDD0FDA4FD62" pageId="37" pageNumber="408" type="diagnosis">
<paragraph id="8BA43689FF983B2B1644FDD0FDA4FD62" blockId="37.[113,752,190,694]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1644FDD0FF40FF02" bold="true" box="[113,244,190,214]" pageId="37" pageNumber="408">Diagnosis.</emphasis>
This new genus is externally similar to
<taxonomicName id="4C1B4D0AFF983B2B1644FDB1FECBFF22" authority="Reuter, 1875" authorityName="Reuter" authorityYear="1875" box="[113,383,222,246]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B1644FDB1FF54FF22" box="[113,224,223,246]" italics="true" pageId="37" pageNumber="408">Lygocoris</emphasis>
<bibRefCitation id="EF8A4B78FF983B2B16D9FDB1FECBFF22" author="Reuter OM" box="[236,383,222,246]" pageId="37" pageNumber="408" refId="ref24240" refString="Reuter OM (1875) Revisio critica Capsinarum praecipue Scandinaviae et Fenniae. Akademisk Afhandling, Helsingfors, 101 + 190 pp." type="book" year="1875">Reuter, 1875</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFF983B2B1788FDB0FD7DFF22" authority="Knight, 1917" authorityName="Knight" authorityYear="1917" box="[445,713,222,246]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B1788FDB0FD90FF22" box="[445,548,222,246]" italics="true" pageId="37" pageNumber="408">Neolygus</emphasis>
<bibRefCitation id="EF8A4B78FF983B2B1405FDB0FD7DFF22" author="Knight HH" box="[560,713,222,246]" pageId="37" pageNumber="408" pagination="555 - 645" refId="ref23971" refString="Knight HH (1917) A revision of the genus Lygus as it occurs on America North of Mexico, with biological data on the species from New York. Cornell University Agricultural Experiment Station Bulletin, 391: 555 - 645." type="journal article" year="1917">Knight, 1917</bibRefCitation>
</taxonomicName>
, as evidenced by previous placements of two known species (
<taxonomicName id="4C1B4D0AFF983B2B164FFC71FEE4FEE2" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[122,336,286,310]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFF983B2B164FFC71FEE4FEE2" box="[122,336,286,310]" italics="true" pageId="37" pageNumber="408">Lygocoris vittulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFF983B2B17BCFC70FDC8FEE2" authorityName="Lu &amp; Zheng" authorityYear="2004" box="[393,636,286,310]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFF983B2B17BCFC70FDC8FEE2" box="[393,636,286,310]" italics="true" pageId="37" pageNumber="408">Neolygus meridionalis</emphasis>
</taxonomicName>
which are herein transferred to
<taxonomicName id="4C1B4D0AFF983B2B1760FC50FE5AFE82" authorityName="Yasunaga" authorityYear="2024" box="[341,494,318,342]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF983B2B1760FC50FE5AFE82" box="[341,494,318,342]" italics="true" pageId="37" pageNumber="408">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B17C9FC51FDC0FE82" box="[508,628,318,342]" pageId="37" pageNumber="408" rank="genus">new genus</taxonomicNameLabel>
). However,
<taxonomicName id="4C1B4D0AFF983B2B1644FC30FEBDFEA2" authorityName="Yasunaga" authorityYear="2024" box="[113,265,350,374]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFF983B2B1644FC30FEBDFEA2" box="[113,265,350,374]" italics="true" pageId="37" pageNumber="408">Paralygocoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF983B2B1720FC31FE3CFEA3" box="[277,392,351,375]" pageId="37" pageNumber="408" rank="genus">new genus</taxonomicNameLabel>
, is distinct in having the following diagnostic characters: Body pale brown or ivory brown, with darkened clypeus, antennomere I, posterior pronotum, anterior-mesal scutellum, inner part of clavus and apical part of corium; tibiae with dark brown spines; hypophysis of right paramere reduced, small; vesica bilobate, with a single, elongate, apically curved spicule, lacking other noticeable sclerite; female genital chamber, along with sclerotised rings, narrow; posterior wall without lateral lobe; dorsal structure basally fused with interramal lobes; and interramal lobe spinulate only along posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1644FFB0FDA4F802" pageId="37" pageNumber="408" type="description">
<paragraph id="8BA43689FF983B2B1644FFB0FDA4F802" blockId="37.[113,752,734,2006]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1644FFB0FEBEFD22" bold="true" box="[113,266,734,758]" pageId="37" pageNumber="408">Description.</emphasis>
Body, elongate oval, moderate in size, subparallel-sided; not sexually dimorphic in general shape (usually female is more oval); basic colouration pale or ivory brown, with similar dark pattern as in
<figureCitation id="13202A0CFF983B2B1415FE50FF33FCA2" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="37" pageNumber="408">Fig. 12H, J, K, M, O</figureCitation>
; dorsal surface shining, with uniformly distributed, pale, simple, reclining setae.
<emphasis id="B96FEA9BFF983B2B1749FE10FE76FC42" bold="true" box="[380,450,894,918]" pageId="37" pageNumber="408">Head:</emphasis>
Shining, vertical in lateral view, almost glabrous; eyes large, contiguous to anterior margin of pronotal collar; vertex narrow, shallowly sulcate mesally, without noticeable basal transverse carina.
<emphasis id="B96FEA9BFF983B2B14B2FEB0FD44FC22" bold="true" box="[647,752,990,1014]" pageId="37" pageNumber="408">Antenna:</emphasis>
Widely darkened, slightly shorter than body, almost linear; antennomere I always dark, about as long as IV; antennomere II sometimes with pale base, longer than basal width of pronotum or labium; antennomeres III and IV filiform.
<emphasis id="B96FEA9BFF983B2B1644F910FF60FB42" bold="true" box="[113,212,1150,1174]" pageId="37" pageNumber="408">Labium:</emphasis>
Relatively short, reaching but not exceeding apex of mesocoxa, obviously shorter than metafemur.
<emphasis id="B96FEA9BFF983B2B14A7F9F0FD44FB62" bold="true" box="[658,752,1182,1206]" pageId="37" pageNumber="408">Thorax:</emphasis>
Pronotum shiny, always darkened posteriorly (except for pale posterior margin), with rather sparsely distributed, short, semi-erect setae, lacking noticeable punctures; calli reduced, weakly demarcated; collar shagreened, narrow, flat, about as thick as antennomere III; propleuron not margined nor carinate; metathoracic scent efferent system subtriangular, with relatively flat peritreme; scutellum smooth, more or less darkened mesad, impunctate, flat.
<emphasis id="B96FEA9BFF983B2B17E8F8F0FDDBFA62" bold="true" box="[477,623,1438,1462]" pageId="37" pageNumber="408">Hemelytron:</emphasis>
Moderately shining, with similar dark pattern on clavus and corium, shallowly and irregularly rugose, with uniformly distributed, pale, simple, reclining setae; lateral margin of exocorium (embolium) slightly rounded.
<emphasis id="B96FEA9BFF983B2B17F3FB70FDB1F9E2" bold="true" box="[454,517,1566,1590]" pageId="37" pageNumber="408">Legs:</emphasis>
Moderate in length; metafemur apically with two brown or reddish brown rings; tibial spines dark brown to fuscous; meta-tarsomere II longer than III; pretarsal structures as in
<figureCitation id="13202A0CFF983B2B1473FB10FD07F942" box="[582,691,1662,1686]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="37" pageNumber="408">Figs. 31K</figureCitation>
,
<specimenCount id="9D1DFD00FF983B2B14F5FB10FD44F942" box="[704,752,1662,1686]" count="33" pageId="37" pageNumber="408" type="generic">
<figureCitation id="13202A0CFF983B2B14F5FB10FD58F942" box="[704,748,1662,1686]" captionStart="Fig" captionStartId="48.[136,172,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@48.[136,1474,251,1929]" captionTargetPageId="48" captionText="Fig. 33. Scanning electron micrographs of Paralygocoris balicus, new species. A, B, head and thorax, left lateral- dorsal views; C, metathoracic scent efferent system; D, head and pronotum, dorsal view; E, metatarsus of each sex; F, pretarsal structure of metaleg; G, pygophore, dorsal view; H, left paramere; I, J, vesica; K, genital chamber; L, sclerotised ring; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683032" httpUri="https://zenodo.org/record/14683032/files/figure.png" pageId="37" pageNumber="408">33F</figureCitation>
.
</specimenCount>
<emphasis id="B96FEA9BFF983B2B1644FBF0FEAAF962" bold="true" box="[113,286,1694,1718]" pageId="37" pageNumber="408">Male genitalia:</emphasis>
Left paramere L-shaped, similar to that of
<taxonomicName id="4C1B4D0AFF983B2B1644FBD0FF60F902" authorityName="Knight" authorityYear="1917" box="[113,212,1726,1750]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B1644FBD0FF60F902" box="[113,212,1726,1750]" italics="true" pageId="37" pageNumber="408">Neolygus</emphasis>
</taxonomicName>
(e.g.,
<figureCitation id="13202A0CFF983B2B172FFBD0FE29F902" box="[282,413,1726,1750]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="37" pageNumber="408">Fig. 16A, B</figureCitation>
); hypophysis of right paramere reduced, small; vesica bilobate, with a single, elongate, apically curved spicule and long spinulate lobe, lacking other noticeable lobal-sclerite; phallotheca rather expanded, with a subapical, thin, semi-circular fin-like process.
<emphasis id="B96FEA9BFF983B2B14A8FA50FF61F8A2" bold="true" pageId="37" pageNumber="408">Female genitalia</emphasis>
: Genital chamber, along with sclerotised rings, narrow (
<figureCitation id="13202A0CFF983B2B16FFFA10FE9CF842" box="[202,296,1918,1942]" captionStart="Fig" captionStartId="26.[136,171,1900,1922]" captionTargetBox="[136,1474,324,1884]" captionTargetId="figure-8@26.[136,1474,324,1884]" captionTargetPageId="26" captionText="Fig. 16. Male (AD) and female (EG) genitalia of Paralygocoris balicus, new species. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="37" pageNumber="408">Fig. 16F</figureCitation>
); posterior wall without lateral lobe; dorsal structure basally fused with interramal lobes; interramal lobe spinulate only along posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518FDD0FAA0FEC2" pageId="37" pageNumber="408" type="etymology">
<paragraph id="8BA43689FF983B2B1518FDD0FAA0FEC2" blockId="37.[813,1451,190,278]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1518FDD0FC06FF02" bold="true" box="[813,946,190,214]" pageId="37" pageNumber="408">Etymology.</emphasis>
Named for the superficial resemblance to a mirine genus
<taxonomicName id="4C1B4D0AFF983B2B158AFDB1FBCEFF22" authority="Reuter" authorityName="Reuter" authorityYear="1875" box="[959,1146,223,246]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B158AFDB1FB9EFF22" box="[959,1066,223,246]" italics="true" pageId="37" pageNumber="408">Lygocoris</emphasis>
Reuter
</taxonomicName>
, with a Greek prefix,
<typeStatus id="54A0882BFF983B2B1352FDB0FA13FF22" box="[1383,1447,222,246]" pageId="37" pageNumber="408" type="paratype">para-</typeStatus>
(similar, apart from, etc.); gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF983B2B1518FC50FC08FC62" pageId="37" pageNumber="408" type="discussion">
<paragraph id="8BA43689FF983B2B1518FC50FABAFD62" blockId="37.[813,1452,318,694]" pageId="37" pageNumber="408">
<emphasis id="B96FEA9BFF983B2B1518FC50FC07FE82" bold="true" box="[813,947,318,342]" pageId="37" pageNumber="408">Discussion.</emphasis>
This unique genus is established for three thermophilic members inhabiting the Oriental Region and Wallacea, and is distinct from superficially similar genera (
<taxonomicName id="4C1B4D0AFF983B2B1500FCF1FC44FE62" authority="Reuter" authorityName="Reuter" authorityYear="1875" box="[821,1008,415,438]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B1500FCF1FC14FE62" box="[821,928,415,438]" italics="true" pageId="37" pageNumber="408">Lygocoris</emphasis>
Reuter
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFF983B2B121CFCF0FB55FE62" authority="Knight" authorityName="Knight" authorityYear="1917" box="[1065,1249,414,438]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B121CFCF0FB38FE62" box="[1065,1164,414,438]" italics="true" pageId="37" pageNumber="408">Neolygus</emphasis>
Knight
</taxonomicName>
) by the diagnostic characters mentioned above. As mentioned in some related works (e.g.,
<bibRefCitation id="EF8A4B78FF983B2B1586FCB1FB14FE22" author="Yasunaga T &amp; Duwal RK &amp; Schwartz MD" box="[947,1184,478,503]" pageId="37" pageNumber="408" pagination="44 - 54" refId="ref24720" refString="Yasunaga T, Duwal RK &amp; Schwartz MD (2012) First record and a new species of Neolygus Knight (Heteroptera: Miridae: Mirinae) from Nepal, with a checklist of Old World species of Neolygus. Zootaxa, 3413: 44 - 54." type="journal article" year="2012">Yasunaga et al., 2012</bibRefCitation>
; Yasunaga, 2023, 2024), a large number of known species as well as undescribed ones (liable to be placed in such genera) occurring in the Oriental Region and Wallacea may be representatives of
<emphasis id="B96FEA9BFF983B2B1518FF30FB8EFDA2" box="[813,1082,606,630]" italics="true" pageId="37" pageNumber="408">
<taxonomicName id="4C1B4D0AFF983B2B1518FF30FC66FDA2" authorityName="Yasunaga, Schwartz &amp; Chérot" authorityYear="2018" box="[813,978,606,630]" class="Insecta" family="Miridae" genus="Miyamotolygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">Miyamotolygus</taxonomicName>
,
<taxonomicName id="4C1B4D0AFF983B2B15EEFF30FB8EFDA2" authorityName="Carvalho" authorityYear="1987" box="[987,1082,606,630]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">Prolygus</taxonomicName>
</emphasis>
, and other (new) genera. In tropical Asia, genuine members of
<taxonomicName id="4C1B4D0AFF983B2B1247FF11FB6BFD42" authorityName="Reuter" authorityYear="1875" box="[1138,1247,639,662]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B1247FF11FB6BFD42" box="[1138,1247,639,662]" italics="true" pageId="37" pageNumber="408">Lygocoris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFF983B2B132EFF10FA34FD42" authorityName="Knight" authorityYear="1917" box="[1307,1408,638,662]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="37" pageNumber="408" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFF983B2B132EFF10FA34FD42" box="[1307,1408,638,662]" italics="true" pageId="37" pageNumber="408">Neolygus</emphasis>
</taxonomicName>
are apparently restricted to cool montane zones.
</paragraph>
<paragraph id="8BA43689FF983B2B1518FFB0FC08FC62" blockId="37.[813,1452,734,950]" pageId="37" pageNumber="408">Although many of the above-mentioned taxa have the similar-shaped left parameres, each group is distinct in the pattern of vesical sclerites and female genitalic structures, which is currently considered homoplasy. Further continuing endeavour is required to clarify relevant taxa as much as possible in order to elucidate their plausible phylogenetic relationships.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,199 @@
<document id="5170A31B80E3F76DA940BC5D30393727" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF9C3B2F16D1FAF3FBB7F8A2" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris theoi Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="404" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="404" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="838EDB097C25BD7C82D3D03BA2DC64F3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="17EA93C7AD53F4FDAC466C47990F99B9">
<mods:title id="7D29944C477031503E2E67FD25EAE4E6">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="14D5423A75AEE6D7E39A4FD3BA904537" type="personal">
<mods:role id="08404FD039E15F1939E3AC317783D0B4">
<mods:roleTerm id="9F39A495438436B33F73A68B796B5FFA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C7B40410EE48CF1D273D41EAE7F775F7">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="94A25BF3A0AD8EA531B4E694EAE4519D">text</mods:typeOfResource>
<mods:relatedItem id="39829198B41B01239F8D92B96E509333" type="host">
<mods:titleInfo id="6F7F1CBCE975EC3E6D9B9C30CCE4A331">
<mods:title id="4018BBC92E2632564C7791C80B8414EE">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="E8AB14221F16EF6431E974AA4F8C9373">
<mods:date id="E95D1F464814935EF52A34D39DBB8346">2024</mods:date>
<mods:detail id="5934DBB79792DDF9A27615A3BD6B6805" type="pubDate">
<mods:number id="486ED6FDDFD087CD9595620750A74912">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="514FCC3187D21542F63DA1C1CF3FC4B7" type="volume">
<mods:number id="770DC0E0BFB0C7C1E48A53C517F53D81">72</mods:number>
</mods:detail>
<mods:extent id="1981C04AA527E20E7EDB0BC5B84F91BD" unit="page">
<mods:start id="13C79333C67E7CD1EDD9B34BE0CA1942">371</mods:start>
<mods:end id="12FF98CF5B16F8A6CE80990738533A04">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D1CB338D5A4EB343A55CEB3A675EC029">journal article</mods:classification>
<mods:identifier id="2C4312C3527C0583815E344D3FF84A74" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="EF780073832FB87997FB730CDDA48862" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DECAC0B947020FBF6F7B7213F4F1BB44" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="A89AFE9A4AA63FB21F9D1461D005E821" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF9C3B2F16D1FAF3FBB7F8A2" LSID="urn:lsid:plazi:treatment:03B2879FFF9C3B2F16D1FAF3FBB7F8A2" httpUri="http://treatment.plazi.org/id/03B2879FFF9C3B2F16D1FAF3FBB7F8A2" lastPageNumber="404" pageId="33" pageNumber="404">
<subSubSection id="C3016502FF9C3B2F16D1FAF3FDC9F861" box="[228,637,1949,1974]" pageId="33" pageNumber="404" type="nomenclature">
<paragraph id="8BA43689FF9C3B2F16D1FAF3FDC9F861" blockId="33.[228,637,1949,2006]" box="[228,637,1949,1974]" pageId="33" pageNumber="404">
<heading id="D0EC81E5FF9C3B2F16D1FAF3FDC9F861" bold="true" box="[228,637,1949,1974]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="404" reason="2">
<emphasis id="B96FEA9BFF9C3B2F16D1FAF3FDC9F861" bold="true" box="[228,637,1949,1974]" pageId="33" pageNumber="404">
<taxonomicName id="4C1B4D0AFF9C3B2F16D1FAF3FE5DF861" authorityName="Yasunaga" authorityYear="2024" box="[228,489,1949,1973]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="species" species="theoi" status="sp. nov.">
<emphasis id="B96FEA9BFF9C3B2F16D1FAF3FE5DF861" bold="true" box="[228,489,1949,1973]" italics="true" pageId="33" pageNumber="404">Himalisundacoris theoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F17CDFAF0FDC9F861" box="[504,637,1949,1974]" pageId="33" pageNumber="404" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1711FAD0FD89F802" box="[292,573,1982,2006]" pageId="33" pageNumber="404" type="description">
<paragraph id="8BA43689FF9C3B2F1711FAD0FD89F802" blockId="33.[228,637,1949,2006]" box="[292,573,1982,2006]" pageId="33" pageNumber="404">
(
<figureCitation id="13202A0CFF9C3B2F1719FAD0FE37F802" box="[300,387,1982,2006]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="33" pageNumber="404">Figs. 12</figureCitation>
EH, 14FJ, 30)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FA90FB42FEC2" pageId="33" pageNumber="404" type="materials_examined">
<paragraph id="8BA43689FF9C3B2F1644FA90FD5BF7E2" blockId="33.[113,751,2045,2102]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FA90FE9DF7C1" bold="true" box="[113,297,2045,2070]" pageId="33" pageNumber="404">Type material.</emphasis>
<materialsCitation id="3B733CD4FF9C3B2F1700FA90FD5BF7E2" collectingDate="1996-03-13" country="Indonesia" latitude="-8.283334" location="Tabanan" longLatPrecision="1301" longitude="115.166664" pageId="33" pageNumber="404" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<emphasis id="B96FEA9BFF9C3B2F1700FA90FE10F7C2" bold="true" box="[309,420,2046,2070]" pageId="33" pageNumber="404">
<typeStatus id="54A0882BFF9C3B2F1700FA90FE10F7C2" box="[309,420,2046,2070]" pageId="33" pageNumber="404" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFF9C3B2F1436FA93FD19F7C1" bold="true" box="[515,685,2045,2069]" pageId="33" pageNumber="404">
<collectingCountry id="F30C7619FF9C3B2F1436FA93FD13F7C1" box="[515,679,2045,2069]" name="Indonesia" pageId="33" pageNumber="404">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFF9C3B2F148CFA90FD5FF7C2" box="[697,747,2046,2070]" country="Indonesia" name="Bali" pageId="33" pageNumber="404">Bali</collectingRegion>
,
<location id="8EC46052FF9C3B2F1644F570FF64F7E2" LSID="urn:lsid:plazi:treatment:03B2879FFF9C3B2F16D1FAF3FBB7F8A2:8EC46052FF9C3B2F1644F570FF64F7E2" box="[113,208,2078,2102]" country="Indonesia" latitude="-8.283334" longLatPrecision="1301" longitude="115.166664" name="Tabanan" pageId="33" pageNumber="404" stateProvince="Bali">Tabanan</location>
,
<geoCoordinate id="EE2F504EFF9C3B2F16EEF572FE90F7E1" box="[219,292,2076,2101]" degrees="8" direction="south" minutes="17" orientation="latitude" pageId="33" pageNumber="404" precision="925" value="-8.283334">8°17S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF9C3B2F1704F572FE23F7E1" box="[305,407,2076,2101]" degrees="115" direction="east" minutes="10" orientation="longitude" pageId="33" pageNumber="404" precision="925" value="115.166664">115°10E</geoCoordinate>
, local collector,
<date id="FFA51049FF9C3B2F147EF570FD5BF7E2" box="[587,751,2078,2102]" pageId="33" pageNumber="404" value="1996-03-13">
<collectingDate id="EFE1E9A1FF9C3B2F147EF570FD5BF7E2" box="[587,751,2078,2102]" pageId="33" pageNumber="404" value="1996-03-13">13 March 1996</collectingDate>
</date>
</materialsCitation>
</paragraph>
<paragraph id="8BA43689FF9C3B2F1518FDD0FB42FEC2" blockId="33.[813,1452,190,278]" pageId="33" pageNumber="404">
(
<collectionCode id="ED0AAE4CFF9C3B2F1501FDD0FCDFFF02" box="[820,875,190,214]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="33" pageNumber="404">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFF9C3B2F15B6FDD0FC62FF02" box="[899,982,190,214]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="404">AMNH</collectionCode>
_PBI 00378787).
<materialsCitation id="3B733CD4FF9C3B2F12AFFDD0FB46FEC2" collectingDate="1996-03-09" collectingDateMax="1996-03-13" collectingDateMin="1996-03-09" collectionCode="AMNH, TYCN" country="Indonesia" latitude="-8.283334" location="Tabanan" longLatPrecision="1301" longitude="115.166664" pageId="33" pageNumber="404" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Bali" typeStatus="paratype">
<typeStatus id="54A0882BFF9C3B2F12AFFDD0FAB2FF02" box="[1178,1286,190,214]" pageId="33" pageNumber="404" type="paratype">Paratypes</typeStatus>
:
<collectingCountry id="F30C7619FF9C3B2F1326FDD0FA1CFF02" box="[1299,1448,190,214]" name="Indonesia" pageId="33" pageNumber="404">INDONESIA</collectingCountry>
:
<specimenCount id="9D1DFD00FF9C3B2F1518FDB0FC31FF22" box="[813,901,222,246]" count="5" pageId="33" pageNumber="404" type="male">5 males</specimenCount>
,
<specimenCount id="9D1DFD00FF9C3B2F15A6FDB0FBB4FF22" box="[915,1024,222,246]" count="2" pageId="33" pageNumber="404" type="female">2 females</specimenCount>
, same data as for holotype, except for date:
<date id="FFA51049FF9C3B2F155CFD90FB9EFEC2" box="[873,1066,254,278]" pageId="33" pageNumber="404" value="1996-03-09" valueMax="1996-03-13" valueMin="1996-03-09">
<collectingDate id="EFE1E9A1FF9C3B2F155CFD90FB9EFEC2" box="[873,1066,254,278]" pageId="33" pageNumber="404" value="1996-03-09" valueMax="1996-03-13" valueMin="1996-03-09">913 March 1996</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF9C3B2F120EFD90FB25FEC2" box="[1083,1169,254,278]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="404">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFF9C3B2F12A8FD90FB5EFEC2" box="[1181,1258,254,278]" pageId="33" pageNumber="404">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FC53FA93FDC2" pageId="33" pageNumber="404" type="diagnosis">
<paragraph id="8BA43689FF9C3B2F1518FC53FA93FDC2" blockId="33.[813,1452,317,534]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FC53FC11FE81" bold="true" box="[813,933,317,341]" pageId="33" pageNumber="404">Diagnosis.</emphasis>
This new species is most similar in external appearance to
<taxonomicName id="4C1B4D0AFF9C3B2F15FFFC30FB9DFEA2" authorityName="Yasunaga" authorityYear="2024" box="[970,1065,350,374]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="species" species="takaii" status="new species">
<emphasis id="B96FEA9BFF9C3B2F15FFFC30FB9DFEA2" box="[970,1065,350,374]" italics="true" pageId="33" pageNumber="404">H. takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F1202FC30FB08FEA2" box="[1079,1212,350,374]" pageId="33" pageNumber="404" rank="species">new species</taxonomicNameLabel>
, from which
<taxonomicName id="4C1B4D0AFF9C3B2F1378FC30FA10FEA2" authorityName="Yasunaga" authorityYear="2024" box="[1357,1444,350,374]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="species" species="theoi" status="new species">
<emphasis id="B96FEA9BFF9C3B2F1378FC30FA10FEA2" box="[1357,1444,350,374]" italics="true" pageId="33" pageNumber="404">H. theoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F1518FC10FC09FE42" box="[813,957,382,406]" pageId="33" pageNumber="404" rank="species">new species</taxonomicNameLabel>
, can be easily distinguished by the basal transverse carina obliterated (
<figureCitation id="13202A0CFF9C3B2F1256FCF0FB71FE62" box="[1123,1221,414,438]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30A</figureCitation>
); antennomere II with a pale subbasal annulation; pronotum with a pair of dark, oval maculae and immaculate lateral margins; thoracic pleura entirely pale; and apex of each tibia darkened.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FF53FBD8F942" pageId="33" pageNumber="404" type="description">
<paragraph id="8BA43689FF9C3B2F1518FF53FC53F982" blockId="33.[813,1452,573,1622]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FF53FC0CFD81" bold="true" box="[813,952,573,597]" pageId="33" pageNumber="404">Description.</emphasis>
Body oval; basic colouration pale olive brown with widely darkened hemelytra (
<figureCitation id="13202A0CFF9C3B2F129CFF30FABAFDA2" box="[1193,1294,606,630]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="33" pageNumber="404">Fig. 12E</figureCitation>
); dorsum oily shiny; dark areas or maculae reduced in female (
<figureCitation id="13202A0CFF9C3B2F130EFF10FA14FD42" box="[1339,1440,638,662]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="33" pageNumber="404">Fig. 12G</figureCitation>
). Head shiny pale brown; basal transverse carina obliterated (
<figureCitation id="13202A0CFF9C3B2F1500FFD0FC17FD02" box="[821,931,702,726]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30A</figureCitation>
); clypeus and lorum entirely shiny fuscous. Antenna dark brown; antennomere I pale brown, obscured mesially, with dark base and apex; antennomere II with a pale subbasal annulation; base of antennomere III yellowish brown. Labium pale reddish brown, slightly exceeding apex of metacoxa; base of segment I and apical half of segment IV infuscate. Pronotum pale olive brown, with a pair of dark, oval maculae (continuing to calli) at middle (
<figureCitation id="13202A0CFF9C3B2F12E9FEF0FA8FFC62" box="[1244,1339,926,950]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="33" pageNumber="404">Fig. 12E</figureCitation>
) (maculae in female reduced and calli with a pair of dark spots as in
<figureCitation id="13202A0CFF9C3B2F1518FEB0FC3FFC22" box="[813,907,990,1014]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="33" pageNumber="404">Fig. 12F</figureCitation>
); pleura uniformly dark brown; metathoracic scent efferent system creamy yellow, relatively large (
<figureCitation id="13202A0CFF9C3B2F1376FE90FCF7FBE2" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30B, D</figureCitation>
); scutellum pale creamy brown, sometimes partly tinged with green, with a pair of dark, elongate-oval maculae on anterior half. Hemelytron oily shiny, with sparsely distributed, pale, simple, reclining setae (
<figureCitation id="13202A0CFF9C3B2F1244F910FB63FB42" box="[1137,1239,1150,1174]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30C</figureCitation>
); clavus and apical half of corium dark brown; cuneus yellowish brown, with darkened basal half and apex; membrane pale greyish brown, semi-transparent, with brown veins. All coxae and legs pale brown; metafemur with two brown, faint, subapical rings; apical part of each tarsomere III slightly darkened; pretarsal structure as in
<figureCitation id="13202A0CFF9C3B2F15FFF850FB93FA82" box="[970,1063,1342,1366]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">
Fig.
<specimenCount id="9D1DFD00FF9C3B2F15CEF850FB93FA82" box="[1019,1063,1342,1366]" count="30" pageId="33" pageNumber="404" type="generic">30F</specimenCount>
</figureCitation>
. Abdomen pale brown; male genital segment (pygophore) darkened apically. Male genitalia (
<figureCitation id="13202A0CFF9C3B2F1342F830FCF3FA42" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="33" pageNumber="404">Figs. 14</figureCitation>
FH, 30GL): Hypophysis of left paramere relatively narrow, with anchor-shaped apex (
<figureCitation id="13202A0CFF9C3B2F1294F8F0FB4EFA62" box="[1185,1274,1438,1462]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30I</figureCitation>
); right paramere with small, blunt-tipped hypophysis (
<figureCitation id="13202A0CFF9C3B2F12F1F8D0FA9EFA02" box="[1220,1322,1470,1494]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="33" pageNumber="404">Fig. 14G</figureCitation>
); secondary gonopore comparatively large, thick-rimmed (
<figureCitation id="13202A0CFF9C3B2F1328F8B0FA14FA22" box="[1309,1440,1502,1526]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30KL</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFF9C3B2F15C0F890FBC2F9C1" box="[1013,1142,1534,1558]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="33" pageNumber="404">Figs. 14I, J</figureCitation>
,
<figureCitation id="13202A0CFF9C3B2F12B0F890FB6AF9C2" box="[1157,1246,1534,1558]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">30MO</figureCitation>
): Sclerotised rings small (
<figureCitation id="13202A0CFF9C3B2F1541FB70FC7FF9E2" box="[884,971,1566,1590]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="33" pageNumber="404">Fig. 14I</figureCitation>
); posterior wall with rather narrow interramal lobes (
<figureCitation id="13202A0CFF9C3B2F1543FB50FC6FF982" box="[886,987,1598,1622]" captionStart="Fig" captionStartId="44.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@44.[135,1473,251,1929]" captionTargetPageId="44" captionText="Fig. 30. Scanning electron micrographs of Himalisundacoris theoi, new species. A, head and pronotum, anterior-left lateral view; B, thoracic pleura, left lateral view; C, scutellum and anterior hemelytron, dorsal view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, pygophore, caudal view; H, ditto, dorsal view; I, left paramere; J, K, vesica; L, secondary gonopore; M, sclerotised ring; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683024" httpUri="https://zenodo.org/record/14683024/files/figure.png" pageId="33" pageNumber="404">Fig. 30N</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FF9C3B2F1518FB10FBD8F942" blockId="33.[813,1132,1662,1686]" box="[813,1132,1662,1686]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FB10FC6FF942" bold="true" box="[813,987,1662,1686]" pageId="33" pageNumber="404">Measurements.</emphasis>
See
<tableCitation id="C6990332FF9C3B2F1226FB10FBD2F942" box="[1043,1126,1662,1686]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="33" pageNumber="404">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FBD0FB44F921" pageId="33" pageNumber="404" type="etymology">
<paragraph id="8BA43689FF9C3B2F1518FBD0FB44F921" blockId="33.[813,1451,1726,1782]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FBD0FC1BF902" bold="true" box="[813,943,1726,1750]" pageId="33" pageNumber="404">Etymology.</emphasis>
Named after Theo, the grandson of Dr. Michael D. Schwartz; a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FA73FBC9F8E2" box="[813,1149,1821,1846]" pageId="33" pageNumber="404" type="distribution">
<paragraph id="8BA43689FF9C3B2F1518FA73FBC9F8E2" blockId="33.[813,1149,1821,1846]" box="[813,1149,1821,1846]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FA73FC0BF8E1" bold="true" box="[813,959,1821,1845]" pageId="33" pageNumber="404">Distribution.</emphasis>
<collectingCountry id="F30C7619FF9C3B2F15F2FA70FB9BF8E2" box="[967,1071,1822,1846]" name="Indonesia" pageId="33" pageNumber="404">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFF9C3B2F1275FA70FBC5F8E2" box="[1088,1137,1822,1846]" country="Indonesia" name="Bali" pageId="33" pageNumber="404">Bali</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1518FA33FBB7F8A2" box="[813,1027,1885,1910]" pageId="33" pageNumber="404" type="biology_ecology">
<paragraph id="8BA43689FF9C3B2F1518FA33FBB7F8A2" blockId="33.[813,1027,1885,1910]" box="[813,1027,1885,1910]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1518FA33FC3EF8A1" bold="true" box="[813,906,1885,1909]" pageId="33" pageNumber="404">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,166 @@
<document id="744DD4149E5F64EBC272F9643FAD489B" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFF9C3B2F16D5FD93FEF3F8A2" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris takaii Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="404" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="404" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="557C7B7DA8CF598A46E7FFF41CD7E0D2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4CB74CE5A0C28520FDE71439C66A1723">
<mods:title id="5B12A6852A8839E470D3E0EA025B6B69">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="5E2B320870536A45C29A13F5E7F4D033" type="personal">
<mods:role id="AD1A3954B90568D0E1B47582463A8266">
<mods:roleTerm id="6E6C08A2655A6D5A85300306B375120A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDF25DF72051F44F7E2FCC87C65DEC40">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="8874242810E31535EA7A346FCE9E0284">text</mods:typeOfResource>
<mods:relatedItem id="3A8F7E07C476BDB5002B1C959FB0BFBD" type="host">
<mods:titleInfo id="7F49745E5A8D56B57D7D7D57A63F57BF">
<mods:title id="32196FD533A2B963433ADDF58D9BA23C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="178D05F1063D8234437627772FEA1E4C">
<mods:date id="C6AA86B8C69EC037B96F55DE69197007">2024</mods:date>
<mods:detail id="ECBE26BA3E9EB74092D09D69A9BBA5DF" type="pubDate">
<mods:number id="EC4B97808C96EBCA0DB2B41CD462D7A6">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="67295EA50172F799D20E11E8051F3D83" type="volume">
<mods:number id="7B224214FF3CB66E388324176022B133">72</mods:number>
</mods:detail>
<mods:extent id="5792AE3A8AC2326E03A4A2E0E9C2A858" unit="page">
<mods:start id="59D511263C3657F18C2E723FB74F1ABD">371</mods:start>
<mods:end id="0D91C3CE1BD2E0E943CFB9787D8573CC">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="451F4119C0EA18217D7826E39E84A29F">journal article</mods:classification>
<mods:identifier id="FDA3184FBAF90CED9D52A7EB5E8D9860" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="E99B3F04E59490192DC2375CD40308EB" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="4BEE24BFC943DB2DDA37C6597C640DEA" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="44C330D894B81352FB95AEB11D3960CD" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFF9C3B2F16D5FD93FEF3F8A2" ID-DOI="http://doi.org/10.5281/zenodo.14683040" ID-Zenodo-Dep="14683040" LSID="urn:lsid:plazi:treatment:03B2879FFF9C3B2F16D5FD93FEF3F8A2" httpUri="http://treatment.plazi.org/id/03B2879FFF9C3B2F16D5FD93FEF3F8A2" lastPageNumber="404" pageId="33" pageNumber="404">
<subSubSection id="C3016502FF9C3B2F16D5FD93FD35FEC1" box="[224,641,253,278]" pageId="33" pageNumber="404" type="nomenclature">
<paragraph id="8BA43689FF9C3B2F16D5FD93FD35FEC1" blockId="33.[224,641,253,310]" box="[224,641,253,278]" pageId="33" pageNumber="404">
<heading id="D0EC81E5FF9C3B2F16D5FD93FD35FEC1" bold="true" box="[224,641,253,278]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="404" reason="2">
<emphasis id="B96FEA9BFF9C3B2F16D5FD93FD35FEC1" bold="true" box="[224,641,253,278]" pageId="33" pageNumber="404">
<taxonomicName id="4C1B4D0AFF9C3B2F16D5FD93FE58FEC1" authorityName="Yasunaga" authorityYear="2024" box="[224,492,253,277]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="species" species="takaii" status="sp. nov.">
<emphasis id="B96FEA9BFF9C3B2F16D5FD93FE58FEC1" bold="true" box="[224,492,253,277]" italics="true" pageId="33" pageNumber="404">Himalisundacoris takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9C3B2F17CEFD90FD35FEC1" box="[507,641,253,278]" pageId="33" pageNumber="404" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F173EFC70FDE2FEE2" box="[267,598,286,310]" pageId="33" pageNumber="404" type="description">
<paragraph id="8BA43689FF9C3B2F173EFC70FDE2FEE2" blockId="33.[224,641,253,310]" box="[267,598,286,310]" pageId="33" pageNumber="404">
(
<figureCitation id="13202A0CFF9C3B2F1726FC70FE28FEE1" box="[275,412,286,310]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="33" pageNumber="404">Figs. 11E, F</figureCitation>
,
<figureCitation id="13202A0CFF9C3B2F179CFC70FE76FEE2" box="[425,450,286,310]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="33" pageNumber="404">15</figureCitation>
IK, 31AG)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FC30FDA6FE02" pageId="33" pageNumber="404" type="materials_examined">
<paragraph id="8BA43689FF9C3B2F1644FC30FDA6FE02" blockId="33.[113,752,349,470]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FC30FE92FEA1" bold="true" box="[113,294,349,374]" pageId="33" pageNumber="404">Type material.</emphasis>
<materialsCitation id="3B733CD4FF9C3B2F1707FC30FDA6FE02" collectingDate="1999-12-22" collectingDateMax="1999-12-25" collectingDateMin="1999-12-22" collectionCode="ZRC, AMNH" collectorName="M. Takai" country="Indonesia" latitude="-5.3166666" location="Lompobattan" longLatPrecision="1305" longitude="119.933334" pageId="33" pageNumber="404" specimenCode="PBI 00378786" specimenCount="1" stateProvince="Sulawesi Selatan" typeStatus="holotype">
<emphasis id="B96FEA9BFF9C3B2F1707FC30FE2BFEA2" bold="true" box="[306,415,350,374]" pageId="33" pageNumber="404">
<typeStatus id="54A0882BFF9C3B2F1707FC30FE2BFEA2" box="[306,415,350,374]" pageId="33" pageNumber="404" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFF9C3B2F17C9FC33FD17FEA1" bold="true" box="[508,675,349,373]" pageId="33" pageNumber="404">
<collectingCountry id="F30C7619FF9C3B2F17C9FC33FD2AFEA1" box="[508,670,349,373]" name="Indonesia" pageId="33" pageNumber="404">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFF9C3B2F149BFC30FF62FE42" country="Indonesia" name="Sulawesi Selatan" pageId="33" pageNumber="404">South Sulawesi</collectingRegion>
,
<location id="8EC46052FF9C3B2F16D0FC10FECDFE42" LSID="urn:lsid:plazi:treatment:03B2879FFF9C3B2F16D5FD93FEF3F8A2:8EC46052FF9C3B2F16D0FC10FECDFE42" box="[229,377,382,406]" country="Indonesia" latitude="-5.3166666" longLatPrecision="1305" longitude="119.933334" name="Lompobattan" pageId="33" pageNumber="404" stateProvince="Sulawesi Selatan">Lompobattan</location>
,
<locationDeviation id="9A162FABFF9C3B2F17BDFC10FD2CFE42" box="[392,664,382,406]" location="Makassar" pageId="33" pageNumber="404">
<quantity id="4CE39B6CFF9C3B2F17BDFC10FE5DFE42" box="[392,489,382,406]" metricMagnitude="4" metricUnit="m" metricValue="4.828032" pageId="33" pageNumber="404" unit="mi" value="30.0">30 miles</quantity>
E of Makassar
</locationDeviation>
(Ujung Pandang),
<geoCoordinate id="EE2F504EFF9C3B2F16DCFCF2FE80FE61" box="[233,308,412,437]" degrees="5" direction="south" minutes="19" orientation="latitude" pageId="33" pageNumber="404" precision="925" value="-5.3166666">5°19S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFF9C3B2F1776FCF2FE1FFE61" box="[323,427,412,437]" degrees="119" direction="east" minutes="56" orientation="longitude" pageId="33" pageNumber="404" precision="925" value="119.933334">119°56E</geoCoordinate>
,
<collectorName id="26EE535FFF9C3B2F178FFCF0FD90FE62" box="[442,548,414,438]" pageId="33" pageNumber="404">M. Takai</collectorName>
,
<date id="FFA51049FF9C3B2F1407FCF0FF12FE02" pageId="33" pageNumber="404" value="1999-12-22" valueMax="1999-12-25" valueMin="1999-12-22">
<collectingDate id="EFE1E9A1FF9C3B2F1407FCF0FF12FE02" pageId="33" pageNumber="404" value="1999-12-22" valueMax="1999-12-25" valueMin="1999-12-22">2225 December 1999</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFF9C3B2F1683FCD0FF59FE02" box="[182,237,446,470]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="33" pageNumber="404">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFF9C3B2F1731FCD0FEE3FE02" box="[260,343,446,470]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="404">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FF9C3B2F1750FCD0FDB3FE02" box="[357,519,446,470]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="33" pageNumber="404">PBI 00378786</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FC93FEC4FD42" pageId="33" pageNumber="404" type="diagnosis">
<paragraph id="8BA43689FF9C3B2F1644FC93FEC4FD42" blockId="33.[113,752,509,662]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FC93FF52FDC1" bold="true" box="[113,230,509,533]" pageId="33" pageNumber="404">Diagnosis.</emphasis>
This new species is currently known only by a single male specimen and distinguished from other congeners by the uniformly dark brown antennomere II, and darkened lateral parts of pronotum, entire thoracic pleura, clavus and abdomen (
<figureCitation id="13202A0CFF9C3B2F16D6FF10FED7FD42" box="[227,355,638,662]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="33" pageNumber="404">Fig. 11E, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FFD3FE05F9A2" pageId="33" pageNumber="404" type="description">
<paragraph id="8BA43689FF9C3B2F1644FFD3FDF2F9E2" blockId="33.[113,753,701,1590]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FFD3FEFDFD02" bold="true" box="[113,329,701,726]" pageId="33" pageNumber="404">Description. Male</emphasis>
: Body oval, slightly elongate; basic colouration pale olive brown with widely darkened pronotum, scutellum, clavus and ventral surface (
<figureCitation id="13202A0CFF9C3B2F143BFF90FD3FFCC2" box="[526,651,766,790]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="33" pageNumber="404">Fig. 11E, F</figureCitation>
); dorsum polished, oily shiny. Head pale reddish brown, smooth; vertex with narrow, weak basal transverse carina that is interrupted mesally (
<figureCitation id="13202A0CFF9C3B2F1767FE30FE03FCA2" box="[338,439,862,886]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">Fig. 31A</figureCitation>
); clypeus and lorum entirely shiny fuscous. Antenna dark brown; antennomere I partly pale mesially; antennomere II uniformly darkened; extreme base of antennomere III creamy yellow. Labium pale brown, reaching but not exceeding apex of metacoxa; apical 2/3 of segment IV darkened. Pronotum pale olive brown, darkened laterally, almost glabrous; pleura uniformly dark brown; metathoracic scent efferent system pale brown; scutellum pale olive brown, with a pair of dark, triangular maculae. Hemelytron oily shiny, with uniformly distributed, pale, simple, reclining setae (
<figureCitation id="13202A0CFF9C3B2F1746F9F0FE63FB62" box="[371,471,1182,1206]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">Fig. 31A</figureCitation>
); whole clavus and apical half of corium dark brown; cuneus yellowish brown, with darkened extreme apex; membrane pale greyish brown, semi-transparent, with slightly darker veins. All coxae reddish brown; legs pale brown; metafemur with reddish basal half and two subapical brown rings; each tarsomere III slightly darkened; meta-tarsomere II as long as III; parempodia rather short (
<figureCitation id="13202A0CFF9C3B2F168FF810FE95FA42" box="[186,289,1406,1430]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">Fig. 31C</figureCitation>
). Abdomen uniformly shiny dark brown. Male genitalia (
<figureCitation id="13202A0CFF9C3B2F1714F8F0FECEFA62" box="[289,378,1438,1462]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="33" pageNumber="404">Figs. 15</figureCitation>
IK, 31DG): Hypophysis of left paramere inflated, elongate, with anchor-shaped apex (
<figureCitation id="13202A0CFF9C3B2F14F0F8D0FF14FA22" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">Fig. 31E</figureCitation>
); right paramere bulbous, short, with small hypophysis (
<figureCitation id="13202A0CFF9C3B2F164CF890FF5EF9C2" box="[121,234,1534,1558]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="33" pageNumber="404">Figs. 15K</figureCitation>
,
<figureCitation id="13202A0CFF9C3B2F16CCF890FE92F9C2" box="[249,294,1534,1558]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">31F</figureCitation>
); notches of vesical lobal-sclerites rather small (
<figureCitation id="13202A0CFF9C3B2F168EFB70FEABF9E2" box="[187,287,1566,1590]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="33" pageNumber="404">Figs. 15I</figureCitation>
,
<figureCitation id="13202A0CFF9C3B2F1718FB70FEEAF9E2" box="[301,350,1566,1590]" captionStart="Fig" captionStartId="46.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@46.[136,1474,251,1929]" captionTargetPageId="46" captionText="Fig. 31. Scanning electron micrographs of Himalisundacoris takaii, new species, holotype male (AG) and Paralygocoris meridionalis, female (HO). A, Head and anterior thorax, dorsal-left lateral, view; B, I, thoracic pleura, left lateral view; C, J, metatarsus; D, pygophore, dorsal view; E, apical part of left paramere; J, right paramere; G, vesica; H, head and pronotum, anterior-left lateral view; K, pretarsal structure of metaleg; L, M, posterior wall; N, sclerotised ring; O, apex of ovipositor (gonapophysis I)." figureDoi="http://doi.org/10.5281/zenodo.14683028" httpUri="https://zenodo.org/record/14683028/files/figure.png" pageId="33" pageNumber="404">31G</figureCitation>
).
<emphasis id="B96FEA9BFF9C3B2F1746FB70FE72F9E2" bold="true" box="[371,454,1566,1590]" pageId="33" pageNumber="404">Female</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BA43689FF9C3B2F1644FB30FE05F9A2" blockId="33.[113,433,1630,1654]" box="[113,433,1630,1654]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FB30FEABF9A2" bold="true" box="[113,287,1630,1654]" pageId="33" pageNumber="404">Measurements.</emphasis>
See
<tableCitation id="C6990332FF9C3B2F1762FB30FE1EF9A2" box="[343,426,1630,1654]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="33" pageNumber="404">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FBF0FE3DF921" pageId="33" pageNumber="404" type="etymology">
<paragraph id="8BA43689FF9C3B2F1644FBF0FE3DF921" blockId="33.[113,752,1694,1782]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FBF0FF47F962" bold="true" box="[113,243,1694,1718]" pageId="33" pageNumber="404">Etymology.</emphasis>
Named after Mr. Mikio Takai, who collected and provided numerous Asian specimens of the
<taxonomicName id="4C1B4D0AFF9C3B2F14B5FBD0FD6CF902" box="[640,728,1726,1750]" class="Insecta" family="Miridae" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="404" phylum="Arthropoda" rank="family">Miridae</taxonomicName>
; a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FA73FDA0F8E2" box="[113,532,1821,1846]" pageId="33" pageNumber="404" type="distribution">
<paragraph id="8BA43689FF9C3B2F1644FA73FDA0F8E2" blockId="33.[113,532,1821,1846]" box="[113,532,1821,1846]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FA73FEB7F8E1" bold="true" box="[113,259,1821,1845]" pageId="33" pageNumber="404">Distribution.</emphasis>
<collectingCountry id="F30C7619FF9C3B2F1739FA70FEC0F8E2" box="[268,372,1822,1846]" name="Indonesia" pageId="33" pageNumber="404">Indonesia</collectingCountry>
(S.
<collectingRegion id="49DFF86BFF9C3B2F1796FA70FDBCF8E2" box="[419,520,1822,1846]" country="Indonesia" name="Sulawesi Utara" pageId="33" pageNumber="404">Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FF9C3B2F1644FA33FEF3F8A2" box="[113,327,1885,1910]" pageId="33" pageNumber="404" type="biology_ecology">
<paragraph id="8BA43689FF9C3B2F1644FA33FEF3F8A2" blockId="33.[113,327,1885,1910]" box="[113,327,1885,1910]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FA33FF7AF8A1" bold="true" box="[113,206,1885,1909]" pageId="33" pageNumber="404">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,185 @@
<document id="D6D8EA180378E7B170464BEB74F78343" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA03B1316E8FCD3FBB7FF01" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris palolo Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="400" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="400" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8AE16DF79CD8FBC928B8C4885F2DE5F4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7F086987980597E48391D482DDC85AFD">
<mods:title id="C5E2702A92717679F991F6176CD50B24">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="2C871580301B0CEB6301BFF87B5AAAAF" type="personal">
<mods:role id="7FC7F28A681475FF3A0A30B7F66B9848">
<mods:roleTerm id="9B04C8ABEA99C202F6769E3975B1EEB8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6279CB609566C65367432B8FA04E45D4">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="76819083A7DB66F9E6313F02BB55F442">text</mods:typeOfResource>
<mods:relatedItem id="D4A39D1241C754FCDCAD0C8580576818" type="host">
<mods:titleInfo id="AA7182C1790004B2D4D4EAE4EFB3387B">
<mods:title id="558008D7FB132F2E3C67D45FCFB3E6B5">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="EB04E910C1B18A8712E0257FEEA1E171">
<mods:date id="A8DD3F98D9B184ACF71B675B0B564118">2024</mods:date>
<mods:detail id="611DEB529DBBB4EB6C27B4A27CFCE06E" type="pubDate">
<mods:number id="484EC1EE0EFE6871B84613154FFBE8FB">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="8E08A99927832658742DED4B9350A6D9" type="volume">
<mods:number id="CE9B60431B034819BBFAB2E5FEBF7BE9">72</mods:number>
</mods:detail>
<mods:extent id="BBDCF6B97C49CB72F6878794EFA0169D" unit="page">
<mods:start id="525CCC1F2689530E530CD392F749A4C5">371</mods:start>
<mods:end id="8F9324839B40E38270257A76DB379714">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E4CB4B913E774A7D1AC27774F48EAE53">journal article</mods:classification>
<mods:identifier id="91D692A47C804624B693A0815192776F" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="A9CE5FBF6D8F730CACE95BF0D9220FCE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C37FF126FCC0BF4232302668F56407AE" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="3E2D6F9DF170A153CD68E160F8237C73" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA03B1316E8FCD3FBB7FF01" ID-DOI="http://doi.org/10.5281/zenodo.14683026" ID-Zenodo-Dep="14683026" LSID="urn:lsid:plazi:treatment:03B2879FFFA03B1316E8FCD3FBB7FF01" httpUri="http://treatment.plazi.org/id/03B2879FFFA03B1316E8FCD3FBB7FF01" lastPageNumber="400" pageId="29" pageNumber="400">
<subSubSection id="C3016502FFA03B1316E8FCD3FD30FE01" box="[221,644,445,469]" pageId="29" pageNumber="400" type="nomenclature">
<paragraph id="8BA43689FFA03B1316E8FCD3FD30FE01" blockId="29.[221,644,445,501]" box="[221,644,445,469]" pageId="29" pageNumber="400">
<heading id="D0EC81E5FFA03B1316E8FCD3FD30FE01" bold="true" box="[221,644,445,469]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="400" reason="2">
<emphasis id="B96FEA9BFFA03B1316E8FCD3FD30FE01" bold="true" box="[221,644,445,469]" pageId="29" pageNumber="400">
<taxonomicName id="4C1B4D0AFFA03B1316E8FCD3FE5BFE01" authorityName="Yasunaga" authorityYear="2024" box="[221,495,445,469]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="400" phylum="Arthropoda" rank="species" species="palolo" status="sp. nov.">
<emphasis id="B96FEA9BFFA03B1316E8FCD3FE5BFE01" bold="true" box="[221,495,445,469]" italics="true" pageId="29" pageNumber="400">Himalisundacoris palolo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA03B1317CBFCD3FD30FE01" box="[510,644,445,469]" pageId="29" pageNumber="400" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131728FCB3FDF0FE21" box="[285,580,477,501]" pageId="29" pageNumber="400" type="description">
<paragraph id="8BA43689FFA03B131728FCB3FDF0FE21" blockId="29.[221,644,445,501]" box="[285,580,477,501]" pageId="29" pageNumber="400">
(
<figureCitation id="13202A0CFFA03B131710FCB3FECFFE21" box="[293,379,477,501]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="29" pageNumber="400">Figs. 11</figureCitation>
AD, 15AC, 28)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FF73FF67FD61" pageId="29" pageNumber="400" type="materials_examined">
<paragraph id="8BA43689FFA03B131644FF73FF67FD61" blockId="29.[113,753,541,693]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FF73FE95FDE1" bold="true" box="[113,289,541,565]" pageId="29" pageNumber="400">Type material.</emphasis>
<materialsCitation id="3B733CD4FFA03B131719FF73FD58FDA1" collectingDate="1988-04-28" collectionCode="UV, T, ZRC, AMNH" country="Indonesia" county="Palu" latitude="-1.2166667" location="Yasunaga" longLatPrecision="1308" longitude="120.05" municipality="Palolo" pageId="29" pageNumber="400" specimenCode="PBI 00378784" specimenCount="1" stateProvince="Sulawesi Tengah" typeStatus="holotype">
<emphasis id="B96FEA9BFFA03B131719FF73FE23FDE1" bold="true" box="[300,407,541,565]" pageId="29" pageNumber="400">
<typeStatus id="54A0882BFFA03B131719FF73FE23FDE1" box="[300,407,541,565]" pageId="29" pageNumber="400" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFA03B1317C5FF73FD20FDE1" bold="true" box="[496,660,541,565]" pageId="29" pageNumber="400">
<collectingCountry id="F30C7619FFA03B1317C5FF73FD3BFDE1" box="[496,655,541,565]" name="Indonesia" pageId="29" pageNumber="400">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFA03B1314AAFF73FF60FD81" country="Indonesia" name="Sulawesi Tengah" pageId="29" pageNumber="400">Central Sulawesi</collectingRegion>
,
<collectingCounty id="62C54E05FFA03B1316D7FF53FEA0FD81" box="[226,276,573,597]" pageId="29" pageNumber="400">Palu</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FFA03B131716FF53FEDEFD81" box="[291,362,573,597]" pageId="29" pageNumber="400">Palolo</collectingMunicipality>
,
<geoCoordinate id="EE2F504EFFA03B13174DFF52FE75FD81" box="[376,449,572,597]" degrees="1" direction="south" minutes="13" orientation="latitude" pageId="29" pageNumber="400" precision="925" value="-1.2166667">1°13S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFA03B1317E5FF52FD82FD81" box="[464,566,572,597]" degrees="120" direction="east" minutes="03" orientation="longitude" pageId="29" pageNumber="400" precision="925" value="120.05">120°03E</geoCoordinate>
,
<collectionCode id="ED0AAE4CFFA03B131471FF50FDDFFD81" box="[580,619,574,597]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="29" pageNumber="400">UV</collectionCode>
lighting,
<collectionCode id="ED0AAE4CFFA03B1314ECFF50FD58FD81" box="[729,748,574,597]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="400">T</collectionCode>
.
<location id="8EC46052FFA03B131644FF30FF69FDA2" LSID="urn:lsid:plazi:treatment:03B2879FFFA03B1316E8FCD3FBB7FF01:8EC46052FFA03B131644FF30FF69FDA2" box="[113,221,606,630]" country="Indonesia" county="Palu" latitude="-1.2166667" longLatPrecision="1308" longitude="120.05" municipality="Palolo" name="Yasunaga" pageId="29" pageNumber="400" stateProvince="Sulawesi Tengah">Yasunaga</location>
,
<date id="FFA51049FFA03B1316DFFF33FE30FDA1" box="[234,388,605,629]" pageId="29" pageNumber="400" value="1988-04-28">
<collectingDate id="EFE1E9A1FFA03B1316DFFF33FE30FDA1" box="[234,388,605,629]" pageId="29" pageNumber="400" value="1988-04-28">28 April 1988</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA03B1317A1FF33FE7FFDA1" box="[404,459,605,629]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="29" pageNumber="400">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFA03B1317D7FF33FD81FDA1" box="[482,565,605,629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="29" pageNumber="400">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFA03B131476FF33FD52FDA1" box="[579,742,605,629]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="29" pageNumber="400">PBI 00378784</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFA03B131644FF10FF7BFD61" collectingDate="1988-04-28" collectionCode="TYCN" country="Indonesia" county="Palu" latitude="-1.2166667" location="Yasunaga" longLatPrecision="1308" longitude="120.05" municipality="Palolo" pageId="29" pageNumber="400" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<typeStatus id="54A0882BFFA03B131644FF10FF67FD42" box="[113,211,638,662]" pageId="29" pageNumber="400" type="paratype">Paratype</typeStatus>
: INDONESIA:
<specimenCount id="9D1DFD00FFA03B13174FFF13FE6CFD41" box="[378,472,637,661]" count="1" pageId="29" pageNumber="400" type="female">1 female</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFA03B13164CFFF3FF7CFD61" box="[121,200,669,693]" pageId="29" pageNumber="400">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FFB3FEA5FC81" pageId="29" pageNumber="400" type="diagnosis">
<paragraph id="8BA43689FFA03B131644FFB3FEA5FC81" blockId="29.[113,752,733,853]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FFB3FF53FD21" bold="true" box="[113,231,733,757]" pageId="29" pageNumber="400">Diagnosis.</emphasis>
This new species is recognised readily by the pale orange-brown basic colouration and entirely shiny fuscous abdomen, in addition to the characters provided in the above key.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FE13FE05F8A1" pageId="29" pageNumber="400" type="description">
<paragraph id="8BA43689FFA03B131644FE13FEE7F8E1" blockId="29.[113,752,893,1845]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FE13FF48FC41" bold="true" box="[113,252,893,917]" pageId="29" pageNumber="400">Description.</emphasis>
Body oval, slightly elongate; basic colouration pale orange-brown, with uniformly shiny chocolate brown abdomen (
<figureCitation id="13202A0CFFA03B1316D2FED3FEC6FC01" box="[231,370,957,981]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="29" pageNumber="400">Fig. 11AD</figureCitation>
). Head pale orange-brown; partly suffused with red; vertex smooth, without basal transverse carina (
<figureCitation id="13202A0CFFA03B1316F4FE93FE91FBC1" box="[193,293,1021,1045]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28D</figureCitation>
); lorum, jugum, and buccula suffused with red; clypeus fuscous. Antenna dark brown; antennomere I pale brown, with darkened base and apex; antennomere II with pale basal ½–⅔ and fuscous extreme base (female); base of antennomere III yellowish brown. Labium pale reddish brown, reaching middle part of metacoxa; apical ¾ of segment IV dark reddish brown. Pronotum uniformly pale, almost glabrous; pleura shiny pale orange-brown; metathoracic scent efferent system with rather small peritreme (
<figureCitation id="13202A0CFFA03B13164CF873FF68FAE1" box="[121,220,1309,1333]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28C</figureCitation>
); scutellum uniformly pale, nearly glabrous (
<figureCitation id="13202A0CFFA03B1314F0F873FF12FA81" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28A</figureCitation>
). Hemelytron pale orange-brown, with uniformly distributed, pale, simple, reclining setae; basal ⅔ and apical margin of cuneus sanguineous; membrane pale smoky brown, semi-transparent, with brown veins. All coxae and legs pale brown; metafemur with two or three reddish rings apically; apical half of each tarsomere III slightly darkened; pretarsal structure as in
<figureCitation id="13202A0CFFA03B131728F893FE34F9C1" box="[285,384,1533,1557]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28F</figureCitation>
; parempodia narrow, lanceolate. Abdomen entirely fuscous. Male genitalia (
<figureCitation id="13202A0CFFA03B131441FB73FD5FF9E1" box="[628,747,1565,1589]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="29" pageNumber="400">Figs. 15A</figureCitation>
,
<figureCitation id="13202A0CFFA03B131644FB53FF08F981" box="[113,188,1597,1621]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">28GJ</figureCitation>
): Hypophysis of left paramere with sharp apex (
<figureCitation id="13202A0CFFA03B1314F0FB53FF10F9A1" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28H</figureCitation>
); right paramere tiny, bulbous, flattened (
<figureCitation id="13202A0CFFA03B13144FFB33FD57F9A1" box="[634,739,1629,1653]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Figs. 28I</figureCitation>
); vesical lobal-sclerites with weak notches (
<figureCitation id="13202A0CFFA03B13140BFB13FD19F941" box="[574,685,1661,1685]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="29" pageNumber="400">Figs. 15A</figureCitation>
,
<figureCitation id="13202A0CFFA03B13148FFB13FD50F941" box="[698,740,1661,1685]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">28J</figureCitation>
); secondary gonopore comparatively large. Female genitalia (
<figureCitation id="13202A0CFFA03B13164CFBD3FEBFF901" box="[121,267,1725,1749]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="29" pageNumber="400">Figs. 15B, C</figureCitation>
,
<figureCitation id="13202A0CFFA03B13172CFBD3FED9F901" box="[281,365,1725,1749]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">28KO</figureCitation>
): Sclerotised rings relatively large, elongate-oval (
<figureCitation id="13202A0CFFA03B131726FBB3FE34F921" box="[275,384,1757,1781]" captionStart="Fig" captionStartId="25.[113,148,1676,1698]" captionTargetBox="[112,1450,189,1661]" captionTargetId="figure-12@25.[112,1450,189,1661]" captionTargetPageId="25" captionText="Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of Himalisundacoris palolo, new species (AC), H. schwartzi, new species (DH) and H. takaii, new species (IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="29" pageNumber="400">Figs. 15B</figureCitation>
,
<figureCitation id="13202A0CFFA03B1317B8FBB3FE0AF921" box="[397,446,1757,1781]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">28K</figureCitation>
); dorsal margin of posterior wall with a pair of fields of spinules which form weak lateral lobes (
<figureCitation id="13202A0CFFA03B13168FFA73FEF2F8E1" box="[186,326,1821,1845]" captionStart="Fig" captionStartId="42.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@42.[135,1473,251,1929]" captionTargetPageId="42" captionText="Fig. 28. Scanning electron micrographs of Himalisundacoris palolo, new species. A, B, head and anterior thorax, dorsal-left lateral views; C, metathoracic scent efferent system; D, head and anterior pronotum, left lateral view; anterior body, dorsal view; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, vesica; K, genital chamber; L, apex of ovipositor (gonapophysis I); MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683020" httpUri="https://zenodo.org/record/14683020/files/figure.png" pageId="29" pageNumber="400">Fig. 28M, N</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFA03B131644FA33FE05F8A1" blockId="29.[113,433,1885,1909]" box="[113,433,1885,1909]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FA33FEABF8A1" bold="true" box="[113,287,1885,1909]" pageId="29" pageNumber="400">Measurements.</emphasis>
See
<tableCitation id="C6990332FFA03B131762FA33FE1EF8A1" box="[343,426,1885,1909]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="29" pageNumber="400">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FAF3FE7CF801" pageId="29" pageNumber="400" type="etymology">
<paragraph id="8BA43689FFA03B131644FAF3FE7CF801" blockId="29.[113,751,1949,2005]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FAF3FF4EF861" bold="true" box="[113,250,1949,1973]" pageId="29" pageNumber="400">Etymology.</emphasis>
Named for the
<typeStatus id="54A0882BFFA03B13178EFAF0FE5FF862" box="[443,491,1950,1974]" pageId="29" pageNumber="400">type</typeStatus>
locality, Palolo of C.
<collectingRegion id="49DFF86BFFA03B131644FAD3FF61F801" box="[113,213,1981,2005]" country="Indonesia" name="Sulawesi Utara" pageId="29" pageNumber="400">Sulawesi</collectingRegion>
; a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FA93FDA3F7C1" box="[113,535,2045,2069]" pageId="29" pageNumber="400" type="distribution">
<paragraph id="8BA43689FFA03B131644FA93FDA3F7C1" blockId="29.[113,535,2045,2069]" box="[113,535,2045,2069]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FA93FEB7F7C1" bold="true" box="[113,259,2045,2069]" pageId="29" pageNumber="400">Distribution.</emphasis>
<collectingCountry id="F30C7619FFA03B131739FA93FEC0F7C1" box="[268,372,2045,2069]" name="Indonesia" pageId="29" pageNumber="400">Indonesia</collectingCountry>
(C.
<collectingRegion id="49DFF86BFFA03B131793FA93FDBFF7C1" box="[422,523,2045,2069]" country="Indonesia" name="Sulawesi Utara" pageId="29" pageNumber="400">Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FDD3FBB7FF01" box="[813,1027,189,213]" pageId="29" pageNumber="400" type="biology_ecology">
<paragraph id="8BA43689FFA03B131518FDD3FBB7FF01" blockId="29.[813,1027,189,213]" box="[813,1027,189,213]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FDD3FC3EFF01" bold="true" box="[813,906,189,213]" pageId="29" pageNumber="400">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,234 @@
<document id="23A5EF52F11C671296D846F61F7F030A" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA03B2F15B2FD93FEF3FF02" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris schwartzi Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="404" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="400" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B2A6B14A11C735748F858CBE68F5082E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0B563E815CB9AD99C8D58A40D90E66A6">
<mods:title id="AB476F3959BEE0991D69F533BC9A67ED">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="EB8166FC905FF2E5584A1F293D9D5ABB" type="personal">
<mods:role id="BE0E586931CD750169918D6D536A4280">
<mods:roleTerm id="62CAA4726193E66AB2C0514CB7CE23BA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05FA154A10274FDDA2680045F057DD0F">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="CE72A8022B2834E47772AD405332B820">text</mods:typeOfResource>
<mods:relatedItem id="1051F60FC86D84A1E7C28632A463894E" type="host">
<mods:titleInfo id="30A37651BA3A6240711703C2053F29D7">
<mods:title id="45707B3DAC38958F4E5DEAEB75725404">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="A4FB0A4E820634A366C4BAA212076D5E">
<mods:date id="664F66E431839F830F6281EF9425970E">2024</mods:date>
<mods:detail id="4C48711DBC203741233A85214E8570ED" type="pubDate">
<mods:number id="B3BCC7763BE35205C8CD8AED0DD457E7">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="E47DF01C31EBCDE737B224AECD7D8CA4" type="volume">
<mods:number id="9444AD49658F4C9BF3CEC71627989BB8">72</mods:number>
</mods:detail>
<mods:extent id="81D4061BAC7892563F181304C3F094B5" unit="page">
<mods:start id="CDD47BB6E549BA64F621404FCF6619E8">371</mods:start>
<mods:end id="1E2F58A947B1F45DA6B636B4B222169E">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A80E06DFEE0A44F273568347D046F635">journal article</mods:classification>
<mods:identifier id="F3A1DE9325D055B7415C03071E378A5D" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="8C53EF1CA593B76EC30ECD75B6936B47" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="D730D87B403D1AAE29F5A4546D6680AA" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="F87CEB050EC4EAF9B12DEFF491568ED3" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA03B2F15B2FD93FEF3FF02" ID-DOI="http://doi.org/10.5281/zenodo.14683034" ID-Zenodo-Dep="14683034" LSID="urn:lsid:plazi:treatment:03B2879FFFA03B2F15B2FD93FEF3FF02" httpUri="http://treatment.plazi.org/id/03B2879FFFA03B2F15B2FD93FEF3FF02" lastPageId="33" lastPageNumber="404" pageId="29" pageNumber="400">
<subSubSection id="C3016502FFA03B1315B2FD93FAE5FEC1" box="[903,1361,253,277]" pageId="29" pageNumber="400" type="nomenclature">
<paragraph id="8BA43689FFA03B1315B2FD93FAE5FEC1" blockId="29.[903,1361,253,309]" box="[903,1361,253,277]" pageId="29" pageNumber="400">
<heading id="D0EC81E5FFA03B1315B2FD93FAE5FEC1" bold="true" box="[903,1361,253,277]" centered="true" fontSize="10" level="2" pageId="29" pageNumber="400" reason="2">
<emphasis id="B96FEA9BFFA03B1315B2FD93FAE5FEC1" bold="true" box="[903,1361,253,277]" pageId="29" pageNumber="400">
<taxonomicName id="4C1B4D0AFFA03B1315B2FD93FB09FEC1" authorityName="Yasunaga" authorityYear="2024" box="[903,1213,253,277]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="400" phylum="Arthropoda" rank="species" species="schwartzi" status="sp. nov.">
<emphasis id="B96FEA9BFFA03B1315B2FD93FB09FEC1" bold="true" box="[903,1213,253,277]" italics="true" pageId="29" pageNumber="400">Himalisundacoris schwartzi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA03B1312F9FD93FAE5FEC1" box="[1228,1361,253,277]" pageId="29" pageNumber="400" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B1315E9FC73FB48FEE1" box="[988,1276,285,309]" pageId="29" pageNumber="400" type="description">
<paragraph id="8BA43689FFA03B1315E9FC73FB48FEE1" blockId="29.[903,1361,253,309]" box="[988,1276,285,309]" pageId="29" pageNumber="400">
(
<figureCitation id="13202A0CFFA03B1315D1FC73FB8EFEE1" box="[996,1082,285,309]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="29" pageNumber="400">Figs. 11</figureCitation>
GJ, 15DH, 29)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FC33FC3AFDC1" pageId="29" pageNumber="400" type="materials_examined">
<paragraph id="8BA43689FFA03B131518FC33FC3AFDC1" blockId="29.[813,1451,349,533]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FC33FC56FEA1" bold="true" box="[813,994,349,373]" pageId="29" pageNumber="400">Type material.</emphasis>
<materialsCitation id="3B733CD4FFA03B1315D8FC33FB8DFE01" collectingDate="1989-12-07" collectionCode="T, ZRC, AMNH" country="Indonesia" county="Brastagi" elevation="1400" latitude="3.18" location="Yasunaga" longLatPrecision="784" longitude="98.5" municipality="North of Toba Lake" pageId="29" pageNumber="400" specimenCode="PBI 00378785" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="B96FEA9BFFA03B1315D8FC33FBEEFEA1" bold="true" box="[1005,1114,349,373]" pageId="29" pageNumber="400">
<typeStatus id="54A0882BFFA03B1315D8FC33FBEEFEA1" box="[1005,1114,349,373]" pageId="29" pageNumber="400" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFA03B131282FC33FAEBFEA1" bold="true" box="[1207,1375,349,373]" pageId="29" pageNumber="400">
<collectingCountry id="F30C7619FFA03B131282FC33FAEDFEA1" box="[1207,1369,349,373]" name="Indonesia" pageId="29" pageNumber="400">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFA03B13135FFC33FC3EFE41" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="400">North Sumatra</collectingRegion>
:
<collectingCounty id="62C54E05FFA03B1315A2FC13FC47FE41" box="[919,1011,381,405]" pageId="29" pageNumber="400">Brastagi</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FFA03B131234FC13FB56FE41" box="[1025,1250,381,405]" pageId="29" pageNumber="400">North of Toba Lake</collectingMunicipality>
,
<geoCoordinate id="EE2F504EFFA03B1312C5FC13FAF4FE41" box="[1264,1344,381,405]" degrees="3.18" direction="north" orientation="latitude" pageId="29" pageNumber="400" precision="555" value="3.18">3.18°N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFA03B13137BFC13FA13FE41" box="[1358,1447,381,405]" degrees="98.50" direction="east" orientation="longitude" pageId="29" pageNumber="400" precision="555" value="98.5">98.50°E</geoCoordinate>
,
<quantity id="4CE39B6CFFA03B131518FCF3FC32FE61" box="[813,902,413,437]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="400" unit="m" value="1400.0">
<elevation id="0036D1BAFFA03B131518FCF3FC32FE61" box="[813,902,413,437]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="29" pageNumber="400" unit="m" value="1400.0">1,400 m</elevation>
</quantity>
, at FL light,
<collectionCode id="ED0AAE4CFFA03B13123BFCF0FB95FE61" box="[1038,1057,414,437]" name="Tavera, Department of Geology and Geophysics" pageId="29" pageNumber="400">T</collectionCode>
.
<location id="8EC46052FFA03B131219FCF0FB2CFE62" LSID="urn:lsid:plazi:treatment:03B2879FFFA03B2F15B2FD93FEF3FF02:8EC46052FFA03B131219FCF0FB2CFE62" box="[1068,1176,414,438]" country="Indonesia" county="Brastagi" latitude="3.18" longLatPrecision="784" longitude="98.5" municipality="North of Toba Lake" name="Yasunaga" pageId="29" pageNumber="400" stateProvince="Sumatera Utara">Yasunaga</location>
,
<date id="FFA51049FFA03B131297FCF0FAEBFE61" box="[1186,1375,413,437]" pageId="29" pageNumber="400" value="1989-12-07">
<collectingDate id="EFE1E9A1FFA03B131297FCF0FAEBFE61" box="[1186,1375,413,437]" pageId="29" pageNumber="400" value="1989-12-07">7 December 1989</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA03B131358FCF3FA10FE61" box="[1389,1444,413,437]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="29" pageNumber="400">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFA03B131501FCD3FC33FE01" box="[820,903,445,469]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="29" pageNumber="400">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFA03B1315A1FCD3FB87FE01" box="[916,1075,445,469]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="29" pageNumber="400">PBI 00378785</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFA03B131277FCD0FC3FFDC1" collectingDate="1989-12-05" collectionCode="TYCN" country="Indonesia" county="Brastagi" elevation="1400" latitude="3.18" location="Yasunaga" longLatPrecision="784" longitude="98.5" municipality="North of Toba Lake" pageId="29" pageNumber="400" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara" typeStatus="paratype">
<typeStatus id="54A0882BFFA03B131277FCD0FB17FE02" box="[1090,1187,446,470]" pageId="29" pageNumber="400" type="paratype">Paratype</typeStatus>
: INDONESIA:
<specimenCount id="9D1DFD00FFA03B13137FFCD3FA13FE01" box="[1354,1447,445,469]" count="1" pageId="29" pageNumber="400" type="female">1 female</specimenCount>
, same data as for holotype, except for date
<date id="FFA51049FFA03B1312DBFCB3FA1FFE21" box="[1262,1451,477,501]" pageId="29" pageNumber="400" value="1989-12-05">
<collectingDate id="EFE1E9A1FFA03B1312DBFCB3FA1FFE21" box="[1262,1451,477,501]" pageId="29" pageNumber="400" value="1989-12-05">5 December 1989</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA03B131501FC93FC37FDC1" box="[820,899,509,533]" pageId="29" pageNumber="400">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FF53FAD8FD01" pageId="29" pageNumber="400" type="diagnosis">
<paragraph id="8BA43689FFA03B131518FF53FAD8FD01" blockId="29.[813,1452,573,725]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FF53FC15FD81" bold="true" box="[813,929,573,597]" pageId="29" pageNumber="400">Diagnosis.</emphasis>
This new species is recognised by the following characters: Pale orange-brown basic colouration; entirely pale creamy brown scutellum; darkened female clavus and posterior corium; and pale reddish-brown abdomen, in addition to the characters provided in the above key.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FF93FBD8F8A1" pageId="29" pageNumber="400" type="description">
<paragraph id="8BA43689FFA03B131518FF93FBF5F8E1" blockId="29.[813,1452,765,1845]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FF93FC0CFCC1" bold="true" box="[813,952,765,789]" pageId="29" pageNumber="400">Description.</emphasis>
Body oval, slightly elongate; basic colouration pale orange-brown (
<figureCitation id="13202A0CFFA03B131233FE73FBDEFCE1" box="[1030,1130,797,821]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="29" pageNumber="400">Fig. 11G</figureCitation>
); in female, clavus and apical part of corium darkened (
<figureCitation id="13202A0CFFA03B131202FE53FB39FC81" box="[1079,1165,829,853]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="29" pageNumber="400">Fig. 11I</figureCitation>
). Head pale orange-brown; vertex smooth, with faintly delimitated basal transverse carina (
<figureCitation id="13202A0CFFA03B131500FE13FC21FC41" box="[821,917,893,917]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29B</figureCitation>
); lorum, jugum, and buccula reddish brown; apical half of clypeus fuscous. Antenna dark brown; antennomere I pale brown, with darkened base and apex; antennomere II with pale subbasal annulation; base of antennomere III yellowish brown. Labium pale reddish brown, reaching middle part of metacoxa; apical part of segment IV dark reddish brown. Pronotum uniformly pale, almost glabrous (
<figureCitation id="13202A0CFFA03B131500F933FC22FBA1" box="[821,918,1117,1141]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29A</figureCitation>
); calli with a pair of reddish spots in female; pleura shiny pale reddish-brown; metathoracic scent efferent system pale brown, with small, semi-circular peritreme (
<figureCitation id="13202A0CFFA03B13130EF9F3FA2BFB61" box="[1339,1439,1181,1205]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29D</figureCitation>
); scutellum pale creamy brown. Hemelytron pale orange-brown, with sparsely distributed, pale, simple, semi-erect setae; clavus and apical half of corium darkened in female; basal half and apex of cuneus sanguineous; membrane pale greyish brown, semi-transparent, with brown veins. All coxae and legs pale brown; metafemur with two or three reddish annulations apically; apex of each tibia dark reddish brown; apical half of each tarsomere III darkened; meta-tarsomere II as long as III (
<figureCitation id="13202A0CFFA03B1315F9F8D3FB9AFA01" box="[972,1070,1469,1493]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29E</figureCitation>
); pretarsal structure as in
<figureCitation id="13202A0CFFA03B13137DF8D3FA13FA01" box="[1352,1447,1469,1493]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29F</figureCitation>
; parempodia relatively short. Abdomen almost uniformly pale reddish brown. Male genitalia (
<figureCitation id="13202A0CFFA03B131283F893FAF4F9C1" box="[1206,1344,1533,1557]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="29" pageNumber="400">Figs. 13DF</figureCitation>
,
<figureCitation id="13202A0CFFA03B131378F893FA15F9C1" box="[1357,1441,1533,1557]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">29GK</figureCitation>
): Hypophysis of left paramere with sharp apex (
<figureCitation id="13202A0CFFA03B13133DFB73FADDF9E1" box="[1288,1385,1565,1589]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29H</figureCitation>
); right paramere ovoid, with short, pointed hypophysis (
<figureCitation id="13202A0CFFA03B131373FB53FA13F981" box="[1350,1447,1597,1621]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="29" pageNumber="400">Figs.13F</figureCitation>
,
<figureCitation id="13202A0CFFA03B131518FB33FCE7F9A1" box="[813,851,1629,1653]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">29I</figureCitation>
); vesica with distinctly notched lobal-sclerite (
<figureCitation id="13202A0CFFA03B131309FB33FA13F9A1" box="[1340,1447,1629,1653]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="29" pageNumber="400">Figs. 13D</figureCitation>
,
<figureCitation id="13202A0CFFA03B131518FB13FCCDF941" box="[813,889,1661,1685]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">29J, K</figureCitation>
); secondary gonopore comparatively large. Female genitalia (
<figureCitation id="13202A0CFFA03B1315ADFBF3FBA8F961" box="[920,1052,1693,1717]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="29" pageNumber="400">Figs. 13H, I</figureCitation>
,
<figureCitation id="13202A0CFFA03B13121DFBF3FBCDF961" box="[1064,1145,1693,1717]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">29LO</figureCitation>
): Sclerotised rings relatively large, subtriangular (
<figureCitation id="13202A0CFFA03B131237FBD3FBEBF901" box="[1026,1119,1725,1749]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29L</figureCitation>
); dorsal margin of posterior wall with sparsely distributed, scaly micro-structures, lacking field of spinules (
<figureCitation id="13202A0CFFA03B131580FB93FBAFF8C1" box="[949,1051,1789,1813]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29N</figureCitation>
); interramal lobes mesally separated to each other (
<figureCitation id="13202A0CFFA03B1315FEFA73FB81F8E1" box="[971,1077,1821,1845]" captionStart="Fig" captionStartId="43.[113,148,1959,1981]" captionTargetBox="[113,1451,265,1943]" captionTargetId="figure-11@43.[113,1451,265,1943]" captionTargetPageId="43" captionText="Fig. 29. Scanning electron micrographs of Himalisundacoris schwartzi, new species. A, B, head and pronotum, dorsal-left lateral views; C, pronotum, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, metatarsus; F, pretarsal structure of metaleg; G, H, left paramere; I, right paramere; J, K, vesica; L, sclerotised ring; MO, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683022" httpUri="https://zenodo.org/record/14683022/files/figure.png" pageId="29" pageNumber="400">Fig. 29M</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFA03B131518FA33FBD8F8A1" blockId="29.[813,1132,1885,1909]" box="[813,1132,1885,1909]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FA33FC6FF8A1" bold="true" box="[813,987,1885,1909]" pageId="29" pageNumber="400">Measurements.</emphasis>
See
<tableCitation id="C6990332FFA03B131227FA33FBD2F8A1" box="[1042,1126,1885,1909]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="29" pageNumber="400">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FAF3FBF1F801" pageId="29" pageNumber="400" type="etymology">
<paragraph id="8BA43689FFA03B131518FAF3FBF1F801" blockId="29.[813,1451,1949,2005]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FAF3FC1BF861" bold="true" box="[813,943,1949,1973]" pageId="29" pageNumber="400">Etymology.</emphasis>
Named to honour Dr. Michael D. Schwartz; a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131518FA93FB78F7C1" box="[813,1228,2045,2069]" pageId="29" pageNumber="400" type="distribution">
<paragraph id="8BA43689FFA03B131518FA93FB78F7C1" blockId="29.[813,1228,2045,2069]" box="[813,1228,2045,2069]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131518FA93FC0BF7C1" bold="true" box="[813,959,2045,2069]" pageId="29" pageNumber="400">Distribution.</emphasis>
<collectingCountry id="F30C7619FFA03B1315F2FA93FB9BF7C1" box="[967,1071,2045,2069]" name="Indonesia" pageId="29" pageNumber="400">Indonesia</collectingCountry>
(N.
<collectingRegion id="49DFF86BFFA03B131256FA93FB75F7C1" box="[1123,1217,2045,2069]" country="Indonesia" name="Sumatera Utara" pageId="29" pageNumber="400">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="DF646601FFA33B1016BDFAF7FB04F7D2" ID-DOI="http://doi.org/10.5281/zenodo.14682996" ID-Zenodo-Dep="14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="30" pageNumber="401" startId="30.[136,171,1945,1967]" targetBox="[136,1476,250,1930]" targetPageId="30" targetType="figure">
<paragraph id="8BA43689FFA33B1016BDFAF7FB04F7D2" blockId="30.[136,1474,1945,2054]" pageId="30" pageNumber="401">
Fig. 19. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFFA33B101405FAF7FD47F87A" authorityName="Yasunaga" authorityYear="2024" box="[560,755,1945,1966]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="401" phylum="Arthropoda" rank="species" species="cheroti" status="sp. nov.">
<emphasis id="B96FEA9BFFA33B101405FAF7FD47F87A" box="[560,755,1945,1966]" italics="true" pageId="30" pageNumber="401">Chelyolygus cheroti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA33B101534FAF4FCCDF87B" box="[769,889,1945,1967]" pageId="30" pageNumber="401" rank="species">new species</taxonomicNameLabel>
. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall.
</paragraph>
</caption>
<caption id="DF646601FFA23B111644FAF6FE48F7D0" ID-DOI="http://doi.org/10.5281/zenodo.14682998" ID-Zenodo-Dep="14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="31" pageNumber="402" startId="31.[113,148,1944,1966]" targetBox="[113,1451,251,1929]" targetPageId="31" targetType="figure">
<paragraph id="8BA43689FFA23B111644FAF6FE48F7D0" blockId="31.[113,1451,1944,2053]" pageId="31" pageNumber="402">
Fig. 20. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFFA23B11142FFAF6FCA8F879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[538,796,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="402" phylum="Arthropoda" rank="species" species="bipuncticollis">
<emphasis id="B96FEA9BFFA23B11142FFAF6FCA8F879" box="[538,796,1944,1965]" italics="true" pageId="31" pageNumber="402">Chelyolygus bipuncticollis</emphasis>
</taxonomicName>
(male) and
<taxonomicName id="4C1B4D0AFFA23B1115A2FAF7FB9EF879" authorityName="Yasunaga" authorityYear="2024" box="[919,1066,1944,1966]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="402" phylum="Arthropoda" rank="species" species="tabananicus" status="new species">
<emphasis id="B96FEA9BFFA23B1115A2FAF7FB9EF879" box="[919,1066,1944,1966]" italics="true" pageId="31" pageNumber="402">C. tabananicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA23B11120CFAF7FB1BF87A" box="[1081,1199,1944,1966]" pageId="31" pageNumber="402" rank="species">new species</taxonomicNameLabel>
(holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure.
</paragraph>
</caption>
<caption id="DF646601FF9D3B2E16BDFAF9FD0CF7D7" ID-DOI="http://doi.org/10.5281/zenodo.14683000" ID-Zenodo-Dep="14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="32" pageNumber="403" startId="32.[136,171,1943,1965]" targetBox="[136,1474,252,1927]" targetPageId="32" targetType="figure">
<paragraph id="8BA43689FF9D3B2E16BDFAF9FD0CF7D7" blockId="32.[136,1475,1943,2051]" pageId="32" pageNumber="403">
Fig. 21. Scanning electron micrographs of
<taxonomicName id="4C1B4D0AFF9D3B2E1412FAF9FC94F878" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[551,800,1943,1964]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="403" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFF9D3B2E1412FAF9FC94F878" box="[551,800,1943,1964]" italics="true" pageId="32" pageNumber="403">Chelyolygus fuhoshoensis</emphasis>
</taxonomicName>
(AG) and
<taxonomicName id="4C1B4D0AFF9D3B2E15A3FAF9FBBFF879" authorityName="Yasunaga" authorityYear="2024" box="[918,1035,1943,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="403" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFF9D3B2E15A3FAF9FBBFF879" box="[918,1035,1943,1965]" italics="true" pageId="32" pageNumber="403">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FF9D3B2E122DFAF9FB3FF879" box="[1048,1163,1943,1965]" pageId="32" pageNumber="403" rank="species">new species</taxonomicNameLabel>
(HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view.
</paragraph>
</caption>
<subSubSection id="C3016502FF9C3B2F1644FDD3FEF3FF02" box="[113,327,189,214]" pageId="33" pageNumber="404" type="biology_ecology">
<paragraph id="8BA43689FF9C3B2F1644FDD3FEF3FF02" blockId="33.[113,327,189,214]" box="[113,327,189,214]" pageId="33" pageNumber="404">
<emphasis id="B96FEA9BFF9C3B2F1644FDD3FF7AFF01" bold="true" box="[113,206,189,213]" pageId="33" pageNumber="404">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,251 @@
<document id="ACBE9AEBBD83AF4D57CEA29B1D9BBCE4" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA53B131585FEB3FEF3FE41" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris omiae Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="400" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="395" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="FB4D622F8C9A870EAA1C3FC0EDA90163" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="95D31C4FE16950BE90B83E161AA9BEB7">
<mods:title id="3DA9CDC18D692762F69C8CAFE428CF99">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="9AB6F560C818AE3DE42378CD5BC1E7FC" type="personal">
<mods:role id="3C781B0D017659ABC9700B35A9CC7AA6">
<mods:roleTerm id="AA14F47A33F666ECD98D5363773F8C64">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FD99745035AD7B8CCCFF128820FA83D">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="8EAF7336791C9F219DA941A15C732962">text</mods:typeOfResource>
<mods:relatedItem id="B951A598E8348DAC4E628EA5645C33C7" type="host">
<mods:titleInfo id="3A3E94F82365B2996948A21076032098">
<mods:title id="D4D3AABFBA5AD8BBB85D2888B962D7F8">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="B2EC6E7863357D73A49361982FEC8DC7">
<mods:date id="17FB3CB60F6D209BC813B55954FA9B59">2024</mods:date>
<mods:detail id="A18A51E66C0A146E8231E9EF3892BC49" type="pubDate">
<mods:number id="E479E78DB6AE04CCC6F5DD91F9A7C8EB">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="157EB0B1E1E54565F768DBF669E42503" type="volume">
<mods:number id="F2B7186CFCB996954B0DE046F2A17DB7">72</mods:number>
</mods:detail>
<mods:extent id="535743FAB2887917362C56FA879A1DEA" unit="page">
<mods:start id="99ADFFAD818758CB43E53D41917F1478">371</mods:start>
<mods:end id="C3246CA2F785CF300E23CDD63F80A400">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F8EC7B22DDCAFE5D58EA456B9C66F836">journal article</mods:classification>
<mods:identifier id="5373FC02BE278D92D9A1AAFA18D8E8DC" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="F9DC597D949C20A3DE3FF2B0F9DEA6F3" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E610D647D2253BE9DB9344533FD08CB2" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="5D2F87DF0E84666EE501A834272BED55" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA53B131585FEB3FEF3FE41" LSID="urn:lsid:plazi:treatment:03B2879FFFA53B131585FEB3FEF3FE41" httpUri="http://treatment.plazi.org/id/03B2879FFFA53B131585FEB3FEF3FE41" lastPageId="29" lastPageNumber="400" pageId="24" pageNumber="395">
<subSubSection id="C3016502FFA53B161585FEB3FAE1FC21" box="[944,1365,989,1013]" pageId="24" pageNumber="395" type="nomenclature">
<paragraph id="8BA43689FFA53B161585FEB3FAE1FC21" blockId="24.[944,1365,989,1045]" box="[944,1365,989,1013]" pageId="24" pageNumber="395">
<heading id="D0EC81E5FFA53B161585FEB3FAE1FC21" bold="true" box="[944,1365,989,1013]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="395" reason="2">
<emphasis id="B96FEA9BFFA53B161585FEB3FAE1FC21" bold="true" box="[944,1365,989,1013]" pageId="24" pageNumber="395">
<taxonomicName id="4C1B4D0AFFA53B161585FEB3FB75FC21" authorityName="Yasunaga" authorityYear="2024" box="[944,1217,989,1013]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="omiae" status="sp. nov.">
<emphasis id="B96FEA9BFFA53B161585FEB3FB75FC21" bold="true" box="[944,1217,989,1013]" italics="true" pageId="24" pageNumber="395">Himalisundacoris omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA53B1612E5FEB3FAE1FC21" box="[1232,1365,989,1013]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161585FE93FAE1FBC1" box="[944,1365,1021,1045]" pageId="24" pageNumber="395" type="description">
<paragraph id="8BA43689FFA53B161585FE93FAE1FBC1" blockId="24.[944,1365,989,1045]" box="[944,1365,1021,1045]" pageId="24" pageNumber="395">
(
<figureCitation id="13202A0CFFA53B16158DFE93FBBBFBC1" box="[952,1039,1021,1045]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="24" pageNumber="395">Figs. 12</figureCitation>
AD, 14AE, 26JO, 27JO)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161576F953FBBFFB01" pageId="24" pageNumber="395" type="materials_examined">
<paragraph id="8BA43689FFA53B161576F953FBBFFB01" blockId="24.[835,1474,1085,1237]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576F953FC4FFB81" bold="true" box="[835,1019,1085,1109]" pageId="24" pageNumber="395">Type material.</emphasis>
<materialsCitation id="3B733CD4FFA53B161232F953FA09FB41" collectingDate="1996-08-08" collectingDateMax="1996-08-12" collectingDateMin="1996-08-08" collectionCode="ZRC, AMNH" country="Indonesia" latitude="-8.266666" location="Bedugul Area" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" pageId="24" pageNumber="395" specimenCode="PBI 00378783" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<emphasis id="B96FEA9BFFA53B161232F953FBC2FB81" bold="true" box="[1031,1142,1085,1109]" pageId="24" pageNumber="395">
<typeStatus id="54A0882BFFA53B161232F953FBC2FB81" box="[1031,1142,1085,1109]" pageId="24" pageNumber="395" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFA53B1612E0F953FACBFB81" bold="true" box="[1237,1407,1085,1109]" pageId="24" pageNumber="395">
<collectingCountry id="F30C7619FFA53B1612E0F953FACDFB81" box="[1237,1401,1085,1109]" name="Indonesia" pageId="24" pageNumber="395">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFA53B1613BEF953FA09FB81" box="[1419,1469,1085,1109]" country="Indonesia" name="Bali" pageId="24" pageNumber="395">Bali</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFA53B161576F933FC16FBA1" box="[835,930,1117,1141]" pageId="24" pageNumber="395">Tabanan</collectingMunicipality>
,
<location id="8EC46052FFA53B16159AF933FBFEFBA1" LSID="urn:lsid:plazi:treatment:03B2879FFFA53B131585FEB3FEF3FE41:8EC46052FFA53B16159AF933FBFEFBA1" box="[943,1098,1117,1141]" country="Indonesia" latitude="-8.266666" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" name="Bedugul Area" pageId="24" pageNumber="395" stateProvince="Bali">Bedugul Area</location>
,
<geoCoordinate id="EE2F504EFFA53B161263F932FB14FBA1" box="[1110,1184,1116,1141]" degrees="8" direction="south" minutes="16" orientation="latitude" pageId="24" pageNumber="395" precision="925" value="-8.266666">8°16S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFA53B16129BF932FAA0FBA1" box="[1198,1300,1116,1141]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="24" pageNumber="395" precision="925" value="115.15">115°09E</geoCoordinate>
, local collector,
<date id="FFA51049FFA53B161576F913FB93FB41" box="[835,1063,1149,1173]" pageId="24" pageNumber="395" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">
<collectingDate id="EFE1E9A1FFA53B161576F913FB93FB41" box="[835,1063,1149,1173]" pageId="24" pageNumber="395" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">812 August 1996</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA53B161208F913FBCDFB41" box="[1085,1145,1149,1173]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="395">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFA53B1612A3F913FB44FB41" box="[1174,1264,1149,1173]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="395">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFA53B1612CAF913FA01FB41" box="[1279,1461,1149,1173]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="395">PBI 00378783</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFA53B161576F9F0FBB3FB01" collectingDate="1996-08-08" collectingDateMax="1996-08-12" collectingDateMin="1996-08-08" collectionCode="TYCN" country="Indonesia" latitude="-8.266666" location="Bedugul Area" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" pageId="24" pageNumber="395" specimenCount="3" specimenCount-female="3" stateProvince="Bali" typeStatus="paratype">
<typeStatus id="54A0882BFFA53B161576F9F0FC0CFB62" box="[835,952,1182,1206]" pageId="24" pageNumber="395" type="paratype">Paratypes</typeStatus>
: INDONESIA:
<specimenCount id="9D1DFD00FFA53B161243F9F3FB58FB61" box="[1142,1260,1181,1205]" count="3" pageId="24" pageNumber="395" type="female">3 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFA53B161584F9D3FBB4FB01" box="[945,1024,1213,1237]" pageId="24" pageNumber="395">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161576F993FA77FA01" pageId="24" pageNumber="395" type="diagnosis">
<paragraph id="8BA43689FFA53B161576F993FA77FA01" blockId="24.[835,1475,1277,1493]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576F993FC0CFAC1" bold="true" box="[835,952,1277,1301]" pageId="24" pageNumber="395">Diagnosis.</emphasis>
This new species is recognised by the following characters: Oily shiny, pale milky olive dorsum with reddish brown band at subapical part of corium; sharp apex of right paramere hypophysis (
<figureCitation id="13202A0CFFA53B161244F833FB67FAA1" box="[1137,1235,1373,1397]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="24" pageNumber="395">Fig. 14B</figureCitation>
); rounded apex of left paramere hypophysis (
<figureCitation id="13202A0CFFA53B161206F813FB21FA41" box="[1075,1173,1405,1429]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">Fig. 27K</figureCitation>
); relatively developed lobal-sclerites and large secondary gonopore (
<figureCitation id="13202A0CFFA53B1612DDF8F3FAE5FA61" box="[1256,1361,1437,1461]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="24" pageNumber="395">Figs. 14C</figureCitation>
,
<figureCitation id="13202A0CFFA53B16136EF8F3FA3EFA61" box="[1371,1418,1437,1461]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27L</figureCitation>
); and large spinules on dorsal margin of posterior wall (
<figureCitation id="13202A0CFFA53B16137AF8D3FA02FA01" box="[1359,1462,1469,1493]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">Fig. 27M</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B171576F893FBD8F7DF" lastPageId="25" lastPageNumber="396" pageId="24" pageNumber="395" type="description">
<paragraph id="8BA43689FFA53B171576F893FA91F81F" blockId="24.[835,1474,1533,2101]" lastBlockId="25.[813,1452,1811,1995]" lastPageId="25" lastPageNumber="396" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576F893FC7AF9C1" bold="true" box="[835,974,1533,1557]" pageId="24" pageNumber="395">Description.</emphasis>
Body generally pale milky olive, oval; dorsum oily shiny, well-polished (
<figureCitation id="13202A0CFFA53B161248FB73FB5EF9E1" box="[1149,1258,1565,1589]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26C</figureCitation>
). Head shiny pale brown, partly suffused with red; basal transverse carina of vertex almost obliterated (
<figureCitation id="13202A0CFFA53B161257FB33FB56F9A1" box="[1122,1250,1629,1653]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26J, K</figureCitation>
); lorum and clypeus fuscous. Antenna dark brown; antennomere I pale brown with darkened base and apex; antennomere II with a pale, narrow annulation near base (male)/ with pale basal half and fuscous extreme base (female); base of antennomere III creamy yellowish brown. Labium pale reddish brown, reaching apex of metacoxa; base of segment I and apical ¾ of segment IV darkened. Pronotum uniformly pale, almost glabrous; calli slightly tinged with red; pleura shiny pale reddish brown; metathoracic scent efferent system creamy yellow, with rather flat peritreme (
<figureCitation id="13202A0CFFA53B161288FAF3FA9CF861" box="[1213,1320,1949,1973]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26M</figureCitation>
); mesoscutum and scutellum with sparsely distributed, short setae (
<figureCitation id="13202A0CFFA53B16136CFAD3FA03F801" box="[1369,1463,1981,2005]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26L</figureCitation>
). Hemelytron oily shiny, smooth, with sparsely distributed, pale, short, simple setae (
<figureCitation id="13202A0CFFA53B161264FA93FB18F7C1" box="[1105,1196,2045,2069]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26J</figureCitation>
); subapical part of clavus, apical ⅓ of corium, and basal ½ and apex of cuneus reddish brown; membrane pale smoky brown, semi-transparent, with brown veins. All coxae and legs shiny pale brown; metafemur with two or three reddish rings apically; each tarsomere III darkened; pretarsal structure as in
<figureCitation id="13202A0CFFA43B1714B2FA1DFD58F85F" box="[647,748,1907,1931]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="25" pageNumber="396">Fig. 26O</figureCitation>
; parempodia relatively narrow. Abdomen pale brown; male genital segment (pygophore) shiny fuscous (
<figureCitation id="13202A0CFFA43B17144DFADDFD57F81F" box="[632,739,1971,1995]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="25" pageNumber="396">Fig. 12B</figureCitation>
); ventral median part more or less darkened in female (
<figureCitation id="13202A0CFFA43B1714F0FABDFF10F7DF" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="25" pageNumber="396">Fig. 12D</figureCitation>
). Male genitalia (
<figureCitation id="13202A0CFFA43B171758FA9DFE7CF7DF" box="[365,456,2035,2059]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="25" pageNumber="396">Figs. 14</figureCitation>
AC, 27JL): Hypophysis of left paramere with somewhat rounded apex (
<figureCitation id="13202A0CFFA43B171448F57DFD57F7FF" box="[637,739,2067,2091]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="25" pageNumber="396">Fig. 27K</figureCitation>
); right paramere with short, pointed hypophysis (
<figureCitation id="13202A0CFFA43B17130FFA7DFA2AF8FF" box="[1338,1438,1811,1835]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="25" pageNumber="396">Fig. 14B</figureCitation>
); vesical lobal-sclerites with rather developed notches (
<figureCitation id="13202A0CFFA43B171343FA5DFCEDF8BF" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="25" pageNumber="396">Figs. 14F</figureCitation>
,
<figureCitation id="13202A0CFFA43B171557FA3DFC25F8BF" box="[866,913,1875,1899]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="25" pageNumber="396">27L</figureCitation>
); secondary gonopore comparatively large. Female genitalia (
<figureCitation id="13202A0CFFA43B1715ADFA1DFB92F85E" box="[920,1062,1907,1931]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="25" pageNumber="396">Figs. 14D, E</figureCitation>
,
<figureCitation id="13202A0CFFA43B171207FA1DFB3EF85F" box="[1074,1162,1907,1931]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="25" pageNumber="396">27MO</figureCitation>
): Sclerotised rings narrow, subtriangular (
<figureCitation id="13202A0CFFA43B1715F9FAFDFB87F87F" box="[972,1075,1939,1963]" captionStart="Fig" captionStartId="23.[113,148,1904,1926]" captionTargetBox="[113,1451,316,1889]" captionTargetId="figure-11@23.[113,1451,316,1889]" captionTargetPageId="23" captionText="Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of Himalisundacoris omiae, new species (AE) and H. theoi, new species (FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="25" pageNumber="396">Figs. 14I</figureCitation>
,
<figureCitation id="13202A0CFFA43B171274FAFDFBC7F87F" box="[1089,1139,1939,1963]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="25" pageNumber="396">27O</figureCitation>
); dorsal margin of posterior wall with relatively large spinules (
<figureCitation id="13202A0CFFA43B17129AFADDFAADF81F" box="[1199,1305,1971,1995]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="25" pageNumber="396">Fig. 27M</figureCitation>
).
</paragraph>
<caption id="DF646601FFA43B171644FBE2FD0DF90F" ID-DOI="http://doi.org/10.5281/zenodo.14682986" ID-Zenodo-Dep="14682986" httpUri="https://zenodo.org/record/14682986/files/figure.png" pageId="25" pageNumber="396" startId="25.[113,148,1676,1698]" targetBox="[112,1450,189,1661]" targetPageId="25" targetType="figure">
<paragraph id="8BA43689FFA43B171644FBE2FD0DF90F" blockId="25.[113,1452,1676,1756]" pageId="25" pageNumber="396">
Fig. 15. Male (A, DF, IK) and female (B, C, G, H) genitalia of
<taxonomicName id="4C1B4D0AFFA43B1714DAFBE2FC6BF975" authorityName="Yasunaga" authorityYear="2024" box="[751,991,1676,1697]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="396" phylum="Arthropoda" rank="species" species="palolo" status="sp. nov.">
<emphasis id="B96FEA9BFFA43B1714DAFBE2FC6BF975" box="[751,991,1676,1697]" italics="true" pageId="25" pageNumber="396">Himalisundacoris palolo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA43B1715D9FBE3FBD4F976" box="[1004,1120,1676,1698]" pageId="25" pageNumber="396" rank="species">new species</taxonomicNameLabel>
(AC),
<taxonomicName id="4C1B4D0AFFA43B171285FBE3FA9EF975" authorityName="Yasunaga" authorityYear="2024" box="[1200,1322,1676,1698]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="396" phylum="Arthropoda" rank="species" species="schwartzi" status="new species">
<emphasis id="B96FEA9BFFA43B171285FBE3FA9EF975" box="[1200,1322,1676,1698]" italics="true" pageId="25" pageNumber="396">H. schwartzi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA43B171302FBE3FA1FF976" box="[1335,1451,1676,1698]" pageId="25" pageNumber="396" rank="species">new species</taxonomicNameLabel>
(DH) and
<taxonomicName id="4C1B4D0AFFA43B1716EBFBC4FE86F96A" authorityName="Yasunaga" authorityYear="2024" box="[222,306,1705,1727]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="396" phylum="Arthropoda" rank="species" species="takaii" status="new species">
<emphasis id="B96FEA9BFFA43B1716EBFBC4FE86F96A" box="[222,306,1705,1727]" italics="true" pageId="25" pageNumber="396">H. takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA43B17170BFBC7FE05F96B" box="[318,433,1705,1727]" pageId="25" pageNumber="396" rank="species">new species</taxonomicNameLabel>
(IK). A, D, I, Vesica; B, genital chamber; C, H, posterior wall; E, J, left paramere; F, K, right paramere; G, apex of ovipositor (gonapophysis I). Scale bars 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BA43689FFA43B171518FA9DFBD8F7DF" blockId="25.[813,1132,2035,2059]" box="[813,1132,2035,2059]" pageId="25" pageNumber="396">
<emphasis id="B96FEA9BFFA43B171518FA9DFC6FF7DF" bold="true" box="[813,987,2035,2059]" pageId="25" pageNumber="396">Measurements.</emphasis>
See
<tableCitation id="C6990332FFA43B171227FA9DFBD2F7DF" box="[1042,1126,2035,2059]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="25" pageNumber="396">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF646601FFA73B1416BDFA02FED0F86F" ID-DOI="http://doi.org/10.5281/zenodo.14682988" ID-Zenodo-Dep="14682988" httpUri="https://zenodo.org/record/14682988/files/figure.png" pageId="26" pageNumber="397" startId="26.[136,171,1900,1922]" targetBox="[136,1474,324,1884]" targetPageId="26" targetType="figure">
<paragraph id="8BA43689FFA73B1416BDFA02FED0F86F" blockId="26.[136,1475,1899,1979]" pageId="26" pageNumber="397">
Fig. 16. Male (AD) and female (EG) genitalia of
<taxonomicName id="4C1B4D0AFFA73B14144AFA02FCE0F855" authorityName="Yasunaga" authorityYear="2024" box="[639,852,1900,1921]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="397" phylum="Arthropoda" rank="species" species="balicus" status="sp. nov.">
<emphasis id="B96FEA9BFFA73B14144AFA02FCE0F855" box="[639,852,1900,1921]" italics="true" pageId="26" pageNumber="397">Paralygocoris balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA73B141554FA02FC6CF856" box="[865,984,1900,1922]" pageId="26" pageNumber="397" rank="species">new species</taxonomicNameLabel>
. A, B, left paramere; C, right paramere; D, vesica; G, ditto, secondary gonopore and adjacent structures; E, posterior wall; F, genital chamber; G, apical parts of ovipositors (gonapophyses I). Scale bars 0.2 mm.
</paragraph>
</caption>
<caption id="DF646601FFA63B151644FA04FD0EF86D" ID-DOI="http://doi.org/10.5281/zenodo.14682990" ID-Zenodo-Dep="14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="27" pageNumber="398" startId="27.[113,148,1898,1920]" targetBox="[113,1451,330,1882]" targetPageId="27" targetType="figure">
<paragraph id="8BA43689FFA63B151644FA04FD0EF86D" blockId="27.[113,1452,1898,1977]" pageId="27" pageNumber="398">
Fig. 17. Male (AE) and female (FH) genitalia of
<taxonomicName id="4C1B4D0AFFA63B15144CFA04FCD0F8AB" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[633,868,1898,1919]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="398" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFFA63B15144CFA04FCD0F8AB" box="[633,868,1898,1919]" italics="true" pageId="27" pageNumber="398">Paralygocoris vittulatus</emphasis>
</taxonomicName>
(AF) and
<taxonomicName id="4C1B4D0AFFA63B1515E8FA04FBC0F8AB" authorityName="Lu &amp; Zheng" authorityYear="2004" box="[989,1140,1898,1919]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="27" pageNumber="398" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFFA63B1515E8FA04FBC0F8AB" box="[989,1140,1898,1919]" italics="true" pageId="27" pageNumber="398">P. meridionalis</emphasis>
</taxonomicName>
(GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm.
</paragraph>
</caption>
<caption id="DF646601FFA13B1216B2FAFDFC26F836" ID-DOI="http://doi.org/10.5281/zenodo.14682992" ID-Zenodo-Dep="14682992" httpUri="https://zenodo.org/record/14682992/files/figure.png" pageId="28" pageNumber="399" startId="28.[135,170,1939,1961]" targetBox="[135,1473,285,1924]" targetPageId="28" targetType="figure">
<paragraph id="8BA43689FFA13B1216B2FAFDFC26F836" blockId="28.[135,1474,1939,2019]" pageId="28" pageNumber="399">
Fig. 18. Male (AD, H) and female (EG, I, J) genitalia of
<taxonomicName id="4C1B4D0AFFA13B1214D1FAFDFC15F87C" authorityName="Yasunaga" authorityYear="2024" box="[740,929,1939,1960]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="399" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFFA13B1214D1FAFDFC15F87C" box="[740,929,1939,1960]" italics="true" pageId="28" pageNumber="399">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA13B121585FAFAFB92F87D" box="[944,1062,1939,1961]" pageId="28" pageNumber="399" rank="species">new species</taxonomicNameLabel>
(AG) and
<taxonomicName id="4C1B4D0AFFA13B121296FAFAFAB5F87D" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[1187,1281,1940,1961]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="399" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFFA13B121296FAFAFAB5F87D" box="[1187,1281,1940,1961]" italics="true" pageId="28" pageNumber="399">S. sauteri</emphasis>
</taxonomicName>
(HJ). A, Apex of pygophore with parameres, dorsal view; B, apex of left paramere; C, apical part of right paramere; D, H, vesica; E, genital chamber; F, I, posterior wall; G, J, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3016502FFA03B131644FDD3FE46FEC1" pageId="29" pageNumber="400" type="etymology">
<paragraph id="8BA43689FFA03B131644FDD3FE46FEC1" blockId="29.[113,752,189,277]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FDD3FF42FF01" bold="true" box="[113,246,189,213]" pageId="29" pageNumber="400">Etymology.</emphasis>
Named after Omi, the granddaughter of Dr. Michael D. Schwartz, long-time friend and colleague of the author; a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FC53FE75FE81" box="[113,449,317,341]" pageId="29" pageNumber="400" type="distribution">
<paragraph id="8BA43689FFA03B131644FC53FE75FE81" blockId="29.[113,449,317,341]" box="[113,449,317,341]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FC53FEB7FE81" bold="true" box="[113,259,317,341]" pageId="29" pageNumber="400">Distribution.</emphasis>
<collectingCountry id="F30C7619FFA03B131739FC53FEC0FE81" box="[268,372,317,341]" name="Indonesia" pageId="29" pageNumber="400">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFFA03B1317B1FC53FE01FE81" box="[388,437,317,341]" country="Indonesia" name="Bali" pageId="29" pageNumber="400">Bali</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA03B131644FC13FEF3FE41" box="[113,327,381,405]" pageId="29" pageNumber="400" type="biology_ecology">
<paragraph id="8BA43689FFA03B131644FC13FEF3FE41" blockId="29.[113,327,381,405]" box="[113,327,381,405]" pageId="29" pageNumber="400">
<emphasis id="B96FEA9BFFA03B131644FC13FF7AFE41" bold="true" box="[113,206,381,405]" pageId="29" pageNumber="400">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,228 @@
<document id="6B6ECD42627F66C044A1CFC591801ED6" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA53B1616DDFE55FABFFC61" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris duwalae Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="395" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="395" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="58516DE4AFA28351CF329077A2231105" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AAA1B8C987C8285DC94284AF52A37172">
<mods:title id="1C12C0CEEBD7DB6F55C70178C328292E">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="3828187B8F9E975F16BD6DA984E1C9A5" type="personal">
<mods:role id="816386917FD95175A02E742270E04D3F">
<mods:roleTerm id="23F381068B81708786B793ECD0F7A298">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DCE0E3814444B16D8E371951323905A7">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="CBD3FAD93E710323898B6F927518C45D">text</mods:typeOfResource>
<mods:relatedItem id="A531ED5929F31D2CEF854D1874588863" type="host">
<mods:titleInfo id="2127FDC8373E9303C483D649C95303E2">
<mods:title id="93DB257EFC37EFAF3B5A7557C949EB9A">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="51E90801B688F4D25B74CFBDD4B27CA6">
<mods:date id="E3CB50ED59DF70B70D53F43676CC68AF">2024</mods:date>
<mods:detail id="11F3232D52103F3E29DC9666A0E8E775" type="pubDate">
<mods:number id="82935EC64215F1B38C1BEF848226D8E6">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="F97628AD526149DDB067A5B61998572D" type="volume">
<mods:number id="49574E7FD20E2CE51F0DEDF9A616D47E">72</mods:number>
</mods:detail>
<mods:extent id="AD9FD5FF07C9F2D5BBF2F6B48428E478" unit="page">
<mods:start id="927446BB1091FFD117D18B61DFD21D5F">371</mods:start>
<mods:end id="C87C024493A8644E0AFFCBEDDD31119C">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C0D64DC4E44DD9EA34E6C4A480213E3B">journal article</mods:classification>
<mods:identifier id="B50985F1E316D2A27699ADDAA87BEF44" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="8E0754A5920A788C22289CB2ED818742" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F47AA597D7D36C5663B1789900974B4B" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="33F085F45FA2C92E5C3E1B5F8467D150" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA53B1616DDFE55FABFFC61" ID-DOI="http://doi.org/10.5281/zenodo.14683016" ID-Zenodo-Dep="14683016" LSID="urn:lsid:plazi:treatment:03B2879FFFA53B1616DDFE55FABFFC61" httpUri="http://treatment.plazi.org/id/03B2879FFFA53B1616DDFE55FABFFC61" lastPageNumber="395" pageId="24" pageNumber="395">
<subSubSection id="C3016502FFA53B1616DDFE55FD13FC87" box="[232,679,827,852]" pageId="24" pageNumber="395" type="nomenclature">
<paragraph id="8BA43689FFA53B1616DDFE55FD13FC87" blockId="24.[232,679,827,884]" box="[232,679,827,852]" pageId="24" pageNumber="395">
<heading id="D0EC81E5FFA53B1616DDFE55FD13FC87" bold="true" box="[232,679,827,852]" centered="true" fontSize="10" level="2" pageId="24" pageNumber="395" reason="2">
<emphasis id="B96FEA9BFFA53B1616DDFE55FD13FC87" bold="true" box="[232,679,827,852]" pageId="24" pageNumber="395">
<taxonomicName id="4C1B4D0AFFA53B1616DDFE55FDA6FC87" authorityName="Yasunaga" authorityYear="2024" box="[232,530,827,851]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFFA53B1616DDFE55FDA6FC87" bold="true" box="[232,530,827,851]" italics="true" pageId="24" pageNumber="395">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA53B161414FE52FD13FC87" box="[545,679,827,852]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B16172FFE35FDC1FCA0" box="[282,629,859,884]" pageId="24" pageNumber="395" type="description">
<paragraph id="8BA43689FFA53B16172FFE35FDC1FCA0" blockId="24.[232,679,827,884]" box="[282,629,859,884]" pageId="24" pageNumber="395">
(
<figureCitation id="13202A0CFFA53B161714FE35FE2AFCA7" box="[289,414,859,883]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="24" pageNumber="395">Figs. 7E, F</figureCitation>
,
<figureCitation id="13202A0CFFA53B16179EFE32FE7CFCA0" box="[427,456,860,884]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">13</figureCitation>
,
<figureCitation id="13202A0CFFA53B1617E0FE35FDA8FCA7" box="[469,540,859,883]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">26AI</figureCitation>
,
<figureCitation id="13202A0CFFA53B16141DFE32FDDAFCA0" box="[552,622,860,884]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27AI</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B1616BDFEF2FE44FBA0" pageId="24" pageNumber="395" type="materials_examined">
<paragraph id="8BA43689FFA53B1616BDFEF2FE44FBA0" blockId="24.[136,775,923,1140]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B1616BDFEF2FE8AFC67" bold="true" box="[136,318,923,948]" pageId="24" pageNumber="395">Type material.</emphasis>
<materialsCitation id="3B733CD4FFA53B16177FFEF5FDCBFBC0" collectingDate="2006-08-04" collectionCode="NMTU, AMNH" country="Nepal" county="Mt. Pulchoki" elevation="1650" latitude="27.579168" location="Kathmandu Valley" longLatPrecision="20" longitude="85.37889" municipality="Godawari" pageId="24" pageNumber="395" specimenCode="PBI 00378782" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A0882BFFA53B16177FFEF5FE0CFC67" box="[330,440,923,947]" pageId="24" pageNumber="395" type="holotype">Holotype</typeStatus>
: male,
<emphasis id="B96FEA9BFFA53B161425FEF5FDCCFC67" bold="true" box="[528,632,923,947]" pageId="24" pageNumber="395">
<collectingCountry id="F30C7619FFA53B161425FEF5FDC7FC67" box="[528,627,923,947]" name="Nepal" pageId="24" pageNumber="395">NEPAL</collectingCountry>
:
</emphasis>
<location id="8EC46052FFA53B1614B1FEF5FF67FC07" LSID="urn:lsid:plazi:treatment:03B2879FFFA53B1616DDFE55FABFFC61:8EC46052FFA53B1614B1FEF5FF67FC07" country="Nepal" county="Mt. Pulchoki" latitude="27.579168" longLatPrecision="20" longitude="85.37889" municipality="Godawari" name="Kathmandu Valley" pageId="24" pageNumber="395">Kathmandu Valley</location>
:
<location id="8EC46052FFA53B1616EBFED5FE83FC07" LSID="urn:lsid:plazi:treatment:03B2879FFFA53B1616DDFE55FABFFC61:8EC46052FFA53B1616EBFED5FE83FC07" box="[222,311,955,979]" country="Nepal" county="Mt. Pulchoki" latitude="27.579168" longLatPrecision="20" longitude="85.37889" municipality="Godawari" name="Lalitpur" pageId="24" pageNumber="395">Lalitpur</location>
,
<collectingMunicipality id="6BC0ACF3FFA53B161777FED5FE1AFC07" box="[322,430,955,979]" pageId="24" pageNumber="395">Godawari</collectingMunicipality>
,
<collectingCounty id="62C54E05FFA53B16178CFED2FDF2FC07" box="[441,582,955,979]" pageId="24" pageNumber="395">Mt. Pulchoki</collectingCounty>
(
<quantity id="4CE39B6CFFA53B161461FED5FD4EFC00" box="[596,762,955,980]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="24" pageNumber="395" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">
<elevation id="0036D1BAFFA53B161461FED5FD4EFC00" box="[596,762,955,980]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="24" pageNumber="395" unit="m" value="1650.0" valueMax="1800.0" valueMin="1500.0">1,5001,800 m</elevation>
</quantity>
),
<geoCoordinate id="EE2F504EFFA53B1616BDFEB4FEBEFC27" box="[136,266,986,1011]" degrees="27" direction="north" minutes="34" orientation="latitude" pageId="24" pageNumber="395" precision="15" seconds="45" value="27.579168">27°3445″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFA53B161723FEB4FE20FC27" box="[278,404,986,1011]" degrees="85" direction="east" minutes="22" orientation="longitude" pageId="24" pageNumber="395" precision="15" seconds="44" value="85.37889">85°2244″E</geoCoordinate>
,
<collectedFrom id="38077574FFA53B161795FEB2FD59FC20" box="[416,749,987,1012]" pageId="24" pageNumber="395">
on
<taxonomicName id="4C1B4D0AFFA53B1617F6FEB2FD59FC20" authority=", T. Yasunaga" authorityName="T. Yasunaga" box="[451,749,987,1012]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="395" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFFA53B1617F6FEB2FDE0FC27" box="[451,596,987,1011]" italics="true" pageId="24" pageNumber="395">Urtica dioica</emphasis>
, T. Yasunaga
</taxonomicName>
</collectedFrom>
,
<date id="FFA51049FFA53B1614CCFEB2FEBAFBC7" pageId="24" pageNumber="395" value="2006-08-04">
<collectingDate id="EFE1E9A1FFA53B1614CCFEB2FEBAFBC7" pageId="24" pageNumber="395" value="2006-08-04">4 August 2006</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA53B16172CFE92FEDEFBC0" box="[281,362,1020,1044]" pageId="24" pageNumber="395">NMTU</collectionCode>
) (
<collectionCode id="ED0AAE4CFFA53B161748FE92FE7BFBC0" box="[381,463,1020,1044]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="395">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFA53B1617E8FE92FDCCFBC0" box="[477,632,1020,1044]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="395">PBI 00378782</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFA53B1614B2FE92FF7CFB80" collectingDate="2006-08-04" collectionCode="AMNH, TYCN, ZRC" country="Nepal" county="Mt. Pulchoki" elevation="1650" latitude="27.579168" location="Kathmandu Valley" longLatPrecision="20" longitude="85.37889" municipality="Godawari" pageId="24" pageNumber="395" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54A0882BFFA53B1614B2FE92FD45FBC0" box="[647,753,1020,1044]" pageId="24" pageNumber="395" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D1DFD00FFA53B1614CCFE92FF7DFBE7" count="2" pageId="24" pageNumber="395" type="male">2 males</specimenCount>
,
<specimenCount id="9D1DFD00FFA53B1616E7F972FE8DFBE7" box="[210,313,1051,1076]" count="2" pageId="24" pageNumber="395" type="female">2 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFA53B16146CF972FD1BFBE0" box="[601,687,1052,1076]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="24" pageNumber="395">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFFA53B16148DF972FCB7FBE0" box="[696,771,1052,1076]" pageId="24" pageNumber="395">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFFA53B1616BDF952FF74FB80" box="[136,192,1084,1108]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="24" pageNumber="395">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B733CD4FFA53B1616E1F952FE59FBA0" collectingDate="2006-08-02" collectionCode="TYCN" country="Nepal" county="Mt. Pulchoki" elevation="1650" latitude="27.579168" location="Kathmandu Valley" longLatPrecision="20" longitude="85.37889" municipality="Godawari" pageId="24" pageNumber="395" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1DFD00FFA53B1616E1F952FEAAFB87" box="[212,286,1083,1108]" count="1" pageId="24" pageNumber="395" type="male">1 male</specimenCount>
,
<specimenCount id="9D1DFD00FFA53B16171FF952FE21FB87" box="[298,405,1083,1108]" count="2" pageId="24" pageNumber="395" type="female">2 females</specimenCount>
, same data as for holotype, except for date
<date id="FFA51049FFA53B1616D0F932FE32FBA7" box="[229,390,1115,1140]" pageId="24" pageNumber="395" value="2006-08-02">
<collectingDate id="EFE1E9A1FFA53B1616D0F932FE32FBA7" box="[229,390,1115,1140]" pageId="24" pageNumber="395" value="2006-08-02">2 August 2006</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFA53B1617A3F932FE51FBA0" box="[406,485,1116,1140]" pageId="24" pageNumber="395">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B1616BDF9F5FD7CFAE7" pageId="24" pageNumber="395" type="diagnosis">
<paragraph id="8BA43689FFA53B1616BDF9F5FD7CFAE7" blockId="24.[136,775,1179,1331]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B1616BDF9F5FF49FB67" bold="true" box="[136,253,1179,1203]" pageId="24" pageNumber="395">Diagnosis.</emphasis>
This new species is recognised by the following characters: Moderate size; yellowish brown, oily shiny, dorsum, with darkened clavus and posterior corium (
<figureCitation id="13202A0CFFA53B1614EEF9B5FF71FAC0" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="24" pageNumber="395">Fig. 7E, F</figureCitation>
); and bright greenish or yellowish white scutellum, in addition to the characters provided in the above key.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B1616BDF835FB37FD41" pageId="24" pageNumber="395" type="description">
<paragraph id="8BA43689FFA53B1616BDF835FBB1FD81" blockId="24.[136,776,1371,2099]" lastBlockId="24.[835,1475,189,597]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B1616BDF835FEA7FAA7" bold="true" box="[136,275,1371,1395]" pageId="24" pageNumber="395">Description.</emphasis>
Body oval, slightly elongate; basic colouration pale brown with widely darkened hemelytra (
<figureCitation id="13202A0CFFA53B1614B6F815FD4DFA40" box="[643,761,1403,1428]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="24" pageNumber="395">Fig. 7E, F</figureCitation>
); dorsum polished, impunctate (
<figureCitation id="13202A0CFFA53B1617FBF8F5FD85FA67" box="[462,561,1435,1459]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26C</figureCitation>
). Head pale whitish brown, smooth, with fuscous apical half (including jugum, lorum, buccula, and clypeus); vertex with faint, weak basal transverse carina (
<figureCitation id="13202A0CFFA53B16176FF895FE70F9C7" box="[346,452,1531,1555]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26B</figureCitation>
); frons (in fresh specimens) tinged with red. Antenna dark brown; antennomere I partly and irregularly pale at middle; antennomere II shorter than basal width of pronotum; extreme base of antennomere III creamy yellow. Labium pale reddish brown, reaching anterior margin of metacoxa; apical half of segment IV darkened. Pronotum pale brown, more or less darkened posteriorly, almost glabrous; calli (in fresh specimens) with a pair of dark small spots, partly suffused with red; pleura uniformly pale reddish brown; metathoracic scent efferent system as in
<figureCitation id="13202A0CFFA53B1616BDFA55FF59F887" box="[136,237,1851,1875]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26G</figureCitation>
; mesoscutum and anterior half of scutellum with sparsely distributed, short setae (
<figureCitation id="13202A0CFFA53B1617CBFA35FDD3F8A7" box="[510,615,1883,1907]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26D</figureCitation>
); mesoscutum dark reddish brown; scutellum greenish or yellowish white, with fuscous extreme apex. Hemelytron dark brown, oily shiny, smooth, with uniformly distributed, pale, short, simple setae (
<figureCitation id="13202A0CFFA53B1616FAFAB5FE80F827" box="[207,308,2011,2035]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26C</figureCitation>
); basal ⅔ of clavus pale brown; subapical part of cuneus narrowly pale; membrane pale smoky brown, semi-transparent, with dark brown veins. All coxae and legs pale brown; each femur with two red, subapical rings; metafemur with broader reddish annulation at middle; apical part of each tarsomere III darkened; pretarsal structure as in
<figureCitation id="13202A0CFFA53B161576FC73FC28FEE1" box="[835,924,285,309]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="24" pageNumber="395">Fig. 26I</figureCitation>
. Abdomen pale reddish brown; apical half of male genital segment (pygophore) fuscous. Male genitalia (
<figureCitation id="13202A0CFFA53B1613B9FC53FCEFFEA1" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13</figureCitation>
AG, 27AE): Hypophysis of left paramere elongate (
<figureCitation id="13202A0CFFA53B1613A2FC33FCC0FE41" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Fig. 13A</figureCitation>
), with barb-like apex (
<figureCitation id="13202A0CFFA53B16125CFC13FB60FE41" box="[1129,1236,381,405]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13B</figureCitation>
,
<figureCitation id="13202A0CFFA53B1612D4FC13FAA4FE41" box="[1249,1296,381,405]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27B</figureCitation>
); right paramere bulbous, ovoid, with short, pointed hypophysis (
<figureCitation id="13202A0CFFA53B161365FCF3FCE2FE01" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13D, E</figureCitation>
,
<figureCitation id="13202A0CFFA53B161550FCD3FC23FE01" box="[869,919,445,469]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27C</figureCitation>
); vesica with three notched lobal-sclerites (
<figureCitation id="13202A0CFFA53B1613BEFCD3FCDBFE21" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13F</figureCitation>
,
<figureCitation id="13202A0CFFA53B161549FCB3FC1AFE21" box="[892,942,477,501]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27D</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFFA53B1612B3FCB3FAB9FE21" box="[1158,1293,477,501]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13H, I</figureCitation>
,
<figureCitation id="13202A0CFFA53B16132EFCB3FAEBFE21" box="[1307,1375,477,501]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27FI</figureCitation>
): Genital chamber circular, with elongate oval sclerotised rings (
<figureCitation id="13202A0CFFA53B1613B9FC93FCDEFDE1" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13I</figureCitation>
,
<figureCitation id="13202A0CFFA53B16154FFF73FC1CFDE1" box="[890,936,541,565]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27F</figureCitation>
); posterior wall with widened interramal lobes (
<figureCitation id="13202A0CFFA53B16157EFF53FC0EFD81" box="[843,954,573,597]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="24" pageNumber="395">Figs. 13H</figureCitation>
,
<figureCitation id="13202A0CFFA53B1615F2FF53FC4CFD81" box="[967,1016,573,597]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="24" pageNumber="395">27H</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFA53B161576FF13FB37FD41" blockId="24.[835,1155,637,661]" box="[835,1155,637,661]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576FF13FC46FD41" bold="true" box="[835,1010,637,661]" pageId="24" pageNumber="395">Measurements.</emphasis>
See
<tableCitation id="C6990332FFA53B16121CFF13FBC8FD41" box="[1065,1148,637,661]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="24" pageNumber="395">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161576FFD3FBB4FCC1" pageId="24" pageNumber="395" type="etymology">
<paragraph id="8BA43689FFA53B161576FFD3FBB4FCC1" blockId="24.[835,1474,701,789]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576FFD3FC71FD01" bold="true" box="[835,965,701,725]" pageId="24" pageNumber="395">Etymology.</emphasis>
Named to honour Dr. Ram Keshari Duwal, a Nepalese heteropterist (now working at CNC); a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161576FE53FAC5FC81" box="[835,1393,829,853]" pageId="24" pageNumber="395" type="distribution">
<paragraph id="8BA43689FFA53B161576FE53FAC5FC81" blockId="24.[835,1393,829,853]" box="[835,1393,829,853]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576FE53FC62FC81" bold="true" box="[835,982,829,853]" pageId="24" pageNumber="395">Distribution.</emphasis>
<collectingCountry id="F30C7619FFA53B1615EBFE53FBAAFC81" box="[990,1054,829,853]" name="Nepal" pageId="24" pageNumber="395">Nepal</collectingCountry>
(Lalitpur, Kathmandu Valley).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA53B161576FE13FABFFC61" pageId="24" pageNumber="395" type="biology_ecology">
<paragraph id="8BA43689FFA53B161576FE13FABFFC61" blockId="24.[835,1474,893,949]" pageId="24" pageNumber="395">
<emphasis id="B96FEA9BFFA53B161576FE13FC14FC41" bold="true" box="[835,928,893,917]" pageId="24" pageNumber="395">Biology.</emphasis>
All available specimens of this new species were found from
<taxonomicName id="4C1B4D0AFFA53B1615F2FEF0FBECFC61" box="[967,1112,925,949]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="395" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFFA53B1615F2FEF0FBECFC61" box="[967,1112,925,949]" italics="true" pageId="24" pageNumber="395">Urtica dioica</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C1B4D0AFFA53B1612B2FEF3FAB4FC61" box="[1159,1280,925,949]" class="Magnoliopsida" family="Urticaceae" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="395" phylum="Tracheophyta" rank="family">Urticaceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,219 @@
<document id="4DD02816A1610429F8C319112C3E5989" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA83B161518FA13FCB1FCC7" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris Yasunaga, 2024, new genus" docType="key" docVersion="2" lastPageNumber="395" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="392" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5695DC6DB89068733870523AC429FFDA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4232F27B73A919BBD24D8BD53C22095E">
<mods:title id="86C1C5BD2B87FCF35137B52150571DEE">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="C59C8722D4DD3E7FEF3A22E15262ED40" type="personal">
<mods:role id="3F56988B9EB0330B8926D255A697A5E4">
<mods:roleTerm id="E09EBAA6A500DF842F9383685BFB8A5A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7DF2EB63A5D2E53C0202EA6A548FE084">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="CB89997F8FDD5FA22C092401B0F00BD9">text</mods:typeOfResource>
<mods:relatedItem id="3DE71FF0938ED9002FAD882DD7877F11" type="host">
<mods:titleInfo id="46FE522E1823DC30235AF3857CC3563A">
<mods:title id="52C6409C980ED1432D23A00F8711D66F">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7DB84293F0E2C37A502CB1508692C496">
<mods:date id="3E9DC1D2248367199473A97CBDC980DE">2024</mods:date>
<mods:detail id="24843959E25135F6AC5A17E565B618F8" type="pubDate">
<mods:number id="58EF57E2707FC1BA7EA4F30DED220F28">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="CD713C93B6EF6B4A66BE3AF8700468B6" type="volume">
<mods:number id="DB5B528E3E1986C59098B2A68952B25F">72</mods:number>
</mods:detail>
<mods:extent id="A3306679EDF6940B9061B9D9EE9C0DC7" unit="page">
<mods:start id="A8F72F3539EB9BC239D99277FA470716">371</mods:start>
<mods:end id="01ACA9443022F142C629A748664AE62B">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E56252A893E391EB8A5FBBD9FEB76B2A">journal article</mods:classification>
<mods:identifier id="A0D58B35C3CA1ABCDBFC47AB00836A45" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="B0871B9EADEBD53C444AA11D2E4386FE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="469C2BBC7D50AD11CE4C06A161EB634A" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="9C17BED0FD5BF7BF0E8F072F87A9F34E" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA83B161518FA13FCB1FCC7" ID-DOI="http://doi.org/10.5281/zenodo.14683008" ID-Zenodo-Dep="14683008" LSID="urn:lsid:plazi:treatment:03B2879FFFA83B161518FA13FCB1FCC7" httpUri="http://treatment.plazi.org/id/03B2879FFFA83B161518FA13FCB1FCC7" lastPageId="24" lastPageNumber="395" pageId="21" pageNumber="392">
<subSubSection id="C3016502FFA83B1B1518FA13FAF0F841" box="[813,1348,1917,1941]" pageId="21" pageNumber="392" type="nomenclature">
<paragraph id="8BA43689FFA83B1B1518FA13FAF0F841" blockId="21.[813,1348,1917,1941]" box="[813,1348,1917,1941]" pageId="21" pageNumber="392">
<heading id="D0EC81E5FFA83B1B1518FA13FAF0F841" bold="true" box="[813,1348,1917,1941]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="392" reason="2">
<emphasis id="B96FEA9BFFA83B1B1518FA13FAF0F841" bold="true" box="[813,1348,1917,1941]" pageId="21" pageNumber="392">
Key to species of
<taxonomicName id="4C1B4D0AFFA83B1B15CDFA13FB09F841" authorityName="Yasunaga" authorityYear="2024" box="[1016,1213,1917,1941]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B15CDFA13FB09F841" bold="true" box="[1016,1213,1917,1941]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B12F8FA13FAF0F841" box="[1229,1348,1917,1941]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B16151BFAD2FCB1FCC7" lastPageId="24" lastPageNumber="395" pageId="21" pageNumber="392" type="key">
<keyStep id="30EF2E2CFFA83B1B151BFAD2FA1EF7F3" pageId="21" pageNumber="392">
<paragraph id="8BA43689FFA83B1B151BFAD2FA1EF83A" blockId="21.[813,1452,1980,2088]" pageId="21" pageNumber="392">
<keyLead id="30EA95BCFFA83B1B151BFAD2FA1EF83A" pageId="21" pageNumber="392">
1. Scutellum pale, with two dark, distinct spots anteriorly (
<figureCitation id="13202A0CFFA83B1B134EFAD2FC3EF83A" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="21" pageNumber="392">Figs. 7E, F</figureCitation>
,
<figureCitation id="13202A0CFFA83B1B15A3FAB7FC61F83A" box="[918,981,2009,2030]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="21" pageNumber="392">11GI</figureCitation>
,
<figureCitation id="13202A0CFFA83B1B15D4FAB7FBB8F83A" box="[993,1036,2009,2030]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="21" pageNumber="392">12A</figureCitation>
) .................................................................2
</keyLead>
</paragraph>
<paragraph id="8BA43689FFA83B1B1518FA98FA1EF7F3" blockId="21.[813,1452,1980,2088]" pageId="21" pageNumber="392">
<keyLead id="30EA95BCFFA83B1B1518FA98FA1EF7F3" pageId="21" pageNumber="392">
Scutellum uniformly creamy yellow, pale brown or brown, without dark spot or macula (
<figureCitation id="13202A0CFFA83B1B1243F57CFB62F7F3" box="[1142,1238,2066,2088]" captionStart="Fig" captionStartId="19.[113,148,1757,1779]" captionTargetBox="[135,1437,204,1724]" captionTargetId="figure-11@19.[113,1451,189,1741]" captionTargetPageId="19" captionText="Fig. 11. Habitus images of Himalisundacoris species from Sumatra and Sulawesi, Indonesia. A, B, H. palolo, new species, holotype male; C, D, ditto, female; E, F, H. takaii, new species, holotype male; G, H, H. schwartzi, new species, holotype male; I, J, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="21" pageNumber="392">Figs. 11E</figureCitation>
,
<figureCitation id="13202A0CFFA83B1B12D6F57CFA9CF7F3" box="[1251,1320,2066,2087]" captionStart="Fig" captionStartId="20.[135,170,1961,1983]" captionTargetBox="[150,1461,271,1930]" captionTargetId="figure-7@20.[135,1473,263,1945]" captionTargetPageId="20" captionText="Fig. 12. Habitus images of Himalisundacoris species from Bali, Indonesia (AH) and Paralygocoris species (HL). A, B, H. omiae, new species, holotype male; C, D, ditto, female; E, F, H. theoi, new species, male; G, H, ditto, female; H, I. P. balicus, new species, male; J, ditto, female; K, L, P. meridionalis, female; M, N, P. vittulatus, male; O, ditto, female." figureDoi="http://doi.org/10.5281/zenodo.14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="21" pageNumber="392">12EG</figureCitation>
)..................3
</keyLead>
</paragraph>
</keyStep>
<caption id="DF646601FFAB3B1816B2FABAFAA1F7D2" ID-DOI="http://doi.org/10.5281/zenodo.14682980" ID-Zenodo-Dep="14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="22" pageNumber="393" startId="22.[135,170,2004,2026]" targetBox="[135,1473,258,1989]" targetPageId="22" targetType="figure">
<paragraph id="8BA43689FFAB3B1816B2FABAFAA1F7D2" blockId="22.[135,1474,2004,2055]" pageId="22" pageNumber="393">
Fig. 13. Male (AG) and female (HI) genitalia of
<taxonomicName id="4C1B4D0AFFAB3B181449FABAFC34F83D" authorityName="Yasunaga" authorityYear="2024" box="[636,896,2004,2025]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="22" pageNumber="393" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFFAB3B181449FABAFC34F83D" box="[636,896,2004,2025]" italics="true" pageId="22" pageNumber="393">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAB3B1815BBFABBFBB1F83E" box="[910,1029,2004,2026]" pageId="22" pageNumber="393" rank="species">new species</taxonomicNameLabel>
. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm.
</paragraph>
</caption>
<caption id="DF646601FFAA3B191644FA1EFD1BF86B" ID-DOI="http://doi.org/10.5281/zenodo.14682982" ID-Zenodo-Dep="14682982" httpUri="https://zenodo.org/record/14682982/files/figure.png" pageId="23" pageNumber="394" startId="23.[113,148,1904,1926]" targetBox="[113,1451,316,1889]" targetPageId="23" targetType="figure">
<paragraph id="8BA43689FFAA3B191644FA1EFD1BF86B" blockId="23.[113,1452,1904,1984]" pageId="23" pageNumber="394">
Fig. 14. Male (AC, FH) and female (D, E, I, J) genitalia of
<taxonomicName id="4C1B4D0AFFAA3B1914D2FA1EFC63F852" authorityName="Yasunaga" authorityYear="2024" box="[743,983,1904,1926]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="394" phylum="Arthropoda" rank="species" species="omiae" status="sp. nov.">
<emphasis id="B96FEA9BFFAA3B1914D2FA1EFC63F852" box="[743,983,1904,1926]" italics="true" pageId="23" pageNumber="394">Himalisundacoris omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAA3B1915D3FA1FFBE8F852" box="[998,1116,1904,1926]" pageId="23" pageNumber="394" rank="species">new species</taxonomicNameLabel>
(AE) and
<taxonomicName id="4C1B4D0AFFAA3B1912E3FA1FFA92F851" authorityName="Yasunaga" authorityYear="2024" box="[1238,1318,1904,1926]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="394" phylum="Arthropoda" rank="species" species="theoi" status="new species">
<emphasis id="B96FEA9BFFAA3B1912E3FA1FFA92F851" box="[1238,1318,1904,1926]" italics="true" pageId="23" pageNumber="394">H. theoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAA3B191300FA1FFA1FF852" box="[1333,1451,1904,1926]" pageId="23" pageNumber="394" rank="species">new species</taxonomicNameLabel>
(FJ). A, F, left paramere; B, G, right paramere; C, H, vesica; G, ditto, secondary gonopore and adjacent structures; D, J, posterior wall; E, genital chamber; I, sclerotised ring. Scale bars 0.1 mm.
</paragraph>
</caption>
<keyStep id="30EF2E2CFFA53B1616BDFDD3FCB1FE92" pageId="24" pageNumber="395">
<paragraph id="8BA43689FFA53B1616BDFDD3FCB1FF24" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<materialsCitation id="3B733CD4FFA53B1616BDFDD3FCB1FF24" collectionCode="II" country="Indonesia" location="Sulawesi" pageId="24" pageNumber="395" specimenCount="1" stateProvince="Sulawesi Utara">
<keyLead id="30EA95BCFFA53B1616BDFDD3FCB1FF24" pageId="24" pageNumber="395">
2. Antennomere
<collectionCode id="ED0AAE4CFFA53B161706FDD0FEF7FF07" box="[307,323,190,211]" pageId="24" pageNumber="395">II</collectionCode>
uniformly dark brown; thoracic pleurites widely darkened;
<collectingCountry id="F30C7619FFA53B161721FDB4FEC5FF24" box="[276,369,218,240]" name="Indonesia" pageId="24" pageNumber="395">Indonesia</collectingCountry>
(S.
<collectingRegion id="49DFF86BFFA53B1617A9FDB4FE40FF24" box="[412,500,218,240]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="395">Sulawesi</collectingRegion>
)........
<taxonomicName id="4C1B4D0AFFA53B161418FDB5FD37FF3B" authorityName="Yasunaga" authorityYear="2024" box="[557,643,218,240]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="takaii" status="new species">
<emphasis id="B96FEA9BFFA53B161418FDB5FD37FF3B" box="[557,643,218,240]" italics="true" pageId="24" pageNumber="395">H. takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FDB5FCB1FF24" box="[656,773,218,240]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</keyLead>
</materialsCitation>
</paragraph>
<paragraph id="8BA43689FFA53B1616BDFD99FCB1FE92" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFD99FCB1FE92" pageId="24" pageNumber="395">
<materialsCitation id="3B733CD4FFA53B16169BFD99FCB1FE92" collectionCode="II" country="Indonesia" location="Bali" pageId="24" pageNumber="395" specimenCount="1" stateProvince="Bali">
Sub-basal part of antennomere
<collectionCode id="ED0AAE4CFFA53B1617FAFD99FE6BFED8" box="[463,479,247,268]" pageId="24" pageNumber="395">II</collectionCode>
with a pale annulation; thoracic pleurites pale brown;
<collectingCountry id="F30C7619FFA53B1617B1FC7AFE55FEFE" box="[388,481,276,298]" name="Indonesia" pageId="24" pageNumber="395">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFFA53B1617C5FC7AFDAEFEFE" box="[496,538,276,298]" country="Indonesia" name="Bali" pageId="24" pageNumber="395">Bali</collectingRegion>
) ..................................... ................................................................
<emphasis id="B96FEA9BFFA53B161406FC5FFD3DFE91" box="[563,649,304,326]" italics="true" pageId="24" pageNumber="395">
<taxonomicName id="4C1B4D0AFFA53B161406FC5FFD31FE91" authorityName="Yasunaga" authorityYear="2024" box="[563,645,304,326]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="theoi" status="new species">H. theoi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FC5FFCB1FE92" box="[656,773,304,326]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</materialsCitation>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFA53B1616BDFC23FCB1FE02" pageId="24" pageNumber="395">
<paragraph id="8BA43689FFA53B1616BDFC23FCB1FE49" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFC23FCB1FE49" pageId="24" pageNumber="395">
3. Vertex with faint, weak basal transverse carina (
<figureCitation id="13202A0CFFA53B1614ABFC23FD4FFEB6" box="[670,763,333,355]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="24" pageNumber="395">Fig. 22B</figureCitation>
); pronotum bicolourous, darkened posteriorly;
<collectingCountry id="F30C7619FFA53B1614B4FC04FD08FE54" box="[641,700,362,384]" name="Nepal" pageId="24" pageNumber="395">Nepal</collectingCountry>
(warm temperate zone) ...............................
<taxonomicName id="4C1B4D0AFFA53B161427FCE6FD36FE48" authorityName="Yasunaga" authorityYear="2024" box="[530,642,391,413]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="duwalae" status="new species">
<emphasis id="B96FEA9BFFA53B161427FCE6FD36FE48" box="[530,642,391,413]" italics="true" pageId="24" pageNumber="395">H. duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FCE9FCB1FE49" box="[656,773,391,413]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFA53B1616BDFCCAFCB1FE02" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFCCAFCB1FE02" pageId="24" pageNumber="395">
Basal carina of vertex obliterated (
<figureCitation id="13202A0CFFA53B1617CBFCCAFDD5FE6D" box="[510,609,420,442]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="24" pageNumber="395">Figs. 24D</figureCitation>
,
<figureCitation id="13202A0CFFA53B161459FCCAFD23FE6E" box="[620,663,420,442]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="24" pageNumber="395">25B</figureCitation>
); pronotum unicolourously pale .................................................................4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFA53B1616BDFCB3FCB1FDB2" pageId="24" pageNumber="395">
<paragraph id="8BA43689FFA53B1616BDFCB3FCB1FDF9" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFCB3FCB1FDF9" pageId="24" pageNumber="395">
4. General colouration pale milky brown (partly tinged with olive); metafemur obviously longer than labium;
<collectingRegion id="49DFF86BFFA53B16147EFC94FDC7FDC4" box="[587,627,506,528]" country="Indonesia" name="Bali" pageId="24" pageNumber="395">Bali</collectingRegion>
........................ ..............................................................
<taxonomicName id="4C1B4D0AFFA53B16141CFF76FD37FDF9" authorityName="Yasunaga" authorityYear="2024" box="[553,643,536,557]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="omiae" status="new species">
<emphasis id="B96FEA9BFFA53B16141CFF76FD37FDF9" box="[553,643,536,557]" italics="true" pageId="24" pageNumber="395">H. omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FF79FCB1FDF9" box="[656,773,535,557]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFA53B1616BDFF5AFCB1FDB2" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFF5AFCB1FDB2" pageId="24" pageNumber="395"> General colouration pale orange-brown; metafemur as long as or shorter than labium.............................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFA53B1616BDFF03FCB1FCC7" pageId="24" pageNumber="395">
<paragraph id="8BA43689FFA53B1616BDFF03FCB1FD69" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFF03FCB1FD69" pageId="24" pageNumber="395">
5. Basal ⅓–½ of antennomere II pale, with darkened extreme base; abdomen uniformly shiny chocolate brown; C.
<collectingRegion id="49DFF86BFFA53B16144AFFE4FD62FD74" box="[639,726,650,672]" country="Indonesia" name="Sulawesi Utara" pageId="24" pageNumber="395">Sulawesi</collectingRegion>
....... ..............................................................
<emphasis id="B96FEA9BFFA53B161413FFC6FD3DFD68" box="[550,649,679,701]" italics="true" pageId="24" pageNumber="395">
<taxonomicName id="4C1B4D0AFFA53B161413FFC6FD31FD68" authorityName="Yasunaga" authorityYear="2024" box="[550,645,679,701]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="palolo" status="new species">H. palolo</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FFC9FCB1FD69" box="[656,773,679,701]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFA53B1616BDFFAAFCB1FCC7" blockId="24.[136,775,189,787]" pageId="24" pageNumber="395">
<keyLead id="30EA95BCFFA53B1616BDFFAAFCB1FCC7" pageId="24" pageNumber="395">
<materialsCitation id="3B733CD4FFA53B16169BFFAAFCB1FCC7" collectionCode="II" country="Indonesia" location="Sumatra" pageId="24" pageNumber="395" specimenCount="1" stateProvince="Sumatera Utara">
Basal part of antennomere
<collectionCode id="ED0AAE4CFFA53B161796FFAAFE07FD0D" box="[419,435,708,729]" pageId="24" pageNumber="395">II</collectionCode>
dark, with pale annulation; abdomen pale reddish brown;
<collectingCountry id="F30C7619FFA53B16174DFF8EFE61FD22" box="[376,469,736,758]" name="Indonesia" pageId="24" pageNumber="395">Indonesia</collectingCountry>
(N.
<collectingRegion id="49DFF86BFFA53B161431FF8FFDE2FD22" box="[516,598,737,758]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="395">Sumatra</collectingRegion>
)............................ ........................................................
<emphasis id="B96FEA9BFFA53B161432FF90FD3DFCC6" box="[519,649,765,787]" italics="true" pageId="24" pageNumber="395">
<taxonomicName id="4C1B4D0AFFA53B161432FF90FD30FCC6" authorityName="Yasunaga" authorityYear="2024" box="[519,644,765,787]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="395" phylum="Arthropoda" rank="species" species="schwartzi" status="new species">H. schwartzi</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25C57E0FFA53B1614A5FF90FCB1FCC7" box="[656,773,765,787]" pageId="24" pageNumber="395" rank="species">new species</taxonomicNameLabel>
</materialsCitation>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,200 @@
<document id="CF862B8E9F216373F209DCFC1BC10CD2" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFA83B1B173EFF53FB21F882" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Himalisundacoris Yasunaga, 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="392" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="392" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="3FCD22F4D15BD396C8D097A8E0204737" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="60F9781A19EB9A39A628E0723EADE33E">
<mods:title id="5B209CE8174810BFE17DBE62024F71E0">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="7D223CB4A5580AEE6FFD0DCCD5F38900" type="personal">
<mods:role id="74D9B67FB074D5C9EE8099EA4DAF9822">
<mods:roleTerm id="7D14357AC4B1661B011333D19FC9E806">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41E25E7DB6248037075C279259F655E1">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="0304DF82F7D882C12036EABDEC41FF18">text</mods:typeOfResource>
<mods:relatedItem id="EB800C6158AA268E9A346194D84FFEBE" type="host">
<mods:titleInfo id="ACAD1A4AE65EF5F839F1A2A3B1029240">
<mods:title id="2A3779C8A880762B3261B28ADBB776EB">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="6BEC454ABDF2FD47467D4C32318A1647">
<mods:date id="195CB8712943F8624F016BBF3470237C">2024</mods:date>
<mods:detail id="7A8E2AAE411DF4C24A24FCDF890CA023" type="pubDate">
<mods:number id="585F15E84B7817DB2E6EBF6ECB87DB47">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="1E5048334354D1EBF5192F71A2E0E17B" type="volume">
<mods:number id="4EC67DD52A2A7F3AB3B1FFE32B6D013E">72</mods:number>
</mods:detail>
<mods:extent id="B21847238F605C5C27C381F16E887B1F" unit="page">
<mods:start id="A7DE4BEB295CAA05CAEA2478A2A5314F">371</mods:start>
<mods:end id="A960B47D75191D32607C12015C452E92">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5B1FBE83524BC0D3F1A7D580AD8628CD">journal article</mods:classification>
<mods:identifier id="82D04A3C7B753E3557C5D8946F45B78E" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="D6447AD18A8919BCC83080139410D358" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DF165AC3F3EAD1D24FEAA60E72625690" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="1E1AD3C502EAF60AD0CC32B968D9A54B" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFA83B1B173EFF53FB21F882" ID-DOI="http://doi.org/10.5281/zenodo.14683002" ID-Zenodo-Dep="14683002" LSID="urn:lsid:plazi:treatment:03B2879FFFA83B1B173EFF53FB21F882" httpUri="http://treatment.plazi.org/id/03B2879FFFA83B1B173EFF53FB21F882" lastPageNumber="392" pageId="21" pageNumber="392">
<subSubSection id="C3016502FFA83B1B173EFF53FDE3FD81" box="[267,599,573,597]" pageId="21" pageNumber="392" type="nomenclature">
<paragraph id="8BA43689FFA83B1B173EFF53FDE3FD81" blockId="21.[267,599,573,597]" box="[267,599,573,597]" pageId="21" pageNumber="392">
<heading id="D0EC81E5FFA83B1B173EFF53FDE3FD81" bold="true" box="[267,599,573,597]" centered="true" fontSize="10" level="2" pageId="21" pageNumber="392" reason="2">
<emphasis id="B96FEA9BFFA83B1B173EFF53FDE3FD81" bold="true" box="[267,599,573,597]" pageId="21" pageNumber="392">
<taxonomicName id="4C1B4D0AFFA83B1B173EFF53FE64FD81" authorityName="Yasunaga" authorityYear="2024" box="[267,464,573,597]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B173EFF53FE64FD81" bold="true" box="[267,464,573,597]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B17EAFF53FDE3FD81" box="[479,599,573,597]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644FF13FD79FD41" box="[113,717,637,661]" pageId="21" pageNumber="392" type="type_taxon">
<paragraph id="8BA43689FFA83B1B1644FF13FD79FD41" blockId="21.[113,717,637,661]" box="[113,717,637,661]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FF13FEBDFD41" bold="true" box="[113,265,637,661]" pageId="21" pageNumber="392">
<typeStatus id="54A0882BFFA83B1B1644FF13FF1FFD41" box="[113,171,637,661]" pageId="21" pageNumber="392">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1B4D0AFFA83B1B1727FF13FD87FD41" authorityName="Yasunaga" authorityYear="2024" box="[274,563,637,661]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFFA83B1B1727FF13FD87FD41" box="[274,563,637,661]" italics="true" pageId="21" pageNumber="392">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B1476FF10FD7CFD41" box="[579,712,637,661]" pageId="21" pageNumber="392" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644FFD3FE77FB61" pageId="21" pageNumber="392" type="diagnosis">
<paragraph id="8BA43689FFA83B1B1644FFD3FE77FB61" blockId="21.[113,753,701,1205]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FFD3FF52FD01" bold="true" box="[113,230,701,725]" pageId="21" pageNumber="392">Diagnosis.</emphasis>
Based on the similar surface and male genitalic structures,
<taxonomicName id="4C1B4D0AFFA83B1B16DCFFB3FE1CFD21" authorityName="Yasunaga" authorityYear="2024" box="[233,424,733,757]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B16DCFFB3FE1CFD21" box="[233,424,733,757]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B1782FFB0FD99FD22" box="[439,557,734,758]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
, is assumed to be most closely related to
<taxonomicName id="4C1B4D0AFFA83B1B1748FF93FD58FCC1" authority="Zheng &amp; Lin, 2002" authorityName="Zheng &amp; Lin" authorityYear="2002" box="[381,748,765,789]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFA83B1B1748FF93FDB6FCC1" box="[381,514,765,789]" italics="true" pageId="21" pageNumber="392">Sabactiopus</emphasis>
<bibRefCitation id="EF8A4B78FFA83B1B1438FF93FD58FCC1" author="Zheng LY &amp; Lin CS" box="[525,748,765,789]" pageId="21" pageNumber="392" pagination="75 - 81" refId="ref24959" refString="Zheng LY &amp; Lin CS (2002) Sabactiopus gen. nov. with a redescription of Sabactus institutus Distant, 1910 (Hemiptera: Miridae: Mirinae). Formosan Entomologist, 22: 75 - 81." type="journal article" year="2002">Zheng &amp; Lin, 2002</bibRefCitation>
</taxonomicName>
, from which
<taxonomicName id="4C1B4D0AFFA83B1B16C2FE73FE02FCE1" authorityName="Yasunaga" authorityYear="2024" box="[247,438,797,821]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B16C2FE73FE02FCE1" box="[247,438,797,821]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B17F0FE70FD8FFCE2" box="[453,571,798,822]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
, is distinguished readily by the following unique characters: Body generally larger; vertex smooth, lacking basal transverse carina (that is faintly recognisable in some species); left paramere with flattened sensory lobe and distinctly hooked (or shovel-headed or spade-shaped) apex; right paramere very tiny, with bulbous sensory lobe; interramal lobe with densely distributed, scaly microstructures.
<taxonomicName id="4C1B4D0AFFA83B1B17C4FE93FD06FBC1" authorityName="Yasunaga" authorityYear="2024" box="[497,690,1021,1045]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B17C4FE93FD06FBC1" box="[497,690,1021,1045]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B14F6FE90FF00FBE2" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
, is also at first sight reminiscent of the European genus
<taxonomicName id="4C1B4D0AFFA83B1B1683F950FE2FFB81" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" box="[182,411,1085,1109]" class="Insecta" family="Miridae" genus="Liocoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFA83B1B1683F950FEBBFB81" box="[182,271,1086,1109]" italics="true" pageId="21" pageNumber="392">Liocoris</emphasis>
<bibRefCitation id="EF8A4B78FFA83B1B1720F953FE2FFB81" author="Fieber FX" box="[277,411,1085,1109]" pageId="21" pageNumber="392" pagination="289 - 327" refId="ref23813" refString="Fieber FX (1858) Criterien zur generischen Theilung der Phytocoriden (Capsini auct.). Wiener entomologische Monatschrift, 2: 289 - 327, 329 - 347, 388, 1 pl." type="journal article" year="1858">Fieber, 1858</bibRefCitation>
</taxonomicName>
(e.g., similar colour pattern and body shape, and shiny dorsum); however, the latter has the punctate dorsum and completely different genitalic structures (cf.
<bibRefCitation id="EF8A4B78FFA83B1B16ABF9F3FE03FB61" author="Schwartz MD &amp; Foottit RG" box="[158,439,1181,1205]" pageId="21" pageNumber="392" pagination="1 - 428" refId="ref24370" refString="Schwartz MD &amp; Foottit RG (1998) Revision of the Nearctic species of the genus Lygus Hahn, with review of the Palearctic species (Heteroptera: Miridae). Memoirs on Entomology International 10. Associated Publishers, Gainesville, i - vii + 1 - 428 pp." type="book chapter" year="1998">Schwartz &amp; Foottit, 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644F9B3FA91FD41" pageId="21" pageNumber="392" type="description">
<paragraph id="8BA43689FFA83B1B1644F9B3FA91FD41" blockId="21.[113,753,1245,2101]" lastBlockId="21.[813,1452,189,661]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644F9B3FEBDFB21" bold="true" box="[113,265,1245,1269]" pageId="21" pageNumber="392">Description.</emphasis>
Body small to moderate in size, ovoid, relatively stout, not sexually dimorphic in general shape; basic colouration pale brown to fuscous; dorsal surface polished, oily shiny, impunctate, with sparsely distributed, pale, simple, semi-erect setae but nearly glabrous in some species.
<emphasis id="B96FEA9BFFA83B1B16FFF813FEA4FA41" bold="true" box="[202,272,1405,1429]" pageId="21" pageNumber="392">Head:</emphasis>
Smooth, shining, somewhat oblique in lateral view, with sparsely distributed, short setae; eyes contiguous to anterior margin of pronotal collar; vertex shallowly sulcate or concaved mesally, without basal transverse carina; clypeus infuscate.
<emphasis id="B96FEA9BFFA83B1B16EAF893FEFDF9C1" bold="true" box="[223,329,1533,1557]" pageId="21" pageNumber="392">Antenna:</emphasis>
Slightly longer than or as long as body, almost linear; antennomeres I and II similar in colour pattern to those of
<taxonomicName id="4C1B4D0AFFA83B1B16D2FB53FEDFF981" authorityName="Yasunaga" authorityYear="2024" box="[231,363,1597,1621]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B16D2FB53FEDFF981" box="[231,363,1597,1621]" italics="true" pageId="21" pageNumber="392">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B174DFB50FE5FF982" box="[376,491,1598,1622]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
, darkened apically (or at apex); antennomere II more or less darkened at base, relatively short, shorter than antennomeres III+IV antennomeres III and IV filiform; antennomere III as long as or longer than width of head across eyes.
<emphasis id="B96FEA9BFFA83B1B17E3FBD3FD8EF901" bold="true" box="[470,570,1725,1749]" pageId="21" pageNumber="392">Labium:</emphasis>
Relatively long, reaching or slightly exceeding apex of metacoxa.
<emphasis id="B96FEA9BFFA83B1B14A6FBB3FD44F921" bold="true" box="[659,752,1757,1781]" pageId="21" pageNumber="392">Thorax:</emphasis>
Pronotum shiny, highly polished, almost glabrous or with sparsely distributed, short, semi-erect setae, impunctate; calli not clearly demarcated; collar shagreened, narrow, about as thick as antennomere III; thoracic pleura shining; propleuron not margined nor carinate; metathoracic scent efferent system, with small peritreme (
<figureCitation id="13202A0CFFA83B1B1430FAF3FDC3F861" box="[517,631,1949,1973]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="21" pageNumber="392">Fig. 2B, K</figureCitation>
); scutellum polished, impunctate, weakly arched.
<emphasis id="B96FEA9BFFA83B1B1415FAD3FD01F801" bold="true" box="[544,693,1981,2005]" pageId="21" pageNumber="392">Hemelytron:</emphasis>
Oily shiny, variable in colour, impunctate, declivous at cuneal fracture, with sparsely (sometimes rather densely) distributed, pale, simple, semi-erect setae.
<emphasis id="B96FEA9BFFA83B1B178BF573FE48F7E1" bold="true" box="[446,508,2077,2101]" pageId="21" pageNumber="392">Legs:</emphasis>
Relatively long; tibial spines pale brown or reddish brown; meta-tarsomere II about as long as III; pretarsal structures as in
<figureCitation id="13202A0CFFA83B1B131CFDB3FA13FF21" box="[1321,1447,221,245]" captionStart="Fig" captionStartId="39.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@39.[113,1451,251,1929]" captionTargetPageId="39" captionText="Fig. 26. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, Left lateral habitus; B, head and anterior pronotum, dorsal view; C, dorsal habitus; D, scutellum, clavus and corium, dorsal view; E, apical forewing, dorsal view; F, thoracic pleura, left lateral view; G, M, metathoracic scent efferent system; H, N, metatarsus; I, O, pretarsal structure of metaleg; J, head and thorax, left lateral view; K, head and anterior pronotum, frontal view; L, scutellum and clavus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683014" httpUri="https://zenodo.org/record/14683014/files/figure.png" pageId="21" pageNumber="392">Fig. 26I, O</figureCitation>
; parempodium relatively short.
<emphasis id="B96FEA9BFFA83B1B12B7FD93FA86FEC1" bold="true" box="[1154,1330,253,277]" pageId="21" pageNumber="392">Male genitalia:</emphasis>
Parameres highly asymmetrical; left paramere with hypophysis flattened, distinctly hooked (or shovel-headed or spade-shaped) apically and sensory lobe more or less projected basally; right paramere very tiny, with bulbous sensory lobe and simple, small hypophysis; vesica with at least two spinulate lobal-sclerites, without spicule; secondary gonopore thick-rimmed, small, rounded; apex of phallotheca smooth.
<emphasis id="B96FEA9BFFA83B1B136CFCB3FC3BFDC1" bold="true" pageId="21" pageNumber="392">Female genitalia</emphasis>
: Genital chamber with relatively wide dorsal sac; sclerotised rings narrow, thin-rimmed, separated to each other mesally; posterior wall lacking dorsal structure or lateral lobe; interramal lobe relatively developed, densely covered with scaly microstructures instead of spinules.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1518FFD3FB74FCE1" pageId="21" pageNumber="392" type="etymology">
<paragraph id="8BA43689FFA83B1B1518FFD3FB74FCE1" blockId="21.[813,1451,701,821]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1518FFD3FC1BFD01" bold="true" box="[813,943,701,725]" pageId="21" pageNumber="392">Etymology.</emphasis>
Named for distribution pattern of the congeners of this new genus, known widely from the Himalayas and Sundaland, combined with Greek noun, koris (= heteropteran bug); gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1518FE33FB21F882" pageId="21" pageNumber="392" type="discussion">
<paragraph id="8BA43689FFA83B1B1518FE33FBF0FA41" blockId="21.[813,1452,861,1429]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1518FE33FC06FCA1" bold="true" box="[813,946,861,885]" pageId="21" pageNumber="392">Discussion.</emphasis>
This new genus is assumed to be sister to
<taxonomicName id="4C1B4D0AFFA83B1B1518FE13FB3AFC41" authority="Zheng &amp; Lin, 2002" authorityName="Zheng &amp; Lin" authorityYear="2002" box="[813,1166,893,917]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFA83B1B1518FE13FC1BFC41" box="[813,943,893,917]" italics="true" pageId="21" pageNumber="392">Sabactiopus</emphasis>
<bibRefCitation id="EF8A4B78FFA83B1B158DFE13FB3AFC41" author="Zheng LY &amp; Lin CS" box="[952,1166,893,917]" pageId="21" pageNumber="392" pagination="75 - 81" refId="ref24959" refString="Zheng LY &amp; Lin CS (2002) Sabactiopus gen. nov. with a redescription of Sabactus institutus Distant, 1910 (Hemiptera: Miridae: Mirinae). Formosan Entomologist, 22: 75 - 81." type="journal article" year="2002">Zheng &amp; Lin, 2002</bibRefCitation>
</taxonomicName>
(see below). Both genera share the following characters: Dorsal surface shiny, well-polished, almost impunctate; head smooth, only with sparse vestiture; clypeus infuscate; pronotum and scutellum shiny, without puncture nor wrinkle; vesica widely membranous, with two spinulate lobal-sclerites, lacking spicule; secondary gonopore rounded, thick-rimmed; posterior wall lacking dorsal structure nor lateral lobe. However,
<taxonomicName id="4C1B4D0AFFA83B1B1338F933FA24FBA1" authorityName="Zheng &amp; Lin" authorityYear="2002" box="[1293,1424,1117,1141]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFA83B1B1338F933FA24FBA1" box="[1293,1424,1117,1141]" italics="true" pageId="21" pageNumber="392">Sabactiopus</emphasis>
</taxonomicName>
is separable from
<taxonomicName id="4C1B4D0AFFA83B1B15D4F913FB1DFB41" authorityName="Yasunaga" authorityYear="2024" box="[993,1193,1149,1173]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B15D4F913FB1DFB41" box="[993,1193,1149,1173]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B128EF910FA80FB42" box="[1211,1332,1150,1174]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
in having definitive characters, including presence of basal transverse carina on vertex (
<figureCitation id="13202A0CFFA83B1B15DEF9D3FBC2FB01" box="[1003,1142,1213,1237]" captionStart="Fig" captionStartId="49.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@49.[113,1451,251,1929]" captionTargetPageId="49" captionText="Fig. 34. Scanning electron micrographs of Paralygocoris vittulatus (AI), Sabactiopus schuhi, new species (JL) and S. sauteri (MO). A, head and pronotum, left lateral view; B, scutellum and hemelytron, left lateral view; C, metathoracic scent efferent system; D, metatarsus; E, pygophore, dorsal view; F, vesica; G, sclerotised ring; H, I, posterior wall; J, left lateral habitus; K, anterior body, dorsal view; L, O, thoracic pleura, left lateral view; M, anterior body, left lateral view; N, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683036" httpUri="https://zenodo.org/record/14683036/files/figure.png" pageId="21" pageNumber="392">Fig. 34K, M</figureCitation>
); antennomere III as long as or slightly longer than IV; scutellum fuscous, surrounded by creamy yellow margin (
<figureCitation id="13202A0CFFA83B1B125FF993FB58FAC1" box="[1130,1260,1277,1301]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="21" pageNumber="392">Fig. 7GH</figureCitation>
); right paramere conventional in shape and size (
<figureCitation id="13202A0CFFA83B1B1297F873FA9BFAE1" box="[1186,1327,1309,1333]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="21" pageNumber="392">Fig. 35E, K</figureCitation>
); narrower sclerotised ring (
<figureCitation id="13202A0CFFA83B1B15D8F853FBE3FA81" box="[1005,1111,1341,1365]" captionStart="Fig" captionStartId="50.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-58@50.[136,1474,265,1943]" captionTargetPageId="50" captionText="Fig. 35. Scanning electron micrographs of Sabactiopus schuhi, new species (AI) and S. sauteri (JO). A, J, metathoracic scent efferent system; B, M, metatarsus; C, pretarsal structure of metaleg; D, left paramere; E, right paramere; F, L, vesica; G, sclerotised ring; H, I, N, O, pygophore with parameres, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14683038" httpUri="https://zenodo.org/record/14683038/files/figure.png" pageId="21" pageNumber="392">Fig. 35G</figureCitation>
); and interramal lobe densely covered with spinules instead of scaly microstructures (cf.
<figureCitation id="13202A0CFFA83B1B1518F813FC0BFA41" box="[813,959,1405,1429]" captionStart="Fig" captionStartId="40.[136,171,1959,1981]" captionTargetBox="[136,1474,265,1943]" captionTargetId="figure-7@40.[136,1474,265,1943]" captionTargetPageId="40" captionText="Fig. 27. Scanning electron micrographs of Himalisundacoris duwalae, new species (AI) and H. omiae, new species (JO). A, B, K, Apical part of left paramere; C, right paramere; D, L, vesica; E, secondary gonopore; F, genital chamber; G, ovipositor (gonapophysis I); H, JN, posterior wall; I, posterior wall and ovipositor (gonapophysis II); O, sclerotised ring." figureDoi="http://doi.org/10.5281/zenodo.14683018" httpUri="https://zenodo.org/record/14683018/files/figure.png" pageId="21" pageNumber="392">Figs. 27H, M</figureCitation>
vs. 35I, O).
</paragraph>
<paragraph id="8BA43689FFA83B1B1518F8D3FB21F882" blockId="21.[813,1452,1469,1878]" pageId="21" pageNumber="392">
Incidentally, the extremely tiny, shortened right paramere possessed by
<taxonomicName id="4C1B4D0AFFA83B1B158EF8B3FBCEFA21" authorityName="Yasunaga" authorityYear="2024" box="[955,1146,1501,1525]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B158EF8B3FBCEFA21" box="[955,1146,1501,1525]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B12B3F8B0FB41FA22" box="[1158,1269,1502,1526]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
is not commonly found in the tribe
<taxonomicName id="4C1B4D0AFFA83B1B15CCF893FBF5F9C1" box="[1017,1089,1533,1557]" pageId="21" pageNumber="392" rank="tribe" tribe="Mirini">Mirini</taxonomicName>
. So far as I am aware, members of
<taxonomicName id="4C1B4D0AFFA83B1B157AFB73FB22F9E1" authority="Fieber, 1858" authorityName="Fieber" authorityYear="1858" box="[847,1174,1565,1589]" class="Insecta" family="Miridae" genus="Charagochilus" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFA83B1B157AFB73FC43F9E1" box="[847,1015,1565,1589]" italics="true" pageId="21" pageNumber="392">Charagochilus</emphasis>
<bibRefCitation id="EF8A4B78FFA83B1B1236FB73FB22F9E1" author="Fieber FX" box="[1027,1174,1565,1589]" pageId="21" pageNumber="392" pagination="289 - 327" refId="ref23813" refString="Fieber FX (1858) Criterien zur generischen Theilung der Phytocoriden (Capsini auct.). Wiener entomologische Monatschrift, 2: 289 - 327, 329 - 347, 388, 1 pl." type="journal article" year="1858">Fieber, 1858</bibRefCitation>
</taxonomicName>
have similarly reduced right parameres. However,
<taxonomicName id="4C1B4D0AFFA83B1B1246FB53FAC9F981" box="[1139,1405,1597,1621]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1246FB53FAA9F981" box="[1139,1309,1597,1621]" italics="true" pageId="21" pageNumber="392">Charagochilus</emphasis>
species
</taxonomicName>
are characterised by the heavily punctate dorsum, and totally different structures of the vesica and female genitalia (cf. Yasunaga, 2023). Therefore, the remarkable asymmetry of the parameres shared by the two genera is simply interpreted as a homoplasy. Additional undescribed congeners are remaining in
<collectingCountry id="F30C7619FFA83B1B15F4FB93FB9DF8C1" box="[961,1065,1789,1813]" name="Indonesia" pageId="21" pageNumber="392">Indonesia</collectingCountry>
(Schwartz MD, pers. comm.), and further investigation is encouraged to clarify the whole fauna of
<taxonomicName id="4C1B4D0AFFA83B1B157EFA53FBBEF881" authorityName="Yasunaga" authorityYear="2024" box="[843,1034,1853,1877]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="392" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFA83B1B157EFA53FBBEF881" box="[843,1034,1853,1877]" italics="true" pageId="21" pageNumber="392">Himalisundacoris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA83B1B122FFA50FB25F882" box="[1050,1169,1854,1878]" pageId="21" pageNumber="392" rank="genus">new genus</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,283 @@
<document id="4B578DB634139B0638639F2308756241" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFAF3B1B15A5FEB3FD74FDC1" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Euphorbialygus rufobrunneus Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="392" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="389" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="46924BA58D2B22A6D01956F0282D652D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0D53F538EBFCAC4DB12CA8FF73DA44E0">
<mods:title id="2BDF7C1DEA1669D3C14BB72CE3DAD4A8">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="2511FB36EFCAA498E3EF007893F59AA8" type="personal">
<mods:role id="D650785805790AAE2F6A8A9BF5DDDC17">
<mods:roleTerm id="6D8FE4BE50D8599EA6C7405DEE4CF151">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07C0F0DDE0DBFE572DB66396CF012070">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="95A85EB36406E20826DA2BED6D1AA15E">text</mods:typeOfResource>
<mods:relatedItem id="D3FC1BC04653041B1033EDD048781A94" type="host">
<mods:titleInfo id="FB2C922E7CFEB1777C856F27D634D9F7">
<mods:title id="4A544EF9F7D321BEFBEC61BE85614986">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="B828510048D3034BE0FD557F027EA3CB">
<mods:date id="E3DE135994540701532ED6D99373B83B">2024</mods:date>
<mods:detail id="D765AB5C44EBCD3ACA5FD7FA22F7A3FF" type="pubDate">
<mods:number id="4B6E21254FC6AD79E2B6F5C9B7ABBFDB">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="E51A7FA10F44B29A733AA3A7A05AA812" type="volume">
<mods:number id="5446F74FDAE7CDA3F4D1CB79941C2F93">72</mods:number>
</mods:detail>
<mods:extent id="50D5D54D8D0AB17E928BF983625343CE" unit="page">
<mods:start id="3471FE6017DB0238AF71BE0793C742E8">371</mods:start>
<mods:end id="5592140F2485828A492A88BD8B440A41">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="57DA67F878A4B2CD623AE49849900F87">journal article</mods:classification>
<mods:identifier id="5D0A67FF8752E49A89AFA45FF5A02AAE" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="61173BFA246F1EB6A9784319AF4753A3" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="1337E3FAC9FC4D1B7FC5755BFFE27A8A" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="2CDF445A40A821238B19AADB71BACDD2" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFAF3B1B15A5FEB3FD74FDC1" LSID="urn:lsid:plazi:treatment:03B2879FFFAF3B1B15A5FEB3FD74FDC1" httpUri="http://treatment.plazi.org/id/03B2879FFFAF3B1B15A5FEB3FD74FDC1" lastPageId="21" lastPageNumber="392" pageId="18" pageNumber="389">
<subSubSection id="C3016502FFAF3B1C15A5FEB3FAC2FC21" box="[912,1398,989,1013]" pageId="18" pageNumber="389" type="nomenclature">
<paragraph id="8BA43689FFAF3B1C15A5FEB3FAC2FC21" blockId="18.[912,1398,989,1045]" box="[912,1398,989,1013]" pageId="18" pageNumber="389">
<heading id="D0EC81E5FFAF3B1C15A5FEB3FAC2FC21" bold="true" box="[912,1398,989,1013]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="389" reason="2">
<emphasis id="B96FEA9BFFAF3B1C15A5FEB3FAC2FC21" bold="true" box="[912,1398,989,1013]" pageId="18" pageNumber="389">
<taxonomicName id="4C1B4D0AFFAF3B1C15A5FEB3FB56FC21" authorityName="Yasunaga" authorityYear="2024" box="[912,1250,989,1013]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFAF3B1C15A5FEB3FB56FC21" bold="true" box="[912,1250,989,1013]" italics="true" pageId="18" pageNumber="389">Euphorbialygus rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C12C4FEB3FAC2FC21" box="[1265,1398,989,1013]" pageId="18" pageNumber="389" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C15F3FE93FA8BFBC1" box="[966,1343,1021,1045]" pageId="18" pageNumber="389" type="description">
<paragraph id="8BA43689FFAF3B1C15F3FE93FA8BFBC1" blockId="18.[912,1398,989,1045]" box="[966,1343,1021,1045]" pageId="18" pageNumber="389">
(
<figureCitation id="13202A0CFFAF3B1C15FBFE93FBDAFBC1" box="[974,1134,1021,1045]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Figs. 8E, F, H</figureCitation>
,
<figureCitation id="13202A0CFFAF3B1C124FFE93FB32FBC1" box="[1146,1158,1021,1045]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="18" pageNumber="389">9</figureCitation>
FH, 10GI, 25)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1576F953FBFDFAC1" pageId="18" pageNumber="389" type="materials_examined">
<paragraph id="8BA43689FFAF3B1C1576F953FBFDFAC1" blockId="18.[835,1475,1085,1301]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576F953FC47FB81" bold="true" box="[835,1011,1085,1109]" pageId="18" pageNumber="389">Type material.</emphasis>
<materialsCitation id="3B733CD4FFAF3B1C15CBF953FA0BFB01" collectingDate="2005-11-07" collectingDateMax="2005-11-09" collectingDateMin="2005-11-07" collectionCode="T, M, B, NMTU, AMNH" country="Nepal" county="Machan Resort" elevation="270" latitude="7.535" location="Makawanpur Dist." longLatPrecision="21" longitude="84.73805" municipality="Chitwan National Park" pageId="18" pageNumber="389" specimenCode="PBI 378781" specimenCount="1" typeStatus="holotype">
<emphasis id="B96FEA9BFFAF3B1C15CBF953FBDDFB81" bold="true" box="[1022,1129,1085,1109]" pageId="18" pageNumber="389">
<typeStatus id="54A0882BFFAF3B1C15CBF953FBDDFB81" box="[1022,1129,1085,1109]" pageId="18" pageNumber="389" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFAF3B1C12F7F953FA93FB81" bold="true" box="[1218,1319,1085,1109]" pageId="18" pageNumber="389">
<collectingCountry id="F30C7619FFAF3B1C12F7F953FA96FB81" box="[1218,1314,1085,1109]" name="Nepal" pageId="18" pageNumber="389">NEPAL</collectingCountry>
:
</emphasis>
<location id="8EC46052FFAF3B1C1307F953FCC9FBA1" LSID="urn:lsid:plazi:treatment:03B2879FFFAF3B1B15A5FEB3FD74FDC1:8EC46052FFAF3B1C1307F953FCC9FBA1" country="Nepal" county="Machan Resort" latitude="7.535" longLatPrecision="21" longitude="84.73805" municipality="Chitwan National Park" name="Makawanpur Dist." pageId="18" pageNumber="389">Makawanpur Dist.</location>
,
<collectingMunicipality id="6BC0ACF3FFAF3B1C15B8F933FB29FBA1" box="[909,1181,1117,1141]" pageId="18" pageNumber="389">Chitwan National Park</collectingMunicipality>
,
<collectingCounty id="62C54E05FFAF3B1C1299F933FAE8FBA1" box="[1196,1372,1117,1141]" pageId="18" pageNumber="389">Machan Resort</collectingCounty>
[current Parsa Wildlife Reserve],
<geoCoordinate id="EE2F504EFFAF3B1C127AF912FB77FB41" box="[1103,1219,1148,1173]" degrees="7" direction="north" minutes="32" orientation="latitude" pageId="18" pageNumber="389" precision="15" seconds="06" value="7.535">7°3206″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFAF3B1C12FBF912FAF8FB41" box="[1230,1356,1148,1173]" degrees="84" direction="east" minutes="44" orientation="longitude" pageId="18" pageNumber="389" precision="15" seconds="17" value="84.73805">84°4417″E</geoCoordinate>
,
<quantity id="4CE39B6CFFAF3B1C1362F913FA28FB41" box="[1367,1436,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="389" unit="m" value="270.0">
<elevation id="0036D1BAFFAF3B1C1362F913FA28FB41" box="[1367,1436,1149,1173]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="18" pageNumber="389" unit="m" value="270.0">270 m</elevation>
</quantity>
, on flowers of
<taxonomicName id="4C1B4D0AFFAF3B1C1582F9F3FB89FB61" box="[951,1085,1181,1205]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96FEA9BFFAF3B1C1582F9F3FBA2FB61" box="[951,1046,1181,1205]" italics="true" pageId="18" pageNumber="389">Mallotus</emphasis>
sp.
</taxonomicName>
,
<date id="FFA51049FFAF3B1C127CF9F3FA9DFB61" box="[1097,1321,1181,1205]" pageId="18" pageNumber="389" value="2005-11-07" valueMax="2005-11-09" valueMin="2005-11-07">
<collectingDate id="EFE1E9A1FFAF3B1C127CF9F3FA9DFB61" box="[1097,1321,1181,1205]" pageId="18" pageNumber="389" value="2005-11-07" valueMax="2005-11-09" valueMin="2005-11-07">79 November 2005</collectingDate>
</date>
,
<collectionCode id="ED0AAE4CFFAF3B1C1301F9F0FAF3FB61" box="[1332,1351,1182,1205]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="389">T</collectionCode>
. Yasunaga,
<collectionCode id="ED0AAE4CFFAF3B1C1576F9D0FCEAFB01" box="[835,862,1214,1237]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="18" pageNumber="389" type="Herbarium">M</collectionCode>
. Takai &amp;
<collectionCode id="ED0AAE4CFFAF3B1C15E7F9D0FC53FB01" box="[978,999,1214,1237]" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="18" pageNumber="389" type="Herbarium">B</collectionCode>
. Shishido (
<collectionCode id="ED0AAE4CFFAF3B1C1256F9D3FB01FB01" box="[1123,1205,1213,1237]" pageId="18" pageNumber="389">NMTU</collectionCode>
) (
<collectionCode id="ED0AAE4CFFAF3B1C12FBF9D3FA95FB01" box="[1230,1313,1213,1237]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFAF3B1C131AF9D3FA03FB01" box="[1327,1463,1213,1237]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">PBI 378781</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFAF3B1C1576F9B0FBF1FAC1" collectingDate="2005-11-07" collectingDateMax="2005-11-09" collectingDateMin="2005-11-07" collectionCode="AMNH, TYCN, ZRC" country="Nepal" county="Machan Resort" elevation="270" latitude="7.535" location="Makawanpur Dist." longLatPrecision="21" longitude="84.73805" municipality="Chitwan National Park" pageId="18" pageNumber="389" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="paratype">
<typeStatus id="54A0882BFFAF3B1C1576F9B0FC04FB22" box="[835,944,1246,1270]" pageId="18" pageNumber="389" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D1DFD00FFAF3B1C158BF9B3FBA2FB21" box="[958,1046,1245,1269]" count="5" pageId="18" pageNumber="389" type="male">5 males</specimenCount>
,
<specimenCount id="9D1DFD00FFAF3B1C1210F9B3FB20FB21" box="[1061,1172,1245,1269]" count="3" pageId="18" pageNumber="389" type="female">3 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFAF3B1C157EF993FC15FAC1" box="[843,929,1277,1301]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFFAF3B1C1598F993FC4CFAC1" box="[941,1016,1277,1301]" pageId="18" pageNumber="389">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFFAF3B1C1230F993FB8AFAC1" box="[1029,1086,1277,1301]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="389">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1576F853FB75F9E1" pageId="18" pageNumber="389" type="diagnosis">
<paragraph id="8BA43689FFAF3B1C1576F853FB75F9E1" blockId="18.[835,1474,1341,1589]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576F853FC0DFA81" bold="true" box="[835,953,1341,1365]" pageId="18" pageNumber="389">Diagnosis.</emphasis>
<taxonomicName id="4C1B4D0AFFAF3B1C15F6F853FABEFA81" authorityName="Yasunaga" authorityYear="2024" box="[963,1290,1341,1365]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFAF3B1C15F6F853FABEFA81" box="[963,1290,1341,1365]" italics="true" pageId="18" pageNumber="389">Euphorbialygus rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C132EF850FA16FA81" box="[1307,1442,1341,1365]" pageId="18" pageNumber="389" rank="species">new species</taxonomicNameLabel>
, is distinguished from
<taxonomicName id="4C1B4D0AFFAF3B1C1222F830FB28FAA1" authorityName="Yasunaga" authorityYear="2024" box="[1047,1180,1373,1397]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFAF3B1C1222F830FB28FAA1" box="[1047,1180,1373,1397]" italics="true" pageId="18" pageNumber="389">E. nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C1299F830FA85FAA1" box="[1196,1329,1373,1397]" pageId="18" pageNumber="389" rank="species">new species</taxonomicNameLabel>
, by its larger size; densely distributed, micro-setae on pronotal collar; larger and triangular dark spot on cuneus; longer hypophysis and weakly inflated sensory lobe of left paramere; shorter flagellate lobal-sclerite on vesica; smaller paired process on posterior wall; and more densely distributed scaly microstructures on interramal lobe.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1B1576FB33FE05FF01" lastPageId="21" lastPageNumber="392" pageId="18" pageNumber="389" type="description">
<paragraph id="8BA43689FFAF3B1D1576FB33FA13F7CA" blockId="18.[835,1475,1629,2101]" lastBlockId="19.[813,1452,1862,2078]" lastPageId="19" lastPageNumber="390" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576FB33FC60F9A1" bold="true" box="[835,980,1629,1653]" pageId="18" pageNumber="389">Description.</emphasis>
Body generally dark reddish brown, oval, comparatively large in size; dorsal surface weakly shining (
<figureCitation id="13202A0CFFAF3B1C157EFBF3FC70F961" box="[843,964,1693,1717]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8A, C</figureCitation>
). Head shiny brown, with sparsely distributed, short, semi-erect setae; jugum and lorum pale brown, partly suffused with red; clypeus dark brown. Antenna dark brown; antennomere I pale reddish brown, with basal ⅓–½ darkened; antennomere II with pale basal ⅓–½ and basal dark ring; extreme base of antennomere III creamy yellow. Labium shiny pale brown, reaching but not exceeding apex of metacoxa; segment IV darkened. Pronotum shining, uniformly castaneous; collar densely furnished with micro-setae or pruinosity (
<figureCitation id="13202A0CFFAF3B1C1210FAD3FB3DF801" box="[1061,1161,1981,2005]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="18" pageNumber="389">Fig. 25B</figureCitation>
) pleura pale brown; ventral margin of propleuron, posterior margin of mesepisternum and metathoracic scent efferent system creamy yellow; scutellum castaneous, without noticeable spot or macula (
<figureCitation id="13202A0CFFAF3B1C1368F573FA01F7E1" box="[1373,1461,2077,2101]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8E</figureCitation>
). Hemelytra weakly shining, reddish chocolate brown; cuneus sanguineous, with a dark, semi-circular spot contiguous to lateral margin (cf.
<figureCitation id="13202A0CFFAE3B1D1773FAE8FE09F84B" box="[326,445,1926,1951]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="19" pageNumber="390">Fig. 8E, F</figureCitation>
); membrane smoky brown, with pale red veins. All coxae and legs pale brown, partly suffused with red; apical part of each femur more or less darkened; each tarsomere III dark brown; pretarsal structures as in
<figureCitation id="13202A0CFFAE3B1D1685F568FEA1F7CA" box="[176,277,2054,2078]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">Fig. 25G</figureCitation>
. Abdomen pale brown, partly with reddish spots. Male genitalia (
<figureCitation id="13202A0CFFAE3B1D1750F548FE04F7EB" box="[357,432,2086,2111]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="19" pageNumber="390">Figs. 9</figureCitation>
FH, 25HJ): Hypophysis of left paramere stout, long (
<figureCitation id="13202A0CFFAE3B1D120FFA28FB3CF88B" box="[1082,1160,1862,1887]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="19" pageNumber="390">Fig. 9F</figureCitation>
,
<figureCitation id="13202A0CFFAE3B1D12A7FA28FB0DF88A" box="[1170,1209,1862,1886]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">25I</figureCitation>
). Vesical spicule thick, curved subapically; flagellate lobal-sclerite comparatively short (
<figureCitation id="13202A0CFFAE3B1D1542FAE8FC67F84B" box="[887,979,1926,1951]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="19" pageNumber="390">Fig. 9H</figureCitation>
,
<figureCitation id="13202A0CFFAE3B1D15D6FAE8FB86F84B" box="[995,1074,1926,1951]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">25J, K</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFFAE3B1D132CFAE8FAC2F84B" box="[1305,1398,1926,1951]" captionStart="Fig" captionStartId="17.[113,148,1674,1696]" captionTargetBox="[113,1449,189,1659]" captionTargetId="figure-67@17.[113,1449,189,1659]" captionTargetPageId="17" captionText="Fig. 10. Female genitalia of Euphorbialygus nanglong, new species (AF) and E. rufobrunneus, new species (GI). A, Genital chamber with ovipositors (gonapophyses II), ventral view; C, G, genital chamber, ventral view; D, ditto, dorsal view; E, sclerotised ring; F, I, posterior wall, anterior view; apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682972" httpUri="https://zenodo.org/record/14682972/files/figure.png" pageId="19" pageNumber="390">Figs. 10</figureCitation>
GI, 25LO): Sclerotised rings relatively enlarged (
<figureCitation id="13202A0CFFAE3B1D1303FAC8FA13F86B" box="[1334,1447,1958,1983]" captionStart="Fig" captionStartId="17.[113,148,1674,1696]" captionTargetBox="[113,1449,189,1659]" captionTargetId="figure-67@17.[113,1449,189,1659]" captionTargetPageId="17" captionText="Fig. 10. Female genitalia of Euphorbialygus nanglong, new species (AF) and E. rufobrunneus, new species (GI). A, Genital chamber with ovipositors (gonapophyses II), ventral view; C, G, genital chamber, ventral view; D, ditto, dorsal view; E, sclerotised ring; F, I, posterior wall, anterior view; apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682972" httpUri="https://zenodo.org/record/14682972/files/figure.png" pageId="19" pageNumber="390">Figs. 10G</figureCitation>
,
<figureCitation id="13202A0CFFAE3B1D1518FAA8FCD6F80A" box="[813,866,1990,2014]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">25M</figureCitation>
); anterior paired process on posterior wall weak, reduced (
<figureCitation id="13202A0CFFAE3B1D1500FA88FC2AF82A" box="[821,926,2022,2046]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">Fig. 25N</figureCitation>
); interramal lobes narrow, with rather densely distributed scaly microstructures on distal half (
<figureCitation id="13202A0CFFAE3B1D1300F568FA2FF7CA" box="[1333,1435,2054,2078]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="19" pageNumber="390">Fig. 25O</figureCitation>
).
</paragraph>
<caption id="DF646601FFAE3B1D1644FBB3FA19F8DB" ID-DOI="http://doi.org/10.5281/zenodo.14682976" ID-Zenodo-Dep="14682976" httpUri="https://zenodo.org/record/14682976/files/figure.png" pageId="19" pageNumber="390" startId="19.[113,148,1757,1779]" targetBox="[135,1437,204,1724]" targetPageId="19" targetType="figure">
<paragraph id="8BA43689FFAE3B1D1644FBB3FA19F8DB" blockId="19.[113,1453,1757,1807]" pageId="19" pageNumber="390">
Fig. 11. Habitus images of
<taxonomicName id="4C1B4D0AFFAE3B1D17BDFBB3FD31F927" box="[392,645,1757,1779]" pageId="19" pageNumber="390">
<emphasis id="B96FEA9BFFAE3B1D17BDFBB3FD82F926" box="[392,566,1757,1778]" italics="true" pageId="19" pageNumber="390">Himalisundacoris</emphasis>
species
</taxonomicName>
from Sumatra and Sulawesi, Indonesia. A, B,
<taxonomicName id="4C1B4D0AFFAE3B1D1257FBB3FB74F926" authorityName="Yasunaga" authorityYear="2024" box="[1122,1216,1757,1778]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="390" phylum="Arthropoda" rank="species" species="palolo" status="new species">
<emphasis id="B96FEA9BFFAE3B1D1257FBB3FB74F926" box="[1122,1216,1757,1778]" italics="true" pageId="19" pageNumber="390">H. palolo</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAE3B1D12E5FBB3FAFEF927" box="[1232,1354,1757,1779]" pageId="19" pageNumber="390" rank="species">new species</taxonomicNameLabel>
, holotype male; C, D, ditto, female; E, F,
<taxonomicName id="4C1B4D0AFFAE3B1D1792FB94FE49F8DA" authorityName="Yasunaga" authorityYear="2024" box="[423,509,1785,1807]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="390" phylum="Arthropoda" rank="species" species="takaii" status="new species">
<emphasis id="B96FEA9BFFAE3B1D1792FB94FE49F8DA" box="[423,509,1785,1807]" italics="true" pageId="19" pageNumber="390">H. takaii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAE3B1D143CFB94FD34F8DB" box="[521,640,1785,1807]" pageId="19" pageNumber="390" rank="species">new species</taxonomicNameLabel>
, holotype male; G, H,
<taxonomicName id="4C1B4D0AFFAE3B1D156AFB94FC6EF8DA" authorityName="Yasunaga" authorityYear="2024" box="[863,986,1785,1807]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="390" phylum="Arthropoda" rank="species" species="schwartzi" status="new species">
<emphasis id="B96FEA9BFFAE3B1D156AFB94FC6EF8DA" box="[863,986,1785,1807]" italics="true" pageId="19" pageNumber="390">H. schwartzi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAE3B1D15D2FB94FBEAF8DB" box="[999,1118,1785,1807]" pageId="19" pageNumber="390" rank="species">new species</taxonomicNameLabel>
, holotype male; I, J, ditto, female.
</paragraph>
</caption>
<caption id="DF646601FFA93B1A16B2FAC7FC50F82C" ID-DOI="http://doi.org/10.5281/zenodo.14682978" ID-Zenodo-Dep="14682978" httpUri="https://zenodo.org/record/14682978/files/figure.png" pageId="20" pageNumber="391" startId="20.[135,170,1961,1983]" targetBox="[150,1461,271,1930]" targetPageId="20" targetType="figure">
<paragraph id="8BA43689FFA93B1A16B2FAC7FC50F82C" blockId="20.[135,1474,1961,2040]" pageId="20" pageNumber="391">
Fig. 12. Habitus images of
<taxonomicName id="4C1B4D0AFFA93B1A17A7FAC7FD3EF86B" box="[402,650,1961,1983]" pageId="20" pageNumber="391">
<emphasis id="B96FEA9BFFA93B1A17A7FAC7FD8AF86A" box="[402,574,1961,1982]" italics="true" pageId="20" pageNumber="391">Himalisundacoris</emphasis>
species
</taxonomicName>
from Bali, Indonesia (AH) and
<taxonomicName id="4C1B4D0AFFA93B1A15E4FAC7FB13F86B" box="[977,1191,1961,1983]" pageId="20" pageNumber="391">
<emphasis id="B96FEA9BFFA93B1A15E4FAC7FBEEF86A" box="[977,1114,1961,1982]" italics="true" pageId="20" pageNumber="391">Paralygocoris</emphasis>
species
</taxonomicName>
(HL). A, B,
<taxonomicName id="4C1B4D0AFFA93B1A1306FAC7FA39F86A" authorityName="Yasunaga" authorityYear="2024" box="[1331,1421,1961,1982]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="391" phylum="Arthropoda" rank="species" species="omiae" status="new species">
<emphasis id="B96FEA9BFFA93B1A1306FAC7FA39F86A" box="[1331,1421,1961,1982]" italics="true" pageId="20" pageNumber="391">H. omiae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA93B1A13AFFAC7FF7BF80F" pageId="20" pageNumber="391" rank="species">new species</taxonomicNameLabel>
, holotype male; C, D, ditto, female; E, F,
<taxonomicName id="4C1B4D0AFFA93B1A145BFAA8FD09F80E" authorityName="Yasunaga" authorityYear="2024" box="[622,701,1989,2011]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="391" phylum="Arthropoda" rank="species" species="theoi" status="new species">
<emphasis id="B96FEA9BFFA93B1A145BFAA8FD09F80E" box="[622,701,1989,2011]" italics="true" pageId="20" pageNumber="391">H. theoi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA93B1A14FEFAA8FCF6F80F" box="[715,834,1989,2011]" pageId="20" pageNumber="391" rank="species">new species</taxonomicNameLabel>
, male; G, H, ditto, female; H, I.
<taxonomicName id="4C1B4D0AFFA93B1A12B1FAA8FB51F80E" authorityName="Yasunaga" authorityYear="2024" box="[1156,1253,1989,2011]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="391" phylum="Arthropoda" rank="species" species="balicus" status="new species">
<emphasis id="B96FEA9BFFA93B1A12B1FAA8FB51F80E" box="[1156,1253,1989,2011]" italics="true" pageId="20" pageNumber="391">P. balicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFA93B1A12C6FAA8FADFF80F" box="[1267,1387,1989,2011]" pageId="20" pageNumber="391" rank="species">new species</taxonomicNameLabel>
, male; J, ditto, female; K, L,
<taxonomicName id="4C1B4D0AFFA93B1A1779FA8DFE56F823" authorityName="Lu &amp; Zheng" authorityYear="2004" box="[332,482,2018,2040]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="391" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFFA93B1A1779FA8DFE56F823" box="[332,482,2018,2040]" italics="true" pageId="20" pageNumber="391">P. meridionalis</emphasis>
</taxonomicName>
, female; M, N,
<taxonomicName id="4C1B4D0AFFA93B1A14B7FA8DFD43F823" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[642,759,2018,2040]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="391" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFFA93B1A14B7FA8DFD43F823" box="[642,759,2018,2040]" italics="true" pageId="20" pageNumber="391">P. vittulatus</emphasis>
</taxonomicName>
, male; O, ditto, female.
</paragraph>
</caption>
<paragraph id="8BA43689FFA83B1B1644FDD3FE05FF01" blockId="21.[113,433,189,213]" box="[113,433,189,213]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FDD3FE94FF01" bold="true" box="[113,288,189,213]" pageId="21" pageNumber="392">Measurements.</emphasis>
See
<tableCitation id="C6990332FFA83B1B1762FDD3FE1FFF01" box="[343,427,189,213]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="21" pageNumber="392">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644FD93FE82FE81" pageId="21" pageNumber="392" type="etymology">
<paragraph id="8BA43689FFA83B1B1644FD93FE82FE81" blockId="21.[113,753,253,341]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FD93FF45FEC1" bold="true" box="[113,241,253,277]" pageId="21" pageNumber="392">Etymology.</emphasis>
From Latin, rufo (or ruber = red) + brunneus (brown), referring to the general colouration of this new species; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644FC13FDE7FE41" box="[113,595,381,405]" pageId="21" pageNumber="392" type="distribution">
<paragraph id="8BA43689FFA83B1B1644FC13FDE7FE41" blockId="21.[113,595,381,405]" box="[113,595,381,405]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FC13FEB0FE41" bold="true" box="[113,260,381,405]" pageId="21" pageNumber="392">Distribution.</emphasis>
<collectingCountry id="F30C7619FFA83B1B1739FC13FEF8FE41" box="[268,332,381,405]" name="Nepal" pageId="21" pageNumber="392">Nepal</collectingCountry>
(Makawanpur District).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFA83B1B1644FCD3FD74FDC1" pageId="21" pageNumber="392" type="biology_ecology">
<paragraph id="8BA43689FFA83B1B1644FCD3FD74FDC1" blockId="21.[113,753,445,533]" pageId="21" pageNumber="392">
<emphasis id="B96FEA9BFFA83B1B1644FCD3FF7AFE01" bold="true" box="[113,206,445,469]" pageId="21" pageNumber="392">Biology.</emphasis>
All available specimens were collected by sweep-netting the inflorescences of
<taxonomicName id="4C1B4D0AFFA83B1B1785FCB3FD83FE21" box="[432,567,477,501]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="392" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96FEA9BFFA83B1B1785FCB3FDBBFE21" box="[432,527,477,501]" italics="true" pageId="21" pageNumber="392">Mallotus</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="4C1B4D0AFFA83B1B1472FCB3FD5DFE21" box="[583,745,477,501]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="392" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) (
<figureCitation id="13202A0CFFA83B1B164CFC93FF65FDC1" box="[121,209,509,533]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="21" pageNumber="392">Fig. 8H</figureCitation>
). No other information is currently available.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,243 @@
<document id="78302833305BE029621C57E90FA0BA92" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFAF3B1C16D9FC93FAA5FC61" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Euphorbialygus nanglong Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="389" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="389" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2568E24F0F7D1E3A743865FDE37D5A99" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4D505AF4F5C2FD02AFC8BCB6E5D8DAB3">
<mods:title id="C478CB0E233DF2D00E682CF914B31546">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="A57D02E75F62EC3EA4009335856CF9C6" type="personal">
<mods:role id="018B38CF6DD2C19A00B235CCABA404DD">
<mods:roleTerm id="6F67A880D15F5ABACFD1EB76650501FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FCF4DD65245FC97733A24C308021BA2">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="68006CFBDE6AD7AEFE6F68D65FFCAC74">text</mods:typeOfResource>
<mods:relatedItem id="027118369BC436133821F7C796A2CF0E" type="host">
<mods:titleInfo id="FD05E9E32E43B8A7B4106F4987697F4B">
<mods:title id="8B41B75B43CDEAE9AA6D8070DF633B87">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="FA5A17CB72311549ED27FFC8B8F678E0">
<mods:date id="F40E383854289E043ED1DB091821E9A0">2024</mods:date>
<mods:detail id="94AA9C359BBF78D75A08CA24C9478944" type="pubDate">
<mods:number id="67B8191781BA1E2B2B7BE372A9CC9DD8">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="86288F9192DF427A1C10953A4EEA6294" type="volume">
<mods:number id="DFF4A6135D2A0F0CCBC4E8E65483DBE4">72</mods:number>
</mods:detail>
<mods:extent id="32940C64E857ED5449315E8B5AE96966" unit="page">
<mods:start id="4CC39F2F36570E8CA4E621A7DFCE0672">371</mods:start>
<mods:end id="076818994180B951BAFDD23AABE7CE33">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CAA0D9130FA70F5F30979F9083E5447C">journal article</mods:classification>
<mods:identifier id="8D42B9EE7A626DBBEE22FC6F28D418C7" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="FC5FC9733E2AC1DAD2A2C680790D9518" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DF804686BB1C2E99F7575516B036AEB2" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="DDC3300F938924179E7085BA5A10BACD" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFAF3B1C16D9FC93FAA5FC61" ID-DOI="http://doi.org/10.5281/zenodo.14682994" ID-Zenodo-Dep="14682994" LSID="urn:lsid:plazi:treatment:03B2879FFFAF3B1C16D9FC93FAA5FC61" httpUri="http://treatment.plazi.org/id/03B2879FFFAF3B1C16D9FC93FAA5FC61" lastPageNumber="389" pageId="18" pageNumber="389">
<subSubSection id="C3016502FFAF3B1C16D9FC93FD17FDC1" box="[236,675,509,533]" pageId="18" pageNumber="389" type="nomenclature">
<paragraph id="8BA43689FFAF3B1C16D9FC93FD17FDC1" blockId="18.[236,675,509,565]" box="[236,675,509,533]" pageId="18" pageNumber="389">
<heading id="D0EC81E5FFAF3B1C16D9FC93FD17FDC1" bold="true" box="[236,675,509,533]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="389" reason="2">
<emphasis id="B96FEA9BFFAF3B1C16D9FC93FD17FDC1" bold="true" box="[236,675,509,533]" pageId="18" pageNumber="389">
<taxonomicName id="4C1B4D0AFFAF3B1C16D9FC93FDBAFDC1" authorityName="Yasunaga" authorityYear="2024" box="[236,526,509,533]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFAF3B1C16D9FC93FDBAFDC1" bold="true" box="[236,526,509,533]" italics="true" pageId="18" pageNumber="389">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C142BFC93FD17FDC1" box="[542,675,509,533]" pageId="18" pageNumber="389" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1730FF73FD3FFDE1" box="[261,651,541,565]" pageId="18" pageNumber="389" type="description">
<paragraph id="8BA43689FFAF3B1C1730FF73FD3FFDE1" blockId="18.[236,675,509,565]" box="[261,651,541,565]" pageId="18" pageNumber="389">
(
<figureCitation id="13202A0CFFAF3B1C1738FF73FE05FDE1" box="[269,433,541,565]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Figs. 8AD, G</figureCitation>
,
<figureCitation id="13202A0CFFAF3B1C178BFF73FE7EFDE1" box="[446,458,541,565]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="18" pageNumber="389">9</figureCitation>
AE, 10AF, 24)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C16BDFF33FE32FC41" pageId="18" pageNumber="389" type="materials_examined">
<paragraph id="8BA43689FFAF3B1C16BDFF33FE32FC41" blockId="18.[136,775,605,917]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C16BDFF33FE8DFDA1" bold="true" box="[136,313,605,629]" pageId="18" pageNumber="389">Type material.</emphasis>
<materialsCitation id="3B733CD4FFAF3B1C1771FF33FE54FD21" collectingDate="2012-12-31" collectionCode="T, DOAT, AMNH" country="Thailand" latitude="14.327639" location="Nang Rong Waterfall" longLatPrecision="1" longitude="101.31872" municipality="Sarika" pageId="18" pageNumber="389" specimenCode="PBI 378780" specimenCount="1" stateProvince="Nakhon Nayok" typeStatus="holotype">
<emphasis id="B96FEA9BFFAF3B1C1771FF33FE1BFDA1" bold="true" box="[324,431,605,629]" pageId="18" pageNumber="389">
<typeStatus id="54A0882BFFAF3B1C1771FF33FE1BFDA1" box="[324,431,605,629]" pageId="18" pageNumber="389" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFAF3B1C143CFF33FD11FDA1" bold="true" box="[521,677,605,629]" pageId="18" pageNumber="389">
<collectingCountry id="F30C7619FFAF3B1C143CFF33FD14FDA1" box="[521,672,605,629]" name="Thailand" pageId="18" pageNumber="389">THAILAND</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFAF3B1C1485FF33FF61FD41" country="Thailand" name="Nakhon Nayok" pageId="18" pageNumber="389">Nakhon Nayok</collectingRegion>
:
<collectingMunicipality id="6BC0ACF3FFAF3B1C16D1FF13FE9EFD41" box="[228,298,637,661]" pageId="18" pageNumber="389">Sarika</collectingMunicipality>
near
<location id="8EC46052FFAF3B1C175AFF10FDD7FD41" LSID="urn:lsid:plazi:treatment:03B2879FFFAF3B1C16D9FC93FAA5FC61:8EC46052FFAF3B1C175AFF10FDD7FD41" box="[367,611,637,662]" country="Thailand" latitude="14.327639" longLatPrecision="1" longitude="101.31872" municipality="Sarika" name="Nang Rong Waterfall" pageId="18" pageNumber="389" stateProvince="Nakhon Nayok">Nang Rong Waterfall</location>
along stream,
<geoCoordinate id="EE2F504EFFAF3B1C16BDFFF3FEAFFD61" box="[136,283,668,693]" degrees="14" direction="north" minutes="19" orientation="latitude" pageId="18" pageNumber="389" precision="1" seconds="39.5" value="14.327639">14°1939.5″N</geoCoordinate>
<geoCoordinate id="EE2F504EFFAF3B1C1711FFF3FE77FD61" box="[292,451,668,693]" degrees="101" direction="east" minutes="19" orientation="longitude" pageId="18" pageNumber="389" precision="1" seconds="07.4" value="101.31872">101°1907.4″E</geoCoordinate>
, on inflorescences and flower buds of
<taxonomicName id="4C1B4D0AFFAF3B1C16E9FFD0FE2BFD01" box="[220,415,702,725]" class="Magnoliopsida" family="Euphorbiaceae" genus="Homonoia" kingdom="Plantae" order="Malpighiales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B96FEA9BFFAF3B1C16E9FFD0FE2BFD01" box="[220,415,702,725]" italics="true" pageId="18" pageNumber="389">Homonoia riparia</emphasis>
</taxonomicName>
,
<date id="FFA51049FFAF3B1C179EFFD3FDC3FD01" box="[427,631,701,725]" pageId="18" pageNumber="389" value="2012-12-31">
<collectingDate id="EFE1E9A1FFAF3B1C179EFFD3FDC3FD01" box="[427,631,701,725]" pageId="18" pageNumber="389" value="2012-12-31">31 December 2012</collectingDate>
</date>
,
<collectionCode id="ED0AAE4CFFAF3B1C14B4FFD0FD20FD01" box="[641,660,702,725]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="389">T</collectionCode>
. Yasunaga (
<collectionCode id="ED0AAE4CFFAF3B1C16BAFFB3FF68FD21" box="[143,220,733,757]" pageId="18" pageNumber="389">DOAT</collectionCode>
) (
<collectionCode id="ED0AAE4CFFAF3B1C16C7FFB3FEF1FD21" box="[242,325,733,757]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFAF3B1C1766FFB3FE6DFD21" box="[339,473,733,757]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">PBI 378780</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFAF3B1C17DEFFB0FE6CFCA1" collectingDate="2012-12-31" collectionCode="AMNH, TYCN, ZRC, UV" country="Thailand" elevation="410" latitude="14.5075" location="Sakaerat Environmental Research Station" longLatPrecision="21" longitude="101.9275" municipality="Wang Nam Khiao" pageId="18" pageNumber="389" specimenCount="11" specimenCount-female="7" specimenCount-male="4" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<typeStatus id="54A0882BFFAF3B1C17DEFFB0FDE3FD22" box="[491,599,734,758]" pageId="18" pageNumber="389" type="paratype">Paratypes</typeStatus>
: THAILAND:
<specimenCount id="9D1DFD00FFAF3B1C14CCFFB3FF7DFCC1" count="3" pageId="18" pageNumber="389" type="male">3 males</specimenCount>
,
<specimenCount id="9D1DFD00FFAF3B1C16E7FF90FE8DFCC1" box="[210,313,765,789]" count="7" pageId="18" pageNumber="389" type="female">7 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFAF3B1C146CFF93FD1BFCC1" box="[601,687,765,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="18" pageNumber="389">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFFAF3B1C148DFF93FCB7FCC1" box="[696,771,765,789]" pageId="18" pageNumber="389">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFFAF3B1C16BDFE73FF76FCE1" box="[136,194,797,821]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="18" pageNumber="389">ZRC</collectionCode>
);
<specimenCount id="9D1DFD00FFAF3B1C16EFFE73FE9FFCE1" box="[218,299,797,821]" count="1" pageId="18" pageNumber="389" type="male">1 male</specimenCount>
,
<collectingRegion id="49DFF86BFFAF3B1C170FFE73FD95FCE1" box="[314,545,797,821]" country="Thailand" name="Nakhon Ratchasima" pageId="18" pageNumber="389">Nakhon Ratchasima</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFAF3B1C1405FE70FCB6FCE1" box="[560,770,797,822]" pageId="18" pageNumber="389">Wang Nam Khiao</collectingMunicipality>
,
<location id="8EC46052FFAF3B1C16BDFE53FDDFFC81" LSID="urn:lsid:plazi:treatment:03B2879FFFAF3B1C16D9FC93FAA5FC61:8EC46052FFAF3B1C16BDFE53FDDFFC81" box="[136,619,829,853]" latitude="14.5075" longLatPrecision="21" longitude="101.9275" municipality="Wang Nam Khiao" name="Sakaerat Environmental Research Station" pageId="18" pageNumber="389" stateProvince="Nakhon Ratchasima">Sakaerat Environmental Research Station</location>
,
<geoCoordinate id="EE2F504EFFAF3B1C144EFE53FCB6FC81" box="[635,770,828,853]" degrees="14" direction="north" minutes="30" orientation="latitude" pageId="18" pageNumber="389" precision="15" seconds="27" value="14.5075">14°3027″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFAF3B1C16BDFE33FEADFCA1" box="[136,281,860,885]" degrees="101" direction="east" minutes="55" orientation="longitude" pageId="18" pageNumber="389" precision="15" seconds="39" value="101.9275">101°5539″E</geoCoordinate>
,
<quantity id="4CE39B6CFFAF3B1C171DFE33FEC5FCA1" box="[296,369,861,885]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="389" unit="m" value="410.0">
<elevation id="0036D1BAFFAF3B1C171DFE33FEC5FCA1" box="[296,369,861,885]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="389" unit="m" value="410.0">410 m</elevation>
</quantity>
alt.,
<collectionCode id="ED0AAE4CFFAF3B1C1784FE30FE6CFCA1" box="[433,472,862,885]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="18" pageNumber="389">UV</collectionCode>
</materialsCitation>
lighting,
<collectionCode id="ED0AAE4CFFAF3B1C147EFE30FDEBFCA1" box="[587,607,862,885]" name="Tavera, Department of Geology and Geophysics" pageId="18" pageNumber="389">T</collectionCode>
.
<materialsCitation id="3B733CD4FFAF3B1C1458FE30FE37FC41" collectingDate="2008-10-28" collectionCode="TYCN" country="Thailand" pageId="18" pageNumber="389" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
Yasunaga,
<date id="FFA51049FFAF3B1C14D9FE33FEA8FC41" pageId="18" pageNumber="389" value="2008-10-28">
<collectingDate id="EFE1E9A1FFAF3B1C14D9FE33FEA8FC41" pageId="18" pageNumber="389" value="2008-10-28">28 October 2008</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFAF3B1C1719FE13FECFFC41" box="[300,379,893,917]" pageId="18" pageNumber="389">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C16BDFED3FEE7FAE1" pageId="18" pageNumber="389" type="diagnosis">
<paragraph id="8BA43689FFAF3B1C16BDFED3FEE7FAE1" blockId="18.[136,776,957,1333]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C16BDFED3FF49FC01" bold="true" box="[136,253,957,981]" pageId="18" pageNumber="389">Diagnosis.</emphasis>
This new species is recognised by the characters mentioned in the generic diagnosis but can be distinguished from the other congener,
<emphasis id="B96FEA9BFFAF3B1C179CFE90FDD0FBC1" box="[425,612,1021,1045]" italics="true" pageId="18" pageNumber="389">
<taxonomicName id="4C1B4D0AFFAF3B1C179CFE90FDD4FBC1" authorityName="Yasunaga" authorityYear="2024" box="[425,608,1021,1045]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="rufobrunneus">E. rufobrunneus</taxonomicName>
,
</emphasis>
by its smaller size; apically situated, dark, ovoid spot on cuneus; shorter hypophysis and triangularly projected sensory lobe of left paramere; sharp, longer flagellate lobal-sclerite on vesica; distinct paired process on posterior wall; and sparsely distributed scaly microstructures on interramal lobe. The final (5th) instar immature form is recognised by its ovoid body shape; greyish dorsum speckled with reddish brown maculae; and antenna and leg with reddish brown annulations or spots (
<figureCitation id="13202A0CFFAF3B1C16DBF873FEF2FAE1" box="[238,326,1309,1333]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C16BDF833FB37FD81" pageId="18" pageNumber="389" type="description">
<paragraph id="8BA43689FFAF3B1C16BDF833FB5AFDC1" blockId="18.[136,776,1373,2101]" lastBlockId="18.[835,1475,189,533]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C16BDF833FEA1FAA1" bold="true" box="[136,277,1373,1397]" pageId="18" pageNumber="389">Description.</emphasis>
Body generally reddish brown or brick-red, oval, comparatively small; dorsal surface weakly shining (
<figureCitation id="13202A0CFFAF3B1C16A5F8F3FEA7FA61" box="[144,275,1437,1461]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8A, C</figureCitation>
). Head shiny reddish brown, with sparsely distributed, short, semi-erect setae; jugum and lorum whitish; clypeus dark reddish brown (
<figureCitation id="13202A0CFFAF3B1C1426F8B3FDD1FA21" box="[531,613,1501,1525]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8B</figureCitation>
). Antenna dark brown; antennomere I pale reddish brown, with basal ⅓–½ darkened; antennomere II with pale basal half and basal dark ring; extreme base of antennomere III creamy yellow. Labium shiny pale brown, reaching or slightly exceeding apex of metacoxa; base of segments III and almost whole IV darkened (
<figureCitation id="13202A0CFFAF3B1C1716FBF3FECEF961" box="[291,378,1693,1717]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8B</figureCitation>
). Pronotum shining, uniformly pale reddish brown; pleura reddish brown, partly speckled with reddish suffusions (but fading to uniformly pale brown in dry-preserved specimens); collar smooth, without micro-setae or pruinosity (
<figureCitation id="13202A0CFFAF3B1C1713FA73FE18F8E1" box="[294,428,1821,1845]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="18" pageNumber="389">Fig. 24B, C</figureCitation>
); ventral margin of propleuron, posterior margin of mesepisternum and metathoracic scent efferent system creamy yellow; scutellum matte, with a pair of obscure maculae (
<figureCitation id="13202A0CFFAF3B1C1743FA13FE40F841" box="[374,500,1917,1941]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8A, C</figureCitation>
). Hemelytra matte, with somewhat darkened apical part of clavus and inner part of corium; cuneus pale reddish brown, with a dark, circular spot at posterolateral margin; membrane smoky brown, tinged with red anteriorly, with pale veins. All coxae and legs pale brown, partly suffused with red; apical part of each femur more or less speckled with brown spots or maculae; each tarsomere III dark brown; pretarsal structures as in
<figureCitation id="13202A0CFFAF3B1C13ADFDB3FCC1FEC1" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="18" pageNumber="389">Fig. 24H</figureCitation>
. Abdomen pale reddish brown, partly with sanguineous spots. Male genitalia (
<figureCitation id="13202A0CFFAF3B1C1202FC73FB36FEE1" box="[1079,1154,285,309]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="18" pageNumber="389">Figs. 9</figureCitation>
AE, 24IK): Hypophysis of left paramere short, with hooked apex (
<figureCitation id="13202A0CFFAF3B1C1335FC53FAE8FE81" box="[1280,1372,317,341]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="18" pageNumber="389">Fig. 9A</figureCitation>
). Vesical spicule relatively slender, weakly curved apically; flagellate lobal-sclerite long (
<figureCitation id="13202A0CFFAF3B1C122BFC13FBC1FE41" box="[1054,1141,381,405]" captionStart="Fig" captionStartId="16.[136,171,1550,1572]" captionTargetBox="[135,1473,189,1534]" captionTargetId="figure-42@16.[135,1473,189,1534]" captionTargetPageId="16" captionText="Fig. 9. Male genitalia of Euphorbialygus nanglong, new species (AE) and E. rufobrunneus, new species (FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="18" pageNumber="389">Fig. 9E</figureCitation>
,
<figureCitation id="13202A0CFFAF3B1C12B1FC13FB18FE41" box="[1156,1196,381,405]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="18" pageNumber="389">24I</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFFAF3B1C13B9FC13FCE8FE61" captionStart="Fig" captionStartId="17.[113,148,1674,1696]" captionTargetBox="[113,1449,189,1659]" captionTargetId="figure-67@17.[113,1449,189,1659]" captionTargetPageId="17" captionText="Fig. 10. Female genitalia of Euphorbialygus nanglong, new species (AF) and E. rufobrunneus, new species (GI). A, Genital chamber with ovipositors (gonapophyses II), ventral view; C, G, genital chamber, ventral view; D, ditto, dorsal view; E, sclerotised ring; F, I, posterior wall, anterior view; apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682972" httpUri="https://zenodo.org/record/14682972/files/figure.png" pageId="18" pageNumber="389">Figs. 10</figureCitation>
AF, 24LO): Sclerotised rings ovoid (
<figureCitation id="13202A0CFFAF3B1C12C0FCF3FAE7FE61" box="[1269,1363,413,437]" captionStart="Fig" captionStartId="17.[113,148,1674,1696]" captionTargetBox="[113,1449,189,1659]" captionTargetId="figure-67@17.[113,1449,189,1659]" captionTargetPageId="17" captionText="Fig. 10. Female genitalia of Euphorbialygus nanglong, new species (AF) and E. rufobrunneus, new species (GI). A, Genital chamber with ovipositors (gonapophyses II), ventral view; C, G, genital chamber, ventral view; D, ditto, dorsal view; E, sclerotised ring; F, I, posterior wall, anterior view; apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682972" httpUri="https://zenodo.org/record/14682972/files/figure.png" pageId="18" pageNumber="389">Fig. 10E</figureCitation>
); posterior wall with a pair of distinct processes anteriorly (
<figureCitation id="13202A0CFFAF3B1C1364FCD3FA02FE01" box="[1361,1462,445,469]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="18" pageNumber="389">Fig. 24N</figureCitation>
); interramal lobes narrow, with sparsely distributed scaly microstructures on distal ⅓ (
<figureCitation id="13202A0CFFAF3B1C1249FC93FB56FDC1" box="[1148,1250,509,533]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="18" pageNumber="389">Fig. 24O</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFAF3B1C1576FF53FB37FD81" blockId="18.[835,1155,573,597]" box="[835,1155,573,597]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576FF53FC46FD81" bold="true" box="[835,1010,573,597]" pageId="18" pageNumber="389">Measurements.</emphasis>
See
<tableCitation id="C6990332FFAF3B1C121CFF53FBC9FD81" box="[1065,1149,573,597]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="18" pageNumber="389">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1576FF13FB4CFD61" pageId="18" pageNumber="389" type="etymology">
<paragraph id="8BA43689FFAF3B1C1576FF13FB4CFD61" blockId="18.[835,1474,637,693]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576FF13FC77FD41" bold="true" box="[835,963,637,661]" pageId="18" pageNumber="389">Etymology.</emphasis>
Named after the
<typeStatus id="54A0882BFFAF3B1C1245FF10FB29FD42" box="[1136,1181,638,662]" pageId="18" pageNumber="389">type</typeStatus>
locality, Nanglong (
<collectingRegion id="49DFF86BFFAF3B1C1358FF13FC39FD61" country="Thailand" name="Nakhon Nayok" pageId="18" pageNumber="389">Nakhon Nayok</collectingRegion>
,
<collectingCountry id="F30C7619FFAF3B1C15AFFFF3FC4AFD61" box="[922,1022,669,693]" name="Thailand" pageId="18" pageNumber="389">Thailand</collectingCountry>
); a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1576FFB3FA77FD21" box="[835,1475,733,757]" pageId="18" pageNumber="389" type="distribution">
<paragraph id="8BA43689FFAF3B1C1576FFB3FA77FD21" blockId="18.[835,1475,733,757]" box="[835,1475,733,757]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576FFB3FC64FD21" bold="true" box="[835,976,733,757]" pageId="18" pageNumber="389">Distribution.</emphasis>
<collectingCountry id="F30C7619FFAF3B1C15E0FFB3FB85FD21" box="[981,1073,733,757]" name="Thailand" pageId="18" pageNumber="389">Thailand</collectingCountry>
(
<collectingRegion id="49DFF86BFFAF3B1C1209FFB3FB68FD21" box="[1084,1244,733,757]" country="Thailand" name="Nakhon Nayok" pageId="18" pageNumber="389">Nakhon Nayok</collectingRegion>
,
<collectingRegion id="49DFF86BFFAF3B1C12D1FFB3FA0DFD21" box="[1252,1465,733,757]" country="Thailand" name="Nakhon Ratchasima" pageId="18" pageNumber="389">Nakhon Ratchasima</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAF3B1C1576FE73FAA5FC61" pageId="18" pageNumber="389" type="biology_ecology">
<paragraph id="8BA43689FFAF3B1C1576FE73FAA5FC61" blockId="18.[835,1475,797,949]" pageId="18" pageNumber="389">
<emphasis id="B96FEA9BFFAF3B1C1576FE73FC17FCE1" bold="true" box="[835,931,797,821]" pageId="18" pageNumber="389">Biology.</emphasis>
In
<collectingRegion id="49DFF86BFFAF3B1C15FAFE73FBC8FCE1" box="[975,1148,797,821]" country="Thailand" name="Nakhon Nayok" pageId="18" pageNumber="389">Nakhon Nayok</collectingRegion>
, central
<collectingCountry id="F30C7619FFAF3B1C12D4FE73FAF2FCE1" box="[1249,1350,797,821]" name="Thailand" pageId="18" pageNumber="389">Thailand</collectingCountry>
, the adults including a few teneral (freshly emerging) individuals and final instar nymph (
<figureCitation id="13202A0CFFAF3B1C1267FE33FB1EFCA1" box="[1106,1194,861,885]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8D</figureCitation>
) were obtained from the inflorescence and flower-buds of
<taxonomicName id="4C1B4D0AFFAF3B1C1283FE10FA77FC41" authority="Lour." authorityName="Lour." box="[1206,1475,894,917]" class="Magnoliopsida" family="Euphorbiaceae" genus="Homonoia" kingdom="Plantae" order="Malpighiales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B96FEA9BFFAF3B1C1283FE10FAC9FC41" box="[1206,1405,894,917]" italics="true" pageId="18" pageNumber="389">Homonoia riparia</emphasis>
Lour.
</taxonomicName>
(
<taxonomicName id="4C1B4D0AFFAF3B1C157FFEF3FC58FC61" box="[842,1004,925,949]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) along a stream (
<figureCitation id="13202A0CFFAF3B1C1299FEF3FAB0FC61" box="[1196,1284,925,949]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="18" pageNumber="389">Fig. 8G</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,324 @@
<document id="166AF245479885D42975232C221DBE9E" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB03B0316EBFED3FA88F921" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus wolskiorum Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="384" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="384" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A5DBF05A316057402D98136F31AC9A08" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="343C046E19CBA92F1624D120FAE3BF83">
<mods:title id="AA22394A3B356B264854988AC31A3D8E">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="C641B70C73294AD51505140096D3B942" type="personal">
<mods:role id="C1488DA1F06AC8AC01A0054C38219C7E">
<mods:roleTerm id="AD92389D503DB28401751D5432B5113C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="88B99A9F7902A182AC3A224DEB45E636">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="EC52F4DC6983E880A9856F25BD410D51">text</mods:typeOfResource>
<mods:relatedItem id="EA60AD7F0587F88BDC88C8FADF1D4086" type="host">
<mods:titleInfo id="E9438DEAA18BA148565B81D8D29E7E2C">
<mods:title id="42338016B0B8BB58B897E1FE9BF179E4">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="FC8BB89D816FBB087D15DB71D776D391">
<mods:date id="3F72A918660861DF8B121D9C0B82E214">2024</mods:date>
<mods:detail id="FAAE0CF8009E45DA9D11B939409B23B2" type="pubDate">
<mods:number id="D6A0E0F8A9F72D753E2BA6DFFFD3F266">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="B2C01BC114FA4FBD90B7ACC5544BEE86" type="volume">
<mods:number id="D9228C7861336012A37652BB5217AE6C">72</mods:number>
</mods:detail>
<mods:extent id="94C15A3CC84DED5DD5AACD61F1F6A624" unit="page">
<mods:start id="07A73DD4F39C04D4ADBF0C0AC6A3E637">371</mods:start>
<mods:end id="F538B50DD00C389891795EE82DA84524">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B42D1D94681322C801CA7D6E2302C31A">journal article</mods:classification>
<mods:identifier id="A2FDDB3BE4783DC957BB0EF3CBFE7C01" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="48CF6C04561C72F483113A18231447EE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C4349258540510D5F6A12972EBB43E83" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="48358C4ACA5AF2AEC16DE6753A45ADDF" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB03B0316EBFED3FA88F921" ID-DOI="http://doi.org/10.5281/zenodo.14682984" ID-Zenodo-Dep="14682984" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921" httpUri="http://treatment.plazi.org/id/03B2879FFFB03B0316EBFED3FA88F921" lastPageNumber="384" pageId="13" pageNumber="384">
<subSubSection id="C3016502FFB03B0316EBFED3FD30FC01" box="[222,644,957,981]" pageId="13" pageNumber="384" type="nomenclature">
<paragraph id="8BA43689FFB03B0316EBFED3FD30FC01" blockId="13.[222,644,957,1013]" box="[222,644,957,981]" pageId="13" pageNumber="384">
<heading id="D0EC81E5FFB03B0316EBFED3FD30FC01" bold="true" box="[222,644,957,981]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="384" reason="2">
<emphasis id="B96FEA9BFFB03B0316EBFED3FD30FC01" bold="true" box="[222,644,957,981]" pageId="13" pageNumber="384">
<taxonomicName id="4C1B4D0AFFB03B0316EBFED3FE5BFC01" authorityName="Yasunaga" authorityYear="2024" box="[222,495,957,981]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="wolskiorum" status="sp. nov.">
<emphasis id="B96FEA9BFFB03B0316EBFED3FE5BFC01" bold="true" box="[222,495,957,981]" italics="true" pageId="13" pageNumber="384">Chelyolygus wolskiorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB03B0317CBFED3FD30FC01" box="[510,644,957,981]" pageId="13" pageNumber="384" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031719FEB3FD82FC21" box="[300,566,989,1013]" pageId="13" pageNumber="384" type="description">
<paragraph id="8BA43689FFB03B031719FEB3FD82FC21" blockId="13.[222,644,957,1013]" box="[300,566,989,1013]" pageId="13" pageNumber="384">
(
<figureCitation id="13202A0CFFB03B031701FEB3FECFFC21" box="[308,379,989,1013]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="13" pageNumber="384">Figs. 1</figureCitation>
GH, 3AF, 23)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644F973FDFFF9E1" pageId="13" pageNumber="384" type="materials_examined">
<paragraph id="8BA43689FFB03B031644F973FDFFF9E1" blockId="13.[113,753,1053,1589]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644F973FE96FBE1" bold="true" box="[113,290,1053,1077]" pageId="13" pageNumber="384">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB03B031718F973FEF9FB61" collectingDate="2008-09-16" collectionCode="UV, T, AMNH, DOAT" country="Thailand" county="Wang Nam Khiao" elevation="410" latitude="14.5075" location="Yasunaga" longLatPrecision="21" longitude="101.9275" municipality="Sakaerat Environmental Research Station" pageId="13" pageNumber="384" specimenCode="PBI 00380777" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="holotype">
<emphasis id="B96FEA9BFFB03B031718F973FE2CFBE1" bold="true" box="[301,408,1053,1077]" pageId="13" pageNumber="384">
<typeStatus id="54A0882BFFB03B031718F973FE2CFBE1" box="[301,408,1053,1077]" pageId="13" pageNumber="384" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFB03B0317C7F973FD3BFBE1" bold="true" box="[498,655,1053,1077]" pageId="13" pageNumber="384">
<collectingCountry id="F30C7619FFB03B0317C7F973FD3DFBE1" box="[498,649,1053,1077]" name="Thailand" pageId="13" pageNumber="384">THAILAND</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFB03B0314ACF973FF40FB81" country="Thailand" name="Nakhon Ratchasima" pageId="13" pageNumber="384">Nakhon Ratchasima</collectingRegion>
,
<collectingCounty id="62C54E05FFB03B031736F950FE60FB81" box="[259,468,1085,1110]" pageId="13" pageNumber="384">Wang Nam Khiao</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FFB03B0317D6F953FE93FBA1" pageId="13" pageNumber="384">Sakaerat Environmental Research Station</collectingMunicipality>
,
<geoCoordinate id="EE2F504EFFB03B031704F933FE07FBA1" box="[305,435,1116,1141]" degrees="14" direction="north" minutes="30" orientation="latitude" pageId="13" pageNumber="384" precision="15" seconds="27" value="14.5075">14°3027″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB03B031788F933FDFCFBA1" box="[445,584,1116,1141]" degrees="101" direction="east" minutes="55" orientation="longitude" pageId="13" pageNumber="384" precision="15" seconds="39" value="101.9275">101°5539″E</geoCoordinate>
,
<quantity id="4CE39B6CFFB03B031466F933FD22FBA1" box="[595,662,1117,1141]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="384" unit="m" value="410.0">
<elevation id="0036D1BAFFB03B031466F933FD22FBA1" box="[595,662,1117,1141]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="384" unit="m" value="410.0">410 m</elevation>
</quantity>
alt.,
<collectionCode id="ED0AAE4CFFB03B0314FFF930FD45FBA1" box="[714,753,1118,1141]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="13" pageNumber="384">UV</collectionCode>
lighting,
<collectionCode id="ED0AAE4CFFB03B0316EDF910FF58FB41" box="[216,236,1150,1173]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="384">T</collectionCode>
.
<location id="8EC46052FFB03B0316CFF910FEDCFB42" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B0316CFF910FEDCFB42" box="[250,360,1150,1174]" country="Thailand" county="Wang Nam Khiao" latitude="14.5075" longLatPrecision="21" longitude="101.9275" municipality="Sakaerat Environmental Research Station" name="Yasunaga" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Yasunaga</location>
,
<date id="FFA51049FFB03B031742F913FDE6FB41" box="[375,594,1149,1173]" pageId="13" pageNumber="384" value="2008-09-16">
<collectingDate id="EFE1E9A1FFB03B031742F913FDE6FB41" box="[375,594,1149,1173]" pageId="13" pageNumber="384" value="2008-09-16">16 September 2008</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB03B031451F913FD0CFB41" box="[612,696,1149,1173]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="384">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB03B0314F3F913FF69FB61" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="384">PBI 00380777</specimenCode>
) (
<collectionCode id="ED0AAE4CFFB03B0316C0F9F3FEF1FB61" box="[245,325,1181,1205]" pageId="13" pageNumber="384">DOAT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB03B03176DF9F0FF7BFAC1" collectingDate="2000-05-08" collectingDateMax="2000-05-11" collectingDateMin="2000-05-08" collectionCode="TYCN" collectorName="S. Nagashima" country="Thailand" county="Doi Suthep" latitude="18.833334" location="Univ. Exp. Forest" longLatPrecision="1273" longitude="98.9" municipality="Chiang Khian" pageId="13" pageNumber="384" specimenCount="1" stateProvince="Chiang Mai" typeStatus="paratype">
<typeStatus id="54A0882BFFB03B03176DF9F0FE70FB62" box="[344,452,1182,1206]" pageId="13" pageNumber="384" type="paratype">Paratypes</typeStatus>
: THAILAND:
<collectingRegion id="49DFF86BFFB03B03145CF9F3FD59FB61" box="[617,749,1181,1205]" country="Thailand" name="Chiang Mai" pageId="13" pageNumber="384">Chiang Mai</collectingRegion>
,
<collectingCounty id="62C54E05FFB03B031644F9D3FF5BFB01" box="[113,239,1213,1237]" pageId="13" pageNumber="384">Doi Suthep</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FFB03B0316CEF9D3FE20FB01" box="[251,404,1213,1237]" pageId="13" pageNumber="384">Chiang Khian</collectingMunicipality>
,
<collectingRegion id="49DFF86BFFB03B031795F9D3FD94FB01" box="[416,544,1213,1237]" country="Thailand" name="Chiang Mai" pageId="13" pageNumber="384">Chiang Mai</collectingRegion>
<location id="8EC46052FFB03B03141CF9D3FD58FB01" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B03141CF9D3FD58FB01" box="[553,748,1213,1237]" country="Thailand" county="Doi Suthep" latitude="18.833334" longLatPrecision="1273" longitude="98.9" municipality="Chiang Khian" name="Univ. Exp. Forest" pageId="13" pageNumber="384" stateProvince="Chiang Mai">Univ. Exp. Forest</location>
,
<geoCoordinate id="EE2F504EFFB03B031644F9B2FF7FFB21" box="[113,203,1244,1269]" degrees="18" direction="north" minutes="50" orientation="latitude" pageId="13" pageNumber="384" precision="925" value="18.833334">18°50N</geoCoordinate>
<geoCoordinate id="EE2F504EFFB03B0316E7F9B2FE9EFB21" box="[210,298,1244,1269]" degrees="98" direction="east" minutes="54" orientation="longitude" pageId="13" pageNumber="384" precision="925" value="98.9">98°54E</geoCoordinate>
,
<collectingMethod id="525A4E9EFFB03B031700F9B3FE2DFB21" box="[309,409,1245,1269]" pageId="13" pageNumber="384">light trap</collectingMethod>
,
<collectorName id="26EE535FFFB03B031791F9B3FD88FB21" box="[420,572,1245,1269]" pageId="13" pageNumber="384">S. Nagashima</collectorName>
,
<date id="FFA51049FFB03B031472F9B3FD44FB21" box="[583,752,1245,1270]" pageId="13" pageNumber="384" value="2000-05-08" valueMax="2000-05-11" valueMin="2000-05-08">
<collectingDate id="EFE1E9A1FFB03B031472F9B3FD44FB21" box="[583,752,1245,1270]" pageId="13" pageNumber="384" value="2000-05-08" valueMax="2000-05-11" valueMin="2000-05-08">811 May 2000</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB03B03164CF993FF73FAC1" box="[121,199,1277,1301]" pageId="13" pageNumber="384">TYCN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B733CD4FFB03B0316EDF993FD59FA81" collectionCode="T" country="Thailand" latitude="14.463638" location="Sakaerat Silvicultural Research Station" longLatPrecision="1" longitude="101.87092" municipality="Wang Nam Khiao" pageId="13" pageNumber="384" specimenCount="2" specimenCount-female="2" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB03B0316EDF993FEF4FAC1" box="[216,320,1277,1301]" count="2" pageId="13" pageNumber="384" type="female">2 females</specimenCount>
,
<collectingRegion id="49DFF86BFFB03B03177CF993FD96FAC1" box="[329,546,1277,1301]" country="Thailand" name="Nakhon Ratchasima" pageId="13" pageNumber="384">Nakhon Ratchasima</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFB03B03141EF990FD59FAC1" box="[555,749,1277,1302]" pageId="13" pageNumber="384">Wang Nam Khiao</collectingMunicipality>
,
<location id="8EC46052FFB03B031644F873FD89FAE1" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B031644F873FD89FAE1" box="[113,573,1309,1333]" country="Thailand" latitude="14.463638" longLatPrecision="1" longitude="101.87092" municipality="Wang Nam Khiao" name="Sakaerat Silvicultural Research Station" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Sakaerat Silvicultural Research Station</location>
,
<geoCoordinate id="EE2F504EFFB03B031478F872FD5FFAE1" box="[589,747,1308,1333]" degrees="14" direction="north" minutes="27" orientation="latitude" pageId="13" pageNumber="384" precision="1" seconds="49.1" value="14.463638">14°2749.1″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB03B031644F852FEA4FA81" box="[113,272,1340,1365]" degrees="101" direction="east" minutes="52" orientation="longitude" pageId="13" pageNumber="384" precision="1" seconds="15.3" value="101.87092">101°5215.3″E</geoCoordinate>
, sweeping inflorescence of a broadleaf tree,
<collectionCode id="ED0AAE4CFFB03B0314EFF850FD59FA81" box="[730,749,1342,1365]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="384">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB03B031644F830FE03FAA1" box="[113,439,1373,1398]" collectingDate="2013-09-26" country="Thailand" location="Yasunaga" pageId="13" pageNumber="384" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<location id="8EC46052FFB03B031644F830FF69FAA2" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B031644F830FF69FAA2" box="[113,221,1374,1398]" country="Thailand" name="Yasunaga" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Yasunaga</location>
,
<date id="FFA51049FFB03B0316D2F833FE03FAA1" box="[231,439,1373,1397]" pageId="13" pageNumber="384" value="2013-09-26">
<collectingDate id="EFE1E9A1FFB03B0316D2F833FE03FAA1" box="[231,439,1373,1397]" pageId="13" pageNumber="384" value="2013-09-26">26 September 2013</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B733CD4FFB03B0317F5F833FF06FA61" collectingDate="2013-09-27" collectionCode="TYCN, ZRC" country="Thailand" pageId="13" pageNumber="384" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB03B0317F5F833FDA8FAA1" box="[448,540,1373,1397]" count="1" pageId="13" pageNumber="384" type="female">1 female</specimenCount>
developed from 5th instar nymph and emerged on
<date id="FFA51049FFB03B031788F813FD3BFA41" box="[445,655,1405,1429]" pageId="13" pageNumber="384" value="2013-09-27">
<collectingDate id="EFE1E9A1FFB03B031788F813FD3BFA41" box="[445,655,1405,1429]" pageId="13" pageNumber="384" value="2013-09-27">27 September 2013</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB03B0314AAF813FD58FA41" box="[671,748,1405,1429]" pageId="13" pageNumber="384">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFFB03B031644F8F3FF1EFA61" box="[113,170,1437,1461]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="13" pageNumber="384">ZRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B733CD4FFB03B0316F5F8F3FE75FA21" collectionCode="UV" country="Thailand" elevation="410" latitude="14.5075" location="Sakaerat Environmental Research Station" longLatPrecision="21" longitude="101.9275" municipality="Wang Nam Khiao" pageId="13" pageNumber="384" specimenCount="1" specimenCount-female="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB03B0316F5F8F3FE90FA61" box="[192,292,1437,1461]" count="1" pageId="13" pageNumber="384" type="female">1 female</specimenCount>
,
<collectingRegion id="49DFF86BFFB03B031707F8F3FDA5FA61" box="[306,529,1437,1461]" country="Thailand" name="Nakhon Ratchasima" pageId="13" pageNumber="384">Nakhon Ratchasima</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFB03B03142AF8F0FD58FA61" box="[543,748,1437,1462]" pageId="13" pageNumber="384">Wang Nam Khiao</collectingMunicipality>
,
<location id="8EC46052FFB03B031644F8D3FDE1FA01" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B031644F8D3FDE1FA01" box="[113,597,1469,1493]" country="Thailand" latitude="14.5075" longLatPrecision="21" longitude="101.9275" municipality="Wang Nam Khiao" name="Sakaerat Environmental Research Station" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Sakaerat Environmental Research Station</location>
,
<geoCoordinate id="EE2F504EFFB03B031451F8D3FD5FFA01" box="[612,747,1468,1493]" degrees="14" direction="north" minutes="30" orientation="latitude" pageId="13" pageNumber="384" precision="15" seconds="27" value="14.5075">14°3027″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB03B031644F8B3FEB6FA21" box="[113,258,1500,1525]" degrees="101" direction="east" minutes="55" orientation="longitude" pageId="13" pageNumber="384" precision="15" seconds="39" value="101.9275">101°5539″E</geoCoordinate>
,
<quantity id="4CE39B6CFFB03B031727F8B3FEEEFA21" box="[274,346,1501,1525]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="384" unit="m" value="410.0">
<elevation id="0036D1BAFFB03B031727F8B3FEEEFA21" box="[274,346,1501,1525]" metricMagnitude="2" metricUnit="m" metricValue="4.1" pageId="13" pageNumber="384" unit="m" value="410.0">410 m</elevation>
</quantity>
alt.,
<collectionCode id="ED0AAE4CFFB03B0317AFF8B0FE75FA21" box="[410,449,1502,1525]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="13" pageNumber="384">UV</collectionCode>
</materialsCitation>
lighting,
<collectionCode id="ED0AAE4CFFB03B031401F8B0FDFCFA21" box="[564,584,1502,1525]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="384">T</collectionCode>
.
<materialsCitation id="3B733CD4FFB03B031462F8B0FE47F9C2" collectingDate="2009-01-24" collectionCode="TYCN" country="Thailand" location="Yasunaga" pageId="13" pageNumber="384" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<location id="8EC46052FFB03B031462F8B0FD73FA22" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B031462F8B0FD73FA22" box="[599,711,1502,1526]" country="Thailand" name="Yasunaga" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Yasunaga</location>
,
<date id="FFA51049FFB03B0314E0F8B3FEB6F9C1" pageId="13" pageNumber="384" value="2009-01-24">
<collectingDate id="EFE1E9A1FFB03B0314E0F8B3FEB6F9C1" pageId="13" pageNumber="384" value="2009-01-24">24 January 2009</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB03B031727F893FED5F9C1" box="[274,353,1533,1557]" pageId="13" pageNumber="384">TYCN</collectionCode>
). VIETNAM
</materialsCitation>
:
<materialsCitation id="3B733CD4FFB03B0317CAF893FF61F9E1" collectionCode="T" collectorName="Ba Vi" country="Thailand" location="Ha Tay Prov." pageId="13" pageNumber="384" specimenCount="1" specimenCount-male="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB03B0317CAF893FDFEF9C1" box="[511,586,1533,1557]" count="1" pageId="13" pageNumber="384" type="male">1 male</specimenCount>
,
<location id="8EC46052FFB03B031462F890FD59F9C1" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B031462F890FD59F9C1" box="[599,749,1534,1558]" country="Thailand" name="Ha Tay Prov." pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Ha Tay Prov.</location>
,
<collectorName id="26EE535FFFB03B031644FB70FF01F9E1" box="[113,181,1565,1589]" pageId="13" pageNumber="384">Ba Vi</collectorName>
,
<collectionCode id="ED0AAE4CFFB03B0316F7FB70FF61F9E1" box="[194,213,1566,1589]" name="Tavera, Department of Geology and Geophysics" pageId="13" pageNumber="384">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB03B0316D4FB73FDF3F9E1" box="[225,583,1565,1589]" collectingDate="2000-07-29" collectionCode="TUAK" country="Thailand" location="Ishikawa" pageId="13" pageNumber="384" specimenCount="1" stateProvince="Nakhon Ratchasima" typeStatus="paratype">
<location id="8EC46052FFB03B0316D4FB73FEF0F9E1" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B0316EBFED3FA88F921:8EC46052FFB03B0316D4FB73FEF0F9E1" box="[225,324,1565,1589]" country="Thailand" name="Ishikawa" pageId="13" pageNumber="384" stateProvince="Nakhon Ratchasima">Ishikawa</location>
,
<date id="FFA51049FFB03B031764FB73FE6AF9E1" box="[337,478,1565,1589]" pageId="13" pageNumber="384" value="2000-07-29">
<collectingDate id="EFE1E9A1FFB03B031764FB73FE6AF9E1" box="[337,478,1565,1589]" pageId="13" pageNumber="384" value="2000-07-29">29 July 2000</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB03B0317DAFB73FD8BF9E1" box="[495,575,1565,1589]" pageId="13" pageNumber="384">TUAK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644FB33FD45F8E1" pageId="13" pageNumber="384" type="diagnosis">
<paragraph id="8BA43689FFB03B031644FB33FD45F8E1" blockId="13.[113,753,1629,1845]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FB33FF53F9A1" bold="true" box="[113,231,1629,1653]" pageId="13" pageNumber="384">Diagnosis.</emphasis>
This new species is recognised readily by the following characters: Pale reddish brown (male)/ ivory brown (female) general colouration (
<figureCitation id="13202A0CFFB03B0317CDFBF3FDDFF961" box="[504,619,1693,1718]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7A, B</figureCitation>
); small size; darkened apex of clypeus; annulated antennomere I; dark, short stripes along veins on hemelytron; circular spot on mesial cuneus; and semi-circular dorsal structure and narrow, less-spinulate, subtriangular interramal lobe on posterior wall.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644FA33FBD8FB01" pageId="13" pageNumber="384" type="description">
<paragraph id="8BA43689FFB03B031644FA33FBB8FB41" blockId="13.[113,753,1885,2101]" lastBlockId="13.[813,1452,188,1173]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FA33FF4BF8A1" bold="true" box="[113,255,1885,1909]" pageId="13" pageNumber="384">Description.</emphasis>
Body generally pale reddish brown (male)/ ivory brown (female) (
<figureCitation id="13202A0CFFB03B031742FA13FE42F841" box="[375,502,1917,1942]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7A, B</figureCitation>
), ovoid, small; dorsal surface shining, impunctate. Head shiny pale brown, with sparsely distributed, semi-erect setae; apical half of clypeus fuscous; vertex with basal transverse carina about as thick as pronotal collar (
<figureCitation id="13202A0CFFB03B0317A3FA93FE4DF7C1" box="[406,505,2045,2069]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">Fig. 23B</figureCitation>
). Antenna dark brown; antennomere I pale, with dark apex, sometimes weakly darkened at middle; antennomere II with pale basal ⅓ and more or less darkened base, longer than labium; extreme base of antennomere III creamy yellow. Labium shiny pale brown, reaching but not exceeding apex of metacoxa; apical half of segment IV reddish brown. Pronotum shining, uniformly pale brown, impunctate, with sparsely distributed, short, reclining setae (
<figureCitation id="13202A0CFFB03B031210FC13FB1AFE41" box="[1061,1198,381,405]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">Fig. 23A, B</figureCitation>
); calli sometimes with a pair of dark spots; pleura pale brown; metathoracic scent efferent system creamy yellow, with small, semi-circular peritreme (
<figureCitation id="13202A0CFFB03B031593FCB3FBBAFE21" box="[934,1038,477,501]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">Fig. 23D</figureCitation>
); scutellum polished, with dark apex and a dark, narrow, mesal stripe in male (
<figureCitation id="13202A0CFFB03B0312DDFC93FA88FDC1" box="[1256,1340,509,533]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7A</figureCitation>
) or brown faint line in female (
<figureCitation id="13202A0CFFB03B031226FF73FBDDFDE1" box="[1043,1129,541,565]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7B</figureCitation>
), impunctate, with uniformly distributed, short, semi-erect setae. Hemelytron shining, with unique striped pattern as in
<figureCitation id="13202A0CFFB03B031261FF33FB06FDA1" box="[1108,1202,605,629]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="13" pageNumber="384">Figs. 1G</figureCitation>
,
<figureCitation id="13202A0CFFB03B03128BFF33FAB4FDA1" box="[1214,1280,605,629]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="13" pageNumber="384">5A, B</figureCitation>
, with uniformly distributed, semi-erect or reclining setae (
<figureCitation id="13202A0CFFB03B0312E3FF13FAE5FD41" box="[1238,1361,637,661]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">Fig. 23A, E</figureCitation>
); cuneus usually with a dark, circular spot; membrane smoky brown. All coxae and legs pale brown; trochanters brown; each tarsomere III dark brown; pretarsal structures as in
<figureCitation id="13202A0CFFB03B031373FFB3FA13FD21" box="[1350,1447,733,757]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="13" pageNumber="384">Fig. 19G</figureCitation>
; parempodia relatively long, lanceolate. Abdomen pale brown, partly infuscate. Male genitalia (
<figureCitation id="13202A0CFFB03B0312BEFE73FB67FCE1" box="[1163,1235,797,821]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Figs. 3</figureCitation>
AD, 23HK): Left paramere stout, C-shaped, with thick hypophysis (
<figureCitation id="13202A0CFFB03B03137CFE53FA13FC81" box="[1353,1447,829,853]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Figs. 3B</figureCitation>
,
<figureCitation id="13202A0CFFB03B031518FE33FCE0FCA1" box="[813,852,861,885]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">23I</figureCitation>
); right paramere with short, tapered hypophysis (
<figureCitation id="13202A0CFFB03B031360FE33FA13FCA1" box="[1365,1447,861,885]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Fig. 3C</figureCitation>
,
<figureCitation id="13202A0CFFB03B031518FE13FCE2FC41" box="[813,854,893,917]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">23J</figureCitation>
). Vesica with two long, stout, weakly curved spicules (
<figureCitation id="13202A0CFFB03B031500FEF3FC24FC61" box="[821,912,925,949]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Fig. 3D</figureCitation>
,
<figureCitation id="13202A0CFFB03B031595FEF3FC67FC61" box="[928,979,925,949]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">23K</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFFB03B03128CFEF3FA89FC61" box="[1209,1341,925,949]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Figs. 3E, F</figureCitation>
,
<figureCitation id="13202A0CFFB03B031378FEF3FA14FC61" box="[1357,1440,925,949]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">23LO</figureCitation>
): Genital chamber narrow, with elongate-oval, thin-rimmed sclerotised rings that are relatively contiguous to each other mesally (
<figureCitation id="13202A0CFFB03B0315B8FE93FC6AFBC1" box="[909,990,1021,1045]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Fig. 3E</figureCitation>
); posterior wall simple, with a row of spines along posterior margin of interramal lobes and anterior interramal sclerite (
<figureCitation id="13202A0CFFB03B0315CFF953FBE0FB81" box="[1018,1108,1085,1109]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="13" pageNumber="384">Figs. 3F</figureCitation>
,
<figureCitation id="13202A0CFFB03B03126BF953FB1AFB81" box="[1118,1198,1085,1109]" captionStart="Fig" captionStartId="35.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@35.[113,1451,251,1929]" captionTargetPageId="35" captionText="Fig. 23. Scanning electron micrographs of Chelyolygus wolskiorum, new species. A, Dorsal habitus; B, head and anterior pronotum, dorsal view; C, head and thorax, left lateral view; D, thoracic pleura, left lateral view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; G, pretarsal structure of metaleg; H, apex of pygophore, dorsal view; I, left paramere; J, apical part of right paramere; K, vesica; L, genital chamber; M, apex of ovipositor (gonapophysis I); N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683006" httpUri="https://zenodo.org/record/14683006/files/figure.png" pageId="13" pageNumber="384">23N, O</figureCitation>
); dorsal structure small, sack-like; interramal lobes narrow, subtriangular, situated near lateral margins.
</paragraph>
<paragraph id="8BA43689FFB03B031518F9D3FBD8FB01" blockId="13.[813,1132,1213,1237]" box="[813,1132,1213,1237]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518F9D3FC6FFB01" bold="true" box="[813,987,1213,1237]" pageId="13" pageNumber="384">Measurements.</emphasis>
See
<tableCitation id="C6990332FFB03B031226F9D3FBD2FB01" box="[1043,1126,1213,1237]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="13" pageNumber="384">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031518F993FC32FA61" pageId="13" pageNumber="384" type="etymology">
<paragraph id="8BA43689FFB03B031518F993FC32FA61" blockId="13.[813,1452,1277,1461]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518F993FC1BFAC1" bold="true" box="[813,943,1277,1301]" pageId="13" pageNumber="384">Etymology.</emphasis>
Named after my Polish colleague and friend, Andrzej Wolski (who regrettably passed away in
<date id="FFA51049FFB03B031302F870FA10FAE1" box="[1335,1444,1309,1334]" pageId="13" pageNumber="384" value="2024-05">May 2024</date>
) and his wife Tamara; I was able to collect much material and confirm the immature forms of this new species while performing fieldwork in central
<collectingCountry id="F30C7619FFB03B0312BCF813FB5CFA41" box="[1161,1256,1405,1429]" name="Thailand" pageId="13" pageNumber="384">Thailand</collectingCountry>
with Andrzej and Tamara.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031518F8B3FC48F9C1" pageId="13" pageNumber="384" type="distribution">
<paragraph id="8BA43689FFB03B031518F8B3FC48F9C1" blockId="13.[813,1452,1501,1557]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518F8B3FC0BFA21" bold="true" box="[813,959,1501,1525]" pageId="13" pageNumber="384">Distribution.</emphasis>
<collectingCountry id="F30C7619FFB03B0315F3F8B3FB91FA21" box="[966,1061,1501,1525]" name="Thailand" pageId="13" pageNumber="384">Thailand</collectingCountry>
(
<collectingRegion id="49DFF86BFFB03B031201F8B3FB0CFA21" box="[1076,1208,1501,1525]" country="Thailand" name="Chiang Mai" pageId="13" pageNumber="384">Chiang Mai</collectingRegion>
,
<collectingRegion id="49DFF86BFFB03B0312F1F8B3FA15FA21" box="[1220,1441,1501,1525]" country="Thailand" name="Nakhon Ratchasima" pageId="13" pageNumber="384">Nakhon Ratchasima</collectingRegion>
),
<collectingCountry id="F30C7619FFB03B031518F893FC3DF9C1" box="[813,905,1533,1557]" name="Vietnam" pageId="13" pageNumber="384">Vietnam</collectingCountry>
(Ha Tay).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031518FB53FA88F921" pageId="13" pageNumber="384" type="biology_ecology">
<paragraph id="8BA43689FFB03B031518FB53FA88F921" blockId="13.[813,1452,1597,1781]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518FB53FC3EF981" bold="true" box="[813,906,1597,1621]" pageId="13" pageNumber="384">Biology.</emphasis>
A female adult (
<figureCitation id="13202A0CFFB03B03120BFB53FB24F981" box="[1086,1168,1597,1621]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7B</figureCitation>
) and final instar immature form (
<figureCitation id="13202A0CFFB03B031558FB33FC09F9A1" box="[877,957,1629,1653]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="13" pageNumber="384">Fig. 7C</figureCitation>
, successfully developing into an adult female in captivity) were captured by sweep-netting the inflorescences of an undetermined evergreen broadleaf tree (possibly
<taxonomicName id="4C1B4D0AFFB03B031518FBD3FC5DF901" box="[813,1001,1725,1749]" class="Magnoliopsida" family="Dipterocarpaceae" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="family">Dipterocarpaceae</taxonomicName>
). Some adults (including the
<typeStatus id="54A0882BFFB03B031325FBD3FADFF901" box="[1296,1387,1725,1749]" pageId="13" pageNumber="384" type="holotype">holotype</typeStatus>
male) were occasionally attracted to UV light at night.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,289 @@
<document id="9DB1D6D2F54528BB1EBCBB09A8C5A270" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB03B1C15E5FA73FE9DFE01" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Euphorbialygus Yasunaga, 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="389" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="384" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="92E34181256D31A1924D0DFC7378B713" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07952962434CD5F52B5C4612ECC493E8">
<mods:title id="0B1E09BC19A07BF5BE503C2CD3B58311">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="5351CC2C2C1F4281EE973DCEE486F948" type="personal">
<mods:role id="80B6A5B50FFF51747EC6D644FB4F8D21">
<mods:roleTerm id="C3B2BB3C5DB47189FAF919CCB90BB8AA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EE0C733A5B60BC7FB7A3B2B2EA146D19">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="23B990A918E6F6337A294A77DAB1047D">text</mods:typeOfResource>
<mods:relatedItem id="2CDBF16137B7A7D2A4014B6C1B50E9A4" type="host">
<mods:titleInfo id="E257107A01A16670643A6165F37D62A7">
<mods:title id="4D743B64C354C694468F9258E7C646CC">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="DA62BEB998645E8E727B421C44000CB5">
<mods:date id="6CA62E1EF7EAE1A90FF1F69E90BB8A09">2024</mods:date>
<mods:detail id="3BB850995A1CCE8B92A73C3037C4F20B" type="pubDate">
<mods:number id="1D6129314075635020159BDA610E1CD3">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="4A9D09D3C34B8A37DF9D2FC63E5AE515" type="volume">
<mods:number id="BB0C9CC588EA39FAC77E0432917376B4">72</mods:number>
</mods:detail>
<mods:extent id="EA46571AB38B550C64801D8135F0BC7F" unit="page">
<mods:start id="3989D4F9E4C56B6E2D82AE54DCAEB7E5">371</mods:start>
<mods:end id="279F36B5EE59E2D12F4FC2CCF4867172">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="47F5B3ED2960F12D40766852F0F55D69">journal article</mods:classification>
<mods:identifier id="393D4045B646860777EE238DBCB083D1" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="DB154B751928B71954DA58672B5C77B3" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A98B5CA5BF6A850BE0AA223AAAED2D80" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="747E876EFCA2D566186EFC9C4A550CC6" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB03B1C15E5FA73FE9DFE01" LSID="urn:lsid:plazi:treatment:03B2879FFFB03B1C15E5FA73FE9DFE01" httpUri="http://treatment.plazi.org/id/03B2879FFFB03B1C15E5FA73FE9DFE01" lastPageId="18" lastPageNumber="389" pageId="13" pageNumber="384">
<subSubSection id="C3016502FFB03B0315E5FA73FABCF8E1" box="[976,1288,1821,1845]" pageId="13" pageNumber="384" type="nomenclature">
<paragraph id="8BA43689FFB03B0315E5FA73FABCF8E1" blockId="13.[976,1288,1821,1845]" box="[976,1288,1821,1845]" pageId="13" pageNumber="384">
<heading id="D0EC81E5FFB03B0315E5FA73FABCF8E1" bold="true" box="[976,1288,1821,1845]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="384" reason="2">
<emphasis id="B96FEA9BFFB03B0315E5FA73FABCF8E1" bold="true" box="[976,1288,1821,1845]" pageId="13" pageNumber="384">
<taxonomicName id="4C1B4D0AFFB03B0315E5FA73FB34F8E1" authorityName="Yasunaga" authorityYear="2024" box="[976,1152,1821,1845]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFB03B0315E5FA73FB34F8E1" bold="true" box="[976,1152,1821,1845]" italics="true" pageId="13" pageNumber="384">Euphorbialygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB03B0312A5FA73FABCF8E1" box="[1168,1288,1821,1845]" pageId="13" pageNumber="384" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031518FA33FACAF8A1" box="[813,1406,1885,1909]" pageId="13" pageNumber="384" type="type_taxon">
<paragraph id="8BA43689FFB03B031518FA33FACAF8A1" blockId="13.[813,1406,1885,1909]" box="[813,1406,1885,1909]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518FA33FC71F8A1" bold="true" box="[813,965,1885,1909]" pageId="13" pageNumber="384">
<typeStatus id="54A0882BFFB03B031518FA33FCD3F8A1" box="[813,871,1885,1909]" pageId="13" pageNumber="384">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1B4D0AFFB03B0315F8FA33FB51F8A1" authorityName="Yasunaga" authorityYear="2024" box="[973,1253,1885,1909]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFB03B0315F8FA33FB51F8A1" box="[973,1253,1885,1909]" italics="true" pageId="13" pageNumber="384">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB03B0312C1FA30FACDF8A1" box="[1268,1401,1885,1909]" pageId="13" pageNumber="384" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B001518FAF3FA76F861" lastPageId="14" lastPageNumber="385" pageId="13" pageNumber="384" type="diagnosis">
<paragraph id="8BA43689FFB03B001518FAF3FA76F861" blockId="13.[813,1452,1949,2101]" lastBlockId="14.[835,1474,1821,1973]" lastPageId="14" lastPageNumber="385" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031518FAF3FC1EF861" bold="true" box="[813,938,1949,1973]" pageId="13" pageNumber="384">Diagnosis.</emphasis>
<taxonomicName id="4C1B4D0AFFB03B031583FAF3FBD9F861" authorityName="Yasunaga" authorityYear="2024" box="[950,1133,1949,1973]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFB03B031583FAF3FBD9F861" box="[950,1133,1949,1973]" italics="true" pageId="13" pageNumber="384">Euphorbialygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB03B0312B5FAF0FAB4F862" box="[1152,1280,1950,1974]" pageId="13" pageNumber="384" rank="genus">new genus</taxonomicNameLabel>
, is recognised primarily by the following characters: Ovoid body; castaneous or red-brown basic colouration; reduced basal transverse carina on vertex; dense silvery, lanceolate setae mixed with pale, simple setae on scutellum and hemelytra; rather matte hemelytra with darker spot at median to lateral parts of cuneus; and unique male and female genitalic structures as described below. Most similar in external appearance to
<taxonomicName id="4C1B4D0AFFB33B0016BDFA13FDD3F841" authority="Yasunaga, Schwartz &amp; Cherot" authorityName="Yasunaga, Schwartz &amp; Cherot" authorityYear="2002" box="[136,615,1917,1941]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFB33B0016BDFA13FEB8F841" box="[136,268,1917,1941]" italics="true" pageId="14" pageNumber="385">Apolygopsis</emphasis>
Yasunaga, Schwartz &amp; Chérot
</taxonomicName>
and
<emphasis id="B96FEA9BFFB33B001496FA13FCB3F841" box="[675,775,1917,1941]" italics="true" pageId="14" pageNumber="385">Apolygus</emphasis>
<bibRefCitation id="EF8A4B78FFB33B0016BDFAF3FEB9F861" author="China WE" box="[136,269,1949,1973]" pageId="14" pageNumber="385" refId="ref23774" refString="China WE (1941) A new subgeneric name for Lygus Reuter 1875 nec Hahn 1833 (Hemipt. Heteropt.). Proceedings of the Royal Entomological Society of London (B), 10: 60." type="journal volume" year="1941">
<collectingCountry id="F30C7619FFB33B0016BDFAF3FF7EF861" box="[136,202,1949,1973]" name="China" pageId="14" pageNumber="385">China</collectingCountry>
, 1941
</bibRefCitation>
; the present new genus can be distinguished by the following combination of characters: Basal transverse carina on vertex obliterated or slightly keeled; scutellum and hemelytron with densely distributed, silvery, lanceolate setae mixed with pale, simple setae; pale brown tibial spines; short, duck head shaped hypophysis of left paramere; tiny hypophysis of right paramere; three distinct long spicules on vesica; ovoid sclerotised rings that are mesally separated to each other; and simple form of posterior wall anteriorly with a pair of small, pointed processes between interramal lobes..
</paragraph>
</subSubSection>
<caption id="DF646601FFB33B0016B2FB32FDE3F931" ID-DOI="http://doi.org/10.5281/zenodo.14682964" ID-Zenodo-Dep="14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="14" pageNumber="385" startId="14.[135,170,1628,1650]" targetBox="[136,1474,189,1613]" targetPageId="14" targetType="figure">
<paragraph id="8BA43689FFB33B0016B2FB32FDE3F931" blockId="14.[135,1474,1628,1765]" pageId="14" pageNumber="385">
Fig. 7. Habitus images and habitats (J, L) of
<taxonomicName id="4C1B4D0AFFB33B001406FB32FC94F9A5" authorityName="Yasunaga" authorityYear="2024" box="[563,800,1628,1649]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="wolskiorum" status="sp. nov.">
<emphasis id="B96FEA9BFFB33B001406FB32FC94F9A5" box="[563,800,1628,1649]" italics="true" pageId="14" pageNumber="385">Chelyolygus wolskiorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B001519FB33FC2BF9A6" box="[812,927,1628,1650]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
(AC),
<taxonomicName id="4C1B4D0AFFB33B0015DAFB33FBD7F9A6" authorityName="Yasunaga" authorityYear="2024" box="[1007,1123,1629,1650]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFFB33B0015DAFB33FBD7F9A6" box="[1007,1123,1629,1650]" italics="true" pageId="14" pageNumber="385">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B00125AFB33FB56F9A6" box="[1135,1250,1628,1650]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
(D),
<taxonomicName id="4C1B4D0AFFB33B001323FB32FF6CF95A" authorityName="Yasunaga" authorityYear="2024" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFFB33B001323FB32FF6CF95A" italics="true" pageId="14" pageNumber="385">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B0016D2FB14FEE9F95B" box="[231,349,1657,1679]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
(E, F),
<taxonomicName id="4C1B4D0AFFB33B00179AFB17FDD8F95A" authorityName="Yasunaga" authorityYear="2024" box="[431,620,1657,1678]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFFB33B00179AFB17FDD8F95A" box="[431,620,1657,1678]" italics="true" pageId="14" pageNumber="385">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B00144EFB14FD44F95B" box="[635,752,1657,1679]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
(G, H) and
<taxonomicName id="4C1B4D0AFFB33B00155AFB17FC79F95B" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[879,973,1657,1679]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFFB33B00155AFB17FC79F95B" box="[879,973,1657,1679]" italics="true" pageId="14" pageNumber="385">S. sauteri</emphasis>
</taxonomicName>
(K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with
<taxonomicName id="4C1B4D0AFFB33B0015FDFBF9FBFEF97F" box="[968,1098,1686,1708]" class="Magnoliopsida" family="Urticaceae" genus="Urtica" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="385" phylum="Tracheophyta" rank="species" species="dioica">
<emphasis id="B96FEA9BFFB33B0015FDFBF9FBFEF97F" box="[968,1098,1686,1708]" italics="true" pageId="14" pageNumber="385">Urtica dioica</emphasis>
</taxonomicName>
, on which
<taxonomicName id="4C1B4D0AFFB33B001282FBF8FA0FF97F" authorityName="Yasunaga" authorityYear="2024" box="[1207,1467,1686,1707]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="duwalae" status="sp. nov.">
<emphasis id="B96FEA9BFFB33B001282FBF8FA0FF97F" box="[1207,1467,1686,1707]" italics="true" pageId="14" pageNumber="385">Himalisundacoris duwalae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B0016B2FBDDFEB4F91D" box="[135,256,1715,1737]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="4C1B4D0AFFB33B00170DFBDDFE41F91C" authorityName="Yasunaga" authorityYear="2024" box="[312,501,1715,1736]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="schuhi" status="sp. nov.">
<emphasis id="B96FEA9BFFB33B00170DFBDDFE41F91C" box="[312,501,1715,1736]" italics="true" pageId="14" pageNumber="385">Sabactiopus schuhi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB33B001431FBDDFDC9F91D" box="[516,637,1715,1737]" pageId="14" pageNumber="385" rank="species">new species</taxonomicNameLabel>
, coexist (Lalitpur, Nepal); L,
<taxonomicName id="4C1B4D0AFFB33B00159CFBDDFBEBF91C" box="[937,1119,1715,1736]" class="Magnoliopsida" family="Styracaceae" genus="Styrax" kingdom="Plantae" order="Ericales" pageId="14" pageNumber="385" phylum="Tracheophyta" rank="species" species="formosanus">
<emphasis id="B96FEA9BFFB33B00159CFBDDFBEBF91C" box="[937,1119,1715,1736]" italics="true" pageId="14" pageNumber="385">Styrax formosanus</emphasis>
</taxonomicName>
inflorescences, one of the breeding hosts of
<taxonomicName id="4C1B4D0AFFB33B0016E9FBA1FE28F931" baseAuthorityName="Poppius" baseAuthorityYear="1912" box="[220,412,1743,1765]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="385" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="B96FEA9BFFB33B0016E9FBA1FE28F931" box="[220,412,1743,1765]" italics="true" pageId="14" pageNumber="385">Sabactiopus sauteri</emphasis>
</taxonomicName>
(Nantou, Taiwan).
</paragraph>
</caption>
<subSubSection id="C3016502FFB33B1F1576FAB2FDAAF79D" lastPageId="17" lastPageNumber="388" pageId="14" pageNumber="385" type="description">
<paragraph id="8BA43689FFB33B1F1576FAB2FDAAF79D" blockId="14.[835,1474,2012,2101]" lastBlockId="17.[113,753,1809,2121]" lastPageId="17" lastPageNumber="388" pageId="14" pageNumber="385">
<emphasis id="B96FEA9BFFB33B001576FAB2FC63F820" bold="true" box="[835,983,2012,2036]" pageId="14" pageNumber="385">Description.</emphasis>
Body oval, relatively small (
<quantity id="4CE39B6CFFB33B001300FAB3FA75F820" box="[1333,1473,2013,2037]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="14" pageNumber="385" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.24.2 mm</quantity>
in total length); general colouration reddish brown or castaneous; dorsal surface weakly shining, partly matte, with uniformly distributed, pale, simple, reclining setae and densely distributed, silvery, lanceolate setae (
<figureCitation id="13202A0CFFAD3B1E14FBFBF9FF01F904" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="16" pageNumber="387">Figs. 24E</figureCitation>
,
<figureCitation id="13202A0CFFAD3B1E16F4FBD9FF44F91B" box="[193,240,1719,1743]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="16" pageNumber="387">25E</figureCitation>
); significant sexual dimorphism not recognisable.
<emphasis id="B96FEA9BFFAD3B1E16BDFBB6FF7AF924" bold="true" box="[136,206,1752,1776]" pageId="16" pageNumber="387">Head:</emphasis>
Vertical (
<figureCitation id="13202A0CFFAD3B1E170DFBB9FE17F924" box="[312,419,1751,1776]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="16" pageNumber="387">Figs. 24B</figureCitation>
,
<figureCitation id="13202A0CFFAD3B1E179BFBB9FE54F93B" box="[430,480,1751,1775]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="16" pageNumber="387">25A</figureCitation>
); eyes large; vertex narrow, weakly concaved mesially, lacking basal transverse carina; clypeus almost flat.
<emphasis id="B96FEA9BFFAD3B1E176FFA79FE71F8FB" bold="true" box="[346,453,1815,1839]" pageId="16" pageNumber="387">Antenna:</emphasis>
Almost linear, slightly shorter than body, generally slender, not incrassate; antennomere I reddish brown, with darkened basal half; antennomeres IIIV dark brown; antennomere II with pale reddish basal half and darkened extreme base, slightly longer (male)/ shorter (female) than basal width of pronotum; antennomeres III and IV filiform.
<emphasis id="B96FEA9BFFAD3B1E1725FAB9FEC0F83B" bold="true" box="[272,372,2007,2031]" pageId="16" pageNumber="387">Labium:</emphasis>
Relatively slender, reaching but not exceeding apex of metacoxa, slightly shorter than metafemur.
<emphasis id="B96FEA9BFFAD3B1E16BDF576FF51F7E4" bold="true" box="[136,229,2072,2096]" pageId="16" pageNumber="387">Thorax:</emphasis>
Pronotum shining, with uniformly distributed, pale, simple, reclining setae, sparsely and shallowly punctate, without silvery setae; calli weakly demarcated; collar flat, about as thick as antennomere III, with sparse, short, upright setae; scutellum pale brown, shallowly and transversely rugose; pleura pale brown; metathoracic scent efferent system creamy yellow, subtriangular, with narrow, somewhat convex peritreme (
<figureCitation id="13202A0CFFAD3B1E1224FA59FBCBF884" box="[1041,1151,1847,1872]" captionStart="Fig" captionStartId="36.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1929]" captionTargetId="figure-7@36.[135,1473,251,1929]" captionTargetPageId="36" captionText="Fig. 24. Scanning electron micrographs of Euphorbialygus nanglong, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, left lateral view; C, head, dorsal view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, pygophore, left lateral view; G, metatarsus; H, pretarsal structure of metaleg; I, vesica; J, left paramere; K, right paramere; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683010" httpUri="https://zenodo.org/record/14683010/files/figure.png" pageId="16" pageNumber="387">Figs. 24C</figureCitation>
,
<figureCitation id="13202A0CFFAD3B1E12B8FA59FB0BF89B" box="[1165,1215,1847,1871]" captionStart="Fig" captionStartId="38.[136,171,1944,1966]" captionTargetBox="[136,1474,251,1929]" captionTargetId="figure-7@38.[136,1474,251,1929]" captionTargetPageId="38" captionText="Fig. 25. Scanning electron micrographs of Euphorbialygus rufobrunneus, new species. A, head and thorax, left lateral view; B, head and anterior pronotum, dorsal view; C, metathoracic scent efferent system; D, anterior body, dorsal view; E, scutellum, clavus and corium, dorsal view; F, metatarsus; H, pretarsal structure of metaleg; H, pygophore, dorsal view; I, left paramere; J, K, vesica; L, genital chamber and ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14683012" httpUri="https://zenodo.org/record/14683012/files/figure.png" pageId="16" pageNumber="387">25G</figureCitation>
).
<emphasis id="B96FEA9BFFAD3B1E12E1FA56FAD2F884" bold="true" box="[1236,1382,1848,1872]" pageId="16" pageNumber="387">Hemelytron:</emphasis>
Weakly shining, matte, with uniformly distributed, minute, circular punctures and with both pale simple setae and silvery setae; cuneus more or less suffused with red.
<emphasis id="B96FEA9BFFAD3B1E12C2FAF6FA82F864" bold="true" box="[1271,1334,1944,1968]" pageId="16" pageNumber="387">Legs:</emphasis>
Pale brown; each femur with several obscure annulations; tibial spines pale reddish brown, rather short; meta-tarsomere I about half as long as II or III; parempodia lanceolate.
<emphasis id="B96FEA9BFFAD3B1E1323FA96FA76F7DB" bold="true" box="[1302,1474,2039,2064]" pageId="16" pageNumber="387">Male genitalia:</emphasis>
Left paramere L-shaped, with developed, with elongate hypophysis terminated in apical hook; right paramere almost straight, with tiny hypophysis; vesica with two spicules and a flagellate lobal-sclerite; seminal duct weakly inflated apically; secondary gonopore circular; phallotheca smooth.
<emphasis id="B96FEA9BFFAC3B1F14A8FA1FFF60F87D" bold="true" pageId="17" pageNumber="388">Female genitalia</emphasis>
: Sclerotised rings oval, thick-rimmed, separated from each other mesially; posterior wall relatively simple in form, anteriorly with a pair of small, pointed processes between interramal lobes, lacking dorsal structure nor lateral lobe; interramal lobe narrow, with sparsely distributed, scaly microstructures along posterior margin.
</paragraph>
</subSubSection>
<caption id="DF646601FFB23B011644FAB7FEE8F7FD" ID-DOI="http://doi.org/10.5281/zenodo.14682966" ID-Zenodo-Dep="14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="15" pageNumber="386" startId="15.[113,148,2009,2031]" targetBox="[113,1451,215,1995]" targetPageId="15" targetType="figure">
<paragraph id="8BA43689FFB23B011644FAB7FEE8F7FD" blockId="15.[113,1452,2009,2089]" pageId="15" pageNumber="386">
Fig. 8. Habitus images and habitats (G, H) of
<taxonomicName id="4C1B4D0AFFB23B011474FAB7FC8AF83A" authorityName="Yasunaga" authorityYear="2024" box="[577,830,2009,2030]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="386" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFB23B011474FAB7FC8AF83A" box="[577,830,2009,2030]" italics="true" pageId="15" pageNumber="386">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB23B011579FAB4FC76F83B" box="[844,962,2009,2031]" pageId="15" pageNumber="386" rank="species">new species</taxonomicNameLabel>
from Nakhon Nayok, Thailand (AD, G) and
<taxonomicName id="4C1B4D0AFFB23B0113A2FAB4FF46F7DF" authorityName="Yasunaga" authorityYear="2024" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="15" pageNumber="386" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFB23B0113A2FAB4FF46F7DF" italics="true" pageId="15" pageNumber="386">E. rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB23B0116CAFA99FEC7F7D8" box="[255,371,2038,2060]" pageId="15" pageNumber="386" rank="species">new species</taxonomicNameLabel>
from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G,
<taxonomicName id="4C1B4D0AFFB23B011373FA99FF01F7FD" class="Magnoliopsida" family="Euphorbiaceae" genus="Homonoia" kingdom="Plantae" order="Malpighiales" pageId="15" pageNumber="386" phylum="Tracheophyta" rank="species" species="riparia">
<emphasis id="B96FEA9BFFB23B011373FA99FF01F7FD" italics="true" pageId="15" pageNumber="386">Homonoia riparia</emphasis>
</taxonomicName>
; H,
<taxonomicName id="4C1B4D0AFFB23B0116D6F57DFEE8F7FD" box="[227,348,2067,2089]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="15" pageNumber="386" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96FEA9BFFB23B0116D6F57DFE8CF7FC" box="[227,312,2067,2088]" italics="true" pageId="15" pageNumber="386">Mallotus</emphasis>
sp.
</taxonomicName>
</paragraph>
</caption>
<caption id="DF646601FFAD3B1E16BDFB60FD70F994" ID-DOI="http://doi.org/10.5281/zenodo.14682970" ID-Zenodo-Dep="14682970" httpUri="https://zenodo.org/record/14682970/files/figure.png" pageId="16" pageNumber="387" startId="16.[136,171,1550,1572]" targetBox="[135,1473,189,1534]" targetPageId="16" targetType="figure">
<paragraph id="8BA43689FFAD3B1E16BDFB60FD70F994" blockId="16.[136,1475,1550,1600]" pageId="16" pageNumber="387">
Fig. 9. Male genitalia of
<taxonomicName id="4C1B4D0AFFAD3B1E174BFB60FDCEF9F7" authorityName="Yasunaga" authorityYear="2024" box="[382,634,1550,1571]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="387" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFAD3B1E174BFB60FDCEF9F7" box="[382,634,1550,1571]" italics="true" pageId="16" pageNumber="387">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAD3B1E14BDFB60FD48F9F0" box="[648,764,1550,1572]" pageId="16" pageNumber="387" rank="species">new species</taxonomicNameLabel>
(AE) and
<taxonomicName id="4C1B4D0AFFAD3B1E1547FB60FBA4F9F7" authorityName="Yasunaga" authorityYear="2024" box="[882,1040,1550,1571]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="16" pageNumber="387" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFAD3B1E1547FB60FBA4F9F7" box="[882,1040,1550,1571]" italics="true" pageId="16" pageNumber="387">E. rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAD3B1E122BFB60FB26F9F0" box="[1054,1170,1550,1572]" pageId="16" pageNumber="387" rank="species">new species</taxonomicNameLabel>
(FH). A, B, F, left paramere; C, D, G, right paramere; E, H, vesica. Scale bars 0.2 mm.
</paragraph>
</caption>
<caption id="DF646601FFAC3B1F1644FBE4FC0BF90D" ID-DOI="http://doi.org/10.5281/zenodo.14682972" ID-Zenodo-Dep="14682972" httpUri="https://zenodo.org/record/14682972/files/figure.png" pageId="17" pageNumber="388" startId="17.[113,148,1674,1696]" targetBox="[113,1449,189,1659]" targetPageId="17" targetType="figure">
<paragraph id="8BA43689FFAC3B1F1644FBE4FC0BF90D" blockId="17.[113,1452,1674,1754]" pageId="17" pageNumber="388">
Fig. 10. Female genitalia of
<taxonomicName id="4C1B4D0AFFAC3B1F17BFFBE4FD33F94B" authorityName="Yasunaga" authorityYear="2024" box="[394,647,1674,1695]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="17" pageNumber="388" phylum="Tracheophyta" rank="species" species="nanglong" status="sp. nov.">
<emphasis id="B96FEA9BFFAC3B1F17BFFBE4FD33F94B" box="[394,647,1674,1695]" italics="true" pageId="17" pageNumber="388">Euphorbialygus nanglong</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAC3B1F14A1FBE5FCBDF974" box="[660,777,1674,1696]" pageId="17" pageNumber="388" rank="species">new species</taxonomicNameLabel>
(AF) and
<taxonomicName id="4C1B4D0AFFAC3B1F154AFBE5FBA9F94B" authorityName="Yasunaga" authorityYear="2024" box="[895,1053,1674,1696]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="17" pageNumber="388" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFAC3B1F154AFBE5FBA9F94B" box="[895,1053,1674,1696]" italics="true" pageId="17" pageNumber="388">E. rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAC3B1F121EFBE5FB14F974" box="[1067,1184,1674,1696]" pageId="17" pageNumber="388" rank="species">new species</taxonomicNameLabel>
(GI). A, Genital chamber with ovipositors (gonapophyses II), ventral view; C, G, genital chamber, ventral view; D, ditto, dorsal view; E, sclerotised ring; F, I, posterior wall, anterior view; apex of ovipositor (gonapophysis I). Scale bars 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3016502FFAC3B1F1518FA7FFC14F85C" pageId="17" pageNumber="388" type="etymology">
<paragraph id="8BA43689FFAC3B1F1518FA7FFC14F85C" blockId="17.[813,1452,1808,1928]" pageId="17" pageNumber="388">
<emphasis id="B96FEA9BFFAC3B1F1518FA7FFC1BF8FD" bold="true" box="[813,943,1809,1833]" pageId="17" pageNumber="388">Etymology.</emphasis>
Named for the plant associations of the known congeners with
<taxonomicName id="4C1B4D0AFFAC3B1F15DCFA5EFB26F89C" box="[1001,1170,1840,1864]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="17" pageNumber="388" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
broadleaf angiosperms, combined with the mirine generic name
<taxonomicName id="4C1B4D0AFFAC3B1F12E3FA3FFAEDF8BC" authority="Hahn" authorityName="Hahn" authorityYear="1833" box="[1238,1369,1872,1896]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFAC3B1F12E3FA3FFAA2F8BC" box="[1238,1302,1873,1896]" italics="true" pageId="17" pageNumber="388">Lygus</emphasis>
Hahn
</taxonomicName>
; gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFAC3B1C1518FADEFE9DFE01" lastPageId="18" lastPageNumber="389" pageId="17" pageNumber="388" type="discussion">
<paragraph id="8BA43689FFAC3B1C1518FADEFF63FEE2" blockId="17.[813,1451,1968,2120]" lastBlockId="18.[136,775,189,310]" lastPageId="18" lastPageNumber="389" pageId="17" pageNumber="388">
<emphasis id="B96FEA9BFFAC3B1F1518FADEFC18F81C" bold="true" box="[813,940,1968,1992]" pageId="17" pageNumber="388">Discussion.</emphasis>
Judging from the patterns of vesical sclerites,
<taxonomicName id="4C1B4D0AFFAC3B1F1518FABEFC52F83C" authorityName="Yasunaga" authorityYear="2024" box="[813,998,2000,2024]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="17" pageNumber="388" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFAC3B1F1518FABEFC52F83C" box="[813,998,2000,2024]" italics="true" pageId="17" pageNumber="388">Euphorbialygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAC3B1F15CFFABFFBCFF83D" box="[1018,1147,2001,2025]" pageId="17" pageNumber="388" rank="genus">new genus</taxonomicNameLabel>
, appears to be related to
<taxonomicName id="4C1B4D0AFFAC3B1F1518FA9EFAB8F7DC" authority="Yasunaga, Schwartz &amp; Cherot" authorityName="Yasunaga, Schwartz &amp; Cherot" authorityYear="2002" box="[813,1292,2032,2057]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="388" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFAC3B1F1518FA9EFC05F7DC" box="[813,945,2032,2056]" italics="true" pageId="17" pageNumber="388">Apolygopsis</emphasis>
Yasunaga, Schwartz &amp; Chérot
</taxonomicName>
and
<emphasis id="B96FEA9BFFAC3B1F1372FA9EFA1FF7DC" box="[1351,1451,2032,2056]" italics="true" pageId="17" pageNumber="388">Apolygus</emphasis>
<collectingCountry id="F30C7619FFAC3B1F1518F57EFCC7F7FC" box="[813,883,2064,2088]" name="China" pageId="17" pageNumber="388">China</collectingCountry>
. Nonetheless, presence of the silvery, lanceolate setae on the dorsum and the form of the female genitalia (e.g., ovoid, mesally separated sclerotised rings and wider interramal lobes on posterior wall) in
<taxonomicName id="4C1B4D0AFFAF3B1C1417FDB3FD78FF21" authorityName="Yasunaga" authorityYear="2024" box="[546,716,221,245]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFAF3B1C1417FDB3FD78FF21" box="[546,716,221,245]" italics="true" pageId="18" pageNumber="389">Euphorbialygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C14EFFDB0FF7DFEC2" pageId="18" pageNumber="389" rank="genus">new genus</taxonomicNameLabel>
, are evidently different from those of the two known genera.
</paragraph>
<paragraph id="8BA43689FFAF3B1C16BDFC33FE9DFE01" blockId="18.[136,775,349,469]" pageId="18" pageNumber="389">
<taxonomicName id="4C1B4D0AFFAF3B1C16BDFC33FEF5FEA1" authorityName="Yasunaga" authorityYear="2024" box="[136,321,349,373]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFAF3B1C16BDFC33FEF5FEA1" box="[136,321,349,373]" italics="true" pageId="18" pageNumber="389">Euphorbialygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFAF3B1C1760FC30FE62FEA2" box="[341,470,350,374]" pageId="18" pageNumber="389" rank="genus">new genus</taxonomicNameLabel>
, currently comprises two thermophilic members, associated with
<taxonomicName id="4C1B4D0AFFAF3B1C1419FC13FD7EFE41" box="[556,714,381,405]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="18" pageNumber="389" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
hosts and distributed in the southern foothills of the Himalayas and Indochina.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,280 @@
<document id="3946E296A147E5CA59F780FA2962358C" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB53B05158BFC13FF51FD41" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus parsanus Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="382" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="379" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="12A618B9CEED0A6072DDBAF8607168FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32927AD897462BAAF256473F85139772">
<mods:title id="7F0B388E38FF7617C45A34D7EBA52025">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="1F17145875079BBBF5D487DEA89ADE44" type="personal">
<mods:role id="21795A77E197104BB4F550ABD2F9685D">
<mods:roleTerm id="0159D8C9EB9F4BDC661612C6FD97278B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="22F0E9D6989F256C3DDD3014C0CE169D">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="D97AEB9F8457A69D7C8876D3EBA50D9E">text</mods:typeOfResource>
<mods:relatedItem id="A70A48E1E6DDD21C398C634818ADDA75" type="host">
<mods:titleInfo id="2DE991DDA8F3B694A9EA204FEFFB0B92">
<mods:title id="BAB0FFDDADC0D043EEF0D41A55133DD7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="6549250082412B70C64750D01CA0F329">
<mods:date id="867578039401F26BE77BD2EFAB4E736B">2024</mods:date>
<mods:detail id="A690E219EA0C11D57666117F0C31FDC4" type="pubDate">
<mods:number id="19AD73051C176655B52E8699A26F9C67">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="344DE602ED5676F90BAF80BF973E415B" type="volume">
<mods:number id="6333B037817F17E28D97400C8A4E6EE1">72</mods:number>
</mods:detail>
<mods:extent id="7832E7BA9BEC7109637E1C3CD83B927B" unit="page">
<mods:start id="2CE77060B9DCCD30DEC27304FFA765B7">371</mods:start>
<mods:end id="33C681E6224D454E8AF4E85A4B031C42">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2EDFC92936E4D50A431D5482EA44E2E6">journal article</mods:classification>
<mods:identifier id="8B501C756AE4A5192C2F960E14EE3766" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="8F48F9334628608E79546034A243ED77" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DFF8C767C39A3A700A060810C2804938" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="64012DD42BAAE122451B45C3296973E6" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB53B05158BFC13FF51FD41" ID-DOI="http://doi.org/10.5281/zenodo.14682968" ID-Zenodo-Dep="14682968" LSID="urn:lsid:plazi:treatment:03B2879FFFB53B05158BFC13FF51FD41" httpUri="http://treatment.plazi.org/id/03B2879FFFB53B05158BFC13FF51FD41" lastPageId="11" lastPageNumber="382" pageId="8" pageNumber="379">
<subSubSection id="C3016502FFB53B06158BFC13FAF3FE41" box="[958,1351,381,405]" pageId="8" pageNumber="379" type="nomenclature">
<paragraph id="8BA43689FFB53B06158BFC13FAF3FE41" blockId="8.[955,1354,381,437]" box="[958,1351,381,405]" pageId="8" pageNumber="379">
<heading id="D0EC81E5FFB53B06158BFC13FAF3FE41" bold="true" box="[958,1351,381,405]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="379" reason="2">
<emphasis id="B96FEA9BFFB53B06158BFC13FAF3FE41" bold="true" box="[958,1351,381,405]" pageId="8" pageNumber="379">
<taxonomicName id="4C1B4D0AFFB53B06158BFC13FB07FE41" authorityName="Yasunaga" authorityYear="2024" box="[958,1203,381,405]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="parsanus" status="sp. nov.">
<emphasis id="B96FEA9BFFB53B06158BFC13FB07FE41" bold="true" box="[958,1203,381,405]" italics="true" pageId="8" pageNumber="379">Chelyolygus parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB53B0612F7FC13FAF3FE41" box="[1218,1351,381,405]" pageId="8" pageNumber="379" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB53B06158EFCF3FAFEFE61" box="[955,1354,413,437]" pageId="8" pageNumber="379" type="description">
<paragraph id="8BA43689FFB53B06158EFCF3FAFEFE61" blockId="8.[955,1354,381,437]" box="[955,1354,413,437]" pageId="8" pageNumber="379">
(
<figureCitation id="13202A0CFFB53B0615F6FCF3FBBAFE61" box="[963,1038,413,437]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Figs. 4</figureCitation>
FJ, 5EH, 21HO, 22AD)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB53B061576FCB3FB7EFD61" pageId="8" pageNumber="379" type="materials_examined">
<paragraph id="8BA43689FFB53B061576FCB3FB7EFD61" blockId="8.[835,1474,477,693]" pageId="8" pageNumber="379">
<emphasis id="B96FEA9BFFB53B061576FCB3FC47FE21" bold="true" box="[835,1011,477,501]" pageId="8" pageNumber="379">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB53B0615CBFCB3FC0EFD41" collectingDate="2005-11-07" collectingDateMax="2005-11-09" collectingDateMin="2005-11-07" collectionCode="T, NMTU, AMNH" collectorName="Yasunaga, M &amp; B. Shishido" country="Nepal" county="Machan Resort" elevation="270" latitude="27.535" location="Makawanpur Dist." longLatPrecision="20" longitude="84.73805" municipality="Chitwan National Park" pageId="8" pageNumber="379" specimenCode="PBI 00378792" specimenCount="1" typeStatus="holotype">
<emphasis id="B96FEA9BFFB53B0615CBFCB3FBDDFE21" bold="true" box="[1022,1129,477,501]" pageId="8" pageNumber="379">
<typeStatus id="54A0882BFFB53B0615CBFCB3FBDDFE21" box="[1022,1129,477,501]" pageId="8" pageNumber="379" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFB53B0612F7FCB3FA93FE21" bold="true" box="[1218,1319,477,501]" pageId="8" pageNumber="379">
<collectingCountry id="F30C7619FFB53B0612F7FCB3FA96FE21" box="[1218,1314,477,501]" name="Nepal" pageId="8" pageNumber="379">NEPAL</collectingCountry>
:
</emphasis>
<location id="8EC46052FFB53B061307FCB3FCCAFDC1" LSID="urn:lsid:plazi:treatment:03B2879FFFB53B05158BFC13FF51FD41:8EC46052FFB53B061307FCB3FCCAFDC1" country="Nepal" county="Machan Resort" latitude="27.535" longLatPrecision="20" longitude="84.73805" municipality="Chitwan National Park" name="Makawanpur Dist." pageId="8" pageNumber="379">Makawanpur Dist.</location>
:
<collectingMunicipality id="6BC0ACF3FFB53B0615BBFC93FB29FDC1" box="[910,1181,509,533]" pageId="8" pageNumber="379">Chitwan National Park</collectingMunicipality>
,
<collectingCounty id="62C54E05FFB53B061299FC93FAE8FDC1" box="[1196,1372,509,533]" pageId="8" pageNumber="379">Machan Resort</collectingCounty>
[current Parsa Wildlife Reserve],
<geoCoordinate id="EE2F504EFFB53B06125EFF72FB40FDE1" box="[1131,1268,540,565]" degrees="27" direction="north" minutes="32" orientation="latitude" pageId="8" pageNumber="379" precision="15" seconds="06" value="27.535">27°3206″N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB53B061336FF72FA3DFDE1" box="[1283,1417,540,565]" degrees="84" direction="east" minutes="44" orientation="longitude" pageId="8" pageNumber="379" precision="15" seconds="17" value="84.73805">84°4417″E</geoCoordinate>
,
<quantity id="4CE39B6CFFB53B0613ACFF73FCEEFD81" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="379" unit="m" value="270.0">
<elevation id="0036D1BAFFB53B0613ACFF73FCEEFD81" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="379" unit="m" value="270.0">270 m</elevation>
</quantity>
, on flowers of
<taxonomicName id="4C1B4D0AFFB53B06123DFF53FB26FD81" box="[1032,1170,573,597]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="379" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96FEA9BFFB53B06123DFF53FBDCFD81" box="[1032,1128,573,597]" italics="true" pageId="8" pageNumber="379">Mallotus</emphasis>
sp.
</taxonomicName>
,
<collectionCode id="ED0AAE4CFFB53B061294FF50FB00FD81" box="[1185,1204,574,597]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="379">T</collectionCode>
.
<collectorName id="26EE535FFFB53B0612F7FF50FAECFD81" box="[1218,1368,574,598]" pageId="8" pageNumber="379">Yasunaga, M</collectorName>
. Takai &amp;
<collectorName id="26EE535FFFB53B061576FF30FC73FDA1" box="[835,967,605,629]" pageId="8" pageNumber="379">B. Shishido</collectorName>
,
<date id="FFA51049FFB53B0615E3FF33FB0EFDA1" box="[982,1210,605,629]" pageId="8" pageNumber="379" value="2005-11-07" valueMax="2005-11-09" valueMin="2005-11-07">
<collectingDate id="EFE1E9A1FFB53B0615E3FF33FB0EFDA1" box="[982,1210,605,629]" pageId="8" pageNumber="379" value="2005-11-07" valueMax="2005-11-09" valueMin="2005-11-07">79 November 2005</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB53B0612F9FF33FAAAFDA1" box="[1228,1310,605,629]" pageId="8" pageNumber="379">NMTU</collectionCode>
) (
<collectionCode id="ED0AAE4CFFB53B061302FF33FA3FFDA1" box="[1335,1419,605,629]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="379">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB53B0613ADFF33FC07FD41" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="8" pageNumber="379">PBI 00378792</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB53B0615F2FF10FB72FD61" collectingDate="2005-11-07" collectingDateMax="2005-11-09" collectingDateMin="2005-11-07" collectionCode="TYCN, ZRC" collectorName="Yasunaga, M &amp; B. Shishido" country="Nepal" county="Machan Resort" elevation="270" latitude="27.535" location="Makawanpur Dist." longLatPrecision="20" longitude="84.73805" municipality="Chitwan National Park" pageId="8" pageNumber="379" specimenCount="10" specimenCount-female="8" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54A0882BFFB53B0615F2FF10FB87FD42" box="[967,1075,638,662]" pageId="8" pageNumber="379" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B96FEA9BFFB53B06120AFF13FB17FD41" bold="true" box="[1087,1187,637,661]" pageId="8" pageNumber="379">
<collectingCountry id="F30C7619FFB53B06120AFF13FB29FD41" box="[1087,1181,637,661]" name="Nepal" pageId="8" pageNumber="379">NEPAL</collectingCountry>
:
</emphasis>
<specimenCount id="9D1DFD00FFB53B061299FF13FAB0FD41" box="[1196,1284,637,661]" count="2" pageId="8" pageNumber="379" type="male">2 males</specimenCount>
,
<specimenCount id="9D1DFD00FFB53B061324FF13FACAFD41" box="[1297,1406,637,661]" count="8" pageId="8" pageNumber="379" type="female">8 females</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFB53B061219FFF3FBCDFD61" box="[1068,1145,669,693]" pageId="8" pageNumber="379">TYCN</collectionCode>
,
<collectionCode id="ED0AAE4CFFB53B0612B3FFF3FB0BFD61" box="[1158,1215,669,693]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="8" pageNumber="379">ZRC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB53B061576FFB3FB1EFBC1" pageId="8" pageNumber="379" type="diagnosis">
<paragraph id="8BA43689FFB53B061576FFB3FB1EFBC1" blockId="8.[835,1474,733,1045]" pageId="8" pageNumber="379">
<emphasis id="B96FEA9BFFB53B061576FFB3FC09FD21" bold="true" box="[835,957,733,757]" pageId="8" pageNumber="379">Diagnosis.</emphasis>
<taxonomicName id="4C1B4D0AFFB53B0615FDFFB3FB76FD21" authorityName="Yasunaga" authorityYear="2024" box="[968,1218,733,757]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="parsanus" status="sp. nov.">
<emphasis id="B96FEA9BFFB53B0615FDFFB3FB76FD21" box="[968,1218,733,757]" italics="true" pageId="8" pageNumber="379">Chelyolygus parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB53B0612E1FFB0FAEBFD21" box="[1236,1375,733,757]" pageId="8" pageNumber="379" rank="species">new species</taxonomicNameLabel>
, is most closely related to
<taxonomicName id="4C1B4D0AFFB53B0615CFFF90FB15FCC1" authorityName="Yasunaga, Schwartz &amp; Chérot" authorityYear="2018" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[1018,1185,765,789]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="379" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFFB53B0615CFFF90FB15FCC1" box="[1018,1185,765,789]" italics="true" pageId="8" pageNumber="379">C. fuhoshoensis</emphasis>
</taxonomicName>
but can be distinguished by the characters mentioned in the above key and the following characters: Relatively smaller body with shorter antennomere II, labium and metafemur; paler maculae on scutellum and hemelytron; shorter hypophysis of left paramere (
<figureCitation id="13202A0CFFB53B061354FE13FA09FC41" box="[1377,1469,893,917]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Figs. 5F</figureCitation>
,
<figureCitation id="13202A0CFFB53B061576FEF3FCC2FC61" box="[835,886,925,949]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">22A</figureCitation>
); generally shorter vesical spicules (
<figureCitation id="13202A0CFFB53B061324FEF3FAC2FC61" box="[1297,1398,925,949]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Figs. 5H</figureCitation>
,
<figureCitation id="13202A0CFFB53B0613B0FEF3FA01FC61" box="[1413,1461,925,949]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">22B</figureCitation>
); smaller sclerotised rings (
<figureCitation id="13202A0CFFB53B061268FED3FB19FC01" box="[1117,1197,957,981]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Fig. 4J</figureCitation>
); and posterior wall with narrower dorsal structure and sparse spinules on anterior interramal sclerite (
<figureCitation id="13202A0CFFB53B061222FE93FB29FBC1" box="[1047,1181,1021,1045]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">Fig. 22C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB53B051576F953FE05FE81" lastPageId="11" lastPageNumber="382" pageId="8" pageNumber="379" type="description">
<paragraph id="8BA43689FFB53B051576F953FE6CFEC1" blockId="8.[835,1475,1085,2101]" lastBlockId="11.[113,752,189,277]" lastPageId="11" lastPageNumber="382" pageId="8" pageNumber="379">
<emphasis id="B96FEA9BFFB53B061576F953FC67FB81" bold="true" box="[835,979,1085,1109]" pageId="8" pageNumber="379">Description.</emphasis>
Body generally stramineous brown, ovoid, moderate in size; dorsal surface shining, with brown to dark brown stripes and maculae as in
<figureCitation id="13202A0CFFB53B0612E3F913FAE5FB41" box="[1238,1361,1149,1173]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Figs. 4F, G</figureCitation>
,
<figureCitation id="13202A0CFFB53B061369F910FA34FB41" box="[1372,1408,1150,1173]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="8" pageNumber="379">7D</figureCitation>
. Head shiny pale brown, with sparsely distributed, short, semi-erect setae; clypeus fuscous; vertex with basal transverse carina concaved mesad, about as thick as pronotal collar (
<figureCitation id="13202A0CFFB53B0613A0F9B3FCD8FAC1" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21J</figureCitation>
). Antenna pale brown; antennomere I with dark apex; apical ¼–⅓ of antennomere II darkened; antennomeres III and IV dark brown, except for pale base of antennomere III. Labium shiny pale brown, reaching but not exceeding middle of metacoxa; apical part of segment IV dark reddish brown. Pronotum shiny pale brown, with sparsely distributed, short, reclining setae and minute punctures (
<figureCitation id="13202A0CFFB53B06130EF8D3FA00FA01" box="[1339,1460,1469,1493]" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21I, J</figureCitation>
); calli usually with a pair of dark spots; pleura pale brown; metathoracic scent efferent system creamy yellow, with small, triangular peritreme (
<figureCitation id="13202A0CFFB53B061243FB73FB6EF9E1" box="[1142,1242,1565,1589]" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21L</figureCitation>
); scutellum polished, with brown mesal stripe and dark apex, finely and shallowly punctate, with uniformly distributed, short, semi-erect setae (
<figureCitation id="13202A0CFFB53B0615B9FB13FC5DF941" box="[908,1001,1661,1685]" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21I</figureCitation>
). Hemelytron shining, stramineous brown, with striped pattern as in
<figureCitation id="13202A0CFFB53B061255FBF3FB54F961" box="[1120,1248,1693,1717]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Figs. 4F, G</figureCitation>
,
<figureCitation id="13202A0CFFB53B0612DBFBF0FAA4F961" box="[1262,1296,1694,1717]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="8" pageNumber="379">7D</figureCitation>
, with uniformly distributed, semi-erect or reclining setae (
<figureCitation id="13202A0CFFB53B061327FBD3FACDF901" box="[1298,1401,1725,1749]" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21H</figureCitation>
); apex of cuneus narrowly brown; membrane pale smoky brown. All coxae and legs pale brown; metafemur apically with two brown rings; apex of each tarsomere III dark brown; pretarsal structures as in
<figureCitation id="13202A0CFFB53B061268FA53FB70F881" box="[1117,1220,1853,1877]" captionStart="Fig" captionStartId="32.[136,171,1943,1965]" captionTargetBox="[136,1474,252,1927]" captionTargetId="figure-7@32.[136,1474,252,1927]" captionTargetPageId="32" captionText="Fig. 21. Scanning electron micrographs of Chelyolygus fuhoshoensis (AG) and C. parsanus, new species (HO). A, Head and pronotum, dorsal view; B, H, dorsal habitus; C, J, anterior body, left lateral view; D, L, metathoracic scent efferent system; E, M, metatarsus; F, N, pretarsal structure of metaleg; G, O, male genital segment (pygophore) with parameres, dorsal view; I, pronotum, scutellum and anterior corium, dorsal view; K, thoracic pleura, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.14683000" httpUri="https://zenodo.org/record/14683000/files/figure.png" pageId="8" pageNumber="379">Fig. 21N</figureCitation>
; parempodia relatively short, narrow. Abdomen pale brown. Male genitalia (
<figureCitation id="13202A0CFFB53B0613B9FA33FCFBF841" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Figs. 5</figureCitation>
EH, 21O, 22A, B): Left paramere stout, L-shaped, with short hypophysis (
<figureCitation id="13202A0CFFB53B061233FAF3FBD4F861" box="[1030,1120,1949,1973]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Figs. 5F</figureCitation>
,
<figureCitation id="13202A0CFFB53B06125EFAF3FB29F861" box="[1131,1181,1949,1973]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">22A</figureCitation>
); right paramere with short, tapered hypophysis (
<figureCitation id="13202A0CFFB53B061204FAD3FB39F801" box="[1073,1165,1981,2005]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Fig. 5G</figureCitation>
). Vesica with two apically tapered spicules (
<figureCitation id="13202A0CFFB53B0615CEFAB3FBFBF821" box="[1019,1103,2013,2037]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Fig. 5H</figureCitation>
,
<figureCitation id="13202A0CFFB53B06126FFAB3FB3DF821" box="[1114,1161,2013,2037]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">22B</figureCitation>
,
<figureCitation id="13202A0CFFB53B0612A6FAB3FB09F821" box="[1171,1213,2013,2037]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[135,170,1927,1949]" captionStartId-1="3.[113,148,1878,1900]" captionTargetBox-0="[135,1464,325,1884]" captionTargetBox-1="[113,1451,352,1863]" captionTargetId-0="figure-7@2.[135,1473,297,1912]" captionTargetId-1="figure-11@3.[113,1451,352,1863]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." captionText-1="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.14682948" figureDoi-1="http://doi.org/10.5281/zenodo.14682952" httpUri-0="https://zenodo.org/record/14682948/files/figure.png" httpUri-1="https://zenodo.org/record/14682952/files/figure.png" pageId="8" pageNumber="379">12</figureCitation>
); lobal-sclerite (
<figureCitation id="13202A0CFFB53B06135FFAB3FA0AF821" box="[1386,1470,2013,2037]" captionStart="Fig" captionStartId="10.[135,170,1945,1967]" captionTargetBox="[135,1473,327,1930]" captionTargetId="figure-7@10.[135,1473,308,1930]" captionTargetPageId="10" captionText="Fig. 5. Male genitalia of Chelyolygus fuhoshoensis (AD) and C. parsanus, new species (EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="8" pageNumber="379">Fig. 5H</figureCitation>
,
<figureCitation id="13202A0CFFB53B061576FA93FCE4F7C1" box="[835,848,2045,2069]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="8" pageNumber="379">3</figureCitation>
) relatively short. Female genitalia (
<figureCitation id="13202A0CFFB53B0612EAFA93FAE6F7C1" box="[1247,1362,2045,2069]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Figs. 4I, J</figureCitation>
,
<figureCitation id="13202A0CFFB53B061354FA93FA02F7C1" box="[1377,1462,2045,2069]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="8" pageNumber="379">22C, D</figureCitation>
): sclerotised rings elongate ovoid, narrow (
<figureCitation id="13202A0CFFB53B0612CDF573FAF7F7E1" box="[1272,1347,2077,2101]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="8" pageNumber="379">Fig. 4J</figureCitation>
); ovipositor (gonapophysis I) as in
<figureCitation id="13202A0CFFB63B051740FDD3FE77FF01" box="[373,451,189,213]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="11" pageNumber="382">Fig. 4I</figureCitation>
; posterior wall with small, rounded dorsal structure and sparse spinules on anterior interramal sclerite (
<figureCitation id="13202A0CFFB63B051770FD93FE7FFEC1" box="[325,459,253,277]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="11" pageNumber="382">Fig. 22C, D</figureCitation>
).
</paragraph>
<caption id="DF646601FFB43B071644FAF6FA28F81E" ID-DOI="http://doi.org/10.5281/zenodo.14682958" ID-Zenodo-Dep="14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="9" pageNumber="380" startId="9.[113,148,1944,1966]" targetBox="[113,1451,328,1928]" targetPageId="9" targetType="figure">
<paragraph id="8BA43689FFB43B071644FAF6FA28F81E" blockId="9.[113,1452,1944,1995]" pageId="9" pageNumber="380">
Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of
<taxonomicName id="4C1B4D0AFFB43B071526FAF6FBBFF879" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[787,1035,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFFB43B071526FAF6FBBFF879" box="[787,1035,1944,1965]" italics="true" pageId="9" pageNumber="380">Chelyolygus fuhoshoensis</emphasis>
</taxonomicName>
(AE) and
<taxonomicName id="4C1B4D0AFFB43B07124EFAF6FB5BF879" authorityName="Yasunaga" authorityYear="2024" box="[1147,1263,1944,1965]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="380" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFFB43B07124EFAF6FB5BF879" box="[1147,1263,1944,1965]" italics="true" pageId="9" pageNumber="380">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB43B0712CFFAF6FAD9F87A" box="[1274,1389,1944,1966]" pageId="9" pageNumber="380" rank="species">new species</taxonomicNameLabel>
(FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures.
</paragraph>
</caption>
<caption id="DF646601FFB73B0416B2FAF7FB09F81F" ID-DOI="http://doi.org/10.5281/zenodo.14682960" ID-Zenodo-Dep="14682960" httpUri="https://zenodo.org/record/14682960/files/figure.png" pageId="10" pageNumber="381" startId="10.[135,170,1945,1967]" targetBox="[135,1473,327,1930]" targetPageId="10" targetType="figure">
<paragraph id="8BA43689FFB73B0416B2FAF7FB09F81F" blockId="10.[135,1474,1945,1996]" pageId="10" pageNumber="381">
Fig. 5. Male genitalia of
<taxonomicName id="4C1B4D0AFFB73B0417B6FAF7FDCBF87A" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[387,639,1945,1966]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFFB73B0417B6FAF7FDCBF87A" box="[387,639,1945,1966]" italics="true" pageId="10" pageNumber="381">Chelyolygus fuhoshoensis</emphasis>
</taxonomicName>
(AD) and
<taxonomicName id="4C1B4D0AFFB73B0414CFFAF4FCC5F87B" authorityName="Yasunaga" authorityYear="2024" box="[762,881,1946,1967]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="381" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFFB73B0414CFFAF4FCC5F87B" box="[762,881,1946,1967]" italics="true" pageId="10" pageNumber="381">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB73B0415B5FAF4FC42F87B" box="[896,1014,1945,1967]" pageId="10" pageNumber="381" rank="species">new species</taxonomicNameLabel>
(EH). A, E, Genital segment (pygophore) in dorsal (A) and ventral (E) views; B, F, left paramere; C, G, right paramere; D, H, vesica. Scale bars 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA43689FFB63B051644FC53FE05FE81" blockId="11.[113,433,317,341]" box="[113,433,317,341]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FC53FE94FE81" bold="true" box="[113,288,317,341]" pageId="11" pageNumber="382">Measurements.</emphasis>
See
<tableCitation id="C6990332FFB63B051762FC53FE1FFE81" box="[343,427,317,341]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="11" pageNumber="382">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644FC13FE77FE61" pageId="11" pageNumber="382" type="etymology">
<paragraph id="8BA43689FFB63B051644FC13FE77FE61" blockId="11.[113,751,381,437]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FC13FF4DFE41" bold="true" box="[113,249,381,405]" pageId="11" pageNumber="382">Etymology.</emphasis>
Named for the
<typeStatus id="54A0882BFFB63B05178CFC10FE5DFE42" box="[441,489,382,406]" pageId="11" pageNumber="382">type</typeStatus>
locality, Parsa Nature Reserve in S.
<collectingCountry id="F30C7619FFB63B051739FCF3FEFBFE61" box="[268,335,413,437]" name="Nepal" pageId="11" pageNumber="382">Nepal</collectingCountry>
; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644FCB3FDE7FE21" box="[113,595,477,501]" pageId="11" pageNumber="382" type="distribution">
<paragraph id="8BA43689FFB63B051644FCB3FDE7FE21" blockId="11.[113,595,477,501]" box="[113,595,477,501]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FCB3FEB0FE21" bold="true" box="[113,260,477,501]" pageId="11" pageNumber="382">Distribution.</emphasis>
<collectingCountry id="F30C7619FFB63B051739FCB3FEF8FE21" box="[268,332,477,501]" name="Nepal" pageId="11" pageNumber="382">Nepal</collectingCountry>
(Makawanpur District).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644FF73FF51FD41" pageId="11" pageNumber="382" type="biology_ecology">
<paragraph id="8BA43689FFB63B051644FF73FF51FD41" blockId="11.[113,752,541,661]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FF73FF67FDE1" bold="true" box="[113,211,541,565]" pageId="11" pageNumber="382">Biology.</emphasis>
This new species was found to co-occur with
<taxonomicName id="4C1B4D0AFFB63B051644FF53FE1BFD81" authorityName="Yasunaga" authorityYear="2024" box="[113,431,573,597]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="11" pageNumber="382" phylum="Tracheophyta" rank="species" species="rufobrunneus" status="sp. nov.">
<emphasis id="B96FEA9BFFB63B051644FF53FE1BFD81" box="[113,431,573,597]" italics="true" pageId="11" pageNumber="382">Euphorbialygus rufobrunneus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB63B05178EFF50FD8FFD81" box="[443,571,573,597]" pageId="11" pageNumber="382" rank="species">new species</taxonomicNameLabel>
, on inflorescence of
<taxonomicName id="4C1B4D0AFFB63B0516BAFF33FEA0FDA1" box="[143,276,605,629]" class="Magnoliopsida" family="Euphorbiaceae" genus="Mallotus" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="382" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B96FEA9BFFB63B0516BAFF33FF5AFDA1" box="[143,238,605,629]" italics="true" pageId="11" pageNumber="382">Mallotus</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13202A0CFFB63B051716FF33FECDFDA1" box="[291,377,605,629]" captionStart="Fig" captionStartId="15.[113,148,2009,2031]" captionTargetBox="[113,1451,215,1995]" captionTargetId="figure-11@15.[113,1451,215,1995]" captionTargetPageId="15" captionText="Fig. 8. Habitus images and habitats (G, H) of Euphorbialygus nanglong, new species from Nakhon Nayok, Thailand (AD, G) and E. rufobrunneus, new species from Chitwan National Park, Nepal (E, F, H). A, B, Male; C, E, F, female; D, 5th instar nymph; G, Homonoia riparia; H, Mallotus sp." figureDoi="http://doi.org/10.5281/zenodo.14682966" httpUri="https://zenodo.org/record/14682966/files/figure.png" pageId="11" pageNumber="382">Fig. 8H</figureCitation>
). However, no immature form was confirmed.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,233 @@
<document id="AF057E0919C3FC861B92C5998A958A10" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB63B0315A0FF53FF65FC42" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus virgulatellus Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="384" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="382" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E07377F686B641FEA424A86FCDB6ABEB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C0F8A7B4E3F78B2D506F1B30212B4FD6">
<mods:title id="FE84A6FBEDCB06459982E06D5851D827">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="57E22B34062EE251A2B2BC7F2044F3B2" type="personal">
<mods:role id="1FB58352AAB8B105F71E6BB008B613CB">
<mods:roleTerm id="75D5EB23A2342A776B27322C42743AFD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70AB43164BCE615F8BC8D30679CF9848">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="1C401371E871B96B9BCE1B87988FB8D1">text</mods:typeOfResource>
<mods:relatedItem id="45D3995F3F4AF0FDAB3E947A03588729" type="host">
<mods:titleInfo id="889F1E3D5EE9EBEB6233C8490B80E237">
<mods:title id="4A95A26B9B1834DA66596C35918E22AC">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1B0A5B8EE39EC7D8FB82E3A601A04D5C">
<mods:date id="F79E3E0FFE35221932EF4C76E855604D">2024</mods:date>
<mods:detail id="AF62880D0DE1E64E934466FC1B446CB3" type="pubDate">
<mods:number id="13F00E8AB76736AC040A9BE76E25CFF2">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="2BC6745027CF9E371F6D399D97D016BD" type="volume">
<mods:number id="62A268A68FD4EC1EA9A3A530ED830853">72</mods:number>
</mods:detail>
<mods:extent id="C3F50A0B685D2C36B094DD5D303B63F3" unit="page">
<mods:start id="70DB68FF4D63B348B7ABF60C16CA0EE2">371</mods:start>
<mods:end id="11269E8FC6BA69AFDAAB7FB12C3A0C65">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A0C8F315229B56F0893B44CD2386C4F9">journal article</mods:classification>
<mods:identifier id="0B73D4B9E2A6360BFA7A7F858D498EBC" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="7C7766629EE494DC56F90FA70375BF44" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="637DE317D27A07DABAFE9BC26C8094E0" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="57D4E45C5B7C3620E91C3D9CD458088C" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB63B0315A0FF53FF65FC42" LSID="urn:lsid:plazi:treatment:03B2879FFFB63B0315A0FF53FF65FC42" httpUri="http://treatment.plazi.org/id/03B2879FFFB63B0315A0FF53FF65FC42" lastPageId="13" lastPageNumber="384" pageId="11" pageNumber="382">
<subSubSection id="C3016502FFB63B0515A0FF53FAF7FD81" box="[917,1347,573,597]" pageId="11" pageNumber="382" type="nomenclature">
<paragraph id="8BA43689FFB63B0515A0FF53FAF7FD81" blockId="11.[917,1347,573,629]" box="[917,1347,573,597]" pageId="11" pageNumber="382">
<heading id="D0EC81E5FFB63B0515A0FF53FAF7FD81" bold="true" box="[917,1347,573,597]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="382" reason="2">
<emphasis id="B96FEA9BFFB63B0515A0FF53FAF7FD81" bold="true" box="[917,1347,573,597]" pageId="11" pageNumber="382">
<taxonomicName id="4C1B4D0AFFB63B0515A0FF53FB1BFD81" authorityName="Yasunaga" authorityYear="2024" box="[917,1199,573,597]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="382" phylum="Arthropoda" rank="species" species="virgulatellus" status="sp. nov.">
<emphasis id="B96FEA9BFFB63B0515A0FF53FB1BFD81" bold="true" box="[917,1199,573,597]" italics="true" pageId="11" pageNumber="382">Chelyolygus virgulatellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB63B05128BFF53FAF7FD81" box="[1214,1347,573,597]" pageId="11" pageNumber="382" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051226FF33FB70FDA1" box="[1043,1220,605,629]" pageId="11" pageNumber="382" type="description">
<paragraph id="8BA43689FFB63B051226FF33FB70FDA1" blockId="11.[917,1347,573,629]" box="[1043,1220,605,629]" pageId="11" pageNumber="382">
(
<figureCitation id="13202A0CFFB63B05122EFF33FBDCFDA1" box="[1051,1128,605,629]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Figs. 6</figureCitation>
,
<figureCitation id="13202A0CFFB63B051243FF33FB09FDA1" box="[1142,1213,605,629]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="11" pageNumber="382">22JO</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051518FFF3FBACFC41" pageId="11" pageNumber="382" type="materials_examined">
<paragraph id="8BA43689FFB63B051518FFF3FBACFC41" blockId="11.[813,1452,669,917]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FFF3FC6CFD61" bold="true" box="[813,984,669,693]" pageId="11" pageNumber="382">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB63B0515D4FFF3FA88FCC1" collectingDate="2023-08-16" collectionCode="T, ZRC, AMNH" country="Singapore" county="Yasunaga" latitude="1.3570833" location="Windsor Nature Park" longLatPrecision="1" longitude="103.81833" municipality="Island Club Road" pageId="11" pageNumber="382" specimenCode="PBI 00378779" specimenCount="1" typeStatus="holotype">
<emphasis id="B96FEA9BFFB63B0515D4FFF3FBFDFD61" bold="true" box="[993,1097,669,693]" pageId="11" pageNumber="382">
<typeStatus id="54A0882BFFB63B0515D4FFF3FBFDFD61" box="[993,1097,669,693]" pageId="11" pageNumber="382" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFB63B0512ABFFF3FAF2FD61" bold="true" box="[1182,1350,669,693]" pageId="11" pageNumber="382">
<collectingCountry id="F30C7619FFB63B0512ABFFF3FAF5FD61" box="[1182,1345,669,693]" name="Singapore" pageId="11" pageNumber="382">SINGAPORE</collectingCountry>
:
</emphasis>
<location id="8EC46052FFB63B05137AFFF3FC1AFD01" LSID="urn:lsid:plazi:treatment:03B2879FFFB63B0315A0FF53FF65FC42:8EC46052FFB63B05137AFFF3FC1AFD01" country="Singapore" county="Yasunaga" latitude="1.3570833" longLatPrecision="1" longitude="103.81833" municipality="Island Club Road" name="Windsor Nature Park" pageId="11" pageNumber="382">Windsor Nature Park</location>
,
<location id="8EC46052FFB63B051582FFD3FC5FFD01" LSID="urn:lsid:plazi:treatment:03B2879FFFB63B0315A0FF53FF65FC42:8EC46052FFB63B051582FFD3FC5FFD01" box="[951,1003,701,725]" country="Singapore" county="Yasunaga" latitude="1.3570833" longLatPrecision="1" longitude="103.81833" municipality="Island Club Road" name="Trail" pageId="11" pageNumber="382">Trail</location>
along
<collectingMunicipality id="6BC0ACF3FFB63B051205FFD3FB5EFD01" box="[1072,1258,701,725]" pageId="11" pageNumber="382">Island Club Road</collectingMunicipality>
, sweeping flowers of a broadleaf tree,
<geoCoordinate id="EE2F504EFFB63B0515C6FFB2FBC2FD21" box="[1011,1142,732,757]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="382" precision="1" seconds="25.5" value="1.3570833">1°2125.5″N</geoCoordinate>
<geoCoordinate id="EE2F504EFFB63B05124EFFB2FAACFD21" box="[1147,1304,732,757]" degrees="103" direction="east" minutes="49" orientation="longitude" pageId="11" pageNumber="382" precision="1" seconds="06.0" value="103.81833">103°4906.0″E</geoCoordinate>
,
<collectionCode id="ED0AAE4CFFB63B051314FFB0FA80FD21" box="[1313,1332,734,757]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="382">T</collectionCode>
.
<collectingCounty id="62C54E05FFB63B051308FFB0FA13FD22" box="[1341,1447,734,758]" pageId="11" pageNumber="382">Yasunaga</collectingCounty>
,
<date id="FFA51049FFB63B051518FF93FC6CFCC1" box="[813,984,765,790]" pageId="11" pageNumber="382" value="2023-08-16">
<collectingDate id="EFE1E9A1FFB63B051518FF93FC6CFCC1" box="[813,984,765,790]" pageId="11" pageNumber="382" value="2023-08-16">16 August 2023</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB63B0515D2FF93FBAAFCC1" box="[999,1054,765,789]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="382">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFB63B051201FF93FB33FCC1" box="[1076,1159,765,789]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="382">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB63B0512A1FF93FA81FCC1" box="[1172,1333,765,789]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="382">PBI 00378779</specimenCode>
)
</materialsCitation>
.
<typeStatus id="54A0882BFFB63B051372FF90FA1DFCC2" box="[1351,1449,766,790]" pageId="11" pageNumber="382" type="paratype">Paratype</typeStatus>
: SINGAPORE:
<materialsCitation id="3B733CD4FFB63B0515E5FE73FA20FCE1" box="[976,1428,797,821]" collectingDate="2023-08-16" collectionCode="TYCN" country="Singapore" county="Yasunaga" latitude="1.3570833" location="Windsor Nature Park" longLatPrecision="1" longitude="103.81833" municipality="Island Club Road" pageId="11" pageNumber="382" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB63B0515E5FE73FBADFCE1" box="[976,1049,797,821]" count="1" pageId="11" pageNumber="382" type="male">1 male</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFB63B05130BFE73FA39FCE1" box="[1342,1421,797,821]" pageId="11" pageNumber="382">TYCN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B733CD4FFB63B0513ABFE73FBA0FC41" collectingDate="2024-03-18" collectionCode="ZRC" collectorName="E. H. Yep" country="Singapore" county="Yasunaga" latitude="1.3589722" location="Windsor Nature Park" longLatPrecision="1" longitude="103.82655" municipality="Island Club Road" pageId="11" pageNumber="382" specimenCode="BDP0372659" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1DFD00FFB63B0513ABFE73FCCCFC81" count="1" pageId="11" pageNumber="382" type="female">1 female</specimenCount>
, same locality,
<collectedFrom id="38077574FFB63B051228FE50FA9AFC81" box="[1053,1326,829,853]" pageId="11" pageNumber="382">
on
<taxonomicName id="4C1B4D0AFFB63B05120AFE53FA9AFC81" box="[1087,1326,829,853]" class="Magnoliopsida" family="Sapindaceae" genus="Nephelium" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="382" phylum="Tracheophyta" rank="species" species="lappaceum">
<emphasis id="B96FEA9BFFB63B05120AFE53FA9AFC81" box="[1087,1326,829,853]" italics="true" pageId="11" pageNumber="382">Nephelium lappaceum</emphasis>
</taxonomicName>
</collectedFrom>
(rambutan) tree,
<geoCoordinate id="EE2F504EFFB63B051554FE32FC52FCA1" box="[865,998,860,885]" degrees="1" direction="north" minutes="21" orientation="latitude" pageId="11" pageNumber="382" precision="1" seconds="32.3" value="1.3589722">1°2132.3″N</geoCoordinate>
<geoCoordinate id="EE2F504EFFB63B0515D9FE32FB38FCA1" box="[1004,1164,860,885]" degrees="103" direction="east" minutes="49" orientation="longitude" pageId="11" pageNumber="382" precision="1" seconds="35.6" value="103.82655">103°4935.6″E</geoCoordinate>
,
<date id="FFA51049FFB63B0512A3FE33FA8DFCA1" box="[1174,1337,861,885]" pageId="11" pageNumber="382" value="2024-03-18">
<collectingDate id="EFE1E9A1FFB63B0512A3FE33FA8DFCA1" box="[1174,1337,861,885]" pageId="11" pageNumber="382" value="2024-03-18">18 March 2024</collectingDate>
</date>
,
<collectorName id="26EE535FFFB63B051376FE30FA1FFCA1" box="[1347,1451,862,885]" pageId="11" pageNumber="382">E. H. Yep</collectorName>
(
<collectionCode id="ED0AAE4CFFB63B051501FE13FCDEFC41" box="[820,874,893,917]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="382">ZRC</collectionCode>
_
<specimenCode id="DBBD9EF2FFB63B05154DFE13FBB9FC41" box="[888,1037,893,917]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="382">BDP0372659</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051518FED3FA11FB41" pageId="11" pageNumber="382" type="diagnosis">
<paragraph id="8BA43689FFB63B051518FED3FA11FB41" blockId="11.[813,1452,957,1173]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FED3FC15FC01" bold="true" box="[813,929,957,981]" pageId="11" pageNumber="382">Diagnosis.</emphasis>
This new species is recognised by the following characters: Widely pale olive-green general colouration (
<figureCitation id="13202A0CFFB63B051500FE93FC67FBC1" box="[821,979,1021,1046]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="11" pageNumber="382">Fig. 7A, C, F</figureCitation>
); small size; pale base of antennomeres I and II; brown, narrow, continuous stripes along veins on hemelytron; small, sharply pointed hypophysis of right paramere; and presence of two additional lobal-sclerites on vesica, in addition to the characters mentioned in the key.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B031518F9D3FE05FE41" lastPageId="13" lastPageNumber="384" pageId="11" pageNumber="382" type="description">
<paragraph id="8BA43689FFB63B031518F9D3FDF4FE81" blockId="11.[813,1452,1213,2101]" lastBlockId="13.[113,753,189,341]" lastPageId="13" lastPageNumber="384" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518F9D3FC0FFB01" bold="true" box="[813,955,1213,1237]" pageId="11" pageNumber="382">Description.</emphasis>
Body generally pale olive green (fading to yellowish in dry-preserved specimen as in
<figureCitation id="13202A0CFFB63B0512D9F9B3FAEDFB21" box="[1260,1369,1245,1269]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Fig. 6F, G</figureCitation>
), small, stout, tortoise-shaped (
<figureCitation id="13202A0CFFB63B051213F993FB2AFAC1" box="[1062,1182,1277,1301]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Fig. 6A, F</figureCitation>
); dorsal surface shining, shallowly and minutely punctate, with uniformly distributed, pale, short, semi-erect setae. Head shiny pale green (
<figureCitation id="13202A0CFFB63B05137EF853FA2BFA81" box="[1355,1439,1341,1365]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Fig. 6D</figureCitation>
); vertex about as wide as an eye in dorsal view, with narrow, continuous basal transverse carina; apical ¼ of clypeus infuscate (
<figureCitation id="13202A0CFFB63B051597F8F3FC4EFA61" box="[930,1018,1437,1461]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Fig. 6E</figureCitation>
). Antenna dark brown; antennomere I pale brown, with fuscous apex and slightly darkened basal ⅓; antennomere II with pale base, about as long as labium, slightly shorter than basal width of pronotum. Labium pale reddish brown, slightly exceeding apex of mesocoxa; apical half of segment IV infuscate (
<figureCitation id="13202A0CFFB63B051243FB53FB5EF981" box="[1142,1258,1597,1621]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Fig. 6C, F</figureCitation>
). Pronotum shiny pale olive; calli pale green; thoracic pleura and scutellum creamy yellow, partly tinged with green; metathoracic scent efferent system as in
<figureCitation id="13202A0CFFB63B05123AFBF3FBDAF961" box="[1039,1134,1693,1717]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="11" pageNumber="382">
Fig.
<specimenCount id="9D1DFD00FFB63B051275FBF3FBDAF961" box="[1088,1134,1693,1717]" count="22" pageId="11" pageNumber="382" type="generic">22L</specimenCount>
</figureCitation>
. Hemelytra pale olive-brown, with dark, narrow, continuous stripes along veins and sutures, on clavus, corium and exocorium (embolium); apical inner corner of corium (or paracuneus) with brown, U-shaped macula; apex of cuneus narrowly darkened; membrane smoky brown, with pale veins. All coxae and legs pale creamy brown; metafemur with two brown apical rings that are interrupted posteriad; tibial spines reddish brown, each with a brown small spot; apical half of each tarsomere III dark brown; pretarsal structure as in
<figureCitation id="13202A0CFFB63B051274FAD3FB1CF801" box="[1089,1192,1981,2005]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="11" pageNumber="382">Fig. 22O</figureCitation>
; parempodia relatively long and wide. Ventral surface of abdomen uniformly pale olive green. Male genitalia (
<figureCitation id="13202A0CFFB63B051251FA93FB6AF7C1" box="[1124,1246,2045,2069]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="11" pageNumber="382">Figs. 6HJ</figureCitation>
,
<figureCitation id="13202A0CFFB63B0512D9FA93FA97F7C1" box="[1260,1315,2045,2069]" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" pageId="11" pageNumber="382">22M</figureCitation>
): Pygophore with triangular projection at base of left paramere (
<figureCitation id="13202A0CFFB63B03134AF573FF1CFF01" captionStart="Fig" captionStartId="34.[135,170,1944,1966]" captionTargetBox="[135,1473,251,1928]" captionTargetId="figure-7@34.[135,1473,251,1928]" captionTargetPageId="34" captionText="Fig. 22. Scanning electron micrographs of Chelyolygus fuhoshoensis (AD), C. parsanus, new species (FI), C. virgulatellus, new species (JO) and Apolygopsis furvocarinatus (E). A, F, Left paramere; B, G, vesica; CE, H, I, posterior wall; J, anterior body, dorsal view; K, left lateral habitus; L, M, apex of pygophore, left lateral view; N, metatarsus; O, pretarsal structure of metaleg. Abbreviations: DOS = dorsal structure, IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.14683004" httpUri="https://zenodo.org/record/14683004/files/figure.png" lastPageId="13" lastPageNumber="384" pageId="11" pageNumber="382">Fig. 22M</figureCitation>
); right paramere with short, pointed hypophysis (
<figureCitation id="13202A0CFFB03B0314F0FDD3FF22FF21" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="13" pageNumber="384">Fig. 4H</figureCitation>
); left paramere C-shaped, with rather elongate process on apex of hypophysis (
<figureCitation id="13202A0CFFB03B031745FD93FE0FFEC1" box="[368,443,253,277]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="13" pageNumber="384">Fig. 4I</figureCitation>
). Vesica with two stout lobal sclerites in addition to two slender, weakly curved spicules (
<figureCitation id="13202A0CFFB03B03164CFC53FF7CFE81" box="[121,200,317,341]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="13" pageNumber="384">Fig. 4J</figureCitation>
). Female genitalia: Not examined.
</paragraph>
<caption id="DF646601FFB13B0216B2FAF9FE62F832" ID-DOI="http://doi.org/10.5281/zenodo.14682962" ID-Zenodo-Dep="14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="12" pageNumber="383" startId="12.[135,170,1943,1965]" targetBox="[135,1473,281,1907]" targetPageId="12" targetType="figure">
<paragraph id="8BA43689FFB13B0216B2FAF9FE62F832" blockId="12.[135,1474,1943,2022]" pageId="12" pageNumber="383">
Fig. 6. Habitus images of
<taxonomicName id="4C1B4D0AFFB13B0217B8FAF9FD31F878" authorityName="Yasunaga" authorityYear="2024" box="[397,645,1943,1964]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="383" phylum="Arthropoda" rank="species" species="virgulatellus" status="sp. nov.">
<emphasis id="B96FEA9BFFB13B0217B8FAF9FD31F878" box="[397,645,1943,1964]" italics="true" pageId="12" pageNumber="383">Chelyolygus virgulatellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB13B0214A1FAF9FCB8F879" box="[660,780,1943,1965]" pageId="12" pageNumber="383" rank="species">new species</taxonomicNameLabel>
, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap.
</paragraph>
</caption>
<paragraph id="8BA43689FFB03B031644FC13FE05FE41" blockId="13.[113,433,381,405]" box="[113,433,381,405]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FC13FE94FE41" bold="true" box="[113,288,381,405]" pageId="13" pageNumber="384">Measurements.</emphasis>
See
<tableCitation id="C6990332FFB03B031762FC13FE1FFE41" box="[343,427,381,405]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="13" pageNumber="384">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644FCD3FE39FDC1" pageId="13" pageNumber="384" type="etymology">
<paragraph id="8BA43689FFB03B031644FCD3FE39FDC1" blockId="13.[113,753,445,533]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FCD3FF49FE01" bold="true" box="[113,253,445,469]" pageId="13" pageNumber="384">Etymology.</emphasis>
From Latin virgulatus (= striped) with diminutive suffix, referring to stripes on hemelytra of this new species; an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644FF53FD21FD81" box="[113,661,573,597]" pageId="13" pageNumber="384" type="distribution">
<paragraph id="8BA43689FFB03B031644FF53FD21FD81" blockId="13.[113,661,573,597]" box="[113,661,573,597]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FF53FEB0FD81" bold="true" box="[113,260,573,597]" pageId="13" pageNumber="384">Distribution.</emphasis>
<collectingCountry id="F30C7619FFB03B031739FF53FECCFD81" box="[268,376,573,597]" name="Singapore" pageId="13" pageNumber="384">Singapore</collectingCountry>
(tropical rainforest zone).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB03B031644FF13FF65FC42" pageId="13" pageNumber="384" type="biology_ecology">
<paragraph id="8BA43689FFB03B031644FF13FF65FC42" blockId="13.[113,753,637,918]" pageId="13" pageNumber="384">
<emphasis id="B96FEA9BFFB03B031644FF13FF65FD41" bold="true" box="[113,209,637,661]" pageId="13" pageNumber="384">Biology.</emphasis>
<specimenCount id="9D1DFD00FFB03B0316E9FF10FEF9FD41" box="[220,333,637,661]" count="2" pageId="13" pageNumber="384" type="male">Two male</specimenCount>
(one teneral) adult individuals were collected by sweeping inflorescences of a broadleaf tree, most probably identical to
<taxonomicName id="4C1B4D0AFFB03B0317ABFFD3FEACFD21" authority="(Reinw. ex Blume) Hassk." authorityName="Hassk." baseAuthorityName="Blume" class="Magnoliopsida" family="Euphorbiaceae" genus="Claoxylon" kingdom="Plantae" order="Malpighiales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="species" species="indicum">
<emphasis id="B96FEA9BFFB03B0317ABFFD3FDD9FD01" box="[414,621,701,725]" italics="true" pageId="13" pageNumber="384">Claoxylon indicum</emphasis>
(Reinw. ex Blume) Hassk.
</taxonomicName>
(
<taxonomicName id="4C1B4D0AFFB03B03171CFFB3FE7BFD21" box="[297,463,733,757]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
) (Yap EH, pers. comm.). The female specimen was found from a rambutan tree,
<taxonomicName id="4C1B4D0AFFB03B031644FE73FEEDFCE1" box="[113,345,797,821]" class="Magnoliopsida" family="Sapindaceae" genus="Nephelium" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="species" species="lappaceum">
<emphasis id="B96FEA9BFFB03B031644FE73FEEDFCE1" box="[113,345,797,821]" italics="true" pageId="13" pageNumber="384">Nephelium lappaceum</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C1B4D0AFFB03B0317B5FE73FDBCFCE1" box="[384,520,797,821]" class="Magnoliopsida" family="Sapindaceae" kingdom="Plantae" order="Sapindales" pageId="13" pageNumber="384" phylum="Tracheophyta" rank="family">Sapindaceae</taxonomicName>
). No other information is currently available. The collection records suggest
<taxonomicName id="4C1B4D0AFFB03B0314E2FE50FF49FCA1" authorityName="Yasunaga" authorityYear="2024" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="384" phylum="Arthropoda" rank="species" species="virgulatellus" status="new species">
<emphasis id="B96FEA9BFFB03B0314E2FE50FF49FCA1" italics="true" pageId="13" pageNumber="384">C. virgulatellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB03B03173AFE30FE2FFCA1" box="[271,411,861,885]" pageId="13" pageNumber="384" rank="species">new species</taxonomicNameLabel>
, has two or more generations per year.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,165 @@
<document id="9755BE1E7DA059A1C8A650A2DF700544" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB63B0516EEFFD3FBB7FDC1" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus tabananicus Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="382" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="382" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="9659B1834B203E5E42898C3A0B2DB30A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EED26DB21ED67351EBF92D75892E7AF5">
<mods:title id="4C13E591641CE53067202E4B59B8434A">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="76D9A8632E4BE17F2F49603555F9C8F2" type="personal">
<mods:role id="563D3C33ECE3C9EA96941BBE63D8DAF3">
<mods:roleTerm id="DD8F0F23782330E36277D01897E2F473">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BA16A5336ADBD160A2F8779357B99F5">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="675DA766D610918515916F7C8ADB02BC">text</mods:typeOfResource>
<mods:relatedItem id="D49647EFB1452060BDDA76CE1B93038A" type="host">
<mods:titleInfo id="33777432898F3EECBD731BE76986E430">
<mods:title id="3C08D51CA74709072ED19A8EF199C59E">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="EE9AB352D172A31E75D2C06F9E9E8262">
<mods:date id="0129C3AFCC3D31A3724ED3020A1A7FCC">2024</mods:date>
<mods:detail id="E8F30FAF37BC6D5EA5BF1B825FF8BA2B" type="pubDate">
<mods:number id="9ED8F682B97E141D8927DFC39F5B72D0">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="C44759CD54D615AA0BA5C91240084ACA" type="volume">
<mods:number id="FDC2A765B6F2C4F9155EFC7CF91A418E">72</mods:number>
</mods:detail>
<mods:extent id="2F49976B05168607F5AA722683100301" unit="page">
<mods:start id="C62D24B06C9F68DAD6A56B8BB4562C8C">371</mods:start>
<mods:end id="63207B424D99E28AAB9842D144D2B537">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="067F57795A115D5592DB2F8ADA3170F8">journal article</mods:classification>
<mods:identifier id="582B95F76D72A9254F28078BBBDCBBA3" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="C377836D308E6DFC4F7574B7B2E8B438" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E6F0D11CD82BA23BB7D192E927681576" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="4717F1516D29AEB400E36B4781DEA4EF" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB63B0516EEFFD3FBB7FDC1" ID-DOI="http://doi.org/10.5281/zenodo.14682974" ID-Zenodo-Dep="14682974" LSID="urn:lsid:plazi:treatment:03B2879FFFB63B0516EEFFD3FBB7FDC1" httpUri="http://treatment.plazi.org/id/03B2879FFFB63B0516EEFFD3FBB7FDC1" lastPageNumber="382" pageId="11" pageNumber="382">
<subSubSection id="C3016502FFB63B0516EEFFD3FD33FD01" box="[219,647,701,725]" pageId="11" pageNumber="382" type="nomenclature">
<paragraph id="8BA43689FFB63B0516EEFFD3FD33FD01" blockId="11.[219,647,701,757]" box="[219,647,701,725]" pageId="11" pageNumber="382">
<heading id="D0EC81E5FFB63B0516EEFFD3FD33FD01" bold="true" box="[219,647,701,725]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="382" reason="2">
<emphasis id="B96FEA9BFFB63B0516EEFFD3FD33FD01" bold="true" box="[219,647,701,725]" pageId="11" pageNumber="382">
<taxonomicName id="4C1B4D0AFFB63B0516EEFFD3FE46FD01" authorityName="Yasunaga" authorityYear="2024" box="[219,498,701,725]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="382" phylum="Arthropoda" rank="species" species="tabananicus" status="sp. nov.">
<emphasis id="B96FEA9BFFB63B0516EEFFD3FE46FD01" bold="true" box="[219,498,701,725]" italics="true" pageId="11" pageNumber="382">Chelyolygus tabananicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB63B051434FFD3FD33FD01" box="[513,647,701,725]" pageId="11" pageNumber="382" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051726FFB3FDFBFD21" box="[275,591,733,757]" pageId="11" pageNumber="382" type="description">
<paragraph id="8BA43689FFB63B051726FFB3FDFBFD21" blockId="11.[219,647,701,757]" box="[275,591,733,757]" pageId="11" pageNumber="382">
(
<figureCitation id="13202A0CFFB63B05172FFFB3FE29FD21" box="[282,413,733,757]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="11" pageNumber="382">Figs. 1A, B</figureCitation>
,
<figureCitation id="13202A0CFFB63B05179FFFB3FE03FD21" box="[426,439,733,757]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="11" pageNumber="382">2</figureCitation>
FH, 20GO)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644FE73FD12FCA1" pageId="11" pageNumber="382" type="materials_examined">
<paragraph id="8BA43689FFB63B051644FE73FD12FCA1" blockId="11.[113,752,797,886]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FE73FE90FCE1" bold="true" box="[113,292,797,821]" pageId="11" pageNumber="382">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB63B05171BFE73FD12FCA1" collectingDate="1996-08-08" collectingDateMax="1996-08-12" collectingDateMin="1996-08-08" collectionCode="ZRC, AMNH" country="Indonesia" latitude="-8.266666" location="Bedugul Area" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" pageId="11" pageNumber="382" specimenCode="PBI 00378778" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<emphasis id="B96FEA9BFFB63B05171BFE73FE2EFCE1" bold="true" box="[302,410,797,821]" pageId="11" pageNumber="382">
<typeStatus id="54A0882BFFB63B05171BFE73FE2EFCE1" box="[302,410,797,821]" pageId="11" pageNumber="382" type="holotype">Holotype</typeStatus>
</emphasis>
: female,
<emphasis id="B96FEA9BFFB63B05143EFE73FD04FCE1" bold="true" box="[523,688,797,821]" pageId="11" pageNumber="382">
<collectingCountry id="F30C7619FFB63B05143EFE73FD1FFCE1" box="[523,683,797,821]" name="Indonesia" pageId="11" pageNumber="382">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFB63B05148EFE73FD5FFCE1" box="[699,747,797,821]" country="Indonesia" name="Bali" pageId="11" pageNumber="382">Bali</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFB63B051644FE53FF64FC81" box="[113,208,829,853]" pageId="11" pageNumber="382">Tabanan</collectingMunicipality>
,
<location id="8EC46052FFB63B0516E8FE53FECCFC82" LSID="urn:lsid:plazi:treatment:03B2879FFFB63B0516EEFFD3FBB7FDC1:8EC46052FFB63B0516E8FE53FECCFC82" box="[221,376,829,854]" country="Indonesia" latitude="-8.266666" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" name="Bedugul Area" pageId="11" pageNumber="382" stateProvince="Bali">Bedugul Area</location>
,
<geoCoordinate id="EE2F504EFFB63B0517B1FE52FE7AFC81" box="[388,462,828,853]" degrees="8" direction="south" minutes="16" orientation="latitude" pageId="11" pageNumber="382" precision="925" value="-8.266666">8°16S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB63B0517E9FE52FDF6FC81" box="[476,578,828,853]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="11" pageNumber="382" precision="925" value="115.15">115°09E</geoCoordinate>
, local collector,
<date id="FFA51049FFB63B051644FE33FE8EFCA1" box="[113,314,861,886]" pageId="11" pageNumber="382" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">
<collectingDate id="EFE1E9A1FFB63B051644FE33FE8EFCA1" box="[113,314,861,886]" pageId="11" pageNumber="382" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">812 August 1996</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB63B05177FFE33FE35FCA1" box="[330,385,861,885]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="382">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFB63B0517ADFE33FE5FFCA1" box="[408,491,861,885]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="382">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB63B0517CCFE33FD2FFCA1" box="[505,667,861,885]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="382">PBI 00378778</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644FEF3FDC0FB41" pageId="11" pageNumber="382" type="diagnosis">
<paragraph id="8BA43689FFB63B051644FEF3FDC0FB41" blockId="11.[113,752,925,1173]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644FEF3FF5CFC61" bold="true" box="[113,232,925,949]" pageId="11" pageNumber="382">Diagnosis.</emphasis>
This new species is currently known only by a single female specimen and recognised readily by the following characters: Pale ivory brown general colouration; relatively large size; weak basal transverse carina on vertex; darkened apex of clypeus, annulated antennomere I; dark, short, interrupted stripes along veins on hemelytra; and semi-circular dorsal structure and narrow, less-spinulate, subtriangular interramal lobe on posterior wall.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051644F9D3FBD8FEE1" pageId="11" pageNumber="382" type="description">
<paragraph id="8BA43689FFB63B051644F9D3FA32FF21" blockId="11.[113,753,1213,2101]" lastBlockId="11.[813,1451,189,245]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051644F9D3FE76FB01" bold="true" box="[113,450,1213,1237]" pageId="11" pageNumber="382">
Description.
<typeStatus id="54A0882BFFB63B051730F9D3FED9FB01" box="[261,365,1213,1237]" pageId="11" pageNumber="382" type="holotype">Holotype</typeStatus>
female
</emphasis>
: Body generally pale ivory brown (
<figureCitation id="13202A0CFFB63B0516F6F9B3FE81FB21" box="[195,309,1245,1269]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="11" pageNumber="382">Fig. 1A, B</figureCitation>
); dorsal surface shining, impunctate, with sparsely distributed, pale, short, reclining setae. Head shiny creamy brown; basal carina on vertex reduced, flat; apex of clypeus and ventral half of buccula infuscate. Antenna dark brown, relatively short; antennomere I pale brown, with obscure basal half and darkened extreme apex; antennomere II pale basal half (except for dark base), shorter than basal width of pronotum; base of antennomere III pale. Labium pale reddish brown, reaching apex of metacoxa; apical ¼ of segment IV darkened. Pronotum polished, with a pair of small, circular spots on calli; collar about as thick as base of antennomere II; thoracic pleura milky brown; metathoracic scent efferent system as in
<figureCitation id="13202A0CFFB63B051784FB33FDA5F9A1" box="[433,529,1629,1653]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">Fig. 20I</figureCitation>
; scutellum shining, tinged with red at apex. Hemelytra with rather uniformly distributed, short, reclining setae; clavus and corium with dark, short, fragmentary stripes; paracuneus with a dark, small spot; apex of cuneus fuscous; membrane pale greyish brown, semi-transparent, with brown veins. All coxae and legs pale brown; metafemur with two brown, apical rings that are obliterated posteriad; tibia with small dark spots at bases of spines; apical part of each tarsomere III dark brown; pretarsal structure as in
<figureCitation id="13202A0CFFB63B05173DFA13FEDDF841" box="[264,361,1917,1941]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">Fig. 20K</figureCitation>
. Abdomen pale brown, speckled with reddish brown ventrally. Female genitalia (2FH, 20LO): Sclerotised rings ovoid, rather small, separated to each other mesally (
<figureCitation id="13202A0CFFB63B0516E6FAB3FE9AF821" box="[211,302,2013,2037]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="11" pageNumber="382">Figs. 2F</figureCitation>
,
<figureCitation id="13202A0CFFB63B05170CFAB3FEDCF821" box="[313,360,2013,2037]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">20L</figureCitation>
); Posterior wall with a row of spines on anterior part of interramal sclerite interrupted mesally (
<figureCitation id="13202A0CFFB63B05164CF573FF6EF7E1" box="[121,218,2077,2101]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="11" pageNumber="382">Figs. 2G</figureCitation>
,
<figureCitation id="13202A0CFFB63B0516D2F573FEAAF7E1" box="[231,286,2077,2101]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">20M</figureCitation>
); dorsal structure wide, semi-circular, with densely distributed spinules (
<figureCitation id="13202A0CFFB63B05125FFDD3FB45FF01" box="[1130,1265,189,213]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">Fig. 20N, O</figureCitation>
); interramal lobe small, subtriangular (
<figureCitation id="13202A0CFFB63B051226FDB3FBC0FF21" box="[1043,1140,221,245]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="11" pageNumber="382">Figs. 2G</figureCitation>
,
<figureCitation id="13202A0CFFB63B0512B4FDB3FB0CFF21" box="[1153,1208,221,245]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="11" pageNumber="382">20M</figureCitation>
).
<emphasis id="B96FEA9BFFB63B0512F9FDB3FAB2FF21" bold="true" box="[1228,1286,221,245]" pageId="11" pageNumber="382">Male</emphasis>
: Unknown.
</paragraph>
<paragraph id="8BA43689FFB63B051518FC73FBD8FEE1" blockId="11.[813,1132,285,309]" box="[813,1132,285,309]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FC73FC6FFEE1" bold="true" box="[813,987,285,309]" pageId="11" pageNumber="382">Measurements.</emphasis>
See
<tableCitation id="C6990332FFB63B051227FC73FBD2FEE1" box="[1042,1126,285,309]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="11" pageNumber="382">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051518FC33FB8EFE41" pageId="11" pageNumber="382" type="etymology">
<paragraph id="8BA43689FFB63B051518FC33FB8EFE41" blockId="11.[813,1451,349,405]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FC33FC1BFEA1" bold="true" box="[813,943,349,373]" pageId="11" pageNumber="382">Etymology.</emphasis>
Named for the
<typeStatus id="54A0882BFFB63B051255FC30FB3AFEA2" box="[1120,1166,350,374]" pageId="11" pageNumber="382">type</typeStatus>
locality, Tabanan District of
<collectingRegion id="49DFF86BFFB63B05157EFC13FCC3FE41" box="[843,887,381,405]" country="Indonesia" name="Bali" pageId="11" pageNumber="382">Bali</collectingRegion>
Island; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051518FCD3FB73FE01" box="[813,1223,445,469]" pageId="11" pageNumber="382" type="distribution">
<paragraph id="8BA43689FFB63B051518FCD3FB73FE01" blockId="11.[813,1223,445,469]" box="[813,1223,445,469]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FCD3FC0BFE01" bold="true" box="[813,959,445,469]" pageId="11" pageNumber="382">Distribution.</emphasis>
<collectingCountry id="F30C7619FFB63B0515F2FCD3FB9BFE01" box="[967,1071,445,469]" name="Indonesia" pageId="11" pageNumber="382">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFFB63B05120AFCD3FBD9FE01" box="[1087,1133,445,469]" country="Indonesia" name="Bali" pageId="11" pageNumber="382">Bali</collectingRegion>
Island).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB63B051518FC93FBB7FDC1" box="[813,1027,509,533]" pageId="11" pageNumber="382" type="biology_ecology">
<paragraph id="8BA43689FFB63B051518FC93FBB7FDC1" blockId="11.[813,1027,509,533]" box="[813,1027,509,533]" pageId="11" pageNumber="382">
<emphasis id="B96FEA9BFFB63B051518FC93FC3EFDC1" bold="true" box="[813,906,509,533]" pageId="11" pageNumber="382">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,866 @@
<document id="ABF9B158D1EFC4A6335510960B8511B8" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB93B0815FFF813FBADF86A" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus cheroti Yasunaga, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="377" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="375" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CD22721AC380519EB533B0D6A491C7F2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="210F705BCC3E1FD8725225BBEB35F09F">
<mods:title id="D54E061837DDDB1549FEC6583BD08D8F">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="20A64F0894104E3C150AEB56EE847FF1" type="personal">
<mods:role id="1A2728234663D4F4F815530ED250BA1B">
<mods:roleTerm id="6D43F7118AEBA2D21F459F18451FB89A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C8EBD3E41667E9E9397A943CE11F847">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="EBDBF09047A5108DF5338801CC6C6349">text</mods:typeOfResource>
<mods:relatedItem id="59D3120C615F2A7E296E3CBCE3043B42" type="host">
<mods:titleInfo id="C84100DFD6C0818B766830AB44FDDB07">
<mods:title id="31A19698045328831966D710EE2FC6B0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D1C82F93101EBB1C205A05F71EA5E4D2">
<mods:date id="169660C8D90F91EB3242D8E925E6682E">2024</mods:date>
<mods:detail id="851A429EFF088786FD473B1DA818E53D" type="pubDate">
<mods:number id="8E9766669339CF4534C389270C1A4B4C">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="7844C29299871E91649A1424AF28FD15" type="volume">
<mods:number id="47D09FE20535F1BF629BB2E8CFBA1116">72</mods:number>
</mods:detail>
<mods:extent id="1DEED80B441405D4D1B8F7B5DE8DD605" unit="page">
<mods:start id="877384A1BCDF38BBFE6013700A291420">371</mods:start>
<mods:end id="14D69336DE9CE1CB564BE96066442846">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F71C920280165C9FE16D58FB05CB020F">journal article</mods:classification>
<mods:identifier id="78706E8757D4F0A86CFD4D37BB87424D" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="E5F9CDFE461D77EF8CB2B36FFA9B33AF" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E7C01D8903E827F331A715D57D4F1B7A" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="5299766D8AD1A7A8598637F3EDE56C83" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB93B0815FFF813FBADF86A" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0815FFF813FBADF86A" httpUri="http://treatment.plazi.org/id/03B2879FFFB93B0815FFF813FBADF86A" lastPageId="6" lastPageNumber="377" pageId="4" pageNumber="375">
<subSubSection id="C3016502FFB93B0A15FFF813FA88FA41" box="[970,1340,1405,1429]" pageId="4" pageNumber="375" type="nomenclature">
<paragraph id="8BA43689FFB93B0A15FFF813FA88FA41" blockId="4.[970,1340,1405,1461]" box="[970,1340,1405,1429]" pageId="4" pageNumber="375">
<heading id="D0EC81E5FFB93B0A15FFF813FA88FA41" bold="true" box="[970,1340,1405,1429]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="375" reason="2">
<emphasis id="B96FEA9BFFB93B0A15FFF813FA88FA41" bold="true" box="[970,1340,1405,1429]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A15FFF813FB1AFA41" bold="true" box="[970,1198,1405,1429]" italics="true" pageId="4" pageNumber="375">
<taxonomicName id="4C1B4D0AFFB93B0A15FFF813FB1CFA41" authorityName="Yasunaga" authorityYear="2024" box="[970,1192,1405,1429]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="cheroti" status="sp. nov.">Chelyolygus cheroti</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A25C57E0FFB93B0A1283F813FA88FA41" box="[1206,1340,1405,1429]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1235F8F3FAB1FA61" box="[1024,1285,1437,1461]" pageId="4" pageNumber="375" type="description">
<paragraph id="8BA43689FFB93B0A1235F8F3FAB1FA61" blockId="4.[970,1340,1405,1461]" box="[1024,1285,1437,1461]" pageId="4" pageNumber="375">
(
<figureCitation id="13202A0CFFB93B0A123DF8F3FBFBFA61" box="[1032,1103,1437,1461]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Figs. 1</figureCitation>
CF, 3GL, 19)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1576F8B3FB20F9A1" pageId="4" pageNumber="375" type="materials_examined">
<paragraph id="8BA43689FFB93B0A1576F8B3FB20F9A1" blockId="4.[835,1474,1501,1653]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576F8B3FC4FFA21" bold="true" box="[835,1019,1501,1525]" pageId="4" pageNumber="375">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB93B0A123DF8B3FA09F9E1" collectingDate="1996-08-08" collectingDateMax="1996-08-12" collectingDateMin="1996-08-08" collectionCode="ZRC, AMNH" country="Indonesia" latitude="-8.266666" location="Bedugul Area" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" pageId="4" pageNumber="375" specimenCode="PBI 00378776" specimenCount="1" stateProvince="Bali" typeStatus="holotype">
<emphasis id="B96FEA9BFFB93B0A123DF8B3FBC3FA21" bold="true" box="[1032,1143,1501,1525]" pageId="4" pageNumber="375">
<typeStatus id="54A0882BFFB93B0A123DF8B3FBC3FA21" box="[1032,1143,1501,1525]" pageId="4" pageNumber="375" type="holotype">Holotype</typeStatus>
</emphasis>
: male,
<emphasis id="B96FEA9BFFB93B0A12E0F8B3FACBFA21" bold="true" box="[1237,1407,1501,1525]" pageId="4" pageNumber="375">
<collectingCountry id="F30C7619FFB93B0A12E0F8B3FACDFA21" box="[1237,1401,1501,1525]" name="Indonesia" pageId="4" pageNumber="375">INDONESIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49DFF86BFFB93B0A13BEF8B3FA09FA21" box="[1419,1469,1501,1525]" country="Indonesia" name="Bali" pageId="4" pageNumber="375">Bali</collectingRegion>
,
<collectingMunicipality id="6BC0ACF3FFB93B0A1576F893FC16F9C1" box="[835,930,1533,1557]" pageId="4" pageNumber="375">Tabanan</collectingMunicipality>
,
<location id="8EC46052FFB93B0A159AF893FBFEF9C2" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0815FFF813FBADF86A:8EC46052FFB93B0A159AF893FBFEF9C2" box="[943,1098,1533,1558]" country="Indonesia" latitude="-8.266666" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" name="Bedugul Area" pageId="4" pageNumber="375" stateProvince="Bali">Bedugul Area</location>
,
<geoCoordinate id="EE2F504EFFB93B0A1263F892FB14F9C1" box="[1110,1184,1532,1557]" degrees="8" direction="south" minutes="16" orientation="latitude" pageId="4" pageNumber="375" precision="925" value="-8.266666">8°16S</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB93B0A129BF892FAA0F9C1" box="[1198,1300,1532,1557]" degrees="115" direction="east" minutes="09" orientation="longitude" pageId="4" pageNumber="375" precision="925" value="115.15">115°09E</geoCoordinate>
, local collector,
<date id="FFA51049FFB93B0A1576FB73FB93F9E1" box="[835,1063,1565,1590]" pageId="4" pageNumber="375" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">
<collectingDate id="EFE1E9A1FFB93B0A1576FB73FB93F9E1" box="[835,1063,1565,1590]" pageId="4" pageNumber="375" value="1996-08-08" valueMax="1996-08-12" valueMin="1996-08-08">812 August 1996</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB93B0A1208FB73FBCDF9E1" box="[1085,1145,1565,1589]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="375">ZRC</collectionCode>
) (
<collectionCode id="ED0AAE4CFFB93B0A12A3FB73FB44F9E1" box="[1174,1264,1565,1589]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="375">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB93B0A12CAFB73FA01F9E1" box="[1279,1461,1565,1589]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="375">PBI 00378776</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB93B0A1576FB50FB3BF9A1" collectingDate="1996-08-08" collectingDateMax="1996-08-12" collectingDateMin="1996-08-08" collectionCode="AMNH, TYCN" country="Indonesia" latitude="-8.266666" location="Bedugul Area" longLatPrecision="1301" longitude="115.15" municipality="Tabanan" pageId="4" pageNumber="375" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Bali" typeStatus="paratype">
<typeStatus id="54A0882BFFB93B0A1576FB50FC1BF982" box="[835,943,1598,1622]" pageId="4" pageNumber="375" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B96FEA9BFFB93B0A1589FB53FBE7F981" bold="true" box="[956,1107,1597,1621]" pageId="4" pageNumber="375">
<collectingCountry id="F30C7619FFB93B0A1589FB53FBE7F981" box="[956,1107,1597,1621]" name="Indonesia" pageId="4" pageNumber="375">INDONESIA</collectingCountry>
</emphasis>
:
<specimenCount id="9D1DFD00FFB93B0A1256FB53FB0FF981" box="[1123,1211,1597,1621]" count="2" pageId="4" pageNumber="375" type="male">2 males</specimenCount>
,
<specimenCount id="9D1DFD00FFB93B0A12FDFB53FA9DF981" box="[1224,1321,1597,1621]" count="1" pageId="4" pageNumber="375" type="female">1 female</specimenCount>
, same data as for holotype (
<collectionCode id="ED0AAE4CFFB93B0A15EDFB33FB9AF9A1" box="[984,1070,1629,1653]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="375">AMNH</collectionCode>
,
<collectionCode id="ED0AAE4CFFB93B0A120EFB33FB3CF9A1" box="[1083,1160,1629,1653]" pageId="4" pageNumber="375">TYCN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0B1576FBF3FEA0FED8" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375" type="diagnosis">
<paragraph id="8BA43689FFB93B0A1576FBF3FBF8F7C1" blockId="4.[835,1475,1693,2069]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FBF3FC0CF961" bold="true" box="[835,952,1693,1717]" pageId="4" pageNumber="375">Diagnosis.</emphasis>
This new species is recognised primarily by the following characters: Dark general colouration (
<figureCitation id="13202A0CFFB93B0A1370FBD3FA01F901" box="[1349,1461,1725,1749]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1CF</figureCitation>
); medium size; widely darkened apical parts of antennomeres I and II; dark mesal stripe on scutellum; widely fuscous hemelytra; partly striped anterior clavus; distinctly bifurcate hypophysis of right paramere (
<figureCitation id="13202A0CFFB93B0A12A2FA53FB59F881" box="[1175,1261,1853,1877]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="4" pageNumber="375">Fig. 3H</figureCitation>
) with a thumb-like process at apex of sensory lobe (
<figureCitation id="13202A0CFFB93B0A128AFA33FAABF8A1" box="[1215,1311,1885,1909]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="4" pageNumber="375">Fig. 17J</figureCitation>
); two slender, weakly curved spicules on vesica; ovoid, rather enlarged, thick-rimmed sclerotised rings relatively contiguous to each other mesally (
<figureCitation id="13202A0CFFB93B0A15DDFAD3FBFDF801" box="[1000,1097,1981,2005]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="4" pageNumber="375">Figs. 3K</figureCitation>
,
<figureCitation id="13202A0CFFB93B0A1262FAD3FB39F801" box="[1111,1165,1981,2005]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="4" pageNumber="375">13M</figureCitation>
); wide, flat dorsal structure; and relatively wide, subtriangular interramal lobe on posterior wall (
<figureCitation id="13202A0CFFB93B0A15B7FA93FC54F7C1" box="[898,992,2045,2069]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="4" pageNumber="375">Figs. 3L</figureCitation>
,
<figureCitation id="13202A0CFFB93B0A15D8FA93FB8BF7C1" box="[1005,1087,2045,2069]" captionStart="Fig" captionStartId="22.[135,170,2004,2026]" captionTargetBox="[135,1473,258,1989]" captionTargetId="figure-7@22.[135,1473,249,1989]" captionTargetPageId="22" captionText="Fig. 13. Male (AG) and female (HI) genitalia of Himalisundacoris duwalae, new species. AC, left paramere; D, E, right paramere; F, vesica; G, ditto, secondary gonopore and adjacent structures; H, posterior wall; I, genital chamber. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682980" httpUri="https://zenodo.org/record/14682980/files/figure.png" pageId="4" pageNumber="375">13NO</figureCitation>
).
</paragraph>
<caption id="DF646601FFB83B0B1644FDD3FA18FF07" box="[113,1452,189,211]" pageId="5" pageNumber="376" startId="5.[113,168,189,211]" targetType="table">
<paragraph id="8BA43689FFB83B0B1644FDD3FA18FF07" blockId="5.[113,1452,189,268]" box="[113,1452,189,211]" pageId="5" pageNumber="376">Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:</paragraph>
</caption>
<paragraph id="8BA43689FFB83B0B1644FDB4FEA0FED8" blockId="5.[113,1452,189,268]" pageId="5" pageNumber="376">
male; PRN: pronotum, PA:
<typeStatus id="54A0882BFFB83B0B17B2FDB5FE68FF24" box="[391,476,219,240]" pageId="5" pageNumber="376" type="paratype">paratype</typeStatus>
, TB: tibia, VTX: vertex (interocular space), W: width. Bold letters showing new taxa; *shrinking antennomere IV.
</paragraph>
</subSubSection>
<paragraph id="8BA43689FFB83B0B1435FC5BFA2FF7CE" pageId="5" pageNumber="376">
<table id="F91BC429FFB8C4F116B5FC5BFA2FF7CE" box="[128,1435,309,2074]" colsContinueIn="6.[151,1459,203,1175]" gridcols="14" gridrows="16" pageId="5" pageNumber="376">
<tr id="352B34CBFFB8C4F116B5FC5BFA2FFE9E" box="[128,1435,309,330]" gridrow="0" pageId="5" pageNumber="376" rowspan-0="1" rowspan-1="1">
<th id="76FA5DB7FFB8C4F11435FC5BFD81FE9E" box="[512,565,309,330]" gridcol="2" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1435FC5BFD81FE9E" bold="true" box="[512,565,309,330]" pageId="5" pageNumber="376">Body</emphasis>
</th>
<th id="76FA5DB7FFB8C4F11460FC5BFD38FE9E" box="[597,652,309,330]" gridcol="3" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1460FC5BFD38FE9E" bold="true" box="[597,652,309,330]" pageId="5" pageNumber="376">Head</emphasis>
</th>
<th id="76FA5DB7FFB8C4F1149EFC5BFD6AFE9E" box="[683,734,309,330]" gridcol="4" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B149EFC5BFD6AFE9E" bold="true" box="[683,734,309,330]" pageId="5" pageNumber="376">VTX</emphasis>
</th>
<th id="76FA5DB7FFB8C4F114CAFC5BFC84FE9E" box="[767,816,309,330]" gridcol="5" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B14CAFC5BFC84FE9E" bold="true" box="[767,816,309,330]" pageId="5" pageNumber="376">PRN</emphasis>
</th>
<th id="76FA5DB7FFB8C4F11567FC5BFC35FE9E" box="[850,897,309,330]" gridcol="6" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1567FC5BFC35FE9E" bold="true" box="[850,897,309,330]" pageId="5" pageNumber="376">Max</emphasis>
</th>
<th id="76FA5DB7FFB8C4F11595FC5BFB1EFE9E" box="[928,1194,309,330]" colspan="4" colspanRight="3" gridcol="7" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B15E3FC5BFBCEFE9E" bold="true" box="[982,1146,309,330]" pageId="5" pageNumber="376">Antennomere L</emphasis>
</th>
<th id="76FA5DB7FFB8C4F112FAFC5BFAB2FE9E" box="[1231,1286,309,330]" gridcol="11" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B12FAFC5BFAB2FE9E" bold="true" box="[1231,1286,309,330]" pageId="5" pageNumber="376">LBM</emphasis>
</th>
<th id="76FA5DB7FFB8C4F11312FC5BFA2FFE9E" box="[1319,1435,309,330]" colspan="2" colspanRight="1" gridcol="12" gridrow="0" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B131EFC5BFA21FE9E" bold="true" box="[1323,1429,309,330]" pageId="5" pageNumber="376">Metaleg L</emphasis>
</th>
</tr>
<tr id="352B34CBFFB8C4F116B5FC0AFA2FFEAD" box="[128,1435,356,377]" gridrow="1" pageId="5" pageNumber="376" rowspan-0="1" rowspan-1="1">
<td id="76FA5DB7FFB8C4F11435FC0AFD81FEAD" box="[512,565,356,377]" gridcol="2" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1426FC0AFD97FEAD" bold="true" box="[531,547,356,377]" pageId="5" pageNumber="376">L</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11460FC0AFD38FEAD" box="[597,652,356,377]" gridcol="3" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1450FC0AFDC9FEAD" bold="true" box="[613,637,356,377]" pageId="5" pageNumber="376">W</emphasis>
</td>
<td id="76FA5DB7FFB8C4F1149EFC0AFD6AFEAD" box="[683,734,356,377]" gridcol="4" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B148CFC0AFD65FEAD" bold="true" box="[697,721,356,377]" pageId="5" pageNumber="376">W</emphasis>
</td>
<td id="76FA5DB7FFB8C4F114CAFC0AFC84FEAD" box="[767,816,356,377]" gridcol="5" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1539FC0AFC90FEAD" bold="true" box="[780,804,356,377]" pageId="5" pageNumber="376">W</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11567FC0AFC35FEAD" box="[850,897,356,377]" gridcol="6" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1568FC0AFCC1FEAD" bold="true" box="[861,885,356,377]" pageId="5" pageNumber="376">W</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11595FC0AFC7EFEAD" box="[928,970,356,377]" gridcol="7" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1585FC0AFC0DFEAD" bold="true" box="[944,953,356,377]" pageId="5" pageNumber="376">I</emphasis>
</td>
<td id="76FA5DB7FFB8C4F115DDFC0AFBA6FEAD" box="[1000,1042,356,377]" gridcol="8" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B15C6FC0AFBB2FEAD" bold="true" box="[1011,1030,356,377]" pageId="5" pageNumber="376">II</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11205FC0AFBEEFEAD" box="[1072,1114,356,377]" gridcol="9" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1202FC0AFBE7FEAD" bold="true" box="[1079,1107,356,377]" pageId="5" pageNumber="376">III</emphasis>
</td>
<td id="76FA5DB7FFB8C4F112B5FC0AFB1EFEAD" box="[1152,1194,356,377]" gridcol="10" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B12B2FC0AFB16FEAD" bold="true" box="[1159,1186,356,377]" pageId="5" pageNumber="376">IV</emphasis>
</td>
<td id="76FA5DB7FFB8C4F112FAFC0AFAB2FEAD" box="[1231,1286,356,377]" gridcol="11" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B12D6FC0AFB47FEAD" bold="true" box="[1251,1267,356,377]" pageId="5" pageNumber="376">L</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11312FC0AFAE5FEAD" box="[1319,1361,356,377]" gridcol="12" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B131CFC0AFAFAFEAD" bold="true" box="[1321,1358,356,377]" pageId="5" pageNumber="376">FM</emphasis>
</td>
<td id="76FA5DB7FFB8C4F11344FC0AFA2FFEAD" box="[1393,1435,356,377]" gridcol="13" gridrow="1" pageId="5" pageNumber="376">
<emphasis id="B96FEA9BFFB83B0B1343FC0AFA22FEAD" bold="true" box="[1398,1430,356,377]" pageId="5" pageNumber="376">TB</emphasis>
</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FCF4FA2FFE24" box="[128,1435,410,496]" gridrow="2" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FCF4FEF6FE24" box="[128,322,410,496]" gridcol="0" gridrow="2" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FCF4FEF6FE7B" authorityName="Yasunaga" authorityYear="2024" box="[128,322,410,431]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="cheroti">
<emphasis id="B96FEA9BFFB83B0B16B5FCF4FEF6FE7B" box="[128,322,410,431]" italics="true" pageId="5" pageNumber="376">Chelyolygus cheroti</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FCF4FE67FE24" box="[403,467,410,496]" gridcol="1" gridrow="2" pageId="5" pageNumber="376">HO-M PA-M PA-M</td>
<td id="76FA5DB7FFB8C4F11435FCF4FD81FE24" box="[512,565,410,496]" gridcol="2" gridrow="2" pageId="5" pageNumber="376">4.17 3.87 4.41</td>
<td id="76FA5DB7FFB8C4F11460FCF4FD38FE24" box="[597,652,410,496]" gridcol="3" gridrow="2" pageId="5" pageNumber="376">1.03 1.01 1.02</td>
<td id="76FA5DB7FFB8C4F1149EFCF4FD6AFE24" box="[683,734,410,496]" gridcol="4" gridrow="2" pageId="5" pageNumber="376">0.32 0.30 0.32</td>
<td id="76FA5DB7FFB8C4F114CAFCF4FC84FE24" box="[767,816,410,496]" gridcol="5" gridrow="2" pageId="5" pageNumber="376">1.64 1.53 1.52</td>
<td id="76FA5DB7FFB8C4F11567FCF4FC35FE24" box="[850,897,410,496]" gridcol="6" gridrow="2" pageId="5" pageNumber="376">1.94 2.12 1.95</td>
<td id="76FA5DB7FFB8C4F11595FCF4FC7EFE24" box="[928,970,410,496]" gridcol="7" gridrow="2" pageId="5" pageNumber="376">0.62 0.63 0.62</td>
<td id="76FA5DB7FFB8C4F115DDFCF4FBA6FE24" box="[1000,1042,410,496]" gridcol="8" gridrow="2" pageId="5" pageNumber="376">1.79 1.80 1.80</td>
<td id="76FA5DB7FFB8C4F11205FCF4FBEEFE24" box="[1072,1114,410,496]" gridcol="9" gridrow="2" pageId="5" pageNumber="376">0.89 0.75 0.87</td>
<td id="76FA5DB7FFB8C4F112B5FCF4FB1EFE24" box="[1152,1194,410,496]" gridcol="10" gridrow="2" pageId="5" pageNumber="376">0.62 0.63 0.65</td>
<td id="76FA5DB7FFB8C4F112FAFCF4FAB2FE24" box="[1231,1286,410,496]" gridcol="11" gridrow="2" pageId="5" pageNumber="376">1.95 1.98 2.15</td>
<td id="76FA5DB7FFB8C4F11312FCF4FAE5FE24" box="[1319,1361,410,496]" gridcol="12" gridrow="2" pageId="5" pageNumber="376">1.67 1.61 1.67</td>
<td id="76FA5DB7FFB8C4F11344FCF4FA2FFE24" box="[1393,1435,410,496]" gridcol="13" gridrow="2" pageId="5" pageNumber="376">2.40 2.30 2.37</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FF6FFA2FFDC3" box="[128,1435,513,535]" gridrow="3" pageId="5" pageNumber="376" rowspan-0="1">
<td id="76FA5DB7FFB8C4F117A6FF6FFE67FDC3" box="[403,467,513,535]" gridcol="1" gridrow="3" pageId="5" pageNumber="376">PA-F</td>
<td id="76FA5DB7FFB8C4F11435FF6FFD81FDC3" box="[512,565,513,535]" gridcol="2" gridrow="3" pageId="5" pageNumber="376">4.66</td>
<td id="76FA5DB7FFB8C4F11460FF6FFD38FDC3" box="[597,652,513,535]" gridcol="3" gridrow="3" pageId="5" pageNumber="376">1.05</td>
<td id="76FA5DB7FFB8C4F1149EFF6FFD6AFDC3" box="[683,734,513,535]" gridcol="4" gridrow="3" pageId="5" pageNumber="376">0.41</td>
<td id="76FA5DB7FFB8C4F114CAFF6FFC84FDC3" box="[767,816,513,535]" gridcol="5" gridrow="3" pageId="5" pageNumber="376">1.79</td>
<td id="76FA5DB7FFB8C4F11567FF6FFC35FDC3" box="[850,897,513,535]" gridcol="6" gridrow="3" pageId="5" pageNumber="376">2.22</td>
<td id="76FA5DB7FFB8C4F11595FF6FFC7EFDC3" box="[928,970,513,535]" gridcol="7" gridrow="3" pageId="5" pageNumber="376">0.62</td>
<td id="76FA5DB7FFB8C4F115DDFF6FFBA6FDC3" box="[1000,1042,513,535]" gridcol="8" gridrow="3" pageId="5" pageNumber="376">1.74</td>
<td id="76FA5DB7FFB8C4F11205FF6FFBEEFDC3" box="[1072,1114,513,535]" gridcol="9" gridrow="3" pageId="5" pageNumber="376">0.90</td>
<td id="76FA5DB7FFB8C4F112B5FF6FFB1EFDC3" box="[1152,1194,513,535]" gridcol="10" gridrow="3" pageId="5" pageNumber="376">0.72</td>
<td id="76FA5DB7FFB8C4F112FAFF6FFAB2FDC3" box="[1231,1286,513,535]" gridcol="11" gridrow="3" pageId="5" pageNumber="376">1.97</td>
<td id="76FA5DB7FFB8C4F11312FF6FFAE5FDC3" box="[1319,1361,513,535]" gridcol="12" gridrow="3" pageId="5" pageNumber="376">1.65</td>
<td id="76FA5DB7FFB8C4F11344FF6FFA2FFDC3" box="[1393,1435,513,535]" gridcol="13" gridrow="3" pageId="5" pageNumber="376">2.55</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FF46FA2FFD69" box="[128,1435,552,701]" gridrow="4" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FF46FEF6FD69" box="[128,322,552,701]" gridcol="0" gridrow="4" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FF47FF41FDEA" authorityName="Yasunaga" authorityYear="2024" box="[128,245,553,574]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="parsanus">
<emphasis id="B96FEA9BFFB83B0B16B5FF47FF41FDEA" box="[128,245,553,574]" italics="true" pageId="5" pageNumber="376">C. parsanus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FF46FE67FD69" box="[403,467,552,701]" gridcol="1" gridrow="4" pageId="5" pageNumber="376">M M F F F</td>
<td id="76FA5DB7FFB8C4F11435FF46FD81FD69" box="[512,565,552,701]" gridcol="2" gridrow="4" pageId="5" pageNumber="376">3.72 3.80 3.87 3.99 4.09</td>
<td id="76FA5DB7FFB8C4F11460FF46FD38FD69" box="[597,652,552,701]" gridcol="3" gridrow="4" pageId="5" pageNumber="376">0.92 0.96 0.98 0.96 0.96</td>
<td id="76FA5DB7FFB8C4F1149EFF46FD6AFD69" box="[683,734,552,701]" gridcol="4" gridrow="4" pageId="5" pageNumber="376">0.26 0.29 0.32 0.33 0.32</td>
<td id="76FA5DB7FFB8C4F114CAFF46FC84FD69" box="[767,816,552,701]" gridcol="5" gridrow="4" pageId="5" pageNumber="376">1.41 1.50 1.64 1.52 1.50</td>
<td id="76FA5DB7FFB8C4F11567FF46FC35FD69" box="[850,897,552,701]" gridcol="6" gridrow="4" pageId="5" pageNumber="376">1.67 1.73 1.77 1.73 1.82</td>
<td id="76FA5DB7FFB8C4F11595FF46FC7EFD69" box="[928,970,552,701]" gridcol="7" gridrow="4" pageId="5" pageNumber="376">0.45 0.48 0.48 0.48 0.45</td>
<td id="76FA5DB7FFB8C4F115DDFF46FBA6FD69" box="[1000,1042,552,701]" gridcol="8" gridrow="4" pageId="5" pageNumber="376">1.47 1.62 1.40 1.41 1.35</td>
<td id="76FA5DB7FFB8C4F11205FF46FBEEFD69" box="[1072,1114,552,701]" gridcol="9" gridrow="4" pageId="5" pageNumber="376">0.78 0.92 0.75</td>
<td id="76FA5DB7FFB8C4F112B5FF46FB1EFD69" box="[1152,1194,552,701]" gridcol="10" gridrow="4" pageId="5" pageNumber="376"> 0.65 0.48</td>
<td id="76FA5DB7FFB8C4F112FAFF46FAB2FD69" box="[1231,1286,552,701]" gridcol="11" gridrow="4" pageId="5" pageNumber="376">1.35 1.35 1.35 1.40 1.38</td>
<td id="76FA5DB7FFB8C4F11312FF46FAE5FD69" box="[1319,1361,552,701]" gridcol="12" gridrow="4" pageId="5" pageNumber="376">1.35 1.62 1.52 1.50</td>
<td id="76FA5DB7FFB8C4F11344FF46FA2FFD69" box="[1393,1435,552,701]" gridcol="13" gridrow="4" pageId="5" pageNumber="376">2.10 2.25 2.12 2.10</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FFB8FA2FFD38" box="[128,1435,726,748]" gridrow="5" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FFB8FEF6FD38" box="[128,322,726,748]" gridcol="0" gridrow="5" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FFB9FEA6FD3F" authorityName="Yasunaga" authorityYear="2024" box="[128,274,726,748]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="tabananicus">
<emphasis id="B96FEA9BFFB83B0B16B5FFB9FEA6FD3F" box="[128,274,726,748]" italics="true" pageId="5" pageNumber="376">C. tabananicus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FFB8FE67FD38" box="[403,467,726,748]" gridcol="1" gridrow="5" pageId="5" pageNumber="376">HO-F</td>
<td id="76FA5DB7FFB8C4F11435FFB8FD81FD38" box="[512,565,726,748]" gridcol="2" gridrow="5" pageId="5" pageNumber="376">4.29</td>
<td id="76FA5DB7FFB8C4F11460FFB8FD38FD38" box="[597,652,726,748]" gridcol="3" gridrow="5" pageId="5" pageNumber="376">1.07</td>
<td id="76FA5DB7FFB8C4F1149EFFB8FD6AFD38" box="[683,734,726,748]" gridcol="4" gridrow="5" pageId="5" pageNumber="376">0.38</td>
<td id="76FA5DB7FFB8C4F114CAFFB8FC84FD38" box="[767,816,726,748]" gridcol="5" gridrow="5" pageId="5" pageNumber="376">1.80</td>
<td id="76FA5DB7FFB8C4F11567FFB8FC35FD38" box="[850,897,726,748]" gridcol="6" gridrow="5" pageId="5" pageNumber="376">2.24</td>
<td id="76FA5DB7FFB8C4F11595FFB8FC7EFD38" box="[928,970,726,748]" gridcol="7" gridrow="5" pageId="5" pageNumber="376">0.53</td>
<td id="76FA5DB7FFB8C4F115DDFFB8FBA6FD38" box="[1000,1042,726,748]" gridcol="8" gridrow="5" pageId="5" pageNumber="376">1.56</td>
<td id="76FA5DB7FFB8C4F11205FFB8FBEEFD38" box="[1072,1114,726,748]" gridcol="9" gridrow="5" pageId="5" pageNumber="376">0.87</td>
<td id="76FA5DB7FFB8C4F112B5FFB8FB1EFD38" box="[1152,1194,726,748]" gridcol="10" gridrow="5" pageId="5" pageNumber="376">0.56</td>
<td id="76FA5DB7FFB8C4F112FAFFB8FAB2FD38" box="[1231,1286,726,748]" gridcol="11" gridrow="5" pageId="5" pageNumber="376">1.67</td>
<td id="76FA5DB7FFB8C4F11312FFB8FAE5FD38" box="[1319,1361,726,748]" gridcol="12" gridrow="5" pageId="5" pageNumber="376">1.65</td>
<td id="76FA5DB7FFB8C4F11344FFB8FA2FFD38" box="[1393,1435,726,748]" gridcol="13" gridrow="5" pageId="5" pageNumber="376">2.48</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FE6BFA2FFCEF" box="[128,1435,773,827]" gridrow="6" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FE6BFEF6FCEF" box="[128,322,773,827]" gridcol="0" gridrow="6" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FE68FEA2FCCE" authorityName="Yasunaga" authorityYear="2024" box="[128,278,773,795]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="virgulatellus">
<emphasis id="B96FEA9BFFB83B0B16B5FE68FEA2FCCE" box="[128,278,773,795]" italics="true" pageId="5" pageNumber="376">C. virgulatellus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FE6BFE67FCEF" box="[403,467,773,827]" gridcol="1" gridrow="6" pageId="5" pageNumber="376">HO-M PA-M</td>
<td id="76FA5DB7FFB8C4F11435FE6BFD81FCEF" box="[512,565,773,827]" gridcol="2" gridrow="6" pageId="5" pageNumber="376">3.45 3.43</td>
<td id="76FA5DB7FFB8C4F11460FE6BFD38FCEF" box="[597,652,773,827]" gridcol="3" gridrow="6" pageId="5" pageNumber="376">0.96 0.95</td>
<td id="76FA5DB7FFB8C4F1149EFE6BFD6AFCEF" box="[683,734,773,827]" gridcol="4" gridrow="6" pageId="5" pageNumber="376">0.30 0.29</td>
<td id="76FA5DB7FFB8C4F114CAFE6BFC84FCEF" box="[767,816,773,827]" gridcol="5" gridrow="6" pageId="5" pageNumber="376">1.49 1.49</td>
<td id="76FA5DB7FFB8C4F11567FE6BFC35FCEF" box="[850,897,773,827]" gridcol="6" gridrow="6" pageId="5" pageNumber="376">1.73 1.67</td>
<td id="76FA5DB7FFB8C4F11595FE6BFC7EFCEF" box="[928,970,773,827]" gridcol="7" gridrow="6" pageId="5" pageNumber="376">0.53 0.53</td>
<td id="76FA5DB7FFB8C4F115DDFE6BFBA6FCEF" box="[1000,1042,773,827]" gridcol="8" gridrow="6" pageId="5" pageNumber="376">1.47 1.47</td>
<td id="76FA5DB7FFB8C4F11205FE6BFBEEFCEF" box="[1072,1114,773,827]" gridcol="9" gridrow="6" pageId="5" pageNumber="376">0.86 0.80</td>
<td id="76FA5DB7FFB8C4F112B5FE6BFB1EFCEF" box="[1152,1194,773,827]" gridcol="10" gridrow="6" pageId="5" pageNumber="376">0.48 0.48</td>
<td id="76FA5DB7FFB8C4F112FAFE6BFAB2FCEF" box="[1231,1286,773,827]" gridcol="11" gridrow="6" pageId="5" pageNumber="376">1.46 1.47</td>
<td id="76FA5DB7FFB8C4F11312FE6BFAE5FCEF" box="[1319,1361,773,827]" gridcol="12" gridrow="6" pageId="5" pageNumber="376">1.49 1.49</td>
<td id="76FA5DB7FFB8C4F11344FE6BFA2FFCEF" box="[1393,1435,773,827]" gridcol="13" gridrow="6" pageId="5" pageNumber="376">1.97 2.07</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FE3AFA2FFBDC" box="[128,1435,852,1032]" gridrow="7" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FE3AFEF6FBDC" box="[128,322,852,1032]" gridcol="0" gridrow="7" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FE3AFEB9FCBD" authorityName="Yasunaga" authorityYear="2024" box="[128,269,852,873]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="wolskiorum">
<emphasis id="B96FEA9BFFB83B0B16B5FE3AFEB9FCBD" box="[128,269,852,873]" italics="true" pageId="5" pageNumber="376">C. wolskiorum</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FE3AFE67FBDC" box="[403,467,852,1032]" gridcol="1" gridrow="7" pageId="5" pageNumber="376">HO-M PA-M PA-M PA-F PA-F PA-F</td>
<td id="76FA5DB7FFB8C4F11435FE3AFD81FBDC" box="[512,565,852,1032]" gridcol="2" gridrow="7" pageId="5" pageNumber="376">3.19 3.50 3.19 4.14 3.43 4.17</td>
<td id="76FA5DB7FFB8C4F11460FE3AFD38FBDC" box="[597,652,852,1032]" gridcol="3" gridrow="7" pageId="5" pageNumber="376">0.98 0.96 0.93 0.99 0.95 0.99</td>
<td id="76FA5DB7FFB8C4F1149EFE3AFD6AFBDC" box="[683,734,852,1032]" gridcol="4" gridrow="7" pageId="5" pageNumber="376">0.26 0.26 0.24 0.33 0.30 0.32</td>
<td id="76FA5DB7FFB8C4F114CAFE3AFC84FBDC" box="[767,816,852,1032]" gridcol="5" gridrow="7" pageId="5" pageNumber="376">1.37 1.37 1.46 1.67 1.50 1.65</td>
<td id="76FA5DB7FFB8C4F11567FE3AFC35FBDC" box="[850,897,852,1032]" gridcol="6" gridrow="7" pageId="5" pageNumber="376">1.55 1.53 1.64 1.86 1.82 1.88</td>
<td id="76FA5DB7FFB8C4F11595FE3AFC7EFBDC" box="[928,970,852,1032]" gridcol="7" gridrow="7" pageId="5" pageNumber="376">0.47 0.48 0.47 0.51 0.45 0.47</td>
<td id="76FA5DB7FFB8C4F115DDFE3AFBA6FBDC" box="[1000,1042,852,1032]" gridcol="8" gridrow="7" pageId="5" pageNumber="376">1.50 1.49 1.47 1.56 1.37 1.40</td>
<td id="76FA5DB7FFB8C4F11205FE3AFBEEFBDC" box="[1072,1114,852,1032]" gridcol="9" gridrow="7" pageId="5" pageNumber="376">0.81 0.87 0.89 0.81 0.83</td>
<td id="76FA5DB7FFB8C4F112B5FE3AFB1EFBDC" box="[1152,1194,852,1032]" gridcol="10" gridrow="7" pageId="5" pageNumber="376">0.59 0.54 </td>
<td id="76FA5DB7FFB8C4F112FAFE3AFAB2FBDC" box="[1231,1286,852,1032]" gridcol="11" gridrow="7" pageId="5" pageNumber="376">1.44 1.37 1.37 1.65 1.49 1.44</td>
<td id="76FA5DB7FFB8C4F11312FE3AFAE5FBDC" box="[1319,1361,852,1032]" gridcol="12" gridrow="7" pageId="5" pageNumber="376">1.47 1.47 1.38 1.55 1.47 1.58</td>
<td id="76FA5DB7FFB8C4F11344FE3AFA2FFBDC" box="[1393,1435,852,1032]" gridcol="13" gridrow="7" pageId="5" pageNumber="376">2.06 2.10 2.07 2.10 2.01 2.13</td>
</tr>
<tr id="352B34CBFFB8C4F116B5F94FFA2FFBE3" box="[128,1435,1057,1079]" gridrow="8" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5F94FFEF6FBE3" box="[128,322,1057,1079]" gridcol="0" gridrow="8" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5F94CFE94FBE2" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[128,288,1057,1079]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="bipuncticollis">
<emphasis id="B96FEA9BFFB83B0B16B5F94CFE94FBE2" box="[128,288,1057,1079]" italics="true" pageId="5" pageNumber="376">C. bipuncticollis</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6F94FFE67FBE3" box="[403,467,1057,1079]" gridcol="1" gridrow="8" pageId="5" pageNumber="376">M</td>
<td id="76FA5DB7FFB8C4F11435F94FFD81FBE3" box="[512,565,1057,1079]" gridcol="2" gridrow="8" pageId="5" pageNumber="376">4.05</td>
<td id="76FA5DB7FFB8C4F11460F94FFD38FBE3" box="[597,652,1057,1079]" gridcol="3" gridrow="8" pageId="5" pageNumber="376">1.09</td>
<td id="76FA5DB7FFB8C4F1149EF94FFD6AFBE3" box="[683,734,1057,1079]" gridcol="4" gridrow="8" pageId="5" pageNumber="376">0.38</td>
<td id="76FA5DB7FFB8C4F114CAF94FFC84FBE3" box="[767,816,1057,1079]" gridcol="5" gridrow="8" pageId="5" pageNumber="376">1.56</td>
<td id="76FA5DB7FFB8C4F11567F94FFC35FBE3" box="[850,897,1057,1079]" gridcol="6" gridrow="8" pageId="5" pageNumber="376">1.92</td>
<td id="76FA5DB7FFB8C4F11595F94FFC7EFBE3" box="[928,970,1057,1079]" gridcol="7" gridrow="8" pageId="5" pageNumber="376">0.58</td>
<td id="76FA5DB7FFB8C4F115DDF94FFBA6FBE3" box="[1000,1042,1057,1079]" gridcol="8" gridrow="8" pageId="5" pageNumber="376">1.73</td>
<td id="76FA5DB7FFB8C4F11205F94FFBEEFBE3" box="[1072,1114,1057,1079]" gridcol="9" gridrow="8" pageId="5" pageNumber="376">0.77</td>
<td id="76FA5DB7FFB8C4F112B5F94FFB1EFBE3" box="[1152,1194,1057,1079]" gridcol="10" gridrow="8" pageId="5" pageNumber="376"></td>
<td id="76FA5DB7FFB8C4F112FAF94FFAB2FBE3" box="[1231,1286,1057,1079]" gridcol="11" gridrow="8" pageId="5" pageNumber="376">1.56</td>
<td id="76FA5DB7FFB8C4F11312F94FFAE5FBE3" box="[1319,1361,1057,1079]" gridcol="12" gridrow="8" pageId="5" pageNumber="376">1.57</td>
<td id="76FA5DB7FFB8C4F11344F94FFA2FFBE3" box="[1393,1435,1057,1079]" gridcol="13" gridrow="8" pageId="5" pageNumber="376">2.11</td>
</tr>
<tr id="352B34CBFFB8C4F116B5F93EFA2FFAD1" box="[128,1435,1104,1285]" gridrow="9" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5F93EFEF6FAD1" box="[128,322,1104,1285]" gridcol="0" gridrow="9" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5F93FFEADFBB1" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[128,281,1104,1126]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="fuhoshoensis">
<emphasis id="B96FEA9BFFB83B0B16B5F93FFEADFBB1" box="[128,281,1104,1126]" italics="true" pageId="5" pageNumber="376">C. fuhoshoensis</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6F93EFE67FAD1" box="[403,467,1104,1285]" gridcol="1" gridrow="9" pageId="5" pageNumber="376">M M F F HO-M PA-M</td>
<td id="76FA5DB7FFB8C4F11435F93EFD81FAD1" box="[512,565,1104,1285]" gridcol="2" gridrow="9" pageId="5" pageNumber="376">4.17 4.26 3.97 4.12 3.26 3.58</td>
<td id="76FA5DB7FFB8C4F11460F93EFD38FAD1" box="[597,652,1104,1285]" gridcol="3" gridrow="9" pageId="5" pageNumber="376">1.01 0.99 1.02 1.04 0.90 0.96</td>
<td id="76FA5DB7FFB8C4F1149EF93EFD6AFAD1" box="[683,734,1104,1285]" gridcol="4" gridrow="9" pageId="5" pageNumber="376">0.27 0.29 0.35 0.32 0.19 0.18</td>
<td id="76FA5DB7FFB8C4F114CAF93EFC84FAD1" box="[767,816,1104,1285]" gridcol="5" gridrow="9" pageId="5" pageNumber="376">1.55 1.56 1.65 1.67 1.25 1.40</td>
<td id="76FA5DB7FFB8C4F11567F93EFC35FAD1" box="[850,897,1104,1285]" gridcol="6" gridrow="9" pageId="5" pageNumber="376">1.74 1.82 1.94 1.83 1.43 1.55</td>
<td id="76FA5DB7FFB8C4F11595F93EFC7EFAD1" box="[928,970,1104,1285]" gridcol="7" gridrow="9" pageId="5" pageNumber="376">0.53 0.51 0.53 0.48 0.45 0.47</td>
<td id="76FA5DB7FFB8C4F115DDF93EFBA6FAD1" box="[1000,1042,1104,1285]" gridcol="8" gridrow="9" pageId="5" pageNumber="376">1.62 1.68 1.53 1.44 1.40 1.50</td>
<td id="76FA5DB7FFB8C4F11205F93EFBEEFAD1" box="[1072,1114,1104,1285]" gridcol="9" gridrow="9" pageId="5" pageNumber="376">0.84 1.04 0.90 0.89 0.78 0.81</td>
<td id="76FA5DB7FFB8C4F112B5F93EFB1EFAD1" box="[1152,1194,1104,1285]" gridcol="10" gridrow="9" pageId="5" pageNumber="376">0.51 0.57 0.56 0.57 0.54 0.56</td>
<td id="76FA5DB7FFB8C4F112FAF93EFAB2FAD1" box="[1231,1286,1104,1285]" gridcol="11" gridrow="9" pageId="5" pageNumber="376">1.40 1.47 1.49 1.49 1.32 1.35</td>
<td id="76FA5DB7FFB8C4F11312F93EFAE5FAD1" box="[1319,1361,1104,1285]" gridcol="12" gridrow="9" pageId="5" pageNumber="376">1.53 1.59 1.62 1.50 1.35 1.52</td>
<td id="76FA5DB7FFB8C4F11344F93EFA2FFAD1" box="[1393,1435,1104,1285]" gridcol="13" gridrow="9" pageId="5" pageNumber="376">2.16 2.25 2.25 2.19 1.92 2.10</td>
</tr>
<tr id="352B34CBFFB8C4F116B5F870FA2FFA07" box="[128,1435,1310,1491]" gridrow="10" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5F870FEF6FA07" box="[128,322,1310,1491]" gridcol="0" gridrow="10" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5F870FF6FFA84" authorityName="Yasunaga" authorityYear="2024" box="[128,281,1310,1360]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="376" phylum="Tracheophyta" rank="species" species="nanglong">
<emphasis id="B96FEA9BFFB83B0B16B5F870FF6FFA84" box="[128,281,1310,1360]" italics="true" pageId="5" pageNumber="376">Euphorbialygus nanglong</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6F870FE67FA07" box="[403,467,1310,1491]" gridcol="1" gridrow="10" pageId="5" pageNumber="376">PA-M PA-F PA-F PA-F HO-M PA-M</td>
<td id="76FA5DB7FFB8C4F11435F870FD81FA07" box="[512,565,1310,1491]" gridcol="2" gridrow="10" pageId="5" pageNumber="376">3.43 3.80 4.04 3.87 3.43 3.53</td>
<td id="76FA5DB7FFB8C4F11460F870FD38FA07" box="[597,652,1310,1491]" gridcol="3" gridrow="10" pageId="5" pageNumber="376">0.95 0.89 0.93 0.89 0.90 0.87</td>
<td id="76FA5DB7FFB8C4F1149EF870FD6AFA07" box="[683,734,1310,1491]" gridcol="4" gridrow="10" pageId="5" pageNumber="376">0.20 0.27 0.29 0.27 0.18 0.20</td>
<td id="76FA5DB7FFB8C4F114CAF870FC84FA07" box="[767,816,1310,1491]" gridcol="5" gridrow="10" pageId="5" pageNumber="376">1.37 1.37 1.49 1.49 1.34 1.31</td>
<td id="76FA5DB7FFB8C4F11567F870FC35FA07" box="[850,897,1310,1491]" gridcol="6" gridrow="10" pageId="5" pageNumber="376">1.56 1.65 1.79 1.76 1.53 1.50</td>
<td id="76FA5DB7FFB8C4F11595F870FC7EFA07" box="[928,970,1310,1491]" gridcol="7" gridrow="10" pageId="5" pageNumber="376">0.45 0.48 0.50 0.45 0.49 0.45</td>
<td id="76FA5DB7FFB8C4F115DDF870FBA6FA07" box="[1000,1042,1310,1491]" gridcol="8" gridrow="10" pageId="5" pageNumber="376">1.43 1.34 1.47 1.35 1.47 1.35</td>
<td id="76FA5DB7FFB8C4F11205F870FBEEFA07" box="[1072,1114,1310,1491]" gridcol="9" gridrow="10" pageId="5" pageNumber="376">0.80 0.86 0.92 0.78 0.87 0.84</td>
<td id="76FA5DB7FFB8C4F112B5F870FB1EFA07" box="[1152,1194,1310,1491]" gridcol="10" gridrow="10" pageId="5" pageNumber="376">0.53 0.57 0.57 0.51 0.62 0.60</td>
<td id="76FA5DB7FFB8C4F112FAF870FAB2FA07" box="[1231,1286,1310,1491]" gridcol="11" gridrow="10" pageId="5" pageNumber="376">1.31 1.38 1.38 1.34 1.32 1.29</td>
<td id="76FA5DB7FFB8C4F11312F870FAE5FA07" box="[1319,1361,1310,1491]" gridcol="12" gridrow="10" pageId="5" pageNumber="376">1.38 1.43 1.53 1.46 1.35 1.38</td>
<td id="76FA5DB7FFB8C4F11344F870FA2FFA07" box="[1393,1435,1310,1491]" gridcol="13" gridrow="10" pageId="5" pageNumber="376">1.95 1.95 2.18 2.07 1.94 1.95</td>
</tr>
<tr id="352B34CBFFB8C4F116B5F882FA2FF955" box="[128,1435,1516,1665]" gridrow="11" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5F882FEF6F955" box="[128,322,1516,1665]" gridcol="0" gridrow="11" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5F882FEA9F9D5" authorityName="Yasunaga" authorityYear="2024" box="[128,285,1516,1537]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Euphorbialygus" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="376" phylum="Tracheophyta" rank="species" species="rufobrunneus">
<emphasis id="B96FEA9BFFB83B0B16B5F882FEA9F9D5" box="[128,285,1516,1537]" italics="true" pageId="5" pageNumber="376">E. rufobrunneus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6F882FE67F955" box="[403,467,1516,1665]" gridcol="1" gridrow="11" pageId="5" pageNumber="376">PA-M PA-F PA-F HO-M PA-M</td>
<td id="76FA5DB7FFB8C4F11435F882FD81F955" box="[512,565,1516,1665]" gridcol="2" gridrow="11" pageId="5" pageNumber="376">3.50 4.12 4.17 4.17 3.68</td>
<td id="76FA5DB7FFB8C4F11460F882FD38F955" box="[597,652,1516,1665]" gridcol="3" gridrow="11" pageId="5" pageNumber="376">0.89 0.95 0.92 0.90 0.90</td>
<td id="76FA5DB7FFB8C4F1149EF882FD6AF955" box="[683,734,1516,1665]" gridcol="4" gridrow="11" pageId="5" pageNumber="376">0.21 0.30 0.30 0.26 0.29</td>
<td id="76FA5DB7FFB8C4F114CAF882FC84F955" box="[767,816,1516,1665]" gridcol="5" gridrow="11" pageId="5" pageNumber="376">1.34 1.56 1.53 1.70 1.62</td>
<td id="76FA5DB7FFB8C4F11567F882FC35F955" box="[850,897,1516,1665]" gridcol="6" gridrow="11" pageId="5" pageNumber="376">1.55 1.92 1.85 2.10 1.85</td>
<td id="76FA5DB7FFB8C4F11595F882FC7EF955" box="[928,970,1516,1665]" gridcol="7" gridrow="11" pageId="5" pageNumber="376">0.47 0.51 0.48 0.60 0.57</td>
<td id="76FA5DB7FFB8C4F115DDF882FBA6F955" box="[1000,1042,1516,1665]" gridcol="8" gridrow="11" pageId="5" pageNumber="376">1.44 1.50 1.38 1.55 1.46</td>
<td id="76FA5DB7FFB8C4F11205F882FBEEF955" box="[1072,1114,1516,1665]" gridcol="9" gridrow="11" pageId="5" pageNumber="376">0.81 0.89 0.90 0.95 0.86</td>
<td id="76FA5DB7FFB8C4F112B5F882FB1EF955" box="[1152,1194,1516,1665]" gridcol="10" gridrow="11" pageId="5" pageNumber="376">0.50 0.50 0.72 0.75</td>
<td id="76FA5DB7FFB8C4F112FAF882FAB2F955" box="[1231,1286,1516,1665]" gridcol="11" gridrow="11" pageId="5" pageNumber="376">1.34 1.41 1.47 1.65 1.38</td>
<td id="76FA5DB7FFB8C4F11312F882FAE5F955" box="[1319,1361,1516,1665]" gridcol="12" gridrow="11" pageId="5" pageNumber="376">1.46 1.59 1.53 1.65 1.50</td>
<td id="76FA5DB7FFB8C4F11344F882FA2FF955" box="[1393,1435,1516,1665]" gridcol="13" gridrow="11" pageId="5" pageNumber="376">2.00 2.22 2.22 2.25 2.10</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FBF7FA2FF8FB" box="[128,1435,1689,1839]" gridrow="12" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FBF7FEF6F8FB" box="[128,322,1689,1839]" gridcol="0" gridrow="12" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FBF4FF65F91F" authorityName="Yasunaga" authorityYear="2024" box="[128,300,1690,1739]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="duwalae">
<emphasis id="B96FEA9BFFB83B0B16B5FBF4FF65F91F" box="[128,300,1690,1739]" italics="true" pageId="5" pageNumber="376">Himalisundacoris duwalae</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FBF7FE67F8FB" box="[403,467,1689,1839]" gridcol="1" gridrow="12" pageId="5" pageNumber="376">PA-M PA-F PA-F PA-F HO-M</td>
<td id="76FA5DB7FFB8C4F11435FBF7FD81F8FB" box="[512,565,1689,1839]" gridcol="2" gridrow="12" pageId="5" pageNumber="376">3.87 3.92 4.21 4.31 3.72</td>
<td id="76FA5DB7FFB8C4F11460FBF7FD38F8FB" box="[597,652,1689,1839]" gridcol="3" gridrow="12" pageId="5" pageNumber="376">0.92 0.92 0.93 0.95 0.93</td>
<td id="76FA5DB7FFB8C4F1149EFBF7FD6AF8FB" box="[683,734,1689,1839]" gridcol="4" gridrow="12" pageId="5" pageNumber="376">0.27 0.32 0.33 0.35 0.26</td>
<td id="76FA5DB7FFB8C4F114CAFBF7FC84F8FB" box="[767,816,1689,1839]" gridcol="5" gridrow="12" pageId="5" pageNumber="376">1.67 1.80 1.88 1.82 1.50</td>
<td id="76FA5DB7FFB8C4F11567FBF7FC35F8FB" box="[850,897,1689,1839]" gridcol="6" gridrow="12" pageId="5" pageNumber="376">1.88 1.98 2.15 2.13 1.73</td>
<td id="76FA5DB7FFB8C4F11595FBF7FC7EF8FB" box="[928,970,1689,1839]" gridcol="7" gridrow="12" pageId="5" pageNumber="376">0.57 0.60 0.60 0.59 0.68</td>
<td id="76FA5DB7FFB8C4F115DDFBF7FBA6F8FB" box="[1000,1042,1689,1839]" gridcol="8" gridrow="12" pageId="5" pageNumber="376">1.50 1.59 1.61 1.52 1.80</td>
<td id="76FA5DB7FFB8C4F11205FBF7FBEEF8FB" box="[1072,1114,1689,1839]" gridcol="9" gridrow="12" pageId="5" pageNumber="376">0.92 0.90 0.96 0.89 </td>
<td id="76FA5DB7FFB8C4F112B5FBF7FB1EF8FB" box="[1152,1194,1689,1839]" gridcol="10" gridrow="12" pageId="5" pageNumber="376"> 0.74 0.72 </td>
<td id="76FA5DB7FFB8C4F112FAFBF7FAB2F8FB" box="[1231,1286,1689,1839]" gridcol="11" gridrow="12" pageId="5" pageNumber="376">1.55 1.50 1.55 1.52 1.43</td>
<td id="76FA5DB7FFB8C4F11312FBF7FAE5F8FB" box="[1319,1361,1689,1839]" gridcol="12" gridrow="12" pageId="5" pageNumber="376">1.55 1.53 1.74 1.65 1.58</td>
<td id="76FA5DB7FFB8C4F11344FBF7FA2FF8FB" box="[1393,1435,1689,1839]" gridcol="13" gridrow="12" pageId="5" pageNumber="376">2.24 2.25 2.33 2.37 2.25</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FA29FA2FF8A9" box="[128,1435,1863,1917]" gridrow="13" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FA29FEF6F8A9" box="[128,322,1863,1917]" gridcol="0" gridrow="13" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FA26FF6EF889" authorityName="Yasunaga" authorityYear="2024" box="[128,218,1864,1885]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="omiae">
<emphasis id="B96FEA9BFFB83B0B16B5FA26FF6EF889" box="[128,218,1864,1885]" italics="true" pageId="5" pageNumber="376">H. omiae</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FA29FE67F8A9" box="[403,467,1863,1917]" gridcol="1" gridrow="13" pageId="5" pageNumber="376">PA-F PA-F</td>
<td id="76FA5DB7FFB8C4F11435FA29FD81F8A9" box="[512,565,1863,1917]" gridcol="2" gridrow="13" pageId="5" pageNumber="376">4.41 4.17</td>
<td id="76FA5DB7FFB8C4F11460FA29FD38F8A9" box="[597,652,1863,1917]" gridcol="3" gridrow="13" pageId="5" pageNumber="376">0.96 0.93</td>
<td id="76FA5DB7FFB8C4F1149EFA29FD6AF8A9" box="[683,734,1863,1917]" gridcol="4" gridrow="13" pageId="5" pageNumber="376">0.32 0.33</td>
<td id="76FA5DB7FFB8C4F114CAFA29FC84F8A9" box="[767,816,1863,1917]" gridcol="5" gridrow="13" pageId="5" pageNumber="376">1.68 1.70</td>
<td id="76FA5DB7FFB8C4F11567FA29FC35F8A9" box="[850,897,1863,1917]" gridcol="6" gridrow="13" pageId="5" pageNumber="376">1.98 1.88</td>
<td id="76FA5DB7FFB8C4F11595FA29FC7EF8A9" box="[928,970,1863,1917]" gridcol="7" gridrow="13" pageId="5" pageNumber="376">0.68 0.63</td>
<td id="76FA5DB7FFB8C4F115DDFA29FBA6F8A9" box="[1000,1042,1863,1917]" gridcol="8" gridrow="13" pageId="5" pageNumber="376">1.65 1.65</td>
<td id="76FA5DB7FFB8C4F11205FA29FBEEF8A9" box="[1072,1114,1863,1917]" gridcol="9" gridrow="13" pageId="5" pageNumber="376">1.05 1.08</td>
<td id="76FA5DB7FFB8C4F112B5FA29FB1EF8A9" box="[1152,1194,1863,1917]" gridcol="10" gridrow="13" pageId="5" pageNumber="376">0.86 0.90</td>
<td id="76FA5DB7FFB8C4F112FAFA29FAB2F8A9" box="[1231,1286,1863,1917]" gridcol="11" gridrow="13" pageId="5" pageNumber="376">1.55 1.55</td>
<td id="76FA5DB7FFB8C4F11312FA29FAE5F8A9" box="[1319,1361,1863,1917]" gridcol="12" gridrow="13" pageId="5" pageNumber="376">1.79 1.65</td>
<td id="76FA5DB7FFB8C4F11344FA29FA2FF8A9" box="[1393,1435,1863,1917]" gridcol="13" gridrow="13" pageId="5" pageNumber="376">2.40 2.40</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FAF8FA2FF818" box="[128,1435,1942,1996]" gridrow="14" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FAF8FEF6F818" box="[128,322,1942,1996]" gridcol="0" gridrow="14" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FAF9FF68F87F" authorityName="Yasunaga" authorityYear="2024" box="[128,220,1942,1964]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="palolo">
<emphasis id="B96FEA9BFFB83B0B16B5FAF9FF68F87F" box="[128,220,1942,1964]" italics="true" pageId="5" pageNumber="376">H. palolo</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FAF8FE67F818" box="[403,467,1942,1996]" gridcol="1" gridrow="14" pageId="5" pageNumber="376">HO-M PA-F</td>
<td id="76FA5DB7FFB8C4F11435FAF8FD81F818" box="[512,565,1942,1996]" gridcol="2" gridrow="14" pageId="5" pageNumber="376">3.55 3.68</td>
<td id="76FA5DB7FFB8C4F11460FAF8FD38F818" box="[597,652,1942,1996]" gridcol="3" gridrow="14" pageId="5" pageNumber="376">0.86 0.93</td>
<td id="76FA5DB7FFB8C4F1149EFAF8FD6AF818" box="[683,734,1942,1996]" gridcol="4" gridrow="14" pageId="5" pageNumber="376">0.27 0.32</td>
<td id="76FA5DB7FFB8C4F114CAFAF8FC84F818" box="[767,816,1942,1996]" gridcol="5" gridrow="14" pageId="5" pageNumber="376">1.31 1.58</td>
<td id="76FA5DB7FFB8C4F11567FAF8FC35F818" box="[850,897,1942,1996]" gridcol="6" gridrow="14" pageId="5" pageNumber="376">1.65 1.95</td>
<td id="76FA5DB7FFB8C4F11595FAF8FC7EF818" box="[928,970,1942,1996]" gridcol="7" gridrow="14" pageId="5" pageNumber="376">0.60 0.69</td>
<td id="76FA5DB7FFB8C4F115DDFAF8FBA6F818" box="[1000,1042,1942,1996]" gridcol="8" gridrow="14" pageId="5" pageNumber="376">1.50 1.67</td>
<td id="76FA5DB7FFB8C4F11205FAF8FBEEF818" box="[1072,1114,1942,1996]" gridcol="9" gridrow="14" pageId="5" pageNumber="376">0.80 0.98</td>
<td id="76FA5DB7FFB8C4F112B5FAF8FB1EF818" box="[1152,1194,1942,1996]" gridcol="10" gridrow="14" pageId="5" pageNumber="376">0.77 0.83</td>
<td id="76FA5DB7FFB8C4F112FAFAF8FAB2F818" box="[1231,1286,1942,1996]" gridcol="11" gridrow="14" pageId="5" pageNumber="376">1.65 1.58</td>
<td id="76FA5DB7FFB8C4F11312FAF8FAE5F818" box="[1319,1361,1942,1996]" gridcol="12" gridrow="14" pageId="5" pageNumber="376">1.35 1.58</td>
<td id="76FA5DB7FFB8C4F11344FAF8FA2FF818" box="[1393,1435,1942,1996]" gridcol="13" gridrow="14" pageId="5" pageNumber="376">1.95 2.21</td>
</tr>
<tr id="352B34CBFFB8C4F116B5FA8BFA2FF7CE" box="[128,1435,2021,2074]" gridrow="15" pageId="5" pageNumber="376">
<th id="76FA5DB7FFB8C4F116B5FA8BFEF6F7CE" box="[128,322,2021,2074]" gridcol="0" gridrow="15" pageId="5" pageNumber="376">
<taxonomicName id="4C1B4D0AFFB83B0B16B5FA88FF4FF82E" authorityName="Yasunaga" authorityYear="2024" box="[128,251,2021,2043]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="376" phylum="Arthropoda" rank="species" species="schwartzi">
<emphasis id="B96FEA9BFFB83B0B16B5FA88FF4FF82E" box="[128,251,2021,2043]" italics="true" pageId="5" pageNumber="376">H. schwartzi</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFB8C4F117A6FA8BFE67F7CE" box="[403,467,2021,2074]" gridcol="1" gridrow="15" pageId="5" pageNumber="376">HO-M PA-F</td>
<td id="76FA5DB7FFB8C4F11435FA8BFD81F7CE" box="[512,565,2021,2074]" gridcol="2" gridrow="15" pageId="5" pageNumber="376">4.02 3.72</td>
<td id="76FA5DB7FFB8C4F11460FA8BFD38F7CE" box="[597,652,2021,2074]" gridcol="3" gridrow="15" pageId="5" pageNumber="376">0.90 0.87</td>
<td id="76FA5DB7FFB8C4F1149EFA8BFD6AF7CE" box="[683,734,2021,2074]" gridcol="4" gridrow="15" pageId="5" pageNumber="376">0.26 0.30</td>
<td id="76FA5DB7FFB8C4F114CAFA8BFC84F7CE" box="[767,816,2021,2074]" gridcol="5" gridrow="15" pageId="5" pageNumber="376">1.40 1.50</td>
<td id="76FA5DB7FFB8C4F11567FA8BFC35F7CE" box="[850,897,2021,2074]" gridcol="6" gridrow="15" pageId="5" pageNumber="376">1.58 1.70</td>
<td id="76FA5DB7FFB8C4F11595FA8BFC7EF7CE" box="[928,970,2021,2074]" gridcol="7" gridrow="15" pageId="5" pageNumber="376">0.63 0.72</td>
<td id="76FA5DB7FFB8C4F115DDFA8BFBA6F7CE" box="[1000,1042,2021,2074]" gridcol="8" gridrow="15" pageId="5" pageNumber="376">1.67 1.40</td>
<td id="76FA5DB7FFB8C4F11205FA8BFBEEF7CE" box="[1072,1114,2021,2074]" gridcol="9" gridrow="15" pageId="5" pageNumber="376">0.95 0.99</td>
<td id="76FA5DB7FFB8C4F112B5FA8BFB1EF7CE" box="[1152,1194,2021,2074]" gridcol="10" gridrow="15" pageId="5" pageNumber="376">0.90 0.65</td>
<td id="76FA5DB7FFB8C4F112FAFA8BFAB2F7CE" box="[1231,1286,2021,2074]" gridcol="11" gridrow="15" pageId="5" pageNumber="376">1.88 1.50</td>
<td id="76FA5DB7FFB8C4F11312FA8BFAE5F7CE" box="[1319,1361,2021,2074]" gridcol="12" gridrow="15" pageId="5" pageNumber="376">1.46 1.53</td>
<td id="76FA5DB7FFB8C4F11344FA8BFA2FF7CE" box="[1393,1435,2021,2074]" gridcol="13" gridrow="15" pageId="5" pageNumber="376">2.25 2.24</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA43689FFBB3B081422FDA5FA07FB43" pageId="6" pageNumber="377">
<table id="F91BC429FFBBC4F116A2FDA5FA07FB43" box="[151,1459,203,1175]" colsContinueFrom="5.[128,1435,309,2074]" gridcols="14" gridrows="16" pageId="6" pageNumber="377">
<tr id="352B34CBFFBBC4F116A2FDA5FA07FF34" box="[151,1459,203,224]" gridrow="0" pageId="6" pageNumber="377" rowspan-0="1" rowspan-1="1">
<th id="76FA5DB7FFBBC4F11422FDA5FDF8FF34" box="[535,588,203,224]" gridcol="2" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081422FDA5FDF8FF34" bold="true" box="[535,588,203,224]" pageId="6" pageNumber="377">Body</emphasis>
</th>
<th id="76FA5DB7FFBBC4F11458FDA5FD10FF34" box="[621,676,203,224]" gridcol="3" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081458FDA5FD10FF34" bold="true" box="[621,676,203,224]" pageId="6" pageNumber="377">Head</emphasis>
</th>
<th id="76FA5DB7FFBBC4F114F6FDA5FD42FF34" box="[707,758,203,224]" gridcol="4" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0814F6FDA5FD42FF34" bold="true" box="[707,758,203,224]" pageId="6" pageNumber="377">VTX</emphasis>
</th>
<th id="76FA5DB7FFBBC4F11522FDA5FCFCFF34" box="[791,840,203,224]" gridcol="5" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081522FDA5FCFCFF34" bold="true" box="[791,840,203,224]" pageId="6" pageNumber="377">PRN</emphasis>
</th>
<th id="76FA5DB7FFBBC4F1155CFDA5FC2CFF34" box="[873,920,203,224]" gridcol="6" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B08155CFDA5FC2CFF34" bold="true" box="[873,920,203,224]" pageId="6" pageNumber="377">Max</emphasis>
</th>
<th id="76FA5DB7FFBBC4F11582FDA5FB7CFF34" box="[951,1224,203,224]" colspan="4" colspanRight="3" gridcol="7" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0815DBFDA5FB25FF34" bold="true" box="[1006,1169,203,224]" pageId="6" pageNumber="377">Antennomere L</emphasis>
</th>
<th id="76FA5DB7FFBBC4F112D2FDA5FAAAFF34" box="[1255,1310,203,224]" gridcol="11" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0812D2FDA5FAAAFF34" bold="true" box="[1255,1310,203,224]" pageId="6" pageNumber="377">LBM</emphasis>
</th>
<th id="76FA5DB7FFBBC4F1130BFDA5FA07FF34" box="[1342,1459,203,224]" colspan="2" colspanRight="1" gridcol="12" gridrow="0" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081376FDA5FA19FF34" bold="true" box="[1347,1453,203,224]" pageId="6" pageNumber="377">Metaleg L</emphasis>
</th>
</tr>
<tr id="352B34CBFFBBC4F116A2FD94FA07FEDB" box="[151,1459,250,271]" gridrow="1" pageId="6" pageNumber="377" rowspan-0="1" rowspan-1="1">
<td id="76FA5DB7FFBBC4F11422FD94FDF8FEDB" box="[535,588,250,271]" gridcol="2" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B08141FFD94FD8EFEDB" bold="true" box="[554,570,250,271]" pageId="6" pageNumber="377">L</emphasis>
</td>
<td id="76FA5DB7FFBBC4F11458FD94FD10FEDB" box="[621,676,250,271]" gridcol="3" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081449FD94FD20FEDB" bold="true" box="[636,660,250,271]" pageId="6" pageNumber="377">W</emphasis>
</td>
<td id="76FA5DB7FFBBC4F114F6FD94FD42FEDB" box="[707,758,250,271]" gridcol="4" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0814E5FD94FD5CFEDB" bold="true" box="[720,744,250,271]" pageId="6" pageNumber="377">W</emphasis>
</td>
<td id="76FA5DB7FFBBC4F11522FD94FCFCFEDB" box="[791,840,250,271]" gridcol="5" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081516FD94FC8FFEDB" bold="true" box="[803,827,250,271]" pageId="6" pageNumber="377">W</emphasis>
</td>
<td id="76FA5DB7FFBBC4F1155CFD94FC2CFEDB" box="[873,920,250,271]" gridcol="6" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081540FD94FC39FEDB" bold="true" box="[885,909,250,271]" pageId="6" pageNumber="377">W</emphasis>
</td>
<td id="76FA5DB7FFBBC4F11582FD94FC56FEDB" box="[951,994,250,271]" gridcol="7" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0815FDFD94FC65FEDB" bold="true" box="[968,977,250,271]" pageId="6" pageNumber="377">I</emphasis>
</td>
<td id="76FA5DB7FFBBC4F115CAFD94FB9EFEDB" box="[1023,1066,250,271]" gridcol="8" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B08123EFD94FBAAFEDB" bold="true" box="[1035,1054,250,271]" pageId="6" pageNumber="377">II</emphasis>
</td>
<td id="76FA5DB7FFBBC4F1127DFD94FBC6FEDB" box="[1096,1138,250,271]" gridcol="9" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B08127AFD94FBDFFEDB" bold="true" box="[1103,1131,250,271]" pageId="6" pageNumber="377">III</emphasis>
</td>
<td id="76FA5DB7FFBBC4F112A4FD94FB7CFEDB" box="[1169,1224,250,271]" gridcol="10" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0812AAFD94FB0EFEDB" bold="true" box="[1183,1210,250,271]" pageId="6" pageNumber="377">IV</emphasis>
</td>
<td id="76FA5DB7FFBBC4F112D2FD94FAAAFEDB" box="[1255,1310,250,271]" gridcol="11" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0812CFFD94FABEFEDB" bold="true" box="[1274,1290,250,271]" pageId="6" pageNumber="377">L</emphasis>
</td>
<td id="76FA5DB7FFBBC4F1130BFD94FADDFEDB" box="[1342,1385,250,271]" gridcol="12" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081374FD94FAD2FEDB" bold="true" box="[1345,1382,250,271]" pageId="6" pageNumber="377">FM</emphasis>
</td>
<td id="76FA5DB7FFBBC4F113BCFD94FA07FEDB" box="[1417,1459,250,271]" gridcol="13" gridrow="1" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0813BBFD94FA1AFEDB" bold="true" box="[1422,1454,250,271]" pageId="6" pageNumber="377">TB</emphasis>
</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FC5FFA07FE93" box="[151,1459,305,327]" gridrow="2" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2FC5FFEDAFE93" box="[151,366,305,327]" gridcol="0" gridrow="2" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2FC5CFF59FE92" authorityName="Yasunaga" authorityYear="2024" box="[151,237,305,327]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="takaii">
<emphasis id="B96FEA9BFFBB3B0816A2FC5CFF59FE92" box="[151,237,305,327]" italics="true" pageId="6" pageNumber="377">H. takaii</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABFC5FFE43FE93" box="[414,503,305,327]" gridcol="1" gridrow="2" pageId="6" pageNumber="377">HO-M</td>
<td id="76FA5DB7FFBBC4F11422FC5FFDF8FE93" box="[535,588,305,327]" gridcol="2" gridrow="2" pageId="6" pageNumber="377">3.92</td>
<td id="76FA5DB7FFBBC4F11458FC5FFD10FE93" box="[621,676,305,327]" gridcol="3" gridrow="2" pageId="6" pageNumber="377">0.90</td>
<td id="76FA5DB7FFBBC4F114F6FC5FFD42FE93" box="[707,758,305,327]" gridcol="4" gridrow="2" pageId="6" pageNumber="377">0.27</td>
<td id="76FA5DB7FFBBC4F11522FC5FFCFCFE93" box="[791,840,305,327]" gridcol="5" gridrow="2" pageId="6" pageNumber="377">1.53</td>
<td id="76FA5DB7FFBBC4F1155CFC5FFC2CFE93" box="[873,920,305,327]" gridcol="6" gridrow="2" pageId="6" pageNumber="377">1.83</td>
<td id="76FA5DB7FFBBC4F11582FC5FFC56FE93" box="[951,994,305,327]" gridcol="7" gridrow="2" pageId="6" pageNumber="377">0.59</td>
<td id="76FA5DB7FFBBC4F115CAFC5FFB9EFE93" box="[1023,1066,305,327]" gridcol="8" gridrow="2" pageId="6" pageNumber="377">1.52</td>
<td id="76FA5DB7FFBBC4F1127DFC5FFBC6FE93" box="[1096,1138,305,327]" gridcol="9" gridrow="2" pageId="6" pageNumber="377">0.87</td>
<td id="76FA5DB7FFBBC4F112A4FC5FFB7CFE93" box="[1169,1224,305,327]" gridcol="10" gridrow="2" pageId="6" pageNumber="377">0.86</td>
<td id="76FA5DB7FFBBC4F112D2FC5FFAAAFE93" box="[1255,1310,305,327]" gridcol="11" gridrow="2" pageId="6" pageNumber="377">1.65</td>
<td id="76FA5DB7FFBBC4F1130BFC5FFADDFE93" box="[1342,1385,305,327]" gridcol="12" gridrow="2" pageId="6" pageNumber="377">1.50</td>
<td id="76FA5DB7FFBBC4F113BCFC5FFA07FE93" box="[1417,1459,305,327]" gridcol="13" gridrow="2" pageId="6" pageNumber="377">2.28</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FC3FFA07FE52" box="[151,1459,337,390]" gridrow="3" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFC3FFE43FE52" box="[414,503,337,390]" gridcol="1" gridrow="3" pageId="6" pageNumber="377">HO-M PA-M</td>
<td id="76FA5DB7FFBBC4F11422FC3FFDF8FE52" box="[535,588,337,390]" gridcol="2" gridrow="3" pageId="6" pageNumber="377">3.19 3.43</td>
<td id="76FA5DB7FFBBC4F11458FC3FFD10FE52" box="[621,676,337,390]" gridcol="3" gridrow="3" pageId="6" pageNumber="377">0.89 0.87</td>
<td id="76FA5DB7FFBBC4F114F6FC3FFD42FE52" box="[707,758,337,390]" gridcol="4" gridrow="3" pageId="6" pageNumber="377">0.27 0.27</td>
<td id="76FA5DB7FFBBC4F11522FC3FFCFCFE52" box="[791,840,337,390]" gridcol="5" gridrow="3" pageId="6" pageNumber="377">1.46 1.46</td>
<td id="76FA5DB7FFBBC4F1155CFC3FFC2CFE52" box="[873,920,337,390]" gridcol="6" gridrow="3" pageId="6" pageNumber="377">1.59 1.62</td>
<td id="76FA5DB7FFBBC4F11582FC3FFC56FE52" box="[951,994,337,390]" gridcol="7" gridrow="3" pageId="6" pageNumber="377">0.74 0.72</td>
<td id="76FA5DB7FFBBC4F115CAFC3FFB9EFE52" box="[1023,1066,337,390]" gridcol="8" gridrow="3" pageId="6" pageNumber="377">1.53 1.53</td>
<td id="76FA5DB7FFBBC4F1127DFC3FFBC6FE52" box="[1096,1138,337,390]" gridcol="9" gridrow="3" pageId="6" pageNumber="377">1.08 1.13</td>
<td id="76FA5DB7FFBBC4F112A4FC3FFB7CFE52" box="[1169,1224,337,390]" gridcol="10" gridrow="3" pageId="6" pageNumber="377">0.57 0.75</td>
<td id="76FA5DB7FFBBC4F112D2FC3FFAAAFE52" box="[1255,1310,337,390]" gridcol="11" gridrow="3" pageId="6" pageNumber="377">1.46 1.50</td>
<td id="76FA5DB7FFBBC4F1130BFC3FFADDFE52" box="[1342,1385,337,390]" gridcol="12" gridrow="3" pageId="6" pageNumber="377">1.65 1.58</td>
<td id="76FA5DB7FFBBC4F113BCFC3FFA07FE52" box="[1417,1459,337,390]" gridcol="13" gridrow="3" pageId="6" pageNumber="377">2.30 2.28</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FCF1FA07FE01" box="[151,1459,415,469]" gridrow="4" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2FCF1FEDAFE01" box="[151,366,415,469]" gridcol="0" gridrow="4" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2FCCEFF52FE60" authorityName="Yasunaga" authorityYear="2024" box="[151,230,415,437]" class="Insecta" family="Miridae" genus="Himalisundacoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="theoi">
<emphasis id="B96FEA9BFFBB3B0816A2FCCEFF52FE60" box="[151,230,415,437]" italics="true" pageId="6" pageNumber="377">H. theoi</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABFCF1FE43FE01" box="[414,503,415,469]" gridcol="1" gridrow="4" pageId="6" pageNumber="377">PA-M PA-F</td>
<td id="76FA5DB7FFBBC4F11422FCF1FDF8FE01" box="[535,588,415,469]" gridcol="2" gridrow="4" pageId="6" pageNumber="377">3.31 3.99</td>
<td id="76FA5DB7FFBBC4F11458FCF1FD10FE01" box="[621,676,415,469]" gridcol="3" gridrow="4" pageId="6" pageNumber="377">0.84 0.93</td>
<td id="76FA5DB7FFBBC4F114F6FCF1FD42FE01" box="[707,758,415,469]" gridcol="4" gridrow="4" pageId="6" pageNumber="377">0.27 0.35</td>
<td id="76FA5DB7FFBBC4F11522FCF1FCFCFE01" box="[791,840,415,469]" gridcol="5" gridrow="4" pageId="6" pageNumber="377">1.35 1.76</td>
<td id="76FA5DB7FFBBC4F1155CFCF1FC2CFE01" box="[873,920,415,469]" gridcol="6" gridrow="4" pageId="6" pageNumber="377">1.50 1.89</td>
<td id="76FA5DB7FFBBC4F11582FCF1FC56FE01" box="[951,994,415,469]" gridcol="7" gridrow="4" pageId="6" pageNumber="377">0.65 0.66</td>
<td id="76FA5DB7FFBBC4F115CAFCF1FB9EFE01" box="[1023,1066,415,469]" gridcol="8" gridrow="4" pageId="6" pageNumber="377">1.46 1.40</td>
<td id="76FA5DB7FFBBC4F1127DFCF1FBC6FE01" box="[1096,1138,415,469]" gridcol="9" gridrow="4" pageId="6" pageNumber="377">1.07 1.04</td>
<td id="76FA5DB7FFBBC4F112A4FCF1FB7CFE01" box="[1169,1224,415,469]" gridcol="10" gridrow="4" pageId="6" pageNumber="377">0.63 0.65</td>
<td id="76FA5DB7FFBBC4F112D2FCF1FAAAFE01" box="[1255,1310,415,469]" gridcol="11" gridrow="4" pageId="6" pageNumber="377">1.49 1.68</td>
<td id="76FA5DB7FFBBC4F1130BFCF1FADDFE01" box="[1342,1385,415,469]" gridcol="12" gridrow="4" pageId="6" pageNumber="377">1.49 1.64</td>
<td id="76FA5DB7FFBBC4F113BCFCF1FA07FE01" box="[1417,1459,415,469]" gridcol="13" gridrow="4" pageId="6" pageNumber="377">2.22 2.40</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FCB1FA07FDE0" box="[151,1459,479,564]" gridrow="5" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFCB1FE43FDE0" box="[414,503,479,564]" gridcol="1" gridrow="5" pageId="6" pageNumber="377">PA-F HO-M PA-M</td>
<td id="76FA5DB7FFBBC4F11422FCB1FDF8FDE0" box="[535,588,479,564]" gridcol="2" gridrow="5" pageId="6" pageNumber="377">3.68 4.41 4.70</td>
<td id="76FA5DB7FFBBC4F11458FCB1FD10FDE0" box="[621,676,479,564]" gridcol="3" gridrow="5" pageId="6" pageNumber="377">0.92 1.07 1.05</td>
<td id="76FA5DB7FFBBC4F114F6FCB1FD42FDE0" box="[707,758,479,564]" gridcol="4" gridrow="5" pageId="6" pageNumber="377">0.33 0.29 0.29</td>
<td id="76FA5DB7FFBBC4F11522FCB1FCFCFDE0" box="[791,840,479,564]" gridcol="5" gridrow="5" pageId="6" pageNumber="377">1.64 1.50 1.52</td>
<td id="76FA5DB7FFBBC4F1155CFCB1FC2CFDE0" box="[873,920,479,564]" gridcol="6" gridrow="5" pageId="6" pageNumber="377">1.83 1.65 1.64</td>
<td id="76FA5DB7FFBBC4F11582FCB1FC56FDE0" box="[951,994,479,564]" gridcol="7" gridrow="5" pageId="6" pageNumber="377">0.66 0.60 0.62</td>
<td id="76FA5DB7FFBBC4F115CAFCB1FB9EFDE0" box="[1023,1066,479,564]" gridcol="8" gridrow="5" pageId="6" pageNumber="377">1.49 2.07 2.12</td>
<td id="76FA5DB7FFBBC4F1127DFCB1FBC6FDE0" box="[1096,1138,479,564]" gridcol="9" gridrow="5" pageId="6" pageNumber="377">1.13 1.20 1.23</td>
<td id="76FA5DB7FFBBC4F112A4FCB1FB7CFDE0" box="[1169,1224,479,564]" gridcol="10" gridrow="5" pageId="6" pageNumber="377">0.75 0.52* 0.62</td>
<td id="76FA5DB7FFBBC4F112D2FCB1FAAAFDE0" box="[1255,1310,479,564]" gridcol="11" gridrow="5" pageId="6" pageNumber="377">1.55 1.50 1.49</td>
<td id="76FA5DB7FFBBC4F1130BFCB1FADDFDE0" box="[1342,1385,479,564]" gridcol="12" gridrow="5" pageId="6" pageNumber="377">1.76 1.80 1.82</td>
<td id="76FA5DB7FFBBC4F113BCFCB1FA07FDE0" box="[1417,1459,479,564]" gridcol="13" gridrow="5" pageId="6" pageNumber="377">2.51 2.73 2.73</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FF23FA07FD57" box="[151,1459,589,643]" gridrow="6" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2FF23FEDAFD57" box="[151,366,589,643]" gridcol="0" gridrow="6" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2FF23FEDAFDB6" authorityName="Yasunaga" authorityYear="2024" box="[151,366,589,610]" class="Insecta" family="Miridae" genus="Paralygocoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="balicus">
<emphasis id="B96FEA9BFFBB3B0816A2FF23FEDAFDB6" box="[151,366,589,610]" italics="true" pageId="6" pageNumber="377">Paralygocoris balicus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABFF23FE43FD57" box="[414,503,589,643]" gridcol="1" gridrow="6" pageId="6" pageNumber="377">PA-M PA-F</td>
<td id="76FA5DB7FFBBC4F11422FF23FDF8FD57" box="[535,588,589,643]" gridcol="2" gridrow="6" pageId="6" pageNumber="377">4.58 4.95</td>
<td id="76FA5DB7FFBBC4F11458FF23FD10FD57" box="[621,676,589,643]" gridcol="3" gridrow="6" pageId="6" pageNumber="377">1.05 1.10</td>
<td id="76FA5DB7FFBBC4F114F6FF23FD42FD57" box="[707,758,589,643]" gridcol="4" gridrow="6" pageId="6" pageNumber="377">0.28 0.39</td>
<td id="76FA5DB7FFBBC4F11522FF23FCFCFD57" box="[791,840,589,643]" gridcol="5" gridrow="6" pageId="6" pageNumber="377">1.52 1.79</td>
<td id="76FA5DB7FFBBC4F1155CFF23FC2CFD57" box="[873,920,589,643]" gridcol="6" gridrow="6" pageId="6" pageNumber="377">1.67 1.98</td>
<td id="76FA5DB7FFBBC4F11582FF23FC56FD57" box="[951,994,589,643]" gridcol="7" gridrow="6" pageId="6" pageNumber="377">0.60 0.63</td>
<td id="76FA5DB7FFBBC4F115CAFF23FB9EFD57" box="[1023,1066,589,643]" gridcol="8" gridrow="6" pageId="6" pageNumber="377">2.10 2.09</td>
<td id="76FA5DB7FFBBC4F1127DFF23FBC6FD57" box="[1096,1138,589,643]" gridcol="9" gridrow="6" pageId="6" pageNumber="377">1.17 1.22</td>
<td id="76FA5DB7FFBBC4F112A4FF23FB7CFD57" box="[1169,1224,589,643]" gridcol="10" gridrow="6" pageId="6" pageNumber="377">0.49* 0.65</td>
<td id="76FA5DB7FFBBC4F112D2FF23FAAAFD57" box="[1255,1310,589,643]" gridcol="11" gridrow="6" pageId="6" pageNumber="377">1.53 1.53</td>
<td id="76FA5DB7FFBBC4F1130BFF23FADDFD57" box="[1342,1385,589,643]" gridcol="12" gridrow="6" pageId="6" pageNumber="377">1.82 1.97</td>
<td id="76FA5DB7FFBBC4F113BCFF23FA07FD57" box="[1417,1459,589,643]" gridcol="13" gridrow="6" pageId="6" pageNumber="377">2.70 3.00</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FFE3FA07FD17" box="[151,1459,653,707]" gridrow="7" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFFE3FE43FD17" box="[414,503,653,707]" gridcol="1" gridrow="7" pageId="6" pageNumber="377">PA-F PA-F</td>
<td id="76FA5DB7FFBBC4F11422FFE3FDF8FD17" box="[535,588,653,707]" gridcol="2" gridrow="7" pageId="6" pageNumber="377">4.43 5.02</td>
<td id="76FA5DB7FFBBC4F11458FFE3FD10FD17" box="[621,676,653,707]" gridcol="3" gridrow="7" pageId="6" pageNumber="377">1.05 1.10</td>
<td id="76FA5DB7FFBBC4F114F6FFE3FD42FD17" box="[707,758,653,707]" gridcol="4" gridrow="7" pageId="6" pageNumber="377">0.33 0.36</td>
<td id="76FA5DB7FFBBC4F11522FFE3FCFCFD17" box="[791,840,653,707]" gridcol="5" gridrow="7" pageId="6" pageNumber="377">1.65 1.70</td>
<td id="76FA5DB7FFBBC4F1155CFFE3FC2CFD17" box="[873,920,653,707]" gridcol="6" gridrow="7" pageId="6" pageNumber="377">1.82 1.97</td>
<td id="76FA5DB7FFBBC4F11582FFE3FC56FD17" box="[951,994,653,707]" gridcol="7" gridrow="7" pageId="6" pageNumber="377">0.62 0.60</td>
<td id="76FA5DB7FFBBC4F115CAFFE3FB9EFD17" box="[1023,1066,653,707]" gridcol="8" gridrow="7" pageId="6" pageNumber="377">1.95 1.98</td>
<td id="76FA5DB7FFBBC4F1127DFFE3FBC6FD17" box="[1096,1138,653,707]" gridcol="9" gridrow="7" pageId="6" pageNumber="377">1.23 1.23</td>
<td id="76FA5DB7FFBBC4F112A4FFE3FB7CFD17" box="[1169,1224,653,707]" gridcol="10" gridrow="7" pageId="6" pageNumber="377">0.62 0.72</td>
<td id="76FA5DB7FFBBC4F112D2FFE3FAAAFD17" box="[1255,1310,653,707]" gridcol="11" gridrow="7" pageId="6" pageNumber="377">1.56 1.58</td>
<td id="76FA5DB7FFBBC4F1130BFFE3FADDFD17" box="[1342,1385,653,707]" gridcol="12" gridrow="7" pageId="6" pageNumber="377">1.91 1.86</td>
<td id="76FA5DB7FFBBC4F113BCFFE3FA07FD17" box="[1417,1459,653,707]" gridcol="13" gridrow="7" pageId="6" pageNumber="377">2.96 2.85</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FFBAFA07FD25" box="[151,1459,724,753]" gridrow="8" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2FFBAFEDAFD25" box="[151,366,724,753]" gridcol="0" gridrow="8" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2FFB2FE99FD24" authorityName="Lu &amp; Zheng" authorityYear="2004" box="[151,301,731,753]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B96FEA9BFFBB3B0816A2FFB2FE99FD24" box="[151,301,731,753]" italics="true" pageId="6" pageNumber="377">P. meridionalis</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABFFBAFE43FD25" box="[414,503,724,753]" gridcol="1" gridrow="8" pageId="6" pageNumber="377">F (Thai)</td>
<td id="76FA5DB7FFBBC4F11422FFBAFDF8FD25" box="[535,588,724,753]" gridcol="2" gridrow="8" pageId="6" pageNumber="377">4.66</td>
<td id="76FA5DB7FFBBC4F11458FFBAFD10FD25" box="[621,676,724,753]" gridcol="3" gridrow="8" pageId="6" pageNumber="377">1.08</td>
<td id="76FA5DB7FFBBC4F114F6FFBAFD42FD25" box="[707,758,724,753]" gridcol="4" gridrow="8" pageId="6" pageNumber="377">0.39</td>
<td id="76FA5DB7FFBBC4F11522FFBAFCFCFD25" box="[791,840,724,753]" gridcol="5" gridrow="8" pageId="6" pageNumber="377">1.59</td>
<td id="76FA5DB7FFBBC4F1155CFFBAFC2CFD25" box="[873,920,724,753]" gridcol="6" gridrow="8" pageId="6" pageNumber="377">1.68</td>
<td id="76FA5DB7FFBBC4F11582FFBAFC56FD25" box="[951,994,724,753]" gridcol="7" gridrow="8" pageId="6" pageNumber="377">0.62</td>
<td id="76FA5DB7FFBBC4F115CAFFBAFB9EFD25" box="[1023,1066,724,753]" gridcol="8" gridrow="8" pageId="6" pageNumber="377">1.95</td>
<td id="76FA5DB7FFBBC4F1127DFFBAFBC6FD25" box="[1096,1138,724,753]" gridcol="9" gridrow="8" pageId="6" pageNumber="377">1.29</td>
<td id="76FA5DB7FFBBC4F112A4FFBAFB7CFD25" box="[1169,1224,724,753]" gridcol="10" gridrow="8" pageId="6" pageNumber="377">0.52*</td>
<td id="76FA5DB7FFBBC4F112D2FFBAFAAAFD25" box="[1255,1310,724,753]" gridcol="11" gridrow="8" pageId="6" pageNumber="377">1.35</td>
<td id="76FA5DB7FFBBC4F1130BFFBAFADDFD25" box="[1342,1385,724,753]" gridcol="12" gridrow="8" pageId="6" pageNumber="377">1.85</td>
<td id="76FA5DB7FFBBC4F113BCFFBAFA07FD25" box="[1417,1459,724,753]" gridcol="13" gridrow="8" pageId="6" pageNumber="377">2.82</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FF95FA07FCC5" box="[151,1459,763,785]" gridrow="9" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFF95FE43FCC5" box="[414,503,763,785]" gridcol="1" gridrow="9" pageId="6" pageNumber="377">F</td>
<td id="76FA5DB7FFBBC4F11422FF95FDF8FCC5" box="[535,588,763,785]" gridcol="2" gridrow="9" pageId="6" pageNumber="377">4.75</td>
<td id="76FA5DB7FFBBC4F11458FF95FD10FCC5" box="[621,676,763,785]" gridcol="3" gridrow="9" pageId="6" pageNumber="377">1.07</td>
<td id="76FA5DB7FFBBC4F114F6FF95FD42FCC5" box="[707,758,763,785]" gridcol="4" gridrow="9" pageId="6" pageNumber="377">0.36</td>
<td id="76FA5DB7FFBBC4F11522FF95FCFCFCC5" box="[791,840,763,785]" gridcol="5" gridrow="9" pageId="6" pageNumber="377">1.55</td>
<td id="76FA5DB7FFBBC4F1155CFF95FC2CFCC5" box="[873,920,763,785]" gridcol="6" gridrow="9" pageId="6" pageNumber="377">1.65</td>
<td id="76FA5DB7FFBBC4F11582FF95FC56FCC5" box="[951,994,763,785]" gridcol="7" gridrow="9" pageId="6" pageNumber="377">0.72</td>
<td id="76FA5DB7FFBBC4F115CAFF95FB9EFCC5" box="[1023,1066,763,785]" gridcol="8" gridrow="9" pageId="6" pageNumber="377">1.95</td>
<td id="76FA5DB7FFBBC4F1127DFF95FBC6FCC5" box="[1096,1138,763,785]" gridcol="9" gridrow="9" pageId="6" pageNumber="377">1.35</td>
<td id="76FA5DB7FFBBC4F112A4FF95FB7CFCC5" box="[1169,1224,763,785]" gridcol="10" gridrow="9" pageId="6" pageNumber="377"></td>
<td id="76FA5DB7FFBBC4F112D2FF95FAAAFCC5" box="[1255,1310,763,785]" gridcol="11" gridrow="9" pageId="6" pageNumber="377">1.37</td>
<td id="76FA5DB7FFBBC4F1130BFF95FADDFCC5" box="[1342,1385,763,785]" gridcol="12" gridrow="9" pageId="6" pageNumber="377">1.85</td>
<td id="76FA5DB7FFBBC4F113BCFF95FA07FCC5" box="[1417,1459,763,785]" gridcol="13" gridrow="9" pageId="6" pageNumber="377">2.90</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FE76FA07FCBF" box="[151,1459,792,875]" gridrow="10" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFE76FE43FCBF" box="[414,503,792,875]" gridcol="1" gridrow="10" pageId="6" pageNumber="377">(Malaya) F (Malaya)</td>
<td id="76FA5DB7FFBBC4F11422FE76FDF8FCBF" box="[535,588,792,875]" gridcol="2" gridrow="10" pageId="6" pageNumber="377">4.83</td>
<td id="76FA5DB7FFBBC4F11458FE76FD10FCBF" box="[621,676,792,875]" gridcol="3" gridrow="10" pageId="6" pageNumber="377">1.08</td>
<td id="76FA5DB7FFBBC4F114F6FE76FD42FCBF" box="[707,758,792,875]" gridcol="4" gridrow="10" pageId="6" pageNumber="377">0.38</td>
<td id="76FA5DB7FFBBC4F11522FE76FCFCFCBF" box="[791,840,792,875]" gridcol="5" gridrow="10" pageId="6" pageNumber="377">1.65</td>
<td id="76FA5DB7FFBBC4F1155CFE76FC2CFCBF" box="[873,920,792,875]" gridcol="6" gridrow="10" pageId="6" pageNumber="377">1.73</td>
<td id="76FA5DB7FFBBC4F11582FE76FC56FCBF" box="[951,994,792,875]" gridcol="7" gridrow="10" pageId="6" pageNumber="377">0.72</td>
<td id="76FA5DB7FFBBC4F115CAFE76FB9EFCBF" box="[1023,1066,792,875]" gridcol="8" gridrow="10" pageId="6" pageNumber="377">1.98</td>
<td id="76FA5DB7FFBBC4F1127DFE76FBC6FCBF" box="[1096,1138,792,875]" gridcol="9" gridrow="10" pageId="6" pageNumber="377">1.40</td>
<td id="76FA5DB7FFBBC4F112A4FE76FB7CFCBF" box="[1169,1224,792,875]" gridcol="10" gridrow="10" pageId="6" pageNumber="377">0.84</td>
<td id="76FA5DB7FFBBC4F112D2FE76FAAAFCBF" box="[1255,1310,792,875]" gridcol="11" gridrow="10" pageId="6" pageNumber="377">1.37</td>
<td id="76FA5DB7FFBBC4F1130BFE76FADDFCBF" box="[1342,1385,792,875]" gridcol="12" gridrow="10" pageId="6" pageNumber="377">1.95</td>
<td id="76FA5DB7FFBBC4F113BCFE76FA07FCBF" box="[1417,1459,792,875]" gridcol="13" gridrow="10" pageId="6" pageNumber="377">2.87</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FE12FA07FC46" box="[151,1459,892,914]" gridrow="11" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2FE12FEDAFC46" box="[151,366,892,914]" gridcol="0" gridrow="11" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2FE13FEB9FC45" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1914" box="[151,269,892,914]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="vittulatus">
<emphasis id="B96FEA9BFFBB3B0816A2FE13FEB9FC45" box="[151,269,892,914]" italics="true" pageId="6" pageNumber="377">P. vittulatus</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABFE12FE43FC46" box="[414,503,892,914]" gridcol="1" gridrow="11" pageId="6" pageNumber="377">M</td>
<td id="76FA5DB7FFBBC4F11422FE12FDF8FC46" box="[535,588,892,914]" gridcol="2" gridrow="11" pageId="6" pageNumber="377">4.97</td>
<td id="76FA5DB7FFBBC4F11458FE12FD10FC46" box="[621,676,892,914]" gridcol="3" gridrow="11" pageId="6" pageNumber="377">1.05</td>
<td id="76FA5DB7FFBBC4F114F6FE12FD42FC46" box="[707,758,892,914]" gridcol="4" gridrow="11" pageId="6" pageNumber="377">0.28</td>
<td id="76FA5DB7FFBBC4F11522FE12FCFCFC46" box="[791,840,892,914]" gridcol="5" gridrow="11" pageId="6" pageNumber="377">1.62</td>
<td id="76FA5DB7FFBBC4F1155CFE12FC2CFC46" box="[873,920,892,914]" gridcol="6" gridrow="11" pageId="6" pageNumber="377">1.94</td>
<td id="76FA5DB7FFBBC4F11582FE12FC56FC46" box="[951,994,892,914]" gridcol="7" gridrow="11" pageId="6" pageNumber="377">0.72</td>
<td id="76FA5DB7FFBBC4F115CAFE12FB9EFC46" box="[1023,1066,892,914]" gridcol="8" gridrow="11" pageId="6" pageNumber="377">2.10</td>
<td id="76FA5DB7FFBBC4F1127DFE12FBC6FC46" box="[1096,1138,892,914]" gridcol="9" gridrow="11" pageId="6" pageNumber="377">1.34</td>
<td id="76FA5DB7FFBBC4F112A4FE12FB7CFC46" box="[1169,1224,892,914]" gridcol="10" gridrow="11" pageId="6" pageNumber="377">0.72</td>
<td id="76FA5DB7FFBBC4F112D2FE12FAAAFC46" box="[1255,1310,892,914]" gridcol="11" gridrow="11" pageId="6" pageNumber="377">1.80</td>
<td id="76FA5DB7FFBBC4F1130BFE12FADDFC46" box="[1342,1385,892,914]" gridcol="12" gridrow="11" pageId="6" pageNumber="377">1.85</td>
<td id="76FA5DB7FFBBC4F113BCFE12FA07FC46" box="[1417,1459,892,914]" gridcol="13" gridrow="11" pageId="6" pageNumber="377">2.85</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FEF2FA07FC25" box="[151,1459,924,1009]" gridrow="12" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFEF2FE43FC25" box="[414,503,924,1009]" gridcol="1" gridrow="12" pageId="6" pageNumber="377">M M F</td>
<td id="76FA5DB7FFBBC4F11422FEF2FDF8FC25" box="[535,588,924,1009]" gridcol="2" gridrow="12" pageId="6" pageNumber="377">5.02 4.90 5.64</td>
<td id="76FA5DB7FFBBC4F11458FEF2FD10FC25" box="[621,676,924,1009]" gridcol="3" gridrow="12" pageId="6" pageNumber="377">1.08 1.07 1.09</td>
<td id="76FA5DB7FFBBC4F114F6FEF2FD42FC25" box="[707,758,924,1009]" gridcol="4" gridrow="12" pageId="6" pageNumber="377">0.29 0.27 0.34</td>
<td id="76FA5DB7FFBBC4F11522FEF2FCFCFC25" box="[791,840,924,1009]" gridcol="5" gridrow="12" pageId="6" pageNumber="377">1.71 1.70 1.83</td>
<td id="76FA5DB7FFBBC4F1155CFEF2FC2CFC25" box="[873,920,924,1009]" gridcol="6" gridrow="12" pageId="6" pageNumber="377">1.95 1.95 2.10</td>
<td id="76FA5DB7FFBBC4F11582FEF2FC56FC25" box="[951,994,924,1009]" gridcol="7" gridrow="12" pageId="6" pageNumber="377">0.71 0.69 0.72</td>
<td id="76FA5DB7FFBBC4F115CAFEF2FB9EFC25" box="[1023,1066,924,1009]" gridcol="8" gridrow="12" pageId="6" pageNumber="377">2.12 2.19 2.07</td>
<td id="76FA5DB7FFBBC4F1127DFEF2FBC6FC25" box="[1096,1138,924,1009]" gridcol="9" gridrow="12" pageId="6" pageNumber="377">1.35 1.37 1.35</td>
<td id="76FA5DB7FFBBC4F112A4FEF2FB7CFC25" box="[1169,1224,924,1009]" gridcol="10" gridrow="12" pageId="6" pageNumber="377">0.68 0.71 0.66</td>
<td id="76FA5DB7FFBBC4F112D2FEF2FAAAFC25" box="[1255,1310,924,1009]" gridcol="11" gridrow="12" pageId="6" pageNumber="377">1.82 1.73 1.91</td>
<td id="76FA5DB7FFBBC4F1130BFEF2FADDFC25" box="[1342,1385,924,1009]" gridcol="12" gridrow="12" pageId="6" pageNumber="377">1.92 1.88 2.04</td>
<td id="76FA5DB7FFBBC4F113BCFEF2FA07FC25" box="[1417,1459,924,1009]" gridcol="13" gridrow="12" pageId="6" pageNumber="377">2.97 2.85 3.03</td>
</tr>
<tr id="352B34CBFFBBC4F116A2FE95FA07FBC5" box="[151,1459,1019,1041]" gridrow="13" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABFE95FE43FBC5" box="[414,503,1019,1041]" gridcol="1" gridrow="13" pageId="6" pageNumber="377">F</td>
<td id="76FA5DB7FFBBC4F11422FE95FDF8FBC5" box="[535,588,1019,1041]" gridcol="2" gridrow="13" pageId="6" pageNumber="377">5.19</td>
<td id="76FA5DB7FFBBC4F11458FE95FD10FBC5" box="[621,676,1019,1041]" gridcol="3" gridrow="13" pageId="6" pageNumber="377">1.07</td>
<td id="76FA5DB7FFBBC4F114F6FE95FD42FBC5" box="[707,758,1019,1041]" gridcol="4" gridrow="13" pageId="6" pageNumber="377">0.36</td>
<td id="76FA5DB7FFBBC4F11522FE95FCFCFBC5" box="[791,840,1019,1041]" gridcol="5" gridrow="13" pageId="6" pageNumber="377">1.80</td>
<td id="76FA5DB7FFBBC4F1155CFE95FC2CFBC5" box="[873,920,1019,1041]" gridcol="6" gridrow="13" pageId="6" pageNumber="377">2.09</td>
<td id="76FA5DB7FFBBC4F11582FE95FC56FBC5" box="[951,994,1019,1041]" gridcol="7" gridrow="13" pageId="6" pageNumber="377">0.68</td>
<td id="76FA5DB7FFBBC4F115CAFE95FB9EFBC5" box="[1023,1066,1019,1041]" gridcol="8" gridrow="13" pageId="6" pageNumber="377">1.95</td>
<td id="76FA5DB7FFBBC4F1127DFE95FBC6FBC5" box="[1096,1138,1019,1041]" gridcol="9" gridrow="13" pageId="6" pageNumber="377">1.28</td>
<td id="76FA5DB7FFBBC4F112A4FE95FB7CFBC5" box="[1169,1224,1019,1041]" gridcol="10" gridrow="13" pageId="6" pageNumber="377">0.69</td>
<td id="76FA5DB7FFBBC4F112D2FE95FAAAFBC5" box="[1255,1310,1019,1041]" gridcol="11" gridrow="13" pageId="6" pageNumber="377">1.89</td>
<td id="76FA5DB7FFBBC4F1130BFE95FADDFBC5" box="[1342,1385,1019,1041]" gridcol="12" gridrow="13" pageId="6" pageNumber="377">1.91</td>
<td id="76FA5DB7FFBBC4F113BCFE95FA07FBC5" box="[1417,1459,1019,1041]" gridcol="13" gridrow="13" pageId="6" pageNumber="377">2.85</td>
</tr>
<tr id="352B34CBFFBBC4F116A2F94CFA07FBEC" box="[151,1459,1058,1080]" gridrow="14" pageId="6" pageNumber="377">
<th id="76FA5DB7FFBBC4F116A2F94CFEDAFBEC" box="[151,366,1058,1080]" gridcol="0" gridrow="14" pageId="6" pageNumber="377">
<taxonomicName id="4C1B4D0AFFBB3B0816A2F94CFEE7FBE3" authorityName="Yasunaga" authorityYear="2024" box="[151,339,1058,1079]" class="Insecta" family="Miridae" genus="Sabactiopus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="377" phylum="Arthropoda" rank="species" species="schuhi">
<emphasis id="B96FEA9BFFBB3B0816A2F94CFEE7FBE3" box="[151,339,1058,1079]" italics="true" pageId="6" pageNumber="377">Sabactiopus schuhi</emphasis>
</taxonomicName>
</th>
<td id="76FA5DB7FFBBC4F117ABF94CFE43FBEC" box="[414,503,1058,1080]" gridcol="1" gridrow="14" pageId="6" pageNumber="377">HO-M</td>
<td id="76FA5DB7FFBBC4F11422F94CFDF8FBEC" box="[535,588,1058,1080]" gridcol="2" gridrow="14" pageId="6" pageNumber="377">3.11</td>
<td id="76FA5DB7FFBBC4F11458F94CFD10FBEC" box="[621,676,1058,1080]" gridcol="3" gridrow="14" pageId="6" pageNumber="377">0.78</td>
<td id="76FA5DB7FFBBC4F114F6F94CFD42FBEC" box="[707,758,1058,1080]" gridcol="4" gridrow="14" pageId="6" pageNumber="377">0.26</td>
<td id="76FA5DB7FFBBC4F11522F94CFCFCFBEC" box="[791,840,1058,1080]" gridcol="5" gridrow="14" pageId="6" pageNumber="377">1.35</td>
<td id="76FA5DB7FFBBC4F1155CF94CFC2CFBEC" box="[873,920,1058,1080]" gridcol="6" gridrow="14" pageId="6" pageNumber="377">1.59</td>
<td id="76FA5DB7FFBBC4F11582F94CFC56FBEC" box="[951,994,1058,1080]" gridcol="7" gridrow="14" pageId="6" pageNumber="377">0.44</td>
<td id="76FA5DB7FFBBC4F115CAF94CFB9EFBEC" box="[1023,1066,1058,1080]" gridcol="8" gridrow="14" pageId="6" pageNumber="377">1.31</td>
<td id="76FA5DB7FFBBC4F1127DF94CFBC6FBEC" box="[1096,1138,1058,1080]" gridcol="9" gridrow="14" pageId="6" pageNumber="377">0.75</td>
<td id="76FA5DB7FFBBC4F112A4F94CFB7CFBEC" box="[1169,1224,1058,1080]" gridcol="10" gridrow="14" pageId="6" pageNumber="377">0.72</td>
<td id="76FA5DB7FFBBC4F112D2F94CFAAAFBEC" box="[1255,1310,1058,1080]" gridcol="11" gridrow="14" pageId="6" pageNumber="377">1.37</td>
<td id="76FA5DB7FFBBC4F1130BF94CFADDFBEC" box="[1342,1385,1058,1080]" gridcol="12" gridrow="14" pageId="6" pageNumber="377"></td>
<td id="76FA5DB7FFBBC4F113BCF94CFA07FBEC" box="[1417,1459,1058,1080]" gridcol="13" gridrow="14" pageId="6" pageNumber="377"></td>
</tr>
<tr id="352B34CBFFBBC4F116A2F92CFA07FB43" box="[151,1459,1090,1175]" gridrow="15" pageId="6" pageNumber="377" rowspan-0="1">
<td id="76FA5DB7FFBBC4F117ABF92CFE43FB43" box="[414,503,1090,1175]" gridcol="1" gridrow="15" pageId="6" pageNumber="377">PA-M PA-M PA-F</td>
<td id="76FA5DB7FFBBC4F11422F92CFDF8FB43" box="[535,588,1090,1175]" gridcol="2" gridrow="15" pageId="6" pageNumber="377">3.16 3.43 3.55</td>
<td id="76FA5DB7FFBBC4F11458F92CFD10FB43" box="[621,676,1090,1175]" gridcol="3" gridrow="15" pageId="6" pageNumber="377">0.83 0.80 0.83</td>
<td id="76FA5DB7FFBBC4F114F6F92CFD42FB43" box="[707,758,1090,1175]" gridcol="4" gridrow="15" pageId="6" pageNumber="377">0.26 0.24 0.30</td>
<td id="76FA5DB7FFBBC4F11522F92CFCFCFB43" box="[791,840,1090,1175]" gridcol="5" gridrow="15" pageId="6" pageNumber="377">1.41 1.35 1.55</td>
<td id="76FA5DB7FFBBC4F1155CF92CFC2CFB43" box="[873,920,1090,1175]" gridcol="6" gridrow="15" pageId="6" pageNumber="377">1.64 1.59 1.80</td>
<td id="76FA5DB7FFBBC4F11582F92CFC56FB43" box="[951,994,1090,1175]" gridcol="7" gridrow="15" pageId="6" pageNumber="377">0.45 0.45 0.45</td>
<td id="76FA5DB7FFBBC4F115CAF92CFB9EFB43" box="[1023,1066,1090,1175]" gridcol="8" gridrow="15" pageId="6" pageNumber="377">1.41 1.31 1.31</td>
<td id="76FA5DB7FFBBC4F1127DF92CFBC6FB43" box="[1096,1138,1090,1175]" gridcol="9" gridrow="15" pageId="6" pageNumber="377">0.77 0.75 0.86</td>
<td id="76FA5DB7FFBBC4F112A4F92CFB7CFB43" box="[1169,1224,1090,1175]" gridcol="10" gridrow="15" pageId="6" pageNumber="377">0.63 0.71</td>
<td id="76FA5DB7FFBBC4F112D2F92CFAAAFB43" box="[1255,1310,1090,1175]" gridcol="11" gridrow="15" pageId="6" pageNumber="377">1.32 1.38 1.53</td>
<td id="76FA5DB7FFBBC4F1130BF92CFADDFB43" box="[1342,1385,1090,1175]" gridcol="12" gridrow="15" pageId="6" pageNumber="377"> 1.20 1.50</td>
<td id="76FA5DB7FFBBC4F113BCF92CFA07FB43" box="[1417,1459,1090,1175]" gridcol="13" gridrow="15" pageId="6" pageNumber="377"> 1.95 2.10</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3016502FFBB3B0816BDF988FB37F96A" pageId="6" pageNumber="377" type="description">
<paragraph id="8BA43689FFBB3B0816BDF988FAEDF9AA" blockId="6.[136,775,1254,2014]" lastBlockId="6.[835,1475,1254,1662]" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B0816BDF988FEAAFB2A" bold="true" box="[136,286,1254,1278]" pageId="6" pageNumber="377">Description.</emphasis>
Body widely darkened posteriorly, with remaining parts pale brown or pale orange brown, oval, tortoise-shaped (
<figureCitation id="13202A0CFFBB3B081702F848FEC4FAEA" box="[311,368,1318,1342]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="6" pageNumber="377">Fig. 1</figureCitation>
CF); dorsum shining, with uniformly and rather sparsely distributed, pale, simple, reclining setae on hemelytron (
<figureCitation id="13202A0CFFBB3B081741F808FE62FAAA" box="[372,470,1382,1406]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">Fig. 19E</figureCitation>
). Head shiny pale brown or orange brown, with fuscous clypeus (
<figureCitation id="13202A0CFFBB3B081423F8E8FD33FA4A" box="[534,647,1414,1438]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="6" pageNumber="377">Fig. 1D, F</figureCitation>
); lorum and buccula slightly darkened; vertex about as wide as an eye in dorsal view, with basal transverse carina about as thick as pronotal collar (
<figureCitation id="13202A0CFFBB3B081769F888FE77FA2A" box="[348,451,1510,1534]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">Fig. 19A</figureCitation>
). Antenna dark brown; basal ¼–⅓ of antennomere I, and bases of II and III usually pale brown; antennomere II shorter than labium. Labium shiny pale brown, partly suffused with red, reaching or slightly exceeding apex of metacoxa (
<figureCitation id="13202A0CFFBB3B0817F4FB08FD85F9AA" box="[449,561,1638,1662]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="6" pageNumber="377">Fig. 1D, F</figureCitation>
). Pronotum shining, polished, widely pale brown, with a pair of dark spots on calli, sparsely bearing short, semi-erect setae laterally; pleura pale brown; metathoracic scent efferent system creamy yellow, rather rounded (
<figureCitation id="13202A0CFFBB3B0817B0FB88FE5CF92A" box="[389,488,1766,1790]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">Fig. 19D</figureCitation>
); scutellum broadly striped mesally, smooth, nearly flat, with sparsely distributed, pale, short, reclining setae (
<figureCitation id="13202A0CFFBB3B081781FA48FDA2F8EA" box="[436,534,1830,1854]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">Fig. 19E</figureCitation>
). Hemelytron shining, widely darkened posteriorly; anterior clavus and corium with striped pattern; apical half of cuneus yellowish brown; membrane pale smoky brown. All coxae and legs pale brown, partly tinged with red; trochanters and legs creamy yellow; metafemur chocolate brown, except for pale extreme base; all tarsi more or less darkened; pretarsal structures as in
<figureCitation id="13202A0CFFBB3B081557F868FC7CFACA" box="[866,968,1286,1310]" captionStart="Fig" captionStartId="27.[113,148,1898,1920]" captionTargetBox="[113,1451,330,1882]" captionTargetId="figure-11@27.[113,1451,330,1882]" captionTargetPageId="27" captionText="Fig. 17. Male (AE) and female (FH) genitalia of Paralygocoris vittulatus (AF) and P. meridionalis (GH). A, Apex of pygophore (genital segment) with parameres, dorsal view; B, right paramere; C, left paramere; D, E, vesica; F, genital chamber; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682990" httpUri="https://zenodo.org/record/14682990/files/figure.png" pageId="6" pageNumber="377">Fig. 17G</figureCitation>
; parempodia wide. Abdomen fuscous, except for male genital segment (pygophore) pale reddish brown. Male genitalia (
<figureCitation id="13202A0CFFBB3B0815C4F828FBDFFA8A" box="[1009,1131,1350,1374]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Figs. 3GJ</figureCitation>
,
<figureCitation id="13202A0CFFBB3B08124CF828FB78FA8A" box="[1145,1228,1350,1374]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">19HK</figureCitation>
): Left paramere rather L-shaped (
<figureCitation id="13202A0CFFBB3B08158EF808FBABFAAA" box="[955,1055,1382,1406]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Figs. 3G</figureCitation>
,
<figureCitation id="13202A0CFFBB3B08121AF808FBD5FAAA" box="[1071,1121,1382,1406]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">19G</figureCitation>
); right paramere with bifurcate hypophysis (
<figureCitation id="13202A0CFFBB3B0815FBF8E8FB90FA4A" box="[974,1060,1414,1438]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Fig. 3H</figureCitation>
) and a thumb-like process at apex of sensory lobe (
<figureCitation id="13202A0CFFBB3B0815EFF8C8FB82FA6A" box="[986,1078,1446,1470]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">Fig. 19J</figureCitation>
). Vesica with two slender, relatively shirt, weakly curved spicules (
<figureCitation id="13202A0CFFBB3B0812AAF8A8FB44FA0A" box="[1183,1264,1478,1502]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Fig. 3I</figureCitation>
). Female genitalia (
<figureCitation id="13202A0CFFBB3B08157EF888FC78FA2A" box="[843,972,1510,1534]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Figs. 3K, L</figureCitation>
,
<figureCitation id="13202A0CFFBB3B0815ECF888FB84FA2A" box="[985,1072,1510,1534]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">19MO</figureCitation>
): Genital chamber with ovoid, rather enlarged, thick-rimmed sclerotised rings that are relatively contiguous to each other mesally (
<figureCitation id="13202A0CFFBB3B081285FB48FABBF9EA" box="[1200,1295,1574,1598]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Figs. 3K</figureCitation>
,
<figureCitation id="13202A0CFFBB3B08132FFB48FAE4F9EA" box="[1306,1360,1574,1598]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">19M</figureCitation>
); posterior wall with wide, flat dorsal structure; and relatively wide, subtriangular interramal lobe (
<figureCitation id="13202A0CFFBB3B0812B8FB08FB5FF9AA" box="[1165,1259,1638,1662]" captionStart="Fig" captionStartId="7.[113,148,1922,1944]" captionTargetBox="[141,1451,302,1907]" captionTargetId="figure-12@7.[113,1451,302,1907]" captionTargetPageId="7" captionText="Fig. 3. Male (AD, GJ) and female (E, F, K, L) genitalia of Chelyolygus wolskiorum, new species (AF) and C. cheroti, new species (GL). A, Apex of pygophore; B, G, left paramere; C, H, right paramere; D, I, J, vesica; E, K, genital chamber with sclerotised rings and adjacent structures; F, L, posterior wall. Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682954" httpUri="https://zenodo.org/record/14682954/files/figure.png" pageId="6" pageNumber="377">Figs. 3L</figureCitation>
,
<figureCitation id="13202A0CFFBB3B0812CCFB08FAF8F9AA" box="[1273,1356,1638,1662]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="6" pageNumber="377">19N, O</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFBB3B081576FBC8FB37F96A" blockId="6.[835,1155,1702,1726]" box="[835,1155,1702,1726]" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081576FBC8FC46F96A" bold="true" box="[835,1010,1702,1726]" pageId="6" pageNumber="377">Measurements.</emphasis>
See
<tableCitation id="C6990332FFBB3B08121CFBC8FBC8F96A" box="[1065,1148,1702,1726]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="6" pageNumber="377">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBB3B081576FB88FCCDF8EA" pageId="6" pageNumber="377" type="etymology">
<paragraph id="8BA43689FFBB3B081576FB88FCCDF8EA" blockId="6.[835,1474,1766,1854]" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081576FB88FC71F92A" bold="true" box="[835,965,1766,1790]" pageId="6" pageNumber="377">Etymology.</emphasis>
Named after Dr. Frédéric Chérot, an eminent Belgian heteropterist and miridologist; a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBB3B081576FA08FB20F8AA" box="[835,1172,1894,1918]" pageId="6" pageNumber="377" type="distribution">
<paragraph id="8BA43689FFBB3B081576FA08FB20F8AA" blockId="6.[835,1172,1894,1918]" box="[835,1172,1894,1918]" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081576FA08FC62F8AA" bold="true" box="[835,982,1894,1918]" pageId="6" pageNumber="377">Distribution.</emphasis>
<collectingCountry id="F30C7619FFBB3B0815EBFA08FBF2F8AA" box="[990,1094,1894,1918]" name="Indonesia" pageId="6" pageNumber="377">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFFBB3B081263FA08FB33F8AA" box="[1110,1159,1894,1918]" country="Indonesia" name="Bali" pageId="6" pageNumber="377">Bali</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBB3B081576FAC8FBADF86A" box="[835,1049,1958,1982]" pageId="6" pageNumber="377" type="biology_ecology">
<paragraph id="8BA43689FFBB3B081576FAC8FBADF86A" blockId="6.[835,1049,1958,1982]" box="[835,1049,1958,1982]" pageId="6" pageNumber="377">
<emphasis id="B96FEA9BFFBB3B081576FAC8FC14F86A" bold="true" box="[835,928,1958,1982]" pageId="6" pageNumber="377">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,265 @@
<document id="ED6939DA74F1DD0BB56C6ED77BB81264" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB93B0A168CF81EFB2AFA81" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus bipuncticollis Yasunaga 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="375" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="375" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="2A841002EB279E9867AC07C6BDB512E6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07FA8B8FF3A5A4E0F26DBB7BE34919C2">
<mods:title id="FE5139A38857D9ABD6D7FD55337566F8">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="25B054984952CE2B58521AAB1AE47C0A" type="personal">
<mods:role id="C147EB8AAF78929E11E9F50B5AFA6644">
<mods:roleTerm id="5060C3ED7FD25AC254D280CA09D1412B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46D6D4383F817ADD847F94F306C32805">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="3E0E54D6C95C316E07EA5C02BC12F9AB">text</mods:typeOfResource>
<mods:relatedItem id="C6AE0AF66F9461E2F2FD7F11A3CFC61C" type="host">
<mods:titleInfo id="029D9EE244F7DCD23E44F7470B167982">
<mods:title id="D62785FA42B812A23E554BBEFC375C92">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CE32467A00BD15990F92E43B0D754E87">
<mods:date id="D0E6A1AB9FEBA0B8D63703570F573120">2024</mods:date>
<mods:detail id="4D4D7F1E56F470E805D7ECFD3ACADB55" type="pubDate">
<mods:number id="E5A84A13CB849E595EB8B8638C8E0BB9">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="7F97D03E02FBCC0FF2AE4ED7709D3FB8" type="volume">
<mods:number id="3C4D97693EE9F40A4E70230FBD97091D">72</mods:number>
</mods:detail>
<mods:extent id="1A620DC28B0D4E83D77B3ECF455D77A2" unit="page">
<mods:start id="EA10EECA1B97531E8A055F15ED67706A">371</mods:start>
<mods:end id="CBDBB52FDBD21B1E8594F921567659A0">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A8F6F80FB6887BEF42322D0498B454B2">journal article</mods:classification>
<mods:identifier id="F3F2206B32CA0999DB63DC977E735DCE" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="8772F9C6F8DFD0E5993F57F0F4FD0B9B" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A7FC3F34B48A0156C37152CC203C565D" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="05366C5A92F1EF3AD4E6207D9B30C1FB" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB93B0A168CF81EFB2AFA81" ID-DOI="http://doi.org/10.5281/zenodo.14682956" ID-Zenodo-Dep="14682956" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0A168CF81EFB2AFA81" httpUri="http://treatment.plazi.org/id/03B2879FFFB93B0A168CF81EFB2AFA81" lastPageNumber="375" pageId="4" pageNumber="375">
<subSubSection id="C3016502FFB93B0A168CF81EFDBAFA7C" pageId="4" pageNumber="375" type="nomenclature">
<paragraph id="8BA43689FFB93B0A168CF81EFDBAFA7C" blockId="4.[185,725,1392,1481]" pageId="4" pageNumber="375">
<heading id="D0EC81E5FFB93B0A168CF81EFDBAFA7C" bold="true" centered="true" fontSize="10" level="2" pageId="4" pageNumber="375" reason="2">
<emphasis id="B96FEA9BFFB93B0A168CF81EFDBAFA7C" bold="true" pageId="4" pageNumber="375">
<taxonomicName id="4C1B4D0AFFB93B0A168CF81EFD2EFA5C" authority="(Poppius, 1915)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[185,666,1392,1416]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis" status="comb. nov.">
<emphasis id="B96FEA9BFFB93B0A168CF81EFE69FA5C" bold="true" box="[185,477,1392,1416]" italics="true" pageId="4" pageNumber="375">Chelyolygus bipuncticollis</emphasis>
(
<bibRefCitation id="EF8A4B78FFB93B0A17D8F81EFD26FA5C" author="Poppius B" box="[493,658,1392,1416]" pageId="4" pageNumber="375" pagination="1 - 80" refId="ref24198" refString="Poppius B (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8): 1 - 80 (1914)." type="journal article" year="1915">Poppius, 1915</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A1492F81FFDBAFA7C" pageId="4" pageNumber="375" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1707F8DEFDE8FA1D" box="[306,604,1456,1481]" pageId="4" pageNumber="375" type="description">
<paragraph id="8BA43689FFB93B0A1707F8DEFDE8FA1D" blockId="4.[185,725,1392,1481]" box="[306,604,1456,1481]" pageId="4" pageNumber="375">
(
<figureCitation id="13202A0CFFB93B0A170FF8DEFE1EFA1C" box="[314,426,1456,1481]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Figs. 1I, J</figureCitation>
,
<figureCitation id="13202A0CFFB93B0A178DF8DFFE71FA1D" box="[440,453,1457,1481]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">2</figureCitation>
AD, 20AF)
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A16BDF89EFE8EF9AC" pageId="4" pageNumber="375" type="reference_group">
<paragraph id="8BA43689FFB93B0A16BDF89EFDB8F9D1" blockId="4.[136,775,1519,1656]" box="[136,524,1519,1541]" pageId="4" pageNumber="375">
<treatmentCitationGroup id="AB0B11A7FFB93B0A16BDF89EFDB8F9D1" box="[136,524,1519,1541]" pageId="4" pageNumber="375">
<treatmentCitation id="0ABA1098FFB93B0A16BDF89EFDBCF9D1" author="Poppius B" box="[136,520,1519,1541]" page="25" pageId="4" pageNumber="375" year="1915">
<taxonomicName id="4C1B4D0AFFB93B0A16BDF89EFDBCF9D1" authority="Poppius, 1915: 25" authorityName="Poppius" authorityPageNumber="25" authorityYear="1915" box="[136,520,1519,1541]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis">
<emphasis id="B96FEA9BFFB93B0A16BDF89EFEF8F9D0" box="[136,332,1519,1541]" italics="true" pageId="4" pageNumber="375">Lygus bipuncticollis</emphasis>
<bibRefCitation id="EF8A4B78FFB93B0A1761F881FDBCF9D1" author="Poppius B" box="[340,520,1519,1541]" pageId="4" pageNumber="375" pagination="1 - 80" refId="ref24198" refString="Poppius B (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8): 1 - 80 (1914)." type="journal article" year="1915">Poppius, 1915: 25</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFB63FE81F9EB" blockId="4.[136,775,1519,1656]" pageId="4" pageNumber="375">
<treatmentCitationGroup id="AB0B11A7FFB93B0A16BDFB63FE81F9EB" pageId="4" pageNumber="375">
<taxonomicName id="4C1B4D0AFFB93B0A16BDFB63FE55F9F5" ID-CoL="6R389" authorityName="Lu &amp; Zheng" authorityYear="1998" box="[136,481,1548,1570]" class="Insecta" family="Miridae" genus="Lygocoris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis" subGenus="Neolygus">
<emphasis id="B96FEA9BFFB93B0A16BDFB63FF5CF9F6" box="[136,232,1549,1570]" italics="true" pageId="4" pageNumber="375">Lygocoris</emphasis>
(
<emphasis id="B96FEA9BFFB93B0A16C3FB62FEFBF9F5" box="[246,335,1548,1569]" italics="true" pageId="4" pageNumber="375">Neolygus</emphasis>
)
<emphasis id="B96FEA9BFFB93B0A176BFB62FE55F9F5" box="[350,481,1548,1569]" italics="true" pageId="4" pageNumber="375">bipuncticollis</emphasis>
</taxonomicName>
:
<treatmentCitation id="0ABA1098FFB93B0A17D8FB62FD65F9F6" author="Lu N &amp; Zheng LY" box="[493,721,1548,1570]" page="186" pageId="4" pageNumber="375" year="1998">
<bibRefCitation id="EF8A4B78FFB93B0A17D8FB62FD65F9F6" author="Lu N &amp; Zheng LY" box="[493,721,1548,1570]" pageId="4" pageNumber="375" pagination="185 - 189" refId="ref24014" refString="Lu N &amp; Zheng LY (1998) Identity of some ' Lygus ' species described from Taiwan by Poppius (Heteroptera: Miridae). Tijdschrift voor Entomologie, 140: 185 - 189." type="journal article" year="1998">Lu &amp; Zheng, 1998: 186</bibRefCitation>
</treatmentCitation>
(new combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFB2BFE8EF9AC" blockId="4.[136,775,1519,1656]" pageId="4" pageNumber="375">
<treatmentCitationGroup id="AB0B11A7FFB93B0A16BDFB2BFE8EF9AC" pageId="4" pageNumber="375">
<taxonomicName id="4C1B4D0AFFB93B0A16BDFB2BFEDDF98E" ID-CoL="747MX" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[136,361,1605,1626]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis">
<emphasis id="B96FEA9BFFB93B0A16BDFB2BFEDDF98E" box="[136,361,1605,1626]" italics="true" pageId="4" pageNumber="375">Neolygus bipuncticollis</emphasis>
</taxonomicName>
:
<treatmentCitation id="0ABA1098FFB93B0A1740FB2BFDFBF98F" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[373,591,1605,1627]" page="388" pageId="4" pageNumber="375" year="2004">
<bibRefCitation id="EF8A4B78FFB93B0A1740FB2BFDFBF98F" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[373,591,1605,1627]" pageId="4" pageNumber="375" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng et al., 2004: 388</bibRefCitation>
</treatmentCitation>
(new combination, redescription).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A16BDFBCFFE29F84C" pageId="4" pageNumber="375" type="materials_examined">
<paragraph id="8BA43689FFB93B0A16BDFBCFFE29F84C" blockId="4.[136,776,1696,1945]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A16BDFBCFFEF6F96C" bold="true" box="[136,322,1696,1721]" pageId="4" pageNumber="375">Type material.</emphasis>
<materialsCitation id="3B733CD4FFB93B0A177BFBCFFCB0F8CC" collectingDate="1912-07" collectorName="H. Sauter &amp; Senckenberg German Entomological Institute" country="Taiwan" county="Male" latitude="22.0" location="Koshun" longLatPrecision="1262" longitude="120.75" municipality="Kankau" pageId="4" pageNumber="375" specimenCount="1" stateProvince="Pingtung" typeStatus="holotype">
<emphasis id="B96FEA9BFFB93B0A177BFBCFFE0AF96D" bold="true" box="[334,446,1697,1721]" pageId="4" pageNumber="375">
<typeStatus id="54A0882BFFB93B0A177BFBCFFE0AF96D" box="[334,446,1697,1721]" pageId="4" pageNumber="375" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCounty id="62C54E05FFB93B0A17E6FBCEFDA5F96C" box="[467,529,1696,1720]" pageId="4" pageNumber="375">Male</collectingCounty>
,
<emphasis id="B96FEA9BFFB93B0A1414FBCEFD16F96C" bold="true" box="[545,674,1696,1720]" pageId="4" pageNumber="375">
<collectingCountry id="F30C7619FFB93B0A1414FBCEFD28F96C" box="[545,668,1696,1720]" name="Taiwan" pageId="4" pageNumber="375">TAIWAN</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BC0ACF3FFB93B0A149BFBCEFCB2F96C" box="[686,774,1696,1720]" pageId="4" pageNumber="375">Kankau</collectingMunicipality>
(
<location id="8EC46052FFB93B0A16BAFBAEFF51F90C" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0A168CF81EFB2AFA81:8EC46052FFB93B0A16BAFBAEFF51F90C" box="[143,229,1728,1752]" country="Taiwan" county="Male" latitude="22.0" longLatPrecision="1262" longitude="120.75" municipality="Kankau" name="Koshun" pageId="4" pageNumber="375" stateProvince="Pingtung">Koshun</location>
) [= currently Hengchun of
<collectingRegion id="49DFF86BFFB93B0A1439FBAEFDD8F90C" box="[524,620,1728,1752]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="375">Pingtung</collectingRegion>
Co.,
<geoCoordinate id="EE2F504EFFB93B0A1492FBD1FCB7F90C" box="[679,771,1727,1752]" degrees="22" direction="north" minutes="00" orientation="latitude" pageId="4" pageNumber="375" precision="925" value="22.0">22°00N</geoCoordinate>
,
<geoCoordinate id="EE2F504EFFB93B0A16BDFBB1FF44F92C" box="[136,240,1759,1784]" degrees="120" direction="east" minutes="45" orientation="longitude" pageId="4" pageNumber="375" precision="925" value="120.75">120°45E</geoCoordinate>
],
<collectorName id="26EE535FFFB93B0A1736FB8FFED8F92D" box="[259,364,1761,1785]" pageId="4" pageNumber="375">H. Sauter</collectorName>
,
<date id="FFA51049FFB93B0A174DFB8EFE55F92D" box="[376,481,1760,1785]" pageId="4" pageNumber="375" value="1912-07">
<collectingDate id="EFE1E9A1FFB93B0A174DFB8EFE55F92D" box="[376,481,1760,1785]" pageId="4" pageNumber="375" value="1912-07">July 1912</collectingDate>
</date>
(deposited in
<collectorName id="26EE535FFFB93B0A1448FB8EFDA5F8CC" pageId="4" pageNumber="375">Senckenberg German Entomological Institute</collectorName>
) [image examined]
</materialsCitation>
.
<materialsCitation id="3B733CD4FFB93B0A16BDFA4EFE2DF84C" collectingDate="2012-12-25" collectionCode="NMNS, AMNH" collectorName="J. F. Tsai" country="Taiwan" county="Kenting National Park" latitude="22.033333" location="Badenixi Stream" longLatPrecision="1261" longitude="120.833336" municipality="Xianglin Village" pageId="4" pageNumber="375" specimenCode="PBI 00378775" specimenCount="1" specimenCount-male="1" stateProvince="Pingtung">
Additional material:
<emphasis id="B96FEA9BFFB93B0A1749FA4EFE4EF8EC" bold="true" box="[380,506,1824,1848]" pageId="4" pageNumber="375">
<collectingCountry id="F30C7619FFB93B0A1749FA4EFE41F8EC" box="[380,501,1824,1848]" name="Taiwan" pageId="4" pageNumber="375">TAIWAN</collectingCountry>
:
</emphasis>
<specimenCount id="9D1DFD00FFB93B0A1433FA4FFDE3F8EC" box="[518,599,1824,1849]" count="1" pageId="4" pageNumber="375" type="male">1 male</specimenCount>
,
<collectingRegion id="49DFF86BFFB93B0A1452FA4EFD78F8EC" box="[615,716,1824,1848]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="375">Pingtung</collectingRegion>
Co.,
<collectingCounty id="62C54E05FFB93B0A16BDFA2EFECAF88C" box="[136,382,1856,1880]" pageId="4" pageNumber="375">Kenting National Park</collectingCounty>
,
<collectingMunicipality id="6BC0ACF3FFB93B0A17BFFA2EFDF4F88C" box="[394,576,1856,1880]" pageId="4" pageNumber="375">Xianglin Village</collectingMunicipality>
,
<location id="8EC46052FFB93B0A1479FA2EFCB7F88D" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0A168CF81EFB2AFA81:8EC46052FFB93B0A1479FA2EFCB7F88D" box="[588,771,1856,1881]" country="Taiwan" county="Kenting National Park" latitude="22.033333" longLatPrecision="1261" longitude="120.833336" municipality="Xianglin Village" name="Badenixi Stream" pageId="4" pageNumber="375" stateProvince="Pingtung">Badenixi Stream</location>
,
<geoCoordinate id="EE2F504EFFB93B0A16BDFA31FF55F8AC" box="[136,225,1887,1912]" degrees="22" direction="north" minutes="02" orientation="latitude" pageId="4" pageNumber="375" precision="925" value="22.033333">22°02N</geoCoordinate>
<geoCoordinate id="EE2F504EFFB93B0A16DFFA31FEE4F8AC" box="[234,336,1887,1912]" degrees="120" direction="east" minutes="50" orientation="longitude" pageId="4" pageNumber="375" precision="925" value="120.833336">120°50E</geoCoordinate>
,
<collectorName id="26EE535FFFB93B0A1769FA0FFE70F8AC" box="[348,452,1888,1912]" pageId="4" pageNumber="375">J. F. Tsai</collectorName>
,
<date id="FFA51049FFB93B0A17E4FA0EFD2BF8AD" box="[465,671,1888,1913]" pageId="4" pageNumber="375" value="2012-12-25">
<collectingDate id="EFE1E9A1FFB93B0A17E4FA0EFD2BF8AD" box="[465,671,1888,1913]" pageId="4" pageNumber="375" value="2012-12-25">25 December 2012</collectingDate>
</date>
(
<collectionCode id="ED0AAE4CFFB93B0A149AFA0FFD4BF8AD" box="[687,767,1889,1913]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="4" pageNumber="375">NMNS</collectionCode>
) (
<collectionCode id="ED0AAE4CFFB93B0A16BAFAEFFF56F84D" box="[143,226,1921,1945]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="375">AMNH</collectionCode>
_
<specimenCode id="DBBD9EF2FFB93B0A16C5FAEFFE26F84C" box="[240,402,1920,1945]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="375">PBI 00378775</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A16BDFAAEFBE9FE21" pageId="4" pageNumber="375" type="diagnosis">
<paragraph id="8BA43689FFB93B0A16BDFAAEFBE9FE21" blockId="4.[136,775,1984,2072]" lastBlockId="4.[835,1475,189,502]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A16BDFAAEFEB4F80C" bold="true" box="[136,256,1984,2008]" pageId="4" pageNumber="375">Diagnosis.</emphasis>
This new species is currently known by
<specimenCount id="9D1DFD00FFB93B0A14EBFAAFFF08F82C" count="2" pageId="4" pageNumber="375" type="male">two male</specimenCount>
specimens mentioned above and recognised readily by the following characters: Orange-yellow basic colouration; oily-shiny, highly polished, impunctate and hairless dorsum (
<figureCitation id="13202A0CFFB93B0A1579FDB3FC00FF21" box="[844,948,221,245]" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="4" pageNumber="375">Fig. 20A</figureCitation>
) with characteristic fuscous spots or maculae on pronotum, scutellum and hemelytron (
<figureCitation id="13202A0CFFB93B0A133AFD93FACCFEC1" box="[1295,1400,253,277]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1I, J</figureCitation>
); fully darkened clypeus; darkened apical ¼ of pale antennomere I; and unique form of male genitalia as described below. Further external features were described by
<bibRefCitation id="EF8A4B78FFB93B0A132BFC33FA76FEA1" author="Poppius B" box="[1310,1474,349,373]" pageId="4" pageNumber="375" pagination="1 - 80" refId="ref24198" refString="Poppius B (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8): 1 - 80 (1914)." type="journal article" year="1915">Poppius (1915)</bibRefCitation>
and images of the
<typeStatus id="54A0882BFFB93B0A1233FC13FBD7FE41" box="[1030,1123,381,405]" pageId="4" pageNumber="375" type="holotype">holotype</typeStatus>
male are available on a database (
<bibRefCitation id="EF8A4B78FFB93B0A157EFCF3FAF5FE61" author="National Museum of Natural Science" box="[843,1345,413,437]" pageId="4" pageNumber="375" refId="ref24112" refString="National Museum of Natural Science (2023) Integrated insect types database of Taiwanese species. http: // twinsecttype. nmns. edu. tw / specimens / (Accessed 25 August 2024)." type="url" year="2023">National Museum of Natural Science, 2023</bibRefCitation>
). The male genitalia are herein described for the first time, but the female is yet to be found.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1576FF73FB37FCE1" pageId="4" pageNumber="375" type="description">
<paragraph id="8BA43689FFB93B0A1576FF73FB88FD21" blockId="4.[835,1475,541,758]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FF73FC66FDE1" bold="true" box="[835,978,541,565]" pageId="4" pageNumber="375">Description.</emphasis>
Male genitalia (
<figureCitation id="13202A0CFFB93B0A12A5FF73FABAFDE1" box="[1168,1294,541,565]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">Fig. 2AH</figureCitation>
): Left paramere C-shaped, similar in general shape to that of
<taxonomicName id="4C1B4D0AFFB93B0A130AFF53FC26FDA1" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A130AFF53FA76FD81" box="[1343,1474,573,597]" italics="true" pageId="4" pageNumber="375">Apolygopsis</emphasis>
species
</taxonomicName>
(but apex of hypophysis lacking lateral process) (
<figureCitation id="13202A0CFFB93B0A157EFF13FC0AFD41" box="[843,958,637,661]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">Fig. 2D, E</figureCitation>
); right paramere with bifurcate hypophysis (
<figureCitation id="13202A0CFFB93B0A13A2FF13FC3CFD61" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">Fig. 2G, H</figureCitation>
) and apical protuberance on sensory lobe (
<figureCitation id="13202A0CFFB93B0A1357FFF3FA02FD61" box="[1378,1462,669,693]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">Fig. 2F</figureCitation>
). Vesica with two stout spicules and small, rounded secondary gonopore (
<figureCitation id="13202A0CFFB93B0A158CFFB3FB84FD21" box="[953,1072,733,757]" captionStart="Fig" captionStartId="3.[113,148,1878,1900]" captionTargetBox="[113,1451,352,1863]" captionTargetId="figure-11@3.[113,1451,352,1863]" captionTargetPageId="3" captionText="Fig. 2. Male (AE) and female (FH) genitalia of Chelyolygus bipuncticollis (AE) and C. tabananicus, new species (FH). A, B, Vesica; C, left paramere; D, E, right paramere; F, genital chamber with sclerotised rings and adjacent structures; G, posterior wall; H, apex of ovipositor (gonapophysis I). Scale bars 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682952" httpUri="https://zenodo.org/record/14682952/files/figure.png" pageId="4" pageNumber="375">Fig. 2AC</figureCitation>
).
</paragraph>
<paragraph id="8BA43689FFB93B0A1576FE73FB37FCE1" blockId="4.[835,1155,797,821]" box="[835,1155,797,821]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FE73FC46FCE1" bold="true" box="[835,1010,797,821]" pageId="4" pageNumber="375">Measurements.</emphasis>
See
<tableCitation id="C6990332FFB93B0A121CFE73FBC9FCE1" box="[1065,1149,797,821]" captionStart="Table 1" captionStartId="5.[113,168,189,211]" captionText="Table 1. Measurements for the species treated. Abbreviations — F: female, FM: femur, HO: holotype, L: length, LBM: labium, M:" pageId="4" pageNumber="375">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1576FE33FB04FCA1" box="[835,1200,861,885]" pageId="4" pageNumber="375" type="distribution">
<paragraph id="8BA43689FFB93B0A1576FE33FB04FCA1" blockId="4.[835,1200,861,885]" box="[835,1200,861,885]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FE33FC62FCA1" bold="true" box="[835,982,861,885]" pageId="4" pageNumber="375">Distribution.</emphasis>
<collectingCountry id="F30C7619FFB93B0A15EBFE33FB9AFCA1" box="[990,1070,861,885]" name="Taiwan" pageId="4" pageNumber="375">Taiwan</collectingCountry>
(
<collectingRegion id="49DFF86BFFB93B0A120BFE33FB11FCA1" box="[1086,1189,861,885]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="375">Pingtung</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1576FEF3FBADFC61" box="[835,1049,925,949]" pageId="4" pageNumber="375" type="biology_ecology">
<paragraph id="8BA43689FFB93B0A1576FEF3FBADFC61" blockId="4.[835,1049,925,949]" box="[835,1049,925,949]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FEF3FC14FC61" bold="true" box="[835,928,925,949]" pageId="4" pageNumber="375">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A1576FEB3FB2AFA81" pageId="4" pageNumber="375" type="discussion">
<paragraph id="8BA43689FFB93B0A1576FEB3FB2AFA81" blockId="4.[835,1474,989,1365]" pageId="4" pageNumber="375">
<emphasis id="B96FEA9BFFB93B0A1576FEB3FC00FC21" bold="true" box="[835,948,989,1013]" pageId="4" pageNumber="375">Remarks.</emphasis>
This Taiwanese taxon has been regarded as a member of the speciose Holarctic genus
<taxonomicName id="4C1B4D0AFFB93B0A132CFE93FACBFBC1" authorityName="Knight" authorityYear="1917" box="[1305,1407,1021,1045]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFB93B0A132CFE93FACBFBC1" box="[1305,1407,1021,1045]" italics="true" pageId="4" pageNumber="375">Neolygus</emphasis>
</taxonomicName>
since the treatment by
<bibRefCitation id="EF8A4B78FFB93B0A15C9F973FB64FBE1" author="Zheng LY &amp; Lu N &amp; Liu G &amp; Xu B" box="[1020,1232,1053,1077]" pageId="4" pageNumber="375" refId="ref24998" refString="Zheng LY, Lu N, Liu G &amp; Xu B (2004) Hemiptera, Miridae, Mirinae. Fauna Sinica, Insecta, Vol. 33. Science Press, Beijing, xix + 797 pp., 8 pls." type="book" year="2004">Zheng et al. (2004)</bibRefCitation>
. However, the present examination on the surface structures and male genitalia warrants its placement in
<taxonomicName id="4C1B4D0AFFB93B0A1246F933FB49FBA1" authorityName="Yasunaga" authorityYear="2024" box="[1139,1277,1117,1141]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFB93B0A1246F933FB49FBA1" box="[1139,1277,1117,1141]" italics="true" pageId="4" pageNumber="375">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A1325F930FA3AFBA2" box="[1296,1422,1118,1142]" pageId="4" pageNumber="375" rank="genus">new genus</taxonomicNameLabel>
. For instance, the highly polished, nearly glabrous dorsal surface is not exhibited in
<taxonomicName id="4C1B4D0AFFB93B0A1225F9F3FBC7FB61" authorityName="Knight" authorityYear="1917" box="[1040,1139,1181,1205]" class="Insecta" family="Miridae" genus="Neolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFB93B0A1225F9F3FBC7FB61" box="[1040,1139,1181,1205]" italics="true" pageId="4" pageNumber="375">Neolygus</emphasis>
</taxonomicName>
, and the principal morphology of the male and female genitalia is significantly different between the two genera (cf.
<bibRefCitation id="EF8A4B78FFB93B0A12B5F9B0FA86FB21" author="Yasunaga T" box="[1152,1330,1245,1270]" pageId="4" pageNumber="375" pagination="435 - 448" refId="ref24458" refString="Yasunaga T (1991) A revision of the plant bug genus Lygocoris Reuter from Japan, Part I (Heteroptera, Miridae, Lygus - complex). Japanese Journal of Entomology, 59: 435 - 448." type="journal article" year="1991">Yasunaga, 1991</bibRefCitation>
, 2023, 2024;
<bibRefCitation id="EF8A4B78FFB93B0A1576F990FBFDFAC1" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" box="[835,1097,1277,1302]" pageId="4" pageNumber="375" pagination="357 - 387" refId="ref24824" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2018) Review of the plant bug genus Prolygus and related mirine taxa from eastern Asia (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae, 58: 357 - 387." type="journal article" year="2018">Yasunaga et al., 2018</bibRefCitation>
). Based on these morphological evidences, a new combination,
<taxonomicName id="4C1B4D0AFFB93B0A1294F873FC19FA81" authority="(Poppius)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis">
<emphasis id="B96FEA9BFFB93B0A1294F873FA76FAE1" box="[1185,1474,1309,1333]" italics="true" pageId="4" pageNumber="375">Chelyolygus bipuncticollis</emphasis>
(Poppius)
</taxonomicName>
, is herein established.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,200 @@
<document id="C362B9C153272FB66D25BCC9D747712F" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFB93B0A16BDFC73FCB1FA9C" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus Yasunaga, 2024, new genus" docType="key" docVersion="2" lastPageNumber="375" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="375" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="3B2B2A42966AA0531531F26EA01141C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="557F0A4E3D17C95620EA570424783A36">
<mods:title id="26702AB01D57FBBFB7328CCDC4A68663">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="17CEA6B52F63551C58FE9BDB52884D33" type="personal">
<mods:role id="8670BB6301447E945AA25FBACD966F18">
<mods:roleTerm id="2572676C98D8CB9C23070A846DA6561B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0A22DFD1ACEEB938363B478C593CD369">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="12B3A3819F9A56C193F6E9ECB917D293">text</mods:typeOfResource>
<mods:relatedItem id="D0F63E710D1A719C9E0F7DBA97F0BEF8" type="host">
<mods:titleInfo id="41B09AF313DD75618DF9849E22ABD401">
<mods:title id="60441648B397D24B7BC9A1146A6B352D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="15D06BB3C26E640B1C557596BF4A9714">
<mods:date id="19F4EF940098C33FA76E876917B576DF">2024</mods:date>
<mods:detail id="1ADF8241C418A429CA4606EEBD8F59D9" type="pubDate">
<mods:number id="E95A18A78F59657F2589F986A7B35311">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="74E6B6F7A6D7BA5599BCE37F7B79194D" type="volume">
<mods:number id="1B3F3FD96323EA7E8FD1BAA42F404BA1">72</mods:number>
</mods:detail>
<mods:extent id="2DB088F08DE27CFB7F239C51AC361F7C" unit="page">
<mods:start id="962BBBA8919E40392D3B7C329ADC9708">371</mods:start>
<mods:end id="35072797A5878C41870CC303BAF68B1A">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E7D451F567764B227E8D9F08C4F0481A">journal article</mods:classification>
<mods:identifier id="28A63BDB87DACC2C0ADA9A07D7DE39A5" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="DCCD8F6C4456A6C94455E2FAE4A824C6" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="38DCA2016F3DFA0D99559BCE5C08BD0C" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="97640007AB38264A436565D6A25EF4E1" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFB93B0A16BDFC73FCB1FA9C" ID-DOI="http://doi.org/10.5281/zenodo.14682950" ID-Zenodo-Dep="14682950" LSID="urn:lsid:plazi:treatment:03B2879FFFB93B0A16BDFC73FCB1FA9C" httpUri="http://treatment.plazi.org/id/03B2879FFFB93B0A16BDFC73FCB1FA9C" lastPageNumber="375" pageId="4" pageNumber="375">
<subSubSection id="C3016502FFB93B0A16BDFC73FDD5FEE1" box="[136,609,285,309]" pageId="4" pageNumber="375" type="nomenclature">
<paragraph id="8BA43689FFB93B0A16BDFC73FDD5FEE1" blockId="4.[136,609,285,309]" box="[136,609,285,309]" pageId="4" pageNumber="375">
<heading id="D0EC81E5FFB93B0A16BDFC73FDD5FEE1" bold="true" box="[136,609,285,309]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="375" reason="2">
<emphasis id="B96FEA9BFFB93B0A16BDFC73FDD5FEE1" bold="true" box="[136,609,285,309]" pageId="4" pageNumber="375">
Key to species of
<taxonomicName id="4C1B4D0AFFB93B0A1761FC73FE6FFEE1" authorityName="Yasunaga" authorityYear="2024" box="[340,475,285,309]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFB93B0A1761FC73FE6FFEE1" bold="true" box="[340,475,285,309]" italics="true" pageId="4" pageNumber="375">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A17DFFC73FDD5FEE1" box="[490,609,285,309]" pageId="4" pageNumber="375" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFB93B0A16BFFC32FCB1FA9C" pageId="4" pageNumber="375" type="key">
<keyStep id="30EF2E2CFFB93B0A16BFFC32FCB1FDD5" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BFFC32FCB1FE7F" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BFFC32FCB1FE7F" pageId="4" pageNumber="375">
1. Scutellum with a dark, broad, median stripe in whole length; posterior part of corium uniformly fuscous (
<figureCitation id="13202A0CFFB93B0A14B3FC17FD4EFE5A" box="[646,762,377,399]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1C, E</figureCitation>
);
<collectingCountry id="F30C7619FFB93B0A169BFCFBFEBFFE7F" box="[174,267,405,427]" name="Indonesia" pageId="4" pageNumber="375">Indonesia</collectingCountry>
(
<collectingRegion id="49DFF86BFFB93B0A172FFCFBFEF0FE7F" box="[282,324,405,427]" country="Indonesia" name="Bali" pageId="4" pageNumber="375">Bali</collectingRegion>
) ...................................
<taxonomicName id="4C1B4D0AFFB93B0A1414FCF8FD36FE7E" authorityName="Yasunaga" authorityYear="2024" box="[545,642,405,427]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="cheroti" status="new species">
<emphasis id="B96FEA9BFFB93B0A1414FCF8FD36FE7E" box="[545,642,405,427]" italics="true" pageId="4" pageNumber="375">C. cheroti</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A14A5FCF8FCB1FE7F" box="[656,773,405,427]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFCDDFCB1FDD5" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFCDDFCB1FDD5" pageId="4" pageNumber="375">
Scutellum pale, or with a triangular mark anteriorly (
<figureCitation id="13202A0CFFB93B0A1483FCDCFD4BFE13" box="[694,767,434,456]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1E</figureCitation>
) or with faint, brown, narrow, mesal stripe (
<figureCitation id="13202A0CFFB93B0A1464FCA1FD2AFE30" box="[593,670,463,485]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1G</figureCitation>
); posterior part of corium with noticeable pale portions .........................2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFB93B0A16BDFF67FCB1FD1F" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BDFF67FCB1FDA1" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFF67FCB1FDA1" pageId="4" pageNumber="375">
2. Basic colouration pale orange-brown (
<figureCitation id="13202A0CFFB93B0A1423FF67FDDEFDCA" box="[534,618,521,543]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1I, J</figureCitation>
); both pronotum and hemelytra almost glabrous, very shiny; basal half of cuneus fuscous;
<collectingCountry id="F30C7619FFB93B0A1733FF2CFEFAFD8C" box="[262,334,578,600]" name="Taiwan" pageId="4" pageNumber="375">Taiwan</collectingCountry>
(
<collectingRegion id="49DFF86BFFB93B0A1769FF2CFE02FD8C" box="[348,438,578,600]" country="Taiwan" name="Pingtung" pageId="4" pageNumber="375">Pingtung</collectingRegion>
) ...................................................... ...............
<taxonomicName id="4C1B4D0AFFB93B0A1739FF31FDE6FDA1" authority="(Poppius, 1915)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[268,594,607,629]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="bipuncticollis" status="new combination">
<emphasis id="B96FEA9BFFB93B0A1739FF31FE19FDA0" box="[268,429,607,628]" italics="true" pageId="4" pageNumber="375">C. bipuncticollis</emphasis>
(
<bibRefCitation id="EF8A4B78FFB93B0A178EFF31FDFFFDA1" author="Poppius B" box="[443,587,607,629]" pageId="4" pageNumber="375" pagination="1 - 80" refId="ref24198" refString="Poppius B (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8): 1 - 80 (1914)." type="journal article" year="1915">Poppius, 1915</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A146BFF0EFCB1FDA1" box="[606,773,607,629]" pageId="4" pageNumber="375" rank="species">new combination</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFF12FCB1FD1F" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFF12FCB1FD1F" pageId="4" pageNumber="375"> Basic colouration pale brown, or milky- or ivory-brown; pronotum and/or hemelytra setose; base of cuneus always pale (median part of cuneus sometimes darkened)........................3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFB93B0A16BDFFBCFCB2FCF6" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BDFFBCFCB1FCD0" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFFBCFCB1FCD0" pageId="4" pageNumber="375">3. Apex of scutellum and inner margin of clavus (at least anterior half) infuscate..........................................................................4</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFE62FCB2FCF6" blockId="4.[136,776,348,1352]" box="[136,774,780,802]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFE62FCB2FCF6" box="[136,774,780,802]" pageId="4" pageNumber="375"> Entire scutellum and inner margin of clavus pale.................6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFB93B0A16BDFE47FCB1FC66" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BDFE47FCB1FCAC" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFE47FCB1FCAC" pageId="4" pageNumber="375">
4. Base of antennomere II darkened; cuneus sometimes with a circular spot medially (
<figureCitation id="13202A0CFFB93B0A17A1FE2BFDB5FC8F" box="[404,513,837,859]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="4" pageNumber="375">Fig. 7A, B</figureCitation>
); Indochina (
<collectingCountry id="F30C7619FFB93B0A14B0FE2BFD68FC8F" box="[645,732,837,859]" name="Thailand" pageId="4" pageNumber="375">Thailand</collectingCountry>
and
<collectingCountry id="F30C7619FFB93B0A169BFE0CFEB6FCAC" box="[174,258,866,888]" name="Vietnam" pageId="4" pageNumber="375">Vietnam</collectingCountry>
) ......................................
<taxonomicName id="4C1B4D0AFFB93B0A17C0FE0DFD36FCA3" authorityName="Yasunaga" authorityYear="2024" box="[501,642,866,888]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="wolskiorum" status="new species">
<emphasis id="B96FEA9BFFB93B0A17C0FE0DFD36FCA3" box="[501,642,866,888]" italics="true" pageId="4" pageNumber="375">C. wolskiorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A14A5FE0DFCB1FCAC" box="[656,773,866,888]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDFEEEFCB1FC66" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFEEEFCB1FC66" pageId="4" pageNumber="375"> Base of antennomere II pale; cuneus entirely pale, sometimes with obscure inner margin ......................................................5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFB93B0A16BDFED2FCB1FBB6" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BDFED2FCB1FBDF" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDFED2FCB1FBDF" pageId="4" pageNumber="375">
5. Male antennomere II dark brown, with pale basal ¼–¼ (
<figureCitation id="13202A0CFFB93B0A14D5FED2FF53FC3A" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="4" pageNumber="375">Fig. 4A, B</figureCitation>
); female metafemur shorter than basal width of pronotum;
<collectingCountry id="F30C7619FFB93B0A169BFE9BFF42FBDF" box="[174,246,1013,1035]" name="Taiwan" pageId="4" pageNumber="375">Taiwan</collectingCountry>
....
<taxonomicName id="4C1B4D0AFFB93B0A1726FE98FDE6FBDF" authority="(Poppius, 1915)" authorityName="Yasunaga" authorityYear="2024" baseAuthorityName="Poppius" baseAuthorityYear="1915" box="[275,594,1013,1035]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="fuhoshoensis" status="new combination">
<emphasis id="B96FEA9BFFB93B0A1726FE98FE19FBDE" box="[275,429,1013,1035]" italics="true" pageId="4" pageNumber="375">C. fuhoshoensis</emphasis>
(
<bibRefCitation id="EF8A4B78FFB93B0A178EFE9BFDFFFBDF" author="Poppius B" box="[443,587,1013,1035]" pageId="4" pageNumber="375" pagination="1 - 80" refId="ref24198" refString="Poppius B (1915) H. Sauter's Formosa-Ausbeute: Nabidae, Anthocoridae, Termatophylidae, Miridae, Isometopidae und Ceratocombidae (Hemiptera). Archiv fur Naturgeschichte, 80 A (8): 1 - 80 (1914)." type="journal article" year="1915">Poppius, 1915</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A146BFE98FCB1FBDF" box="[606,773,1013,1035]" pageId="4" pageNumber="375" rank="species">new combination</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDF97DFCB1FBB6" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDF97DFCB1FBB6" pageId="4" pageNumber="375">
Male antennomere II pale brown, with darkened apical ¼–⅓ (
<figureCitation id="13202A0CFFB93B0A1680F941FEA9FB90" box="[181,285,1071,1093]" captionStart="Fig" captionStartId="9.[113,148,1944,1966]" captionTargetBox="[113,1451,328,1928]" captionTargetId="figure-11@9.[113,1451,309,1928]" captionTargetPageId="9" captionText="Fig. 4. Habitus images (AC, FH) and female genitalia (D, E, I, J) of Chelyolygus fuhoshoensis (AE) and C. parsanus, new species (FJ). A, B, F, G, Male; C, H, female; D, I, ovipositor (gonapophysis I); E, J, genital chamber with sclerotised rings and adjacent structures." figureDoi="http://doi.org/10.5281/zenodo.14682958" httpUri="https://zenodo.org/record/14682958/files/figure.png" pageId="4" pageNumber="375">Fig. 4F, G</figureCitation>
); female metafemur equal in length to basal width of pronotum;
<collectingCountry id="F30C7619FFB93B0A1700F922FEDAFBB6" box="[309,366,1100,1122]" name="Nepal" pageId="4" pageNumber="375">Nepal</collectingCountry>
(lowland) ........
<taxonomicName id="4C1B4D0AFFB93B0A1438F922FD36FBB6" authorityName="Yasunaga" authorityYear="2024" box="[525,642,1100,1122]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="parsanus" status="new species">
<emphasis id="B96FEA9BFFB93B0A1438F922FD36FBB6" box="[525,642,1100,1122]" italics="true" pageId="4" pageNumber="375">C. parsanus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A14A5F922FCB1FBB6" box="[656,773,1100,1122]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30EF2E2CFFB93B0A16BDF907FCB1FA9C" pageId="4" pageNumber="375">
<paragraph id="8BA43689FFB93B0A16BDF907FCB1FB01" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDF907FCB1FB01" pageId="4" pageNumber="375">
6. Dorsum with clear striped pattern; base of antennomere II pale (
<figureCitation id="13202A0CFFB93B0A16DCF9EBFEEDFB4F" box="[233,345,1157,1179]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="4" pageNumber="375">Fig. 6B, F</figureCitation>
); corium and clavus with brown, narrow, continuous stripes along wing veins and fractures (
<figureCitation id="13202A0CFFB93B0A1487F9CCFF0AFB00" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="4" pageNumber="375">Fig. 6A, F</figureCitation>
);
<collectingCountry id="F30C7619FFB93B0A16E7F9D1FE87FB01" box="[210,307,1215,1237]" name="Singapore" pageId="4" pageNumber="375">Singapore</collectingCountry>
..............................
<taxonomicName id="4C1B4D0AFFB93B0A17DEF9D1FD36FB00" authorityName="Yasunaga" authorityYear="2024" box="[491,642,1215,1236]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="virgulatellus" status="new species">
<emphasis id="B96FEA9BFFB93B0A17DEF9D1FD36FB00" box="[491,642,1215,1236]" italics="true" pageId="4" pageNumber="375">C. virgulatellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A14A5F9AEFCB1FB01" box="[656,773,1215,1237]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BA43689FFB93B0A16BDF9B2FCB1FA9C" blockId="4.[136,776,348,1352]" pageId="4" pageNumber="375">
<keyLead id="30EA95BCFFB93B0A16BDF9B2FCB1FA9C" pageId="4" pageNumber="375">
Dorsum widely whitish, without clear striped pattern; base of antennomere II darkened; corium and clavus with greyish brown, interrupted, fragmentary stripes (
<figureCitation id="13202A0CFFB93B0A140EF87BFD3CFAFF" box="[571,648,1301,1323]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="4" pageNumber="375">Fig. 1A</figureCitation>
);
<collectingRegion id="49DFF86BFFB93B0A14A8F87BFD71FAFF" box="[669,709,1301,1323]" country="Indonesia" name="Bali" pageId="4" pageNumber="375">Bali</collectingRegion>
.......... ....................................................
<taxonomicName id="4C1B4D0AFFB93B0A17DAF85DFD36FA93" authorityName="Yasunaga" authorityYear="2024" box="[495,642,1330,1352]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="species" species="tabananicus" status="new species">
<emphasis id="B96FEA9BFFB93B0A17DAF85DFD36FA93" box="[495,642,1330,1352]" italics="true" pageId="4" pageNumber="375">C. tabananicus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFB93B0A14A5F85DFCB1FA9C" box="[656,773,1330,1352]" pageId="4" pageNumber="375" rank="species">new species</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,182 @@
<document id="AEE0A96B8B54157DD1B1EA2210C94512" ID-DOI="10.26107/RBZ-2024-0030" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682946" ID-ZooBank="1F1754CC-6835-4B66-AB5C-FA22AC85D481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171824069" checkinUser="felipe" docAuthor="Yasunaga, Tomohide" docDate="2024" docId="03B2879FFFBC3B0A171CF9D3FD86FF21" docLanguage="en" docName="RafflesBZool.72.371-423.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chelyolygus Yasunaga, 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="372" masterDocId="FF8BFFE7FFBD3B0E1635FD6EFFB4FFD4" masterDocTitle="New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)" masterLastPageNumber="423" masterPageNumber="371" pageNumber="372" updateTime="1737175705380" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B8F3DF7C53C7DB564B68C9530D89ED43" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3AEFF90CB39306C61D200B0B986C3608">
<mods:title id="CBB66987B5261FD98757EE12D515ED0C">New genera and species of the Lygus-complex ranging from the Himalaya-Oriental to the Wallacea region (Insecta: Heteroptera: Miridae: Mirinae)</mods:title>
</mods:titleInfo>
<mods:name id="F7F48D751B0CFFB0C466F64E9518BC6F" type="personal">
<mods:role id="E5A11D68E0EB388015F0C6D03B363549">
<mods:roleTerm id="B20726D81075FFC82B06D431F305CFB0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5588F2FDFA986914B546E1E118DA976F">Yasunaga, Tomohide</mods:namePart>
</mods:name>
<mods:typeOfResource id="9679CC1B0837AAA9C0E6810DA0DB41E9">text</mods:typeOfResource>
<mods:relatedItem id="B3846013A70C3DA436CF39F3718AC5CC" type="host">
<mods:titleInfo id="3572D52C10948B66EB3382D5332F3197">
<mods:title id="C75E8461CA0E8257BE4A29E10AEF6A14">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0B94B2B1C5FBDC5DA01F61B51574AE46">
<mods:date id="22B4912E00AB1988BB9432643A733A73">2024</mods:date>
<mods:detail id="01D741DD4AB331D7A80EAB141F812845" type="pubDate">
<mods:number id="C3C99D93B56FC91E446ED2A895F1B627">2024-10-30</mods:number>
</mods:detail>
<mods:detail id="D89AFD67BEDC4060C93C0000DCB37460" type="volume">
<mods:number id="56125868EB6E5831B33EF4CB747B7292">72</mods:number>
</mods:detail>
<mods:extent id="B2DF9CA7F5BD37EF62D1CFF911966C95" unit="page">
<mods:start id="E56D48D01D1B95EBAFBEAF396C3721DD">371</mods:start>
<mods:end id="36CAF1D665716CC93C6DA1254D2D76DB">423</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="19158E6FCC0022B306A15BF33A46BC17">journal article</mods:classification>
<mods:identifier id="F84218708DDD472D8F2A06E292C97AE2" type="DOI">10.26107/RBZ-2024-0030</mods:identifier>
<mods:identifier id="4B44523A9E3E149AF53046AFC1CE73DE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="22D744C05400B2C58D0DA7C169710FD7" type="Zenodo-Dep">14682946</mods:identifier>
<mods:identifier id="128D58F526E8D008CDD372EB0F085F84" type="ZooBank">1F1754CC-6835-4B66-AB5C-FA22AC85D481</mods:identifier>
</mods:mods>
<treatment id="03B2879FFFBC3B0A171CF9D3FD86FF21" ID-DOI="http://doi.org/10.5281/zenodo.14682944" ID-Zenodo-Dep="14682944" LSID="urn:lsid:plazi:treatment:03B2879FFFBC3B0A171CF9D3FD86FF21" httpUri="http://treatment.plazi.org/id/03B2879FFFBC3B0A171CF9D3FD86FF21" lastPageId="4" lastPageNumber="372" pageId="1" pageNumber="372">
<subSubSection id="C3016502FFBC3B0F171CF9D3FD83FB01" box="[297,567,1213,1237]" pageId="1" pageNumber="372" type="nomenclature">
<paragraph id="8BA43689FFBC3B0F171CF9D3FD83FB01" blockId="1.[297,567,1213,1237]" box="[297,567,1213,1237]" pageId="1" pageNumber="372">
<heading id="D0EC81E5FFBC3B0F171CF9D3FD83FB01" bold="true" box="[297,567,1213,1237]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="372" reason="2">
<emphasis id="B96FEA9BFFBC3B0F171CF9D3FD83FB01" bold="true" box="[297,567,1213,1237]" pageId="1" pageNumber="372">
<taxonomicName id="4C1B4D0AFFBC3B0F171CF9D3FE04FB01" authorityName="Yasunaga" authorityYear="2024" box="[297,432,1213,1237]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFBC3B0F171CF9D3FE04FB01" bold="true" box="[297,432,1213,1237]" italics="true" pageId="1" pageNumber="372">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F17F5F9D3FD83FB01" box="[448,567,1213,1237]" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBC3B0F1644F993FD00FAC1" box="[113,692,1277,1301]" pageId="1" pageNumber="372" type="type_taxon">
<paragraph id="8BA43689FFBC3B0F1644F993FD00FAC1" blockId="1.[113,692,1277,1301]" box="[113,692,1277,1301]" pageId="1" pageNumber="372">
<emphasis id="B96FEA9BFFBC3B0F1644F993FEBDFAC1" bold="true" box="[113,265,1277,1301]" pageId="1" pageNumber="372">
<typeStatus id="54A0882BFFBC3B0F1644F993FF1FFAC1" box="[113,171,1277,1301]" pageId="1" pageNumber="372">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C1B4D0AFFBC3B0F1727F993FDAEFAC1" authorityName="Yasunaga" authorityYear="2024" box="[274,538,1277,1301]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="species" species="wolskiorum" status="sp. nov.">
<emphasis id="B96FEA9BFFBC3B0F1727F993FDAEFAC1" box="[274,538,1277,1301]" italics="true" pageId="1" pageNumber="372">Chelyolygus wolskiorum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F141FF990FD1BFAC1" box="[554,687,1277,1301]" pageId="1" pageNumber="372" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBC3B0F1644F853FD44F8C1" pageId="1" pageNumber="372" type="diagnosis">
<paragraph id="8BA43689FFBC3B0F1644F853FD44F8C1" blockId="1.[113,752,1341,1813]" pageId="1" pageNumber="372">
<emphasis id="B96FEA9BFFBC3B0F1644F853FF53FA81" bold="true" box="[113,231,1341,1365]" pageId="1" pageNumber="372">Diagnosis.</emphasis>
<taxonomicName id="4C1B4D0AFFBC3B0F16C4F853FEC2FA81" authorityName="Yasunaga" authorityYear="2024" box="[241,374,1341,1365]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFBC3B0F16C4F853FEC2FA81" box="[241,374,1341,1365]" italics="true" pageId="1" pageNumber="372">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F17B2F850FE4AFA82" box="[391,510,1342,1366]" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
, can be distinguished from other members of the
<taxonomicName id="4C1B4D0AFFBC3B0F17A0F830FE61FAA1" authorityName="Hahn" authorityYear="1833" box="[405,469,1374,1397]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F17A0F830FE61FAA1" box="[405,469,1374,1397]" italics="true" pageId="1" pageNumber="372">Lygus</emphasis>
</taxonomicName>
-complex by the following combination of characters: Body relatively small (
<quantity id="4CE39B6CFFBC3B0F14ABF813FF32FA61" metricMagnitude="-2" metricUnit="m" metricValue="9.906" metricValueMax="11.938" metricValueMin="7.874" pageId="1" pageNumber="372" unit="in" value="3.9" valueMax="4.7" valueMin="3.1">3.14.7 in</quantity>
total length), ovoid to disk-like, tortoise-shaped; basic colouration variable, pale ivory white to fuscous (
<figureCitation id="13202A0CFFBC3B0F14A9F8D3FD5CFA01" box="[668,744,1469,1493]" captionStart="Fig" captionStartId="2.[135,170,1927,1949]" captionTargetBox="[135,1464,325,1884]" captionTargetId="figure-7@2.[135,1473,297,1912]" captionTargetPageId="2" captionText="Fig. 1. Habitus images of Chelyolygus species. A, B, Chelyolygus tabananicus, new species, holotype female in dorsal (A) and ventral (B) views; C, D, C. cheroti, new species, male; E, F, ditto, female; G, C. wolskiorum, new species, female in ventral view; H, ditto, holotype male in ventral view; I, C. bipuncticollis, male in dorsal view; J, ditto in right lateral view." figureDoi="http://doi.org/10.5281/zenodo.14682948" httpUri="https://zenodo.org/record/14682948/files/figure.png" pageId="1" pageNumber="372">Figs. 1</figureCitation>
,
<figureCitation id="13202A0CFFBC3B0F1644F8B3FF00FA21" box="[113,180,1501,1525]" captionStart="Fig" captionStartId="12.[135,170,1943,1965]" captionTargetBox="[135,1473,281,1907]" captionTargetId="figure-8@12.[135,1473,281,1927]" captionTargetPageId="12" captionText="Fig. 6. Habitus images of Chelyolygus virgulatellus, new species, living (A, B, D) or fresh (CE) and dry-preserved specimens (F, G), and male genitalia (HJ) of holotype male (AC, I, J) and paratype female (F, G). H, Right paramere; I, left paramere; J, vesica. Images (F, G) courtesy of Mr. E. H. Yap." figureDoi="http://doi.org/10.5281/zenodo.14682962" httpUri="https://zenodo.org/record/14682962/files/figure.png" pageId="1" pageNumber="372">6A, F</figureCitation>
,
<figureCitation id="13202A0CFFBC3B0F16F7F8B0FE99FA21" box="[194,301,1501,1526]" captionStart="Fig" captionStartId="14.[135,170,1628,1650]" captionTargetBox="[136,1474,189,1613]" captionTargetId="figure-8@14.[136,1474,189,1613]" captionTargetPageId="14" captionText="Fig. 7. Habitus images and habitats (J, L) of Chelyolygus wolskiorum, new species (AC), C. parsanus, new species (D), Himalisundacoris duwalae, new species (E, F), Sabactiopus schuhi, new species (G, H) and S. sauteri (K). A, E, G, K, Adult male; B, D, F, H, I, adult female; C, final (5th) instar immature form; J, deciduous-broadleaf mixed forest with Urtica dioica, on which Himalisundacoris duwalae, new species, and Sabactiopus schuhi, new species, coexist (Lalitpur, Nepal); L, Styrax formosanus inflorescences, one of the breeding hosts of Sabactiopus sauteri (Nantou, Taiwan)." figureDoi="http://doi.org/10.5281/zenodo.14682964" httpUri="https://zenodo.org/record/14682964/files/figure.png" pageId="1" pageNumber="372">7A, B, D</figureCitation>
); dorsum shining, smooth, with sparsely (usually uniformly on hemelytra) distributed, simple, short setae, lacking noticeable puncture; vertex with flat, continuous basal transverse carina that is about as thick as pronotal collar; pronotum oily and shiny, sometimes nearly glabrous; parameres stout, with developed and often bifurcated apical parts of hypophyses; vesica with two distinct spicules; sclerotised rings oval, small, separated to each other mesally; posterior wall with a row of spinules on anterior interramal sclerite and with small dorsal structure and interramal lobes.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBC3B0F1644FA53FB67FB01" pageId="1" pageNumber="372" type="description">
<paragraph id="8BA43689FFBC3B0F1644FA53FB67FB01" blockId="1.[113,752,1853,2102]" lastBlockId="1.[813,1452,189,1237]" pageId="1" pageNumber="372">
<emphasis id="B96FEA9BFFBC3B0F1644FA53FF48F881" bold="true" box="[113,252,1853,1877]" pageId="1" pageNumber="372">Description.</emphasis>
Body small to moderate in size, ovoid to disk-like, tortoise-shaped, not sexually dimorphic in general shape but pattern in male sometimes darker; basic colouration varying from ivory white to widely fuscous; dorsal surface shining, with uniformly distributed, pale, simple, semi-erect setae but partly glabrous in some species.
<emphasis id="B96FEA9BFFBC3B0F1472FAB3FD38F821" bold="true" box="[583,652,2013,2037]" pageId="1" pageNumber="372">Head:</emphasis>
Smooth, shining, rather vertical in lateral view, almost glabrous; eyes contiguous to anterior margin of pronotal collar; vertex weakly sulcate mesally, with flat, continuous basal transverse carina that is about as thick as pronotal collar.
<emphasis id="B96FEA9BFFBC3B0F1375FDB3FA1FFF21" bold="true" box="[1344,1451,221,245]" pageId="1" pageNumber="372">Antenna:</emphasis>
Shorter than body, almost linear; antennomeres I and II always with dark apices (or apical parts), about as thick as protibia; antennomere II in female shorter than basal width of pronotum; antennomeres III and IV filiform.
<emphasis id="B96FEA9BFFBC3B0F1372FC33FA18FEA1" bold="true" box="[1351,1452,349,373]" pageId="1" pageNumber="372">Labium:</emphasis>
Relatively long and slender, reaching or slightly exceeding apex of metacoxa.
<emphasis id="B96FEA9BFFBC3B0F15DAFCF3FBF8FE61" bold="true" box="[1007,1100,413,437]" pageId="1" pageNumber="372">Thorax:</emphasis>
Pronotum polished, inflated, with sparsely distributed, short, reclining setae, lacking noticeable punctures, in some species glabrous; calli with a pair of dark spots, flat, not clearly demarcated; collar narrow; propleuron not margined nor carinate; metathoracic scent efferent system subtriangular, ovoid dorsally, with narrow, more or less produced peritreme; scutellum polished, flat.
<emphasis id="B96FEA9BFFBC3B0F132FFF33FA18FDA1" bold="true" box="[1306,1452,605,629]" pageId="1" pageNumber="372">Hemelytron:</emphasis>
Shining, variable in colour but as a rule with striped pattern on clavus and corium, almost impunctate, declivous at cuneal fracture, usually with uniformly distributed, pale, simple, reclining setae; lateral margin of exocorium (embolium) rounded.
<emphasis id="B96FEA9BFFBC3B0F15AFFF93FC6FFCC1" bold="true" box="[922,987,765,789]" pageId="1" pageNumber="372">Legs:</emphasis>
Moderate in length; meta-tarsomere III longer than I or II; pretarsal structures as in
<figureCitation id="13202A0CFFBC3B0F1335FE73FADFFCE1" box="[1280,1387,797,821]" captionStart="Fig" captionStartId="30.[136,171,1945,1967]" captionTargetBox="[136,1476,250,1930]" captionTargetId="figure-8@30.[136,1476,250,1930]" captionTargetPageId="30" captionText="Fig. 19. Scanning electron micrographs of Chelyolygus cheroti, new species. A, Head and anterior pronotum, dorsal view; B, ditto, left lateral view; C, scutellum and anterior hemelytron, left lateral view; D, metathoracic scent efferent system; E, scutellum, clavus and corium, dorsal view; F, metatarsi of each sex; G, pretarsal structure of metaleg; H, apex of pygophore, ventral view; I, left paramere; J, right paramere; K, vesica; L, apex of ovipositor (gonapophysis I); M, genital chamber; N, O, posterior wall." figureDoi="http://doi.org/10.5281/zenodo.14682996" httpUri="https://zenodo.org/record/14682996/files/figure.png" pageId="1" pageNumber="372">Figs. 19G</figureCitation>
,
<figureCitation id="13202A0CFFBC3B0F1342FE73FCD2FC81" captionStart="Fig" captionStartId="31.[113,148,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@31.[113,1451,251,1929]" captionTargetPageId="31" captionText="Fig. 20. Scanning electron micrographs of Chelyolygus bipuncticollis (male) and C. tabananicus, new species (holotype female). A, G, Head and anterior thorax, left lateral view; B, head, frontal view; C, ditto, dorsal view; D, E, K, pretarsal structure; F, metatarsus; H, head and pronotum, frontal-left lateral view; I, metathoracic scent efferent system; J, mesotarsus; L, genital chamber, dorsal view; M, N, posterior wall; O, ditto, dorsal structure." figureDoi="http://doi.org/10.5281/zenodo.14682998" httpUri="https://zenodo.org/record/14682998/files/figure.png" pageId="1" pageNumber="372">20D, E, K</figureCitation>
; parempodia usually short, with thick bases.
<emphasis id="B96FEA9BFFBC3B0F1345FE53FC2CFCA1" bold="true" pageId="1" pageNumber="372">Male genitalia:</emphasis>
Pygophore relatively short. Parameres generally stout, sometimes with developed and bifurcate apical parts of hypophyses; left paramere C- or L-shaped, with elongate apical process of hypophysis; hypophysis of right paramere often bifurcate apically or sometimes terminated in a sharp point. Vesica with two distinct spicules, with one or two additional lobal-sclerites in some species; gonopore small, circular; apex of phallotheca simple, narrow.
<emphasis id="B96FEA9BFFBC3B0F1366F953FC3BFBA1" bold="true" pageId="1" pageNumber="372">Female genitalia</emphasis>
: Genital chamber with oval, small sclerotised rings separated to each other mesally; posterior wall with a row of spinules on anterior interramal sclerite; dorsal structure and interramal lobes relatively narrow.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBC3B0F1518F993FC45FAA1" pageId="1" pageNumber="372" type="etymology">
<paragraph id="8BA43689FFBC3B0F1518F993FC45FAA1" blockId="1.[813,1452,1277,1397]" pageId="1" pageNumber="372">
<emphasis id="B96FEA9BFFBC3B0F1518F993FC1BFAC1" bold="true" box="[813,943,1277,1301]" pageId="1" pageNumber="372">Etymology.</emphasis>
From Greek noun, chelys (= turtle), combined with mirine genus
<taxonomicName id="4C1B4D0AFFBC3B0F15C9F870FB36FAE1" authority="Hahn" authorityName="Hahn" authorityYear="1833" box="[1020,1154,1309,1333]" class="Insecta" family="Miridae" genus="Lygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F15C9F870FB88FAE1" box="[1020,1084,1310,1333]" italics="true" pageId="1" pageNumber="372">Lygus</emphasis>
Hahn
</taxonomicName>
, referring to the ovoid and stout body shape that is reminiscent of a tortoise-shell; gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C3016502FFBC3B0A1518F8F3FD86FF21" lastPageId="4" lastPageNumber="375" pageId="1" pageNumber="372" type="discussion">
<paragraph id="8BA43689FFBC3B0A1518F8F3FD86FF21" blockId="1.[813,1452,1437,2101]" lastBlockId="4.[136,775,189,245]" lastPageId="4" lastPageNumber="375" pageId="1" pageNumber="372">
<emphasis id="B96FEA9BFFBC3B0F1518F8F3FC18FA61" bold="true" box="[813,940,1437,1461]" pageId="1" pageNumber="372">Discussion.</emphasis>
<taxonomicName id="4C1B4D0AFFBC3B0F1580F8F3FB8DFA61" authorityName="Yasunaga" authorityYear="2024" box="[949,1081,1437,1461]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFBC3B0F1580F8F3FB8DFA61" box="[949,1081,1437,1461]" italics="true" pageId="1" pageNumber="372">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F127CF8F0FB74FA62" box="[1097,1216,1438,1462]" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
, is herein established to accommodate five representatives which have distinct characters diagnosed above. In addition to the ovoid or tortoise-shaped body shape, it is defined principally by the unique male and female genitalic structures, such as two spicules (and sometimes one or two additional lobal-sclerites) on the vesica and a row of spinules on anterior part of interramal sclerite and narrow interramal lobe and dorsal structure on the posterior wall. Currently, any plausible sister-group is not posited for
<taxonomicName id="4C1B4D0AFFBC3B0F12DFFBD3FADAF901" authorityName="Yasunaga" authorityYear="2024" box="[1258,1390,1725,1749]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFBC3B0F12DFFBD3FADAF901" box="[1258,1390,1725,1749]" italics="true" pageId="1" pageNumber="372">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F134AFBD0FCC5F922" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
. The ovoid or disk-like body form is reminiscent of
<taxonomicName id="4C1B4D0AFFBC3B0F157BFB93FACEF8C1" authority="Yasunaga, Schwartz &amp; Cherot, 2002" authorityName="Yasunaga, Schwartz &amp; Cherot" authorityYear="2002" box="[846,1402,1789,1814]" class="Insecta" family="Miridae" genus="Apolygopsis" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F157BFB93FC67F8C1" box="[846,979,1789,1813]" italics="true" pageId="1" pageNumber="372">Apolygopsis</emphasis>
<bibRefCitation id="EF8A4B78FFBC3B0F15E8FB90FACEF8C1" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" box="[989,1402,1789,1814]" pageId="1" pageNumber="372" pagination="1 - 26" refId="ref24768" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2002) New genera, species, synonymies, and combinations in the &quot; Lygus - complex &quot; from Japan, with discussion of Peltidolygus Poppius and Warrisia Carvalho (Heteroptera: Miridae: Mirinae). American Museum Novitates, 3378: 1 - 26." type="journal article" year="2002">Yasunaga, Schwartz &amp; Chérot, 2002</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1B4D0AFFBC3B0F1518FA73FBD3F8E1" authority="Horvath, 1889" authorityName="Horvath" authorityYear="1889" box="[813,1127,1821,1845]" class="Insecta" family="Miridae" genus="Chilocrates" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F1518FA73FC06F8E1" box="[813,946,1821,1845]" italics="true" pageId="1" pageNumber="372">Chilocrates</emphasis>
<bibRefCitation id="EF8A4B78FFBC3B0F1588FA73FBD3F8E1" author="Horvath G" box="[957,1127,1821,1845]" pageId="1" pageNumber="372" pagination="29 - 40" refId="ref23872" refString="Horvath G (1889) Analecta ad Cognitionem Heteropterotum Himalayensium conscripsit. Termeszetrajzi Fuzetek, 11: 29 - 40." type="journal article" year="1889">Horváth, 1889</bibRefCitation>
</taxonomicName>
, but the colour pattern and genitalic structure of
<taxonomicName id="4C1B4D0AFFBC3B0F123BFA53FB26F881" authorityName="Yasunaga" authorityYear="2024" box="[1038,1170,1853,1877]" class="Insecta" family="Miridae" genus="Chelyolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B96FEA9BFFBC3B0F123BFA53FB26F881" box="[1038,1170,1853,1877]" italics="true" pageId="1" pageNumber="372">Chelyolygus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25C57E0FFBC3B0F12ABFA50FAA6F882" box="[1182,1298,1854,1878]" pageId="1" pageNumber="372" rank="genus">new genus</taxonomicNameLabel>
, are obviously distinct from those of the latter two genera (
<bibRefCitation id="EF8A4B78FFBC3B0F1314FA33FC6EF841" author="Yasunaga T &amp; Schwartz MD" pageId="1" pageNumber="372" pagination="152 - 162" refId="ref24585" refString="Yasunaga T &amp; Schwartz MD (2000) Restored status of the genus Chilocrates Horvath (Heteroptera: Miridae), with a review of the species and nomenclatural changes. Journal of the New York Entomological Society, 108: 152 - 162." type="journal article" year="2000">Yasunaga &amp; Schwartz, 2000</bibRefCitation>
;
<bibRefCitation id="EF8A4B78FFBC3B0F15DDFA10FB6AF841" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" box="[1000,1246,1917,1942]" pageId="1" pageNumber="372" pagination="1 - 26" refId="ref24768" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2002) New genera, species, synonymies, and combinations in the &quot; Lygus - complex &quot; from Japan, with discussion of Peltidolygus Poppius and Warrisia Carvalho (Heteroptera: Miridae: Mirinae). American Museum Novitates, 3378: 1 - 26." type="journal article" year="2002">Yasunaga et al., 2002</bibRefCitation>
; Yasunaga, 2023). Based on similarity in the striped pattern on the hemelytra and vesical sclerites,
<taxonomicName id="4C1B4D0AFFBC3B0F122FFAD3FB53F801" authority="Carvalho" authorityName="Carvalho" authorityYear="1987" box="[1050,1255,1981,2005]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F122FFAD3FBCFF801" box="[1050,1147,1981,2005]" italics="true" pageId="1" pageNumber="372">Prolygus</emphasis>
Carvalho
</taxonomicName>
is assumed to be one of the closer taxa. However,
<taxonomicName id="4C1B4D0AFFBC3B0F129AFAB3FAA6F821" authorityName="Carvalho" authorityYear="1987" box="[1199,1298,2013,2037]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="372" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFBC3B0F129AFAB3FAA6F821" box="[1199,1298,2013,2037]" italics="true" pageId="1" pageNumber="372">Prolygus</emphasis>
</taxonomicName>
is distinct in having the elongate, conventional mirine shape (
<figureCitation id="13202A0CFFBC3B0F1376FA93FA10F7C1" box="[1347,1444,2045,2069]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="1" pageNumber="372">Fig. 32L</figureCitation>
) and female posterior wall with developed, wide interramal lobes and lateral lobes (
<figureCitation id="13202A0CFFB93B0A17B8FDD3FDA7FF01" box="[397,531,189,213]" captionStart="Fig" captionStartId="47.[113,149,1944,1966]" captionTargetBox="[113,1451,251,1929]" captionTargetId="figure-11@47.[113,1451,251,1929]" captionTargetPageId="47" captionText="Fig. 32. Scanning electron micrographs of Paralygocoris alboscutellatus (AF), Prolygus disciger (GK), P. papuanus (LO) and Apolygopsis furvocarinatus (E). A, B, G, Anterior body, left lateral view; C, H, thoracic pleura, left lateral view; D, N, genital chamber; E, F, K, O; posterior wall; I, Left paramere; J, M, vesica; L, dorsal habitus. Abbreviations: IRL = interramal lobe, IRS = interramal sclerite, LLB = lateral lobe" figureDoi="http://doi.org/10.5281/zenodo.14683030" httpUri="https://zenodo.org/record/14683030/files/figure.png" pageId="4" pageNumber="375">Fig. 32K, O</figureCitation>
) (see
<bibRefCitation id="EF8A4B78FFB93B0A146DFDD0FF74FF21" author="Yasunaga T &amp; Schwartz MD &amp; Cherot F" pageId="4" pageNumber="375" pagination="357 - 387" refId="ref24824" refString="Yasunaga T, Schwartz MD &amp; Cherot F (2018) Review of the plant bug genus Prolygus and related mirine taxa from eastern Asia (Hemiptera: Heteroptera: Miridae). Acta Entomologica Musei Nationalis Pragae, 58: 357 - 387." type="journal article" year="2018">Yasunaga et al., 2018</bibRefCitation>
, for typical features of
<taxonomicName id="4C1B4D0AFFB93B0A17F7FDB3FD96FF21" authorityName="Carvalho" authorityYear="1987" box="[450,546,221,245]" class="Insecta" family="Miridae" genus="Prolygus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="375" phylum="Arthropoda" rank="genus">
<emphasis id="B96FEA9BFFB93B0A17F7FDB3FD96FF21" box="[450,546,221,245]" italics="true" pageId="4" pageNumber="375">Prolygus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,918 @@
<document id="F05995BC254F56EC4C9F35CE3E33B1A2" ID-DOI="10.26107/RBZ-2024-0019" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683134" ID-ZooBank="1EEDBDEA-E973-4A2F-A1F7-07DDD5063386" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172000403" checkinUser="felipe" docAuthor="Gonggoli, Ade Damara, , Misbahul Munir, , Fajar Kaprawi, , Tom Kirschey, Hamidy, &amp; Amir" docDate="2024" docId="03C987A0FFA5FF85C694FBBEC495FC47" docLanguage="en" docName="RafflesBZool.72.219-234.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Zhangixalus faritsalhadii Gonggoli, , , , Hamidy &amp; Amir, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="229" masterDocId="FFF0FFD8FFA0FF8FC64DFFB8C715FFC9" masterDocTitle="A new species of tree frog (Amphibia, Anura, Rhacophoridae) from Central Java, Indonesia" masterLastPageNumber="234" masterPageNumber="219" pageNumber="224" updateTime="1737175833791" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="0692587F323D7B55E9914A91601E5790" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="077F61D135F4456245319FB27B8E1B4F">
<mods:title id="3B534648F939EFF82EEC2C62B8D8B56C">A new species of tree frog (Amphibia, Anura, Rhacophoridae) from Central Java, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="F670672C25D8CE0A92B3D7C954E63757" type="personal">
<mods:role id="64728F79BF0719D93BB6B003926D68CE">
<mods:roleTerm id="FC6D35D04F0B920C672FE019F307A164">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5CFA801654B6442A04764BDC1268A4C7">Gonggoli, Ade Damara</mods:namePart>
</mods:name>
<mods:name id="D5C33BBC749784533ABAF2D2CB336CC9" type="personal">
<mods:role id="13B99BFFD2F77A3DB0157D86021A7850">
<mods:roleTerm id="3C1FB590C503DBD418DCA2DF914D2AA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="420B5D7FB8AF6607C95F6573536F02C8">, Misbahul Munir</mods:namePart>
</mods:name>
<mods:name id="6918474DCC3664975F7382D93A8BF6F0" type="personal">
<mods:role id="B75FFC840F5921F31C82D176B4C70BFF">
<mods:roleTerm id="8FF18E3ED19D68853400B35E5DC455CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CE72AF75B24356EB2574372820F3E8BB">, Fajar Kaprawi</mods:namePart>
</mods:name>
<mods:name id="83863D045FB5A8FD3477F9D67E4FE98D" type="personal">
<mods:role id="E13616B515993FB2345D5F3F983BD4C7">
<mods:roleTerm id="FD7792C8F1DB22292872408C0E7A4AF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D079C3C32A446A7BACE5C2BB6F7E2B85">, Tom Kirschey</mods:namePart>
</mods:name>
<mods:name id="3B7D3E8B057F9117B7E6A137368C8C5F" type="personal">
<mods:role id="F4340CFE3BA251C60581DAFEE2612803">
<mods:roleTerm id="E800FFA20B99F59C6C425AE0AD663110">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09969F13E40A257E6BAFE5CAD5981596">Hamidy,</mods:namePart>
</mods:name>
<mods:name id="82D29450F78688F2810D54352FB1A155" type="personal">
<mods:role id="F3BF7D0975305D633227DE1EF63D19F6">
<mods:roleTerm id="BF27C7886827E22048E80B3262A164D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="64305FF17150EDAC6E03AAFAA34001DD">Amir</mods:namePart>
</mods:name>
<mods:typeOfResource id="8BC5AECEA86041272C14133F2F4DABC0">text</mods:typeOfResource>
<mods:relatedItem id="BFF2A741F23436F867D6C5024DE8B935" type="host">
<mods:titleInfo id="BA5A694203E339ED856582A0E39DF03B">
<mods:title id="D66C4AC91B0E0A7EC5B7D4113E6857B7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7B13F0C92D5A37F1E2A7F05AA5C7B885">
<mods:date id="3EEF7F55F2245B7F9349059FE98AEB21">2024</mods:date>
<mods:detail id="2161CE61B2D493C36D78E36D0764C35E" type="pubDate">
<mods:number id="71F7FEED3F5277038CAA1D1D446D67E2">2024-08-08</mods:number>
</mods:detail>
<mods:detail id="E6F6C292BD98333B819BD480C518274F" type="volume">
<mods:number id="F32025BB4C5CDAD2515D7F24C3556953">72</mods:number>
</mods:detail>
<mods:extent id="A1EEB3069A41ED7BB5BC22E4FFCE2E0A" unit="page">
<mods:start id="20CF5495A41B04B3642DB895F092AA6F">219</mods:start>
<mods:end id="A3376E48C62E2BB2DBA29E057C87F4BF">234</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7E8A4A2D2D75D2F71CF74298C87B6011">journal article</mods:classification>
<mods:identifier id="322316A17F4F7F9FBCDE843AB7BD1A80" type="DOI">10.26107/RBZ-2024-0019</mods:identifier>
<mods:identifier id="8F0E63E23F324F3BF174847B4779DDB5" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="5A34451D133828DD98EB17F9D95D05A2" type="Zenodo-Dep">14683134</mods:identifier>
<mods:identifier id="930FF732899FFC7AB9F0547ED72D2DBF" type="ZooBank">1EEDBDEA-E973-4A2F-A1F7-07DDD5063386</mods:identifier>
</mods:mods>
<treatment id="03C987A0FFA5FF85C694FBBEC495FC47" LSID="urn:lsid:plazi:treatment:03C987A0FFA5FF85C694FBBEC495FC47" httpUri="http://treatment.plazi.org/id/03C987A0FFA5FF85C694FBBEC495FC47" lastPageId="10" lastPageNumber="229" pageId="5" pageNumber="224">
<subSubSection id="C37A653DFFA5FF8AC694FBBEC59CFBD6" box="[217,649,1030,1055]" pageId="5" pageNumber="224" type="nomenclature">
<paragraph id="8BDF36B6FFA5FF8AC694FBBEC59CFBD6" blockId="5.[217,649,1030,1087]" box="[217,649,1030,1055]" pageId="5" pageNumber="224">
<heading id="D09781DAFFA5FF8AC694FBBEC59CFBD6" bold="true" box="[217,649,1030,1055]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="224" reason="2">
<emphasis id="B914EAA4FFA5FF8AC694FBBEC59CFBD6" bold="true" box="[217,649,1030,1055]" pageId="5" pageNumber="224">
<taxonomicName id="4C604D35FFA5FF8AC694FBBEC6E1FBD7" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[217,500,1030,1054]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="224" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFA5FF8AC694FBBEC6E1FBD7" bold="true" box="[217,500,1030,1054]" italics="true" pageId="5" pageNumber="224">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFA5FF8AC44EFBBFC59CFBD6" box="[515,649,1031,1055]" pageId="5" pageNumber="224" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37A653DFFA5FF8AC739FB9FC6F1FBB2" pageId="5" pageNumber="224" type="description">
<paragraph id="8BDF36B6FFA5FF8AC739FB9FC6FBFBF6" blockId="5.[217,649,1030,1087]" box="[372,494,1063,1087]" pageId="5" pageNumber="224">
(
<figureCitation id="135B2A33FFA5FF8AC731FB9FC6DCFBF6" box="[380,457,1063,1087]" captionStart="Fig" captionStartId="6.[136,171,1098,1120]" captionTargetBox="[136,1474,189,1083]" captionTargetId="figure-8@6.[133,1477,184,1086]" captionTargetPageId="6" captionText="Fig. 4. A, Zhangixalus faritsalhadii, new species, male holotype in life, MZB Amph 32885; B, female paratype, MZB Amph 32887; C, MZB Amph 32889, female paratype in ventral view; D, MZB Amph 32886 and MZB Amph 32889 in amplexus. Photographs by Misbahul Munir (AC) and Fajar Kaprawi (D)." pageId="5" pageNumber="224">Figs. 4</figureCitation>
,
<figureCitation id="135B2A33FFA5FF8AC79AFB9FC6F0FBF6" box="[471,485,1063,1087]" captionStart="Fig" captionStartId="7.[113,148,1967,1989]" captionTargetBox="[119,1451,288,1929]" captionTargetId="figure-12@7.[103,1472,263,1945]" captionTargetPageId="7" captionText="Fig. 5. Zhangixalus faritsalhadii, new species, from Mount Slamet, Java, Indonesia. AF, an adult male holotype, MZB Amph 32885; GL, an adult female paratype, MZB Amph 32887. Scale bar = 10 mm. Photographs by Misbahul Munir." pageId="5" pageNumber="224">5</figureCitation>
)
</paragraph>
<paragraph id="8BDF36B6FFA5FF8AC63CFBDEC6F1FBB2" blockId="5.[113,484,1126,1148]" box="[113,484,1126,1148]" pageId="5" pageNumber="224">
<taxonomicName id="4C604D35FFA5FF8AC63CFBDEC60DFBB2" box="[113,280,1126,1147]" class="Amphibia" family="Rhacophoridae" genus="Rhacophorus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="224" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B914EAA4FFA5FF8AC63CFBDEC7E4FBB2" box="[113,241,1126,1147]" italics="true" pageId="5" pageNumber="224">Rhacophorus</emphasis>
sp.
</taxonomicName>
:
<bibRefCitation id="EFF14B47FFA5FF8AC76EFBDEC6F1FBB2" author="Alhadi F &amp; Kaprawi F &amp; Hamidy A &amp; Kirschey T" box="[291,484,1126,1148]" pageId="5" pageNumber="224" refId="ref8869" refString="Alhadi F, Kaprawi F, Hamidy A &amp; Kirschey T (2021) Amfibi Pulau Jawa-Panduan Bergambar dan Identifikasi. Perkumpulan Amfibi Reptil Sumatra &amp; NABU, 213 pp." type="book" year="2021">Alhadi et al. (2021)</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A653DFFA5FF8AC63CFB1CC662F995" pageId="5" pageNumber="224" type="materials_examined">
<paragraph id="8BDF36B6FFA5FF8AC63CFB1CC60AFA95" blockId="5.[113,753,1188,1372]" pageId="5" pageNumber="224">
<materialsCitation id="3B083CEBFFA5FF8AC63CFB1CC60AFA95" collectingDate="2022-07-04" collectionCode="MZB" collectorName="Fajar Kaprawi &amp; Eki Abdul Kholik &amp; Tarwo" country="Indonesia" county="Banyumas Regency" elevation="773" latitude="-7.318147" location="Amph" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" pageId="5" pageNumber="224" specimenCount="1" stateProvince="Jawa Tengah" typeStatus="holotype">
<emphasis id="B914EAA4FFA5FF8AC63CFB1CC7F5FB75" bold="true" box="[113,224,1188,1212]" pageId="5" pageNumber="224">
<typeStatus id="54DB8814FFA5FF8AC63CFB1CC7CEFB75" box="[113,219,1188,1212]" pageId="5" pageNumber="224" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED71AE73FFA5FF8AC6ABFB1CC635FB72" box="[230,288,1188,1211]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="224" type="Museum">MZB</collectionCode>
<location id="8EBF606DFFA5FF8AC76AFB1CC67FFB75" LSID="urn:lsid:plazi:treatment:03C987A0FFA5FF85C694FBBEC495FC47:8EBF606DFFA5FF8AC76AFB1CC67FFB75" box="[295,362,1188,1212]" country="Indonesia" county="Banyumas Regency" latitude="-7.318147" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" name="Amph" pageId="5" pageNumber="224" stateProvince="Jawa Tengah">Amph</location>
32885, an adult male collected from
<location id="8EBF606DFFA5FF8AC63CFB7CC7FFFB15" LSID="urn:lsid:plazi:treatment:03C987A0FFA5FF85C694FBBEC495FC47:8EBF606DFFA5FF8AC63CFB7CC7FFFB15" box="[113,234,1220,1244]" country="Indonesia" county="Banyumas Regency" latitude="-7.318147" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" name="Mt. Slamet" pageId="5" pageNumber="224" stateProvince="Jawa Tengah">Mt. Slamet</location>
,
<location id="8EBF606DFFA5FF8AC6B9FB7CC64DFB15" LSID="urn:lsid:plazi:treatment:03C987A0FFA5FF85C694FBBEC495FC47:8EBF606DFFA5FF8AC6B9FB7CC64DFB15" box="[244,344,1220,1244]" country="Indonesia" county="Banyumas Regency" latitude="-7.318147" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" name="Kalipagu" pageId="5" pageNumber="224" stateProvince="Jawa Tengah">Kalipagu</location>
,
<location id="8EBF606DFFA5FF8AC72EFB7CC50EFB15" LSID="urn:lsid:plazi:treatment:03C987A0FFA5FF85C694FBBEC495FC47:8EBF606DFFA5FF8AC72EFB7CC50EFB15" box="[355,539,1220,1244]" country="Indonesia" county="Banyumas Regency" latitude="-7.318147" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" name="Ketenger Village" pageId="5" pageNumber="224" stateProvince="Jawa Tengah">Ketenger Village</location>
,
<collectingMunicipality id="6BBBACCCFFA5FF8AC46BFB7CC5F9FB15" box="[550,748,1220,1244]" pageId="5" pageNumber="224">Baturaden District</collectingMunicipality>
,
<collectingCounty id="62BE4E3AFFA5FF8AC63CFB5CC64FFB35" box="[113,346,1252,1276]" pageId="5" pageNumber="224">Banyumas Regency</collectingCounty>
,
<collectingRegion id="49A4F854FFA5FF8AC727FB5CC564FB35" box="[362,625,1252,1276]" country="Indonesia" name="Jawa Tengah" pageId="5" pageNumber="224">Central Java Province</collectingRegion>
,
<collectingCountry id="F3777626FFA5FF8AC4CCFB5CC5FAFB35" box="[641,751,1252,1276]" name="Indonesia" pageId="5" pageNumber="224">Indonesia</collectingCountry>
(
<geoCoordinate id="EE545071FFA5FF8AC637FABAC603FAD2" box="[122,278,1282,1307]" degrees="7" direction="south" minutes="19" orientation="latitude" pageId="5" pageNumber="224" precision="1" seconds="05.33" value="-7.318147">7°1905.33″S</geoCoordinate>
,
<geoCoordinate id="EE545071FFA5FF8AC76BFABAC6CAFAD2" box="[294,479,1282,1307]" degrees="109" direction="east" minutes="12" orientation="longitude" pageId="5" pageNumber="224" precision="1" seconds="17.46" value="109.20485">109°1217.46″E</geoCoordinate>
;
<elevation id="004DD185FFA5FF8AC7A2FABCC5B8FAD2" box="[495,685,1284,1308]" metricMagnitude="2" metricUnit="m" metricValue="7.7299999999999995" pageId="5" pageNumber="224" unit="m" value="773.0">
elevation
<quantity id="4C989B53FFA5FF8AC42EFABCC5B8FAD2" box="[611,685,1284,1308]" metricMagnitude="2" metricUnit="m" metricValue="7.7299999999999995" pageId="5" pageNumber="224" unit="m" value="773.0">773 m</quantity>
</elevation>
a.s.l) collected by
<collectorName id="26955360FFA5FF8AC6B2FA9CC688FAF5" box="[255,413,1316,1340]" pageId="5" pageNumber="224">Fajar Kaprawi</collectorName>
,
<collectorName id="26955360FFA5FF8AC7E4FA9CC565FAF5" box="[425,624,1316,1340]" pageId="5" pageNumber="224">Eki Abdul Kholik</collectorName>
, and
<collectorName id="26955360FFA5FF8AC4E6FA9CC5E4FAF2" box="[683,753,1316,1339]" pageId="5" pageNumber="224">Tarwo</collectorName>
on
<date id="FFDE1076FFA5FF8AC6DAFAFCC60EFA95" box="[151,283,1348,1372]" pageId="5" pageNumber="224" value="2022-07-04">
<collectingDate id="EF9AE99EFFA5FF8AC6DAFAFCC60EFA95" box="[151,283,1348,1372]" pageId="5" pageNumber="224" value="2022-07-04">4 July 2022</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDF36B6FFA5FF8AC63CFA3CC662F995" blockId="5.[113,754,1412,1628]" pageId="5" pageNumber="224">
<materialsCitation id="3B083CEBFFA5FF8AC63CFA3CC6A3FA35" collectingDate="2022-07-04" collectionCode="MZB" collectorName="Fajar Kaprawi &amp; Eki Abdul Kholik &amp; Tarwo" country="Indonesia" county="Banyumas Regency" elevation="773" latitude="-7.318147" location="Amph" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" pageId="5" pageNumber="224" specimenCount="9" specimenCount-adult="4" stateProvince="Jawa Tengah" typeStatus="paratype">
<emphasis id="B914EAA4FFA5FF8AC63CFA3CC7E7FA55" bold="true" box="[113,242,1412,1436]" pageId="5" pageNumber="224">
<typeStatus id="54DB8814FFA5FF8AC63CFA3CC7F8FA55" box="[113,237,1412,1436]" pageId="5" pageNumber="224" type="paratype">Paratypes</typeStatus>
.
</emphasis>
A total of
<specimenCount id="9D66FD3FFFA5FF8AC732FA3CC527FA55" box="[383,562,1412,1436]" count="5" pageId="5" pageNumber="224" type="generic">five specimens</specimenCount>
:
<specimenCount id="9D66FD3FFFA5FF8AC40CFA3CC5BBFA55" box="[577,686,1412,1436]" count="1" pageId="5" pageNumber="224" type="adult">one adult</specimenCount>
male (
<collectionCode id="ED71AE73FFA5FF8AC634FA1CC7A1FA75" box="[121,180,1444,1468]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="224" type="Museum">MZB</collectionCode>
Amph 32886) and
<specimenCount id="9D66FD3FFFA5FF8AC7C6FA1CC6EBFA75" box="[395,510,1444,1468]" count="3" pageId="5" pageNumber="224" type="adult">three adult</specimenCount>
females (
<collectionCode id="ED71AE73FFA5FF8AC427FA1CC5B0FA75" box="[618,677,1444,1468]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="224" type="Museum">MZB</collectionCode>
Amph 3288789) collected by the same collectors and from the same locality as the holotype
</materialsCitation>
;
<materialsCitation id="3B083CEBFFA5FF8AC789FA5CC666F995" collectingDate="2020-02-10" collectionCode="MZB" collectorName="Farits Alhadi &amp; Fajar Kaprawi &amp; Jarian Permana &amp; Karso" country="Indonesia" county="Banyumas Regency" elevation="773" latitude="-7.318147" location="Amph" longLatPrecision="1" longitude="109.20485" municipality="Baturaden District" pageId="5" pageNumber="224" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Tengah" typeStatus="paratype">
and
<specimenCount id="9D66FD3FFFA5FF8AC7B9FA5CC54DFA35" box="[500,600,1508,1532]" count="1" pageId="5" pageNumber="224" type="male">one male</specimenCount>
(
<collectionCode id="ED71AE73FFA5FF8AC424FA5CC5B1FA35" box="[617,676,1508,1532]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="5" pageNumber="224" type="Museum">MZB</collectionCode>
Amph 32890) collected by
<collectorName id="26955360FFA5FF8AC716F9BCC6E5F9D5" box="[347,496,1540,1564]" pageId="5" pageNumber="224">Farits Alhadi</collectorName>
,
<collectorName id="26955360FFA5FF8AC7B3F9BCC5B4F9D5" box="[510,673,1540,1564]" pageId="5" pageNumber="224">Fajar Kaprawi</collectorName>
,
<collectorName id="26955360FFA5FF8AC4E2F9BCC7C7F9F2" pageId="5" pageNumber="224">Jarian Permana</collectorName>
, and
<collectorName id="26955360FFA5FF8AC75CF99CC644F9F2" box="[273,337,1572,1595]" pageId="5" pageNumber="224">Karso</collectorName>
on
<date id="FFDE1076FFA5FF8AC7CDF99CC553F9F5" box="[384,582,1572,1596]" pageId="5" pageNumber="224" value="2020-02-10">
<collectingDate id="EF9AE99EFFA5FF8AC7CDF99CC553F9F5" box="[384,582,1572,1596]" pageId="5" pageNumber="224" value="2020-02-10">10 February 2020</collectingDate>
</date>
from the same locality as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37A653DFFA5FF8AC63EF93CC7D4F815" pageId="5" pageNumber="224" type="etymology">
<paragraph id="8BDF36B6FFA5FF8AC63EF93CC7DEF8F2" blockId="5.[112,753,1668,1851]" pageId="5" pageNumber="224">
<emphasis id="B914EAA4FFA5FF8AC63EF93CC7E6F955" bold="true" box="[115,243,1668,1692]" pageId="5" pageNumber="224">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C604D35FFA5FF8AC789F93CC544F955" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[452,593,1668,1692]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="224" phylum="Chordata" rank="species" species="faritsalhadii">
<emphasis id="B914EAA4FFA5FF8AC79DF93CC544F955" box="[464,593,1668,1692]" italics="true" pageId="5" pageNumber="224">faritsalhadii</emphasis>
</taxonomicName>
” is an eponym, dedicated to the deceased Farits Alhadi, a herpetologist in
<collectingCountry id="F3777626FFA5FF8AC63CF97CC7CCF915" box="[113,217,1732,1756]" name="Indonesia" pageId="5" pageNumber="224">Indonesia</collectingCountry>
who collected the first known specimen of this species that is now part of the
<typeStatus id="54DB8814FFA5FF8AC780F95CC6EEF935" box="[461,507,1764,1788]" pageId="5" pageNumber="224">type</typeStatus>
series and the first to discover that it could possibly be a new undescribed species on
<collectingRegion id="49A4F854FFA5FF8AC6D8F89CC7DDF8F2" box="[149,200,1828,1851]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="224">Java</collectingRegion>
.
</paragraph>
<paragraph id="8BDF36B6FFA5FF8AC63CF8DBC5FAF8B5" blockId="5.[113,751,1891,1916]" box="[113,751,1891,1916]" pageId="5" pageNumber="224">
<emphasis id="B914EAA4FFA5FF8AC63CF8DBC6F9F8B5" bold="true" box="[113,492,1891,1916]" pageId="5" pageNumber="224">Suggested English common name.</emphasis>
Farits Alhadis tree frog.
</paragraph>
<paragraph id="8BDF36B6FFA5FF8AC63CF81BC7D4F815" blockId="5.[113,752,1955,2012]" pageId="5" pageNumber="224">
<emphasis id="B914EAA4FFA5FF8AC63CF81BC509F875" bold="true" box="[113,540,1955,1980]" pageId="5" pageNumber="224">Suggested Indonesian common name.</emphasis>
Katak-pohon Farits Alhadi.
</paragraph>
</subSubSection>
<subSubSection id="C37A653DFFA5FF8AC63CF7BBC366FAF6" pageId="5" pageNumber="224" type="diagnosis">
<paragraph id="8BDF36B6FFA5FF8AC63CF7BBC5E5F7F2" blockId="5.[113,752,2051,2108]" pageId="5" pageNumber="224">
<emphasis id="B914EAA4FFA5FF8AC63CF7BBC7FBF7D2" bold="true" box="[113,238,2051,2075]" pageId="5" pageNumber="224">Diagnosis.</emphasis>
Based on the molecular analyses, the new species is nested within the clade of the genus
<taxonomicName id="4C604D35FFA5FF8AC42BF79CC5FCF7F5" authorityName="Li, Jiang, Ren &amp; Jiang" authorityYear="2019" box="[614,745,2084,2108]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="224" phylum="Chordata" rank="genus">
<emphasis id="B914EAA4FFA5FF8AC42BF79CC5FCF7F5" box="[614,745,2084,2108]" italics="true" pageId="5" pageNumber="224">Zhangixalus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF36B6FFA5FF8AC560FC9FC366FAF6" blockId="5.[811,1452,807,1343]" pageId="5" pageNumber="224">
<taxonomicName id="4C604D35FFA5FF8AC560FC9FC32FFCF6" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[813,1082,807,831]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="224" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFA5FF8AC560FC9FC32FFCF6" box="[813,1082,807,831]" italics="true" pageId="5" pageNumber="224">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFA5FF8AC20BFC90C3DCFCF6" box="[1094,1225,807,831]" pageId="5" pageNumber="224" rank="species">new species</taxonomicNameLabel>
, can be differentiated from its congeners using the following combination of morphological characters: (1) relatively medium-sized body (SVL
<quantity id="4C989B53FFA5FF8AC5FEFC3FC341FC56" box="[947,1108,903,927]" metricMagnitude="-2" metricUnit="m" metricValue="3.915" metricValueMax="4.07" metricValueMin="3.76" pageId="5" pageNumber="224" unit="mm" value="39.15" valueMax="40.7" valueMin="37.6">37.640.7 mm</quantity>
in males; SVL
<quantity id="4C989B53FFA5FF8AC346FC3FC2BEFC56" box="[1291,1451,903,927]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="5.45" metricValueMin="5.07" pageId="5" pageNumber="224" unit="mm" value="52.6" valueMax="54.5" valueMin="50.7">50.754.5 mm</quantity>
in females); (2) yellowish-green dorsum colouration with numerous dark spot and irregular blotches cream; (3) surfaces of dorsum smooth, covered with small and low spicules; (4) snout in lateral view slope with blunt edge; (5) fringes of skin along limbs and tarsal projection; (6) extensive webbing of toes; (7) finger webbing in males between first to second fingers, and that between second to third fingers reduced, only reaching the distal subarticular tubercle, whereas that in females much more extensive, reaching the base of the disc; (8) webbing between third to fifth toes red; (9) well-developed bilobed supracloacal dermal ridge and tarsal dermal ridge; (10) presence of vomerine teeth; (11) and absence of vocal sac opening.
</paragraph>
</subSubSection>
<subSubSection id="C37A653DFFA5FF85C560FADFC495FC47" lastPageId="10" lastPageNumber="229" pageId="5" pageNumber="224" type="description">
<paragraph id="8BDF36B6FFA5FF8AC560FADFC2B9F876" blockId="5.[812,1453,1383,1983]" pageId="5" pageNumber="224">
<emphasis id="B914EAA4FFA5FF8AC560FADFC359FA49" bold="true" box="[813,1100,1383,1408]" pageId="5" pageNumber="224">
Description of
<typeStatus id="54DB8814FFA5FF8AC5ADFAD0C352FA49" box="[992,1095,1384,1408]" pageId="5" pageNumber="224" type="holotype">holotype</typeStatus>
.
</emphasis>
MZB Amph 32885 (
<figureCitation id="135B2A33FFA5FF8AC30CFADFC2B3FA49" box="[1345,1446,1383,1408]" captionStart="Fig" captionStartId="6.[136,171,1098,1120]" captionTargetBox="[136,1474,189,1083]" captionTargetId="figure-8@6.[133,1477,184,1086]" captionTargetPageId="6" captionText="Fig. 4. A, Zhangixalus faritsalhadii, new species, male holotype in life, MZB Amph 32885; B, female paratype, MZB Amph 32887; C, MZB Amph 32889, female paratype in ventral view; D, MZB Amph 32886 and MZB Amph 32889 in amplexus. Photographs by Misbahul Munir (AC) and Fajar Kaprawi (D)." pageId="5" pageNumber="224">Figs. 4A</figureCitation>
,
<figureCitation id="135B2A33FFA5FF8AC560FA3FC478FA56" box="[813,877,1415,1439]" captionStart="Fig" captionStartId="7.[113,148,1967,1989]" captionTargetBox="[119,1451,288,1929]" captionTargetId="figure-12@7.[103,1472,263,1945]" captionTargetPageId="7" captionText="Fig. 5. Zhangixalus faritsalhadii, new species, from Mount Slamet, Java, Indonesia. AF, an adult male holotype, MZB Amph 32885; GL, an adult female paratype, MZB Amph 32887. Scale bar = 10 mm. Photographs by Misbahul Munir." pageId="5" pageNumber="224">5AF</figureCitation>
), an adult male SVL
<quantity id="4C989B53FFA5FF8AC219FA30C3A2FA56" box="[1108,1207,1416,1440]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="5" pageNumber="224" unit="mm" value="40.7">40.7 mm</quantity>
; head longer (HL
<quantity id="4C989B53FFA5FF8AC330FA30C44FFA76" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="5" pageNumber="224" unit="mm" value="14.2">14.2 mm</quantity>
, 34.8 % SVL) than wider (HW
<quantity id="4C989B53FFA5FF8AC2E1FA10C218FA76" box="[1196,1293,1448,1472]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="5" pageNumber="224" unit="mm" value="13.8">13.8 mm</quantity>
, 33.9 % SVL); snout longer (SL
<quantity id="4C989B53FFA5FF8AC5A3FA7FC350FA16" box="[1006,1093,1479,1503]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="224" unit="mm" value="6.5">6.5 mm</quantity>
, 15.9 % SVL) than eye distance (ED
<quantity id="4C989B53FFA5FF8AC512FA50C4A6FA36" box="[863,947,1512,1536]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="224" unit="mm" value="4.4">4.4 mm</quantity>
, 10.7 % SVL), laterally sloped with blunt edge, and projecting over lower jaw; canthus rostralis distinct; lore oblique, concave; nostril slightly protuberant, without flap of skin, closer to the tip of the snout (SNL
<quantity id="4C989B53FFA5FF8AC302F9F0C2B2F996" box="[1359,1447,1608,1632]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="224" unit="mm" value="3.1">3.1 mm</quantity>
, 7.6 % SVL) than to the eye (NEL
<quantity id="4C989B53FFA5FF8AC2FAF9D0C21BF9B6" box="[1207,1294,1640,1664]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="224" unit="mm" value="3.2">3.2 mm</quantity>
, 7.8 % SVL); internarial distance (IND
<quantity id="4C989B53FFA5FF8AC21CF930C3BFF956" box="[1105,1194,1672,1696]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="224" unit="mm" value="4.4">4.4 mm</quantity>
, 10.7 % SVL) smaller than interorbital distance (IOD
<quantity id="4C989B53FFA5FF8AC2D3F91FC3ECF976" box="[1182,1273,1703,1727]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="5" pageNumber="224" unit="mm" value="5.3">5.3 mm</quantity>
, 12.9 % SVL); interorbital distance wider than the eyelid (UEW
<quantity id="4C989B53FFA5FF8AC31DF970C2B2F916" box="[1360,1447,1736,1760]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="224" unit="mm" value="3.1">3.1 mm</quantity>
, 7.7 % SVL); pineal spot absent; pupil of eye horizontal; tympanum distinct, circular, diameter (TD
<quantity id="4C989B53FFA5FF8AC352F8B0C26FF8D6" box="[1311,1402,1800,1824]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="224" unit="mm" value="3.8">3.8 mm</quantity>
, 9.4 % SVL) approximately half eye length; choanae circular; vomerine teeth in straight transverse in two series between the choanae and on a level with their anterior borders, not touching the choanae and separated from each other by about two individual ridges; and vocal sac opening absent.
</paragraph>
<paragraph id="8BDF36B6FFA5FF89C560F85FC412F9C0" blockId="5.[813,1452,2023,2112]" lastBlockId="6.[136,776,1233,1545]" lastPageId="6" lastPageNumber="225" pageId="5" pageNumber="224">
Forelimb slender; relative finger length I &lt;II &lt;IV &lt;III, second finger (Fin2L
<quantity id="4C989B53FFA5FF8AC26EF7B0C369F7D6" box="[1059,1148,2056,2080]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="224" unit="mm" value="4.2">4.2 mm</quantity>
, 10.2 % SVL) longer than the first finger (Fin1L
<quantity id="4C989B53FFA5FF8AC26DF790C363F7F6" box="[1056,1142,2088,2112]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="5" pageNumber="224" unit="mm" value="3.4">3.4 mm</quantity>
, 8.4 % SVL); tips of fingers expanded into large discs (3FDW
<quantity id="4C989B53FFA6FF89C45EFB69C57BFB20" box="[531,622,1233,1257]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="6" pageNumber="225" unit="mm" value="2.3">2.3 mm</quantity>
, 5.7 % SVL) with circummarginal and transverse ventral grooves; fingers incomplete webbing, webbing between first and second fingers reduced, it only reaches distal subarticular tubercle on both fingers, webbing formula I 22 II 12 III 11 IV; nuptial excrescences on first finger, from the base of the first finger to the level of the subarticular tubercle; supernumerary tubercles present, but indistinct; prepollex absent; inner palmar tubercle oval (IPTL
<quantity id="4C989B53FFA6FF89C78DFA69C502FA20" box="[448,535,1489,1513]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="225" unit="mm" value="2.6">2.6 mm</quantity>
, 6.3 % SVL), middle palmar tubercle indistinct, and outer palmar tubercle oval.
</paragraph>
<caption id="DF1F663EFFA6FF89C6C5FBF2C6E2FB50" pageId="6" pageNumber="225" startId="6.[136,171,1098,1120]" targetBox="[136,1474,189,1083]" targetPageId="6" targetType="figure">
<paragraph id="8BDF36B6FFA6FF89C6C5FBF2C6E2FB50" blockId="6.[136,1476,1098,1178]" pageId="6" pageNumber="225">
Fig. 4. A,
<taxonomicName id="4C604D35FFA6FF89C6A2FBF2C6F1FB96" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[239,484,1098,1119]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="225" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFA6FF89C6A2FBF2C6F1FB96" box="[239,484,1098,1119]" italics="true" pageId="6" pageNumber="225">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFA6FF89C7BFFBF3C57FFBA9" box="[498,618,1098,1120]" pageId="6" pageNumber="225" rank="species">new species</taxonomicNameLabel>
, male holotype in life, MZB Amph 32885; B, female paratype, MZB Amph 32887; C, MZB Amph 32889, female paratype in ventral view; D, MZB Amph 32886 and MZB Amph 32889 in amplexus. Photographs by Misbahul Munir (AC) and Fajar Kaprawi (D).
</paragraph>
</caption>
<paragraph id="8BDF36B6FFA6FF89C6C5F989C41DF860" blockId="6.[134,777,1585,1961]" pageId="6" pageNumber="225">
Hindlimb slender; relative length of toes I &lt;II &lt;III &lt;V &lt;IV, tibiotarsal articulation of adpressed limb reaching eye; thigh (FML
<quantity id="4C989B53FFA6FF89C75DF9C9C661F940" box="[272,372,1649,1673]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="6" pageNumber="225" unit="mm" value="18.1">18.1 mm</quantity>
, 44.5 % SVL) shorter than tibia (TL
<quantity id="4C989B53FFA6FF89C6C7F929C7FBF960" box="[138,238,1681,1705]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="6" pageNumber="225" unit="mm" value="19.1">19.1 mm</quantity>
, 46.8 % SVL); foot (FL
<quantity id="4C989B53FFA6FF89C448F929C57CF960" box="[517,617,1681,1705]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="6" pageNumber="225" unit="mm" value="16.3">16.3 mm</quantity>
, 40.1 % SVL) shorter than tibia; tips of toes expanded into a round disc with circummarginal grooves (4TDW
<quantity id="4C989B53FFA6FF89C790F969C53BF920" box="[477,558,1745,1769]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="225" unit="mm" value="1.8">1.8 mm</quantity>
, 4.5 % SVL), smaller than disc of fingers; toe webbing more extensive, toe webbing formula I 11 II 11 III 11 IV 11 V; subarticular tubercles well-developed, rounded; supernumerary tubercles indistinct; inner metatarsal tubercles (IMTL
<quantity id="4C989B53FFA6FF89C7BCF8E9C550F8A0" box="[497,581,1873,1897]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="225" unit="mm" value="1.6">1.6 mm</quantity>
, 4.0 % SVL) oval and low, length more than half length of first toe (Toe1L
<quantity id="4C989B53FFA6FF89C6C5F829C7F7F860" box="[136,226,1937,1961]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="225" unit="mm" value="2.7">2.7 mm</quantity>
, 6.7 % SVL); outer metatarsal tubercles absent.
</paragraph>
<paragraph id="8BDF36B6FFA6FF89C6C5F86AC22DFA40" blockId="6.[135,777,2001,2089]" lastBlockId="6.[835,1476,1233,1417]" pageId="6" pageNumber="225">Dorsum smooth, covered with small and low spicules, also occurs on eyelid and densely in head; abdomen, chest, throat, thigh, cloaca, and flank prominently granular; flanks wrinkled; an oblique thickened skin forms a low supratympanic fold, from the eye and indistinctly ending above arm insertion, slightly touching the tympanic annulus; dorsolateral fold absent; fringe of skin present along the edge of forearm and the outer edge of fifth to tibiotarsal articulation; dermal ridge at tibiotarsal curved, and supracloaca bilobed.</paragraph>
<paragraph id="8BDF36B6FFA6FF89C50EFA09C2D4F840" blockId="6.[834,1476,1457,1929]" pageId="6" pageNumber="225">
<emphasis id="B914EAA4FFA6FF89C50EFA09C48FFA00" bold="true" box="[835,922,1457,1481]" pageId="6" pageNumber="225">Colour.</emphasis>
In life, dorsum yellowish-green with numerous dark spots and irregular cream blotches, dark spots prominent in eyelid, head, and nostril regions; lower lips cream; from the anterior eye to above the nostril, extending to the tip of the snout with a cream line; limbs without crossbars; abdomen, thighs, ventral of tarsus, tibia, lower arms, and upper arms translucent; throat, flanks, elbows, and knees covered with white spot covering the granular; eyes with black periphery, white sclera, dark pupils, and thin pale yellowish network of iris; webbing between third to fifth toes red, yellowish-green on remaining toes; tips of fingers and toes bluishgreen; inguinal region, ventral elbows and knees bluish. In preservative, dorsum green fading to cream and cream blotches become faded, but dark spots remain unchanged; red webbing on toes fading to light red and eyelids turn black.
</paragraph>
<paragraph id="8BDF36B6FFA6FF89C50EF809C2D7F7E0" blockId="6.[835,1474,1969,2089]" pageId="6" pageNumber="225">
<emphasis id="B914EAA4FFA6FF89C50EF809C4D7F800" bold="true" box="[835,962,1969,1993]" pageId="6" pageNumber="225">Variations.</emphasis>
The morphometric variations are listed in
<tableCitation id="C6E2030DFFA6FF89C3C8F809C441F820" captionStart="Table 2" captionStartId="4.[111,166,493,516]" captionText="Table 2. Morphometric variation of selected characters of Zhanigxalus faritsalhadii, new species, with other Sundaland species of Zhangixalus in ratios (R) values, with T-test values. ZF: Zhangixalus faritsalhadii, new species; ZP: Zhangixalus prominanus." pageId="6" pageNumber="225">Table 2</tableCitation>
. Males had larger RHL, RHW, RSNL, RNEL, RED, RTD, RIND, RUEW, RIPTL, and RIMTL than females, while females had larger SVL, RIOD, RBL, RLAL, Rfin1L,
</paragraph>
<caption id="DF1F663EFFA7FF88C63CF817C391F82B" pageId="7" pageNumber="226" startId="7.[113,148,1967,1989]" targetBox="[119,1451,288,1929]" targetPageId="7" targetType="figure">
<paragraph id="8BDF36B6FFA7FF88C63CF817C391F82B" blockId="7.[113,1452,1967,2018]" pageId="7" pageNumber="226">
Fig. 5.
<taxonomicName id="4C604D35FFA7FF88C6F6F817C6A4F80D" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[187,433,1967,1988]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="7" pageNumber="226" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFA7FF88C6F6F817C6A4F80D" box="[187,433,1967,1988]" italics="true" pageId="7" pageNumber="226">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFA7FF88C78DF808C52DF80C" box="[448,568,1967,1989]" pageId="7" pageNumber="226" rank="species">new species</taxonomicNameLabel>
, from Mount Slamet, Java, Indonesia. AF, an adult male holotype, MZB Amph 32885; GL, an adult female paratype, MZB Amph 32887. Scale bar = 10 mm. Photographs by Misbahul Munir.
</paragraph>
</caption>
<caption id="DF1F663EFFA8FF87C6CAF97DC699F931" pageId="8" pageNumber="227" startId="8.[135,170,1733,1755]" targetBox="[138,1476,188,1717]" targetPageId="8" targetType="figure">
<paragraph id="8BDF36B6FFA8FF87C6CAF97DC699F931" blockId="8.[135,1474,1733,1784]" pageId="8" pageNumber="227">
Fig. 6. Comparison of morphological characters between male species of
<taxonomicName id="4C604D35FFA8FF87C52DF97DC4C2F913" authorityName="Li, Jiang, Ren &amp; Jiang" authorityYear="2019" box="[864,983,1733,1754]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="8" pageNumber="227" phylum="Chordata" rank="genus">
<emphasis id="B914EAA4FFA8FF87C52DF97DC4C2F913" box="[864,983,1733,1754]" italics="true" pageId="8" pageNumber="227">Zhangixalus</emphasis>
</taxonomicName>
from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli.
</paragraph>
</caption>
<paragraph id="8BDF36B6FFA8FF87C6C5F897C23BF801" blockId="8.[135,775,1839,2087]" lastBlockId="8.[835,1476,1839,1992]" pageId="8" pageNumber="227">
Rfin2L, Rfin3L, Rfin4L, RFin1DW, RToe3DW, RFML, RTL, RTSL, RFL, RToe1L, RToe2L, RToe3L, RToe4L, RToe5L, RToe1WD, RToe3DW, and RToe4DW than males. A male
<typeStatus id="54DB8814FFA8FF87C6C4F828C7FCF861" box="[137,233,1936,1960]" pageId="8" pageNumber="227" type="paratype">paratype</typeStatus>
(MZB Amph 32890) had a smaller body than others. All types of females had a smoother dorsum with no spicules compared to males. In females, webbing on first to second and second to third fingers was more extensive than that for males. Finger webbing formula for females: I 21 II 11 III 11 IV (
<figureCitation id="135B2A33FFA8FF87C271F897C386F88E" box="[1084,1171,1839,1863]" captionStart="Fig" captionStartId="7.[113,148,1967,1989]" captionTargetBox="[119,1451,288,1929]" captionTargetId="figure-12@7.[103,1472,263,1945]" captionTargetPageId="7" captionText="Fig. 5. Zhangixalus faritsalhadii, new species, from Mount Slamet, Java, Indonesia. AF, an adult male holotype, MZB Amph 32885; GL, an adult female paratype, MZB Amph 32887. Scale bar = 10 mm. Photographs by Misbahul Munir." pageId="8" pageNumber="227">Fig. 5K</figureCitation>
). The tibiotarsal articulation varies among females, but does not exceed the nose and eyes; that for MZB Amph 32887 reaches the nose, that for MZB Amph 32888 reaches between the nose and eyes, and that for MZB Amph 32889 reaches the eyes.
</paragraph>
<paragraph id="8BDF36B6FFA8FF87C50EF857C2D6F7EE" blockId="8.[835,1475,2031,2087]" pageId="8" pageNumber="227">
<emphasis id="B914EAA4FFA8FF87C50EF857C4E1F7CE" bold="true" box="[835,1012,2031,2055]" pageId="8" pageNumber="227">Comparisons.</emphasis>
Based on the phylogenetic tree and geographical proximity that separated Sundaland species (
<emphasis id="B914EAA4FFA8FF87C3E0F7A8C2D6F7EE" box="[1453,1475,2064,2087]" italics="true" pageId="8" pageNumber="227">Z.</emphasis>
</paragraph>
<caption id="DF1F663EFFA9FF86C622FDB5C3CCFDED" box="[111,1241,525,548]" pageId="9" pageNumber="228" startId="9.[111,166,525,548]" targetType="table">
<paragraph id="8BDF36B6FFA9FF86C622FDB5C3CCFDED" blockId="9.[111,1241,525,548]" box="[111,1241,525,548]" pageId="9" pageNumber="228">
Table 3. Morphometric variation of selected characters of
<taxonomicName id="4C604D35FFA9FF86C4E2FDB5C4DDFDED" baseAuthorityName="Harvey, Pemberton &amp; Smith" baseAuthorityYear="2002" box="[687,968,525,548]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="achantharrhena">
<emphasis id="B914EAA4FFA9FF86C4E2FDB5C4DDFDED" box="[687,968,525,548]" italics="true" pageId="9" pageNumber="228">Zhangixalus achantharrhena</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D35FFA9FF86C5B5FDB5C3C6FDED" baseAuthorityName="Boulenger" baseAuthorityYear="1892" box="[1016,1235,525,548]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="dulitensis">
<emphasis id="B914EAA4FFA9FF86C5B5FDB5C3C6FDED" box="[1016,1235,525,548]" italics="true" pageId="9" pageNumber="228">Zhangixalus dulitensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BDF36B6FFA9FF86C7AFFDECC080F8EC" pageId="9" pageNumber="228">
<table id="F960C416FFA90070C633FDECC0A2F8EC" box="[126,1975,596,1829]" gridcols="9" gridrows="31" pageId="9" pageNumber="228">
<tr id="355034F4FFA90070C633FDECC0A2FDA2" box="[126,1975,596,619]" gridrow="0" pageId="9" pageNumber="228" rowspan-0="1" rowspan-4="1" rowspan-5="1" rowspan-8="1">
<th id="76815D88FFA90070C73DFDECC491FDA2" box="[368,900,596,619]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="9" pageNumber="228">
<taxonomicName id="4C604D35FFA9FF86C7AFFDECC41DFDA2" baseAuthorityName="Harvey, Pemberton &amp; Smith" baseAuthorityYear="2002" box="[482,776,596,619]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="achantharrhena">
<emphasis id="B914EAA4FFA9FF86C7AFFDECC41DFDA2" bold="true" box="[482,776,596,619]" italics="true" pageId="9" pageNumber="228">Zhangixalus achantharrhena</emphasis>
</taxonomicName>
</th>
<th id="76815D88FFA90070C33CFDECC1E8FDA2" box="[1393,1789,596,619]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="9" pageNumber="228">
<taxonomicName id="4C604D35FFA9FF86C33CFDECC140FDA2" authority="Characters" authorityName="Characters" box="[1393,1621,596,619]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="228" phylum="Chordata" rank="species" species="dulitensis">
<emphasis id="B914EAA4FFA9FF86C33CFDECC140FDA2" bold="true" box="[1393,1621,596,619]" italics="true" pageId="9" pageNumber="228">Zhangixalus dulitensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="355034F4FFA90070C633FD3BC0A2FD53" box="[126,1975,643,666]" gridrow="1" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FD3BC61AFD53" box="[126,271,643,666]" gridcol="0" gridrow="1" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C6D1FD3BC61AFD53" bold="true" box="[156,271,643,666]" pageId="9" pageNumber="228">Characters</emphasis>
</th>
<td id="76815D88FFA90070C73DFD3BC5B3FD53" box="[368,678,643,666]" colspan="2" colspanRight="1" gridcol="1" gridrow="1" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C78EFD3BC54EFD53" bold="true" box="[451,603,643,666]" pageId="9" pageNumber="228">Males (n = 13)</emphasis>
</td>
<td id="76815D88FFA90070C543FD3BC359FD53" box="[782,1100,643,666]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C529FD3BC313FD53" bold="true" box="[868,1030,643,666]" pageId="9" pageNumber="228">Females (n = 6)</emphasis>
</td>
<td id="76815D88FFA90070C284FD3BC101FD53" box="[1225,1556,643,666]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C36CFD3BC2B9FD53" bold="true" box="[1313,1452,643,666]" pageId="9" pageNumber="228">Males (n = 9)</emphasis>
</td>
<td id="76815D88FFA90070C019FD3BC0A2FD53" box="[1620,1975,643,666]" colspan="2" colspanRight="1" gridcol="7" gridrow="1" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C084FD3BC07EFD53" bold="true" box="[1737,1899,643,666]" pageId="9" pageNumber="228">Females (n = 2)</emphasis>
</td>
</tr>
<tr id="355034F4FFA90070C633FD0AC0A2FD00" box="[126,1975,690,713]" gridrow="2" pageId="9" pageNumber="228" rowspan-0="1">
<td id="76815D88FFA90070C73DFD0AC6C1FD00" box="[368,468,690,713]" gridcol="1" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C7CEFD0AC6D5FD00" bold="true" box="[387,448,690,713]" pageId="9" pageNumber="228">Males</emphasis>
</td>
<td id="76815D88FFA90070C408FD0AC5B3FD00" box="[581,678,690,713]" gridcol="2" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C406FD0AC58AFD00" bold="true" box="[587,671,690,713]" pageId="9" pageNumber="228">Females</emphasis>
</td>
<td id="76815D88FFA90070C543FD0AC491FD00" box="[782,900,690,713]" gridcol="3" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C543FD0AC491FD00" bold="true" box="[782,900,690,713]" pageId="9" pageNumber="228">Mean ± SD</emphasis>
</td>
<td id="76815D88FFA90070C5A6FD0AC359FD00" box="[1003,1100,690,713]" gridcol="4" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C5B6FD0AC329FD00" bold="true" box="[1019,1084,690,713]" pageId="9" pageNumber="228">Range</emphasis>
</td>
<td id="76815D88FFA90070C284FD0AC255FD00" box="[1225,1344,690,713]" gridcol="5" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C284FD0AC255FD00" bold="true" box="[1225,1344,690,713]" pageId="9" pageNumber="228">Mean ± SD</emphasis>
</td>
<td id="76815D88FFA90070C33CFD0AC101FD00" box="[1393,1556,690,713]" gridcol="6" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C38EFD0AC111FD00" bold="true" box="[1475,1540,690,713]" pageId="9" pageNumber="228">Range</emphasis>
</td>
<td id="76815D88FFA90070C019FD0AC1E8FD00" box="[1620,1789,690,713]" gridcol="7" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C0CBFD0AC1E8FD00" bold="true" box="[1670,1789,690,713]" pageId="9" pageNumber="228">Mean ± SD</emphasis>
</td>
<td id="76815D88FFA90070C100FD0AC0A2FD00" box="[1869,1975,690,713]" gridcol="8" gridrow="2" pageId="9" pageNumber="228">
<emphasis id="B914EAA4FFA9FF86C13BFD0AC0A2FD00" bold="true" box="[1910,1975,690,713]" pageId="9" pageNumber="228">Range</emphasis>
</td>
</tr>
<tr id="355034F4FFA90070C633FD50C0A2FD36" box="[126,1975,744,767]" gridrow="3" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FD50C61AFD36" box="[126,271,744,767]" gridcol="0" gridrow="3" pageId="9" pageNumber="228">SVL</th>
<td id="76815D88FFA90070C73DFD50C6C1FD36" box="[368,468,744,767]" gridcol="1" gridrow="3" pageId="9" pageNumber="228">35.0 ± 1.1</td>
<td id="76815D88FFA90070C408FD50C5B3FD36" box="[581,678,744,767]" gridcol="2" gridrow="3" pageId="9" pageNumber="228">33.537.9</td>
<td id="76815D88FFA90070C543FD50C491FD36" box="[782,900,744,767]" gridcol="3" gridrow="3" pageId="9" pageNumber="228">43.3 ± 1.8</td>
<td id="76815D88FFA90070C5A6FD50C359FD36" box="[1003,1100,744,767]" gridcol="4" gridrow="3" pageId="9" pageNumber="228">39.644.6</td>
<td id="76815D88FFA90070C284FD50C255FD36" box="[1225,1344,744,767]" gridcol="5" gridrow="3" pageId="9" pageNumber="228">35.9 ± 1.6</td>
<td id="76815D88FFA90070C33CFD50C101FD36" box="[1393,1556,744,767]" gridcol="6" gridrow="3" pageId="9" pageNumber="228">33.638.7</td>
<td id="76815D88FFA90070C019FD50C1E8FD36" box="[1620,1789,744,767]" gridcol="7" gridrow="3" pageId="9" pageNumber="228">46.4 ± 0.6</td>
<td id="76815D88FFA90070C100FD50C0A2FD36" box="[1869,1975,744,767]" gridcol="8" gridrow="3" pageId="9" pageNumber="228">46.046.9</td>
</tr>
<tr id="355034F4FFA90070C633FCA8C0A2FCEE" box="[126,1975,784,807]" gridrow="4" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FCA8C61AFCEE" box="[126,271,784,807]" gridcol="0" gridrow="4" pageId="9" pageNumber="228">RHL</th>
<td id="76815D88FFA90070C73DFCA8C6C1FCEE" box="[368,468,784,807]" gridcol="1" gridrow="4" pageId="9" pageNumber="228">33.7 ± 1.1</td>
<td id="76815D88FFA90070C408FCA8C5B3FCEE" box="[581,678,784,807]" gridcol="2" gridrow="4" pageId="9" pageNumber="228">31.435.5</td>
<td id="76815D88FFA90070C543FCA8C491FCEE" box="[782,900,784,807]" gridcol="3" gridrow="4" pageId="9" pageNumber="228">33.5 ± 0.4</td>
<td id="76815D88FFA90070C5A6FCA8C359FCEE" box="[1003,1100,784,807]" gridcol="4" gridrow="4" pageId="9" pageNumber="228">32.934.0</td>
<td id="76815D88FFA90070C284FCA8C255FCEE" box="[1225,1344,784,807]" gridcol="5" gridrow="4" pageId="9" pageNumber="228">32.5 ± 0.5</td>
<td id="76815D88FFA90070C33CFCA8C101FCEE" box="[1393,1556,784,807]" gridcol="6" gridrow="4" pageId="9" pageNumber="228">31.532.9</td>
<td id="76815D88FFA90070C019FCA8C1E8FCEE" box="[1620,1789,784,807]" gridcol="7" gridrow="4" pageId="9" pageNumber="228">32.8 ± 0.3</td>
<td id="76815D88FFA90070C100FCA8C0A2FCEE" box="[1869,1975,784,807]" gridcol="8" gridrow="4" pageId="9" pageNumber="228">32.633.0</td>
</tr>
<tr id="355034F4FFA90070C633FC8FC0A2FC87" box="[126,1975,823,846]" gridrow="5" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FC8FC61AFC87" box="[126,271,823,846]" gridcol="0" gridrow="5" pageId="9" pageNumber="228">RHW</th>
<td id="76815D88FFA90070C73DFC8FC6C1FC87" box="[368,468,823,846]" gridcol="1" gridrow="5" pageId="9" pageNumber="228">33.2 ± 0.6</td>
<td id="76815D88FFA90070C408FC8FC5B3FC87" box="[581,678,823,846]" gridcol="2" gridrow="5" pageId="9" pageNumber="228">32.034.5</td>
<td id="76815D88FFA90070C543FC8FC491FC87" box="[782,900,823,846]" gridcol="3" gridrow="5" pageId="9" pageNumber="228">32.6 ± 0.4</td>
<td id="76815D88FFA90070C5A6FC8FC359FC87" box="[1003,1100,823,846]" gridcol="4" gridrow="5" pageId="9" pageNumber="228">32.133.2</td>
<td id="76815D88FFA90070C284FC8FC255FC87" box="[1225,1344,823,846]" gridcol="5" gridrow="5" pageId="9" pageNumber="228">29.7 ± 0.4</td>
<td id="76815D88FFA90070C33CFC8FC101FC87" box="[1393,1556,823,846]" gridcol="6" gridrow="5" pageId="9" pageNumber="228">29.030.1</td>
<td id="76815D88FFA90070C019FC8FC1E8FC87" box="[1620,1789,823,846]" gridcol="7" gridrow="5" pageId="9" pageNumber="228">29.5 ± 0.3</td>
<td id="76815D88FFA90070C100FC8FC0A2FC87" box="[1869,1975,823,846]" gridcol="8" gridrow="5" pageId="9" pageNumber="228">29.229.7</td>
</tr>
<tr id="355034F4FFA90070C633FCE6C0A2FCBC" box="[126,1975,862,885]" gridrow="6" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FCE6C61AFCBC" box="[126,271,862,885]" gridcol="0" gridrow="6" pageId="9" pageNumber="228">RSL</th>
<td id="76815D88FFA90070C73DFCE6C6C1FCBC" box="[368,468,862,885]" gridcol="1" gridrow="6" pageId="9" pageNumber="228">14.7 ± 0.4</td>
<td id="76815D88FFA90070C408FCE6C5B3FCBC" box="[581,678,862,885]" gridcol="2" gridrow="6" pageId="9" pageNumber="228">14.315.9</td>
<td id="76815D88FFA90070C543FCE6C491FCBC" box="[782,900,862,885]" gridcol="3" gridrow="6" pageId="9" pageNumber="228">15.1 ± 0.1</td>
<td id="76815D88FFA90070C5A6FCE6C359FCBC" box="[1003,1100,862,885]" gridcol="4" gridrow="6" pageId="9" pageNumber="228">14.915.4</td>
<td id="76815D88FFA90070C284FCE6C255FCBC" box="[1225,1344,862,885]" gridcol="5" gridrow="6" pageId="9" pageNumber="228">13.9 ± 0.1</td>
<td id="76815D88FFA90070C33CFCE6C101FCBC" box="[1393,1556,862,885]" gridcol="6" gridrow="6" pageId="9" pageNumber="228">13.714.2</td>
<td id="76815D88FFA90070C019FCE6C1E8FCBC" box="[1620,1789,862,885]" gridcol="7" gridrow="6" pageId="9" pageNumber="228">15.2 ± 0.0</td>
<td id="76815D88FFA90070C100FCE6C0A2FCBC" box="[1869,1975,862,885]" gridcol="8" gridrow="6" pageId="9" pageNumber="228">15.115.2</td>
</tr>
<tr id="355034F4FFA90070C633FC3EC0A2FC54" box="[126,1975,902,925]" gridrow="7" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FC3EC61AFC54" box="[126,271,902,925]" gridcol="0" gridrow="7" pageId="9" pageNumber="228">RSNL</th>
<td id="76815D88FFA90070C73DFC3EC6C1FC54" box="[368,468,902,925]" gridcol="1" gridrow="7" pageId="9" pageNumber="228">6.2 ± 0.2</td>
<td id="76815D88FFA90070C408FC3EC5B3FC54" box="[581,678,902,925]" gridcol="2" gridrow="7" pageId="9" pageNumber="228">5.76.5</td>
<td id="76815D88FFA90070C543FC3EC491FC54" box="[782,900,902,925]" gridcol="3" gridrow="7" pageId="9" pageNumber="228">5.9 ± 0.2</td>
<td id="76815D88FFA90070C5A6FC3EC359FC54" box="[1003,1100,902,925]" gridcol="4" gridrow="7" pageId="9" pageNumber="228">5.76.1</td>
<td id="76815D88FFA90070C284FC3EC255FC54" box="[1225,1344,902,925]" gridcol="5" gridrow="7" pageId="9" pageNumber="228">5.9 ± 0.4</td>
<td id="76815D88FFA90070C33CFC3EC101FC54" box="[1393,1556,902,925]" gridcol="6" gridrow="7" pageId="9" pageNumber="228">5.46.4</td>
<td id="76815D88FFA90070C019FC3EC1E8FC54" box="[1620,1789,902,925]" gridcol="7" gridrow="7" pageId="9" pageNumber="228">5.9 ± 0.1</td>
<td id="76815D88FFA90070C100FC3EC0A2FC54" box="[1869,1975,902,925]" gridcol="8" gridrow="7" pageId="9" pageNumber="228">5.86.0</td>
</tr>
<tr id="355034F4FFA90070C633FC15C0A2FC0D" box="[126,1975,941,964]" gridrow="8" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FC15C61AFC0D" box="[126,271,941,964]" gridcol="0" gridrow="8" pageId="9" pageNumber="228">RNEL</th>
<td id="76815D88FFA90070C73DFC15C6C1FC0D" box="[368,468,941,964]" gridcol="1" gridrow="8" pageId="9" pageNumber="228">7.5 ± 0.3</td>
<td id="76815D88FFA90070C408FC15C5B3FC0D" box="[581,678,941,964]" gridcol="2" gridrow="8" pageId="9" pageNumber="228">6.98.0</td>
<td id="76815D88FFA90070C543FC15C491FC0D" box="[782,900,941,964]" gridcol="3" gridrow="8" pageId="9" pageNumber="228">8.8 ± 0.5</td>
<td id="76815D88FFA90070C5A6FC15C359FC0D" box="[1003,1100,941,964]" gridcol="4" gridrow="8" pageId="9" pageNumber="228">7.89.2</td>
<td id="76815D88FFA90070C284FC15C255FC0D" box="[1225,1344,941,964]" gridcol="5" gridrow="8" pageId="9" pageNumber="228">7.3 ± 0.3</td>
<td id="76815D88FFA90070C33CFC15C101FC0D" box="[1393,1556,941,964]" gridcol="6" gridrow="8" pageId="9" pageNumber="228">6.87.8</td>
<td id="76815D88FFA90070C019FC15C1E8FC0D" box="[1620,1789,941,964]" gridcol="7" gridrow="8" pageId="9" pageNumber="228">8.2 ± 0.0</td>
<td id="76815D88FFA90070C100FC15C0A2FC0D" box="[1869,1975,941,964]" gridcol="8" gridrow="8" pageId="9" pageNumber="228">8.28.3</td>
</tr>
<tr id="355034F4FFA90070C633FC6CC0A2FC22" box="[126,1975,980,1003]" gridrow="9" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FC6CC61AFC22" box="[126,271,980,1003]" gridcol="0" gridrow="9" pageId="9" pageNumber="228">RED</th>
<td id="76815D88FFA90070C73DFC6CC6C1FC22" box="[368,468,980,1003]" gridcol="1" gridrow="9" pageId="9" pageNumber="228">11.7 ± 0.4</td>
<td id="76815D88FFA90070C408FC6CC5B3FC22" box="[581,678,980,1003]" gridcol="2" gridrow="9" pageId="9" pageNumber="228">11.312.7</td>
<td id="76815D88FFA90070C543FC6CC491FC22" box="[782,900,980,1003]" gridcol="3" gridrow="9" pageId="9" pageNumber="228">11.8 ± 0.1</td>
<td id="76815D88FFA90070C5A6FC6CC359FC22" box="[1003,1100,980,1003]" gridcol="4" gridrow="9" pageId="9" pageNumber="228">11.512.0</td>
<td id="76815D88FFA90070C284FC6CC255FC22" box="[1225,1344,980,1003]" gridcol="5" gridrow="9" pageId="9" pageNumber="228">12.7 ± 0.1</td>
<td id="76815D88FFA90070C33CFC6CC101FC22" box="[1393,1556,980,1003]" gridcol="6" gridrow="9" pageId="9" pageNumber="228">12.412.9</td>
<td id="76815D88FFA90070C019FC6CC1E8FC22" box="[1620,1789,980,1003]" gridcol="7" gridrow="9" pageId="9" pageNumber="228">12.3 ± 0.2</td>
<td id="76815D88FFA90070C100FC6CC0A2FC22" box="[1869,1975,980,1003]" gridcol="8" gridrow="9" pageId="9" pageNumber="228">12.112.5</td>
</tr>
<tr id="355034F4FFA90070C633FC44C0A2FBDA" box="[126,1975,1020,1043]" gridrow="10" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FC44C61AFBDA" box="[126,271,1020,1043]" gridcol="0" gridrow="10" pageId="9" pageNumber="228">RTD</th>
<td id="76815D88FFA90070C73DFC44C6C1FBDA" box="[368,468,1020,1043]" gridcol="1" gridrow="10" pageId="9" pageNumber="228">8.0 ± 0.2</td>
<td id="76815D88FFA90070C408FC44C5B3FBDA" box="[581,678,1020,1043]" gridcol="2" gridrow="10" pageId="9" pageNumber="228">7.48.1</td>
<td id="76815D88FFA90070C543FC44C491FBDA" box="[782,900,1020,1043]" gridcol="3" gridrow="10" pageId="9" pageNumber="228">8.0 ± 0.2</td>
<td id="76815D88FFA90070C5A6FC44C359FBDA" box="[1003,1100,1020,1043]" gridcol="4" gridrow="10" pageId="9" pageNumber="228">7.78.3</td>
<td id="76815D88FFA90070C284FC44C255FBDA" box="[1225,1344,1020,1043]" gridcol="5" gridrow="10" pageId="9" pageNumber="228">7.5 ± 0.2</td>
<td id="76815D88FFA90070C33CFC44C101FBDA" box="[1393,1556,1020,1043]" gridcol="6" gridrow="10" pageId="9" pageNumber="228">7.17.9</td>
<td id="76815D88FFA90070C019FC44C1E8FBDA" box="[1620,1789,1020,1043]" gridcol="7" gridrow="10" pageId="9" pageNumber="228">7.4 ± 0.4</td>
<td id="76815D88FFA90070C100FC44C0A2FBDA" box="[1869,1975,1020,1043]" gridcol="8" gridrow="10" pageId="9" pageNumber="228">7.17.7</td>
</tr>
<tr id="355034F4FFA90070C633FB9BC0A2FBF3" box="[126,1975,1059,1082]" gridrow="11" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FB9BC61AFBF3" box="[126,271,1059,1082]" gridcol="0" gridrow="11" pageId="9" pageNumber="228">RIND</th>
<td id="76815D88FFA90070C73DFB9BC6C1FBF3" box="[368,468,1059,1082]" gridcol="1" gridrow="11" pageId="9" pageNumber="228">10.3 ± 0.4</td>
<td id="76815D88FFA90070C408FB9BC5B3FBF3" box="[581,678,1059,1082]" gridcol="2" gridrow="11" pageId="9" pageNumber="228">9.510.9</td>
<td id="76815D88FFA90070C543FB9BC491FBF3" box="[782,900,1059,1082]" gridcol="3" gridrow="11" pageId="9" pageNumber="228">9.8 ± 0.2</td>
<td id="76815D88FFA90070C5A6FB9BC359FBF3" box="[1003,1100,1059,1082]" gridcol="4" gridrow="11" pageId="9" pageNumber="228">9.610.2</td>
<td id="76815D88FFA90070C284FB9BC255FBF3" box="[1225,1344,1059,1082]" gridcol="5" gridrow="11" pageId="9" pageNumber="228">11.1 ± 0.3</td>
<td id="76815D88FFA90070C33CFB9BC101FBF3" box="[1393,1556,1059,1082]" gridcol="6" gridrow="11" pageId="9" pageNumber="228">10.611.5</td>
<td id="76815D88FFA90070C019FB9BC1E8FBF3" box="[1620,1789,1059,1082]" gridcol="7" gridrow="11" pageId="9" pageNumber="228">10.8 ± 0.2</td>
<td id="76815D88FFA90070C100FB9BC0A2FBF3" box="[1869,1975,1059,1082]" gridcol="8" gridrow="11" pageId="9" pageNumber="228">10.711.0</td>
</tr>
<tr id="355034F4FFA90070C633FBF2C0A2FBA8" box="[126,1975,1098,1121]" gridrow="12" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FBF2C61AFBA8" box="[126,271,1098,1121]" gridcol="0" gridrow="12" pageId="9" pageNumber="228">RICD</th>
<td id="76815D88FFA90070C73DFBF2C6C1FBA8" box="[368,468,1098,1121]" gridcol="1" gridrow="12" pageId="9" pageNumber="228">19.4 ± 1.0</td>
<td id="76815D88FFA90070C408FBF2C5B3FBA8" box="[581,678,1098,1121]" gridcol="2" gridrow="12" pageId="9" pageNumber="228">17.620.7</td>
<td id="76815D88FFA90070C543FBF2C491FBA8" box="[782,900,1098,1121]" gridcol="3" gridrow="12" pageId="9" pageNumber="228">19.2 ± 0.4</td>
<td id="76815D88FFA90070C5A6FBF2C359FBA8" box="[1003,1100,1098,1121]" gridcol="4" gridrow="12" pageId="9" pageNumber="228">18.519.6</td>
<td id="76815D88FFA90070C284FBF2C255FBA8" box="[1225,1344,1098,1121]" gridcol="5" gridrow="12" pageId="9" pageNumber="228">19.7 ± 0.5</td>
<td id="76815D88FFA90070C33CFBF2C101FBA8" box="[1393,1556,1098,1121]" gridcol="6" gridrow="12" pageId="9" pageNumber="228">18.520.3</td>
<td id="76815D88FFA90070C019FBF2C1E8FBA8" box="[1620,1789,1098,1121]" gridcol="7" gridrow="12" pageId="9" pageNumber="228">21.0 ± 0.0</td>
<td id="76815D88FFA90070C100FBF2C0A2FBA8" box="[1869,1975,1098,1121]" gridcol="8" gridrow="12" pageId="9" pageNumber="228">21.0</td>
</tr>
<tr id="355034F4FFA90070C633FBCAC0A2FB40" box="[126,1975,1138,1161]" gridrow="13" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FBCAC61AFB40" box="[126,271,1138,1161]" gridcol="0" gridrow="13" pageId="9" pageNumber="228">RIOD</th>
<td id="76815D88FFA90070C73DFBCAC6C1FB40" box="[368,468,1138,1161]" gridcol="1" gridrow="13" pageId="9" pageNumber="228">13.9 ± 0.4</td>
<td id="76815D88FFA90070C408FBCAC5B3FB40" box="[581,678,1138,1161]" gridcol="2" gridrow="13" pageId="9" pageNumber="228">12.915.0</td>
<td id="76815D88FFA90070C543FBCAC491FB40" box="[782,900,1138,1161]" gridcol="3" gridrow="13" pageId="9" pageNumber="228">14.2 ± 0.5</td>
<td id="76815D88FFA90070C5A6FBCAC359FB40" box="[1003,1100,1138,1161]" gridcol="4" gridrow="13" pageId="9" pageNumber="228">13.114.8</td>
<td id="76815D88FFA90070C284FBCAC255FB40" box="[1225,1344,1138,1161]" gridcol="5" gridrow="13" pageId="9" pageNumber="228">12.3 ± 0.3</td>
<td id="76815D88FFA90070C33CFBCAC101FB40" box="[1393,1556,1138,1161]" gridcol="6" gridrow="13" pageId="9" pageNumber="228">12.012.8</td>
<td id="76815D88FFA90070C019FBCAC1E8FB40" box="[1620,1789,1138,1161]" gridcol="7" gridrow="13" pageId="9" pageNumber="228">13.3 ± 0.0</td>
<td id="76815D88FFA90070C100FBCAC0A2FB40" box="[1869,1975,1138,1161]" gridcol="8" gridrow="13" pageId="9" pageNumber="228">13.313.4</td>
</tr>
<tr id="355034F4FFA90070C633FB21C0A2FB79" box="[126,1975,1177,1200]" gridrow="14" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FB21C61AFB79" box="[126,271,1177,1200]" gridcol="0" gridrow="14" pageId="9" pageNumber="228">RUEW</th>
<td id="76815D88FFA90070C73DFB21C6C1FB79" box="[368,468,1177,1200]" gridcol="1" gridrow="14" pageId="9" pageNumber="228">7.8 ± 0.5</td>
<td id="76815D88FFA90070C408FB21C5B3FB79" box="[581,678,1177,1200]" gridcol="2" gridrow="14" pageId="9" pageNumber="228">7.18.9</td>
<td id="76815D88FFA90070C543FB21C491FB79" box="[782,900,1177,1200]" gridcol="3" gridrow="14" pageId="9" pageNumber="228">7.5 ± 0.2</td>
<td id="76815D88FFA90070C5A6FB21C359FB79" box="[1003,1100,1177,1200]" gridcol="4" gridrow="14" pageId="9" pageNumber="228">7.38.0</td>
<td id="76815D88FFA90070C284FB21C255FB79" box="[1225,1344,1177,1200]" gridcol="5" gridrow="14" pageId="9" pageNumber="228">8.1 ± 0.2</td>
<td id="76815D88FFA90070C33CFB21C101FB79" box="[1393,1556,1177,1200]" gridcol="6" gridrow="14" pageId="9" pageNumber="228">7.68.3</td>
<td id="76815D88FFA90070C019FB21C1E8FB79" box="[1620,1789,1177,1200]" gridcol="7" gridrow="14" pageId="9" pageNumber="228">7.9 ± 0.1</td>
<td id="76815D88FFA90070C100FB21C0A2FB79" box="[1869,1975,1177,1200]" gridcol="8" gridrow="14" pageId="9" pageNumber="228">7.78.0</td>
</tr>
<tr id="355034F4FFA90070C633FB78C0A2FB1E" box="[126,1975,1216,1239]" gridrow="15" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FB78C61AFB1E" box="[126,271,1216,1239]" gridcol="0" gridrow="15" pageId="9" pageNumber="228">RBH</th>
<td id="76815D88FFA90070C73DFB78C6C1FB1E" box="[368,468,1216,1239]" gridcol="1" gridrow="15" pageId="9" pageNumber="228">21.3 ± 0.6</td>
<td id="76815D88FFA90070C408FB78C5B3FB1E" box="[581,678,1216,1239]" gridcol="2" gridrow="15" pageId="9" pageNumber="228">20.322.4</td>
<td id="76815D88FFA90070C543FB78C491FB1E" box="[782,900,1216,1239]" gridcol="3" gridrow="15" pageId="9" pageNumber="228">21.2 ± 0.4</td>
<td id="76815D88FFA90070C5A6FB78C359FB1E" box="[1003,1100,1216,1239]" gridcol="4" gridrow="15" pageId="9" pageNumber="228">20.521.7</td>
<td id="76815D88FFA90070C284FB78C255FB1E" box="[1225,1344,1216,1239]" gridcol="5" gridrow="15" pageId="9" pageNumber="228">20.8 ± 0.4</td>
<td id="76815D88FFA90070C33CFB78C101FB1E" box="[1393,1556,1216,1239]" gridcol="6" gridrow="15" pageId="9" pageNumber="228">20.021.3</td>
<td id="76815D88FFA90070C019FB78C1E8FB1E" box="[1620,1789,1216,1239]" gridcol="7" gridrow="15" pageId="9" pageNumber="228">18.8 ± 0.4</td>
<td id="76815D88FFA90070C100FB78C0A2FB1E" box="[1869,1975,1216,1239]" gridcol="8" gridrow="15" pageId="9" pageNumber="228">18.519.1</td>
</tr>
<tr id="355034F4FFA90070C633FB50C0A2FB36" box="[126,1975,1256,1279]" gridrow="16" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FB50C61AFB36" box="[126,271,1256,1279]" gridcol="0" gridrow="16" pageId="9" pageNumber="228">RLAL</th>
<td id="76815D88FFA90070C73DFB50C6C1FB36" box="[368,468,1256,1279]" gridcol="1" gridrow="16" pageId="9" pageNumber="228">50.5 ± 0.8</td>
<td id="76815D88FFA90070C408FB50C5B3FB36" box="[581,678,1256,1279]" gridcol="2" gridrow="16" pageId="9" pageNumber="228">49.151.9</td>
<td id="76815D88FFA90070C543FB50C491FB36" box="[782,900,1256,1279]" gridcol="3" gridrow="16" pageId="9" pageNumber="228">51.2 ± 0.3</td>
<td id="76815D88FFA90070C5A6FB50C359FB36" box="[1003,1100,1256,1279]" gridcol="4" gridrow="16" pageId="9" pageNumber="228">50.751.6</td>
<td id="76815D88FFA90070C284FB50C255FB36" box="[1225,1344,1256,1279]" gridcol="5" gridrow="16" pageId="9" pageNumber="228">46.4 ± 0.9</td>
<td id="76815D88FFA90070C33CFB50C101FB36" box="[1393,1556,1256,1279]" gridcol="6" gridrow="16" pageId="9" pageNumber="228">44.747.9</td>
<td id="76815D88FFA90070C019FB50C1E8FB36" box="[1620,1789,1256,1279]" gridcol="7" gridrow="16" pageId="9" pageNumber="228">46.9 ± 0.5</td>
<td id="76815D88FFA90070C100FB50C0A2FB36" box="[1869,1975,1256,1279]" gridcol="8" gridrow="16" pageId="9" pageNumber="228">46.547.3</td>
</tr>
<tr id="355034F4FFA90070C633FAB7C0A2FAEF" box="[126,1975,1295,1318]" gridrow="17" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FAB7C61AFAEF" box="[126,271,1295,1318]" gridcol="0" gridrow="17" pageId="9" pageNumber="228">RFin1L</th>
<td id="76815D88FFA90070C73DFAB7C6C1FAEF" box="[368,468,1295,1318]" gridcol="1" gridrow="17" pageId="9" pageNumber="228">8.0 ± 0.3</td>
<td id="76815D88FFA90070C408FAB7C5B3FAEF" box="[581,678,1295,1318]" gridcol="2" gridrow="17" pageId="9" pageNumber="228">7.38.8</td>
<td id="76815D88FFA90070C543FAB7C491FAEF" box="[782,900,1295,1318]" gridcol="3" gridrow="17" pageId="9" pageNumber="228">9.1 ± 0.5</td>
<td id="76815D88FFA90070C5A6FAB7C359FAEF" box="[1003,1100,1295,1318]" gridcol="4" gridrow="17" pageId="9" pageNumber="228">8.19.6</td>
<td id="76815D88FFA90070C284FAB7C255FAEF" box="[1225,1344,1295,1318]" gridcol="5" gridrow="17" pageId="9" pageNumber="228">8.0 ± 0.3</td>
<td id="76815D88FFA90070C33CFAB7C101FAEF" box="[1393,1556,1295,1318]" gridcol="6" gridrow="17" pageId="9" pageNumber="228">7.58.6</td>
<td id="76815D88FFA90070C019FAB7C1E8FAEF" box="[1620,1789,1295,1318]" gridcol="7" gridrow="17" pageId="9" pageNumber="228">8.6 ± 0.2</td>
<td id="76815D88FFA90070C100FAB7C0A2FAEF" box="[1869,1975,1295,1318]" gridcol="8" gridrow="17" pageId="9" pageNumber="228">8.58.8</td>
</tr>
<tr id="355034F4FFA90070C633FA8EC0A2FA84" box="[126,1975,1334,1357]" gridrow="18" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FA8EC61AFA84" box="[126,271,1334,1357]" gridcol="0" gridrow="18" pageId="9" pageNumber="228">RFin3L</th>
<td id="76815D88FFA90070C73DFA8EC6C1FA84" box="[368,468,1334,1357]" gridcol="1" gridrow="18" pageId="9" pageNumber="228">17.5 ± 0.3</td>
<td id="76815D88FFA90070C408FA8EC5B3FA84" box="[581,678,1334,1357]" gridcol="2" gridrow="18" pageId="9" pageNumber="228">16.918.0</td>
<td id="76815D88FFA90070C543FA8EC491FA84" box="[782,900,1334,1357]" gridcol="3" gridrow="18" pageId="9" pageNumber="228">18.6 ± 0.3</td>
<td id="76815D88FFA90070C5A6FA8EC359FA84" box="[1003,1100,1334,1357]" gridcol="4" gridrow="18" pageId="9" pageNumber="228">17.918.9</td>
<td id="76815D88FFA90070C284FA8EC255FA84" box="[1225,1344,1334,1357]" gridcol="5" gridrow="18" pageId="9" pageNumber="228">16.8 ± 0.2</td>
<td id="76815D88FFA90070C33CFA8EC101FA84" box="[1393,1556,1334,1357]" gridcol="6" gridrow="18" pageId="9" pageNumber="228">16.417.2</td>
<td id="76815D88FFA90070C019FA8EC1E8FA84" box="[1620,1789,1334,1357]" gridcol="7" gridrow="18" pageId="9" pageNumber="228">17.6 ± 0.1</td>
<td id="76815D88FFA90070C100FA8EC0A2FA84" box="[1869,1975,1334,1357]" gridcol="8" gridrow="18" pageId="9" pageNumber="228">17.517.6</td>
</tr>
<tr id="355034F4FFA90070C633FAE6C0A2FABC" box="[126,1975,1374,1397]" gridrow="19" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FAE6C61AFABC" box="[126,271,1374,1397]" gridcol="0" gridrow="19" pageId="9" pageNumber="228">RFin3DW</th>
<td id="76815D88FFA90070C73DFAE6C6C1FABC" box="[368,468,1374,1397]" gridcol="1" gridrow="19" pageId="9" pageNumber="228">5.7 ± 0.3</td>
<td id="76815D88FFA90070C408FAE6C5B3FABC" box="[581,678,1374,1397]" gridcol="2" gridrow="19" pageId="9" pageNumber="228">5.26.3</td>
<td id="76815D88FFA90070C543FAE6C491FABC" box="[782,900,1374,1397]" gridcol="3" gridrow="19" pageId="9" pageNumber="228">6.7 ± 0.2</td>
<td id="76815D88FFA90070C5A6FAE6C359FABC" box="[1003,1100,1374,1397]" gridcol="4" gridrow="19" pageId="9" pageNumber="228">6.47.0</td>
<td id="76815D88FFA90070C284FAE6C255FABC" box="[1225,1344,1374,1397]" gridcol="5" gridrow="19" pageId="9" pageNumber="228">6.6 ± 0.2</td>
<td id="76815D88FFA90070C33CFAE6C101FABC" box="[1393,1556,1374,1397]" gridcol="6" gridrow="19" pageId="9" pageNumber="228">6.37.0</td>
<td id="76815D88FFA90070C019FAE6C1E8FABC" box="[1620,1789,1374,1397]" gridcol="7" gridrow="19" pageId="9" pageNumber="228">6.0 ± 0.1</td>
<td id="76815D88FFA90070C100FAE6C0A2FABC" box="[1869,1975,1374,1397]" gridcol="8" gridrow="19" pageId="9" pageNumber="228">5.96.1</td>
</tr>
<tr id="355034F4FFA90070C633FA3DC0A2FA55" box="[126,1975,1413,1436]" gridrow="20" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FA3DC61AFA55" box="[126,271,1413,1436]" gridcol="0" gridrow="20" pageId="9" pageNumber="228">RIPTL</th>
<td id="76815D88FFA90070C73DFA3DC6C1FA55" box="[368,468,1413,1436]" gridcol="1" gridrow="20" pageId="9" pageNumber="228">4.4 ± 0.2</td>
<td id="76815D88FFA90070C408FA3DC5B3FA55" box="[581,678,1413,1436]" gridcol="2" gridrow="20" pageId="9" pageNumber="228">4.14.8</td>
<td id="76815D88FFA90070C543FA3DC491FA55" box="[782,900,1413,1436]" gridcol="3" gridrow="20" pageId="9" pageNumber="228">5.4 ± 0.2</td>
<td id="76815D88FFA90070C5A6FA3DC359FA55" box="[1003,1100,1413,1436]" gridcol="4" gridrow="20" pageId="9" pageNumber="228">5.05.6</td>
<td id="76815D88FFA90070C284FA3DC255FA55" box="[1225,1344,1413,1436]" gridcol="5" gridrow="20" pageId="9" pageNumber="228">6.4 ± 0.1</td>
<td id="76815D88FFA90070C33CFA3DC101FA55" box="[1393,1556,1413,1436]" gridcol="6" gridrow="20" pageId="9" pageNumber="228">6.16.7</td>
<td id="76815D88FFA90070C019FA3DC1E8FA55" box="[1620,1789,1413,1436]" gridcol="7" gridrow="20" pageId="9" pageNumber="228">6.0 ± 0.0</td>
<td id="76815D88FFA90070C100FA3DC0A2FA55" box="[1869,1975,1413,1436]" gridcol="8" gridrow="20" pageId="9" pageNumber="228">5.96.0</td>
</tr>
<tr id="355034F4FFA90070C633FA14C0A2FA0A" box="[126,1975,1452,1475]" gridrow="21" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FA14C61AFA0A" box="[126,271,1452,1475]" gridcol="0" gridrow="21" pageId="9" pageNumber="228">RFML</th>
<td id="76815D88FFA90070C73DFA14C6C1FA0A" box="[368,468,1452,1475]" gridcol="1" gridrow="21" pageId="9" pageNumber="228">50.7 ± 1.3</td>
<td id="76815D88FFA90070C408FA14C5B3FA0A" box="[581,678,1452,1475]" gridcol="2" gridrow="21" pageId="9" pageNumber="228">48.653.5</td>
<td id="76815D88FFA90070C543FA14C491FA0A" box="[782,900,1452,1475]" gridcol="3" gridrow="21" pageId="9" pageNumber="228">49.4 ± 1.5</td>
<td id="76815D88FFA90070C5A6FA14C359FA0A" box="[1003,1100,1452,1475]" gridcol="4" gridrow="21" pageId="9" pageNumber="228">47.751.7</td>
<td id="76815D88FFA90070C284FA14C255FA0A" box="[1225,1344,1452,1475]" gridcol="5" gridrow="21" pageId="9" pageNumber="228">48.0 ± 2.0</td>
<td id="76815D88FFA90070C33CFA14C101FA0A" box="[1393,1556,1452,1475]" gridcol="6" gridrow="21" pageId="9" pageNumber="228">44.850.9</td>
<td id="76815D88FFA90070C019FA14C1E8FA0A" box="[1620,1789,1452,1475]" gridcol="7" gridrow="21" pageId="9" pageNumber="228">52.2 ± 0.7</td>
<td id="76815D88FFA90070C100FA14C0A2FA0A" box="[1869,1975,1452,1475]" gridcol="8" gridrow="21" pageId="9" pageNumber="228">51.752.7</td>
</tr>
<tr id="355034F4FFA90070C633FA6CC0A2FA22" box="[126,1975,1492,1515]" gridrow="22" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FA6CC61AFA22" box="[126,271,1492,1515]" gridcol="0" gridrow="22" pageId="9" pageNumber="228">RTL</th>
<td id="76815D88FFA90070C73DFA6CC6C1FA22" box="[368,468,1492,1515]" gridcol="1" gridrow="22" pageId="9" pageNumber="228">52.7 ± 1.4</td>
<td id="76815D88FFA90070C408FA6CC5B3FA22" box="[581,678,1492,1515]" gridcol="2" gridrow="22" pageId="9" pageNumber="228">50.855.0</td>
<td id="76815D88FFA90070C543FA6CC491FA22" box="[782,900,1492,1515]" gridcol="3" gridrow="22" pageId="9" pageNumber="228">52.8 ± 0.9</td>
<td id="76815D88FFA90070C5A6FA6CC359FA22" box="[1003,1100,1492,1515]" gridcol="4" gridrow="22" pageId="9" pageNumber="228">51.854.4</td>
<td id="76815D88FFA90070C284FA6CC255FA22" box="[1225,1344,1492,1515]" gridcol="5" gridrow="22" pageId="9" pageNumber="228">51.0 ± 1.8</td>
<td id="76815D88FFA90070C33CFA6CC101FA22" box="[1393,1556,1492,1515]" gridcol="6" gridrow="22" pageId="9" pageNumber="228">48.253.9</td>
<td id="76815D88FFA90070C019FA6CC1E8FA22" box="[1620,1789,1492,1515]" gridcol="7" gridrow="22" pageId="9" pageNumber="228">54.4 ± 0.3</td>
<td id="76815D88FFA90070C100FA6CC0A2FA22" box="[1869,1975,1492,1515]" gridcol="8" gridrow="22" pageId="9" pageNumber="228">54.254.7</td>
</tr>
<tr id="355034F4FFA90070C633FA43C0A2F9DB" box="[126,1975,1531,1554]" gridrow="23" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633FA43C61AF9DB" box="[126,271,1531,1554]" gridcol="0" gridrow="23" pageId="9" pageNumber="228">RTSL</th>
<td id="76815D88FFA90070C73DFA43C6C1F9DB" box="[368,468,1531,1554]" gridcol="1" gridrow="23" pageId="9" pageNumber="228">28.7 ± 0.7</td>
<td id="76815D88FFA90070C408FA43C5B3F9DB" box="[581,678,1531,1554]" gridcol="2" gridrow="23" pageId="9" pageNumber="228">27.229.9</td>
<td id="76815D88FFA90070C543FA43C491F9DB" box="[782,900,1531,1554]" gridcol="3" gridrow="23" pageId="9" pageNumber="228">28.3 ± 0.4</td>
<td id="76815D88FFA90070C5A6FA43C359F9DB" box="[1003,1100,1531,1554]" gridcol="4" gridrow="23" pageId="9" pageNumber="228">27.929.3</td>
<td id="76815D88FFA90070C284FA43C255F9DB" box="[1225,1344,1531,1554]" gridcol="5" gridrow="23" pageId="9" pageNumber="228">26.4 ± 0.5</td>
<td id="76815D88FFA90070C33CFA43C101F9DB" box="[1393,1556,1531,1554]" gridcol="6" gridrow="23" pageId="9" pageNumber="228">25.427.2</td>
<td id="76815D88FFA90070C019FA43C1E8F9DB" box="[1620,1789,1531,1554]" gridcol="7" gridrow="23" pageId="9" pageNumber="228">28.2 ± 0.8</td>
<td id="76815D88FFA90070C100FA43C0A2F9DB" box="[1869,1975,1531,1554]" gridcol="8" gridrow="23" pageId="9" pageNumber="228">27.628.8</td>
</tr>
<tr id="355034F4FFA90070C633F99AC0A2F9F0" box="[126,1975,1570,1593]" gridrow="24" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F99AC61AF9F0" box="[126,271,1570,1593]" gridcol="0" gridrow="24" pageId="9" pageNumber="228">RFL</th>
<td id="76815D88FFA90070C73DF99AC6C1F9F0" box="[368,468,1570,1593]" gridcol="1" gridrow="24" pageId="9" pageNumber="228">43.4 ± 1.0</td>
<td id="76815D88FFA90070C408F99AC5B3F9F0" box="[581,678,1570,1593]" gridcol="2" gridrow="24" pageId="9" pageNumber="228">40.845.0</td>
<td id="76815D88FFA90070C543F99AC491F9F0" box="[782,900,1570,1593]" gridcol="3" gridrow="24" pageId="9" pageNumber="228">47.0 ± 0.3</td>
<td id="76815D88FFA90070C5A6F99AC359F9F0" box="[1003,1100,1570,1593]" gridcol="4" gridrow="24" pageId="9" pageNumber="228">46.647.4</td>
<td id="76815D88FFA90070C284F99AC255F9F0" box="[1225,1344,1570,1593]" gridcol="5" gridrow="24" pageId="9" pageNumber="228">44.1 ± 1.0</td>
<td id="76815D88FFA90070C33CF99AC101F9F0" box="[1393,1556,1570,1593]" gridcol="6" gridrow="24" pageId="9" pageNumber="228">42.946.2</td>
<td id="76815D88FFA90070C019F99AC1E8F9F0" box="[1620,1789,1570,1593]" gridcol="7" gridrow="24" pageId="9" pageNumber="228">49.2 ± 0.4</td>
<td id="76815D88FFA90070C100F99AC0A2F9F0" box="[1869,1975,1570,1593]" gridcol="8" gridrow="24" pageId="9" pageNumber="228">48.949.5</td>
</tr>
<tr id="355034F4FFA90070C633F9F1C0A2F9A9" box="[126,1975,1609,1632]" gridrow="25" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F9F1C61AF9A9" box="[126,271,1609,1632]" gridcol="0" gridrow="25" pageId="9" pageNumber="228">RToe1L</th>
<td id="76815D88FFA90070C73DF9F1C6C1F9A9" box="[368,468,1609,1632]" gridcol="1" gridrow="25" pageId="9" pageNumber="228">7.7 ± 0.2</td>
<td id="76815D88FFA90070C408F9F1C5B3F9A9" box="[581,678,1609,1632]" gridcol="2" gridrow="25" pageId="9" pageNumber="228">7.48.1</td>
<td id="76815D88FFA90070C543F9F1C491F9A9" box="[782,900,1609,1632]" gridcol="3" gridrow="25" pageId="9" pageNumber="228">8.3 ± 0.2</td>
<td id="76815D88FFA90070C5A6F9F1C359F9A9" box="[1003,1100,1609,1632]" gridcol="4" gridrow="25" pageId="9" pageNumber="228">8.08.7</td>
<td id="76815D88FFA90070C284F9F1C255F9A9" box="[1225,1344,1609,1632]" gridcol="5" gridrow="25" pageId="9" pageNumber="228">7.7 ± 0.3</td>
<td id="76815D88FFA90070C33CF9F1C101F9A9" box="[1393,1556,1609,1632]" gridcol="6" gridrow="25" pageId="9" pageNumber="228">7.08.3</td>
<td id="76815D88FFA90070C019F9F1C1E8F9A9" box="[1620,1789,1609,1632]" gridcol="7" gridrow="25" pageId="9" pageNumber="228">8.0 ± 0.0</td>
<td id="76815D88FFA90070C100F9F1C0A2F9A9" box="[1869,1975,1609,1632]" gridcol="8" gridrow="25" pageId="9" pageNumber="228">7.98.0</td>
</tr>
<tr id="355034F4FFA90070C633F9C9C0A2F941" box="[126,1975,1649,1672]" gridrow="26" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F9C9C61AF941" box="[126,271,1649,1672]" gridcol="0" gridrow="26" pageId="9" pageNumber="228">RToe4L</th>
<td id="76815D88FFA90070C73DF9C9C6C1F941" box="[368,468,1649,1672]" gridcol="1" gridrow="26" pageId="9" pageNumber="228">24.4 ± 0.5</td>
<td id="76815D88FFA90070C408F9C9C5B3F941" box="[581,678,1649,1672]" gridcol="2" gridrow="26" pageId="9" pageNumber="228">23.625.3</td>
<td id="76815D88FFA90070C543F9C9C491F941" box="[782,900,1649,1672]" gridcol="3" gridrow="26" pageId="9" pageNumber="228">26.1 ± 0.3</td>
<td id="76815D88FFA90070C5A6F9C9C359F941" box="[1003,1100,1649,1672]" gridcol="4" gridrow="26" pageId="9" pageNumber="228">25.626.7</td>
<td id="76815D88FFA90070C284F9C9C255F941" box="[1225,1344,1649,1672]" gridcol="5" gridrow="26" pageId="9" pageNumber="228">24.7 ± 0.5</td>
<td id="76815D88FFA90070C33CF9C9C101F941" box="[1393,1556,1649,1672]" gridcol="6" gridrow="26" pageId="9" pageNumber="228">24.025.6</td>
<td id="76815D88FFA90070C019F9C9C1E8F941" box="[1620,1789,1649,1672]" gridcol="7" gridrow="26" pageId="9" pageNumber="228">27.2 ± 0.5</td>
<td id="76815D88FFA90070C100F9C9C0A2F941" box="[1869,1975,1649,1672]" gridcol="8" gridrow="26" pageId="9" pageNumber="228">26.827.6</td>
</tr>
<tr id="355034F4FFA90070C633F920C0A2F966" box="[126,1975,1688,1711]" gridrow="27" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F920C61AF966" box="[126,271,1688,1711]" gridcol="0" gridrow="27" pageId="9" pageNumber="228">RToe1DW</th>
<td id="76815D88FFA90070C73DF920C6C1F966" box="[368,468,1688,1711]" gridcol="1" gridrow="27" pageId="9" pageNumber="228">3.1 ± 0.1</td>
<td id="76815D88FFA90070C408F920C5B3F966" box="[581,678,1688,1711]" gridcol="2" gridrow="27" pageId="9" pageNumber="228">3.03.4</td>
<td id="76815D88FFA90070C543F920C491F966" box="[782,900,1688,1711]" gridcol="3" gridrow="27" pageId="9" pageNumber="228">3.6 ± 0.1</td>
<td id="76815D88FFA90070C5A6F920C359F966" box="[1003,1100,1688,1711]" gridcol="4" gridrow="27" pageId="9" pageNumber="228">3.43.7</td>
<td id="76815D88FFA90070C284F920C255F966" box="[1225,1344,1688,1711]" gridcol="5" gridrow="27" pageId="9" pageNumber="228">2.9 ± 0.1</td>
<td id="76815D88FFA90070C33CF920C101F966" box="[1393,1556,1688,1711]" gridcol="6" gridrow="27" pageId="9" pageNumber="228">2.73.0</td>
<td id="76815D88FFA90070C019F920C1E8F966" box="[1620,1789,1688,1711]" gridcol="7" gridrow="27" pageId="9" pageNumber="228">2.6 ± 0.1</td>
<td id="76815D88FFA90070C100F920C0A2F966" box="[1869,1975,1688,1711]" gridcol="8" gridrow="27" pageId="9" pageNumber="228">2.42.7</td>
</tr>
<tr id="355034F4FFA90070C633F907C0A2F91F" box="[126,1975,1727,1750]" gridrow="28" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F907C61AF91F" box="[126,271,1727,1750]" gridcol="0" gridrow="28" pageId="9" pageNumber="228">RToe3DW</th>
<td id="76815D88FFA90070C73DF907C6C1F91F" box="[368,468,1727,1750]" gridcol="1" gridrow="28" pageId="9" pageNumber="228">3.7 ± 0.2</td>
<td id="76815D88FFA90070C408F907C5B3F91F" box="[581,678,1727,1750]" gridcol="2" gridrow="28" pageId="9" pageNumber="228">3.34.1</td>
<td id="76815D88FFA90070C543F907C491F91F" box="[782,900,1727,1750]" gridcol="3" gridrow="28" pageId="9" pageNumber="228">4.9 ± 0.0</td>
<td id="76815D88FFA90070C5A6F907C359F91F" box="[1003,1100,1727,1750]" gridcol="4" gridrow="28" pageId="9" pageNumber="228">4.85.0</td>
<td id="76815D88FFA90070C284F907C255F91F" box="[1225,1344,1727,1750]" gridcol="5" gridrow="28" pageId="9" pageNumber="228">4.1 ± 0.3</td>
<td id="76815D88FFA90070C33CF907C101F91F" box="[1393,1556,1727,1750]" gridcol="6" gridrow="28" pageId="9" pageNumber="228">3.84.9</td>
<td id="76815D88FFA90070C019F907C1E8F91F" box="[1620,1789,1727,1750]" gridcol="7" gridrow="28" pageId="9" pageNumber="228">3.8 ± 0.0</td>
<td id="76815D88FFA90070C100F907C0A2F91F" box="[1869,1975,1727,1750]" gridcol="8" gridrow="28" pageId="9" pageNumber="228">3.73.8</td>
</tr>
<tr id="355034F4FFA90070C633F95FC0A2F937" box="[126,1975,1767,1790]" gridrow="29" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F95FC61AF937" box="[126,271,1767,1790]" gridcol="0" gridrow="29" pageId="9" pageNumber="228">RToe4DW</th>
<td id="76815D88FFA90070C73DF95FC6C1F937" box="[368,468,1767,1790]" gridcol="1" gridrow="29" pageId="9" pageNumber="228">4.3 ± 0.2</td>
<td id="76815D88FFA90070C408F95FC5B3F937" box="[581,678,1767,1790]" gridcol="2" gridrow="29" pageId="9" pageNumber="228">3.94.7</td>
<td id="76815D88FFA90070C543F95FC491F937" box="[782,900,1767,1790]" gridcol="3" gridrow="29" pageId="9" pageNumber="228">5.2 ± 0.1</td>
<td id="76815D88FFA90070C5A6F95FC359F937" box="[1003,1100,1767,1790]" gridcol="4" gridrow="29" pageId="9" pageNumber="228">5.05.4</td>
<td id="76815D88FFA90070C284F95FC255F937" box="[1225,1344,1767,1790]" gridcol="5" gridrow="29" pageId="9" pageNumber="228">5.0 ± 0.5</td>
<td id="76815D88FFA90070C33CF95FC101F937" box="[1393,1556,1767,1790]" gridcol="6" gridrow="29" pageId="9" pageNumber="228">4.05.5</td>
<td id="76815D88FFA90070C019F95FC1E8F937" box="[1620,1789,1767,1790]" gridcol="7" gridrow="29" pageId="9" pageNumber="228">4.4 ± 0.0</td>
<td id="76815D88FFA90070C100F95FC0A2F937" box="[1869,1975,1767,1790]" gridcol="8" gridrow="29" pageId="9" pageNumber="228">4.44.5</td>
</tr>
<tr id="355034F4FFA90070C633F8B6C0A2F8EC" box="[126,1975,1806,1829]" gridrow="30" pageId="9" pageNumber="228">
<th id="76815D88FFA90070C633F8B6C61AF8EC" box="[126,271,1806,1829]" gridcol="0" gridrow="30" pageId="9" pageNumber="228">RIMTL</th>
<td id="76815D88FFA90070C73DF8B6C6C1F8EC" box="[368,468,1806,1829]" gridcol="1" gridrow="30" pageId="9" pageNumber="228">3.9 ± 0.3</td>
<td id="76815D88FFA90070C408F8B6C5B3F8EC" box="[581,678,1806,1829]" gridcol="2" gridrow="30" pageId="9" pageNumber="228">3.44.3</td>
<td id="76815D88FFA90070C543F8B6C491F8EC" box="[782,900,1806,1829]" gridcol="3" gridrow="30" pageId="9" pageNumber="228">3.8 ± 0.0</td>
<td id="76815D88FFA90070C5A6F8B6C359F8EC" box="[1003,1100,1806,1829]" gridcol="4" gridrow="30" pageId="9" pageNumber="228">3.74.0</td>
<td id="76815D88FFA90070C284F8B6C255F8EC" box="[1225,1344,1806,1829]" gridcol="5" gridrow="30" pageId="9" pageNumber="228">2.9 ± 0.2</td>
<td id="76815D88FFA90070C33CF8B6C101F8EC" box="[1393,1556,1806,1829]" gridcol="6" gridrow="30" pageId="9" pageNumber="228">2.63.2</td>
<td id="76815D88FFA90070C019F8B6C1E8F8EC" box="[1620,1789,1806,1829]" gridcol="7" gridrow="30" pageId="9" pageNumber="228">4.0 ± 0.1</td>
<td id="76815D88FFA90070C100F8B6C0A2F8EC" box="[1869,1975,1806,1829]" gridcol="8" gridrow="30" pageId="9" pageNumber="228">3.94.1</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDF36B6FFAAFF85C6C5FF05C7CEFD7C" blockId="10.[135,776,189,693]" pageId="10" pageNumber="229">
<taxonomicName id="4C604D35FFAAFF85C6C5FF05C61CFF1C" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[136,265,189,213]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFAAFF85C6C5FF05C61CFF1C" box="[136,265,189,213]" italics="true" pageId="10" pageNumber="229">faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C759FF06C686FF1C" box="[276,403,189,213]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
,
<taxonomicName id="4C604D35FFAAFF85C7D7FF06C524FF1C" baseAuthorityName="Smith" baseAuthorityYear="1924" box="[410,561,190,213]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="prominanus">
<emphasis id="B914EAA4FFAAFF85C7D7FF06C524FF1C" box="[410,561,190,213]" italics="true" pageId="10" pageNumber="229">Z. prominanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C604D35FFAAFF85C473FF06C415FF1C" baseAuthorityName="Harvey, Pemberton &amp; Smith" baseAuthorityYear="2002" box="[574,768,189,213]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="achantharrhena">
<emphasis id="B914EAA4FFAAFF85C473FF06C558FF1C" box="[574,589,190,213]" italics="true" pageId="10" pageNumber="229">Z</emphasis>
.
<emphasis id="B914EAA4FFAAFF85C415FF05C415FF1C" box="[600,768,189,213]" italics="true" pageId="10" pageNumber="229">achantharrhena</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C604D35FFAAFF85C6F4FF66C657FF3C" box="[185,322,221,245]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="dulitensis">
<emphasis id="B914EAA4FFAAFF85C6F4FF66C657FF3C" box="[185,322,221,245]" italics="true" pageId="10" pageNumber="229">Z. dulitensis</emphasis>
</taxonomicName>
) from Asian mainland species, we only compared
<taxonomicName id="4C604D35FFAAFF85C74FFF46C6A4FEDC" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[258,433,253,277]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="new species">
<emphasis id="B914EAA4FFAAFF85C74FFF46C6A4FEDC" box="[258,433,253,277]" italics="true" pageId="10" pageNumber="229">Z. faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C789FF46C547FEDC" box="[452,594,253,277]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, with the other three Sundaland species. Morphologically,
<taxonomicName id="4C604D35FFAAFF85C42DFEA6C415FEFC" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[608,768,285,309]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="new species">
<emphasis id="B914EAA4FFAAFF85C42DFEA6C415FEFC" box="[608,768,285,309]" italics="true" pageId="10" pageNumber="229">Z. faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C6C5FE86C61BFE9C" box="[136,270,317,341]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, is more similar to
<taxonomicName id="4C604D35FFAAFF85C7ABFE86C591FE9C" baseAuthorityName="Smith" baseAuthorityYear="1924" box="[486,644,318,341]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="prominanus">
<emphasis id="B914EAA4FFAAFF85C7ABFE86C591FE9C" box="[486,644,318,341]" italics="true" pageId="10" pageNumber="229">Z. prominanus</emphasis>
</taxonomicName>
, but can be distinguished from the latter with its smaller body (SVL
<quantity id="4C989B53FFAAFF85C6C5FEC5C631FE5C" box="[136,292,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="3.915" metricValueMax="4.07" metricValueMin="3.76" pageId="10" pageNumber="229" unit="mm" value="39.15" valueMax="40.7" valueMin="37.6">37.640.7 mm</quantity>
vs.
<quantity id="4C989B53FFAAFF85C71EFEC5C6FAFE5C" box="[339,495,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="5.145" metricValueMax="5.85" metricValueMin="4.4399999999999995" pageId="10" pageNumber="229" unit="mm" value="51.45" valueMax="58.5" valueMin="44.4">44.458.5 mm</quantity>
in males; SVL
<quantity id="4C989B53FFAAFF85C4D1FEC5C7A7FE7C" metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="5.45" metricValueMin="5.07" pageId="10" pageNumber="229" unit="mm" value="52.6" valueMax="54.5" valueMin="50.7">50.754.5 mm</quantity>
vs.
<quantity id="4C989B53FFAAFF85C691FE25C661FE7C" box="[220,372,413,437]" metricMagnitude="-2" metricUnit="m" metricValue="6.55" metricValueMax="7.05" metricValueMin="6.05" pageId="10" pageNumber="229" unit="mm" value="65.5" valueMax="70.5" valueMin="60.5">60.570.5 mm</quantity>
in females); dorsum with low spicules which are not prominent in males (vs. appears prominently and elevated;
<figureCitation id="135B2A33FFAAFF85C750FE65C69CFE3C" box="[285,393,477,501]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6I, J</figureCitation>
); males snout slope with blunt edge in lateral view (vs. slope with pointed edge;
<figureCitation id="135B2A33FFAAFF85C43BFE45C5ECFDDC" box="[630,761,509,533]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6A, B</figureCitation>
); supracloaca dermal ridge broader and shorter (vs. longer and narrower;
<figureCitation id="135B2A33FFAAFF85C6B8FD85C66DFD9C" box="[245,376,573,597]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6M, N</figureCitation>
); less distribution of melanophores at kness (vs. dense;
<figureCitation id="135B2A33FFAAFF85C70FFDE5C6D4FDBC" box="[322,449,605,629]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6Q, R</figureCitation>
); males with reduced webbing on second to third fingers (vs. extensive web finger;
<figureCitation id="135B2A33FFAAFF85C49FFDC5C7DAFD7C" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6U, V</figureCitation>
).
</paragraph>
<caption id="DF1F663EFFAAFF85C50EFD14C227FD16" pageId="10" pageNumber="229" startId="10.[835,870,684,706]" targetBox="[835,1475,189,669]" targetPageId="10" targetType="figure">
<paragraph id="8BDF36B6FFAAFF85C50EFD14C227FD16" blockId="10.[835,1476,684,735]" pageId="10" pageNumber="229">
Fig. 7. Natural habitat of
<taxonomicName id="4C604D35FFAAFF85C20EFD14C22CFD08" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[1091,1337,684,705]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFAAFF85C20EFD14C22CFD08" box="[1091,1337,684,705]" italics="true" pageId="10" pageNumber="229">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C30AFD15C2D5FD0B" box="[1351,1472,684,706]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, at the type locality. Photograph by Fajar Kaprawi.
</paragraph>
</caption>
<paragraph id="8BDF36B6FFAAFF85C6C5FD65C6CFFC3C" blockId="10.[135,776,733,1013]" pageId="10" pageNumber="229">
<taxonomicName id="4C604D35FFAAFF85C6C5FD65C6BFFD3C" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[136,426,733,757]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFAAFF85C6C5FD65C6BFFD3C" box="[136,426,733,757]" italics="true" pageId="10" pageNumber="229">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C7F0FD66C55FFD3C" box="[445,586,733,757]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="4C604D35FFAAFF85C4BCFD66C627FCDC" baseAuthorityName="Harvey, Pemberton &amp; Smith" baseAuthorityYear="2002" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="achantharrhena">
<emphasis id="B914EAA4FFAAFF85C4BCFD66C627FCDC" italics="true" pageId="10" pageNumber="229">Z. achantharrhena</emphasis>
</taxonomicName>
by having a larger body size (SVL
<quantity id="4C989B53FFAAFF85C4D3FD45C7A4FCFC" metricMagnitude="-2" metricUnit="m" metricValue="3.915" metricValueMax="4.07" metricValueMin="3.76" pageId="10" pageNumber="229" unit="mm" value="39.15" valueMax="40.7" valueMin="37.6">37.640.7 mm</quantity>
vs.
<quantity id="4C989B53FFAAFF85C6A9FCA5C691FCFC" box="[228,388,797,821]" metricMagnitude="-2" metricUnit="m" metricValue="3.57" metricValueMax="3.79" metricValueMin="3.35" pageId="10" pageNumber="229" unit="mm" value="35.7" valueMax="37.9" valueMin="33.5">33.537.9 mm</quantity>
in males; SVL
<quantity id="4C989B53FFAAFF85C471FCA5C5C9FCFC" box="[572,732,797,821]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="5.45" metricValueMin="5.07" pageId="10" pageNumber="229" unit="mm" value="52.6" valueMax="54.5" valueMin="50.7">50.754.5 mm</quantity>
vs.
<quantity id="4C989B53FFAAFF85C6C5FC85C63CFC9C" box="[136,297,829,853]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" metricValueMax="4.46" metricValueMin="3.96" pageId="10" pageNumber="229" unit="mm" value="42.1" valueMax="44.6" valueMin="39.6">39.644.6 mm</quantity>
in females; see
<tableCitation id="C6E2030DFFAAFF85C7ABFC85C528FC9C" box="[486,573,829,853]" captionStart="Table 3" captionStartId="9.[111,166,525,548]" captionText="Table 3. Morphometric variation of selected characters of Zhangixalus achantharrhena and Zhangixalus dulitensis." pageId="10" pageNumber="229">Table 3</tableCitation>
); more developed webbing on fingers (vs. reduced;
<figureCitation id="135B2A33FFAAFF85C7B0FCE5C59FFCBC" box="[509,650,861,885]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6U, W</figureCitation>
); vocal sac opening absent (vs. present); dorsum with small spicules (vs. relatively large;
<figureCitation id="135B2A33FFAAFF85C775FC25C6A4FC7C" box="[312,433,925,949]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6I, K</figureCitation>
); less distributed melanophores on knees (vs. dense;
<figureCitation id="135B2A33FFAAFF85C734FC05C6E9FC1C" box="[377,508,957,981]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6Q, S</figureCitation>
); dark spots on dorsum scattered (vs. relatively dense).
</paragraph>
<paragraph id="8BDF36B6FFAAFF85C6C5FBA5C6FCFABC" blockId="10.[135,777,1053,1397]" pageId="10" pageNumber="229">
<taxonomicName id="4C604D35FFAAFF85C6C5FBA5C6BFFBFC" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" box="[136,426,1053,1077]" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFAAFF85C6C5FBA5C6BFFBFC" box="[136,426,1053,1077]" italics="true" pageId="10" pageNumber="229">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C7F0FBA6C55FFBFC" box="[445,586,1053,1077]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="4C604D35FFAAFF85C4BCFBA6C7FBFB9C" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="dulitensis">
<emphasis id="B914EAA4FFAAFF85C4BCFBA6C7FBFB9C" italics="true" pageId="10" pageNumber="229">Z. dulitensis</emphasis>
</taxonomicName>
by having a larger body size (SVL
<quantity id="4C989B53FFAAFF85C421FB85C413FB9C" box="[620,774,1085,1109]" metricMagnitude="-2" metricUnit="m" metricValue="3.915" metricValueMax="4.07" metricValueMin="3.76" pageId="10" pageNumber="229" unit="mm" value="39.15" valueMax="40.7" valueMin="37.6">37.640.7 mm</quantity>
vs.
<quantity id="4C989B53FFAAFF85C6E1FBE5C651FBBC" box="[172,324,1117,1141]" metricMagnitude="-2" metricUnit="m" metricValue="3.615" metricValueMax="3.87" metricValueMin="3.36" pageId="10" pageNumber="229" unit="mm" value="36.15" valueMax="38.7" valueMin="33.6">33.638.7 mm</quantity>
in males;
<quantity id="4C989B53FFAAFF85C7E3FBE5C553FBBC" box="[430,582,1117,1141]" metricMagnitude="-2" metricUnit="m" metricValue="5.26" metricValueMax="5.45" metricValueMin="5.07" pageId="10" pageNumber="229" unit="mm" value="52.6" valueMax="54.5" valueMin="50.7">50.754.5 mm</quantity>
vs. 46.0
<quantity id="4C989B53FFAAFF85C4E6FBE5C412FBBC" box="[683,775,1117,1141]" metricMagnitude="-2" metricUnit="m" metricValue="4.6899999999999995" pageId="10" pageNumber="229" unit="mm" value="46.9">46.9 mm</quantity>
in females; see
<tableCitation id="C6E2030DFFAAFF85C77FFBC5C693FB5C" box="[306,390,1149,1173]" captionStart="Table 3" captionStartId="9.[111,166,525,548]" captionText="Table 3. Morphometric variation of selected characters of Zhangixalus achantharrhena and Zhangixalus dulitensis." pageId="10" pageNumber="229">Table 3</tableCitation>
); web colouration between third to fifth toes red (vs. only between fourth to fifth toes); vocal sac opening absent (vs. present); shorter supracloaca dermal ridge (vs. longer;
<figureCitation id="135B2A33FFAAFF85C709FB65C6D6FB3C" box="[324,451,1245,1269]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6M, P</figureCitation>
); males snout slope with blunt edge in lateral view (vs. slope with sharply pointed edge;
<figureCitation id="135B2A33FFAAFF85C6C5FAA5C61FFAFC" box="[136,266,1309,1333]" captionStart="Fig" captionStartId="8.[135,170,1733,1755]" captionTargetBox="[138,1476,188,1717]" captionTargetId="figure-8@8.[120,1492,165,1728]" captionTargetPageId="8" captionText="Fig. 6. Comparison of morphological characters between male species of Zhangixalus from Sundaland. Scale bar = 5 mm. Photographs by Ade Damara Gonggoli." pageId="10" pageNumber="229">Figs. 6A, D</figureCitation>
); well-developed fringe of skin along the edge of the forearm and the outer edge of fifth to tibiotarsal articulation (vs. less developed).
</paragraph>
<paragraph id="8BDF36B6FFAAFF85C6C5FA25C495FC47" blockId="10.[135,777,1437,2101]" lastBlockId="10.[835,1474,790,910]" pageId="10" pageNumber="229">
<emphasis id="B914EAA4FFAAFF85C6C5FA25C6AFFA7C" bold="true" box="[136,442,1437,1461]" pageId="10" pageNumber="229">Range and natural history.</emphasis>
The type series of
<taxonomicName id="4C604D35FFAAFF85C4CEFA25C618FA1C" authorityName="Gonggoli &amp; &amp; &amp; &amp; Hamidy &amp; Amir" authorityYear="2024" class="Amphibia" family="Rhacophoridae" genus="Zhangixalus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="faritsalhadii" status="sp. nov.">
<emphasis id="B914EAA4FFAAFF85C4CEFA25C618FA1C" italics="true" pageId="10" pageNumber="229">Zhangixalus faritsalhadii</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A22757DFFFAAFF85C753FA06C6B0FA1C" box="[286,421,1469,1493]" pageId="10" pageNumber="229" rank="species">new species</taxonomicNameLabel>
, is so far known only from the highlands of Mount Slamet. The type series was found in Rasamala (
<taxonomicName id="4C604D35FFAAFF85C778FA45C6ECF9DC" box="[309,505,1533,1557]" class="Magnoliopsida" family="Altingiaceae" genus="Altingia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="10" pageNumber="229" phylum="Tracheophyta" rank="species" species="excelsa">
<emphasis id="B914EAA4FFAAFF85C778FA45C6ECF9DC" box="[309,505,1533,1557]" italics="true" pageId="10" pageNumber="229">Altingia excelsa</emphasis>
</taxonomicName>
) forest near swamps, approximately
<quantity id="4C989B53FFAAFF85C761F9A5C670F9FC" box="[300,357,1565,1589]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="229" unit="m" value="10.0">10 m</quantity>
from the main river (
<figureCitation id="135B2A33FFAAFF85C417F9A5C58AF9FC" box="[602,671,1565,1589]" captionStart="Fig" captionStartId="10.[835,870,684,706]" captionTargetBox="[835,1475,189,669]" captionTargetId="figure-26@10.[835,1475,189,669]" captionTargetPageId="10" captionText="Fig. 7. Natural habitat of Zhangixalus faritsalhadii, new species, at the type locality. Photograph by Fajar Kaprawi." pageId="10" pageNumber="229">Fig. 7</figureCitation>
). A male
<typeStatus id="54DB8814FFAAFF85C6C5F986C7F1F99F" box="[136,228,1598,1622]" pageId="10" pageNumber="229" type="paratype">paratype</typeStatus>
(MZB Amph 32890) was collected while it was calling from a palm tree, approximately
<quantity id="4C989B53FFAAFF85C47EF9E5C57CF9BC" box="[563,617,1629,1653]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="229" unit="m" value="15.0">15 m</quantity>
above ground, during rainy season in
<date id="FFDE1076FFAAFF85C730F9C5C508F95C" box="[381,541,1661,1685]" pageId="10" pageNumber="229" value="2020-02">February 2020</date>
. During the transition season from the dry to the rainy season in
<date id="FFDE1076FFAAFF85C429F925C5C7F97C" box="[612,722,1693,1717]" pageId="10" pageNumber="229" value="2022-07">July 2022</date>
, two pairs of amplectants of the type series were found on a rattan plant (MZB Amph 32886 and MZB Amph 32889;
<figureCitation id="135B2A33FFAAFF85C6C5F945C7C8F8DC" box="[136,221,1789,1813]" captionStart="Fig" captionStartId="6.[136,171,1098,1120]" captionTargetBox="[136,1474,189,1083]" captionTargetId="figure-8@6.[133,1477,184,1086]" captionTargetPageId="6" captionText="Fig. 4. A, Zhangixalus faritsalhadii, new species, male holotype in life, MZB Amph 32885; B, female paratype, MZB Amph 32887; C, MZB Amph 32889, female paratype in ventral view; D, MZB Amph 32886 and MZB Amph 32889 in amplexus. Photographs by Misbahul Munir (AC) and Fajar Kaprawi (D)." pageId="10" pageNumber="229">Fig. 4D</figureCitation>
) and on tree leaves (MZB Amph 32885 and MZB Amph 32888), while
<specimenCount id="9D66FD3FFFAAFF85C73AF8A6C6E5F8FC" box="[375,496,1821,1845]" count="1" pageId="10" pageNumber="229" type="female">one female</specimenCount>
(MZB Amph 32889) not far from the same locality was also found. The habitat was surrounded by small puddles formed from water running off the slope. This new species may be in other lowland areas of Mount Slamet. A record dated
<date id="FFDE1076FFAAFF85C791F825C588F87C" box="[476,669,1949,1973]" pageId="10" pageNumber="229" value="2020-02-18">18 February 2020</date>
appeared in a citizen science database at the exact location (
<bibRefCitation id="EFF14B47FFAAFF85C4E1F805C7D7F83C" author="Khafizh" pageId="10" pageNumber="229" refId="ref10530" refString="Khafizh (2020) iNaturalist: Zhangixalus sp. https: // www. inaturalist. org / observations / 38781391 (Accessed 5 November 2023)." type="url" year="2020">Khafizh, 2020</bibRefCitation>
). Tadpole and acoustic information are unknown. The following frog species are sympatric with the new species:
<taxonomicName id="4C604D35FFAAFF85C6C7F7A5C59AF7FC" authority="(Gravenhorst, 1829)" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[138,655,2077,2101]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="leucomystax">
<emphasis id="B914EAA4FFAAFF85C6C7F7A5C6B4F7FC" box="[138,417,2077,2101]" italics="true" pageId="10" pageNumber="229">Polypedates leucomystax</emphasis>
(
<bibRefCitation id="EFF14B47FFAAFF85C7FCF7A5C592F7FC" author="Gravenhorst JLC" box="[433,647,2077,2101]" pageId="10" pageNumber="229" refId="ref9834" refString="Gravenhorst JLC (1829) Deliciae Musei Zoologici Vratislaviensis. Fasciculus primus. Chelonios et Batrachia. Leipzig: Leopold Voss." type="book" year="1829">Gravenhorst, 1829</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C604D35FFAAFF85C4D3F7A5C38CFCE7" authority="(Schlegel, 1837)" baseAuthorityName="Schlegel" baseAuthorityYear="1837" box="[670,1177,790,2101]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="aurifasciatus">
<emphasis id="B914EAA4FFAAFF85C4D3F7A5C4C7FCE7" box="[670,978,790,2101]" italics="true" pageId="10" pageNumber="229">Philautus aurifasciatus</emphasis>
(
<bibRefCitation id="EFF14B47FFAAFF85C5A8FCAEC387FCE7" author="Schlegel H" box="[997,1170,790,814]" pageId="10" pageNumber="229" refId="ref11830" refString="Schlegel H (1837) Abbildungen neuer oder unvollstandig bekannter Amphibien, nach der Natur oder dem Leben entworfen, herausgegeben und mit einem erlauternden Texte begleitet. Arnz &amp; Comp, Dusseldorf, pl. 107." type="journal volume" year="1837">Schlegel, 1837</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C604D35FFAAFF85C2E4FCAFC4E2FC87" authority="(Boettger, 1892)" baseAuthorityName="Boettger" baseAuthorityYear="1892" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="microdiscus">
<emphasis id="B914EAA4FFAAFF85C2E4FCAFC2D7FCE7" box="[1193,1474,790,814]" italics="true" pageId="10" pageNumber="229">Limnonectes microdiscus</emphasis>
(
<bibRefCitation id="EFF14B47FFAAFF85C506FC8EC4FAFC87" author="Boettger O" box="[843,1007,822,846]" pageId="10" pageNumber="229" pagination="65 - 164" refId="ref8993" refString="Boettger O (1892) Listen von Kriechtieren und Lurchen aus dem tropischen Asien und aus Papuasien. Bericht des Offenbacher Vereins fur Naturkunde, 1892: 65 - 164." type="journal article" year="1892">Boettger, 1892</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="4C604D35FFAAFF85C24EFC8EC205FC87" baseAuthorityName="Schlegel" baseAuthorityYear="1840" box="[1027,1296,822,846]" class="Amphibia" family="Rhacophoridae" genus="Rhacophorus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="reinwardtii">
<emphasis id="B914EAA4FFAAFF85C24EFC8EC205FC87" box="[1027,1296,822,846]" italics="true" pageId="10" pageNumber="229">Rhacophorus reinwardtii</emphasis>
</taxonomicName>
;
<taxonomicName id="4C604D35FFAAFF85C352FC8EC353FCA7" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="hasseltii">
<emphasis id="B914EAA4FFAAFF85C352FC8EC489FCA7" italics="true" pageId="10" pageNumber="229">Leptobrachium hasseltii</emphasis>
<bibRefCitation id="EFF14B47FFAAFF85C5EBFCEEC353FCA7" author="Tschudi JJV" box="[934,1094,854,878]" pageId="10" pageNumber="229" refId="ref12265" refString="Tschudi JJV (1838) Classification der Batrachier mit Berucksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Petitpierre, Neuchatel, 99 pp." type="book" year="1838">Tschudi, 1838</bibRefCitation>
</taxonomicName>
; and
<taxonomicName id="4C604D35FFAAFF85C2C8FCEEC469FC47" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="10" pageNumber="229" phylum="Chordata" rank="species" species="achatina">
<emphasis id="B914EAA4FFAAFF85C2C8FCEEC24EFCA7" box="[1157,1371,854,878]" italics="true" pageId="10" pageNumber="229">Microhyla achatina</emphasis>
<bibRefCitation id="EFF14B47FFAAFF85C328FCEEC469FC47" author="Tschudi JJV" pageId="10" pageNumber="229" refId="ref12265" refString="Tschudi JJV (1838) Classification der Batrachier mit Berucksichtigung der fossilen Thiere dieser Abtheilung der Reptilien. Petitpierre, Neuchatel, 99 pp." type="book" year="1838">Tschudi, 1838</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,432 @@
<document id="84D87D3803B493FD41B3ACA6752E6DF3" ID-DOI="10.26107/RBZ-2024-0026" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683110" ID-ZooBank="B8F6DA4A-6776-43C8-9688-53589751DD44" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171913440" checkinUser="felipe" docAuthor="Tseng, Wei-Zhe, Cheng, &amp; Ren-Chung" docDate="2024" docId="03E087ADFF808B58ED3659C0FA44F7E4" docLanguage="en" docName="RafflesBZool.72.324-334.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Eucryptorrhynchus scrobiculatus" docType="treatment" docVersion="2" lastPageNumber="327" masterDocId="FFD9FFD5FF828B5BEE0D5E60FF85FFCE" masterDocTitle="Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)" masterLastPageNumber="334" masterPageNumber="324" pageNumber="326" updateTime="1737175838455" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A3B1C84E0E14149E8D0F1E2CA11A80B0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AB01193664B1761D12C833DD20598D0E">
<mods:title id="F56C980AF60583AC9E87F208C8908F1E">Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)</mods:title>
</mods:titleInfo>
<mods:name id="45C00DD28AB5055341F85A6FFD4494D0" type="personal">
<mods:role id="781CE382D17A9CBEB10CD880DA40A176">
<mods:roleTerm id="11A258F81E156174A89A20A9B0A287F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C73862B84FF13E053697DB97BAE20442">Tseng, Wei-Zhe</mods:namePart>
</mods:name>
<mods:name id="DB2DF35F8E48F000B64BF096793D0134" type="personal">
<mods:role id="A0C65421A8E034C27627C4631D3FE3CA">
<mods:roleTerm id="8BFC95BFF5C51D7A482C4C1FC86EE966">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2FC11C87591BCDFB113663C274B22A2">Cheng,</mods:namePart>
</mods:name>
<mods:name id="0E236A20E1246184058E858E1F2158BE" type="personal">
<mods:role id="D297730F04DB57C7885B5CD7DA5D37C7">
<mods:roleTerm id="FAB4FA5676A8BEBB8714E37C7C706C00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EEDF6C1C45A8538E86DBBE3E2B6DFD2E">Ren-Chung</mods:namePart>
</mods:name>
<mods:typeOfResource id="4889EC9CBB07375BCD51D119EADA16BC">text</mods:typeOfResource>
<mods:relatedItem id="58679BEBA5549B50F58681B1F87897EC" type="host">
<mods:titleInfo id="70355F51F099EFA6CBCBFBE1F8FC3B8D">
<mods:title id="AF7CB9F77EDBA9DA5E859C6747E3E2D2">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="BA34F4E34DCFF70E0D807FAD2AF2935D">
<mods:date id="0F481498F7E3B79A645315377F050492">2024</mods:date>
<mods:detail id="AC41E0CE8802F124B7D68231B13E2CB0" type="pubDate">
<mods:number id="4AAB90FB688F7CCCD1C890247CB91A6A">2024-09-20</mods:number>
</mods:detail>
<mods:detail id="CAF186D0EF9617477107DED5F24B8BE7" type="volume">
<mods:number id="4FCADDBEAB2D20EF081E12C8C5072055">72</mods:number>
</mods:detail>
<mods:extent id="24747D341A734C22EEF6C3F7B37CB599" unit="page">
<mods:start id="C74CC933F0BB30812FE502AC2AE67FB4">324</mods:start>
<mods:end id="5990034B59CB39DF33826293EED9ABE1">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="444FEC47F737B1F6D1E89E79A50AED8E">journal article</mods:classification>
<mods:identifier id="19D23CFD7840B42BBB4B1145A24827B7" type="DOI">10.26107/RBZ-2024-0026</mods:identifier>
<mods:identifier id="9A101B0CBECAB2FEBD12731B3ACA8141" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="47ECC48B7D4DD6F423E383C1255D974D" type="Zenodo-Dep">14683110</mods:identifier>
<mods:identifier id="2C4B4BE61AC5E7065E398DB8361D2CC3" type="ZooBank">B8F6DA4A-6776-43C8-9688-53589751DD44</mods:identifier>
</mods:mods>
<treatment id="03E087ADFF808B58ED3659C0FA44F7E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4" httpUri="http://treatment.plazi.org/id/03E087ADFF808B58ED3659C0FA44F7E4" lastPageId="3" lastPageNumber="327" pageId="2" pageNumber="326">
<subSubSection id="C3536530FF808B59ED3659C0FA18F876" box="[827,1437,1952,1976]" pageId="2" pageNumber="326" type="nomenclature">
<paragraph id="8BF636BBFF808B59ED3659C0FA18F876" blockId="2.[827,1437,1952,2009]" box="[827,1437,1952,1976]" pageId="2" pageNumber="326">
<heading id="D0BE81D7FF808B59ED3659C0FA18F876" bold="true" box="[827,1437,1952,1976]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="326" reason="2">
<taxonomicName id="4C494D38FF808B59ED3659C0FA18F876" ID-CoL="3BYMD" authority="(Motschulsky, 1854)" baseAuthorityName="Motschulsky" baseAuthorityYear="1854" box="[827,1437,1952,1976]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="326" phylum="Arthropoda" rank="species" species="scrobiculatus">
<emphasis id="B93DEAA9FF808B59ED3659C0FA18F876" bold="true" box="[827,1437,1952,1976]" pageId="2" pageNumber="326">
<emphasis id="B93DEAA9FF808B59ED3659C0FB28F876" bold="true" box="[827,1197,1952,1976]" italics="true" pageId="2" pageNumber="326">Eucryptorrhynchus scrobiculatus</emphasis>
(
<bibRefCitation id="EFD84B4AFF808B59EAB059C0FA13F876" author="Motschulsky V de" box="[1213,1430,1952,1976]" pageId="2" pageNumber="326" pagination="44 - 51" refId="ref7692" refString="Motschulsky V de (1854) Diagnoses de coleopteres nouveaux, trouves par M. M. Tatarinoff et Gaschkewitsch aux environs de Pekin. Etudes Entomologiques, 2: 44 - 51." type="journal article" year="1854">Motschulsky, 1854</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF808B59EDF659A0FB58F817" box="[1019,1245,1984,2009]" pageId="2" pageNumber="326" type="description">
<paragraph id="8BF636BBFF808B59EDF659A0FB58F817" blockId="2.[827,1437,1952,2009]" box="[1019,1245,1984,2009]" pageId="2" pageNumber="326">
(
<figureCitation id="13722A3EFF808B59EA0E59A0FB00F816" box="[1027,1157,1984,2009]" captionStart="Fig" captionStartId="3.[136,171,1352,1374]" captionTargetBox="[160,1445,215,1305]" captionTargetId="figure-48@3.[136,1474,189,1336]" captionTargetPageId="3" captionText="Fig. 1. Habitus of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, ventral views; E. brandti (Harold, 1880): C, dorsal and D, ventral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, ventral views." figureDoi="http://doi.org/10.5281/zenodo.14683112" httpUri="https://zenodo.org/record/14683112/files/figure.png" pageId="2" pageNumber="326">Figs. 1A, B</figureCitation>
,
<figureCitation id="13722A3EFF808B59EA9F59A1FB51F817" box="[1170,1236,1985,2009]" captionStart="Fig" captionStartId="4.[113,148,953,975]" captionTargetBox="[113,1451,189,938]" captionTargetId="figure-52@4.[113,1451,189,938]" captionTargetPageId="4" captionText="Fig. 2. Male genitalia of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, lateral views; E. brandti (Harold, 1880): C, dorsal and D, lateral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, lateral views." figureDoi="http://doi.org/10.5281/zenodo.14683114" httpUri="https://zenodo.org/record/14683114/files/figure.png" pageId="2" pageNumber="326">2A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF808B58ED205660FB0BF804" lastPageId="3" lastPageNumber="327" pageId="2" pageNumber="326" type="materials_examined">
<paragraph id="8BF636BBFF808B59ED205660FA2EF7F6" blockId="2.[812,1451,2048,2105]" pageId="2" pageNumber="326">
<emphasis id="B93DEAA9FF808B59ED205660FB0CF7D6" bold="true" box="[813,1161,2048,2072]" pageId="2" pageNumber="326">
Material examined.
<collectingCountry id="F35E762BFF808B59EA025660FB01F7D6" box="[1039,1156,2048,2072]" name="Taiwan" pageId="2" pageNumber="326">TAIWAN</collectingCountry>
:
</emphasis>
<specimenCount id="9D4FFD32FF808B59EA835661FB6FF7D6" box="[1166,1258,2048,2073]" count="1" pageId="2" pageNumber="326" type="female">1 female</specimenCount>
(ex. 15; accession number: PP697477) (
<collectionCode id="ED58AE7EFF808B59EA165641FBEAF7F7" box="[1051,1135,2081,2105]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="2" pageNumber="326">NMNS</collectionCode>
), Wuling Farm [
<emphasis id="B93DEAA9FF808B59EB225640FA1EF7F8" bold="true" box="[1327,1435,2080,2102]" pageId="2" pageNumber="326">武Dãae</emphasis>
],
</paragraph>
<caption id="DF366633FF818B58EE855B28FADBFAB4" ID-DOI="http://doi.org/10.5281/zenodo.14683112" ID-Zenodo-Dep="14683112" httpUri="https://zenodo.org/record/14683112/files/figure.png" pageId="3" pageNumber="327" startId="3.[136,171,1352,1374]" targetBox="[160,1445,215,1305]" targetPageId="3" targetType="figure">
<paragraph id="8BF636BBFF818B58EE855B28FADBFAB4" blockId="3.[136,1475,1352,1402]" pageId="3" pageNumber="327">
Fig. 1. Habitus of
<taxonomicName id="4C494D38FF818B58EF4E5B28FDEBFA93" box="[323,622,1352,1374]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="327" phylum="Arthropoda" rank="species" species="undefined-E">
<emphasis id="B93DEAA9FF818B58EF4E5B28FE78FA93" box="[323,509,1352,1373]" italics="true" pageId="3" pageNumber="327">Eucryptorrhynchus</emphasis>
species.
<emphasis id="B93DEAA9FF818B58EC515B28FDEBFA93" box="[604,622,1352,1373]" italics="true" pageId="3" pageNumber="327">E</emphasis>
</taxonomicName>
<emphasis id="B93DEAA9FF818B58EC605B28FD7EFA93" box="[621,763,1352,1373]" italics="true" pageId="3" pageNumber="327">. scrobiculatus</emphasis>
(
<bibRefCitation id="EFD84B4AFF818B58ED065B28FC4CFA90" author="Motschulsky V de" box="[779,969,1352,1374]" pageId="3" pageNumber="327" pagination="44 - 51" refId="ref7692" refString="Motschulsky V de (1854) Diagnoses de coleopteres nouveaux, trouves par M. M. Tatarinoff et Gaschkewitsch aux environs de Pekin. Etudes Entomologiques, 2: 44 - 51." type="journal article" year="1854">Motschulsky, 1854</bibRefCitation>
) from Taiwan: A, dorsal and B, ventral views;
<taxonomicName id="4C494D38FF818B58EBA05B28FEEFFAB4" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="327" phylum="Arthropoda" rank="species" species="brandti">
<emphasis id="B93DEAA9FF818B58EBA05B28FF4AFAB7" italics="true" pageId="3" pageNumber="327">E. brandti</emphasis>
(
<bibRefCitation id="EFD84B4AFF818B58EED05B04FEE6FAB4" author="Harold E von" box="[221,355,1380,1402]" pageId="3" pageNumber="327" pagination="148 - 171" refId="ref6994" refString="Harold E von (1880) Einige neue Coleopteren. Mittheilungen des Munchner Entomologischen Vereins, 4: 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
)
</taxonomicName>
: C, dorsal and D, ventral views;
<taxonomicName id="4C494D38FF818B58ECBF5B05FBA3FAB4" authority="Devi, Dey &amp; Ray, 2016" authorityName="Devi, Dey &amp; Ray" authorityYear="2016" box="[690,1062,1380,1402]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="327" phylum="Arthropoda" rank="species" species="khasiensis">
<emphasis id="B93DEAA9FF818B58ECBF5B05FCB4FAB7" box="[690,817,1380,1402]" italics="true" pageId="3" pageNumber="327">E. khasiensis</emphasis>
<bibRefCitation id="EFD84B4AFF818B58ED3A5B04FBA3FAB4" author="Devi SR &amp; Dey D &amp; Ray DC" box="[823,1062,1380,1402]" pageId="3" pageNumber="327" pagination="70 - 78" refId="ref6690" refString="Devi SR, Dey D &amp; Ray DC (2016) First record of genus Eucryptorrhynchus (Coleoptera: Curculionidae: Cryptorhynchinae) from India along with description of a new species. Oriental Insects, 51: 70 - 78." type="journal article" year="2016">Devi, Dey &amp; Ray, 2016</bibRefCitation>
</taxonomicName>
: E, dorsal and F, ventral views.
</paragraph>
</caption>
<paragraph id="8BF636BBFF818B58EE855BD2FB0BF804" blockId="3.[134,777,1458,2090]" lastBlockId="3.[834,1475,1458,1994]" pageId="3" pageNumber="327">
Heping District, Taichung City, coll. W-Z Tseng,
<date id="FFF7107BFF818B58ECB85BD2FF44FA27" pageId="3" pageNumber="327" value="2015-07-23">23 July 2015</date>
;
<materialsCitation id="3B213CE6FF818B58EEC05BB2FEF9F9E4" collectingDate="2018-08-11" collectionCode="NMNS" collectorName="Lai" county="Hsinchu County" latitude="24.682503" location="Jianshih Township" longLatPrecision="1" longitude="121.29563" municipality="Mamei" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D4FFD32FF818B58EEC05BB2FE92FA24" box="[205,279,1490,1514]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(
<collectionCode id="ED58AE7EFF818B58EF2B5BB2FEFDFA24" box="[294,376,1490,1514]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EF815BB2FE5DFA24" box="[396,472,1490,1514]" pageId="3" pageNumber="327">Mamei</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EFE45BB2FD9BFA26" bold="true" box="[489,542,1490,1512]" pageId="3" pageNumber="327">Ḅħ</emphasis>
],
<location id="8E966060FF818B58EC3B5BB2FC87FA24" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EC3B5BB2FC87FA24" box="[566,770,1490,1514]" county="Hsinchu County" latitude="24.682503" longLatPrecision="1" longitude="121.29563" municipality="Mamei" name="Jianshih Township" pageId="3" pageNumber="327">Jianshih Township</location>
,
<collectingCounty id="62974E37FF818B58EE875B92FECCF9C4" box="[138,329,1522,1546]" pageId="3" pageNumber="327">Hsinchu County</collectingCounty>
,
<geoCoordinate id="EE7D507CFF818B58EF545B91FE60F9C7" box="[345,485,1521,1545]" degrees="24.682502" direction="north" orientation="latitude" pageId="3" pageNumber="327" precision="1" value="24.682503">N24.682502</geoCoordinate>
<geoCoordinate id="EE7D507CFF818B58EFFC5B91FD09F9C7" box="[497,652,1521,1545]" degrees="121.295629" direction="east" orientation="longitude" pageId="3" pageNumber="327" precision="1" value="121.29563">E121.295629</geoCoordinate>
, coll. Y-T
<collectorName id="26BC536DFF818B58EE855872FF35F9E4" box="[136,176,1554,1578]" pageId="3" pageNumber="327">Lai</collectorName>
,
<date id="FFF7107BFF818B58EEB25872FEF9F9E4" box="[191,380,1554,1578]" pageId="3" pageNumber="327" value="2018-08-11">
<collectingDate id="EFB3E993FF818B58EEB25872FEF9F9E4" box="[191,380,1554,1578]" pageId="3" pageNumber="327" value="2018-08-11">11 August 2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EF865872FE97F944" collectingDate="2019-06-07" collectionCode="NMNS" collectorName="Ho" county="Newjiayang" latitude="24.247667" location="Taichung City" longLatPrecision="1" longitude="121.22869" municipality="Heping District" pageId="3" pageNumber="327" specimenCount="1">
<specimenCount id="9D4FFD32FF818B58EF865872FE40F9E7" box="[395,453,1554,1578]" count="1" pageId="3" pageNumber="327" type="generic">1 ex.</specimenCount>
(ex. 13; accession number: PP697493) (
<collectionCode id="ED58AE7EFF818B58EF075852FED9F984" box="[266,348,1586,1610]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingCounty id="62974E37FF818B58EF605852FE68F984" box="[365,493,1586,1610]" pageId="3" pageNumber="327">Newjiayang</collectingCounty>
[
<emphasis id="B93DEAA9FF818B58EFF75852FDCFF986" bold="true" box="[506,586,1586,1608]" pageId="3" pageNumber="327">Ǜ佳ọ</emphasis>
],
<collectingMunicipality id="6B92ACC1FF818B58EC535852FC86F984" box="[606,771,1586,1610]" pageId="3" pageNumber="327">Heping District</collectingMunicipality>
,
<location id="8E966060FF818B58EE8A5832FEADF9A4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EE8A5832FEADF9A4" box="[135,296,1618,1642]" county="Newjiayang" latitude="24.247667" longLatPrecision="1" longitude="121.22869" municipality="Heping District" name="Taichung City" pageId="3" pageNumber="327">Taichung City</location>
,
<geoCoordinate id="EE7D507CFF818B58EF3A5832FE38F9A4" box="[311,445,1618,1642]" degrees="24.247667" direction="north" orientation="latitude" pageId="3" pageNumber="327" precision="1" value="24.247667">N24.247667</geoCoordinate>
<geoCoordinate id="EE7D507CFF818B58EFC55832FDD9F9A4" box="[456,604,1618,1642]" degrees="121.228694" direction="east" orientation="longitude" pageId="3" pageNumber="327" precision="1" value="121.22869">E121.228694</geoCoordinate>
, coll. B-H
<collectorName id="26BC536DFF818B58ECD05832FC85F9A7" box="[733,768,1618,1641]" pageId="3" pageNumber="327">Ho</collectorName>
,
<date id="FFF7107BFF818B58EE855812FE97F944" box="[136,274,1650,1674]" pageId="3" pageNumber="327" value="2019-06-07">
<collectingDate id="EFB3E993FF818B58EE855812FE97F944" box="[136,274,1650,1674]" pageId="3" pageNumber="327" value="2019-06-07">7 June 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EF2D5812FE06F924" collectingDate="2019-06-07" collectionCode="NMNS" collectorName="Ho" county="Hualien County" latitude="24.1874" location="Xiulin Township" longLatPrecision="7" longitude="121.3395" municipality="Kuanyun Mountain" pageId="3" pageNumber="327" specimenCount="1">
<specimenCount id="9D4FFD32FF818B58EF2D5812FED2F947" box="[288,343,1650,1674]" count="1" pageId="3" pageNumber="327" type="generic">1 ex.</specimenCount>
(ex. 14; accession number: PP697494) (
<collectionCode id="ED58AE7EFF818B58EE9D58F2FF63F964" box="[144,230,1682,1706]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EEF058F2FE5EF964" box="[253,475,1682,1706]" pageId="3" pageNumber="327">Kuanyun Mountain</collectingMunicipality>
Villa [
<emphasis id="B93DEAA9FF818B58EC3E58F2FD26F966" bold="true" box="[563,675,1682,1704]" pageId="3" pageNumber="327">üṟ山¶</emphasis>
],
<location id="8E966060FF818B58ECB258F2FF7FF904" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ECB258F2FF7FF904" county="Hualien County" latitude="24.1874" longLatPrecision="7" longitude="121.3395" municipality="Kuanyun Mountain" name="Xiulin Township" pageId="3" pageNumber="327">Xiulin Township</location>
,
<collectingCounty id="62974E37FF818B58EF0758D2FE47F904" box="[266,450,1714,1738]" pageId="3" pageNumber="327">Hualien County</collectingCounty>
,
<geoCoordinate id="EE7D507CFF818B58EFDC58D2FDC5F904" box="[465,576,1714,1738]" degrees="24.1874" direction="north" orientation="latitude" pageId="3" pageNumber="327" precision="5" value="24.1874">N24.1874</geoCoordinate>
<geoCoordinate id="EE7D507CFF818B58EC4758D1FD42F907" box="[586,711,1713,1737]" degrees="121.3395" direction="east" orientation="longitude" pageId="3" pageNumber="327" precision="5" value="121.3395">E121.3395</geoCoordinate>
, coll. B-H
<collectorName id="26BC536DFF818B58EECC58B2FF60F927" box="[193,229,1746,1769]" pageId="3" pageNumber="327">Ho</collectorName>
,
<date id="FFF7107BFF818B58EEF958B2FE06F924" box="[244,387,1746,1770]" pageId="3" pageNumber="327" value="2019-06-07">
<collectingDate id="EFB3E993FF818B58EEF958B2FE06F924" box="[244,387,1746,1770]" pageId="3" pageNumber="327" value="2019-06-07">7 June 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EF9F58B2FE21F884" collectingDate="2019-08-06" collectionCode="NMNS" collectorName="Yeh" county="Yilan County" latitude="24.394508" location="Datung Township" longLatPrecision="1" longitude="121.36284" municipality="Sihyuan Wind Gap" pageId="3" pageNumber="327" specimenCount="3">
<specimenCount id="9D4FFD32FF818B58EF9F58B2FE4EF927" box="[402,459,1746,1770]" count="3" pageId="3" pageNumber="327" type="generic">3 ex.</specimenCount>
(ex. 12; accession number: PP697492) (
<collectionCode id="ED58AE7EFF818B58EF045892FEDFF8C4" box="[265,346,1778,1802]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EF675892FDB5F8C4" box="[362,560,1778,1802]" pageId="3" pageNumber="327">Sihyuan Wind Gap</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EC305892FD23F8C6" bold="true" box="[573,678,1778,1800]" pageId="3" pageNumber="327">思fi埡口</emphasis>
],
<location id="8E966060FF818B58ECB75892FF71F8E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ECB75892FF71F8E4" county="Yilan County" latitude="24.394508" longLatPrecision="1" longitude="121.36284" municipality="Sihyuan Wind Gap" name="Datung Township" pageId="3" pageNumber="327">Datung Township</location>
,
<collectingCounty id="62974E37FF818B58EF0F5972FE1CF8E4" box="[258,409,1810,1834]" pageId="3" pageNumber="327">Yilan County</collectingCounty>
,
<geoCoordinate id="EE7D507CFF818B58EFAA5971FDA9F8E7" box="[423,556,1809,1833]" degrees="24.394508" direction="north" orientation="latitude" pageId="3" pageNumber="327" precision="1" value="24.394508">N24.394508</geoCoordinate>
<geoCoordinate id="EE7D507CFF818B58EC3B5972FD4CF8E4" box="[566,713,1810,1834]" degrees="121.362836" direction="east" orientation="longitude" pageId="3" pageNumber="327" precision="1" value="121.36284">E121.362836</geoCoordinate>
, coll. L-W
<collectorName id="26BC536DFF818B58EECE5952FF77F884" box="[195,242,1842,1866]" pageId="3" pageNumber="327">Yeh</collectorName>
,
<date id="FFF7107BFF818B58EEF25952FE21F884" box="[255,420,1842,1866]" pageId="3" pageNumber="327" value="2019-08-06">
<collectingDate id="EFB3E993FF818B58EEF25952FE21F884" box="[255,420,1842,1866]" pageId="3" pageNumber="327" value="2019-08-06">6 August 2019</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B213CE6FF818B58EFBF5951FE20F867" collectingDate="2015-08-04" collectionCode="TARI" country="China" county="Rongshui Miao Autonomous County" location="Liuzhou" municipality="Yangmeiao" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi">
<emphasis id="B93DEAA9FF818B58EFBF5951FD00F887" bold="true" box="[434,645,1841,1865]" pageId="3" pageNumber="327">
<collectingCountry id="F35E762BFF818B58EFBF5951FD89F887" box="[434,524,1841,1865]" name="China" pageId="3" pageNumber="327">CHINA</collectingCountry>
,
<collectingRegion id="498DF859FF818B58EC175951FD05F887" box="[538,640,1841,1865]" country="China" name="Guangxi" pageId="3" pageNumber="327">Guangxi</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF818B58EC865952FD50F884" box="[651,725,1842,1866]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(ex. 1; accession number: PP692124) (
<collectionCode id="ED58AE7EFF818B58EC0B5932FDCFF8A4" box="[518,586,1874,1898]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EC6D5932FD5EF8A4" box="[608,731,1874,1898]" pageId="3" pageNumber="327">Yangmeiao</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58ECE15932FF38F846" bold="true" pageId="3" pageNumber="327">杨 梅坳</emphasis>
],
<collectingCounty id="62974E37FF818B58EED85912FDE0F844" box="[213,613,1906,1930]" pageId="3" pageNumber="327">Rongshui Miao Autonomous County</collectingCounty>
,
<location id="8E966060FF818B58EC7C5912FD48F844" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EC7C5912FD48F844" box="[625,717,1906,1930]" country="China" county="Rongshui Miao Autonomous County" municipality="Yangmeiao" name="Liuzhou" pageId="3" pageNumber="327" stateProvince="Guangxi">Liuzhou</location>
, coll. Y-T
<location id="8E966060FF818B58EEB659F2FF78F864" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EEB659F2FF78F864" box="[187,253,1938,1962]" country="China" county="Rongshui Miao Autonomous County" municipality="Yangmeiao" name="Wang" pageId="3" pageNumber="327" stateProvince="Guangxi">Wang</location>
,
<date id="FFF7107BFF818B58EF0A59F2FE20F867" box="[263,421,1937,1962]" pageId="3" pageNumber="327" value="2015-08-04">
<collectingDate id="EFB3E993FF818B58EF0A59F2FE20F867" box="[263,421,1937,1962]" pageId="3" pageNumber="327" value="2015-08-04">4 August 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EFA259F2FDC6F827" collectingDate="2015-08-17" collectionCode="TARI" country="China" county="Xing'an County" location="Guilin" municipality="Maoershan" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi">
<specimenCount id="9D4FFD32FF818B58EFA259F2FE70F864" box="[431,501,1938,1962]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(ex. 3; accession number: PP697478) (
<collectionCode id="ED58AE7EFF818B58EF0059D2FED5F804" box="[269,336,1970,1994]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EF6E59D2FE5CF804" box="[355,473,1970,1994]" pageId="3" pageNumber="327">Maoershan</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EFE659D2FDBEF806" bold="true" box="[491,571,1970,1992]" pageId="3" pageNumber="327">Ê儿山</emphasis>
],
<collectingCounty id="62974E37FF818B58EC5E59D2FC86F804" box="[595,771,1970,1994]" pageId="3" pageNumber="327">Xingan County</collectingCounty>
,
<location id="8E966060FF818B58EE8459B2FF55F824" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EE8459B2FF55F824" box="[137,208,2002,2026]" country="China" county="Xing'an County" municipality="Maoershan" name="Guilin" pageId="3" pageNumber="327" stateProvince="Guangxi">Guilin</location>
, coll. Y-T
<location id="8E966060FF818B58EF4859B2FE02F824" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EF4859B2FE02F824" box="[325,391,2002,2026]" country="China" county="Xing'an County" municipality="Maoershan" name="Wang" pageId="3" pageNumber="327" stateProvince="Guangxi">Wang</location>
,
<date id="FFF7107BFF818B58EF9E59B2FDC6F827" box="[403,579,2001,2026]" pageId="3" pageNumber="327" value="2015-08-17">
<collectingDate id="EFB3E993FF818B58EF9E59B2FDC6F827" box="[403,579,2001,2026]" pageId="3" pageNumber="327" value="2015-08-17">17 August 2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B213CE6FF818B58EC4059B1FB58FA04" collectingDate="2016-07-23" collectionCode="TARI" country="China" county="Miao Autonomous County" location="Guanling Buyei" municipality="Huixiangping" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="B93DEAA9FF818B58EC4059B1FD3DF827" bold="true" box="[589,696,2001,2025]" pageId="3" pageNumber="327">
<collectingRegion id="498DF859FF818B58EC4059B1FD36F827" box="[589,691,2001,2025]" country="China" name="Guizhou" pageId="3" pageNumber="327">Guizhou</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF818B58ECB359B2FC82F824" box="[702,775,2002,2026]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(ex. 4; accession number: PP697479) (
<collectionCode id="ED58AE7EFF818B58EC2D5992FDE6F7C4" box="[544,611,2034,2058]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EC785992FC8DF7C4" box="[629,776,2034,2058]" pageId="3" pageNumber="327">Huixiangping</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EE9C5672FF64F7E6" bold="true" box="[145,225,2066,2088]" pageId="3" pageNumber="327">回⁂坪</emphasis>
],
<location id="8E966060FF818B58EEF45672FE23F7E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EEF45672FE23F7E4" box="[249,422,2066,2090]" country="China" county="Miao Autonomous County" municipality="Huixiangping" name="Guanling Buyei" pageId="3" pageNumber="327" stateProvince="Guizhou">Guanling Buyei</location>
and
<collectingCounty id="62974E37FF818B58EFD35672FC86F7E4" box="[478,771,2066,2090]" pageId="3" pageNumber="327">Miao Autonomous County</collectingCounty>
,
<location id="8E966060FF818B58ED4E5BD2FC1DFA04" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ED4E5BD2FC1DFA04" box="[835,920,1458,1482]" country="China" county="Miao Autonomous County" municipality="Huixiangping" name="Anshun" pageId="3" pageNumber="327" stateProvince="Guizhou">Anshun</location>
, coll. Y-T
<location id="8E966060FF818B58EA0A5BD2FBCCFA04" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EA0A5BD2FBCCFA04" box="[1031,1097,1458,1482]" country="China" county="Miao Autonomous County" municipality="Huixiangping" name="Wang" pageId="3" pageNumber="327" stateProvince="Guizhou">Wang</location>
,
<date id="FFF7107BFF818B58EA5F5BD2FB58FA04" box="[1106,1245,1458,1482]" pageId="3" pageNumber="327" value="2016-07-23">
<collectingDate id="EFB3E993FF818B58EA5F5BD2FB58FA04" box="[1106,1245,1458,1482]" pageId="3" pageNumber="327" value="2016-07-23">23 July 2016</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B213CE6FF818B58EAEE5BD2FA3AF9E4" collectingDate="2016-06-10" collectionCode="TARI" country="China" county="Autonomous County" location="Gongshan" municipality="Sengang" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<emphasis id="B93DEAA9FF818B58EAEE5BD2FACDFA04" bold="true" box="[1251,1352,1458,1482]" pageId="3" pageNumber="327">
<collectingRegion id="498DF859FF818B58EAEE5BD2FAC6FA04" box="[1251,1347,1458,1482]" country="China" name="Yunnan" pageId="3" pageNumber="327">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF818B58EB435BD2FA10FA04" box="[1358,1429,1458,1482]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(ex. 9; accession number: PP697480) (
<collectionCode id="ED58AE7EFF818B58EABA5BB2FB7FFA24" box="[1207,1274,1490,1514]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EB035BB2FAEFFA24" box="[1294,1386,1490,1514]" pageId="3" pageNumber="327">Sengang</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EB705BB2FA37FA26" bold="true" box="[1405,1458,1490,1512]" pageId="3" pageNumber="327">AEDz</emphasis>
],
<location id="8E966060FF818B58ED4E5B92FC2AF9C4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ED4E5B92FC2AF9C4" box="[835,943,1522,1546]" country="China" county="Autonomous County" municipality="Sengang" name="Gongshan" pageId="3" pageNumber="327" stateProvince="Yunnan">Gongshan</location>
Derung-Nu
<collectingCounty id="62974E37FF818B58EA355B92FA9FF9C4" box="[1080,1306,1522,1546]" pageId="3" pageNumber="327">Autonomous County</collectingCounty>
,
<location id="8E966060FF818B58EB2B5B92FAF9F9C4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EB2B5B92FAF9F9C4" box="[1318,1404,1522,1546]" country="China" county="Autonomous County" municipality="Sengang" name="Nujiang" pageId="3" pageNumber="327" stateProvince="Yunnan">Nujiang</location>
of the
<location id="8E966060FF818B58ED4E5872FBF3F9E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ED4E5872FBF3F9E4" box="[835,1142,1554,1578]" country="China" county="Autonomous County" municipality="Sengang" name="Lisu Autonomous Prefecture" pageId="3" pageNumber="327" stateProvince="Yunnan">Lisu Autonomous Prefecture</location>
, coll. Y-T
<location id="8E966060FF818B58EAE95872FAA3F9E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EAE95872FAA3F9E4" box="[1252,1318,1554,1578]" country="China" county="Autonomous County" municipality="Sengang" name="Wang" pageId="3" pageNumber="327" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF818B58EB3D5872FA3AF9E4" box="[1328,1471,1554,1578]" pageId="3" pageNumber="327" value="2016-06-10">
<collectingDate id="EFB3E993FF818B58EB3D5872FA3AF9E4" box="[1328,1471,1554,1578]" pageId="3" pageNumber="327" value="2016-06-10">10 June 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58ED4F5852FBE7F964" collectingDate="2014-06-12" collectionCode="TARI" country="China" county="Autonomous County" location="Gongshan" municipality="Naqialuo" pageId="3" pageNumber="327" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D4FFD32FF818B58ED4F5852FC21F984" box="[834,932,1586,1610]" count="1" pageId="3" pageNumber="327" type="female">1 female</specimenCount>
(ex. 11; accession number: PP697481) (
<collectionCode id="ED58AE7EFF818B58EB7D5852FA31F984" box="[1392,1460,1586,1610]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58ED4E5832FC2CF9A4" box="[835,937,1618,1642]" pageId="3" pageNumber="327">Naqialuo</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EDB35832FB94F9A6" bold="true" box="[958,1041,1618,1640]" pageId="3" pageNumber="327">DZ恰洛</emphasis>
],
<location id="8E966060FF818B58EA205832FB18F9A4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EA205832FB18F9A4" box="[1069,1181,1618,1642]" country="China" county="Autonomous County" municipality="Naqialuo" name="Gongshan" pageId="3" pageNumber="327" stateProvince="Yunnan">Gongshan</location>
Derung-Nu
<collectingCounty id="62974E37FF818B58EB3F5832FC11F944" pageId="3" pageNumber="327">Autonomous County</collectingCounty>
,
<location id="8E966060FF818B58EDAF5812FC7DF944" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EDAF5812FC7DF944" box="[930,1016,1650,1674]" country="China" county="Autonomous County" municipality="Naqialuo" name="Nujiang" pageId="3" pageNumber="327" stateProvince="Yunnan">Nujiang</location>
of the
<location id="8E966060FF818B58EA475812FA03F944" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EA475812FA03F944" box="[1098,1414,1650,1674]" country="China" county="Autonomous County" municipality="Naqialuo" name="Lisu Autonomous Prefecture" pageId="3" pageNumber="327" stateProvince="Yunnan">Lisu Autonomous Prefecture</location>
, coll. Y-T
<location id="8E966060FF818B58ED7758F2FC39F964" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ED7758F2FC39F964" box="[890,956,1682,1706]" country="China" county="Autonomous County" municipality="Naqialuo" name="Wang" pageId="3" pageNumber="327" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF818B58EDC758F2FBE7F964" box="[970,1122,1682,1706]" pageId="3" pageNumber="327" value="2014-06-12">
<collectingDate id="EFB3E993FF818B58EDC758F2FBE7F964" box="[970,1122,1682,1706]" pageId="3" pageNumber="327" value="2014-06-12">12 June 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EA7D58F2FA45F8E4" collectingDate="2016-06-15" collectionCode="TARI" country="China" county="Autonomous County" location="Gongshan" municipality="Nageluo" pageId="3" pageNumber="327" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Yunnan">
<specimenCount id="9D4FFD32FF818B58EA7D58F2FB4DF964" box="[1136,1224,1682,1706]" count="4" pageId="3" pageNumber="327" type="male">4 males</specimenCount>
,
<specimenCount id="9D4FFD32FF818B58EADB58F2FAB3F964" box="[1238,1334,1682,1706]" count="1" pageId="3" pageNumber="327" type="female">1 female</specimenCount>
(ex. 5, 6, 7, 8, 10; accession number: PP697485, PP697486, PP697484, PP697483, PP697482) (
<collectionCode id="ED58AE7EFF818B58EA3158B2FBFBF924" box="[1084,1150,1746,1770]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="3" pageNumber="327">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF818B58EA8258B2FB6DF924" box="[1167,1256,1746,1770]" pageId="3" pageNumber="327">Nageluo</collectingMunicipality>
[
<emphasis id="B93DEAA9FF818B58EAF858B2FAC1F926" bold="true" box="[1269,1348,1746,1768]" pageId="3" pageNumber="327">Oi格洛</emphasis>
],
<location id="8E966060FF818B58EB5A58B2FA47F924" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EB5A58B2FA47F924" box="[1367,1474,1746,1770]" country="China" county="Autonomous County" municipality="Nageluo" name="Gongshan" pageId="3" pageNumber="327" stateProvince="Yunnan">Gongshan</location>
Derung-Nu
<collectingCounty id="62974E37FF818B58EDC25892FB40F8C4" box="[975,1221,1778,1802]" pageId="3" pageNumber="327">Autonomous County</collectingCounty>
,
<location id="8E966060FF818B58EAD85892FAB4F8C4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EAD85892FAB4F8C4" box="[1237,1329,1778,1802]" country="China" county="Autonomous County" municipality="Nageluo" name="Nujiang" pageId="3" pageNumber="327" stateProvince="Yunnan">Nujiang</location>
of the
<location id="8E966060FF818B58EB835892FBD5F8E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EB835892FBD5F8E4" country="China" county="Autonomous County" municipality="Nageluo" name="Lisu Autonomous Prefecture" pageId="3" pageNumber="327" stateProvince="Yunnan">Lisu Autonomous Prefecture</location>
, coll. Y-T
<location id="8E966060FF818B58EADC5972FA90F8E4" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58EADC5972FA90F8E4" box="[1233,1301,1810,1834]" country="China" county="Autonomous County" municipality="Nageluo" name="Wang" pageId="3" pageNumber="327" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF818B58EB2E5971FA45F8E4" box="[1315,1472,1809,1834]" pageId="3" pageNumber="327" value="2016-06-15">
<collectingDate id="EFB3E993FF818B58EB2E5971FA45F8E4" box="[1315,1472,1809,1834]" pageId="3" pageNumber="327" value="2016-06-15">15 June 2016</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B213CE6FF818B58ED4E5951FBDAF8A4" collectingDate="2017-06" collectionCode="NMNS" country="Vietnam" location="Ha Giang Province" pageId="3" pageNumber="327" specimenCount="1" specimenCount-male="1" stateProvince="Ha Giang">
<emphasis id="B93DEAA9FF818B58ED4E5951FC55F887" bold="true" box="[835,976,1841,1865]" pageId="3" pageNumber="327">
<collectingCountry id="F35E762BFF818B58ED4E5951FC4EF887" box="[835,971,1841,1865]" name="Vietnam" pageId="3" pageNumber="327">VIETNAM</collectingCountry>
:
</emphasis>
<specimenCount id="9D4FFD32FF818B58EDD65952FBADF884" box="[987,1064,1842,1866]" count="1" pageId="3" pageNumber="327" type="male">1 male</specimenCount>
(
<collectionCode id="ED58AE7EFF818B58EA365952FB0AF884" box="[1083,1167,1842,1866]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingRegion id="498DF859FF818B58EAAB5952FA06F884" box="[1190,1411,1842,1866]" country="Vietnam" name="Ha Giang" pageId="3" pageNumber="327">Hà Giang Province</collectingRegion>
, coll. local collector,
<date id="FFF7107BFF818B58EDE05932FBDAF8A4" box="[1005,1119,1874,1898]" pageId="3" pageNumber="327" value="2017-06">
<collectingDate id="EFB3E993FF818B58EDE05932FBDAF8A4" box="[1005,1119,1874,1898]" pageId="3" pageNumber="327" value="2017-06">June 2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF818B58EA615932FB0EF804" collectingDate="2019-04-10" collectionCode="NMNS" collectorName="S. Alekseenko" country="Vietnam" county="Phan Xi Pang Mountain" latitude="22.349443" location="District" longLatPrecision="1" longitude="103.770836" municipality="Tam Du" pageId="3" pageNumber="327" specimenCount="1" stateProvince="Lai Chau">
<specimenCount id="9D4FFD32FF818B58EA615932FB26F8A7" box="[1132,1187,1874,1898]" count="1" pageId="3" pageNumber="327" type="generic">1 ex.</specimenCount>
(ex. 2; accession number: PP697495) (
<collectionCode id="ED58AE7EFF818B58EDC65912FB98F844" box="[971,1053,1906,1930]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="3" pageNumber="327">NMNS</collectionCode>
),
<collectingCounty id="62974E37FF818B58EA3D5912FAB3F844" box="[1072,1334,1906,1930]" pageId="3" pageNumber="327">Phan Xi Păng Mountain</collectingCounty>
,
<collectingMunicipality id="6B92ACC1FF818B58EB4C5912FA1FF844" box="[1345,1434,1906,1930]" pageId="3" pageNumber="327">Tam Đư</collectingMunicipality>
ờng
<location id="8E966060FF818B58ED4E59F2FC13F864" LSID="urn:lsid:plazi:treatment:03E087ADFF808B58ED3659C0FA44F7E4:8E966060FF818B58ED4E59F2FC13F864" box="[835,918,1938,1962]" country="Vietnam" county="Phan Xi Pang Mountain" latitude="22.349443" longLatPrecision="1" longitude="103.770836" municipality="Tam Du" name="District" pageId="3" pageNumber="327" stateProvince="Lai Chau">District</location>
,
<collectingRegion id="498DF859FF818B58ED9259F2FBE3F864" box="[927,1126,1938,1962]" country="Vietnam" name="Lai Chau" pageId="3" pageNumber="327">Lai Châu Province</collectingRegion>
,
<geoCoordinate id="EE7D507CFF818B58EA7D59F2FB71F864" box="[1136,1268,1938,1962]" degrees="22.349444" direction="north" orientation="latitude" pageId="3" pageNumber="327" precision="1" value="22.349443">N22.349444</geoCoordinate>
<geoCoordinate id="EE7D507CFF818B58EAF459F2FA0EF864" box="[1273,1419,1938,1962]" degrees="103.770833" direction="east" orientation="longitude" pageId="3" pageNumber="327" precision="1" value="103.770836">E103.770833</geoCoordinate>
, coll.
<collectorName id="26BC536DFF818B58ED4F59D2FC64F804" box="[834,993,1970,1994]" pageId="3" pageNumber="327">S. Alekseenko</collectorName>
,
<date id="FFF7107BFF818B58EDE059D2FB0EF804" box="[1005,1163,1970,1994]" pageId="3" pageNumber="327" value="2019-04-10">
<collectingDate id="EFB3E993FF818B58EDE059D2FB0EF804" box="[1005,1163,1970,1994]" pageId="3" pageNumber="327" value="2019-04-10">10 April 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF818B58ED4E5991FA44F7E4" pageId="3" pageNumber="327" type="distribution">
<paragraph id="8BF636BBFF818B58ED4E5991FA44F7E4" blockId="3.[835,1474,2033,2090]" pageId="3" pageNumber="327">
<emphasis id="B93DEAA9FF818B58ED4E5991FC54F7C7" bold="true" box="[835,977,2033,2057]" pageId="3" pageNumber="327">Distribution.</emphasis>
<collectingCountry id="F35E762BFF818B58EDDB5992FB91F7C4" box="[982,1044,2034,2058]" name="China" pageId="3" pageNumber="327">China</collectingCountry>
(
<collectingRegion id="498DF859FF818B58EA2C5992FBE3F7C4" box="[1057,1126,2034,2058]" country="China" name="Anhui" pageId="3" pageNumber="327">Anhui</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EA635992FB38F7C4" box="[1134,1213,2034,2058]" country="China" name="Beijing" pageId="3" pageNumber="327">Beijing</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EACB5992FA8EF7C4" box="[1222,1291,2034,2058]" country="China" name="Fujian" pageId="3" pageNumber="327">Fujian</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EB195992FADCF7C4" box="[1300,1369,2034,2058]" country="China" name="Gansu" pageId="3" pageNumber="327">Gansu</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EB6C5992FA3BF7C4" box="[1377,1470,2034,2058]" country="China" name="Guizhou" pageId="3" pageNumber="327">Guizhou</collectingRegion>
,
<collectingRegion id="498DF859FF818B58ED4E5672FC00F7E4" box="[835,901,2066,2090]" country="China" name="Hebei" pageId="3" pageNumber="327">Hebei</collectingRegion>
,
<collectingRegion id="498DF859FF818B58ED9E5672FC5EF7E7" box="[915,987,2066,2089]" country="China" name="Henan" pageId="3" pageNumber="327">Henan</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EDE75672FBABF7E4" box="[1002,1070,2066,2090]" country="China" name="Hubei" pageId="3" pageNumber="327">Hubei</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EA365672FB00F7E7" box="[1083,1157,2066,2089]" country="China" name="Hunan" pageId="3" pageNumber="327">Hunan</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EA9E5672FB63F7E4" box="[1171,1254,2066,2090]" country="China" name="Jiangsu" pageId="3" pageNumber="327">Jiangsu</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EAF95672FAD2F7E4" box="[1268,1367,2066,2090]" country="China" name="Liaoning" pageId="3" pageNumber="327">Liaoning</collectingRegion>
,
<collectingRegion id="498DF859FF818B58EB685672FA3BF7E4" box="[1381,1470,2066,2090]" country="China" name="Qinghai" pageId="3" pageNumber="327">Qinghai</collectingRegion>
,
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,73 @@
<document id="456420BC4FF1BB10F7560E29BBB75E1F" ID-DOI="10.26107/RBZ-2024-0026" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683110" ID-ZooBank="B8F6DA4A-6776-43C8-9688-53589751DD44" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171913440" checkinUser="felipe" docAuthor="Tseng, Wei-Zhe, Cheng, &amp; Ren-Chung" docDate="2024" docId="03E087ADFF868B5FED205963FA2FF85E" docLanguage="en" docName="RafflesBZool.72.324-334.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Eucryptorrhynchus" docType="key" docVersion="2" lastPageNumber="328" masterDocId="FFD9FFD5FF828B5BEE0D5E60FF85FFCE" masterDocTitle="Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)" masterLastPageNumber="334" masterPageNumber="324" pageNumber="328" updateTime="1737175838455" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="32E7E006D1F9444441B627EC8A24EA67" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E0C3EBF262CF9DD5D6FB87EF4207464C">
<mods:title id="CF134AAF4BD85AB8BDA9C2FE702F90EB">Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)</mods:title>
</mods:titleInfo>
<mods:name id="7900FE6C131D23ECB921A896AE5245AA" type="personal">
<mods:role id="1D1E542572840FFE5FCA30AE67BFEA53">
<mods:roleTerm id="E2E4E0960E3D3D4E9E6D02081B319863">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="422FACA53F7C471E945FA473666528C9">Tseng, Wei-Zhe</mods:namePart>
</mods:name>
<mods:name id="693638BE11DD5DB4BF75BC8D458BBD7F" type="personal">
<mods:role id="DC5CACDE4302C23EA1D400FF75B6A0B7">
<mods:roleTerm id="B2C7C15BAD5627206722F33FD6CAF586">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1ADAAA507FDFDEC91AFC84BFEAD8249">Cheng,</mods:namePart>
</mods:name>
<mods:name id="7FB198764F580C2C638357EBECA5F129" type="personal">
<mods:role id="08D327527AB1B8AF8A825B7EF1243F7B">
<mods:roleTerm id="72C379325CED631C9210BF466B84B8EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D09588BEAF52BFDBBFCB4BC0F459C7A3">Ren-Chung</mods:namePart>
</mods:name>
<mods:typeOfResource id="5C30B07F4ED2A2B64C6B7690E86F16D6">text</mods:typeOfResource>
<mods:relatedItem id="7AF24DE1A55DDAAA5CC7BCEA0D8D0182" type="host">
<mods:titleInfo id="BA64DDEE1221C1DBBA8D839D379A0B28">
<mods:title id="10F3F01EE4D1EC980F3F1B1E69B92CDE">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="6613D1A8294CF2E64F6A3038EC7E150D">
<mods:date id="C435E2281239A8051741A44137501498">2024</mods:date>
<mods:detail id="9E7C188B4A7833D2F50481F6588C585E" type="pubDate">
<mods:number id="43225D92D6846A1800D6865FBD1A3A4A">2024-09-20</mods:number>
</mods:detail>
<mods:detail id="EAE10768781A6C19ACFD7F63D56CE95C" type="volume">
<mods:number id="D4E8F004B7BC18E86CA493E61B246468">72</mods:number>
</mods:detail>
<mods:extent id="862C566BD0FA1165AA94639867D9C774" unit="page">
<mods:start id="9215F90ABF851CE98C89CED462BD37E9">324</mods:start>
<mods:end id="49D59DA0947D8BFD2B142135080B500B">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4EF789950C7BCE9E39274BBEF291C66C">journal article</mods:classification>
<mods:identifier id="4CEC439F184AD3F691C2D5BC1CB83244" type="DOI">10.26107/RBZ-2024-0026</mods:identifier>
<mods:identifier id="0CA15615AB58F188C5662DC143D139E9" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="AC66AF5DAC3F002C8C330E57F5A42CD2" type="Zenodo-Dep">14683110</mods:identifier>
<mods:identifier id="05B3AC3F54FE1566F4E9BC55956E06DF" type="ZooBank">B8F6DA4A-6776-43C8-9688-53589751DD44</mods:identifier>
</mods:mods>
<treatment id="03E087ADFF868B5FED205963FA2FF85E" ID-DOI="http://doi.org/10.5281/zenodo.14683136" ID-Zenodo-Dep="14683136" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FED205963FA2FF85E" httpUri="http://treatment.plazi.org/id/03E087ADFF868B5FED205963FA2FF85E" lastPageNumber="328" pageId="4" pageNumber="328">
<subSubSection id="C3536530FF868B5FED205963FA24F8D5" box="[813,1441,1795,1819]" pageId="4" pageNumber="328" type="nomenclature">
<paragraph id="8BF636BBFF868B5FED205963FA24F8D5" blockId="4.[813,1441,1795,1819]" box="[813,1441,1795,1819]" pageId="4" pageNumber="328">
<heading id="D0BE81D7FF868B5FED205963FA24F8D5" bold="true" box="[813,1441,1795,1819]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="328" reason="2">
<emphasis id="B93DEAA9FF868B5FED205963FA24F8D5" bold="true" box="[813,1441,1795,1819]" pageId="4" pageNumber="328">
Key to the species of
<taxonomicName id="4C494D38FF868B5FEA2B5963FA24F8D5" ID-CoL="8HJGJ" authority="(Heller, 1937)" baseAuthorityName="Heller" baseAuthorityYear="1937" box="[1062,1441,1795,1819]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B93DEAA9FF868B5FEA2B5963FB78F8D5" bold="true" box="[1062,1277,1795,1819]" italics="true" pageId="4" pageNumber="328">Eucryptorrhynchus</emphasis>
(
<bibRefCitation id="EFD84B4AFF868B5FEB065963FA1FF8D5" author="Heller KM" box="[1291,1434,1795,1819]" pageId="4" pageNumber="328" pagination="51 - 78" refId="ref7096" refString="Heller KM (1937) Coleoptera Javana, mit Berucksichtigung der ihnen nahestehenden Formen anderer Herkunft. Stettiner Entomologische Zeitung, 98: 51 - 78." type="journal article" year="1937">Heller, 1937</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FED205922FA2FF85E" pageId="4" pageNumber="328" type="key">
<paragraph id="8BF636BBFF868B5FED205922FA2FF85E" blockId="4.[813,1454,1858,2081]" pageId="4" pageNumber="328">
<keyLead id="30B8958EFF868B5FED205922FA2FF85E" pageId="4" pageNumber="328">
<keyStep id="30BD2E1EFF868B5FED205922FA2FF85E" pageId="4" pageNumber="328">1. Pronotum sparsely covered with whitish or reddish-brown vestiture, with two patches of reddish-brown, erect scales on the middle................................................................................2</keyStep>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,133 @@
<document id="8818A86FA80BB4D75B6AB7746D245B32" ID-DOI="10.26107/RBZ-2024-0026" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683110" ID-ZooBank="B8F6DA4A-6776-43C8-9688-53589751DD44" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171913440" checkinUser="felipe" docAuthor="Tseng, Wei-Zhe, Cheng, &amp; Ren-Chung" docDate="2024" docId="03E087ADFF868B5FED375AE3FB2FF915" docLanguage="en" docName="RafflesBZool.72.324-334.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Eucryptorrhynchus khasiensis Devi, Dey &amp; Ray 2016" docType="treatment" docVersion="2" lastPageNumber="328" masterDocId="FFD9FFD5FF828B5BEE0D5E60FF85FFCE" masterDocTitle="Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)" masterLastPageNumber="334" masterPageNumber="324" pageNumber="328" updateTime="1737175838455" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="E1E75EE8847BE173DE076C7CD41BBE3A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D65B56CBC06099C2F4AC1D4FEABCA220">
<mods:title id="06C6790E72AC922B6F6A4F7A8FD7B12B">Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)</mods:title>
</mods:titleInfo>
<mods:name id="E9AF94EEEE1FF4A3495730B96C606401" type="personal">
<mods:role id="F7526830C54757F314FD62CF9CEE610F">
<mods:roleTerm id="B2E76D6E5EB0C305D39550330A25FFDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F80C8815E05E922F86B9FA5E87FD84C">Tseng, Wei-Zhe</mods:namePart>
</mods:name>
<mods:name id="FE8DE744284FD4125D2C3F19AD2C91D7" type="personal">
<mods:role id="6D41332FB299018F16F16896030DBB71">
<mods:roleTerm id="95777E8C9BA791866F47F5E9EFF7508A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8DCFAE524A8D8306BF85524F67C3B443">Cheng,</mods:namePart>
</mods:name>
<mods:name id="F4157FDA6D3F25E690375B7AFFDA5802" type="personal">
<mods:role id="A43C2FFCBB80D77D296D8D8E5346F6CC">
<mods:roleTerm id="AD72374753BAD8F4B42AE806EC714B28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E25456D60AA8E0F28541388371C696D">Ren-Chung</mods:namePart>
</mods:name>
<mods:typeOfResource id="CEE7240EF7EABB7085D37329123874B9">text</mods:typeOfResource>
<mods:relatedItem id="0BA32F6D4ABF6BBB42CAF5AF16AA38FA" type="host">
<mods:titleInfo id="D983A1DA53BDFB156C0AADB78D2C50C1">
<mods:title id="5F7004C2252168E7B03C71E65AC1E1D0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="2628B8B4DCF677E17158087EE086A793">
<mods:date id="3C38ABD70D6454A50360AC09E8C3F6F7">2024</mods:date>
<mods:detail id="C95D049518BF7399728208E6408CA351" type="pubDate">
<mods:number id="A426BC7D9A2F16BF86132DEB8A5C3048">2024-09-20</mods:number>
</mods:detail>
<mods:detail id="4BCFE59B7510749C38FF4C7B2B6F88F0" type="volume">
<mods:number id="1FA59AD28E64C9868BAD8EA9F2CD8890">72</mods:number>
</mods:detail>
<mods:extent id="CC8B69E584E5B4EB6667BC573E43FDBE" unit="page">
<mods:start id="B04D5C4FF5A624603EC9E269EECA74C0">324</mods:start>
<mods:end id="61EAA20FFC4AB68755D1B17173080274">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A50C47F9A732F027AF7ED37B8BF27D19">journal article</mods:classification>
<mods:identifier id="CE3551530941D602582D0ACF34C60CC4" type="DOI">10.26107/RBZ-2024-0026</mods:identifier>
<mods:identifier id="048102B06C057E3B271385DE44D2C161" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DCF02A55E66FF460F08B76A0F6C4E874" type="Zenodo-Dep">14683110</mods:identifier>
<mods:identifier id="828BB8AD455E5B66061CA6297E83311A" type="ZooBank">B8F6DA4A-6776-43C8-9688-53589751DD44</mods:identifier>
</mods:mods>
<treatment id="03E087ADFF868B5FED375AE3FB2FF915" ID-DOI="http://doi.org/10.5281/zenodo.14683132" ID-Zenodo-Dep="14683132" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FED375AE3FB2FF915" httpUri="http://treatment.plazi.org/id/03E087ADFF868B5FED375AE3FB2FF915" lastPageNumber="328" pageId="4" pageNumber="328">
<subSubSection id="C3536530FF868B5FED375AE3FA1BFB55" box="[826,1438,1155,1179]" pageId="4" pageNumber="328" type="nomenclature">
<paragraph id="8BF636BBFF868B5FED375AE3FA1BFB55" blockId="4.[826,1438,1155,1211]" box="[826,1438,1155,1179]" pageId="4" pageNumber="328">
<heading id="D0BE81D7FF868B5FED375AE3FA1BFB55" bold="true" box="[826,1438,1155,1179]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="328" reason="2">
<taxonomicName id="4C494D38FF868B5FED375AE3FA1BFB55" authority="Devi, Dey &amp; Ray, 2016" authorityName="Devi, Dey &amp; Ray" authorityYear="2016" box="[826,1438,1155,1179]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="khasiensis">
<emphasis id="B93DEAA9FF868B5FED375AE3FA1BFB55" bold="true" box="[826,1438,1155,1179]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FED375AE3FB0EFB55" bold="true" box="[826,1163,1155,1179]" italics="true" pageId="4" pageNumber="328">Eucryptorrhynchus khasiensis</emphasis>
<bibRefCitation id="EFD84B4AFF868B5FEA995AE3FA1BFB55" author="Devi SR &amp; Dey D &amp; Ray DC" box="[1172,1438,1155,1179]" pageId="4" pageNumber="328" pagination="70 - 78" refId="ref6690" refString="Devi SR, Dey D &amp; Ray DC (2016) First record of genus Eucryptorrhynchus (Coleoptera: Curculionidae: Cryptorhynchinae) from India along with description of a new species. Oriental Insects, 51: 70 - 78." type="journal article" year="2016">Devi, Dey &amp; Ray, 2016</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FEA0C5AC3FB52FB75" box="[1025,1239,1187,1211]" pageId="4" pageNumber="328" type="description">
<paragraph id="8BF636BBFF868B5FEA0C5AC3FB52FB75" blockId="4.[826,1438,1155,1211]" box="[1025,1239,1187,1211]" pageId="4" pageNumber="328">
(
<figureCitation id="13722A3EFF868B5FEA045AC3FB00FB75" box="[1033,1157,1187,1211]" captionStart="Fig" captionStartId="3.[136,171,1352,1374]" captionTargetBox="[160,1445,215,1305]" captionTargetId="figure-48@3.[136,1474,189,1336]" captionTargetPageId="3" captionText="Fig. 1. Habitus of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, ventral views; E. brandti (Harold, 1880): C, dorsal and D, ventral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, ventral views." figureDoi="http://doi.org/10.5281/zenodo.14683112" httpUri="https://zenodo.org/record/14683112/files/figure.png" pageId="4" pageNumber="328">Figs. 1E, F</figureCitation>
,
<figureCitation id="13722A3EFF868B5FEA9F5AC3FB4AFB75" box="[1170,1231,1187,1211]" captionStart="Fig" captionStartId="4.[113,148,953,975]" captionTargetBox="[113,1451,189,938]" captionTargetId="figure-52@4.[113,1451,189,938]" captionTargetPageId="4" captionText="Fig. 2. Male genitalia of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, lateral views; E. brandti (Harold, 1880): C, dorsal and D, lateral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, lateral views." figureDoi="http://doi.org/10.5281/zenodo.14683114" httpUri="https://zenodo.org/record/14683114/files/figure.png" pageId="4" pageNumber="328">2E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FED205A83FB74FAF5" pageId="4" pageNumber="328" type="materials_examined">
<paragraph id="8BF636BBFF868B5FED205A83FB74FAF5" blockId="4.[811,1452,1251,1340]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FED205A83FB9CFB35" bold="true" box="[813,1049,1251,1275]" pageId="4" pageNumber="328">Material examined.</emphasis>
<materialsCitation id="3B213CE6FF868B5FEA295A83FB68FAF5" collectingDate="2015-09-08" collectionCode="TARI" collectorName="Mang City" country="China" location="Yunnan" pageId="4" pageNumber="328" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan">
<emphasis id="B93DEAA9FF868B5FEA295A83FB7EFB35" bold="true" box="[1060,1275,1251,1275]" pageId="4" pageNumber="328">
<collectingCountry id="F35E762BFF868B5FEA295A83FB04FB35" box="[1060,1153,1251,1275]" name="China" pageId="4" pageNumber="328">CHINA</collectingCountry>
,
<collectingRegion id="498DF859FF868B5FEA9F5A83FB73FB35" box="[1170,1270,1251,1275]" country="China" name="Yunnan" pageId="4" pageNumber="328">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF868B5FEB0B5A83FAD0FB35" box="[1286,1365,1251,1275]" count="1" pageId="4" pageNumber="328" type="male">1 male</specimenCount>
(ex. 1; accession number: PP697491) (
<collectionCode id="ED58AE7EFF868B5FEA795B63FB33FAD5" box="[1140,1206,1283,1307]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="328">TARI</collectionCode>
), Bangda [
<emphasis id="B93DEAA9FF868B5FEB285B63FADFFAD7" bold="true" box="[1317,1370,1283,1305]" pageId="4" pageNumber="328">fflṁ</emphasis>
],
<collectorName id="26BC536DFF868B5FEB635B64FCD9FAF5" pageId="4" pageNumber="328">Mang City</collectorName>
, coll. Y-T Wang,
<date id="FFF7107BFF868B5FEA295B43FB68FAF5" box="[1060,1261,1315,1339]" pageId="4" pageNumber="328" value="2015-09-08">
<collectingDate id="EFB3E993FF868B5FEA295B43FB68FAF5" box="[1060,1261,1315,1339]" pageId="4" pageNumber="328" value="2015-09-08">8 September 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FED205B03FBB2FA55" pageId="4" pageNumber="328" type="distribution">
<paragraph id="8BF636BBFF868B5FED205B03FBB2FA55" blockId="4.[813,1451,1379,1435]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FED205B03FC3AFAB5" bold="true" box="[813,959,1379,1403]" pageId="4" pageNumber="328">Distribution.</emphasis>
<collectingCountry id="F35E762BFF868B5FEDC45B03FB85FAB5" box="[969,1024,1379,1403]" name="India" pageId="4" pageNumber="328">India</collectingCountry>
(Khasi hills) (
<bibRefCitation id="EFD84B4AFF868B5FEAAE5B03FAE1FAB5" author="Devi SR &amp; Dey D &amp; Ray DC" box="[1187,1380,1379,1403]" pageId="4" pageNumber="328" pagination="70 - 78" refId="ref6690" refString="Devi SR, Dey D &amp; Ray DC (2016) First record of genus Eucryptorrhynchus (Coleoptera: Curculionidae: Cryptorhynchinae) from India along with description of a new species. Oriental Insects, 51: 70 - 78." type="journal article" year="2016">Devi et al., 2016</bibRefCitation>
). New record:
<collectingCountry id="F35E762BFF868B5FED8C5BE3FC44FA55" box="[897,961,1411,1435]" name="China" pageId="4" pageNumber="328">China</collectingCountry>
(
<collectingRegion id="498DF859FF868B5FEDDC5BE3FBA9FA55" box="[977,1068,1411,1435]" country="China" name="Yunnan" pageId="4" pageNumber="328">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FED205BA3FB2FF915" pageId="4" pageNumber="328" type="key">
<paragraph id="8BF636BBFF868B5FED205BA3FB2FF915" blockId="4.[811,1452,1475,1755]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FED205BA3FB81FA15" bold="true" box="[813,1028,1475,1499]" pageId="4" pageNumber="328">Identification key.</emphasis>
We provided a revised identification key that covers five of the six described
<taxonomicName id="4C494D38FF868B5FEAD05B83FC04F9D5" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93DEAA9FF868B5FEAD05B83FA2EFA35" box="[1245,1451,1507,1531]" italics="true" pageId="4" pageNumber="328">Eucryptorrhynchus</emphasis>
species.
</taxonomicName>
For the morphological comparisons of two species,
<taxonomicName id="4C494D38FF868B5FED205844FC31F9F5" baseAuthorityName="Heller" baseAuthorityYear="1915" box="[813,948,1572,1595]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="papuanus">
<emphasis id="B93DEAA9FF868B5FED205844FC31F9F5" box="[813,948,1572,1595]" italics="true" pageId="4" pageNumber="328">E. papuanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C494D38FF868B5FEDE75844FBD8F9F5" authorityName="Pancini" authorityYear="2023" box="[1002,1117,1572,1595]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="cenninii">
<emphasis id="B93DEAA9FF868B5FEDE75844FBD8F9F5" box="[1002,1117,1572,1595]" italics="true" pageId="4" pageNumber="328">E. cenninii</emphasis>
</taxonomicName>
, we had to rely on the original descriptions due to the unavailability of specimens for examination. Additionally, we were unable to include
<taxonomicName id="4C494D38FF868B5FEB995804FC2FF955" baseAuthorityName="Heller" baseAuthorityYear="1922" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="B93DEAA9FF868B5FEB995804FC2FF955" italics="true" pageId="4" pageNumber="328">E. intermedius</emphasis>
</taxonomicName>
in the key, due to the unavailability of specimens for examination and the limited information available in the original description of this species.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,227 @@
<document id="9E98669FFBE9745181E053E6D8768846" ID-DOI="10.26107/RBZ-2024-0026" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683110" ID-ZooBank="B8F6DA4A-6776-43C8-9688-53589751DD44" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171913440" checkinUser="felipe" docAuthor="Tseng, Wei-Zhe, Cheng, &amp; Ren-Chung" docDate="2024" docId="03E087ADFF868B5FEECC5BC3FAB0FB95" docLanguage="en" docName="RafflesBZool.72.324-334.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Eucryptorrhynchus brandti" docType="treatment" docVersion="2" lastPageNumber="328" masterDocId="FFD9FFD5FF828B5BEE0D5E60FF85FFCE" masterDocTitle="Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)" masterLastPageNumber="334" masterPageNumber="324" pageNumber="328" updateTime="1737175838455" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="4D18035314945051F28D57639984211E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD072981449738138F506358807F2BA7">
<mods:title id="D1E90ADAC12956C98803D631F1BB2F2D">Phylogenetic analyses ofEucryptorrhynchus (Coleoptera: Curculionidae) reveal a cryptic species within eurytopic E. scrobiculatus (Motschulsky, 1854)</mods:title>
</mods:titleInfo>
<mods:name id="64D294CB426335872E8C5B328751C14E" type="personal">
<mods:role id="7CF74EE98D1CBF82B8E0B80CC5F7C9D8">
<mods:roleTerm id="C00E8E856FEAF95888D3A23F93BF0197">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="094B3744D82178421B191267493B7405">Tseng, Wei-Zhe</mods:namePart>
</mods:name>
<mods:name id="24F07F1CA4E6AE2FC2296DE20AB4180E" type="personal">
<mods:role id="25D7F228C7AFF02C7CE1C962547C285D">
<mods:roleTerm id="7966411512CFB4A18C23795E4A8ABFDB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25C73B36CFE5E734E5CC1F6C68E1635D">Cheng,</mods:namePart>
</mods:name>
<mods:name id="8D65B370D944586A46AB6607A7778DAD" type="personal">
<mods:role id="AA03F5897ABC3F547071A63F01980761">
<mods:roleTerm id="F0E8DFA6CF4A7BDD13B2C5549606D02F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8F2A9E2AA98C8860172B249423DA3BA">Ren-Chung</mods:namePart>
</mods:name>
<mods:typeOfResource id="8AA2E4565E6452A882DA0DD9B0ADAFFA">text</mods:typeOfResource>
<mods:relatedItem id="F95D3A07D52240872105B1744ED3E7C5" type="host">
<mods:titleInfo id="846D15868C7B84D1BA15FC1D856F7812">
<mods:title id="059C5A39F37DFF547BE90652DA584B07">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="625E0398919A52FB4034205D589B2FE1">
<mods:date id="0BE6A1DA24CD0972D0CF48AFB8E9334C">2024</mods:date>
<mods:detail id="599C59D9968094EECE709A2C4992EFBA" type="pubDate">
<mods:number id="B0DC442C400F80FC4F3E0C85827521CA">2024-09-20</mods:number>
</mods:detail>
<mods:detail id="3BE1C5B87902AB64CDDE42FF86B298DC" type="volume">
<mods:number id="694BC23DD24EAE6EACE59057930EB04B">72</mods:number>
</mods:detail>
<mods:extent id="8CB157A0B349BE1C65E1C8927D45DA01" unit="page">
<mods:start id="C01926F0461E1A37765DF3ECF5C66755">324</mods:start>
<mods:end id="954C0E74C2DADF2A4088783C328DA966">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3D6F9E2F31787014606ED8730722B65D">journal article</mods:classification>
<mods:identifier id="8C477EB6AC78EC9369E7BC1E32E29D5E" type="DOI">10.26107/RBZ-2024-0026</mods:identifier>
<mods:identifier id="A412783F06E425269EB48B03DC2596F8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="25E6C64679DA4929EF5C8B3FC3E4CC6A" type="Zenodo-Dep">14683110</mods:identifier>
<mods:identifier id="79F2EA35A6F719D284CB1A8ABAEAA413" type="ZooBank">B8F6DA4A-6776-43C8-9688-53589751DD44</mods:identifier>
</mods:mods>
<treatment id="03E087ADFF868B5FEECC5BC3FAB0FB95" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95" httpUri="http://treatment.plazi.org/id/03E087ADFF868B5FEECC5BC3FAB0FB95" lastPageNumber="328" pageId="4" pageNumber="328">
<subSubSection id="C3536530FF868B5FEECC5BC3FD25FA75" box="[193,672,1443,1467]" pageId="4" pageNumber="328" type="nomenclature">
<paragraph id="8BF636BBFF868B5FEECC5BC3FD25FA75" blockId="4.[193,672,1443,1499]" box="[193,672,1443,1467]" pageId="4" pageNumber="328">
<heading id="D0BE81D7FF868B5FEECC5BC3FD25FA75" bold="true" box="[193,672,1443,1467]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="328" reason="2">
<taxonomicName id="4C494D38FF868B5FEECC5BC3FD25FA75" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[193,672,1443,1467]" class="Insecta" family="Curculionidae" genus="Eucryptorrhynchus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="328" phylum="Arthropoda" rank="species" species="brandti">
<emphasis id="B93DEAA9FF868B5FEECC5BC3FD25FA75" bold="true" box="[193,672,1443,1467]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FEECC5BC3FE75FA75" bold="true" box="[193,496,1443,1467]" italics="true" pageId="4" pageNumber="328">Eucryptorrhynchus brandti</emphasis>
(
<bibRefCitation id="EFD84B4AFF868B5FEC0D5BC3FD1FFA75" author="Harold E von" box="[512,666,1443,1467]" pageId="4" pageNumber="328" pagination="148 - 171" refId="ref6994" refString="Harold E von (1880) Einige neue Coleopteren. Mittheilungen des Munchner Entomologischen Vereins, 4: 148 - 171." type="journal article" year="1880">Harold, 1880</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FEF4D5BA3FDA7FA15" box="[320,546,1475,1499]" pageId="4" pageNumber="328" type="description">
<paragraph id="8BF636BBFF868B5FEF4D5BA3FDA7FA15" blockId="4.[193,672,1443,1499]" box="[320,546,1475,1499]" pageId="4" pageNumber="328">
(
<figureCitation id="13722A3EFF868B5FEF455BA3FE4CFA15" box="[328,457,1475,1499]" captionStart="Fig" captionStartId="3.[136,171,1352,1374]" captionTargetBox="[160,1445,215,1305]" captionTargetId="figure-48@3.[136,1474,189,1336]" captionTargetPageId="3" captionText="Fig. 1. Habitus of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, ventral views; E. brandti (Harold, 1880): C, dorsal and D, ventral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, ventral views." figureDoi="http://doi.org/10.5281/zenodo.14683112" httpUri="https://zenodo.org/record/14683112/files/figure.png" pageId="4" pageNumber="328">Figs. 1C, D</figureCitation>
,
<figureCitation id="13722A3EFF868B5FEFDA5BA3FD9FFA15" box="[471,538,1475,1499]" captionStart="Fig" captionStartId="4.[113,148,953,975]" captionTargetBox="[113,1451,189,938]" captionTargetId="figure-52@4.[113,1451,189,938]" captionTargetPageId="4" captionText="Fig. 2. Male genitalia of Eucryptorrhynchus species. E. scrobiculatus (Motschulsky, 1854) from Taiwan: A, dorsal and B, lateral views; E. brandti (Harold, 1880): C, dorsal and D, lateral views; E. khasiensis Devi, Dey &amp; Ray, 2016: E, dorsal and F, lateral views." figureDoi="http://doi.org/10.5281/zenodo.14683114" httpUri="https://zenodo.org/record/14683114/files/figure.png" pageId="4" pageNumber="328">2C, D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FEE7C5863FDC6F855" pageId="4" pageNumber="328" type="materials_examined">
<paragraph id="8BF636BBFF868B5FEE7C5863FDC6F855" blockId="4.[112,753,1539,1948]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FEE7C5863FED6F9D5" bold="true" box="[113,339,1539,1563]" pageId="4" pageNumber="328">Material examined.</emphasis>
<materialsCitation id="3B213CE6FF868B5FEF505863FEE4F955" collectingDate="2016-07-23" collectionCode="TARI" collectorName="Anshun" country="China" county="Guanling Buyei and Miao Autonomous County" location="Guanling Buyei" municipality="Huixiangping" pageId="4" pageNumber="328" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Guizhou">
<emphasis id="B93DEAA9FF868B5FEF505863FDB4F9D5" bold="true" box="[349,561,1539,1563]" pageId="4" pageNumber="328">
<collectingCountry id="F35E762BFF868B5FEF505863FE32F9D5" box="[349,439,1539,1563]" name="China" pageId="4" pageNumber="328">CHINA</collectingCountry>
,
<collectingRegion id="498DF859FF868B5FEFCB5863FDA9F9D5" box="[454,556,1539,1563]" country="China" name="Guizhou" pageId="4" pageNumber="328">Guizhou</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF868B5FEC365863FD03F9D5" box="[571,646,1539,1563]" count="1" pageId="4" pageNumber="328" type="male">1 male</specimenCount>
<specimenCount id="9D4FFD32FF868B5FEC9D5863FD74F9D5" box="[656,753,1539,1563]" count="1" pageId="4" pageNumber="328" type="female">1 female</specimenCount>
(ex. 1, 2; accession number: PP697490, PP697489) (
<collectionCode id="ED58AE7EFF868B5FEE775823FF4DF995" box="[122,200,1603,1627]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="328">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF868B5FEEE95823FE0AF995" box="[228,399,1603,1627]" pageId="4" pageNumber="328">Huixiangping</collectingMunicipality>
[
<emphasis id="B93DEAA9FF868B5FEFA45823FD87F997" bold="true" box="[425,514,1603,1625]" pageId="4" pageNumber="328">回 ⁂ 坪</emphasis>
],
<collectingCounty id="62974E37FF868B5FEC2B5823FE7CF9B5" pageId="4" pageNumber="328">
<location id="8E966060FF868B5FEC2B5823FD6BF995" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEC2B5823FD6BF995" box="[550,750,1603,1627]" country="China" county="Guanling Buyei and Miao Autonomous County" municipality="Huixiangping" name="Guanling Buyei" pageId="4" pageNumber="328" stateProvince="Guizhou">Guanling Buyei</location>
and
<location id="8E966060FF868B5FEEA45803FE7CF9B5" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEEA45803FE7CF9B5" box="[169,505,1635,1659]" country="China" county="Guanling Buyei and Miao Autonomous County" municipality="Huixiangping" name="Miao Autonomous County" pageId="4" pageNumber="328" stateProvince="Guizhou">Miao Autonomous County</location>
</collectingCounty>
,
<collectorName id="26BC536DFF868B5FEC075803FDEFF9B5" box="[522,618,1635,1659]" pageId="4" pageNumber="328">Anshun</collectorName>
, coll. Y-T
<location id="8E966060FF868B5FEE7C58E4FF33F952" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEE7C58E4FF33F952" box="[113,182,1668,1692]" country="China" county="Guanling Buyei and Miao Autonomous County" municipality="Huixiangping" name="Wang" pageId="4" pageNumber="328" stateProvince="Guizhou">Wang</location>
,
<date id="FFF7107BFF868B5FEECB58E3FEE4F955" box="[198,353,1667,1691]" pageId="4" pageNumber="328" value="2016-07-23">
<collectingDate id="EFB3E993FF868B5FEECB58E3FEE4F955" box="[198,353,1667,1691]" pageId="4" pageNumber="328" value="2016-07-23">23 July 2016</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B213CE6FF868B5FEF7D58E3FEDEF915" collectingDate="2014-06-30" collectionCode="TARI" collectorName="Diqing Tibetan Autonomous Prefecture" country="China" location="Wang" municipality="Dadi" pageId="4" pageNumber="328" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<emphasis id="B93DEAA9FF868B5FEF7D58E3FE5CF955" bold="true" box="[368,473,1667,1691]" pageId="4" pageNumber="328">
<collectingRegion id="498DF859FF868B5FEF7D58E3FE51F955" box="[368,468,1667,1691]" country="China" name="Yunnan" pageId="4" pageNumber="328">Yunnan</collectingRegion>
:
</emphasis>
<specimenCount id="9D4FFD32FF868B5FEFE958E3FDCCF955" box="[484,585,1667,1691]" count="1" pageId="4" pageNumber="328" type="female">1 female</specimenCount>
(
<collectionCode id="ED58AE7EFF868B5FEC5158E3FD26F955" box="[604,675,1667,1691]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="328">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF868B5FECB758E3FD6AF955" box="[698,751,1667,1691]" pageId="4" pageNumber="328">Dadi</collectingMunicipality>
[
<emphasis id="B93DEAA9FF868B5FEE7758C3FF2AF977" bold="true" box="[122,175,1699,1721]" pageId="4" pageNumber="328">巴®</emphasis>
],
<collectorName id="26BC536DFF868B5FEEC458C3FDF8F975" box="[201,637,1699,1723]" pageId="4" pageNumber="328">Diqing Tibetan Autonomous Prefecture</collectorName>
, coll. Y-T
<location id="8E966060FF868B5FEE7C58A4FF31F912" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEE7C58A4FF31F912" box="[113,180,1732,1756]" country="China" municipality="Dadi" name="Wang" pageId="4" pageNumber="328" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF868B5FEECF58A3FEDEF915" box="[194,347,1731,1755]" pageId="4" pageNumber="328" value="2014-06-30">
<collectingDate id="EFB3E993FF868B5FEECF58A3FEDEF915" box="[194,347,1731,1755]" pageId="4" pageNumber="328" value="2014-06-30">30 June 2014</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF868B5FEF6758A3FF2EF8F5" collectingDate="2016-07-08" collectionCode="TARI" collectorName="Diqing Tibetan Autonomous Prefecture" country="China" location="Wang" municipality="Wangzanka Hotel" pageId="4" pageNumber="328" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan">
<specimenCount id="9D4FFD32FF868B5FEF6758A3FE4FF915" box="[362,458,1731,1755]" count="1" pageId="4" pageNumber="328" type="female">1 female</specimenCount>
(ex. 4; accession number: PP693364) (
<collectionCode id="ED58AE7EFF868B5FEEFE5883FEB3F935" box="[243,310,1763,1787]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="328">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF868B5FEF4B5883FD87F935" box="[326,514,1763,1787]" pageId="4" pageNumber="328">Wangzanka Hotel</collectingMunicipality>
[
<emphasis id="B93DEAA9FF868B5FEC025883FD16F937" bold="true" box="[527,659,1763,1785]" pageId="4" pageNumber="328">ffi咱ϮΘ店</emphasis>
],
<collectorName id="26BC536DFF868B5FECAA5883FE52F8D5" pageId="4" pageNumber="328">Diqing Tibetan Autonomous Prefecture</collectorName>
, coll. Y-T
<location id="8E966060FF868B5FEC5A5964FD1FF8D2" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEC5A5964FD1FF8D2" box="[599,666,1796,1820]" country="China" municipality="Wangzanka Hotel" name="Wang" pageId="4" pageNumber="328" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF868B5FECA45963FF2EF8F5" pageId="4" pageNumber="328" value="2016-07-08">
<collectingDate id="EFB3E993FF868B5FECA45963FF2EF8F5" pageId="4" pageNumber="328" value="2016-07-08">8 July 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B213CE6FF868B5FEEB75943FDC5F855" collectingDate="2016-06-15" collectionCode="TARI" collectorName="Lisu Autonomous Prefecture" country="China" county="Autonomous County" location="Gongshan" municipality="Nageluo" pageId="4" pageNumber="328" specimenCount="2" specimenCount-female="2" stateProvince="Yunnan">
<specimenCount id="9D4FFD32FF868B5FEEB75943FEA0F8F5" box="[186,293,1827,1851]" count="2" pageId="4" pageNumber="328" type="female">2 females</specimenCount>
(ex. 3, 5; accession number: PP697487, PP697488) (
<collectionCode id="ED58AE7EFF868B5FEEF55923FEBEF895" box="[248,315,1859,1883]" country="Taiwan" httpUri="http://grbio.org/cool/a13e-ta1z" name="Taiwan Agricultural Research Institute" pageId="4" pageNumber="328">TARI</collectionCode>
),
<collectingMunicipality id="6B92ACC1FF868B5FEF425923FE2CF895" box="[335,425,1859,1883]" pageId="4" pageNumber="328">Nageluo</collectingMunicipality>
[
<emphasis id="B93DEAA9FF868B5FEFB65923FD8EF897" bold="true" box="[443,523,1859,1881]" pageId="4" pageNumber="328">Oi格洛</emphasis>
],
<location id="8E966060FF868B5FEC2E5923FD0AF895" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEC2E5923FD0AF895" box="[547,655,1859,1883]" country="China" county="Autonomous County" municipality="Nageluo" name="Gongshan" pageId="4" pageNumber="328" stateProvince="Yunnan">Gongshan</location>
Derung-Nu
<collectingCounty id="62974E37FF868B5FEE975903FEF8F8B5" box="[154,381,1891,1915]" pageId="4" pageNumber="328">Autonomous County</collectingCounty>
,
<location id="8E966060FF868B5FEF875903FE65F8B5" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEF875903FE65F8B5" box="[394,480,1891,1915]" country="China" county="Autonomous County" municipality="Nageluo" name="Nujiang" pageId="4" pageNumber="328" stateProvince="Yunnan">Nujiang</location>
of the
<collectorName id="26BC536DFF868B5FEC225903FF64F855" pageId="4" pageNumber="328">Lisu Autonomous Prefecture</collectorName>
, coll. Y-T
<location id="8E966060FF868B5FEF5659E4FE18F852" LSID="urn:lsid:plazi:treatment:03E087ADFF868B5FEECC5BC3FAB0FB95:8E966060FF868B5FEF5659E4FE18F852" box="[347,413,1924,1948]" country="China" county="Autonomous County" municipality="Nageluo" name="Wang" pageId="4" pageNumber="328" stateProvince="Yunnan">Wang</location>
,
<date id="FFF7107BFF868B5FEFA759E3FDC5F855" box="[426,576,1923,1947]" pageId="4" pageNumber="328" value="2016-06-15">
<collectingDate id="EFB3E993FF868B5FEFA759E3FDC5F855" box="[426,576,1923,1947]" pageId="4" pageNumber="328" value="2016-06-15">15 June 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3536530FF868B5FEE7C59A3FAB0FB95" pageId="4" pageNumber="328" type="distribution">
<paragraph id="8BF636BBFF868B5FEE7C59A3FD75F7D5" blockId="4.[113,754,1987,2075]" pageId="4" pageNumber="328">
<emphasis id="B93DEAA9FF868B5FEE7C59A3FE91F815" bold="true" box="[113,276,1987,2011]" pageId="4" pageNumber="328">Distribution.</emphasis>
<collectingCountry id="F35E762BFF868B5FEF2D59A3FEE0F815" box="[288,357,1987,2011]" name="China" pageId="4" pageNumber="328">China</collectingCountry>
(
<collectingRegion id="498DF859FF868B5FEF7759A3FE4DF815" box="[378,456,1987,2011]" country="China" name="Anhui" pageId="4" pageNumber="328">Anhui</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEFD459A3FDB6F815" box="[473,563,1987,2011]" country="China" name="Beijing" pageId="4" pageNumber="328">Beijing</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEC4859A3FD17F815" box="[581,658,1987,2011]" country="China" name="Gansu" pageId="4" pageNumber="328">Gansu</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FECAE59A3FD6EF815" box="[675,747,1987,2011]" country="China" name="Hebei" pageId="4" pageNumber="328">Hebei</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEE7E5983FE81F835" box="[115,260,2019,2043]" country="China" name="Heilongjiang" pageId="4" pageNumber="328">Heilongjiang</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEF1F5984FEDEF835" box="[274,347,2020,2043]" country="China" name="Henan" pageId="4" pageNumber="328">Henan</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEF675983FE2AF835" box="[362,431,2019,2043]" country="China" name="Hubei" pageId="4" pageNumber="328">Hubei</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEFB05983FD94F835" box="[445,529,2019,2043]" country="China" name="Jiangsu" pageId="4" pageNumber="328">Jiangsu</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEC2D5983FD00F835" box="[544,645,2019,2043]" country="China" name="Liaoning" pageId="4" pageNumber="328">Liaoning</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEC995983FD6BF835" box="[660,750,2019,2043]" country="China" name="Sichuan" pageId="4" pageNumber="328">Sichuan</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEE7F5663FF4EF7D5" box="[114,203,2051,2075]" country="China" name="Shaanxi" pageId="4" pageNumber="328">Shaanxi</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEED95663FEBFF7D5" box="[212,314,2051,2075]" country="China" name="Shanghai" pageId="4" pageNumber="328">Shanghai</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEF495663FE34F7D5" box="[324,433,2051,2075]" country="China" name="Shandong" pageId="4" pageNumber="328">Shandong</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEFB65663FD8CF7D5" box="[443,521,2051,2075]" country="China" name="Shanxi" pageId="4" pageNumber="328">Shanxi</collectingRegion>
),
<collectingCountry id="F35E762BFF868B5FEC165664FDDFF7D5" box="[539,602,2052,2075]" name="Japan" pageId="4" pageNumber="328">Japan</collectingCountry>
,
<collectingCountry id="F35E762BFF868B5FEC695663FD68F7D5" box="[612,749,2051,2075]" name="North Korea" pageId="4" pageNumber="328">North Korea</collectingCountry>
,
</paragraph>
<paragraph id="8BF636BBFF868B5FED215A43FAB0FB95" blockId="4.[812,1450,1059,1115]" pageId="4" pageNumber="328">
<collectingCountry id="F35E762BFF868B5FED215A43FCF0FBF5" box="[812,885,1059,1083]" name="Russia" pageId="4" pageNumber="328">Russia</collectingCountry>
(Far East),
<collectingCountry id="F35E762BFF868B5FEA0F5A43FB0AFBF5" box="[1026,1167,1059,1083]" name="South Korea" pageId="4" pageNumber="328">South Korea</collectingCountry>
(
<bibRefCitation id="EFD84B4AFF868B5FEAAC5A43FCE2FB95" author="Alonso-Zarazaga MA &amp; Barrios H &amp; Borovec R &amp; Bouchard P &amp; Caldara R &amp; Colonnelli E &amp; Gultekin L &amp; Hlavac P &amp; Korotyaev B &amp; Lyal CHC &amp; Machado A &amp; Meregalli M &amp; Pierotti H &amp; Ren L &amp; Sanchez-Ruiz M &amp; Sforzi A &amp; Silfverberg H &amp; Skuhrovec J &amp; Tryzna M &amp; Velazquez de Castro AJ &amp; Yunakov NN" pageId="4" pageNumber="328" refId="ref6096" refString="Alonso-Zarazaga MA, Barrios H, Borovec R, Bouchard P, Caldara R, Colonnelli E, Gultekin L, Hlavac P, Korotyaev B, Lyal CHC, Machado A, Meregalli M, Pierotti H, Ren L, Sanchez-Ruiz M, Sforzi A, Silfverberg H, Skuhrovec J, Tryzna M, Velazquez de Castro AJ &amp; Yunakov NN (2017) Cooperative Catalogue of Palaearctic Coleoptera Curculionoidea. Monografias Electronicas SEA 8, Zaragoza, Spain, 729 pp." type="book" year="2017">Alonso-Zarazaga et al., 2017</bibRefCitation>
). New records:
<collectingCountry id="F35E762BFF868B5FEA195A23FBD1FB95" box="[1044,1108,1091,1115]" name="China" pageId="4" pageNumber="328">China</collectingCountry>
(
<collectingRegion id="498DF859FF868B5FEA6E5A23FB46FB95" box="[1123,1219,1091,1115]" country="China" name="Guizhou" pageId="4" pageNumber="328">Guizhou</collectingRegion>
,
<collectingRegion id="498DF859FF868B5FEADD5A23FAACFB95" box="[1232,1321,1091,1115]" country="China" name="Yunnan" pageId="4" pageNumber="328">Yunnan</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

File diff suppressed because one or more lines are too long

View file

@ -0,0 +1,172 @@
<document id="D379E959EC3669EE35BF1C6FF87DB174" ID-DOI="10.26107/RBZ-2024-0023" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682920" ID-ZooBank="1A56A449-CC19-40DD-AF92-0604AD10850C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171908861" checkinUser="felipe" docAuthor="Savatenalinton, Sukonthip" docDate="2024" docId="03EE87F9FFCBDC3B7B62F84BFA27F7B5" docLanguage="en" docName="RafflesBZool.72.280-293.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Gurayacypris danielopoli Savatenalinton 2024" docType="treatment" docVersion="2" lastPageNumber="291" masterDocId="FFD7FF81FFC1DC30784FFFCBFF89FFA6" masterDocTitle="A new species of the genus Gurayacypris Battish, 1987 (Crustacea: Ostracoda: Notodromadidae) from a rice field in northeast Thailand" masterLastPageNumber="293" masterPageNumber="280" pageNumber="290" updateTime="1737175572129" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8D4A40F5838DBAEFEA22B625DF84FF66" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8905B91C7ABB162CE09ADCB1FC7A2B00">
<mods:title id="4B326BBC6E04D1A10B1CEAE706467928">A new species of the genus Gurayacypris Battish, 1987 (Crustacea: Ostracoda: Notodromadidae) from a rice field in northeast Thailand</mods:title>
</mods:titleInfo>
<mods:name id="6071F4C83E8D9A06F21C7489D09C96B1" type="personal">
<mods:role id="1C46A8D547AD3C1774432B6BDD7F2420">
<mods:roleTerm id="A52030BE8CB6326F8B0D27B0FEF2FE2E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="769512FA48D2431DC5CF45AE15CEB533">Savatenalinton, Sukonthip</mods:namePart>
</mods:name>
<mods:typeOfResource id="81116110E2739C568D1876106ABE05DD">text</mods:typeOfResource>
<mods:relatedItem id="ED924E0054CBB7CCD3960F21039B41F9" type="host">
<mods:titleInfo id="132D88991B43EC7C94E0685BA498FC30">
<mods:title id="5B6FD2FF8BFAFF0D65DF4BA5E210F5F3">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="A9DF76E257AE310365AE11D8C4CD4304">
<mods:date id="5A2FE42A96A14768112A92720826D24F">2024</mods:date>
<mods:detail id="0FE0FE1F6362C1B4D691689BE6561006" type="pubDate">
<mods:number id="616A594185810209C9E571D9BCC92130">2024-08-21</mods:number>
</mods:detail>
<mods:detail id="E031F260179D781085EFD51C8F684D1C" type="volume">
<mods:number id="1DC5B5AE6553F815191B52F5970A8574">72</mods:number>
</mods:detail>
<mods:extent id="164F82D476F54821D88D1A359491A1CE" unit="page">
<mods:start id="D71C3942E21B953C1717DE35976CD004">280</mods:start>
<mods:end id="4F9EBE427F6DF311090B7684436DEEB5">293</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FA69DC959740409690F6E1D1674103A5">journal article</mods:classification>
<mods:identifier id="72FA5394EFDDCC864CD2202FE1EDC3F2" type="DOI">10.26107/RBZ-2024-0023</mods:identifier>
<mods:identifier id="1C4E6A11E989DCBAFE34FB226B1C304D" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="B38FDF957DB6CBBD537B7C0BCF329BA3" type="Zenodo-Dep">14682920</mods:identifier>
<mods:identifier id="096A249787E1C8202B2583DB95560EFB" type="ZooBank">1A56A449-CC19-40DD-AF92-0604AD10850C</mods:identifier>
</mods:mods>
<treatment id="03EE87F9FFCBDC3B7B62F84BFA27F7B5" LSID="urn:lsid:plazi:treatment:03EE87F9FFCBDC3B7B62F84BFA27F7B5" httpUri="http://treatment.plazi.org/id/03EE87F9FFCBDC3B7B62F84BFA27F7B5" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290">
<subSubSection id="C35D6564FFCBDC3A7B62F84BFA22F83F" box="[813,1451,1920,1945]" pageId="10" pageNumber="290" type="nomenclature">
<paragraph id="8BF836EFFFCBDC3A7B62F84BFA22F83F" blockId="10.[813,1452,1920,2105]" box="[813,1451,1920,1945]" pageId="10" pageNumber="290">
<heading id="D0B08183FFCBDC3A7B62F84BFA22F83F" bold="true" box="[813,1451,1920,1945]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="290" reason="2">
<emphasis id="B933EAFDFFCBDC3A7B62F84BFA22F83F" bold="true" box="[813,1451,1920,1945]" pageId="10" pageNumber="290">
Comparisons between
<taxonomicName id="4C474D6CFFCBDC3A7C75F84AFA22F83F" authorityName="Savatenalinton" authorityYear="2024" box="[1082,1451,1920,1945]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="290" phylum="Arthropoda" rank="species" species="danielopoli" status="new species and">
<emphasis id="B933EAFDFFCBDC3A7C75F84AFB52F83E" bold="true" box="[1082,1243,1920,1945]" italics="true" pageId="10" pageNumber="290">G. danielopoli</emphasis>
, new species and
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35D6564FFCBDC3B7B62F86AFA27F7B5" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290" type="description">
<paragraph id="8BF836EFFFCBDC3A7B62F86AFC5DF81E" blockId="10.[813,1452,1920,2105]" box="[813,980,1952,1977]" pageId="10" pageNumber="290">
<taxonomicName id="4C474D6CFFCBDC3A7B62F86AFC5DF81E" authorityName="Battish" authorityYear="1987" box="[813,980,1952,1977]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="290" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCBDC3A7B62F86AFC5DF81E" bold="true" box="[813,980,1952,1977]" italics="true" pageId="10" pageNumber="290">G. kangraensis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF836EFFFCBDC3B7B62F80AFED5F8D5" blockId="10.[813,1452,1920,2105]" lastBlockId="11.[136,777,1403,1907]" lastPageId="11" lastPageNumber="291" pageId="10" pageNumber="290">
<taxonomicName id="4C474D6CFFCBDC3A7B62F80AFBC6F87F" authorityName="Savatenalinton" authorityYear="2024" box="[813,1103,1985,2009]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="290" phylum="Arthropoda" rank="species" species="danielopoli" status="sp. nov.">
<emphasis id="B933EAFDFFCBDC3A7B62F80AFBC6F87F" box="[813,1103,1985,2009]" italics="true" pageId="10" pageNumber="290">Gurayacypris danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCBDC3A7C2EF80AFB64F87F" box="[1121,1261,1985,2009]" pageId="10" pageNumber="290" rank="species">new species</taxonomicNameLabel>
, differs from
<taxonomicName id="4C474D6CFFCBDC3A7DDEF80AFC39F85F" authorityName="Battish" authorityYear="1987" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="290" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCBDC3A7DDEF80AFC39F85F" italics="true" pageId="10" pageNumber="290">G. kangraensis</emphasis>
</taxonomicName>
in many aspects of its valves and soft parts. In the male
<taxonomicName id="4C474D6CFFCBDC3A7BFAF7CAFBDBF7BF" authorityName="Savatenalinton" authorityYear="2024" box="[949,1106,2049,2073]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="10" pageNumber="290" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCBDC3A7BFAF7CAFBDBF7BF" box="[949,1106,2049,2073]" italics="true" pageId="10" pageNumber="290">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCBDC3A7C2CF7CAFB63F7BF" box="[1123,1258,2049,2073]" pageId="10" pageNumber="290" rank="species">new species</taxonomicNameLabel>
, the Cp in dorsal view is tumid and the greatest width is post mid-length whereas in
<taxonomicName id="4C474D6CFFCADC3B7947FAB7FE24FA35" authorityName="Battish" authorityYear="1987" box="[264,429,1403,1427]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7947FAB7FE24FA35" box="[264,429,1403,1427]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
, it is elliptical and the greatest width is at mid-length. The posterior extremity of the new species possesses a tiny pointed tip, but in
<taxonomicName id="4C474D6CFFCADC3B7A10FA77FC89FA75" authorityName="Battish" authorityYear="1987" box="[607,768,1467,1491]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7A10FA77FC89FA75" box="[607,768,1467,1491]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
, this part is rounded. The female Cp shape in lateral view also demonstrates the difference between these two species.
<taxonomicName id="4C474D6CFFCADC3B78C7F9D7FE16F995" authorityName="Savatenalinton" authorityYear="2024" box="[136,415,1563,1587]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="sp. nov.">
<emphasis id="B933EAFDFFCADC3B78C7F9D7FE16F995" box="[136,415,1563,1587]" italics="true" pageId="11" pageNumber="291">Gurayacypris danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B79FFF9D7FDBEF995" box="[432,567,1563,1587]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
, has a subglobular shape with a strongly arched dorsal margin while in
<taxonomicName id="4C474D6CFFCADC3B7AA3F9F7FE80F9D5" authorityName="Battish" authorityYear="1987" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7AA3F9F7FE80F9D5" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
, the shape is subquadrate with a slightly arched dorsal margin. Although both species have a flange (spur) on the female postero-ventral part, the aspect of this structure is different. It is subtriangular with a pointed tip in the Thai species, but it is tiny and subquadrate, with a blunt tip in the Indian species. The Cp size of
<taxonomicName id="4C474D6CFFCADC3B7999F937FDF9F8B5" authorityName="Savatenalinton" authorityYear="2024" box="[470,624,1787,1811]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCADC3B7999F937FDF9F8B5" box="[470,624,1787,1811]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B7A30F937FC8AF8B5" box="[639,771,1787,1811]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
, (806832 μm in male, 794800 μm in female) is also larger than
<taxonomicName id="4C474D6CFFCADC3B788FF8F7FEEDF8F5" authorityName="Battish" authorityYear="1987" box="[192,356,1851,1875]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B788FF8F7FEEDF8F5" box="[192,356,1851,1875]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
(760 μm in male, 700 μm in female) (see
<bibRefCitation id="EFD64B1EFFCADC3B78F3F890FED9F8D5" author="Battish SK" box="[188,336,1883,1907]" pageId="11" pageNumber="291" pagination="127 - 135" refId="ref6179" refString="Battish SK (1987) A new recent genus and species of Notodromadinid ostracod from India. Researches on Crustacea, 16: 127 - 135." type="journal article" year="1987">Battish, 1987</bibRefCitation>
).
</paragraph>
<caption id="DF386667FFCADC3B78C8FADAFC95FAE2" ID-DOI="http://doi.org/10.5281/zenodo.14682940" ID-Zenodo-Dep="14682940" httpUri="https://zenodo.org/record/14682940/files/figure.png" pageId="11" pageNumber="291" startId="11.[135,170,1297,1319]" targetBox="[135,1473,189,1282]" targetPageId="11" targetType="figure">
<paragraph id="8BF836EFFFCADC3B78C8FADAFC95FAE2" blockId="11.[135,1474,1297,1348]" pageId="11" pageNumber="291">
Fig. 10. Distribution map of Oriental
<taxonomicName id="4C474D6CFFCADC3B79A4FADAFD05FA81" authorityName="Kaufmann" authorityYear="1900" box="[491,652,1297,1319]" class="Ostracoda" family="Notodromadidae" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="subFamily" subFamily="Notodromadinae">Notodromadinae</taxonomicName>
(for numbers, see table 1; some sites are only approximately plotted due to limitations of locality information;
<emphasis id="B933EAFDFFCADC3B793CFAE7FE0CFAE5" bold="true" box="[371,389,1324,1347]" pageId="11" pageNumber="291">N</emphasis>
, Locality unknown or with limited data).
</paragraph>
</caption>
<paragraph id="8BF836EFFFCADC3B78C7F857FA27F7B5" blockId="11.[136,777,1947,2067]" lastBlockId="11.[835,1476,1403,2067]" pageId="11" pageNumber="291">
In addition, the differences can be seen in the morphology of the soft parts, especially the female and male T1 palp, the male T2, and the Hp. The female T1 palp of both species is large, but it is smaller in
<taxonomicName id="4C474D6CFFCADC3B79D0F837FDB2F7B5" authorityName="Savatenalinton" authorityYear="2024" box="[415,571,2043,2067]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCADC3B79D0F837FDB2F7B5" box="[415,571,2043,2067]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B7A04F837FD58F7B5" box="[587,721,2043,2067]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
. The tip of this palp bears two small and unequal bristles in
<taxonomicName id="4C474D6CFFCADC3B7DE7FAB7FC4DFA15" authorityName="Battish" authorityYear="1987" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7DE7FAB7FC4DFA15" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
while
<taxonomicName id="4C474D6CFFCADC3B7C42FA57FB2FFA15" authorityName="Savatenalinton" authorityYear="2024" box="[1037,1190,1435,1459]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli">
<emphasis id="B933EAFDFFCADC3B7C42FA57FB2FFA15" box="[1037,1190,1435,1459]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
has a broad, dome-shaped seta with pointed apical tip. The T1 left palp of male of
<taxonomicName id="4C474D6CFFCADC3B7DE6FA77FC41FA55" authorityName="Battish" authorityYear="1987" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7DE6FA77FC41FA55" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
possesses a pointed projection on the distal margin of the basal segment, whereas there is no projection on this part in
<taxonomicName id="4C474D6CFFCADC3B7BA9F9D7FB09F995" authorityName="Savatenalinton" authorityYear="2024" box="[998,1152,1563,1587]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCADC3B7BA9F9D7FB09F995" box="[998,1152,1563,1587]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B7CC0F9D7FA9DF995" box="[1167,1300,1563,1587]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
. The h3 seta on the male T2 is long in
<taxonomicName id="4C474D6CFFCADC3B7C0AF9F7FB63F9F5" authorityName="Battish" authorityYear="1987" box="[1093,1258,1595,1619]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7C0AF9F7FB63F9F5" box="[1093,1258,1595,1619]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
, but it is shorter in the new species. In
<taxonomicName id="4C474D6CFFCADC3B7C51F997FB33F9D5" authorityName="Savatenalinton" authorityYear="2024" box="[1054,1210,1627,1651]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCADC3B7C51F997FB33F9D5" box="[1054,1210,1627,1651]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B7C84F997FAD9F9D5" box="[1227,1360,1627,1651]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
, the Hp is subovate and the ms is short (not protruding beyond edge of ls), with a gently curved dorsal margin, while in
<taxonomicName id="4C474D6CFFCADC3B7DE7F957FC4DF975" authorityName="Battish" authorityYear="1987" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7DE7F957FC4DF975" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
the ms extends over the margin of the ls and has a strongly curved dorsal margin. Additionally, the ls of the new species is subtriangular with a pointed tip directed outward but, in
<taxonomicName id="4C474D6CFFCADC3B7BBEF8D7FB1AF895" authorityName="Battish" authorityYear="1987" box="[1009,1171,1819,1843]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7BBEF8D7FB1AF895" box="[1009,1171,1819,1843]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
, this part bears two circular lobes; one is small and appearing on the inner side, another one is large and located on the outer side. The number of bristles on the masticatory processes of the female T1 is seven in
<taxonomicName id="4C474D6CFFCADC3B7BE9F857FBC0F815" authorityName="Battish" authorityYear="1987" box="[934,1097,1947,1971]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="kangraensis">
<emphasis id="B933EAFDFFCADC3B7BE9F857FBC0F815" box="[934,1097,1947,1971]" italics="true" pageId="11" pageNumber="291">G. kangraensis</emphasis>
</taxonomicName>
whereas it is
<quantity id="4CBF9B0AFFCADC3B7CAFF850FA91F815" box="[1248,1304,1947,1971]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="11" pageNumber="291" unit="in" value="14.0">14 in</quantity>
<taxonomicName id="4C474D6CFFCADC3B7D6FF857FA32F815" authorityName="Savatenalinton" authorityYear="2024" box="[1312,1467,1947,1971]" class="Ostracoda" family="Notodromadidae" genus="Gurayacypris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Podocopida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="danielopoli" status="new species">
<emphasis id="B933EAFDFFCADC3B7D6FF857FA32F815" box="[1312,1467,1947,1971]" italics="true" pageId="11" pageNumber="291">G. danielopoli</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2005786FFCADC3B7B0CF877FC44F875" box="[835,973,1979,2003]" pageId="11" pageNumber="291" rank="species">new species</taxonomicNameLabel>
. Nevertheless, this character should be used with caution as these bristles are small and densely arranged which could affect the observation of the actual number.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,602 @@
<document id="6F6CDAC157E303A786EDD88D97F117C0" ID-DOI="10.26107/RBZ-2024-0033" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682896" ID-ZooBank="56BAA238-5F15-4BE9-933C-0777E8C08C0B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171772636" checkinUser="felipe" docAuthor="Macharoenboon, Kongkit, , Varitsara Manonai, Jeratthitikul, &amp; Ekgachai" docDate="2024" docId="03EFD249FF98FFB8FF41FD4B34E4FACF" docLanguage="en" docName="RafflesBZool.72.450-468.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Caridina sumatrensis De Man 1892" docType="treatment" docVersion="2" lastPageNumber="464" masterDocId="FFD6AA31FF93FFB6FFBAFFB63615FFBA" masterDocTitle="Caridina maeklongensis, a new landlocked freshwater shrimp species (Crustacea: Decapoda: Atyidae) from the Mae Klong Basin, Thailand" masterLastPageNumber="468" masterPageNumber="450" pageNumber="461" updateTime="1737175469701" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="46C1177A81835D2D5FCD6F11024F8C82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9FB6E68C7613203DA95BE2B594D3D4BF">
<mods:title id="FA07EFFDC35042B1CC8DC74CFDB59899">Caridina maeklongensis, a new landlocked freshwater shrimp species (Crustacea: Decapoda: Atyidae) from the Mae Klong Basin, Thailand</mods:title>
</mods:titleInfo>
<mods:name id="176DF30B1D63E0C4F6AB55541CBBB605" type="personal">
<mods:role id="C5D46B02CD87D0DB7C56E2804F58F414">
<mods:roleTerm id="759D568A14B09CE55253B73BE51AE261">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A80F0CA8909B13998BD8D075D3A2972B">Macharoenboon, Kongkit</mods:namePart>
</mods:name>
<mods:name id="E96C8D34AADE0A240083D5A251629BAE" type="personal">
<mods:role id="3180F0C1B2D4A7209760E502EB9FDACC">
<mods:roleTerm id="54AD5A46C48217CD37B23AA301693220">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A6CB12FA07C2E1030B6420AA4A8959FE">, Varitsara Manonai</mods:namePart>
</mods:name>
<mods:name id="1052472E24621549924609E1369F3509" type="personal">
<mods:role id="9DA9312800FF9A03008D31927DE73C51">
<mods:roleTerm id="FF9B5D4003BE4DB186369E1C325CA92D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8B4418B7154451A46836EDE9FD423249">Jeratthitikul,</mods:namePart>
</mods:name>
<mods:name id="06F40B1F9773F1A847A5E89DFA39AD8D" type="personal">
<mods:role id="F8DC703E5AD9A4898E5468A8CE6BACD7">
<mods:roleTerm id="3B8D66C25EEB6875BB64D3952F8237BF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="538F4E39D24D4E9335E001C00B5B18CA">Ekgachai</mods:namePart>
</mods:name>
<mods:typeOfResource id="60A03D1B59EB61655E732851E6C2D0C9">text</mods:typeOfResource>
<mods:relatedItem id="8C5D2DE11E8E31C1D4AB12A0D69E2750" type="host">
<mods:titleInfo id="876BA068822F3E714772BA5EAC547B7D">
<mods:title id="FD5D1331A74C1ADAB3A8182F8B576C75">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="E6E7E62525D9B125A705D88B23182D5B">
<mods:date id="492AEF0662B63CD6F8B07B1735DDDCA2">2024</mods:date>
<mods:detail id="402445A0FFB5A2CC6C3144CCEA3082F7" type="pubDate">
<mods:number id="2980946AC38C9C0CDEB15789A1E1F22C">2024-11-13</mods:number>
</mods:detail>
<mods:detail id="646888352A56C0E1F1A4340CB6D898E1" type="volume">
<mods:number id="620CCFFF8879E225A156E5518B3183D6">72</mods:number>
</mods:detail>
<mods:extent id="B1F87C2A4AFF46C6D8FA16D7CF523215" unit="page">
<mods:start id="4C77728D69D253D4E126615ABE0A956B">450</mods:start>
<mods:end id="9C991B349164DDB678F02BEF53DD94E9">468</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="B29C5FBAE29FE14C850D99973403AB4A">journal article</mods:classification>
<mods:identifier id="96C5959A6942AC0C83931DC102CFC788" type="DOI">10.26107/RBZ-2024-0033</mods:identifier>
<mods:identifier id="EBF9F1DF91F78A855E657C447A846A01" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F093B303FAC4ECB9C9522BC464B614E5" type="Zenodo-Dep">14682896</mods:identifier>
<mods:identifier id="73504404F1B4903A146A4B9CEFD98AE5" type="ZooBank">56BAA238-5F15-4BE9-933C-0777E8C08C0B</mods:identifier>
</mods:mods>
<treatment id="03EFD249FF98FFB8FF41FD4B34E4FACF" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF" httpUri="http://treatment.plazi.org/id/03EFD249FF98FFB8FF41FD4B34E4FACF" lastPageId="14" lastPageNumber="464" pageId="11" pageNumber="461">
<subSubSection id="C35C30D4FF98FFBDFF41FD4B3481FCAF" box="[251,660,765,789]" pageId="11" pageNumber="461" type="nomenclature">
<paragraph id="8BF9635FFF98FFBDFF41FD4B3481FCAF" blockId="11.[251,660,765,821]" box="[251,660,765,789]" pageId="11" pageNumber="461">
<heading id="D0B1D433FF98FFBDFF41FD4B3481FCAF" bold="true" box="[251,660,765,789]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="461" reason="2">
<taxonomicName id="4C4618DCFF98FFBDFF41FD4B3481FCAF" ID-CoL="8VKFZ" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[251,660,765,789]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF98FFBDFF41FD4B3481FCAF" bold="true" box="[251,660,765,789]" pageId="11" pageNumber="461">
<emphasis id="B932BF4DFF98FFBDFF41FD4B37FFFCAF" bold="true" box="[251,490,765,789]" italics="true" pageId="11" pageNumber="461">Caridina sumatrensis</emphasis>
<bibRefCitation id="EFD71EAEFF98FFBDFE48FD4B3481FCAF" author="De Man JG" box="[498,660,765,789]" pageId="11" pageNumber="461" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C30D4FF98FFBDFED1FCAB3436FC8F" box="[363,547,797,821]" pageId="11" pageNumber="461" type="description">
<paragraph id="8BF9635FFF98FFBDFED1FCAB3436FC8F" blockId="11.[251,660,765,821]" box="[363,547,797,821]" pageId="11" pageNumber="461">
(
<figureCitation id="137D7FDAFF98FFBDFEC8FCAB3415FC8F" box="[370,512,797,821]" captionStart="Fig" captionStartId="8.[113,148,1079,1101]" captionTargetBox="[113,1451,188,1063]" captionTargetId="figure-58@8.[113,1451,188,1063]" captionTargetPageId="8" captionText="Fig. 4. Living habitus specimens of Caridina species from the Mae Klong Basin. AD, Caridina maeklongensis, new species: A, paratype male, MUMNH-CAR064-4; B, paratype female, MUMNH-CAR064-1; C, paratype female, MUMNH-CAR064-15; and D, paratype female, MUMNH-CAR064-13. E, F, C. sumatrensis: E, male, MUMNH-CAR706-P2; and F, female, MUMNH-CAR706-P1." pageId="11" pageNumber="461">Figure 4E, F</figureCitation>
,
<figureCitation id="137D7FDAFF98FFBDFDB7FCAB340EFC8F" box="[525,539,797,821]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="11" pageNumber="461">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C30D4FF98FFBDFF32FCEA3434FB2B" pageId="11" pageNumber="461" type="reference_group">
<paragraph id="8BF9635FFF98FFBDFF32FCEA3759FC5E" blockId="11.[136,776,860,1170]" pageId="11" pageNumber="461">
<treatmentCitationGroup id="AB564471FF98FFBDFF32FCEA3759FC5E" pageId="11" pageNumber="461">
<treatmentCitation id="0AE7454EFF98FFBDFF32FCEA3483FCC8" author="De Man JG" box="[136,662,860,882]" page="375" pageId="11" pageNumber="461" year="1892">
<taxonomicName id="4C4618DCFF98FFBDFF32FCEA3483FCC8" ID-CoL="8VKFZ" authority="De Man, 1892: 375" authorityName="De Man" authorityPageNumber="375" authorityYear="1892" box="[136,662,860,882]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="variety" species="weberi" variety="sumatrensis">
<emphasis id="B932BF4DFF98FFBDFF32FCEA3732FCCB" box="[136,295,860,881]" italics="true" pageId="11" pageNumber="461">Caridina weberi</emphasis>
var.
<emphasis id="B932BF4DFF98FFBDFEE1FCEB37DBFCC8" box="[347,462,861,882]" italics="true" pageId="11" pageNumber="461">sumatrensis</emphasis>
<bibRefCitation id="EFD71EAEFF98FFBDFE6FFCEA3483FCC8" author="De Man JG" box="[469,662,860,882]" pageId="11" pageNumber="461" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892: 375</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 22, fig.
<quantity id="4CBECEBAFF98FFBDFF14FCCF36C1FC34" box="[174,212,889,910]" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" pageId="11" pageNumber="461" unit="g" value="23.0">23g</quantity>
, Type locality: Deli,
<collectingRegion id="4982ADBDFF98FFBDFE16FCCF37EAFC34" box="[428,511,889,910]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="461">Sumatra</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFDB1FCCF347EFC35" box="[523,619,889,911]" name="Indonesia" pageId="11" pageNumber="461">Indonesia</collectingCountry>
, Type material:
<typeStatus id="54FDDDFDFF98FFBDFF14FC203705FC11" box="[174,272,918,939]" pageId="11" pageNumber="461" type="lectotype">Lectotype</typeStatus>
, ZMA De 102603, ovigerous female (CL
<quantity id="4CBECEBAFF98FFBDFD09FC233515FC11" box="[691,768,917,939]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="11" pageNumber="461" unit="mm" value="5.3">5.3 mm</quantity>
) (designated by
<bibRefCitation id="EFD71EAEFF98FFBDFEF6FC043429FC7D" author="Cai Y &amp; Naiyanetr P" box="[332,572,946,968]" pageId="11" pageNumber="461" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
).
<treatmentCitation id="0AE7454EFF98FFBDFDF5FC053516FC7D" author="Kemp S" box="[591,771,946,968]" page="99" pageId="11" pageNumber="461" year="1918">
<bibRefCitation id="EFD71EAEFF98FFBDFDF5FC053516FC7D" author="Kemp S" box="[591,771,946,968]" pageId="11" pageNumber="461" pagination="81 - 102" refId="ref14666" refString="Kemp S (1918 a) Crustacea Decapoda of the Inle Lake Basin. Records of the Indian Museum, 14: 81 - 102." type="journal article" year="1918">Kemp, 1918a: 99</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE7454EFF98FFBDFF14FC79375DFC5E" author="Pillai RS" box="[174,328,975,997]" page="43" pageId="11" pageNumber="461" year="1964">
<bibRefCitation id="EFD71EAEFF98FFBDFF14FC79375DFC5E" author="Pillai RS" box="[174,328,975,997]" pageId="11" pageNumber="461" pagination="42 - 47" refId="ref15689" refString="Pillai RS (1964) Four species of Caridina from Travacore, including of a new variety. Journal of the Marine Biological Association of India, 6 (1): 42 - 47." type="journal article" year="1964">Pillai, 1964: 43</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF9635FFF98FFBDFF32FC5A3439FB81" blockId="11.[136,776,860,1170]" pageId="11" pageNumber="461">
<treatmentCitationGroup id="AB564471FF98FFBDFF32FC5A3439FB81" pageId="11" pageNumber="461">
<taxonomicName id="4C4618DCFF98FFBDFF32FC5A37A6FBB8" ID-CoL="8VKFZ" authorityName="De Man" authorityYear="1892" box="[136,435,1004,1026]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="subSpecies" species="weberi" subSpecies="sumatrensis">
<emphasis id="B932BF4DFF98FFBDFF32FC5A37A6FBB8" box="[136,435,1004,1026]" italics="true" pageId="11" pageNumber="461">Caridina weberi sumatrensis</emphasis>
</taxonomicName>
<treatmentCitation id="0AE7454EFF98FFBDFE6EFC5A34B4FBBB" author="Bouvier EL" box="[468,673,1004,1026]" page="132" pageId="11" pageNumber="461" year="1904">
<bibRefCitation id="EFD71EAEFF98FFBDFE6EFC5A34B4FBBB" author="Bouvier EL" box="[468,673,1004,1026]" pageId="11" pageNumber="461" pagination="129 - 138" refId="ref12797" refString="Bouvier EL (1904) Crevettes de la famille de Atyides: Especes qui font partie des collections du Museum d'histoire Naturelle. Bulletin du Museum d'Histoire Naturelle, 10: 129 - 138." type="journal article" year="1904">Bouvier, 1904: 132</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE7454EFF98FFBDFD15FC5A3711FBA5" author="Bouvier EL" page="75" pageId="11" pageNumber="461" year="1905">
<bibRefCitation id="EFD71EAEFF98FFBDFD15FC5A3711FBA5" author="Bouvier EL" pageId="11" pageNumber="461" pagination="57 - 134" refId="ref12832" refString="Bouvier EL (1905) Observations nouvelles sur les crevettes de la famille des Atyides. Bulletin Scientifique de la France et de la Belgique, 39: 57 - 134." type="journal article" year="1905">Bouvier, 1905: 75</bibRefCitation>
</treatmentCitation>
, 83;
<treatmentCitation id="0AE7454EFF98FFBDFE89FBBF37FDFBA4" author="Kemp S" box="[307,488,1033,1055]" page="292" pageId="11" pageNumber="461" year="1918">
<bibRefCitation id="EFD71EAEFF98FFBDFE89FBBF37FDFBA4" author="Kemp S" box="[307,488,1033,1055]" pageId="11" pageNumber="461" refId="ref14692" refString="Kemp S (1918 b) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. In: Annandale N (ed.) Zoological results of a tour in the Far East. Memoirs of the Asiatic Society of Bengal. Volume 6. The Asiatic Society, Calcutta, pp. 217 - 297." type="book" year="1918">Kemp, 1918b: 292</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AE7454EFF98FFBDFE4BFBBF34B8FBA4" author="Bouvier EL" box="[497,685,1033,1055]" page="247" pageId="11" pageNumber="461" year="1925">
<bibRefCitation id="EFD71EAEFF98FFBDFE4BFBBF34B8FBA4" author="Bouvier EL" box="[497,685,1033,1055]" pageId="11" pageNumber="461" pagination="1 - 370" refId="ref12928" refString="Bouvier EL (1925) Recherches sur la morphologie, les variations, la distribution geographique des crevettes des la famille des Atyides. Encyclopedie Entomologique, 4: 1 - 370." type="journal article" year="1925">Bouvier, 1925: 247</bibRefCitation>
</treatmentCitation>
, fig. 567;
<treatmentCitation id="0AE7454EFF98FFBDFF14FB933771FB81" author="Johnson DS" box="[174,356,1061,1083]" page="46" pageId="11" pageNumber="461" year="1961">
<bibRefCitation id="EFD71EAEFF98FFBDFF14FB933771FB81" author="Johnson DS" box="[174,356,1061,1083]" pageId="11" pageNumber="461" pagination="44 - 79" refId="ref14466" refString="Johnson DS (1961) A synopsis of the decapoda Caridea and Stenopodidea of Singapore, with notes on their distribution and a key to the genera of Caridea occurring in Malayan waters. Bulletin of the National Museum of Singapore, 30: 44 - 79." type="journal article" year="1961">Johnson, 1961: 46</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="EFD71EAEFF98FFBDFED5FB93343DFB81" author="Kazmi Q &amp; Siddiqui FA &amp; Kazmi MA" box="[367,552,1061,1083]" pageId="11" pageNumber="461" pagination="43 - 46" refId="ref14618" refString="Kazmi Q, Siddiqui FA &amp; Kazmi MA (2002) Range extension of Caridina weberi sumatrensis De Man, 1892 (Decapoda, Caridea, Atyidae) to Sindh waters (Karachi, Pakistan). Turkish Journal of Zoology, 26: 43 - 46." type="journal article" year="2002">Kazmi et al., 2002</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF9635FFF98FFBDFF32FBF43434FB2B" blockId="11.[136,776,860,1170]" pageId="11" pageNumber="461">
<treatmentCitationGroup id="AB564471FF98FFBDFF32FBF43434FB2B" pageId="11" pageNumber="461">
<taxonomicName id="4C4618DCFF98FFBDFF32FBF4374EFBE2" ID-CoL="8VKFZ" authorityName="De Man" authorityYear="1892" box="[136,347,1090,1112]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF98FFBDFF32FBF4374EFBE2" box="[136,347,1090,1112]" italics="true" pageId="11" pageNumber="461">Caridina sumatrensis</emphasis>
</taxonomicName>
<treatmentCitation id="0AE7454EFF98FFBDFEC2FBF5347BFBED" author="Wowor D &amp; Cai Y &amp; Ng PKL" box="[376,622,1090,1112]" page="343" pageId="11" pageNumber="461" year="2004">
<bibRefCitation id="EFD71EAEFF98FFBDFEC2FBF5347BFBED" author="Wowor D &amp; Cai Y &amp; Ng PKL" box="[376,622,1090,1112]" pageId="11" pageNumber="461" pagination="337 - 357" refId="ref16444" refString="Wowor D, Cai Y &amp; Ng PKL (2004) Crustacea: Decapoda, Caridea. In: Yule CM &amp; Sen YH (eds.) The Freshwater Invertebrates of the Malaysian Region. Academy of Sciences Malaysia, Kuala Lumpur, pp. 337 - 357." type="book chapter" year="2004">Wowor et al., 2004: 343</bibRefCitation>
</treatmentCitation>
, fig. 6p, q;
<treatmentCitation id="0AE7454EFF98FFBDFD5FFBF43796FBCF" author="Cai Y &amp; Shokita S" page="246" pageId="11" pageNumber="461" year="2006">
<bibRefCitation id="EFD71EAEFF98FFBDFD5FFBF43796FBCF" author="Cai Y &amp; Shokita S" pageId="11" pageNumber="461" pagination="245 - 270" refId="ref13226" refString="Cai Y &amp; Shokita S (2006) Report on a collection of freshwater shrimps (Crustacea: Decapoda: Caridea) from the Philippines, with descriptions of four new species. Raffles Bulletin of Zoology, 54 (2): 245 - 270." type="journal article" year="2006">Cai &amp; Shokita, 2006: 246</bibRefCitation>
</treatmentCitation>
, fig. 6CF; Cai et al., 2007: 285, fig. 7;
<treatmentCitation id="0AE7454EFF98FFBDFF14FBCA37D7FB2B" author="Cai Y &amp; Naiyanetr P" box="[174,450,1148,1170]" page="774" pageId="11" pageNumber="461" year="2024">
<bibRefCitation id="EFD71EAEFF98FFBDFF14FBCA37D7FB2B" author="Cai Y &amp; Naiyanetr P" box="[174,450,1148,1170]" pageId="11" pageNumber="461" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024: 774</bibRefCitation>
</treatmentCitation>
, fig. 2, 3.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35C30D4FF98FFBDFF32FB0C37FBF8E8" pageId="11" pageNumber="461" type="materials_examined">
<paragraph id="8BF9635FFF98FFBDFF32FB0C37FBF8E8" blockId="11.[135,775,1210,1874]" pageId="11" pageNumber="461">
<emphasis id="B932BF4DFF98FFBDFF32FB0C376BFB68" bold="true" box="[136,382,1210,1234]" pageId="11" pageNumber="461">Materials examined.</emphasis>
<specimenCount id="9D40A8D6FF98FFBDFE33FB0C37E2FB68" box="[393,503,1210,1234]" count="9" pageId="11" pageNumber="461" type="female">9 females</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFD83FB0C34F7FB68" box="[569,738,1210,1234]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" metricValueMax="4.49" metricValueMin="3.91" pageId="11" pageNumber="461" unit="mm" value="4.2" valueMax="4.49" valueMin="3.91">3.914.49 mm</quantity>
),
<materialsCitation id="3B2E6902FF98FFBDFD43FB0C3516FAE8" collectingDate="2019-04-19" collectorName="W. Siriwut &amp; W. Manonai" country="Thailand" latitude="13.8679" location="Mae Klong River" longLatPrecision="7" longitude="99.8167" municipality="Tha Maka District" pageId="11" pageNumber="461" specimenCode="MUMNH-CAR053" specimenCount="3" specimenCount-male="3" stateProvince="Kanchanaburi">
<specimenCount id="9D40A8D6FF98FFBDFD43FB0C36DDFB48" count="3" pageId="11" pageNumber="461" type="male">3 males</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFEB0FB6C37A5FB48" box="[266,432,1242,1266]" metricMagnitude="-3" metricUnit="m" metricValue="2.7249999999999996" metricValueMax="2.94" metricValueMin="2.51" pageId="11" pageNumber="461" unit="mm" value="2.7249999999999996" valueMax="2.94" valueMin="2.51">2.512.94 mm</quantity>
)(
<specimenCode id="DBE0CB24FF98FFBDFE04FB6C348FFB48" box="[446,666,1242,1266]" collectionCode="MUMNH-CAR" pageId="11" pageNumber="461">MUMNH-CAR053</specimenCode>
, used for measurement in description),
<location id="8E993584FF98FFBDFE71FB4D3499FAA8" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF:8E993584FF98FFBDFE71FB4D3499FAA8" box="[459,652,1274,1298]" country="Thailand" latitude="13.8679" longLatPrecision="7" longitude="99.8167" municipality="Tha Maka District" name="Mae Klong River" pageId="11" pageNumber="461" stateProvince="Kanchanaburi">Mae Klong River</location>
,
<collectingMunicipality id="6B9DF925FF98FFBDFD22FB4C36F7FA88" pageId="11" pageNumber="461">Tha Maka District</collectingMunicipality>
,
<collectingRegion id="4982ADBDFF98FFBDFF48FAAC3414FA88" box="[242,513,1306,1330]" country="Thailand" name="Kanchanaburi" pageId="11" pageNumber="461">Kanchanaburi Province</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFDABFAAC346DFA88" box="[529,632,1306,1330]" name="Thailand" pageId="11" pageNumber="461">Thailand</collectingCountry>
,
<geoCoordinate id="EE720598FF98FFBDFD3DFAAC3516FA88" box="[647,771,1306,1330]" degrees="13.8679" direction="north" orientation="latitude" pageId="11" pageNumber="461" precision="5" value="13.8679">13.8679°N</geoCoordinate>
,
<geoCoordinate id="EE720598FF98FFBDFF32FA8C36E8FAE8" box="[136,253,1338,1362]" degrees="99.8167" direction="east" orientation="longitude" pageId="11" pageNumber="461" precision="5" value="99.8167">99.8167°E</geoCoordinate>
, coll.
<collectorName id="26B30689FF98FFBDFE87FA8D37A6FAE8" box="[317,435,1338,1362]" pageId="11" pageNumber="461">W. Siriwut</collectorName>
&amp;
<collectorName id="26B30689FF98FFBDFE6FFA8D344BFAE8" box="[469,606,1338,1362]" pageId="11" pageNumber="461">W. Manonai</collectorName>
,
<date id="FFF8459FFF98FFBDFDD3FA8C3516FAE8" box="[617,771,1338,1362]" pageId="11" pageNumber="461" value="2019-04-19">
<collectingDate id="EFBCBC77FF98FFBDFDD3FA8C3516FAE8" box="[617,771,1338,1362]" pageId="11" pageNumber="461" value="2019-04-19">19 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2E6902FF98FFBDFF32FAEC373CFA68" collectingDate="2019-04-19" collectorName="W. Siriwut &amp; W. Manonai" country="Thailand" latitude="13.6318" location="Mae Klong River" longLatPrecision="7" longitude="99.8162" municipality="Photharam District" pageId="11" pageNumber="461" specimenCode="MUMNH-CAR054" specimenCount="1" specimenCount-female="1" stateProvince="Ratchaburi">
<specimenCount id="9D40A8D6FF98FFBDFF32FAEC36FCFAC8" box="[136,233,1370,1394]" count="1" pageId="11" pageNumber="461" type="female">1 female</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFE93FAEC3798FAC8" box="[297,397,1370,1394]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="461" unit="mm" value="4.1">4.10 mm</quantity>
)(
<specimenCode id="DBE0CB24FF98FFBDFE26FAEC3460FAC8" box="[412,629,1370,1394]" collectionCode="MUMNH-CAR" pageId="11" pageNumber="461">MUMNH-CAR054</specimenCode>
),
<location id="8E993584FF98FFBDFD30FAED36D3FA28" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF:8E993584FF98FFBDFD30FAED36D3FA28" country="Thailand" latitude="13.6318" longLatPrecision="7" longitude="99.8162" municipality="Photharam District" name="Mae Klong River" pageId="11" pageNumber="461" stateProvince="Ratchaburi">Mae Klong River</location>
,
<collectingMunicipality id="6B9DF925FF98FFBDFF6EFACC37B0FA28" box="[212,421,1402,1426]" pageId="11" pageNumber="461">Photharam District</collectingMunicipality>
,
<collectingRegion id="4982ADBDFF98FFBDFE09FACC3481FA28" box="[435,660,1402,1426]" country="Thailand" name="Ratchaburi" pageId="11" pageNumber="461">Ratchaburi Province</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFD1BFACC3516FA28" box="[673,771,1402,1426]" name="Thailand" pageId="11" pageNumber="461">Thailand</collectingCountry>
,
<geoCoordinate id="EE720598FF98FFBDFF32FA2C3717FA08" box="[136,258,1434,1458]" degrees="13.6318" direction="north" orientation="latitude" pageId="11" pageNumber="461" precision="5" value="13.6318">13.6318°N</geoCoordinate>
,
<geoCoordinate id="EE720598FF98FFBDFEAAFA2C3792FA08" box="[272,391,1434,1458]" degrees="99.8162" direction="east" orientation="longitude" pageId="11" pageNumber="461" precision="5" value="99.8162">99.8162°E</geoCoordinate>
, coll.
<collectorName id="26B30689FF98FFBDFE6AFA2D345FFA08" box="[464,586,1434,1458]" pageId="11" pageNumber="461">W. Siriwut</collectorName>
&amp;
<collectorName id="26B30689FF98FFBDFDCEFA2D3517FA08" box="[628,770,1434,1458]" pageId="11" pageNumber="461">W. Manonai</collectorName>
,
<date id="FFF8459FFF98FFBDFF32FA0C373CFA68" box="[136,297,1466,1490]" pageId="11" pageNumber="461" value="2019-04-19">
<collectingDate id="EFBCBC77FF98FFBDFF32FA0C373CFA68" box="[136,297,1466,1490]" pageId="11" pageNumber="461" value="2019-04-19">19 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2E6902FF98FFBDFE82FA0C37F7F988" collectingDate="2019-04-19" collectorName="W. Siriwut &amp; W. Manonai" country="Thailand" latitude="13.7152" location="Mae Klong River" longLatPrecision="7" longitude="99.8492" municipality="Photharam District" pageId="11" pageNumber="461" specimenCode="MUMNH-CAR055" specimenCount="4" specimenCount-female="4" stateProvince="Ratchaburi">
<specimenCount id="9D40A8D6FF98FFBDFE82FA0C37B6FA68" box="[312,419,1466,1490]" count="4" pageId="11" pageNumber="461" type="female">4 females</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFE59FA0C3491FA68" box="[483,644,1466,1490]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.21" metricValueMin="3.29" pageId="11" pageNumber="461" unit="mm" value="3.75" valueMax="4.21" valueMin="3.29">3.294.21 mm</quantity>
)(
<specimenCode id="DBE0CB24FF98FFBDFD28FA0C36FEFA48" collectionCode="MUMNH-CAR" pageId="11" pageNumber="461">MUMNH-CAR055</specimenCode>
),
<location id="8E993584FF98FFBDFF41FA6D37A0FA48" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF:8E993584FF98FFBDFF41FA6D37A0FA48" box="[251,437,1498,1522]" country="Thailand" latitude="13.7152" longLatPrecision="7" longitude="99.8492" municipality="Photharam District" name="Mae Klong River" pageId="11" pageNumber="461" stateProvince="Ratchaburi">Mae Klong River</location>
,
<collectingMunicipality id="6B9DF925FF98FFBDFE04FA6C349CFA48" box="[446,649,1498,1522]" pageId="11" pageNumber="461">Photharam District</collectingMunicipality>
,
<collectingRegion id="4982ADBDFF98FFBDFD28FA6C36FCF9A8" country="Thailand" name="Ratchaburi" pageId="11" pageNumber="461">Ratchaburi Province</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFF48FA4C3741F9A8" box="[242,340,1530,1554]" name="Thailand" pageId="11" pageNumber="461">Thailand</collectingCountry>
,
<geoCoordinate id="EE720598FF98FFBDFEE7FA4C37C1F9A8" box="[349,468,1530,1554]" degrees="13.7152" direction="north" orientation="latitude" pageId="11" pageNumber="461" precision="5" value="13.7152">13.7152°N</geoCoordinate>
,
<geoCoordinate id="EE720598FF98FFBDFE64FA4C3446F9A8" box="[478,595,1530,1554]" degrees="99.8492" direction="east" orientation="longitude" pageId="11" pageNumber="461" precision="5" value="99.8492">99.8492°E</geoCoordinate>
, coll.
<collectorName id="26B30689FF98FFBDFD2BFA4D3513F9A8" box="[657,774,1530,1554]" pageId="11" pageNumber="461">W. Siriwut</collectorName>
&amp;
<collectorName id="26B30689FF98FFBDFF1CF9AD3727F988" box="[166,306,1562,1586]" pageId="11" pageNumber="461">W. Manonai</collectorName>
,
<date id="FFF8459FFF98FFBDFEFBF9AC37F7F988" box="[321,482,1562,1586]" pageId="11" pageNumber="461" value="2019-04-19">
<collectingDate id="EFBCBC77FF98FFBDFEFBF9AC37F7F988" box="[321,482,1562,1586]" pageId="11" pageNumber="461" value="2019-04-19">19 April 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2E6902FF98FFBDFE4AF9AC3516F928" collectingDate="2019-04-19" collectorName="W. Siriwut &amp; W. Manonai" country="Thailand" latitude="13.7393" location="Mae Klong River" longLatPrecision="7" longitude="99.842" municipality="Photharam District" pageId="11" pageNumber="461" specimenCode="MUMNH-CAR056" specimenCount="3" specimenCount-female="3" stateProvince="Ratchaburi">
<specimenCount id="9D40A8D6FF98FFBDFE4AF9AC3449F988" box="[496,604,1562,1586]" count="3" pageId="11" pageNumber="461" type="female">3 females</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFD21F9AC36A1F9E8" metricMagnitude="-3" metricUnit="m" metricValue="4.01" metricValueMax="4.29" metricValueMin="3.73" pageId="11" pageNumber="461" unit="mm" value="4.01" valueMax="4.29" valueMin="3.73">3.734.29 mm</quantity>
)(
<specimenCode id="DBE0CB24FF98FFBDFF79F98C37B5F9E8" box="[195,416,1594,1618]" collectionCode="MUMNH-CAR" pageId="11" pageNumber="461">MUMNH-CAR056</specimenCode>
),
<location id="8E993584FF98FFBDFE0CF98D3494F9E8" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF:8E993584FF98FFBDFE0CF98D3494F9E8" box="[438,641,1594,1618]" country="Thailand" latitude="13.7393" longLatPrecision="7" longitude="99.842" municipality="Photharam District" name="Mae Klong River" pageId="11" pageNumber="461" stateProvince="Ratchaburi">Mae Klong River</location>
,
<collectingMunicipality id="6B9DF925FF98FFBDFD2AF98C36F0F9C8" pageId="11" pageNumber="461">Photharam District</collectingMunicipality>
,
<collectingRegion id="4982ADBDFF98FFBDFF4DF9EC37E4F9C8" box="[247,497,1626,1650]" country="Thailand" name="Ratchaburi" pageId="11" pageNumber="461">Ratchaburi Province</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFDB8F9EC347BF9C8" box="[514,622,1626,1650]" name="Thailand" pageId="11" pageNumber="461">Thailand</collectingCountry>
,
<geoCoordinate id="EE720598FF98FFBDFDC5F9EC3514F9C8" box="[639,769,1626,1650]" degrees="13.7393" direction="north" orientation="latitude" pageId="11" pageNumber="461" precision="5" value="13.7393">13.7393°N</geoCoordinate>
,
<geoCoordinate id="EE720598FF98FFBDFF32F9CC36E8F928" box="[136,253,1658,1682]" degrees="99.8420" direction="east" orientation="longitude" pageId="11" pageNumber="461" precision="5" value="99.842">99.8420°E</geoCoordinate>
, coll.
<collectorName id="26B30689FF98FFBDFE87F9CD37A6F928" box="[317,435,1658,1682]" pageId="11" pageNumber="461">W. Siriwut</collectorName>
&amp;
<collectorName id="26B30689FF98FFBDFE6FF9CD344BF928" box="[469,606,1658,1682]" pageId="11" pageNumber="461">W. Manonai</collectorName>
,
<date id="FFF8459FFF98FFBDFDD3F9CC3516F928" box="[617,771,1658,1682]" pageId="11" pageNumber="461" value="2019-04-19">
<collectingDate id="EFBCBC77FF98FFBDFDD3F9CC3516F928" box="[617,771,1658,1682]" pageId="11" pageNumber="461" value="2019-04-19">19 April 2019</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D40A8D6FF98FFBDFF32F92C36EFF908" box="[136,250,1690,1714]" count="4" pageId="11" pageNumber="461" type="female">4 females</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFE84F92C37FDF908" box="[318,488,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="3.825" metricValueMax="4.08" metricValueMin="3.57" pageId="11" pageNumber="461" unit="mm" value="3.825" valueMax="4.08" valueMin="3.57">3.574.08 mm</quantity>
) (used for measurement in description);
<materialsCitation id="3B2E6902FF98FFBDFE87F90C37FFF8E8" collectingDate="2019-08-08" collectorName="E. Jeratthitikul &amp; K. Macharoenboon" country="Thailand" latitude="13.6537" location="Klong Bang Tanod River" longLatPrecision="7" longitude="99.8116" municipality="Photharam District" pageId="11" pageNumber="461" specimenCode="MUMNH-CAR706" specimenCount="4" specimenCount-male="4" stateProvince="Ratchaburi">
<specimenCount id="9D40A8D6FF98FFBDFE87F90C3781F968" box="[317,404,1722,1746]" count="4" pageId="11" pageNumber="461" type="male">4 males</specimenCount>
(CL
<quantity id="4CBECEBAFF98FFBDFE6FF90C346DF968" box="[469,632,1722,1746]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.93" metricValueMin="2.41" pageId="11" pageNumber="461" unit="mm" value="2.67" valueMax="2.93" valueMin="2.41">2.412.93 mm</quantity>
) (
<specimenCode id="DBE0CB24FF98FFBDFD2BF90C36F8F948" collectionCode="MUMNH-CAR" pageId="11" pageNumber="461">MUMNH-CAR706</specimenCode>
, used for measurement in description),
<location id="8E993584FF98FFBDFD7BF96C3759F8A8" LSID="urn:lsid:plazi:treatment:03EFD249FF98FFB8FF41FD4B34E4FACF:8E993584FF98FFBDFD7BF96C3759F8A8" country="Thailand" latitude="13.6537" longLatPrecision="7" longitude="99.8116" municipality="Photharam District" name="Klong Bang Tanod River" pageId="11" pageNumber="461" stateProvince="Ratchaburi">Klong Bang Tanod River</location>
,
<collectingMunicipality id="6B9DF925FF98FFBDFEEFF94C340AF8A8" box="[341,543,1786,1810]" pageId="11" pageNumber="461">Photharam District</collectingMunicipality>
,
<collectingRegion id="4982ADBDFF98FFBDFD92F94C3516F8A8" box="[552,771,1786,1810]" country="Thailand" name="Ratchaburi" pageId="11" pageNumber="461">Ratchaburi Province</collectingRegion>
,
<collectingCountry id="F35123CFFF98FFBDFF3DF8AC36FCF888" box="[135,233,1818,1842]" name="Thailand" pageId="11" pageNumber="461">Thailand</collectingCountry>
,
<geoCoordinate id="EE720598FF98FFBDFF49F8AC377FF888" box="[243,362,1818,1842]" degrees="13.6537" direction="north" orientation="latitude" pageId="11" pageNumber="461" precision="5" value="13.6537">13.6537°N</geoCoordinate>
,
<geoCoordinate id="EE720598FF98FFBDFECCF8AC37FFF888" box="[374,490,1818,1842]" degrees="99.8116" direction="east" orientation="longitude" pageId="11" pageNumber="461" precision="5" value="99.8116">99.8116°E</geoCoordinate>
, coll.
<collectorName id="26B30689FF98FFBDFD91F8AD34DDF888" box="[555,712,1818,1842]" pageId="11" pageNumber="461">E. Jeratthitikul</collectorName>
&amp;
<collectorName id="26B30689FF98FFBDFD51F8AD372CF8E8" pageId="11" pageNumber="461">K. Macharoenboon</collectorName>
,
<date id="FFF8459FFF98FFBDFEFFF88C37FFF8E8" box="[325,490,1850,1874]" pageId="11" pageNumber="461" value="2019-08-08">
<collectingDate id="EFBCBC77FF98FFBDFEFFF88C37FFF8E8" box="[325,490,1850,1874]" pageId="11" pageNumber="461" value="2019-08-08">8 August 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF3933D7FF98FFBDFCF9FB6D3247FAFD" pageId="11" pageNumber="461" startId="11.[835,870,1243,1265]" targetBox="[838,1475,189,1226]" targetPageId="11" targetType="figure">
<paragraph id="8BF9635FFF98FFBDFCF9FB6D3247FAFD" blockId="11.[835,1475,1243,1351]" pageId="11" pageNumber="461">
Fig. 5. Map of the Mae Klong Basin showing sampling localities of
<taxonomicName id="4C4618DCFF98FFBDFCDBFB4E3241FAB7" authorityName="Macharoenboon &amp; &amp; Jeratthitikul &amp; Ekgachai" authorityYear="2024" box="[865,1108,1272,1293]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="species" species="maeklongensis" status="sp. nov.">
<emphasis id="B932BF4DFF98FFBDFCDBFB4E3241FAB7" box="[865,1108,1272,1293]" italics="true" pageId="11" pageNumber="461">Caridina maeklongensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2010236FF98FFBDFBDEFB4E32C8FAB4" box="[1124,1245,1272,1294]" pageId="11" pageNumber="461" rank="species">new species</taxonomicNameLabel>
(black circles) and
<taxonomicName id="4C4618DCFF98FFBDFA16FB4E35ADFA90" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="461" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF98FFBDFA16FB4E35ADFA90" italics="true" pageId="11" pageNumber="461">C. sumatrensis</emphasis>
</taxonomicName>
(grey circle), and the localities where both species were absent (white circles).
</paragraph>
</caption>
<subSubSection id="C35C30D4FF98FFBBFF32F8CC3276FAAC" lastPageId="13" lastPageNumber="463" pageId="11" pageNumber="461" type="description">
<paragraph id="8BF9635FFF98FFBDFF32F8CC3512F788" blockId="11.[136,777,1914,2099]" pageId="11" pageNumber="461">
<emphasis id="B932BF4DFF98FFBDFF32F8CC3512F828" bold="true" box="[136,775,1914,1938]" pageId="11" pageNumber="461">Description. Cephalothorax and cephalic appendage.</emphasis>
CL
<quantity id="4CBECEBAFF98FFBDFF0CF82C374FF808" box="[182,346,1946,1970]" metricMagnitude="-3" metricUnit="m" metricValue="4.085" metricValueMax="4.6" metricValueMin="3.57" pageId="11" pageNumber="461" unit="mm" value="4.085" valueMax="4.6" valueMin="3.57">3.574.60 mm</quantity>
(median =
<quantity id="4CBECEBAFF98FFBDFE5CF82C345AF808" box="[486,591,1946,1970]" metricMagnitude="-3" metricUnit="m" metricValue="4.08" pageId="11" pageNumber="461" unit="mm" value="4.08">4.08 mm</quantity>
, n = 13), width
<quantity id="4CBECEBAFF98FFBDFF32F80C3731F868" box="[136,292,1978,2002]" metricMagnitude="-3" metricUnit="m" metricValue="3.065" metricValueMax="3.48" metricValueMin="2.65" pageId="11" pageNumber="461" unit="mm" value="3.065" valueMax="3.48" valueMin="2.65">2.653.48 mm</quantity>
(median =
<quantity id="4CBECEBAFF98FFBDFE1EF80C3412F868" box="[420,519,1978,2002]" metricMagnitude="-3" metricUnit="m" metricValue="2.92" pageId="11" pageNumber="461" unit="mm" value="2.92">2.92 mm</quantity>
, n = 13). Rostrum with basal ridge, slightly bent down, frequently reaching near the end of antennular peduncle (at least reaching beyond the end of second segment of antennular peduncle), 0.500.60
</paragraph>
<paragraph id="8BF9635FFF98FFBDFCFFFAC832E5F8EC" blockId="11.[834,1476,1406,1878]" pageId="11" pageNumber="461">
(median = 0.55, n = 13) times as long as CL (
<figureCitation id="137D7FDAFF98FFBDFAFEFAC833A2FA2D" box="[1348,1463,1406,1431]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="11" pageNumber="461">Fig 6A, B</figureCitation>
). Rostral formula based on 13 individuals: (46) + 1721 / 36. Antennal spine placed below inferior orbital angle. Pterygostomian margin subrectangular. Eye well-developed, anterior end reaching to 0.590.84 (median = 0.71, n = 13) of first segment of antennular peduncle. Antennular peduncle 0.460.55 (median = 0.51, n = 13) times as long as carapace, first segment 1.602.15 (median = 1.80, n = 13) times as long as second segment, second segment 1.031.33 (median = 1.16, n = 13) times as long as third segment. Tooth on distolateral margin of first segment of antennular peduncle prominent. Stylocerite reaching to 0.590.74 (median = 0.70, n = 13) of first segment of antennular peduncle. Scaphocerite 2.502.97 (median = 2.75, n = 13) times as long as wide, distal margin with short plumose setae.
</paragraph>
<paragraph id="8BF9635FFF98FFBBFCF9F8C83477FEAF" blockId="11.[835,1475,1918,2102]" lastBlockId="13.[136,774,188,277]" lastPageId="13" lastPageNumber="463" pageId="11" pageNumber="461">
<emphasis id="B932BF4DFF98FFBDFCF9F8C83209F82C" bold="true" box="[835,1052,1918,1942]" pageId="11" pageNumber="461">Branchial formula.</emphasis>
Podobranch on second maxilliped well-developed. Third maxilliped possesses one small and one large arthrobranch. Pleurobranchs present on all pereopods. Third maxilliped with epipod. Ultimate segment of endopod with a row of strong spinules at proximal ⅔ of posterior margin, ending with one large claw, 57 spiniform setae on distal ⅓ of posterior margin, 0.740.91 (median = 0.83, n = 10) times as long as penultimate segment. Exopod long and slender, with a tuft of long setae at tip.
</paragraph>
<caption id="DF3933D7FF9FFFBAFFCBF8C03508F7A1" pageId="12" pageNumber="462" startId="12.[113,148,1910,1932]" targetBox="[116,1454,229,1901]" targetPageId="12" targetType="figure">
<paragraph id="8BF9635FFF9FFFBAFFCBF8C03508F7A1" blockId="12.[113,1452,1910,2076]" pageId="12" pageNumber="462">
Fig. 6.
<taxonomicName id="4C4618DCFF9FFFBAFF0EF8C03403F831" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[180,534,1910,1932]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="462" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9FFFBAFF0EF8C03791F836" box="[180,388,1910,1932]" italics="true" pageId="12" pageNumber="462">Caridina sumatrensis</emphasis>
<bibRefCitation id="EFD71EAEFF9FFFBAFE33F8C03403F831" author="De Man JG" box="[393,534,1910,1931]" pageId="12" pageNumber="462" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNHCAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BF9635FFF9EFFBBFF32FE8B3513FECF" blockId="13.[136,775,317,373]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFF32FE8B3713FEEF" bold="true" box="[136,262,317,341]" pageId="13" pageNumber="463">Pereopods.</emphasis>
Epipod present on first four pereopods. Chelae of first and second pereopods well-developed (
<figureCitation id="137D7FDAFF9EFFBBFD3CFEEB34ECFECF" box="[646,761,349,373]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6E, F</figureCitation>
).
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFF32FE2B374FFD6F" blockId="13.[136,775,413,725]" pageId="13" pageNumber="463">
First pereopod short (
<figureCitation id="137D7FDAFF9EFFBBFECCFE2B37DEFE0F" box="[374,459,413,437]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6E</figureCitation>
); chela 2.112.66 (median = 2.18, n = 11) times as long as wide, 1.311.62 (median = 1.48, n = 11) times as long as carpus; tips of fingers rounded, with tuft of setae near tip; dactylus 0.911.26 (median = 1.07, n = 11) times as long as palm; carpus excavated distally, 1.401.83 (median = 1.67, n = 11) times as long as wide, 0.781.10 (median = 0.96, n = 11) times as long as merus; merus 2.442.93 (median = 2.62, n = 11) times as long as wide, 1.152.14 (median = 1.84, n = 11) times as long as ischium.
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFF32FD4B3459FB8F" blockId="13.[136,777,765,1077]" pageId="13" pageNumber="463">
Second pereopod more slender than first pereopod (
<figureCitation id="137D7FDAFF9EFFBBFD60FD4B36BDFC8F" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6F</figureCitation>
); chela long, 2.713.40 (median = 2.84, n = 11) times as long as wide, 0.650.87 (median = 0.74, n = 11) times as long as carpus, tips of fingers round, with tuft of setae near tip; dactylus 1.172.20 (median = 1.80, n = 11) times as long as palm; carpus slender, 4.636.69 (median = 5.72, n = 11) times as long as wide, 0.430.75 (median = 0.54, n = 11) times as long as merus; merus 5.007.41 (median = 5.83, n = 11) times as long as wide, 1.242.15 (median = 1.54, n = 11) times as long as ischium.
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFF32FBEB351DF98F" blockId="13.[135,776,1117,1589]" pageId="13" pageNumber="463">
Third pereopod not sexually dimorphic (
<figureCitation id="137D7FDAFF9EFFBBFDFBFBEB348CFBCF" box="[577,665,1117,1141]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6G</figureCitation>
); dactylus with 46 spiniform setae on flexor margin (
<figureCitation id="137D7FDAFF9EFFBBFD21FBCB34EEFB2F" box="[667,763,1149,1173]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6H</figureCitation>
), 2.233.13 (median = 2.65, n = 9) times as long as wide (including terminal claw), terminating with one large claw; propodus with numerous spiniform setae on lateral and posterior margin, 8.4911.53 (median = 9.71, n = 9) times as long as wide, 4.345.67 (median = 4.50, n = 9) times as long as dactylus; carpus with 35 spiniform setae on posterior margin of outer surface, the distal seta largest, the other setae minute, 4.135.36 (median = 4.69, n = 9) times as long as wide, 0.500.69 (median = 0.66, n = 9) times as long as propodus; merus with 45 large spiniform setae on posterior margin of outer surface, 6.256.96 (median = 6.67, n = 9) times as long as wide, 1.882.26 (median = 2.09, n = 9) times as long as carpus; ischium with one spiniform seta.
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFF32F9EB34EFF78F" blockId="13.[136,777,1629,2101]" pageId="13" pageNumber="463">
Fifth pereopod slender (
<figureCitation id="137D7FDAFF9EFFBBFE12F9EB37E8F9CF" box="[424,509,1629,1653]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6I</figureCitation>
); dactylus with 3243 spiniform setae on flexor margin (
<figureCitation id="137D7FDAFF9EFFBBFE4FF9CB344FF92F" box="[501,602,1661,1685]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6J, n</figureCitation>
= 9), 2.763.96 (median = 3.41, n = 9) times as long as wide (including terminal claw), terminating with one large claw; propodus with numerous spiniform setae on posterior margin, 12.3914.45 (median = 13.61, n = 9) times as long as wide, 4.315.79 (median = 4.70, n = 9) times as long as dactylus; carpus with 35 spiniform setae on posterior margin of outer surface, the distal seta largest, the other setae minute, 4.655.78 (median = 5.05, n = 9) times as long as wide, 0.450.56 (median = 0.52, n = 9) times as long as propodus; merus with 24 (mode = 2, n = 9) large setae on posterior margin of outer surface, 6.127.45 (median = 6.91, n = 9) times as long as wide, 1.481.82 (median = 1.61, n = 9) times as long as carpus; ischium without spiniform setae.
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFCF9FF0B32A4FE6F" blockId="13.[834,1476,189,469]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCF9FF0B35A4FF6F" bold="true" box="[835,945,189,213]" pageId="13" pageNumber="463">Pleopods.</emphasis>
Endopod of male first pleopod subtriangular (
<figureCitation id="137D7FDAFF9EFFBBFA2DFF0B3596FF4F" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6O, P</figureCitation>
), wider proximally, 1.732.27 (median = 1.94, n = 7) times as long as width, 0.320.46 (median = 0.41, n = 7) times exopod length, with elongated appendix interna. Appendix masculina of second male pleopod rod-shaped (
<figureCitation id="137D7FDAFF9EFFBBFADFFE8B354DFECF" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6Q, R</figureCitation>
), with numerous setae, 0.740.81 (median = 0.77, n = 7) times as long as endopod (including distal setae). Appendix interna slender, reaching 0.550.64 (median = 0.61, n = 7) times appendix masculina length.
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFCF9FE4B35E7FCCF" blockId="13.[834,1476,509,885]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCF9FE4B35AEFDAF" bold="true" box="[835,955,509,533]" pageId="13" pageNumber="463">Abdomen.</emphasis>
Sixth abdominal somite 0.320.50 (median = 0.43, n = 10) times carapace length, 1.101.54 (median = 1.29, n = 10) times as long as fifth somite, 0.600.99 (median = 0.82, n = 10) times as long as telson (
<figureCitation id="137D7FDAFF9EFFBBFAE4FDEB33A3FDCF" box="[1374,1462,605,629]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6A</figureCitation>
). Telson 2.332.96 (median = 2.71, n = 10) times as long as wide, with 34 pairs of dorsal spiniform setae and one pair of dorsolateral spiniform setae (
<figureCitation id="137D7FDAFF9EFFBBFB18FD0B32EFFD6F" box="[1186,1274,701,725]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6K</figureCitation>
). Distal margin of telson subtriangular, with a posteromedian projection, and 69 moveable plumose setae, subequal in length (
<figureCitation id="137D7FDAFF9EFFBBFADBFD4B33A3FCAF" box="[1377,1462,765,789]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6L</figureCitation>
). Preanal carina subtriangular, high, without a spine (
<figureCitation id="137D7FDAFF9EFFBBFAD9FCAB33A2FC8F" box="[1379,1463,797,821]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6N</figureCitation>
). Uropodal diaeresis with 1522 short moveable spiniform setae (
<figureCitation id="137D7FDAFF9EFFBBFC33FCEB35F3FCCF" box="[905,998,861,885]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6M</figureCitation>
).
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFCF9FC2B35C2FC4F" blockId="13.[835,1474,925,1013]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCF9FC2B3596FC0F" bold="true" box="[835,899,925,949]" pageId="13" pageNumber="463">Eggs.</emphasis>
Ovigerous females with numerous eggs (
<figureCitation id="137D7FDAFF9EFFBBFADCFC2B33A8FC0F" box="[1382,1469,925,949]" captionStart="Fig" captionStartId="7.[136,171,1151,1173]" captionTargetBox="[136,1479,191,1135]" captionTargetId="figure-8@7.[126,1485,180,1154]" captionTargetPageId="7" captionText="Fig. 3. Caridina maeklongensis, new species. A, first pereopod; B, second pereopod; C, third pereopod; D, dactylus of third pereopod; E, fifth pereopod; F, dactylus of fifth pereopod; G, male first pleopod; H, endopod of male first pleopod; I, male second pleopod; J, appendix masculina and appendix interna of male second pleopod. Drawings were made from paratype female, MUMNH-CAR064-7 (AF) and paratype male, MUMNH-CAR064-4 (GJ). Scale bars: D, F, J = 0.1 mm; AC, E, GI = 0.5 mm." pageId="13" pageNumber="463">Fig. 3F</figureCitation>
,
<figureCitation id="137D7FDAFF9EFFBBFCF9FC0B357CFC6F" box="[835,873,957,981]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">6A</figureCitation>
). Size of eye-developed eggs 0.410.46 ×
<quantity id="4CBECEBAFF9EFFBBFAEEFC0B3579FC4F" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="13" pageNumber="463" unit="mm" value="0.265" valueMax="0.29" valueMin="0.24">0.240.29 mm</quantity>
(n = 30).
</paragraph>
<paragraph id="8BF9635FFF9EFFBBFCFFFBAB3276FAAC" blockId="13.[834,1475,1053,1302]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCFFFBAB35F9FB8F" bold="true" box="[837,1004,1053,1077]" pageId="13" pageNumber="463">Colour in life.</emphasis>
Female and male shrimps exhibit distinct colouration. For specimens from the Mae Klong River, male body is translucent, decorated with scattered small dark spots (
<figureCitation id="137D7FDAFF9EFFBBFCF0FBCB35B4FB2F" box="[842,929,1149,1173]" captionStart="Fig" captionStartId="8.[113,148,1079,1101]" captionTargetBox="[113,1451,188,1063]" captionTargetId="figure-58@8.[113,1451,188,1063]" captionTargetPageId="8" captionText="Fig. 4. Living habitus specimens of Caridina species from the Mae Klong Basin. AD, Caridina maeklongensis, new species: A, paratype male, MUMNH-CAR064-4; B, paratype female, MUMNH-CAR064-1; C, paratype female, MUMNH-CAR064-15; and D, paratype female, MUMNH-CAR064-13. E, F, C. sumatrensis: E, male, MUMNH-CAR706-P2; and F, female, MUMNH-CAR706-P1." pageId="13" pageNumber="463">Fig. 4E</figureCitation>
). Colour of female shrimps is frequently dark to light blue, decorated with scattered small dark spots (
<figureCitation id="137D7FDAFF9EFFBBFA2DFB2B3576FB6F" captionStart="Fig" captionStartId="8.[113,148,1079,1101]" captionTargetBox="[113,1451,188,1063]" captionTargetId="figure-58@8.[113,1451,188,1063]" captionTargetPageId="8" captionText="Fig. 4. Living habitus specimens of Caridina species from the Mae Klong Basin. AD, Caridina maeklongensis, new species: A, paratype male, MUMNH-CAR064-4; B, paratype female, MUMNH-CAR064-1; C, paratype female, MUMNH-CAR064-15; and D, paratype female, MUMNH-CAR064-13. E, F, C. sumatrensis: E, male, MUMNH-CAR706-P2; and F, female, MUMNH-CAR706-P1." pageId="13" pageNumber="463">Fig. 4F</figureCitation>
). Rostrum is yellowish-orange. Each abdominal somite furnished dorsally with conspicuous yellowish-orange bands. Eggs are brown to orange.
</paragraph>
</subSubSection>
<subSubSection id="C35C30D4FF9EFFBBFCFFFA8B32CCF98F" pageId="13" pageNumber="463" type="distribution">
<paragraph id="8BF9635FFF9EFFBBFCFFFA8B32CCF98F" blockId="13.[834,1475,1341,1589]" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCFFFA8B324BFAEF" bold="true" box="[837,1118,1341,1365]" pageId="13" pageNumber="463">Distribution and habitat.</emphasis>
<collectingCountry id="F35123CFFF9EFFBBFBE5FA8B32D3FAEF" box="[1119,1222,1341,1365]" name="Indonesia" pageId="13" pageNumber="463">Indonesia</collectingCountry>
(
<collectingRegion id="4982ADBDFF9EFFBBFB6EFA8B3327FAEF" box="[1236,1330,1341,1365]" country="Indonesia" name="Sumatera Utara" pageId="13" pageNumber="463">Sumatra</collectingRegion>
),
<collectingCountry id="F35123CFFF9EFFBBFAF9FA8B33ABFAEF" box="[1347,1470,1341,1365]" name="Philippines" pageId="13" pageNumber="463">Philippines</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFCF8FAEB35A7FACF" box="[834,946,1373,1397]" name="Singapore" pageId="13" pageNumber="463">Singapore</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFC7AFAEB3230FACF" box="[960,1061,1373,1397]" name="Malaysia" pageId="13" pageNumber="463">Malaysia</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFB89FAEB3280FACF" box="[1075,1173,1373,1397]" name="Thailand" pageId="13" pageNumber="463">Thailand</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFB1EFAE8331AFACC" box="[1188,1295,1374,1398]" name="Myanmar" pageId="13" pageNumber="463">Myanmar</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFAA7FAEB3342FACF" box="[1309,1367,1373,1397]" name="India" pageId="13" pageNumber="463">India</collectingCountry>
,
<collectingCountry id="F35123CFFF9EFFBBFADFFAEB33D5FACF" box="[1381,1472,1373,1397]" name="Pakistan" pageId="13" pageNumber="463">Pakistan</collectingCountry>
(
<bibRefCitation id="EFD71EAEFF9EFFBBFCF1FACB35CBFA2F" author="Kemp S" box="[843,990,1405,1429]" pageId="13" pageNumber="463" pagination="81 - 102" refId="ref14666" refString="Kemp S (1918 a) Crustacea Decapoda of the Inle Lake Basin. Records of the Indian Museum, 14: 81 - 102." type="journal article" year="1918">Kemp, 1918a</bibRefCitation>
, b;
<bibRefCitation id="EFD71EAEFF9EFFBBFBB9FACB32DFFA2F" author="Kazmi Q &amp; Siddiqui FA &amp; Kazmi MA" box="[1027,1226,1405,1429]" pageId="13" pageNumber="463" pagination="43 - 46" refId="ref14618" refString="Kazmi Q, Siddiqui FA &amp; Kazmi MA (2002) Range extension of Caridina weberi sumatrensis De Man, 1892 (Decapoda, Caridea, Atyidae) to Sindh waters (Karachi, Pakistan). Turkish Journal of Zoology, 26: 43 - 46." type="journal article" year="2002">Kazmi et al., 2002</bibRefCitation>
; Cai et al., 2007;
<bibRefCitation id="EFD71EAEFF9EFFBBFA38FACB35E4FA0F" author="Cai Y &amp; Naiyanetr P" pageId="13" pageNumber="463" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
; this study). Based on the field survey in the Mae Klong Basin in this study,
<taxonomicName id="4C4618DCFF9EFFBBFB25FA08332BFA6F" authorityName="De Man" authorityYear="1892" box="[1183,1342,1470,1493]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9EFFBBFB25FA08332BFA6F" box="[1183,1342,1470,1493]" italics="true" pageId="13" pageNumber="463">C. sumatrensis</emphasis>
</taxonomicName>
is restricted to the lower part of the basin (
<figureCitation id="137D7FDAFF9EFFBBFB32FA6B32DFFA4F" box="[1160,1226,1501,1525]" captionStart="Fig" captionStartId="11.[835,870,1243,1265]" captionTargetBox="[838,1475,189,1226]" captionTargetId="figure-26@11.[1028,1215,703,769]" captionTargetPageId="11" captionText="Fig. 5. Map of the Mae Klong Basin showing sampling localities of Caridina maeklongensis, new species (black circles) and C. sumatrensis (grey circle), and the localities where both species were absent (white circles)." pageId="13" pageNumber="463">Fig. 5</figureCitation>
), and commonly found in lowland rivers and their tributaries. They are found living on aquatic vegetation near the banks.
</paragraph>
</subSubSection>
<subSubSection id="C35C30D4FF9EFFB8FCFFF9EB34E4FACF" lastPageId="14" lastPageNumber="464" pageId="13" pageNumber="463" type="discussion">
<paragraph id="8BF9635FFF9EFFB8FCFFF9EB378AFE0F" blockId="13.[834,1476,1629,2102]" lastBlockId="14.[112,753,189,437]" lastPageId="14" lastPageNumber="464" pageId="13" pageNumber="463">
<emphasis id="B932BF4DFF9EFFBBFCFFF9EB35ADF9CF" bold="true" box="[837,952,1629,1653]" pageId="13" pageNumber="463">Remarks.</emphasis>
The morphology of
<taxonomicName id="4C4618DCFF9EFFBBFB11F9E83341F9CF" authorityName="De Man" authorityYear="1892" box="[1195,1364,1630,1653]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="463" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9EFFBBFB11F9E83341F9CF" box="[1195,1364,1630,1653]" italics="true" pageId="13" pageNumber="463">C. sumatrensis</emphasis>
</taxonomicName>
from the Mae Klong Basin in our collection closely matches with the original description (
<bibRefCitation id="EFD71EAEFF9EFFBBFBEDF92B32E8F90F" author="De Man JG" box="[1111,1277,1693,1717]" pageId="13" pageNumber="463" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
), redescription of
<typeStatus id="54FDDDFDFF9EFFBBFCF9F9083564F96C" box="[835,881,1726,1750]" pageId="13" pageNumber="463">type</typeStatus>
materials (
<bibRefCitation id="EFD71EAEFF9EFFBBFC4BF90B32EDF96F" author="Cai Y &amp; Naiyanetr P" box="[1009,1272,1725,1749]" pageId="13" pageNumber="463" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
), and descriptions in previous taxonomic literature (e.g.,
<bibRefCitation id="EFD71EAEFF9EFFBBFB58F9683362F94F" author="Kemp S" box="[1250,1399,1757,1781]" pageId="13" pageNumber="463" refId="ref14692" refString="Kemp S (1918 b) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. In: Annandale N (ed.) Zoological results of a tour in the Far East. Memoirs of the Asiatic Society of Bengal. Volume 6. The Asiatic Society, Calcutta, pp. 217 - 297." type="book" year="1918">Kemp, 1918b</bibRefCitation>
;
<bibRefCitation id="EFD71EAEFF9EFFBBFA39F96B3568F8AF" author="Pillai RS" pageId="13" pageNumber="463" pagination="42 - 47" refId="ref15689" refString="Pillai RS (1964) Four species of Caridina from Travacore, including of a new variety. Journal of the Marine Biological Association of India, 6 (1): 42 - 47." type="journal article" year="1964">Pillai, 1964</bibRefCitation>
; Cai et al., 2007). Particularly, the characteristic of the slightly bent downward rostrum with a ridge at the base in our samples (
<figureCitation id="137D7FDAFF9EFFBBFC1CF88B3202F8EF" box="[934,1047,1853,1877]" captionStart="Fig" captionStartId="12.[113,148,1910,1932]" captionTargetBox="[116,1454,229,1901]" captionTargetId="figure-136@12.[99,1467,216,1912]" captionTargetPageId="12" captionText="Fig. 6. Caridina sumatrensis De Man, 1892. A, ovigerous female; B, cephalothorax and cephalic appendage; C, scaphocerite; D, antennular peduncle; E, first pereopod; F, second pereopod; G, third pereopod; H, dactylus of third pereopod; I, fifth pereopod; J, dactylus of fifth pereopod; K, telson; L, distal end of telson; M, uropodal dieresis; N, preanal carina; O, male first pleopod; P, endopod of male first pleopod; Q, male second pleopod; R, appendix masculina of male second pleopod. Drawings were made from ovigerous female, MUMNH- CAR706-P1 (AM) and male, MUMNH-CAR706-P2 (OR) collected from the Mae Klong Basin, Thailand. Scale bars: A, B, E, F, G, I, K = 1 mm; C, D, M, O, Q = 0.5 mm; H, J, L, N, P, R = 0.25 mm." pageId="13" pageNumber="463">Fig. 6A, B</figureCitation>
) is in close agreement with illustrations by previous studies (i.e.,
<bibRefCitation id="EFD71EAEFF9EFFBBFBE6F8E83315F8CF" author="De Man JG" box="[1116,1280,1885,1909]" pageId="13" pageNumber="463" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
; Cai et al., 2007;
<bibRefCitation id="EFD71EAEFF9EFFBBFCF9F8CB322CF82F" author="Cai Y &amp; Naiyanetr P" box="[835,1081,1917,1941]" pageId="13" pageNumber="463" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
). Populations from Mae Klong Basin share a similar number of rostral teeth with those from the original description by
<bibRefCitation id="EFD71EAEFF9EFFBBFB8DF80832CEF86F" author="De Man JG" box="[1079,1243,1981,2005]" pageId="13" pageNumber="463" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man (1892)</bibRefCitation>
, including the number of dorsal teeth (1721 vs 1620), ventral teeth (36 both in the Mae Klong population and the original description), and postorbital teeth (46 vs 56). The egg size of the Mae Klong population is also congruent with previous reports (0.410.46 ×
<quantity id="4CBECEBAFF9DFFB8FF32FF6B3730FF4F" box="[136,293,221,245]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="14" pageNumber="464" unit="mm" value="0.265" valueMax="0.29" valueMin="0.24">0.240.29 mm</quantity>
vs 0.460.47 ×
<quantity id="4CBECEBAFF9DFFB8FE66FF6B346CFF4F" box="[476,633,221,245]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="2.9" metricValueMin="2.8" pageId="14" pageNumber="464" unit="mm" value="0.285" valueMax="0.29" valueMin="0.28">0.280.29 mm</quantity>
by
<bibRefCitation id="EFD71EAEFF9DFFB8FD1DFF6836A2FEAF" author="Kemp S" pageId="14" pageNumber="464" refId="ref14692" refString="Kemp S (1918 b) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. In: Annandale N (ed.) Zoological results of a tour in the Far East. Memoirs of the Asiatic Society of Bengal. Volume 6. The Asiatic Society, Calcutta, pp. 217 - 297." type="book" year="1918">Kemp, 1918b</bibRefCitation>
; 0.46 ×
<quantity id="4CBECEBAFF9DFFB8FEA2FF4B376EFEAF" box="[280,379,253,277]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="14" pageNumber="464" unit="mm" value="0.28">0.28 mm</quantity>
by
<bibRefCitation id="EFD71EAEFF9DFFB8FE10FF4B3438FEAF" author="Pillai RS" box="[426,557,253,277]" pageId="14" pageNumber="464" pagination="42 - 47" refId="ref15689" refString="Pillai RS (1964) Four species of Caridina from Travacore, including of a new variety. Journal of the Marine Biological Association of India, 6 (1): 42 - 47." type="journal article" year="1964">Pillai, 1964</bibRefCitation>
; and 0.400.42 ×
<quantity id="4CBECEBAFF9DFFB8FFCBFEAB3707FE8F" box="[113,274,285,309]" metricMagnitude="-4" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.5" metricValueMin="2.2" pageId="14" pageNumber="464" unit="mm" value="0.235" valueMax="0.25" valueMin="0.22">0.220.25 mm</quantity>
by
<bibRefCitation id="EFD71EAEFF9DFFB8FEFBFEAB3453FE8F" author="Cai Y &amp; Naiyanetr P" box="[321,582,285,309]" pageId="14" pageNumber="464" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
). However, the Mae Klong population possesses a slightly lower number of spiniform setae on the P5 dactylus (3243 vs 3657 by
<bibRefCitation id="EFD71EAEFF9DFFB8FFCBFEC83712FE2F" author="Kemp S" box="[113,263,381,405]" pageId="14" pageNumber="464" refId="ref14692" refString="Kemp S (1918 b) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. In: Annandale N (ed.) Zoological results of a tour in the Far East. Memoirs of the Asiatic Society of Bengal. Volume 6. The Asiatic Society, Calcutta, pp. 217 - 297." type="book" year="1918">Kemp, 1918b</bibRefCitation>
; more than 40 by Cai et al., 2007; and 4249 by
<bibRefCitation id="EFD71EAEFF9DFFB8FF2CFE2B3786FE0F" author="Cai Y &amp; Naiyanetr P" box="[150,403,413,437]" pageId="14" pageNumber="464" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
).
</paragraph>
<paragraph id="8BF9635FFF9DFFB8FFCBFE6B346BFC4F" blockId="14.[112,754,477,1013]" pageId="14" pageNumber="464">
Originally,
<taxonomicName id="4C4618DCFF9DFFB8FF55FE68379AFE4F" authorityName="De Man" authorityYear="1892" box="[239,399,478,501]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FF55FE68379AFE4F" box="[239,399,478,501]" italics="true" pageId="14" pageNumber="464">C. sumatrensis</emphasis>
</taxonomicName>
was proposed as a variant form of
<taxonomicName id="4C4618DCFF9DFFB8FF34FE4836E0FDAF" authorityName="De Man" authorityYear="1892" box="[142,245,509,533]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="B932BF4DFF9DFFB8FF34FE4836E0FDAF" box="[142,245,509,533]" italics="true" pageId="14" pageNumber="464">C. weberi</emphasis>
</taxonomicName>
, as “
<taxonomicName id="4C4618DCFF9DFFB8FE90FE4B36BFFD8F" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="variety" species="weberi" variety="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FE90FE4B37CCFDAF" box="[298,473,509,533]" italics="true" pageId="14" pageNumber="464">Caridina weberi</emphasis>
var.
<emphasis id="B932BF4DFF9DFFB8FDB7FE483499FDAF" box="[525,652,510,533]" italics="true" pageId="14" pageNumber="464">sumatrensis</emphasis>
<bibRefCitation id="EFD71EAEFF9DFFB8FD29FE4836BFFD8F" author="De Man JG" pageId="14" pageNumber="464" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
”.
<bibRefCitation id="EFD71EAEFF9DFFB8FF07FDA8379BFD8F" author="Wowor D &amp; Cai Y &amp; Ng PKL" box="[189,398,541,565]" pageId="14" pageNumber="464" pagination="337 - 357" refId="ref16444" refString="Wowor D, Cai Y &amp; Ng PKL (2004) Crustacea: Decapoda, Caridea. In: Yule CM &amp; Sen YH (eds.) The Freshwater Invertebrates of the Malaysian Region. Academy of Sciences Malaysia, Kuala Lumpur, pp. 337 - 357." type="book chapter" year="2004">Wowor et al. (2004)</bibRefCitation>
later elevated it as a valid species. This classification has been widely accepted in subsequent works (
<bibRefCitation id="EFD71EAEFF9DFFB8FF07FDEB3786FDCF" author="Cai Y &amp; Shokita S" box="[189,403,605,629]" pageId="14" pageNumber="464" pagination="245 - 270" refId="ref13226" refString="Cai Y &amp; Shokita S (2006) Report on a collection of freshwater shrimps (Crustacea: Decapoda: Caridea) from the Philippines, with descriptions of four new species. Raffles Bulletin of Zoology, 54 (2): 245 - 270." type="journal article" year="2006">Cai &amp; Shokita, 2006</bibRefCitation>
; Cai et al., 2007;
<bibRefCitation id="EFD71EAEFF9DFFB8FDFBFDEB36BCFD2F" author="Cai Y &amp; Naiyanetr P" pageId="14" pageNumber="464" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
).
<taxonomicName id="4C4618DCFF9DFFB8FF00FDCB3789FD2F" authorityName="De Man" authorityYear="1892" box="[186,412,637,661]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FF00FDCB3789FD2F" box="[186,412,637,661]" italics="true" pageId="14" pageNumber="464">Caridina sumatrensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C4618DCFF9DFFB8FD9BFDC83493FD2F" authorityName="De Man" authorityYear="1892" box="[545,646,637,661]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="B932BF4DFF9DFFB8FD9BFDC83493FD2F" box="[545,646,637,661]" italics="true" pageId="14" pageNumber="464">C. weberi</emphasis>
</taxonomicName>
in several points (see summary in
<tableCitation id="C6C456E4FF9DFFB8FEC0FD2B37D8FD0F" box="[378,461,669,693]" captionStart="Table 3" captionStartId="9.[111,166,525,548]" captionText="Table 3. Comparison of morphological characters between C. weberi, C. sumatrensis, C. babaulti, and C. maeklongensis, new species" pageId="14" pageNumber="464">Table 3</tableCitation>
), including having a ridge at the base of rostrum (vs absent), longer rostrum (reaching near the end of third segment of antennular peduncle vs reaching to the base or near the middle of second segment of antennular peduncle), more dorsal teeth (1620 vs 1019, usually 1314), more postorbital teeth (46 vs 02), shorter antennular peduncle (0.460.55 as long as CL vs 0.64 as long as CL), stouter P5 dactylus (2.763.96 vs 3.95.2 as long as wide), and the distal end of telson subtriangular (vs rounded) (
<bibRefCitation id="EFD71EAEFF9DFFB8FF59FC0B3792FC6F" author="De Man JG" box="[227,391,957,981]" pageId="14" pageNumber="464" pagination="15 - 29" refId="ref13508" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse eniner Reise in Niederlandisch Ost-Indien. Volume 2. E. J. Brill, Leiden, pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
; Cai et al., 2007,
<bibRefCitation id="EFD71EAEFF9DFFB8FDF5FC0B36E5FC4F" author="de Mazancourt V &amp; Boseto D &amp; Marquet G &amp; Keith P" pageId="14" pageNumber="464" pagination="1 - 86" refId="ref13715" refString="de Mazancourt V, Boseto D, Marquet G &amp; Keith P (2020) Solomon's Gold Mine: Description or redescription of 24 species of Caridina (Crustacea: Decapoda: Atyidae) freshwater shrimps from the Solomon Islands, including 11 new species. European Journal of Taxonomy, 696: 1 - 86." type="journal article" year="2020">de Mazancourt et al., 2020</bibRefCitation>
;
<bibRefCitation id="EFD71EAEFF9DFFB8FF46FC6B37ECFC4F" author="Cai Y &amp; Naiyanetr P" box="[252,505,989,1013]" pageId="14" pageNumber="464" pagination="767 - 804" refId="ref13052" refString="Cai Y &amp; Naiyanetr P (2024) The freshwater shrimps of the family Atyidae (Decapoda, Caridea) of Thailand. Crustaceana, 97 (5 - 9): 767 - 804." type="journal article" year="2024">Cai &amp; Naiyanetr, 2024</bibRefCitation>
; this study).
</paragraph>
<paragraph id="8BF9635FFF9DFFB8FFC9FBAB34E4FACF" blockId="14.[112,753,1053,1397]" pageId="14" pageNumber="464">
<taxonomicName id="4C4618DCFF9DFFB8FFC9FBAB374BFB8F" authorityName="De Man" authorityYear="1892" box="[115,350,1053,1077]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FFC9FBAB374BFB8F" box="[115,350,1053,1077]" italics="true" pageId="14" pageNumber="464">Caridina sumatrensis</emphasis>
</taxonomicName>
shares similarities with
<emphasis id="B932BF4DFF9DFFB8FDD4FBA834E4FB8F" box="[622,753,1053,1077]" italics="true" pageId="14" pageNumber="464">
<taxonomicName id="4C4618DCFF9DFFB8FDD4FBA834F8FB8F" authorityName="Bouvier" authorityYear="1918" box="[622,749,1053,1077]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="babaulti">C. babaulti</taxonomicName>
.
</emphasis>
However, several key morphological characteristics can be used to separate them. The P2 chela of
<taxonomicName id="4C4618DCFF9DFFB8FDBBFBE83488FBCF" authorityName="De Man" authorityYear="1892" box="[513,669,1118,1141]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FDBBFBE83488FBCF" box="[513,669,1118,1141]" italics="true" pageId="14" pageNumber="464">C. sumatrensis</emphasis>
</taxonomicName>
is more slender than those of
<taxonomicName id="4C4618DCFF9DFFB8FEF2FBC837A9FB2F" authorityName="Bouvier" authorityYear="1918" box="[328,444,1149,1173]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="babaulti">
<emphasis id="B932BF4DFF9DFFB8FEF2FBC837A9FB2F" box="[328,444,1149,1173]" italics="true" pageId="14" pageNumber="464">C. babaulti</emphasis>
</taxonomicName>
(2.713.40 vs 2.52.9), while the P3 dactylus of
<taxonomicName id="4C4618DCFF9DFFB8FE83FB2837C2FB0F" authorityName="De Man" authorityYear="1892" box="[313,471,1182,1205]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FE83FB2837C2FB0F" box="[313,471,1182,1205]" italics="true" pageId="14" pageNumber="464">C. sumatrensis</emphasis>
</taxonomicName>
has fewer spiniform setae (46 vs 710).
<taxonomicName id="4C4618DCFF9DFFB8FEB1FB0B37F8FB6F" authorityName="De Man" authorityYear="1892" box="[267,493,1213,1237]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FEB1FB0B37F8FB6F" box="[267,493,1213,1237]" italics="true" pageId="14" pageNumber="464">Caridina sumatrensis</emphasis>
</taxonomicName>
possesses a conspicuous posteromedian projection on the distal end of the telson. This feature is absent in
<taxonomicName id="4C4618DCFF9DFFB8FEFCFB4837ABFAAF" authorityName="Bouvier" authorityYear="1918" box="[326,446,1277,1301]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="babaulti">
<emphasis id="B932BF4DFF9DFFB8FEFCFB4837ABFAAF" box="[326,446,1277,1301]" italics="true" pageId="14" pageNumber="464">C. babaulti</emphasis>
</taxonomicName>
. In addition,
<taxonomicName id="4C4618DCFF9DFFB8FDE8FB4834E4FAAF" authorityName="De Man" authorityYear="1892" box="[594,753,1278,1301]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B932BF4DFF9DFFB8FDE8FB4834E4FAAF" box="[594,753,1278,1301]" italics="true" pageId="14" pageNumber="464">C. sumatrensis</emphasis>
</taxonomicName>
produces small-sized eggs, whereas the eggs of
<taxonomicName id="4C4618DCFF9DFFB8FDC2FAA834E4FA8F" authorityName="Bouvier" authorityYear="1918" box="[632,753,1309,1333]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="464" phylum="Arthropoda" rank="species" species="babaulti">
<emphasis id="B932BF4DFF9DFFB8FDC2FAA834E4FA8F" box="[632,753,1309,1333]" italics="true" pageId="14" pageNumber="464">C. babaulti</emphasis>
</taxonomicName>
are much larger (0.410.46 ×
<quantity id="4CBECEBAFF9DFFB8FE05FA8B3448FAEF" box="[447,605,1341,1365]" metricMagnitude="-4" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="14" pageNumber="464" unit="mm" value="0.265" valueMax="0.29" valueMin="0.24">0.240.29 mm</quantity>
vs 0.91.0 ×
<quantity id="4CBECEBAFF9DFFB8FFCBFAEB3705FACF" box="[113,272,1373,1397]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" metricValueMax="6.2" metricValueMin="5.0" pageId="14" pageNumber="464" unit="mm" value="0.56" valueMax="0.62" valueMin="0.5">0.500.62 mm</quantity>
) (
<bibRefCitation id="EFD71EAEFF9DFFB8FE9DFAEB37D3FACF" author="Bouvier EL" box="[295,454,1373,1397]" pageId="14" pageNumber="464" pagination="386 - 393" refId="ref12864" refString="Bouvier EL (1918) Sur quelques Crustaces decapodes recueillis par M. Guy Babault dans les eaux douce de Inde anglaise. Bulletin du Museum National d'Histoire Naturelle, Paris, 24: 386 - 393." type="journal article" year="1918">Bouvier, 1918</bibRefCitation>
;
<bibRefCitation id="EFD71EAEFF9DFFB8FE69FAEB34F0FACF" author="Pandya P &amp; Richard J" box="[467,741,1373,1397]" pageId="14" pageNumber="464" pagination="470 - 482" refId="ref15640" refString="Pandya P &amp; Richard J (2019) Report of Caridina babaulti Bouvier, 1918 (Crustacea: Decapoda: Caridea: Atyidae) and description of a new species Caridina kutchi sp. nov. from Gujarat, India. Zootaxa, 4568: 470 - 482." type="journal article" year="2019">Pandya &amp; Richard, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,167 @@
<document id="FD214A87D36B408AFA12D1AF8DFB8139" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB1FF98440D546D6C548BFA" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Arctosa C. L. Koch 1847" docType="treatment" docVersion="2" lastPageNumber="236" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="236" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BACC42B238B1068725C5FF7FE4E4C067" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E1193263DB7D445BC48856AF49586D87">
<mods:title id="2D932620D8CC33CB0EA6B48B811CF9CC">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="D2B6EF8217C0AF9724DF60713D96F0B7" type="personal">
<mods:role id="BCD13C386DA51FA3C995A0E27B842314">
<mods:roleTerm id="33344EFDA5170CC12488C2EF43F40FE2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8840C8E0FF62C993FA0F16D04DF2B95E">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2E4A2128DE34ADABC265E214B67BE260">text</mods:typeOfResource>
<mods:relatedItem id="7EB330522D053023390B31FD41D64483" type="host">
<mods:titleInfo id="1FB680EBF2BFC75A4F4C6AF93E6E2BBB">
<mods:title id="3CA7EF4B2D91D38FE8696C4B2EBD0863">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="B3CF7F69BBFD027C69062155D4F3F4D9">
<mods:date id="83C333395658521EF84426D081D00BB1">2024</mods:date>
<mods:detail id="122D395C6383A00060CFF4DCB954CCD6" type="pubDate">
<mods:number id="828B5C545CAC29AAD68DFD37853E127B">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="99088402FFBC79085D04033F473AFF28" type="volume">
<mods:number id="6085E49F481231AF9EF09D7AFD9A6767">72</mods:number>
</mods:detail>
<mods:extent id="C41B0D0289CFA6C57D9276553B5B6E0C" unit="page">
<mods:start id="B520BE00490AF3FE8ED1512E3C41128E">235</mods:start>
<mods:end id="503CDB6B523A56021DD6203082B9B1DD">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FCBA9514FD4D71A050605085510B222F">journal article</mods:classification>
<mods:identifier id="AA76D1E43AE8839DA6317928EE574F79" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="26741BB22626E93E4758655C0D674D30" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="BB255140C56B0FD77D16C984BE5A8B72" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="48913B3468BCF39F48DF2D7CE8C071D8" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB1FF98440D546D6C548BFA" ID-DOI="http://doi.org/10.5281/zenodo.14683148" ID-Zenodo-Dep="14683148" LSID="urn:lsid:plazi:treatment:03F187BDFFB1FF98440D546D6C548BFA" httpUri="http://treatment.plazi.org/id/03F187BDFFB1FF98440D546D6C548BFA" lastPageNumber="236" pageId="1" pageNumber="236">
<subSubSection id="C3426520FFB1FF98440D546D6F2E8E3A" box="[287,577,829,853]" pageId="1" pageNumber="236" type="nomenclature">
<paragraph id="8BE736ABFFB1FF98440D546D6F2E8E3A" blockId="1.[287,577,829,853]" box="[287,577,829,853]" pageId="1" pageNumber="236">
<heading id="D0AF81C7FFB1FF98440D546D6F2E8E3A" bold="true" box="[287,577,829,853]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="236" reason="2">
<taxonomicName id="4C584D28FFB1FF98440D546D6F2E8E3A" ID-CoL="32DS" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[287,577,829,853]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB1FF98440D546D6F2E8E3A" bold="true" box="[287,577,829,853]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98440D546D6C1C8E3A" bold="true" box="[287,371,829,853]" italics="true" pageId="1" pageNumber="236">Arctosa</emphasis>
C. L.
<bibRefCitation id="EFC94B5AFFB1FF9844D2546D6F2E8E3A" author="Koch CL" box="[448,577,829,853]" pageId="1" pageNumber="236" refId="ref9676" refString="Koch CL (1847) Die Arachniden. J. L. Lotzbeck, Nurnberg, Vierzehnter Band, pp. 89 - 210, pl. 481 - 504 (f. 1343 - 1412); Funfzehnter Band, pp. 1 - 136, pl. 505 - 540 (f. 1413 - 1504); Sechszehnter und letzter Band, pp. 1 - 80, pl. 541 - 563 (f. 1505 - 1550), Index 64 pp." type="book" year="1847">Koch, 1847</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF984563542D6F9F8EFA" box="[113,752,893,917]" pageId="1" pageNumber="236" type="type_taxon">
<paragraph id="8BE736ABFFB1FF984563542D6F9F8EFA" blockId="1.[113,752,893,917]" box="[113,752,893,917]" pageId="1" pageNumber="236">
<materialsCitation id="3B303CF6FFB1FF984563542D6F838EFA" box="[113,748,893,917]" country="France" location="France" pageId="1" pageNumber="236" specimenCount="1" typeStatus="holotype">
<emphasis id="B92CEAB9FFB1FF984563542D6C698EFA" bold="true" box="[113,262,893,917]" pageId="1" pageNumber="236">
<typeStatus id="54E38809FFB1FF984563542D6DC48EFA" box="[113,171,893,917]" pageId="1" pageNumber="236">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C584D28FFB1FF98441E542E6F328EFA" authority="Fabricius, 1777" authorityName="Fabricius" authorityYear="1777" box="[268,605,893,917]" class="Arachnida" family="Agelenidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="cinerea">
<emphasis id="B92CEAB9FFB1FF98441E542E6CDF8EFA" box="[268,432,894,917]" italics="true" pageId="1" pageNumber="236">Aranea cinerea</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF9844A7542D6F328EFA" author="Fabricius JC" box="[437,605,893,917]" pageId="1" pageNumber="236" refId="ref9577" refString="Fabricius JC (1777) Genera insectorum eorumque characteres naturales, secundum numerum, figuram, situm et proportionem omnium partium oris; adjecta mantissa specierum nuper detectarum. Michael Friedrich Bartsch, Chilonii [= Kiel], 310 pp." type="book" year="1777">Fabricius, 1777</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F34F763BFFB1FF9847B2542E6F838EFA" box="[672,748,894,917]" name="France" pageId="1" pageNumber="236">France</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF98456354ED6C548BFA" pageId="1" pageNumber="236" type="discussion">
<paragraph id="8BE736ABFFB1FF98456354ED6F3089BA" blockId="1.[113,754,957,1237]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98456354ED6D998EBA" bold="true" box="[113,246,957,981]" pageId="1" pageNumber="236">Comments.</emphasis>
<taxonomicName id="4C584D28FFB1FF98441354EE6F4D8EBA" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[257,546,957,981]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB1FF98441354EE6C398EBA" box="[257,342,958,981]" italics="true" pageId="1" pageNumber="236">Arctosa</emphasis>
C.L.
<bibRefCitation id="EFC94B5AFFB1FF98448E54ED6F4D8EBA" author="Koch CL" box="[412,546,957,981]" pageId="1" pageNumber="236" refId="ref9676" refString="Koch CL (1847) Die Arachniden. J. L. Lotzbeck, Nurnberg, Vierzehnter Band, pp. 89 - 210, pl. 481 - 504 (f. 1343 - 1412); Funfzehnter Band, pp. 1 - 136, pl. 505 - 540 (f. 1413 - 1504); Sechszehnter und letzter Band, pp. 1 - 80, pl. 541 - 563 (f. 1505 - 1550), Index 64 pp." type="book" year="1847">Koch, 1847</bibRefCitation>
</taxonomicName>
, with 168 species and two subspecies (
<bibRefCitation id="EFC94B5AFFB1FF984471548D6C818E9A" author="WSC" box="[355,494,989,1013]" pageId="1" pageNumber="236" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC, 2024</bibRefCitation>
), is one of the largest genera of
<taxonomicName id="4C584D28FFB1FF9845CF54AD6C21897A" authorityName="Sundevall" authorityYear="1833" box="[221,334,1021,1045]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
. It is a poorly defined and undoubtedly polyphyletic genus (
<bibRefCitation id="EFC94B5AFFB1FF98444B534D6F40895A" author="Wang LY &amp; Marusik YM &amp; Zhang ZS" box="[345,559,1053,1077]" pageId="1" pageNumber="236" pagination="53 - 68" refId="ref11188" refString="Wang LY, Marusik YM &amp; Zhang ZS (2012) Notes on three poorly known Arctosa species from China (Araneae: Lycosidae). Zootaxa, 3404: 53 - 68." type="journal article" year="2012">Wang et al., 2012</bibRefCitation>
).
<taxonomicName id="4C584D28FFB1FF984754534E6FF3895A" authorityName="C. L. Koch" authorityYear="1847" box="[582,668,1054,1077]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB1FF984754534E6FF3895A" box="[582,668,1054,1077]" italics="true" pageId="1" pageNumber="236">Arctosa</emphasis>
</taxonomicName>
has an almost worldwide distribution, although it does not occur in
<collectingCountry id="F34F763BFFB1FF98459C530D6D9C891A" box="[142,243,1117,1141]" name="Australia" pageId="1" pageNumber="236">Australia</collectingCountry>
. The genus is relatively well studied in Europe (
<bibRefCitation id="EFC94B5AFFB1FF98456B532D6CF089FA" author="Lugetti G &amp; Tongiorgi P" box="[121,415,1149,1173]" pageId="1" pageNumber="236" pagination="165 - 229" refId="ref10062" refString="Lugetti G &amp; Tongiorgi P (1965) Revisione delle specie italiane dei generi Arctosa C. L. Koch e Tricca Simon con note su una Acantholycosa delle Alpi Giulie (Araneae-Lycosidae). Redia, 49: 165 - 229." type="journal article" year="1965">Lugetti &amp; Tongiorgi, 1965</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB1FF9844BE532D6F3789FA" author="Almquist S" box="[428,600,1149,1173]" pageId="1" pageNumber="236" pagination="1 - 284" refId="ref9149" refString="Almquist S (2005) Swedish Araneae, part 1 - families Atypidae to Hahniidae (Linyphiidae excluded). Insect Systematics &amp; Evolution, Supplement 62: 1 - 284." type="journal article" year="2005">Almquist, 2005</bibRefCitation>
), the Nearctic (
<bibRefCitation id="EFC94B5AFFB1FF98456853CD6CE389DA" author="Dondale CD &amp; Redner JH" box="[122,396,1181,1205]" pageId="1" pageNumber="236" pagination="1 - 30" refId="ref9491" refString="Dondale CD &amp; Redner JH (1983) Revision of the wolf spiders of the genus Arctosa C. L. Koch in North and Central America (Araneae: Lycosidae). Journal of Arachnology, 11: 1 - 30." type="journal article" year="1983">Dondale &amp; Redner, 1983</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB1FF98448553CD6FCD89DA" author="Paquin P &amp; Duperre N" box="[407,674,1181,1205]" pageId="1" pageNumber="236" pagination="1 - 251" refId="ref10328" refString="Paquin P &amp; Duperre N (2003) Guide d'identification des araignees de Quebec. Fabreries, Supplement 11: 1 - 251." type="journal article" year="2003">Paquin &amp; Dupérré, 2003</bibRefCitation>
),
<collectingCountry id="F34F763BFFB1FF9847A753CE6F9D89DA" box="[693,754,1182,1205]" name="Japan" pageId="1" pageNumber="236">Japan</collectingCountry>
(
<bibRefCitation id="EFC94B5AFFB1FF98456B53ED6C6089BA" author="Tanaka H" box="[121,271,1213,1237]" pageId="1" pageNumber="236" pagination="222 - 248" refId="ref10868" refString="Tanaka H (2009) Lycosidae. In: Ono H (ed.) The spiders of Japan with keys to the families and genera and illustrations of the species. Tokai University Press, Kanagawa, pp. 222 - 248." type="book chapter" year="2009">Tanaka, 2009</bibRefCitation>
) and
<collectingCountry id="F34F763BFFB1FF98445C53EE6CE089BA" box="[334,399,1214,1237]" name="South Korea" pageId="1" pageNumber="236">Korea</collectingCountry>
(
<bibRefCitation id="EFC94B5AFFB1FF98448D53ED6F3B89BA" author="Namkung J" box="[415,596,1213,1237]" pageId="1" pageNumber="236" refId="ref10190" refString="Namkung J (2003) The Spiders of Korea, 2 nd. ed. Kyo-Hak Publishing Co., Seoul, 648 pp." type="book" year="2003">Namkung, 2003</bibRefCitation>
).
</paragraph>
<paragraph id="8BE736ABFFB1FF98456353AD6C5D88DA" blockId="1.[113,753,1277,1461]" pageId="1" pageNumber="236">
Almost one-fifth (32) of all named
<taxonomicName id="4C584D28FFB1FF9844ED53AE6FC6887A" box="[511,681,1277,1301]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF9844ED53AE6F3D887A" box="[511,594,1278,1301]" italics="true" pageId="1" pageNumber="236">Arctosa</emphasis>
species
</taxonomicName>
occur in
<collectingCountry id="F34F763BFFB1FF984580524D6DBB885A" box="[146,212,1309,1333]" name="China" pageId="1" pageNumber="236">China</collectingCountry>
(
<bibRefCitation id="EFC94B5AFFB1FF9845FA524D6C1B885A" author="WSC" box="[232,372,1309,1333]" pageId="1" pageNumber="236" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC, 2024</bibRefCitation>
). Chinese species have not been revised, and data regarding their taxonomy and distribution is scattered throughout numerous different papers (
<bibRefCitation id="EFC94B5AFFB1FF984789520D6DBB88FA" author="Song DX &amp; Zhu MS &amp; Chen J" pageId="1" pageNumber="236" refId="ref10744" refString="Song DX, Zhu MS &amp; Chen J (1999) The spiders of China. Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp." type="book" year="1999">Song et al., 1999</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB1FF9845F2522D6CE288FA" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" box="[224,397,1405,1429]" pageId="1" pageNumber="236" refId="ref11534" refString="Yin CM, Peng XJ, Yan HM., Bao YH, Xu X, Tang G, Zhou QS &amp; Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">Yin et al., 2012</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB1FF98448B522E6F3388FA" author="Wang LY &amp; Marusik YM &amp; Zhang ZS" box="[409,604,1405,1430]" pageId="1" pageNumber="236" pagination="53 - 68" refId="ref11188" refString="Wang LY, Marusik YM &amp; Zhang ZS (2012) Notes on three poorly known Arctosa species from China (Araneae: Lycosidae). Zootaxa, 3404: 53 - 68." type="journal article" year="2012">Wang et al., 2012</bibRefCitation>
,
<bibRefCitation id="EFC94B5AFFB1FF98477A522D6FCE88FA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[616,673,1405,1429]" pageId="1" pageNumber="236" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">2021</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB1FF9847BF522D6D9F88DA" author="Zhang F &amp; Peng JY &amp; Zhang BS" pageId="1" pageNumber="236" refId="ref11648" refString="Zhang F, Peng JY &amp; Zhang BS (2022) Spiders of Mt. Xiaowutai. Science Press, Beijing, 387 pp." type="book" year="2022">Zhang et al., 2022</bibRefCitation>
, etc.).
</paragraph>
<paragraph id="8BE736ABFFB1FF984561528D6C548BFA" blockId="1.[112,752,1501,1685]" pageId="1" pageNumber="236">
The diversity of
<taxonomicName id="4C584D28FFB1FF984409528E6C02889A" authorityName="C. L. Koch" authorityYear="1847" box="[283,365,1502,1525]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB1FF984409528E6C02889A" box="[283,365,1502,1525]" italics="true" pageId="1" pageNumber="236">Arctosa</emphasis>
</taxonomicName>
in Southeast Asia was poorly studied until recently. Currently, five species of this genus are known from this region:
<taxonomicName id="4C584D28FFB1FF984438514E6F808B5A" authority="Barrion &amp; Litsinger, 1995" authorityName="Barrion &amp; Litsinger" authorityYear="1995" box="[298,751,1565,1589]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="tanakai">
<emphasis id="B92CEAB9FFB1FF984438514E6CBC8B5A" box="[298,467,1565,1589]" italics="true" pageId="1" pageNumber="236">Arctosa tanakai</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF9844C8514D6F808B5A" author="Barrion AT &amp; Litsinger JA" box="[474,751,1565,1589]" pageId="1" pageNumber="236" refId="ref9181" refString="Barrion AT &amp; Litsinger JA (1995) Riceland spiders of South and Southeast Asia. CAB International, Wallingford, UK, 700 pp." type="book" year="1995">Barrion &amp; Litsinger, 1995</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="F34F763BFFB1FF98456A516D6D998B3A" box="[120,246,1597,1621]" name="Philippines" pageId="1" pageNumber="236">Philippines</collectingCountry>
),
<taxonomicName id="4C584D28FFB1FF98441B516E6CFB8B3A" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[265,404,1597,1621]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF98441B516E6CFB8B3A" box="[265,404,1597,1621]" italics="true" pageId="1" pageNumber="236">A. delaportei</emphasis>
</taxonomicName>
,
<taxonomicName id="4C584D28FFB1FF9844B3516E6F548B3A" baseAuthorityName="Bosenberg &amp; Strand" baseAuthorityYear="1906" box="[417,571,1597,1621]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="depectinata">
<emphasis id="B92CEAB9FFB1FF9844B3516E6F548B3A" box="[417,571,1597,1621]" italics="true" pageId="1" pageNumber="236">A. depectinata</emphasis>
</taxonomicName>
,
<emphasis id="B92CEAB9FFB1FF98475B516E6F808B3A" box="[585,751,1597,1621]" italics="true" pageId="1" pageNumber="236">
<taxonomicName id="4C584D28FFB1FF98475B516E6F848B3A" baseAuthorityName="Schenkel" baseAuthorityYear="1963" box="[585,747,1597,1621]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="kiangsiensis">A. kiangsiensis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C584D28FFB1FF98458F510E6C478B1A" authorityName="Yin, Wang, Xie &amp; Peng" authorityYear="1993" box="[157,296,1630,1653]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="springiosa">
<emphasis id="B92CEAB9FFB1FF98458F510E6C478B1A" box="[157,296,1630,1653]" italics="true" pageId="1" pageNumber="236">A. springiosa</emphasis>
</taxonomicName>
(
<collectingCountry id="F34F763BFFB1FF984426510D6C058B1A" box="[308,362,1629,1653]" name="Laos" pageId="1" pageNumber="236">Laos</collectingCountry>
) (
<bibRefCitation id="EFC94B5AFFB1FF98446C510D6FFE8B1A" author="Barrion AT &amp; Litsinger JA" box="[382,657,1629,1653]" pageId="1" pageNumber="236" refId="ref9181" refString="Barrion AT &amp; Litsinger JA (1995) Riceland spiders of South and Southeast Asia. CAB International, Wallingford, UK, 700 pp." type="book" year="1995">Barrion &amp; Litsinger, 1995</bibRefCitation>
,
<bibRefCitation id="EFC94B5AFFB1FF984788510D6C5F8BFA" author="Omelko MM &amp; Marusik YM" pageId="1" pageNumber="236" pagination="397 - 406" refId="ref10251" refString="Omelko MM &amp; Marusik YM (2022) First data on Arctosa wolf spiders (Araneae: Lycosidae) from Laos with description of a new species. Raffles Bulletin of Zoology, 70: 397 - 406." type="journal article" year="2022">Omelko &amp; Marusik, 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,219 @@
<document id="528A383FB04893C64AB798E2426FAF01" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB1FF9B45A051ED6F898852" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Arctosa delaportei Omelko &amp; Marusik 2022" docType="treatment" docVersion="2" lastPageNumber="237" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="236" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C5F4D0C74A2C917FFF8CB8F0CEE03C38" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B6112E0AF23E67B60DAAB17D8C417771">
<mods:title id="C839BF93ADB69CC85B90BFFDB29E3C56">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="485E0A745988BB4725F54F8AED5F9DA1" type="personal">
<mods:role id="786C3338FFC087D7EA7ECADDAC22F409">
<mods:roleTerm id="71A8C1E28A40805A90857919BD868E7D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="280A08FB305FCB883F988AAC9B20B0CA">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A0B963F278110FD2408A38E45F9BBC4C">text</mods:typeOfResource>
<mods:relatedItem id="521935EBEC8D01448FFA3A12C87794C8" type="host">
<mods:titleInfo id="CA4E9C5FB768D06291C76C8CB0C1A398">
<mods:title id="7A3B48AA34FC6C3F833C2B48F9242FB4">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="24E9CF9F65204A6FFFBB1333C2E00248">
<mods:date id="F5D8F4C9BB98BA1FCF894C03AA4A7C02">2024</mods:date>
<mods:detail id="2AED069555A8FC69D82EFC1E86D1DB55" type="pubDate">
<mods:number id="170827ECFF4F24F1B60B9DFA7CED2308">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="8F2F74AF70D521DEA9C269F6B514D3E7" type="volume">
<mods:number id="9443494DA6EE75D6B016965DE7C07E91">72</mods:number>
</mods:detail>
<mods:extent id="81F62D48C56EAEE165CA8D2739EEB0C9" unit="page">
<mods:start id="5132B033754E4AFED09A8A53A072AE6E">235</mods:start>
<mods:end id="DF4342C53430BBE671FCC6BD8AB7D5DD">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A7A61696BBA68AF3AD1FECED8771496D">journal article</mods:classification>
<mods:identifier id="071C4ED8DA8166F1CA2596D9B8F8B2D1" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="58C1F4D507C0DBAA96A46D099EA8BC93" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="72C6CBD993C3CB53FD39BB29EEC37E34" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="8A1A3D218B36EFC24C0C1CB1FCD59760" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB1FF9B45A051ED6F898852" LSID="urn:lsid:plazi:treatment:03F187BDFFB1FF9B45A051ED6F898852" httpUri="http://treatment.plazi.org/id/03F187BDFFB1FF9B45A051ED6F898852" lastPageId="2" lastPageNumber="237" pageId="1" pageNumber="236">
<subSubSection id="C3426520FFB1FF9845A051ED6F7B8B9A" pageId="1" pageNumber="236" type="nomenclature">
<paragraph id="8BE736ABFFB1FF9845A051ED6FC18BBA" blockId="1.[178,686,1725,1781]" box="[178,686,1725,1749]" pageId="1" pageNumber="236">
<heading id="D0AF81C7FFB1FF9845A051ED6FC18BBA" bold="true" box="[178,686,1725,1749]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="236" reason="2">
<taxonomicName id="4C584D28FFB1FF9845A051ED6FC18BBA" ID-CoL="C67QP" authority="Omelko &amp; Marusik, 2022" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[178,686,1725,1749]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF9845A051ED6FC18BBA" bold="true" box="[178,686,1725,1749]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF9845A051ED6C118BBA" bold="true" box="[178,382,1725,1749]" italics="true" pageId="1" pageNumber="236">Arctosa delaportei</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF98449451ED6FC18BBA" author="Omelko MM &amp; Marusik YM" box="[390,686,1725,1749]" pageId="1" pageNumber="236" pagination="397 - 406" refId="ref10251" refString="Omelko MM &amp; Marusik YM (2022) First data on Arctosa wolf spiders (Araneae: Lycosidae) from Laos with description of a new species. Raffles Bulletin of Zoology, 70: 397 - 406." type="journal article" year="2022">Omelko &amp; Marusik, 2022</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE736ABFFB1FF98445E518D6F7B8B9A" blockId="1.[178,686,1725,1781]" box="[332,532,1757,1781]" pageId="1" pageNumber="236">(Figs. 15, 5253)</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF984563504C6C528A20" pageId="1" pageNumber="236" type="reference_group">
<paragraph id="8BE736ABFFB1FF984563504C6C528A20" blockId="1.[113,751,1820,1871]" pageId="1" pageNumber="236">
<treatmentCitationGroup id="AB481185FFB1FF984563504C6C528A20" pageId="1" pageNumber="236">
<treatmentCitation id="0AF910BAFFB1FF984563504C6FEE8A5E" author="Omelko MM &amp; Marusik YM" box="[113,641,1820,1842]" page="398" pageId="1" pageNumber="236" year="2022">
<taxonomicName id="4C584D28FFB1FF984563504C6FEE8A5E" ID-CoL="C67QP" authority="Omelko &amp; Marusik, 2022: 398" authorityName="Omelko &amp; Marusik" authorityPageNumber="398" authorityYear="2022" box="[113,641,1820,1842]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF984563504C6C5E8A5E" box="[113,305,1820,1841]" italics="true" pageId="1" pageNumber="236">Arctosa delaportei</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF984429504C6FEE8A5E" author="Omelko MM &amp; Marusik YM" box="[315,641,1820,1842]" pageId="1" pageNumber="236" pagination="397 - 406" refId="ref10251" refString="Omelko MM &amp; Marusik YM (2022) First data on Arctosa wolf spiders (Araneae: Lycosidae) from Laos with description of a new species. Raffles Bulletin of Zoology, 70: 397 - 406." type="journal article" year="2022">Omelko &amp; Marusik, 2022: 398</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, f. 1, 78, 1517, 26, male.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF98456350276F3B8A80" pageId="1" pageNumber="236" type="materials_examined">
<paragraph id="8BE736ABFFB1FF98456350276F3B8A80" blockId="1.[113,753,1911,2031]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98456350276C248AE0" bold="true" box="[113,331,1911,1935]" pageId="1" pageNumber="236">Material examined</emphasis>
.
<materialsCitation id="3B303CF6FFB1FF98444B50276F3F8A80" collectingDate="2016-05-22" collectingDateMax="2016-05-26" collectingDateMin="2016-05-22" collectionCode="ZMMU" collectorName="M. M. Omelko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="1" pageNumber="236" specimenCount="1" specimenCount-female="1" stateProvince="Viangchan">
<specimenCount id="9D5EFD22FFB1FF98444B50276CD78AE0" box="[345,440,1911,1935]" count="1" pageId="1" pageNumber="236" type="female">1 female</specimenCount>
(
<collectionCode id="ED49AE6EFFB1FF9844D550276F4F8AE0" box="[455,544,1911,1935]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="236" type="Museum">ZMMU</collectionCode>
),
<collectingCountry id="F34F763BFFB1FF98472150276F148AE0" box="[563,635,1911,1935]" name="Laos" pageId="1" pageNumber="236">LAOS</collectingCountry>
,
<collectingRegion id="499CF849FFB1FF98479A50276DDA8AC1" country="Laos" name="Viangchan" pageId="1" pageNumber="236">Vientiane Prov.</collectingRegion>
, env. of
<location id="8E876070FFB1FF98443A50C76F5F8AC0" LSID="urn:lsid:plazi:treatment:03F187BDFFB1FF9B45A051ED6F898852:8E876070FFB1FF98443A50C76F5F8AC0" box="[296,560,1943,1967]" country="Laos" latitude="18.614773" longLatPrecision="1" longitude="102.40885" name="Nam-Lik Eco-Village" pageId="1" pageNumber="236" stateProvince="Viangchan">Nam-Lik Eco-Village</location>
,
<geoCoordinate id="EE6C506CFFB1FF98475350C56F9F8AC1" box="[577,752,1941,1966]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="1" pageNumber="236" precision="1" seconds="53.18" value="18.614773">18°3653.18″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB1FF98456150E56C4C8AA1" box="[115,291,1973,1998]" degrees="102" direction="east" minutes="24" orientation="longitude" pageId="1" pageNumber="236" precision="1" seconds="31.87" value="102.40885">102°2431.87″E</geoCoordinate>
,
<collectingMethod id="52194EBCFFB1FF98442050E76CDC8AA1" box="[306,435,1975,1999]" pageId="1" pageNumber="236">pitfall traps</collectingMethod>
at forests edge near paddy field, coll.
<collectorName id="26AD537DFFB1FF9845FA50876CE98A80" box="[232,390,2007,2031]" pageId="1" pageNumber="236">M.M. Omelko</collectorName>
,
<date id="FFE6106BFFB1FF98448050876F3F8A80" box="[402,592,2007,2031]" pageId="1" pageNumber="236" value="2016-05-22" valueMax="2016-05-26" valueMin="2016-05-22">
<collectingDate id="EFA2E983FFB1FF98448050876F3F8A80" box="[402,592,2007,2031]" pageId="1" pageNumber="236" value="2016-05-22" valueMax="2016-05-26" valueMin="2016-05-22">2226 May 2016</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF98463F57ED69E58FDA" pageId="1" pageNumber="236" type="diagnosis">
<paragraph id="8BE736ABFFB1FF98463F57ED69E58FDA" blockId="1.[813,1452,189,694]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98463F57ED6ECE8DBA" bold="true" box="[813,929,189,213]" pageId="1" pageNumber="236">Diagnosis.</emphasis>
By the trapezoidal septum with short, wide stem and general conformation of epigyne, female of
<taxonomicName id="4C584D28FFB1FF984045578E6EF08C7A" authorityName="Omelko &amp; Marusik" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF984045578E6EF08C7A" italics="true" pageId="1" pageNumber="236">Arctosa delaportei</emphasis>
</taxonomicName>
is related to that of
<taxonomicName id="4C584D28FFB1FF98418757AE6E0A8C5A" authority="Chen &amp; Song, 1999" authorityName="Chen &amp; Song" authorityYear="1999" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="gougu">
<emphasis id="B92CEAB9FFB1FF98418757AE69938C7A" box="[1173,1276,254,277]" italics="true" pageId="1" pageNumber="236">A. gougu</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF98401557AD6E0A8C5A" author="Chen J &amp; Song DX" pageId="1" pageNumber="236" pagination="138 - 143" refId="ref9309" refString="Chen J &amp; Song DX (1999) On some species of the genus Arctosa from China (Araneae: Lycosidae). Acta Zootaxonomica Sinica, 24: 138 - 143." type="journal article" year="1999">Chen &amp; Song, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C584D28FFB1FF984661564E69698C5A" authorityName="Yin, Wang, Xie &amp; Peng" authorityYear="1993" box="[883,1030,286,309]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="springiosa">
<emphasis id="B92CEAB9FFB1FF984661564E69698C5A" box="[883,1030,286,309]" italics="true" pageId="1" pageNumber="236">A. springiosa</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C584D28FFB1FF98415A564E6E098C3A" authority="Yin, Peng &amp; Bao, 1997" authorityName="Yin, Peng &amp; Bao" authorityYear="1997" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="ziyunensis">
<emphasis id="B92CEAB9FFB1FF98415A564E69B88C5A" box="[1096,1239,286,309]" italics="true" pageId="1" pageNumber="236">A. ziyunensis</emphasis>
<bibRefCitation id="EFC94B5AFFB1FF9841F0564D6E098C3A" author="Yin CM &amp; Peng XJ &amp; Bao YH" pageId="1" pageNumber="236" pagination="1 - 5" refId="ref11384" refString="Yin CM, Peng XJ &amp; Bao YH (1997) Two new species of the genus Arctosa (Araneae, Lycosidae) from China. Acta Arachnologica Sinica, 6: 1 - 5." type="journal article" year="1997">Yin, Peng &amp; Bao, 1997</bibRefCitation>
</taxonomicName>
. Besides the similarity in its epigynes structure, the new species is close to
<taxonomicName id="4C584D28FFB1FF98413B560E69D78C1A" authorityName="Yin, Wang, Xie &amp; Peng" authorityYear="1993" box="[1065,1208,350,373]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="springiosa">
<emphasis id="B92CEAB9FFB1FF98413B560E69D78C1A" box="[1065,1208,350,373]" italics="true" pageId="1" pageNumber="236">A. springiosa</emphasis>
</taxonomicName>
in having very similar dark body colouration in dorsal view (cf. Fig. 1 and fig. 13B in
<bibRefCitation id="EFC94B5AFFB1FF98469156CE690C8CDA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[899,1123,413,438]" pageId="1" pageNumber="236" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
. All these species are known from
<collectingCountry id="F34F763BFFB1FF98467B56ED6EC68CBA" box="[873,937,445,469]" name="China" pageId="1" pageNumber="236">China</collectingCountry>
(
<taxonomicName id="4C584D28FFB1FF9846A856EE69248CBA" authorityName="Yin, Wang, Xie &amp; Peng" authorityYear="1993" box="[954,1099,446,469]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="springiosa">
<emphasis id="B92CEAB9FFB1FF9846A856EE69248CBA" box="[954,1099,446,469]" italics="true" pageId="1" pageNumber="236">A. springiosa</emphasis>
</taxonomicName>
from
<collectingCountry id="F34F763BFFB1FF98419D56ED69A08CBA" box="[1167,1231,445,469]" name="China" pageId="1" pageNumber="236">China</collectingCountry>
and
<collectingCountry id="F34F763BFFB1FF98401556ED68508CBA" box="[1287,1343,445,469]" name="Laos" pageId="1" pageNumber="236">Laos</collectingCountry>
). Females of
<taxonomicName id="4C584D28FFB1FF98465F568E6EB38C9A" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[845,988,477,501]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF98465F568E6EB38C9A" box="[845,988,477,501]" italics="true" pageId="1" pageNumber="236">A. delaportei</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="4C584D28FFB1FF984005568E68148C9A" authorityName="Chen &amp; Song" authorityYear="1999" box="[1303,1403,478,501]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="gougu">
<emphasis id="B92CEAB9FFB1FF984005568E68148C9A" box="[1303,1403,478,501]" italics="true" pageId="1" pageNumber="236">A. gougu</emphasis>
</taxonomicName>
and
<taxonomicName id="4C584D28FFB1FF98463F56AE6ED68F7A" authorityName="Yin, Peng &amp; Bao" authorityYear="1997" box="[813,953,510,533]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="ziyunensis">
<emphasis id="B92CEAB9FFB1FF98463F56AE6ED68F7A" box="[813,953,510,533]" italics="true" pageId="1" pageNumber="236">A. ziyunensis</emphasis>
</taxonomicName>
by the presence of large round glands (Gl) on its copulatory ducts (vs. absent; cf. Figs. 34 and fig. 1A in
<bibRefCitation id="EFC94B5AFFB1FF984659556E69698F3A" author="Pan F &amp; Zheng G &amp; Li SQ" box="[843,1030,573,597]" pageId="1" pageNumber="236" pagination="404 - 414" refId="ref10290" refString="Pan F, Zheng G &amp; Li SQ (2016) Wolf spiders (Araneae: Lycosidae) from Xishuangbanna Tropical Botanical Garden, China. Zoological Systematics, 41 (4): 404 - 414." type="journal article" year="2016">Pan et al. (2016)</bibRefCitation>
, fig. 406d in
<bibRefCitation id="EFC94B5AFFB1FF98418E556D680E8F3A" author="Yin CM &amp; Peng XJ &amp; Yan HM &amp; Bao YH &amp; Xu X &amp; Tang G &amp; Zhou QS &amp; Liu P" box="[1180,1377,573,597]" pageId="1" pageNumber="236" refId="ref11534" refString="Yin CM, Peng XJ, Yan HM., Bao YH, Xu X, Tang G, Zhou QS &amp; Liu P (2012) Fauna Hunan: Araneae in Hunan, China. Hunan Science and Technology Press, Changsha, 1590 pp." type="book" year="2012">Yin et al. (2012))</bibRefCitation>
. It can be easily differentiated from
<taxonomicName id="4C584D28FFB1FF98417E550E69918F1A" authorityName="Yin, Wang, Xie &amp; Peng" authorityYear="1993" box="[1132,1278,606,629]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="springiosa">
<emphasis id="B92CEAB9FFB1FF98417E550E69918F1A" box="[1132,1278,606,629]" italics="true" pageId="1" pageNumber="236">A. springiosa</emphasis>
</taxonomicName>
by the strongly curved copulatory ducts (CD); (vs. straight; cf. Fig. 5 and fig. 13H in
<bibRefCitation id="EFC94B5AFFB1FF9846BE55CE69E98FDA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[940,1158,669,694]" pageId="1" pageNumber="236" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB1FF9B463F558D6F618991" lastPageId="2" lastPageNumber="237" pageId="1" pageNumber="236" type="description">
<paragraph id="8BE736ABFFB1FF98463F558D68E58EDA" blockId="1.[811,1452,733,949]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98463F558D6ED98F9A" bold="true" box="[813,950,733,757]" pageId="1" pageNumber="236">Description.</emphasis>
Female (Figs. 12). Total length 5.47. Carapace 2.51 long, 1.58 wide. Opisthosoma 2.81 long, 1.65 wide. Colouration. Carapace dark brown, lacking distinct pattern, slightly lighter in central part. Lateral bands absent. Fovea almost invisible. Clypeus brown, chelicerae brown. Chelicera with 3 pro- and 3 retromarginal teeth. Labium and endites brown with yellowish outer edge. Sternum dark brown.
</paragraph>
<paragraph id="8BE736ABFFB1FF98463F548E6928893A" blockId="1.[813,1452,989,1109]" pageId="1" pageNumber="236">Eye sizes and interdistances: AME 0.09, ALE 0.08, PME 0.21, PLE 0.15; AMEAME 0.06, AMEALE 0.02, PME PME 0.10, PMEPLE 0.12, AMEPME 0.05, ALEPME 0.05; clypeus height 0.08.</paragraph>
<paragraph id="8BE736ABFFB1FF98463F532D69F1881A" blockId="1.[812,1452,1149,1397]" pageId="1" pageNumber="236">
Palpal femur dark brown; rest of segments brown. For palp and legs measurements see
<tableCitation id="C6DA0310FFB1FF98417553CD69D189DA" box="[1127,1214,1181,1205]" captionStart="Table 1" captionStartId="2.[835,890,1222,1244]" captionText="Table 1. Length of palp and leg segments for the female of Arctosa delaportei" pageId="1" pageNumber="236">Table 1</tableCitation>
. Coxae IIV yellow; femora I dark brown with yellow distal part, IIIV dark brown with yellow proximal part; patellae IIV dark brown; tibiae IIV dark brown with yellow ring in middle; metatarsi IIII dark brown with poorly visible annulation, IV dark brown with clearly visible annulation; tarsi IIV light brown. For spination of legs III see
<tableCitation id="C6DA0310FFB1FF984157520D69F5881A" box="[1093,1178,1373,1397]" captionStart="Table 2" captionStartId="2.[835,890,1664,1686]" captionText="Table 2. Leg spination for the female of Arctosa delaportei" pageId="1" pageNumber="236">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8BE736ABFFB1FF98463F52CD6E998B7A" blockId="1.[813,1452,1437,1557]" pageId="1" pageNumber="236">Abdomen dorsally dark grey, with poorly visible cardiac mark and irregular spots of short white setae. Lateral sides of the opisthosoma greyish. Ventral part dark brown. Spinnerets yellow.</paragraph>
<paragraph id="8BE736ABFFB1FF98463F516D695F8A7A" blockId="1.[813,1453,1597,1813]" pageId="1" pageNumber="236">Epigyne as shown in Figs. 35. Septum (Se) more or less trapezoidal in shape, covered with short, thick setae, septal stem (SS) short and wide. Hoods (Ho) large. Receptacles (Re) large, drop shaped, spaced by a distance 2 times their width. Copulatory ducts (CD) comparatively short, curved with pair of large round glands (Gl) clearly visible even on non-macerated epigyne.</paragraph>
<paragraph id="8BE736ABFFB1FF98463F506D68C3855A" blockId="1.[811,1453,1853,2101]" pageId="1" pageNumber="236">
<emphasis id="B92CEAB9FFB1FF98463F506D6E1C8A3A" bold="true" box="[813,883,1853,1877]" pageId="1" pageNumber="236">Notes.</emphasis>
<taxonomicName id="4C584D28FFB1FF98466B506E692E8A3A" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[889,1089,1853,1877]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="236" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB1FF98466B506E692E8A3A" box="[889,1089,1853,1877]" italics="true" pageId="1" pageNumber="236">Arctosa delaportei</emphasis>
</taxonomicName>
was described on the basis of
<specimenCount id="9D5EFD22FFB1FF984091506E6E058A1A" count="2" pageId="1" pageNumber="236" type="male">two males</specimenCount>
only, the female was reported as unknown (
<bibRefCitation id="EFC94B5AFFB1FF98402B500D6EA18AFA" author="Omelko MM &amp; Marusik YM" pageId="1" pageNumber="236" pagination="397 - 406" refId="ref10251" refString="Omelko MM &amp; Marusik YM (2022) First data on Arctosa wolf spiders (Araneae: Lycosidae) from Laos with description of a new species. Raffles Bulletin of Zoology, 70: 397 - 406." type="journal article" year="2022">Omelko &amp; Marusik, 2022</bibRefCitation>
). While further studying spiders collected in
<collectingCountry id="F34F763BFFB1FF98463F50CE6E0C8ADA" box="[813,867,1950,1973]" name="Laos" pageId="1" pageNumber="236">Laos</collectingCountry>
, I unexpectedly found a single female of this species which was lost among other wolf spiders during initial sorting. Correctness of male/female matching is confirmed by: 1) the fact that the female was collected in the same locality and habitat as the males 2) both sexes were collected during the
</paragraph>
<paragraph id="8BE736ABFFB2FF9B47245722697C8DE5" blockId="2.[566,1043,114,138]" box="[566,1043,114,138]" pageId="2" pageNumber="237">
<emphasis id="B92CEAB9FFB2FF9B472457226EBA8DE6" box="[566,981,114,137]" italics="true" pageId="2" pageNumber="237">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2024
</paragraph>
<paragraph id="8BE736ABFFB2FF9B459A530C68AC89E1" blockId="2.[136,1475,1116,1166]" pageId="2" pageNumber="237">
Figs. 15. Habitus (12) and epigyne (35) of
<taxonomicName id="4C584D28FFB2FF9B474E530C6E7D891E" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[604,786,1116,1137]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="237" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFB2FF9B474E530C6E7D891E" box="[604,786,1116,1137]" italics="true" pageId="2" pageNumber="237">Arctosa delaportei</emphasis>
</taxonomicName>
, female. 1, 5 dorsal; 2, 3, 4 ventral. Scale:
<quantity id="4CA09B4EFFB2FF9B41E3530C6845891E" box="[1265,1322,1116,1137]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="237" unit="mm" value="1.0">1 mm</quantity>
(12),
<quantity id="4CA09B4EFFB2FF9B4064530C68AD891E" box="[1398,1474,1116,1137]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="237" unit="mm" value="0.2">0.2 mm</quantity>
(35). Abbreviations: CD copulatory duct, FD fertilisation duct, Gl gland, Ho hood, Re receptacle, Se septum, SS septal stem.
</paragraph>
<paragraph id="8BE736ABFFB2FF9B459A53966F618991" blockId="2.[136,775,1222,1278]" pageId="2" pageNumber="237">same period of the year (summer) and 3) the female has the exact same colouration as the male.</paragraph>
</subSubSection>
<caption id="DF276623FFB2FF9B465153966EC98998" pageId="2" pageNumber="237" startId="2.[835,890,1222,1244]" targetType="table">
<paragraph id="8BE736ABFFB2FF9B465153966EC98998" blockId="2.[835,1474,1222,1271]" pageId="2" pageNumber="237">
Table 1. Length of palp and leg segments for the female of
<taxonomicName id="4C584D28FFB2FF9B406553966EC98998" authorityName="Omelko &amp; Marusik" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="237" phylum="Arthropoda" rank="species" species="delaportei">Arctosa delaportei</taxonomicName>
</paragraph>
</caption>
<subSubSection id="C3426520FFB2FF9B459A52756F898852" box="[136,742,1317,1342]" pageId="2" pageNumber="237" type="distribution">
<paragraph id="8BE736ABFFB2FF9B459A52756F898852" blockId="2.[136,742,1317,1342]" box="[136,742,1317,1342]" pageId="2" pageNumber="237">
<emphasis id="B92CEAB9FFB2FF9B459A52756C7C8852" bold="true" box="[136,275,1317,1341]" pageId="2" pageNumber="237">Distribution</emphasis>
.
<typeStatus id="54E38809FFB2FF9B443152766C358851" box="[291,346,1318,1342]" pageId="2" pageNumber="237">Type</typeStatus>
locality in
<collectingCountry id="F34F763BFFB2FF9B44CA52766F638852" box="[472,524,1318,1341]" name="Laos" pageId="2" pageNumber="237">Laos</collectingCountry>
only (
<figureCitation id="13632A2EFFB2FF9B474752766FB58852" box="[597,730,1317,1342]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="2" pageNumber="237">Figs. 5253</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,479 @@
<document id="077BB0879C33E2B8F5E2A8923D6CE652" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB3FF9F45EF516D6EB38A55" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Pardosa zhishengi Omelko, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="241" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="238" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BFDD147D63A10DEA30BD0D5926A682C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6769F7E9DA7226C32BFAA83BC368D22B">
<mods:title id="AE634231963EB786F48DD3BE0B9D459D">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="DD080FAAC43C4D6B4427097BE601A6B8" type="personal">
<mods:role id="7BB5BBD4DF3D31A5843F7B6F0F3B528C">
<mods:roleTerm id="E9FD29033CD4CFFB996A0555A15204EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B026EB49513A5B96311675F43AA0FBCB">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7DDA328C22D502D3A7188A77B0C8BEA3">text</mods:typeOfResource>
<mods:relatedItem id="981E0E17E76A8B1B1BB958389166A202" type="host">
<mods:titleInfo id="EFD6EA282E677E34FE2C4B9C6F7333CD">
<mods:title id="3952F63C4CD22A77229754ED219718CE">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="24C87911F8BD97D676105640A2E38895">
<mods:date id="46ACF5B323563BC4864E67E03CB180D2">2024</mods:date>
<mods:detail id="56456B0629EEA63A5DD812C8FD9CFD98" type="pubDate">
<mods:number id="27F2E59467969A8DA8FE332846CC0049">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="300211A30A2006C7B2EFCF01661EF30F" type="volume">
<mods:number id="6A1579D5CEA914AB8D96CF42C8C50318">72</mods:number>
</mods:detail>
<mods:extent id="385CA4D2F3041FA9A23E340F4F82600E" unit="page">
<mods:start id="2D3B52BC3A9EB360FC0A0F07B288EB6D">235</mods:start>
<mods:end id="B91EB1715EB0D672D8C148B5B0A9C6BE">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="657D7CE6A2E2CDC56F8CF88DFA675BD6">journal article</mods:classification>
<mods:identifier id="9D74A70990BA34826894E10A66FEEE99" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="035550FF0D66BD6733AC73EB068A83E8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="FA1C95B616C13231476F9C8C11D2EDB3" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="58C2E41275AF4E40557670E08D6F1862" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB3FF9F45EF516D6EB38A55" LSID="urn:lsid:plazi:treatment:03F187BDFFB3FF9F45EF516D6EB38A55" httpUri="http://treatment.plazi.org/id/03F187BDFFB3FF9F45EF516D6EB38A55" lastPageId="6" lastPageNumber="241" pageId="3" pageNumber="238">
<subSubSection id="C3426520FFB3FF9A45EF516D6F0B8B3A" box="[253,612,1597,1621]" pageId="3" pageNumber="238" type="nomenclature">
<paragraph id="8BE736ABFFB3FF9A45EF516D6F0B8B3A" blockId="3.[253,612,1597,1653]" box="[253,612,1597,1621]" pageId="3" pageNumber="238">
<heading id="D0AF81C7FFB3FF9A45EF516D6F0B8B3A" bold="true" box="[253,612,1597,1621]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="238" reason="2">
<emphasis id="B92CEAB9FFB3FF9A45EF516D6F0B8B3A" bold="true" box="[253,612,1597,1621]" pageId="3" pageNumber="238">
<taxonomicName id="4C584D28FFB3FF9A45EF516D6CA68B3A" authorityName="Omelko" authorityYear="2024" box="[253,457,1597,1621]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB3FF9A45EF516D6CA68B3A" bold="true" box="[253,457,1597,1621]" italics="true" pageId="3" pageNumber="238">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB3FF9A44CA516D6F0F8B3A" box="[472,608,1597,1621]" pageId="3" pageNumber="238" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB3FF9A4457510D6F748B1A" box="[325,539,1629,1653]" pageId="3" pageNumber="238" type="description">
<paragraph id="8BE736ABFFB3FF9A4457510D6F748B1A" blockId="3.[253,612,1597,1653]" box="[325,539,1629,1653]" pageId="3" pageNumber="238">
(
<figureCitation id="13632A2EFFB3FF9A445F510D6CAC8B1A" box="[333,451,1629,1653]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStartId-0="4.[136,180,1264,1286]" captionStartId-1="5.[113,157,2006,2028]" captionStartId-2="6.[135,179,1240,1262]" captionTargetBox-0="[136,1474,189,1249]" captionTargetBox-1="[139,1433,208,1991]" captionTargetBox-2="[135,1473,189,1225]" captionTargetId-0="figure-35@4.[136,1474,189,1249]" captionTargetId-1="figure-10@5.[134,1433,189,1991]" captionTargetId-2="figure-7@6.[135,1473,189,1225]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figs. 69. Habitus of Pardosa zhishengi, new species. 6 male, dorsal; 7 ditto, ventral; 8 female, dorsal; 9 ditto, ventral. Scale: 2 mm." captionText-1="Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of Pardosa zhishengi, new species. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles." captionText-2="Figs. 1921. Epigyne of Pardosa zhishengi, new species. 19, 20 ventral; 21 dorsal. Scale: 0.1 mm. Abbreviations: CD copulatory duct, Ho hood, Re receptacle, SB septal base, SS septal stem, SW septal “wings”." figureDoi-0="http://doi.org/10.5281/zenodo.14683142" figureDoi-1="http://doi.org/10.5281/zenodo.14683144" figureDoi-2="http://doi.org/10.5281/zenodo.14683146" httpUri-0="https://zenodo.org/record/14683142/files/figure.png" httpUri-1="https://zenodo.org/record/14683144/files/figure.png" httpUri-2="https://zenodo.org/record/14683146/files/figure.png" pageId="3" pageNumber="238">Figs. 621</figureCitation>
,
<figureCitation id="13632A2EFFB3FF9A44C2510D6F7B8B1A" box="[464,532,1629,1653]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="3" pageNumber="238">5253</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB3FF9A456351CD6C6A8A5A" pageId="3" pageNumber="238" type="materials_examined">
<paragraph id="8BE736ABFFB3FF9A456351CD6C6A8A5A" blockId="3.[113,753,1693,1845]" pageId="3" pageNumber="238">
<emphasis id="B92CEAB9FFB3FF9A456351CD6C328BDA" bold="true" box="[113,349,1693,1717]" pageId="3" pageNumber="238">Material examined.</emphasis>
<materialsCitation id="3B303CF6FFB3FF9A447A51CD6F828B9A" collectingDate="2013-11-01" collectingDateMax="2013-11-06" collectingDateMin="2013-11-01" collectionCode="ZMMU" collectorName="M. M. Omelko" country="Laos" latitude="15.193723" location="Tad" longLatPrecision="1" longitude="106.09925" municipality="Male" pageId="3" pageNumber="238" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
<typeStatus id="54E38809FFB3FF9A447A51CD6CBB8BDA" box="[360,468,1693,1717]" pageId="3" pageNumber="238" type="holotype">Holotype</typeStatus>
:
<collectingMunicipality id="6B83ACD1FFB3FF9A44F151CD6F738BDA" box="[483,540,1693,1717]" pageId="3" pageNumber="238">Male</collectingMunicipality>
(
<collectionCode id="ED49AE6EFFB3FF9A473C51CD6FE58BDA" box="[558,650,1693,1717]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="238" type="Museum">ZMMU</collectionCode>
),
<collectingCountry id="F34F763BFFB3FF9A47B351CD6F848BDA" box="[673,747,1693,1717]" name="Laos" pageId="3" pageNumber="238">LAOS</collectingCountry>
,
<collectingRegion id="499CF849FFB3FF9A456051ED6C5C8BBA" box="[114,307,1725,1749]" country="Laos" name="Champasak" pageId="3" pageNumber="238">Champasak Prov.</collectingRegion>
, env. of
<location id="8E876070FFB3FF9A448151ED6CD38BBA" LSID="urn:lsid:plazi:treatment:03F187BDFFB3FF9F45EF516D6EB38A55:8E876070FFB3FF9A448151ED6CD38BBA" box="[403,444,1725,1749]" country="Laos" latitude="15.193723" longLatPrecision="1" longitude="106.09925" municipality="Male" name="Tad" pageId="3" pageNumber="238" stateProvince="Champasak">Tad</location>
E-Tu resort,
<geoCoordinate id="EE6C506CFFB3FF9A474251EC6F9E8BBA" box="[592,753,1724,1749]" degrees="15" direction="north" minutes="11" orientation="latitude" pageId="3" pageNumber="238" precision="1" seconds="37.40" value="15.193723">15°1137.40″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB3FF9A4563518D6C758B9A" box="[113,282,1756,1781]" degrees="106" direction="east" minutes="5" orientation="longitude" pageId="3" pageNumber="238" precision="1" seconds="57.28" value="106.09925">106° 557.28″E</geoCoordinate>
, coll.
<collectorName id="26AD537DFFB3FF9A444C518E6C928B9A" box="[350,509,1757,1781]" pageId="3" pageNumber="238">M.M. Omelko</collectorName>
,
<date id="FFE6106BFFB3FF9A471B518D6F828B9A" box="[521,749,1757,1781]" pageId="3" pageNumber="238" value="2013-11-01" valueMax="2013-11-06" valueMin="2013-11-01">
<collectingDate id="EFA2E983FFB3FF9A471B518D6F828B9A" box="[521,749,1757,1781]" pageId="3" pageNumber="238" value="2013-11-01" valueMax="2013-11-06" valueMin="2013-11-01">16 November 2013</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B303CF6FFB3FF9A456351AE6C6E8A5A" collectionCode="ZMMU" country="Laos" pageId="3" pageNumber="238" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Champasak" typeStatus="paratype">
<typeStatus id="54E38809FFB3FF9A456351AE6DB28A79" box="[113,221,1790,1814]" pageId="3" pageNumber="238" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5EFD22FFB3FF9A45F851AD6C2D8A7A" box="[234,322,1789,1813]" count="2" pageId="3" pageNumber="238" type="male">2 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB3FF9A444251AD6CDF8A7A" box="[336,432,1789,1813]" count="1" pageId="3" pageNumber="238" type="female">1 female</specimenCount>
(
<collectionCode id="ED49AE6EFFB3FF9A44D351AD6F758A7A" box="[449,538,1789,1813]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="3" pageNumber="238" type="Museum">ZMMU</collectionCode>
), same place, dates and collector
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB3FF9A4563500D6C648ABA" pageId="3" pageNumber="238" type="etymology">
<paragraph id="8BE736ABFFB3FF9A4563500D6C648ABA" blockId="3.[113,752,1885,2005]" pageId="3" pageNumber="238">
<emphasis id="B92CEAB9FFB3FF9A4563500D6D998A1A" bold="true" box="[113,246,1885,1909]" pageId="3" pageNumber="238">Etymology.</emphasis>
The specific name is a patronym in honour of my colleague Zhi-Sheng Zhang (
<collectingRegion id="499CF849FFB3FF9A4713502D6F128AFA" box="[513,637,1917,1941]" country="China" name="Chongqing" pageId="3" pageNumber="238">Chongqing</collectingRegion>
,
<collectingCountry id="F34F763BFFB3FF9A4798502D6FA18AFA" box="[650,718,1917,1941]" name="China" pageId="3" pageNumber="238">China</collectingCountry>
), a prominent taxonomist for his contribution to the study of Asian spiders.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB3FF9A456350AD6E818FBA" pageId="3" pageNumber="238" type="diagnosis">
<paragraph id="8BE736ABFFB3FF9A456350AD6E818FBA" blockId="3.[113,752,2045,2102]" lastBlockId="3.[811,1453,189,725]" pageId="3" pageNumber="238">
<emphasis id="B92CEAB9FFB3FF9A456350AD6D82857A" bold="true" box="[113,237,2045,2069]" pageId="3" pageNumber="238">Diagnosis.</emphasis>
By general appearance and structure of its copulatory organs (shape of cymbium, palea, tegular apophysis, and septum) both sexes of
<taxonomicName id="4C584D28FFB3FF9A41A057ED681C8DBA" authorityName="Omelko" authorityYear="2024" box="[1202,1395,189,213]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB3FF9A41A057ED681C8DBA" box="[1202,1395,189,213]" italics="true" pageId="3" pageNumber="238">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB3FF9A406D57EE6E148D9A" pageId="3" pageNumber="238" rank="species">new species</taxonomicNameLabel>
, are similar to
<taxonomicName id="4C584D28FFB3FF9A410D578E69B18D9A" authorityName="Wang &amp; Zhang" authorityYear="2014" box="[1055,1246,221,245]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="238" phylum="Arthropoda" rank="species" species="parathompsoni">
<emphasis id="B92CEAB9FFB3FF9A410D578E69B18D9A" box="[1055,1246,221,245]" italics="true" pageId="3" pageNumber="238">P. parathompsoni</emphasis>
</taxonomicName>
known from south
<collectingCountry id="F34F763BFFB3FF9A463F57AD6E028C7A" box="[813,877,253,277]" name="China" pageId="3" pageNumber="238">China</collectingCountry>
and
<collectingCountry id="F34F763BFFB3FF9A468D57AD6EB68C7A" box="[927,985,253,277]" name="India" pageId="3" pageNumber="238">India</collectingCountry>
. Males of new species may be distinguished by the following features: 1) sperm duct (SD) is bent at an angle of almost 90° in ventral view (vs. almost straight, cf.
<figureCitation id="13632A2EFFB3FF9A463C560D6EDF8C1A" box="[814,944,349,373]" captionStart="Figs" captionStartId="5.[113,157,2006,2028]" captionTargetBox="[139,1433,208,1991]" captionTargetId="figure-10@5.[134,1433,189,1991]" captionTargetPageId="5" captionText="Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of Pardosa zhishengi, new species. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles." figureDoi="http://doi.org/10.5281/zenodo.14683144" httpUri="https://zenodo.org/record/14683144/files/figure.png" pageId="3" pageNumber="238">Figs. 10, 13</figureCitation>
and figs. 2D, F in
<bibRefCitation id="EFC94B5AFFB3FF9A4191560E68E68C1A" author="Wang D &amp; Zhang ZS" box="[1155,1417,349,374]" pageId="3" pageNumber="238" pagination="227 - 240" refId="ref11148" refString="Wang D &amp; Zhang ZS (2014) Two new species and a new synonym in the Pardosa nebulosa - group (Lycosidae: Pardosa) from China. Zootaxa, 3856 (2): 227 - 240." type="journal article" year="2014">Wang &amp; Zhang (2014))</bibRefCitation>
, 2) tegular apophysis (TA) pointing prolaterally (vs. anterioprolaterally,
<figureCitation id="13632A2EFFB3FF9A46A556CD69568CDA" box="[951,1081,413,437]" captionStart="Figs" captionStartId="5.[113,157,2006,2028]" captionTargetBox="[139,1433,208,1991]" captionTargetId="figure-10@5.[134,1433,189,1991]" captionTargetPageId="5" captionText="Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of Pardosa zhishengi, new species. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles." figureDoi="http://doi.org/10.5281/zenodo.14683144" httpUri="https://zenodo.org/record/14683144/files/figure.png" pageId="3" pageNumber="238">Figs. 10, 13</figureCitation>
and fig. 2D, F in
<bibRefCitation id="EFC94B5AFFB3FF9A401356CE6E118CBA" author="Wang D &amp; Zhang ZS" pageId="3" pageNumber="238" pagination="227 - 240" refId="ref11148" refString="Wang D &amp; Zhang ZS (2014) Two new species and a new synonym in the Pardosa nebulosa - group (Lycosidae: Pardosa) from China. Zootaxa, 3856 (2): 227 - 240." type="journal article" year="2014">Wang &amp; Zhang (2014))</bibRefCitation>
, 3) terminal apophysis (Tr) long (vs. short, cf.
<figureCitation id="13632A2EFFB3FF9A406456ED6EF58C9A" captionStart="Figs" captionStartId="5.[113,157,2006,2028]" captionTargetBox="[139,1433,208,1991]" captionTargetId="figure-10@5.[134,1433,189,1991]" captionTargetPageId="5" captionText="Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of Pardosa zhishengi, new species. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles." figureDoi="http://doi.org/10.5281/zenodo.14683144" httpUri="https://zenodo.org/record/14683144/files/figure.png" pageId="3" pageNumber="238">Figs. 14, 17, 18</figureCitation>
and figs. 1C, 2 GH in
<bibRefCitation id="EFC94B5AFFB3FF9A41B1568E68C78C9A" author="Wang D &amp; Zhang ZS" box="[1187,1448,477,502]" pageId="3" pageNumber="238" pagination="227 - 240" refId="ref11148" refString="Wang D &amp; Zhang ZS (2014) Two new species and a new synonym in the Pardosa nebulosa - group (Lycosidae: Pardosa) from China. Zootaxa, 3856 (2): 227 - 240." type="journal article" year="2014">Wang &amp; Zhang (2014))</bibRefCitation>
. Females of new species may be distinguished by: 1) base of septum (SB) with semi-transparent “wings” (SW) on its anterior edge (vs. lacking such structures, cf.
<figureCitation id="13632A2EFFB3FF9A4039556D68C28F3A" box="[1323,1453,573,597]" captionStart="Figs" captionStartId="6.[135,179,1240,1262]" captionTargetBox="[135,1473,189,1225]" captionTargetId="figure-7@6.[135,1473,189,1225]" captionTargetPageId="6" captionText="Figs. 1921. Epigyne of Pardosa zhishengi, new species. 19, 20 ventral; 21 dorsal. Scale: 0.1 mm. Abbreviations: CD copulatory duct, Ho hood, Re receptacle, SB septal base, SS septal stem, SW septal “wings”." figureDoi="http://doi.org/10.5281/zenodo.14683146" httpUri="https://zenodo.org/record/14683146/files/figure.png" pageId="3" pageNumber="238">Figs. 1920</figureCitation>
and figs. 1D, 2I in
<bibRefCitation id="EFC94B5AFFB3FF9A4113550E68688F1A" author="Wang D &amp; Zhang ZS" box="[1025,1287,605,630]" pageId="3" pageNumber="238" pagination="227 - 240" refId="ref11148" refString="Wang D &amp; Zhang ZS (2014) Two new species and a new synonym in the Pardosa nebulosa - group (Lycosidae: Pardosa) from China. Zootaxa, 3856 (2): 227 - 240." type="journal article" year="2014">Wang &amp; Zhang (2014))</bibRefCitation>
, 2) receptacles (Re) with a strong constriction in the middle part (vs. of almost uniform width, cf.
<figureCitation id="13632A2EFFB3FF9A415B55CD69F98FDA" box="[1097,1174,669,693]" captionStart="Figs" captionStartId="6.[135,179,1240,1262]" captionTargetBox="[135,1473,189,1225]" captionTargetId="figure-7@6.[135,1473,189,1225]" captionTargetPageId="6" captionText="Figs. 1921. Epigyne of Pardosa zhishengi, new species. 19, 20 ventral; 21 dorsal. Scale: 0.1 mm. Abbreviations: CD copulatory duct, Ho hood, Re receptacle, SB septal base, SS septal stem, SW septal “wings”." figureDoi="http://doi.org/10.5281/zenodo.14683146" httpUri="https://zenodo.org/record/14683146/files/figure.png" pageId="3" pageNumber="238">Fig. 21</figureCitation>
and figs. 1E, 2J in
<bibRefCitation id="EFC94B5AFFB3FF9A407955CE6E858FBA" author="Wang D &amp; Zhang ZS" pageId="3" pageNumber="238" pagination="227 - 240" refId="ref11148" refString="Wang D &amp; Zhang ZS (2014) Two new species and a new synonym in the Pardosa nebulosa - group (Lycosidae: Pardosa) from China. Zootaxa, 3856 (2): 227 - 240." type="journal article" year="2014">Wang &amp; Zhang (2014))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB3FF9F463E55AD69A98BB5" lastPageId="6" lastPageNumber="241" pageId="3" pageNumber="238" type="description">
<paragraph id="8BE736ABFFB3FF9A463E55AD6967897A" blockId="3.[811,1452,765,1045]" pageId="3" pageNumber="238">
<emphasis id="B92CEAB9FFB3FF9A463E55AD6ED88E7A" bold="true" box="[812,951,765,789]" pageId="3" pageNumber="238">Description.</emphasis>
Male (
<typeStatus id="54E38809FFB3FF9A411355AD690D8E7A" box="[1025,1122,765,789]" pageId="3" pageNumber="238" type="holotype">holotype</typeStatus>
) (
<figureCitation id="13632A2EFFB3FF9A416455AD69B18E7A" box="[1142,1246,765,789]" captionStart="Figs" captionStartId="4.[136,180,1264,1286]" captionTargetBox="[136,1474,189,1249]" captionTargetId="figure-35@4.[136,1474,189,1249]" captionTargetPageId="4" captionText="Figs. 69. Habitus of Pardosa zhishengi, new species. 6 male, dorsal; 7 ditto, ventral; 8 female, dorsal; 9 ditto, ventral. Scale: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14683142" httpUri="https://zenodo.org/record/14683142/files/figure.png" pageId="3" pageNumber="238">Figs. 67</figureCitation>
). Total length 6.12. Carapace 3.21 long, 2.38 wide. Opisthosoma 2.89 long, 1.65 wide. Colouration. Carapace black with yellow median band widened near eye field. Widened part of median band with couple of black spots. Eye field black. Lateral bands yellow, distinct. Fovea thin, black. Clypeus and chelicerae black. Labium and endites black with yellow edges. Sternum black with yellow longitudinal stripe and three pairs of yellow spots near its edges.
</paragraph>
<paragraph id="8BE736ABFFB3FF9A463E536E692989DA" blockId="3.[812,1451,1085,1205]" pageId="3" pageNumber="238">Eye sizes and interdistances: AME 0.14, ALE 0.12, PME 0.31, PLE 0.26; AMEAME 0.08, AMEALE 0.04, PME PME 0.27, PMEPLE 0.33, AMEPME 0.12, ALEPME 0.16; clypeus height 0.16.</paragraph>
<paragraph id="8BE736ABFFB3FF9A463E538D6EDF8B7A" blockId="3.[811,1452,1245,1557]" pageId="3" pageNumber="238">
Palpal femur black; patella dark brown; tibia and cymbium black. For palp and leg measurements, see
<tableCitation id="C6DA0310FFB3FF9A41EA53AD6823887A" box="[1272,1356,1277,1301]" captionStart="Table 3" captionStartId="4.[135,190,1341,1363]" captionText="Table 3. Length of palp and leg segments for the male of Pardosa zhishengi, new species." pageId="3" pageNumber="238">Table 3</tableCitation>
. Coxae I light brown ventrally and black laterally, IIIV light brown; femora I dorsally brown, laterally and ventrally black with yellow proximal and distal parts, femora I dorsally and ventrally brown, laterally black with yellow proximal and distal parts; IIIIV brown with irregular black spots; patellae IIV light brown; tibiae and metatarsi IIV yellowish, without annulation; tarsi IIV yellowish. For leg spination, see
<tableCitation id="C6DA0310FFB3FF9A464552AD6EC38B7A" box="[855,940,1533,1557]" captionStart="Table 4" captionStartId="4.[836,891,1341,1363]" captionText="Table 4. Leg spination for the male of Pardosa zhishengi, new species." pageId="3" pageNumber="238">Table 4</tableCitation>
.
</paragraph>
<paragraph id="8BE736ABFFB3FF9A463E516D69AA8BBA" blockId="3.[812,1451,1597,1749]" pageId="3" pageNumber="238">Dorsal part of the opisthosoma black, with black cardiac mark bordered with yellow stripes and series of yellow spots posteriorly. Lateral sides of the opisthosoma black with lots of white spots. Ventral part light brown with grey longitudinal stripe. Spinnerets brown.</paragraph>
<paragraph id="8BE736ABFFB3FF9A463E51AD6EC7857A" blockId="3.[812,1453,1789,2069]" pageId="3" pageNumber="238">
Palp as shown in
<figureCitation id="13632A2EFFB3FF9A46CC51AD69338A7A" box="[990,1116,1789,1813]" captionStart="Figs" captionStartId="5.[113,157,2006,2028]" captionTargetBox="[139,1433,208,1991]" captionTargetId="figure-10@5.[134,1433,189,1991]" captionTargetPageId="5" captionText="Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of Pardosa zhishengi, new species. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles." figureDoi="http://doi.org/10.5281/zenodo.14683144" httpUri="https://zenodo.org/record/14683144/files/figure.png" pageId="3" pageNumber="238">Figs. 1018</figureCitation>
. Cymbium ca. 1.92 times longer than bulb. Tegulum (Te) with several wrinkles (TW) on its retrolateral side. Subtegulum (St) oval, located prolaterally. Tegular apophysis (TA) large, widened proximally with pointed tip curved ventrally. Terminal apophysis (Tr) long, crescent-like, sharply pointed. Conductor (Co) thin, semi-transparent. Palea (Pa) large, chitinised prolaterally with semi-transparent peak (Pe). Embolus (Em) long, thin, wavy near its tip.
</paragraph>
<paragraph id="8BE736ABFFB4FF9D47245722697C8DE5" blockId="4.[566,1043,114,138]" box="[566,1043,114,138]" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D472457226EBA8DE6" box="[566,981,114,137]" italics="true" pageId="4" pageNumber="239">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2024
</paragraph>
<caption id="DF276623FFB4FF9D459A53A068AC8869" ID-DOI="http://doi.org/10.5281/zenodo.14683142" ID-Zenodo-Dep="14683142" box="[136,1475,1264,1286]" httpUri="https://zenodo.org/record/14683142/files/figure.png" pageId="4" pageNumber="239" startId="4.[136,180,1264,1286]" targetBox="[136,1474,189,1249]" targetPageId="4" targetType="figure">
<paragraph id="8BE736ABFFB4FF9D459A53A068AC8869" blockId="4.[136,1475,1264,1286]" box="[136,1475,1264,1286]" pageId="4" pageNumber="239">
Figs. 69. Habitus of
<taxonomicName id="4C584D28FFB4FF9D444553A06F66886A" authorityName="Omelko" authorityYear="2024" box="[343,521,1264,1285]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB4FF9D444553A06F66886A" box="[343,521,1264,1285]" italics="true" pageId="4" pageNumber="239">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB4FF9D470753A16FE58869" box="[533,650,1264,1286]" pageId="4" pageNumber="239" rank="species">new species</taxonomicNameLabel>
. 6 male, dorsal; 7 ditto, ventral; 8 female, dorsal; 9 ditto, ventral. Scale: 2 mm.
</paragraph>
</caption>
<caption id="DF276623FFB4FF9D4595526D6C03881F" pageId="4" pageNumber="239" startId="4.[135,190,1341,1363]" targetType="table">
<paragraph id="8BE736ABFFB4FF9D4595526D6C03881F" blockId="4.[135,774,1341,1392]" pageId="4" pageNumber="239">
Table 3. Length of palp and leg segments for the male of
<taxonomicName id="4C584D28FFB4FF9D47A7526D6D8D8800" authorityName="Omelko" authorityYear="2024" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB4FF9D47A7526D6D8D8800" italics="true" pageId="4" pageNumber="239">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB4FF9D45E2520A6C08881F" box="[240,359,1370,1392]" pageId="4" pageNumber="239" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<caption id="DF276623FFB4FF9D4656526D6EFF881F" pageId="4" pageNumber="239" startId="4.[836,891,1341,1363]" targetType="table">
<paragraph id="8BE736ABFFB4FF9D4656526D6EFF881F" blockId="4.[836,1474,1341,1392]" pageId="4" pageNumber="239">
Table 4. Leg spination for the male of
<taxonomicName id="4C584D28FFB4FF9D41C1526D68E4883D" authorityName="Omelko" authorityYear="2024" box="[1235,1419,1341,1362]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="239" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB4FF9D41C1526D68E4883D" box="[1235,1419,1341,1362]" italics="true" pageId="4" pageNumber="239">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB4FF9D4088526E6EE3881F" pageId="4" pageNumber="239" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BE736ABFFB4FF9D45EF52F26F888BC5" pageId="4" pageNumber="239">
<table id="F958C40BFFB40066458452F16F878BC5" box="[150,744,1441,1706]" gridcols="7" gridrows="6" pageId="4" pageNumber="239">
<tr id="356834E9FFB40066458452F16F8788D8" box="[150,744,1441,1463]" gridrow="0" pageId="4" pageNumber="239" rowspan-0="1">
<th id="76B95D95FFB4006645E752F16C7088D8" box="[245,287,1441,1463]" gridcol="1" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D45EF52F26C7988D8" bold="true" box="[253,278,1442,1463]" pageId="4" pageNumber="239">Fe</emphasis>
</th>
<th id="76B95D95FFB40066445F52F16C1888D8" box="[333,375,1441,1463]" gridcol="2" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D444652F26C0088D8" bold="true" box="[340,367,1442,1463]" pageId="4" pageNumber="239">Pa</emphasis>
</th>
<th id="76B95D95FFB4006644B652F16CA188D8" box="[420,462,1441,1463]" gridcol="3" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D44BC52F16CAA88D8" bold="true" box="[430,453,1441,1463]" pageId="4" pageNumber="239">Ti</emphasis>
</th>
<th id="76B95D95FFB4006644EE52F16F4988D8" box="[508,550,1441,1463]" gridcol="4" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D471052F26F4E88D8" bold="true" box="[514,545,1442,1463]" pageId="4" pageNumber="239">Mt</emphasis>
</th>
<th id="76B95D95FFB40066474652F16F1188D8" box="[596,638,1441,1463]" gridcol="5" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D474952F26F1888D8" bold="true" box="[603,631,1442,1463]" pageId="4" pageNumber="239">Ta</emphasis>
</th>
<th id="76B95D95FFB4006647A352F16F8788D8" box="[689,744,1441,1463]" gridcol="6" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D47A352F26F8788D8" bold="true" box="[689,744,1442,1463]" pageId="4" pageNumber="239">Total</emphasis>
</th>
</tr>
<tr id="356834E9FFB40066458452886F878881" box="[150,744,1496,1518]" gridrow="1" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066458452886DAE8881" box="[150,193,1496,1518]" gridcol="0" gridrow="1" pageId="4" pageNumber="239">Palp</th>
<td id="76B95D95FFB4006645E752886C708881" box="[245,287,1496,1518]" gridcol="1" gridrow="1" pageId="4" pageNumber="239">1.35</td>
<td id="76B95D95FFB40066445F52886C188881" box="[333,375,1496,1518]" gridcol="2" gridrow="1" pageId="4" pageNumber="239">0.66</td>
<td id="76B95D95FFB4006644B652886CA18881" box="[420,462,1496,1518]" gridcol="3" gridrow="1" pageId="4" pageNumber="239">0.88</td>
<td id="76B95D95FFB4006644EE52886F498881" box="[508,550,1496,1518]" gridcol="4" gridrow="1" pageId="4" pageNumber="239"></td>
<td id="76B95D95FFB40066474652886F118881" box="[596,638,1496,1518]" gridcol="5" gridrow="1" pageId="4" pageNumber="239">1.09</td>
<td id="76B95D95FFB4006647A352886F878881" box="[689,744,1496,1518]" gridcol="6" gridrow="1" pageId="4" pageNumber="239">3.98</td>
</tr>
<tr id="356834E9FFB40066458451576F878B72" box="[150,744,1543,1565]" gridrow="2" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066458451576DAE8B72" box="[150,193,1543,1565]" gridcol="0" gridrow="2" pageId="4" pageNumber="239">I</th>
<td id="76B95D95FFB4006645E751576C708B72" box="[245,287,1543,1565]" gridcol="1" gridrow="2" pageId="4" pageNumber="239">2.42</td>
<td id="76B95D95FFB40066445F51576C188B72" box="[333,375,1543,1565]" gridcol="2" gridrow="2" pageId="4" pageNumber="239">1.06</td>
<td id="76B95D95FFB4006644B651576CA18B72" box="[420,462,1543,1565]" gridcol="3" gridrow="2" pageId="4" pageNumber="239">2.18</td>
<td id="76B95D95FFB4006644EE51576F498B72" box="[508,550,1543,1565]" gridcol="4" gridrow="2" pageId="4" pageNumber="239">2.07</td>
<td id="76B95D95FFB40066474651576F118B72" box="[596,638,1543,1565]" gridcol="5" gridrow="2" pageId="4" pageNumber="239">1.46</td>
<td id="76B95D95FFB4006647A351576F878B72" box="[689,744,1543,1565]" gridcol="6" gridrow="2" pageId="4" pageNumber="239">9.19</td>
</tr>
<tr id="356834E9FFB40066458451666F878B23" box="[150,744,1590,1612]" gridrow="3" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066458451666DAE8B23" box="[150,193,1590,1612]" gridcol="0" gridrow="3" pageId="4" pageNumber="239">II</th>
<td id="76B95D95FFB4006645E751666C708B23" box="[245,287,1590,1612]" gridcol="1" gridrow="3" pageId="4" pageNumber="239">2.33</td>
<td id="76B95D95FFB40066445F51666C188B23" box="[333,375,1590,1612]" gridcol="2" gridrow="3" pageId="4" pageNumber="239">1.03</td>
<td id="76B95D95FFB4006644B651666CA18B23" box="[420,462,1590,1612]" gridcol="3" gridrow="3" pageId="4" pageNumber="239">1.99</td>
<td id="76B95D95FFB4006644EE51666F498B23" box="[508,550,1590,1612]" gridcol="4" gridrow="3" pageId="4" pageNumber="239">1.94</td>
<td id="76B95D95FFB40066474651666F118B23" box="[596,638,1590,1612]" gridcol="5" gridrow="3" pageId="4" pageNumber="239">1.31</td>
<td id="76B95D95FFB4006647A351666F878B23" box="[689,744,1590,1612]" gridcol="6" gridrow="3" pageId="4" pageNumber="239">8.6</td>
</tr>
<tr id="356834E9FFB40066458451356F878B14" box="[150,744,1637,1659]" gridrow="4" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066458451356DAE8B14" box="[150,193,1637,1659]" gridcol="0" gridrow="4" pageId="4" pageNumber="239">III</th>
<td id="76B95D95FFB4006645E751356C708B14" box="[245,287,1637,1659]" gridcol="1" gridrow="4" pageId="4" pageNumber="239">2.28</td>
<td id="76B95D95FFB40066445F51356C188B14" box="[333,375,1637,1659]" gridcol="2" gridrow="4" pageId="4" pageNumber="239">0.94</td>
<td id="76B95D95FFB4006644B651356CA18B14" box="[420,462,1637,1659]" gridcol="3" gridrow="4" pageId="4" pageNumber="239">1.91</td>
<td id="76B95D95FFB4006644EE51356F498B14" box="[508,550,1637,1659]" gridcol="4" gridrow="4" pageId="4" pageNumber="239">2.15</td>
<td id="76B95D95FFB40066474651356F118B14" box="[596,638,1637,1659]" gridcol="5" gridrow="4" pageId="4" pageNumber="239">1.19</td>
<td id="76B95D95FFB4006647A351356F878B14" box="[689,744,1637,1659]" gridcol="6" gridrow="4" pageId="4" pageNumber="239">8.47</td>
</tr>
<tr id="356834E9FFB40066458451C46F878BC5" box="[150,744,1684,1706]" gridrow="5" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066458451C46DAE8BC5" box="[150,193,1684,1706]" gridcol="0" gridrow="5" pageId="4" pageNumber="239">IV</th>
<td id="76B95D95FFB4006645E751C46C708BC5" box="[245,287,1684,1706]" gridcol="1" gridrow="5" pageId="4" pageNumber="239">2.92</td>
<td id="76B95D95FFB40066445F51C46C188BC5" box="[333,375,1684,1706]" gridcol="2" gridrow="5" pageId="4" pageNumber="239">1.03</td>
<td id="76B95D95FFB4006644B651C46CA18BC5" box="[420,462,1684,1706]" gridcol="3" gridrow="5" pageId="4" pageNumber="239">2.57</td>
<td id="76B95D95FFB4006644EE51C46F498BC5" box="[508,550,1684,1706]" gridcol="4" gridrow="5" pageId="4" pageNumber="239">3.49</td>
<td id="76B95D95FFB40066474651C46F118BC5" box="[596,638,1684,1706]" gridcol="5" gridrow="5" pageId="4" pageNumber="239">1.67</td>
<td id="76B95D95FFB4006647A351C46F878BC5" box="[689,744,1684,1706]" gridcol="6" gridrow="5" pageId="4" pageNumber="239">11.68</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFB4FF9D46A052F268D88B1D" pageId="4" pageNumber="239">
<table id="F958C40BFFB40066464152F168D88B1C" box="[851,1463,1441,1651]" gridcols="5" gridrows="5" pageId="4" pageNumber="239">
<tr id="356834E9FFB40066464152F168D888D8" box="[851,1463,1441,1463]" gridrow="0" pageId="4" pageNumber="239" rowspan-0="1">
<th id="76B95D95FFB40066469A52F16E9988D8" box="[904,1014,1441,1463]" gridcol="1" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D46A052F26EA488D8" bold="true" box="[946,971,1442,1463]" pageId="4" pageNumber="239">Fe</emphasis>
</th>
<th id="76B95D95FFB40066410152F1690888D8" box="[1043,1127,1441,1463]" gridcol="2" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D412252F2692488D8" bold="true" box="[1072,1099,1442,1463]" pageId="4" pageNumber="239">Pa</emphasis>
</th>
<th id="76B95D95FFB40066419B52F1687E88D8" box="[1161,1297,1441,1463]" gridcol="3" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D41D352F169B788D8" bold="true" box="[1217,1240,1441,1463]" pageId="4" pageNumber="239">Ti</emphasis>
</th>
<th id="76B95D95FFB40066403552F168D888D8" box="[1319,1463,1441,1463]" gridcol="4" gridrow="0" pageId="4" pageNumber="239">
<emphasis id="B92CEAB9FFB4FF9D407252F2681088D8" bold="true" box="[1376,1407,1442,1463]" pageId="4" pageNumber="239">Mt</emphasis>
</th>
</tr>
<tr id="356834E9FFB400664641528068D88889" box="[851,1463,1488,1510]" gridrow="1" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066464152806E038889" box="[851,876,1488,1510]" gridcol="0" gridrow="1" pageId="4" pageNumber="239">I</th>
<td id="76B95D95FFB40066469A52806E998889" box="[904,1014,1488,1510]" gridcol="1" gridrow="1" pageId="4" pageNumber="239">3d 2p 2(3)r</td>
<td id="76B95D95FFB400664101528069088889" box="[1043,1127,1488,1510]" gridcol="2" gridrow="1" pageId="4" pageNumber="239">1p 1r</td>
<td id="76B95D95FFB40066419B5280687E8889" box="[1161,1297,1488,1510]" gridcol="3" gridrow="1" pageId="4" pageNumber="239">2p 2r 2-2v</td>
<td id="76B95D95FFB400664035528068D88889" box="[1319,1463,1488,1510]" gridcol="4" gridrow="1" pageId="4" pageNumber="239">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFB40066464152AF68D88B7A" box="[851,1463,1535,1557]" gridrow="2" pageId="4" pageNumber="239">
<th id="76B95D95FFB40066464152AF6E038B7A" box="[851,876,1535,1557]" gridcol="0" gridrow="2" pageId="4" pageNumber="239">II</th>
<td id="76B95D95FFB40066469A52AF6E998B7A" box="[904,1014,1535,1557]" gridcol="1" gridrow="2" pageId="4" pageNumber="239">3d 2p 2r</td>
<td id="76B95D95FFB40066410152AF69088B7A" box="[1043,1127,1535,1557]" gridcol="2" gridrow="2" pageId="4" pageNumber="239">1p 1r</td>
<td id="76B95D95FFB40066419B52AF687E8B7A" box="[1161,1297,1535,1557]" gridcol="3" gridrow="2" pageId="4" pageNumber="239">2p 2r 2-2v</td>
<td id="76B95D95FFB40066403552AF68D88B7A" box="[1319,1463,1535,1557]" gridcol="4" gridrow="2" pageId="4" pageNumber="239">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFB400664641517E68D88B2B" box="[851,1463,1582,1604]" gridrow="3" pageId="4" pageNumber="239">
<th id="76B95D95FFB400664641517E6E038B2B" box="[851,876,1582,1604]" gridcol="0" gridrow="3" pageId="4" pageNumber="239">III</th>
<td id="76B95D95FFB40066469A517E6E998B2B" box="[904,1014,1582,1604]" gridcol="1" gridrow="3" pageId="4" pageNumber="239">3d 2p 2r</td>
<td id="76B95D95FFB400664101517E69088B2B" box="[1043,1127,1582,1604]" gridcol="2" gridrow="3" pageId="4" pageNumber="239">2d 1p 1r</td>
<td id="76B95D95FFB40066419B517E687E8B2B" box="[1161,1297,1582,1604]" gridcol="3" gridrow="3" pageId="4" pageNumber="239">2d 2p 2r 2-2v</td>
<td id="76B95D95FFB400664035517E68D88B2B" box="[1319,1463,1582,1604]" gridcol="4" gridrow="3" pageId="4" pageNumber="239">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFB400664641510D68D88B1C" box="[851,1463,1629,1651]" gridrow="4" pageId="4" pageNumber="239">
<th id="76B95D95FFB400664641510D6E038B1C" box="[851,876,1629,1651]" gridcol="0" gridrow="4" pageId="4" pageNumber="239">IV</th>
<td id="76B95D95FFB40066469A510D6E998B1C" box="[904,1014,1629,1651]" gridcol="1" gridrow="4" pageId="4" pageNumber="239">3d 2p 1r</td>
<td id="76B95D95FFB400664101510D69088B1C" box="[1043,1127,1629,1651]" gridcol="2" gridrow="4" pageId="4" pageNumber="239">2d 1p 1r</td>
<td id="76B95D95FFB40066419B510D687E8B1C" box="[1161,1297,1629,1651]" gridcol="3" gridrow="4" pageId="4" pageNumber="239">2d 2p 2r 2-2v</td>
<td id="76B95D95FFB400664035510D68D88B1C" box="[1319,1463,1629,1651]" gridcol="4" gridrow="4" pageId="4" pageNumber="239">2p 2r 1-1-1-1v</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFB4FF9D459A51A86CD38ADF" blockId="4.[135,775,1784,1968]" pageId="4" pageNumber="239">
Female (one of the
<typeStatus id="54E38809FFB4FF9D444751A86CD28A7F" box="[341,445,1784,1808]" pageId="4" pageNumber="239" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="13632A2EFFB4FF9D44C051A86F578A7F" box="[466,568,1784,1808]" captionStart="Figs" captionStartId="4.[136,180,1264,1286]" captionTargetBox="[136,1474,189,1249]" captionTargetId="figure-35@4.[136,1474,189,1249]" captionTargetPageId="4" captionText="Figs. 69. Habitus of Pardosa zhishengi, new species. 6 male, dorsal; 7 ditto, ventral; 8 female, dorsal; 9 ditto, ventral. Scale: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14683142" httpUri="https://zenodo.org/record/14683142/files/figure.png" pageId="4" pageNumber="239">Figs. 89</figureCitation>
). Total length 6.78. Carapace 3.39 long, 2.44 wide. Opisthosoma 3.61 long, 2.09 wide. Colouration. Carapace dorsally like in male but much lighter. Clypeus yellow, chelicerae light brown. Labium grey with light brown outer edge. Endites light brown. Sternum light brown without pattern.
</paragraph>
<paragraph id="8BE736ABFFB4FF9D459A50896E69857F" blockId="4.[136,775,2008,2065]" pageId="4" pageNumber="239">Eye sizes and interdistances: AME 0.12, ALE 0.10, PME 0.36, PLE 0.31; AMEAME 0.12, AMEALE 0.05, PME</paragraph>
<paragraph id="8BE736ABFFB4FF9D4651519269328B95" blockId="4.[835,1474,1729,1786]" pageId="4" pageNumber="239">PME 0.30, PMEPLE 0.39, AMEPME 0.15, ALEPME 0.12; clypeus height 0.21.</paragraph>
<paragraph id="8BE736ABFFB4FF9D4651507168798AB6" blockId="4.[835,1475,1825,2009]" pageId="4" pageNumber="239">
Palpal femur light brown with grey spots distally; other segments light brown. For palp and legs measurements see
<tableCitation id="C6DA0310FFB4FF9D465750316EF68A16" box="[837,921,1889,1913]" captionStart="Table 5" captionStartId="6.[136,191,1346,1368]" captionText="Table 5. Length of palp and leg segments for the female of Pardosa zhishengi, new species." pageId="4" pageNumber="239">Table 5</tableCitation>
. Coxae IIV light brown; femora IIV light brown with greyish spots; patellae IIV light brown; tibiae and metatarsi IIV light brown, without annulation; tarsi IIV light brown. For leg spination see
<tableCitation id="C6DA0310FFB4FF9D41AE5091687D8AB6" box="[1212,1298,1985,2009]" captionStart="Table 6" captionStartId="6.[136,191,1789,1811]" captionText="Table 6. Leg spination for the female of Pardosa zhishengi, new species." pageId="4" pageNumber="239">Table 6</tableCitation>
.
</paragraph>
<caption id="DF276623FFB5FF9C456350866F21852C" ID-DOI="http://doi.org/10.5281/zenodo.14683144" ID-Zenodo-Dep="14683144" httpUri="https://zenodo.org/record/14683144/files/figure.png" pageId="5" pageNumber="240" startId="5.[113,157,2006,2028]" targetBox="[139,1433,208,1991]" targetPageId="5" targetType="figure">
<paragraph id="8BE736ABFFB5FF9C456350866F21852C" blockId="5.[113,1454,2006,2115]" pageId="5" pageNumber="240">
Figs. 1018. Male palp (1011), bulb (1215) and embolic division (1618) of
<taxonomicName id="4C584D28FFB5FF9C466C508669408A84" authorityName="Omelko" authorityYear="2024" box="[894,1071,2006,2027]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB5FF9C466C508669408A84" box="[894,1071,2006,2027]" italics="true" pageId="5" pageNumber="240">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB5FF9C412F508769DB8A83" box="[1085,1204,2006,2028]" pageId="5" pageNumber="240" rank="species">new species</taxonomicNameLabel>
. 10, 13, 16 ventral; 11, 14 retrolateral; 12 prolateral; 15, 17 anterior; 18 ventro-retrolateral. Scale: 0.2 mm (1015), 0.1 mm (1618). Abbreviations: Co conductor, Em embolus, Pa palea, Pe paleal peack, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, Tr terminal apophysis, TW tegular wrinkles.
</paragraph>
</caption>
<paragraph id="8BE736ABFFB6FF9F47245722697C8DE5" blockId="6.[566,1043,114,138]" box="[566,1043,114,138]" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F472457226EBA8DE6" box="[566,981,114,137]" italics="true" pageId="6" pageNumber="241">RAFFLES BULLETIN OF ZOOLOGY</emphasis>
2024
</paragraph>
<caption id="DF276623FFB6FF9F45955388683C8864" ID-DOI="http://doi.org/10.5281/zenodo.14683146" ID-Zenodo-Dep="14683146" httpUri="https://zenodo.org/record/14683146/files/figure.png" pageId="6" pageNumber="241" startId="6.[135,179,1240,1262]" targetBox="[135,1473,189,1225]" targetPageId="6" targetType="figure">
<paragraph id="8BE736ABFFB6FF9F4595538869148981" blockId="6.[135,1363,1240,1291]" box="[135,1147,1240,1262]" pageId="6" pageNumber="241">
Figs. 1921. Epigyne of
<taxonomicName id="4C584D28FFB6FF9F446E53886F5F8982" authorityName="Omelko" authorityYear="2024" box="[380,560,1240,1261]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB6FF9F446E53886F5F8982" box="[380,560,1240,1261]" italics="true" pageId="6" pageNumber="241">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB6FF9F472F53896FDA8981" box="[573,693,1240,1262]" pageId="6" pageNumber="241" rank="species">new species</taxonomicNameLabel>
. 19, 20 ventral; 21 dorsal. Scale: 0.1 mm.
</paragraph>
<paragraph id="8BE736ABFFB6FF9F459553A5683C8864" blockId="6.[135,1363,1240,1291]" box="[135,1363,1269,1291]" pageId="6" pageNumber="241">Abbreviations: CD copulatory duct, Ho hood, Re receptacle, SB septal base, SS septal stem, SW septal “wings”.</paragraph>
</caption>
<caption id="DF276623FFB6FF9F459A52126C03881A" pageId="6" pageNumber="241" startId="6.[136,191,1346,1368]" targetType="table">
<paragraph id="8BE736ABFFB6FF9F459A52126C03881A" blockId="6.[136,774,1346,1397]" pageId="6" pageNumber="241">
Table 5. Length of palp and leg segments for the female of
<taxonomicName id="4C584D28FFB6FF9F47A752126D8C881B" authorityName="Omelko" authorityYear="2024" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB6FF9F47A752126D8C881B" italics="true" pageId="6" pageNumber="241">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB6FF9F45E252306C07881A" box="[240,360,1375,1397]" pageId="6" pageNumber="241" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<paragraph id="8BE736ABFFB6FF9F4651521268AC88F5" blockId="6.[835,1475,1346,1435]" pageId="6" pageNumber="241">Dorsal part of the opisthosoma like in males but much lighter. Lateral sides of the opisthosoma light brown with grey spots. Ventral part light yellow. Spinnerets light brown.</paragraph>
<paragraph id="8BE736ABFFB6FF9F45E552F76F898BC1" pageId="6" pageNumber="241">
<table id="F958C40BFFB60066458552F66F898BC0" box="[151,742,1446,1711]" gridcols="7" gridrows="6" pageId="6" pageNumber="241">
<tr id="356834E9FFB60066458552F66F8988D3" box="[151,742,1446,1468]" gridrow="0" pageId="6" pageNumber="241" rowspan-0="1">
<th id="76B95D95FFB6006645FD52F66C7688D3" box="[239,281,1446,1468]" gridcol="1" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F45E552F76C7F88D3" bold="true" box="[247,272,1447,1468]" pageId="6" pageNumber="241">Fe</emphasis>
</th>
<th id="76B95D95FFB60066445452F66C1F88D3" box="[326,368,1446,1468]" gridcol="2" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F445C52F76C0688D3" bold="true" box="[334,361,1447,1468]" pageId="6" pageNumber="241">Pa</emphasis>
</th>
<th id="76B95D95FFB60066448C52F66CA788D3" box="[414,456,1446,1468]" gridcol="3" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F44BA52F66CD088D3" bold="true" box="[424,447,1446,1468]" pageId="6" pageNumber="241">Ti</emphasis>
</th>
<th id="76B95D95FFB6006644E452F66F4F88D3" box="[502,544,1446,1468]" gridcol="4" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F44E952F76F7588D3" bold="true" box="[507,538,1447,1468]" pageId="6" pageNumber="241">Mt</emphasis>
</th>
<th id="76B95D95FFB60066475F52F66F1888D3" box="[589,631,1446,1468]" gridcol="5" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F474652F76F1F88D3" bold="true" box="[596,624,1447,1468]" pageId="6" pageNumber="241">Ta</emphasis>
</th>
<th id="76B95D95FFB6006647BD52F66F8988D3" box="[687,742,1446,1468]" gridcol="6" gridrow="0" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F47BD52F76F8988D3" bold="true" box="[687,742,1447,1468]" pageId="6" pageNumber="241">Total</emphasis>
</th>
</tr>
<tr id="356834E9FFB600664585528D6F89889C" box="[151,742,1501,1523]" gridrow="1" pageId="6" pageNumber="241">
<th id="76B95D95FFB600664585528D6DAD889C" box="[151,194,1501,1523]" gridcol="0" gridrow="1" pageId="6" pageNumber="241">Palp</th>
<td id="76B95D95FFB6006645FD528D6C76889C" box="[239,281,1501,1523]" gridcol="1" gridrow="1" pageId="6" pageNumber="241">1.18</td>
<td id="76B95D95FFB600664454528D6C1F889C" box="[326,368,1501,1523]" gridcol="2" gridrow="1" pageId="6" pageNumber="241">0.58</td>
<td id="76B95D95FFB60066448C528D6CA7889C" box="[414,456,1501,1523]" gridcol="3" gridrow="1" pageId="6" pageNumber="241">0.82</td>
<td id="76B95D95FFB6006644E4528D6F4F889C" box="[502,544,1501,1523]" gridcol="4" gridrow="1" pageId="6" pageNumber="241"></td>
<td id="76B95D95FFB60066475F528D6F18889C" box="[589,631,1501,1523]" gridcol="5" gridrow="1" pageId="6" pageNumber="241">1.18</td>
<td id="76B95D95FFB6006647BD528D6F89889C" box="[687,742,1501,1523]" gridcol="6" gridrow="1" pageId="6" pageNumber="241">3.76</td>
</tr>
<tr id="356834E9FFB600664585515C6F898B4D" box="[151,742,1548,1570]" gridrow="2" pageId="6" pageNumber="241">
<th id="76B95D95FFB600664585515C6DAD8B4D" box="[151,194,1548,1570]" gridcol="0" gridrow="2" pageId="6" pageNumber="241">I</th>
<td id="76B95D95FFB6006645FD515C6C768B4D" box="[239,281,1548,1570]" gridcol="1" gridrow="2" pageId="6" pageNumber="241">2.53</td>
<td id="76B95D95FFB600664454515C6C1F8B4D" box="[326,368,1548,1570]" gridcol="2" gridrow="2" pageId="6" pageNumber="241">1.17</td>
<td id="76B95D95FFB60066448C515C6CA78B4D" box="[414,456,1548,1570]" gridcol="3" gridrow="2" pageId="6" pageNumber="241">2.21</td>
<td id="76B95D95FFB6006644E4515C6F4F8B4D" box="[502,544,1548,1570]" gridcol="4" gridrow="2" pageId="6" pageNumber="241">1.83</td>
<td id="76B95D95FFB60066475F515C6F188B4D" box="[589,631,1548,1570]" gridcol="5" gridrow="2" pageId="6" pageNumber="241">1.37</td>
<td id="76B95D95FFB6006647BD515C6F898B4D" box="[687,742,1548,1570]" gridcol="6" gridrow="2" pageId="6" pageNumber="241">9.11</td>
</tr>
<tr id="356834E9FFB600664585516B6F898B3E" box="[151,742,1595,1617]" gridrow="3" pageId="6" pageNumber="241">
<th id="76B95D95FFB600664585516B6DAD8B3E" box="[151,194,1595,1617]" gridcol="0" gridrow="3" pageId="6" pageNumber="241">II</th>
<td id="76B95D95FFB6006645FD516B6C768B3E" box="[239,281,1595,1617]" gridcol="1" gridrow="3" pageId="6" pageNumber="241">2.56</td>
<td id="76B95D95FFB600664454516B6C1F8B3E" box="[326,368,1595,1617]" gridcol="2" gridrow="3" pageId="6" pageNumber="241">1.08</td>
<td id="76B95D95FFB60066448C516B6CA78B3E" box="[414,456,1595,1617]" gridcol="3" gridrow="3" pageId="6" pageNumber="241">2.03</td>
<td id="76B95D95FFB6006644E4516B6F4F8B3E" box="[502,544,1595,1617]" gridcol="4" gridrow="3" pageId="6" pageNumber="241">1.81</td>
<td id="76B95D95FFB60066475F516B6F188B3E" box="[589,631,1595,1617]" gridcol="5" gridrow="3" pageId="6" pageNumber="241">1.31</td>
<td id="76B95D95FFB6006647BD516B6F898B3E" box="[687,742,1595,1617]" gridcol="6" gridrow="3" pageId="6" pageNumber="241">8.79</td>
</tr>
<tr id="356834E9FFB600664585513A6F898BEF" box="[151,742,1642,1664]" gridrow="4" pageId="6" pageNumber="241">
<th id="76B95D95FFB600664585513A6DAD8BEF" box="[151,194,1642,1664]" gridcol="0" gridrow="4" pageId="6" pageNumber="241">III</th>
<td id="76B95D95FFB6006645FD513A6C768BEF" box="[239,281,1642,1664]" gridcol="1" gridrow="4" pageId="6" pageNumber="241">2.30</td>
<td id="76B95D95FFB600664454513A6C1F8BEF" box="[326,368,1642,1664]" gridcol="2" gridrow="4" pageId="6" pageNumber="241">0.96</td>
<td id="76B95D95FFB60066448C513A6CA78BEF" box="[414,456,1642,1664]" gridcol="3" gridrow="4" pageId="6" pageNumber="241">1.94</td>
<td id="76B95D95FFB6006644E4513A6F4F8BEF" box="[502,544,1642,1664]" gridcol="4" gridrow="4" pageId="6" pageNumber="241">2.10</td>
<td id="76B95D95FFB60066475F513A6F188BEF" box="[589,631,1642,1664]" gridcol="5" gridrow="4" pageId="6" pageNumber="241">1.25</td>
<td id="76B95D95FFB6006647BD513A6F898BEF" box="[687,742,1642,1664]" gridcol="6" gridrow="4" pageId="6" pageNumber="241">8.55</td>
</tr>
<tr id="356834E9FFB60066458551C96F898BC0" box="[151,742,1689,1711]" gridrow="5" pageId="6" pageNumber="241">
<th id="76B95D95FFB60066458551C96DAD8BC0" box="[151,194,1689,1711]" gridcol="0" gridrow="5" pageId="6" pageNumber="241">IV</th>
<td id="76B95D95FFB6006645FD51C96C768BC0" box="[239,281,1689,1711]" gridcol="1" gridrow="5" pageId="6" pageNumber="241">3.08</td>
<td id="76B95D95FFB60066445451C96C1F8BC0" box="[326,368,1689,1711]" gridcol="2" gridrow="5" pageId="6" pageNumber="241">1.09</td>
<td id="76B95D95FFB60066448C51C96CA78BC0" box="[414,456,1689,1711]" gridcol="3" gridrow="5" pageId="6" pageNumber="241">2.74</td>
<td id="76B95D95FFB6006644E451C96F4F8BC0" box="[502,544,1689,1711]" gridcol="4" gridrow="5" pageId="6" pageNumber="241">3.64</td>
<td id="76B95D95FFB60066475F51C96F188BC0" box="[589,631,1689,1711]" gridcol="5" gridrow="5" pageId="6" pageNumber="241">1.79</td>
<td id="76B95D95FFB6006647BD51C96F898BC0" box="[687,742,1689,1711]" gridcol="6" gridrow="5" pageId="6" pageNumber="241">12.34</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFB6FF9F4651529269A98BB5" blockId="6.[835,1474,1474,1754]" pageId="6" pageNumber="241">
Epigyne as shown in
<figureCitation id="13632A2EFFB6FF9F4121529269D888B4" box="[1075,1207,1474,1499]" captionStart="Figs" captionStartId="6.[135,179,1240,1262]" captionTargetBox="[135,1473,189,1225]" captionTargetId="figure-7@6.[135,1473,189,1225]" captionTargetPageId="6" captionText="Figs. 1921. Epigyne of Pardosa zhishengi, new species. 19, 20 ventral; 21 dorsal. Scale: 0.1 mm. Abbreviations: CD copulatory duct, Ho hood, Re receptacle, SB septal base, SS septal stem, SW septal “wings”." figureDoi="http://doi.org/10.5281/zenodo.14683146" httpUri="https://zenodo.org/record/14683146/files/figure.png" pageId="6" pageNumber="241">Figs. 1921</figureCitation>
. Septum anchor-shaped, septal base (SB) with kind of semi-transparent “wings” (SW) on its anterior edge, long stem (SS) narrowing posteriorly and widening anteriorly. Hoods (Ho) triangular, widely spaced (distance between hoods 1.4 times of hood length). Receptacles (Re) large, kidney-shaped with strong constriction in their middle part, spaced by a distance ca. 2.5 times their width. Copulatory ducts (CD) shorter than receptacles length, slightly curved.
</paragraph>
</subSubSection>
<caption id="DF276623FFB6FF9F459A51AD6DBB8A5F" pageId="6" pageNumber="241" startId="6.[136,191,1789,1811]" targetType="table">
<paragraph id="8BE736ABFFB6FF9F459A51AD6DBB8A5F" blockId="6.[136,775,1789,1840]" pageId="6" pageNumber="241">
Table 6. Leg spination for the female of
<taxonomicName id="4C584D28FFB6FF9F470F51AD6FBE8A7D" authorityName="Omelko" authorityYear="2024" box="[541,721,1789,1810]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFB6FF9F470F51AD6FBE8A7D" box="[541,721,1789,1810]" italics="true" pageId="6" pageNumber="241">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFB6FF9F47CD51AE6DBF8A5F" pageId="6" pageNumber="241" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<subSubSection id="C3426520FFB6FF9F465150526EB38A55" pageId="6" pageNumber="241" type="distribution">
<paragraph id="8BE736ABFFB6FF9F465150526EB38A55" blockId="6.[835,1474,1794,1850]" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F465150526EA08A75" bold="true" box="[835,975,1794,1818]" pageId="6" pageNumber="241">Distribution</emphasis>
. Only known from the
<typeStatus id="54E38809FFB6FF9F41CE505368658A74" box="[1244,1290,1795,1819]" pageId="6" pageNumber="241">type</typeStatus>
locality in
<collectingCountry id="F34F763BFFB6FF9F409C505368AD8A75" box="[1422,1474,1795,1818]" name="Laos" pageId="6" pageNumber="241">Laos</collectingCountry>
(
<figureCitation id="13632A2EFFB6FF9F465950726EBF8A55" box="[843,976,1826,1850]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="6" pageNumber="241">Figs. 5253</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,105 @@
<document id="60054364128717410E4D019079D9F913" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB6FF9E46FA50326CD38543" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Trochosa C. L. Koch 1847" docType="treatment" docVersion="2" lastPageNumber="241" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="241" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="506970E2189C51FBBBF52D93500D71E3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C0BCA637E16E233E02CD3A059A67B37">
<mods:title id="311A58B060ECF3CE61B4AC3380D57773">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="B24E1473C9D195BD1E35744B44763487" type="personal">
<mods:role id="4449ADC3921943DC77DDEE0A1788BC2B">
<mods:roleTerm id="45E6E438AB69E4D32F026B95AC343EF9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C85CA580941A76C43F1B385788E9F05A">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2B2526C184586DE439D5D6FDDB3CB572">text</mods:typeOfResource>
<mods:relatedItem id="F70EB3468969583C8612B4E27996DD8C" type="host">
<mods:titleInfo id="92DFF93355A9B23EFC17A5B769D9B69F">
<mods:title id="7378334E0516F7EE93393493107CB501">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1867AD157794E6B338C999D160E31151">
<mods:date id="7D3FC4A82B2B94287320EF5704D7342F">2024</mods:date>
<mods:detail id="A73EBC44B1F1767E26E90F2E766E035C" type="pubDate">
<mods:number id="9E05635DF86ABB3BB033628074CD2842">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="6D79DD87927C517283CCC9751188B404" type="volume">
<mods:number id="C60593BE2E5ADCB5B8EFAE78D19719A7">72</mods:number>
</mods:detail>
<mods:extent id="6FD8AACC1DED5E6C2E6849D7021252E0" unit="page">
<mods:start id="0CE32FC9A0AAC34A798414DD6E422897">235</mods:start>
<mods:end id="94948A0238E3AB04709A6DBF19A70448">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8BB6A7BCE0EAB091EDF478097E0563A6">journal article</mods:classification>
<mods:identifier id="39DCC9A065AA31895B5B5766129D044B" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="0541C6CA8590137BB1A2FCEE8F242154" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="AD0D21C6B3CD4E3BDAE4E14B7215A920" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="51604CF189601293D1533068CF7125A9" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB6FF9E46FA50326CD38543" LSID="urn:lsid:plazi:treatment:03F187BDFFB6FF9E46FA50326CD38543" httpUri="http://treatment.plazi.org/id/03F187BDFFB6FF9E46FA50326CD38543" lastPageId="7" lastPageNumber="241" pageId="6" pageNumber="241">
<subSubSection id="C3426520FFB6FF9F46FA503268728A15" box="[1000,1309,1890,1915]" pageId="6" pageNumber="241" type="nomenclature">
<paragraph id="8BE736ABFFB6FF9F46FA503268728A15" blockId="6.[1000,1309,1890,1915]" box="[1000,1309,1890,1915]" pageId="6" pageNumber="241">
<heading id="D0AF81C7FFB6FF9F46FA503268728A15" bold="true" box="[1000,1309,1890,1915]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="241" reason="2">
<taxonomicName id="4C584D28FFB6FF9F46FA503268728A15" ID-CoL="82S4" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[1000,1309,1890,1915]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB6FF9F46FA503268728A15" bold="true" box="[1000,1309,1890,1915]" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F46FA5032693F8A15" bold="true" box="[1000,1104,1890,1914]" italics="true" pageId="6" pageNumber="241">Trochosa</emphasis>
C. L.
<bibRefCitation id="EFC94B5AFFB6FF9F418E503368728A15" author="Koch CL" box="[1180,1309,1890,1915]" pageId="6" pageNumber="241" refId="ref9676" refString="Koch CL (1847) Die Arachniden. J. L. Lotzbeck, Nurnberg, Vierzehnter Band, pp. 89 - 210, pl. 481 - 504 (f. 1343 - 1412); Funfzehnter Band, pp. 1 - 136, pl. 505 - 540 (f. 1413 - 1504); Sechszehnter und letzter Band, pp. 1 - 80, pl. 541 - 563 (f. 1505 - 1550), Index 64 pp." type="book" year="1847">Koch, 1847</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB6FF9F465150F36EDB8AB5" pageId="6" pageNumber="241" type="type_taxon">
<paragraph id="8BE736ABFFB6FF9F465150F36EDB8AB5" blockId="6.[835,1474,1954,2010]" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F465150F36E8F8AD5" bold="true" box="[835,992,1954,1979]" pageId="6" pageNumber="241">
<typeStatus id="54E38809FFB6FF9F465150F36E118AD4" box="[835,894,1955,1979]" pageId="6" pageNumber="241">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C584D28FFB6FF9F46F950F3683C8AD4" authority="De Geer, 1778" authorityName="De Geer" authorityYear="1778" box="[1003,1363,1954,1979]" class="Arachnida" family="Agelenidae" genus="Aranea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="ruricola">
<emphasis id="B92CEAB9FFB6FF9F46F950F369F08AD5" box="[1003,1183,1954,1978]" italics="true" pageId="6" pageNumber="241">Aranea ruricola</emphasis>
<bibRefCitation id="EFC94B5AFFB6FF9F41BB50F3683C8AD4" author="De Geer C" box="[1193,1363,1955,1979]" pageId="6" pageNumber="241" pagination="176 - 324" refId="ref9408" refString="De Geer C (1778) Des araignees. In: Memoires pour servir a l'histoire des insectes. Tome septieme. Pierre Hesselberg, Stockholm, pp. 176 - 324." type="book chapter" year="1778">De Geer, 1778</bibRefCitation>
</taxonomicName>
, from the Palearctic.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB6FF9E46515F526CD38543" lastPageId="7" lastPageNumber="242" pageId="6" pageNumber="241" type="discussion">
<paragraph id="8BE736ABFFB6FF9E46515F526CD38543" blockId="6.[835,1475,2050,2107]" lastBlockId="7.[112,754,1652,2093]" lastPageId="7" lastPageNumber="242" pageId="6" pageNumber="241">
<emphasis id="B92CEAB9FFB6FF9F46515F526EA98575" bold="true" box="[835,966,2050,2074]" pageId="6" pageNumber="241">Comments.</emphasis>
<taxonomicName id="4C584D28FFB6FF9F46C25F5268688574" authority="C. L. Koch, 1847" authorityName="C. L. Koch" authorityYear="1847" box="[976,1287,2050,2075]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB6FF9F46C25F52695A8575" box="[976,1077,2050,2074]" italics="true" pageId="6" pageNumber="241">Trochosa</emphasis>
C. L.
<bibRefCitation id="EFC94B5AFFB6FF9F41965F5268688574" author="Koch CL" box="[1156,1287,2050,2075]" pageId="6" pageNumber="241" refId="ref9676" refString="Koch CL (1847) Die Arachniden. J. L. Lotzbeck, Nurnberg, Vierzehnter Band, pp. 89 - 210, pl. 481 - 504 (f. 1343 - 1412); Funfzehnter Band, pp. 1 - 136, pl. 505 - 540 (f. 1413 - 1504); Sechszehnter und letzter Band, pp. 1 - 80, pl. 541 - 563 (f. 1505 - 1550), Index 64 pp." type="book" year="1847">Koch, 1847</bibRefCitation>
</taxonomicName>
, with 93 species (
<bibRefCitation id="EFC94B5AFFB6FF9F46595F736EA18554" author="WSC" box="[843,974,2083,2107]" pageId="6" pageNumber="241" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC, 2024</bibRefCitation>
), is a relatively large genus within
<taxonomicName id="4C584D28FFB6FF9F405C5F7268D08555" authorityName="Sundevall" authorityYear="1833" box="[1358,1471,2082,2106]" class="Arachnida" family="Lycosidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="family">Lycosidae</taxonomicName>
. The majority of species in this genus (33) are known from the Afrotropical realm, with only 17 species inhabiting the Indomalayan realm. Indomalayan
<taxonomicName id="4C584D28FFB7FF9E44C351E46F5B8BA3" authorityName="C. L. Koch" authorityYear="1847" box="[465,564,1716,1740]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB7FF9E44C351E46F5B8BA3" box="[465,564,1716,1740]" italics="true" pageId="7" pageNumber="242">Trochosa</emphasis>
</taxonomicName>
are recorded from
<collectingCountry id="F34F763BFFB7FF9E456251846DC88B83" box="[112,167,1748,1772]" name="India" pageId="7" pageNumber="242">India</collectingCountry>
and
<collectingCountry id="F34F763BFFB7FF9E45CA51846C778B83" box="[216,280,1748,1772]" name="Nepal" pageId="7" pageNumber="242">Nepal</collectingCountry>
(6 species combined),
<collectingCountry id="F34F763BFFB7FF9E471B51846F1E8B83" box="[521,625,1748,1772]" name="Indonesia" pageId="7" pageNumber="242">Indonesia</collectingCountry>
(5 species), southern
<collectingCountry id="F34F763BFFB7FF9E45C651A46C7B8A63" box="[212,276,1780,1804]" name="China" pageId="7" pageNumber="242">China</collectingCountry>
(3 species), the
<collectingCountry id="F34F763BFFB7FF9E44D451A46F518A63" box="[454,574,1780,1804]" name="Philippines" pageId="7" pageNumber="242">Philippines</collectingCountry>
(2 species), and 1 species,
<taxonomicName id="4C584D28FFB7FF9E45F750446FE68A43" authority="Schenkel, 1963" authorityName="Schenkel" authorityYear="1963" box="[229,649,1812,1836]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB7FF9E45F750446CB98A43" box="[229,470,1812,1836]" italics="true" pageId="7" pageNumber="242">Trochosa ruricoloides</emphasis>
<bibRefCitation id="EFC94B5AFFB7FF9E44CD50446FE68A43" author="Schenkel E" box="[479,649,1812,1836]" pageId="7" pageNumber="242" pagination="1 - 481" refId="ref10677" refString="Schenkel E (1963) Ostasiatische Spinnen aus dem Museum d'Histoire naturelle de Paris. Memoires du Museum National d'Histoire Naturelle de Paris (A, Zool.), 25: 1 - 481." type="journal article" year="1963">Schenkel, 1963</bibRefCitation>
</taxonomicName>
, is found throughout the entire Indomalayan realm. Until now, no species of this genus have been reported from
<collectingCountry id="F34F763BFFB7FF9E479350056FD68A03" box="[641,697,1877,1900]" name="Laos" pageId="7" pageNumber="242">Laos</collectingCountry>
. The Indomalayan
<taxonomicName id="4C584D28FFB7FF9E441450246C048AE3" authorityName="C. L. Koch" authorityYear="1847" box="[262,363,1908,1932]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB7FF9E441450246C048AE3" box="[262,363,1908,1932]" italics="true" pageId="7" pageNumber="242">Trochosa</emphasis>
</taxonomicName>
remain poorly studied, with nearly half of the species (8) known only from one sex, primarily females. This may lead to identification challenges, as females in this genus are frequently almost indistinguishable. Almost all species from this region (14 out of 17) have been illustrated in 12 publications.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,166 @@
<document id="5399D7E3CAA125CC70399F933DAACFE8" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB7FF91462A51246C028A04" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Trochosa bannaensis Yin &amp; Chen 1995" docType="treatment" docVersion="2" lastPageNumber="243" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="242" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="67FE44EB6140C6F7445F738FC3E89F9C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="294D839EE5016BFCEF31F409FDED73C4">
<mods:title id="CA504BBC181CA0FB3C5B515461FA27B5">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="CBBBAE63E39CA9E00948BC617E266E69" type="personal">
<mods:role id="7781053A679C9BD7D1A531ADCD406ED1">
<mods:roleTerm id="D423DAE1BA464CB00E3D4505ECA14DCE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED7F1C69970F62C45244380EB3FFC796">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="B7D759CC1933D02AAE8F3FA6DDE5BE5E">text</mods:typeOfResource>
<mods:relatedItem id="E13ED9B65D67CC4CA5537E8A8464B08D" type="host">
<mods:titleInfo id="9C8BCB149033EF35E460E5E3B3135CEC">
<mods:title id="D080ED2A7F816A270EFDDE8EB4887774">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1CD69D443F4279421BCE58598ECF35B8">
<mods:date id="D64C572DD24112632E709E7E578BA621">2024</mods:date>
<mods:detail id="71EA5C00117685735FBB1B10A5CA3963" type="pubDate">
<mods:number id="D72361033D863ACC9AC005386CBEA1E2">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="B77EDEAABF33FC81900FD055F7D630FC" type="volume">
<mods:number id="59E6D5DC9CEB3F95C3C2147F95ED2663">72</mods:number>
</mods:detail>
<mods:extent id="3EE9A82A14BEB275E66A92EFBE95FA02" unit="page">
<mods:start id="A267ED8E9B62EE4EFBD8EB4321AB8516">235</mods:start>
<mods:end id="267C08D93A1C890E4E4D90BC1EB61372">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E2BD99F5E6A0EE16CFB290CF96C4D222">journal article</mods:classification>
<mods:identifier id="A9FED393F94E540255B4C6B5E8CAC4D7" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="983F27F5575EDE10110729DD2DB567D5" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E171B6E83FD877F1BB24FF5376AC6859" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="693C651FEB6E4326393AD892E5416998" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB7FF91462A51246C028A04" LSID="urn:lsid:plazi:treatment:03F187BDFFB7FF91462A51246C028A04" httpUri="http://treatment.plazi.org/id/03F187BDFFB7FF91462A51246C028A04" lastPageId="8" lastPageNumber="243" pageId="7" pageNumber="242">
<subSubSection id="C3426520FFB7FF9E462A512469E78BC3" pageId="7" pageNumber="242" type="nomenclature">
<paragraph id="8BE736ABFFB7FF9E462A512469E78BC3" blockId="7.[824,1442,1652,1740]" pageId="7" pageNumber="242">
<heading id="D0AF81C7FFB7FF9E462A512469E78BC3" bold="true" centered="true" fontSize="10" level="2" pageId="7" pageNumber="242" reason="2">
<emphasis id="B92CEAB9FFB7FF9E462A512469E78BC3" bold="true" pageId="7" pageNumber="242">
<taxonomicName id="4C584D28FFB7FF9E462A5124686B8BE3" ID-CoL="597M4" authority="Yin &amp; Chen, 1995" authorityName="Yin &amp; Chen" authorityYear="1995" box="[824,1284,1652,1676]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="bannaensis">
<emphasis id="B92CEAB9FFB7FF9E462A5124694B8BE3" bold="true" box="[824,1060,1652,1676]" italics="true" pageId="7" pageNumber="242">Trochosa bannaensis</emphasis>
Yin &amp; Chen, 1995
</taxonomicName>
, in
<bibRefCitation id="EFC94B5AFFB7FF9E4022512469E78BC3" author="Yin CM &amp; Bao YH &amp; Wang JF" pageId="7" pageNumber="242" pagination="23 - 36" refId="ref11348" refString="Yin CM, Bao YH &amp; Wang JF (1995) An advanced study on the genus Trochosa from China (Araneae: Lycosidae). Acta Arachnologica Sinica, 4: 23 - 36." type="journal article" year="1995">Yin et al., 1995</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB7FF9E46E851E469B28BA3" box="[1018,1245,1716,1740]" pageId="7" pageNumber="242" type="description">
<paragraph id="8BE736ABFFB7FF9E46E851E469B28BA3" blockId="7.[824,1442,1652,1740]" box="[1018,1245,1716,1740]" pageId="7" pageNumber="242">
(
<figureCitation id="13632A2EFFB7FF9E411051E469EA8BA3" box="[1026,1157,1716,1740]" captionStart="Figs" captionStartId="7.[113,157,1547,1569]" captionTargetBox="[121,1440,204,1525]" captionTargetId="figure-10@7.[113,1451,195,1525]" captionTargetPageId="7" captionText="Figs. 2226. Male (22), female (23), male palp (2425) and epigyne (26) of Trochosa bannaensis. 22, 23 dorsal; 24, 26 ventral; 25 retrolateral. Scale: 1 mm (2223), 0.2 mm (2426)." figureDoi="http://doi.org/10.5281/zenodo.14683150" httpUri="https://zenodo.org/record/14683150/files/figure.png" pageId="7" pageNumber="242">Figs. 2226</figureCitation>
,
<figureCitation id="13632A2EFFB7FF9E418051E469B98BA3" box="[1170,1238,1716,1740]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="7" pageNumber="242">5253</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB7FF9E463F51A369818A14" pageId="7" pageNumber="242" type="reference_group">
<paragraph id="8BE736ABFFB7FF9E463F51A36EF48A4A" blockId="7.[813,1452,1779,1916]" pageId="7" pageNumber="242">
<treatmentCitationGroup id="AB481185FFB7FF9E463F51A36EF48A4A" pageId="7" pageNumber="242">
<taxonomicName id="4C584D28FFB7FF9E463F51A36EDB8A67" ID-CoL="597M4" authorityName="Yin &amp; Chen" authorityYear="1995" box="[813,948,1779,1800]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="bannaensis">
<emphasis id="B92CEAB9FFB7FF9E463F51A36EDB8A67" box="[813,948,1779,1800]" italics="true" pageId="7" pageNumber="242">T. bannaensis</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AF910BAFFB7FF9E46D051A369FE8A67" author="Pan F &amp; Zheng G &amp; Li SQ" box="[962,1169,1779,1801]" page="410" pageId="7" pageNumber="242" year="2016">
<bibRefCitation id="EFC94B5AFFB7FF9E46D051A369FE8A67" author="Pan F &amp; Zheng G &amp; Li SQ" box="[962,1169,1779,1801]" pageId="7" pageNumber="242" pagination="404 - 414" refId="ref10290" refString="Pan F, Zheng G &amp; Li SQ (2016) Wolf spiders (Araneae: Lycosidae) from Xishuangbanna Tropical Botanical Garden, China. Zoological Systematics, 41 (4): 404 - 414." type="journal article" year="2016">Pan et al., 2016: 410</bibRefCitation>
</treatmentCitation>
, fig. 4AE, 5AD, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE736ABFFB7FF9E463F507D6E938A30" blockId="7.[813,1452,1779,1916]" pageId="7" pageNumber="242">
<treatmentCitationGroup id="AB481185FFB7FF9E463F507D6E938A30" pageId="7" pageNumber="242">
<taxonomicName id="4C584D28FFB7FF9E463F507D6EDD8A2E" ID-CoL="597M4" authorityName="Yin &amp; Chen" authorityYear="1995" box="[813,946,1836,1858]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="bannaensis">
<emphasis id="B92CEAB9FFB7FF9E463F507D6EDD8A2E" box="[813,946,1836,1858]" italics="true" pageId="7" pageNumber="242">T. bannaensis</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AF910BAFFB7FF9E46AC507D69E28A2D" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[958,1165,1836,1858]" page="57" pageId="7" pageNumber="242" year="2021">
<bibRefCitation id="EFC94B5AFFB7FF9E46AC507D69E28A2D" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[958,1165,1836,1858]" pageId="7" pageNumber="242" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al., 2021: 57</bibRefCitation>
</treatmentCitation>
, fig. 58AH, 59AC, 60AD, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE736ABFFB7FF9E463F503669818A14" blockId="7.[813,1452,1779,1916]" box="[813,1262,1894,1916]" pageId="7" pageNumber="242">
See the full list of references in
<bibRefCitation id="EFC94B5AFFB7FF9E4179503669858A14" author="WSC" box="[1131,1258,1894,1915]" pageId="7" pageNumber="242" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB7FF91463F50F46F838A63" lastPageId="8" lastPageNumber="243" pageId="7" pageNumber="242" type="materials_examined">
<paragraph id="8BE736ABFFB7FF9E463F50F468C38573" blockId="7.[812,1452,1956,2076]" pageId="7" pageNumber="242">
<emphasis id="B92CEAB9FFB7FF9E463F50F4697F8AD3" bold="true" box="[813,1040,1956,1980]" pageId="7" pageNumber="242">Material examined.</emphasis>
<materialsCitation id="3B303CF6FFB7FF9E410850F468C38573" collectionCode="FEFU" collectorName="M. M. Omelko &amp; Omelko, A. A. &amp; Komisarenko &amp; May" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="7" pageNumber="242" specimenCount="53" specimenCount-female="1" specimenCount-male="52" stateProvince="Viangchan">
<specimenCount id="9D5EFD22FFB7FF9E410850F469EF8AD3" box="[1050,1152,1956,1980]" count="52" pageId="7" pageNumber="242" type="male">52 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB7FF9E419D50F4699F8AD3" box="[1167,1264,1956,1980]" count="1" pageId="7" pageNumber="242" type="female">1 female</specimenCount>
(
<collectionCode id="ED49AE6EFFB7FF9E401350F468278AD3" box="[1281,1352,1956,1980]" pageId="7" pageNumber="242">FEFU</collectionCode>
),
<collectingCountry id="F34F763BFFB7FF9E404C50F468C88AD3" box="[1374,1447,1956,1980]" name="Laos" pageId="7" pageNumber="242">LAOS</collectingCountry>
,
<collectingRegion id="499CF849FFB7FF9E463F50946EBE8AB3" box="[813,977,1988,2012]" country="Laos" name="Viangchan" pageId="7" pageNumber="242">Vientiane Prov.</collectingRegion>
, env. of
<location id="8E876070FFB7FF9E4131509468688AB3" LSID="urn:lsid:plazi:treatment:03F187BDFFB7FF91462A51246C028A04:8E876070FFB7FF9E4131509468688AB3" box="[1059,1287,1988,2012]" country="Laos" latitude="18.614773" longLatPrecision="1" longitude="102.40885" name="Nam-Lik Eco-Village" pageId="7" pageNumber="242" stateProvince="Viangchan">Nam-Lik Eco-Village</location>
,
<geoCoordinate id="EE6C506CFFB7FF9E4002509368C38AB3" box="[1296,1452,1987,2012]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="7" pageNumber="242" precision="1" seconds="53.18" value="18.614773">18°3653.18″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB7FF9E463E50B36EB88A93" box="[812,983,2019,2044]" degrees="102" direction="east" minutes="24" orientation="longitude" pageId="7" pageNumber="242" precision="1" seconds="31.87" value="102.40885">102°2431.87″E</geoCoordinate>
,
<collectingMethod id="52194EBCFFB7FF9E46F250B469388A93" box="[992,1111,2020,2044]" pageId="7" pageNumber="242">pitfall traps</collectingMethod>
in secondary forest and near rice field, coll.
<collectorName id="26AD537DFFB7FF9E46B05F55692E8573" box="[930,1089,2052,2076]" pageId="7" pageNumber="242">M.M. Omelko</collectorName>
, A.A.,
<collectorName id="26AD537DFFB7FF9E41825F54684C8573" box="[1168,1315,2052,2076]" pageId="7" pageNumber="242">Komisarenko</collectorName>
, 23
<collectorName id="26AD537DFFB7FF9E40405F5468EB8573" box="[1362,1412,2052,2076]" pageId="7" pageNumber="242">May</collectorName>
12
</materialsCitation>
</paragraph>
<caption id="DF276623FFB8FF91459552FA6EC588B3" ID-DOI="http://doi.org/10.5281/zenodo.14683152" ID-Zenodo-Dep="14683152" httpUri="https://zenodo.org/record/14683152/files/figure.png" pageId="8" pageNumber="243" startId="8.[135,179,1450,1472]" targetBox="[140,1465,189,1434]" targetPageId="8" targetType="figure">
<paragraph id="8BE736ABFFB8FF91459552FA6EC588B3" blockId="8.[135,1473,1450,1500]" pageId="8" pageNumber="243">
Figs. 2731. Male (27), female (28), male palp (2930, cymbial claw is broken off) and epigyne (31) of
<taxonomicName id="4C584D28FFB8FF91419952FA680C88D0" authorityName="Schenkel" authorityYear="1963" box="[1163,1379,1450,1471]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB8FF91419952FA680C88D0" box="[1163,1379,1450,1471]" italics="true" pageId="8" pageNumber="243">Trochosa ruricoloides</emphasis>
</taxonomicName>
. 27, 28 dorsal; 29, 31 ventral; 30 retrolateral. Scale: 1 mm (2728), 0.2 mm (2931).
</paragraph>
</caption>
<paragraph id="8BE736ABFFB8FF91459A51446C8C8B23" blockId="8.[136,775,1556,1612]" pageId="8" pageNumber="243">
June 2016;
<materialsCitation id="3B303CF6FFB8FF91441151446CB08B23" collectingDate="2017-06-04" collectingDateMax="2017-06-26" collectingDateMin="2017-06-04" collectionCode="FEFU" pageId="8" pageNumber="243" specimenCount="11" specimenCount-female="7" specimenCount-male="4">
<specimenCount id="9D5EFD22FFB8FF91441151446C378B43" box="[259,344,1556,1580]" count="4" pageId="8" pageNumber="243" type="male">4 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB8FF91447151446CA58B43" box="[355,458,1556,1580]" count="7" pageId="8" pageNumber="243" type="female">7 females</specimenCount>
(
<collectionCode id="ED49AE6EFFB8FF9144CA51446F708B43" box="[472,543,1556,1580]" pageId="8" pageNumber="243">FEFU</collectionCode>
), same place, habitats and collectors,
<date id="FFE6106BFFB8FF91443C51646CB08B23" box="[302,479,1588,1612]" pageId="8" pageNumber="243" value="2017-06-04" valueMax="2017-06-26" valueMin="2017-06-04">
<collectingDate id="EFA2E983FFB8FF91443C51646CB08B23" box="[302,479,1588,1612]" pageId="8" pageNumber="243" value="2017-06-04" valueMax="2017-06-26" valueMin="2017-06-04">426 June 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE736ABFFB8FF91459A51246F838A63" blockId="8.[136,776,1652,1804]" pageId="8" pageNumber="243">
<emphasis id="B92CEAB9FFB8FF91459A51246DA08BE3" bold="true" box="[136,207,1652,1676]" pageId="8" pageNumber="243">Notes.</emphasis>
I do not provide a redescription here because this species was well illustrated recently (see references above). It is important to note that the shape of the tegular apophysis slightly varies from male to male—such differences may lead to mistaken inferences of new undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB8FF91459850636C028A04" pageId="8" pageNumber="243" type="distribution">
<paragraph id="8BE736ABFFB8FF91459850636C028A04" blockId="8.[137,773,1843,1900]" pageId="8" pageNumber="243">
<emphasis id="B92CEAB9FFB8FF91459850636C498A24" bold="true" box="[138,294,1843,1867]" pageId="8" pageNumber="243">Distribution.</emphasis>
<collectingCountry id="F34F763BFFB8FF91443D50646C1D8A23" box="[303,370,1844,1868]" name="China" pageId="8" pageNumber="243">China</collectingCountry>
(
<collectingRegion id="499CF849FFB8FF91449450646CB38A23" box="[390,476,1844,1868]" country="China" name="Hainan" pageId="8" pageNumber="243">Hainan</collectingRegion>
,
<collectingRegion id="499CF849FFB8FF9144FE50646F288A23" box="[492,583,1844,1868]" country="China" name="Yunnan" pageId="8" pageNumber="243">Yunnan</collectingRegion>
) and
<collectingCountry id="F34F763BFFB8FF91479F50646FAC8A24" box="[653,707,1844,1867]" name="Laos" pageId="8" pageNumber="243">Laos</collectingCountry>
(new record;
<figureCitation id="13632A2EFFB8FF9145CF50046C0E8A04" box="[221,353,1875,1900]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="8" pageNumber="243">Figs. 5253</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,226 @@
<document id="259BDE0E24775BA0FDF3A81965AF07D0" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB8FF9045E650C36FF88C9A" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Trochosa ruricoloides Schenkel 1963" docType="treatment" docVersion="2" lastPageNumber="244" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="243" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BCF8B54A2225BBDA78693BF2D18B1579" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="40E8ECFE8B4B031077385D6466405C63">
<mods:title id="BF472AC59A31A8CFAE877C6367C5FA25">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="4BE13F88EAE44BEBCFBB7D882F497EEA" type="personal">
<mods:role id="091BD81B98278965551FA8D32FF4C8A6">
<mods:roleTerm id="175768941D8309D836E0FE05F52721D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD10DC98288C0E74D2079AE28BFC2BF3">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="910CDD24ECAD16E0F5584A1186F13208">text</mods:typeOfResource>
<mods:relatedItem id="0F164A91DE2D5195DB48C367F008DC1E" type="host">
<mods:titleInfo id="C723DFDCF226687ED369A0D40AD0E07C">
<mods:title id="05D9D263A2AAE3F781008D40BD0D3BD7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="ECC9A7B5FF4ECE76DFA8BD0BD1FCADFA">
<mods:date id="2FE6691503D77B75B2335205116FB485">2024</mods:date>
<mods:detail id="1509D2D247C8546E9B29E70B513B01C9" type="pubDate">
<mods:number id="417B1B9DDAF456F6E199219F3CAD79DA">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="98E37674CC7B7F51D022D22B8B051466" type="volume">
<mods:number id="D0137FCC6AB0B30BAF5ADDC4C32D9DEA">72</mods:number>
</mods:detail>
<mods:extent id="5D885D1AC3E4DF6D9805E694782D3742" unit="page">
<mods:start id="209E73DB510BE7D6BCBE6A541F0E9ECB">235</mods:start>
<mods:end id="6FD043C0FCBCB05C5604D86B3C29C27D">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3D771D9DEA0409310ED1F39FAB1ED6B7">journal article</mods:classification>
<mods:identifier id="A9EB274177F67FAFAB1BAD6E31E931AE" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="A9025A56001D2219A5018B8864D577AF" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="5838F2E3B70A135AA87891E81E4038A8" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="420A1031E7B9646228A89E957270FBA8" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB8FF9045E650C36FF88C9A" LSID="urn:lsid:plazi:treatment:03F187BDFFB8FF9045E650C36FF88C9A" httpUri="http://treatment.plazi.org/id/03F187BDFFB8FF9045E650C36FF88C9A" lastPageId="9" lastPageNumber="244" pageId="8" pageNumber="243">
<subSubSection id="C3426520FFB8FF9145E650C36FF48AC4" box="[244,667,1939,1963]" pageId="8" pageNumber="243" type="nomenclature">
<paragraph id="8BE736ABFFB8FF9145E650C36FF48AC4" blockId="8.[244,667,1939,1996]" box="[244,667,1939,1963]" pageId="8" pageNumber="243">
<heading id="D0AF81C7FFB8FF9145E650C36FF48AC4" bold="true" box="[244,667,1939,1963]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="243" reason="2">
<taxonomicName id="4C584D28FFB8FF9145E650C36FF48AC4" ID-CoL="597P8" authority="Schenkel, 1963" authorityName="Schenkel" authorityYear="1963" box="[244,667,1939,1963]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB8FF9145E650C36FF48AC4" bold="true" box="[244,667,1939,1963]" pageId="8" pageNumber="243">
<emphasis id="B92CEAB9FFB8FF9145E650C36C878AC4" bold="true" box="[244,488,1939,1963]" italics="true" pageId="8" pageNumber="243">Trochosa ruricoloides</emphasis>
<bibRefCitation id="EFC94B5AFFB8FF9144E250C36FF48AC4" author="Schenkel E" box="[496,667,1939,1963]" pageId="8" pageNumber="243" pagination="1 - 481" refId="ref10677" refString="Schenkel E (1963) Ostasiatische Spinnen aus dem Museum d'Histoire naturelle de Paris. Memoires du Museum National d'Histoire Naturelle de Paris (A, Zool.), 25: 1 - 481." type="journal article" year="1963">Schenkel, 1963</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB8FF91444450E46F568AA4" box="[342,569,1971,1996]" pageId="8" pageNumber="243" type="description">
<paragraph id="8BE736ABFFB8FF91444450E46F568AA4" blockId="8.[244,667,1939,1996]" box="[342,569,1971,1996]" pageId="8" pageNumber="243">
(
<figureCitation id="13632A2EFFB8FF91444F50E46C8E8AA3" box="[349,481,1972,1996]" captionStart="Figs" captionStartId="8.[135,179,1450,1472]" captionTargetBox="[140,1465,189,1434]" captionTargetId="figure-7@8.[135,1473,189,1434]" captionTargetPageId="8" captionText="Figs. 2731. Male (27), female (28), male palp (2930, cymbial claw is broken off) and epigyne (31) of Trochosa ruricoloides. 27, 28 dorsal; 29, 31 ventral; 30 retrolateral. Scale: 1 mm (2728), 0.2 mm (2931)." figureDoi="http://doi.org/10.5281/zenodo.14683152" httpUri="https://zenodo.org/record/14683152/files/figure.png" pageId="8" pageNumber="243">Figs. 2731</figureCitation>
,
<figureCitation id="13632A2EFFB8FF9144FC50E36F5D8AA4" box="[494,562,1971,1995]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="8" pageNumber="243">5253</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB8FF91459A50A3686A8B0D" pageId="8" pageNumber="243" type="reference_group">
<paragraph id="8BE736ABFFB8FF91459A50A36C37854A" blockId="8.[136,774,2034,2085]" pageId="8" pageNumber="243">
<treatmentCitationGroup id="AB481185FFB8FF91459A50A36C37854A" pageId="8" pageNumber="243">
<taxonomicName id="4C584D28FFB8FF91459A50A36C718568" ID-CoL="597P8" authorityName="Schenkel" authorityYear="1963" box="[136,286,2034,2056]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB8FF91459A50A36C718568" box="[136,286,2034,2056]" italics="true" pageId="8" pageNumber="243">T. ruricoloides</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AF910BAFFB8FF91443D50A26F598567" author="Marusik YM &amp; Nadolny AA &amp; Koponen S" box="[303,566,2034,2056]" page="485" pageId="8" pageNumber="243" year="2020">
<bibRefCitation id="EFC94B5AFFB8FF91443D50A26F598567" author="Marusik YM &amp; Nadolny AA &amp; Koponen S" box="[303,566,2034,2056]" pageId="8" pageNumber="243" pagination="482 - 489" refId="ref10153" refString="Marusik YM, Nadolny AA &amp; Koponen S (2020) Redescription of Trochosa urbana (Araneae: Lycosidae) with notes on its distribution. Arachnology, 18 (5): 482 - 489." type="journal article" year="2020">Marusik et al., 2020: 485</bibRefCitation>
</treatmentCitation>
, fig. 1de, 2ef, 4d, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE736ABFFB8FF9146515144697D8B29" blockId="8.[835,1474,1555,1635]" pageId="8" pageNumber="243">
<treatmentCitationGroup id="AB481185FFB8FF9146515144697D8B29" pageId="8" pageNumber="243">
<taxonomicName id="4C584D28FFB8FF91465151446EBF8B46" ID-CoL="597P8" authorityName="Schenkel" authorityYear="1963" box="[835,976,1556,1577]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB8FF91465151446EBF8B46" box="[835,976,1556,1577]" italics="true" pageId="8" pageNumber="243">T. ruricoloides</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AF910BAFFB8FF9146CE514469C78B45" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[988,1192,1556,1578]" page="64" pageId="8" pageNumber="243" year="2021">
<bibRefCitation id="EFC94B5AFFB8FF9146CE514469C78B45" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[988,1192,1556,1578]" pageId="8" pageNumber="243" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al., 2021: 64</bibRefCitation>
</treatmentCitation>
, fig. 63AH, 64AE, 65AD, male and female.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE736ABFFB8FF914651511D686A8B0D" blockId="8.[835,1474,1555,1635]" box="[835,1285,1613,1635]" pageId="8" pageNumber="243">
See the full list of references in
<bibRefCitation id="EFC94B5AFFB8FF914190511D686E8B0D" author="WSC" box="[1154,1281,1613,1634]" pageId="8" pageNumber="243" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB8FF90465151DB6C278C3A" lastPageId="9" lastPageNumber="244" pageId="8" pageNumber="243" type="materials_examined">
<paragraph id="8BE736ABFFB8FF91465151DB69B58ACB" blockId="8.[834,1474,1675,1956]" pageId="8" pageNumber="243">
<emphasis id="B92CEAB9FFB8FF91465151DB69748BCC" bold="true" box="[835,1051,1675,1699]" pageId="8" pageNumber="243">Material examined.</emphasis>
<materialsCitation id="3B303CF6FFB8FF91410D51DB6EF48A6C" collectingDate="2013-11-01" collectingDateMax="2013-11-06" collectingDateMin="2013-11-01" collectionCode="FEFU" collectorName="M. M. Omelko" country="Laos" latitude="15.193723" location="Tad" longLatPrecision="1" longitude="106.09925" municipality="Male" pageId="8" pageNumber="243" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Champasak">
<collectingMunicipality id="6B83ACD1FFB8FF91410D51DB693A8BCC" box="[1055,1109,1675,1699]" pageId="8" pageNumber="243">Male</collectingMunicipality>
(
<collectionCode id="ED49AE6EFFB8FF91417351DC69C98BCB" box="[1121,1190,1676,1700]" pageId="8" pageNumber="243">FEFU</collectionCode>
),
<collectingCountry id="F34F763BFFB8FF9141A451DC69928BCB" box="[1206,1277,1676,1700]" name="Laos" pageId="8" pageNumber="243">LAOS</collectingCountry>
,
<collectingRegion id="499CF849FFB8FF91401451DB68D08BCC" box="[1286,1471,1675,1699]" country="Laos" name="Champasak" pageId="8" pageNumber="243">Champasak Prov.</collectingRegion>
, env. of
<location id="8E876070FFB8FF91468951FB6EAA8BAC" LSID="urn:lsid:plazi:treatment:03F187BDFFB8FF9045E650C36FF88C9A:8E876070FFB8FF91468951FB6EAA8BAC" box="[923,965,1707,1731]" country="Laos" latitude="15.193723" longLatPrecision="1" longitude="106.09925" municipality="Male" name="Tad" pageId="8" pageNumber="243" stateProvince="Champasak">Tad</location>
E-Tu resort,
<geoCoordinate id="EE6C506CFFB8FF91417351FA686B8BAC" box="[1121,1284,1706,1731]" degrees="15" direction="north" minutes="11" orientation="latitude" pageId="8" pageNumber="243" precision="1" seconds="37.40" value="15.193723">15°1137.40″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB8FF91401D51FB68D38BAC" box="[1295,1468,1706,1731]" degrees="106" direction="east" minutes="5" orientation="longitude" pageId="8" pageNumber="243" precision="1" seconds="57.28" value="106.09925">106° 557.28″E</geoCoordinate>
, coll.
<collectorName id="26AD537DFFB8FF914691519C69418B8C" box="[899,1070,1739,1763]" pageId="8" pageNumber="243">M.M. Omelko</collectorName>
,
<date id="FFE6106BFFB8FF91412C519B685A8B8B" box="[1086,1333,1739,1764]" pageId="8" pageNumber="243" value="2013-11-01" valueMax="2013-11-06" valueMin="2013-11-01">
<collectingDate id="EFA2E983FFB8FF91412C519B685A8B8B" box="[1086,1333,1739,1764]" pageId="8" pageNumber="243" value="2013-11-01" valueMax="2013-11-06" valueMin="2013-11-01">16 November 2013</collectingDate>
</date>
;
<specimenCount id="9D5EFD22FFB8FF914057519C68CA8B8C" box="[1349,1445,1739,1764]" count="2" pageId="8" pageNumber="243" type="male">2 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB8FF9140A7519B6EF48A6C" count="5" pageId="8" pageNumber="243" type="female">5 females</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B303CF6FFB8FF9146B851BB69138A0B" collectingDate="2013-11-19" collectingDateMax="2013-11-24" collectingDateMin="2013-11-19" collectionCode="FEFU" collectorName="M. M. Omelko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="8" pageNumber="243" specimenCount="15" specimenCount-female="2" specimenCount-male="13" stateProvince="Viangchan">
<collectingRegion id="499CF849FFB8FF9146B851BB69318A6C" box="[938,1118,1771,1795]" country="Laos" name="Viangchan" pageId="8" pageNumber="243">Vientiane Prov.</collectingRegion>
, env. of
<location id="8E876070FFB8FF9141D551BB68D28A6C" LSID="urn:lsid:plazi:treatment:03F187BDFFB8FF9045E650C36FF88C9A:8E876070FFB8FF9141D551BB68D28A6C" box="[1223,1469,1771,1795]" country="Laos" latitude="18.614773" longLatPrecision="1" longitude="102.40885" name="Nam-Lik Eco-Village" pageId="8" pageNumber="243" stateProvince="Viangchan">Nam-Lik Eco-Village</location>
,
<geoCoordinate id="EE6C506CFFB8FF914651505A6E8D8A4C" box="[835,994,1802,1827]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="8" pageNumber="243" precision="1" seconds="53.18" value="18.614773">18°3653.18″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB8FF9146FA505A69FB8A4C" box="[1000,1172,1802,1827]" degrees="102" direction="east" minutes="24" orientation="longitude" pageId="8" pageNumber="243" precision="1" seconds="31.87" value="102.40885">102°2431.87″E</geoCoordinate>
,
<collectingMethod id="52194EBCFFB8FF91418F505B687A8A4C" box="[1181,1301,1803,1827]" pageId="8" pageNumber="243">pitfall traps</collectingMethod>
at meadows and rice paddies, coll.
<collectorName id="26AD537DFFB8FF914102507C69DF8A2C" box="[1040,1200,1835,1859]" pageId="8" pageNumber="243">M.M. Omelko</collectorName>
,
<date id="FFE6106BFFB8FF9141AC507C68D18A2B" box="[1214,1470,1835,1860]" pageId="8" pageNumber="243" value="2013-11-19" valueMax="2013-11-24" valueMin="2013-11-19">
<collectingDate id="EFA2E983FFB8FF9141AC507C68D18A2B" box="[1214,1470,1835,1860]" pageId="8" pageNumber="243" value="2013-11-19" valueMax="2013-11-24" valueMin="2013-11-19">1924 November 2013</collectingDate>
</date>
;
<specimenCount id="9D5EFD22FFB8FF914651501C6EC78A0C" box="[835,936,1867,1892]" count="13" pageId="8" pageNumber="243" type="male">13 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB8FF9146A7501C69718A0C" box="[949,1054,1867,1892]" count="2" pageId="8" pageNumber="243" type="female">2 females</specimenCount>
(
<collectionCode id="ED49AE6EFFB8FF91413D501C69198A0B" box="[1071,1142,1868,1892]" pageId="8" pageNumber="243">FEFU</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B303CF6FFB8FF914198501C682B8AEB" collectingDate="2016-05-17" collectingDateMax="2016-06-12" collectingDateMin="2016-05-17" collectionCode="FEFU" country="Laos" pageId="8" pageNumber="243" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Viangchan">
same place and collector,
<date id="FFE6106BFFB8FF9140B5501C697E8AEC" pageId="8" pageNumber="243" value="2016-05-17" valueMax="2016-06-12" valueMin="2016-05-17">
<collectingDate id="EFA2E983FFB8FF9140B5501C697E8AEC" pageId="8" pageNumber="243" value="2016-05-17" valueMax="2016-06-12" valueMin="2016-05-17">17 May12 June 2016</collectingDate>
</date>
;
<specimenCount id="9D5EFD22FFB8FF914108503C69148AEC" box="[1050,1147,1899,1924]" count="10" pageId="8" pageNumber="243" type="male">10 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB8FF914196503B69858AEC" box="[1156,1258,1899,1923]" count="5" pageId="8" pageNumber="243" type="female">5 females</specimenCount>
(
<collectionCode id="ED49AE6EFFB8FF9141E4503C68528AEB" box="[1270,1341,1900,1924]" pageId="8" pageNumber="243">FEFU</collectionCode>
)
</materialsCitation>
, same place and collector,
<date id="FFE6106BFFB8FF9146CD50DC69B98ACB" box="[991,1238,1932,1956]" pageId="8" pageNumber="243" value="2017-05-30" valueMax="2017-06-27" valueMin="2017-05-30">30 May27 June 2017</date>
.
</paragraph>
<paragraph id="8BE736ABFFB8FF904651509C6C278C3A" blockId="8.[835,1475,1995,2084]" lastBlockId="9.[113,752,189,341]" lastPageId="9" lastPageNumber="244" pageId="8" pageNumber="243">
<emphasis id="B92CEAB9FFB8FF914651509C6EE08A8B" bold="true" box="[835,911,1996,2020]" pageId="8" pageNumber="243">Notes.</emphasis>
I do not provide a redescription here because this species was well illustrated recently (see references above). A single male collected in
<collectingRegion id="499CF849FFB8FF9141D65F5B68C7854C" box="[1220,1448,2059,2083]" country="Laos" name="Champasak" pageId="8" pageNumber="243">Champasak Province</collectingRegion>
is noticeably larger than those from
<collectingRegion id="499CF849FFB9FF9044FF57ED6FD08DBA" box="[493,703,189,213]" country="Laos" name="Viangchan" pageId="9" pageNumber="244">Vientiane Province</collectingRegion>
and its palp structure is slightly different. There is a possibility that it represents an undescribed
<taxonomicName id="4C584D28FFB9FF9044FB57AD6FDE8C7A" box="[489,689,253,277]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92CEAB9FFB9FF9044FB57AD6F3F8C7A" box="[489,592,253,277]" italics="true" pageId="9" pageNumber="244">Trochosa</emphasis>
species.
</taxonomicName>
<figureCitation id="13632A2EFFB9FF9047A857AD6D8B8C5A" captionStart="Figs" captionStartId="8.[135,179,1450,1472]" captionTargetBox="[140,1465,189,1434]" captionTargetId="figure-7@8.[135,1473,189,1434]" captionTargetPageId="8" captionText="Figs. 2731. Male (27), female (28), male palp (2930, cymbial claw is broken off) and epigyne (31) of Trochosa ruricoloides. 27, 28 dorsal; 29, 31 ventral; 30 retrolateral. Scale: 1 mm (2728), 0.2 mm (2931)." figureDoi="http://doi.org/10.5281/zenodo.14683152" httpUri="https://zenodo.org/record/14683152/files/figure.png" pageId="9" pageNumber="244">Figs. 27, 2930</figureCitation>
show a typical male of
<taxonomicName id="4C584D28FFB9FF904714564E6FDF8C5A" authorityName="Schenkel" authorityYear="1963" box="[518,688,285,309]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFB9FF904714564E6FDF8C5A" box="[518,688,285,309]" italics="true" pageId="9" pageNumber="244">T. ruricoloides</emphasis>
</taxonomicName>
from
<collectingRegion id="499CF849FFB9FF904563566D6C2B8C3A" box="[113,324,317,341]" country="Laos" name="Viangchan" pageId="9" pageNumber="244">Vientiane Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF904563562D6FF88C9A" pageId="9" pageNumber="244" type="distribution">
<paragraph id="8BE736ABFFB9FF904563562D6FF88C9A" blockId="9.[113,753,381,501]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF904563562D6C678CFA" bold="true" box="[113,264,381,405]" pageId="9" pageNumber="244">Distribution.</emphasis>
<collectingCountry id="F34F763BFFB9FF904400562D6C228CFA" box="[274,333,381,405]" name="India" pageId="9" pageNumber="244">India</collectingCountry>
,
<collectingCountry id="F34F763BFFB9FF90444E562D6CF28CFA" box="[348,413,381,405]" name="China" pageId="9" pageNumber="244">China</collectingCountry>
(
<collectingRegion id="499CF849FFB9FF9044A2562D6F6C8CFA" box="[432,515,381,405]" country="China" name="Hainan" pageId="9" pageNumber="244">Hainan</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF904700562D6FFB8CFA" box="[530,660,381,405]" country="China" name="Guangdong" pageId="9" pageNumber="244">Guangdong</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9047B1562D6F838CFA" box="[675,748,381,405]" country="China" name="Fujian" pageId="9" pageNumber="244">Fujian</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF90456056CE6DA68CDA" box="[114,201,414,437]" country="China" name="Yunnan" pageId="9" pageNumber="244">Yunnan</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9045C456CD6C498CDA" box="[214,294,413,437]" country="China" name="Jiangxi" pageId="9" pageNumber="244">Jiangxi</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF90442656CD6CF98CDA" box="[308,406,413,437]" country="China" name="Zhejiang" pageId="9" pageNumber="244">Zhejiang</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9044B156CE6C828CDA" box="[419,493,414,437]" country="China" name="Hunan" pageId="9" pageNumber="244">Hunan</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9044E856CD6F518CDA" box="[506,574,413,437]" country="China" name="Hubei" pageId="9" pageNumber="244">Hubei</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF90475956CD6FCB8CDA" box="[587,676,413,437]" country="China" name="Sichuan" pageId="9" pageNumber="244">Sichuan</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9047A056CD6F828CDA" box="[690,749,413,437]" country="China" name="Xizang" pageId="9" pageNumber="244">Tibet</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF90456356ED6DA18CBA" box="[113,206,445,469]" country="China" name="Shaanxi" pageId="9" pageNumber="244">Shaanxi</collectingRegion>
),
<collectingCountry id="F34F763BFFB9FF9045F756ED6C558CBA" box="[229,314,445,469]" name="Taiwan" pageId="9" pageNumber="244">Taiwan</collectingCountry>
,
<collectingCountry id="F34F763BFFB9FF90445A56ED6CC38CBA" box="[328,428,445,469]" name="Thailand" pageId="9" pageNumber="244">Thailand</collectingCountry>
,
<collectingCountry id="F34F763BFFB9FF9044A956ED6F4D8CBA" box="[443,546,445,469]" name="Malaysia" pageId="9" pageNumber="244">Malaysia</collectingCountry>
,
<collectingCountry id="F34F763BFFB9FF90472356ED6FF08CBA" box="[561,671,445,469]" name="Indonesia" pageId="9" pageNumber="244">Indonesia</collectingCountry>
,
<collectingCountry id="F34F763BFFB9FF9047BC56EE6D928C9A" name="Papua New Guinea" pageId="9" pageNumber="244">Papua New Guinea</collectingCountry>
, and
<collectingCountry id="F34F763BFFB9FF90442A568E6C038C9A" box="[312,364,478,501]" name="Laos" pageId="9" pageNumber="244">Laos</collectingCountry>
(new record;
<figureCitation id="13632A2EFFB9FF904715568D6FE48C9A" box="[519,651,477,501]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="9" pageNumber="244">Figs. 5253</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,187 @@
<document id="D61D79DC12915082501C77CFF741DD31" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB9FF90443F554D681F8D9A" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Wadicosa Zyuzin 1985" docType="treatment" docVersion="2" lastPageNumber="244" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="244" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="04F6C8424E710C9BFCEF1F2EC75ED5A7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7EBF4A68B241D36D435B75778F033794">
<mods:title id="D3F2EC87E8EB8A042AE7A086B8A7D462">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="117D9F2E1E34136DF9BFB3C412F97234" type="personal">
<mods:role id="A9262AA06A00D4F78109EBD76220AEFB">
<mods:roleTerm id="0C997F42E572002CECB35F52D7B8E123">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67437F8923CCF69625B570D27BEC0419">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DAF1D7B37B052A6F214C58D85B49F2D0">text</mods:typeOfResource>
<mods:relatedItem id="5A91FDC931886F806341C8024A6886C4" type="host">
<mods:titleInfo id="9D9E82AF37A607C80C4154937585CE1B">
<mods:title id="ADEC73468C9ED7BC08CE2057BF9A6FE9">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="40F9A6D5D0551634585DE65F29AB81B7">
<mods:date id="64FC7E4ECC0F7C57EF976A19183585D3">2024</mods:date>
<mods:detail id="0941C623EDD0720B6BE513C052120606" type="pubDate">
<mods:number id="909F22B4CB31F3D6A9AC603B013042B3">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="EC07AD91C1D3F72551EEACC5D2EABA85" type="volume">
<mods:number id="B289301CB4B9E2619F2AD11692F2E326">72</mods:number>
</mods:detail>
<mods:extent id="4B43B645B7324652ECB1ECD5410E47AF" unit="page">
<mods:start id="85160A652F2266B51EDCC788AF1465BE">235</mods:start>
<mods:end id="E720097899C8480895C2245D29F20E6E">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9C4DA53BA760E8DE275E756E8DAFC496">journal article</mods:classification>
<mods:identifier id="1B7ED7F0D333055F91516C6369A08E0F" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="951EC3C8F6F043ACEDA70092874DF1E4" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="BE24B62F9970F04EF94BA5003BC769B1" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="C7CFCD90E23281F6660E92EB685B608D" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB9FF90443F554D681F8D9A" ID-DOI="http://doi.org/10.5281/zenodo.14683162" ID-Zenodo-Dep="14683162" LSID="urn:lsid:plazi:treatment:03F187BDFFB9FF90443F554D681F8D9A" httpUri="http://treatment.plazi.org/id/03F187BDFFB9FF90443F554D681F8D9A" lastPageNumber="244" pageId="9" pageNumber="244">
<subSubSection id="C3426520FFB9FF90443F554D6F5B8F5A" box="[301,564,541,565]" pageId="9" pageNumber="244" type="nomenclature">
<paragraph id="8BE736ABFFB9FF90443F554D6F5B8F5A" blockId="9.[301,564,541,565]" box="[301,564,541,565]" pageId="9" pageNumber="244">
<heading id="D0AF81C7FFB9FF90443F554D6F5B8F5A" bold="true" box="[301,564,541,565]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="244" reason="2">
<taxonomicName id="4C584D28FFB9FF90443F554D6F5B8F5A" ID-CoL="87D8" authority="Zyuzin, 1985" authorityName="Zyuzin" authorityYear="1985" box="[301,564,541,565]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF90443F554D6F5B8F5A" bold="true" box="[301,564,541,565]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90443F554D6CF78F5A" bold="true" box="[301,408,541,565]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
<bibRefCitation id="EFC94B5AFFB9FF9044B2554D6F5B8F5A" author="Zyuzin AA" box="[416,564,541,565]" pageId="9" pageNumber="244" pagination="40 - 51" refId="ref11713" refString="Zyuzin AA (1985) Generic and subfamilial criteria in the systematics of the spider family Lycosidae (Aranei), with the description of a new genus and two new subfamilies. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 139: 40 - 51" type="journal article" year="1985">Zyuzin, 1985</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF904563550D6C498FFA" pageId="9" pageNumber="244" type="type_taxon">
<paragraph id="8BE736ABFFB9FF904563550D6C498FFA" blockId="9.[113,752,605,661]" pageId="9" pageNumber="244">
<materialsCitation id="3B303CF6FFB9FF904563550D6C4E8FFA" location="West Asia" pageId="9" pageNumber="244" specimenCount="1" typeStatus="holotype">
<emphasis id="B92CEAB9FFB9FF904563550D6C658F1A" bold="true" box="[113,266,605,629]" pageId="9" pageNumber="244">
<typeStatus id="54E38809FFB9FF904563550D6DC48F1A" box="[113,171,605,629]" pageId="9" pageNumber="244">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C584D28FFB9FF904401550E6F838F1A" authority="O. Pickard-Cambridge, 1872" authorityName="O. Pickard-Cambridge" authorityYear="1872" box="[275,748,605,629]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF904401550E6CC88F1A" box="[275,423,605,629]" italics="true" pageId="9" pageNumber="244">Lycosa fidelis</emphasis>
O.
<bibRefCitation id="EFC94B5AFFB9FF9044C1550D6F838F1A" author="Pickard-Cambridge O" box="[467,748,605,629]" pageId="9" pageNumber="244" pagination="212 - 354" refId="ref10387" refString="Pickard-Cambridge O (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40 (1): 212 - 354." type="journal article" year="1872">Pickard-Cambridge, 1872</bibRefCitation>
</taxonomicName>
, from
<location id="8E876070FFB9FF9045BC552E6C4E8FFA" LSID="urn:lsid:plazi:treatment:03F187BDFFB9FF90443F554D681F8D9A:8E876070FFB9FF9045BC552E6C4E8FFA" box="[174,289,637,661]" name="West Asia" pageId="9" pageNumber="244">West Asia</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF90456355ED681F8D9A" pageId="9" pageNumber="244" type="discussion">
<paragraph id="8BE736ABFFB9FF90456355ED6CB5891A" blockId="9.[112,753,701,1141]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90456355ED6D9D8FBA" bold="true" box="[113,242,701,725]" pageId="9" pageNumber="244">Comments.</emphasis>
<taxonomicName id="4C584D28FFB9FF9045E455ED6C318FBA" authorityName="Zyuzin" authorityYear="1985" box="[246,350,701,725]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF9045E455ED6C318FBA" box="[246,350,701,725]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
is a comparatively small genus of wolf spiders consisting of 17 named species (
<bibRefCitation id="EFC94B5AFFB9FF904701558D6FFE8F9A" author="WSC" box="[531,657,733,757]" pageId="9" pageNumber="244" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC, 2024</bibRefCitation>
) and one species from Borneo which was described but not formally named (
<bibRefCitation id="EFC94B5AFFB9FF9045D9544D6CFE8E5A" author="Zehethofer K" box="[203,401,797,821]" pageId="9" pageNumber="244" pagination="161 - 165" refId="ref11614" refString="Zehethofer K (1998) Zum Vorkommen einer Wadicosa - Art in Sabah, Nord-Borneo (Araneae: Lycosidae). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 85: 161 - 165." type="journal article" year="1998">Zehethofer, 1998</bibRefCitation>
).
<taxonomicName id="4C584D28FFB9FF9044B5544D6F7D8E5A" authorityName="Zyuzin" authorityYear="1985" box="[423,530,797,821]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF9044B5544D6F7D8E5A" box="[423,530,797,821]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
was established by
<bibRefCitation id="EFC94B5AFFB9FF904560546D6C7F8E3A" author="Zyuzin AA" box="[114,272,829,853]" pageId="9" pageNumber="244" pagination="40 - 51" refId="ref11713" refString="Zyuzin AA (1985) Generic and subfamilial criteria in the systematics of the spider family Lycosidae (Aranei), with the description of a new genus and two new subfamilies. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, Leningrad, 139: 40 - 51" type="journal article" year="1985">Zyuzin (1985)</bibRefCitation>
as the only genus of Wadicosinae Zуuzin, 1985 with the
<typeStatus id="54E38809FFB9FF904405540E6C2A8E19" box="[279,325,862,886]" pageId="9" pageNumber="244">type</typeStatus>
species
<taxonomicName id="4C584D28FFB9FF9044BA540E6F838E1A" authority="Lucas, 1846" authorityName="Lucas" authorityYear="1846" box="[424,748,861,885]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="venatrix">
<emphasis id="B92CEAB9FFB9FF9044BA540E6F378E1A" box="[424,600,862,885]" italics="true" pageId="9" pageNumber="244">Lycosa venatrix</emphasis>
<bibRefCitation id="EFC94B5AFFB9FF904770540E6F838E1A" author="Lucas H" box="[610,748,861,885]" pageId="9" pageNumber="244" pagination="89 - 271" refId="ref10010" refString="Lucas H (1846) Histoire naturelle des animaux articules. In: Exploration scientifique de l'Algerie pendant les annees 1840, 1841, 1842 publiee par ordre du Gouvernement et avec le concours d'une commission academique. Paris, Sciences physiques, Zoologie 1, pp. 89 - 271." type="journal article" year="1846">Lucas, 1846</bibRefCitation>
</taxonomicName>
. The genus diagnosis was amended by
<bibRefCitation id="EFC94B5AFFB9FF90471F542D6DD78EDA" author="Kronestedt T &amp; Zyuzin AA" pageId="9" pageNumber="244" pagination="813 - 828" refId="ref9908" refString="Kronestedt T &amp; Zyuzin AA (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescription of the species. Contributions to Natural History, 12: 813 - 828." type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009)</bibRefCitation>
together with the fixation of
<taxonomicName id="4C584D28FFB9FF9044F154CE6C458EBA" authority="O. Pickard-Cambridge, 1872" authorityName="O. Pickard-Cambridge" authorityYear="1872" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF9044F154CE6F1D8EDA" box="[483,626,925,949]" italics="true" pageId="9" pageNumber="244">Lycosa fidelis</emphasis>
O.
<bibRefCitation id="EFC94B5AFFB9FF90478554CD6C458EBA" author="Pickard-Cambridge O" pageId="9" pageNumber="244" pagination="212 - 354" refId="ref10387" refString="Pickard-Cambridge O (1872) General list of the spiders of Palestine and Syria, with descriptions of numerous new species, and characters of two new genera. Proceedings of the Zoological Society of London, 40 (1): 212 - 354." type="journal article" year="1872">Pickard-Cambridge, 1872</bibRefCitation>
</taxonomicName>
as a new
<typeStatus id="54E38809FFB9FF90448154EE6CAE8EB9" box="[403,449,958,982]" pageId="9" pageNumber="244">type</typeStatus>
species. Males of
<taxonomicName id="4C584D28FFB9FF90479554ED6F9F8EBA" authorityName="Zyuzin" authorityYear="1985" box="[647,752,957,981]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF90479554ED6F9F8EBA" box="[647,752,957,981]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
can be recognised by the palpal tegulum having an anterior retrolateral process. Some species also have a posterior retrolateral process. Females have epigyne with oval or rounded foveolae (pockets). For the full list of characters, see
<bibRefCitation id="EFC94B5AFFB9FF90458E530D6CB9891A" author="Kronestedt T &amp; Zyuzin AA" box="[156,470,1117,1141]" pageId="9" pageNumber="244" pagination="813 - 828" refId="ref9908" refString="Kronestedt T &amp; Zyuzin AA (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescription of the species. Contributions to Natural History, 12: 813 - 828." type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE736ABFFB9FF90456353CE6FCC8B7A" blockId="9.[113,753,1181,1557]" pageId="9" pageNumber="244">
Most species are restricted in their distribution to the Afrotropical and Indomalayan realms (ten and four, respectively). One species is known from the Palearctic; three species, including the
<typeStatus id="54E38809FFB9FF9044BA53AE6CB98879" box="[424,470,1278,1302]" pageId="9" pageNumber="244">type</typeStatus>
species
<taxonomicName id="4C584D28FFB9FF90472553AD6F85887A" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[567,746,1277,1301]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF90472553AD6F85887A" box="[567,746,1277,1301]" italics="true" pageId="9" pageNumber="244">Wadicosa fidelis</emphasis>
</taxonomicName>
, are known across the Palearctic and Indomalayan realms. The actual range boundaries of
<taxonomicName id="4C584D28FFB9FF9044E2526D6F0F883A" authorityName="Zyuzin" authorityYear="1985" box="[496,608,1341,1365]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF9044E2526D6F0F883A" box="[496,608,1341,1365]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
distribution remain unclear; however, they are evidently much broader than currently known. As mentioned earlier, one unnamed species is known from Borneo, and another poorly known species, currently placed in
<taxonomicName id="4C584D28FFB9FF9044B652ED6C9188BA" authorityName="C. L. Koch" authorityYear="1847" box="[420,510,1469,1493]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF9044B652ED6C9188BA" box="[420,510,1469,1493]" italics="true" pageId="9" pageNumber="244">Pardosa</emphasis>
</taxonomicName>
under the name of
<taxonomicName id="4C584D28FFB9FF9047CB52EE6C59889A" authority="(Dahl, 1908)" authorityName="Omelko" authorityYear="2024" baseAuthorityName="Dahl" baseAuthorityYear="1908" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="bleyi">
<emphasis id="B92CEAB9FFB9FF9047CB52EE6DCA889A" italics="true" pageId="9" pageNumber="244">P. bleyi</emphasis>
(
<bibRefCitation id="EFC94B5AFFB9FF9045A6528D6C41889A" author="Dahl F" box="[180,302,1501,1525]" pageId="9" pageNumber="244" pagination="175 - 678" refId="ref9364" refString="Dahl F (1908) Die Lycosiden oder Wolfspinnen Deutschlands und ihre Stellung im Haushalt der Natur. Nach statistischen Untersuchungen dargestellt. Nova Acta, Abhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher, 88 (3): 175 - 678." type="journal article" year="1908">Dahl, 1908</bibRefCitation>
)
</taxonomicName>
but undoubtedly belonging to
<taxonomicName id="4C584D28FFB9FF904792528D6F86889A" authorityName="Zyuzin" authorityYear="1985" box="[640,745,1501,1525]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF904792528D6F86889A" box="[640,745,1501,1525]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
, is described from
<collectingCountry id="F34F763BFFB9FF90442552AE6F668B7A" box="[311,521,1533,1557]" name="Papua New Guinea" pageId="9" pageNumber="244">Papua New Guinea</collectingCountry>
(
<bibRefCitation id="EFC94B5AFFB9FF90470B52AD6FF88B7A" author="Dahl F" box="[537,663,1533,1557]" pageId="9" pageNumber="244" pagination="175 - 678" refId="ref9364" refString="Dahl F (1908) Die Lycosiden oder Wolfspinnen Deutschlands und ihre Stellung im Haushalt der Natur. Nach statistischen Untersuchungen dargestellt. Nova Acta, Abhandlungen der Kaiserlichen Leopoldinisch-Carolinischen Deutschen Akademie der Naturforscher, 88 (3): 175 - 678." type="journal article" year="1908">Dahl, 1908</bibRefCitation>
).
</paragraph>
<paragraph id="8BE736ABFFB9FF904563516D681F8D9A" blockId="9.[112,753,1597,2069]" lastBlockId="9.[812,1451,189,245]" pageId="9" pageNumber="244">
Almost all
<taxonomicName id="4C584D28FFB9FF9045F6516D6C228B3A" authorityName="Zyuzin" authorityYear="1985" box="[228,333,1597,1621]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="genus">
<emphasis id="B92CEAB9FFB9FF9045F6516D6C228B3A" box="[228,333,1597,1621]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
</taxonomicName>
are known from both sexes. Exceptions are
<taxonomicName id="4C584D28FFB9FF90458F510E6F728B1A" authority="Kronestedt, 2015" authorityName="Kronestedt" authorityYear="2015" box="[157,541,1629,1653]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="russellsmithi">
<emphasis id="B92CEAB9FFB9FF90458F510E6C3D8B1A" box="[157,338,1629,1653]" italics="true" pageId="9" pageNumber="244">W. russellsmithi</emphasis>
<bibRefCitation id="EFC94B5AFFB9FF90444F510D6F728B1A" author="Kronestedt T" box="[349,541,1629,1653]" pageId="9" pageNumber="244" pagination="1 - 19" refId="ref9789" refString="Kronestedt T (2015) Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa. European Journal of Taxonomy, 132: 1 - 19." type="journal article" year="2015">Kronestedt, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C584D28FFB9FF904748510E6FCE8BFA" authority="Ahmed, Anam, Saikia, Manthen &amp; Saikia, 2014" authorityName="Ahmed, Anam, Saikia, Manthen &amp; Saikia" authorityYear="2014" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="prasantae">
<emphasis id="B92CEAB9FFB9FF904748510E6F9F8B1A" box="[602,752,1630,1653]" italics="true" pageId="9" pageNumber="244">W. prasantae</emphasis>
<bibRefCitation id="EFC94B5AFFB9FF904563512D6FCE8BFA" author="Ahmed M &amp; Anam J &amp; Saikia MK &amp; Manthen SV &amp; Saikia PK" box="[113,673,1661,1686]" pageId="9" pageNumber="244" pagination="60 - 65" refId="ref8993" refString="Ahmed M, Anam J, Saikia MK, Manthen SV &amp; Saikia PK (2014) New records spider species under Wadicosa genus (Sub-order: Araneae; Family: Lycosidae) from agricultural field of Sonitpur District, Assam, India. Journal on New Biological Reports, 3 (1): 60 - 65." type="journal article" year="2014">Ahmed, Anam, Saikia, Manthen &amp; Saikia, 2014</bibRefCitation>
</taxonomicName>
which were described using females only. Most species are welldescribed/redescribed and depicted thanks to
<bibRefCitation id="EFC94B5AFFB9FF90476A51ED6DD88B9A" author="Kronestedt T" pageId="9" pageNumber="244" pagination="1 - 19" refId="ref9789" refString="Kronestedt T (2015) Species of Wadicosa (Araneae, Lycosidae): transfer of two species from Pardosa and description of three new species from Africa. European Journal of Taxonomy, 132: 1 - 19." type="journal article" year="2015">Kronestedt (2015</bibRefCitation>
,
<bibRefCitation id="EFC94B5AFFB9FF9045D5518D6C6C8B9A" author="Kronestedt T" box="[199,259,1757,1781]" pageId="9" pageNumber="244" pagination="295 - 300" refId="ref9829" refString="Kronestedt T (2017) Species of Wadicosa (Araneae, Lycosidae): a new species close to W. quadrifera (Gravely) from the Western Ghats, India. Zootaxa, 4300 (2): 295 - 300." type="journal article" year="2017">2017</bibRefCitation>
,
<bibRefCitation id="EFC94B5AFFB9FF904401518D6C228B9A" author="Kronestedt T" box="[275,333,1757,1781]" pageId="9" pageNumber="244" pagination="531 - 548" refId="ref9872" refString="Kronestedt T (2023) Species of Wadicosa (Araneae, Lycosidae): transfer of four species from Africa currently placed in Pardosa. Zootaxa, 5227 (5): 531 - 548." type="journal article" year="2023">2023</bibRefCitation>
) and Kronestedt &amp; Zуuzin (2009). However, a few
<taxonomicName id="4C584D28FFB9FF90442851AD6F698A7A" box="[314,518,1789,1813]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90442851AD6CC68A7A" box="[314,425,1789,1813]" italics="true" pageId="9" pageNumber="244">Wadicosa</emphasis>
species
</taxonomicName>
are poorly studied, being known only from the original descriptions. One such species is
<taxonomicName id="4C584D28FFB9FF9045F1506E6C078A3A" authorityName="Yin, Peng &amp; Zhang" authorityYear="1997" box="[227,360,1853,1877]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF9045F1506E6C078A3A" box="[227,360,1853,1877]" italics="true" pageId="9" pageNumber="244">W. daliensis</emphasis>
</taxonomicName>
which was only known from south
<collectingCountry id="F34F763BFFB9FF904563500D6DDD8A1A" box="[113,178,1885,1909]" name="China" pageId="9" pageNumber="244">China</collectingCountry>
(
<collectingRegion id="499CF849FFB9FF9045D6500D6C798A1A" box="[196,278,1885,1909]" country="China" name="Hainan" pageId="9" pageNumber="244">Hainan</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF904437500D6CC98A1A" box="[293,422,1885,1909]" country="China" name="Guangdong" pageId="9" pageNumber="244">Guangdong</collectingRegion>
,
<collectingRegion id="499CF849FFB9FF9044A6500E6F668A1A" box="[436,521,1886,1909]" country="China" name="Yunnan" pageId="9" pageNumber="244">Yunnan</collectingRegion>
Provinces) until the present study. Despite this species being depicted in several publications (
<bibRefCitation id="EFC94B5AFFB9FF90441950CD6CBC8ADA" author="Yin CM &amp; Peng XJ &amp; Zhang YJ" box="[267,467,1949,1973]" pageId="9" pageNumber="244" pagination="99 - 101" refId="ref11420" refString="Yin CM, Peng XJ &amp; Zhang YJ (1997 a) One new species of the genus Wadicosa (Araneae, Lycosidae) from China. Acta Arachnologica Sinica, 6: 99 - 101." type="journal article" year="1997">Yin et al., 1997a</bibRefCitation>
;
<bibRefCitation id="EFC94B5AFFB9FF9044F050CD6FC68ADA" author="Yin CM &amp; Peng XJ &amp; Xie LP &amp; Bao YH &amp; Wang JF" box="[482,681,1949,1973]" pageId="9" pageNumber="244" refId="ref11457" refString="Yin CM, Peng XJ, Xie LP, Bao YH &amp; Wang JF (1997 b) Lycosids in China (Arachnida: Araneae). Hunan Normal University Press, Changsha, 317 pp." type="book" year="1997">Yin et al., 1997b</bibRefCitation>
; Song et. al., 1999) all its images were taken from the original description. Because these images are very schematic, bulb and embolic division have never been studied before and the original diagnosis was made in Chinese only, I decided to redescribe this species in detail and re-diagnose it.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,647 @@
<document id="5384F3789C291353E478FFEA8558C7A8" ID-DOI="10.26107/RBZ-2024-0020" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683140" ID-ZooBank="4CE93702-0F79-43A8-8876-1FE76944C010" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172002389" checkinUser="felipe" docAuthor="Omelko, Mikhail M." docDate="2024" docId="03F187BDFFB9FF97467854666F058EFA" docLanguage="en" docName="RafflesBZool.72.235-251.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Wadicosa daliensis Yin, Peng &amp; Zhang 1997" docType="treatment" docVersion="2" lastPageNumber="249" masterDocId="FFC8FFC5FFB0FF99451257506D6F8D6F" masterDocTitle="New data on wolf spiders (Araneae: Lycosidae) from Laos" masterLastPageNumber="251" masterPageNumber="235" pageNumber="244" updateTime="1737175888817" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A66AFD494F46BE4CDA811DB33B54163B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="24C6F2DBD68DAB3CC1359B83A636EE24">
<mods:title id="94F47F5E46E43DF21FEE1477ED5D6845">New data on wolf spiders (Araneae: Lycosidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="063C4D56956FE3E26DF1ABEBC76BB372" type="personal">
<mods:role id="34C46804D9C050972C5723C1E720752C">
<mods:roleTerm id="3380B2E1633F816571490D62A6A2CE4D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EBB290F0E57EACEEA2D1C93D946D9F76">Omelko, Mikhail M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8BE8BE118428343FA99E307F25DAD0D">text</mods:typeOfResource>
<mods:relatedItem id="3660095329632BBB6F4B3AF90C9D8F1E" type="host">
<mods:titleInfo id="0E8EE2AF47DE3DCECB7456D58B5FED75">
<mods:title id="B38F025760AC8A850D1E277087AA64BD">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="9A0AFF45DF40EC5AA48489FE38AB957C">
<mods:date id="F62D0E37E4BE7B88CB8BCE7684524A37">2024</mods:date>
<mods:detail id="5612860FF2507533FAEC96AD9B1B561C" type="pubDate">
<mods:number id="3315A3077424DBEC753574120C5DC9EF">2024-08-12</mods:number>
</mods:detail>
<mods:detail id="8AC42ADDF3FE585DFB13C4296D826129" type="volume">
<mods:number id="1F841216FB662947FF866E2CC3EA9579">72</mods:number>
</mods:detail>
<mods:extent id="383148FBD6DE020B1DEA4E625E709B22" unit="page">
<mods:start id="954164F97CAE8F5A270D4E2FB0D5A4D6">235</mods:start>
<mods:end id="D939F171D31FF9E7D5A161DA107B3215">251</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D178D13C734A27161DD2E786E735D731">journal article</mods:classification>
<mods:identifier id="E3A5405DCECDA81716D32CF70E26E277" type="DOI">10.26107/RBZ-2024-0020</mods:identifier>
<mods:identifier id="634CB3AF4FFA3FE4B09933FE87E37E20" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="BA6E0A7DA56FA907C32727BBBE57730C" type="Zenodo-Dep">14683140</mods:identifier>
<mods:identifier id="1F194460EB15CC32881C570EBE217C6A" type="ZooBank">4CE93702-0F79-43A8-8876-1FE76944C010</mods:identifier>
</mods:mods>
<treatment id="03F187BDFFB9FF97467854666F058EFA" LSID="urn:lsid:plazi:treatment:03F187BDFFB9FF97467854666F058EFA" httpUri="http://treatment.plazi.org/id/03F187BDFFB9FF97467854666F058EFA" lastPageId="14" lastPageNumber="249" pageId="9" pageNumber="244">
<subSubSection id="C3426520FFB9FF904678546668028E20" box="[874,1389,822,847]" pageId="9" pageNumber="244" type="nomenclature">
<paragraph id="8BE736ABFFB9FF904678546668028E20" blockId="9.[874,1389,822,879]" box="[874,1389,822,847]" pageId="9" pageNumber="244">
<heading id="D0AF81C7FFB9FF904678546668028E20" bold="true" box="[874,1389,822,847]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="244" reason="2">
<taxonomicName id="4C584D28FFB9FF904678546668028E20" ID-CoL="5BSRH" authority="Yin, Peng &amp; Zhang, 1997" authorityName="Yin, Peng &amp; Zhang" authorityYear="1997" box="[874,1389,822,847]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF904678546668028E20" bold="true" box="[874,1389,822,847]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF904678546669528E21" bold="true" box="[874,1085,822,846]" italics="true" pageId="9" pageNumber="244">Wadicosa daliensis</emphasis>
<bibRefCitation id="EFC94B5AFFB9FF904154546768028E20" author="Yin CM &amp; Peng XJ &amp; Zhang YJ" box="[1094,1389,823,847]" pageId="9" pageNumber="244" pagination="99 - 101" refId="ref11420" refString="Yin CM, Peng XJ &amp; Zhang YJ (1997 a) One new species of the genus Wadicosa (Araneae, Lycosidae) from China. Acta Arachnologica Sinica, 6: 99 - 101." type="journal article" year="1997">Yin, Peng &amp; Zhang, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF904131540769DB8E00" box="[1059,1204,855,879]" pageId="9" pageNumber="244" type="description">
<paragraph id="8BE736ABFFB9FF904131540769DB8E00" blockId="9.[874,1389,822,879]" box="[1059,1204,855,879]" pageId="9" pageNumber="244">
(
<figureCitation id="13632A2EFFB9FF904139540769C28E00" box="[1067,1197,855,879]" captionStart-0="Figs" captionStart-1="Figs" captionStart-2="Figs" captionStart-3="Figs" captionStartId-0="10.[136,180,1940,1962]" captionStartId-1="12.[136,180,1977,1999]" captionStartId-2="13.[113,157,1230,1252]" captionStartId-3="13.[114,159,1980,2002]" captionTargetBox-0="[146,1455,318,1909]" captionTargetBox-1="[177,1430,206,1961]" captionTargetBox-2="[113,1451,274,1205]" captionTargetBox-3="[113,1451,1365,1964]" captionTargetId-0="figure-7@10.[136,1474,312,1924]" captionTargetId-1="figure-7@12.[174,1436,190,1962]" captionTargetId-2="figure-10@13.[113,1451,274,1215]" captionTargetId-3="figure-66@13.[113,1451,1364,1964]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="13" captionText-0="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." captionText-1="Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of Wadicosa daliensis. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis." captionText-2="Figs. 4951. Epigyne of Wadicosa daliensis. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm. Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem." captionText-3="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi-0="http://doi.org/10.5281/zenodo.14683154" figureDoi-1="http://doi.org/10.5281/zenodo.14683156" figureDoi-2="http://doi.org/10.5281/zenodo.14683158" figureDoi-3="http://doi.org/10.5281/zenodo.14683160" httpUri-0="https://zenodo.org/record/14683154/files/figure.png" httpUri-1="https://zenodo.org/record/14683156/files/figure.png" httpUri-2="https://zenodo.org/record/14683158/files/figure.png" httpUri-3="https://zenodo.org/record/14683160/files/figure.png" pageId="9" pageNumber="244">Figs. 3253</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF90463E54C669828EA8" pageId="9" pageNumber="244" type="reference_group">
<paragraph id="8BE736ABFFB9FF90463E54C668138EC4" blockId="9.[812,1404,917,968]" box="[812,1404,917,939]" pageId="9" pageNumber="244">
<taxonomicName id="4C584D28FFB9FF90463E54C669018EC4" ID-CoL="5BSRH" authority="Yin et al., 1997: 99" authorityName="Yin" authorityPageNumber="99" authorityYear="1997" box="[812,1134,917,939]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF90463E54C66ECC8EC5" box="[812,931,917,939]" italics="true" pageId="9" pageNumber="244">W. daliensis</emphasis>
Yin et al., 1997: 99
</taxonomicName>
, fig. 16, male and female.
</paragraph>
<paragraph id="8BE736ABFFB9FF90463E54E269828EA8" blockId="9.[812,1404,917,968]" box="[812,1261,946,968]" pageId="9" pageNumber="244">
See the full list of references in
<bibRefCitation id="EFC94B5AFFB9FF90417954E269858EA8" author="WSC" box="[1131,1258,946,967]" pageId="9" pageNumber="244" refId="ref11273" refString="WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern. http: // wsc. nmbe. ch (Accessed 26 April 2024)" type="url" year="2024">WSC (2024)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF90463E54BD696F89AA" pageId="9" pageNumber="244" type="materials_examined">
<paragraph id="8BE736ABFFB9FF90463E54BD696F89AA" blockId="9.[811,1452,1005,1221]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90463E54BD6965896A" bold="true" box="[812,1034,1005,1029]" pageId="9" pageNumber="244">Material examined</emphasis>
.
<materialsCitation id="3B303CF6FFB9FF90410954BE6E0A89EA" collectingDate="2013-11-07" collectingDateMax="2013-11-25" collectingDateMin="2013-11-07" collectionCode="FEFU" collectorName="M. M. Omelko" country="Laos" latitude="18.614773" location="Nam-Lik Eco-Village" longLatPrecision="1" longitude="102.40885" pageId="9" pageNumber="244" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Viangchan">
<specimenCount id="9D5EFD22FFB9FF90410954BE691A896A" box="[1051,1141,1005,1029]" count="7" pageId="9" pageNumber="244" type="male">7 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB9FF90419154BD6980896A" box="[1155,1263,1005,1029]" count="3" pageId="9" pageNumber="244" type="female">3 females</specimenCount>
(
<collectionCode id="ED49AE6EFFB9FF90401354BD6826896A" box="[1281,1353,1005,1029]" pageId="9" pageNumber="244">FEFU</collectionCode>
),
<collectingCountry id="F34F763BFFB9FF90404C54BD68C8896A" box="[1374,1447,1005,1029]" name="Laos" pageId="9" pageNumber="244">LAOS</collectingCountry>
,
<collectingRegion id="499CF849FFB9FF90463E535D6EBF894A" box="[812,976,1037,1061]" country="Laos" name="Viangchan" pageId="9" pageNumber="244">Vientiane Prov.</collectingRegion>
, env. of
<location id="8E876070FFB9FF904131535D6868894A" LSID="urn:lsid:plazi:treatment:03F187BDFFB9FF97467854666F058EFA:8E876070FFB9FF904131535D6868894A" box="[1059,1287,1037,1061]" country="Laos" latitude="18.614773" longLatPrecision="1" longitude="102.40885" name="Nam-Lik Eco-Village" pageId="9" pageNumber="244" stateProvince="Viangchan">Nam-Lik Eco-Village</location>
,
<geoCoordinate id="EE6C506CFFB9FF90401D535C68C3894A" box="[1295,1452,1036,1061]" degrees="18" direction="north" minutes="36" orientation="latitude" pageId="9" pageNumber="244" precision="1" seconds="53.18" value="18.614773">18°3653.18″N</geoCoordinate>
<geoCoordinate id="EE6C506CFFB9FF90463E537C6EB6892A" box="[812,985,1068,1093]" degrees="102" direction="east" minutes="24" orientation="longitude" pageId="9" pageNumber="244" precision="1" seconds="31.87" value="102.40885">102°2431.87″E</geoCoordinate>
,
<collectingMethod id="52194EBCFFB9FF9046F4537D684A892A" box="[998,1317,1069,1093]" pageId="9" pageNumber="244">hand picking and pitfall traps</collectingMethod>
at river and lake banks, along roads, coll.
<collectorName id="26AD537DFFB9FF90414D531E6997890A" box="[1119,1272,1101,1125]" pageId="9" pageNumber="244">M.M. Omelko</collectorName>
,
<date id="FFE6106BFFB9FF904013531D6E0A89EA" pageId="9" pageNumber="244" value="2013-11-07" valueMax="2013-11-25" valueMin="2013-11-07">
<collectingDate id="EFA2E983FFB9FF904013531D6E0A89EA" pageId="9" pageNumber="244" value="2013-11-07" valueMax="2013-11-25" valueMin="2013-11-07">725 November 2013</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5EFD22FFB9FF904661533D6EAC89EA" box="[883,963,1133,1157]" count="1" pageId="9" pageNumber="244" type="male">1 male</specimenCount>
, same place and collector,
<date id="FFE6106BFFB9FF9041EE533D68CA89EA" box="[1276,1445,1133,1157]" pageId="9" pageNumber="244" value="2016-06-04" valueMax="2016-06-09" valueMin="2016-06-04">49 June 2016</date>
;
<materialsCitation id="3B303CF6FFB9FF90463E53DD6E9389AA" collectingDate="2017-06-14" collectingDateMax="2017-07-25" collectingDateMin="2017-06-14" collectionCode="FEFU" country="Laos" pageId="9" pageNumber="244" specimenCount="10" specimenCount-female="1" specimenCount-male="9">
<specimenCount id="9D5EFD22FFB9FF90463E53DD6EE989CA" box="[812,902,1165,1189]" count="9" pageId="9" pageNumber="244" type="male">9 males</specimenCount>
,
<specimenCount id="9D5EFD22FFB9FF90468753DD6E9889CA" box="[917,1015,1165,1189]" count="1" pageId="9" pageNumber="244" type="female">1 female</specimenCount>
(
<collectionCode id="ED49AE6EFFB9FF90411B53DD693E89CA" box="[1033,1105,1165,1189]" pageId="9" pageNumber="244">FEFU</collectionCode>
), same place and collector,
<date id="FFE6106BFFB9FF90408253DD6E9389AA" pageId="9" pageNumber="244" value="2017-06-14" valueMax="2017-07-25" valueMin="2017-06-14">
<collectingDate id="EFA2E983FFB9FF90408253DD6E9389AA" pageId="9" pageNumber="244" value="2017-06-14" valueMax="2017-07-25" valueMin="2017-06-14">14 June25 July 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF90463E53BD69318ACA" pageId="9" pageNumber="244" type="diagnosis">
<paragraph id="8BE736ABFFB9FF90463E53BD69318ACA" blockId="9.[811,1452,1261,1958]" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90463E53BD6EF3886A" bold="true" box="[812,924,1261,1285]" pageId="9" pageNumber="244">Diagnosis</emphasis>
. By general appearance, palp and structure of its copulatory organs (shape of tegulum, tegular apophysis, embolus, fovea, and receptacle)
<taxonomicName id="4C584D28FFB9FF9041BF527D68E3882A" box="[1197,1420,1325,1349]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF9041BF527D68E3882A" box="[1197,1420,1325,1349]" italics="true" pageId="9" pageNumber="244">Wadicosa daliensis</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C584D28FFB9FF9046E7521E6933880A" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[1013,1116,1357,1381]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF9046E7521E6933880A" box="[1013,1116,1357,1381]" italics="true" pageId="9" pageNumber="244">W. fidelis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C584D28FFB9FF904185521E6E0088EA" authority="(Tanaka, 1985)" baseAuthorityName="Tanaka" baseAuthorityYear="1985" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFB9FF904185521E682D880A" box="[1175,1346,1357,1381]" italics="true" pageId="9" pageNumber="244">W. okinawensis</emphasis>
(
<bibRefCitation id="EFC94B5AFFB9FF904046521D6E0988EA" author="Tanaka H" pageId="9" pageNumber="244" pagination="51 - 87" refId="ref10801" refString="Tanaka H (1985) Descriptions of new species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, 33: 51 - 87." type="journal article" year="1985">Tanaka, 1985</bibRefCitation>
)
</taxonomicName>
. Males of
<taxonomicName id="4C584D28FFB9FF9046F8523E691C88EA" box="[1002,1139,1389,1413]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF9046F8523E691C88EA" box="[1002,1139,1389,1413]" italics="true" pageId="9" pageNumber="244">W. daliensis</emphasis>
</taxonomicName>
may be differentiated from
<taxonomicName id="4C584D28FFB9FF90463E52DE6EF988CA" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[812,918,1421,1445]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF90463E52DE6EF988CA" box="[812,918,1421,1445]" italics="true" pageId="9" pageNumber="244">W. fidelis</emphasis>
</taxonomicName>
by: 1) a large posterior retrolateral process of tegulum (PP; vs. small, cf.
<figureCitation id="13632A2EFFB9FF90414D52FD698E88AA" box="[1119,1249,1453,1477]" captionStart="Figs" captionStartId="12.[136,180,1977,1999]" captionTargetBox="[177,1430,206,1961]" captionTargetId="figure-7@12.[174,1436,190,1962]" captionTargetPageId="12" captionText="Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of Wadicosa daliensis. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.14683156" httpUri="https://zenodo.org/record/14683156/files/figure.png" pageId="9" pageNumber="244">Figs. 4143</figureCitation>
and figs. 1,
<quantity id="4CA09B4EFFB9FF90406352FD68C488AA" box="[1393,1451,1453,1477]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="9" pageNumber="244" unit="in" value="12.0">12 in</quantity>
<bibRefCitation id="EFC94B5AFFB9FF90463E529D6916888A" author="Kronestedt T &amp; Zyuzin AA" box="[812,1145,1485,1509]" pageId="9" pageNumber="244" pagination="813 - 828" refId="ref9908" refString="Kronestedt T &amp; Zyuzin AA (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescription of the species. Contributions to Natural History, 12: 813 - 828." type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009))</bibRefCitation>
, 2) tegular apophysis (TA) strongly curved, widened medially in ventral view (vs. slightly curved, widened distally, cf.
<figureCitation id="13632A2EFFB9FF9041AC515D68518B4A" box="[1214,1342,1549,1573]" captionStart="Figs" captionStartId="12.[136,180,1977,1999]" captionTargetBox="[177,1430,206,1961]" captionTargetId="figure-7@12.[174,1436,190,1962]" captionTargetPageId="12" captionText="Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of Wadicosa daliensis. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.14683156" httpUri="https://zenodo.org/record/14683156/files/figure.png" pageId="9" pageNumber="244">Figs. 39, 42</figureCitation>
and fig.
<quantity id="4CA09B4EFFB9FF90408C515D6E2E8B2A" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="244" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFC94B5AFFB9FF904658517D69E18B2A" author="Kronestedt T &amp; Zyuzin AA" box="[842,1166,1581,1605]" pageId="9" pageNumber="244" pagination="813 - 828" refId="ref9908" refString="Kronestedt T &amp; Zyuzin AA (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescription of the species. Contributions to Natural History, 12: 813 - 828." type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009))</bibRefCitation>
; from
<taxonomicName id="4C584D28FFB9FF9041CA517E68108B2A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[1240,1407,1581,1605]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFB9FF9041CA517E68108B2A" box="[1240,1407,1581,1605]" italics="true" pageId="9" pageNumber="244">W. okinawensis</emphasis>
</taxonomicName>
by: 1) dark carapace and abdomen coloration (vs. very light, cf.
<figureCitation id="13632A2EFFB9FF904641513D6EB68BEA" box="[851,985,1645,1669]" captionStart="Figs" captionStartId="10.[136,180,1940,1962]" captionTargetBox="[146,1455,318,1909]" captionTargetId="figure-7@10.[136,1474,312,1924]" captionTargetPageId="10" captionText="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." figureDoi="http://doi.org/10.5281/zenodo.14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="9" pageNumber="244">Figs. 32, 34</figureCitation>
and figs. 69A in
<bibRefCitation id="EFC94B5AFFB9FF9041B7513E68E98BEA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[1189,1414,1645,1670]" pageId="9" pageNumber="244" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
, 2) serrated retrolateral edge of tegulum (Ed; vs. smooth, cf.
<figureCitation id="13632A2EFFB9FF90409351DD6E298BAA" captionStart="Figs" captionStartId="12.[136,180,1977,1999]" captionTargetBox="[177,1430,206,1961]" captionTargetId="figure-7@12.[174,1436,190,1962]" captionTargetPageId="12" captionText="Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of Wadicosa daliensis. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.14683156" httpUri="https://zenodo.org/record/14683156/files/figure.png" pageId="9" pageNumber="244">Fig. 43</figureCitation>
and Figs. 69D, 70B in
<bibRefCitation id="EFC94B5AFFB9FF90415951FE684A8BAA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[1099,1317,1709,1734]" pageId="9" pageNumber="244" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
. Females of
<taxonomicName id="4C584D28FFB9FF90463C519E6EDB8B8A" box="[814,948,1741,1765]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFB9FF90463C519E6EDB8B8A" box="[814,948,1741,1765]" italics="true" pageId="9" pageNumber="244">W. daliensis</emphasis>
</taxonomicName>
can be differentiated from those of
<taxonomicName id="4C584D28FFB9FF904057519E68C48B8A" baseAuthorityName="O. Pickard-Cambridge" baseAuthorityYear="1872" box="[1349,1451,1741,1765]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis id="B92CEAB9FFB9FF904057519E68C48B8A" box="[1349,1451,1741,1765]" italics="true" pageId="9" pageNumber="244">W. fidelis</emphasis>
</taxonomicName>
by U-shaped receptacles (vs. rounded, cf.
<figureCitation id="13632A2EFFB9FF9041ED51BD68218A6A" box="[1279,1358,1773,1797]" captionStart="Figs" captionStartId="13.[113,157,1230,1252]" captionTargetBox="[113,1451,274,1205]" captionTargetId="figure-10@13.[113,1451,274,1215]" captionTargetPageId="13" captionText="Figs. 4951. Epigyne of Wadicosa daliensis. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm. Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem." figureDoi="http://doi.org/10.5281/zenodo.14683158" httpUri="https://zenodo.org/record/14683158/files/figure.png" pageId="9" pageNumber="244">Fig. 51</figureCitation>
and fig.
<quantity id="4CA09B4EFFB9FF90463E505D6E098A4A" box="[812,870,1805,1829]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" pageId="9" pageNumber="244" unit="in" value="17.0">17 in</quantity>
<bibRefCitation id="EFC94B5AFFB9FF904662505D69D98A4A" author="Kronestedt T &amp; Zyuzin AA" box="[880,1206,1805,1829]" pageId="9" pageNumber="244" pagination="813 - 828" refId="ref9908" refString="Kronestedt T &amp; Zyuzin AA (2009) Fixation of Lycosa fidelis O. Pickard-Cambridge, 1872 as the type species for the genus Wadicosa Zyuzin, 1985 (Araneae: Lycosidae), with a redescription of the species. Contributions to Natural History, 12: 813 - 828." type="journal article" year="2009">Kronestedt &amp; Zyuzin (2009))</bibRefCitation>
; from
<taxonomicName id="4C584D28FFB9FF904010505E68C48A4A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[1282,1451,1805,1829]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFB9FF904010505E68C48A4A" box="[1282,1451,1805,1829]" italics="true" pageId="9" pageNumber="244">W. okinawensis</emphasis>
</taxonomicName>
by: 1) septal base (SB) width/height ratio ca. 2.9 (vs. 2.1, cf.
<figureCitation id="13632A2EFFB9FF904642501D6EAA8A0A" box="[848,965,1869,1893]" captionStart="Figs" captionStartId="13.[113,157,1230,1252]" captionTargetBox="[113,1451,274,1205]" captionTargetId="figure-10@13.[113,1451,274,1215]" captionTargetPageId="13" captionText="Figs. 4951. Epigyne of Wadicosa daliensis. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm. Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem." figureDoi="http://doi.org/10.5281/zenodo.14683158" httpUri="https://zenodo.org/record/14683158/files/figure.png" pageId="9" pageNumber="244">Fig. 4950</figureCitation>
and fig. 69G in
<bibRefCitation id="EFC94B5AFFB9FF904168501E683D8A0A" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[1146,1362,1869,1894]" pageId="9" pageNumber="244" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
, 2) oval foveolae (Fv) (vs. almost round, cf.
<figureCitation id="13632A2EFFB9FF9041DA503D68238AEA" box="[1224,1356,1901,1925]" captionStart="Figs" captionStartId="13.[113,157,1230,1252]" captionTargetBox="[113,1451,274,1205]" captionTargetId="figure-10@13.[113,1451,274,1215]" captionTargetPageId="13" captionText="Figs. 4951. Epigyne of Wadicosa daliensis. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm. Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem." figureDoi="http://doi.org/10.5281/zenodo.14683158" httpUri="https://zenodo.org/record/14683158/files/figure.png" pageId="9" pageNumber="244">Figs. 4950</figureCitation>
and fig. 69G in
<bibRefCitation id="EFC94B5AFFB9FF90469250DE69358ACA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[896,1114,1933,1958]" pageId="9" pageNumber="244" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFB9FF97463E509D6FA98E5A" lastPageId="14" lastPageNumber="249" pageId="9" pageNumber="244" type="description">
<paragraph id="8BE736ABFFB9FF92463E509D6C0789F0" blockId="9.[811,1451,1997,2085]" lastBlockId="11.[112,753,999,1183]" lastPageId="11" lastPageNumber="246" pageId="9" pageNumber="244">
<emphasis id="B92CEAB9FFB9FF90463E509D6EDA8A8A" bold="true" box="[812,949,1997,2021]" pageId="9" pageNumber="244">Description.</emphasis>
Male (
<figureCitation id="13632A2EFFB9FF9046EF509D69128A8A" box="[1021,1149,1997,2021]" captionStart="Figs" captionStartId="10.[136,180,1940,1962]" captionTargetBox="[146,1455,318,1909]" captionTargetId="figure-7@10.[136,1474,312,1924]" captionTargetPageId="10" captionText="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." figureDoi="http://doi.org/10.5281/zenodo.14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="9" pageNumber="244">Figs. 3234</figureCitation>
). Total length 5.41. Carapace 3.01 long, 2.29 wide. Opisthosoma 2.42 long, 1.67 wide. Colouration. Carapace dark brown covered with more or less dense (cf.
<figureCitation id="13632A2EFFBBFF92440B54B76CA58E90" box="[281,458,999,1023]" captionStart="Figs" captionStartId="10.[136,180,1940,1962]" captionTargetBox="[146,1455,318,1909]" captionTargetId="figure-7@10.[136,1474,312,1924]" captionTargetPageId="10" captionText="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." figureDoi="http://doi.org/10.5281/zenodo.14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="11" pageNumber="246">Figs. 32 and 34</figureCitation>
) white setae, with yellow, star-like median band, eye field black. Lateral bands brown, indistinct, divided into several spots. Fovea thin, dark brown. Clypeus black, chelicerae dark brown, labium black. Endites dark brown with yellow outer edges. Sternum black with indistinct brown spots.
</paragraph>
<caption id="DF276623FFBAFF93459A50C46C888AA9" ID-DOI="http://doi.org/10.5281/zenodo.14683154" ID-Zenodo-Dep="14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="10" pageNumber="245" startId="10.[136,180,1940,1962]" targetBox="[146,1455,318,1909]" targetPageId="10" targetType="figure">
<paragraph id="8BE736ABFFBAFF93459A50C46C888AA9" blockId="10.[136,1475,1940,1991]" pageId="10" pageNumber="245">
Figs. 3237. Male (3234), female (3536) and male palp (37) of
<taxonomicName id="4C584D28FFBAFF93461750C46ED08AC6" box="[773,959,1940,1961]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBAFF93461750C46ED08AC6" box="[773,959,1940,1961]" italics="true" pageId="10" pageNumber="245">Wadicosa daliensis</emphasis>
</taxonomicName>
. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37).
</paragraph>
</caption>
<caption id="DF276623FFBBFF92456357ED6DA98D80" pageId="11" pageNumber="246" startId="11.[113,168,189,211]" targetType="table">
<paragraph id="8BE736ABFFBBFF92456357ED6DA98D80" blockId="11.[113,752,189,239]" pageId="11" pageNumber="246">
Table 7. Length of palp and leg segments for the male of
<taxonomicName id="4C584D28FFBBFF92478357ED6DA98D80" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="daliensis">Wadicosa daliensis</taxonomicName>
</paragraph>
</caption>
<caption id="DF276623FFBBFF92463F57ED6E878D80" pageId="11" pageNumber="246" startId="11.[813,871,189,211]" targetType="table">
<paragraph id="8BE736ABFFBBFF92463F57ED6E878D80" blockId="11.[813,1451,189,239]" pageId="11" pageNumber="246">
Table 9. Length of palp and leg segments for the female of
<taxonomicName id="4C584D28FFBBFF92463F578A6E878D80" box="[813,1000,218,239]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="daliensis">Wadicosa daliensis</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BE736ABFFBBFF9245FA56726FBC8F46" pageId="11" pageNumber="246">
<table id="F958C40BFFBB0066459356716FBB8F46" box="[129,724,289,553]" gridcols="7" gridrows="6" pageId="11" pageNumber="246">
<tr id="356834E9FFBB0066459356716FBB8C58" box="[129,724,289,311]" gridrow="0" pageId="11" pageNumber="246" rowspan-0="1">
<th id="76B95D95FFBB006645F256716C658C58" box="[224,266,289,311]" gridcol="1" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9245FA56726C6E8C58" bold="true" box="[232,257,290,311]" pageId="11" pageNumber="246">Fe</emphasis>
</th>
<th id="76B95D95FFBB0066442A56716C0D8C58" box="[312,354,289,311]" gridcol="2" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92442D56726C358C58" bold="true" box="[319,346,290,311]" pageId="11" pageNumber="246">Pa</emphasis>
</th>
<th id="76B95D95FFBB0066449D56716CD68C58" box="[399,441,289,311]" gridcol="3" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92448B56716CDF8C58" bold="true" box="[409,432,289,311]" pageId="11" pageNumber="246">Ti</emphasis>
</th>
<th id="76B95D95FFBB006644F556716F7E8C58" box="[487,529,289,311]" gridcol="4" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9244FF56726F638C58" bold="true" box="[493,524,290,311]" pageId="11" pageNumber="246">Mt</emphasis>
</th>
<th id="76B95D95FFBB0066472D56716F068C58" box="[575,617,289,311]" gridcol="5" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92475456726F0D8C58" bold="true" box="[582,610,290,311]" pageId="11" pageNumber="246">Ta</emphasis>
</th>
<th id="76B95D95FFBB0066478F56716FBB8C58" box="[669,724,289,311]" gridcol="6" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92478F56726FBB8C58" bold="true" box="[669,724,290,311]" pageId="11" pageNumber="246">Total</emphasis>
</th>
</tr>
<tr id="356834E9FFBB0066459356086FBB8C01" box="[129,724,344,366]" gridrow="1" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066459356086DC38C01" box="[129,172,344,366]" gridcol="0" gridrow="1" pageId="11" pageNumber="246">Palp</th>
<td id="76B95D95FFBB006645F256086C658C01" box="[224,266,344,366]" gridcol="1" gridrow="1" pageId="11" pageNumber="246">1.09</td>
<td id="76B95D95FFBB0066442A56086C0D8C01" box="[312,354,344,366]" gridcol="2" gridrow="1" pageId="11" pageNumber="246">0.52</td>
<td id="76B95D95FFBB0066449D56086CD68C01" box="[399,441,344,366]" gridcol="3" gridrow="1" pageId="11" pageNumber="246">0.66</td>
<td id="76B95D95FFBB006644F556086F7E8C01" box="[487,529,344,366]" gridcol="4" gridrow="1" pageId="11" pageNumber="246"></td>
<td id="76B95D95FFBB0066472D56086F068C01" box="[575,617,344,366]" gridcol="5" gridrow="1" pageId="11" pageNumber="246">1.23</td>
<td id="76B95D95FFBB0066478F56086FBB8C01" box="[669,724,344,366]" gridcol="6" gridrow="1" pageId="11" pageNumber="246">3.5</td>
</tr>
<tr id="356834E9FFBB0066459356D76FBB8CF2" box="[129,724,391,413]" gridrow="2" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066459356D76DC38CF2" box="[129,172,391,413]" gridcol="0" gridrow="2" pageId="11" pageNumber="246">I</th>
<td id="76B95D95FFBB006645F256D76C658CF2" box="[224,266,391,413]" gridcol="1" gridrow="2" pageId="11" pageNumber="246">2.27</td>
<td id="76B95D95FFBB0066442A56D76C0D8CF2" box="[312,354,391,413]" gridcol="2" gridrow="2" pageId="11" pageNumber="246">0.98</td>
<td id="76B95D95FFBB0066449D56D76CD68CF2" box="[399,441,391,413]" gridcol="3" gridrow="2" pageId="11" pageNumber="246">2.15</td>
<td id="76B95D95FFBB006644F556D76F7E8CF2" box="[487,529,391,413]" gridcol="4" gridrow="2" pageId="11" pageNumber="246">2.03</td>
<td id="76B95D95FFBB0066472D56D76F068CF2" box="[575,617,391,413]" gridcol="5" gridrow="2" pageId="11" pageNumber="246">1.36</td>
<td id="76B95D95FFBB0066478F56D76FBB8CF2" box="[669,724,391,413]" gridcol="6" gridrow="2" pageId="11" pageNumber="246">8.79</td>
</tr>
<tr id="356834E9FFBB0066459356E66FBB8CA3" box="[129,724,438,460]" gridrow="3" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066459356E66DC38CA3" box="[129,172,438,460]" gridcol="0" gridrow="3" pageId="11" pageNumber="246">II</th>
<td id="76B95D95FFBB006645F256E66C658CA3" box="[224,266,438,460]" gridcol="1" gridrow="3" pageId="11" pageNumber="246">2.21</td>
<td id="76B95D95FFBB0066442A56E66C0D8CA3" box="[312,354,438,460]" gridcol="2" gridrow="3" pageId="11" pageNumber="246">0.90</td>
<td id="76B95D95FFBB0066449D56E66CD68CA3" box="[399,441,438,460]" gridcol="3" gridrow="3" pageId="11" pageNumber="246">1.86</td>
<td id="76B95D95FFBB006644F556E66F7E8CA3" box="[487,529,438,460]" gridcol="4" gridrow="3" pageId="11" pageNumber="246">2.02</td>
<td id="76B95D95FFBB0066472D56E66F068CA3" box="[575,617,438,460]" gridcol="5" gridrow="3" pageId="11" pageNumber="246">1.27</td>
<td id="76B95D95FFBB0066478F56E66FBB8CA3" box="[669,724,438,460]" gridcol="6" gridrow="3" pageId="11" pageNumber="246">8.26</td>
</tr>
<tr id="356834E9FFBB0066459356B56FBB8C94" box="[129,724,485,507]" gridrow="4" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066459356B56DC38C94" box="[129,172,485,507]" gridcol="0" gridrow="4" pageId="11" pageNumber="246">III</th>
<td id="76B95D95FFBB006645F256B56C658C94" box="[224,266,485,507]" gridcol="1" gridrow="4" pageId="11" pageNumber="246">2.04</td>
<td id="76B95D95FFBB0066442A56B56C0D8C94" box="[312,354,485,507]" gridcol="2" gridrow="4" pageId="11" pageNumber="246">0.92</td>
<td id="76B95D95FFBB0066449D56B56CD68C94" box="[399,441,485,507]" gridcol="3" gridrow="4" pageId="11" pageNumber="246">1.59</td>
<td id="76B95D95FFBB006644F556B56F7E8C94" box="[487,529,485,507]" gridcol="4" gridrow="4" pageId="11" pageNumber="246">2.22</td>
<td id="76B95D95FFBB0066472D56B56F068C94" box="[575,617,485,507]" gridcol="5" gridrow="4" pageId="11" pageNumber="246">1.12</td>
<td id="76B95D95FFBB0066478F56B56FBB8C94" box="[669,724,485,507]" gridcol="6" gridrow="4" pageId="11" pageNumber="246">7.89</td>
</tr>
<tr id="356834E9FFBB0066459355436FBB8F46" box="[129,724,531,553]" gridrow="5" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066459355436DC38F46" box="[129,172,531,553]" gridcol="0" gridrow="5" pageId="11" pageNumber="246">IV</th>
<td id="76B95D95FFBB006645F255436C658F46" box="[224,266,531,553]" gridcol="1" gridrow="5" pageId="11" pageNumber="246">2.76</td>
<td id="76B95D95FFBB0066442A55436C0D8F46" box="[312,354,531,553]" gridcol="2" gridrow="5" pageId="11" pageNumber="246">1.02</td>
<td id="76B95D95FFBB0066449D55436CD68F46" box="[399,441,531,553]" gridcol="3" gridrow="5" pageId="11" pageNumber="246">2.43</td>
<td id="76B95D95FFBB006644F555436F7E8F46" box="[487,529,531,553]" gridcol="4" gridrow="5" pageId="11" pageNumber="246">3.82</td>
<td id="76B95D95FFBB0066472D55436F068F46" box="[575,617,531,553]" gridcol="5" gridrow="5" pageId="11" pageNumber="246">1.64</td>
<td id="76B95D95FFBB0066478F55436FBB8F46" box="[669,724,531,553]" gridcol="6" gridrow="5" pageId="11" pageNumber="246">11.67</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFBBFF92468B567268E78F45" pageId="11" pageNumber="246">
<table id="F958C40BFFBB0066462E567168E68F45" box="[828,1417,289,554]" gridcols="7" gridrows="6" pageId="11" pageNumber="246">
<tr id="356834E9FFBB0066462E567168E68C58" box="[828,1417,289,311]" gridrow="0" pageId="11" pageNumber="246" rowspan-0="1">
<th id="76B95D95FFBB0066468356716ED48C58" box="[913,955,289,311]" gridcol="1" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92468B56726EDD8C58" bold="true" box="[921,946,290,311]" pageId="11" pageNumber="246">Fe</emphasis>
</th>
<th id="76B95D95FFBB006646FA5671697D8C58" box="[1000,1042,289,311]" gridcol="2" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9246E2567269648C58" bold="true" box="[1008,1035,290,311]" pageId="11" pageNumber="246">Pa</emphasis>
</th>
<th id="76B95D95FFBB00664152567169058C58" box="[1088,1130,289,311]" gridcol="3" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92415B5671690F8C58" bold="true" box="[1097,1120,289,311]" pageId="11" pageNumber="246">Ti</emphasis>
</th>
<th id="76B95D95FFBB00664185567169AE8C58" box="[1175,1217,289,311]" gridcol="4" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92418F567269D38C58" bold="true" box="[1181,1212,290,311]" pageId="11" pageNumber="246">Mt</emphasis>
</th>
<th id="76B95D95FFBB006641FC567168778C58" box="[1262,1304,289,311]" gridcol="5" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9241E75672687E8C58" bold="true" box="[1269,1297,290,311]" pageId="11" pageNumber="246">Ta</emphasis>
</th>
<th id="76B95D95FFBB00664040567168E68C58" box="[1362,1417,289,311]" gridcol="6" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF924040567268E68C58" bold="true" box="[1362,1417,290,311]" pageId="11" pageNumber="246">Total</emphasis>
</th>
</tr>
<tr id="356834E9FFBB0066462E560868E68C01" box="[828,1417,344,366]" gridrow="1" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E56086E088C01" box="[828,871,344,366]" gridcol="0" gridrow="1" pageId="11" pageNumber="246">Palp</th>
<td id="76B95D95FFBB0066468356086ED48C01" box="[913,955,344,366]" gridcol="1" gridrow="1" pageId="11" pageNumber="246">1.11</td>
<td id="76B95D95FFBB006646FA5608697D8C01" box="[1000,1042,344,366]" gridcol="2" gridrow="1" pageId="11" pageNumber="246">0.55</td>
<td id="76B95D95FFBB00664152560869058C01" box="[1088,1130,344,366]" gridcol="3" gridrow="1" pageId="11" pageNumber="246">0.75</td>
<td id="76B95D95FFBB00664185560869AE8C01" box="[1175,1217,344,366]" gridcol="4" gridrow="1" pageId="11" pageNumber="246"></td>
<td id="76B95D95FFBB006641FC560868778C01" box="[1262,1304,344,366]" gridcol="5" gridrow="1" pageId="11" pageNumber="246">1.02</td>
<td id="76B95D95FFBB00664040560868E68C01" box="[1362,1417,344,366]" gridcol="6" gridrow="1" pageId="11" pageNumber="246">3.43</td>
</tr>
<tr id="356834E9FFBB0066462E56D768E68CF2" box="[828,1417,391,413]" gridrow="2" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E56D76E088CF2" box="[828,871,391,413]" gridcol="0" gridrow="2" pageId="11" pageNumber="246">I</th>
<td id="76B95D95FFBB0066468356D76ED48CF2" box="[913,955,391,413]" gridcol="1" gridrow="2" pageId="11" pageNumber="246">2.30</td>
<td id="76B95D95FFBB006646FA56D7697D8CF2" box="[1000,1042,391,413]" gridcol="2" gridrow="2" pageId="11" pageNumber="246">1.02</td>
<td id="76B95D95FFBB0066415256D769058CF2" box="[1088,1130,391,413]" gridcol="3" gridrow="2" pageId="11" pageNumber="246">2.14</td>
<td id="76B95D95FFBB0066418556D769AE8CF2" box="[1175,1217,391,413]" gridcol="4" gridrow="2" pageId="11" pageNumber="246">1.87</td>
<td id="76B95D95FFBB006641FC56D768778CF2" box="[1262,1304,391,413]" gridcol="5" gridrow="2" pageId="11" pageNumber="246">1.32</td>
<td id="76B95D95FFBB0066404056D768E68CF2" box="[1362,1417,391,413]" gridcol="6" gridrow="2" pageId="11" pageNumber="246">8.65</td>
</tr>
<tr id="356834E9FFBB0066462E56E668E68CA3" box="[828,1417,438,460]" gridrow="3" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E56E66E088CA3" box="[828,871,438,460]" gridcol="0" gridrow="3" pageId="11" pageNumber="246">II</th>
<td id="76B95D95FFBB0066468356E66ED48CA3" box="[913,955,438,460]" gridcol="1" gridrow="3" pageId="11" pageNumber="246">2.13</td>
<td id="76B95D95FFBB006646FA56E6697D8CA3" box="[1000,1042,438,460]" gridcol="2" gridrow="3" pageId="11" pageNumber="246">1.06</td>
<td id="76B95D95FFBB0066415256E669058CA3" box="[1088,1130,438,460]" gridcol="3" gridrow="3" pageId="11" pageNumber="246">1.87</td>
<td id="76B95D95FFBB0066418556E669AE8CA3" box="[1175,1217,438,460]" gridcol="4" gridrow="3" pageId="11" pageNumber="246">1.81</td>
<td id="76B95D95FFBB006641FC56E668778CA3" box="[1262,1304,438,460]" gridcol="5" gridrow="3" pageId="11" pageNumber="246">1.20</td>
<td id="76B95D95FFBB0066404056E668E68CA3" box="[1362,1417,438,460]" gridcol="6" gridrow="3" pageId="11" pageNumber="246">8.07</td>
</tr>
<tr id="356834E9FFBB0066462E56B568E68C94" box="[828,1417,485,507]" gridrow="4" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E56B56E088C94" box="[828,871,485,507]" gridcol="0" gridrow="4" pageId="11" pageNumber="246">III</th>
<td id="76B95D95FFBB0066468356B56ED48C94" box="[913,955,485,507]" gridcol="1" gridrow="4" pageId="11" pageNumber="246">2.06</td>
<td id="76B95D95FFBB006646FA56B5697D8C94" box="[1000,1042,485,507]" gridcol="2" gridrow="4" pageId="11" pageNumber="246">0.87</td>
<td id="76B95D95FFBB0066415256B569058C94" box="[1088,1130,485,507]" gridcol="3" gridrow="4" pageId="11" pageNumber="246">1.55</td>
<td id="76B95D95FFBB0066418556B569AE8C94" box="[1175,1217,485,507]" gridcol="4" gridrow="4" pageId="11" pageNumber="246">2.06</td>
<td id="76B95D95FFBB006641FC56B568778C94" box="[1262,1304,485,507]" gridcol="5" gridrow="4" pageId="11" pageNumber="246">1.09</td>
<td id="76B95D95FFBB0066404056B568E68C94" box="[1362,1417,485,507]" gridcol="6" gridrow="4" pageId="11" pageNumber="246">7.63</td>
</tr>
<tr id="356834E9FFBB0066462E554468E68F45" box="[828,1417,532,554]" gridrow="5" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E55446E088F45" box="[828,871,532,554]" gridcol="0" gridrow="5" pageId="11" pageNumber="246">IV</th>
<td id="76B95D95FFBB0066468355446ED48F45" box="[913,955,532,554]" gridcol="1" gridrow="5" pageId="11" pageNumber="246">2.96</td>
<td id="76B95D95FFBB006646FA5544697D8F45" box="[1000,1042,532,554]" gridcol="2" gridrow="5" pageId="11" pageNumber="246">0.99</td>
<td id="76B95D95FFBB00664152554469058F45" box="[1088,1130,532,554]" gridcol="3" gridrow="5" pageId="11" pageNumber="246">2.49</td>
<td id="76B95D95FFBB00664185554469AE8F45" box="[1175,1217,532,554]" gridcol="4" gridrow="5" pageId="11" pageNumber="246">3.82</td>
<td id="76B95D95FFBB006641FC554468778F45" box="[1262,1304,532,554]" gridcol="5" gridrow="5" pageId="11" pageNumber="246">1.60</td>
<td id="76B95D95FFBB00664040554468E68F45" box="[1362,1417,532,554]" gridcol="6" gridrow="5" pageId="11" pageNumber="246">11.86</td>
</tr>
</table>
</paragraph>
<caption id="DF276623FFBBFF92456355286FC08FE2" box="[113,687,632,654]" pageId="11" pageNumber="246" startId="11.[113,168,632,654]" targetType="table">
<paragraph id="8BE736ABFFBBFF92456355286FC08FE2" blockId="11.[113,687,632,654]" box="[113,687,632,654]" pageId="11" pageNumber="246">
Table 8. Leg spination for the male of
<taxonomicName id="4C584D28FFBBFF9244E155286FC08FE2" box="[499,687,632,653]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="daliensis">Wadicosa daliensis</taxonomicName>
</paragraph>
</caption>
<caption id="DF276623FFBBFF92463F552868E68FE2" box="[813,1417,632,654]" pageId="11" pageNumber="246" startId="11.[813,868,632,654]" targetType="table">
<paragraph id="8BE736ABFFBBFF92463F552868E68FE2" blockId="11.[813,1417,632,654]" box="[813,1417,632,654]" pageId="11" pageNumber="246">
Table 10. Leg spination for the female of
<taxonomicName id="4C584D28FFBBFF9241DF552868E68FE2" box="[1229,1417,632,653]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="246" phylum="Arthropoda" rank="species" species="daliensis">Wadicosa daliensis</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BE736ABFFBBFF9245CB55906F8C8EF7" pageId="11" pageNumber="246">
<table id="F958C40BFFBB0066456B55EF6F8C8EF6" box="[121,739,703,921]" gridcols="5" gridrows="5" pageId="11" pageNumber="246">
<tr id="356834E9FFBB0066456B55EF6F8C8FBA" box="[121,739,703,725]" gridrow="0" pageId="11" pageNumber="246" rowspan-0="1">
<th id="76B95D95FFBB006645BC55EF6C728FBA" box="[174,285,703,725]" gridcol="1" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9245CB55906D9D8FBA" bold="true" box="[217,242,704,725]" pageId="11" pageNumber="246">Fe</emphasis>
</th>
<th id="76B95D95FFBB0066442A55EF6CE48FBA" box="[312,395,703,725]" gridcol="2" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92444655906C008FBA" bold="true" box="[340,367,704,725]" pageId="11" pageNumber="246">Pa</emphasis>
</th>
<th id="76B95D95FFBB006644B055EF6F518FBA" box="[418,574,703,725]" gridcol="3" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9244F755EF6C938FBA" bold="true" box="[485,508,703,725]" pageId="11" pageNumber="246">Ti</emphasis>
</th>
<th id="76B95D95FFBB0066474155EF6F8C8FBA" box="[595,739,703,725]" gridcol="4" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92479E55906FC48FBA" bold="true" box="[652,683,704,725]" pageId="11" pageNumber="246">Mt</emphasis>
</th>
</tr>
<tr id="356834E9FFBB0066456B55A66F8C8E63" box="[121,739,758,780]" gridrow="1" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066456B55A66DFD8E63" box="[121,146,758,780]" gridcol="0" gridrow="1" pageId="11" pageNumber="246">I</th>
<td id="76B95D95FFBB006645BC55A66C728E63" box="[174,285,758,780]" gridcol="1" gridrow="1" pageId="11" pageNumber="246">3d 2p 3r</td>
<td id="76B95D95FFBB0066442A55A66CE48E63" box="[312,395,758,780]" gridcol="2" gridrow="1" pageId="11" pageNumber="246">1d 1p 1r</td>
<td id="76B95D95FFBB006644B055A66F518E63" box="[418,574,758,780]" gridcol="3" gridrow="1" pageId="11" pageNumber="246">2p 2r 2-2v</td>
<td id="76B95D95FFBB0066474155A66F8C8E63" box="[595,739,758,780]" gridcol="4" gridrow="1" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066456B54756F8C8E54" box="[121,739,805,827]" gridrow="2" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066456B54756DFD8E54" box="[121,146,805,827]" gridcol="0" gridrow="2" pageId="11" pageNumber="246">II</th>
<td id="76B95D95FFBB006645BC54756C728E54" box="[174,285,805,827]" gridcol="1" gridrow="2" pageId="11" pageNumber="246">3d 2p 3r</td>
<td id="76B95D95FFBB0066442A54756CE48E54" box="[312,395,805,827]" gridcol="2" gridrow="2" pageId="11" pageNumber="246">2d 1p 1r</td>
<td id="76B95D95FFBB006644B054756F518E54" box="[418,574,805,827]" gridcol="3" gridrow="2" pageId="11" pageNumber="246">2p 2r 2-2v</td>
<td id="76B95D95FFBB0066474154756F8C8E54" box="[595,739,805,827]" gridcol="4" gridrow="2" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066456B54046F8C8E05" box="[121,739,852,874]" gridrow="3" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066456B54046DFD8E05" box="[121,146,852,874]" gridcol="0" gridrow="3" pageId="11" pageNumber="246">III</th>
<td id="76B95D95FFBB006645BC54046C728E05" box="[174,285,852,874]" gridcol="1" gridrow="3" pageId="11" pageNumber="246">3d 2p 2(3)r</td>
<td id="76B95D95FFBB0066442A54046CE48E05" box="[312,395,852,874]" gridcol="2" gridrow="3" pageId="11" pageNumber="246">2d 1p 1r</td>
<td id="76B95D95FFBB006644B054046F518E05" box="[418,574,852,874]" gridcol="3" gridrow="3" pageId="11" pageNumber="246">2d 2p 2r 2-2v</td>
<td id="76B95D95FFBB0066474154046F8C8E05" box="[595,739,852,874]" gridcol="4" gridrow="3" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066456B54D36F8C8EF6" box="[121,739,899,921]" gridrow="4" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066456B54D36DFD8EF6" box="[121,146,899,921]" gridcol="0" gridrow="4" pageId="11" pageNumber="246">IV</th>
<td id="76B95D95FFBB006645BC54D36C728EF6" box="[174,285,899,921]" gridcol="1" gridrow="4" pageId="11" pageNumber="246">3d 2p 1r</td>
<td id="76B95D95FFBB0066442A54D36CE48EF6" box="[312,395,899,921]" gridcol="2" gridrow="4" pageId="11" pageNumber="246">2d 1p 1r</td>
<td id="76B95D95FFBB006644B054D36F518EF6" box="[418,574,899,921]" gridcol="3" gridrow="4" pageId="11" pageNumber="246">2d 2p 2r 2-1-1v</td>
<td id="76B95D95FFBB0066474154D36F8C8EF6" box="[595,739,899,921]" gridcol="4" gridrow="4" pageId="11" pageNumber="246">2p 2r 1-1-1-1v</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFBBFF92468C559068F88EF7" pageId="11" pageNumber="246">
<table id="F958C40BFFBB0066462E559068F88EF6" box="[828,1431,704,921]" gridcols="5" gridrows="5" pageId="11" pageNumber="246">
<tr id="356834E9FFBB0066462E559068F88FB9" box="[828,1431,704,726]" gridrow="0" pageId="11" pageNumber="246" rowspan-0="1">
<th id="76B95D95FFBB0066469355906EBA8FB9" box="[897,981,704,726]" gridcol="1" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF92468C55906ED88FBA" bold="true" box="[926,951,704,725]" pageId="11" pageNumber="246">Fe</emphasis>
</th>
<th id="76B95D95FFBB0066411A559069338FB9" box="[1032,1116,704,726]" gridcol="2" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9241375590692F8FBA" bold="true" box="[1061,1088,704,725]" pageId="11" pageNumber="246">Pa</emphasis>
</th>
<th id="76B95D95FFBB00664197559068638FB9" box="[1157,1292,704,726]" gridcol="3" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9241AF559069BB8FB9" bold="true" box="[1213,1236,704,726]" pageId="11" pageNumber="246">Ti</emphasis>
</th>
<th id="76B95D95FFBB0066403D559068F88FB9" box="[1327,1431,704,726]" gridcol="4" gridrow="0" pageId="11" pageNumber="246">
<emphasis id="B92CEAB9FFBBFF9240465590681C8FBA" bold="true" box="[1364,1395,704,725]" pageId="11" pageNumber="246">Mt</emphasis>
</th>
</tr>
<tr id="356834E9FFBB0066462E55A668F88E63" box="[828,1431,758,780]" gridrow="1" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E55A66E3A8E63" box="[828,853,758,780]" gridcol="0" gridrow="1" pageId="11" pageNumber="246">I</th>
<td id="76B95D95FFBB0066469355A66EBA8E63" box="[897,981,758,780]" gridcol="1" gridrow="1" pageId="11" pageNumber="246">3d 2p 3r</td>
<td id="76B95D95FFBB0066411A55A669338E63" box="[1032,1116,758,780]" gridcol="2" gridrow="1" pageId="11" pageNumber="246">1p 1r</td>
<td id="76B95D95FFBB0066419755A668638E63" box="[1157,1292,758,780]" gridcol="3" gridrow="1" pageId="11" pageNumber="246">2p 1r 2-2v</td>
<td id="76B95D95FFBB0066403D55A668F88E63" box="[1327,1431,758,780]" gridcol="4" gridrow="1" pageId="11" pageNumber="246">1p 1r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066462E547568F88E54" box="[828,1431,805,827]" gridrow="2" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E54756E3A8E54" box="[828,853,805,827]" gridcol="0" gridrow="2" pageId="11" pageNumber="246">II</th>
<td id="76B95D95FFBB0066469354756EBA8E54" box="[897,981,805,827]" gridcol="1" gridrow="2" pageId="11" pageNumber="246">3d 2p 3r</td>
<td id="76B95D95FFBB0066411A547569338E54" box="[1032,1116,805,827]" gridcol="2" gridrow="2" pageId="11" pageNumber="246">1p 1r</td>
<td id="76B95D95FFBB00664197547568638E54" box="[1157,1292,805,827]" gridcol="3" gridrow="2" pageId="11" pageNumber="246">2p 2r 2-2v</td>
<td id="76B95D95FFBB0066403D547568F88E54" box="[1327,1431,805,827]" gridcol="4" gridrow="2" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066462E540468F88E05" box="[828,1431,852,874]" gridrow="3" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E54046E3A8E05" box="[828,853,852,874]" gridcol="0" gridrow="3" pageId="11" pageNumber="246">III</th>
<td id="76B95D95FFBB0066469354046EBA8E05" box="[897,981,852,874]" gridcol="1" gridrow="3" pageId="11" pageNumber="246">3d 3p 3r</td>
<td id="76B95D95FFBB0066411A540469338E05" box="[1032,1116,852,874]" gridcol="2" gridrow="3" pageId="11" pageNumber="246">2d 1p 1r</td>
<td id="76B95D95FFBB00664197540468638E05" box="[1157,1292,852,874]" gridcol="3" gridrow="3" pageId="11" pageNumber="246">2d 2p 2r 2-2v</td>
<td id="76B95D95FFBB0066403D540468F88E05" box="[1327,1431,852,874]" gridcol="4" gridrow="3" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
<tr id="356834E9FFBB0066462E54D368F88EF6" box="[828,1431,899,921]" gridrow="4" pageId="11" pageNumber="246">
<th id="76B95D95FFBB0066462E54D36E3A8EF6" box="[828,853,899,921]" gridcol="0" gridrow="4" pageId="11" pageNumber="246">IV</th>
<td id="76B95D95FFBB0066469354D36EBA8EF6" box="[897,981,899,921]" gridcol="1" gridrow="4" pageId="11" pageNumber="246">3d 2p 1r</td>
<td id="76B95D95FFBB0066411A54D369338EF6" box="[1032,1116,899,921]" gridcol="2" gridrow="4" pageId="11" pageNumber="246">2d 1p 1r</td>
<td id="76B95D95FFBB0066419754D368638EF6" box="[1157,1292,899,921]" gridcol="3" gridrow="4" pageId="11" pageNumber="246">2d 2p 2r 2-2v</td>
<td id="76B95D95FFBB0066403D54D368F88EF6" box="[1327,1431,899,921]" gridcol="4" gridrow="4" pageId="11" pageNumber="246">2p 2r 2-2v</td>
</tr>
</table>
</paragraph>
<paragraph id="8BE736ABFFBBFF92456353986CE38850" blockId="11.[113,752,1223,1343]" pageId="11" pageNumber="246">Eye sizes and interdistances: AME 0.14, ALE 0.11, PME 0.33, PLE 0.26; AMEAME 0.10, AMEALE 0.03, PME PME 0.33, PMEPLE 0.35, AMEPME 0.10, ALEPME 0.09; clypeus height 0.22.</paragraph>
<paragraph id="8BE736ABFFBBFF92456352376D9A8B10" blockId="11.[113,753,1383,1663]" pageId="11" pageNumber="246">
Palpal femur black with yellow distal part; patella and tibia yellow covered with white setae; cymbium black with white setae proximally. For palp and legs measurements see
<tableCitation id="C6DA0310FFBBFF92458C52976D9A88B0" box="[158,245,1479,1503]" captionStart="Table 7" captionStartId="11.[113,168,189,211]" captionText="Table 7. Length of palp and leg segments for the male of Wadicosa daliensis" pageId="11" pageNumber="246">Table 7</tableCitation>
. Coxae III dark grey, IIIIV yellow; femora I dark brown dorsally and laterally, brown ventrally, IIIV light brown with grey annulation; patellae IIV light brown; tibiae and metatarsi IIV light brown with poorly visible greyish annulation; tarsi IIV light brown. For leg spination see
<tableCitation id="C6DA0310FFBBFF92458E51376D9E8B10" box="[156,241,1639,1663]" captionStart="Table 8" captionStartId="11.[113,168,632,654]" captionText="Table 8. Leg spination for the male of Wadicosa daliensis" pageId="11" pageNumber="246">Table 8</tableCitation>
.
</paragraph>
<paragraph id="8BE736ABFFBBFF92456351F76F9E8A30" blockId="11.[113,755,1703,1887]" pageId="11" pageNumber="246">
Dorsal part of the opisthosoma grey with dark brown cardiac mark and series of yellow spots. In some specimens these spots large, merging with each other (cf.
<figureCitation id="13632A2EFFBBFF92472451B76F8B8B90" box="[566,740,1767,1791]" captionStart="Figs" captionStartId="10.[136,180,1940,1962]" captionTargetBox="[146,1455,318,1909]" captionTargetId="figure-7@10.[136,1474,312,1924]" captionTargetPageId="10" captionText="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." figureDoi="http://doi.org/10.5281/zenodo.14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="11" pageNumber="246">Figs. 32 and 34</figureCitation>
). Lateral sides of the opisthosoma greyish with lots of yellow spots. Ventral part light brown with 4 sub-longitudinal lines consisting of tiny dark brown spots. Spinnerets light brown.
</paragraph>
<paragraph id="8BE736ABFFBBFF92456350D768C38990" blockId="11.[113,753,1927,2111]" lastBlockId="11.[811,1452,999,1279]" pageId="11" pageNumber="246">
Palp as shown in
<figureCitation id="13632A2EFFBBFF92442450D76CD58AF0" box="[310,442,1927,1951]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="10.[136,180,1940,1962]" captionStartId-1="12.[136,180,1977,1999]" captionTargetBox-0="[146,1455,318,1909]" captionTargetBox-1="[177,1430,206,1961]" captionTargetId-0="figure-7@10.[136,1474,312,1924]" captionTargetId-1="figure-7@12.[174,1436,190,1962]" captionTargetPageId-0="10" captionTargetPageId-1="12" captionText-0="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." captionText-1="Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of Wadicosa daliensis. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis." figureDoi-0="http://doi.org/10.5281/zenodo.14683154" figureDoi-1="http://doi.org/10.5281/zenodo.14683156" httpUri-0="https://zenodo.org/record/14683154/files/figure.png" httpUri-1="https://zenodo.org/record/14683156/files/figure.png" pageId="11" pageNumber="246">Figs. 3748</figureCitation>
. Tibia 1.6 times longer than wide. Cymbium 1.5 times longer than bulb and 1.5 times longer than tibia. Tegulum (Te) with heavily sclerotised lanceolate posterior retrolateral process (PP), conical anterior process (AP); retrolaterally with serrated edge (Ed). Subtegulum (St) prolaterally positioned, partly hidden by large shell-like cuticular cover (CC). Tegular apophysis (TA) consists of two branches. Upper branch (UB) long, crescent-like in ventral view, strongly flattened in anterior view, with triangular extension (AE) medially. UB proximally covered with tiny spinulae barely visible in light microscope. Lower branch (LB) small, triangular. Conductor (Co) large, heavily sclerotised with hook-shaped tip in retrolateral view. Embolus (Em) long, forming large bend over tegular apophysis. Apical part of embolus widened, semi-transparent; embolic tip acute.
</paragraph>
<paragraph id="8BE736ABFFBBFF92463F527768C488F0" blockId="11.[811,1452,1319,1439]" pageId="11" pageNumber="246">
Female (
<figureCitation id="13632A2EFFBBFF92469A527769668850" box="[904,1033,1319,1343]" captionStart="Figs" captionStartId="10.[136,180,1940,1962]" captionTargetBox="[146,1455,318,1909]" captionTargetId="figure-7@10.[136,1474,312,1924]" captionTargetPageId="10" captionText="Figs. 3237. Male (3234), female (3536) and male palp (37) of Wadicosa daliensis. 32, 34, 35 dorsal; 33, 36 ventral; 37 prolateral. Scale: 1 mm (3236), 0.2 mm (37)." figureDoi="http://doi.org/10.5281/zenodo.14683154" httpUri="https://zenodo.org/record/14683154/files/figure.png" pageId="11" pageNumber="246">Figs. 3536</figureCitation>
). Total length 6.04. Carapace 2.88 long, 2.36 wide. Opisthosoma 3.13 long, 2.09 wide. Colouration. Carapace and opisthosoma like in male but somewhat lighter, white setae less visible. Clypeus and chelicerae light brown.
</paragraph>
<paragraph id="8BE736ABFFBBFF92463F529869288B50" blockId="11.[813,1452,1479,1599]" pageId="11" pageNumber="246">Eye sizes and interdistances: AME 0.16, ALE 0.11, PME 0.33, PLE 0.28; AMEAME 0.10, AMEALE 0.04, PME PME 0.31, PMEPLE 0.39, AMEPME 0.12, ALEPME 0.11; clypeus height 0.26.</paragraph>
<paragraph id="8BE736ABFFBBFF92463F513768118A70" blockId="11.[813,1452,1639,1823]" pageId="11" pageNumber="246">
All segments of palp light brown. For palp and legs measurements see
<tableCitation id="C6DA0310FFBBFF9246E851D7693E8BF0" box="[1018,1105,1671,1695]" captionStart="Table 9" captionStartId="11.[813,871,189,211]" captionText="Table 9. Length of palp and leg segments for the female of Wadicosa daliensis" pageId="11" pageNumber="246">Table 9</tableCitation>
. Coxae IIV yellowish. femora IIV dorsally and laterally light brown with grey annulation, ventrally yellow; patellae IIV light brown; tibiae and metatarsi IIV light brown with poorly visible annulation; tarsi IIV light brown. For leg spination see
<tableCitation id="C6DA0310FFBBFF924005505768148A70" box="[1303,1403,1799,1823]" captionStart="Table 10" captionStartId="11.[813,868,632,654]" captionText="Table 10. Leg spination for the female of Wadicosa daliensis" pageId="11" pageNumber="246">Table 10</tableCitation>
.
</paragraph>
<paragraph id="8BE736ABFFBBFF92463F501768618A10" blockId="11.[813,1452,1863,1919]" pageId="11" pageNumber="246">Lateral sides of the opisthosoma yellow with tiny grey spots. Ventral part yellow. Spinnerets light brown.</paragraph>
<paragraph id="8BE736ABFFBBFF92463F50F769828550" blockId="11.[813,1452,1959,2111]" pageId="11" pageNumber="246">
Epigyne as shown in
<figureCitation id="13632A2EFFBBFF92410450F769F78AD0" box="[1046,1176,1959,1983]" captionStart="Figs" captionStartId="13.[113,157,1230,1252]" captionTargetBox="[113,1451,274,1205]" captionTargetId="figure-10@13.[113,1451,274,1215]" captionTargetPageId="13" captionText="Figs. 4951. Epigyne of Wadicosa daliensis. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm. Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem." figureDoi="http://doi.org/10.5281/zenodo.14683158" httpUri="https://zenodo.org/record/14683158/files/figure.png" pageId="11" pageNumber="246">Figs. 4951</figureCitation>
. Fovea (Fo) ca. 2.2 times wider than long. Receptacles (Re) long, turned U-shaped. Septum (Se) triangular with wide base (SB) and short, poorly visible stem (SS). Foveolae (Fv; pockets) oval, ca. 2 times longer than wide, with large hoods (Ho).
</paragraph>
<caption id="DF276623FFBCFF95459A50E96923852C" ID-DOI="http://doi.org/10.5281/zenodo.14683156" ID-Zenodo-Dep="14683156" httpUri="https://zenodo.org/record/14683156/files/figure.png" pageId="12" pageNumber="247" startId="12.[136,180,1977,1999]" targetBox="[177,1430,206,1961]" targetPageId="12" targetType="figure">
<paragraph id="8BE736ABFFBCFF95459A50E96923852C" blockId="12.[136,1474,1977,2115]" pageId="12" pageNumber="247">
Figs. 3848. Male palp (3840), bulb (4144) and embolic division (4548) of
<taxonomicName id="4C584D28FFBCFF95466D50E969568AA1" box="[895,1081,1977,1998]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="247" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBCFF95466D50E969568AA1" box="[895,1081,1977,1998]" italics="true" pageId="12" pageNumber="247">Wadicosa daliensis</emphasis>
</taxonomicName>
. 38, 41 prolateral; 39, 42, 45 ventral; 40, 43, 46 retrolateral; 44, 47 anterior; 48 prolatero-anterior. Scale: 0.2 mm (3844), 0.1 mm (4548). Abbreviations: AE extension of tegular apophysis, AP anterior retrolateral process of tegulum, CC cuticular cover of bulb, Co conductor, Ed retrolateral edge of tegulum, Em embolus, LB lower branch of tegular apophysis, PP posterior retrolateral process of tegulum, SD sperm duct, St subtegulum, TA tegular apophysis, Te tegulum, UB upper branch of tegular apophysis.
</paragraph>
</caption>
<caption id="DF276623FFBDFF944563539E6F728872" ID-DOI="http://doi.org/10.5281/zenodo.14683158" ID-Zenodo-Dep="14683158" httpUri="https://zenodo.org/record/14683158/files/figure.png" pageId="13" pageNumber="248" startId="13.[113,157,1230,1252]" targetBox="[113,1451,274,1205]" targetPageId="13" targetType="figure">
<paragraph id="8BE736ABFFBDFF944563539E6993898B" blockId="13.[113,1450,1230,1310]" box="[113,1276,1230,1252]" pageId="13" pageNumber="248">
Figs. 4951. Epigyne of
<taxonomicName id="4C584D28FFBDFF944477539E6F4F898C" box="[357,544,1230,1251]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBDFF944477539E6F4F898C" box="[357,544,1230,1251]" italics="true" pageId="13" pageNumber="248">Wadicosa daliensis</emphasis>
</taxonomicName>
. 49 ventral, intact; 50 ventral, macerated; 51 dorsal. Scale: 0.2 mm.
</paragraph>
<paragraph id="8BE736ABFFBDFF94456353BB6F728872" blockId="13.[113,1450,1230,1310]" pageId="13" pageNumber="248">Abbreviations: FD fertilisation duct, Fv foveola (pocket), Fo fovea, Ho hood, Re receptacle, SB septal base, Se septum (outlined by dashed line), SS septal stem.</paragraph>
</caption>
<caption id="DF276623FFBDFF94456050EC68088A81" ID-DOI="http://doi.org/10.5281/zenodo.14683160" ID-Zenodo-Dep="14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="13" pageNumber="248" startId="13.[114,159,1980,2002]" targetBox="[113,1451,1365,1964]" targetPageId="13" targetType="figure">
<paragraph id="8BE736ABFFBDFF94456050EC68088A81" blockId="13.[113,1451,1980,2031]" pageId="13" pageNumber="248">
Figs. 5253. Collecting localities of (1)
<taxonomicName id="4C584D28FFBDFF94471C50EC6EB38ABE" authority="Omelko &amp; Marusik, 2022" authorityName="Omelko &amp; Marusik" authorityYear="2022" box="[526,988,1980,2002]" class="Arachnida" family="Lycosidae" genus="Arctosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="delaportei">
<emphasis id="B92CEAB9FFBDFF94471C50EC6FA68ABE" box="[526,713,1980,2001]" italics="true" pageId="13" pageNumber="248">Arctosa delaportei</emphasis>
<bibRefCitation id="EFC94B5AFFBDFF9447C150EC6EB38ABE" author="Omelko MM &amp; Marusik YM" box="[723,988,1980,2002]" pageId="13" pageNumber="248" pagination="397 - 406" refId="ref10251" refString="Omelko MM &amp; Marusik YM (2022) First data on Arctosa wolf spiders (Araneae: Lycosidae) from Laos with description of a new species. Raffles Bulletin of Zoology, 70: 397 - 406." type="journal article" year="2022">Omelko &amp; Marusik, 2022</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C584D28FFBDFF9446F850EC68E88ABD" authority="Yin &amp; Chen, 1995" authorityName="Yin &amp; Chen" authorityYear="1995" box="[1002,1415,1980,2002]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="bannaensis">
<emphasis id="B92CEAB9FFBDFF9446F850EC69D08ABE" box="[1002,1215,1980,2001]" italics="true" pageId="13" pageNumber="248">Trochosa bannaensis</emphasis>
Yin &amp; Chen, 1995
</taxonomicName>
,
<taxonomicName id="4C584D28FFBDFF94408550ED6CEC8A80" authority="Schenkel, 1963" authorityName="Schenkel" authorityYear="1963" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFBDFF94408550ED6D898A81" italics="true" pageId="13" pageNumber="248">T. ruricoloides</emphasis>
<bibRefCitation id="EFC94B5AFFBDFF9445FC50896CEC8A80" author="Schenkel E" box="[238,387,2009,2031]" pageId="13" pageNumber="248" pagination="1 - 481" refId="ref10677" refString="Schenkel E (1963) Ostasiatische Spinnen aus dem Museum d'Histoire naturelle de Paris. Memoires du Museum National d'Histoire Naturelle de Paris (A, Zool.), 25: 1 - 481." type="journal article" year="1963">Schenkel, 1963</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C584D28FFBDFF94448350896E398A81" authority="Yin, Peng &amp; Zhang, 1997" authorityName="Yin, Peng &amp; Zhang" authorityYear="1997" box="[401,854,2009,2031]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBDFF94448350896F228A81" box="[401,589,2009,2030]" italics="true" pageId="13" pageNumber="248">Wadicosa daliensis</emphasis>
<bibRefCitation id="EFC94B5AFFBDFF94474750896E398A81" author="Yin CM &amp; Peng XJ &amp; Zhang YJ" box="[597,854,2009,2031]" pageId="13" pageNumber="248" pagination="99 - 101" refId="ref11420" refString="Yin CM, Peng XJ &amp; Zhang YJ (1997 a) One new species of the genus Wadicosa (Araneae, Lycosidae) from China. Acta Arachnologica Sinica, 6: 99 - 101." type="journal article" year="1997">Yin, Peng &amp; Zhang, 1997</bibRefCitation>
</taxonomicName>
, (2)
<taxonomicName id="4C584D28FFBDFF944695508969558A81" authorityName="Omelko" authorityYear="2024" box="[903,1082,2009,2030]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="zhishengi" status="sp. nov.">
<emphasis id="B92CEAB9FFBDFF944695508969558A81" box="[903,1082,2009,2030]" italics="true" pageId="13" pageNumber="248">Pardosa zhishengi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21F57C2FFBDFF944155508969AD8A80" box="[1095,1218,2009,2031]" pageId="13" pageNumber="248" rank="species">new species</taxonomicNameLabel>
.,
<taxonomicName id="4C584D28FFBDFF9441C0508A680E8A81" authorityName="Schenkel" authorityYear="1963" box="[1234,1377,2009,2031]" class="Arachnida" family="Lycosidae" genus="Trochosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="248" phylum="Arthropoda" rank="species" species="ruricoloides">
<emphasis id="B92CEAB9FFBDFF9441C0508A680E8A81" box="[1234,1377,2009,2031]" italics="true" pageId="13" pageNumber="248">T. ruricoloides</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BE736ABFFBEFF97459A57ED6FA98E5A" blockId="14.[135,775,189,821]" pageId="14" pageNumber="249">
<emphasis id="B92CEAB9FFBEFF97459A57ED6DA08DBA" bold="true" box="[136,207,189,213]" pageId="14" pageNumber="249">Notes.</emphasis>
As mentioned above, one of the species closest to
<taxonomicName id="4C584D28FFBEFF97459A578D6C0E8D9A" authorityName="Yin, Peng &amp; Zhang" authorityYear="1997" box="[136,353,221,245]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBEFF97459A578D6C0E8D9A" box="[136,353,221,245]" italics="true" pageId="14" pageNumber="249">Wadicosa daliensis</emphasis>
</taxonomicName>
is
<taxonomicName id="4C584D28FFBEFF97449A578E6F598D9A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[392,566,221,245]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF97449A578E6F598D9A" box="[392,566,221,245]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
. This species was described from southern
<collectingCountry id="F34F763BFFBEFF9744BB57AD6C878C7B" box="[425,488,253,276]" name="Japan" pageId="14" pageNumber="249">Japan</collectingCountry>
(Nansei islands;
<bibRefCitation id="EFC94B5AFFBEFF9747A757AD6DB98C5A" author="Tanaka H" pageId="14" pageNumber="249" pagination="51 - 87" refId="ref10801" refString="Tanaka H (1985) Descriptions of new species of the Lycosidae (Araneae) from Japan. Acta Arachnologica, 33: 51 - 87." type="journal article" year="1985">Tanaka (1985))</bibRefCitation>
and later found in
<collectingCountry id="F34F763BFFBEFF974486564D6CBC8C5A" box="[404,467,285,309]" name="China" pageId="14" pageNumber="249">China</collectingCountry>
(
<collectingRegion id="499CF849FFBEFF9744CD564D6F408C5A" box="[479,559,285,309]" country="China" name="Hainan" pageId="14" pageNumber="249">Hainan</collectingRegion>
;
<bibRefCitation id="EFC94B5AFFBEFF97472A564D6E6B8C5A" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[568,772,285,309]" pageId="14" pageNumber="249" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021))</bibRefCitation>
. The original black and white drawings of the copulatory organs of
<taxonomicName id="4C584D28FFBEFF9745EF560E6CC48C1A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[253,427,349,373]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF9745EF560E6CC48C1A" box="[253,427,349,373]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
are highly schematic, and the colour photographs are of small size (
<bibRefCitation id="EFC94B5AFFBEFF97471F562D6FC88CFA" author="Chikuni Y" box="[525,679,381,405]" pageId="14" pageNumber="249" refId="ref9342" refString="Chikuni Y (1989) Pictorial encyclopedia of spiders in Japan. Kaisei-sha Publishing Co., Tokyo, 310 pp." type="book" year="1989">Chikuni, 1989</bibRefCitation>
), making a careful comparison of the photos of my specimens of
<taxonomicName id="4C584D28FFBEFF9747F856CE6D868CBA" authorityName="Yin, Peng &amp; Zhang" authorityYear="1997" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="daliensis">
<emphasis id="B92CEAB9FFBEFF9747F856CE6D868CBA" italics="true" pageId="14" pageNumber="249">W. daliensis</emphasis>
</taxonomicName>
with the images of
<taxonomicName id="4C584D28FFBEFF9744C056EE6F108CBA" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[466,639,445,469]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF9744C056EE6F108CBA" box="[466,639,445,469]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F34F763BFFBEFF9747DB56ED6E688CBB" box="[713,775,445,468]" name="Japan" pageId="14" pageNumber="249">Japan</collectingCountry>
impossible. For this reason, I only compare them with photos of
<taxonomicName id="4C584D28FFBEFF9745B556AE6C218F7A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[167,334,509,533]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF9745B556AE6C218F7A" box="[167,334,509,533]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F34F763BFFBEFF97448156AD6CB98F7A" box="[403,470,509,533]" name="China" pageId="14" pageNumber="249">China</collectingCountry>
. However, live photographs of
<taxonomicName id="4C584D28FFBEFF9745B6554E6C278F5A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[164,328,541,565]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF9745B6554E6C278F5A" box="[164,328,541,565]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
from
<collectingCountry id="F34F763BFFBEFF974495554D6CA88F5A" box="[391,455,541,565]" name="China" pageId="14" pageNumber="249">China</collectingCountry>
differ significantly from those from
<collectingCountry id="F34F763BFFBEFF9745D0556D6D908F3B" box="[194,255,573,596]" name="Japan" pageId="14" pageNumber="249">Japan</collectingCountry>
(cf. Figs. 2EF in
<bibRefCitation id="EFC94B5AFFBEFF9744D5556D6FE08F3A" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[455,655,573,597]" pageId="14" pageNumber="249" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021)</bibRefCitation>
and fig.
<quantity id="4CA09B4EFFBEFF9747FE556D6DF28F1A" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="14" pageNumber="249" unit="in" value="35.0">35 in</quantity>
<bibRefCitation id="EFC94B5AFFBEFF9745B0550D6C208F1A" author="Chikuni Y" box="[162,335,605,629]" pageId="14" pageNumber="249" refId="ref9342" refString="Chikuni Y (1989) Pictorial encyclopedia of spiders in Japan. Kaisei-sha Publishing Co., Tokyo, 310 pp." type="book" year="1989">Chikuni (1989))</bibRefCitation>
. Additionally, noticeable differences exist in the structure of the female epigyne, such as the varying shape of the fovea and foveolae (pockets) (cf. fig. 69G in
<bibRefCitation id="EFC94B5AFFBEFF97459A55ED6C358FBA" author="Wang LY &amp; Lu T &amp; Cai DC &amp; Barrion AT &amp; Heong KL &amp; Li SQ &amp; Zhang ZS" box="[136,346,701,725]" pageId="14" pageNumber="249" pagination="16 - 74" refId="ref11222" refString="Wang LY, Lu T, Cai DC, Barrion AT, Heong KL, Li SQ &amp; Zhang ZS (2021) Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae). Zoological Systematics, 46 (1): 16 - 74." type="journal article" year="2021">Wang et al. (2021)</bibRefCitation>
and fig.
<quantity id="4CA09B4EFFBEFF9744D355ED6C958FBA" box="[449,506,701,725]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="14" pageNumber="249" unit="in" value="35.0">35 in</quantity>
<bibRefCitation id="EFC94B5AFFBEFF97471655ED6FD88FBA" author="Chikuni Y" box="[516,695,701,725]" pageId="14" pageNumber="249" refId="ref9342" refString="Chikuni Y (1989) Pictorial encyclopedia of spiders in Japan. Kaisei-sha Publishing Co., Tokyo, 310 pp." type="book" year="1989">Chikuni (1989))</bibRefCitation>
. In my opinion, the spiders identified from
<collectingCountry id="F34F763BFFBEFF974713558D6F2E8F9A" box="[513,577,733,757]" name="China" pageId="14" pageNumber="249">China</collectingCountry>
as
<taxonomicName id="4C584D28FFBEFF974770558E6E698F9A" baseAuthorityName="Tanaka" baseAuthorityYear="1985" box="[610,774,733,757]" class="Arachnida" family="Lycosidae" genus="Wadicosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="249" phylum="Arthropoda" rank="species" species="okinawensis">
<emphasis id="B92CEAB9FFBEFF974770558E6E698F9A" box="[610,774,733,757]" italics="true" pageId="14" pageNumber="249">W. okinawensis</emphasis>
</taxonomicName>
are likely not conspecific with those from
<collectingCountry id="F34F763BFFBEFF97477055AD6FF08E7B" box="[610,671,765,788]" name="Japan" pageId="14" pageNumber="249">Japan</collectingCountry>
and may possibly be described as a new species in the future.
</paragraph>
</subSubSection>
<subSubSection id="C3426520FFBEFF97459A540D6F058EFA" pageId="14" pageNumber="249" type="distribution">
<paragraph id="8BE736ABFFBEFF97459A540D6F058EFA" blockId="14.[136,773,861,917]" pageId="14" pageNumber="249">
<emphasis id="B92CEAB9FFBEFF97459A540D6C488E1A" bold="true" box="[136,295,861,885]" pageId="14" pageNumber="249">Distribution</emphasis>
.
<collectingCountry id="F34F763BFFBEFF97442F540D6CEB8E1A" box="[317,388,861,885]" name="China" pageId="14" pageNumber="249">China</collectingCountry>
(
<collectingRegion id="499CF849FFBEFF974488540D6F448E1A" box="[410,555,861,885]" country="China" name="Guangdong" pageId="14" pageNumber="249">Guangdong</collectingRegion>
,
<collectingRegion id="499CF849FFBEFF97472F540D6FF88E1A" box="[573,663,861,885]" country="China" name="Hainan" pageId="14" pageNumber="249">Hainan</collectingRegion>
,
<collectingRegion id="499CF849FFBEFF9747BB540D6E6A8E1B" box="[681,773,861,884]" country="China" name="Yunnan" pageId="14" pageNumber="249">Yunnan</collectingRegion>
Provinces),
<collectingCountry id="F34F763BFFBEFF974419542D6C508EFB" box="[267,319,893,916]" name="Laos" pageId="14" pageNumber="249">Laos</collectingCountry>
(new record;
<figureCitation id="13632A2EFFBEFF9744C8542D6F318EFA" box="[474,606,893,917]" captionStart="Figs" captionStartId="13.[114,159,1980,2002]" captionTargetBox="[113,1451,1365,1964]" captionTargetId="figure-66@13.[113,1451,1364,1964]" captionTargetPageId="13" captionText="Figs. 5253. Collecting localities of (1) Arctosa delaportei Omelko &amp; Marusik, 2022, Trochosa bannaensis Yin &amp; Chen, 1995, T. ruricoloides Schenkel, 1963, Wadicosa daliensis Yin, Peng &amp; Zhang, 1997, (2) Pardosa zhishengi, new species., T. ruricoloides." figureDoi="http://doi.org/10.5281/zenodo.14683160" httpUri="https://zenodo.org/record/14683160/files/figure.png" pageId="14" pageNumber="249">Figs. 5253</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,217 @@
<document id="9F20E98FB3A2CBFD606B131873943FE2" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C40CE27FE89C6D39FC8FD80" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Takedaplax Ng, Ahyong &amp; T., 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="348" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="347" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="556191188049F2CC91369108568C05DD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EFD0ABF69A3D4B00C6CC32BD5F6B1255">
<mods:title id="EEE6ABBB0C7FDFB8F62ACD6D4A5AE0F9">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="A3E1DC18157530FC1F271346D9E4D1D5" type="personal">
<mods:role id="C859042CCD44B08A26BD0AA601FB9C33">
<mods:roleTerm id="9E8209631805887E1AC30563EE6BA50E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8A775F6E5652383D4A07D9BCE2B7F877">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="14133F0577AD0C344EF6F0D8F6A261B7" type="personal">
<mods:role id="E819538D2BA2B52A062F98F41FD5220D">
<mods:roleTerm id="BEC7CDC6EFBF4C596DBEEBA6CE60123B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="050E9025A1BBE17694D882CC9F0D8F64">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="C4FF777D0E6C4865174C29184BC66FDA" type="personal">
<mods:role id="9136990FAF563C461B0EAD520F8060D9">
<mods:roleTerm id="2653AD3569FD1603698901B028697A5F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CBFC432AD5244B209A0DAED00538A8C">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C455D0E9BACA6E47B28814FAC892964D">text</mods:typeOfResource>
<mods:relatedItem id="EAA88B7542049F858AEB3D7803D2512E" type="host">
<mods:titleInfo id="5C05ABEEB7153A2CEAF1C6458D8DDA97">
<mods:title id="866883D7F4D160B943258CD62C2EE2DB">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="04F88AFACDAF7755DD5D953AA01076D6">
<mods:date id="21D48AAF2722F1B7E80C11D76ECE14F7">2024</mods:date>
<mods:detail id="0B34B0D61C780E121FAA13605FB55B0A" type="pubDate">
<mods:number id="6035FA056FAD4D627519868CBB2F900C">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="38C410AA3FAF71A7433C777DE48789DD" type="volume">
<mods:number id="F392F51E950D8ADE0119ECFBD25FF2AD">72</mods:number>
</mods:detail>
<mods:extent id="E3F880FAD71223D7E13C0FDA3FA7BE61" unit="page">
<mods:start id="712F1E1FA4B3F0227A45BD3A2C07B45D">335</mods:start>
<mods:end id="575DD83CDDE174BC67C1147567FB4E63">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="823D45D8E3270545CDC50BEC760ED180">journal article</mods:classification>
<mods:identifier id="6870A415329F17D1CCF09A0900007233" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="E5C4E6BA929F619D3FB18A1F7BC78942" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="51F56C3FEBBB67528D17A70561A25AF4" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="F8B97BE5CCED2B90F8797F9481BFB692" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C40CE27FE89C6D39FC8FD80" ID-DOI="http://doi.org/10.5281/zenodo.14682791" ID-Zenodo-Dep="14682791" LSID="urn:lsid:plazi:treatment:1D03E8542C40CE27FE89C6D39FC8FD80" httpUri="http://treatment.plazi.org/id/1D03E8542C40CE27FE89C6D39FC8FD80" lastPageId="13" lastPageNumber="348" pageId="12" pageNumber="347">
<subSubSection id="DDB00AC92C40CE26FE89C6D39C14F903" box="[323,587,1757,1781]" pageId="12" pageNumber="347" type="nomenclature">
<paragraph id="951559422C40CE26FE89C6D39C14F903" blockId="12.[323,587,1757,1781]" box="[323,587,1757,1781]" pageId="12" pageNumber="347">
<heading id="CE5DEE2E2C40CE26FE89C6D39C14F903" bold="true" box="[323,587,1757,1781]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="347" reason="2">
<emphasis id="A7DE85502C40CE26FE89C6D39C14F903" bold="true" box="[323,587,1757,1781]" pageId="12" pageNumber="347">
<taxonomicName id="52AA22C12C40CE26FE89C6D39F9BF903" ID-CoL="CL3BY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[323,452,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FE89C6D39F9BF903" bold="true" box="[323,452,1757,1781]" italics="true" pageId="12" pageNumber="347">Takedaplax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FE1EC6D39C14F903" box="[468,587,1757,1781]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FF42C7139F65F843" pageId="12" pageNumber="347" type="type_taxon">
<paragraph id="951559422C40CE26FF42C7139CACF8A3" blockId="12.[136,775,1821,1878]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7139F40F8C3" bold="true" box="[136,287,1821,1845]" pageId="12" pageNumber="347">
<typeStatus id="4A11E7E02C40CE26FF42C7139E9DF8C3" box="[136,194,1821,1845]" pageId="12" pageNumber="347">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C40CE26FEECC7139E9EF8A3" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984" authorityName="Takeda &amp; Kurata" authorityYear="1984" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FEECC7139C62F8C3" box="[294,573,1821,1845]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C40CE26FD8FC7139E9EF8A3" author="Takeda M &amp; Kurata Y" pageId="12" pageNumber="347" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
</taxonomicName>
, by present designation. Gender of genus feminine.
</paragraph>
<paragraph id="951559422C40CE26FF42C7739F65F843" blockId="12.[136,774,1917,1973]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7739F13F863" bold="true" box="[136,332,1917,1941]" pageId="12" pageNumber="347">Included species.</emphasis>
Only
<taxonomicName id="52AA22C12C40CE26FE5FC7739F69F843" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984" authorityName="Takeda &amp; Kurata" authorityYear="1984" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FE5FC7739CF1F863" box="[405,686,1917,1941]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C40CE26FD72C7739F69F843" author="Takeda M &amp; Kurata Y" pageId="12" pageNumber="347" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FF42C7D39B36FBA3" pageId="12" pageNumber="347" type="diagnosis">
<paragraph id="951559422C40CE26FF42C7D39B36FBA3" blockId="12.[136,775,2013,2069]" lastBlockId="12.[835,1475,189,1109]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C7D39EA2F803" bold="true" box="[136,253,2013,2037]" pageId="12" pageNumber="347">Diagnosis.</emphasis>
Carapace rectangular; epigastric, postorbital and hepatic areoles well developed with anterior margin cristate to subcristate; epigastric region gently convex, smooth, setose, not areolate; protogastric, cardiac, mesobranchial and cardiobranchial regions may be raised to form areole in larger specimens, barely or not visible in smaller specimens; intestinal region not prominently raised, no visible intestino-branchial areole; surface adjacent to posterior carapace margin almost flat, not forming ridge, no transverse channel visible; orbit clearly visible in dorsal view, supraorbital margin breaking general carapace outline, forming distinct concavity; eyes freely movable, ocular peduncle relatively long; lateral lobe of frontal margin low, indistinct; anterolateral margin with 4 visible teeth or spines (including external orbital tooth), last one often small; posterolateral margins sinuous, gently converging towards posterior carapace margin. Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges. Posterior margin of epistome entire, without lateral fissures, median part triangular, lateral parts distinctly concave. Third maxilliped ischium subquadrate, length about 1.3× width. Adult male chelipeds almost symmetrical; dorsal surface of carpus with prominent longitudinal areole, separated from swollen lateral fringe by groove; chela relatively slender with external surface covered with tubercles and granules. Ambulatory legs elongate, slender, P4 longest, about 2.0× carapace width; P25 merus without crest on extensor margin, distally unarmed, outer surface smooth, without medio-longitudinal swelling; P4 merus length about 4.3× height, distinctly longer than half carapace width. G1 strongly sinuous, tip recurved.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FC89C4739A44FAA3" pageId="12" pageNumber="347" type="etymology">
<paragraph id="951559422C40CE26FC89C4739A44FAA3" blockId="12.[835,1474,1149,1365]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FC89C4739D9AFB63" bold="true" box="[835,965,1149,1173]" pageId="12" pageNumber="347">Etymology.</emphasis>
The genus is named after Masatsune Takeda, whose work on the genus has been invaluable in the present revision, and we thank him for encouraging us to “finish the job” by revising the generic position of all species previously placed in
<taxonomicName id="52AA22C12C40CE26FBEDC4F39AC6FAE3" authorityName="Tesch" authorityYear="1918" box="[1063,1177,1277,1301]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FBEDC4F39AC6FAE3" box="[1063,1177,1277,1301]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
. The name is derived from his family name, in arbitrary combination with
<taxonomicName id="52AA22C12C40CE26FA83C5139BE4FAC3" authorityName="Tesch" authorityYear="1918" box="[1353,1467,1309,1333]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FA83C5139BE4FAC3" box="[1353,1467,1309,1333]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE27FC89C5739FC8FD80" lastPageId="13" lastPageNumber="348" pageId="12" pageNumber="347" type="discussion">
<paragraph id="951559422C40CE26FC89C5739A55F803" blockId="12.[835,1475,1405,2038]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FC89C5739DEDFA63" bold="true" box="[835,946,1405,1429]" pageId="12" pageNumber="347">Remarks.</emphasis>
The six prominent areolets on the anterior half of the carapace of
<taxonomicName id="52AA22C12C40CE26FBD6C5939B61FA43" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[1052,1342,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FBD6C5939B61FA43" box="[1052,1342,1437,1461]" italics="true" pageId="12" pageNumber="347">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is the only character that links the species to
<taxonomicName id="52AA22C12C40CE26FB73C5B39B74FA23" authorityName="Tesch" authorityYear="1918" box="[1209,1323,1469,1493]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FB73C5B39B74FA23" box="[1209,1323,1469,1493]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
, although the position of the areolets are different. Compared to
<taxonomicName id="52AA22C12C40CE26FA85C5D39B9FFA03" authorityName="Tesch" authorityYear="1918" box="[1359,1472,1501,1525]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C40CE26FA85C5D39B9FFA03" box="[1359,1472,1501,1525]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C40CE26FC89C5F09D8EF9E3" box="[835,977,1533,1557]" pageId="12" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
, however,
<taxonomicName id="52AA22C12C40CE26FB84C5F09B51F9E3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[1102,1294,1533,1557]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C40CE26FB84C5F09B51F9E3" box="[1102,1294,1533,1557]" italics="true" pageId="12" pageNumber="347">L. sextuberculata</emphasis>
</taxonomicName>
has a distinctly wider carapace which appears rectangular in form, in sharp contrast to the more quadrate form. Most markedly, it differs in having the posterior margin of the epistome entire, without any lateral fissures (
<figureCitation id="0D9145C72C40CE26FBFAC6739ACBF963" box="[1072,1172,1661,1685]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="12" pageNumber="347">Figs. 8E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FB6EC6739A96F963" box="[1188,1225,1661,1685]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="12" pageNumber="347">9D</figureCitation>
). In
<taxonomicName id="52AA22C12C40CE26FAC9C6739B25F963" authorityName="Tesch" authorityYear="1918" box="[1283,1402,1661,1685]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C40CE26FAC9C6739B25F963" box="[1283,1402,1661,1685]" italics="true" pageId="12" pageNumber="347">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C40CE26FA4EC6709DD7F943" pageId="12" pageNumber="347" sensu="stricto">sensu stricto</taxonomicNameLabel>
, as well as
<taxonomicName id="52AA22C12C40CE26FBC4C6939AC0F943" authorityName="Deb" authorityYear="1989" box="[1038,1183,1693,1717]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FBC4C6939AC0F943" box="[1038,1183,1693,1717]" italics="true" pageId="12" pageNumber="347">Myopilumnus</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C40CE26FB65C6939B59F943" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[1199,1286,1693,1717]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FB65C6939B59F943" box="[1199,1286,1693,1717]" italics="true" pageId="12" pageNumber="347">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FADDC6909BD1F940" box="[1303,1422,1694,1718]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
, and
<taxonomicName id="52AA22C12C40CE26FC89C6B39D8DF923" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[835,978,1725,1749]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C40CE26FC89C6B39D8DF923" box="[835,978,1725,1749]" italics="true" pageId="12" pageNumber="347">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C40CE26FC29C6B09A05F920" box="[995,1114,1726,1750]" pageId="12" pageNumber="347" rank="genus">new genus</taxonomicNameLabel>
, the posterior epistomial margin is divided into three distinct lobes by well-defined fissures (e.g.,
<figureCitation id="0D9145C72C40CE26FC49C6F39DBDF8E3" box="[899,994,1789,1813]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="12" pageNumber="347">Figs. 3B</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FC3AC6F39A4FF8E3" box="[1008,1040,1789,1813]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="12" pageNumber="347">4E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FBD4C6F39A61F8E3" box="[1054,1086,1789,1813]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="12" pageNumber="347">5E</figureCitation>
,
<figureCitation id="0D9145C72C40CE26FB86C6F39ACEF8E3" box="[1100,1169,1789,1813]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="12" pageNumber="347">6C, D</figureCitation>
). The form of the epistome is the same as in the pilumnid genus
<taxonomicName id="52AA22C12C40CE26FB05C7139D27F8A3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FB05C7139B26F8C3" box="[1231,1401,1821,1845]" italics="true" pageId="12" pageNumber="347">Pseudolitochira</emphasis>
<bibRefCitation id="F13B24B32C40CE26FA4AC7139D27F8A3" author="Ward M" pageId="12" pageNumber="347" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
(sensu
<bibRefCitation id="F13B24B32C40CE26FC1BC7309A21F8A3" author="Ng PKL &amp; Clark PF &amp; Clark B &amp; Kamanli SA" box="[977,1150,1853,1878]" pageId="12" pageNumber="347" pagination="377 - 391" refId="ref14159" refString="Ng PKL, Clark PF, Clark B &amp; Kamanli SA (2021) Pseudolitochira integra (Miers, 1884) (Crustacea: Brachyura: Pilumnidae): redescribed and illustrated from micro-CT scanning the type female. Zootaxa, 4969 (2): 377 - 391." type="journal article" year="2021">Ng et al., 2021</bibRefCitation>
). Species of
<taxonomicName id="52AA22C12C40CE26FADAC7339BE4F8A3" authorityName="Ward" authorityYear="1942" box="[1296,1467,1853,1877]" class="Malacostraca" family="Pilumnidae" genus="Pseudolitochira" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C40CE26FADAC7339BE4F8A3" box="[1296,1467,1853,1877]" italics="true" pageId="12" pageNumber="347">Pseudolitochira</emphasis>
</taxonomicName>
, however, are very setose, with long silky setae that partially or completely obscure the margins and surfaces, and lack any trace of areoles on the carapace, with the regions smooth and hardly demarcated (
<bibRefCitation id="F13B24B32C40CE26FB86C7B39B7BF823" author="Ng PKL &amp; Clark PF" box="[1100,1316,1981,2005]" pageId="12" pageNumber="347" pagination="349 - 364" refId="ref14076" refString="Ng PKL &amp; Clark PF (2022 a) Descriptions of three new Pseudolitochira Ward, 1942 species (Crustacea: Brachyura: Pilumnidae) from the West Pacific. Journal of Natural History, 56 (5 - 8): 349 - 364." type="journal article" year="2022">Ng &amp; Clark, 2022a</bibRefCitation>
, b;
<bibRefCitation id="F13B24B32C40CE26FA85C7B09D23F803" author="Ng PKL &amp; Lin C-W" pageId="12" pageNumber="347" pagination="101 - 114" refId="ref14285" refString="Ng PKL &amp; Lin C-W (2023) Two new species of hairy crabs (Pilumnidae) and the first record of Crinitocinus alcocki (Borradaile, 1900) (Acidopsidae) (Crustacea: Brachyura) from Taiwan. Zootaxa, 5297 (1): 101 - 114." type="journal article" year="2023">Ng &amp; Lin, 2023</bibRefCitation>
;
<bibRefCitation id="F13B24B32C40CE26FC43C7D09DA1F803" author="Ng PKL" box="[905,1022,2013,2038]" pageId="12" pageNumber="347" pagination="120 - 126" refId="ref13924" refString="Ng PKL (2024 a) Pseudolitochira spinosa, a new species of pilumnid crab from Panglao, Philippines, with a note on P. setosa (A. Milne-Edwards, 1873) (Crustacea, Brachyura, Pilumnoidea). Zootaxa, 5512 (1): 120 - 126." type="journal article" year="2024">Ng, 2024a</bibRefCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FFBBC0B09C1FFDE3" blockId="13.[113,753,189,534]" pageId="13" pageNumber="348">
In the general appearance of the carapace, especially in the overall shape and the reduced size of the fourth anterolateral tooth/spine,
<taxonomicName id="52AA22C12C41CE27FE69C0F39C8CFEE3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[419,723,253,277]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FE69C0F39C8CFEE3" box="[419,723,253,277]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="52AA22C12C41CE27FEBBC1139C9DFEC3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[369,706,285,309]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="A7DE85502C41CE27FEBBC1139C6BFEC3" box="[369,564,285,309]" italics="true" pageId="13" pageNumber="348">Viaderiana typica</emphasis>
<bibRefCitation id="F13B24B32C41CE27FDF4C1139C9DFEC3" author="Ward M" box="[574,706,285,309]" pageId="13" pageNumber="348" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="4A11E7E02C41CE27FFBBC1309EC0FEA0" box="[113,159,318,342]" pageId="13" pageNumber="348">type</typeStatus>
species of
<taxonomicName id="52AA22C12C41CE27FED7C1339C7DFEA3" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[285,546,317,341]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C41CE27FED7C1339FCAFEA3" box="[285,405,317,341]" italics="true" pageId="13" pageNumber="348">Viaderiana</emphasis>
<bibRefCitation id="F13B24B32C41CE27FE54C1339C7DFEA3" author="Ward M" box="[414,546,317,341]" pageId="13" pageNumber="348" pagination="49 - 113" refId="ref15262" refString="Ward M (1942) Notes on the Crustacea of the Desjardins Museum, Mauritius Institute, with descriptions of new genera and species. Bulletin of the Mauritius Institute, 2 (2): 49 - 113." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="52AA22C12C41CE27FDE4C1339CAFFEA3" authorityName="Ward" authorityYear="1942" box="[558,752,317,341]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="A7DE85502C41CE27FDE4C1339CAFFEA3" box="[558,752,317,341]" italics="true" pageId="13" pageNumber="348">Viaderiana typica</emphasis>
</taxonomicName>
(and congeners), however, have the extensor margin of the ambulatory merus armed with at least one spine, the regions are never prominently raised to form granules, the ischium of the third maxilliped is distinctly more elongate, and the posterior epistomal margin is divided into three distinct lobes by distinct fissures (see Ng, 2023, 2024b).
</paragraph>
<paragraph id="951559422C41CE27FFBBC2309FC8FD80" blockId="13.[113,752,573,630]" pageId="13" pageNumber="348">
As such,
<taxonomicName id="52AA22C12C41CE27FF19C2339FB6FDA3" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[211,489,573,597]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FF19C2339FB6FDA3" box="[211,489,573,597]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is made the
<typeStatus id="4A11E7E02C41CE27FDA5C2309CC2FDA0" box="[623,669,574,598]" pageId="13" pageNumber="348">type</typeStatus>
species of
<taxonomicName id="52AA22C12C41CE27FF5AC2539F53FD83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[144,268,605,629]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C41CE27FF5AC2539F53FD83" box="[144,268,605,629]" italics="true" pageId="13" pageNumber="348">Takedaplax</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C41CE27FED6C2509FCCFD80" box="[284,403,606,630]" pageId="13" pageNumber="348" rank="genus">new genus</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,317 @@
<document id="AE44976FA763F394CDD93F79386CB61E" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C41CE3AFF4AC2939D57FE63" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Takedaplax sextuberculata Ng &amp; Ahyong &amp; Shane T. 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="350" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="348" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="198A7C27D4FE033F9A99736AA69E433E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8A9851945793B8D806BF44F7D93A00B6">
<mods:title id="7C1342A2A7A84FC79AAF1E86EAAF0D25">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="53F61BB53E30A416ADC508F7B715CA18" type="personal">
<mods:role id="E2C66807B0DB2372696D4D342EB02386">
<mods:roleTerm id="8F0047CFD36F346D8D780C4B2B43D869">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8993FD26D11756D115196A1ACEB0226D">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="77662BFCF1AA08139F2E890406A926E2" type="personal">
<mods:role id="B3394F99685D074167BA3F4ACE64EF66">
<mods:roleTerm id="AA70F65D1A68EBF2B3CD20E82C8944F0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DE85EAE4C57F2412CC2A3315B8DEAC3D">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="C43A8FA54F3B5494067460D7AA5FEEC9" type="personal">
<mods:role id="10C858B44FC8C96BE7A692360BCB0FBC">
<mods:roleTerm id="04A27D4F1B7319451E271A09B9CD4894">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA85B143A7F761F5ECC808693B4EAD6B">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="50F1F471AB9E8A82A363D239599F650E">text</mods:typeOfResource>
<mods:relatedItem id="7B4A4CDF3D8EC848ED01A8659B6FF55F" type="host">
<mods:titleInfo id="272E8E370FE53CA80EB0CBAF6ADF00AF">
<mods:title id="09C827951573E122AD7B2439BC11F4FE">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="582E4F0206371FEC551F662B7F009A5D">
<mods:date id="4514C807BEEA6757C477170D049221D4">2024</mods:date>
<mods:detail id="9D53F12BE949BE91F7BED9C1C41D5299" type="pubDate">
<mods:number id="9424BA58B3DCE294041A81439BE8B764">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="5F00A465A16DBA631B088C84BC463A39" type="volume">
<mods:number id="787FF04BA17901D377B3D809EA6470FF">72</mods:number>
</mods:detail>
<mods:extent id="40C1B2CAB43AE27D8B3A842EACEBF99F" unit="page">
<mods:start id="391C97795628D2A87871493C41FFA8A2">335</mods:start>
<mods:end id="482ABC89D0067A47D2ABB495F353F976">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4A7936B751D28E429E8E14C07A3F23D4">journal article</mods:classification>
<mods:identifier id="B3647B38703D8AC3C70C2C13AB2B75F4" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="28E723963578F53493F01C338CAC1788" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="D4733DC9777E571832A75ECA542825A1" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="9A08A10B8F2573080D380A067EE02C08" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C41CE3AFF4AC2939D57FE63" ID-DOI="http://doi.org/10.5281/zenodo.14682793" ID-Zenodo-Dep="14682793" LSID="urn:lsid:plazi:treatment:1D03E8542C41CE3AFF4AC2939D57FE63" httpUri="http://treatment.plazi.org/id/1D03E8542C41CE3AFF4AC2939D57FE63" lastPageId="16" lastPageNumber="350" pageId="13" pageNumber="348">
<subSubSection id="DDB00AC92C41CE27FF4AC2939C4CFD23" pageId="13" pageNumber="348" type="nomenclature">
<paragraph id="951559422C41CE27FF4AC2939C4CFD23" blockId="13.[128,737,669,757]" pageId="13" pageNumber="348">
<heading id="CE5DEE2E2C41CE27FF4AC2939C4CFD23" bold="true" centered="true" fontSize="10" level="2" pageId="13" pageNumber="348" reason="2">
<emphasis id="A7DE85502C41CE27FF4AC2939C4CFD23" bold="true" pageId="13" pageNumber="348">
<taxonomicName id="52AA22C12C41CE27FF4AC2939C83FD43" ID-CoL="CL2XM" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[128,732,669,693]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C41CE27FF4AC2939FF4FD43" bold="true" box="[128,427,669,693]" italics="true" pageId="13" pageNumber="348">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C41CE27FE71C2939C8BFD43" author="Takeda M &amp; Kurata Y" box="[443,724,669,693]" pageId="13" pageNumber="348" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C41CE27FE84C2B39C4CFD23" box="[334,531,701,725]" pageId="13" pageNumber="348" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FEBEC2D39FB2FD03" box="[372,493,733,757]" pageId="13" pageNumber="348" type="description">
<paragraph id="951559422C41CE27FEBEC2D39FB2FD03" blockId="13.[128,737,669,757]" box="[372,493,733,757]" pageId="13" pageNumber="348">
(
<figureCitation id="0D9145C72C41CE27FEB6C2D39F96FD03" box="[380,457,733,757]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FE1DC2D39FBAFD03" box="[471,485,733,757]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC3129CCEFC9D" pageId="13" pageNumber="348" type="reference_group">
<paragraph id="951559422C41CE27FFBBC3129CCEFC9D" blockId="13.[113,752,796,875]" pageId="13" pageNumber="348">
<treatmentCitationGroup id="B5BA7E6C2C41CE27FFBBC3129CCEFC9D" pageId="13" pageNumber="348">
<treatmentCitation id="140B7F532C41CE27FFBBC3129C26FCC7" author="Takeda M &amp; Kurata Y" box="[113,633,796,818]" page="200" pageId="13" pageNumber="348" year="1984">
<taxonomicName id="52AA22C12C41CE27FFBBC3129C26FCC7" ID-CoL="6QMFQ" authority="Takeda &amp; Kurata, 1984: 200" authorityName="Takeda &amp; Kurata" authorityPageNumber="200" authorityYear="1984" box="[113,633,796,818]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="348" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C41CE27FFBBC3129F37FCC7" box="[113,360,796,817]" italics="true" pageId="13" pageNumber="348">Lophoplax sextuberculata</emphasis>
<bibRefCitation id="F13B24B32C41CE27FEA6C3129C26FCC7" author="Takeda M &amp; Kurata Y" box="[364,633,796,818]" pageId="13" pageNumber="348" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984: 200</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 1416;
<treatmentCitation id="140B7F532C41CE27FF5DC3379F19FCB8" author="Takeda M" box="[151,326,825,847]" page="40" pageId="13" pageNumber="348" year="1993">
<bibRefCitation id="F13B24B32C41CE27FF5DC3379F19FCB8" author="Takeda M" box="[151,326,825,847]" pageId="13" pageNumber="348" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993: 40</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C41CE27FE98C3379C42FCB8" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[338,541,825,847]" page="143" pageId="13" pageNumber="348" year="2008">
<bibRefCitation id="F13B24B32C41CE27FE98C3379C42FCB8" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[338,541,825,847]" pageId="13" pageNumber="348" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C41CE27FDE3C3379EA3FC9D" author="Takeda M &amp; Komatsu H" page="176" pageId="13" pageNumber="348" year="2018">
<bibRefCitation id="F13B24B32C41CE27FDE3C3379EA3FC9D" author="Takeda M &amp; Komatsu H" pageId="13" pageNumber="348" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018: 176</bibRefCitation>
</treatmentCitation>
, fig. 6H;
<treatmentCitation id="140B7F532C41CE27FE90C35B9CD2FC9D" author="Takeda M &amp; Komatsu H" box="[346,653,853,875]" page="167" pageId="13" pageNumber="348" year="2023">
<bibRefCitation id="F13B24B32C41CE27FE90C35B9CD2FC9D" author="Takeda M &amp; Komatsu H" box="[346,653,853,875]" pageId="13" pageNumber="348" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">Takeda &amp; Komatsu, 2023: 167</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC39D9F52FBDA" pageId="13" pageNumber="348" type="materials_examined">
<paragraph id="951559422C41CE27FFBBC39D9F52FBDA" blockId="13.[113,752,915,1068]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC39D9F0CFC5D" bold="true" box="[113,339,915,939]" pageId="13" pageNumber="348">Material examined.</emphasis>
<materialsCitation id="25C2531F2C41CE27FE96C39A9F56FBDA" collectingDate="2008-10-28" collectionCode="R, V" country="Japan" elevation="81" latitude="27.078833" location="Chichi-jima" longLatPrecision="12" longitude="142.15517" municipality="Ogasawara Islands" pageId="13" pageNumber="348" specimenCount="1" specimenCount-male="1">
<specimenCount id="83AC92CB2C41CE27FE96C39A9FF9FC5A" box="[348,422,916,940]" count="1" pageId="13" pageNumber="348" type="male">1 male</specimenCount>
(4.0 ×
<quantity id="5252F4A72C41CE27FE31C39A9C0EFC5A" box="[507,593,916,940]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="348" unit="mm" value="3.2">3.2 mm</quantity>
) (NSMT-Cr S 1253), west of
<location id="90750F992C41CE27FED0C3BA9FC6FC3A" LSID="urn:lsid:plazi:treatment:1D03E8542C41CE3AFF4AC2939D57FE63:90750F992C41CE27FED0C3BA9FC6FC3A" box="[282,409,948,972]" country="Japan" latitude="27.078833" longLatPrecision="12" longitude="142.15517" municipality="Ogasawara Islands" name="Chichi-jima" pageId="13" pageNumber="348">Chichi-jima</location>
[= Island],
<collectingMunicipality id="7571C3382C41CE27FDD4C3BA9CB3FC3A" box="[542,748,948,972]" pageId="13" pageNumber="348">Ogasawara Islands</collectingMunicipality>
,
<collectingCountry id="EDBD19D22C41CE27FFBBC3DA9EEEFC1D" box="[113,177,980,1003]" name="Japan" pageId="13" pageNumber="348">Japan</collectingCountry>
,
<geoCoordinate id="F09E3F852C41CE27FF0AC3DC9F63FC1D" box="[192,316,978,1003]" degrees="27" direction="north" minutes="04.55" orientation="latitude" pageId="13" pageNumber="348" precision="9" value="27.075834">27°04.55N</geoCoordinate>
<geoCoordinate id="F09E3F852C41CE27FE8DC3DC9F91FC1D" box="[327,462,978,1003]" degrees="142" direction="east" minutes="09.16" orientation="longitude" pageId="13" pageNumber="348" precision="9" value="142.15266">142°09.16E</geoCoordinate>
<geoCoordinate id="F09E3F852C41CE27FE16C3DC9C07FC1D" box="[476,600,978,1003]" degrees="27" direction="north" minutes="04.73" orientation="latitude" pageId="13" pageNumber="348" precision="9" value="27.078833">27°04.73N</geoCoordinate>
<geoCoordinate id="F09E3F852C41CE27FDA8C3DC9CB3FC1D" box="[610,748,978,1003]" degrees="142" direction="east" minutes="09.31" orientation="longitude" pageId="13" pageNumber="348" precision="9" value="142.15517">142°09.31E</geoCoordinate>
, 83
<quantity id="5252F4A72C41CE27FF56C3FA9E88FBFD" box="[156,215,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="348" unit="m" value="81.0">
<elevation id="1E87BE712C41CE27FF56C3FA9E88FBFD" box="[156,215,1012,1036]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="13" pageNumber="348" unit="m" value="81.0">81 m</elevation>
</quantity>
,
<collectionCode id="F3BBC1872C41CE27FF2DC3FA9EA6FBFA" box="[231,249,1012,1036]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="348">R</collectionCode>
/
<collectionCode id="F3BBC1872C41CE27FECBC3FA9F4BFBFA" box="[257,276,1012,1036]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="348" type="Museum">V</collectionCode>
Koyo, 2008 cruise, station KY-08-15,
<date id="E1147F822C41CE27FD1FC3FA9F56FBDA" pageId="13" pageNumber="348" value="2008-10-28">
<collectingDate id="F150866A2C41CE27FD1FC3FA9F56FBDA" pageId="13" pageNumber="348" value="2008-10-28">28 October 2008</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE27FFBBC45D9FDDFB9A" box="[113,386,1107,1132]" pageId="13" pageNumber="348" type="diagnosis">
<paragraph id="951559422C41CE27FFBBC45D9FDDFB9A" blockId="13.[113,386,1107,1132]" box="[113,386,1107,1132]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC45D9EB9FB9D" bold="true" box="[113,230,1107,1131]" pageId="13" pageNumber="348">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C41CE25FFBBC49D9F6BF9AE" lastPageId="15" lastPageNumber="350" pageId="13" pageNumber="348" type="description">
<paragraph id="951559422C41CE27FFBBC49D9ACBFE23" blockId="13.[113,753,1171,2060]" lastBlockId="13.[812,1452,189,469]" pageId="13" pageNumber="348">
<emphasis id="A7DE85502C41CE27FFBBC49D9EA3FB5D" bold="true" box="[113,252,1171,1195]" pageId="13" pageNumber="348">Description.</emphasis>
Carapace rectangular; frontal and anterolateral margins with scattered tufts of short and medium-length setae, not brush-like, not concealing margins; regions well demarcated with 6 anterior areoles always large, distinct: epigastric areole with cristate anterior margin; postorbital areole transversely wide, anterior margin subcristate; hepatic areole rounded with subcristate anterior margin; protogastric, cardiac, mesobranchial and cardiobranchial regions gently convex to barely discernible in smaller specimens, gently raised to low areoles in larger specimens; gastro-cardiac with grooves shallow (
<figureCitation id="0D9145C72C41CE27FEBCC5DA9FAAFA1A" box="[374,501,1492,1516]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
); intestinal region not prominently raised, no visible intestino-branchial areole; surface adjacent to posterior carapace margin almost flat, not forming ridge, no transverse channel visible (
<figureCitation id="0D9145C72C41CE27FD47C63A9CBDF9BA" box="[653,738,1588,1612]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
); mesobranchial regions smooth; metabranchial regions covered with granules (
<figureCitation id="0D9145C72C41CE27FEBAC67A9F9AF97A" box="[368,453,1652,1676]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
). Subhepatic and suborbital regions almost smooth; pterygostomial region with scattered small granules (
<figureCitation id="0D9145C72C41CE27FED7C6BA9F2CF93A" box="[285,371,1716,1740]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8D</figureCitation>
). Frontal margin distinctly bilobed, convex, with deep median cleft, lateral lobe very low, just visible in dorsal view (
<figureCitation id="0D9145C72C41CE27FE94C6FA9F90F8FA" box="[350,463,1780,1804]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Supraorbital margin long, sinuous, with distinct submedian lobe, without fissure or cleft (
<figureCitation id="0D9145C72C41CE27FFB3C73A9E8DF8BA" box="[121,210,1844,1868]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B</figureCitation>
). Orbit dorsal, supraorbital margin, eyes clearly visible in dorsal view; eyes freely movable, ocular peduncle relatively long, filling orbit (
<figureCitation id="0D9145C72C41CE27FE08C77A9C1BF87A" box="[450,580,1908,1932]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8B, D</figureCitation>
). Anterolateral margin with 4 visible teeth or spines (including external orbital tooth), external orbital tooth triangular, next 3 teeth acutely triangular to spiniform, last tooth smallest (
<figureCitation id="0D9145C72C41CE27FD09C7DA9EE6F7FA" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Posterolateral margin sinuous, unarmed, margins gently converging towards almost straight posterior carapace margin (
<figureCitation id="0D9145C72C41CE27FC4EC0D39DABFF03" box="[900,1012,221,245]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, B</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C41CE27FB94C0F39AEBFEE3" box="[1118,1204,253,277]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8D</figureCitation>
). Basal antennal article subquadrate; flagellum entering orbital hiatus. Posterior margin of epistome entire, without lateral fissures, median part triangular, lateral parts distinctly concave (8E, 9D). Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges present.
</paragraph>
<paragraph id="951559422C41CE27FCE7C1F39A2BFD63" blockId="13.[813,1453,509,661]" pageId="13" pageNumber="348">
Third maxilliped with ischium relatively short, subquadrate, length 1.3× width, with shallow median oblique sulcus; merus subquadrate, anterolateral angle slightly auriculiform. Exopod relatively stout, tip just reaching to before distal edge of merus (
<figureCitation id="0D9145C72C41CE27FC12C2739A67FD63" box="[984,1080,637,661]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8C</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FB8FC2739A38FD63" box="[1093,1127,637,661]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9C</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C2B39B45FC43" blockId="13.[813,1452,701,949]" pageId="13" pageNumber="348">
Male thoracic sternum surface smooth; sternites 1 and 2 completely fused, wide, lateral margins gently concave to sinuous; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 gently sinuous (
<figureCitation id="0D9145C72C41CE27FB44C3339ABFFCA3" box="[1166,1248,829,853]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8F</figureCitation>
). Sternite 8 clearly visible as subovate plate adjacent to pleonal somite 2 when pleon closed (
<figureCitation id="0D9145C72C41CE27FC03C3739A7CFC63" box="[969,1059,893,917]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">Fig. 9G</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior one-third of somite 5.
</paragraph>
<paragraph id="951559422C41CE27FCE7C3D39BF3FA43" blockId="13.[812,1452,989,1462]" pageId="13" pageNumber="348">
Adult male chelipeds almost symmetrical, relatively slender (
<figureCitation id="0D9145C72C41CE27FCFFC3F39D84FBE3" box="[821,987,1021,1045]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, G, H</figureCitation>
). Basis-ischium fused but suture visible; extensor margins granulated (
<figureCitation id="0D9145C72C41CE27FBA0C4139AE2FBC3" box="[1130,1213,1053,1077]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="13" pageNumber="348">Fig. 1C</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with tubercles and granules; inner distal angle dentiform with low subdistal angle (
<figureCitation id="0D9145C72C41CE27FCBFC4739DB3FB63" box="[885,1004,1149,1173]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8A, H</figureCitation>
). Carpus surface granulate; dorsal surface of carpus with prominent longitudinal areole, separated from swollen lateral fringe by groove; inner distal angle with sharp tooth (
<figureCitation id="0D9145C72C41CE27FC76C4D39A1AFB03" box="[956,1093,1245,1269]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8A, H</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FB9EC4D39A29FB03" box="[1108,1142,1245,1269]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9B</figureCitation>
). Chela with dorsal surface covered with granules; inner surface almost smooth; median and ventral surfaces covered with tubercles and granules; margins of chela with scattered long and short setae, not dense or brush-like; fingers slender, slightly shorter than palm, pigmented brown along distal half; occlusal margins with low teeth, forming distinct gape when closed (
<figureCitation id="0D9145C72C41CE27FAECC5939BFFFA43" box="[1318,1440,1437,1461]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Fig. 8G, H</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C5D39A61F8A3" blockId="13.[813,1452,1501,1877]" pageId="13" pageNumber="348">
Ambulatory legs long, P4 longest, about 2.0× carapace width; P4 merus length about 4.3× height, distinctly longer than half carapace width; outer surface almost smooth; coxa rounded; merus elongate, P25 extensor margin not armed, entire, lined with short and long setae that does not obscure surface; surface of P25 gently convex, without median swelling or ridge; P25 carpus relatively smooth, with very low submarginal ridge, lowest or barely visible in P5; propodus laterally flattened, unarmed, relatively long in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, slightly upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C41CE27FC4EC7339DBAF8A3" box="[900,997,1853,1877]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="13" pageNumber="348">Figs. 8A</figureCitation>
,
<figureCitation id="0D9145C72C41CE27FC38C7339A6EF8A3" box="[1010,1073,1853,1877]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">9E, F</figureCitation>
).
</paragraph>
<paragraph id="951559422C41CE27FCE7C7739BC2F7E3" blockId="13.[813,1451,1917,2069]" pageId="13" pageNumber="348">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely narrower than somite 1, subtrapezoidal; somite 3 broadly subtrapezoidal, as wide as somite 1, somites 4 and 5 trapezoidal, somite 6 and telson missing (
<figureCitation id="0D9145C72C41CE27FAF3C7F39BCEF7E3" box="[1337,1425,2045,2069]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="13" pageNumber="348">Fig. 9G</figureCitation>
).
</paragraph>
<caption id="C1D509CA2C42CE24FF42C7FD9AFDF7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682783" ID-Zenodo-Dep="14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="14" pageNumber="349" startId="14.[136,172,2035,2057]" targetBox="[146,1464,189,2020]" targetPageId="14" targetType="figure">
<paragraph id="951559422C42CE24FF42C7FD9AFDF7B5" blockId="14.[136,1475,2035,2115]" pageId="14" pageNumber="349">
Fig. 8.
<taxonomicName id="52AA22C12C42CE24FF19C7FD9CB6F7FE" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[211,745,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="349" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C42CE24FF19C7FD9F81F7FE" box="[211,478,2035,2056]" italics="true" pageId="14" pageNumber="349">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C42CE24FE24C7FD9CBDF7FE" author="Takeda M &amp; Kurata Y" box="[494,738,2035,2057]" pageId="14" pageNumber="349" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C42CE24FD3DC7FA9DFDF7FF" box="[759,930,2035,2057]" pageId="14" pageNumber="349" rank="species">new combination</taxonomicNameLabel>
, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped.
</paragraph>
</caption>
<caption id="C1D509CA2C43CE25FFBBC52E9F58FA5E" ID-DOI="http://doi.org/10.5281/zenodo.14682785" ID-Zenodo-Dep="14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350" startId="15.[113,149,1312,1334]" targetBox="[114,1452,188,1301]" targetPageId="15" targetType="figure">
<paragraph id="951559422C43CE25FFBBC52E9F58FA5E" blockId="15.[113,1452,1312,1448]" pageId="15" pageNumber="350">
Fig. 9.
<taxonomicName id="52AA22C12C43CE25FF77C52E9C8CFAC3" authority="(Takeda &amp; Kurata, 1984)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Takeda &amp; Kurata" baseAuthorityYear="1984" box="[189,723,1312,1334]" class="Malacostraca" family="Pilumnidae" genus="Takedaplax" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata" status="comb. nov.">
<emphasis id="A7DE85502C43CE25FF77C52E9F97FAC3" box="[189,456,1312,1333]" italics="true" pageId="15" pageNumber="350">Takedaplax sextuberculata</emphasis>
(
<bibRefCitation id="F13B24B32C43CE25FE1DC52E9C93FAC3" author="Takeda M &amp; Kurata Y" box="[471,716,1312,1334]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C43CE25FD2AC52E9DD3FAC0" box="[736,908,1312,1334]" pageId="15" pageNumber="350" rank="species">new combination</taxonomicNameLabel>
. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after
<bibRefCitation id="F13B24B32C43CE25FF27C5339F87FAA4" author="Takeda M &amp; Kurata Y" box="[237,472,1341,1363]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm.
</paragraph>
</caption>
<paragraph id="951559422C43CE25FFBBC5EE9F6BF9AE" blockId="15.[113,752,1504,1624]" pageId="15" pageNumber="350">
G1strongly sinuous, distally recurved, tapering to slightly rounded tip, hooked inwards; subdistal surface with 1 long stout seta (
<figureCitation id="0D9145C72C43CE25FF25C62E9F3BF9CE" box="[239,356,1568,1592]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350">Fig. 9H, I</figureCitation>
). G2 short, about ¼ length of G1, sigmoid (
<figureCitation id="0D9145C72C43CE25FF12C64E9F78F9AE" box="[216,295,1600,1624]" captionStart="Fig" captionStartId="15.[113,149,1312,1334]" captionTargetBox="[114,1452,188,1301]" captionTargetId="figure-8@15.[98,1465,184,1327]" captionTargetPageId="15" captionText="Fig. 9. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination. AC, holotype female (8.2 × 6.2 mm) (NSMT-Cr S 8949) (after Takeda &amp; Kurata, 1984: figs. 1416); DJ, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, carapace; B, dorsal view of left cheliped; C, right third maxilliped; D, epistome; E, right P4; F, right P5; G, pleon (missing somite 6 and telson) showing exposed sternite 8 lateral to somite 2; H, right G1 (ventral view); I, right G1 (dorsal view); J, right G2. Scales: D, G = 0.5 mm; E, F = 1.0 mm; HJ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682785" httpUri="https://zenodo.org/record/14682785/files/figure.png" pageId="15" pageNumber="350">Fig. 9J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C43CE3AFFBBC68E9D57FE63" lastPageId="16" lastPageNumber="351" pageId="15" pageNumber="350" type="discussion">
<paragraph id="951559422C43CE25FFBBC68E9C57F88E" blockId="15.[113,753,1664,1912]" pageId="15" pageNumber="350">
<emphasis id="A7DE85502C43CE25FFBBC68E9EBFF96E" bold="true" box="[113,224,1664,1688]" pageId="15" pageNumber="350">Remarks.</emphasis>
<taxonomicName id="52AA22C12C43CE25FF2FC68E9FA7F96E" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[229,504,1664,1688]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C43CE25FF2FC68E9FA7F96E" box="[229,504,1664,1688]" italics="true" pageId="15" pageNumber="350">Lophoplax sextuberculata</emphasis>
</taxonomicName>
was described from the female
<typeStatus id="4A11E7E02C43CE25FF74C6AE9F45F94E" box="[190,282,1696,1720]" pageId="15" pageNumber="350" type="holotype">holotype</typeStatus>
(8.2 ×
<quantity id="5252F4A72C43CE25FEA8C6AE9FECF94E" box="[354,435,1696,1720]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="350" unit="mm" value="6.2">6.2 mm</quantity>
) and
<specimenCount id="83AC92CB2C43CE25FE20C6AF9C16F94E" box="[490,585,1696,1720]" count="2" pageId="15" pageNumber="350" type="male">two male</specimenCount>
<typeStatus id="4A11E7E02C43CE25FD84C6AF9CECF94F" box="[590,691,1697,1721]" pageId="15" pageNumber="350" type="paratype">paratypes</typeStatus>
(both 5.2 ×
<quantity id="5252F4A72C43CE25FF7CC6CE9F52F92E" box="[182,269,1728,1752]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="350" unit="mm" value="3.8">3.8 mm</quantity>
) collected from inside a fish stomach from Takinoura Bay, Ani-jima [=
<collectingCountry id="EDBD19D22C43CE25FE55C6EE9FBAF90E" box="[415,485,1760,1784]" name="Iceland" pageId="15" pageNumber="350">Island</collectingCountry>
], Ogasawara Islands, and a
<typeStatus id="4A11E7E02C43CE25FF48C70F9E82F8EF" box="[130,221,1793,1817]" pageId="15" pageNumber="350" type="paratype">paratype</typeStatus>
male (5.5 ×
<quantity id="5252F4A72C43CE25FEAAC70E9FEEF8EE" box="[352,433,1792,1816]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="15" pageNumber="350" unit="mm" value="4.5">4.5 mm</quantity>
) from Tanegashima [=
<collectingCountry id="EDBD19D22C43CE25FD6CC70E9CB5F8EE" box="[678,746,1792,1816]" name="Iceland" pageId="15" pageNumber="350">Island</collectingCountry>
] (southern Kyushu) (
<bibRefCitation id="F13B24B32C43CE25FE8CC72E9C19F8CE" author="Takeda M &amp; Kurata Y" box="[326,582,1824,1848]" pageId="15" pageNumber="350" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984</bibRefCitation>
;
<bibRefCitation id="F13B24B32C43CE25FD9AC72E9CBBF8CE" author="Takeda M" box="[592,740,1824,1848]" pageId="15" pageNumber="350" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993</bibRefCitation>
).
<bibRefCitation id="F13B24B32C43CE25FFBBC74E9FFAF8AE" author="Takeda M &amp; Marumura M" box="[113,421,1856,1880]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995)</bibRefCitation>
reported additional specimens from the Kii Peninsula and Ryukyus.
</paragraph>
<paragraph id="951559422C43CE3AFFBBC7AE9D57FE63" blockId="15.[113,753,1952,2104]" lastBlockId="16.[136,776,189,406]" lastPageId="16" lastPageNumber="351" pageId="15" pageNumber="350">
They noted that while the median and posterior surfaces of the carapace were smooth in the
<typeStatus id="4A11E7E02C43CE25FE2BC7CF9C50F82F" box="[481,527,1985,2009]" pageId="15" pageNumber="350">type</typeStatus>
specimens from the Ogasawara Islands (as in the case of the present specimen; see also
<bibRefCitation id="F13B24B32C43CE25FF04C80E9FB6F7EE" author="Takeda M &amp; Komatsu H" box="[206,489,2048,2072]" pageId="15" pageNumber="350" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: fig. 6H); the specimens from Tanegashima [=
<collectingCountry id="EDBD19D22C43CE25FEACC82E9FF5F7CE" box="[358,426,2080,2104]" name="Iceland" pageId="15" pageNumber="350">Island</collectingCountry>
] and Kii Peninsula had seven additional areoles in two transverse rows on the following regions: two on the protogastric, one each on anterior parts of the mesobranchials, one cardiac and one each on the cardiobranchials (see
<bibRefCitation id="F13B24B32C43CE25FBD4C64E9B23F9AE" author="Takeda M &amp; Marumura M" box="[1054,1404,1600,1624]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 89</bibRefCitation>
, 91, fig. 1). They noted that the only specimen they had from the Ryukyus was small (4.8 × 4.0 mm) and had the median and posterior parts of the carapace smooth (
<bibRefCitation id="F13B24B32C43CE25FB0CC6AE9DD6F92E" author="Takeda M &amp; Marumura M" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 91</bibRefCitation>
). As such,
<bibRefCitation id="F13B24B32C43CE25FC33C6CE9B19F92E" author="Takeda M &amp; Marumura M" box="[1017,1350,1728,1752]" pageId="15" pageNumber="350" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995: 91)</bibRefCitation>
observed that “… there may be some individual differences as for the development of the areolets on the posterior part, but it is concluded that
<taxonomicName id="52AA22C12C43CE25FC13C72E9AA9F8CE" authorityName="Takeda &amp; Kurata" authorityYear="1984" box="[985,1270,1824,1848]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="350" phylum="Arthropoda" rank="species" species="sextuberculata">
<emphasis id="A7DE85502C43CE25FC13C72E9AA9F8CE" box="[985,1270,1824,1848]" italics="true" pageId="15" pageNumber="350">Lophoplax sextuberculata</emphasis>
</taxonomicName>
is characterised by having 13 areolets, not 6 mentioned in the original description and scientific name”. Examining the present specimen from the Ogasawara Islands, we note that its protogastric, mesobranchial, cardiac, and cardiobranchial regions are only gently convex and smooth, and barely discernible as areolets in dorsal view (
<figureCitation id="0D9145C72C43CE25FB0CC7EE9B04F80E" box="[1222,1371,2016,2040]" captionStart="Fig" captionStartId="14.[136,172,2035,2057]" captionTargetBox="[146,1464,189,2020]" captionTargetId="figure-7@14.[146,1464,189,2020]" captionTargetPageId="14" captionText="Fig. 8. Takedaplax sextuberculata (Takeda &amp; Kurata, 1984), new combination, male (4.0 × 3.2 mm) (NSMT-Cr S 1253), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and part of pleon; G, left chela; H, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682783" httpUri="https://zenodo.org/record/14682783/files/figure.png" pageId="15" pageNumber="350">Fig. 8A, B, D</figureCitation>
). In the colour figure of this specimen (
<bibRefCitation id="F13B24B32C43CE25FB4CC80E9BF7F7EE" author="Takeda M &amp; Komatsu H" box="[1158,1448,2048,2072]" pageId="15" pageNumber="350" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
: fig. 6H), we note that these regions are also slightly more orangish than the surrounding beige surface as described by
<bibRefCitation id="F13B24B32C5CCE3AFF42C0D39FE8FF03" author="Takeda M &amp; Marumura M" box="[136,439,221,245]" pageId="16" pageNumber="351" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura (1995)</bibRefCitation>
for their specimens. As such, it seems likely that these areolets are present but not always discernible, probably becoming more prominent in larger individuals.
<bibRefCitation id="F13B24B32C5CCE3AFEC0C1309FB7FEA3" author="Ng PKL &amp; Rahayu DL" box="[266,488,317,342]" pageId="16" pageNumber="351" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu (2023)</bibRefCitation>
reported a similar situation for
<taxonomicName id="52AA22C12C5CCE3AFF7AC1539FF9FE83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Ng &amp; Rahayu" baseAuthorityYear="2023" box="[176,422,349,373]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="sordidus">
<emphasis id="A7DE85502C5CCE3AFF7AC1539FF9FE83" box="[176,422,349,373]" italics="true" pageId="16" pageNumber="351">Myopilumnus sordidus</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C5CCE3AFE2AC1509CD2FE83" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[480,653,349,373]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="vermiculatus">
<emphasis id="A7DE85502C5CCE3AFE2AC1509CD2FE83" box="[480,653,349,373]" italics="true" pageId="16" pageNumber="351">M. vermiculatus</emphasis>
</taxonomicName>
, where the areoles are relatively poorly developed in young specimens.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,337 @@
<document id="02106DDE6D591A0B73EA926F0F7FF78F" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C45CE26FC93C3F39D57F943" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Hosekia symmetrinuda Ng &amp; Ahyong &amp; Shane T. 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="347" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="344" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="66F2FE82C9F47BAD2FC089F8C6D61390" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="62CC6B6D79E4C03C21336CC71D6577E1">
<mods:title id="567D2FE8C49EDA20FFAB691F99807578">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="9CB5B5723D871DEA737E7A1B0A536C75" type="personal">
<mods:role id="6DC3D7A4668484C7EDA69F6DB4860E72">
<mods:roleTerm id="BD9BBCE6C3E2C9403BB21E39B00BEDA8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D0DB930DAA74A6F87B9528127EAB420">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="FE889667DBB051EDFF2B62BBC9F2D263" type="personal">
<mods:role id="3DDECC05EF29D1120EA4F970B997315F">
<mods:roleTerm id="71EBE61F2D48EA50FC3E0317C66A7760">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9E1699FA5255A3CBACF750707BEBBC70">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="D0F9CCCA256AF0698D27E820DDBA1E6E" type="personal">
<mods:role id="5CCC1F395A75D505E722C45F484681A1">
<mods:roleTerm id="9D3B66E37161CF866E75B3DBFB92EEBE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CDD9DCDEB57DCDEE697317384AB76B79">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0B57AEF40979F3BEADE23EC6481EDF34">text</mods:typeOfResource>
<mods:relatedItem id="D91C0F399D91243099103E5F0BBA9168" type="host">
<mods:titleInfo id="6C09D40A06CCC82A46A327785DAA13BA">
<mods:title id="8A6738B56F558EEF8CDBBBDEA509F601">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0853554DC15FF8E5B02CD5AF77E27696">
<mods:date id="C72F7263BF1854075355F37D0E827DB2">2024</mods:date>
<mods:detail id="A5DF73BBF39671CCE46891B929A096C7" type="pubDate">
<mods:number id="FEE4332449D0C695AA693CC2B02046FF">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="2894CB0BDFFBD305A263914766521463" type="volume">
<mods:number id="EB027E7F5EEDAFBA8E36B2D2A60BCD47">72</mods:number>
</mods:detail>
<mods:extent id="066022AEB658544DF2691AA3EBED23FE" unit="page">
<mods:start id="B3D7AC169FFABAAA43C7033D63B77F06">335</mods:start>
<mods:end id="8238C7F8B5245003309B1F475A7EF20E">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0EF838BF331A3D00C4C1BCA3A31DB0A3">journal article</mods:classification>
<mods:identifier id="E34B026DC44AEEFA22F4EEDB8647EF58" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="2C705243C551F44CB101DE47CD0114B9" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="1D667330863EAE13DFC7E395D1734F5E" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="87A10EC0F9A691F02B36A983475322EC" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C45CE26FC93C3F39D57F943" LSID="urn:lsid:plazi:treatment:1D03E8542C45CE26FC93C3F39D57F943" httpUri="http://treatment.plazi.org/id/1D03E8542C45CE26FC93C3F39D57F943" lastPageId="12" lastPageNumber="347" pageId="9" pageNumber="344">
<subSubSection id="DDB00AC92C45CE23FC93C3F39AEDFBC3" pageId="9" pageNumber="344" type="nomenclature">
<paragraph id="951559422C45CE23FC93C3F39AEDFBC3" blockId="9.[857,1406,1021,1109]" pageId="9" pageNumber="344">
<heading id="CE5DEE2E2C45CE23FC93C3F39AEDFBC3" bold="true" centered="true" fontSize="10" level="2" pageId="9" pageNumber="344" reason="2">
<emphasis id="A7DE85502C45CE23FC93C3F39AEDFBC3" bold="true" pageId="9" pageNumber="344">
<taxonomicName id="52AA22C12C45CE23FC93C3F39B1DFBE3" ID-CoL="CKZYG" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[857,1346,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C45CE23FC93C3F39A06FBE3" bold="true" box="[857,1113,1021,1045]" italics="true" pageId="9" pageNumber="344">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C45CE23FBA3C3F39B64FBE3" author="Edmondson CH" box="[1129,1339,1021,1045]" pageId="9" pageNumber="344" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C45CE23FA9AC3F39AEDFBC3" pageId="9" pageNumber="344" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FBE5C4339AF7FBA3" box="[1071,1192,1085,1109]" pageId="9" pageNumber="344" type="description">
<paragraph id="951559422C45CE23FBE5C4339AF7FBA3" blockId="9.[857,1406,1021,1109]" box="[1071,1192,1085,1109]" pageId="9" pageNumber="344">
(
<figureCitation id="0D9145C72C45CE23FBFDC4339ADBFBA3" box="[1079,1156,1085,1109]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="9" pageNumber="344">Figs. 6</figureCitation>
,
<figureCitation id="0D9145C72C45CE23FB58C4339AFFFBA3" box="[1170,1184,1085,1109]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="9" pageNumber="344">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C4729A00FAC8" pageId="9" pageNumber="344" type="reference_group">
<paragraph id="951559422C45CE23FCE6C4729AE6FB1E" blockId="9.[812,1452,1148,1343]" pageId="9" pageNumber="344">
<treatmentCitationGroup id="B5BA7E6C2C45CE23FCE6C4729AE6FB1E" pageId="9" pageNumber="344">
<treatmentCitation id="140B7F532C45CE23FCE6C4729BDEFB67" author="Edmondson CH" box="[812,1409,1148,1170]" page="233" pageId="9" pageNumber="344" year="1951">
<taxonomicName id="52AA22C12C45CE23FCE6C4729BDEFB67" ID-CoL="4NV9Q" authority="Edmondson, 1951: 233" authorityName="Edmondson" authorityPageNumber="233" authorityYear="1951" box="[812,1409,1148,1170]" class="Malacostraca" family="Pilumnidae" genus="Pseudocryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="A7DE85502C45CE23FCE6C4729AC6FB67" box="[812,1177,1148,1169]" italics="true" pageId="9" pageNumber="344">Pseudocryptocoeloma symmetrinudus</emphasis>
<bibRefCitation id="F13B24B32C45CE23FB55C4729BDEFB67" author="Edmondson CH" box="[1183,1409,1148,1170]" pageId="9" pageNumber="344" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951: 233</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 34;
<treatmentCitation id="140B7F532C45CE23FCB2C4979A7FFB59" author="Serene R" box="[888,1056,1177,1199]" page="86" pageId="9" pageNumber="344" year="1968">
<bibRefCitation id="F13B24B32C45CE23FCB2C4979A7FFB59" author="Serene R" box="[888,1056,1177,1199]" pageId="9" pageNumber="344" pagination="33 - 120" refId="ref14694" refString="Serene R (1968) The Brachyura of the Indo Pacific Region. In: Prodromus for a Check List of the Non-planctonic Marine Fauna of South East Asia. Special Publication of the Singapore National Academy of Science No. 1: 33 - 120." type="book chapter" year="1968">Serène, 1968: 86</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FBE1C4979AF1FB58" author="Ng PKL" box="[1067,1198,1177,1198]" page="79" pageId="9" pageNumber="344" year="1987">
<bibRefCitation id="F13B24B32C45CE23FBE1C4979AF1FB58" author="Ng PKL" box="[1067,1198,1177,1198]" pageId="9" pageNumber="344" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 79</bibRefCitation>
</treatmentCitation>
, 97;
<treatmentCitation id="140B7F532C45CE23FB2AC4979BF8FB58" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1248,1447,1177,1199]" page="144" pageId="9" pageNumber="344" year="2008">
<bibRefCitation id="F13B24B32C45CE23FB2AC4979BF8FB58" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1248,1447,1177,1199]" pageId="9" pageNumber="344" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 144</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FC98C4B89AC9FB3D" author="Marumura M &amp; Takeda M" box="[850,1174,1205,1227]" page="192" pageId="9" pageNumber="344" year="2012">
<bibRefCitation id="F13B24B32C45CE23FC98C4B89AC9FB3D" author="Marumura M &amp; Takeda M" box="[850,1174,1205,1227]" pageId="9" pageNumber="344" pagination="189 - 197" refId="ref13535" refString="Marumura M &amp; Takeda M (2012) Crab specimens in the Wakayama Prefectural Museum of Nature (Nagai Collection) taxonomic study III. New record species in Japan (3). Teikyo Heisei University Bulletin, 23 (1): 189 - 197." type="journal article" year="2012">Marumura &amp; Takeda, 2012: 192</bibRefCitation>
</treatmentCitation>
, figs. 2A, 3, 4;
<treatmentCitation id="140B7F532C45CE23FAFFC4B89DF5FB11" author="Maenosono T" page="32" pageId="9" pageNumber="344" year="2019">
<bibRefCitation id="F13B24B32C45CE23FAFFC4B89DF5FB11" author="Maenosono T" pageId="9" pageNumber="344" pagination="19 - 44" refId="ref13491" refString="Maenosono T (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48: 19 - 44. [In Japanese with English abstract]" type="journal article" year="2019">Maenosono, 2019: 32</bibRefCitation>
</treatmentCitation>
, figs. 8B, 10, 13B, 14K, L.
</treatmentCitationGroup>
</paragraph>
<paragraph id="951559422C45CE23FCE6C4E19A00FAC8" blockId="9.[812,1452,1148,1343]" pageId="9" pageNumber="344">
<treatmentCitationGroup id="B5BA7E6C2C45CE23FCE6C4E19A00FAC8" pageId="9" pageNumber="344">
<taxonomicName id="52AA22C12C45CE23FCE6C4E19A76FAF2" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[812,1065,1263,1284]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="344" phylum="Arthropoda" rank="species" species="symmetrinudus">
<emphasis id="A7DE85502C45CE23FCE6C4E19A76FAF2" box="[812,1065,1263,1284]" italics="true" pageId="9" pageNumber="344">Lophoplax symmetrinudus</emphasis>
</taxonomicName>
<treatmentCitation id="140B7F532C45CE23FB81C4E19B03FAF2" author="Takeda M &amp; Kurata Y" box="[1099,1372,1263,1285]" page="201" pageId="9" pageNumber="344" year="1984">
<bibRefCitation id="F13B24B32C45CE23FB81C4E19B03FAF2" author="Takeda M &amp; Kurata Y" box="[1099,1372,1263,1285]" pageId="9" pageNumber="344" pagination="195 - 202" refId="ref15050" refString="Takeda M &amp; Kurata Y (1984) Crabs of the Ogasawara Islands. VII. Third report on the species obtained from stomachs of fishes. Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 10 (4): 195 - 202, figs. 1 - 16." type="journal article" year="1984">Takeda &amp; Kurata, 1984: 201</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FAAEC4E19A63FAD7" author="Takeda M &amp; Marumura M" page="90" pageId="9" pageNumber="344" year="1995">
<bibRefCitation id="F13B24B32C45CE23FAAEC4E19A63FAD7" author="Takeda M &amp; Marumura M" pageId="9" pageNumber="344" pagination="87 - 91" refId="ref15107" refString="Takeda M &amp; Marumura M (1995) Emendatory notes on Lophoplax sextuberculata Takeda et Kurata (Crustacea, Decapoda, Brachyura). Bulletin of the National Science Museum, Tokyo, Series A (Zoology), 21 (2): 87 - 91." type="journal article" year="1995">Takeda &amp; Marumura, 1995: 90</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FB82C5029B39FAD4" author="Poore GCB &amp; Ahyong ST" box="[1096,1382,1292,1314]" page="680" pageId="9" pageNumber="344" year="2023">
<bibRefCitation id="F13B24B32C45CE23FB82C5029B39FAD4" author="Poore GCB &amp; Ahyong ST" box="[1096,1382,1292,1314]" pageId="9" pageNumber="344" refId="ref14446" refString="Poore GCB &amp; Ahyong ST (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, Australia, xii + 890 pp." type="book" year="2023">Poore &amp; Ahyong, 2023: 680</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C45CE23FAB8C5029A51FAC8" author="Ng PKL &amp; Rahayu DL" page="444" pageId="9" pageNumber="344" year="2023">
<bibRefCitation id="F13B24B32C45CE23FAB8C5029A51FAC8" author="Ng PKL &amp; Rahayu DL" pageId="9" pageNumber="344" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023: 444</bibRefCitation>
</treatmentCitation>
, fig. 12.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C5699AB0F9C9" pageId="9" pageNumber="344" type="materials_examined">
<paragraph id="951559422C45CE23FCE6C5699AB0F9C9" blockId="9.[812,1451,1383,1599]" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C5699A50FA89" bold="true" box="[812,1039,1383,1407]" pageId="9" pageNumber="344">Material examined.</emphasis>
<materialsCitation id="25C2531F2C45CE23FBD3C5699AB0F9C9" collectingDate="2016-02-10" collectingDateMax="2020-04-07" collectingDateMin="2016-02-10" collectionCode="T" collectorName="Siufaga &amp; Tau &amp; W. Harris &amp; Others &amp; Bise &amp; Motobu &amp; Ryukyus &amp; Maenosono" country="Samoa" county="Odo Beach" location="Ryukyus" municipality="Itoman" pageId="9" pageNumber="344" specimenCode="BPBM 5109" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus id="4A11E7E02C45CE23FBD3C5699ADEFA89" box="[1049,1153,1383,1407]" pageId="9" pageNumber="344" type="holotype">Holotype</typeStatus>
:
<specimenCode id="C50CF1392C45CE23FB44C5699B44FA89" box="[1166,1307,1383,1407]" collectionCode="BPBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="9" pageNumber="344" type="Museum">BPBM 5109</specimenCode>
, male (6.5 × 5.0 mm),
<collectorName id="385F3C942C45CE23FC56C5899DAEFA69" box="[924,1009,1415,1439]" pageId="9" pageNumber="344">Siufaga</collectorName>
,
<collectorName id="385F3C942C45CE23FBCAC5899A72FA68" box="[1024,1069,1415,1438]" pageId="9" pageNumber="344">Tau</collectorName>
,
<collectingCountry id="EDBD19D22C45CE23FBF1C5899AD8FA69" box="[1083,1159,1415,1439]" name="Samoa" pageId="9" pageNumber="344">Samoa</collectingCountry>
, low tide, table reef, coll.
<collectorName id="385F3C942C45CE23FCE6C5A99DC2FA49" box="[812,925,1447,1471]" pageId="9" pageNumber="344">W. Harris</collectorName>
, 1937.
<collectorName id="385F3C942C45CE23FC3BC5A99A62FA49" box="[1009,1085,1447,1471]" pageId="9" pageNumber="344">Others</collectorName>
: RUMF-ZC-5199,
<specimenCount id="83AC92CB2C45CE23FAD8C5A99B02FA49" box="[1298,1373,1447,1471]" count="1" pageId="9" pageNumber="344" type="male">1 male</specimenCount>
(7.6 ×
<quantity id="5252F4A72C45CE23FCE6C5C99DDDFA29" box="[812,898,1479,1503]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="344" unit="mm" value="4.2">4.2 mm</quantity>
),
<collectorName id="385F3C942C45CE23FC5CC5C99D97FA29" box="[918,968,1479,1503]" pageId="9" pageNumber="344">Bise</collectorName>
,
<collectorName id="385F3C942C45CE23FC1EC5C99A74FA29" box="[980,1067,1479,1503]" pageId="9" pageNumber="344">Motobu</collectorName>
,
<collectingRegion id="576E97A02C45CE23FBFDC5C99AC5FA29" box="[1079,1178,1479,1503]" country="Japan" name="Okinawa" pageId="9" pageNumber="344">Okinawa</collectingRegion>
,
<collectorName id="385F3C942C45CE23FB6FC5C99B58FA29" box="[1189,1287,1479,1503]" pageId="9" pageNumber="344">Ryukyus</collectorName>
,
<collectingCountry id="EDBD19D22C45CE23FAD9C5C99B0DFA28" box="[1299,1362,1479,1502]" name="Japan" pageId="9" pageNumber="344">Japan</collectingCountry>
, coll.
<collectionCode id="F3BBC1872C45CE23FA5EC5C99BF8FA28" box="[1428,1447,1479,1502]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="344">T</collectionCode>
.
<collectorName id="385F3C942C45CE23FCE6C5E99DF4FA08" box="[812,939,1511,1534]" pageId="9" pageNumber="344">Maenosono</collectorName>
,
<date id="E1147F822C45CE23FC72C5E99A22FA09" box="[952,1149,1511,1535]" pageId="9" pageNumber="344" value="2016-02-10">
<collectingDate id="F150866A2C45CE23FC72C5E99A22FA09" box="[952,1149,1511,1535]" pageId="9" pageNumber="344" value="2016-02-10">10 February 2016</collectingDate>
</date>
; RUMF-ZC-7536,
<specimenCount id="83AC92CB2C45CE23FA87C5E99BF4FA09" box="[1357,1451,1511,1535]" count="1" pageId="9" pageNumber="344" type="female">1 female</specimenCount>
(9.7 ×
<quantity id="5252F4A72C45CE23FCB0C6099D8BF9E9" box="[890,980,1543,1567]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="9" pageNumber="344" unit="mm" value="6.9">6.9 mm</quantity>
),
<collectingCounty id="7C7421CE2C45CE23FC20C6099A37F9E9" box="[1002,1128,1543,1567]" pageId="9" pageNumber="344">Odo Beach</collectingCounty>
,
<collectingMunicipality id="7571C3382C45CE23FBBCC6099A9AF9E8" box="[1142,1221,1543,1566]" pageId="9" pageNumber="344">Itoman</collectingMunicipality>
,
<collectingRegion id="576E97A02C45CE23FB19C6099B68F9E9" box="[1235,1335,1543,1567]" country="Japan" name="Okinawa" pageId="9" pageNumber="344">Okinawa</collectingRegion>
,
<location id="90750F992C45CE23FA8FC6099BF8F9E9" LSID="urn:lsid:plazi:treatment:1D03E8542C45CE26FC93C3F39D57F943:90750F992C45CE23FA8FC6099BF8F9E9" box="[1349,1447,1543,1567]" country="Samoa" county="Odo Beach" municipality="Itoman" name="Ryukyus" pageId="9" pageNumber="344" stateProvince="Okinawa">Ryukyus</location>
,
<collectingCountry id="EDBD19D22C45CE23FCE6C6299D34F9C8" box="[812,875,1575,1598]" name="Japan" pageId="9" pageNumber="344">Japan</collectingCountry>
, coll.
<collectionCode id="F3BBC1872C45CE23FC7AC6299D9CF9C8" box="[944,963,1575,1598]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="344">T</collectionCode>
.
<location id="90750F992C45CE23FC05C6299A11F9C8" LSID="urn:lsid:plazi:treatment:1D03E8542C45CE26FC93C3F39D57F943:90750F992C45CE23FC05C6299A11F9C8" box="[975,1102,1575,1598]" country="Samoa" county="Odo Beach" municipality="Itoman" name="Maenosono" pageId="9" pageNumber="344" stateProvince="Okinawa">Maenosono</location>
,
<date id="E1147F822C45CE23FB91C6299AB4F9C9" box="[1115,1259,1575,1599]" pageId="9" pageNumber="344" value="2020-04-07">
<collectingDate id="F150866A2C45CE23FB91C6299AB4F9C9" box="[1115,1259,1575,1599]" pageId="9" pageNumber="344" value="2020-04-07">7 April 2020</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE23FCE6C6699A62F989" box="[812,1085,1639,1663]" pageId="9" pageNumber="344" type="diagnosis">
<paragraph id="951559422C45CE23FCE6C6699A62F989" blockId="9.[812,1085,1639,1663]" box="[812,1085,1639,1663]" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C6699DFEF989" bold="true" box="[812,929,1639,1663]" pageId="9" pageNumber="344">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C45CE26FCE6C6A99F30FA03" lastPageId="12" lastPageNumber="347" pageId="9" pageNumber="344" type="description">
<paragraph id="951559422C45CE20FCE6C6A99B27F7FD" blockId="9.[812,1452,1703,2111]" lastBlockId="10.[835,1475,1682,2059]" lastPageId="10" lastPageNumber="345" pageId="9" pageNumber="344">
<emphasis id="A7DE85502C45CE23FCE6C6A99DE4F949" bold="true" box="[812,955,1703,1727]" pageId="9" pageNumber="344">Description.</emphasis>
Carapace transversely ovate-subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like, not concealing margins; regions well demarcated with areoles smooth, prominent: epigastric areoles distinct; protogastric areole well developed, extending obliquely to branchial region, approximately separated by median constriction; cardio-branchial areole transversely ovate, well-developed; gastro-cardiac region depressed, with grooves barely visible (
<figureCitation id="0D9145C72C45CE23FB12C7A99B07F848" box="[1240,1368,1959,1983]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="9" pageNumber="344">Figs. 6A, B</figureCitation>
,
<figureCitation id="0D9145C72C45CE23FAAEC7A99D1DF829" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="9" pageNumber="344">7A, C, D</figureCitation>
); intestino-branchial areoles prominent, transversely ovate, one on each side of low intestinal region; posterior carapace margin cristate, adjacent to prominent intestino-branchial areole, forming narrow channel extending to beginning of sub-branchial region; no ridges on sub-branchial region (
<figureCitation id="0D9145C72C46CE20FF5AC6BC9F4EF93C" box="[144,273,1714,1738]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A, B</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FED4C6BD9FDBF93D" box="[286,388,1715,1739]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7A, C, D</figureCitation>
). Subhepatic and suborbital regions slightly rugose to almost smooth; pterygostomial region with numerous closely packed small granules (
<figureCitation id="0D9145C72C46CE20FD4DC6FC9CA3F8FC" box="[647,764,1778,1802]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6CE</figureCitation>
). Frontal margin bilobed, gently convex, with narrow, small median notch, without lateral lobe (
<figureCitation id="0D9145C72C46CE20FDD4C73C9CFBF8BC" box="[542,676,1842,1866]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6AD</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FD79C73D9CA4F8BD" box="[691,763,1843,1867]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Supraorbital margin short, sinuous, with low lobe on posterior half (formed by anterior margin of hepatic areole), without fissure or cleft (
<figureCitation id="0D9145C72C46CE20FEF9C79C9FCCF85C" box="[307,403,1938,1962]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FE55C79D9FBCF85D" box="[415,483,1939,1963]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Orbit dorsal, supraorbital margin appears confluent with general carapace outline, eyes clearly visible in dorsal view; eyes mobile, ocular peduncle short, filling orbit (
<figureCitation id="0D9145C72C46CE20FE96C7FC9F8AF7FC" box="[348,469,2034,2058]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6BD</figureCitation>
). Anterolateral margin with 3 distinct teeth (including external orbital tooth), first 2 broadly triangular with cristate margin, last tooth lower, more rounded; distinct groove visible adjacent to margin, formed with outer margin of hepatic lobe (
<figureCitation id="0D9145C72C46CE20FAE3C6DC9BD2F91C" box="[1321,1421,1746,1770]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FA56C6DD9D06F8FD" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Posterolateral margin gently convex, smooth, unarmed; margins distinctly converging towards posterior carapace margin (
<figureCitation id="0D9145C72C46CE20FC57C73C9DA4F8BC" box="[925,1019,1842,1866]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C46CE20FBCFC73D9A18F8BD" box="[1029,1095,1843,1867]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="10" pageNumber="345">7C, D</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C46CE20FB60C75C9B46F89D" box="[1194,1305,1874,1899]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Basal antennal article subquadrate; flagellum enters orbital hiatus (
<figureCitation id="0D9145C72C46CE20FA5FC77C9DD8F85D" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Epistome with biconcave posterior margin, median lobe triangular with very short median fissue (
<figureCitation id="0D9145C72C46CE20FAF7C7BC9BEAF83D" box="[1341,1461,1970,1995]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
). Endostome distinct, sloping gradually posteriorly into buccal cavern, endostomial ridges, short, low (
<figureCitation id="0D9145C72C46CE20FB38C7FC9B34F7FD" box="[1266,1387,2034,2059]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345">Fig. 6C, D</figureCitation>
).
</paragraph>
<caption id="C1D509CA2C46CE20FF42C6029B9CF9AD" ID-DOI="http://doi.org/10.5281/zenodo.14682775" ID-Zenodo-Dep="14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="10" pageNumber="345" startId="10.[136,171,1548,1570]" targetBox="[135,1473,189,1532]" targetPageId="10" targetType="figure">
<paragraph id="951559422C46CE20FF42C6029B9CF9AD" blockId="10.[136,1475,1548,1627]" pageId="10" pageNumber="345">
Fig. 6.
<taxonomicName id="52AA22C12C46CE20FF18C6029C20F9D4" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[210,639,1548,1570]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="345" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C46CE20FF18C6029FEDF9D7" box="[210,434,1548,1569]" italics="true" pageId="10" pageNumber="345">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C46CE20FE0BC6029C27F9D4" author="Edmondson CH" box="[449,632,1548,1570]" pageId="10" pageNumber="345" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C46CE20FD46C6029D67F9D4" box="[652,824,1548,1570]" pageId="10" pageNumber="345" rank="species">new combination</taxonomicNameLabel>
, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela.
</paragraph>
</caption>
<caption id="C1D509CA2C47CE21FFBBC6949FA8F8F0" ID-DOI="http://doi.org/10.5281/zenodo.14682777" ID-Zenodo-Dep="14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346" startId="11.[113,148,1690,1712]" targetBox="[113,1451,189,1674]" targetPageId="11" targetType="figure">
<paragraph id="951559422C47CE21FFBBC6949FA8F8F0" blockId="11.[113,1452,1690,1798]" pageId="11" pageNumber="346">
Fig. 7.
<taxonomicName id="52AA22C12C47CE21FF76C6949C33F946" authority="(Edmondson, 1951)" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[188,620,1690,1712]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="346" phylum="Arthropoda" rank="species" species="symmetrinuda" status="comb. nov.">
<emphasis id="A7DE85502C47CE21FF76C6949FC1F959" box="[188,414,1690,1711]" italics="true" pageId="11" pageNumber="346">Hosekia symmetrinuda</emphasis>
(
<bibRefCitation id="F13B24B32C47CE21FE64C6949C3AF946" author="Edmondson CH" box="[430,613,1690,1712]" pageId="11" pageNumber="346" pagination="183 - 243" refId="ref13456" refString="Edmondson CH (1951) Some Central Pacific crustaceans. Occasional Papers of the Bernice P. Bishop Museum, 20 (13): 183 - 243, figs. 1 - 38." type="journal article" year="1951">Edmondson, 1951</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C47CE21FDB0C6949D77F946" box="[634,808,1690,1712]" pageId="11" pageNumber="346" rank="species">new combination</taxonomicNameLabel>
. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae.
</paragraph>
</caption>
<paragraph id="951559422C47CE21FFBBC7339C36F843" blockId="11.[113,753,1853,1973]" pageId="11" pageNumber="346">
Third maxilliped with ischium relatively short, subquadrate, length 1.2× width; merus quadrate, anterolateral angle gently auriculiform. Exopod relatively stout, tip reaching to just before distal edge of merus (
<figureCitation id="0D9145C72C47CE21FE66C7939C71F843" box="[428,558,1949,1973]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Figs. 6D, E</figureCitation>
,
<figureCitation id="0D9145C72C47CE21FDF0C7939C02F843" box="[570,605,1949,1973]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">7B</figureCitation>
).
</paragraph>
<paragraph id="951559422C47CE21FFBBC7D39A09F843" blockId="11.[113,752,2013,2102]" lastBlockId="11.[813,1452,1853,1973]" pageId="11" pageNumber="346">
Male thoracic sternum surface relatively smooth; sternites 1 and 2 completely fused, lateral margins gently concave; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 gently concave towards buccal cavity (
<figureCitation id="0D9145C72C47CE21FB25C7539B2EF883" box="[1263,1393,1885,1909]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Figs. 6D, E</figureCitation>
,
<figureCitation id="0D9145C72C47CE21FAB4C7539BC0F883" box="[1406,1439,1885,1909]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">7E</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior third of somite 5 (
<figureCitation id="0D9145C72C47CE21FC3EC7939A16F843" box="[1012,1097,1949,1973]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="11" pageNumber="346">Fig. 6E</figureCitation>
).
</paragraph>
<paragraph id="951559422C47CE26FCE7C7D39CC1FE23" blockId="11.[813,1452,2013,2102]" lastBlockId="12.[136,775,189,470]" lastPageId="12" lastPageNumber="347" pageId="11" pageNumber="346">
Adult male chelipeds symmetrical, stout (
<figureCitation id="0D9145C72C47CE21FB30C7D39B0CF803" box="[1274,1363,2013,2037]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">Fig. 7A</figureCitation>
). Basis-ischium fused but suture visible, flexor margin with low granules (
<figureCitation id="0D9145C72C47CE21FC56C8139DAAF7C3" box="[924,1013,2077,2101]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="11" pageNumber="346">Fig. 7E</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with small granules; inner distal angle sharp visible but not with subdistal angle not elongate (
<figureCitation id="0D9145C72C40CE26FF5AC0F39EBBFEE3" box="[144,228,253,277]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7E</figureCitation>
). Carpus surface granulate; inner distal angle with low sharp tooth (
<figureCitation id="0D9145C72C40CE26FEF0C1139FD0FEC3" box="[314,399,285,309]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7A</figureCitation>
). Chela with dorsal surface densely covered with granules; inner surface almost smooth; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, shorter than palm, pigmented brown along distal ⅔; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C40CE26FDF4C1B39CCEFE23" box="[574,657,445,469]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="12" pageNumber="347">Fig. 6F</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C1F39D58FCE3" blockId="12.[136,775,509,789]" pageId="12" pageNumber="347">
Ambulatory legs short, P4 longest, about 1.2× carapace width; P4 merus length about 2.22.4× height, less than half carapace width; outer surface slightly rugose to almost smooth; coxa rounded; merus short, extensor margin not cristate, surface of P25 gently convex, without ridge; P25 carpus smooth, with low submarginal ridge, lowest in P5; propodus laterally flattened, unarmed, relatively short in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, tip slightly upturned; dactylo-propodal lock distinct (
<figureCitation id="0D9145C72C40CE26FD69C2F39CA4FCE3" box="[675,763,765,789]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C3339F84FC03" blockId="12.[136,775,829,1013]" pageId="12" pageNumber="347">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somites 36 trapezoidal, gradually decreasing in width. Telson wider than long, rounded-subtriangular, lateral margins weakly convex, longer than somite 6 (
<figureCitation id="0D9145C72C40CE26FEB3C3D39F91FC03" box="[377,462,989,1013]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="951559422C40CE26FF42C4139F25FB83" blockId="12.[136,775,1053,1141]" pageId="12" pageNumber="347">G1 strongly sinuous, distally gently curved outwards, tip mucronate; with short row of setae subdistally. G2 sigmoid, about ¼ length of G1.</paragraph>
<paragraph id="951559422C40CE26FF42C4939CF2FA83" blockId="12.[136,775,1181,1397]" pageId="12" pageNumber="347">Female pleon longitudinally ovate; with 6 somites and telson, all free; not completely covering thoracic sternum; somite 1 broadly subrectangular, just reaching base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somite 3 widest, somite 4 as wide as somite 2; somites 36 trapezoidal, gradually decreasing in width; telson triangular with convex lateral margins, longer than somite 6.</paragraph>
<paragraph id="951559422C40CE26FF42C5939F30FA03" blockId="12.[136,775,1437,1525]" pageId="12" pageNumber="347">
Sternopleonal cavity shallow (
<figureCitation id="0D9145C72C40CE26FE0DC5939C48FA43" box="[455,535,1437,1461]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7F</figureCitation>
; vulva large, obliquely ovate, covering anterior ⅔ on sternite 6, without vulvar projections (
<figureCitation id="0D9145C72C40CE26FEDAC5D39F3BFA03" box="[272,356,1501,1525]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="12" pageNumber="347">Fig. 7F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C40CE26FF42C6139D57F943" pageId="12" pageNumber="347" type="discussion">
<paragraph id="951559422C40CE26FF42C6139D57F943" blockId="12.[136,776,1565,1717]" pageId="12" pageNumber="347">
<emphasis id="A7DE85502C40CE26FF42C6139EA8F9C3" bold="true" box="[136,247,1565,1589]" pageId="12" pageNumber="347">Remarks.</emphasis>
<taxonomicName id="52AA22C12C40CE26FF37C6139FADF9C3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" baseAuthorityName="Edmondson" baseAuthorityYear="1951" box="[253,498,1565,1589]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="347" phylum="Arthropoda" rank="species" species="symmetrinuda">
<emphasis id="A7DE85502C40CE26FF37C6139FADF9C3" box="[253,498,1565,1589]" italics="true" pageId="12" pageNumber="347">Hosekia symmetrinuda</emphasis>
</taxonomicName>
is a reef species currently known only from
<collectingCountry id="EDBD19D22C40CE26FE8EC6339FD2F9A3" box="[324,397,1597,1621]" name="Samoa" pageId="12" pageNumber="347">Samoa</collectingCountry>
and southern
<collectingCountry id="EDBD19D22C40CE26FDEAC6309C3FF9A3" box="[544,608,1598,1621]" name="Japan" pageId="12" pageNumber="347">Japan</collectingCountry>
. The specimens from both locations are nearly identical and we are confident they are conspecific. Nothing else is known about its biology except that it occurs in the intertidal zone (
<bibRefCitation id="F13B24B32C40CE26FDF1C6939CA3F943" author="Maenosono T" box="[571,764,1693,1717]" pageId="12" pageNumber="347" pagination="19 - 44" refId="ref13491" refString="Maenosono T (2019) Report on nine rare pilumnid crabs (Crustacea: Decapoda: Brachyura) collected from southern Japan, including three new records. Fauna Ryukyuana, 48: 19 - 44. [In Japanese with English abstract]" type="journal article" year="2019">Maenosono, 2019</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,274 @@
<document id="C008C946234CC7BD498DD3890D51E508" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C4ACE22FC0AC1739CF3F98B" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Lophoplax takakurai Sakai 1935" docType="treatment" docVersion="2" lastPageNumber="343" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="341" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="3B61758A178648071C443B4D33D8CE64" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="07EE2E85C23C0C1DFDC24E5EF7037196">
<mods:title id="A36049B5263BF5E77EEA304C07437F3C">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="0C69E96BDC8164312CF8EC12502D8B62" type="personal">
<mods:role id="D8D360A27DE5E75848A4E4387935F538">
<mods:roleTerm id="CD8CC12874CFAC6955B8E0529A3EE8CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A433AADE573BC0CBC11EDD4AEA31A830">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="42E521EBE3EF792C24D48482162D01DF" type="personal">
<mods:role id="BA5F30457EF20EE17472D0398500E6BA">
<mods:roleTerm id="15BF9D00B2A516E9481F0DD5C7E565EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="66601228CC0FCE4A8F2E20F3DEBAF185">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="7DACA02E224557782EC7A45BC339B295" type="personal">
<mods:role id="75E7AC1EF9FAECED74414842E3DBE037">
<mods:roleTerm id="8238FC333F98BD40CE209771D46880B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D81C47583789D4AF1E01F477F73B8CC">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="56D14B6FBD19FD266FE5D0969131D086">text</mods:typeOfResource>
<mods:relatedItem id="C974FD475B6A952FBA6151E8A96A6281" type="host">
<mods:titleInfo id="C5EB89A311821037CEE8E5A41F83BD7C">
<mods:title id="905FF736A06690A106087917C8D25D37">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="E04E6DE597AE4E1B07396B6E3EBB8376">
<mods:date id="3C81D6CCA828B21782C99E0597536664">2024</mods:date>
<mods:detail id="659C57ABACD4494F1EC669C060781DBE" type="pubDate">
<mods:number id="E0BC1D76FFCE98ECF6F92730F2A97485">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="9B0EE0030A0B661B2B6611C6DE03091E" type="volume">
<mods:number id="2E4707313DEF47233BFD705E4934B214">72</mods:number>
</mods:detail>
<mods:extent id="85615FE7D63065545B5F17D29749F1F7" unit="page">
<mods:start id="8019DDD1C3852F5F75E68E748EA060FD">335</mods:start>
<mods:end id="BAFAFCBC2D145BDE655B518B886E575F">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="D104F2F0C8A70430ACFE003CE59DCFA3">journal article</mods:classification>
<mods:identifier id="43360BF11906CFF024D4797F22EF6C3F" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="B184317E521C21962E544D016BAFC859" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="37FFDEC33BAAF52267F53915775ABF76" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="924677FCECBEA9CF582FD219D40C2470" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C4ACE22FC0AC1739CF3F98B" ID-DOI="http://doi.org/10.5281/zenodo.14682771" ID-Zenodo-Dep="14682771" LSID="urn:lsid:plazi:treatment:1D03E8542C4ACE22FC0AC1739CF3F98B" httpUri="http://treatment.plazi.org/id/1D03E8542C4ACE22FC0AC1739CF3F98B" lastPageId="8" lastPageNumber="343" pageId="6" pageNumber="341">
<subSubSection id="DDB00AC92C4ACE2CFC0AC1739B1AFE63" box="[960,1349,381,405]" pageId="6" pageNumber="341" type="nomenclature">
<paragraph id="951559422C4ACE2CFC0AC1739B1AFE63" blockId="6.[960,1349,381,437]" box="[960,1349,381,405]" pageId="6" pageNumber="341">
<heading id="CE5DEE2E2C4ACE2CFC0AC1739B1AFE63" bold="true" box="[960,1349,381,405]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="341" reason="2">
<taxonomicName id="52AA22C12C4ACE2CFC0AC1739B1AFE63" ID-CoL="3W3WV" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[960,1349,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC0AC1739B1AFE63" bold="true" box="[960,1349,381,405]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC0AC1739AF4FE63" bold="true" box="[960,1195,381,405]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4ACE2CFB79C1739B1AFE63" author="Sakai T" box="[1203,1349,381,405]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFB8CC1939A9FFE43" box="[1094,1216,413,437]" pageId="6" pageNumber="341" type="description">
<paragraph id="951559422C4ACE2CFB8CC1939A9FFE43" blockId="6.[960,1349,381,437]" box="[1094,1216,413,437]" pageId="6" pageNumber="341">
(
<figureCitation id="0D9145C72C4ACE2CFB84C1939AC4FE43" box="[1102,1179,413,437]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Figs. 4</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFB63C1939AE8FE43" box="[1193,1207,413,437]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C1D29B9CFD93" pageId="6" pageNumber="341" type="reference_group">
<paragraph id="951559422C4ACE2CFC89C1D29B9CFD93" blockId="6.[835,1475,476,613]" pageId="6" pageNumber="341">
<treatmentCitationGroup id="B5BA7E6C2C4ACE2CFC89C1D29B9CFD93" pageId="6" pageNumber="341">
<treatmentCitation id="140B7F532C4ACE2CFC89C1D29A9EFE07" author="Sakai T" box="[835,1217,476,498]" page="82" pageId="6" pageNumber="341" year="1935">
<taxonomicName id="52AA22C12C4ACE2CFC89C1D29A9EFE07" ID-CoL="3W3WV" authority="Sakai, 1935 a: 82" authorityName="Sakai" authorityPageNumber="82" authorityYear="1935" box="[835,1217,476,498]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC89C1D29A50FE07" box="[835,1039,476,497]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4ACE2CFBD2C1D29A9EFE07" author="Sakai T" box="[1048,1217,476,498]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, text-fig. 15, pl. 7, fig. 2;
<treatmentCitation id="140B7F532C4ACE2CFCA3C1F79A40FDF8" author="Sakai T" box="[873,1055,505,527]" page="188" pageId="6" pageNumber="341" year="1935">
<bibRefCitation id="F13B24B32C4ACE2CFCA3C1F79A40FDF8" author="Sakai T" box="[873,1055,505,527]" pageId="6" pageNumber="341" refId="ref14541" refString="Sakai T (1935 b) Crabs of Japan. 66 Plates in Life colours with descriptions. Sanseido, Tokyo, 239 pp." type="book" year="1935">Sakai, 1935b: 188</bibRefCitation>
</treatmentCitation>
, frontispiece, fig. 2;
<treatmentCitation id="140B7F532C4ACE2CFB3BC1F79BC3FDF9" author="Sakai T" box="[1265,1436,505,527]" page="567" pageId="6" pageNumber="341" year="1939">
<bibRefCitation id="F13B24B32C4ACE2CFB3BC1F79BC3FDF9" author="Sakai T" box="[1265,1436,505,527]" pageId="6" pageNumber="341" pagination="42 - 111" refId="ref14566" refString="Sakai T (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 [index], pls. 42 - 111." type="book chapter" year="1939">Sakai, 1939: 567</bibRefCitation>
</treatmentCitation>
, pl. 67, fig. l;
<treatmentCitation id="140B7F532C4ACE2CFC05C21B9A29FDDD" author="Sakai T" box="[975,1142,533,555]" page="541" pageId="6" pageNumber="341" year="1976">
<bibRefCitation id="F13B24B32C4ACE2CFC05C21B9A29FDDD" author="Sakai T" box="[975,1142,533,555]" pageId="6" pageNumber="341" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">Sakai, 1976: 541</bibRefCitation>
</treatmentCitation>
pl. 191, fig. 4, text-fig. 288;
<treatmentCitation id="140B7F532C4ACE2CFA55C2189D9FFDB1" author="Ng PKL" page="79" pageId="6" pageNumber="341" year="1987">
<bibRefCitation id="F13B24B32C4ACE2CFA55C2189D9FFDB1" author="Ng PKL" pageId="6" pageNumber="341" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 79</bibRefCitation>
</treatmentCitation>
, 100;
<treatmentCitation id="140B7F532C4ACE2CFC31C23C9AFBFDB1" author="Takeda M" box="[1019,1188,562,584]" page="41" pageId="6" pageNumber="341" year="1993">
<bibRefCitation id="F13B24B32C4ACE2CFC31C23C9AFBFDB1" author="Takeda M" box="[1019,1188,562,584]" pageId="6" pageNumber="341" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda, 1993: 41</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C4ACE2CFB65C23D9B2DFDB1" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1199,1394,562,584]" page="143" pageId="6" pageNumber="341" year="2008">
<bibRefCitation id="F13B24B32C4ACE2CFB65C23D9B2DFDB1" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[1199,1394,562,584]" pageId="6" pageNumber="341" pagination="1 - 286" refId="ref14245" refString="Ng PKL, Guinot D &amp; Davie PJF (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">Ng et al., 2008: 143</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="140B7F532C4ACE2CFAB1C23C9A17FD93" author="Takeda M &amp; Komatsu H" page="176" pageId="6" pageNumber="341" year="2018">
<bibRefCitation id="F13B24B32C4ACE2CFAB1C23C9A17FD93" author="Takeda M &amp; Komatsu H" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018: 176</bibRefCitation>
</treatmentCitation>
, fig. 7;
<treatmentCitation id="140B7F532C4ACE2CFB58C2419BE0FD93" author="Takeda M &amp; Komatsu H" box="[1170,1471,591,613]" page="167" pageId="6" pageNumber="341" year="2023">
<bibRefCitation id="F13B24B32C4ACE2CFB58C2419BE0FD93" author="Takeda M &amp; Komatsu H" box="[1170,1471,591,613]" pageId="6" pageNumber="341" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">Takeda &amp; Komatsu, 2023: 167</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C2839A17FCD3" pageId="6" pageNumber="341" type="materials_examined">
<paragraph id="951559422C4ACE2CFC89C2839A17FCD3" blockId="6.[835,1475,653,805]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C2839A73FD53" bold="true" box="[835,1068,653,677]" pageId="6" pageNumber="341">Material examined.</emphasis>
<materialsCitation id="25C2531F2C4ACE2CFBFCC2839A1BFCD3" collectingDate="2010-07-09" collectionCode="R, V" country="Japan" county="Ogasawara Islands" elevation="42" latitude="27.079334" location="Futami Bay" longLatPrecision="12" longitude="142.1955" municipality="Chichi-jima" pageId="6" pageNumber="341" specimenCount="1" specimenCount-male="1">
<specimenCount id="83AC92CB2C4ACE2CFBFCC2839ADCFD53" box="[1078,1155,653,677]" count="1" pageId="6" pageNumber="341" type="male">1 male</specimenCount>
(5.0 ×
<quantity id="5252F4A72C4ACE2CFB14C2839B68FD53" box="[1246,1335,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="341" unit="mm" value="4.2">4.2 mm</quantity>
) (NSMT-Cr S 1254),
<location id="90750F992C4ACE2CFC67C2A39A6DFD30" LSID="urn:lsid:plazi:treatment:1D03E8542C4ACE22FC0AC1739CF3F98B:90750F992C4ACE2CFC67C2A39A6DFD30" box="[941,1074,685,710]" country="Japan" county="Ogasawara Islands" latitude="27.079334" longLatPrecision="12" longitude="142.1955" municipality="Chichi-jima" name="Futami Bay" pageId="6" pageNumber="341">Futami Bay</location>
,
<collectingMunicipality id="7571C3382C4ACE2CFB8AC2A39A9EFD33" box="[1088,1217,685,709]" pageId="6" pageNumber="341">Chichi-jima</collectingMunicipality>
[= Island],
<collectingCounty id="7C7421CE2C4ACE2CFA81C2A39DC9FD13" pageId="6" pageNumber="341">Ogasawara Islands</collectingCounty>
,
<collectingCountry id="EDBD19D22C4ACE2CFC6DC2C09DB5FD13" box="[935,1002,718,741]" name="Japan" pageId="6" pageNumber="341">Japan</collectingCountry>
,
<geoCoordinate id="F09E3F852C4ACE2CFC30C2C29A24FD13" box="[1018,1147,716,741]" degrees="27" direction="north" minutes="04.77" orientation="latitude" pageId="6" pageNumber="341" precision="9" value="27.0795">27°04.77N</geoCoordinate>
<geoCoordinate id="F09E3F852C4ACE2CFB4DC2C29B4BFD13" box="[1159,1300,716,741]" degrees="142" direction="east" minutes="11.68" orientation="longitude" pageId="6" pageNumber="341" precision="9" value="142.19467">142°11.68E</geoCoordinate>
to
<geoCoordinate id="F09E3F852C4ACE2CFA8BC2C29B9DFD13" box="[1345,1474,716,741]" degrees="27" direction="north" minutes="04.76" orientation="latitude" pageId="6" pageNumber="341" precision="9" value="27.079334">27°04.76N</geoCoordinate>
<geoCoordinate id="F09E3F852C4ACE2CFC8EC2E29D93FCF3" box="[836,972,748,773]" degrees="142" direction="east" minutes="11.73" orientation="longitude" pageId="6" pageNumber="341" precision="9" value="142.1955">142°11.73E</geoCoordinate>
, 42.2
<quantity id="5252F4A72C4ACE2CFBDEC2E39A3DFCF3" box="[1044,1122,749,773]" metricMagnitude="1" metricUnit="m" metricValue="4.16" pageId="6" pageNumber="341" unit="m" value="41.6">
<elevation id="1E87BE712C4ACE2CFBDEC2E39A3DFCF3" box="[1044,1122,749,773]" metricMagnitude="1" metricUnit="m" metricValue="4.16" pageId="6" pageNumber="341" unit="m" value="41.6">41.6 m</elevation>
</quantity>
,
<collectionCode id="F3BBC1872C4ACE2CFBA5C2E39ADEFCF3" box="[1135,1153,749,773]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="341">R</collectionCode>
/
<collectionCode id="F3BBC1872C4ACE2CFB43C2E39AC3FCF3" box="[1161,1180,749,773]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="341" type="Museum">V</collectionCode>
Koyo, 2010 cruise, station KY-10-25,
<date id="E1147F822C4ACE2CFC0AC3039A1BFCD3" box="[960,1092,781,805]" pageId="6" pageNumber="341" value="2010-07-09">
<collectingDate id="F150866A2C4ACE2CFC0AC3039A1BFCD3" box="[960,1092,781,805]" pageId="6" pageNumber="341" value="2010-07-09">9 July 2010</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE2CFC89C3439AB9FAD3" pageId="6" pageNumber="341" type="diagnosis">
<paragraph id="951559422C4ACE2CFC89C3439AB9FAD3" blockId="6.[835,1475,845,1317]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C3439DE7FC93" bold="true" box="[835,952,845,869]" pageId="6" pageNumber="341">Diagnosis.</emphasis>
Carapace frontal margin gently convex, smooth, produced anteriorly, lateral lobe not visible (
<figureCitation id="0D9145C72C4ACE2CFAF2C3639BEBFC73" box="[1336,1460,877,901]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4A, B</figureCitation>
); epigastric areole barely visible; protogastric lobe low, with granular surface (
<figureCitation id="0D9145C72C4ACE2CFBCEC3A39A03FC33" box="[1028,1116,941,965]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4B</figureCitation>
); metabranchial region with low granules (
<figureCitation id="0D9145C72C4ACE2CFC7EC3C39A4FFC13" box="[948,1040,973,997]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4B</figureCitation>
); median lobe on posterior margin of epistome subtruncate in form (
<figureCitation id="0D9145C72C4ACE2CFB5AC3E39AB1FBF3" box="[1168,1262,1005,1029]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Figs. 4E</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFB30C3E39B45FBF3" box="[1274,1306,1005,1029]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">5E</figureCitation>
). Distal margin of ischium of third maxilliped with row of closely set low granules (
<figureCitation id="0D9145C72C4ACE2CFC67C4239A5CFBB3" box="[941,1027,1069,1093]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4C</figureCitation>
). P25 merus extensor margin P25 with low proximal tooth (
<figureCitation id="0D9145C72C4ACE2CFBE4C4439AF5FB93" box="[1070,1194,1101,1125]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5F, G</figureCitation>
). Cheliped carpus dorsal surface with longitudinal areole separated by groove from swollen lateral margin (
<figureCitation id="0D9145C72C4ACE2CFB82C4839AE7FB53" box="[1096,1208,1165,1189]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4A, I</figureCitation>
); proximal basal part of male chela covered with strong tubercles (
<figureCitation id="0D9145C72C4ACE2CFAD1C4A39B2AFB33" box="[1307,1397,1197,1221]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4G</figureCitation>
). Male thoracic sternum anteriorly relatively narrower (
<figureCitation id="0D9145C72C4ACE2CFAF5C4C39BCFFB13" box="[1343,1424,1229,1253]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4F</figureCitation>
). G1 strongly sinuous, distal part of G1 gently recurved (
<figureCitation id="0D9145C72C4ACE2CFAB8C4E39D0FFAD3" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5I, J</figureCitation>
). G2 sigmoid, about ¼ length of G1.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4ACE22FC89C5439CF3F98B" lastPageId="8" lastPageNumber="343" pageId="6" pageNumber="341" type="discussion">
<paragraph id="951559422C4ACE2CFC89C5439A1EF850" blockId="6.[835,1475,1357,1958]" pageId="6" pageNumber="341">
<emphasis id="A7DE85502C4ACE2CFC89C5439DEDFA93" bold="true" box="[835,946,1357,1381]" pageId="6" pageNumber="341">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4ACE2CFC70C5439AC5FA93" authorityName="Sakai" authorityYear="1935" box="[954,1178,1357,1381]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC70C5439AC5FA93" box="[954,1178,1357,1381]" italics="true" pageId="6" pageNumber="341">Lophoplax takakurai</emphasis>
</taxonomicName>
was described on the basis of a female (11.0 × 9.0 mm) collected off Yoshihama, west of Sagami Bay, and later reported from off Mikawa Bay (south of Nagoya) (
<bibRefCitation id="F13B24B32C4ACE2CFBD1C5A39AF4FA33" author="Sakai T" box="[1051,1195,1453,1477]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
, b, 1976).
<bibRefCitation id="F13B24B32C4ACE2CFAE9C5A39B9DFA33" author="Takeda M" box="[1315,1474,1453,1477]" pageId="6" pageNumber="341" refId="ref14889" refString="Takeda M (1993) Catalogue of crab specimens (Crustacea: Decapoda: Brachyura). Part 1. National Science Museum, Tokyo, 93 pp." type="book" year="1993">Takeda (1993)</bibRefCitation>
subsequently recorded the species from Shibushi Bay in
<collectingRegion id="576E97A02C4ACE2CFC8EC5E39D9FF9F3" box="[836,960,1517,1541]" country="Japan" name="Kagoshima" pageId="6" pageNumber="341">Kagoshima</collectingRegion>
, with
<bibRefCitation id="F13B24B32C4ACE2CFC37C5E39B4FF9F3" author="Takeda M &amp; Komatsu H" box="[1021,1296,1517,1541]" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu (2018</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFAD3C5E39B0FF9F3" author="Takeda M &amp; Komatsu H" box="[1305,1360,1517,1541]" pageId="6" pageNumber="341" pagination="161 - 178" refId="ref14998" refString="Takeda M &amp; Komatsu H (2023) Updated checklist of the crabs (Crustacea: Decapoda: Brachyura) from the sea around the Ogasawara Islands, Japan. Bulletin of the National Museum of Nature and Science, Series A, 49 (4): 161 - 178." type="journal article" year="2023">2023</bibRefCitation>
) recording it from the Ogasawara Islands. The whereabouts of the type specimen is not known but appears to be lost. The present specimen matches the type description and figures of
<bibRefCitation id="F13B24B32C4ACE2CFA4DC6439DD0F973" author="Sakai T" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai (1935a</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFC50C6639D8DF973" author="Sakai T" box="[922,978,1645,1669]" pageId="6" pageNumber="341" pagination="42 - 111" refId="ref14566" refString="Sakai T (1939) Studies on the Crabs of Japan. Brachygnatha, Brachyrhyncha, IV. Yokendo, Tokyo, pp. 365 - 741 + 1 - 23 [index], pls. 42 - 111." type="book chapter" year="1939">1939</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4ACE2CFC17C6639A4BF973" author="Sakai T" box="[989,1044,1645,1669]" pageId="6" pageNumber="341" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">1976</bibRefCitation>
) well, especially in the general carapace features and proportions of the ambulatory legs. Significantly,
<bibRefCitation id="F13B24B32C4ACE2CFC89C6A39D84F933" author="Sakai T" box="[835,987,1709,1733]" pageId="6" pageNumber="341" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai (1935a</bibRefCitation>
: text-fig. 15c; present
<figureCitation id="0D9145C72C4ACE2CFB15C6A39B6AF933" box="[1247,1333,1709,1733]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="6" pageNumber="341">Fig. 5B</figureCitation>
) figured the third maxilliped showing a row of closely set granules on the distal margin of the ischium, a character also distinct in the present male specimen (
<figureCitation id="0D9145C72C4ACE2CFB57C7039AA9F8D3" box="[1181,1270,1805,1829]" captionStart="Fig" captionStartId="7.[113,148,2035,2057]" captionTargetBox="[168,1396,189,2020]" captionTargetId="figure-7@7.[168,1396,189,2020]" captionTargetPageId="7" captionText="Fig. 4. Lophoplax takakurai Sakai, 1935a, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped." figureDoi="http://doi.org/10.5281/zenodo.14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="6" pageNumber="341">Fig. 4C</figureCitation>
). As such, we are confident of its identity. For the moment,
<taxonomicName id="52AA22C12C4ACE2CFADCC7209BC4F8B3" authorityName="Sakai" authorityYear="1935" box="[1302,1435,1837,1861]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFADCC7209BC4F8B3" box="[1302,1435,1837,1861]" italics="true" pageId="6" pageNumber="341">L. takakurai</emphasis>
</taxonomicName>
, as defined by
<bibRefCitation id="F13B24B32C4ACE2CFC73C7439A89F893" author="Takeda M &amp; Komatsu H" box="[953,1238,1869,1893]" pageId="6" pageNumber="341" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu (2018)</bibRefCitation>
and the present study, is clearly recognisable, so a
<typeStatus id="4A11E7E02C4ACE2CFB4BC7609A89F870" box="[1153,1238,1902,1926]" pageId="6" pageNumber="341" type="neotype">neotype</typeStatus>
designation does not appear to be necessary.
</paragraph>
<paragraph id="951559422C4ACE22FC89C7C39C82F9EB" blockId="6.[835,1475,1997,2085]" lastBlockId="8.[136,775,1477,1565]" lastPageId="8" lastPageNumber="343" pageId="6" pageNumber="341">
Although
<taxonomicName id="52AA22C12C4ACE2CFC78C7C09A68F813" authorityName="Sakai" authorityYear="1935" box="[946,1079,1997,2021]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4ACE2CFC78C7C09A68F813" box="[946,1079,1997,2021]" italics="true" pageId="6" pageNumber="341">L. takakurai</emphasis>
</taxonomicName>
differs markedly from
<taxonomicName id="52AA22C12C4ACE2CFAF7C7C09B9DF813" authorityName="Tesch" authorityYear="1918" box="[1341,1474,1997,2021]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4ACE2CFAF7C7C09B9DF813" box="[1341,1474,1997,2021]" italics="true" pageId="6" pageNumber="341">L. bicristata</emphasis>
</taxonomicName>
in many characters of the carapace and pereopods, both species have generally similar carapace shape and features. In addition, both species have a stouter G1 (
<figureCitation id="0D9145C72C44CE22FD9DC5CB9C98FA2B" box="[599,711,1477,1501]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="8" pageNumber="343">Figs. 3FI</figureCitation>
,
<figureCitation id="0D9145C72C44CE22FD1BC5CB9CA0FA2B" box="[721,767,1477,1501]" captionStart="Fig" captionStartId="8.[136,171,1314,1336]" captionTargetBox="[134,1474,187,1303]" captionTargetId="figure-8@8.[124,1482,179,1307]" captionTargetPageId="8" captionText="Fig. 5. Lophoplax takakurai Sakai, 1935a. AD, holotype male (11.5 × 9.0 mm) (after Sakai, 1935a: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="8" pageNumber="343">5I, J</figureCitation>
) compared to more typical pilumnids. It is therefore reasonable to retain both species in
<taxonomicName id="52AA22C12C44CE22FE5CC60B9C57F9EB" authorityName="Tesch" authorityYear="1918" box="[406,520,1541,1565]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C44CE22FE5CC60B9C57F9EB" box="[406,520,1541,1565]" italics="true" pageId="8" pageNumber="343">Lophoplax</emphasis>
</taxonomicName>
for the time being.
</paragraph>
<caption id="C1D509CA2C4BCE2DFFBBC7FD9D55F7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682769" ID-Zenodo-Dep="14682769" httpUri="https://zenodo.org/record/14682769/files/figure.png" pageId="7" pageNumber="342" startId="7.[113,148,2035,2057]" targetBox="[168,1396,189,2020]" targetPageId="7" targetType="figure">
<paragraph id="951559422C4BCE2DFFBBC7FD9D55F7B5" blockId="7.[113,1452,2035,2115]" pageId="7" pageNumber="342">
Fig. 4.
<taxonomicName id="52AA22C12C4BCE2DFF7CC7FD9C5BF7FF" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[182,516,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="342" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4BCE2DFF7CC7FD9F20F7FE" box="[182,383,2035,2056]" italics="true" pageId="7" pageNumber="342">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C4BCE2DFE4FC7FD9C5BF7FF" author="Sakai T" box="[389,516,2035,2057]" pageId="7" pageNumber="342" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, dorsal habitus (right side partially brushed); B, carapace (right side denuded); C, left third maxilliped; D, frontal view of cephalothorax; E, epistome; F, anterior thoracic sternum and pleon; G, right chela; H, left chela; I, dorsal view of right cheliped.
</paragraph>
</caption>
<caption id="C1D509CA2C44CE22FF42C52C9D4FFA78" ID-DOI="http://doi.org/10.5281/zenodo.14682773" ID-Zenodo-Dep="14682773" httpUri="https://zenodo.org/record/14682773/files/figure.png" pageId="8" pageNumber="343" startId="8.[136,171,1314,1336]" targetBox="[134,1474,187,1303]" targetPageId="8" targetType="figure">
<paragraph id="951559422C44CE22FF42C52C9D4FFA78" blockId="8.[136,1475,1314,1422]" pageId="8" pageNumber="343">
Fig. 5.
<taxonomicName id="52AA22C12C44CE22FF05C52C9C41FACE" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" box="[207,542,1314,1336]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C44CE22FF05C52C9FC6FAC1" box="[207,409,1314,1335]" italics="true" pageId="8" pageNumber="343">Lophoplax takakurai</emphasis>
<bibRefCitation id="F13B24B32C44CE22FE55C52C9C41FACE" author="Sakai T" box="[415,542,1314,1336]" pageId="8" pageNumber="343" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
. AD, holotype male (11.5 × 9.0 mm) (after
<bibRefCitation id="F13B24B32C44CE22FC1DC52C9A08FACE" author="Sakai T" box="[983,1111,1314,1336]" pageId="8" pageNumber="343" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
: text-fig. 15; pl. 7, fig. 2); EK, male (5.0 × 4.2 mm) (NSMT-Cr S 1254), Japan. A, overall dorsal habitus; B, right third maxilliped; C, left chela; D, female pleon; E, epistome; F, left P4; G, right P5; H, pleon, showing exposed sternite 8 lateral to somite 2; I, right G1 (ventral view); J, right G1 (dorsal view); K, right G2. Scales: E, H = 0.5 mm; F, G = 1.0 mm; IK = 0.2 mm.
</paragraph>
</caption>
<paragraph id="951559422C44CE22FF42C64B9CF3F98B" blockId="8.[136,775,1605,1661]" pageId="8" pageNumber="343">
<taxonomicName id="52AA22C12C44CE22FF42C64B9F38F9AB" authorityName="Sakai" authorityYear="1935" box="[136,359,1605,1629]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="343" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C44CE22FF42C64B9F38F9AB" box="[136,359,1605,1629]" italics="true" pageId="8" pageNumber="343">Lophoplax takakurai</emphasis>
</taxonomicName>
occurs on muddy bottoms
<quantity id="5252F4A72C44CE22FD42C64B9D58F9AB" box="[648,775,1605,1629]" metricMagnitude="1" metricUnit="m" metricValue="7.08" metricValueMax="10.0" metricValueMin="4.16" pageId="8" pageNumber="343" unit="m" value="70.8" valueMax="100.0" valueMin="41.6">41.6100 m</quantity>
in depth (
<bibRefCitation id="F13B24B32C44CE22FF3BC66B9F29F98B" author="Sakai T" box="[241,374,1637,1661]" pageId="8" pageNumber="343" pagination="1 - 16" refId="ref14607" refString="Sakai T (1976) Crabs of Japan and the Adjacent Seas. In three volumes; English Text, pp. xxix + 773 pp., Japanese Text, pp. 1 - 461, Plates volume, pp. 1 - 16, pls. 1 - 251. Kodansha Ltd., Tokyo." type="book chapter" year="1976">Sakai, 1976</bibRefCitation>
;
<bibRefCitation id="F13B24B32C44CE22FE48C66B9CFFF98B" author="Takeda M &amp; Komatsu H" box="[386,672,1637,1661]" pageId="8" pageNumber="343" pagination="153 - 189" refId="ref14949" refString="Takeda M &amp; Komatsu H (2018) Offshore crabs of the family Xanthidae and some related families (Crustacea, Decapoda, Brachyura) from the Ogasawara Islands, Japan. Memoirs of the National Museum of Natural Science, Tokyo, 52: 153 - 189." type="journal article" year="2018">Takeda &amp; Komatsu, 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,269 @@
<document id="FCA763F0EB1ACFDA7CF5C76986F16199" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C4DCE29FC23C5939A6CFD63" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Lophoplax Tesch 1918" docType="treatment" docVersion="2" lastPageNumber="338" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="336" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8DB176D7AC69AD26D19F8E48638ECB15" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BA329E482E3FE2B213BCDF06FB6E9DD6">
<mods:title id="29727D2A0DC1DAE852D6130A155C34CC">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="CD2A20C95EFF95DCC42C79D92099491B" type="personal">
<mods:role id="420C12A05829A5EE25A3A2CC6C011985">
<mods:roleTerm id="350AEDA051362BE53100594C7327A34A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72E7BF83B20C3A10165E039BED95F0DC">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="DC8E7EEA406DCEBBFE4FB4641A3937D4" type="personal">
<mods:role id="C1726E4AFC4870444F947175E598273C">
<mods:roleTerm id="78B36E6B6AD4A06819FB6FF1B35A1B4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B5237E3A86140104D449D5FD267681D">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="420EB8F79B0B6204E0ADD9E94BD512B5" type="personal">
<mods:role id="366375F5209CE53F5F2A83403E7359D5">
<mods:roleTerm id="84E75F154AA8805C437D8CD8D64AB4EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FFB88F02CE53F6CFA05F3DCBD23C4E7A">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0A5E5A384C3044C9D664630ECC540A19">text</mods:typeOfResource>
<mods:relatedItem id="F6A579C1F4C3B671A9F42A6403A4E95E" type="host">
<mods:titleInfo id="C648F10EDD85A1A3EBB34E5AD847C2E8">
<mods:title id="0DC9D6634EDD27AC954062712655510B">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="26057BDD41DF3D6BC7EF301066A3EB6D">
<mods:date id="1D349A53DC6085D681CB838130CF0A04">2024</mods:date>
<mods:detail id="4C697793858B780D239A1EEB48627863" type="pubDate">
<mods:number id="CF01A903DFD37A3C82EA4B30E5D12062">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="0C88E66C3918E4BA4381B274498D8555" type="volume">
<mods:number id="43834B4D50887F44BDA24E7B56803CBC">72</mods:number>
</mods:detail>
<mods:extent id="E3086DE17CEC8FAD3094064BAAF9A2A8" unit="page">
<mods:start id="A7D32E90FFC4D29E6DF3A2F8054B09C1">335</mods:start>
<mods:end id="98923269127A74E2D0C6613225AD3403">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="02DAA15E0D75961A78D6B8AFF4F5E84F">journal article</mods:classification>
<mods:identifier id="F29428A5BE11A5CF605C42BB05D756B7" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="0370172153DDAD4AF64DC54359324013" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E6052FB4476BDC771F1003E02EB12527" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="DAB7FC3E68B57E4B50938A8FC35F0084" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C4DCE29FC23C5939A6CFD63" LSID="urn:lsid:plazi:treatment:1D03E8542C4DCE29FC23C5939A6CFD63" httpUri="http://treatment.plazi.org/id/1D03E8542C4DCE29FC23C5939A6CFD63" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336">
<subSubSection id="DDB00AC92C4DCE2BFC23C5939AB0FA43" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336" type="nomenclature">
<paragraph id="951559422C4DCE2BFC23C5939AB0FA43" blockId="1.[1001,1263,1437,1461]" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336">
<heading id="CE5DEE2E2C4DCE2BFC23C5939AB0FA43" bold="true" box="[1001,1263,1437,1461]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="336" reason="2">
<taxonomicName id="52AA22C12C4DCE2BFC23C5939AB0FA43" ID-CoL="7P7TN" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[1001,1263,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4DCE2BFC23C5939AB0FA43" bold="true" box="[1001,1263,1437,1461]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFC23C5939A3FFA43" bold="true" box="[1001,1120,1437,1461]" italics="true" pageId="1" pageNumber="336">Lophoplax</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFBA2C5939AB0FA43" author="Tesch JJ" box="[1128,1263,1437,1461]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE2BFCE7C5D29B09F9DD" pageId="1" pageNumber="336" type="reference_group">
<paragraph id="951559422C4DCE2BFCE7C5D29B09F9DD" blockId="1.[813,1452,1500,1579]" pageId="1" pageNumber="336">
<treatmentCitationGroup id="B5BA7E6C2C4DCE2BFCE7C5D29B09F9DD" pageId="1" pageNumber="336">
<treatmentCitation id="140B7F532C4DCE2BFCE7C5D29A1CFA04" author="Tesch JJ" box="[813,1091,1500,1522]" page="196" pageId="1" pageNumber="336" year="1918">
<taxonomicName id="52AA22C12C4DCE2BFCE7C5D29A1CFA04" ID-CoL="7P7TN" authority="Tesch, 1918: 196" authorityName="Tesch" authorityPageNumber="196" authorityYear="1918" box="[813,1091,1500,1522]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4DCE2BFCE7C5D29DCBFA07" box="[813,916,1500,1521]" italics="true" pageId="1" pageNumber="336">Lophoplax</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFC51C5D29A1CFA04" author="Tesch JJ" box="[923,1091,1500,1522]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918: 196</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFB44C5D29B45FA07" author="Ng PKL" box="[1166,1306,1500,1521]" page="100" pageId="1" pageNumber="336" year="1987">
<bibRefCitation id="F13B24B32C4DCE2BFB44C5D29B45FA07" author="Ng PKL" box="[1166,1306,1500,1521]" pageId="1" pageNumber="336" pagination="69 - 111" refId="ref13788" refString="Ng PKL (1987) The IndoPacific Pilumnidae II. A revision of the genus Rhizopa Stimpson, 1858 and the status of the Rhizopinae Stimpson, 1858 (Crustacea: Decapoda: Brachyura). IndoMalayan Zoology, 1987, 4 (1): 69 - 111, pl. 1." type="journal article" year="1987">Ng, 1987: 100</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFAAFC5D29DAEF9F8" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" page="595" pageId="1" pageNumber="336" year="2022">
<bibRefCitation id="F13B24B32C4DCE2BFAAFC5D29DAEF9F8" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" pageId="1" pageNumber="336" pagination="595 - 600" refId="ref15186" refString="Trivedi J, Patel K, Mitra S &amp; Ng PKL (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4): 595 - 600." type="journal article" year="2022">Trivedi et al., 2022: 595</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation id="140B7F532C4DCE2BFBF1C5F79B14F9F9" author="Poore GCB &amp; Ahyong ST" box="[1083,1355,1529,1551]" page="667" pageId="1" pageNumber="336" year="2023">
<bibRefCitation id="F13B24B32C4DCE2BFBF1C5F79B14F9F9" author="Poore GCB &amp; Ahyong ST" box="[1083,1355,1529,1551]" pageId="1" pageNumber="336" refId="ref14446" refString="Poore GCB &amp; Ahyong ST (2023) Marine decapod Crustacea: a guide to the families and genera of the world. CSIRO Publishing, Collingwood, Australia, xii + 890 pp." type="book" year="2023">Poore &amp; Ahyong 2023: 667</bibRefCitation>
</treatmentCitation>
, 670, 680 (part), fig. 14.112u;
<treatmentCitation id="140B7F532C4DCE2BFBD3C6189B4DF9DD" author="Ng PKL &amp; Rahayu DL" box="[1049,1298,1557,1579]" page="429" pageId="1" pageNumber="336" year="2023">
<bibRefCitation id="F13B24B32C4DCE2BFBD3C6189B4DF9DD" author="Ng PKL &amp; Rahayu DL" box="[1049,1298,1557,1579]" pageId="1" pageNumber="336" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023: 429</bibRefCitation>
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE2BFCE7C65A9A70F91D" pageId="1" pageNumber="336" type="type_taxon">
<paragraph id="951559422C4DCE2BFCE7C65A9A85F97A" blockId="1.[813,1451,1619,1676]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C65A9D9DF99D" bold="true" box="[813,962,1619,1644]" pageId="1" pageNumber="336">
<typeStatus id="4A11E7E02C4DCE2BFCE7C65A9D38F99A" box="[813,871,1620,1644]" pageId="1" pageNumber="336">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C4DCE2BFC0DC65A9B71F99A" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[967,1326,1620,1644]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4DCE2BFC0DC65A9AFAF99A" box="[967,1189,1620,1644]" italics="true" pageId="1" pageNumber="336">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFB60C65A9B71F99A" author="Tesch JJ" box="[1194,1326,1620,1644]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, by original designation. Gender of genus feminine.
</paragraph>
<paragraph id="951559422C4DCE2BFCE7C6BA9A70F91D" blockId="1.[813,1452,1715,1772]" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C6BA9DA7F93D" bold="true" box="[813,1016,1715,1740]" pageId="1" pageNumber="336">Included species.</emphasis>
<taxonomicName id="52AA22C12C4DCE2BFBC9C6BA9BD9F93A" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[1027,1414,1716,1740]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4DCE2BFBC9C6BA9AB2F93A" box="[1027,1261,1716,1740]" italics="true" pageId="1" pageNumber="336">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFB32C6BA9BD9F93A" author="Tesch JJ" box="[1272,1414,1716,1740]" pageId="1" pageNumber="336" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="52AA22C12C4DCE2BFA5CC6BA9A75F91D" ID-CoL="3W3WV" authority="Sakai, 1935 a" authorityName="Sakai" authorityYear="1935" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4DCE2BFA5CC6BA9DCCF91A" italics="true" pageId="1" pageNumber="336">L. takakurai</emphasis>
<bibRefCitation id="F13B24B32C4DCE2BFC56C6DA9A75F91D" author="Sakai T" box="[924,1066,1747,1772]" pageId="1" pageNumber="336" pagination="63 - 88" refId="ref14481" refString="Sakai T (1935 a) New or rare species of Brachyura, collected by the &quot; Misago &quot; during the Zoological Survey around the Izu- Peninsula. Science Reports of the Tokyo Bunrika Daigaku, Section B, 2 (32): 63 - 88, fig. 1 - 17, pls. 6 - 8." type="journal article" year="1935">Sakai, 1935a</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4DCE29FCE7C71D9F69FC63" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336" type="diagnosis">
<paragraph id="951559422C4DCE29FCE7C71D9F69FC63" blockId="1.[812,1452,1811,2092]" lastBlockId="3.[113,754,189,917]" lastPageId="3" lastPageNumber="338" pageId="1" pageNumber="336">
<emphasis id="A7DE85502C4DCE2BFCE7C71D9DFAF8DD" bold="true" box="[813,933,1811,1835]" pageId="1" pageNumber="336">Diagnosis.</emphasis>
Carapace subquadrate; only protogastric and intestinal areoles visible; epigastric region gently convex, smooth but setose, not areolate; cardiac region raised but not areolate, granulate and setose; hepatic areole absent, with only tubercle present; intestinal region forming small areole, adjacent areas slightly raised, granulate and setose, no visible intestino-branchial areole; surface adjacent to posterior carapace margin gently concave, not raised, not forming ridge, no transverse channel visible; orbit clearly visible in dorsal view, supraorbital margin breaking general carapace outline, forming distinct concavity; eyes freely movable, ocular peduncle relatively long; lateral lobe of frontal margin low, indistinct; anterolateral margin with 4 teeth (including external orbital tooth), with 2 tubercles on posterolateral margin; posterolateral margins gently converging towards posterior carapace margin. Endostome not obliquely recessed posteriorly into buccal cavern, subparallel to adjacent pterygostomial surface, low, short endostomial ridges present. Posterior margin of epistome with distinct median lobe, separated from lateral lobes by distinct fissure. Third maxilliped ischium subquadrate, length about 1.2× width. Adult male chelipeds slightly asymmetrical; dorsal surface of carpus with quadrate areole with raised margins; chela relatively enlarged with external median surface smooth. Ambulatory legs elongate, slender, P4 longest, about 1.8× carapace width; merus without obvious crest on extensor margin, instead P24 lined with low, broad well-spaced setose prominences or low tooth; P5 extensor margin smooth, outer surface smooth, without medio-longitudinal swelling; P4 merus length about 5× height, distinctly longer than half carapace width. G1 sinuous, relatively stout, tip strongly recurved.
</paragraph>
</subSubSection>
<caption id="C1D509CA2C4ECE28FF4DC7939FA0F7B4" ID-DOI="http://doi.org/10.5281/zenodo.14682763" ID-Zenodo-Dep="14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="2" pageNumber="337" startId="2.[135,170,1949,1971]" targetBox="[176,1433,189,1933]" targetPageId="2" targetType="figure">
<paragraph id="951559422C4ECE28FF4DC7939FA0F7B4" blockId="2.[135,1474,1948,2115]" pageId="2" pageNumber="337">
Fig. 1. Terminology of carapace structures (based on
<taxonomicName id="52AA22C12C4ECE28FD5AC7939A47F844" authority="(Stimpson, 1858))" authorityName="Stimpson" authorityYear="1858" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[656,1048,1948,1971]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="sculpta">
<emphasis id="A7DE85502C4ECE28FD5AC7939D00F844" box="[656,863,1949,1970]" italics="true" pageId="2" pageNumber="337">Myopilumnus sculpta</emphasis>
(
<bibRefCitation id="F13B24B32C4ECE28FCA6C7939A56F844" author="Stimpson W" box="[876,1033,1948,1971]" pageId="2" pageNumber="337" pagination="93 - 110" refId="ref14833" refString="Stimpson W (1858) Prodromus descriptionis animalium evertebratorum in expeditione ad Oceanum Pacificum Septentrionalem missa, C. Ringgold et Johanne Rodgers ducibus, observatorum et descriptorum. Pars V. Crustacea Ocypodoidea. Proceedings of the Academy of Natural Sciences of Philadelphia, 10: 93 - 110 [39 - 56]." type="journal article" year="1858">Stimpson, 1858</bibRefCitation>
))
</taxonomicName>
. A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14.
</paragraph>
</caption>
<subSubSection id="DDB00AC92C4FCE29FFBBC3B39A6CFD63" pageId="3" pageNumber="338" type="discussion">
<paragraph id="951559422C4FCE29FFBBC3B39F27F8A3" blockId="3.[113,754,957,1877]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FFBBC3B39EBFFC23" bold="true" box="[113,224,957,981]" pageId="3" pageNumber="338">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4FCE29FF2CC3B39F07FC23" authorityName="Tesch" authorityYear="1918" box="[230,344,957,981]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="A7DE85502C4FCE29FF2CC3B39F07FC23" box="[230,344,957,981]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCED382B2C4FCE29FE95C3B09FBBFC23" box="[351,484,957,981]" pageId="3" pageNumber="338" sensu="stricto">sensu stricto</taxonomicNameLabel>
is distinct in its carapace ornamentation and the length and slenderness of the ambulatory legs.
<taxonomicName id="52AA22C12C4FCE29FEE3C3F39FC4FBE3" authorityName="Tesch" authorityYear="1918" box="[297,411,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FEE3C3F39FC4FBE3" box="[297,411,1021,1045]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
is separable from
<taxonomicName id="52AA22C12C4FCE29FD95C3F39CAFFBE3" authorityName="Deb" authorityYear="1989" box="[607,752,1021,1045]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FD95C3F39CAFFBE3" box="[607,752,1021,1045]" italics="true" pageId="3" pageNumber="338">Myopilumnus</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C4FCE29FF68C4139EA5FBC3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[162,250,1053,1077]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C4FCE29FF68C4139EA5FBC3" box="[162,250,1053,1077]" italics="true" pageId="3" pageNumber="338">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C4FCE29FEC6C4109FDBFBC0" box="[268,388,1054,1078]" pageId="3" pageNumber="338" rank="genus">new genus</taxonomicNameLabel>
, in the following characters: the hepatic surface of the carapace is ornamented only with a tubercle (
<figureCitation id="0D9145C72C4FCE29FF13C4539F6EFB83" box="[217,305,1117,1141]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus with a large, prominent areole; cf.
<figureCitation id="0D9145C72C4FCE29FF5EC4739EADFB63" box="[148,242,1149,1173]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FF37C4739F1FFB63" box="[253,320,1149,1173]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); the intestino-branchial areole is absent with only a low intestinal areole visible (
<figureCitation id="0D9145C72C4FCE29FDF1C4939CCDFB43" box="[571,658,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus intestino-branchial areole large and distinct; cf.
<figureCitation id="0D9145C72C4FCE29FD43C4B39CB3FB23" box="[649,748,1213,1237]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FFBBC4D39EECFB03" box="[113,179,1245,1269]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); the surface adjacent to the posterior carapace margin is relatively wide, not forming a channel as no intestino-branchial areole is present (
<figureCitation id="0D9145C72C4FCE29FE6DC5139FA1FAC3" box="[423,510,1309,1333]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
) (versus with distinct channel between intestino-branchial areole and posterior carapace margin; cf.
<figureCitation id="0D9145C72C4FCE29FE93C5539FE6FA83" box="[345,441,1373,1397]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FE0DC5539C53FA83" box="[455,524,1373,1397]" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
); and the absence of a dorsal crest on the ambulatory merus (
<figureCitation id="0D9145C72C4FCE29FDD4C5739CC3FA63" box="[542,668,1405,1429]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2A, B</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD6DC5739CB7FA63" box="[679,744,1405,1429]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3C, D</figureCitation>
) (versus with prominent crest; cf.
<bibRefCitation id="F13B24B32C4FCE29FE14C5909C9FFA43" author="Ng PKL &amp; Rahayu DL" box="[478,704,1437,1462]" pageId="3" pageNumber="338" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023</bibRefCitation>
: fig. 7GL). The adult male chela is relatively more enlarged, with the external median surface almost smooth (
<figureCitation id="0D9145C72C4FCE29FD5FC5D39ED9F9E3" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2C, D</figureCitation>
) (versus chelae not enlarged and the outer surface of the chela densely covered with granules; cf.
<figureCitation id="0D9145C72C4FCE29FD94C6139CEEF9C3" box="[606,689,1565,1589]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Fig. 6F</figureCitation>
). The ambulatory legs are also comparatively slender (P4 merus length about 5× height) and more elongated (P4 length about 1.8× carapace width), with the merus surface lacking a medio-longitudinal swelling or ridge (
<figureCitation id="0D9145C72C4FCE29FDFBC6939CCAF943" box="[561,661,1693,1717]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD6EC6939CB7F943" box="[676,744,1693,1717]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3C, D</figureCitation>
) (versus median part swollen, P4 merus length 3.03.5× height; cf.
<bibRefCitation id="F13B24B32C4FCE29FF2FC6D09F9FF903" author="Ng PKL &amp; Rahayu DL" box="[229,448,1757,1782]" pageId="3" pageNumber="338" pagination="428 - 454" refId="ref14336" refString="Ng PKL &amp; Rahayu DL (2023) Review of the pilumnid crab genus Lophoplax Tesch, 1918 from the western Pacific, with descriptions of two new species, and the clarification of the identity of Pseudocryptocoeloma parvus Ward, 1936 (Crustacea: Brachyura). Zootaxa, 5244 (5): 428 - 454." type="journal article" year="2023">Ng &amp; Rahayu, 2023</bibRefCitation>
: fig. 7GL).
<taxonomicName id="52AA22C12C4FCE29FD83C6D39CE4F903" authorityName="Tesch" authorityYear="1918" box="[585,699,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FD83C6D39CE4F903" box="[585,699,1757,1781]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
, like
<taxonomicName id="52AA22C12C4FCE29FFB8C6F39F5CF8E3" authorityName="Deb" authorityYear="1989" box="[114,259,1789,1813]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FFB8C6F39F5CF8E3" box="[114,259,1789,1813]" italics="true" pageId="3" pageNumber="338">Myopilumnus</emphasis>
</taxonomicName>
, further differs from
<taxonomicName id="52AA22C12C4FCE29FE3AC6F39C1AF8E3" box="[496,581,1789,1813]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FE3AC6F39C1AF8E3" box="[496,581,1789,1813]" italics="true" pageId="3" pageNumber="338">Hosekia</emphasis>
</taxonomicName>
, in having four instead of three anterolateral carapace teeth (
<figureCitation id="0D9145C72C4FCE29FD94C7139CE0F8C3" box="[606,703,1821,1845]" captionStart="Fig" captionStartId="10.[136,171,1548,1570]" captionTargetBox="[135,1473,189,1532]" captionTargetId="figure-7@10.[135,1473,189,1532]" captionTargetPageId="10" captionText="Fig. 6. Hosekia symmetrinuda (Edmondson, 1951), new combination, holotype male (6.5 × 5.0 mm) (BPBM 5109), Samoa. A, dorsal view of carapace (right side denuded); B, subfrontal view of cephalothorax (right side denuded); C, frontal view of cephalothorax (right side denuded); D, frontal view showing buccal cavity; E, buccal cavity, anterior thoracic sternum and sternopleonal cavity; F, left chela." figureDoi="http://doi.org/10.5281/zenodo.14682775" httpUri="https://zenodo.org/record/14682775/files/figure.png" pageId="3" pageNumber="338">Figs. 6A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FD06C7139EDBF8A3" captionStart="Fig" captionStartId="11.[113,148,1690,1712]" captionTargetBox="[113,1451,189,1674]" captionTargetId="figure-7@11.[113,1451,189,1674]" captionTargetPageId="11" captionText="Fig. 7. Hosekia symmetrinuda (Edmondson, 1951), new combination. AC, E, male (7.6 × 4.2 mm) (RUMF-ZC-5199), Japan; D, E, female (9.7 × 6.9 mm) (RUMF-ZC-7536), Japan. A, dorsal habitus (right side denuded); B, right third maxilliped; C, dorsal view of carapace (right side denuded); D, dorsal view of carapace (not denuded); E, male buccal cavity, anterior thoracic sternum and pleon; F, female sternopleonal cavity and vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682777" httpUri="https://zenodo.org/record/14682777/files/figure.png" pageId="3" pageNumber="338">7C, D</figureCitation>
versus
<figureCitation id="0D9145C72C4FCE29FF10C7339F66F8A3" box="[218,313,1853,1877]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Figs. 1A</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FE8DC7339F33F8A3" box="[327,364,1853,1877]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">2A</figureCitation>
).
</paragraph>
<paragraph id="951559422C4FCE29FFBBC7739D9FFEE3" blockId="3.[113,753,1917,2101]" lastBlockId="3.[813,1451,189,277]" pageId="3" pageNumber="338">
<taxonomicName id="52AA22C12C4FCE29FFBBC7739EB4F863" authorityName="Tesch" authorityYear="1918" box="[113,235,1917,1941]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FFBBC7739EB4F863" box="[113,235,1917,1941]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
superficially resembles
<taxonomicName id="52AA22C12C4FCE29FDDFC7739CF0F863" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[533,687,1917,1941]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C4FCE29FDDFC7739CF0F863" box="[533,687,1917,1941]" italics="true" pageId="3" pageNumber="338">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C4FCE29FD08C7709EECF840" pageId="3" pageNumber="338" rank="genus">new genus</taxonomicNameLabel>
, in the slender ambulatory legs (albeit with P4 merus length about 5× height, versus 4× height) and in having blunt prominences along most of the lateral margins of the carapace, but is immediately distinguished by the position of the orbits: on the anterior margin and fully visible dorsally in
<taxonomicName id="52AA22C12C4FCE29FCE7C0B39DC0FF23" authorityName="Tesch" authorityYear="1918" box="[813,927,189,213]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FCE7C0B39DC0FF23" box="[813,927,189,213]" italics="true" pageId="3" pageNumber="338">Lophoplax</emphasis>
</taxonomicName>
, versus submarginal and scarcely visible dorsally in
<taxonomicName id="52AA22C12C4FCE29FC8DC0D39D8AFF03" box="[839,981,221,245]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FC8DC0D39D8AFF03" box="[839,981,221,245]" italics="true" pageId="3" pageNumber="338">Striacoeloma</emphasis>
</taxonomicName>
(resembling the condition in
<taxonomicName id="52AA22C12C4FCE29FACDC0D39DEBFEE3" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C4FCE29FACDC0D39BF4FF03" box="[1287,1451,221,245]" italics="true" pageId="3" pageNumber="338">Cryptocoeloma</emphasis>
<bibRefCitation id="F13B24B32C4FCE29FCE7C0F39DEBFEE3" author="Miers EJ" box="[813,948,253,277]" pageId="3" pageNumber="338" pagination="18 - 32" refId="ref13583" refString="Miers EJ (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. London, British Museum (Natural History) pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="951559422C4FCE29FCE7C1339A6CFD63" blockId="3.[813,1452,317,661]" pageId="3" pageNumber="338">
<bibRefCitation id="F13B24B32C4FCE29FCE7C1339A7EFEA3" author="Serene R &amp; Soh CL" box="[813,1057,317,341]" pageId="3" pageNumber="338" pagination="1 - 37" refId="ref14789" refString="Serene R &amp; Soh CL (1976) Brachyura collected during the Thai- Danish Expedition (1966). Research Bulletin Phuket Marine Biological Center, 12: 1 - 37, figs. 1 - 28, pls. 1 - 7." type="journal article" year="1976">Serène &amp; Soh (1976: 4</bibRefCitation>
, 21) recorded a small male specimen of “?
<taxonomicName id="52AA22C12C4FCE29FCA7C1539A13FE83" authorityName="Sakai" authorityYear="1935" box="[877,1100,349,373]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4FCE29FCA7C1539A13FE83" box="[877,1100,349,373]" italics="true" pageId="3" pageNumber="338">Lophoplax takakurai</emphasis>
</taxonomicName>
” from the Andaman Sea stating only that “Like
<taxonomicName id="52AA22C12C4FCE29FC20C1739A09FE63" authorityName="Sakai" authorityYear="1935" box="[1002,1110,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="takakurai">
<emphasis id="A7DE85502C4FCE29FC20C1739A09FE63" box="[1002,1110,381,405]" italics="true" pageId="3" pageNumber="338">takakurai</emphasis>
</taxonomicName>
the present specimen has no elevation, but it differs by its anterolateral teeth much less clearly separated one from the other”. On this basis, it seems unlikely to be
<taxonomicName id="52AA22C12C4FCE29FC06C1D39AABFE03" authorityName="Deb" authorityYear="1989" box="[972,1268,477,501]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="andamanicus">
<emphasis id="A7DE85502C4FCE29FC06C1D39AABFE03" box="[972,1268,477,501]" italics="true" pageId="3" pageNumber="338">Myopilumnus andamanicus</emphasis>
</taxonomicName>
, which was also described from the Andaman Sea (
<bibRefCitation id="F13B24B32C4FCE29FB09C1F39B1FFDE3" author="Deb M" box="[1219,1344,509,533]" pageId="3" pageNumber="338" pagination="113 - 116" refId="ref13417" refString="Deb M (1989) Myopilumnus adamanicus n. gen., n. sp., a xanthid crab from Andamans. Journal of the Andaman Science Association, 5 (2): 113 - 116." type="journal article" year="1989">Deb, 1989</bibRefCitation>
; see also
<bibRefCitation id="F13B24B32C4FCE29FCE7C2139A52FDC3" author="Trivedi J &amp; Patel K &amp; Mitra S &amp; Ng PKL" box="[813,1037,541,565]" pageId="3" pageNumber="338" pagination="595 - 600" refId="ref15186" refString="Trivedi J, Patel K, Mitra S &amp; Ng PKL (2022) On the identity of Myopilumnus andamanicus Deb, 1989 (Crustacea: Decapoda: Brachyura: Pilumnidae) from India. Zootaxa, 5194 (4): 595 - 600." type="journal article" year="2022">Trivedi et al., 2022</bibRefCitation>
).
<bibRefCitation id="F13B24B32C4FCE29FBE9C2139B78FDC3" author="Serene R &amp; Soh CL" box="[1059,1319,541,565]" pageId="3" pageNumber="338" pagination="1 - 37" refId="ref14789" refString="Serene R &amp; Soh CL (1976) Brachyura collected during the Thai- Danish Expedition (1966). Research Bulletin Phuket Marine Biological Center, 12: 1 - 37, figs. 1 - 28, pls. 1 - 7." type="journal article" year="1976">Serène &amp; Sohs (1976)</bibRefCitation>
record was subsequently repeated in the checklists of the Thai fauna by
<bibRefCitation id="F13B24B32C4FCE29FC9BC2539A5BFD83" author="Naiyanetr P" box="[849,1028,605,629]" pageId="3" pageNumber="338" refId="ref13683" refString="Naiyanetr P (1980) Crustacean fauna of Thailand (Decapoda and Stomatopoda). Chulalongkorn University, Bangkok, Thailand, unnumbered pages + 73 pp." type="book" year="1980">Naiyanetr (1980</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4FCE29FBDBC2539A16FD83" author="Naiyanetr P" box="[1041,1097,605,629]" pageId="3" pageNumber="338" refId="ref13711" refString="Naiyanetr P (1998) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Environmental Policy and Planning, Bangkok, Thailand, 161 pp." type="book" year="1998">1998</bibRefCitation>
,
<bibRefCitation id="F13B24B32C4FCE29FB9CC2539AD2FD83" author="Naiyanetr P" box="[1110,1165,605,629]" pageId="3" pageNumber="338" refId="ref13748" refString="Naiyanetr P (2007) Checklist of Crustacean Fauna in Thailand (Decapoda, Stomatopoda, Anostraca, Myodocopa and Isopoda). Office of Natural Resources and Environmental Policy and Planning, Bangkok, Thailand, 196 pp." type="book" year="2007">2007</bibRefCitation>
) and
<bibRefCitation id="F13B24B32C4FCE29FB07C2509BF7FD83" author="Ng PKL &amp; Davie PJF" box="[1229,1448,605,630]" pageId="3" pageNumber="338" pagination="369 - 384" refId="ref14209" refString="Ng PKL &amp; Davie PJF (2002) A checklist of the brachyuran crabs of Phuket and western Thailand. Phuket Marine Biological Center Special Publication, 23 (2): 369 - 384." type="journal article" year="2002">Ng &amp; Davie (2002)</bibRefCitation>
; it requires confirmation.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,276 @@
<document id="DA73084188AAD88407B1A3C9489648CC" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C4FCE2CFC7BC2B39BEDFEA3" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Lophoplax bicristata Tesch 1918" docType="treatment" docVersion="2" lastPageNumber="341" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="338" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="11C133E4B3B8F776FD4E559BB91F8300" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F26020AA98ABA4EFCC45A0954F5D8E2">
<mods:title id="1DDC1C336237E11D878A2D59793CA624">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="3BE60F70A95098C04474309A0885589B" type="personal">
<mods:role id="AAB11986B8C565D5EAD749F97854DA36">
<mods:roleTerm id="5154027066DE95A25696F4704BC3D30C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05B95EBD16A73A44B4F1216086398BEB">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="313FE5856B624FAF0D454105466F1628" type="personal">
<mods:role id="0C965B0378DD0762E42DC0E3C740818E">
<mods:roleTerm id="6DEDEF08CBC660C95AA2518778F755B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BBA2D4DA2E5724B4F6114F4AC31A380D">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="D52148307EF0126F3189BBEC43F2A4A5" type="personal">
<mods:role id="AA9F295547828D42136851331039E311">
<mods:roleTerm id="CBCBD8BB05D7617F463E56E88AB8EE36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6CB1006C3E2448599EC210DBE8CCD910">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3F718835826BAA318C08CBC95E015745">text</mods:typeOfResource>
<mods:relatedItem id="087707779293BF0CAA6ACA4C3650746F" type="host">
<mods:titleInfo id="024FA5E99A53956194BF9535DBA3315B">
<mods:title id="FD6582D35B39B77B1AD57CEFE0B3B29B">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="A7216872C27D201137576BCFB05EE831">
<mods:date id="8B4EC7A1036CEEDC7A81837D6E0C1A33">2024</mods:date>
<mods:detail id="31ADB5EFD084D63D225861BDE8C092AE" type="pubDate">
<mods:number id="3123EEB3B6A58B4EC446535E3FE7B128">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="30F6A5DFEEE899E63F482E12FE161079" type="volume">
<mods:number id="97E5519234FADE67A91C0462FB260BF2">72</mods:number>
</mods:detail>
<mods:extent id="182F6DA151188F680C0F691D4C7DDCFF" unit="page">
<mods:start id="5ECAA2B30DFA318B165D81E1E770BEB7">335</mods:start>
<mods:end id="EB56A892445808134F4CBA9F5EFA0109">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7D98B1CA7ED6950A016C86455340B981">journal article</mods:classification>
<mods:identifier id="94882CD47A748409CC9E3AE0BEACCBCE" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="32342D4170E1EAFABDDEC1EE497B587B" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="770EDDBBAB46245BDDE287A932163C94" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="AE65F1D35562A6DE6310CE8325A13124" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C4FCE2CFC7BC2B39BEDFEA3" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3" httpUri="http://treatment.plazi.org/id/1D03E8542C4FCE2CFC7BC2B39BEDFEA3" lastPageId="6" lastPageNumber="341" pageId="3" pageNumber="338">
<subSubSection id="DDB00AC92C4FCE29FC7BC2B39B79FD23" box="[945,1318,701,725]" pageId="3" pageNumber="338" type="nomenclature">
<paragraph id="951559422C4FCE29FC7BC2B39B79FD23" blockId="3.[945,1318,701,757]" box="[945,1318,701,725]" pageId="3" pageNumber="338">
<heading id="CE5DEE2E2C4FCE29FC7BC2B39B79FD23" bold="true" box="[945,1318,701,725]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="338" reason="2">
<taxonomicName id="52AA22C12C4FCE29FC7BC2B39B79FD23" ID-CoL="3W3WR" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[945,1318,701,725]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4FCE29FC7BC2B39B79FD23" bold="true" box="[945,1318,701,725]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FC7BC2B39AC8FD23" bold="true" box="[945,1175,701,725]" italics="true" pageId="3" pageNumber="338">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C4FCE29FB55C2B39B79FD23" author="Tesch JJ" box="[1183,1318,701,725]" pageId="3" pageNumber="338" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FBE5C2D39AF6FD03" box="[1071,1193,733,757]" pageId="3" pageNumber="338" type="description">
<paragraph id="951559422C4FCE29FBE5C2D39AF6FD03" blockId="3.[945,1318,701,757]" box="[1071,1193,733,757]" pageId="3" pageNumber="338">
(
<figureCitation id="0D9145C72C4FCE29FBFDC2D39ADAFD03" box="[1079,1157,733,757]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Figs. 2</figureCitation>
,
<figureCitation id="0D9145C72C4FCE29FB58C2D39AC0FD03" box="[1170,1183,733,757]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FCE7C3139BF5FBE3" pageId="3" pageNumber="338" type="materials_examined">
<paragraph id="951559422C4FCE29FCE7C3139BF5FBE3" blockId="3.[813,1452,797,1045]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C3139A52FCC3" bold="true" box="[813,1037,797,821]" pageId="3" pageNumber="338">Material examined.</emphasis>
<materialsCitation id="25C2531F2C4FCE29FBDFC3109A8CFC43" collectingDate="1899-12-16" collectingDateMax="1899-12-18" collectingDateMin="1899-12-16" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="90" latitude="-5.6083336" location="Nuhu Jaan" longLatPrecision="130" longitude="132.92" municipality="Kei Islands" pageId="3" pageNumber="338" specimenCode="D103002" specimenCount="1" typeStatus="lectotype">
<typeStatus id="4A11E7E02C4FCE29FBDFC3109ADFFCC0" box="[1045,1152,798,822]" pageId="3" pageNumber="338" type="lectotype">Lectotype</typeStatus>
(here designated):
<collectionCode id="F3BBC1872C4FCE29FA99C3109D37FCA0" pageId="3" pageNumber="338">RMNH-ZMA</collectionCode>
<specimenCode id="C50CF1392C4FCE29FCA5C3339D89FCA3" box="[879,982,829,853]" collectionCode="RMNH-ZMA" pageId="3" pageNumber="338">D103002</specimenCode>
, male (5.8 ×
<quantity id="5252F4A72C4FCE29FBAEC3339AE5FCA3" box="[1124,1210,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="338" unit="mm" value="4.3">4.3 mm</quantity>
),
<quantity id="5252F4A72C4FCE29FB04C3339B6EFCA3" box="[1230,1329,829,853]" metricMagnitude="3" metricUnit="m" metricValue="3.7014912" pageId="3" pageNumber="338" unit="mi" value="2.3">2.3 miles</quantity>
from north point of
<location id="90750F992C4FCE29FC45C3539A56FC83" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4FCE29FC45C3539A56FC83" box="[911,1033,861,885]" country="Indonesia" county="Siboga Expedition station" latitude="-5.6083336" longLatPrecision="130" longitude="132.92" municipality="Kei Islands" name="Nuhu Jaan" pageId="3" pageNumber="338">Nuhu Jaan</location>
, northwest of
<collectingMunicipality id="7571C3382C4FCE29FB66C3539B72FC83" box="[1196,1325,861,885]" pageId="3" pageNumber="338">Kei Islands</collectingMunicipality>
,
<collectingCountry id="EDBD19D22C4FCE29FAF6C3539BF7FC83" box="[1340,1448,861,885]" name="Indonesia" pageId="3" pageNumber="338">Indonesia</collectingCountry>
,
<geoCoordinate id="F09E3F852C4FCE29FCE7C3729DD7FC63" box="[813,904,892,917]" degrees="5" direction="south" minutes="36.5" orientation="latitude" pageId="3" pageNumber="338" precision="92" value="-5.6083336">5°36.5S</geoCoordinate>
<geoCoordinate id="F09E3F852C4FCE29FC5AC3729A55FC63" box="[912,1034,892,917]" degrees="132" direction="east" minutes="55.2" orientation="longitude" pageId="3" pageNumber="338" precision="92" value="132.92">132°55.2E</geoCoordinate>
,
<quantity id="5252F4A72C4FCE29FBDDC3739A0EFC63" box="[1047,1105,893,917]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="338" unit="m" value="90.0">
<elevation id="1E87BE712C4FCE29FBDDC3739A0EFC63" box="[1047,1105,893,917]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="338" unit="m" value="90.0">90 m</elevation>
</quantity>
,
<collectingCounty id="7C7421CE2C4FCE29FB94C3739B2BFC63" box="[1118,1396,893,917]" pageId="3" pageNumber="338">Siboga Expedition station</collectingCounty>
260, coll.
<collectorName id="385F3C942C4FCE29FCA8C3909D8EFC43" box="[866,977,925,949]" pageId="3" pageNumber="338">M. Weber</collectorName>
,
<date id="E1147F822C4FCE29FC16C3939A8CFC43" box="[988,1235,925,949]" pageId="3" pageNumber="338" value="1899-12-16" valueMax="1899-12-18" valueMin="1899-12-16">
<collectingDate id="F150866A2C4FCE29FC16C3939A8CFC43" box="[988,1235,925,949]" pageId="3" pageNumber="338" value="1899-12-16" valueMax="1899-12-18" valueMin="1899-12-16">1618 December 1899</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="25C2531F2C4FCE29FB17C3939BF9FBE3" collectingDate="1899-06-10" collectionCode="RMNH-ZMA" collectorName="M. Weber" country="Indonesia" county="Siboga Expedition station" elevation="59" location="Borneo Bank" municipality="Strait of Makassar" pageId="3" pageNumber="338" specimenCode="D103001" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="4A11E7E02C4FCE29FB17C3939B31FC43" box="[1245,1390,925,949]" pageId="3" pageNumber="338" type="paralectotype">Paralectotype</typeStatus>
(here designated):
<collectionCode id="F3BBC1872C4FCE29FC7DC3B09A14FC20" box="[951,1099,958,982]" pageId="3" pageNumber="338">RMNH-ZMA</collectionCode>
<specimenCode id="C50CF1392C4FCE29FB99C3B39AE6FC23" box="[1107,1209,957,981]" collectionCode="RMNH-ZMA" pageId="3" pageNumber="338">D103001</specimenCode>
, male (4.5 ×
<quantity id="5252F4A72C4FCE29FA83C3B39BC0FC23" box="[1353,1439,957,981]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="338" unit="mm" value="3.7">3.7 mm</quantity>
),
<location id="90750F992C4FCE29FCE7C3D09DE0FC03" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4FCE29FCE7C3D09DE0FC03" box="[813,959,989,1013]" country="Indonesia" county="Siboga Expedition station" municipality="Strait of Makassar" name="Borneo Bank" pageId="3" pageNumber="338">Borneo Bank</location>
,
<collectingMunicipality id="7571C3382C4FCE29FC06C3D39AC9FC03" box="[972,1174,989,1013]" pageId="3" pageNumber="338">Strait of Makassar</collectingMunicipality>
,
<collectingCountry id="EDBD19D22C4FCE29FB69C3D39B52FC03" box="[1187,1293,989,1013]" name="Indonesia" pageId="3" pageNumber="338">Indonesia</collectingCountry>
,
<quantity id="5252F4A72C4FCE29FAD0C3D39B0BFC03" box="[1306,1364,989,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="338" unit="m" value="59.0">
<elevation id="1E87BE712C4FCE29FAD0C3D39B0BFC03" box="[1306,1364,989,1013]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="338" unit="m" value="59.0">59 m</elevation>
</quantity>
,
<collectingCounty id="7C7421CE2C4FCE29FAABC3D39DAEFBE3" pageId="3" pageNumber="338">Siboga Expedition station</collectingCounty>
77, coll. coll.
<collectorName id="385F3C942C4FCE29FB59C3F09B5BFBE3" box="[1171,1284,1021,1045]" pageId="3" pageNumber="338">M. Weber</collectorName>
,
<date id="E1147F822C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">
<collectingDate id="F150866A2C4FCE29FADAC3F39BF9FBE3" box="[1296,1446,1021,1045]" pageId="3" pageNumber="338" value="1899-06-10">10 June 1899</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE29FCE7C4339B6BF9E3" pageId="3" pageNumber="338" type="diagnosis">
<paragraph id="951559422C4FCE29FCE7C4339B6BF9E3" blockId="3.[811,1452,1085,1557]" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C4339DC1FBA3" bold="true" box="[813,926,1085,1109]" pageId="3" pageNumber="338">Diagnosis.</emphasis>
Carapace frontal margin almost straight, granulate, not distinctly produced anteriorly, lateral lobe very low but just visible (
<figureCitation id="0D9145C72C4FCE29FC25C4739A2EFB63" box="[1007,1137,1149,1173]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); epigastric areole low but visible; protogastric areolet distinct, smooth (
<figureCitation id="0D9145C72C4FCE29FAE9C4939BC1FB43" box="[1315,1438,1181,1205]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A, B</figureCitation>
); metabranchial region with prominent raised granules (
<figureCitation id="0D9145C72C4FCE29FA4AC4B39D0EFB03" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2A</figureCitation>
); median lobe on posterior margin of epistome obtusely triangular (
<figureCitation id="0D9145C72C4FCE29FC65C4F39A54FAE3" box="[943,1035,1277,1301]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3B</figureCitation>
). Distal margin of ischium of third maxilliped smooth (
<figureCitation id="0D9145C72C4FCE29FBCBC5139A09FAC3" box="[1025,1110,1309,1333]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3A</figureCitation>
). Cheliped carpus dorsal surface with subquadrate areole with crenulate ridged margins (
<figureCitation id="0D9145C72C4FCE29FA4AC5339D2DFA83" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2AC</figureCitation>
); proximal basal part of male chela almost smooth or with very low tubercles (
<figureCitation id="0D9145C72C4FCE29FB96C5739AEBFA63" box="[1116,1204,1405,1429]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2D</figureCitation>
). P24 merus extensor margin distinctly crenulated, with low lobes. P5 merus unarmed (
<figureCitation id="0D9145C72C4FCE29FC6AC5B39A7FFA23" box="[928,1056,1469,1493]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3C, D</figureCitation>
). Male thoracic sternum anteriorly relatively wide (
<figureCitation id="0D9145C72C4FCE29FC16C5D39A6FFA03" box="[988,1072,1501,1525]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="3" pageNumber="338">Fig. 2E</figureCitation>
). G1 gently sinuous, distal strongly recurved (
<figureCitation id="0D9145C72C4FCE29FC51C5F39A5BF9E3" box="[923,1028,1533,1557]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="3" pageNumber="338">Fig. 3FI</figureCitation>
). G2 about ¼ length of G1.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C4FCE2CFCE7C6339BEDFEA3" lastPageId="6" lastPageNumber="341" pageId="3" pageNumber="338" type="description">
<paragraph id="951559422C4FCE2FFCE7C6339A93F7E3" blockId="3.[813,1452,1597,2101]" lastBlockId="5.[813,1453,1725,2069]" lastPageId="5" lastPageNumber="340" pageId="3" pageNumber="338">
<emphasis id="A7DE85502C4FCE29FCE7C6339DE8F9A3" bold="true" box="[813,951,1597,1621]" pageId="3" pageNumber="338">Description.</emphasis>
Carapace subquadrate; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like, not concealing margins; regions well demarcated with few areoles distinct: protogastric areole transversely ovate, distinct; epigastric region low, visible when denuded, not areolate; hepatic region with 1 tubercle, no areole visible; cardiac region raised, covered with granules, setose, not areolate, lateral parts with raised patches of granules, but no visible cardio-branchial areole; gastro-cardiac region depressed, with grooves visible but not deep (
<figureCitation id="0D9145C72C4FCE29FAEFC7539BC1F883" box="[1317,1438,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); intestinal region convex, forming rounded areole, adjacent surfaces gently raised, with small granules, setose, separated from cristate posterior carapace margin by wide space, no obvious channel (
<figureCitation id="0D9145C72C4FCE29FC3DC7D39A29F803" box="[1015,1142,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, B</figureCitation>
); meso- and metabranchial regions each with 1 gently raised area covered with granules; metabranchial region with tubercle just posterior to raised patch of granules (
<figureCitation id="0D9145C72C49CE2FFE84C6B39FF4F923" box="[334,427,1725,1749]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Subhepatic and suborbital regions slightly rugose to almost smooth; pterygostomial region with small granules. Frontal margin weakly bilobed, almost straight, with shallow median concavity, margins granulate; lateral lobe, small, low, just visible in dorsal view (
<figureCitation id="0D9145C72C49CE2FFFB3C7539E8FF883" box="[121,208,1885,1909]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Supraorbital margin short, sinuous, with distinct submedian lobe, without fissure or cleft (
<figureCitation id="0D9145C72C49CE2FFD8FC7739CC0F863" box="[581,671,1917,1941]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Orbit dorsal, supraorbital margin, eyes clearly visible in dorsal view; eyes freely movable, ocular peduncle relatively long, filling orbit (
<figureCitation id="0D9145C72C49CE2FFECAC7D39F06F803" box="[256,345,2013,2037]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Anterolateral margin with 4 small, sharp teeth (including external orbital tooth), external orbital tooth broader, next 3 teeth acutely triangular, tip slightly enlarged (
<figureCitation id="0D9145C72C49CE2FFC52C6D39A53F903" box="[920,1036,1757,1781]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="5" pageNumber="340">Fig. 2A, B</figureCitation>
). Posterolateral margin uneven, with 2 tubercles, each with smaller basal granules; margins gently converging towards posterior carapace margin (
<figureCitation id="0D9145C72C49CE2FFA8EC7139BC1F8C3" box="[1348,1438,1821,1845]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="5" pageNumber="340">Fig. 1A</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally. Basal antennal article subquadrate; flagellum entering orbital hiatus. Epistome with median lobe of posterior margin obtusely triangular, no obvious fissure in frontal view; lateral parts sinuous with small cleft (
<figureCitation id="0D9145C72C49CE2FFA81C7B39BC1F823" box="[1355,1438,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340">Fig. 3B</figureCitation>
). Endostome distinct, sloping gradually posteriorly into buccal cavern, endostomial ridges low, short.
</paragraph>
<caption id="C1D509CA2C48CE2EFF4DC7FD9B55F7B5" ID-DOI="http://doi.org/10.5281/zenodo.14682765" ID-Zenodo-Dep="14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="4" pageNumber="339" startId="4.[135,170,2035,2057]" targetBox="[162,1447,189,2020]" targetPageId="4" targetType="figure">
<paragraph id="951559422C48CE2EFF4DC7FD9B55F7B5" blockId="4.[135,1474,2035,2115]" pageId="4" pageNumber="339">
Fig. 2.
<taxonomicName id="52AA22C12C48CE2EFF04C7FD9C49F7FE" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[206,534,2035,2057]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C48CE2EFF04C7FD9FC8F7FE" box="[206,407,2035,2056]" italics="true" pageId="4" pageNumber="339">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C48CE2EFE57C7FD9C49F7FE" author="Tesch JJ" box="[413,534,2035,2057]" pageId="4" pageNumber="339" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon.
</paragraph>
</caption>
<caption id="C1D509CA2C49CE2FFFBBC6179C11F973" ID-DOI="http://doi.org/10.5281/zenodo.14682767" ID-Zenodo-Dep="14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="5" pageNumber="340" startId="5.[113,148,1561,1583]" targetBox="[115,1452,189,1546]" targetPageId="5" targetType="figure">
<paragraph id="951559422C49CE2FFFBBC6179C11F973" blockId="5.[113,1452,1561,1670]" pageId="5" pageNumber="340">
Fig. 3.
<taxonomicName id="52AA22C12C49CE2FFF76C6179C52F9D8" authority="Tesch, 1918" authorityName="Tesch" authorityYear="1918" box="[188,525,1561,1583]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C49CE2FFF76C6179FD6F9D8" box="[188,393,1561,1582]" italics="true" pageId="5" pageNumber="340">Lophoplax bicristata</emphasis>
<bibRefCitation id="F13B24B32C49CE2FFE58C6179C52F9D8" author="Tesch JJ" box="[402,525,1561,1583]" pageId="5" pageNumber="340" pagination="7 - 18" refId="ref15155" refString="Tesch JJ (1918) Decapoda Brachyura II. Goneplacidae and Pinnotheridae. Siboga Expeditie Monographie, 39 c 1: 149 - 295, pls. 7 - 18." type="journal article" year="1918">Tesch, 1918</bibRefCitation>
</taxonomicName>
, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm.
</paragraph>
</caption>
<paragraph id="951559422C4ACE2CFF42C0B39FC7FEA3" blockId="6.[136,776,189,341]" pageId="6" pageNumber="341">
Third maxilliped with ischium relatively short, subquadrate, length 1.2× width, with shallow median oblique sulcus; merus subquadrate, anterolateral angle gently auriculiform. Exopod relatively stout, tip reaching to just before distal edge of merus (
<figureCitation id="0D9145C72C4ACE2CFEFEC1339FD3FEA3" box="[308,396,317,341]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C1739F15FD63" blockId="6.[136,776,381,661]" pageId="6" pageNumber="341">
Male thoracic sternum surface relatively smooth, slightly rugose; sternites 1 and 2 completely fused, wide, lateral margins gently concave; sternites 3 and 4 fused with only very shallow median groove and lateral notches visible; suture between sternites 2 and 3 almost straight (
<figureCitation id="0D9145C72C4ACE2CFD6CC1F39CA4FDE3" box="[678,763,509,533]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Fig. 2E</figureCitation>
). Sternite 8 clearly visible as subrectangular plate adjacent to pleonal somite 2 when pleon closed (
<figureCitation id="0D9145C72C4ACE2CFDF4C2339CCCFDA3" box="[574,659,573,597]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
). Tubercle of male pleonal locking mechanism on anterior third of thoracic somite 5.
</paragraph>
<paragraph id="951559422C4ACE2CFF42C2B39FBFFB63" blockId="6.[136,776,701,1174]" pageId="6" pageNumber="341">
Adult male chelipeds almost symmetrical, stout (
<figureCitation id="0D9145C72C4ACE2CFD60C2B39EC3FD03" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1A, B</figureCitation>
). Basis-ischium fused but suture visible; extensor margins uneven but unarmed (
<figureCitation id="0D9145C72C4ACE2CFEBAC2F39F9CFCE3" box="[368,451,765,789]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Merus short, surface slightly rugose; flexor and extensor margins with tubercles and granules; inner distal angle dentiform with subdistal angle tuberculate (
<figureCitation id="0D9145C72C4ACE2CFEC2C3539F07FC83" box="[264,344,861,885]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C</figureCitation>
). Carpus surface granulate; dorsal surface with subquadrate areole with crenulate ridged margins; inner distal angle with low sharp tooth (
<figureCitation id="0D9145C72C4ACE2CFDC9C3939C24FC43" box="[515,635,925,949]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1AC</figureCitation>
). Chela with dorsal surface covered with granules; inner surface almost smooth; median and ventral surfaces almost smooth or with very low tubercles; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, slightly shorter than palm, pigmented brown along distal half; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C4ACE2CFE90C4739F8CFB63" box="[346,467,1149,1173]" captionStart="Fig" captionStartId="2.[135,170,1949,1971]" captionTargetBox="[176,1433,189,1933]" captionTargetId="figure-7@2.[176,1433,189,1933]" captionTargetPageId="2" captionText="Fig. 1. Terminology of carapace structures (based on Myopilumnus sculpta (Stimpson, 1858)). A, dorsal view of carapace; B, right lateral view of cephalothorax. Abbreviations: brs = branchiostegite ridge; brt = branchiostegite; car = cardiac region; car-bra = cardio-branchial areole; cha = channel; epi = epibranchial areole; epi-bra = epibranchial region; hep = hepatic areole; int = intestinal region; int-bra = intestino-branchial areole; meo = Milne Edwards opening; mes-bra = mesobranchial region; met-bra = metabranchial region; met-gas = metagastric region; porb = postorbital areole; pos = posterior carapace margin; pot = tubercles on posterolateral margin; pro = protogastric areole; t1t4 = anterolateral teeth 14." figureDoi="http://doi.org/10.5281/zenodo.14682763" httpUri="https://zenodo.org/record/14682763/files/figure.png" pageId="6" pageNumber="341">Fig. 1C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C4B39CCAF9C3" blockId="6.[136,776,1213,1589]" pageId="6" pageNumber="341">
Ambulatory legs long, P4 longest, about 1.8× carapace width; P4 merus length about 5× height, distinctly longer than half carapace width; outer surface slightly rugose to almost smooth; coxa rounded; merus elongate, P24 extensor margin not cristate with low, broad, well-spaced setose prominences, P5 merus extensor margin smooth, fully setose; surface of P25 gently convex, without median swelling or ridge; P25 carpus relatively smooth, with very low submarginal ridge, lowest in P5; propodus laterally flattened, unarmed, relatively long in P24, shortest in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, slightly upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C4ACE2CFE1FC6139C69F9C3" box="[469,566,1565,1589]" captionStart="Fig" captionStartId="4.[135,170,2035,2057]" captionTargetBox="[162,1447,189,2020]" captionTargetId="figure-7@4.[162,1447,189,2020]" captionTargetPageId="4" captionText="Fig. 2. Lophoplax bicristata Tesch, 1918. A, B, DF, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands; C, paralectotype male (4.5 × 3.7 mm) (ZMA D103001), Strait of Makassar. A, dorsal habitus (right side partially brushed); B, dorsal habitus (right side denuded); C, dorsal habitus (not denuded); D, dorsal view of left cheliped; E, left chela; F, anterior thoracic sternum and pleon." figureDoi="http://doi.org/10.5281/zenodo.14682765" httpUri="https://zenodo.org/record/14682765/files/figure.png" pageId="6" pageNumber="341">Figs. 2A</figureCitation>
,
<figureCitation id="0D9145C72C4ACE2CFD89C6139CD7F9C3" box="[579,648,1565,1589]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">3C, D</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C6539FD1F8A3" blockId="6.[136,776,1629,1877]" pageId="6" pageNumber="341">
Male pleon triangular, all somites and telson free; somite 1 broadly rectangular, reaching base of P5 coxae; somite 2 transversely much narrower than somite 1, trapezoidal; somite 3 broadly subtrapezoidal, widest, somites 46 trapezoidal, gradually decreasing in width with somite 6 almost rectangular; telson slightly wider than long, rounded, slightly narrowly distally, lateral margins convex, longer than somite 6 (
<figureCitation id="0D9145C72C4ACE2CFEE7C7339FDDF8A3" box="[301,386,1853,1877]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C7739F80F803" blockId="6.[136,775,1917,2037]" pageId="6" pageNumber="341">
G1 relatively stout, gently sinuous, distally tapering to narrowly rounded tip, strongly hooked inwards; subdistal surface with 2 long stout setae (
<figureCitation id="0D9145C72C4ACE2CFE32C7B39C38F823" box="[504,615,1981,2005]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3FI</figureCitation>
). G2 sigmoid, about ¼ length of G1 (
<figureCitation id="0D9145C72C4ACE2CFE4EC7D39F8CF803" box="[388,467,2013,2037]" captionStart="Fig" captionStartId="5.[113,148,1561,1583]" captionTargetBox="[115,1452,189,1546]" captionTargetId="figure-8@5.[95,1462,176,1556]" captionTargetPageId="5" captionText="Fig. 3. Lophoplax bicristata Tesch, 1918, lectotype male (5.8 × 4.3 mm) (ZMA D103002), Kei Islands. A, right third maxilliped; B, epistome; C, left P4; D, left P5; E, pleon, showing exposed sternite 8 lateral to somite 2; F, left G1 (ventral view); G, distal part of left G1 (ventral view); H, distal part of left G1 (dorsomedial view); I, distal part of left G1 (dorsal view); J, left G2. Scales: A, B = 0.5 mm; CE = 1.0 mm; F, J = 0.25 mm; GI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.14682767" httpUri="https://zenodo.org/record/14682767/files/figure.png" pageId="6" pageNumber="341">Fig. 3J</figureCitation>
).
</paragraph>
<paragraph id="951559422C4ACE2CFF42C8139BEDFEA3" blockId="6.[136,356,2077,2101]" lastBlockId="6.[835,1475,189,341]" pageId="6" pageNumber="341">
Females not known.
<emphasis id="A7DE85502C4ACE2CFC89C0B39DEBFF23" bold="true" box="[835,948,189,213]" pageId="6" pageNumber="341">Remarks.</emphasis>
<taxonomicName id="52AA22C12C4ACE2CFC74C0B39AFBFF23" authorityName="Tesch" authorityYear="1918" box="[958,1188,189,213]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="bicristata">
<emphasis id="A7DE85502C4ACE2CFC74C0B39AFBFF23" box="[958,1188,189,213]" italics="true" pageId="6" pageNumber="341">Lophoplax bicristata</emphasis>
</taxonomicName>
is only known from
<specimenCount id="83AC92CB2C4ACE2CFA53C0B09DDDFF03" count="2" pageId="6" pageNumber="341" type="male">two males</specimenCount>
<materialsCitation id="25C2531F2C4ACE2CFC43C0D39BEDFEA3" country="Indonesia" county="They" elevation="75" location="Kei Islands" municipality="Both" pageId="6" pageNumber="341" specimenCount="1">
collected from relatively near each other (
<location id="90750F992C4ACE2CFA82C0D39B9CFF03" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFA82C0D39B9CFF03" box="[1352,1475,221,245]" country="Indonesia" county="They" municipality="Both" name="Kei Islands" pageId="6" pageNumber="341">Kei Islands</location>
and
<location id="90750F992C4ACE2CFCBFC0F09A54FEE3" LSID="urn:lsid:plazi:treatment:1D03E8542C4FCE2CFC7BC2B39BEDFEA3:90750F992C4ACE2CFCBFC0F09A54FEE3" box="[885,1035,253,277]" country="Indonesia" county="They" municipality="Both" name="Borneo Bank" pageId="6" pageNumber="341">Borneo Bank</location>
,
<collectingCountry id="EDBD19D22C4ACE2CFBD3C0F39AD7FEE3" box="[1049,1160,253,277]" name="Indonesia" pageId="6" pageNumber="341">Indonesia</collectingCountry>
).
<collectingMunicipality id="7571C3382C4ACE2CFB57C0F39A8DFEE3" box="[1181,1234,253,277]" pageId="6" pageNumber="341">Both</collectingMunicipality>
are mature and were covered with silt and mud, suggesting they live on soft substrates.
<collectingCounty id="7C7421CE2C4ACE2CFC76C1339DACFEA3" box="[956,1011,317,341]" pageId="6" pageNumber="341">They</collectingCounty>
were collected from depths of
<quantity id="5252F4A72C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">
<elevation id="1E87BE712C4ACE2CFA81C1339BF1FEA3" box="[1355,1454,317,341]" metricMagnitude="1" metricUnit="m" metricValue="7.45" metricValueMax="9.0" metricValueMin="5.9" pageId="6" pageNumber="341" unit="m" value="74.5" valueMax="90.0" valueMin="59.0">5990 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,236 @@
<document id="A1B8F92611A03078A44D8C8E7B95B730" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C5CCE39FC07C4939E8AFD03" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Striacoeloma tubur Ng, Ahyong &amp; T., 2024, new species" docType="treatment" docVersion="2" lastPageNumber="354" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="351" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BDE67D0CC528E77342C6D5B77156CB75" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B64F567974B0A83017ACA7C5356768B7">
<mods:title id="5145B194804FD586073FFB4D19F5F19C">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="BE351C213B620D7ED9386E6ECBF8985C" type="personal">
<mods:role id="057C8B6684AAC8ECBCCC4A61C8A20F30">
<mods:roleTerm id="1DD62F7EDD167EA02D6C99769332691A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D536352F4CB79EF086AE89023B6DE3B0">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="2350EFBF9E1C51FAB83966EFA8B6974B" type="personal">
<mods:role id="3B62D28F69D01B14FDFB1CC3E2F7875C">
<mods:roleTerm id="1A07188044B2845E1BF5A44EBFE050D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B0DF55B2562BD6B60609450F02AC0C2">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="DABAB3C8BE01A8679A018DCF65E2F1CE" type="personal">
<mods:role id="04E5308E18F2293D43E404E871F72AE0">
<mods:roleTerm id="BBC4BA23406DAA9C62E09A49C4749E5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F7779CF19FDB4E6F494FCBDB03954BB">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="568E31DFE84423585AC11458B018024D">text</mods:typeOfResource>
<mods:relatedItem id="B7224AAB16A8FC07E12E0B39309B4382" type="host">
<mods:titleInfo id="D740DE760BB02F765780A1F70EEFBFF5">
<mods:title id="7AB9B264C0448CEABBB42022129E0038">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CB99D10FC30B08B91D181B76E52A56EB">
<mods:date id="29D9C97020A70A37330042A7AFAE19D5">2024</mods:date>
<mods:detail id="A6066B306280D7089D8C52DB94088BD5" type="pubDate">
<mods:number id="D9FA3795775B59EB6A40614EAC4A44FE">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="69E4A48DE202735C413FAC9F18659A3A" type="volume">
<mods:number id="B72CBDABFB9CBA89936D3BC9E746822E">72</mods:number>
</mods:detail>
<mods:extent id="3600A4F9709715803CC8AAD355EAB954" unit="page">
<mods:start id="658EF403602ED79D51B7A7A9F4197370">335</mods:start>
<mods:end id="A2400380AFF6258E392FCBAEBBC3BCE7">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E873AE713604367CC8DCB0FE0D42CA96">journal article</mods:classification>
<mods:identifier id="25873BA5557DAA31D3AD5C73CFEEDB21" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="E440B6F88AC194447E8257D3ABC6F8F9" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="F8CE29BD2D720CAAB9CF21C96E25E073" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="4FFED01E60050E2B7273EB9030BC89B9" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C5CCE39FC07C4939E8AFD03" LSID="urn:lsid:plazi:treatment:1D03E8542C5CCE39FC07C4939E8AFD03" httpUri="http://treatment.plazi.org/id/1D03E8542C5CCE39FC07C4939E8AFD03" lastPageId="19" lastPageNumber="354" pageId="16" pageNumber="351">
<subSubSection id="DDB00AC92C5CCE3AFC07C4939B66FB43" box="[973,1337,1181,1205]" pageId="16" pageNumber="351" type="nomenclature">
<paragraph id="951559422C5CCE3AFC07C4939B66FB43" blockId="16.[973,1337,1181,1237]" box="[973,1337,1181,1205]" pageId="16" pageNumber="351">
<heading id="CE5DEE2E2C5CCE3AFC07C4939B66FB43" bold="true" box="[973,1337,1181,1205]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="351" reason="2">
<emphasis id="A7DE85502C5CCE3AFC07C4939B66FB43" bold="true" box="[973,1337,1181,1205]" pageId="16" pageNumber="351">
<taxonomicName id="52AA22C12C5CCE3AFC07C4939AFBFB43" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[973,1188,1181,1205]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFC07C4939AFBFB43" bold="true" box="[973,1188,1181,1205]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFB79C4939B66FB43" box="[1203,1337,1181,1205]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFBF3C4B39A92FB23" box="[1081,1229,1213,1237]" pageId="16" pageNumber="351" type="description">
<paragraph id="951559422C5CCE3AFBF3C4B39A92FB23" blockId="16.[973,1337,1181,1237]" box="[1081,1229,1213,1237]" pageId="16" pageNumber="351">
(
<figureCitation id="0D9145C72C5CCE3AFB8BC4B39AC3FB23" box="[1089,1180,1213,1237]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Figs. 10</figureCitation>
,
<figureCitation id="0D9145C72C5CCE3AFB63C4B39A9BFB23" box="[1193,1220,1213,1237]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C4F39DC3FA83" pageId="16" pageNumber="351" type="materials_examined">
<paragraph id="951559422C5CCE3AFC89C4F39DC3FA83" blockId="16.[835,1474,1277,1398]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C4F39A7EFAE3" bold="true" box="[835,1057,1277,1301]" pageId="16" pageNumber="351">Material examined.</emphasis>
<materialsCitation id="25C2531F2C5CCE3AFBEDC4F39DC3FA83" collectionCode="AM" collectorName="Cardno Pty Ltd." country="Australia" elevation="6" latitude="-12.588056" location="Middle Arm" longLatPrecision="21" longitude="130.87138" municipality="Darwin Harbour" pageId="16" pageNumber="351" specimenCode="P107758" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="4A11E7E02C5CCE3AFBEDC4F39AD1FAE3" box="[1063,1166,1277,1301]" pageId="16" pageNumber="351" type="holotype">Holotype</typeStatus>
:
<collectionCode id="F3BBC1872C5CCE3AFB52C4F09A9CFAE0" box="[1176,1219,1278,1302]" country="Australia" name="Australian Museum" pageId="16" pageNumber="351" type="Museum">AM</collectionCode>
<specimenCode id="C50CF1392C5CCE3AFB03C4F39B75FAE3" box="[1225,1322,1277,1301]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="16" pageNumber="351" type="Museum">P107758</specimenCode>
, female (8.6 ×
<quantity id="5252F4A72C5CCE3AFC89C5139DC8FAC3" box="[835,919,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="16" pageNumber="351" unit="mm" value="6.3">6.3 mm</quantity>
),
<location id="90750F992C5CCE3AFC63C5139A6FFAC0" LSID="urn:lsid:plazi:treatment:1D03E8542C5CCE39FC07C4939E8AFD03:90750F992C5CCE3AFC63C5139A6FFAC0" box="[937,1072,1309,1334]" country="Australia" latitude="-12.588056" longLatPrecision="21" longitude="130.87138" municipality="Darwin Harbour" name="Middle Arm" pageId="16" pageNumber="351" stateProvince="Northern Territory">Middle Arm</location>
,
<collectingMunicipality id="7571C3382C5CCE3AFBF0C5139AB3FAC3" box="[1082,1260,1309,1333]" pageId="16" pageNumber="351">Darwin Harbour</collectingMunicipality>
,
<collectingRegion id="576E97A02C5CCE3AFB3FC5139BE1FAC3" box="[1269,1470,1309,1333]" country="Australia" name="Northern Territory" pageId="16" pageNumber="351">Northern Territory</collectingRegion>
,
<collectingCountry id="EDBD19D22C5CCE3AFC89C5339DF5FAA3" box="[835,938,1341,1365]" name="Australia" pageId="16" pageNumber="351">Australia</collectingCountry>
,
<geoCoordinate id="F09E3F852C5CCE3AFC73C5329A69FAA3" box="[953,1078,1340,1365]" degrees="12" direction="south" minutes="35" orientation="latitude" pageId="16" pageNumber="351" precision="15" seconds="17" value="-12.588056">12°3517″S</geoCoordinate>
<geoCoordinate id="F09E3F852C5CCE3AFB8BC5329A90FAA3" box="[1089,1231,1340,1365]" degrees="130" direction="east" minutes="52" orientation="longitude" pageId="16" pageNumber="351" precision="15" seconds="17" value="130.87138">130°5217″E</geoCoordinate>
,
<quantity id="5252F4A72C5CCE3AFB14C5339B77FAA3" box="[1246,1320,1341,1365]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="351" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">
<elevation id="1E87BE712C5CCE3AFB14C5339B77FAA3" box="[1246,1320,1341,1365]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="16" pageNumber="351" unit="m" value="6.0" valueMax="7.0" valueMin="5.0">57 m</elevation>
</quantity>
, coll.
<collectorName id="385F3C942C5CCE3AFAB8C5339DC3FA83" pageId="16" pageNumber="351">Cardno Pty Ltd.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C5939A4AFA03" pageId="16" pageNumber="351" type="etymology">
<paragraph id="951559422C5CCE3AFC89C5939A4AFA03" blockId="16.[835,1474,1437,1525]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C5939D9AFA43" bold="true" box="[835,965,1437,1461]" pageId="16" pageNumber="351">Etymology.</emphasis>
Named
<taxonomicName id="52AA22C12C5CCE3AFBEFC5939A00FA43" box="[1061,1119,1437,1461]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur">
<emphasis id="A7DE85502C5CCE3AFBEFC5939A00FA43" box="[1061,1119,1437,1461]" italics="true" pageId="16" pageNumber="351">tubur</emphasis>
</taxonomicName>
for the tubercle-like protrusions on the extensor margins of the ambulatory legs; used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFC89C6139A0BF9C0" box="[835,1108,1565,1590]" pageId="16" pageNumber="351" type="diagnosis">
<paragraph id="951559422C5CCE3AFC89C6139A0BF9C0" blockId="16.[835,1108,1565,1590]" box="[835,1108,1565,1590]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C6139DE7F9C3" bold="true" box="[835,952,1565,1589]" pageId="16" pageNumber="351">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE39FC89C6539F64FE03" lastPageId="19" lastPageNumber="354" pageId="16" pageNumber="351" type="description">
<paragraph id="951559422C5CCE38FC89C6539F8EF917" blockId="16.[835,1475,1629,2101]" lastBlockId="18.[136,776,1129,1761]" lastPageId="18" lastPageNumber="353" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFC89C6539AEEF983" bold="true" box="[835,1201,1629,1653]" pageId="16" pageNumber="351">
Description of
<typeStatus id="4A11E7E02C5CCE3AFC38C6539A0BF983" box="[1010,1108,1629,1653]" pageId="16" pageNumber="351" type="holotype">holotype</typeStatus>
female.
</emphasis>
Carapace subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like; surface appearing irregularly corrugated, partly eroded; gastric regions prominently raised, forming 2 transverse swellings extending to branchial region, with posterior one reaching to lateral margin; gastro-cardiac region depressed, with grooves visible; cardiac region with transverse swelling, extends almost to posterolateral margin (
<figureCitation id="0D9145C72C5CCE3AFC81C7539D91F883" box="[843,974,1885,1909]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10AC</figureCitation>
); surface adjacent to posterior carapace margin raised, with prominent wide rounded transverse ridge adjacent to the posterior carapace margin, forming channel extending to sub-branchial region (
<figureCitation id="0D9145C72C5CCE3AFBACC7B39AA8F823" box="[1126,1271,1981,2005]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A, B</figureCitation>
). Subhepatic and suborbital regions slightly rugose; pterygostomial region with numerous fine granules (
<figureCitation id="0D9145C72C5CCE3AFB87C7F39AEEF7E3" box="[1101,1201,2045,2069]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10D</figureCitation>
). Frontal margin bilobed, almost straight, lined with granules, with shallow median notch, lateral lobe small but distinct, not visible in dorsal view (
<figureCitation id="0D9145C72C5ECE38FF5AC4879F48FB54" box="[144,279,1161,1186]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10AD</figureCitation>
). Supraorbital margin sinuous, with low lobe on posterior third, with 2 short fissures (
<figureCitation id="0D9145C72C5ECE38FDF3C4A79CE4FB34" box="[569,699,1193,1218]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10B, C</figureCitation>
). Orbit submarginal, supraorbital margin appearing almost confluent with general carapace outline, eyes partially visible in dorsal view; eyes scarcely movable, ocular peduncle short, filling orbit, scarcely movable (
<figureCitation id="0D9145C72C5ECE38FE5BC5279C4BFAB4" box="[401,532,1321,1346]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10C, D</figureCitation>
). Anterolateral margin with 4 distinct teeth (including external orbital tooth), first 2 broad, lobiform, with surface adjacent to them eroded, each with median oblique depression; last 2 dentiform (
<figureCitation id="0D9145C72C5ECE38FD16C5879E82FA34" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A, B</figureCitation>
). Posterolateral margin with 2 distinct tubercles, margins distinctly converging towards posterior carapace margin (
<figureCitation id="0D9145C72C5ECE38FF22C5E79F2BF9F4" box="[232,372,1513,1538]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A, B</figureCitation>
). Antennules rectangular, relatively large; antennules folding laterally (
<figureCitation id="0D9145C72C5ECE38FE33C6079C04F9D4" box="[505,603,1545,1570]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Basal antennal article subquadrate; flagellum entering orbital hiatus (
<figureCitation id="0D9145C72C5ECE38FD11C6279EE5F994" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Epistome with shallow biconcave posterior margin, median lobe subtruncate with short median fissue (
<figureCitation id="0D9145C72C5ECE38FD13C6679EE4F954" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10D</figureCitation>
). Endostome short, recessed posteriorly into buccal cavern oblique to plane of adjacent pterygostomial surface, endostomial ridges low, short.
</paragraph>
<caption id="C1D509CA2C5DCE3BFFBBC7AA9DB5F802" ID-DOI="http://doi.org/10.5281/zenodo.14682787" ID-Zenodo-Dep="14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="17" pageNumber="352" startId="17.[113,148,1956,1978]" targetBox="[113,1451,268,1941]" targetPageId="17" targetType="figure">
<paragraph id="951559422C5DCE3BFFBBC7AA9DB5F802" blockId="17.[113,1452,1956,2036]" pageId="17" pageNumber="352">
Fig. 10.
<taxonomicName id="52AA22C12C5DCE3BFF0CC7AA9FDDF84F" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[198,386,1956,1977]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="352" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5DCE3BFF0CC7AA9FDDF84F" box="[198,386,1956,1977]" italics="true" pageId="17" pageNumber="352">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5DCE3BFE5BC7AB9C2AF84C" box="[401,629,1956,1978]" pageId="17" pageNumber="352" rank="species">new genus and species</taxonomicNameLabel>
, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped.
</paragraph>
</caption>
<caption id="C1D509CA2C5ECE38FF4DC40E9B68FBC4" ID-DOI="http://doi.org/10.5281/zenodo.14682789" ID-Zenodo-Dep="14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353" startId="18.[135,170,1024,1046]" targetBox="[135,1473,189,1008]" targetPageId="18" targetType="figure">
<paragraph id="951559422C5ECE38FF4DC40E9B68FBC4" blockId="18.[135,1474,1023,1074]" pageId="18" pageNumber="353">
Fig. 11.
<taxonomicName id="52AA22C12C5ECE38FF16C40E9FC7FBE3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[220,408,1024,1045]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="353" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5ECE38FF16C40E9FC7FBE3" box="[220,408,1024,1045]" italics="true" pageId="18" pageNumber="353">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5ECE38FE6DC40E9CD4FBE0" box="[423,651,1024,1046]" pageId="18" pageNumber="353" rank="species">new genus and species</taxonomicNameLabel>
, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5.
</paragraph>
</caption>
<paragraph id="951559422C5ECE38FF42C7079D57F894" blockId="18.[136,776,1801,1890]" pageId="18" pageNumber="353">
Third maxilliped ischium short, quadrate, length 1.2× width; merus with anteroexternal angle distinctly auriculiform; exopod stout, tip reaching distal edge of merus (
<figureCitation id="0D9145C72C5ECE38FD53C7479CA3F894" box="[665,764,1865,1890]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FF42C7879C38F7F4" blockId="18.[136,775,1929,2050]" pageId="18" pageNumber="353">
Thoracic sternum surface smooth; sternites 1 and 2 completely fused, lateral margins sinuous; sternites 3 and 4 fused with only lateral notches demarcating sternites; suture between sternites 2 and 3 almost straight (
<figureCitation id="0D9145C72C5ECE38FE3FC7E79C05F7F4" box="[501,602,2025,2050]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FC89C4679B4FF9D4" blockId="18.[835,1475,1129,1570]" pageId="18" pageNumber="353">
Adult chelipeds symmetrical or almost so, not enlarged (
<figureCitation id="0D9145C72C5ECE38FC86C4879DEBFB54" box="[844,948,1161,1186]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A</figureCitation>
). Basis-ischium fused but suture visible, flexor margin with short sharp granules (
<figureCitation id="0D9145C72C5ECE38FB74C4A79B7CFB34" box="[1214,1315,1193,1218]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">Fig. 11A</figureCitation>
). Merus short, surface rugose; flexor and extensor margins with low sharp tubercles and granules; inner distal angle sharp but not elongate, with subdistal tooth (
<figureCitation id="0D9145C72C5ECE38FB6DC5079B44FAD4" box="[1191,1307,1289,1314]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FAE0C5049B02FAD4" box="[1322,1373,1290,1314]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11A</figureCitation>
). Carpus rugose; inner distal angle with strong sharp tooth (
<figureCitation id="0D9145C72C5ECE38FA9EC5279BE8FAB4" box="[1364,1463,1321,1346]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Fig. 10A</figureCitation>
). Chela with dorsal surface rugose and covered with granules, median part smoother; ventral surface near pollex covered with small granules; inner surface smooth; margins of chela with scattered long and short setae, not dense or brush-like; fingers relatively slender, shorter than palm, pigmented brown along distal ⅔; occlusal margins with low teeth, forming small gape when closed (
<figureCitation id="0D9145C72C5ECE38FB92C6079A99F9D4" box="[1112,1222,1545,1570]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FB1EC6049B5CF9D4" box="[1236,1283,1546,1570]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11C</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE38FC89C6479A0AF834" blockId="18.[835,1475,1609,1986]" pageId="18" pageNumber="353">
Ambulatory legs relatively short, P4 longest, about 1.3× carapace width; outer surface gently rugose; coxa with distal angle slightly raised, no visible serrated flange; P24 with outer surface slightly raised medio-longitudinally, indistinct on P5; merus relatively slender, extensor margin not cristate, with evenly, widely spaced row of blunt, prominent nonsetose nodules; P4 merus length about 4× height, length half carapace width; P24 carpus with outer submarginal ridge, absent on P5; propodus laterally flattened, elongate in P24, short in P5; P24 dactylus falciform, unarmed, P5 dactylus shorter, gently upturned; dactylo-propodal lock visible (
<figureCitation id="0D9145C72C5ECE38FC51C7A79A56F834" box="[923,1033,1961,1986]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="18" pageNumber="353">Figs. 10A</figureCitation>
,
<figureCitation id="0D9145C72C5ECE38FBDDC7A49A17F834" box="[1047,1096,1962,1986]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="18" pageNumber="353">11D</figureCitation>
).
</paragraph>
<paragraph id="951559422C5ECE39FC89C7E79C1FFEC3" blockId="18.[835,1474,2025,2113]" lastBlockId="19.[113,752,189,309]" lastPageId="19" lastPageNumber="354" pageId="18" pageNumber="353">
Female pleon longitudinally ovate; with 6 somites and telson, all free; not completely covering thoracic sternum; somite 1 broadly rectangular, just reaches base of P5 coxae; somite 2 transversely narrower than somite 1, trapezoidal; somites 36 trapezoidal, gradually decreasing in width, somite 6 longest; telson semicircular with convex lateral margins, distinctly longer than somite 6 (
<figureCitation id="0D9145C72C5FCE39FE05C1139C6BFEC3" box="[463,564,285,309]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="951559422C5FCE39FFBBC1539FEAFE43" blockId="19.[113,753,349,437]" pageId="19" pageNumber="354">
Sternopleonal cavity shallow (
<figureCitation id="0D9145C72C5FCE39FE17C1539C16FE83" box="[477,585,349,373]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11B</figureCitation>
); vulva large, obliquely ovate, covering most of space on sternite 6, without vulvar projections (
<figureCitation id="0D9145C72C5FCE39FE8FC1939FF7FE43" box="[325,424,413,437]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="19" pageNumber="354">Fig. 11B</figureCitation>
).
</paragraph>
<paragraph id="951559422C5FCE39FFBBC1D39F64FE03" blockId="19.[113,315,477,501]" box="[113,315,477,501]" pageId="19" pageNumber="354">
<emphasis id="A7DE85502C5FCE39FFBBC1D39EEDFE03" bold="true" box="[113,178,477,501]" pageId="19" pageNumber="354">Male.</emphasis>
Not known.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5FCE39FFBBC2139E8AFD03" pageId="19" pageNumber="354" type="discussion">
<paragraph id="951559422C5FCE39FFBBC2139E8AFD03" blockId="19.[113,752,541,757]" pageId="19" pageNumber="354">
<emphasis id="A7DE85502C5FCE39FFBBC2139EBEFDC3" bold="true" box="[113,225,541,565]" pageId="19" pageNumber="354">Remarks.</emphasis>
<taxonomicName id="52AA22C12C5FCE39FF21C2139FE1FDC3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[235,446,541,565]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="tubur" status="new genus and species">
<emphasis id="A7DE85502C5FCE39FF21C2139FE1FDC3" box="[235,446,541,565]" italics="true" pageId="19" pageNumber="354">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5FCE39FE05C2109C8FFDC3" box="[463,720,541,566]" pageId="19" pageNumber="354" rank="species">new genus and species</taxonomicNameLabel>
, is currently known only from the female
<typeStatus id="4A11E7E02C5FCE39FDDEC2339C2AFDA3" box="[532,629,573,597]" pageId="19" pageNumber="354" type="holotype">holotype</typeStatus>
. Nothing is known about the ecology other than it was collected from soft, muddy substrates. The carapace and pereopods were partially encrusted with what appear to be light-brown, possibly ferric, precipitates, suggestive of reducing or anoxic environs.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,197 @@
<document id="B800EC483DF96920B5C361CF91AE3D05" ID-DOI="10.26107/RBZ-2024-0027" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682761" ID-ZooBank="4BFA36EF-2F2A-4162-8810-5E0414E5DE55" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171673110" checkinUser="felipe" docAuthor="Ng, Peter K. L., Ahyong, &amp; Shane T." docDate="2024" docId="1D03E8542C5CCE3AFEF1C1B39A31FB83" docLanguage="en" docName="RafflesBZool.72.335-355.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Striacoeloma Ng, Ahyong &amp; T., 2024, new genus" docType="treatment" docVersion="2" lastPageNumber="351" masterDocId="E13A902C2C4CCE2AFFCAC00E9E5FFFF6" masterDocTitle="Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)" masterLastPageNumber="355" masterPageNumber="335" pageNumber="351" updateTime="1737175223449" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BB675CB1EB40323C1198D5D3BC4DEB84" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="747A0951E59837F24BBA3120168B9C88">
<mods:title id="B5F7177EDB16DC369E95FD191FD9BDDA">Redefining Lophoplax Tesch, 1918: recognising Myopilumnus Deb, 1989, and the establishment of three new genera, Hosekia, Takedaplax, and Striacoeloma (Crustacea: Brachyura: Pilumnidae)</mods:title>
</mods:titleInfo>
<mods:name id="3004CCAF2BF7F1BC39CE14BA89554822" type="personal">
<mods:role id="3F31EEEE12560CF518A9B2BD0FDAEB74">
<mods:roleTerm id="3000C9F27CB1C11BE6389ECE79C40C35">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C87A87F297C7971D0727F2F466C907E">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="5EB06ADB2A7ABA4E23748196C5A8309E" type="personal">
<mods:role id="98F479466FA5B5B9C14C48781AC3E1BA">
<mods:roleTerm id="06816FCC5AC007B7EDFCCC1EE3A81DBC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B2E86E8A5F71E94729AA924DA343FF9">Ahyong,</mods:namePart>
</mods:name>
<mods:name id="7AB7EAB7191B4BBBCDB82848CAF4EA4F" type="personal">
<mods:role id="F611E8C6A25BB9EF028725A0A9C46AA0">
<mods:roleTerm id="8CDB1AB568938DF59B50DFB1620264E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="683F7A3B568B470A9F39A0EB3D5B9A2F">Shane T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="63C899406FB90CA58E7B2881616681A6">text</mods:typeOfResource>
<mods:relatedItem id="FFF10F14F873F76DF376B1B8F1818BCF" type="host">
<mods:titleInfo id="89818A121C928B62BA437B11D7BB55FA">
<mods:title id="D7FCDB1648EF89994BC2B8EC14B0C6DF">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="44AABEA6DD59F8145C37F188BBA20A43">
<mods:date id="630D1DB735CCBE8A78EFDA876D3FA589">2024</mods:date>
<mods:detail id="92C1FE604B157F35A396F24C38D36B6F" type="pubDate">
<mods:number id="BA1962F5542F62F0B51D39E848DEE958">2024-09-25</mods:number>
</mods:detail>
<mods:detail id="5805E452E3016848D56CA65FA54504EA" type="volume">
<mods:number id="A2FCEA51F044653410331A7FA2A2E4D7">72</mods:number>
</mods:detail>
<mods:extent id="9A1003B22BC653A7F2A2FC9D4B62EC1F" unit="page">
<mods:start id="C45104F3AF44B13C28CCC5DA2CAD832B">335</mods:start>
<mods:end id="DA2C43E4E391B28745073AE0765BA675">355</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BB7A7BE700D2CAF93692581E4A62B6BB">journal article</mods:classification>
<mods:identifier id="E453A53513F4924578445E076D87D2DA" type="DOI">10.26107/RBZ-2024-0027</mods:identifier>
<mods:identifier id="09E54F8EDCAA60B5AB17D3DA483617D8" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A7E9DF35289F13F623CFC9E24BD97FC6" type="Zenodo-Dep">14682761</mods:identifier>
<mods:identifier id="0E74612089A14E23EE181E4C5E0ABC64" type="ZooBank">4BFA36EF-2F2A-4162-8810-5E0414E5DE55</mods:identifier>
</mods:mods>
<treatment id="1D03E8542C5CCE3AFEF1C1B39A31FB83" ID-DOI="http://doi.org/10.5281/zenodo.14682795" ID-Zenodo-Dep="14682795" LSID="urn:lsid:plazi:treatment:1D03E8542C5CCE3AFEF1C1B39A31FB83" httpUri="http://treatment.plazi.org/id/1D03E8542C5CCE3AFEF1C1B39A31FB83" lastPageNumber="351" pageId="16" pageNumber="351">
<subSubSection id="DDB00AC92C5CCE3AFEF1C1B39C0CFE23" box="[315,595,445,469]" pageId="16" pageNumber="351" type="nomenclature">
<paragraph id="951559422C5CCE3AFEF1C1B39C0CFE23" blockId="16.[315,595,445,469]" box="[315,595,445,469]" pageId="16" pageNumber="351">
<heading id="CE5DEE2E2C5CCE3AFEF1C1B39C0CFE23" bold="true" box="[315,595,445,469]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="351" reason="2">
<emphasis id="A7DE85502C5CCE3AFEF1C1B39C0CFE23" bold="true" box="[315,595,445,469]" pageId="16" pageNumber="351">
<taxonomicName id="52AA22C12C5CCE3AFEF1C1B39F93FE23" ID-CoL="CL2RM" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[315,460,445,469]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFEF1C1B39F93FE23" bold="true" box="[315,460,445,469]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFE16C1B39C0CFE23" box="[476,595,445,469]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFF42C1F39CA4FD83" pageId="16" pageNumber="351" type="type_taxon">
<paragraph id="951559422C5CCE3AFF42C1F39C46FDC3" blockId="16.[136,775,509,566]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C1F39F7FFDE3" bold="true" box="[136,288,509,533]" pageId="16" pageNumber="351">
<typeStatus id="4A11E7E02C5CCE3AFF42C1F39E9DFDE3" box="[136,194,509,533]" pageId="16" pageNumber="351">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="52AA22C12C5CCE3AFEEDC1F39FA7FDE3" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[295,504,509,533]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFEEDC1F39FA7FDE3" box="[295,504,509,533]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDCCC1F09CD7FDE3" box="[518,648,509,533]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
, by present designation. Gender of genus neuter.
</paragraph>
<paragraph id="951559422C5CCE3AFF42C2539CA4FD83" blockId="16.[136,763,605,629]" box="[136,763,605,629]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C2539F14FD83" bold="true" box="[136,331,605,629]" pageId="16" pageNumber="351">Included species.</emphasis>
Only
<taxonomicName id="52AA22C12C5CCE3AFE5BC2539C3DFD83" ID-CoL="CL3GY" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[401,610,605,629]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="species" species="tubur" status="sp. nov.">
<emphasis id="A7DE85502C5CCE3AFE5BC2539C3DFD83" box="[401,610,605,629]" italics="true" pageId="16" pageNumber="351">Striacoeloma tubur</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDBBC2509CA9FD83" box="[625,758,605,629]" pageId="16" pageNumber="351" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFF42C2939D57F983" pageId="16" pageNumber="351" type="diagnosis">
<paragraph id="951559422C5CCE3AFF42C2939D57F983" blockId="16.[136,776,669,1653]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C2939EA2FD43" bold="true" box="[136,253,669,693]" pageId="16" pageNumber="351">Diagnosis.</emphasis>
Carapace subhexagonal; frontal and anterolateral margins with scattered tufts of short and long setae, not brush-like; surface appearing irregularly transversely corrugated, partly eroded; gastric and cardiac regions prominently raised, forming transverse swellings extending to branchial region, almost to lateral margin; surface adjacent to posterior carapace margin raised, with prominent wide rounded transverse ridge adjacent to the posterior carapace margin, forming channel extending to sub-branchial region; orbit submarginal, supraorbital margin almost confluent with general carapace outline, eyes only partially visible in dorsal view; eyes scarcely movable, ocular peduncle short, filling orbit, scarcely movable; lateral lobe of frontal margin small but distinct; anterolateral margin with 4 distinct teeth (including external orbital tooth), first 2 broad, lobiform, with 2 additional tubercles on posterolateral margin; posterolateral margins distinctly converging towards posterior carapace margin. Epistome with shallow biconcave posterior margin, median lobe subtruncate. Endostome short, recessed posteriorly into buccal cavern oblique to plane of adjacent pterygostomial surface, endostomial ridges low, short. Third maxilliped ischium short, quadrate, length 1.2× width; merus with anterolateral angle distinctly auriculiform. Adult female chelipeds symmetrical or almost so, stout, outer surface of chela with margins rugose and covered with granules, median part smoother; margins not lined with dense brush-like setae. Ambulatory legs short, P4 longest, about 1.3× carapace width; merus slender, length about 4× height, extensor margin not cristate, with evenly, widely spaced row of blunt, prominent nodules, outer surface slightly raised medio-longitudinally; P4 merus length about 4× height, length half carapace width.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFF42C6939C48F8C3" pageId="16" pageNumber="351" type="etymology">
<paragraph id="951559422C5CCE3AFF42C6939C48F8C3" blockId="16.[136,775,1693,1846]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C6939F54F943" bold="true" box="[136,267,1693,1717]" pageId="16" pageNumber="351">Etymology.</emphasis>
The name is an arbitrary combination of the Latin word “stria”, for ridge, in arbitrary combination with part of the genus name,
<taxonomicName id="52AA22C12C5CCE3AFE53C6D39C62F903" authorityName="Miers" authorityYear="1884" box="[409,573,1757,1781]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE53C6D39C62F903" box="[409,573,1757,1781]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
, in recognition of the ridged carapace surface and close relationship to the pilumnid genus. Gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="DDB00AC92C5CCE3AFF42C7539A31FB83" pageId="16" pageNumber="351" type="discussion">
<paragraph id="951559422C5CCE3AFF42C7539A31FB83" blockId="16.[135,775,1885,2101]" lastBlockId="16.[835,1475,189,1142]" pageId="16" pageNumber="351">
<emphasis id="A7DE85502C5CCE3AFF42C7539EA9F883" bold="true" box="[136,246,1885,1909]" pageId="16" pageNumber="351">Remarks.</emphasis>
Whereas
<taxonomicName id="52AA22C12C5CCE3AFE96C7539FB7F883" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[348,488,1885,1909]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFE96C7539FB7F883" box="[348,488,1885,1909]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFE3EC7509C3AF880" box="[500,613,1886,1910]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
, is superficially similar to various planopilumnids (now in the Pseudozioidea
<bibRefCitation id="F13B24B32C5CCE3AFF42C7939F45F843" author="Alcock A" box="[136,282,1949,1973]" pageId="16" pageNumber="351" pagination="67 - 233" refId="ref13260" refString="Alcock A (1898) Materials for a carcinological fauna of India. No. 3. The Brachyura Cyclometopa. Part I. The family Xanthidae. Journal of the Asiatic Society of Bengal, LVII (II): 67 - 233." type="book chapter" year="1898">Alcock, 1898</bibRefCitation>
) (cf.
<bibRefCitation id="F13B24B32C5CCE3AFE9FC7909FE3F843" author="Ng PKL" box="[341,444,1949,1974]" pageId="16" pageNumber="351" pagination="33 - 61" refId="ref13842" refString="Ng PKL (2010) On the Planopilumnidae Serene, 1984 (Crustacea: Brachyura: Pseudozioidea), with diagnoses of two new pilumnoid genera for species previously assigned to Planopilumnus Balss, 1933. Zootaxa, 2392: 33 - 61." type="journal article" year="2010">Ng, 2010</bibRefCitation>
), it is clearly a pilumnid based the form of its carapace and chelipeds. The new genus is morphologically more similar to
<taxonomicName id="52AA22C12C5CCE3AFE2AC7D39D58F803" authority="Miers, 1884" authorityName="Miers" authorityYear="1884" box="[480,775,2013,2037]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE2AC7D39CDCF803" box="[480,643,2013,2037]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
<bibRefCitation id="F13B24B32C5CCE3AFD42C7D39D58F803" author="Miers EJ" box="[648,775,2013,2037]" pageId="16" pageNumber="351" pagination="18 - 32" refId="ref13583" refString="Miers EJ (1884) Crustacea. In: Report on the zoological collections made in the Indo-Pacific Ocean during the voyage of H. M. S. Alert 1881 - 1882. Part I. The collections from Melanesia. London, British Museum (Natural History) pp. 178 - 322, pls. 18 - 32." type="book chapter" year="1884">Miers, 1884</bibRefCitation>
</taxonomicName>
than
<taxonomicName id="52AA22C12C5CCE3AFF0AC7F39F6DF7E3" authorityName="Tesch" authorityYear="1918" box="[192,306,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFF0AC7F39F6DF7E3" box="[192,306,2045,2069]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C5CCE3AFE88C7F39F8CF7E3" authorityName="Deb" authorityYear="1989" box="[322,467,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE88C7F39F8CF7E3" box="[322,467,2045,2069]" italics="true" pageId="16" pageNumber="351">Myopilumnus</emphasis>
</taxonomicName>
, or
<taxonomicName id="52AA22C12C5CCE3AFDCEC7F39C04F7E3" authorityName="Ng &amp; Ahyong &amp; Shane T." authorityYear="2024" box="[516,603,2045,2069]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="A7DE85502C5CCE3AFDCEC7F39C04F7E3" box="[516,603,2045,2069]" italics="true" pageId="16" pageNumber="351">Hosekia</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BCED382B2C5CCE3AFDA6C7F09CBBF7E0" box="[620,740,2046,2070]" pageId="16" pageNumber="351" rank="genus">new genus</taxonomicNameLabel>
. In
<taxonomicName id="52AA22C12C5CCE3AFF42C8139F4CF7C3" box="[136,275,2077,2101]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFF42C8139F4CF7C3" box="[136,275,2077,2101]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
, like
<taxonomicName id="52AA22C12C5CCE3AFE80C8139FB4F7C3" authorityName="Miers" authorityYear="1884" box="[330,491,2077,2101]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFE80C8139FB4F7C3" box="[330,491,2077,2101]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
, the orbits are submarginal, so the supraorbital margin does not form a strong concavity that noticeably breaks the general carapace outline. The eyes too, are relatively smaller in
<taxonomicName id="52AA22C12C5CCE3AFB63C0F39B68FEE3" box="[1193,1335,253,277]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB63C0F39B68FEE3" box="[1193,1335,253,277]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
and scarcely movable, compared with the proportionally larger, freely movable eyes of
<taxonomicName id="52AA22C12C5CCE3AFBCDC1339A24FEA3" authorityName="Tesch" authorityYear="1918" box="[1031,1147,317,341]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFBCDC1339A24FEA3" box="[1031,1147,317,341]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
,
<taxonomicName id="52AA22C12C5CCE3AFB46C1339B7FFEA3" authorityName="Deb" authorityYear="1989" box="[1164,1312,317,341]" class="Malacostraca" family="Pilumnidae" genus="Myopilumnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB46C1339B7FFEA3" box="[1164,1312,317,341]" italics="true" pageId="16" pageNumber="351">Myopilumnus</emphasis>
</taxonomicName>
, and
<taxonomicName id="52AA22C12C5CCE3AFAA8C1339BE4FEA3" box="[1378,1467,317,341]" class="Malacostraca" family="Pilumnidae" genus="Hosekia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFAA8C1339BE4FEA3" box="[1378,1467,317,341]" italics="true" pageId="16" pageNumber="351">Hosekia</emphasis>
</taxonomicName>
. The chelae of
<taxonomicName id="52AA22C12C5CCE3AFC23C1539A24FE83" box="[1001,1147,349,373]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFC23C1539A24FE83" box="[1001,1147,349,373]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
and
<taxonomicName id="52AA22C12C5CCE3AFB7DC1539B00FE83" authorityName="Miers" authorityYear="1884" box="[1207,1375,349,373]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFB7DC1539B00FE83" box="[1207,1375,349,373]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
are also stouter than in
<taxonomicName id="52AA22C12C5CCE3AFC17C1739A10FE63" authorityName="Tesch" authorityYear="1918" box="[989,1103,381,405]" class="Malacostraca" family="Pilumnidae" genus="Lophoplax" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFC17C1739A10FE63" box="[989,1103,381,405]" italics="true" pageId="16" pageNumber="351">Lophoplax</emphasis>
</taxonomicName>
and allies, and both have a shallow biconcave posterior margin of the epistome with the median lobe broader and subtruncate (
<figureCitation id="0D9145C72C5CCE3AFB4BC1B39ABFFE23" box="[1153,1248,445,469]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10D</figureCitation>
) (versus median lobe broadly triangular; Ng et al., 2022: fig. 3F).
<taxonomicName id="52AA22C12C5CCE3AFAE7C1D39BE4FE03" box="[1325,1467,477,501]" class="Malacostraca" family="Pilumnidae" genus="Striacoeloma" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFAE7C1D39BE4FE03" box="[1325,1467,477,501]" italics="true" pageId="16" pageNumber="351">Striacoeloma</emphasis>
</taxonomicName>
, however, differs from
<taxonomicName id="52AA22C12C5CCE3AFBF0C1F39A81FDE3" authorityName="Miers" authorityYear="1884" box="[1082,1246,509,533]" class="Malacostraca" family="Pilumnidae" genus="Cryptocoeloma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="A7DE85502C5CCE3AFBF0C1F39A81FDE3" box="[1082,1246,509,533]" italics="true" pageId="16" pageNumber="351">Cryptocoeloma</emphasis>
</taxonomicName>
in having the dorsal surface of carapace ornamented with transverse, corrugated swellings (
<figureCitation id="0D9145C72C5CCE3AFC7FC2339A6AFDA3" box="[949,1077,573,597]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10AC</figureCitation>
) (versus surface granulate and rugose but without swellings and regions well demarcated; Ng et al., 2022: fig. 2); the frontal and anterolateral margins have tufts of short and long setae but not dense or brush-like (
<figureCitation id="0D9145C72C5CCE3AFA5DC2939DCDFD23" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A, B</figureCitation>
) (versus margins evenly lined with dense short and long setae, appearing brush-like; Ng et al., 2022: figs. 1A, B, 2A, B, 5A, B, 7A, B); the outer surface of the chelae are covered with scattered long and short setae, not dense or brush-like (
<figureCitation id="0D9145C72C5CCE3AFC2CC3339A13FCA3" box="[998,1100,829,853]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10A</figureCitation>
) (versus lined with dense brush-like setae; Ng et al., 2022: figs. 1A, B, 2A, B, 5A, B, 7A, B); the ischium of the third maxilliped is subquadrate and shorter (
<figureCitation id="0D9145C72C5CCE3AFC56C3939DA5FC43" box="[924,1018,925,949]" captionStart="Fig" captionStartId="17.[113,148,1956,1978]" captionTargetBox="[113,1451,268,1941]" captionTargetId="figure-7@17.[113,1451,268,1941]" captionTargetPageId="17" captionText="Fig. 10. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, dorsal habitus (right side denuded); B, dorsal view of carapace (right side denuded); C, subfrontal view of cephalothorax (right side denuded); D, frontal view of cephalothorax (right side denuded); E, right third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14682787" httpUri="https://zenodo.org/record/14682787/files/figure.png" pageId="16" pageNumber="351">Fig. 10E</figureCitation>
) (length 1.2× width versus 1.3×; Ng et al., 2022: fig. 4A); and the ambulatory merus is dentate along the extensor margin (
<figureCitation id="0D9145C72C5CCE3AFBE5C3D39ACDFC03" box="[1071,1170,989,1013]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">Fig. 11D</figureCitation>
) (versus smooth; Ng et al., 2022: figs. 2, 4B); and the vulva is proportionately larger, occupying most of the space of sternite 5 (
<figureCitation id="0D9145C72C5CCE3AFACDC4139B39FBC3" box="[1287,1382,1053,1077]" captionStart="Fig" captionStartId="18.[135,170,1024,1046]" captionTargetBox="[135,1473,189,1008]" captionTargetId="figure-8@18.[135,1473,189,1008]" captionTargetPageId="18" captionText="Fig. 11. Striacoeloma tubur, new genus and species, holotype female (8.6 × 6.3 mm) (AM P107758), Northern Territory, Australia. A, buccal cavity, anterior thoracic sternum and pleon; B, sternopleonal cavity and vulvae; C, right chela; D, right P4 and P5." figureDoi="http://doi.org/10.5281/zenodo.14682789" httpUri="https://zenodo.org/record/14682789/files/figure.png" pageId="16" pageNumber="351">Fig. 11B</figureCitation>
) (versus vulva smaller, occupying two-thirds the space of sternite 5; Ng et al., 2022: fig. 7E, F).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,372 @@
<document id="5EFACFEF30E8D8FD67F0103C41545079" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC1FFCC218A1797EDADFB7B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Trigonocera ubinensis Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="314" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="312" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5004E272C9B2C59A3712188BE949D21A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90C0ECD24311D8013AD5641C4161C89B">
<mods:title id="6D8390232D8A81FEFB70E6C96E639653">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="260F58297FA9DAFFE7EDC6B10E2A43EE" type="personal">
<mods:role id="3114A026FB5EB8330CD8D6C493AD6557">
<mods:roleTerm id="DB9684EDC612CE8AC12D6627C7E9605C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A1DFB3F2F50FA939B5CE2E5970E1B6A6">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="06A85D6CA9163F8F6ED6C43C4D26B90F" type="personal">
<mods:role id="BCA08DB81532BD2DF179925227B31670">
<mods:roleTerm id="AD87B88476319FF25C600CEFC50B0C6E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9704E919156A72F33A28322CFD29CED">Velde,</mods:namePart>
</mods:name>
<mods:name id="8FEECCFACA1A2FF164CCA98DB85F0F18" type="personal">
<mods:role id="F43FD34D402182806AA0BE84B0042F3C">
<mods:roleTerm id="94BC73FDAFB0454A8371F11C85FE34E0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D0820E10045C1F1A04687003151F2C6">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="51E94BCB8631E9278E3B17393FFBF8D7">text</mods:typeOfResource>
<mods:relatedItem id="14B1117FFD3F0A8CDEB3124E394A9C1F" type="host">
<mods:titleInfo id="A39E0B772BCD05CA3E5FBA4A3E456D4D">
<mods:title id="3BA7B351EEF352863E279FA5D85B8271">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="878A98D429849D020C81B21CF3E81CDF">
<mods:date id="B72709D7B118DD7140BEF6E87932A06F">2024</mods:date>
<mods:detail id="B166B18BDD60C060A551FFE9D3E5858A" type="pubDate">
<mods:number id="F24222E8CF79EEB8E12B51B9FE46EBBD">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="8DB33847DFFF4434FB9A00B75296B6B8" type="volume">
<mods:number id="5F8FB059E40B4B5AF95CEF727A5FABF0">72</mods:number>
</mods:detail>
<mods:extent id="A7A37713227166483CC3432350A8DB5D" unit="page">
<mods:start id="30929743E3D4E3E9E61DD4FE7142B9FE">303</mods:start>
<mods:end id="A741E7D3816EE1642CC9A1937CB1B911">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F7DD25FAB7A83FD28CA37998CD67D141">journal article</mods:classification>
<mods:identifier id="5CC453E7CAD72697AEAA81BD784D6D2D" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="4ADAE196BDBA82E32716616FF7CC5AFA" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="8E195BEA7296B53D7E425AA6CDDCF0F3" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="B8561C2E110326187D0BE72EBECE5D51" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFC1FFCC218A1797EDADFB7B" ID-DOI="http://doi.org/10.5281/zenodo.14683096" ID-Zenodo-Dep="14683096" LSID="urn:lsid:plazi:treatment:461287F6FFC1FFCC218A1797EDADFB7B" httpUri="http://treatment.plazi.org/id/461287F6FFC1FFCC218A1797EDADFB7B" lastPageId="11" lastPageNumber="314" pageId="9" pageNumber="312">
<subSubSection id="86A1656BFFC1FFCE218A1797ED0FFAE7" box="[933,1332,1265,1289]" pageId="9" pageNumber="312" type="nomenclature">
<paragraph id="CE0436E0FFC1FFCE218A1797ED0FFAE7" blockId="9.[933,1332,1265,1321]" box="[933,1332,1265,1289]" pageId="9" pageNumber="312">
<heading id="954C818CFFC1FFCE218A1797ED0FFAE7" bold="true" box="[933,1332,1265,1289]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="312" reason="2">
<emphasis id="FCCFEAF2FFC1FFCE218A1797ED0FFAE7" bold="true" box="[933,1332,1265,1289]" pageId="9" pageNumber="312">
<taxonomicName id="09BB4D63FFC1FFCE218A1797ECA4FAE7" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[933,1183,1265,1289]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFC1FFCE218A1797ECA4FAE7" bold="true" box="[933,1183,1265,1289]" italics="true" pageId="9" pageNumber="312">Trigonocera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC1FFCE26811797ED0FFAE7" box="[1198,1332,1265,1289]" pageId="9" pageNumber="312" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC1FFCE26001677EC93FAC7" box="[1071,1192,1297,1321]" pageId="9" pageNumber="312" type="description">
<paragraph id="CE0436E0FFC1FFCE26001677EC93FAC7" blockId="9.[933,1332,1265,1321]" box="[1071,1192,1297,1321]" pageId="9" pageNumber="312">
(
<figureCitation id="56802A65FFC1FFCE26181677ECBEFAC7" box="[1079,1157,1297,1321]" captionStart="Fig" captionStartId="9.[813,852,1160,1182]" captionTargetBox="[813,1451,189,1144]" captionTargetId="figure-495@9.[813,1451,189,1144]" captionTargetPageId="9" captionText="Fig. 8. Trigonocera ubinensis, new species, male habitus (ZRCBDP0276611). Photograph by Rene Ong." figureDoi="http://doi.org/10.5281/zenodo.14683076" httpUri="https://zenodo.org/record/14683076/files/figure.png" pageId="9" pageNumber="312">Figs. 8</figureCitation>
,
<figureCitation id="56802A65FFC1FFCE26BD1677EC9BFAC7" box="[1170,1184,1297,1321]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="9" pageNumber="312">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC1FFCE21021637EC1DF9C7" pageId="9" pageNumber="312" type="diagnosis">
<paragraph id="CE0436E0FFC1FFCE21021637EC1DF9C7" blockId="9.[813,1452,1361,1577]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE21021637EB9EFA87" bold="true" box="[813,933,1361,1385]" pageId="9" pageNumber="312">Diagnosis.</emphasis>
A medium-sized species (about
<quantity id="09439B05FFC1FFCE27311637ED5BFA87" box="[1310,1376,1361,1385]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="312" unit="mm" value="3.0">3 mm</quantity>
long). Scape bare. Males with postpedicel elongate triangular 22.5 times as long as wide, arista 1.5 times longer than postpedicel. A long lower propleural bristle. Tergites 13 yellowish white with apical border brown. Hind coxa and metepimeron yellow. Female with postpedicel rounded, a little shorter than high.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC1FFCE21021537EB95F967" pageId="9" pageNumber="312" type="etymology">
<paragraph id="CE0436E0FFC1FFCE21021537EB95F967" blockId="9.[812,1451,1617,1673]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE21021537EB94F987" bold="true" box="[813,943,1617,1641]" pageId="9" pageNumber="312">Etymology.</emphasis>
The new species is named after the
<typeStatus id="11008842FFC1FFCE27071534ED6DF984" box="[1320,1366,1618,1642]" pageId="9" pageNumber="312">type</typeStatus>
locality Pulau Ubin.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC1FFCC210215D7ED69FE5B" lastPageId="11" lastPageNumber="314" pageId="9" pageNumber="312" type="materials_examined">
<paragraph id="CE0436E0FFC1FFCE210215D7ECFBF8C7" blockId="9.[811,1452,1713,1833]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE210215D7EC39F927" bold="true" box="[813,1026,1713,1737]" pageId="9" pageNumber="312">Material examined</emphasis>
.
<materialsCitation id="7ED33CBDFFC1FFCE262115D7ECFBF8C7" collectingDate="2018-05-17" collectionCode="LKCNH" country="Singapore" latitude="1.40645" location="Pulau Ubin" municipality="Puaka Hill" pageId="9" pageNumber="312" specimenCode="PU14, ZRCBDP0256851" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFC1FFCE262115D7EC4AF927" box="[1038,1137,1713,1737]" pageId="9" pageNumber="312" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFC1FFCE269915D7ED1EF927" box="[1206,1317,1713,1737]" name="Singapore" pageId="9" pageNumber="312">Singapore</collectingCountry>
,
<location id="CB64603BFFC1FFCE270115D7ED93F927" LSID="urn:lsid:plazi:treatment:461287F6FFC1FFCC218A1797EDADFB7B:CB64603BFFC1FFCE270115D7ED93F927" box="[1326,1448,1713,1737]" country="Singapore" latitude="1.40645" municipality="Puaka Hill" name="Pulau Ubin" pageId="9" pageNumber="312">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFC1FFCE210415B7EBA5F907" box="[811,926,1745,1769]" pageId="9" pageNumber="312">Puaka Hill</collectingMunicipality>
(
<specimenCode id="9E1D9E9BFFC1FFCE218015B7EBCBF907" box="[943,1008,1745,1769]" collectionCode="PU" country="USA" name="Princeton University" pageId="9" pageNumber="312">PU14</specimenCode>
),
<date id="BA051020FFC1FFCE262A15B7ECA1F907" box="[1029,1178,1745,1770]" pageId="9" pageNumber="312" value="2018-05-17">
<collectingDate id="AA41E9C8FFC1FFCE262A15B7ECA1F907" box="[1029,1178,1745,1770]" pageId="9" pageNumber="312" value="2018-05-17">17 May 2018</collectingDate>
</date>
, (
<geoCoordinate id="AB8F5027FFC1FFCE268015B6ED7CF907" box="[1199,1351,1744,1769]" degrees="1" direction="north" minutes="24" orientation="latitude" pageId="9" pageNumber="312" precision="1" seconds="23.22" value="1.40645">1°2423.22″N</geoCoordinate>
,103°573 0.60″E), secondary forest (
<specimenCode id="9E1D9E9BFFC1FFCE267D1597ED22F8E7" box="[1106,1305,1777,1801]" collectionCode="ZRCBDP" pageId="9" pageNumber="312">ZRCBDP0256851</specimenCode>
in
<collectionCode id="A8AAAE25FFC1FFCE276F1597ED9AF8E7" box="[1344,1441,1777,1801]" pageId="9" pageNumber="312">LKCNH</collectionCode>
). GenBank accession code: PP893301.
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFC1FFCE21021437EBC4F827" blockId="9.[812,1452,1873,1993]" pageId="9" pageNumber="312">
<materialsCitation id="7ED33CBDFFC1FFCE21021437EB5EF867" collectingDate="2018-04-05" collectionCode="PU" country="Singapore" location="Pulau Ubin" pageId="9" pageNumber="312" specimenCode="PU14" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<emphasis id="FCCFEAF2FFC1FFCE21021437EB9DF887" bold="true" box="[813,934,1873,1897]" pageId="9" pageNumber="312">
<typeStatus id="11008842FFC1FFCE21021437EB9AF887" box="[813,929,1873,1897]" pageId="9" pageNumber="312" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFC1FFCE21831437EC20F887" box="[940,1051,1873,1897]" name="Singapore" pageId="9" pageNumber="312">Singapore</collectingCountry>
,
<specimenCount id="D8BDFD69FFC1FFCE260A1437EC41F887" box="[1061,1146,1873,1897]" count="3" pageId="9" pageNumber="312" type="male">3 males</specimenCount>
,
<location id="CB64603BFFC1FFCE26AB1437ECC6F887" LSID="urn:lsid:plazi:treatment:461287F6FFC1FFCC218A1797EDADFB7B:CB64603BFFC1FFCE26AB1437ECC6F887" box="[1156,1277,1873,1897]" country="Singapore" name="Pulau Ubin" pageId="9" pageNumber="312">Pulau Ubin</location>
(
<specimenCode id="9E1D9E9BFFC1FFCE27241437ED77F887" box="[1291,1356,1873,1897]" collectionCode="PU" country="USA" name="Princeton University" pageId="9" pageNumber="312">PU14</specimenCode>
),
<date id="BA051020FFC1FFCE27701437EB5EF867" pageId="9" pageNumber="312" value="2018-04-05">
<collectingDate id="AA41E9C8FFC1FFCE27701437EB5EF867" pageId="9" pageNumber="312" value="2018-04-05">5 April 2018</collectingDate>
</date>
</materialsCitation>
, secondary forest (Ma9837,
<collectionCode id="A8AAAE25FFC1FFCE26881417ECC7F867" box="[1191,1276,1905,1929]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="9" pageNumber="312" type="Museum">RBINS</collectionCode>
):
<materialsCitation id="7ED33CBDFFC1FFCE273E1417EC46F847" collectionCode="PU, ZRCBDP" country="Singapore" pageId="9" pageNumber="312" specimenCode="PU14, ZRCBDP0256869" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFC1FFCE273E1417ED5BF867" box="[1297,1376,1905,1929]" count="1" pageId="9" pageNumber="312" type="male">1 male</specimenCount>
, Pulau Ubin,
<specimenCode id="9E1D9E9BFFC1FFCE214114F7EB95F847" box="[878,942,1937,1961]" collectionCode="PU" country="USA" name="Princeton University" pageId="9" pageNumber="312">PU14</specimenCode>
,
<specimenCode id="9E1D9E9BFFC1FFCE219814F7EC46F847" box="[951,1149,1937,1961]" collectionCode="ZRCBDP" pageId="9" pageNumber="312">ZRCBDP0256869</specimenCode>
</materialsCitation>
_
<materialsCitation id="7ED33CBDFFC1FFCE26A514F7EBC0F827" collectingDate="2018-05-17" collectionCode="MIPD, PU" country="Singapore" location="Nabilah-Kareen" pageId="9" pageNumber="312" specimenCode="MIPD17, PU14" specimenCount="1" typeStatus="paratype">
<location id="CB64603BFFC1FFCE26A514F7ED0DF847" LSID="urn:lsid:plazi:treatment:461287F6FFC1FFCC218A1797EDADFB7B:CB64603BFFC1FFCE26A514F7ED0DF847" box="[1162,1334,1937,1961]" country="Singapore" name="Nabilah-Kareen" pageId="9" pageNumber="312">Nabilah-Kareen</location>
_
<specimenCode id="9E1D9E9BFFC1FFCE276B14F7EDA5F847" box="[1348,1438,1937,1961]" collectionCode="MIPD" pageId="9" pageNumber="312">MIPD17</specimenCode>
_
<specimenCode id="9E1D9E9BFFC1FFCE210214D7EB51F827" box="[813,874,1969,1993]" collectionCode="PU" country="USA" name="Princeton University" pageId="9" pageNumber="312">PU14</specimenCode>
_
<date id="BA051020FFC1FFCE215714D7EBC0F827" box="[888,1019,1969,1993]" pageId="9" pageNumber="312" value="2018-05-17">
<collectingDate id="AA41E9C8FFC1FFCE215714D7EBC0F827" box="[888,1019,1969,1993]" pageId="9" pageNumber="312" value="2018-05-17">17May2018</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<caption id="9AC46668FFC2FFCD22A717A2EB70FB18" ID-DOI="http://doi.org/10.5281/zenodo.14683078" ID-Zenodo-Dep="14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="10" pageNumber="313" startId="10.[136,171,1220,1242]" targetBox="[139,1475,191,1199]" targetPageId="10" targetType="figure">
<paragraph id="CE0436E0FFC2FFCD22A717A2EB70FB18" blockId="10.[136,1475,1220,1271]" pageId="10" pageNumber="313">
Fig. 9.
<taxonomicName id="09BB4D63FFC2FFCD22FF17A3E990FB37" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[208,427,1220,1242]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="313" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFC2FFCD22FF17A3E990FB37" box="[208,427,1220,1242]" italics="true" pageId="10" pageNumber="313">Trigonocera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC2FFCD239617A3EA0AFB34" box="[441,561,1220,1242]" pageId="10" pageNumber="313" rank="species">new species</taxonomicNameLabel>
, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC2FFCD22A71648EA91FA68" blockId="10.[136,775,1326,1414]" pageId="10" pageNumber="313">
Barcode:
<collectionCode id="A8AAAE25FFC2FFCD232F1648E94FFAA8" box="[256,372,1326,1350]" pageId="10" pageNumber="313">ZRCBDP</collectionCode>
0276611_WanTing_SGDolis025_
<collectionCode id="A8AAAE25FFC2FFCD22A71628E897FA88" box="[136,172,1358,1382]" country="USA" name="Princeton University" pageId="10" pageNumber="313">PU</collectionCode>
14_
<date id="BA051020FFC2FFCD22FD1628E977FA88" box="[210,332,1358,1382]" pageId="10" pageNumber="313" value="2018-04-12">12Apr2018</date>
_imaged; whole mitogenome was done by Darren Yeo. GenBank accession code: PP893303.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD22A716C8E9B5F988" blockId="10.[136,777,1454,1638]" pageId="10" pageNumber="313">
<materialsCitation id="7ED33CBDFFC2FFCD22A716C8E8D8FA08" collectionCode="PU" pageId="10" pageNumber="313" specimenCode="PU14" specimenCount="19">
<emphasis id="FCCFEAF2FFC2FFCD22A716C8E9F0FA28" bold="true" box="[136,459,1454,1478]" pageId="10" pageNumber="313">
<typeStatus id="11008842FFC2FFCD22A716C8E9FDFA28" box="[136,454,1454,1478]" isOtherTypeStatus="true" pageId="10" pageNumber="313">Other material examined</typeStatus>
.
</emphasis>
<specimenCount id="D8BDFD69FFC2FFCD23F816C8EA4EFA28" box="[471,629,1454,1478]" count="19" pageId="10" pageNumber="313" type="generic">19 specimens</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD20AE16C8EAFAFA28" box="[641,705,1454,1478]" collectionCode="PU" country="USA" name="Princeton University" pageId="10" pageNumber="313">PU14</specimenCode>
(
<typeStatus id="11008842FFC2FFCD20FB16C8EB3DFA28" box="[724,774,1454,1478]" pageId="10" pageNumber="313">type</typeStatus>
locality)
</materialsCitation>
,
<materialsCitation id="7ED33CBDFFC2FFCD22DF16A8E985FA08" box="[240,446,1486,1510]" collectionCode="PU" pageId="10" pageNumber="313" specimenCode="PU12" specimenCount="3">
<specimenCount id="D8BDFD69FFC2FFCD22DF16A8E94EFA08" box="[240,373,1486,1510]" count="3" pageId="10" pageNumber="313" type="generic">3 specimens</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD235116A8E985FA08" box="[382,446,1486,1510]" collectionCode="PU" country="USA" name="Princeton University" pageId="10" pageNumber="313">PU12</specimenCode>
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFC2FFCD23E416A8EAB5FA08" box="[459,654,1486,1510]" collectionCode="PU" pageId="10" pageNumber="313" specimenCode="PU13" specimenCount="1">
<specimenCount id="D8BDFD69FFC2FFCD23E416A8EA7DFA08" box="[459,582,1486,1510]" count="1" pageId="10" pageNumber="313" type="generic">1 specimen</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD206116A8EAB5FA08" box="[590,654,1486,1510]" collectionCode="PU" country="USA" name="Princeton University" pageId="10" pageNumber="313">PU13</specimenCode>
</materialsCitation>
; specimen;
<materialsCitation id="7ED33CBDFFC2FFCD22A71688E977F9E8" box="[136,332,1518,1542]" collectionCode="PU" pageId="10" pageNumber="313" specimenCode="PU20" specimenCount="1">
<specimenCount id="D8BDFD69FFC2FFCD22A71688E93FF9E8" box="[136,260,1518,1542]" count="1" pageId="10" pageNumber="313" type="generic">1 specimen</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD23221688E977F9E8" box="[269,332,1518,1542]" collectionCode="PU" country="USA" name="Princeton University" pageId="10" pageNumber="313">PU20</specimenCode>
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFC2FFCD23751688EA12F9E8" box="[346,553,1518,1542]" collectionCode="NSM" pageId="10" pageNumber="313" specimenCode="NSM1" specimenCount="1">
<specimenCount id="D8BDFD69FFC2FFCD23751688E9EEF9E8" box="[346,469,1518,1542]" count="1" pageId="10" pageNumber="313" type="generic">1 specimen</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD23F11688EA12F9E8" box="[478,553,1518,1542]" collectionCode="NSM" country="Canada" httpUri="http://grbio.org/cool/zeys-ppzt" name="Nova Scotia Museum of Natural History" pageId="10" pageNumber="313">NSM1</specimenCode>
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFC2FFCD20191688EB3FF9E8" box="[566,772,1518,1542]" collectionCode="KM" pageId="10" pageNumber="313" specimenCode="KM03" specimenCount="1">
<specimenCount id="D8BDFD69FFC2FFCD20191688EA89F9E8" box="[566,690,1518,1542]" count="1" pageId="10" pageNumber="313" type="generic">1 specimen</specimenCount>
<specimenCode id="9E1D9E9BFFC2FFCD20941688EB3FF9E8" box="[699,772,1518,1542]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="10" pageNumber="313">KM03</specimenCode>
</materialsCitation>
. In total,
<specimenCount id="D8BDFD69FFC2FFCD22C91568E943F9C8" box="[230,376,1550,1574]" count="27" pageId="10" pageNumber="313" type="generic">27 specimens</specimenCount>
were barcoded. They form a cluster of 0.3%, meaning that there is only 1 base pair difference between the haplotypes.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD22A715E8EAFEF948" blockId="10.[136,709,1678,1702]" box="[136,709,1678,1702]" pageId="10" pageNumber="313">
<emphasis id="FCCFEAF2FFC2FFCD22A715E8E8F2F948" bold="true" box="[136,201,1678,1702]" pageId="10" pageNumber="313">Male.</emphasis>
Length: body
<quantity id="09439B05FFC2FFCD234415E8E9CBF948" box="[363,496,1678,1702]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="10" pageNumber="313" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.82.9 mm</quantity>
; wing
<quantity id="09439B05FFC2FFCD201415E8EAFAF948" box="[571,705,1678,1702]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="10" pageNumber="313" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.42.5 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD22A715A8E9EEF828" blockId="10.[136,777,1742,1990]" pageId="10" pageNumber="313">
<emphasis id="FCCFEAF2FFC2FFCD22A715A8E8FDF908" bold="true" box="[136,198,1742,1766]" pageId="10" pageNumber="313">Head</emphasis>
. Frons wide, wider than base of postpedicel. Face wide, parallel-sides a little narrower than frons. A pair of long retro- and lateroclinate ocellars, followed by pair of short proclinate bristles (⅓ length of the anterior bristles). A pair of short postvertical bristle, no vertical bristle. Upper postocular bristles short, black, and uniserial above, below longer, white, and only near mouth a few additional bristles behind the row of postoculars.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD22A71488EC30F9C8" blockId="10.[135,775,2030,2118]" lastBlockId="10.[835,1476,1326,1574]" pageId="10" pageNumber="313">
<emphasis id="FCCFEAF2FFC2FFCD22A71488E8C7F7E8" bold="true" box="[136,252,2030,2054]" pageId="10" pageNumber="313">Antennae.</emphasis>
With brown postpedicel, black scape, and pedicel (
<figureCitation id="56802A65FFC2FFCD22A01B68E8EFF7C8" box="[143,212,2062,2086]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="10" pageNumber="313">Fig. 9</figureCitation>
). Scape a little longer than pedicel. Pedicel with a dorsal bristle a little longer than width of pedicel. Tip of pedicel conical going inside the postpedicel, but no thumb-like projection on pedicel reaching over the postpedicel. Postpedicel elongate triangular (
<quantity id="09439B05FFC2FFCD26B51608ECC2FA68" box="[1178,1273,1390,1414]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="10" pageNumber="313" unit="mm" value="0.31">0.31 mm</quantity>
/
<quantity id="09439B05FFC2FFCD27281608ED58FA68" box="[1287,1379,1390,1414]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="313" unit="mm" value="0.15">0.15mm</quantity>
), 2 times as long as wide at base. Ventral base of postpedicel not overlapping the pedicel (
<figureCitation id="56802A65FFC2FFCD267216C8EC82FA28" box="[1117,1209,1454,1478]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="10" pageNumber="313">Fig. 9A</figureCitation>
). Arista subapical (
<figureCitation id="56802A65FFC2FFCD27BA16C8EB5CFA08" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="10" pageNumber="313">Fig. 9A</figureCitation>
, detail of apex) about 1.5 times as long as postpedicel. Palpus white, with a long black apical bristle and 5 short additional bristles.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD216C1528EC12F928" blockId="10.[835,1474,1614,1734]" pageId="10" pageNumber="313">
<emphasis id="FCCFEAF2FFC2FFCD216C1528EBA6F988" bold="true" box="[835,925,1614,1638]" pageId="10" pageNumber="313">Thorax</emphasis>
. 8 pairs of short acr in anterior half; 4 long dorsocentrals, almost equally long. 2 strong marginal scutellars with a tiny hair at outside. A long propleural black bristle (
<figureCitation id="56802A65FFC2FFCD21F615C8EC27F928" box="[985,1052,1710,1734]" captionStart="Fig" captionStartId="9.[813,852,1160,1182]" captionTargetBox="[813,1451,189,1144]" captionTargetId="figure-495@9.[813,1451,189,1144]" captionTargetPageId="9" captionText="Fig. 8. Trigonocera ubinensis, new species, male habitus (ZRCBDP0276611). Photograph by Rene Ong." figureDoi="http://doi.org/10.5281/zenodo.14683076" httpUri="https://zenodo.org/record/14683076/files/figure.png" pageId="10" pageNumber="313">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFC2FFCD216C1588EBE1F8A8" blockId="10.[835,1475,1774,1862]" pageId="10" pageNumber="313">
<emphasis id="FCCFEAF2FFC2FFCD216C1588EB43F8E8" bold="true" box="[835,888,1774,1798]" pageId="10" pageNumber="313">Legs</emphasis>
. Entirely yellow, except for the brown mid coxa and a pale brownish patch on the base of the hind coxa. All bristles black.
</paragraph>
<paragraph id="CE0436E0FFC2FFCD216C1409ED10F7A8" blockId="10.[834,1476,1902,2119]" pageId="10" pageNumber="313">Fore legs. Coxa with 3 long black anterior bristles in apical half, a few short bristles in basal half, 2 short apical bristles. Femur with 3 preapical posteroventrals as long as femur is wide; 1 short preapical posterior, 3 fine ventrals near middle nearly half as long as femur is wide. Tibia with 1 short dorsal at middle, 1 ad on basal quarter, 1 short preapical dorsal. A pair of very small claws and minute pulvilli.</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E13DBE922FDFB" blockId="11.[113,755,189,533]" pageId="11" pageNumber="314">Mid legs. Coxa with an exterior black bristle at base a little longer than coxa is high and a shorter exterior near apex; anteriorly set with long bristles. Trochanter with 2 strong ventral bristles, longer than trochanter is long. Femur with a row of short anteroventral bristles, half as long as femur is wide. An anterior preapical bristle as long as femur is wide and 2 preapical anteroventrals. Tibia with a short ad near base and strong ad at basal fifth, a strong dorsal near middle and a shorter anteroventral below; 2 strong ventral bristles in apical crown. A pair of very small claws and minute pulvilli.</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E115BE9B6FCDB" blockId="11.[113,753,573,821]" pageId="11" pageNumber="314">Hind legs. Coxa with a long exterior at base, as long as coxa is high and a minute bristle near apex. Trochanter with a long anterodorsal bristle. Femur with about 5 fine anteroventral bristles in apical half, half as long as femur is wide; a strong anterior preapical a little longer than femur is wide. Tibia with 5 short posterodorsal bristles; 2 anterodorsal, 3 short anteroventral bristles. A pair of short claws and minute pulvilli.</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E103BE927FC1B" blockId="11.[112,753,861,1013]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC225E103BE88FFC9B" bold="true" box="[113,180,861,885]" pageId="11" pageNumber="314">Wings</emphasis>
. Hyaline with a faint brownish tinge, veins brownish. Veins
<collectionCode id="A8AAAE25FFC3FFCC22951018E8F7FC7B" box="[186,204,894,917]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="314">R</collectionCode>
<subScript id="523F34A5FFC3FFCC22E210EFE8DDFC79" attach="left" box="[205,230,905,919]" fontSize="6" pageId="11" pageNumber="314">4+5</subScript>
and M
<subScript id="523F34A5FFC3FFCC231510EFE968FC79" attach="left" box="[314,339,905,919]" fontSize="6" pageId="11" pageNumber="314">1+2</subScript>
running parallel in apical half, M
<subScript id="523F34A5FFC3FFCC20F810EFEACBFC79" attach="left" box="[727,752,905,919]" fontSize="6" pageId="11" pageNumber="314">1+2</subScript>
with a faint bent before middle. Squama yellowish white with very long black bristles; haltere white. Metepimeron entirely yellow.
</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E177BE905FAFB" blockId="11.[112,755,1053,1302]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC225E177BE8DBFBDB" bold="true" box="[113,224,1053,1077]" pageId="11" pageNumber="314">Abdomen</emphasis>
. Tergites 13 yellowish white, but apical border of these 3 tergites with a brown rim. Base of tergite 3 also with a brown rim. Tergite 4 with a yellowish spot at side. All tergites with short dorsal bristles and long apical bristles. Tergite 6 lacking bristles. Sternites 15 yellowish white; sternites 24 each with at least 4 moderately long bristles; no bristles on sternite 5. Tergite 8 (
<figureCitation id="56802A65FFC3FFCC23DE17BBEA7FFB1B" box="[497,580,1245,1269]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="11" pageNumber="314">Fig. 9B</figureCitation>
) with 4 equally strong macrosetae.
</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E165BE90BF9BB" blockId="11.[112,753,1341,1621]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC225E165BE912FABB" bold="true" box="[113,297,1341,1365]" pageId="11" pageNumber="314">Male terminalia.</emphasis>
<figureCitation id="56802A65FFC3FFCC2301165BE957FABB" box="[302,364,1341,1365]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="11" pageNumber="314">Fig. 9</figureCitation>
. Cercus yellow including the rounded basal lobe. Apical appendage of cercus at least 3 times as long as basal rounded part of cercus. Dorsal surstylus tubular with an apical bristle (
<figureCitation id="56802A65FFC3FFCC237016FBE98DFA5B" box="[351,438,1437,1461]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="11" pageNumber="314">Fig. 9D</figureCitation>
). Postgonite with rectangular bent near middle, at base swollen with ventral part rounded (
<figureCitation id="56802A65FFC3FFCC225516BBE8EAFA1B" box="[122,209,1501,1525]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="11" pageNumber="314">Fig. 9C</figureCitation>
). Ventral surstylus yellowish brown, with apex a little darkened. Epandrial lobe broad, trapezoid with 2 long subapical bristles and a small bristle on the inside, near middle (
<figureCitation id="56802A65FFC3FFCC22E3155BE91FF9BB" box="[204,292,1597,1621]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="11" pageNumber="314">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E151BEA88F97B" blockId="11.[113,691,1661,1685]" box="[113,691,1661,1685]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC225E151BE8F0F97B" bold="true" box="[113,203,1661,1685]" pageId="11" pageNumber="314">Female.</emphasis>
Length: body
<quantity id="09439B05FFC3FFCC2342151BE9E4F97B" box="[365,479,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="11" pageNumber="314" unit="mm" value="2.95" valueMax="3.0" valueMin="2.9">2.93 mm</quantity>
; wing
<quantity id="09439B05FFC3FFCC2006151BEA94F97B" box="[553,687,1661,1685]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="11" pageNumber="314" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.32.4 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFC3FFCC225E15DBEBD1FF1B" blockId="11.[113,755,1725,2037]" lastBlockId="11.[813,1451,189,245]" pageId="11" pageNumber="314">Postpedicel a little shorter than long, with a rounded apex (not triangular). Postpedicel paler brown than the dark brown scape and pedicel. A pair of very long proclinate vertical bristles present, which is absent in males. Proboscis much stouter than in males with a larger yellowish palpus. Bristling and colouration of legs identical to males. Only mid coxa brown, while hind coxa and metepimeron are yellow. Colouration of tergites (tergite 13 yellowish white with posterior border browned and tergite 4 mainly brown with a yellow patch at side; tergite 5 entirely brown) and sternites (sternites 14 yellowish white, sternite 5 brown) identical to male.</paragraph>
<paragraph id="CE0436E0FFC3FFCC21021278ED69FE5B" blockId="11.[813,1452,285,437]" pageId="11" pageNumber="314">
It is difficult to distinguish the females of these
<taxonomicName id="09BB4D63FFC3FFCC27091278ED90FEDB" authorityName="Becker" authorityYear="1902" box="[1318,1451,286,309]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC3FFCC27091278ED90FEDB" box="[1318,1451,286,309]" italics="true" pageId="11" pageNumber="314">Trigonocera</emphasis>
</taxonomicName>
in a sample from
<taxonomicName id="09BB4D63FFC3FFCC21C2125BEC64FEBB" baseAuthorityName="Grichanov" baseAuthorityYear="2011" box="[1005,1119,317,341]" class="Insecta" family="Dolichopodidae" genus="Diaphorus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC3FFCC21C2125BEC64FEBB" box="[1005,1119,317,341]" italics="true" pageId="11" pageNumber="314">Diaphorus</emphasis>
</taxonomicName>
females. Females have a short pedicel with a rounded apex, while males have a long triangular postpedicel. The
<specimenCount id="D8BDFD69FFC3FFCC26701218ECDFFE7B" box="[1119,1252,381,405]" count="2" pageId="11" pageNumber="314" type="female">two females</specimenCount>
reported here are associated with the males thanks to the barcoding.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC3FFCC210212BBEDADFB7B" pageId="11" pageNumber="314" type="discussion">
<paragraph id="CE0436E0FFC3FFCC210212BBED97FCFB" blockId="11.[811,1454,477,789]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC210212BBEB80FE1B" bold="true" box="[813,955,477,501]" pageId="11" pageNumber="314">Comments.</emphasis>
We refer to the comments made under the previous species
<taxonomicName id="09BB4D63FFC3FFCC260A1298ED3EFDFB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1061,1285,509,533]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFC3FFCC260A1298ED3EFDFB" box="[1061,1285,509,533]" italics="true" pageId="11" pageNumber="314">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC3FFCC27371298ED93FDFB" box="[1304,1448,509,533]" pageId="11" pageNumber="314" rank="species">new species</taxonomicNameLabel>
.
<taxonomicName id="09BB4D63FFC3FFCC21021178EC17FDDB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[813,1068,541,565]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFC3FFCC21021178EC17FDDB" box="[813,1068,541,565]" italics="true" pageId="11" pageNumber="314">Trigonocera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC3FFCC26111178ECF1FDDB" box="[1086,1226,541,565]" pageId="11" pageNumber="314" rank="species">new species</taxonomicNameLabel>
, could be confused with some
<taxonomicName id="09BB4D63FFC3FFCC2184115BEC4CFDBB" box="[939,1143,573,597]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC2184115BEC24FDBB" box="[939,1055,573,597]" italics="true" pageId="11" pageNumber="314">Diaphorus</emphasis>
species
</taxonomicName>
in also having a triangular postpedicel.
<taxonomicName id="09BB4D63FFC3FFCC219B1138ED39FD9B" authority="Olejnicek, 2005" authorityName="Olejnicek" authorityYear="2005" box="[948,1282,605,629]" class="Insecta" family="Dolichopodidae" genus="Diaphorus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="314" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="FCCFEAF2FFC3FFCC219B1138EC74FD9B" box="[948,1103,605,629]" italics="true" pageId="11" pageNumber="314">D. longicornis</emphasis>
<bibRefCitation id="AA2A4B11FFC3FFCC2678113BED39FD9B" author="Olejnicek J" box="[1111,1282,605,629]" pageId="11" pageNumber="314" pagination="499 - 506" refId="ref14752" refString="Olejnicek J (2005) Some new and little known Diaphorus species (Diptera, Dolichopodidae) from the Oriental Region. Biologia, 60 (5): 499 - 506." type="journal article" year="2005">Olejníček, 2005</bibRefCitation>
</taxonomicName>
described from North
<collectingCountry id="B6AC7670FFC3FFCC21401118EB99FD7B" box="[879,930,638,661]" name="Laos" pageId="11" pageNumber="314">Laos</collectingCountry>
is such a species with a long triangular postpedicel 1.5 times as long as high, also with only mid coxa brown, and fore and hind coxa yellow. However, the fore tarsus lacks claws and has pulvilli longer than apical tarsomere and the abdomen lacks yellow patches in the latter species.
</paragraph>
<paragraph id="CE0436E0FFC3FFCC2102105BEDADFB7B" blockId="11.[811,1454,829,1173]" pageId="11" pageNumber="314">
<emphasis id="FCCFEAF2FFC3FFCC2102105BEC19FCBB" bold="true" box="[813,1058,829,853]" pageId="11" pageNumber="314">National distribution.</emphasis>
The new species is not very common in
<collectingCountry id="B6AC7670FFC3FFCC216A103BEB8FFC9B" box="[837,948,861,885]" name="Singapore" pageId="11" pageNumber="314">Singapore</collectingCountry>
.
<materialsCitation id="7ED33CBDFFC3FFCC21931038ED25FBDB" collectionCode="PU" country="Singapore" location="Singapore" pageId="11" pageNumber="314" specimenCode="PU14, PU13, PU20" specimenCount="2">
It is most abundant in the
<typeStatus id="11008842FFC3FFCC26E81038ECCEFC98" box="[1223,1269,862,886]" pageId="11" pageNumber="314">type</typeStatus>
locality on Pulau Ubin (
<specimenCode id="9E1D9E9BFFC3FFCC215F101BEB8AFC7B" box="[880,945,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="11" pageNumber="314">PU14</specimenCode>
), a secondary forest on Puaka Hill. It was also recorded from another secondary forest patch along the road in front of the Endut Senin campsite on Pulau Ubin. Single specimens were found in the back mangroves of Sungei [= River] Puaka (
<specimenCode id="9E1D9E9BFFC3FFCC21ED109BEC3BFBFB" box="[962,1024,1021,1045]" collectionCode="PU" country="USA" name="Princeton University" pageId="11" pageNumber="314">PU13</specimenCode>
) and on Pulau Ubin (
<specimenCode id="9E1D9E9BFFC3FFCC26F7109BED22FBFB" box="[1240,1305,1021,1045]" collectionCode="PU" country="USA" name="Princeton University" pageId="11" pageNumber="314">PU20</specimenCode>
), respectively. Only
<specimenCount id="D8BDFD69FFC3FFCC21471778EC3CFBDB" box="[872,1031,1053,1077]" count="2" pageId="11" pageNumber="314" type="generic">two specimens</specimenCount>
were found on
<collectingCountry id="B6AC7670FFC3FFCC269D177BED25FBDB" box="[1202,1310,1053,1077]" name="Singapore" pageId="11" pageNumber="314">Singapore</collectingCountry>
</materialsCitation>
<materialsCitation id="7ED33CBDFFC3FFCC2709177BEDA9FB7B" collectorName="Kranji Marshes. A &amp; Preah Khan" country="Cambodia" location="Siem Reap" pageId="11" pageNumber="314" specimenCount="1" stateProvince="Siem Reap">
island itself: a single specimen in the swamp forest of Nee Soon and
<specimenCount id="D8BDFD69FFC3FFCC27A91758EBA9FB9B" count="1" pageId="11" pageNumber="314" type="generic">one specimen</specimenCount>
in the swamp forest of
<collectorName id="634E5336FFC3FFCC26B6173BED58FB98" box="[1177,1379,1117,1142]" pageId="11" pageNumber="314">Kranji Marshes. A</collectorName>
single male was found in
<collectorName id="634E5336FFC3FFCC262D171BECBAFB7B" box="[1026,1153,1149,1173]" pageId="11" pageNumber="314">Preah Khan</collectorName>
(
<collectingRegion id="0C7FF802FFC3FFCC26BE171BED36FB7B" box="[1169,1293,1149,1173]" country="Cambodia" name="Siem Reap" pageId="11" pageNumber="314">Siem Reap</collectingRegion>
),
<collectingCountry id="B6AC7670FFC3FFCC270E171BEDA9FB7B" box="[1313,1426,1149,1173]" name="Cambodia" pageId="11" pageNumber="314">Cambodia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,284 @@
<document id="395F3E52AD765CA180ADE98A24B78DEF" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC4FFC9230E1083E97DFE1B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Medetera stigma Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="316" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="315" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CD1B2C1BC36B3E99DA78B6B2A576531A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BE0DE61539A6800FB5567886D42D2E6B">
<mods:title id="DE39DC881575B2978694F3FD8D993658">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="5F4959D6C50CC8BAFF02944D022DF976" type="personal">
<mods:role id="32E3FF4EE380DE8BC19825862C244F2C">
<mods:roleTerm id="97C117D6F4C8EF391E41A59934BE8E2A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B13122A3568942E26F097ADC3E63C43">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="FCFD2AB1B28E2A8F50601FE5A2B34FB9" type="personal">
<mods:role id="B6348B27E0CCC432D3400FF2EE896613">
<mods:roleTerm id="5400ECA8D0BA19552D9AF72257CFE8F7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF3E16D3F22748AF77B03ABD8C6AF5C9">Velde,</mods:namePart>
</mods:name>
<mods:name id="4A166D0B0FAC93ACBABD35DBFB8C00E2" type="personal">
<mods:role id="4F81191FCDBD6CDD1B2C6216957F99B4">
<mods:roleTerm id="46DE48CA81891EFB734D94B2076B6931">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D2461E3A55C0B07EE9B6E2D38598037">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="0C810729893CFC5E510B344379DD4FEC">text</mods:typeOfResource>
<mods:relatedItem id="7A5C46011FF359CBD90670DAF4343240" type="host">
<mods:titleInfo id="1B234B98F1D90AEF15FDDA7152BE3D01">
<mods:title id="DA508E334FF5C198A0BA3CF49B1D363E">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="0C856DAFDB51F6C45DE354146667E4DE">
<mods:date id="60BBEC73559D5378D652CA91692CA4A4">2024</mods:date>
<mods:detail id="5CDA484E79776DB33C26DC05544E1914" type="pubDate">
<mods:number id="4138E91349F9BA10E6C569279DD216E5">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="55D350E2E25B52E36B31A896AE415FF7" type="volume">
<mods:number id="734878714FD1BDF4F9580097443899FC">72</mods:number>
</mods:detail>
<mods:extent id="10D947CB68F82A15524B006B9D32FC49" unit="page">
<mods:start id="4008AA4EE763B6357E8FE21517BB8D78">303</mods:start>
<mods:end id="C58812E9413933B966DB77BA90946DE1">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="EEA223D3E1E87E33950DE9F9B16F9804">journal article</mods:classification>
<mods:identifier id="D94A934F6A4FF2C4AE2136E3C551B462" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="A46B552C91FDC9BBA96AF36E6BC11A7E" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="653D87121C5A154B646B6B81B151DCE6" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="CDB39BD215D3F05FF5FEDF10A928CA89" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFC4FFC9230E1083E97DFE1B" LSID="urn:lsid:plazi:treatment:461287F6FFC4FFC9230E1083E97DFE1B" httpUri="http://treatment.plazi.org/id/461287F6FFC4FFC9230E1083E97DFE1B" lastPageId="14" lastPageNumber="316" pageId="12" pageNumber="315">
<subSubSection id="86A1656BFFC4FFCB230E1083EA54FC10" box="[289,623,997,1022]" pageId="12" pageNumber="315" type="nomenclature">
<paragraph id="CE0436E0FFC4FFCB230E1083EA54FC10" blockId="12.[289,623,997,1054]" box="[289,623,997,1022]" pageId="12" pageNumber="315">
<heading id="954C818CFFC4FFCB230E1083EA54FC10" bold="true" box="[289,623,997,1022]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="315" reason="2">
<emphasis id="FCCFEAF2FFC4FFCB230E1083EA54FC10" bold="true" box="[289,623,997,1022]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB230E1083E9DAFC10" bold="true" box="[289,481,997,1022]" italics="true" pageId="12" pageNumber="315">
<taxonomicName id="09BB4D63FFC4FFCB230E1083E9E0FC10" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[289,475,997,1022]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">Medetera stigma</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="E7FC5789FFC4FFCB23C61080EA54FC10" box="[489,623,998,1022]" pageId="12" pageNumber="315" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCB23511760EA29FBF0" box="[382,530,1030,1054]" pageId="12" pageNumber="315" type="description">
<paragraph id="CE0436E0FFC4FFCB23511760EA29FBF0" blockId="12.[289,623,997,1054]" box="[382,530,1030,1054]" pageId="12" pageNumber="315">
(
<figureCitation id="56802A65FFC4FFCB23AA1760E9DAFBF0" box="[389,481,1030,1054]" captionStart="Fig" captionStartId="12.[136,172,764,786]" captionTargetBox="[136,774,189,749]" captionTargetId="figure-8@12.[136,774,189,749]" captionTargetPageId="12" captionText="Fig. 10. Medetera stigma, new species, holotype male habitus, dorsal (ZRCBDP0317875). Photograph by Isabella Van de Velde." figureDoi="http://doi.org/10.5281/zenodo.14683080" httpUri="https://zenodo.org/record/14683080/files/figure.png" pageId="12" pageNumber="315">Figs. 10</figureCitation>
,
<figureCitation id="56802A65FFC4FFCB23C11760EA32FBF0" box="[494,521,1030,1054]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCB22A71720E8C6FA70" pageId="12" pageNumber="315" type="materials_examined">
<paragraph id="CE0436E0FFC4FFCB22A71720EA93FB30" blockId="12.[134,775,1094,1246]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71720E917FBB0" bold="true" box="[136,300,1094,1118]" pageId="12" pageNumber="315">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFC4FFCB231E1720EA93FB30" collectingDate="2019-03-19" collectionCode="LKCNHM" latitude="1.4181389" longLatPrecision="1" longitude="103.93514" pageId="12" pageNumber="315" specimenCode="PU23, ZRCBDP0317875" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFC4FFCB231E1720E9A8FBB0" box="[305,403,1094,1118]" pageId="12" pageNumber="315" type="holotype">Holotype</typeStatus>
male. Pulau Ubin, Sungei [= River] Sempit,
<collectingMethod id="17FA4EF7FFC4FFCB22F31700E966FB93" box="[220,349,1126,1150]" pageId="12" pageNumber="315">Malaise trap</collectingMethod>
along creek in back mangrove,
<date id="BA051020FFC4FFCB208E1700E885FB70" pageId="12" pageNumber="315" value="2019-03-19">
<collectingDate id="AA41E9C8FFC4FFCB208E1700E885FB70" pageId="12" pageNumber="315" value="2019-03-19">19 March 2019</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFC4FFCB22E417E0E931FB70" box="[203,266,1158,1182]" collectionCode="PU" country="USA" name="Princeton University" pageId="12" pageNumber="315">PU23</specimenCode>
,
<geoCoordinate id="AB8F5027FFC4FFCB233817E2E9A7FB73" box="[279,412,1156,1181]" degrees="1" direction="north" minutes="25" orientation="latitude" pageId="12" pageNumber="315" precision="1" seconds="05.3" value="1.4181389">1°2505.3″N</geoCoordinate>
<geoCoordinate id="AB8F5027FFC4FFCB238A17E2EA7FFB73" box="[421,580,1156,1181]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="12" pageNumber="315" precision="1" seconds="06.5" value="103.93514">103°5606.5″E</geoCoordinate>
, (in 70% ethanol,
<specimenCode id="9E1D9E9BFFC4FFCB22A717C0E963FB50" box="[136,344,1190,1214]" collectionCode="ZRCBDP" pageId="12" pageNumber="315">ZRCBDP0317875</specimenCode>
_Jayanthi_JPP930_
<specimenCode id="9E1D9E9BFFC4FFCB201917C0EA4EFB50" box="[566,629,1190,1214]" collectionCode="PU" country="USA" name="Princeton University" pageId="12" pageNumber="315">PU23</specimenCode>
_
<date id="BA051020FFC4FFCB20AC17C0EB38FB50" box="[643,771,1190,1214]" pageId="12" pageNumber="315" value="2019-03-19">
<collectingDate id="AA41E9C8FFC4FFCB20AC17C0EB38FB50" box="[643,771,1190,1214]" pageId="12" pageNumber="315" value="2019-03-19">19Mar2019</collectingDate>
</date>
;
<collectionCode id="A8AAAE25FFC4FFCB22A617A0E939FB30" box="[137,258,1222,1246]" pageId="12" pageNumber="315">LKCNHM</collectionCode>
). GenBank accession code: PP893304.
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFC4FFCB22A71660E8C6FA70" blockId="12.[136,775,1286,1438]" pageId="12" pageNumber="315">
<materialsCitation id="7ED33CBDFFC4FFCB22A71660E8C1FA70" collectingDate="2018-03-29" collectionCode="LKCNHM" latitude="1.4123555" longLatPrecision="1" longitude="103.96643" pageId="12" pageNumber="315" specimenCode="PU17, ZRCBDP0288845" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFC4FFCB22A71660E8C0FAF0" bold="true" box="[136,251,1286,1310]" pageId="12" pageNumber="315">
<typeStatus id="11008842FFC4FFCB22A71660E8CDFAF0" box="[136,246,1286,1310]" pageId="12" pageNumber="315" type="paratype">Paratype</typeStatus>
.
</emphasis>
Pulau Ubin, male, Sungei [= River] Maman, creek,
<date id="BA051020FFC4FFCB22E31640E957FAD0" box="[204,364,1318,1342]" pageId="12" pageNumber="315" value="2018-03-29">
<collectingDate id="AA41E9C8FFC4FFCB22E31640E957FAD0" box="[204,364,1318,1342]" pageId="12" pageNumber="315" value="2018-03-29">29 March 2018</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFC4FFCB235B1640E988FAD0" box="[372,435,1318,1342]" collectionCode="PU" country="USA" name="Princeton University" pageId="12" pageNumber="315">PU17</specimenCode>
,
<geoCoordinate id="AB8F5027FFC4FFCB23941642EA70FAD3" box="[443,587,1316,1341]" degrees="1" direction="north" minutes="24" orientation="latitude" pageId="12" pageNumber="315" precision="1" seconds="44.48" value="1.4123555">1°2444.48″N</geoCoordinate>
<geoCoordinate id="AB8F5027FFC4FFCB207F1642EAC0FAD3" box="[592,763,1316,1341]" degrees="103" direction="east" minutes="57" orientation="longitude" pageId="12" pageNumber="315" precision="1" seconds="59.17" value="103.96643">103°5759.17″E</geoCoordinate>
), front mangrove (70% ethanol)
<specimenCode id="9E1D9E9BFFC4FFCB20301620EB3BFAB0" box="[543,768,1350,1374]" collectionCode="ZRCBDP" pageId="12" pageNumber="315">ZRCBDP0288845</specimenCode>
,
<specimenCode id="9E1D9E9BFFC4FFCB22A71600E8FFFA90" box="[136,196,1382,1406]" collectionCode="PU" country="USA" name="Princeton University" pageId="12" pageNumber="315">PU17</specimenCode>
_
<date id="BA051020FFC4FFCB22FD1600E96AFA90" box="[210,337,1382,1406]" pageId="12" pageNumber="315" value="2018-03-29">
<collectingDate id="AA41E9C8FFC4FFCB22FD1600E96AFA90" box="[210,337,1382,1406]" pageId="12" pageNumber="315" value="2018-03-29">29Mar2018</collectingDate>
</date>
<collectionCode id="A8AAAE25FFC4FFCB23741600E9EEFA90" box="[347,469,1382,1406]" pageId="12" pageNumber="315">LKCNHM</collectionCode>
). GenBank accession code: PP893305
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCB22A716A0E9E5F9F3" pageId="12" pageNumber="315" type="etymology">
<paragraph id="CE0436E0FFC4FFCB22A716A0E9E5F9F3" blockId="12.[136,776,1478,1566]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A716A0E931FA30" bold="true" box="[136,266,1478,1502]" pageId="12" pageNumber="315">Etymology.</emphasis>
The new species is named
<taxonomicName id="09BB4D63FFC4FFCB201716A1EA45FA30" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[568,638,1479,1502]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma">
<emphasis id="FCCFEAF2FFC4FFCB201716A1EA45FA30" box="[568,638,1479,1502]" italics="true" pageId="12" pageNumber="315">stigma</emphasis>
</taxonomicName>
, referring to the presence of a brown spot at the base of the costa. The name
<taxonomicName id="09BB4D63FFC4FFCB22E51561E92AF9F0" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[202,273,1543,1566]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma">
<emphasis id="FCCFEAF2FFC4FFCB22E51561E92AF9F0" box="[202,273,1543,1566]" italics="true" pageId="12" pageNumber="315">stigma</emphasis>
</taxonomicName>
is used as a noun.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCA22A71520E97FF89D" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315" type="diagnosis">
<paragraph id="CE0436E0FFC4FFCB22A71520EA2AF870" blockId="12.[135,777,1606,1950]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71520E8CDF9B0" bold="true" box="[136,246,1606,1630]" pageId="12" pageNumber="315">Diagnosis</emphasis>
. A small-sized species (about
<quantity id="09439B05FFC4FFCB20681520EA9BF9B0" box="[583,672,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="315" unit="mm" value="2.5">2.5 mm</quantity>
). Base of wings with a brownish streak at both sides of R
<subScript id="523F34A5FFC4FFCB20A01514EAACF96E" attach="left" box="[655,663,1650,1664]" fontSize="6" pageId="12" pageNumber="315">1</subScript>
. Face and palpus metallic green. Antenna yellow, postpedicel brownish on apex. Scutum without yellow colouration. Acrostichals biserial; 9 dorsocentrals, anterior 7 short, posterior 2 bristles long. Fore legs entirely black. Mid legs with trochanter and basal half of femur yellow, apical half femur, tibia and tarsus brown. Hind tibia with a black posterodorsal tooth beneath the preapical comb of bristles. Male terminalia with base of aedeagus not recurved, bent at a right angle. Tip hypandrium lacking denticles. Cercus undivided.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB22A714A0EA5CF833" blockId="12.[136,615,1990,2014]" box="[136,615,1990,2014]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A714A0E8F2F830" bold="true" box="[136,201,1990,2014]" pageId="12" pageNumber="315">Male.</emphasis>
Length: body
<quantity id="09439B05FFC4FFCB234414A0E9F9F833" box="[363,450,1990,2014]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="315" unit="mm" value="2.6">2.6 mm</quantity>
; wing
<quantity id="09439B05FFC4FFCB202314A0EA58F833" box="[524,611,1990,2014]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="315" unit="mm" value="2.1">2.1 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB22A71B60ECCBFD1B" blockId="12.[135,775,2054,2110]" lastBlockId="12.[834,1474,477,757]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71B60E8F7F7F0" bold="true" box="[136,204,2054,2078]" pageId="12" pageNumber="315">Head.</emphasis>
Face wide, with a pruinosity, but shiny metallic green with some bluish reflection at base. Frons very short (shorter
<emphasis id="FCCFEAF2FFC4FFCB216C12BBEBACFE1B" bold="true" box="[835,919,477,501]" pageId="12" pageNumber="315">Thorax</emphasis>
. Brownish black as base colour with a green metallic green shine, covered with a fine pruinosity. All bristles black. Acrostichals biserial, about 8 pairs; 9 dorsocentrals, anterior 7 bristles short, posterior 2 much longer, prescutellar bristle longest. Apical pair of scutellar bristles twice as long as bristles outside. A long white propleural bristle. Squama white with apical border brown, set with a row of pale brown bristles a little longer than squama is wide. Knob of haltere white, but with a brownish cap.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C107BECB8FC3B" blockId="12.[834,1475,797,981]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C107BEBBBFCDB" bold="true" box="[835,896,797,821]" pageId="12" pageNumber="315">Legs.</emphasis>
Fore legs entirely black. Mid legs with tip of coxa pale brown, trochanter, basal half of femur yellow; apical half femur, tibia and tarsus brown. Hind legs with coxa brown, trochanter yellow, basal half femur yellow, but apical half yellowish brown, basal half tibia pale brown, apical half brown, tarsus entirely brown.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C1098ED97FBBB" blockId="12.[834,1474,1021,1109]" pageId="12" pageNumber="315">Fore legs. Coxa with apical bristles yellow, shorter than coxa is wide; anterior bristles short, black. Femur with a posterior row of short brown bristles. No distinct bristles on tibia.</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C171BECB5FADB" blockId="12.[834,1474,1149,1333]" pageId="12" pageNumber="315">Mid legs. Coxa with whitish bristles; exterior bristle near apex coxa. Mid femur swollen on basal half with pale bristles, brown in apical half, half as long as femur is wide. Tibia with a short anterodorsal and posterodorsal bristle at basal quarter, as long as tibia is wide and a preapical ventral a little longer than tibia is wide.</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C163BEDA1F95B" blockId="12.[834,1476,1373,1717]" pageId="12" pageNumber="315">
Hind legs. Coxa near middle with a pale brown bristle, shorter than coxa is long. Femur somewhat dorsoventrally bent in basal third; with 5 long black dorsal bristles near base; 4 anterior bristles on apical third and 3 anteroventral bristles as long as femur is wide; basal half of femur with short ventral whitish bristles. Tibia with a black posterodorsal tooth-like projection composed of 4 closely set bristles, beneath the preapical comb of pale bristles; apical quarter more densely set with short ventral bristles than the basal ¾ of tibia. Basal tarsomere much shorter than second tarsomere. Length ratio tarsomeres
<quantity id="09439B05FFC4FFCB219015FBEC3EF95B" box="[959,1029,1693,1717]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="12" pageNumber="315" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">15 in</quantity>
mm: 0.16: 0.36: 0.22: 0.08: 0.95.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C15BBEDF8F8DB" blockId="12.[835,1475,1757,1845]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C15BBEB8CF91B" bold="true" box="[835,951,1757,1781]" pageId="12" pageNumber="315">Abdomen</emphasis>
. Brown in ground-colour with a faint green metallic shine. Tergites with all bristles brown, marginal bristles a little longer. Sternites with minute white bristles.
</paragraph>
<paragraph id="CE0436E0FFC4FFCA216C143BE8D0F8DD" blockId="12.[834,1475,1885,2101]" lastBlockId="13.[113,753,1723,1843]" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C143BEC2EF89B" bold="true" box="[835,1045,1885,1909]" pageId="12" pageNumber="315">Male terminalia.</emphasis>
<figureCitation id="56802A65FFC4FFCB260E143BEC41F89B" box="[1057,1146,1885,1909]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11</figureCitation>
. Entirely brown. A pair of apicoventral epandrial bristles of equal length inserted on globular papilla. Basal epandrial bristle minute (
<figureCitation id="56802A65FFC4FFCB271714FBED8DF85B" box="[1336,1462,1949,1973]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11B, D</figureCitation>
). Tip hypandrium much widened before tip, lacking denticles (
<figureCitation id="56802A65FFC4FFCB216414BBEB8AF81B" box="[843,945,2013,2037]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11G</figureCitation>
). Base of aedeagus not recurved, bent at a right angle (
<figureCitation id="56802A65FFC4FFCB21A5149BEBD0F7FB" box="[906,1003,2045,2069]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11G</figureCitation>
). Dorsal and ventral surstylus well separated. Dorsal surstylus with an apical slit, bearing a lateral (
<figureCitation id="56802A65FFC4FFCA27B81B7BE886F93D" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315">Fig. 11CE</figureCitation>
) and reflexed appendage at the outside. Postgonite pointed in lateral view (
<figureCitation id="56802A65FFC5FFCA235515BDE9DBF91D" box="[378,480,1755,1779]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11G</figureCitation>
). Cercus undivided (
<figureCitation id="56802A65FFC5FFCA20EA15BDE8F9F8FD" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11F, G</figureCitation>
), with a digitiform ventral appendage near middle (
<figureCitation id="56802A65FFC5FFCA2256147DE8E5F8DD" box="[121,222,1819,1843]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11G</figureCitation>
).
</paragraph>
<caption id="9AC46668FFC5FFCA225B1552E991F96D" ID-DOI="http://doi.org/10.5281/zenodo.14683082" ID-Zenodo-Dep="14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316" startId="13.[116,151,1588,1610]" targetBox="[113,1457,190,1575]" targetPageId="13" targetType="figure">
<paragraph id="CE0436E0FFC5FFCA225B1552E991F96D" blockId="13.[116,1454,1588,1668]" pageId="13" pageNumber="316">
Fig. 11.
<taxonomicName id="09BB4D63FFC5FFCA22E41552E94AF9A4" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[203,369,1588,1610]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">
<emphasis id="FCCFEAF2FFC5FFCA22E41552E94AF9A4" box="[203,369,1588,1610]" italics="true" pageId="13" pageNumber="316">Medetera stigma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC5FFCA23AE1553E9C1F9A4" box="[385,506,1588,1610]" pageId="13" pageNumber="316" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC5FFCA225E143DE97FF89D" blockId="13.[113,324,1883,1907]" box="[113,324,1883,1907]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA225E143DE8F0F89D" bold="true" box="[113,203,1883,1907]" pageId="13" pageNumber="316">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC5FFC9225E14FDE97DFE1B" lastPageId="14" lastPageNumber="317" pageId="13" pageNumber="316" type="discussion">
<paragraph id="CE0436E0FFC5FFCA225E14FDE8FDF7DD" blockId="13.[113,754,1947,2099]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA225E14FDE8C4F85D" bold="true" box="[113,255,1947,1971]" pageId="13" pageNumber="316">Comments.</emphasis>
<taxonomicName id="09BB4D63FFC5FFCA232314FDE9E2F85D" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[268,473,1947,1971]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">
<emphasis id="FCCFEAF2FFC5FFCA232314FDE9E2F85D" box="[268,473,1947,1971]" italics="true" pageId="13" pageNumber="316">Medetera stigma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC5FFCA23C214FAEABFF85D" box="[493,644,1947,1971]" pageId="13" pageNumber="316" rank="species">new species</taxonomicNameLabel>
, is quite characteristic in having at the base of the wings a brownish streak between the costa and R
<subScript id="523F34A5FFC5FFCA23911481E9FDF81B" attach="left" box="[446,454,2023,2037]" fontSize="6" pageId="13" pageNumber="316">1</subScript>
and below R
<subScript id="523F34A5FFC5FFCA207B1481EA67F81B" attach="left" box="[596,604,2023,2037]" fontSize="6" pageId="13" pageNumber="316">1</subScript>
. As far as we know, this characteristic is not observed in other
<taxonomicName id="09BB4D63FFC5FFCA20A3149DE8FDF7DD" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFC5FFCA20A3149DEAC9F7FD" box="[652,754,2043,2067]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="CE0436E0FFC5FFCA210215DDED97F8FD" blockId="13.[813,1452,1723,1812]" pageId="13" pageNumber="316">
<materialsCitation id="7ED33CBDFFC5FFCA210215DDED93F8FD" collectionCode="PU" pageId="13" pageNumber="316" specimenCode="PU23, PU17" specimenCount="2" typeStatus="holotype">
Only
<specimenCount id="D8BDFD69FFC5FFCA214415DAEC30F93D" box="[875,1035,1723,1747]" count="2" pageId="13" pageNumber="316" type="generic">two specimens</specimenCount>
are known: the
<typeStatus id="11008842FFC5FFCA26EE15DDED25F93D" box="[1217,1310,1723,1747]" pageId="13" pageNumber="316" type="holotype">holotype</typeStatus>
from a back mangrove near the mouth of Sungei Sempit (
<specimenCode id="9E1D9E9BFFC5FFCA273015BDED65F91D" box="[1311,1374,1755,1779]" collectionCode="PU" country="USA" name="Princeton University" pageId="13" pageNumber="316">PU23</specimenCode>
) and a
<typeStatus id="11008842FFC5FFCA2102159AEBBCF8FA" box="[813,903,1788,1812]" pageId="13" pageNumber="316" type="paratype">paratype</typeStatus>
from a front mangrove along Sungei Maman (
<specimenCode id="9E1D9E9BFFC5FFCA274F159DED9AF8FD" box="[1376,1441,1787,1811]" collectionCode="PU" country="USA" name="Princeton University" pageId="13" pageNumber="316">PU17</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFC5FFCA2102145AEB8DF85D" blockId="13.[812,1451,1851,1971]" pageId="13" pageNumber="316">
None of the species handled by
<bibRefCitation id="AA2A4B11FFC5FFCA2652145DED2BF8BD" author="Bickel D" box="[1149,1296,1851,1875]" pageId="13" pageNumber="316" pagination="195 - 259" refId="ref14423" refString="Bickel D (1987) A revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39: 195 - 259." type="journal article" year="1987">Bickel (1987)</bibRefCitation>
in his revision of the
<collectingRegion id="0C7FF802FFC5FFCA215A143DEBF7F89D" box="[885,972,1883,1907]" country="Egypt" name="Ash Sharqiyah" pageId="13" pageNumber="316">Oriental</collectingRegion>
and Australasian
<taxonomicName id="09BB4D63FFC5FFCA26BC143DECC2F89D" authorityName="Fischer von Waldheim" authorityYear="1819" box="[1171,1273,1883,1907]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC5FFCA26BC143DECC2F89D" box="[1171,1273,1883,1907]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
</taxonomicName>
and by
<bibRefCitation id="AA2A4B11FFC5FFCA277A143AEBA1F87D" author="Yang D &amp; Zhang LL &amp; Wang MQ &amp; Zhu YJ" pageId="13" pageNumber="316" refId="ref15650" refString="Yang D, Zhang LL, Wang MQ &amp; Zhu YJ (2011) Fauna Sinica Insecta, Vol. 53. Diptera Dolichopodidae. Science Press, Beijing, 1912 pp. [in Chinese, with English summary]" type="book" year="2011">Yang et al. (2011)</bibRefCitation>
in the Fauna sinica, correspond to the present new species.
</paragraph>
<paragraph id="CE0436E0FFC5FFC9210214BDE97DFE1B" blockId="13.[813,1452,2011,2099]" lastBlockId="14.[136,776,189,501]" lastPageId="14" lastPageNumber="317" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA210214BDEB96F81D" bold="true" box="[813,941,2011,2035]" pageId="13" pageNumber="316">Bionomics.</emphasis>
Adults of most
<taxonomicName id="09BB4D63FFC5FFCA264514BDED17F81D" box="[1130,1324,2011,2035]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA264514BDECE9F81D" box="[1130,1234,2011,2035]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
species
</taxonomicName>
are known to dwell on tree trunks where they hunt for small insects. The larvae of most species are predacious and live inside the burrows of bark beetles where they feed on the larvae of these beetles. Having this
<typeStatus id="11008842FFC6FFC923FF13B8E9C5FF18" box="[464,510,222,246]" pageId="14" pageNumber="317">type</typeStatus>
of biology that on first sight is independent of the saline mangrove conditions, it is remarkable that a number of these
<taxonomicName id="09BB4D63FFC6FFC9200B127BEAE6FEDB" box="[548,733,285,309]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC9200B127BEAB1FEDB" box="[548,650,285,309]" italics="true" pageId="14" pageNumber="317">Medetera</emphasis>
species
</taxonomicName>
are exclusive for back mangroves and occur in the terrestrial forests behind the mangroves. At present, four of the eleven species actually known from
<collectingCountry id="B6AC7670FFC6FFC923FC121BEA7AFE7B" box="[467,577,381,405]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
are exclusive for mangroves. Some species are dominant in mangroves and rarely occur in other biotopes, while some are exclusive for terrestrial forests.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,255 @@
<document id="C285B1F1AC3DA1A3E706203814E5A4FB" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC6FFC822DC165BED9CF9FB" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Protomedetera squamata Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="318" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="317" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A0877CEF2E2DE8408F6834D2A500DE8A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4D4F0AC168546EDD9AA612D5111743CA">
<mods:title id="6EB8F9C2E7A24AD54A1A6F3AD3C0A8A4">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="9A161F3F6328123C389DBFA18C5FC185" type="personal">
<mods:role id="4EF4F284CAB48434FC705F1F58725C91">
<mods:roleTerm id="F5BA8D30702A0335E7730FDDB9B3447C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3FE110870D27ABCD51D3A3830A5C6ECB">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="AE5AB5D03FDAD2B576A22E7BADF3DB62" type="personal">
<mods:role id="125ACDFADB137D4CACF2CC74C25D9799">
<mods:roleTerm id="93AA7B080F03F7C1B377A6CF1245AB46">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2735FC1201290483D7BB9FCA6EEE526E">Velde,</mods:namePart>
</mods:name>
<mods:name id="6770FACAE35240D6FD6ABF9A650ACA91" type="personal">
<mods:role id="6FC367DBFAD24284ED4D42C6753E26C1">
<mods:roleTerm id="B33B19935237A314DEE9F377965911FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5FC8EA70122A50FE2E04991CC632372">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="B852ED0B0F226024AF95A7B70D1E6CA5">text</mods:typeOfResource>
<mods:relatedItem id="F1649324F7996940549BEDAB7F9535AE" type="host">
<mods:titleInfo id="6ECC57D2BE16BBEA84FB829561C46FFF">
<mods:title id="EDA7E41A53BC427B7B63EC2D848B4459">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="C8E6676AC786BAF5F6B601D25B7244F4">
<mods:date id="47E145C45C87DEA3709CCD0012E038FC">2024</mods:date>
<mods:detail id="AF8845452A8DA2B77B0B32BDFCE22AE6" type="pubDate">
<mods:number id="A7A22ED55EA7209424DF93EAE9935F36">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="6D00CBFC8A7D347F60FCAC90ADC81DC9" type="volume">
<mods:number id="D89DAA1FB29129DDAEA01C8A2F98C3B3">72</mods:number>
</mods:detail>
<mods:extent id="7AF211A8F4B9DC39ACC62E3DCDCE683B" unit="page">
<mods:start id="EA182DDAB1708CC1D5D1A22EDC7D7BA1">303</mods:start>
<mods:end id="E87478571BE94BED5BDE809E3E232A99">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="39001993F5DA3F7D016F3497D626FDB5">journal article</mods:classification>
<mods:identifier id="CD6E15B705823D0AD5577DFEAE9D2028" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="79C1CBC0856429CFEC091A98309AF5A5" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="801A82D066A1506E8BC46017BB772B31" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="16C37FF5C00965C5C43763A03CE33B34" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFC6FFC822DC165BED9CF9FB" LSID="urn:lsid:plazi:treatment:461287F6FFC6FFC822DC165BED9CF9FB" httpUri="http://treatment.plazi.org/id/461287F6FFC6FFC822DC165BED9CF9FB" lastPageId="15" lastPageNumber="318" pageId="14" pageNumber="317">
<subSubSection id="86A1656BFFC6FFC922DC165BEAA6FABB" box="[243,669,1341,1365]" pageId="14" pageNumber="317" type="nomenclature">
<paragraph id="CE0436E0FFC6FFC922DC165BEAA6FABB" blockId="14.[243,669,1341,1397]" box="[243,669,1341,1365]" pageId="14" pageNumber="317">
<heading id="954C818CFFC6FFC922DC165BEAA6FABB" bold="true" box="[243,669,1341,1365]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="317" reason="2">
<emphasis id="FCCFEAF2FFC6FFC922DC165BEAA6FABB" bold="true" box="[243,669,1341,1365]" pageId="14" pageNumber="317">
<taxonomicName id="09BB4D63FFC6FFC922DC165BEA33FABB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[243,520,1341,1365]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="squamata" status="sp. nov.">
<emphasis id="FCCFEAF2FFC6FFC922DC165BEA33FABB" bold="true" box="[243,520,1341,1365]" italics="true" pageId="14" pageNumber="317">Protomedetera squamata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC6FFC92038165BEAA6FABB" box="[535,669,1341,1365]" pageId="14" pageNumber="317" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC6FFC92350163BEA2BFA9B" box="[383,528,1373,1397]" pageId="14" pageNumber="317" type="description">
<paragraph id="CE0436E0FFC6FFC92350163BEA2BFA9B" blockId="14.[243,669,1341,1397]" box="[383,528,1373,1397]" pageId="14" pageNumber="317">
(
<figureCitation id="56802A65FFC6FFC923A9163BEA33FA9B" box="[390,520,1373,1397]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="14.[837,874,756,778]" captionStartId-1="14.[836,871,1613,1635]" captionStartId-2="15.[113,147,1040,1062]" captionTargetBox-0="[837,1473,189,741]" captionTargetBox-1="[838,1475,867,1597]" captionTargetBox-2="[119,758,191,1023]" captionTargetId-0="figure-428@14.[837,1473,189,741]" captionTargetId-1="figure-444@14.[823,1494,859,1608]" captionTargetId-2="figure-16@15.[90,766,169,1033]" captionTargetPageId-0="14" captionTargetPageId-1="14" captionTargetPageId-2="15" captionText-0="Fig. 12. Protomedetera squamata, new species, habitus male (ZRCBDP0084004). Photograph by Kristy Chang." captionText-1="Fig. 13.Protomedetera squamata, new species, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." captionText-2="Fig.14.Protomedetera squamata, new species, female ovipositor.A, Dorsal; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi-0="http://doi.org/10.5281/zenodo.14683084" figureDoi-1="http://doi.org/10.5281/zenodo.14683086" figureDoi-2="http://doi.org/10.5281/zenodo.14683088" httpUri-0="https://zenodo.org/record/14683084/files/figure.png" httpUri-1="https://zenodo.org/record/14683086/files/figure.png" httpUri-2="https://zenodo.org/record/14683088/files/figure.png" pageId="14" pageNumber="317">Figs. 1214</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC6FFC922A716FBEA8AF91B" pageId="14" pageNumber="317" type="materials_examined">
<paragraph id="CE0436E0FFC6FFC922A716FBE931F9FB" blockId="14.[134,775,1437,1557]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC922A716FBE902FA5B" bold="true" box="[136,313,1437,1461]" pageId="14" pageNumber="317">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFC6FFC9236A16FBE931F9FB" collectingDate="2016-04-07" collectionCode="MT, LKCNHM" country="Singapore" latitude="1.4201111" location="Pulau Ubin" longLatPrecision="1" longitude="103.9351" pageId="14" pageNumber="317" specimenCode="PU01" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFC6FFC9236A16FBE997FA5B" box="[325,428,1437,1461]" pageId="14" pageNumber="317" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFC6FFC923D216FBEA4BFA5B" box="[509,624,1437,1461]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
,
<location id="CB64603BFFC6FFC920AF16FBEB39FA5B" LSID="urn:lsid:plazi:treatment:461287F6FFC6FFC822DC165BED9CF9FB:CB64603BFFC6FFC920AF16FBEB39FA5B" box="[640,770,1437,1461]" country="Singapore" latitude="1.4201111" longLatPrecision="1" longitude="103.9351" name="Pulau Ubin" pageId="14" pageNumber="317">Pulau Ubin</location>
,
<date id="BA051020FFC6FFC922A716D8E925FA3B" box="[136,286,1469,1493]" pageId="14" pageNumber="317" value="2016-04-07">
<collectingDate id="AA41E9C8FFC6FFC922A716D8E925FA3B" box="[136,286,1469,1493]" pageId="14" pageNumber="317" value="2016-04-07">7 April 2016</collectingDate>
</date>
,
<collectionCode id="A8AAAE25FFC6FFC9230216D8E963FA3B" box="[301,344,1470,1493]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="14" pageNumber="317">MT</collectionCode>
,
<specimenCode id="9E1D9E9BFFC6FFC9234716DBE99EFA3B" box="[360,421,1469,1493]" collectionCode="PU" country="USA" name="Princeton University" pageId="14" pageNumber="317">PU01</specimenCode>
(
<geoCoordinate id="AB8F5027FFC6FFC9239716DAEA6AFA3B" box="[440,593,1468,1493]" degrees="1" direction="north" minutes="25" orientation="latitude" pageId="14" pageNumber="317" precision="1" seconds="12.40" value="1.4201111">1°2512.40″N</geoCoordinate>
<geoCoordinate id="AB8F5027FFC6FFC9207316DAEAC4FA3B" box="[604,767,1468,1493]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="14" pageNumber="317" precision="1" seconds="6.35" value="103.9351">103°566.35″E</geoCoordinate>
) terminalia dissected and illustrated, no mitogenome (Ma9210,
<collectionCode id="A8AAAE25FFC6FFC922A9169BE8C4F9FB" box="[134,255,1533,1557]" pageId="14" pageNumber="317">LKCNHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFC6FFC922A7155BEA8AF91B" blockId="14.[136,775,1597,1781]" pageId="14" pageNumber="317">
<materialsCitation id="7ED33CBDFFC6FFC922A7155BE98EF97B" collectingDate="2016-04-07" collectionCode="MT" country="Singapore" location="Pulau Ubin" pageId="14" pageNumber="317" specimenCode="PU01, ZRCBDP0084002" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFC6FFC922A7155BE938F9BB" bold="true" box="[136,259,1597,1621]" pageId="14" pageNumber="317">
<typeStatus id="11008842FFC6FFC922A7155BE8C5F9BB" box="[136,254,1597,1621]" pageId="14" pageNumber="317" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFC6FFC92322155BE99CF9BB" box="[269,423,1597,1621]" name="Singapore" pageId="14" pageNumber="317">SINGAPORE</collectingCountry>
,
<specimenCount id="D8BDFD69FFC6FFC9239A155BEA22F9BB" box="[437,537,1597,1621]" count="1" pageId="14" pageNumber="317" type="female">1 female</specimenCount>
,
<location id="CB64603BFFC6FFC92008155BEA93F9BB" LSID="urn:lsid:plazi:treatment:461287F6FFC6FFC822DC165BED9CF9FB:CB64603BFFC6FFC92008155BEA93F9BB" box="[551,680,1597,1621]" country="Singapore" name="Pulau Ubin" pageId="14" pageNumber="317">Pulau Ubin</location>
,
<date id="BA051020FFC6FFC920991558E8FAF99B" pageId="14" pageNumber="317" value="2016-04-07">
<collectingDate id="AA41E9C8FFC6FFC920991558E8FAF99B" pageId="14" pageNumber="317" value="2016-04-07">7 April 2016</collectingDate>
</date>
,
<collectionCode id="A8AAAE25FFC6FFC922FF1538E8C0F99B" box="[208,251,1630,1653]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="14" pageNumber="317">MT</collectionCode>
,
<specimenCode id="9E1D9E9BFFC6FFC92326153BE97DF99B" box="[265,326,1629,1653]" collectionCode="PU" country="USA" name="Princeton University" pageId="14" pageNumber="317">PU01</specimenCode>
(Ma9211)
<specimenCode id="9E1D9E9BFFC6FFC923E8153BEAA9F99B" box="[455,658,1629,1653]" collectionCode="ZRCBDP" pageId="14" pageNumber="317">ZRCBDP0084002</specimenCode>
; GenBank accession code: PP893307
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFC6FFC923EB151BE8C1F93B" collectingDate="2018-05-17" collectionCode="ZRCBDP, PU" country="Singapore" pageId="14" pageNumber="317" specimenCode="ZRCBDP0255641, PU19" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFC6FFC923EB151BEA1DF97B" box="[452,550,1661,1685]" count="1" pageId="14" pageNumber="317" type="female">1 female</specimenCount>
<specimenCode id="9E1D9E9BFFC6FFC9201F151BEAC2F97B" box="[560,761,1661,1685]" collectionCode="ZRCBDP" pageId="14" pageNumber="317">ZRCBDP0255641</specimenCode>
_
<specimenCode id="9E1D9E9BFFC6FFC922A715FBE8FDF95B" box="[136,198,1693,1717]" collectionCode="PU" country="USA" name="Princeton University" pageId="14" pageNumber="317">PU19</specimenCode>
_
<date id="BA051020FFC6FFC922FB15FBE96DF95B" box="[212,342,1693,1717]" pageId="14" pageNumber="317" value="2018-05-17">
<collectingDate id="AA41E9C8FFC6FFC922FB15FBE96DF95B" box="[212,342,1693,1717]" pageId="14" pageNumber="317" value="2018-05-17">17May2018</collectingDate>
</date>
_20190204; GenBank accession code: PP893308
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFC6FFC9232915DBEA96F91B" collectingDate="2018-03-29" collectionCode="LKCNHM" country="Singapore" pageId="14" pageNumber="317" specimenCode="ZRCBDP0285063, PU19" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFC6FFC9232915DBE95DF93B" box="[262,358,1725,1749]" count="1" pageId="14" pageNumber="317" type="female">1 female</specimenCount>
<specimenCode id="9E1D9E9BFFC6FFC9234115DBEA0DF93B" box="[366,566,1725,1749]" collectionCode="ZRCBDP" pageId="14" pageNumber="317">ZRCBDP0285063</specimenCode>
_
<specimenCode id="9E1D9E9BFFC6FFC9206C15DBEA46F93B" box="[579,637,1725,1749]" collectionCode="PU" country="USA" name="Princeton University" pageId="14" pageNumber="317">PU19</specimenCode>
_
<date id="BA051020FFC6FFC920A315DBEB3CF93B" box="[652,775,1725,1749]" pageId="14" pageNumber="317" value="2018-03-29">
<collectingDate id="AA41E9C8FFC6FFC920A315DBEB3CF93B" box="[652,775,1725,1749]" pageId="14" pageNumber="317" value="2018-03-29">29Mar2018</collectingDate>
</date>
(
<collectionCode id="A8AAAE25FFC6FFC922A015BBE931F91B" box="[143,266,1757,1781]" pageId="14" pageNumber="317">LKCNHM</collectionCode>
): GenBank accession code: PP893309
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="9AC46668FFC6FFC9216B152BEDF8F991" ID-DOI="http://doi.org/10.5281/zenodo.14683086" ID-Zenodo-Dep="14683086" httpUri="https://zenodo.org/record/14683086/files/figure.png" pageId="14" pageNumber="317" startId="14.[836,871,1613,1635]" targetBox="[838,1475,867,1597]" targetPageId="14" targetType="figure">
<paragraph id="CE0436E0FFC6FFC9216B152BEDF8F991" blockId="14.[835,1475,1613,1664]" pageId="14" pageNumber="317">
Fig. 13.
<taxonomicName id="09BB4D63FFC6FFC921BE152BECBFF98D" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[913,1156,1613,1635]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="squamata" status="sp. nov.">
<emphasis id="FCCFEAF2FFC6FFC921BE152BECBFF98D" box="[913,1156,1613,1635]" italics="true" pageId="14" pageNumber="317">Protomedetera squamata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC6FFC926A0152BED3FF98D" box="[1167,1284,1613,1635]" pageId="14" pageNumber="317" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<subSubSection id="86A1656BFFC6FFC922A7147BEADCF8BB" pageId="14" pageNumber="317" type="etymology">
<paragraph id="CE0436E0FFC6FFC922A7147BEADCF8BB" blockId="14.[136,774,1821,1877]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC922A7147BE92CF8DB" bold="true" box="[136,279,1821,1845]" pageId="14" pageNumber="317">Etymology.</emphasis>
The name
<taxonomicName id="09BB4D63FFC6FFC923871478EA21F8DB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[424,538,1822,1845]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="squamata">
<emphasis id="FCCFEAF2FFC6FFC923871478EA21F8DB" box="[424,538,1822,1845]" italics="true" pageId="14" pageNumber="317">squamata</emphasis>
</taxonomicName>
refers to the large squamiform bristles on the apex of the dorsal surstylus.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC6FFC822A7141BEC29FA5B" lastPageId="15" lastPageNumber="318" pageId="14" pageNumber="317" type="diagnosis">
<paragraph id="CE0436E0FFC6FFC922A7141BED86F901" blockId="14.[135,776,1917,2101]" lastBlockId="14.[835,1474,1719,1775]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC922A7141BE8C6F87B" bold="true" box="[136,253,1917,1941]" pageId="14" pageNumber="317">Diagnosis.</emphasis>
Small species with shiny metallic green head and thorax. Antenna yellow. Pedicel with 3 long marginal bristles covering the spheroid postpedicel. Biserial acrostichals and 4 long yellowish brown dorsocentrals. All coxae and femora blackish brown. Tibiae and tarsi yellow, except for a brown dorsal streak on fore tibia and apical tarsomere of all legs brown (almost black on fore tarsus). Surstylus with 2 wide, pale apicoventral squamiform bristles. Cercus small, oval.
</paragraph>
<paragraph id="CE0436E0FFC6FFC9216C1471EBC3F8A0" blockId="14.[835,1474,1815,1871]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC9216C1471EBBFF8C1" bold="true" box="[835,900,1815,1839]" pageId="14" pageNumber="317">Male.</emphasis>
<figureCitation id="56802A65FFC6FFC921A51471EBE2F8C1" box="[906,985,1815,1839]" captionStart="Fig" captionStartId="14.[837,874,756,778]" captionTargetBox="[837,1473,189,741]" captionTargetId="figure-428@14.[837,1473,189,741]" captionTargetPageId="14" captionText="Fig. 12. Protomedetera squamata, new species, habitus male (ZRCBDP0084004). Photograph by Kristy Chang." figureDoi="http://doi.org/10.5281/zenodo.14683084" httpUri="https://zenodo.org/record/14683084/files/figure.png" pageId="14" pageNumber="317">Fig. 12</figureCitation>
. Length: body
<quantity id="09439B05FFC6FFC926571471ECE1F8C0" box="[1144,1242,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="1.74" pageId="14" pageNumber="317" unit="mm" value="1.74">1.74 mm</quantity>
; wing
<quantity id="09439B05FFC6FFC9270E1471EDB9F8C0" box="[1313,1410,1815,1839]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="317" unit="mm" value="1.5">1.50 mm</quantity>
, wing width:
<quantity id="09439B05FFC6FFC921BF1451EBCFF8A0" box="[912,1012,1847,1871]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="14" pageNumber="317" unit="mm" value="0.47">0.47 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFC6FFC8216C1411EBE3FEDB" blockId="14.[834,1476,1911,2095]" lastBlockId="15.[813,1452,189,309]" lastPageId="15" lastPageNumber="318" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC9216C1411EBBCF861" bold="true" box="[835,903,1911,1935]" pageId="14" pageNumber="317">Head.</emphasis>
With frons and face very wide, shiny metallic green. Face triangular, wide below antenna, narrowing down to clypeus, there a little wider than pedicel. Hairs and bristles on head yellowish brown. A pair of strong retroclinate ocellar bristles, a pair of strong but shorter proclinate vertical bristles and a pair of short postvertical bristles. Postocular Fore legs. Coxa densely covered with short pale bristles, at least 2 very long apical bristles. Fore femur with short pale ventrals and 2 distinct posterior preapical bristles. Tibia without apicals.
</paragraph>
<paragraph id="CE0436E0FFC7FFC82102123BEC21FDFB" blockId="15.[812,1453,349,533]" pageId="15" pageNumber="318">Mid legs. Coxa with short anterior bristles and a long pale brownish exterior bristle as long as coxa is long. Femur without ventrals, a tiny anterior preapical and posteroventral preapical. Tibia with a short dorsal near base, a longer at basal third. Ventral apical bristle long (nearly 2.5 times as long as tibia is wide).</paragraph>
<paragraph id="CE0436E0FFC7FFC82102115BED90FD5B" blockId="15.[811,1453,573,693]" pageId="15" pageNumber="318">Hind legs. Coxa with a short brownish exterior bristle near middle. Femur in apical half with a row of pale anteroventral bristles about half as long as femur is wide. Tibia with a long anterodorsal on basal third and a few distinct apical bristles.</paragraph>
<caption id="9AC46668FFC7FFC8225E1776EACBFBAC" ID-DOI="http://doi.org/10.5281/zenodo.14683088" ID-Zenodo-Dep="14683088" httpUri="https://zenodo.org/record/14683088/files/figure.png" pageId="15" pageNumber="318" startId="15.[113,147,1040,1062]" targetBox="[119,758,191,1023]" targetPageId="15" targetType="figure">
<paragraph id="CE0436E0FFC7FFC8225E1776EACBFBAC" blockId="15.[112,753,1040,1090]" pageId="15" pageNumber="318">
Fig. 14.
<taxonomicName id="09BB4D63FFC7FFC822941776E993FBCB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[187,424,1040,1061]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="318" phylum="Arthropoda" rank="species" species="squamata" status="sp. nov.">
<emphasis id="FCCFEAF2FFC7FFC822941776E993FBCB" box="[187,424,1040,1061]" italics="true" pageId="15" pageNumber="318">Protomedetera squamata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC7FFC8239C1776EA1FFBC8" box="[435,548,1040,1062]" pageId="15" pageNumber="318" rank="species">new species</taxonomicNameLabel>
, female ovipositor. A, Dorsal; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC7FFC8225E171CE954FB7C" blockId="15.[113,367,1146,1170]" box="[113,367,1146,1170]" pageId="15" pageNumber="318">bristles short, uniserial.</paragraph>
<paragraph id="CE0436E0FFC7FFC8225E17DFE970F9DC" blockId="15.[112,754,1209,1586]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8225E17DFE8DDFB3F" bold="true" box="[113,230,1209,1233]" pageId="15" pageNumber="318">Antennae.</emphasis>
Scape yellow, short, with an interior projection. Pedicel yellow, cup-like, large, with postpedicel sunken in it; in the apical crown of bristles 3 long inner apical bristles longer than and curved over postpedicel; below the apical row, a second row of short bristles, sometimes with some additional scattered bristles. Postpedicel dusky yellowish, spheroid, with short pubescence and a subapical (shifted toward outside) long arista. All bristles on antenna yellowish brown. Arista black, naked, basal portion of arista short, like a short tubercle on the postpedicel. Palpus brown, small, circular, with a long yellowish brown subapical bristle and a few short bristles.
</paragraph>
<paragraph id="CE0436E0FFC7FFC8225E153CE916F89C" blockId="15.[112,755,1626,1906]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8225E153CE8F6F99C" bold="true" box="[113,205,1626,1650]" pageId="15" pageNumber="318">Thorax.</emphasis>
Shiny metallic green (pollinosity probably absent, not visible in ethanol conserved specimen). Pleura more brownish green. Apical half of mesonotum flat before scutellum. Hairs and bristles on thorax yellowish (dusky). Anterior dorsocentrals short, followed by 4 long dorsocentrals becoming longer toward scutellum, acrostichals present only on basal half of mesonotum, biserial very short, consisting of 8 pairs. A pair of very long crossing scutellars, lacking bristle at outside.
</paragraph>
<paragraph id="CE0436E0FFC7FFC8225E14FCEAF4F7FC" blockId="15.[112,752,1946,2066]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8225E14FCE895F85C" bold="true" box="[113,174,1946,1970]" pageId="15" pageNumber="318">Legs.</emphasis>
Mainly brown but trochanters, tip of all femora, all tibiae and tarsomeres yellow, except for the dorsal fore tibia, which is darkened and tarsomere 5 of all legs brownish (darkest on fore tarsus). Hairs and bristles on legs pale.
</paragraph>
<paragraph id="CE0436E0FFC7FFC8210211BBED97FCFB" blockId="15.[813,1452,733,789]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8210211BBEB4DFD1B" bold="true" box="[813,886,733,757]" pageId="15" pageNumber="318">Wings.</emphasis>
Nearly hyaline, tinged light yellow; veins yellowish brown. Squama pale with a short pale bristle. Haltere white.
</paragraph>
<paragraph id="CE0436E0FFC7FFC82102105BEC54FC9B" blockId="15.[813,1451,829,885]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC82102105BEB98FCBB" bold="true" box="[813,931,829,853]" pageId="15" pageNumber="318">Abdomen.</emphasis>
Dark brown. Hairs and bristles on tergites pale, only marginal bristles longer.
</paragraph>
<paragraph id="CE0436E0FFC7FFC8210210FBEBF1FAFB" blockId="15.[811,1454,925,1301]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8210210FBEBD7FC5B" bold="true" box="[813,1004,925,949]" pageId="15" pageNumber="318">Male terminalia.</emphasis>
<figureCitation id="56802A65FFC7FFC821DB10FBEC7FFC5B" box="[1012,1092,925,949]" captionStart="Fig" captionStartId="14.[836,871,1613,1635]" captionTargetBox="[838,1475,867,1597]" captionTargetId="figure-444@14.[823,1494,859,1608]" captionTargetPageId="14" captionText="Fig. 13.Protomedetera squamata, new species, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683086" httpUri="https://zenodo.org/record/14683086/files/figure.png" pageId="15" pageNumber="318">Fig. 13</figureCitation>
. Foramen basolateral (
<figureCitation id="56802A65FFC7FFC8271310FBEDA4FC5B" box="[1340,1439,925,949]" captionStart="Fig" captionStartId="14.[836,871,1613,1635]" captionTargetBox="[838,1475,867,1597]" captionTargetId="figure-444@14.[823,1494,859,1608]" captionTargetPageId="14" captionText="Fig. 13.Protomedetera squamata, new species, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683086" httpUri="https://zenodo.org/record/14683086/files/figure.png" pageId="15" pageNumber="318">Fig. 13B</figureCitation>
). Cercus pale, very short, oval, not pointed, with a subapical bristle hardly longer than the other bristles. Surstylus large, subrectangular, at the inner side with a rim bearing a strong bristle near middle (
<figureCitation id="56802A65FFC7FFC8263C177BECA4FBDB" box="[1043,1183,1053,1077]" captionStart="Fig" captionStartId="14.[836,871,1613,1635]" captionTargetBox="[838,1475,867,1597]" captionTargetId="figure-444@14.[823,1494,859,1608]" captionTargetPageId="14" captionText="Fig. 13.Protomedetera squamata, new species, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683086" httpUri="https://zenodo.org/record/14683086/files/figure.png" pageId="15" pageNumber="318">Fig. 13A, B</figureCitation>
). Apicoventrally with 2 very large, wide pale squamiform bristles. Tip of surstylus with 3 smaller bristles and 2 longer bristles in a more lateral position (subapical). Apicoventral epandrial lobe lacking, but 3 epandrial bristles present, apical and basal longest. Basoventral epandrial lobe and bristles lacking. Hypandrium short (
<figureCitation id="56802A65FFC7FFC8215F17BBEBEEFB1B" box="[880,981,1245,1269]" captionStart="Fig" captionStartId="14.[836,871,1613,1635]" captionTargetBox="[838,1475,867,1597]" captionTargetId="figure-444@14.[823,1494,859,1608]" captionTargetPageId="14" captionText="Fig. 13.Protomedetera squamata, new species, male terminalia. A, Ventral; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683086" httpUri="https://zenodo.org/record/14683086/files/figure.png" pageId="15" pageNumber="318">Fig. 13A</figureCitation>
). A large vesical structure present at the tip of the phallus.
</paragraph>
<paragraph id="CE0436E0FFC7FFC82102165BEC29FA5B" blockId="15.[813,1453,1341,1461]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC82102165BEBB8FABB" bold="true" box="[813,899,1341,1365]" pageId="15" pageNumber="318">Female</emphasis>
. Length: body
<quantity id="09439B05FFC7FFC82615165BECADFABB" box="[1082,1174,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="318" unit="mm" value="1.5">1.5 mm</quantity>
; wing
<quantity id="09439B05FFC7FFC826C6165BED7EFABB" box="[1257,1349,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="318" unit="mm" value="1.4">1.4 mm</quantity>
. In most respects identical to male. A minute hair is present at both sides of the scutellars that is absent in male. Ovipositor illustrated in
<figureCitation id="56802A65FFC7FFC8219216FBEC35FA5B" box="[957,1038,1437,1461]" captionStart="Fig" captionStartId="15.[113,147,1040,1062]" captionTargetBox="[119,758,191,1023]" captionTargetId="figure-16@15.[90,766,169,1033]" captionTargetPageId="15" captionText="Fig.14.Protomedetera squamata, new species, female ovipositor.A, Dorsal; B, Lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683088" httpUri="https://zenodo.org/record/14683088/files/figure.png" pageId="15" pageNumber="318">Fig. 14</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC7FFC8210216BBED9CF9FB" pageId="15" pageNumber="318" type="discussion">
<paragraph id="CE0436E0FFC7FFC8210216BBED9CF9FB" blockId="15.[813,1452,1501,1558]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8210216BBEB94FA1B" bold="true" box="[813,943,1501,1525]" pageId="15" pageNumber="318">Comments.</emphasis>
The presence of large squamiform bristles on the dorsal surstylus is unique in the genus
<taxonomicName id="09BB4D63FFC7FFC8272D169BED9BF9FB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1282,1440,1533,1557]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC7FFC8272D169BED9BF9FB" box="[1282,1440,1533,1557]" italics="true" pageId="15" pageNumber="318">Protomedetera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,127 @@
<document id="424FD41250647510862C0EA092AC9B08" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC6FFC922B8117BEAB2FAFB" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Protomedetera Tang, Yang &amp; Grootaert 2018" docType="treatment" docVersion="2" lastPageNumber="317" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="317" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="C9E8F54F761787A98F3C62D4D541F880" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14DA90CF597E0F7DF50F1B3BA68DA72E">
<mods:title id="7F1BAF11C65F1501873BF72640EB726E">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="B7F13B6C2C8A8421963819A66242FFA7" type="personal">
<mods:role id="8F5F6BD48007BA32A3C4EDD9A273A77E">
<mods:roleTerm id="DD64568178FADEE136E75B5198814224">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D712D7D159A3069C3090132A9752B0DD">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="45C9C7D9D0B6FEEBCFB42B6463F47821" type="personal">
<mods:role id="941E2D653FC09D28C70E9074CCAEFA0E">
<mods:roleTerm id="CB3C527B4664288121FB2F1EF5506518">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6ABF5C87F489004F39151AFE827A3734">Velde,</mods:namePart>
</mods:name>
<mods:name id="7C6150D26EDCF009A26B606E8ADF49E4" type="personal">
<mods:role id="63F83BE4223DBA73DC97A29C5AACB7DB">
<mods:roleTerm id="B39388F7DBE645DB6F0C20F4A2A19130">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BAC2749B3D7E7B685D42414AA1863F46">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="9F4DEC250D535BDE15BF465C3400ADE7">text</mods:typeOfResource>
<mods:relatedItem id="8CED19A0D473045979116C947BF7CB3E" type="host">
<mods:titleInfo id="F0C4524615685DE1302C4126FE16B4A3">
<mods:title id="C4B11BC127BBFADB4CDAA9F5A909F26A">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="A5B282B46D9A35B41A20BED3378B7929">
<mods:date id="CFF80B542A34FC5115C8470FC369C327">2024</mods:date>
<mods:detail id="225E1B0EB8A7D413476E68692D0438E4" type="pubDate">
<mods:number id="FA5ED87E449D8A928CCFD6C930004BE9">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="10CC253A6EF5B746FE309E6D9B261B61" type="volume">
<mods:number id="7ED9832D7A02AACF410F745526A33616">72</mods:number>
</mods:detail>
<mods:extent id="BA2D6D8DB253A8907A8E5332D13011D8" unit="page">
<mods:start id="A626EBA8CA034FC396310D1481D01C73">303</mods:start>
<mods:end id="CF7AEA0CC605E958C0EE6496E923DEB0">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="86DA34CCEDEDE81F4BAD326142888609">journal article</mods:classification>
<mods:identifier id="D647C3DEE8274923A39C161EA2864E0B" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="3C3668E8CDA4AA40B2133446F538AE92" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="AEC9D3DA8964DD44C0E48C73ACB7E855" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="A8A05242496A762DF33F860664B7856E" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFC6FFC922B8117BEAB2FAFB" ID-DOI="http://doi.org/10.5281/zenodo.14683102" ID-Zenodo-Dep="14683102" LSID="urn:lsid:plazi:treatment:461287F6FFC6FFC922B8117BEAB2FAFB" httpUri="http://treatment.plazi.org/id/461287F6FFC6FFC922B8117BEAB2FAFB" lastPageNumber="317" pageId="14" pageNumber="317">
<subSubSection id="86A1656BFFC6FFC922B8117BEAC3FDDB" box="[151,760,541,565]" pageId="14" pageNumber="317" type="nomenclature">
<paragraph id="CE0436E0FFC6FFC922B8117BEAC3FDDB" blockId="14.[151,760,541,565]" box="[151,760,541,565]" pageId="14" pageNumber="317">
<heading id="954C818CFFC6FFC922B8117BEAC3FDDB" bold="true" box="[151,760,541,565]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="317" reason="2">
<emphasis id="FCCFEAF2FFC6FFC922B8117BEAC3FDDB" bold="true" box="[151,760,541,565]" pageId="14" pageNumber="317">
Genus
<taxonomicName id="09BB4D63FFC6FFC922C7117BEAC3FDDB" ID-CoL="89R97" authority="Tang, Yang &amp; Grootaert, 2018" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[232,760,541,565]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC6FFC922C7117BE9B2FDDB" bold="true" box="[232,393,541,565]" italics="true" pageId="14" pageNumber="317">Protomedetera</emphasis>
<bibRefCitation id="AA2A4B11FFC6FFC923BD117BEAC3FDDB" author="Tang C &amp; Grootaert P &amp; Yang D" box="[402,760,541,565]" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang, Yang &amp; Grootaert, 2018</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC6FFC922A7113BEAB2FAFB" pageId="14" pageNumber="317" type="type_taxon">
<paragraph id="CE0436E0FFC6FFC922A7113BE9CEFD7B" blockId="14.[136,774,605,662]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC922A7113BE91EFD9B" bold="true" box="[136,293,605,629]" pageId="14" pageNumber="317">
<typeStatus id="11008842FFC6FFC922A7113BE8F8FD9B" box="[136,195,605,629]" pageId="14" pageNumber="317">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="09BB4D63FFC6FFC92300113BE938FD7B" ID-CoL="8DHKB" authority="Grootaert &amp; Tang, 2018" authorityName="Grootaert &amp; Tang" authorityYear="2018" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="FCCFEAF2FFC6FFC92300113BEA4EFD9B" box="[303,629,605,629]" italics="true" pageId="14" pageNumber="317">Protomedetera singaporensis</emphasis>
Grootaert &amp;
<bibRefCitation id="AA2A4B11FFC6FFC922A71118E938FD7B" author="Tang C &amp; Grootaert P &amp; Yang D" box="[136,259,637,662]" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang, 2018</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="AA2A4B11FFC6FFC9231C1118E9D5FD7B" author="Tang C &amp; Grootaert P &amp; Yang D" box="[307,494,637,662]" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
)
</paragraph>
<paragraph id="CE0436E0FFC6FFC922A711DBEAB2FAFB" blockId="14.[134,776,701,1302]" pageId="14" pageNumber="317">
<taxonomicName id="09BB4D63FFC6FFC922A711DBE91EFD3B" ID-CoL="89R97" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[136,293,701,725]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC6FFC922A711DBE91EFD3B" box="[136,293,701,725]" italics="true" pageId="14" pageNumber="317">Protomedetera</emphasis>
</taxonomicName>
comprise small species with a body length less than or at most
<quantity id="09439B05FFC6FFC9231E11BBE948FD1B" box="[305,371,733,757]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="317" unit="mm" value="2.0">2 mm</quantity>
. The genus is quite rare in
<collectingCountry id="B6AC7670FFC6FFC920B611BBEB3EFD1B" box="[665,773,733,757]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
and up until now, only two species were known from
<collectingCountry id="B6AC7670FFC6FFC922A7107BE8C2FCDB" box="[136,249,797,821]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
. The
<typeStatus id="11008842FFC6FFC923151078E953FCD8" box="[314,360,798,822]" pageId="14" pageNumber="317">type</typeStatus>
species of the genus,
<taxonomicName id="09BB4D63FFC6FFC92049107BEA19FCBB" ID-CoL="8DHKB" authority="Grootaert &amp; Tang, 2018" authorityName="Grootaert &amp; Tang" authorityYear="2018" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="singaporensis">
<emphasis id="FCCFEAF2FFC6FFC92049107BE927FCBB" italics="true" pageId="14" pageNumber="317">Protomedetera singaporensis</emphasis>
Grootaert &amp;
<bibRefCitation id="AA2A4B11FFC6FFC923861058EA19FCBB" author="Tang C &amp; Grootaert P &amp; Yang D" box="[425,546,829,854]" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang, 2018</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="AA2A4B11FFC6FFC920631058EB3BFCBB" author="Tang C &amp; Grootaert P &amp; Yang D" box="[588,768,829,854]" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
) was described from a single male found in the swamp forest at Nee Soon (
<collectingCountry id="B6AC7670FFC6FFC92333101BE9B5FC7B" box="[284,398,893,917]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
). A second species
<taxonomicName id="09BB4D63FFC6FFC9204C1018E93CFC5B" ID-CoL="8DHK9" authority="Tang et al., 2018" authorityName="Tang" authorityYear="2018" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFC6FFC9204C1018EAF2FC7B" box="[611,713,893,917]" italics="true" pageId="14" pageNumber="317">P. glabra</emphasis>
<bibRefCitation id="AA2A4B11FFC6FFC920FF1018E93CFC5B" author="Tang C &amp; Grootaert P &amp; Yang D" pageId="14" pageNumber="317" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
</taxonomicName>
, found only in anthropogenic habitats, is more common in
<collectingCountry id="B6AC7670FFC6FFC9233F10DBE9BAFC3B" box="[272,385,957,981]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
. This species was attributed to the genus
<taxonomicName id="09BB4D63FFC6FFC922FC10BBE943FC1B" ID-CoL="89R97" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[211,376,989,1013]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC6FFC922FC10BBE943FC1B" box="[211,376,989,1013]" italics="true" pageId="14" pageNumber="317">Protomedetera</emphasis>
</taxonomicName>
, but it has a different morphology and should probably be attributed to another genus. The postpedicel does not bear long apical bristles and the position of the male terminalia on the abdomen is different.
<taxonomicName id="09BB4D63FFC6FFC9208D1758EB3DFBBB" ID-CoL="8DHK9" box="[674,774,1085,1109]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="317" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFC6FFC9208D1758EB3DFBBB" box="[674,774,1085,1109]" italics="true" pageId="14" pageNumber="317">P. glabra</emphasis>
</taxonomicName>
was originally described from
<collectingCountry id="B6AC7670FFC6FFC923F61738EA96FB9B" box="[473,685,1117,1141]" name="Papua New Guinea" pageId="14" pageNumber="317">Papua New Guinea</collectingCountry>
and the specimens found in
<collectingCountry id="B6AC7670FFC6FFC92370171BE9F0FB7B" box="[351,459,1149,1173]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
were considered conspecific. Although the male terminalia of the specimens from
<collectingCountry id="B6AC7670FFC6FFC920E917F8E92BFB3B" name="Papua New Guinea" pageId="14" pageNumber="317">Papua New Guinea</collectingCountry>
and those from
<collectingCountry id="B6AC7670FFC6FFC9239017DBEA10FB3B" box="[447,555,1213,1237]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
are similar, they are distinctly different in other characteristics and hence the taxonomy of this species group needs revision.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,227 @@
<document id="55069D0BF83AD4EEA6B583B7A3E241E6" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC7FFD721B7155BEC13FAD5" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Protomedetera ubinensis Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="319" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="318" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6FA1E1105F7E9FA4A2FE66840BAFF991" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="58115FFCA312CA1CCD4696904FD4E1CA">
<mods:title id="A7C670294F9DA6507508C1F8995E5F0C">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="E1C65B78F51541F511AC3973754A1E22" type="personal">
<mods:role id="6B9E09F0DDC0D7AAA38BCDAAA7A12FD5">
<mods:roleTerm id="DAFFEFBCC342551520D0C2D586B67BAF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D3E7AF71123DB493E1E1D52C9713249">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="9D4F8C231555B705362C7C0E58F4C3CC" type="personal">
<mods:role id="D3F65A3338A7AE96AEBAAFEFC2F9039A">
<mods:roleTerm id="9D4D8180FA30AB757C8D41875FBF8E5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="45D3E313F20DB5FDD35820D11B6A3BD6">Velde,</mods:namePart>
</mods:name>
<mods:name id="80996822911CF1AE61C7CCC91825623E" type="personal">
<mods:role id="5F5DB3ACAC79C8EFF5005AFB0B64F504">
<mods:roleTerm id="F48B4A7A24E1BF4F8262E97D4E211540">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7E84C017235A719AE14B2FE914EF7E34">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="DAA0056475AC23541232163E2A3A5499">text</mods:typeOfResource>
<mods:relatedItem id="85C40BEA8E96D970AB5C72C2D20E43A0" type="host">
<mods:titleInfo id="9F918F148ECB6A52E6EF525B34417EE7">
<mods:title id="3C75FADB85161B6CD94878E6C6DF34C7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D22915ACC5425D0789B2197FB81F38B1">
<mods:date id="B1604980CE9BF51A02FA20E244573194">2024</mods:date>
<mods:detail id="4EC5266FBC2A7221F477CE3039B2646C" type="pubDate">
<mods:number id="9473FF3BA1F30F6B2784C0CFA88C4783">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="94904E9B8BEF4C52E084143FE4C96703" type="volume">
<mods:number id="BCBAEA420C7758433DE69D068992C7D0">72</mods:number>
</mods:detail>
<mods:extent id="196AE6E9695A1DA2CACDDC16262BE1E8" unit="page">
<mods:start id="4F8076D57FE325D3C649C84B97F47FA4">303</mods:start>
<mods:end id="B18FFAAC6973C90B13481FECDAB44BA3">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3DB0F28AC3841CAE60AE8E4945BA9B46">journal article</mods:classification>
<mods:identifier id="5FE67B9C3F35DB51DC3893DAC1870FE8" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="0C5D8CCDC5B211AFAE16C9827F00FD59" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="240B333A1017B4288E6AC9D71E562032" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="1D87A5770AA660970111C596012F1313" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFC7FFD721B7155BEC13FAD5" ID-DOI="http://doi.org/10.5281/zenodo.14683104" ID-Zenodo-Dep="14683104" LSID="urn:lsid:plazi:treatment:461287F6FFC7FFD721B7155BEC13FAD5" httpUri="http://treatment.plazi.org/id/461287F6FFC7FFD721B7155BEC13FAD5" lastPageId="16" lastPageNumber="319" pageId="15" pageNumber="318">
<subSubSection id="86A1656BFFC7FFC821B7155BED7BF9BB" box="[920,1344,1597,1621]" pageId="15" pageNumber="318" type="nomenclature">
<paragraph id="CE0436E0FFC7FFC821B7155BED7BF9BB" blockId="15.[920,1344,1597,1653]" box="[920,1344,1597,1621]" pageId="15" pageNumber="318">
<heading id="954C818CFFC7FFC821B7155BED7BF9BB" bold="true" box="[920,1344,1597,1621]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="318" reason="2">
<emphasis id="FCCFEAF2FFC7FFC821B7155BED7BF9BB" bold="true" box="[920,1344,1597,1621]" pageId="15" pageNumber="318">
<taxonomicName id="09BB4D63FFC7FFC821B7155BEC90F9BB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[920,1195,1597,1621]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="318" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFC7FFC821B7155BEC90F9BB" bold="true" box="[920,1195,1597,1621]" italics="true" pageId="15" pageNumber="318">Protomedetera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC7FFC82694155BED7BF9BB" box="[1211,1344,1597,1621]" pageId="15" pageNumber="318" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC7FFC82613153BECA0F99B" box="[1084,1179,1629,1653]" pageId="15" pageNumber="318" type="description">
<paragraph id="CE0436E0FFC7FFC82613153BECA0F99B" blockId="15.[920,1344,1597,1653]" box="[1084,1179,1629,1653]" pageId="15" pageNumber="318">
(
<figureCitation id="56802A65FFC7FFC8266B153BECA8F99B" box="[1092,1171,1629,1653]" captionStart="Fig" captionStartId="16.[136,171,858,880]" captionTargetBox="[136,777,189,843]" captionTargetId="figure-8@16.[136,777,189,843]" captionTargetPageId="16" captionText="Fig. 15. Protomedetera ubinensis, new species, holotype female (ZRCBDP0085237). Photograph by Rene Ong." figureDoi="http://doi.org/10.5281/zenodo.14683092" httpUri="https://zenodo.org/record/14683092/files/figure.png" pageId="15" pageNumber="318">Fig. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC7FFC8210215FBEC23F81B" pageId="15" pageNumber="318" type="diagnosis">
<paragraph id="CE0436E0FFC7FFC8210215FBEC23F81B" blockId="15.[811,1454,1693,2037]" pageId="15" pageNumber="318">
<emphasis id="FCCFEAF2FFC7FFC8210215FBEB98F95B" bold="true" box="[813,931,1693,1717]" pageId="15" pageNumber="318">Diagnosis.</emphasis>
Small species with shiny metallic green head and thorax. Antenna yellowish brown. Pedicel with 3 long inner marginal bristles bent over the darkened spheroid postpedicel. Palpus brown with a single brown bristle twice as long as palpus. Fore tibia much shorter than fore femur with a long pale ventral preapical bristle. Acr quadriserial, close to but distinctly separated from dorsocentrals. Anterior dorsocentrals very short, biserial, only the 2 prescutellar bristles long. Basal tarsomere of the hind leg that is contrastingly dark in comparison with the tibia and the following tarsomeres.
</paragraph>
</subSubSection>
<caption id="9AC46668FFD8FFD722A7103CEA6CFC62" ID-DOI="http://doi.org/10.5281/zenodo.14683092" ID-Zenodo-Dep="14683092" httpUri="https://zenodo.org/record/14683092/files/figure.png" pageId="16" pageNumber="319" startId="16.[136,171,858,880]" targetBox="[136,777,189,843]" targetPageId="16" targetType="figure">
<paragraph id="CE0436E0FFD8FFD722A7103CEA6CFC62" blockId="16.[136,777,858,909]" pageId="16" pageNumber="319">
Fig. 15.
<taxonomicName id="09BB4D63FFD8FFD722F0103CE9EFFC81" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[223,468,858,879]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFD8FFD722F0103CE9EFFC81" box="[223,468,858,879]" italics="true" pageId="16" pageNumber="319">Protomedetera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD723CC103DEA66FC9E" box="[483,605,858,880]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, holotype female (ZRCBDP0085237). Photograph by Rene Ong.
</paragraph>
</caption>
<subSubSection id="86A1656BFFD8FFD722A710A3E951FC12" pageId="16" pageNumber="319" type="etymology">
<paragraph id="CE0436E0FFD8FFD722A710A3E951FC12" blockId="16.[136,775,964,1020]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD722A710A3E931FC33" bold="true" box="[136,266,965,989]" pageId="16" pageNumber="319">Etymology.</emphasis>
The name of the new species refers to the
<typeStatus id="11008842FFD8FFD720F610A3EB3CFC33" box="[729,775,965,989]" pageId="16" pageNumber="319">type</typeStatus>
locality, Pulau Ubin.
</paragraph>
</subSubSection>
<caption id="9AC46668FFD8FFD72169177FED08FBA2" ID-DOI="http://doi.org/10.5281/zenodo.14683094" ID-Zenodo-Dep="14683094" httpUri="https://zenodo.org/record/14683094/files/figure.png" pageId="16" pageNumber="319" startId="16.[838,875,1049,1071]" targetBox="[836,1472,189,1034]" targetPageId="16" targetType="figure">
<paragraph id="CE0436E0FFD8FFD72169177FED08FBA2" blockId="16.[836,1472,1049,1100]" pageId="16" pageNumber="319">
Fig. 16.
<taxonomicName id="09BB4D63FFD8FFD7218E177FECA6FBC1" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[929,1181,1049,1071]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD8FFD7218E177FECA6FBC1" box="[929,1181,1049,1071]" italics="true" pageId="16" pageNumber="319">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD72681177FED16FBC1" box="[1198,1325,1049,1071]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, male habitus (ZRCBDP0085280). Photograph by Kristy Chang.
</paragraph>
</caption>
<subSubSection id="86A1656BFFD8FFD722A71743E979F892" pageId="16" pageNumber="319" type="materials_examined">
<paragraph id="CE0436E0FFD8FFD722A71743EA99FB53" blockId="16.[134,776,1060,1213]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD722A71743E90BFBD2" bold="true" box="[136,304,1060,1085]" pageId="16" pageNumber="319">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFD8FFD723181742EA99FB53" collectingDate="2016-04-07" collectionCode="MT, LKCNHM" country="Singapore" latitude="1.4198194" location="Pulau Ubin" longLatPrecision="1" longitude="103.94705" pageId="16" pageNumber="319" specimenCode="PU04, ZRCBDP0085237" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFD8FFD723181742E9A1FBD2" box="[311,410,1060,1084]" pageId="16" pageNumber="319" type="holotype">Holotype</typeStatus>
female.
<collectingCountry id="B6AC7670FFD8FFD723D71742EA5CFBD2" box="[504,615,1060,1084]" name="Singapore" pageId="16" pageNumber="319">Singapore</collectingCountry>
,
<location id="CB64603BFFD8FFD7205D1742EAD5FBD2" LSID="urn:lsid:plazi:treatment:461287F6FFC7FFD721B7155BEC13FAD5:CB64603BFFD8FFD7205D1742EAD5FBD2" box="[626,750,1060,1084]" country="Singapore" latitude="1.4198194" longLatPrecision="1" longitude="103.94705" name="Pulau Ubin" pageId="16" pageNumber="319">Pulau Ubin</location>
,
<date id="BA051020FFD8FFD720D61743E8C7FBB2" pageId="16" pageNumber="319" value="2016-04-07">
<collectingDate id="AA41E9C8FFD8FFD720D61743E8C7FBB2" pageId="16" pageNumber="319" value="2016-04-07">7 April 2016</collectingDate>
</date>
,
<collectionCode id="A8AAAE25FFD8FFD723291723E90BFBB2" box="[262,304,1093,1116]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="16" pageNumber="319">MT</collectionCode>
,
<specimenCode id="9E1D9E9BFFD8FFD723161723E94EFBB3" box="[313,373,1093,1117]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU04</specimenCode>
(
<geoCoordinate id="AB8F5027FFD8FFD723AD1725EA2FFBB2" box="[386,532,1091,1116]" degrees="1" direction="north" minutes="25" orientation="latitude" pageId="16" pageNumber="319" precision="1" seconds="11.35" value="1.4198194">1°2511.35″N</geoCoordinate>
<geoCoordinate id="AB8F5027FFD8FFD720361725EAFEFBB2" box="[537,709,1091,1116]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="16" pageNumber="319" precision="1" seconds="49.39" value="103.94705">103°5649.39″E</geoCoordinate>
), back mangrove (
<specimenCode id="9E1D9E9BFFD8FFD722D31702E984FB92" box="[252,447,1124,1148]" collectionCode="ZRCBDP" pageId="16" pageNumber="319">ZRCBDP0085237</specimenCode>
_
<specimenCode id="9E1D9E9BFFD8FFD723E31702EA32FB92" box="[460,521,1124,1148]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU04</specimenCode>
_
<date id="BA051020FFD8FFD7203A1702EAB7FB92" box="[533,652,1124,1148]" pageId="16" pageNumber="319" value="2016-04-07">
<collectingDate id="AA41E9C8FFD8FFD7203A1702EAB7FB92" box="[533,652,1124,1148]" pageId="16" pageNumber="319" value="2016-04-07">07Apr2016</collectingDate>
</date>
_20170417, extracted for barcode and mitogenome by Darren Yeo;
<collectionCode id="A8AAAE25FFD8FFD722A617C3E8C5FB53" box="[137,254,1189,1213]" pageId="16" pageNumber="319">LKCNHM</collectionCode>
) GenBank accession code: PP893310.
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFD8FFD722A71783EA93FAB3" blockId="16.[136,776,1252,1373]" pageId="16" pageNumber="319">
<materialsCitation id="7ED33CBDFFD8FFD722A71783EA9FFAB3" collectingDate="2018-03-29" collectionCode="MT, LKCNHM" country="Singapore" latitude="1.4162333" location="Pulau Ubin" longLatPrecision="1" longitude="103.94616" pageId="16" pageNumber="319" specimenCode="PU19, ZRCBDP0285052" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFD8FFD722A71783E8C2FB13" bold="true" box="[136,249,1253,1277]" pageId="16" pageNumber="319">
<typeStatus id="11008842FFD8FFD722A71783E8CFFB13" box="[136,244,1253,1277]" pageId="16" pageNumber="319" type="paratype">Paratype</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFD8FFD7232B1783E99AFB13" box="[260,417,1253,1277]" name="Singapore" pageId="16" pageNumber="319">SINGAPORE</collectingCountry>
: female,
<location id="CB64603BFFD8FFD720251782EAB7FB12" LSID="urn:lsid:plazi:treatment:461287F6FFC7FFD721B7155BEC13FAD5:CB64603BFFD8FFD720251782EAB7FB12" box="[522,652,1252,1276]" country="Singapore" latitude="1.4162333" longLatPrecision="1" longitude="103.94616" name="Pulau Ubin" pageId="16" pageNumber="319">Pulau Ubin</location>
,
<date id="BA051020FFD8FFD720B51783E8F8FAF3" pageId="16" pageNumber="319" value="2018-03-29">
<collectingDate id="AA41E9C8FFD8FFD720B51783E8F8FAF3" pageId="16" pageNumber="319" value="2018-03-29">29 March 2018</collectingDate>
</date>
,
<collectionCode id="A8AAAE25FFD8FFD722FC1663E8C4FAF2" box="[211,255,1285,1308]" country="Russia" name="Mus. Tinro, Vladyvostok" pageId="16" pageNumber="319">MT</collectionCode>
,
<specimenCode id="9E1D9E9BFFD8FFD723201663E976FAF3" box="[271,333,1285,1309]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU19</specimenCode>
(
<geoCoordinate id="AB8F5027FFD8FFD7234E1665E9C5FAF2" box="[353,510,1283,1308]" degrees="1" direction="north" minutes="24" orientation="latitude" pageId="16" pageNumber="319" precision="1" seconds="58.44" value="1.4162333">1°2458.44″N</geoCoordinate>
<geoCoordinate id="AB8F5027FFD8FFD720261665EA84FAF2" box="[521,703,1283,1308]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="16" pageNumber="319" precision="1" seconds="46.18" value="103.94616">103°5646.18″E</geoCoordinate>
) back mangrove (
<specimenCode id="9E1D9E9BFFD8FFD7232A1642E9EBFAD2" box="[261,464,1316,1340]" collectionCode="ZRCBDP" pageId="16" pageNumber="319">ZRCBDP0285052</specimenCode>
_
<specimenCode id="9E1D9E9BFFD8FFD723F11642EA22FAD2" box="[478,537,1316,1340]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU19</specimenCode>
_
<date id="BA051020FFD8FFD720081642EA98FAD2" box="[551,675,1316,1340]" pageId="16" pageNumber="319" value="2018-03-29">
<collectingDate id="AA41E9C8FFD8FFD720081642EA98FAD2" box="[551,675,1316,1340]" pageId="16" pageNumber="319" value="2018-03-29">29Mar2018</collectingDate>
</date>
; imaged;
<collectionCode id="A8AAAE25FFD8FFD722A71623E93AFAB3" box="[136,257,1349,1373]" pageId="16" pageNumber="319">LKCNHM</collectionCode>
). GenBank accession code: PP893311
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFD8FFD722A716E3E909F8D3" blockId="16.[135,777,1412,1853]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD722A716E3E8D9FA73" bold="true" box="[136,226,1413,1437]" pageId="16" pageNumber="319">Female.</emphasis>
Length: body
<quantity id="09439B05FFD8FFD7235616E3E983FA72" box="[377,440,1413,1437]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="319" unit="mm" value="2.0">2 mm</quantity>
; wing
<quantity id="09439B05FFD8FFD723D316E2EA66FA72" box="[508,605,1412,1436]" metricMagnitude="-3" metricUnit="m" metricValue="1.63" pageId="16" pageNumber="319" unit="mm" value="1.63">1.63 mm</quantity>
. Habitus similar to
<taxonomicName id="09BB4D63FFD8FFD7228D16C3E922FA52" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[162,281,1445,1468]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata" status="new species">
<emphasis id="FCCFEAF2FFD8FFD7228D16C3E922FA52" box="[162,281,1445,1468]" italics="true" pageId="16" pageNumber="319">P. uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD7230A16C3E99EFA52" box="[293,421,1444,1468]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, in having quadriserial acrostichal bristles and only 2 long dorsocentral bristles. Antenna dusky yellow with postpedicel brown. Palpus brown with a single brown bristle twice as long as palpus and a few short hairs. Legs with coxae and femora black. Tibia and tarsi yellow. Only the apical tarsomere of all tarsi dusky (not brown) and tarsomere 1 of hind leg contrastingly darkened. Fore tibia much shorter than fore femur with a long pale ventral preapical bristle. Ratio of fore femur, tibia and tarsomeres: 0.43: 0.33: 0.14: 0.06: 0.048: 0.032: 0.06 (in mm). Hind leg with tibia longer than femur. Ratio of hind femur, tibia and tarsomeres: 0.51: 0.55: 0.12: 0.17: 0.13: 0.06: 0.07 (in mm).
</paragraph>
<paragraph id="CE0436E0FFD8FFD722A71403E979F892" blockId="16.[136,322,1892,1917]" box="[136,322,1892,1917]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD722A71403E8F2F893" bold="true" box="[136,201,1893,1917]" pageId="16" pageNumber="319">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD722A714C2EC13FAD5" pageId="16" pageNumber="319" type="discussion">
<paragraph id="CE0436E0FFD8FFD722A714C2EC13FAD5" blockId="16.[136,775,1956,2076]" lastBlockId="16.[834,1475,1155,1339]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD722A714C2E92AF852" bold="true" box="[136,273,1956,1980]" pageId="16" pageNumber="319">Comments.</emphasis>
<taxonomicName id="09BB4D63FFD8FFD7233214C2EA7BF852" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[285,576,1956,1980]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFD8FFD7233214C2EA7BF852" box="[285,576,1956,1980]" italics="true" pageId="16" pageNumber="319">Protomedetera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD7207B14C3EADFF852" box="[596,740,1956,1980]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, is described from
<specimenCount id="D8BDFD69FFD8FFD7231614A3E98EF832" box="[313,437,1988,2012]" count="2" pageId="16" pageNumber="319" type="female">two female</specimenCount>
specimens only because their identical barcodes differ more than 10% from all other known
<taxonomicName id="09BB4D63FFD8FFD722F61B62E9EEF7F2" box="[217,469,2052,2076]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFD8FFD722F61B62E942F7F2" box="[217,377,2052,2076]" italics="true" pageId="16" pageNumber="319">Protomedetera</emphasis>
species.
</taxonomicName>
In addition, both specimens have the contrastingly darkened hind tarsomere 1, a character not present in other
<taxonomicName id="09BB4D63FFD8FFD7260C17C5ECFAFB55" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1059,1217,1187,1211]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFD8FFD7260C17C5ECFAFB55" box="[1059,1217,1187,1211]" italics="true" pageId="16" pageNumber="319">Protomedetera</emphasis>
</taxonomicName>
. The new species most resembles
<taxonomicName id="09BB4D63FFD8FFD7219817A2EC0FFB35" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[951,1076,1220,1243]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata" status="new species">
<emphasis id="FCCFEAF2FFD8FFD7219817A2EC0FFB35" box="[951,1076,1220,1243]" italics="true" pageId="16" pageNumber="319">P. uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD7266C17A5ECF2FB35" box="[1091,1225,1219,1243]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, in having quadriserial acrostichal bristles and 2 long dorsocentrals, while
<taxonomicName id="09BB4D63FFD8FFD727851782EB90FAF5" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="squamata" status="new species">
<emphasis id="FCCFEAF2FFD8FFD727851782EB90FAF5" italics="true" pageId="16" pageNumber="319">P. squamata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD721951665EC05FAF5" box="[954,1086,1283,1307]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
, has biserial acrostichal bristles and 4 long dorsocentrals.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,492 @@
<document id="51F23C24320743CAB19BBDC6CBA03634" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFCAFFC3231F151BECD3FD3B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chrysotus dot Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="307" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="305" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8B3CB0C96C3942BEF976EBA0CE07581E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9F9284A9A06961DDE637037BB2EB1CB4">
<mods:title id="7F1882515B6A256A0F53899AE9A1698B">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="7EA071ADA7FC87C8468A836156EA3957" type="personal">
<mods:role id="EFE86DE1E6621ECFA10CA0B86E2195EA">
<mods:roleTerm id="93D3BB4D998BEA73931240BF4D36DFDA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4256AEA0B39E5A952C49B103E8EBBCE1">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="133794598D02420F26CF0880CC1E720C" type="personal">
<mods:role id="E9DB6E203A3A5D50459A0A2ACA06233F">
<mods:roleTerm id="85ADC9CD6637CB1072788CCA48AA6DB9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FF7EC112FF455FE2711006276071789">Velde,</mods:namePart>
</mods:name>
<mods:name id="2F1D0702CB78B667EFA1CB71CFA51A25" type="personal">
<mods:role id="1EA13B1DCF93033F413C64B5E189229A">
<mods:roleTerm id="AA60AF6E2AF9620370DE405806CABE03">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="12E723E3B491A55E53A2E1A8433ECD39">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="8FDCFF192C0DF296F3CFA4AB02F986F1">text</mods:typeOfResource>
<mods:relatedItem id="3CC8FF14492BB615D1AA1D52A99543B9" type="host">
<mods:titleInfo id="3BD462AD293827A05A293C0460888B7F">
<mods:title id="F13997BA03B25F159900C1CE80EF60A0">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="096F382A901682B6AEADF0714AA290AD">
<mods:date id="736A669FF045E85A3E63CD15576CE74C">2024</mods:date>
<mods:detail id="976A6943440F1B19DB16BE625EBF7BFC" type="pubDate">
<mods:number id="DA30B7DB43C8973975EFC93D6B4AF79D">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="EB8C905B60EAB684F3FB3174992E9AF6" type="volume">
<mods:number id="76B38B7FF544B7385BD7CE13D0DA03A8">72</mods:number>
</mods:detail>
<mods:extent id="89C09925FA28004B7FA3F327B7B02943" unit="page">
<mods:start id="714642709A8FC5AAA2956D2B3D07D069">303</mods:start>
<mods:end id="D3EE0DEFB43DF7B25CD2AC74BF2DBE3B">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="917268590547F90BEB8DE93FC1734A79">journal article</mods:classification>
<mods:identifier id="477E82875495EE932E7FBB4891788B06" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="663C2BCA5D2C9D23CF275A02072637CD" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="6D9CCDCAA575B13D718C7CE3C0299230" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="439B5579B4C0FCF159549D4FD4CF0F73" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFCAFFC3231F151BECD3FD3B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B" httpUri="http://treatment.plazi.org/id/461287F6FFCAFFC3231F151BECD3FD3B" lastPageId="4" lastPageNumber="307" pageId="2" pageNumber="305">
<subSubSection id="86A1656BFFCAFFC5231F151BEA64F97B" box="[304,607,1661,1685]" pageId="2" pageNumber="305" type="nomenclature">
<paragraph id="CE0436E0FFCAFFC5231F151BEA64F97B" blockId="2.[304,607,1661,1717]" box="[304,607,1661,1685]" pageId="2" pageNumber="305">
<heading id="954C818CFFCAFFC5231F151BEA64F97B" bold="true" box="[304,607,1661,1685]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="305" reason="2">
<emphasis id="FCCFEAF2FFCAFFC5231F151BEA64F97B" bold="true" box="[304,607,1661,1685]" pageId="2" pageNumber="305">
<taxonomicName id="09BB4D63FFCAFFC5231F151BE9F1F97B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[304,458,1661,1685]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCAFFC5231F151BE9F1F97B" bold="true" box="[304,458,1661,1685]" italics="true" pageId="2" pageNumber="305">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCAFFC523F5151BEA64F97B" box="[474,607,1661,1685]" pageId="2" pageNumber="305" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC523A415FBEA3FF95B" box="[395,516,1693,1717]" pageId="2" pageNumber="305" type="description">
<paragraph id="CE0436E0FFCAFFC523A415FBEA3FF95B" blockId="2.[304,607,1661,1717]" box="[395,516,1693,1717]" pageId="2" pageNumber="305">
(
<figureCitation id="56802A65FFCAFFC523BC15FBE9DBF95B" box="[403,480,1693,1717]" captionStart="Fig" captionStartId="2.[835,873,843,865]" captionTargetBox="[835,1474,189,828]" captionTargetId="figure-671@2.[835,1474,189,828]" captionTargetPageId="2" captionText="Fig. 2. Chrysotus dot, new species, habitus paratype male (ZRCBDP0083923). Photograph by Miteshravin S/O Ravindran." figureDoi="http://doi.org/10.5281/zenodo.14683060" httpUri="https://zenodo.org/record/14683060/files/figure.png" pageId="2" pageNumber="305">Figs. 2</figureCitation>
,
<figureCitation id="56802A65FFCAFFC523C115FBE9C0F95B" box="[494,507,1693,1717]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="2" pageNumber="305">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC522A715BBEC41FAE3" pageId="2" pageNumber="305" type="materials_examined">
<paragraph id="CE0436E0FFCAFFC522A715BBE97BF89B" blockId="2.[135,777,1757,1909]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC522A715BBE902F91B" bold="true" box="[136,313,1757,1781]" pageId="2" pageNumber="305">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFCAFFC5236A15BBE97BF89B" collectingDate="2019-03-19" collectionCode="OBS, LKCNHM" collectorName="P. Grootaert &amp; GenBank" country="Singapore" county="Tock Kew" location="Pulau Ubin" municipality="Sungei" pageId="2" pageNumber="305" specimenCode="PU22, ZRCBDP0317760" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFCAFFC5236A15BBE997F91B" box="[325,428,1757,1781]" pageId="2" pageNumber="305" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFCAFFC523D215BBEA4BF91B" box="[509,624,1757,1781]" name="Singapore" pageId="2" pageNumber="305">Singapore</collectingCountry>
,
<location id="CB64603BFFCAFFC520AF15BBEB39F91B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCAFFC520AF15BBEB39F91B" box="[640,770,1757,1781]" country="Singapore" county="Tock Kew" municipality="Sungei" name="Pulau Ubin" pageId="2" pageNumber="305">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFCAFFC522A7159BE8E9F8FB" box="[136,210,1789,1813]" pageId="2" pageNumber="305">Sungei</collectingMunicipality>
[= River]
<collectingCounty id="27654E6CFFCAFFC52367159BE981F8FB" box="[328,442,1789,1813]" pageId="2" pageNumber="305">Tock Kew</collectingCounty>
basin (
<collectionCode id="A8AAAE25FFCAFFC52023159BEA7FF8FB" box="[524,580,1789,1813]" pageId="2" pageNumber="305">OBS</collectionCode>
),
<date id="BA051020FFCAFFC52076159BEB38F8FB" box="[601,771,1789,1813]" pageId="2" pageNumber="305" value="2019-03-19">
<collectingDate id="AA41E9C8FFCAFFC52076159BEB38F8FB" box="[601,771,1789,1813]" pageId="2" pageNumber="305" value="2019-03-19">19 March 2019</collectingDate>
</date>
,
<collectingMethod id="17FA4EF7FFCAFFC522A5147BE936F8DB" box="[138,269,1821,1845]" pageId="2" pageNumber="305">Malaise trap</collectingMethod>
(
<specimenCode id="9E1D9E9BFFCAFFC52334147BE961F8DB" box="[283,346,1821,1845]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
) in back mangroves (
<specimenCode id="9E1D9E9BFFCAFFC52014147BEB3EF8DB" box="[571,773,1821,1845]" collectionCode="ZRCBDP" pageId="2" pageNumber="305">ZRCBDP0317760</specimenCode>
, leg.
<collectorName id="634E5336FFCAFFC5229B1458E90DF8BB" box="[180,310,1853,1877]" pageId="2" pageNumber="305">P. Grootaert</collectorName>
; in ethanol,
<collectionCode id="A8AAAE25FFCAFFC5239D145BEA11F8BB" box="[434,554,1853,1877]" pageId="2" pageNumber="305">LKCNHM</collectionCode>
).
<collectorName id="634E5336FFCAFFC52016145BEAA6F8BB" box="[569,669,1853,1877]" pageId="2" pageNumber="305">GenBank</collectorName>
accession code: PP893289.
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFCAFFC522A714FBED47FB63" blockId="2.[136,775,1949,2101]" lastBlockId="2.[835,1475,949,1165]" pageId="2" pageNumber="305">
<materialsCitation id="7ED33CBDFFCAFFC522A714FBE929F83B" collectionCode="OBS" country="Singapore" location="Pulau Ubin" municipality="Sungei Tock Kew" pageId="2" pageNumber="305" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFCAFFC522A714FBE932F85B" bold="true" box="[136,265,1949,1973]" pageId="2" pageNumber="305">
<typeStatus id="11008842FFCAFFC522A714FBE93FF85B" box="[136,260,1949,1973]" pageId="2" pageNumber="305" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFCAFFC5233A14FBE9B7F85B" box="[277,396,1949,1973]" name="Singapore" pageId="2" pageNumber="305">Singapore</collectingCountry>
,
<location id="CB64603BFFCAFFC523B414FBEA18F85B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCAFFC523B414FBEA18F85B" box="[411,547,1949,1973]" country="Singapore" municipality="Sungei Tock Kew" name="Pulau Ubin" pageId="2" pageNumber="305">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFCAFFC5201C14FBEB3DF85B" box="[563,774,1949,1973]" pageId="2" pageNumber="305">Sungei Tock Kew</collectingMunicipality>
basin (
<collectionCode id="A8AAAE25FFCAFFC522FD14DBE931F83B" box="[210,266,1981,2005]" pageId="2" pageNumber="305">OBS</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="7ED33CBDFFCAFFC5230E14DBE93FF81B" collectingDate="2019-03-19" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="31" specimenCount-female="22" specimenCount-male="9" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5230E14DBE942F83B" box="[289,377,1981,2005]" count="9" pageId="2" pageNumber="305" type="male">9 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC523A814DBEA39F83B" box="[391,514,1981,2005]" count="22" pageId="2" pageNumber="305" type="female">22 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5203F14DBEA74F83B" box="[528,591,1981,2005]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5207214DBEB3DF83B" box="[605,774,1981,2005]" pageId="2" pageNumber="305" value="2019-03-19">
<collectingDate id="AA41E9C8FFCAFFC5207214DBEB3DF83B" box="[605,774,1981,2005]" pageId="2" pageNumber="305" value="2019-03-19">19 March 2019</collectingDate>
</date>
(Ma10640)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5233F14BBEB38F81B" box="[272,771,2013,2037]" collectingDate="2019-03-26" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5233F14BBE940F81B" box="[272,379,2013,2037]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC523A814BBE9FDF81B" box="[391,454,2013,2037]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC523FD14BBEA43F81B" box="[466,632,2013,2037]" pageId="2" pageNumber="305" value="2019-03-26">
<collectingDate id="AA41E9C8FFCAFFC523FD14BBEA43F81B" box="[466,632,2013,2037]" pageId="2" pageNumber="305" value="2019-03-26">26 March 2019</collectingDate>
</date>
(MA10955)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC522A7149BEAD1F7FB" box="[136,746,2045,2069]" collectingDate="2019-04-02" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="15" specimenCount-female="9" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC522A7149BE8DEF7FB" box="[136,229,2045,2069]" count="6" pageId="2" pageNumber="305" type="male">6 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC522DB149BE95CF7FB" box="[244,359,2045,2069]" count="9" pageId="2" pageNumber="305" type="female">9 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC52359149BE983F7FB" box="[374,440,2045,2069]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC523E8149BEA66F7FB" box="[455,605,2045,2069]" pageId="2" pageNumber="305" value="2019-04-02">
<collectingDate id="AA41E9C8FFCAFFC523E8149BEA66F7FB" box="[455,605,2045,2069]" pageId="2" pageNumber="305" value="2019-04-02">2 April 2019</collectingDate>
</date>
(Ma10718)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC520D6149BEAD0F7DB" collectingDate="2019-04-16" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="8" specimenCount-female="2" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC520D6149BE8F6F7DB" count="6" pageId="2" pageNumber="305" type="male">6 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC522F21B7BE96BF7DB" box="[221,336,2077,2101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC523701B7BE99AF7DB" box="[351,417,2077,2101]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5239E1B7BEA6DF7DB" box="[433,598,2077,2101]" pageId="2" pageNumber="305" value="2019-04-16">
<collectingDate id="AA41E9C8FFCAFFC5239E1B7BEA6DF7DB" box="[433,598,2077,2101]" pageId="2" pageNumber="305" value="2019-04-16">16 April 2019</collectingDate>
</date>
(MA10986)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC520D51B7BED28FC23" box="[762,1299,949,2101]" collectingDate="2019-04-24" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC520D51B7BEBA1FC23" box="[762,922,949,2101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5218A10D3EBDEFC23" box="[933,997,949,973]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC521DF10D3ECBCFC23" box="[1008,1159,949,973]" pageId="2" pageNumber="305" value="2019-04-24">
<collectingDate id="AA41E9C8FFCAFFC521DF10D3ECBCFC23" box="[1008,1159,949,973]" pageId="2" pageNumber="305" value="2019-04-24">24 April 2019</collectingDate>
</date>
(MA10972)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5273110D3EC57FC03" collectingDate="2019-05-15" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5273110D3ED4FFC23" box="[1310,1396,949,973]" count="3" pageId="2" pageNumber="305" type="male">3 males</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5275010D3ED84FC23" box="[1407,1471,949,973]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5216C10B3EBE0FC03" box="[835,987,981,1006]" pageId="2" pageNumber="305" value="2019-05-15">
<collectingDate id="AA41E9C8FFCAFFC5216C10B3EBE0FC03" box="[835,987,981,1006]" pageId="2" pageNumber="305" value="2019-05-15">15 May 2019</collectingDate>
</date>
(MA11506)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5265510B3EC7FFBE3" collectingDate="2019-05-28" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5265510B3ECF1FC03" box="[1146,1226,981,1005]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526F610B3ED72FC03" box="[1241,1353,981,1005]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5277810B3EDA3FC03" box="[1367,1432,981,1005]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5278810B3EB8EFBE3" pageId="2" pageNumber="305" value="2019-05-28">
<collectingDate id="AA41E9C8FFCAFFC5278810B3EB8EFBE3" pageId="2" pageNumber="305" value="2019-05-28">28 May 2019</collectingDate>
</date>
(MA11189)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5267D1093EC29FBC3" collectingDate="2019-06-03" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5267D1093EC9BFBE3" box="[1106,1184,1013,1037]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526801093ED26FBE3" box="[1199,1309,1013,1037]" count="1" pageId="2" pageNumber="305" type="female">1 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC527041093ED50FBE3" box="[1323,1387,1013,1037]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC527561093EB40FBC3" pageId="2" pageNumber="305" value="2019-06-03">
<collectingDate id="AA41E9C8FFCAFFC527561093EB40FBC3" pageId="2" pageNumber="305" value="2019-06-03">3 June 2019</collectingDate>
</date>
(MA10982)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5260D1773EC3DFBA3" collectingDate="2019-06-11" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="9" specimenCount-female="4" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5260D1773ECBBFBC3" box="[1058,1152,1045,1069]" count="5" pageId="2" pageNumber="305" type="male">5 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526BF1773ED3EFBC3" box="[1168,1285,1045,1069]" count="4" pageId="2" pageNumber="305" type="female">4 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5273B1773ED63FBC3" box="[1300,1368,1045,1069]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC527481773EB42FBA3" pageId="2" pageNumber="305" value="2019-06-11">
<collectingDate id="AA41E9C8FFCAFFC527481773EB42FBA3" pageId="2" pageNumber="305" value="2019-06-11">11 June 2019</collectingDate>
</date>
(MA11180)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5263D1753EBFCFB83" collectingDate="2019-06-19" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5263D1753EC65FBA3" box="[1042,1118,1077,1101]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526441753ECE3FBA3" box="[1131,1240,1077,1101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC526CB1753ED18FBA3" box="[1252,1315,1077,1101]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5271F1753EDF8FBA3" box="[1328,1475,1077,1101]" pageId="2" pageNumber="305" value="2019-06-19">
<collectingDate id="AA41E9C8FFCAFFC5271F1753EDF8FBA3" box="[1328,1475,1077,1101]" pageId="2" pageNumber="305" value="2019-06-19">19 June 2019</collectingDate>
</date>
(MA10976)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC521FA1733ED84FB83" box="[981,1471,1109,1133]" collectingDate="2019-06-25" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC521FA1733EC03FB83" box="[981,1080,1109,1133]" count="1" pageId="2" pageNumber="305" type="female">1 female</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC526691733ECBDFB83" box="[1094,1158,1109,1133]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC526BB1733ED17FB83" box="[1172,1324,1109,1133]" pageId="2" pageNumber="305" value="2019-06-25">
<collectingDate id="AA41E9C8FFCAFFC526BB1733ED17FB83" box="[1172,1324,1109,1133]" pageId="2" pageNumber="305" value="2019-06-25">25 June 2019</collectingDate>
</date>
, (MA10967)
</materialsCitation>
. Barcodes of
<materialsCitation id="7ED33CBDFFCAFFC521E11713ECD8FB63" box="[974,1251,1141,1165]" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="58" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC521E11713EC5BFB63" box="[974,1120,1141,1165]" count="58" pageId="2" pageNumber="305" type="generic">58 specimens</specimenCount>
from
<specimenCode id="9E1D9E9BFFCAFFC526891713ECD8FB63" box="[1190,1251,1141,1165]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
</materialsCitation>
are available.
</paragraph>
<paragraph id="CE0436E0FFCAFFC5216C17D3EC41FAE3" blockId="2.[835,1474,1205,1293]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C17D3EC53FB23" bold="true" box="[835,1128,1205,1229]" pageId="2" pageNumber="305">Other material examined</emphasis>
. 2,357 barcodes of
<taxonomicName id="09BB4D63FFCAFFC5276C17D0EDBDFB23" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1347,1414,1205,1229]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot" status="new species">
<emphasis id="FCCFEAF2FFCAFFC5276C17D0EDBDFB23" box="[1347,1414,1205,1229]" italics="true" pageId="2" pageNumber="305">C. dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCAFFC527B917D0EBADFB03" pageId="2" pageNumber="305" rank="species">new species</taxonomicNameLabel>
, are available from Pulau Ubin, Pulau [=
<collectingCountry id="B6AC7670FFCAFFC5275D17B3ED80FB03" box="[1394,1467,1237,1261]" name="Iceland" pageId="2" pageNumber="305">Island</collectingCountry>
] Semakau, and Coney Island.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC5216C1653ED2DFA63" pageId="2" pageNumber="305" type="etymology">
<paragraph id="CE0436E0FFCAFFC5216C1653ED2DFA63" blockId="2.[834,1474,1333,1421]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C1653EBFEFAA3" bold="true" box="[835,965,1333,1357]" pageId="2" pageNumber="305">Etymology.</emphasis>
The species is named after the brown spot or dot that is present on the fore femora of the males and females. The English word
<taxonomicName id="09BB4D63FFCAFFC5263F1613EC7BFA63" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1040,1088,1397,1421]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot">“dot</taxonomicName>
” is used as a noun.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC3216C16D3E91BFDDB" lastPageId="4" lastPageNumber="307" pageId="2" pageNumber="305" type="diagnosis">
<paragraph id="CE0436E0FFCAFFC5216C16D3EC85F823" blockId="2.[834,1476,1461,1997]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C16D3EB83FA23" bold="true" box="[835,952,1461,1485]" pageId="2" pageNumber="305">Diagnosis.</emphasis>
A small species (
<quantity id="09439B05FFCAFFC5265716D3ECC0FA23" box="[1144,1275,1461,1485]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
) with a black spot located dorsally on the middle of the fore femur in both sexes. Four dorsocentrals, 5 uniserial acrostichals. Haltere yellowish. Legs yellow, with apical 2 tarsomeres of all legs brown. Male: Antenna yellowish brown, pedicel contrastingly darker brown than the yellowish brown postpedicel. Frons very wide, eyes touching near middle of face, with at least 4 rows of enlarged eye facets on each side of face. Palpus yellow, large, oval, hiding proboscis and shorter in females, lacking black bristling. Apical 2 tarsomeres of all legs brown, slightly dorsoventrally flattened. Pulvilli present on all legs, though minute and narrow. Sternite 8 lacking macrosetae. Cerci yellow, ventral surstylus dark brown. Tip of aedeagus widened with complex structure. Female: palpus brown, not hiding the proboscis, covered with black bristles and a distinct apical. Legs coloured like in males. Apical tarsomeres of all legs brownish but not widened.
</paragraph>
<paragraph id="CE0436E0FFCAFFC5216C1493EDBBF7E3" blockId="2.[835,1408,2037,2061]" box="[835,1408,2037,2061]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C1493EBBFF7E3" bold="true" box="[835,900,2037,2061]" pageId="2" pageNumber="305">Male.</emphasis>
Length: body
<quantity id="09439B05FFCAFFC526091493EC97F7E3" box="[1062,1196,2037,2061]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
; wing
<quantity id="09439B05FFCAFFC526D91493ED47F7E3" box="[1270,1404,2037,2061]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCBFFC423621314ECEBFF64" blockId="3.[333,1232,114,138]" box="[333,1232,114,138]" pageId="3" pageNumber="306">Grootaert &amp; Van de Velde: New long-legged flies from mangroves on Pulau Ubin</paragraph>
<caption id="9AC46668FFCBFFC4225E1776EA7EFBAC" ID-DOI="http://doi.org/10.5281/zenodo.14683064" ID-Zenodo-Dep="14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306" startId="3.[113,148,1040,1062]" targetBox="[95,1465,167,1091]" targetPageId="3" targetType="figure">
<paragraph id="CE0436E0FFCBFFC4225E1776EA7EFBAC" blockId="3.[113,1453,1040,1091]" pageId="3" pageNumber="306">
Fig. 3.
<taxonomicName id="09BB4D63FFCBFFC422971776E904FBCB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[184,319,1040,1061]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCBFFC422971776E904FBCB" box="[184,319,1040,1061]" italics="true" pageId="3" pageNumber="306">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCBFFC423641776E9F9FBC8" box="[331,450,1040,1062]" pageId="3" pageNumber="306" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFCBFFC4225E171CE9D4FADC" blockId="3.[113,753,1146,1330]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E171CE88EFB7C" bold="true" box="[113,181,1146,1170]" pageId="3" pageNumber="306">Head.</emphasis>
Frons very wide, metallic green. Eyes touching near middle of face, with at least 4 rows of enlarged eye facets at each side of face. All bristles black. A pair of long retroclinate, diverging ocellar bristles, a pair of proclinate vertical bristles as long as ocellars. Postoculars, short, fine, uniserial above, not distinct below.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E163FE9D1F9FC" blockId="3.[113,754,1369,1554]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E163FE8DCFA9F" bold="true" box="[113,231,1369,1393]" pageId="3" pageNumber="306">Antennae.</emphasis>
Yellowish brown, pedicel contrastingly darker brown than postpedicel which is more yellowish. Pedicel with dorsal apical bristle nearly as long as pedicel is high. Postpedicel kidney-shaped, arista apical near middle, set with minute hairs. Palpus yellow, very large, oval, hiding proboscis, lacking distinct bristles.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E155CE9B0F95C" blockId="3.[112,753,1594,1714]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E155CE8EAF9BC" bold="true" box="[113,209,1594,1618]" pageId="3" pageNumber="306">Thorax.</emphasis>
Shiny metallic green. All bristles black. Four dorsocentrals almost equally long. Five uniserial acrostichals. A pair of widely separated marginal scutellars 2.5 times as long as scutellum is long.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E15BCEA28F8DC" blockId="3.[112,752,1754,1842]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E15BCE895F91C" bold="true" box="[113,174,1754,1778]" pageId="3" pageNumber="306">Legs.</emphasis>
Yellow, fore femur with a blackish brown spot located dorsally in middle. Mid coxa brown, hind coxae paler brown. Apical 2 tarsomeres of all legs brown.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E143CEACBF85C" blockId="3.[112,752,1882,1970]" pageId="3" pageNumber="306">Fore legs. Coxa with indistinct pale apical bristles. Femur and tibia lacking distinct bristles. Length femur, tibia, tarsomeres (in mm): 0.33: 0.34: 0.12: 0.047: 0.039: 0.032: 0.032.</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E14BCEBEEFB1C" blockId="3.[113,753,2010,2066]" lastBlockId="3.[813,1452,1146,1266]" pageId="3" pageNumber="306">Mid legs. Coxa lacking an exterior bristle, but a long bristle on middle of margin between anterior and dorsal part of the coxa. Femur with a brown posterior preapical bristle. Tibia with a short anterodorsal in basal quarter. Length femur, tibia, tarsomeres (in mm): 0.39: 0.39: 0.19: 0.09: 0.06: 0.032: 0.032.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102167CED23FA3C" blockId="3.[812,1452,1306,1490]" pageId="3" pageNumber="306">Hind legs. A brown exterior bristle near base of coxa, longer than coxa is high and a slightly shorter apical bristle. Trochanter with a long dorsal bristle longer than trochanter. Femur with 3 preapical anteroventral bristles, middle bristle longest, longer than width of femur. Tibia lacking distinct bristles except for a few short apical bristles.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102169CEC71F9DC" blockId="3.[813,1452,1530,1586]" pageId="3" pageNumber="306">Length femur, tibia, tarsomeres (in mm): 0.39: 0.39: 0.17: 0.08: 0.05: 0.32: 0.05.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102153FEB8EF95C" blockId="3.[813,1452,1625,1714]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC42102153FEB43F99F" bold="true" box="[813,888,1625,1649]" pageId="3" pageNumber="306">Wings.</emphasis>
Brownish tinge with pale brown veins. Squama brown with 2 pale brown bristles. Haltere yellowish (not clear white).
</paragraph>
<paragraph id="CE0436E0FFCBFFC4210215BFEBCEF8DC" blockId="3.[812,1452,1753,1842]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4210215BFEB98F91F" bold="true" box="[813,931,1753,1777]" pageId="3" pageNumber="306">Abdomen.</emphasis>
Brown, indistinctly shiny. All bristles black and short. Sternite 8 lacking macrosetae though a row of bristles is present (
<figureCitation id="56802A65FFCBFFC4218B147CEBDCF8DC" box="[932,999,1818,1842]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFCBFFC42102143CEDB3F7FC" blockId="3.[813,1453,1881,2066]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC42102143CEBD5F89F" bold="true" box="[813,1006,1881,1906]" pageId="3" pageNumber="306">Male terminalia.</emphasis>
<figureCitation id="56802A65FFCBFFC421D7143CEC07F89C" box="[1016,1084,1882,1906]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3</figureCitation>
. Cercus yellow, surstylus brown. Epandrial lobe with 3 bristles, the apical bristle as long as lobe is wide, other 2 bristles minute. Ventral surstylus with a strong subapical bristle and a smaller bristle below. Apex of aedeagus widened with a complex structure (
<figureCitation id="56802A65FFCBFFC4270414BCED9AF81C" box="[1323,1441,2010,2034]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3AC</figureCitation>
). Postgonite short, densely set with short hairs (
<figureCitation id="56802A65FFCBFFC42709149CED47F7FC" box="[1318,1404,2042,2066]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A713DBEAC3FF3B" blockId="4.[136,760,189,213]" box="[136,760,189,213]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A713DBE8D9FF3B" bold="true" box="[136,226,189,213]" pageId="4" pageNumber="307">Female.</emphasis>
Length: body
<quantity id="09439B05FFCCFFC323AB13DBEA31FF3B" box="[388,522,189,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="4" pageNumber="307" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
; wing
<quantity id="09439B05FFCCFFC3207B13DBEACFFF3B" box="[596,756,189,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.11" metricValueMin="1.1" pageId="4" pageNumber="307" unit="mm" value="1.105" valueMax="1.11" valueMin="1.1">1.101.11 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7139BE91BFDDB" blockId="4.[135,777,253,565]" pageId="4" pageNumber="307">Similar to males in colouration of legs. Palpus brown and much smaller than in males, with longer black bristles and a distinct apical. Proboscis yellowish, not hidden by palpus, long, about half as long as eye is high. Face as wide as postpedicel, apically protruding and pointed. Postpedicel reniform as in males, but more uniformly brown. Eye facets not enlarged and eyes not as wide as in males so that the row of lower postoculars is visible. Postoculars black above, longer and yellow below. Marginal bristles on tergites longer than in males.</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC322A7113BED82FF1B" pageId="4" pageNumber="307" type="discussion">
<paragraph id="CE0436E0FFCCFFC322A7113BEB3DFC3B" blockId="4.[135,776,605,981]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7113BE937FD9B" bold="true" box="[136,268,605,629]" pageId="4" pageNumber="307">Comments.</emphasis>
This small new species is easily recognised by the yellow legs with a black spot located dorsally in the middle of the fore femora and this is in both sexes. The postpedicel is yellowish brown and somewhat kidney-shaped with an apical arista. The male has a very large oval yellowish palpus, while the female has a smaller brown palpus. It is the most common
<taxonomicName id="09BB4D63FFCCFFC3237B107BEA28FCDB" box="[340,531,797,821]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3237B107BE9FBFCDB" box="[340,448,797,821]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
in back mangroves. It hardly visits front mangroves and was never observed in non-mangrove biotopes where many species of the entirely black
<taxonomicName id="09BB4D63FFCCFFC322E7101BE9BCFC7B" box="[200,391,893,917]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322E7101BE90FFC7B" box="[200,308,893,917]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
are dominant. The genetic variation measured by the number of haplotypes is low, especially in comparison with the terrestrial black
<taxonomicName id="09BB4D63FFCCFFC3201010DBEB3DFC3B" box="[575,774,957,981]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFCCFFC3201010DBEA90FC3B" box="[575,683,957,981]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7109BEAB9FB1B" blockId="4.[136,776,1021,1270]" pageId="4" pageNumber="307">
The tip of the aedeagus in
<taxonomicName id="09BB4D63FFCCFFC32380109BEA78FBFB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[431,579,1021,1045]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCCFFC32380109BEA78FBFB" box="[431,579,1021,1045]" italics="true" pageId="4" pageNumber="307">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC3207D1098EAECFBFB" box="[594,727,1021,1045]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, has a complex structure with folds and a spine-like projection (
<figureCitation id="56802A65FFCCFFC322BF175BE936FBBB" box="[144,269,1085,1109]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="4" pageNumber="307">Fig. 3AC</figureCitation>
), similar to species of the
<taxonomicName id="09BB4D63FFCCFFC3206B1758EB3CFBBB" authorityName="Wei &amp; Yang" authorityYear="2007" box="[580,775,1085,1109]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="leigongshanus">
<emphasis id="FCCFEAF2FFCCFFC3206B1758EB3CFBBB" box="[580,775,1085,1109]" italics="true" pageId="4" pageNumber="307">C. leigongshanus</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="AA2A4B11FFCCFFC32307173BE9A6FB9B" author="Wei L" box="[296,413,1117,1141]" pageId="4" pageNumber="307" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei, 2018</bibRefCitation>
). However, this species group has black femora, no pulvilli on the hind tarsus and sternite 8 with a pair of strong setae. In
<taxonomicName id="09BB4D63FFCCFFC323F817FBEA55FB5B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[471,622,1181,1205]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCCFFC323F817FBEA55FB5B" box="[471,622,1181,1205]" italics="true" pageId="4" pageNumber="307">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC3205117F8EB38FB5B" box="[638,771,1181,1205]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, the femora are yellow, there are small pulvilli present on all legs, and sternite 8 does not bear strong setae.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7167BEA32FA3B" blockId="4.[135,777,1309,1493]" pageId="4" pageNumber="307">
Several other, but less common and yet to be described
<taxonomicName id="09BB4D63FFCCFFC322A7165BE97EFABB" box="[136,325,1341,1365]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7165BE8C8FABB" box="[136,243,1341,1365]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
with yellow legs occur in back mangroves. They do not bear the typical black spot on the fore femora and have a brown triangular or trapezoid postpedicel with a short ventral protuberance. One of these species,
<taxonomicName id="09BB4D63FFCCFFC3205016F8EAC4FA5B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[639,767,1437,1461]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="ubinensis" status="new species">
<emphasis id="FCCFEAF2FFCCFFC3205016F8EAC4FA5B" box="[639,767,1437,1461]" italics="true" pageId="4" pageNumber="307">C. ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC322A716D8E936FA3B" box="[136,269,1469,1493]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, is described hereafter.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7169BE93DF91B" blockId="4.[135,777,1533,1781]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7169BE9BBF9FB" bold="true" box="[136,384,1533,1557]" pageId="4" pageNumber="307">National distribution.</emphasis>
The new species is one of the most common
<taxonomicName id="09BB4D63FFCCFFC322C2157BE962F9DB" authorityName="Meigen" authorityYear="1824" box="[237,345,1565,1589]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCCFFC322C2157BE962F9DB" box="[237,345,1565,1589]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
</taxonomicName>
and dolichopodid species, in
<collectingCountry id="B6AC7670FFCCFFC320B4157BEB3CF9DB" box="[667,775,1565,1589]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
in the back mangroves of Pulau Ubin. It was found all over the island in all 26 back mangrove stations that were sampled with Malaise traps (
<figureCitation id="56802A65FFCCFFC3234F151BE99BF97B" box="[352,416,1661,1685]" captionStart="Fig" captionStartId="1.[113,148,1433,1455]" captionTargetBox="[113,1451,189,1417]" captionTargetId="figure-15@1.[113,1451,189,1417]" captionTargetPageId="1" captionText="Fig. 1. Distribution of the sampling stations with Malaise traps on Pulau Ubin. A, all sites from 2005 to 2019; B, detailed position of traps in the OBS area in 2019." figureDoi="http://doi.org/10.5281/zenodo.14683058" httpUri="https://zenodo.org/record/14683058/files/figure.png" pageId="4" pageNumber="307">Fig. 1</figureCitation>
). In total, 2,784 specimens were collected in the back mangroves and 2,357 specimens were barcoded, showing very little variation in haplotypes (less than 0.7%).
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A71478ED82FF1B" blockId="4.[135,776,1821,2101]" lastBlockId="4.[834,1473,189,246]" pageId="4" pageNumber="307">
<materialsCitation id="7ED33CBDFFCCFFC322A71478ED8EFF1B" collectionCode="SMO" county="Mandai" location="It" municipality="Lim Chu Kang" pageId="4" pageNumber="307" specimenCode="SMN3" specimenCount="484">
In the front mangroves of the replanted mangroves of Pulau Semakau, only a
<collectingMethod id="17FA4EF7FFCCFFC323BD145BE96DF89B" pageId="4" pageNumber="307">single specimen was found in the three Malaise traps</collectingMethod>
that were operational for two full years (
<specimenCode id="9E1D9E9BFFCCFFC322BE141BE8E6F87B" box="[145,221,1917,1941]" collectionCode="SMN" country="China" lsid="urn:lsid:biocol.org:col:14058" name="Simao District National Medical and Pharmaceutical Institute" pageId="4" pageNumber="307" type="Herbarium">SMN3</specimenCode>
), while
<specimenCount id="D8BDFD69FFCCFFC32301141BE9F2F87B" box="[302,457,1917,1941]" count="239" pageId="4" pageNumber="307" type="generic">239 specimens</specimenCount>
were found in the adjacent old mangroves (
<collectionCode id="A8AAAE25FFCCFFC3232714FBE97DF85B" box="[264,326,1949,1973]" pageId="4" pageNumber="307">SMO</collectionCode>
), thus underlining that
<taxonomicName id="09BB4D63FFCCFFC3201A14F8EA4EF85B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[565,629,1949,1973]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="new species">
<emphasis id="FCCFEAF2FFCCFFC3201A14F8EA4EF85B" box="[565,629,1949,1973]" italics="true" pageId="4" pageNumber="307">C. dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC320AD14F8EB38F85B" box="[642,771,1949,1973]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, is really a back mangrove species.
<location id="CB64603BFFCCFFC3202514D8EA21F83B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3202514D8EA21F83B" box="[522,538,1982,2005]" county="Mandai" municipality="Lim Chu Kang" name="It" pageId="4" pageNumber="307">It</location>
was further found in the back mangroves of
<location id="CB64603BFFCCFFC323BC14BBEA1DF81B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC323BC14BBEA1DF81B" box="[403,550,2013,2037]" county="Mandai" municipality="Lim Chu Kang" name="Coney Island" pageId="4" pageNumber="307">Coney Island</location>
(
<specimenCount id="D8BDFD69FFCCFFC3201614BBEAEBF81B" box="[569,720,2013,2037]" count="17" pageId="4" pageNumber="307" type="generic">17 specimens</specimenCount>
in 5 sampled stations),
<location id="CB64603BFFCCFFC32360149BE9CAF7FB" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC32360149BE9CAF7FB" box="[335,497,2045,2069]" county="Mandai" municipality="Lim Chu Kang" name="Berlayer Creek" pageId="4" pageNumber="307">Berlayer Creek</location>
(
<specimenCount id="D8BDFD69FFCCFFC3202D149BEA42F7FB" box="[514,633,2045,2069]" count="1" pageId="4" pageNumber="307" type="generic">1 specimen</specimenCount>
in 2 sampled stations), and
<location id="CB64603BFFCCFFC323371B7BEA64F7DB" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC323371B7BEA64F7DB" box="[280,607,2077,2101]" county="Mandai" municipality="Lim Chu Kang" name="Sungei Buloh Wetland Reserve" pageId="4" pageNumber="307">Sungei Buloh Wetland Reserve</location>
(
<specimenCount id="D8BDFD69FFCCFFC320441B7BEB3CF7DB" box="[619,775,2077,2101]" count="227" pageId="4" pageNumber="307" type="generic">227 specimens</specimenCount>
in 14 sampled stations).
<location id="CB64603BFFCCFFC3267A13DBECE0FF3B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3267A13DBECE0FF3B" box="[1109,1243,189,213]" county="Mandai" municipality="Lim Chu Kang" name="Remarkably" pageId="4" pageNumber="307">Remarkably</location>
, it was not found in the mangroves at
<location id="CB64603BFFCCFFC3262913BBEC4BFF1B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3262913BBEC4BFF1B" box="[1030,1136,221,245]" county="Mandai" municipality="Lim Chu Kang" name="Sarimbun" pageId="4" pageNumber="307">Sarimbun</location>
,
<collectingMunicipality id="2E60AC9AFFCCFFC3265213BBED18FF18" box="[1149,1315,221,246]" pageId="4" pageNumber="307">Lim Chu Kang</collectingMunicipality>
, and
<collectingCounty id="27654E6CFFCCFFC3274F13BBED8EFF1B" box="[1376,1461,221,245]" pageId="4" pageNumber="307">Mandai</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC3216C127BECD3FD3B" pageId="4" pageNumber="307" type="distribution">
<paragraph id="CE0436E0FFCCFFC3216C127BEDFFFDFB" blockId="4.[834,1476,285,533]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C127BECA6FEDB" bold="true" box="[835,1181,285,309]" pageId="4" pageNumber="307">Distribution in Southeast Asia.</emphasis>
The new species was found in all three mangrove sites in
<collectingCountry id="B6AC7670FFCCFFC3265E125BEC82FEBB" box="[1137,1209,317,341]" name="Brunei" pageId="4" pageNumber="307">Brunei</collectingCountry>
we investigated (
<collectingRegion id="0C7FF802FFCCFFC32742125BED86FEBB" box="[1389,1469,317,341]" country="Brunei" name="Tutong" pageId="4" pageNumber="307">Tutong</collectingRegion>
, Berambang, and Labu). They differ by 2.3% in haplotype structure from the
<collectingCountry id="B6AC7670FFCCFFC3260E121BECA8FE7B" box="[1057,1171,381,405]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
populations and a deeper morphological study is needed to show if they represent the same species. Only a single female was found in a mangrove site in southern
<collectingCountry id="B6AC7670FFCCFFC321DF12BBEC74FE1B" box="[1008,1103,477,501]" name="Thailand" pageId="4" pageNumber="307">Thailand</collectingCountry>
(
<collectingRegion id="0C7FF802FFCCFFC3267012BBECD9FE1B" box="[1119,1250,477,501]" country="Thailand" name="Surat Thani" pageId="4" pageNumber="307">Surat Thani</collectingRegion>
, leg. A. Samoh) and genetically it falls entirely in the population of Pulau Ubin.
</paragraph>
<paragraph id="CE0436E0FFCCFFC3216C115BECD3FD3B" blockId="4.[835,1476,573,725]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C115BEBFAFDBB" bold="true" box="[835,961,573,597]" pageId="4" pageNumber="307">Phenology.</emphasis>
The new species is active throughout the year, but with its peak activity at the start of the rainy season in April and the lowest activity in the dryer periods of the year from end June to end September, with a second peak of activity from October to December.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,120 @@
<document id="90AA7F09A5592650756D0062E2E391D8" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFCAFFC52339119BE94FF9BB" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chrysotus Meigen 1824" docType="treatment" docVersion="2" lastPageNumber="305" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="305" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="9D90F9B8DE0CF49FC1D650F62FD21FC0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="49B6B2EFA858706011973113A2D86CE5">
<mods:title id="5F05E186C1E9E5FF469528EBB983DF30">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="E356304E7622DF49710B984D7B0228C5" type="personal">
<mods:role id="D8D59D508130E1396BCB489A51CD7E8E">
<mods:roleTerm id="038DD2D1E350E89D5107447320713D89">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C6100F302C93E75996DA5125CE0EF63D">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="861A274DEA71AA41D2CE6DAD2E688669" type="personal">
<mods:role id="5C6B8616619523E1BC80E3BDB8350307">
<mods:roleTerm id="DDD7BD23CE24C53CE8753444F95E514E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10D9995403989A6590C21C8F5F934859">Velde,</mods:namePart>
</mods:name>
<mods:name id="88BEDFDD0A34A2BDE197BA14888B996D" type="personal">
<mods:role id="79EE9780EBB26BDDB7A08B9BA17DA58C">
<mods:roleTerm id="10FCD9171D30CD8089F1ECE03AA0539A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4F9002519D636571EDC3620937E5E49">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="370EE508CEB75626CC76F0C2EA1DDA6B">text</mods:typeOfResource>
<mods:relatedItem id="E3EEA676E64E38A59FF4CF933DEEE918" type="host">
<mods:titleInfo id="43C2939D83B16D0E21B2F07F17D41B68">
<mods:title id="433DBB49944EE01073946D645635C9D9">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1210C3C73BD7932990CC6A173F2D5176">
<mods:date id="6C5EC6BAB65E8046AD63E83CCD34AC23">2024</mods:date>
<mods:detail id="74F1335B6600E90949E25BD8EF96BF61" type="pubDate">
<mods:number id="B8F6E8926E973FA534AFAC50AC210F1D">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="881B9B28491EBF62BBB4D1DBB35DB5DE" type="volume">
<mods:number id="12C22E5D6968A23716A004BC5A80F175">72</mods:number>
</mods:detail>
<mods:extent id="CC972D8D2D552117B557EA18E6BCD5B7" unit="page">
<mods:start id="26CA5D71735474F90F2ED955E89F5CB2">303</mods:start>
<mods:end id="11889944347EE18BFA7B5B08B2486666">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4F4700676A67EE44AE026B05EA637943">journal article</mods:classification>
<mods:identifier id="9A1A9080257A2CE9434CC2F594BA8721" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="550F001FF575CCE069BFBE47D2CAE106" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="683A4B3293C9C4C112ACA6C78846FF5D" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="1762CE05069F4F86940E95F802D88F1B" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFCAFFC52339119BE94FF9BB" ID-DOI="http://doi.org/10.5281/zenodo.14683072" ID-Zenodo-Dep="14683072" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC52339119BE94FF9BB" httpUri="http://treatment.plazi.org/id/461287F6FFCAFFC52339119BE94FF9BB" lastPageNumber="305" pageId="2" pageNumber="305">
<subSubSection id="86A1656BFFCAFFC52339119BEA43FCFB" box="[278,632,765,789]" pageId="2" pageNumber="305" type="nomenclature">
<paragraph id="CE0436E0FFCAFFC52339119BEA43FCFB" blockId="2.[278,632,765,789]" box="[278,632,765,789]" pageId="2" pageNumber="305">
<heading id="954C818CFFCAFFC52339119BEA43FCFB" bold="true" box="[278,632,765,789]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="305" reason="2">
<emphasis id="FCCFEAF2FFCAFFC52339119BEA43FCFB" bold="true" box="[278,632,765,789]" pageId="2" pageNumber="305">
Genus
<taxonomicName id="09BB4D63FFCAFFC52347119BEA43FCFB" ID-CoL="9G2FY" authority="Meigen, 1824" authorityName="Meigen" authorityYear="1824" box="[360,632,765,789]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCAFFC52347119BE9ECFCFB" bold="true" box="[360,471,765,789]" italics="true" pageId="2" pageNumber="305">Chrysotus</emphasis>
<bibRefCitation id="AA2A4B11FFCAFFC523F0119BEA43FCFB" author="Meigen JW" box="[479,632,765,789]" pageId="2" pageNumber="305" pagination="33 - 41" refId="ref15074" refString="Meigen JW (1824). Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Volume 4. Hamm, xii + 428 pp., pis. 33 - 41." type="book chapter" year="1824">Meigen, 1824</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC522A7105BE94FF9BB" pageId="2" pageNumber="305" type="materials_examined">
<paragraph id="CE0436E0FFCAFFC522A7105BEB32FABB" blockId="2.[134,777,829,1365]" pageId="2" pageNumber="305">
The genus
<taxonomicName id="09BB4D63FFCAFFC5232A105BE949FCBB" authorityName="Meigen" authorityYear="1824" box="[261,370,829,853]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCAFFC5232A105BE949FCBB" box="[261,370,829,853]" italics="true" pageId="2" pageNumber="305">Chrysotus</emphasis>
</taxonomicName>
is very diverse with more than 550 species already known worldwide and more than 200 species known from the Oriental realm (
<bibRefCitation id="AA2A4B11FFCAFFC523FB101BEAB2FC7B" author="Yang D &amp; Zhu YJ &amp; Wang M &amp; Zhang LL" box="[468,649,893,917]" pageId="2" pageNumber="305" refId="ref15691" refString="Yang D, Zhu YJ, Wang M &amp; Zhang LL (2006) World Catalog of Dolichopodidae (Insecta: Diptera). China Agricultural University Press, Beijing, 704 pp." type="book" year="2006">Yang et al., 2006</bibRefCitation>
;
<bibRefCitation id="AA2A4B11FFCAFFC520BD101BEB3AFC7B" author="Wei L" box="[658,769,893,917]" pageId="2" pageNumber="305" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei, 2018</bibRefCitation>
;
<bibRefCitation id="AA2A4B11FFCAFFC522A710FBE908FC5B" author="Liu RS &amp; Zhong Y &amp; Zhang R &amp; Yang D" box="[136,307,925,949]" pageId="2" pageNumber="305" pagination="1 - 6" refId="ref14990" refString="Liu RS, Zhong Y, Zhang R &amp; Yang D (2020) A study of Chrysotus (Diptera: Dolichopodidae) from Sichuan, China with one new species. Entomotaxonomia, 42 (2): 1 - 6." type="journal article" year="2020">Liu et al., 2020</bibRefCitation>
). These small dolichopodid flies of about
<quantity id="09439B05FFCAFFC520D610FBE88AFC3B" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="305" unit="mm" value="2.0">2 mm</quantity>
in body length need microscopic identification and the male terminalia need to be dissected and examined in order to classify them into species groups (
<bibRefCitation id="AA2A4B11FFCAFFC52022109BEA81FBFB" author="Wei LM &amp; Zhang L &amp; Zhou ZX" box="[525,698,1021,1045]" pageId="2" pageNumber="305" pagination="1 - 112" refId="ref15518" refString="Wei LM, Zhang L &amp; Zhou ZX (2015) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of papuanus species group. Oriental Insects, 48 (3 - 4): 1 - 112." type="journal article" year="2015">Wei et al., 2015</bibRefCitation>
). In the Oriental region, many
<taxonomicName id="09BB4D63FFCAFFC523AF177BEA7BFBDB" box="[384,576,1053,1077]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC523AF177BE9D7FBDB" box="[384,492,1053,1077]" italics="true" pageId="2" pageNumber="305">Chrysotus</emphasis>
species
</taxonomicName>
coexist and many species are yet to be described. Identification of females based on morphology is almost impossible because females are all very similar-looking and are hardly ever associated with males in species descriptions. In his recent revision of the
<taxonomicName id="09BB4D63FFCAFFC5228017DBE980FB3B" authorityName="Wei &amp; Yang" authorityYear="2007" box="[175,443,1213,1237]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="leigongshanus">
<emphasis id="FCCFEAF2FFCAFFC5228017DBE980FB3B" box="[175,443,1213,1237]" italics="true" pageId="2" pageNumber="305">Chrysotus leigongshanus</emphasis>
</taxonomicName>
species group mainly based on material collected in the mountainous areas in southern
<collectingCountry id="B6AC7670FFCAFFC520EE17BBEB39FB1B" box="[705,770,1245,1269]" name="China" pageId="2" pageNumber="305">China</collectingCountry>
,
<bibRefCitation id="AA2A4B11FFCAFFC522A9179BE8C7FAFB" author="Wei L" box="[134,252,1277,1301]" pageId="2" pageNumber="305" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei (2018)</bibRefCitation>
treated 59 species and described 56 species as new to science. Of these, 48 species were only known from the
<typeStatus id="11008842FFCAFFC522A5165BE8DCFABB" box="[138,231,1341,1365]" pageId="2" pageNumber="305" type="holotype">holotype</typeStatus>
male, of which none were associated with females.
</paragraph>
<caption id="9AC46668FFCAFFC5216C102DED86FC90" ID-DOI="http://doi.org/10.5281/zenodo.14683060" ID-Zenodo-Dep="14683060" httpUri="https://zenodo.org/record/14683060/files/figure.png" pageId="2" pageNumber="305" startId="2.[835,873,843,865]" targetBox="[835,1474,189,828]" targetPageId="2" targetType="figure">
<paragraph id="CE0436E0FFCAFFC5216C102DED86FC90" blockId="2.[835,1473,843,894]" pageId="2" pageNumber="305">
Fig. 2.
<taxonomicName id="09BB4D63FFCAFFC521B9102DEC16FC8E" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[918,1069,843,864]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCAFFC521B9102DEC16FC8E" box="[918,1069,843,864]" italics="true" pageId="2" pageNumber="305">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCAFFC52610102AECFFFC8F" box="[1087,1220,843,865]" pageId="2" pageNumber="305" rank="species">new species</taxonomicNameLabel>
, habitus paratype male (ZRCBDP0083923). Photograph by Miteshravin S/O Ravindran.
</paragraph>
</caption>
<paragraph id="CE0436E0FFCAFFC522A7161BE94FF9BB" blockId="2.[135,777,1405,1621]" pageId="2" pageNumber="305">
The two species described in this paper do not belong to those from the
<collectingRegion id="0C7FF802FFCAFFC5230316FBE9F4FA5B" box="[300,463,1437,1461]" country="Egypt" name="Ash Sharqiyah" pageId="2" pageNumber="305">Oriental region</collectingRegion>
that were recently described or revised (
<bibRefCitation id="AA2A4B11FFCAFFC5232816DBE985FA3B" author="Wei LM &amp; Zhang L &amp; Zhou ZX" box="[263,446,1469,1493]" pageId="2" pageNumber="305" pagination="1 - 112" refId="ref15518" refString="Wei LM, Zhang L &amp; Zhou ZX (2015) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of papuanus species group. Oriental Insects, 48 (3 - 4): 1 - 112." type="journal article" year="2015">Wei et al., 2015</bibRefCitation>
;
<bibRefCitation id="AA2A4B11FFCAFFC523E416DBEA7AFA3B" author="Wei L" box="[459,577,1469,1493]" pageId="2" pageNumber="305" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei, 2018</bibRefCitation>
;
<bibRefCitation id="AA2A4B11FFCAFFC5206116DBEAC6FA3B" author="Liu RS &amp; Zhong Y &amp; Zhang R &amp; Yang D" box="[590,765,1469,1493]" pageId="2" pageNumber="305" pagination="1 - 6" refId="ref14990" refString="Liu RS, Zhong Y, Zhang R &amp; Yang D (2020) A study of Chrysotus (Diptera: Dolichopodidae) from Sichuan, China with one new species. Entomotaxonomia, 42 (2): 1 - 6." type="journal article" year="2020">Liu et al., 2020</bibRefCitation>
). Moreover, both new species are exclusive mangrove species, a biotope that was never well investigated. Barcodes, which distinguish the new species from other related, cryptic species, are provided.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,427 @@
<document id="71C1D7A717D54FCDAED51CD367DB925C" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFCCFFC121E7119BED64F95C" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chrysotus ubinensis Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="309" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="307" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A3B8E9D6286EA05A15926FA97209A48A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2ED355F3B86BE159ED85A23F4DEE52DD">
<mods:title id="7505F752260017AC73EA9A19065CBD5E">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="CA0476133BCD6219E784DF9A8B985FAF" type="personal">
<mods:role id="6D48ED629F938F088995762185C51015">
<mods:roleTerm id="7C0C71B91634538EE22D38D587BE1C77">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AADBD109209688384609AFF785946117">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="3D799CAB8E37108377632CEDDF2738E4" type="personal">
<mods:role id="0327CB70A3A319FF3E60DD5824A873A7">
<mods:roleTerm id="1492084D5C14307C01066FE1F7DF1274">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AFFD105C1ED73CCAB5E4D797128A6710">Velde,</mods:namePart>
</mods:name>
<mods:name id="D7635949FA5AE3E707D7B50C873E5744" type="personal">
<mods:role id="43A4975017F9E4A22492EE87C790F14D">
<mods:roleTerm id="D708A066A4DA02147AD80AEED77732D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F898984BDD82C169A08EF5565B21E042">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="2925477542F4C1EFB414E94BD9C28DFD">text</mods:typeOfResource>
<mods:relatedItem id="B1FCFD324170BE52CF8B6A3EA5038A6C" type="host">
<mods:titleInfo id="8495636CBA37567F8FEC52B4521A2A8B">
<mods:title id="41C738DBF2E999A4452C3A2096C2181D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="E64DFCA46651E82A1BC4AA78A988A456">
<mods:date id="C99F34B3D3C22DBF7352E106FA2ACE37">2024</mods:date>
<mods:detail id="73721E36B359A3B7A1381FF234089FFF" type="pubDate">
<mods:number id="4B66461AE6C644629164E88601D5C372">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="4DA2BD7E1767D81E12B3D03EF706B0DF" type="volume">
<mods:number id="BE815C55B3E20D3C1F85050658CBA91D">72</mods:number>
</mods:detail>
<mods:extent id="4E74CACAE8855C111E88FF533D63BEEC" unit="page">
<mods:start id="3ED1D8243F1890B6EAC552EA9E3A906C">303</mods:start>
<mods:end id="DF29FC1DCBEEB4556D7709828619F5A4">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2A5DBE06EF52B80FB73E74A17A41D541">journal article</mods:classification>
<mods:identifier id="61942325BBACB3D3D1A8EE529A6B84FF" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="42B6341B01FDC3AFF52E929AB8CA977F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="DF311F4FDF7C8E766B1B95CAD8B2BD60" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="27067CA4688148C625FD7E19177C6BE4" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFCCFFC121E7119BED64F95C" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C" httpUri="http://treatment.plazi.org/id/461287F6FFCCFFC121E7119BED64F95C" lastPageId="6" lastPageNumber="309" pageId="4" pageNumber="307">
<subSubSection id="86A1656BFFCCFFC321E7119BED05FCFB" box="[968,1342,765,789]" pageId="4" pageNumber="307" type="nomenclature">
<paragraph id="CE0436E0FFCCFFC321E7119BED05FCFB" blockId="4.[968,1342,765,821]" box="[968,1342,765,789]" pageId="4" pageNumber="307">
<heading id="954C818CFFCCFFC321E7119BED05FCFB" bold="true" box="[968,1342,765,789]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="307" reason="2">
<emphasis id="FCCFEAF2FFCCFFC321E7119BED05FCFB" bold="true" box="[968,1342,765,789]" pageId="4" pageNumber="307">
<taxonomicName id="09BB4D63FFCCFFC321E7119BEC92FCFB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[968,1193,765,789]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFCCFFC321E7119BEC92FCFB" bold="true" box="[968,1193,765,789]" italics="true" pageId="4" pageNumber="307">Chrysotus ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC32697119BED05FCFB" box="[1208,1342,765,789]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC32669107BECFBFCDB" box="[1094,1216,797,821]" pageId="4" pageNumber="307" type="description">
<paragraph id="CE0436E0FFCCFFC32669107BECFBFCDB" blockId="4.[968,1342,765,821]" box="[1094,1216,797,821]" pageId="4" pageNumber="307">
(
<figureCitation id="56802A65FFCCFFC32661107BECA0FCDB" box="[1102,1179,797,821]" captionStart="Fig" captionStartId="5.[113,148,1057,1079]" captionTargetBox="[128,1444,189,1033]" captionTargetId="figure-482@5.[783,1452,189,1044]" captionTargetPageId="5" captionText="Fig. 4. Chrysotus ubinensis, new species. A, Habitus paratype male (Ma6107). B, Paratype female (Ma6107). Photographs by Jayanthi Puniamoorthy." figureDoi="http://doi.org/10.5281/zenodo.14683066" httpUri="https://zenodo.org/record/14683066/files/figure.png" pageId="4" pageNumber="307">Figs. 4</figureCitation>
,
<figureCitation id="56802A65FFCCFFC32686107BEC8CFCDB" box="[1193,1207,797,821]" captionStart="Fig" captionStartId="6.[136,171,1104,1126]" captionTargetBox="[138,1472,189,1092]" captionTargetId="figure-8@6.[126,1491,174,1104]" captionTargetPageId="6" captionText="Fig. 5. Chrysotus ubinensis, new species, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="4" pageNumber="307">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC3216C103BECCFF91B" pageId="4" pageNumber="307" type="materials_examined">
<paragraph id="CE0436E0FFCCFFC3216C103BEBEAFC5B" blockId="4.[835,1474,861,949]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C103BEBCFFC9B" bold="true" box="[835,1012,861,885]" pageId="4" pageNumber="307">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFCCFFC3262F103BEBEAFC5B" collectingDate="2019-04-16" collectionCode="LKCNHM" country="Singapore" location="Pulau Ubin" municipality="Sungei Tock Kew" pageId="4" pageNumber="307" specimenCode="PU22" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="11008842FFCCFFC3262F103BEC5CFC9B" box="[1024,1127,861,885]" pageId="4" pageNumber="307" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFCCFFC32696103BED17FC9B" box="[1209,1324,861,885]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
,
<location id="CB64603BFFCCFFC32714103BED86FC9B" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC32714103BED86FC9B" box="[1339,1469,861,885]" country="Singapore" municipality="Sungei Tock Kew" name="Pulau Ubin" pageId="4" pageNumber="307">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFCCFFC3216C101BEC33FC7B" box="[835,1032,893,917]" pageId="4" pageNumber="307">Sungei Tock Kew</collectingMunicipality>
,
<specimenCount id="D8BDFD69FFCCFFC3263C101BEC65FC7B" box="[1043,1118,893,917]" count="1" pageId="4" pageNumber="307" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCCFFC32645101BECEEFC7B" box="[1130,1237,893,917]" count="3" pageId="4" pageNumber="307" type="female">3 females</specimenCount>
,
<date id="BA051020FFCCFFC326CF101BED41FC7B" box="[1248,1402,893,917]" pageId="4" pageNumber="307" value="2019-04-16">
<collectingDate id="AA41E9C8FFCCFFC326CF101BED41FC7B" box="[1248,1402,893,917]" pageId="4" pageNumber="307" value="2019-04-16">16 April 2019</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFCCFFC327AA101BEDF9FC7B" box="[1413,1474,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU22</specimenCode>
(
<collectionCode id="A8AAAE25FFCCFFC3216410FBEBFEFC5B" box="[843,965,925,949]" pageId="4" pageNumber="307">LKCNHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFCCFFC3216C10BBED96FBDB" blockId="4.[835,1474,989,1077]" pageId="4" pageNumber="307">
<materialsCitation id="7ED33CBDFFCCFFC3216C10BBED93FBDB" collectingDate="2018-03-19" collectingDateMax="2019-04-16" collectingDateMin="2018-03-19" collectionCode="PU, ZRCBDP" country="Singapore" location="Pulau Ubin" municipality="Sungei Tock Kew" pageId="4" pageNumber="307" specimenCode="PU22, ZRCBDP0317766" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="FCCFEAF2FFCCFFC3216C10BBEBF8FC1B" bold="true" box="[835,963,989,1013]" pageId="4" pageNumber="307">
<typeStatus id="11008842FFCCFFC3216C10BBEB85FC1B" box="[835,958,989,1013]" pageId="4" pageNumber="307" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFCCFFC321E010BBEC7FFC1B" box="[975,1092,989,1013]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
,
<location id="CB64603BFFCCFFC3267B10BBECE1FC1B" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC3267B10BBECE1FC1B" box="[1108,1242,989,1013]" country="Singapore" municipality="Sungei Tock Kew" name="Pulau Ubin" pageId="4" pageNumber="307">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFCCFFC326C510BBED85FC1B" box="[1258,1470,989,1013]" pageId="4" pageNumber="307">Sungei Tock Kew</collectingMunicipality>
,
<specimenCount id="D8BDFD69FFCCFFC3216C109BEB88FBFB" box="[835,947,1021,1045]" count="3" pageId="4" pageNumber="307" type="female">3 females</specimenCount>
,
<date id="BA051020FFCCFFC321EC109BEC5DFBFB" box="[963,1126,1021,1045]" pageId="4" pageNumber="307" value="2019-04-16">
<collectingDate id="AA41E9C8FFCCFFC321EC109BEC5DFBFB" box="[963,1126,1021,1045]" pageId="4" pageNumber="307" value="2019-04-16">16 April 2019</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFCCFFC3265A109BEC8CFBFB" box="[1141,1207,1021,1045]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU22</specimenCode>
; male,
<date id="BA051020FFCCFFC32722109BED85FBFB" box="[1293,1470,1021,1045]" pageId="4" pageNumber="307" value="2018-03-19">
<collectingDate id="AA41E9C8FFCCFFC32722109BED85FBFB" box="[1293,1470,1021,1045]" pageId="4" pageNumber="307" value="2018-03-19">19 March 2018</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFCCFFC3216B177BEC36FBDB" box="[836,1037,1053,1077]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0317766</specimenCode>
: GenBank accession code: PP893290
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFCCFFC3216C173BED81F97B" blockId="4.[834,1476,1117,1685]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C173BEBCDFB9B" bold="true" box="[835,1014,1117,1141]" pageId="4" pageNumber="307">Other material.</emphasis>
<materialsCitation id="7ED33CBDFFCCFFC321D2173BED8DF97B" collectingDate="2012-04-20" collectingDateMax="2019-05-08" collectingDateMin="2012-04-20" collectionCode="PU, ZRCBDP, IDOM, MIPD, SMO, SM, KM" county="GenBank" location="Sarimbun" municipality="Jayanthi" pageId="4" pageNumber="307" specimenCode="PU1, PU2, PU1-2013, PU01-2016, PU01-2019, PU02-2016, PU09, PU10, PU13, PU19, PU20, PU21, PU22, PU23, PU25, PU26, ZRCBDP0084405, IDOM13, PU01, ZRCBDP0257018, MIPD19, SMO2, SM02, 1, SM01, 5, SM03, ZRCBDP0041305, ZRCBDP0084993, IDOM20, ZRCBDP0329050, KM04" specimenCount="215" specimenCount-female="12" specimenCount-male="3">
Pulau Ubin, Chek Jawa,
<specimenCount id="D8BDFD69FFCCFFC32727173BED69FB9B" box="[1288,1362,1117,1141]" count="1" pageId="4" pageNumber="307" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCCFFC32771173BED85FB9B" box="[1374,1470,1117,1141]" count="1" pageId="4" pageNumber="307" type="female">1 female</specimenCount>
,
<date id="BA051020FFCCFFC3216B171BEBEBFB7B" box="[836,976,1149,1173]" pageId="4" pageNumber="307" value="2012-06-16">
<collectingDate id="AA41E9C8FFCCFFC3216B171BEBEBFB7B" box="[836,976,1149,1173]" pageId="4" pageNumber="307" value="2012-06-16">16 June 2012</collectingDate>
</date>
(
<specimenCode id="9E1D9E9BFFCCFFC321F0171BEC2AFB7B" box="[991,1041,1149,1173]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU1</specimenCode>
); Chek Jawa,
<specimenCount id="D8BDFD69FFCCFFC3268B171BECD6FB7B" box="[1188,1261,1149,1173]" count="1" pageId="4" pageNumber="307" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCCFFC326D7171BED5AFB7B" box="[1272,1377,1149,1173]" count="4" pageId="4" pageNumber="307" type="female">4 females</specimenCount>
,
<date id="BA051020FFCCFFC32745171BEB43FB5B" pageId="4" pageNumber="307" value="2012-04-20">
<collectingDate id="AA41E9C8FFCCFFC32745171BEB43FB5B" pageId="4" pageNumber="307" value="2012-04-20">20 April 2012</collectingDate>
</date>
(
<specimenCode id="9E1D9E9BFFCCFFC321A717FBEB86FB5B" box="[904,957,1181,1205]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU2</specimenCode>
). More than
<specimenCount id="D8BDFD69FFCCFFC3266A17FBECDFFB5B" box="[1093,1252,1181,1205]" count="200" pageId="4" pageNumber="307" type="generic">200 specimens</specimenCount>
were barcoded from Pulau Ubin:
<specimenCode id="9E1D9E9BFFCCFFC321EA17DBEC08FB3B" box="[965,1075,1213,1237]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU1-2013</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3261317DBEC8CFB3B" box="[1084,1207,1213,1237]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU01-2016</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC326EF17DBED00FB3B" box="[1216,1339,1213,1237]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU01-2019</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3276B17DBED84FB3B" box="[1348,1471,1213,1237]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU02-2016</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3216D17BBEBBFFB1B" box="[834,900,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU09</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC321BC17BBEBEEFB1B" box="[915,981,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU10</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC321CB17BBEC1DFB1B" box="[996,1062,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU13</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3261A17BBEC4CFB1B" box="[1077,1143,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU19</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC326A817BBECF3FB1B" box="[1159,1224,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU20</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC326F717BBED21FB1B" box="[1240,1306,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU21</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3270617BBED50FB1B" box="[1321,1387,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU22</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3275517BBED87FB1B" box="[1402,1468,1245,1269]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU23</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3216C179BEBBDFAFB" box="[835,902,1277,1301]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU25</specimenCode>
, and
<specimenCode id="9E1D9E9BFFCCFFC321E6179BEC37FAFB" box="[969,1036,1277,1301]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU26</specimenCode>
. Female,
<specimenCode id="9E1D9E9BFFCCFFC326AF179BED6BFAFB" box="[1152,1360,1277,1301]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0084405</specimenCode>
_YuTing_
<specimenCode id="9E1D9E9BFFCCFFC3216C167BEB90FADB" box="[835,939,1309,1333]" collectionCode="IDOM" pageId="4" pageNumber="307">IDOM13</specimenCode>
_
<specimenCode id="9E1D9E9BFFCCFFC32195167BEBC1FADB" box="[954,1018,1309,1333]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU01</specimenCode>
_
<date id="BA051020FFCCFFC32626167BECBCFADB" box="[1033,1159,1309,1333]" pageId="4" pageNumber="307" value="2016-06-23">
<collectingDate id="AA41E9C8FFCCFFC32626167BECBCFADB" box="[1033,1159,1309,1333]" pageId="4" pageNumber="307" value="2016-06-23">23Jun2016</collectingDate>
</date>
: GenBank accession code: PP893291;
<specimenCode id="9E1D9E9BFFCCFFC321E9165BECB4FABB" box="[966,1167,1341,1365]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0257018</specimenCode>
_Nabilah-Kareen_
<specimenCode id="9E1D9E9BFFCCFFC32776165BED8EFABB" box="[1369,1461,1341,1365]" collectionCode="MIPD" pageId="4" pageNumber="307">MIPD19</specimenCode>
_
<specimenCode id="9E1D9E9BFFCCFFC3216C163BEBB8FA9B" box="[835,899,1373,1397]" collectionCode="PU" country="USA" name="Princeton University" pageId="4" pageNumber="307">PU20</specimenCode>
_
<date id="BA051020FFCCFFC321BE163BEC29FA9B" box="[913,1042,1373,1397]" pageId="4" pageNumber="307" value="2018-03-29">
<collectingDate id="AA41E9C8FFCCFFC321BE163BEC29FA9B" box="[913,1042,1373,1397]" pageId="4" pageNumber="307" value="2018-03-29">29Mar2018</collectingDate>
</date>
. GenBank accession code: PP893292; Pulau Semakau, old mangroves,
<specimenCount id="D8BDFD69FFCCFFC32683161BECC3FA7B" box="[1196,1272,1405,1429]" count="1" pageId="4" pageNumber="307" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCCFFC327291618ED49FA7B" box="[1286,1394,1405,1429]" count="7" pageId="4" pageNumber="307" type="female">7 females</specimenCount>
,
<date id="BA051020FFCCFFC32750161BEB43FA5B" pageId="4" pageNumber="307" value="2013-07-04">
<collectingDate id="AA41E9C8FFCCFFC32750161BEB43FA5B" pageId="4" pageNumber="307" value="2013-07-04">4 July 2013</collectingDate>
</date>
(
<specimenCode id="9E1D9E9BFFCCFFC321AA16FBEBF5FA5B" box="[901,974,1437,1461]" collectionCode="SMO" pageId="4" pageNumber="307">SMO2</specimenCode>
) (Ma6107; male terminalia illustrated). 15 from
<specimenCode id="9E1D9E9BFFCCFFC3216C16DBEBBDFA3B" box="[835,902,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">SM02</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC321A016DBEBA7FA3B" box="[911,924,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">1</specimenCode>
from
<specimenCode id="9E1D9E9BFFCCFFC321F616DBEC20FA3B" box="[985,1051,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">SM01</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3260B16DBEC0AFA3B" box="[1060,1073,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">5</specimenCode>
from
<specimenCode id="9E1D9E9BFFCCFFC3264116DBEC8AFA3B" box="[1134,1201,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">SM03</specimenCode>
,
<specimenCode id="9E1D9E9BFFCCFFC3269516DBECFCFA3B" box="[1210,1223,1469,1493]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="4" pageNumber="307">1</specimenCode>
from SMN2;
<location id="CB64603BFFCCFFC3277916DBED84FA3B" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC3277916DBED84FA3B" box="[1366,1471,1469,1493]" county="GenBank" municipality="Jayanthi" name="Sarimbun" pageId="4" pageNumber="307">Sarimbun</location>
,
<specimenCode id="9E1D9E9BFFCCFFC3216C16BBEC23FA1B" box="[835,1048,1501,1525]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0041305</specimenCode>
_RP161_WTH24_SR03_
<date id="BA051020FFCCFFC3271A16BBED86FA1B" box="[1333,1469,1501,1525]" pageId="4" pageNumber="307" value="2014-05-28">
<collectingDate id="AA41E9C8FFCCFFC3271A16BBED86FA1B" box="[1333,1469,1501,1525]" pageId="4" pageNumber="307" value="2014-05-28">28May2014</collectingDate>
</date>
;
<location id="CB64603BFFCCFFC3216B169BEB89F9FB" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC3216B169BEB89F9FB" box="[836,946,1533,1557]" county="GenBank" municipality="Jayanthi" name="GenBank" pageId="4" pageNumber="307">GenBank</location>
accession code: PP893294;
<location id="CB64603BFFCCFFC32733169BEC29F9DB" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC32733169BEC29F9DB" county="GenBank" municipality="Jayanthi" name="Sungei Buloh Wetland Reserve" pageId="4" pageNumber="307">Sungei Buloh Wetland Reserve</location>
,
<specimenCode id="9E1D9E9BFFCCFFC3260C157BECCEF9DB" box="[1059,1269,1565,1589]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0084993</specimenCode>
<location id="CB64603BFFCCFFC3272E157BED04F9DB" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC3272E157BED04F9DB" box="[1281,1343,1565,1589]" county="GenBank" municipality="Jayanthi" name="Minh" pageId="4" pageNumber="307">Minh</location>
_
<specimenCode id="9E1D9E9BFFCCFFC32761157BED8DF9DB" box="[1358,1462,1565,1589]" collectionCode="IDOM" pageId="4" pageNumber="307">IDOM20</specimenCode>
_ SB02_
<date id="BA051020FFCCFFC321A1155BEC2AF9BB" box="[910,1041,1597,1621]" pageId="4" pageNumber="307" value="2016-03-14">
<collectingDate id="AA41E9C8FFCCFFC321A1155BEC2AF9BB" box="[910,1041,1597,1621]" pageId="4" pageNumber="307" value="2016-03-14">14Mar2016</collectingDate>
</date>
;
<location id="CB64603BFFCCFFC32630155BECBDF9BB" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC32630155BECBDF9BB" box="[1055,1158,1597,1621]" county="GenBank" municipality="Jayanthi" name="GenBank" pageId="4" pageNumber="307">GenBank</location>
accession code: PP893295;
<location id="CB64603BFFCCFFC3216B153BEBC0F99B" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCCFFC3216B153BEBC0F99B" box="[836,1019,1629,1653]" county="GenBank" municipality="Jayanthi" name="Kranji Marshes" pageId="4" pageNumber="307">Kranji Marshes</location>
,
<specimenCode id="9E1D9E9BFFCCFFC32625153BECE0F99B" box="[1034,1243,1629,1653]" collectionCode="ZRCBDP" pageId="4" pageNumber="307">ZRCBDP0329050</specimenCode>
_
<collectingMunicipality id="2E60AC9AFFCCFFC326C5153BED7CF99B" box="[1258,1351,1629,1653]" pageId="4" pageNumber="307">Jayanthi</collectingMunicipality>
_MTJ024_
<specimenCode id="9E1D9E9BFFCCFFC3216C151BEBB1F97B" box="[835,906,1661,1685]" collectionCode="KM" country="Russia" name="Kotel'nich Museum" pageId="4" pageNumber="307">KM04</specimenCode>
_
<date id="BA051020FFCCFFC321B7151BEC20F97B" box="[920,1051,1661,1685]" pageId="4" pageNumber="307" value="2019-05-08">
<collectingDate id="AA41E9C8FFCCFFC321B7151BEC20F97B" box="[920,1051,1661,1685]" pageId="4" pageNumber="307" value="2019-05-08">08May2019</collectingDate>
</date>
.
<collectingCounty id="27654E6CFFCCFFC32609151BECB0F97B" box="[1062,1163,1661,1685]" pageId="4" pageNumber="307">GenBank</collectingCounty>
accession code: PP893296
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFCCFFC3216C15D8ECCFF91B" blockId="4.[835,1474,1725,1781]" pageId="4" pageNumber="307">
In total,
<specimenCount id="D8BDFD69FFCCFFC3218A15DBEC7CF93B" box="[933,1095,1725,1749]" count="225" pageId="4" pageNumber="307" type="generic">225 specimens</specimenCount>
were barcoded and all specimens had less than 1% difference in barcode.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC3216C147BED2FF7DB" pageId="4" pageNumber="307" type="diagnosis">
<paragraph id="CE0436E0FFCCFFC3216C147BED2FF7DB" blockId="4.[834,1475,1821,2101]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C147BEB83F8DB" bold="true" box="[835,952,1821,1845]" pageId="4" pageNumber="307">Diagnosis.</emphasis>
A small species (
<quantity id="09439B05FFCCFFC3265B147BECC1F8DB" box="[1140,1274,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="4" pageNumber="307" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.82.2 mm</quantity>
). Male postpedicel trapezoid, a little longer than high, with a distinct ventral projection. Females with a shorter trapezoid postpedicel, a little shorter than high, also with a short ventral projection. Frons and face broad in both sexes, parallel-sided, twice as wide as postpedicel. Legs yellow, but mid and hind coxae brown, apical 23 tarsomeres of all legs brownish (somewhat variable in males, in females only apical tarsomere brown). Sternite 8 with a pair of weak macrosetae.
</paragraph>
</subSubSection>
<caption id="9AC46668FFCDFFC2225E1747E93BFBBA" ID-DOI="http://doi.org/10.5281/zenodo.14683066" ID-Zenodo-Dep="14683066" httpUri="https://zenodo.org/record/14683066/files/figure.png" pageId="5" pageNumber="308" startId="5.[113,148,1057,1079]" targetBox="[128,1444,189,1033]" targetPageId="5" targetType="figure">
<paragraph id="CE0436E0FFCDFFC2225E1747E93BFBBA" blockId="5.[113,1451,1057,1108]" pageId="5" pageNumber="308">
Fig. 4.
<taxonomicName id="09BB4D63FFCDFFC222951747E944FBD8" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[186,383,1057,1078]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFCDFFC222951747E944FBD8" box="[186,383,1057,1078]" italics="true" pageId="5" pageNumber="308">Chrysotus ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCDFFC223A11747EA3DFBD9" box="[398,518,1057,1079]" pageId="5" pageNumber="308" rank="species">new species</taxonomicNameLabel>
. A, Habitus paratype male (Ma6107). B, Paratype female (Ma6107). Photographs by Jayanthi Puniamoorthy.
</paragraph>
</caption>
<subSubSection id="86A1656BFFCDFFC2225E17EDEDB3F94C" pageId="5" pageNumber="308" type="etymology">
<paragraph id="CE0436E0FFCDFFC2225E17EDEA71FB2D" blockId="5.[113,751,1163,1219]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E17EDE8CEFB4D" bold="true" box="[113,245,1163,1187]" pageId="5" pageNumber="308">Etymology.</emphasis>
The species is named after the island, Pulau Ubin, where it was found for the first time.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E178DEA65FAEC" blockId="5.[113,606,1259,1283]" box="[113,606,1259,1283]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E178DE889FAED" bold="true" box="[113,178,1259,1283]" pageId="5" pageNumber="308">Male.</emphasis>
Length: body
<quantity id="09439B05FFCDFFC2237B178DE990FAEC" box="[340,427,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="308" unit="mm" value="1.8">1.8 mm</quantity>
; wing
<quantity id="09439B05FFCDFFC223D9178DEA61FAEC" box="[502,602,1259,1283]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="5" pageNumber="308" unit="mm" value="1.75">1.75 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E164DE9E4FA8D" blockId="5.[113,752,1323,1379]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E164DE88EFAAD" bold="true" box="[113,181,1323,1347]" pageId="5" pageNumber="308">Head.</emphasis>
Frons and face grey dusted, both parallel-sided, wide, two times as wide as postpedicel.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E16EDE9B1F9AD" blockId="5.[113,753,1419,1603]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E16EDE8DDFA4D" bold="true" box="[113,230,1419,1443]" pageId="5" pageNumber="308">Antennae.</emphasis>
Black. Scape as long as pedicel. Pedicel lacking a long dorsal bristle. Postpedicel trapezoid, a little longer than high, with a distinct ventral projection. Vertical bristle short, ¼ length of ocellar bristle (verticals much longer in female). Palpus yellow, apical bristle not distinct from the other small black bristles.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E150DE9ECF8CD" blockId="5.[112,753,1643,1827]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E150DE8F7F96D" bold="true" box="[113,204,1643,1667]" pageId="5" pageNumber="308">Thorax.</emphasis>
Mesoscutum metallic green, grey dusted. All bristles pale brownish. 45 pairs of short acrostichal bristles; 6 dorsocentral bristles, anterior most very short, the following dc becoming longer toward scutellum, prescutellar longest and outside the row. A pair of very long marginal scutellar bristles, no bristle at the outside.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E142DEAFDF86D" blockId="5.[113,753,1867,1923]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E142DE895F88D" bold="true" box="[113,174,1867,1891]" pageId="5" pageNumber="308">Legs.</emphasis>
Yellow, except for brown mid and hind coxae and apical 23 tarsomeres of all legs brownish (not black).
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E14CDEAD4F80D" blockId="5.[112,752,1963,2019]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E14CDE881F82D" bold="true" box="[113,186,1963,1987]" pageId="5" pageNumber="308">Wings.</emphasis>
With a greyish tinge, veins yellowish brown. Squama yellowish with a row of 5 pale brown bristles. Haltere white.
</paragraph>
<paragraph id="CE0436E0FFCDFFC2225E1B6DECA9FAED" blockId="5.[112,753,2059,2115]" lastBlockId="5.[813,1451,1163,1283]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2225E1B6DE8DEF7CD" bold="true" box="[113,229,2059,2083]" pageId="5" pageNumber="308">Abdomen.</emphasis>
Brown in ground-colour with metallic green shine. Sternites with a few long, fine bristles. Male terminalia (
<figureCitation id="56802A65FFCDFFC220EC1B4DEB07FB4D" box="[707,828,1163,2115]" captionStart="Fig" captionStartId="6.[136,171,1104,1126]" captionTargetBox="[138,1472,189,1092]" captionTargetId="figure-8@6.[126,1491,174,1104]" captionTargetPageId="6" captionText="Fig. 5. Chrysotus ubinensis, new species, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="5" pageNumber="308">Fig. 5</figureCitation>
). Eighth sternite with a pair of weak macrosetae. Cercus yellowish, ventral surstylus brown with a black apical spine (
<figureCitation id="56802A65FFCDFFC2211A17ADEB8BFB0D" box="[821,944,1227,1251]" captionStart="Fig" captionStartId="6.[136,171,1104,1126]" captionTargetBox="[138,1472,189,1092]" captionTargetId="figure-8@6.[126,1491,174,1104]" captionTargetPageId="6" captionText="Fig. 5. Chrysotus ubinensis, new species, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="5" pageNumber="308">Fig. 5AC</figureCitation>
). Tip of phallus simple (
<figureCitation id="56802A65FFCDFFC226E717ADED18FB0D" box="[1224,1315,1227,1251]" captionStart="Fig" captionStartId="6.[136,171,1104,1126]" captionTargetBox="[138,1472,189,1092]" captionTargetId="figure-8@6.[126,1491,174,1104]" captionTargetPageId="6" captionText="Fig. 5. Chrysotus ubinensis, new species, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="5" pageNumber="308">Fig. 5A</figureCitation>
). Postgonite conical in lateral view (
<figureCitation id="56802A65FFCDFFC22600178DECBDFAED" box="[1071,1158,1259,1283]" captionStart="Fig" captionStartId="6.[136,171,1104,1126]" captionTargetBox="[138,1472,189,1092]" captionTargetId="figure-8@6.[126,1491,174,1104]" captionTargetPageId="6" captionText="Fig. 5. Chrysotus ubinensis, new species, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="5" pageNumber="308">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFCDFFC22102164DED97FAAC" blockId="5.[813,1452,1323,1347]" box="[813,1452,1323,1347]" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC22102164DEBBCFAAD" bold="true" box="[813,903,1323,1347]" pageId="5" pageNumber="308">Female.</emphasis>
Length: body
<quantity id="09439B05FFCDFFC2260B164DECF9FAAC" box="[1060,1218,1323,1347]" metricMagnitude="-3" metricUnit="m" metricValue="1.97" metricValueMax="2.18" metricValueMin="1.76" pageId="5" pageNumber="308" unit="mm" value="1.97" valueMax="2.18" valueMin="1.76">1.762.18 mm</quantity>
; wing
<quantity id="09439B05FFCDFFC22726164DED93FAAC" box="[1289,1448,1323,1347]" metricMagnitude="-3" metricUnit="m" metricValue="1.615" metricValueMax="1.95" metricValueMin="1.28" pageId="5" pageNumber="308" unit="mm" value="1.615" valueMax="1.95" valueMin="1.28">1.281.95 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCDFFC22102160DEDB3F94C" blockId="5.[811,1453,1387,1699]" pageId="5" pageNumber="308">Face as wide as frons, both parallel-sided and two times as wide as postpedicel. Clypeus protruding. Proboscis much stronger than in males, palpus also larger than in males, yellow with black bristles with a distinct apical bristle. Postpedicel in females also trapezoid, a little shorter than high, with a shorter ventral projection. Vertical bristle longer than in males, proclinate and half as long the brown retroclinate ocellar. A postvertical bristle, upper postoculars uniserial brown and becoming pale and longer below. Apical tarsomere of all legs brownish in contrast to the other tarsomeres.</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCDFFC1210215ADED64F95C" lastPageId="6" lastPageNumber="309" pageId="5" pageNumber="308" type="discussion">
<paragraph id="CE0436E0FFCDFFC1210215ADEB33FA5C" blockId="5.[811,1453,1739,2083]" lastBlockId="6.[136,776,1210,1458]" lastPageId="6" lastPageNumber="309" pageId="5" pageNumber="308">
<emphasis id="FCCFEAF2FFCDFFC2210215ADEB94F90D" bold="true" box="[813,943,1739,1763]" pageId="5" pageNumber="308">Comments.</emphasis>
<taxonomicName id="09BB4D63FFCDFFC2219815ADECAAF90D" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[951,1169,1739,1763]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="308" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFCDFFC2219815ADECAAF90D" box="[951,1169,1739,1763]" italics="true" pageId="5" pageNumber="308">Chrysotus ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCDFFC2268E15ADED1DF90D" box="[1185,1318,1739,1763]" pageId="5" pageNumber="308" rank="species">new species</taxonomicNameLabel>
, is part of a group of six cryptic species in
<collectingCountry id="B6AC7670FFCDFFC2265E158DECE6F8ED" box="[1137,1245,1771,1795]" name="Singapore" pageId="5" pageNumber="308">Singapore</collectingCountry>
that are recognised thanks to the barcoding differing from at least 6.3% to more than 10%. They all have yellow legs, with only the mid and hind coxae brownish, as well as the apical tarsomere that is brownish and, in some species, contrastingly black. All species of this group have a more or less triangular third antennal segment that in the new species is rather large, almost trapezoidal in males with a distinct ventral projection. The third antennal segment is smaller in females and the ventral projection is shorter. In the other species, the third antennal segment is smaller and a ventral projection is indistinct. This means that identification on morphological grounds is difficult and barcoding is necessary, especially because females are more similar than males. On Pulau Ubin, three other species—in addition to the new species—were also found and are very rare. All were females and two species were only found in the secondary forests on Pulau Ubin, Nee Soon, Bukit Timah Nature Reserve, and Kranji Marshes.
</paragraph>
<caption id="9AC46668FFCEFFC122A71736EBE4FB6C" ID-DOI="http://doi.org/10.5281/zenodo.14683068" ID-Zenodo-Dep="14683068" httpUri="https://zenodo.org/record/14683068/files/figure.png" pageId="6" pageNumber="309" startId="6.[136,171,1104,1126]" targetBox="[138,1472,189,1092]" targetPageId="6" targetType="figure">
<paragraph id="CE0436E0FFCEFFC122A71736EBE4FB6C" blockId="6.[136,1474,1104,1155]" pageId="6" pageNumber="309">
Fig. 5.
<taxonomicName id="09BB4D63FFCEFFC122FD1736E9A3FB8B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[210,408,1104,1125]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="309" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFCEFFC122FD1736E9A3FB8B" box="[210,408,1104,1125]" italics="true" pageId="6" pageNumber="309">Chrysotus ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCEFFC123881736EA1AFB88" box="[423,545,1104,1126]" pageId="6" pageNumber="309" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Hypopygium lateral; B, Hypopygium ventral; C, Epandrial lobe and tip ventral surstylus lateral; D, postgonite. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFCEFFC122A716BCEC8AFB1C" blockId="6.[134,778,1498,2098]" lastBlockId="6.[835,1473,1210,1266]" pageId="6" pageNumber="309">
<emphasis id="FCCFEAF2FFCEFFC122A716BCE9B6FA1C" bold="true" box="[136,397,1498,1522]" pageId="6" pageNumber="309">National distribution.</emphasis>
<materialsCitation id="7ED33CBDFFCEFFC123B716BCEC96FB1C" collectionCode="SMO, SMN" country="Singapore" county="The" location="Although" municipality="It" pageId="6" pageNumber="309" specimenCount="200">
Confirmed by the 225 available barcodes,
<taxonomicName id="09BB4D63FFCEFFC122DC169CE94DF9FC" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[243,374,1530,1554]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="309" phylum="Arthropoda" rank="species" species="ubinensis" status="new species">
<emphasis id="FCCFEAF2FFCEFFC122DC169CE94DF9FC" box="[243,374,1530,1554]" italics="true" pageId="6" pageNumber="309">C. ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCEFFC123AE169CEA38F9FC" box="[385,515,1530,1554]" pageId="6" pageNumber="309" rank="species">new species</taxonomicNameLabel>
, is very common in back mangroves on Pulau Ubin (nearly
<specimenCount id="D8BDFD69FFCEFFC123D3157CEAA7F9DC" box="[508,668,1562,1586]" count="200" pageId="6" pageNumber="309" type="generic">200 specimens</specimenCount>
barcoded from 14 of the 26
<collectingMethod id="17FA4EF7FFCEFFC1237E155CEB33F9BF" box="[337,776,1594,1618]" pageId="6" pageNumber="309">stations ever sampled with Malaise traps</collectingMethod>
all over the island). The species is also common in the old back mangroves of Pulau Semakau (
<collectionCode id="A8AAAE25FFCEFFC12066151CEABDF97C" box="[585,646,1658,1682]" pageId="6" pageNumber="309">SMO</collectionCode>
) with more than 20 barcoded specimens, while there was only a single record from the replanted mangroves (
<collectionCode id="A8AAAE25FFCEFFC1200B15DCEA58F93C" box="[548,611,1722,1746]" country="China" lsid="urn:lsid:biocol.org:col:14058" name="Simao District National Medical and Pharmaceutical Institute" pageId="6" pageNumber="309" type="Herbarium">SMN</collectionCode>
), which did not have established back mangroves during the two-year study (20122014).
<location id="CB64603BFFCEFFC12335159CE946F8FC" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCEFFC12335159CE946F8FC" box="[282,381,1786,1810]" country="Singapore" county="The" municipality="It" name="Although" pageId="6" pageNumber="309">Although</location>
many other mangroves were surveyed in
<collectingCountry id="B6AC7670FFCEFFC1228E147CE936F8DC" box="[161,269,1818,1842]" name="Singapore" pageId="6" pageNumber="309">Singapore</collectingCountry>
during the last 15 years, the new species is very rare in other sites; there is a single record from
<location id="CB64603BFFCEFFC120B8145CEB3AF8BC" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCEFFC120B8145CEB3AF8BC" box="[663,769,1850,1874]" country="Singapore" county="The" municipality="It" name="Sarimbun" pageId="6" pageNumber="309">Sarimbun</location>
, a single record from
<location id="CB64603BFFCEFFC1235A143CEAE9F89F" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCEFFC1235A143CEAE9F89F" box="[373,722,1882,1906]" country="Singapore" county="The" municipality="It" name="Sungei Buloh Wetland Reserve" pageId="6" pageNumber="309">Sungei Buloh Wetland Reserve</location>
, and a single record from
<location id="CB64603BFFCEFFC12342141CEA2CF87C" LSID="urn:lsid:plazi:treatment:461287F6FFCCFFC121E7119BED64F95C:CB64603BFFCEFFC12342141CEA2CF87C" box="[365,535,1914,1938]" country="Singapore" county="The" municipality="It" name="Kranji Marshes" pageId="6" pageNumber="309">Kranji Marshes</location>
, which were formerly mangroves but has turned into a freshwater swamp forest.
<collectingMunicipality id="2E60AC9AFFCEFFC122A714DCE8A3F83F" box="[136,152,1978,2001]" pageId="6" pageNumber="309">It</collectingMunicipality>
was also the only record of the species that is not a true back mangrove species. A noteworthy observation is that the haplotypes in the new species do not vary much (less than 0.3%) over all the mangroves in
<collectingCountry id="B6AC7670FFCEFFC120031B7CEAA7F7DC" box="[556,668,2074,2098]" name="Singapore" pageId="6" pageNumber="309">Singapore</collectingCountry>
.
<collectingCounty id="27654E6CFFCEFFC120871B7CEAEAF7DC" box="[680,721,2074,2098]" pageId="6" pageNumber="309">The</collectingCounty>
new species was not yet recognised outside of
<collectingCountry id="B6AC7670FFCEFFC1271F17DCED9BFB3C" box="[1328,1440,1210,1234]" name="Singapore" pageId="6" pageNumber="309">Singapore</collectingCountry>
as morpho-species nor as haplotype
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFCEFFC1216C167CED64F95C" blockId="6.[835,1475,1305,1714]" pageId="6" pageNumber="309">
<emphasis id="FCCFEAF2FFCEFFC1216C167CECFAFADC" bold="true" box="[835,1217,1305,1330]" pageId="6" pageNumber="309">
General comments on
<taxonomicName id="09BB4D63FFCEFFC12664167FEC81FADF" authorityName="Meigen" authorityYear="1824" box="[1099,1210,1305,1329]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCEFFC12664167FEC81FADF" bold="true" box="[1099,1210,1305,1329]" italics="true" pageId="6" pageNumber="309">Chrysotus</emphasis>
</taxonomicName>
.
</emphasis>
The barcoding showed that there are several cryptic species of
<taxonomicName id="09BB4D63FFCEFFC126D0165CED50FABC" authorityName="Meigen" authorityYear="1824" box="[1279,1387,1338,1362]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCEFFC126D0165CED50FABC" box="[1279,1387,1338,1362]" italics="true" pageId="6" pageNumber="309">Chrysotus</emphasis>
</taxonomicName>
present in the same samples. There were at least 10 cryptic species hidden in a
<taxonomicName id="09BB4D63FFCEFFC121FB161CEC7BFA7C" authorityName="Meigen" authorityYear="1824" box="[980,1088,1402,1426]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCEFFC121FB161CEC7BFA7C" box="[980,1088,1402,1426]" italics="true" pageId="6" pageNumber="309">Chrysotus</emphasis>
</taxonomicName>
with black legs, of which the males are morphologically difficult to distinguish. A thorough study of the male terminalia of this group is needed to see whether they belong to species already described (
<bibRefCitation id="AA2A4B11FFCEFFC127A216BCEB44F9FC" author="Wei L" pageId="6" pageNumber="309" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei, 2018</bibRefCitation>
). Association of females, representing 6080% of the population, to males is generally very difficult because firstly, females were generally not described and, secondly, they are very similar in morphology. Genetic barcoding is mandatory to provide reliable identifications so that accurate data will become available for ecological studies.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,375 @@
<document id="9A386E6D4B126C4B5FB4A5D6BC5A4D90" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFCFFFCE22D2152EEAA6F81B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Trigonocera pilifer Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="312" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="310" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="7551282429CAF90BDB5BC3BDA4B97B37" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A197A9AF4078B2B108B173F66D5EBC19">
<mods:title id="193D22311A897D710B248F8C5B474571">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="6BCE9FBABEB61A58DF453BC000FC9167" type="personal">
<mods:role id="8FB990DADEEB04EC82D0DDCD143626C4">
<mods:roleTerm id="B3FB5BE1729830B3CA4082F76B1F0743">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5747990C2FE3F1705FE04C6C84A142E">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="82BAB74A8A6086A2F4AD8B9109F73D4A" type="personal">
<mods:role id="C86502F54822CC77033273BDA64B1A23">
<mods:roleTerm id="372869185539E1818661044C1B7FA254">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7445BBF995876CCD6D80D7A3203C1556">Velde,</mods:namePart>
</mods:name>
<mods:name id="0C12B0B4DBF80CF8E80351AAEDA1C6E2" type="personal">
<mods:role id="6DBA1D973879A2AC9C3000A1F1A7710B">
<mods:roleTerm id="8E5C6B3E70300694578D505695E9E189">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81383836D17DA54D955189918BF853E2">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="B99AA5226EA47C21D232B1E73DCB8E68">text</mods:typeOfResource>
<mods:relatedItem id="BA95BDB08B49B9B06843824C95EFE434" type="host">
<mods:titleInfo id="AFE1F8D0B1FDD5C798CA5508508192A1">
<mods:title id="CA51D8FD788D4BFD7D359DEF6B268CE7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="41C74B9572E344F0D3A33A07177AACD2">
<mods:date id="8D92847F358F6224FE96A5F1C8AE6BCC">2024</mods:date>
<mods:detail id="BB9AB9A2926190E91AD0F4C5925CBB97" type="pubDate">
<mods:number id="CDD44FF76A9A40C9C178332F13B75285">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="ACCCC2288BA29A55E7E9BD0B97A0C30A" type="volume">
<mods:number id="8D51C7160C475B6F51EDEDC9E75702E6">72</mods:number>
</mods:detail>
<mods:extent id="9E8E04A689FDCE5136387E458ABA9A4F" unit="page">
<mods:start id="7515E9BD7692E8EB0854D4F7D65C8FDF">303</mods:start>
<mods:end id="EEA62050605B026FD66A879C34DA4A1D">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="39CA66D8BFC9B5164C964FB18E3A0BED">journal article</mods:classification>
<mods:identifier id="437186A65EEA726B3AA2BD788D41654C" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="0C3D8ADFE04B08C56C600746C804EE07" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="4E23521F442CD8A2C00B6D667E26E46C" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="6A8D12FF8C1EC10BA94BCCCFB481D617" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFCFFFCE22D2152EEAA6F81B" ID-DOI="http://doi.org/10.5281/zenodo.14683090" ID-Zenodo-Dep="14683090" LSID="urn:lsid:plazi:treatment:461287F6FFCFFFCE22D2152EEAA6F81B" httpUri="http://treatment.plazi.org/id/461287F6FFCFFFCE22D2152EEAA6F81B" lastPageId="9" lastPageNumber="312" pageId="7" pageNumber="310">
<subSubSection id="86A1656BFFCFFFC022D2152EEA5EF98E" box="[253,613,1608,1632]" pageId="7" pageNumber="310" type="nomenclature">
<paragraph id="CE0436E0FFCFFFC022D2152EEA5EF98E" blockId="7.[253,613,1608,1664]" box="[253,613,1608,1632]" pageId="7" pageNumber="310">
<heading id="954C818CFFCFFFC022D2152EEA5EF98E" bold="true" box="[253,613,1608,1632]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="310" reason="2">
<emphasis id="FCCFEAF2FFCFFFC022D2152EEA5EF98E" bold="true" box="[253,613,1608,1632]" pageId="7" pageNumber="310">
<taxonomicName id="09BB4D63FFCFFFC022D2152EE9EAF98E" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[253,465,1608,1632]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="310" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFCFFFC022D2152EE9EAF98E" bold="true" box="[253,465,1608,1632]" italics="true" pageId="7" pageNumber="310">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCFFFC023CF152EEA5EF98E" box="[480,613,1608,1632]" pageId="7" pageNumber="310" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCFFFC0235B150EE9D6F96E" box="[372,493,1640,1664]" pageId="7" pageNumber="310" type="description">
<paragraph id="CE0436E0FFCFFFC0235B150EE9D6F96E" blockId="7.[253,613,1608,1664]" box="[372,493,1640,1664]" pageId="7" pageNumber="310">
(
<figureCitation id="56802A65FFCFFFC02353150EE9F2F96E" box="[380,457,1640,1664]" captionStart="Fig" captionStartId="7.[113,148,955,977]" captionTargetBox="[113,1451,189,940]" captionTargetId="figure-15@7.[113,1451,189,940]" captionTargetPageId="7" captionText="Fig. 6. Trigonocera pilifer, new species, male habitus (Ma9841, Pulau Ubin). Photograph by Isabella Van de Velde." figureDoi="http://doi.org/10.5281/zenodo.14683070" httpUri="https://zenodo.org/record/14683070/files/figure.png" pageId="7" pageNumber="310">Figs. 6</figureCitation>
,
<figureCitation id="56802A65FFCFFFC023F8150EE9DEF96E" box="[471,485,1640,1664]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="7" pageNumber="310">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCFFFC0225E15CEEC6FFB8E" pageId="7" pageNumber="310" type="materials_examined">
<paragraph id="CE0436E0FFCFFFC0225E15CEE9DBF8EE" blockId="7.[113,753,1704,1792]" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC0225E15CEE918F92E" bold="true" box="[113,291,1704,1728]" pageId="7" pageNumber="310">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFCFFFC0230115CEE9DBF8EE" collectingDate="2018-03-29" collectionCode="LKCNHM" collectorName="Ma &amp; P. Grootaert" country="Singapore" location="Pulau Ubin" pageId="7" pageNumber="310" specimenCode="PU14" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFCFFFC0230115CEE9AEF92E" box="[302,405,1704,1728]" pageId="7" pageNumber="310" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFCFFFC023C815CEEA61F92E" box="[487,602,1704,1728]" name="Singapore" pageId="7" pageNumber="310">Singapore</collectingCountry>
,
<location id="CB64603BFFCFFFC0204615CEEAD7F92E" LSID="urn:lsid:plazi:treatment:461287F6FFCFFFCE22D2152EEAA6F81B:CB64603BFFCFFFC0204615CEEAD7F92E" box="[617,748,1704,1728]" country="Singapore" name="Pulau Ubin" pageId="7" pageNumber="310">Pulau Ubin</location>
,
<date id="BA051020FFCFFFC0225E15AEE925F90E" box="[113,286,1736,1760]" pageId="7" pageNumber="310" value="2018-03-29">
<collectingDate id="AA41E9C8FFCFFFC0225E15AEE925F90E" box="[113,286,1736,1760]" pageId="7" pageNumber="310" value="2018-03-29">29 March 2018</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFCFFFC0230315AEE957F90E" box="[300,364,1736,1760]" collectionCode="PU" country="USA" name="Princeton University" pageId="7" pageNumber="310">PU14</specimenCode>
, secondary forest (
<collectorName id="634E5336FFCFFFC0201115AEEA5FF90E" box="[574,612,1736,1760]" pageId="7" pageNumber="310">Ma</collectorName>
9841, leg.
<collectorName id="634E5336FFCFFFC020F415AFE8E0F8EE" pageId="7" pageNumber="310">P. Grootaert</collectorName>
; in ethanol
<collectionCode id="A8AAAE25FFCFFFC02373158EE9EEF8EE" box="[348,469,1768,1792]" pageId="7" pageNumber="310">LKCNHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="CE0436E0FFCFFFC0225E144EEC6FFB8E" blockId="7.[112,753,1832,2113]" lastBlockId="7.[813,1452,1032,1120]" pageId="7" pageNumber="310">
<materialsCitation id="7ED33CBDFFCFFFC0225E144EEC6BFB8E" collectingDate="2014-01-23" collectingDateMax="2018-06-07" collectingDateMin="2014-01-23" collectionCode="LKCNHM, CUGE" collectorName="Ma &amp; P. Grootaert &amp; GenBank &amp; Botanic Gardens" country="Singapore" location="Pulau Ubin" pageId="7" pageNumber="310" specimenCode="PU14, PU13, ZRCBDP0257677, ZRCBDP0281269, ZRCBDP0276806, BAP01, ZRCBDP0275497, ZRCBDP0041193, SM03, ZRCBDP0105383" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<emphasis id="FCCFEAF2FFCFFFC0225E144EE8D1F8AE" bold="true" box="[113,234,1832,1856]" pageId="7" pageNumber="310">
<typeStatus id="11008842FFCFFFC0225E144EE8DEF8AE" box="[113,229,1832,1856]" pageId="7" pageNumber="310" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFCFFFC022DF144EE964F8AE" box="[240,351,1832,1856]" name="Singapore" pageId="7" pageNumber="310">Singapore</collectingCountry>
,
<location id="CB64603BFFCFFFC02345144EE9DEF8AE" LSID="urn:lsid:plazi:treatment:461287F6FFCFFFCE22D2152EEAA6F81B:CB64603BFFCFFFC02345144EE9DEF8AE" box="[362,485,1832,1856]" country="Singapore" name="Pulau Ubin" pageId="7" pageNumber="310">Pulau Ubin</location>
,
<specimenCount id="D8BDFD69FFCFFFC023C0144EEA7FF8AE" box="[495,580,1832,1856]" count="5" pageId="7" pageNumber="310" type="male">5 males</specimenCount>
,
<date id="BA051020FFCFFFC02061144EEACBF8AE" box="[590,752,1832,1856]" pageId="7" pageNumber="310" value="2018-03-29">
<collectingDate id="AA41E9C8FFCFFFC02061144EEACBF8AE" box="[590,752,1832,1856]" pageId="7" pageNumber="310" value="2018-03-29">29 March 2018</collectingDate>
</date>
station,
<specimenCode id="9E1D9E9BFFCFFFC022ED142EE8C5F88E" box="[194,254,1864,1888]" collectionCode="PU" country="USA" name="Princeton University" pageId="7" pageNumber="310">PU14</specimenCode>
(
<collectorName id="634E5336FFCFFFC02325142EE914F88E" box="[266,303,1864,1888]" pageId="7" pageNumber="310">Ma</collectorName>
9841, leg.
<collectorName id="634E5336FFCFFFC023B5142FEA27F88E" box="[410,540,1864,1888]" pageId="7" pageNumber="310">P. Grootaert</collectorName>
;
<collectionCode id="A8AAAE25FFCFFFC0200B142EEAA0F88E" box="[548,667,1864,1888]" pageId="7" pageNumber="310">LKCNHM</collectionCode>
).
<specimenCount id="D8BDFD69FFCFFFC02084142EEACBF88E" box="[683,752,1864,1888]" count="1" pageId="7" pageNumber="310" type="male">1 male</specimenCount>
dissected and figured; female,
<specimenCode id="9E1D9E9BFFCFFFC02397140EE9CCF86E" box="[440,503,1896,1920]" collectionCode="PU" country="USA" name="Princeton University" pageId="7" pageNumber="310">PU13</specimenCode>
,
<specimenCode id="9E1D9E9BFFCFFFC0202D140EEAF1F86E" box="[514,714,1896,1920]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0257677</specimenCode>
,
<date id="BA051020FFCFFFC020FA140EE8C7F84E" pageId="7" pageNumber="310" value="2018-03-29">
<collectingDate id="AA41E9C8FFCFFFC020FA140EE8C7F84E" pageId="7" pageNumber="310" value="2018-03-29">29 March 2018</collectingDate>
</date>
;
<collectorName id="634E5336FFCFFFC0232414EEE94AF84E" box="[267,369,1928,1952]" pageId="7" pageNumber="310">GenBank</collectorName>
accession code: PP893297; male,
<specimenCode id="9E1D9E9BFFCFFFC0225D14CEE88FF82E" box="[114,180,1960,1984]" collectionCode="PU" country="USA" name="Princeton University" pageId="7" pageNumber="310">PU13</specimenCode>
,
<specimenCode id="9E1D9E9BFFCFFFC022EA14CEE9A3F82E" box="[197,408,1960,1984]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0281269</specimenCode>
,
<date id="BA051020FFCFFFC0238714CFEA06F82E" box="[424,573,1960,1984]" pageId="7" pageNumber="310" value="2018-06-07">
<collectingDate id="AA41E9C8FFCFFFC0238714CFEA06F82E" box="[424,573,1960,1984]" pageId="7" pageNumber="310" value="2018-06-07">7 June 2018</collectingDate>
</date>
; male,
<collectorName id="634E5336FFCFFFC020B714CEE8F7F80E" pageId="7" pageNumber="310">Botanic Gardens</collectorName>
(
<collectionCode id="A8AAAE25FFCFFFC022F114AEE911F80E" box="[222,298,1992,2016]" pageId="7" pageNumber="310">CUGE</collectionCode>
)
<specimenCode id="9E1D9E9BFFCFFFC0231314AEEA3EF80E" box="[316,517,1992,2016]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0276806</specimenCode>
,
<date id="BA051020FFCFFFC0203C14AEEAD6F80E" box="[531,749,1992,2016]" pageId="7" pageNumber="310" value="2017-11-10">
<collectingDate id="AA41E9C8FFCFFFC0203C14AEEAD6F80E" box="[531,749,1992,2016]" pageId="7" pageNumber="310" value="2017-11-10">10 November 2017</collectingDate>
</date>
; male, Bishan-Ang Mo Kio Park (
<specimenCode id="9E1D9E9BFFCFFFC023E6148EEA22F7EE" box="[457,537,2024,2048]" collectionCode="BAP" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13830" name="Oxford Botanic Garden" pageId="7" pageNumber="310" type="Herbarium">BAP01</specimenCode>
)
<specimenCode id="9E1D9E9BFFCFFFC02009148EEAD6F7EE" box="[550,749,2024,2048]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0275497</specimenCode>
,
<date id="BA051020FFCFFFC0225F1B6EE8C3F7CE" box="[112,248,2056,2080]" pageId="7" pageNumber="310" value="2018-06-03">
<collectingDate id="AA41E9C8FFCFFFC0225F1B6EE8C3F7CE" box="[112,248,2056,2080]" pageId="7" pageNumber="310" value="2018-06-03">3 June 2018</collectingDate>
</date>
; GenBank accession code: PP893298; female, Prince Georges Park,
<specimenCode id="9E1D9E9BFFCFFFC0235F1B4EEA05F7AE" box="[368,574,2088,2112]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0041193</specimenCode>
,
<date id="BA051020FFCFFFC020611B4EEAD0F7AE" box="[590,747,2088,2113]" pageId="7" pageNumber="310" value="2015-05-20">
<collectingDate id="AA41E9C8FFCFFFC020611B4EEAD0F7AE" box="[590,747,2088,2113]" pageId="7" pageNumber="310" value="2015-05-20">20 May 2015</collectingDate>
</date>
; GenBank accession code: PP893299; male, Pulau Semakau (
<specimenCode id="9E1D9E9BFFCFFFC0211A174EEB40FBAE" box="[821,891,1064,1088]" collectionCode="SM" country="Malaysia" name="Sarawak Museum" pageId="7" pageNumber="310">SM03</specimenCode>
)
<specimenCode id="9E1D9E9BFFCFFFC021A1174EEC64FBAE" box="[910,1119,1064,1088]" collectionCode="ZRCBDP" pageId="7" pageNumber="310">ZRCBDP0105383</specimenCode>
,
<date id="BA051020FFCFFFC02640174EED08FBAE" box="[1135,1331,1064,1089]" pageId="7" pageNumber="310" value="2014-01-23">
<collectingDate id="AA41E9C8FFCFFFC02640174EED08FBAE" box="[1135,1331,1064,1089]" pageId="7" pageNumber="310" value="2014-01-23">23 January 2014</collectingDate>
</date>
, GenBank accession code: PP893300
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCFFFC0210217EEECF5FAEE" pageId="7" pageNumber="310" type="diagnosis">
<paragraph id="CE0436E0FFCFFFC0210217EEECF5FAEE" blockId="7.[813,1452,1160,1280]" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC0210217EEEB9FFB4E" bold="true" box="[813,932,1160,1184]" pageId="7" pageNumber="310">Diagnosis.</emphasis>
A medium-sized species (
<quantity id="09439B05FFCFFFC026FC17EEED21FB4E" box="[1235,1306,1160,1184]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="310" unit="mm" value="3.0">3 mm</quantity>
). Scape with bristles above. Propleural bristle short. Fore tarsus with claws and minute pulvilli. Metepimeron and basal tergite yellowish brown, tergites 23 whitish.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCFFFCE2102164EEA05FD1B" lastPageId="9" lastPageNumber="312" pageId="7" pageNumber="310" type="etymology">
<paragraph id="CE0436E0FFCFFFC02102164EED97FA8E" blockId="7.[813,1452,1320,1377]" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC02102164EEB8AFAAE" bold="true" box="[813,945,1320,1344]" pageId="7" pageNumber="310">Etymology.</emphasis>
The new species is named pilifer after the bristles that are present on the first antennal segment (scape).
</paragraph>
<paragraph id="CE0436E0FFCFFFC0210216EEED37FA4E" blockId="7.[813,1292,1416,1440]" box="[813,1292,1416,1440]" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC0210216EEEB56FA4E" bold="true" box="[813,877,1416,1440]" pageId="7" pageNumber="310">Male.</emphasis>
Length: body
<quantity id="09439B05FFCFFFC0262016EEEC5DFA4E" box="[1039,1126,1416,1440]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="310" unit="mm" value="2.9">2.9 mm</quantity>
; wing
<quantity id="09439B05FFCFFFC0269E16EEED33FA4E" box="[1201,1288,1416,1440]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="7" pageNumber="310" unit="mm" value="2.4">2.4 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCFFFC0210216AEECEAF8CE" blockId="7.[813,1453,1480,1824]" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC0210216AEEB51FA0E" bold="true" box="[813,874,1480,1504]" pageId="7" pageNumber="310">Head</emphasis>
. Frons very wide, anteriorly nearly twice as wide as base of postpedicel. Face narrower than frons, face at base as wide as base of postpedicel, narrowing downwards. A pair of long retro- and lateroclinate ocellars, followed by a pair of short, weak proclinate basal ocellar bristles (¼ length of the anterior bristles). A pair of short postvertical bristles. A pair of strong proclinate vertical bristles nearly as long as the anterior ocellars. Upper postocular bristles short, black, and uniserial above, below only a little longer, white, and longest only near mouth. A few pale additional bristles behind the row of postoculars.
</paragraph>
<paragraph id="CE0436E0FFCFFFCF2102142EE942FACD" blockId="7.[811,1454,1864,2112]" lastBlockId="8.[136,774,1259,1315]" lastPageId="8" lastPageNumber="311" pageId="7" pageNumber="310">
<emphasis id="FCCFEAF2FFCFFFC02102142EEB9AF88E" bold="true" box="[813,929,1864,1888]" pageId="7" pageNumber="310">Antennae.</emphasis>
Dark brown (
<figureCitation id="56802A65FFCFFFC02615142EECBDF88E" box="[1082,1158,1864,1888]" captionStart="Fig" captionStartId="7.[113,148,955,977]" captionTargetBox="[113,1451,189,940]" captionTargetId="figure-15@7.[113,1451,189,940]" captionTargetPageId="7" captionText="Fig. 6. Trigonocera pilifer, new species, male habitus (Ma9841, Pulau Ubin). Photograph by Isabella Van de Velde." figureDoi="http://doi.org/10.5281/zenodo.14683070" httpUri="https://zenodo.org/record/14683070/files/figure.png" pageId="7" pageNumber="310">Figs. 6</figureCitation>
,
<figureCitation id="56802A65FFCFFFC026BC142EECECF88E" box="[1171,1239,1864,1888]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="7" pageNumber="310">7AC</figureCitation>
). Scape longer than pedicel, dorsally with a few bristles, half as long as scape. Pedicel with a dorsal bristle as long as pedicel is wide. Inner apical margin projecting inside base of postpedicel. Postpedicel triangular, nearly 2 times as long as wide at base, with the ventral margin widely overlapping the pedicel. Arista subapical, inserted a little beyond middle of dorsal margin of postpedicel. Palpus white, with a black apical bristle rather strong and half as long as palpus, in addition a few fine short hairs.
</paragraph>
<caption id="9AC46668FFC0FFCF22A717E7ED07FB5D" ID-DOI="http://doi.org/10.5281/zenodo.14683074" ID-Zenodo-Dep="14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="8" pageNumber="311" startId="8.[136,171,1153,1175]" targetBox="[136,1475,188,1138]" targetPageId="8" targetType="figure">
<paragraph id="CE0436E0FFC0FFCF22A717E7ED07FB5D" blockId="8.[136,1475,1153,1204]" pageId="8" pageNumber="311">
Fig. 7.
<taxonomicName id="09BB4D63FFC0FFCF22FF17E4E9B7FB78" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[208,396,1153,1175]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="311" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFC0FFCF22FF17E4E9B7FB78" box="[208,396,1153,1175]" italics="true" pageId="8" pageNumber="311">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC0FFCF23B617E4EA29FB79" box="[409,530,1153,1175]" pageId="8" pageNumber="311" rank="species">new species</taxonomicNameLabel>
, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC0FFCF22A7162DEA89F9ED" blockId="8.[135,777,1354,1539]" pageId="8" pageNumber="311">
<emphasis id="FCCFEAF2FFC0FFCF22A7162DE8E7FA8D" bold="true" box="[136,220,1355,1379]" pageId="8" pageNumber="311">Thorax</emphasis>
. 7 pairs of short fine acrostichals in anterior ⅔; 5 long dorsocentrals, anterior 3 equally long, posterior 2 bristles 1.5 times as long as preceding; prescutellar a little outside the row of dorsoventrals. 2 strong marginal scutellars with a tiny hair at outside. A short, fine propleural black bristle (not as prominent as in
<taxonomicName id="09BB4D63FFC0FFCF23A0168AEA2BF9ED" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[399,528,1515,1539]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="311" phylum="Arthropoda" rank="species" species="ubinensis" status="new species">
<emphasis id="FCCFEAF2FFC0FFCF23A0168AEA2BF9ED" box="[399,528,1515,1539]" italics="true" pageId="8" pageNumber="311">T. ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC0FFCF2030168AEA9DF9ED" box="[543,678,1515,1539]" pageId="8" pageNumber="311" rank="species">new species</taxonomicNameLabel>
).
</paragraph>
<paragraph id="CE0436E0FFC0FFCF22A7154DE9ACF98D" blockId="8.[136,774,1579,1635]" pageId="8" pageNumber="311">
<emphasis id="FCCFEAF2FFC0FFCF22A7154DE886F9AD" bold="true" box="[136,189,1579,1603]" pageId="8" pageNumber="311">Legs</emphasis>
. Entirely yellow, except for the brown mid and hind coxae. All bristles black.
</paragraph>
<paragraph id="CE0436E0FFC0FFCF22A715EAEA69F84D" blockId="8.[135,776,1675,1955]" pageId="8" pageNumber="311">Fore legs. Coxa with 3 long black anterior bristles in apical half, a few minute bristles, and 2 short apical bristles. Femur with 2 preapical posteroventrals as long as femur is wide; 1 short preapical posterior, a row of fine ventral bristle, 3 ventrals near middle longest, nearly half as long as femur is wide. Tibia lacking distinct bristles except from the weak bristles in the apical crown; a dense set row of fine pale bristles located ventrally, nearly as long as tibia is wide. A pair of small claws and indistinct pulvilli.</paragraph>
<paragraph id="CE0436E0FFC0FFCF22A714ADEC64FA4D" blockId="8.[136,776,1995,2115]" lastBlockId="8.[835,1474,1259,1443]" pageId="8" pageNumber="311">Mid legs. Coxa with an exterior black bristle at base as long as coxa is high and a shorter exterior near apex; anteriorly set with long bristles. Trochanter with 2 black anterior bristles, longer than trochanter is long, apical most bristle longest. Femur with a row of posteroventral bristles in apical half, as long as femur is wide. No anterior preapical bristle. Tibia with a short anterodorsal near base and 2 shorter dorsal bristles near middle; bristles in apical crown strong but not much longer than width of apex of tibia. A pair of small claws and minute pulvilli.</paragraph>
<paragraph id="CE0436E0FFC0FFCF216C16ADEDF8F94D" blockId="8.[835,1475,1483,1699]" pageId="8" pageNumber="311">Hind legs. Coxa with a long exterior at base, as long as coxa is high and 1 or 2 minute bristles near apex. Trochanter with two long anterodorsal bristles. Femur lacking an anterior preapical, but with a preapical av about as long as femur is wide; no other distinct bristles (no ventrals). Tibia with 2 short dorsal bristles in apical half, and an apical crown of rather short bristles. A pair of short claws and minute pulvilli.</paragraph>
<paragraph id="CE0436E0FFC0FFCF216C15ADEC7DF88D" blockId="8.[834,1475,1739,1891]" pageId="8" pageNumber="311">
<emphasis id="FCCFEAF2FFC0FFCF216C15ADEBB7F90D" bold="true" box="[835,908,1739,1763]" pageId="8" pageNumber="311">Wings.</emphasis>
Hyaline with a faint brownish tinge, veins brownish. Veins R
<subScript id="523F34A5FFC0FFCF21B61591EB89F8EB" attach="left" box="[921,946,1783,1797]" fontSize="6" pageId="8" pageNumber="311">4+5</subScript>
and M
<subScript id="523F34A5FFC0FFCF21D31591EC2EF8EB" attach="left" box="[1020,1045,1783,1797]" fontSize="6" pageId="8" pageNumber="311">1+2</subScript>
running parallel over entire length, M
<subScript id="523F34A5FFC0FFCF27851591EDF9F8EB" attach="left" box="[1450,1474,1783,1797]" fontSize="6" pageId="8" pageNumber="311">1+2</subScript>
straight (lacking a weak bent near middle). Squama dusky with very long black bristles, dorsal paler. Haltere white. Metepimeron brownish.
</paragraph>
<paragraph id="CE0436E0FFC0FFCE216C14EDE96DFF1B" blockId="8.[834,1475,1931,2116]" lastBlockId="9.[113,752,189,245]" lastPageId="9" lastPageNumber="312" pageId="8" pageNumber="311">
<emphasis id="FCCFEAF2FFC0FFCF216C14EDEB87F84D" bold="true" box="[835,956,1931,1955]" pageId="8" pageNumber="311">Abdomen.</emphasis>
Tergite 1 pale brown, tergites 23 yellowish white, but all 3 tergites dorsally with apical border brownish. Tergite 4 with a yellowish spot at side. All tergites with short dorsal bristles and long apical bristles, except tergite 6 which lacks bristles. Sternites 13 yellowish white; sternites 45 brown. Sternites 24 each with very short black bristles, and a pair of somewhat longer apical bristles; no apical bristles on sternite 5.
</paragraph>
<paragraph id="CE0436E0FFC1FFCE225E127BEAE2FD9B" blockId="9.[113,754,285,629]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE225E127BE90FFEDB" bold="true" box="[113,308,285,309]" pageId="9" pageNumber="312">Male terminalia.</emphasis>
<figureCitation id="56802A65FFC1FFCE2311127BE9B8FEDB" box="[318,387,285,309]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7</figureCitation>
. Cercus pale brownish including the rounded basal lobe that bears black bristles, the ventral appendage at least 2 times as long as rounded part, with a few bristles located dorsally, apical pair of bristles strongest (
<figureCitation id="56802A65FFC1FFCE225612FBE8EBFE5B" box="[121,208,413,437]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7F</figureCitation>
). Dorsal surstylus tubular, hidden by the ventral surstylus, bearing a long apical bristle. Ventral surstylus brown, long, narrowing toward tip (
<figureCitation id="56802A65FFC1FFCE23C412BBEA05FE1B" box="[491,574,477,501]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7D</figureCitation>
) with only a few minute hairs near apex. Postgonite with a rectangular bent near base and a rounded, rectangular bent near middle (
<figureCitation id="56802A65FFC1FFCE20EA117BE8A8FDBB" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7E</figureCitation>
). Epandrial lobe, elongate rectangular with a long bristle inside its base and 2 closely set apical bristles (
<figureCitation id="56802A65FFC1FFCE2056113BEAF6FD9B" box="[633,717,605,629]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7F</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFC1FFCE225E11FBEA05FD1B" blockId="9.[113,752,669,757]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE225E11FBE8F0FD5B" bold="true" box="[113,203,669,693]" pageId="9" pageNumber="312">Female.</emphasis>
Unknown. Probably not recognised because it was confused with similar-looking
<taxonomicName id="09BB4D63FFC1FFCE23EE11DBEA08FD3B" baseAuthorityName="Grichanov" baseAuthorityYear="2011" box="[449,563,701,725]" class="Insecta" family="Dolichopodidae" genus="Diaphorus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC1FFCE23EE11DBEA08FD3B" box="[449,563,701,725]" italics="true" pageId="9" pageNumber="312">Diaphorus</emphasis>
</taxonomicName>
females that also have a shorter, non-triangular postpedicel.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC1FFCE225E107BEAA6F81B" pageId="9" pageNumber="312" type="discussion">
<paragraph id="CE0436E0FFC1FFCE225E107BE9DBFB1B" blockId="9.[112,755,797,1269]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE225E107BE8CAFCDB" bold="true" box="[113,241,797,821]" pageId="9" pageNumber="312">Comments.</emphasis>
<taxonomicName id="09BB4D63FFC1FFCE22DA1078E980FCDB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[245,443,797,821]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFC1FFCE22DA1078E980FCDB" box="[245,443,797,821]" italics="true" pageId="9" pageNumber="312">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC1FFCE23E81078EA7EFCDB" box="[455,581,797,821]" pageId="9" pageNumber="312" rank="species">new species</taxonomicNameLabel>
, is quite peculiar in having distinct dorsal bristles on the scape. In the world key recently made by
<bibRefCitation id="AA2A4B11FFC1FFCE235D103BEA0AFC9B" author="Liu R &amp; Wang M &amp; Tang C &amp; Yang D" box="[370,561,861,885]" pageId="9" pageNumber="312" pagination="267 - 273" refId="ref14943" refString="Liu R, Wang M, Tang C &amp; Yang D (2018) A new species of Trigonocera (Diptera: Dolichopodidae, Diaphorinae) from China with an updated key to species worldwide. Entomotaxonomia, 40 (4): 267 - 273." type="journal article" year="2018">Liu et al. (2018)</bibRefCitation>
, the new species would run to
<taxonomicName id="09BB4D63FFC1FFCE23251018EA72FC7B" authority="Becker, 1922" authorityName="Becker" authorityYear="1922" box="[266,585,893,917]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="lucidiventris">
<emphasis id="FCCFEAF2FFC1FFCE23251018E995FC7B" box="[266,430,893,917]" italics="true" pageId="9" pageNumber="312">T. lucidiventris</emphasis>
<bibRefCitation id="AA2A4B11FFC1FFCE2397101BEA72FC7B" author="Becker T" box="[440,585,893,917]" pageId="9" pageNumber="312" pagination="1 - 247" refId="ref14398" refString="Becker T (1922) Dipterologische Studien. Dolichopodidae der Indo-Australischen Region. Capita Zoologica, 1 (4): 1 - 247." type="journal article" year="1922">
Becker,
<quantity id="09439B05FFC1FFCE203B101BEA72FC7B" box="[532,585,893,917]" metricMagnitude="1" metricUnit="m" metricValue="4.88188" pageId="9" pageNumber="312" unit="in" value="1922.0">1922</quantity>
</bibRefCitation>
</taxonomicName>
in having pale basal tergites.
<bibRefCitation id="AA2A4B11FFC1FFCE232710FBE987FC5B" author="Olejnicek J" box="[264,444,925,949]" pageId="9" pageNumber="312" pagination="167 - 173" refId="ref14708" refString="Olejnicek J (2004) A note on the genus Trigonocera Becker, 1902 with description of a new species (Insecta, Diptera, Dolichopodidae). Acta Musei Moraviae, Scientiae Biologicae (Brno), 89: 167 - 173." type="journal article" year="2004">Olejníček (2004)</bibRefCitation>
revised the
<typeStatus id="11008842FFC1FFCE201510F8EA53FC58" box="[570,616,926,950]" pageId="9" pageNumber="312">type</typeStatus>
material and made a re-description and illustrations of this species. The scape does not bear dorsal bristles (
<bibRefCitation id="AA2A4B11FFC1FFCE23DA10BBEA9DFC1B" author="Olejnicek J" box="[501,678,989,1013]" pageId="9" pageNumber="312" pagination="167 - 173" refId="ref14708" refString="Olejnicek J (2004) A note on the genus Trigonocera Becker, 1902 with description of a new species (Insecta, Diptera, Dolichopodidae). Acta Musei Moraviae, Scientiae Biologicae (Brno), 89: 167 - 173." type="journal article" year="2004">Olejníček, 2004</bibRefCitation>
; fig. 1) and the postpedicel is shorter in the new species but elongate triangular in
<taxonomicName id="09BB4D63FFC1FFCE22D11778E99BFBDB" authorityName="Becker" authorityYear="1922" box="[254,416,1053,1077]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="lucidiventris">
<emphasis id="FCCFEAF2FFC1FFCE22D11778E99BFBDB" box="[254,416,1053,1077]" italics="true" pageId="9" pageNumber="312">T. lucidiventris</emphasis>
</taxonomicName>
. The base of the postpedicel is truncate, while in the new species it has a wide ventral lobe over the pedicel (
<figureCitation id="56802A65FFC1FFCE231C173BE997FB9B" box="[307,428,1117,1141]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Fig. 7B, C</figureCitation>
). It is not clear if the pedicel has a thumb-like projection into the base of the postpedicel. The epandrial lobe is much narrower (
<bibRefCitation id="AA2A4B11FFC1FFCE202417FBEA80FB5B" author="Olejnicek J" box="[523,699,1181,1205]" pageId="9" pageNumber="312" pagination="167 - 173" refId="ref14708" refString="Olejnicek J (2004) A note on the genus Trigonocera Becker, 1902 with description of a new species (Insecta, Diptera, Dolichopodidae). Acta Musei Moraviae, Scientiae Biologicae (Brno), 89: 167 - 173." type="journal article" year="2004">Olejníček, 2004</bibRefCitation>
;
<figureCitation id="56802A65FFC1FFCE20E917FBE844FB3B" captionStart="Fig" captionStartId="5.[113,148,1057,1079]" captionTargetBox="[128,1444,189,1033]" captionTargetId="figure-482@5.[783,1452,189,1044]" captionTargetPageId="5" captionText="Fig. 4. Chrysotus ubinensis, new species. A, Habitus paratype male (Ma6107). B, Paratype female (Ma6107). Photographs by Jayanthi Puniamoorthy." figureDoi="http://doi.org/10.5281/zenodo.14683066" httpUri="https://zenodo.org/record/14683066/files/figure.png" pageId="9" pageNumber="312">Fig. 4</figureCitation>
) than in
<taxonomicName id="09BB4D63FFC1FFCE22F717D8E998FB3B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[216,419,1213,1237]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFC1FFCE22F717D8E998FB3B" box="[216,419,1213,1237]" italics="true" pageId="9" pageNumber="312">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC1FFCE239F17D8EA0AFB3B" box="[432,561,1213,1237]" pageId="9" pageNumber="312" rank="species">new species</taxonomicNameLabel>
, but unfortunately the postgonite was not illustrated.
</paragraph>
<caption id="9AC46668FFC1FFCE210217EEECC7FB54" ID-DOI="http://doi.org/10.5281/zenodo.14683076" ID-Zenodo-Dep="14683076" httpUri="https://zenodo.org/record/14683076/files/figure.png" pageId="9" pageNumber="312" startId="9.[813,852,1160,1182]" targetBox="[813,1451,189,1144]" targetPageId="9" targetType="figure">
<paragraph id="CE0436E0FFC1FFCE210217EEECC7FB54" blockId="9.[813,1450,1160,1211]" pageId="9" pageNumber="312">
Fig. 8.
<taxonomicName id="09BB4D63FFC1FFCE21AD17EEEC4CFB73" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[898,1143,1160,1181]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="ubinensis" status="sp. nov.">
<emphasis id="FCCFEAF2FFC1FFCE21AD17EEEC4CFB73" box="[898,1143,1160,1181]" italics="true" pageId="9" pageNumber="312">Trigonocera ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC1FFCE26A517EEED2BFB70" box="[1162,1296,1160,1182]" pageId="9" pageNumber="312" rank="species">new species</taxonomicNameLabel>
, male habitus (ZRCBDP0276611). Photograph by Rene Ong.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC1FFCE225E1678EAEAFA1B" blockId="9.[112,754,1309,1526]" pageId="9" pageNumber="312">
<taxonomicName id="09BB4D63FFC1FFCE225E1678E9CAFADB" authority="Becker, 1922" authorityName="Becker" authorityYear="1922" box="[113,497,1309,1333]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="specialis">
<emphasis id="FCCFEAF2FFC1FFCE225E1678E960FADB" box="[113,347,1309,1333]" italics="true" pageId="9" pageNumber="312">Trigonocera specialis</emphasis>
<bibRefCitation id="AA2A4B11FFC1FFCE234C167BE9CAFADB" author="Becker T" box="[355,497,1309,1333]" pageId="9" pageNumber="312" pagination="1 - 247" refId="ref14398" refString="Becker T (1922) Dipterologische Studien. Dolichopodidae der Indo-Australischen Region. Capita Zoologica, 1 (4): 1 - 247." type="journal article" year="1922">Becker, 1922</bibRefCitation>
</taxonomicName>
that also has pale basal tergites, has enlarged pulvilli on the fore tarsus. According to
<bibRefCitation id="AA2A4B11FFC1FFCE22BF163BE971FA9B" author="Olejnicek J" box="[144,330,1373,1397]" pageId="9" pageNumber="312" pagination="167 - 173" refId="ref14708" refString="Olejnicek J (2004) A note on the genus Trigonocera Becker, 1902 with description of a new species (Insecta, Diptera, Dolichopodidae). Acta Musei Moraviae, Scientiae Biologicae (Brno), 89: 167 - 173." type="journal article" year="2004">Olejníček (2004)</bibRefCitation>
, who also gave a re-description of this species, the scape is bare and the postpedicel is nearly 3 times as long as wide, somewhat onion-shaped at its base with a long narrow apex (
<bibRefCitation id="AA2A4B11FFC1FFCE236C16DBE9CDFA3B" author="Olejnicek J" box="[323,502,1469,1493]" pageId="9" pageNumber="312" pagination="167 - 173" refId="ref14708" refString="Olejnicek J (2004) A note on the genus Trigonocera Becker, 1902 with description of a new species (Insecta, Diptera, Dolichopodidae). Acta Musei Moraviae, Scientiae Biologicae (Brno), 89: 167 - 173." type="journal article" year="2004">Olejníček, 2004</bibRefCitation>
;
<figureCitation id="56802A65FFC1FFCE202C16DBEA7DFA3B" box="[515,582,1469,1493]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="9" pageNumber="312">Fig. 9</figureCitation>
). The epandrial lobe is very narrow bearing a pair of very long bristles.
</paragraph>
<paragraph id="CE0436E0FFC1FFCE225E157BEA12F91B" blockId="9.[113,753,1565,1781]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE225E157BE9A9F9DB" bold="true" box="[113,402,1565,1589]" pageId="9" pageNumber="312">National distribution.</emphasis>
Barcoding allowed for the identification of formerly unrecognised specimens that seem to have a wide distribution not limited to mangroves alone.
<taxonomicName id="09BB4D63FFC1FFCE225D1518E97AF97B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[114,321,1661,1685]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="species" species="pilifer" status="sp. nov.">
<emphasis id="FCCFEAF2FFC1FFCE225D1518E97AF97B" box="[114,321,1661,1685]" italics="true" pageId="9" pageNumber="312">Trigonocera pilifer</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC1FFCE237E1518E9ECF97B" box="[337,471,1661,1685]" pageId="9" pageNumber="312" rank="species">new species</taxonomicNameLabel>
, is presently known from Pulau Ubin, Pulau Semakau, and the more anthropogenic habitats at Prince Georges Park (PGP), Bishan-Ang Mo Kio Park, and the Botanic Gardens (CUGE).
</paragraph>
<paragraph id="CE0436E0FFC1FFCE225E147BEAA6F81B" blockId="9.[112,754,1821,2037]" pageId="9" pageNumber="312">
<emphasis id="FCCFEAF2FFC1FFCE225E147BE8C8F8DB" bold="true" box="[113,243,1821,1845]" pageId="9" pageNumber="312">Comments.</emphasis>
Apart from the triangular postpedicel in males and the attachment of the postpedicel on the pedicel, the differences between
<taxonomicName id="09BB4D63FFC1FFCE2372143BE9E9F89B" baseAuthorityName="Grichanov" baseAuthorityYear="2011" box="[349,466,1885,1909]" class="Insecta" family="Dolichopodidae" genus="Diaphorus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC1FFCE2372143BE9E9F89B" box="[349,466,1885,1909]" italics="true" pageId="9" pageNumber="312">Diaphorus</emphasis>
</taxonomicName>
and
<taxonomicName id="09BB4D63FFC1FFCE203F1438EAA1F89B" authorityName="Becker" authorityYear="1902" box="[528,666,1886,1909]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC1FFCE203F1438EAA1F89B" box="[528,666,1886,1909]" italics="true" pageId="9" pageNumber="312">Trigonocera</emphasis>
</taxonomicName>
remain subtle. In
<taxonomicName id="09BB4D63FFC1FFCE22FA1418E963F87B" authorityName="Becker" authorityYear="1902" box="[213,344,1918,1941]" class="Insecta" family="Dolichopodidae" genus="Trigonocera" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC1FFCE22FA1418E963F87B" box="[213,344,1918,1941]" italics="true" pageId="9" pageNumber="312">Trigonocera</emphasis>
</taxonomicName>
, the postgonite has a dorsal rectangular bent near its middle (see
<figureCitation id="56802A65FFC1FFCE235614FBE9FAF85B" box="[377,449,1949,1973]" captionStart="Fig" captionStartId="8.[136,171,1153,1175]" captionTargetBox="[136,1475,188,1138]" captionTargetId="figure-8@8.[114,1490,169,1144]" captionTargetPageId="8" captionText="Fig. 7. Trigonocera pilifer, new species, male. A, Antenna outside; B, Postpedicel outside; C, Postpedicel inside; D, Dorsal and ventral surstylus; E, Postgonite and dorsal surstylus; F, Hypopygium lateral view. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683074" httpUri="https://zenodo.org/record/14683074/files/figure.png" pageId="9" pageNumber="312">Figs. 7</figureCitation>
and
<figureCitation id="56802A65FFC1FFCE23DB14FBEA38F85B" box="[500,515,1949,1973]" captionStart="Fig" captionStartId="10.[136,171,1220,1242]" captionTargetBox="[139,1475,191,1199]" captionTargetId="figure-8@10.[112,1492,171,1202]" captionTargetPageId="10" captionText="Fig. 9. Trigonocera ubinensis, new species, male. A, Antenna outside with detail of apex postpedicel; B, Eighth sternite; C, Postgonite; D, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683078" httpUri="https://zenodo.org/record/14683078/files/figure.png" pageId="9" pageNumber="312">9</figureCitation>
), while the postgonite is generally not so strongly bent near its middle in
<taxonomicName id="09BB4D63FFC1FFCE205814DBEAD3F83B" baseAuthorityName="Grichanov" baseAuthorityYear="2011" box="[631,744,1981,2005]" class="Insecta" family="Dolichopodidae" genus="Diaphorus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="312" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC1FFCE205814DBEAD3F83B" box="[631,744,1981,2005]" italics="true" pageId="9" pageNumber="312">Diaphorus</emphasis>
</taxonomicName>
. This character, however, needs further verification.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,476 @@
<document id="E39F70D7C745E3669519CF88F277C7E0" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFD8FFD5219D1604EDF9F8DB" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Protomedetera uncinata Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="321" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="319" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BABF8BA54B54C132A94FBF101E6F0A68" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="035E8A8CC3736C49F552824331A47626">
<mods:title id="6D0488E19F4359060F29E64A6F861236">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
</mods:titleInfo>
<mods:name id="AC9385C855A09AEBD7521B16D311A29A" type="personal">
<mods:role id="83BBCF600ECF6B7F83D9DB4FB18FCFE2">
<mods:roleTerm id="5C5D208923755361DBD6950FB5D441CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DF40B4A90177158E5CB061CCF49BCE7E">Grootaert, Patrick</mods:namePart>
</mods:name>
<mods:name id="ADECBE6BACA426AFE3A5C8A69398135C" type="personal">
<mods:role id="D6A18CC3323422899935C85C71E5B87D">
<mods:roleTerm id="B0CD0FB46F8621D0275830B19DCCD461">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F7A5C957DA40B02B6B9E4FB292E6185">Velde,</mods:namePart>
</mods:name>
<mods:name id="1E8147E7EC81F910CA493C241AEE7BC6" type="personal">
<mods:role id="4E9028EBB9F49AE5D3E6545CD13227C7">
<mods:roleTerm id="6B4C618399CDFF2E47F9DB3D2562BAD1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="56165325B5B197D909357D28082895D6">Isabella Van de</mods:namePart>
</mods:name>
<mods:typeOfResource id="E73BFFD7847F613C40E4E91118E7CA96">text</mods:typeOfResource>
<mods:relatedItem id="26A8B480BA6200E11B641816F1646E05" type="host">
<mods:titleInfo id="6454C0089BF560496BCE527B7B9ACF7B">
<mods:title id="37FF690C2FF9C04AECDB3DF7ACFB884A">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="17050A30E49281C5B0841B726F895BBB">
<mods:date id="7E517B3F3E16E8D353013C95568D2BA1">2024</mods:date>
<mods:detail id="A5BFB3AC0EC9EFB6B10CA3C5F9E5D9EB" type="pubDate">
<mods:number id="516DE1308823D7203492B3827818CBF7">2024-09-19</mods:number>
</mods:detail>
<mods:detail id="9CCD03822F7F81A7464DCC31ACBEE889" type="volume">
<mods:number id="8A847C75613ED06F15D6B9C71DDF559B">72</mods:number>
</mods:detail>
<mods:extent id="BB55E9F596C82761B52EDCEB8058EF88" unit="page">
<mods:start id="EDAE0DA814FCF78C77DE0D77F86E6F28">303</mods:start>
<mods:end id="CE977C3901ECAE64A11F4E7DE91578E8">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="34184C8821FCB115AFAA8E2639A45B01">journal article</mods:classification>
<mods:identifier id="10560547A5C71B230123D61676EA1549" type="DOI">10.26107/RBZ-2024-0025</mods:identifier>
<mods:identifier id="4BF295C58352CFF31A47834019E16148" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="3D803CE0CE7E0F0D9C4156328B78D58A" type="Zenodo-Dep">14683054</mods:identifier>
<mods:identifier id="A309DF7165C12326D2DF6F513238EB5C" type="ZooBank">079CC651-8D4D-4B95-B022-388B2FA51FE2</mods:identifier>
</mods:mods>
<treatment id="461287F6FFD8FFD5219D1604EDF9F8DB" ID-DOI="http://doi.org/10.5281/zenodo.14683106" ID-Zenodo-Dep="14683106" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB" httpUri="http://treatment.plazi.org/id/461287F6FFD8FFD5219D1604EDF9F8DB" lastPageId="18" lastPageNumber="321" pageId="16" pageNumber="319">
<subSubSection id="86A1656BFFD8FFD7219D1604ED68FA95" box="[946,1363,1378,1403]" pageId="16" pageNumber="319" type="nomenclature">
<paragraph id="CE0436E0FFD8FFD7219D1604ED68FA95" blockId="16.[946,1363,1378,1435]" box="[946,1363,1378,1403]" pageId="16" pageNumber="319">
<heading id="954C818CFFD8FFD7219D1604ED68FA95" bold="true" box="[946,1363,1378,1403]" centered="true" fontSize="10" level="2" pageId="16" pageNumber="319" reason="2">
<emphasis id="FCCFEAF2FFD8FFD7219D1604ED68FA95" bold="true" box="[946,1363,1378,1403]" pageId="16" pageNumber="319">
<taxonomicName id="09BB4D63FFD8FFD7219D1604EC84FA95" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[946,1215,1378,1403]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD8FFD7219D1604EC84FA95" bold="true" box="[946,1215,1378,1403]" italics="true" pageId="16" pageNumber="319">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD8FFD726E11605ED68FA95" box="[1230,1363,1379,1403]" pageId="16" pageNumber="319" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7261516E5ECF7FA75" box="[1082,1228,1411,1435]" pageId="16" pageNumber="319" type="description">
<paragraph id="CE0436E0FFD8FFD7261516E5ECF7FA75" blockId="16.[946,1363,1378,1435]" box="[1082,1228,1411,1435]" pageId="16" pageNumber="319">
(
<figureCitation id="56802A65FFD8FFD7266D16E5ECFFFA75" box="[1090,1220,1411,1435]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="16.[838,875,1049,1071]" captionStartId-1="17.[113,149,827,849]" captionStartId-2="17.[813,848,1205,1227]" captionTargetBox-0="[836,1472,189,1034]" captionTargetBox-1="[113,749,189,811]" captionTargetBox-2="[816,1451,194,1187]" captionTargetId-0="figure-264@16.[836,1472,189,1034]" captionTargetId-1="figure-15@17.[113,749,189,811]" captionTargetId-2="figure-17@17.[798,1469,176,1197]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="17" captionText-0="Fig. 16. Protomedetera uncinata, new species, male habitus (ZRCBDP0085280). Photograph by Kristy Chang." captionText-1="Fig. 17. Protomedetera uncinata, new species, female habitus (ZRCBDP008400). Photograph by Kristy Chang." captionText-2="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi-0="http://doi.org/10.5281/zenodo.14683094" figureDoi-1="http://doi.org/10.5281/zenodo.14683098" figureDoi-2="http://doi.org/10.5281/zenodo.14683100" httpUri-0="https://zenodo.org/record/14683094/files/figure.png" httpUri-1="https://zenodo.org/record/14683098/files/figure.png" httpUri-2="https://zenodo.org/record/14683100/files/figure.png" pageId="16" pageNumber="319">Figs. 1618</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7216C16A5ED97F935" pageId="16" pageNumber="319" type="diagnosis">
<paragraph id="CE0436E0FFD8FFD7216C16A5ED97F935" blockId="16.[834,1476,1475,1755]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C16A5EB81FA35" bold="true" box="[835,954,1475,1499]" pageId="16" pageNumber="319">Diagnosis.</emphasis>
Small species with shiny metallic green head and thorax. Antenna yellow, but pedicel with almost black apex. Pedicel with 3 long marginal bristles covering the spheroid postpedicel. Anterior half uniformly covered with short bristles; 4-serial acrostichals indistinctly separated from dorsocentrals. Only the 2 prescutellar dorsocentral bristles long. All coxae and femora blackish brown, tibiae and tarsi yellow. Tip hypandrium very wide, forked, and dorsally bent. Surstylus lacking squamiform bristles. Cerci small, oval.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD7216C1465EBF7F8B5" pageId="16" pageNumber="319" type="etymology">
<paragraph id="CE0436E0FFD8FFD7216C1465EBF7F8B5" blockId="16.[835,1475,1795,1883]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C1465EBFDF8F5" bold="true" box="[835,966,1795,1819]" pageId="16" pageNumber="319">Etymology.</emphasis>
The name
<taxonomicName id="09BB4D63FFD8FFD726661462EC9CF8F5" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1097,1191,1796,1819]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="319" phylum="Arthropoda" rank="species" species="uncinata">
<emphasis id="FCCFEAF2FFD8FFD726661462EC9CF8F5" box="[1097,1191,1796,1819]" italics="true" pageId="16" pageNumber="319">uncinata</emphasis>
</taxonomicName>
(Lat. Feminine uncīnāta: bearing hooks) refers to hook-like bifurcate tip of the hypandrium.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFD8FFD5216C14E5E973FA9B" lastPageId="18" lastPageNumber="321" pageId="16" pageNumber="319" type="materials_examined">
<paragraph id="CE0436E0FFD8FFD7216C14E5EB8AF815" blockId="16.[834,1475,1923,2043]" pageId="16" pageNumber="319">
<emphasis id="FCCFEAF2FFD8FFD7216C14E5EC1BF875" bold="true" box="[835,1056,1923,1947]" pageId="16" pageNumber="319">Material examined.</emphasis>
<materialsCitation id="7ED33CBDFFD8FFD7260A14E5EB8AF815" collectingDate="2016-03-16" collectionCode="LKCNHM" collectorName="P. Grootaert &amp; GenBank" country="Singapore" location="Pulau Ubin" pageId="16" pageNumber="319" specimenCode="PU07, ZRCBDP0085280" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFD8FFD7260A14E5ECB3F875" box="[1061,1160,1923,1947]" pageId="16" pageNumber="319" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFD8FFD726E214E5ED07F875" box="[1229,1340,1923,1947]" name="Singapore" pageId="16" pageNumber="319">Singapore</collectingCountry>
,
<location id="CB64603BFFD8FFD7276B14E5ED85F875" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD8FFD7276B14E5ED85F875" box="[1348,1470,1923,1947]" country="Singapore" name="Pulau Ubin" pageId="16" pageNumber="319">Pulau Ubin</location>
, (
<specimenCode id="9E1D9E9BFFD8FFD7216614C5EBB1F855" box="[841,906,1955,1979]" collectionCode="PU" country="USA" name="Princeton University" pageId="16" pageNumber="319">PU07</specimenCode>
)
<date id="BA051020FFD8FFD721B314C5EC77F855" box="[924,1100,1955,1979]" pageId="16" pageNumber="319" value="2016-03-16">
<collectingDate id="AA41E9C8FFD8FFD721B314C5EC77F855" box="[924,1100,1955,1979]" pageId="16" pageNumber="319" value="2016-03-16">16 March 2016</collectingDate>
</date>
,
<collectingMethod id="17FA4EF7FFD8FFD7267514C5ECDEF854" box="[1114,1253,1955,1979]" pageId="16" pageNumber="319">Malaise trap</collectingMethod>
(
<specimenCode id="9E1D9E9BFFD8FFD726D814C5EDFAF855" box="[1271,1473,1955,1979]" collectionCode="ZRCBDP" pageId="16" pageNumber="319">ZRCBDP0085280</specimenCode>
in
<collectionCode id="A8AAAE25FFD8FFD7214F14A5EBECF835" box="[864,983,1987,2011]" pageId="16" pageNumber="319">LKCNHM</collectionCode>
; leg.
<collectorName id="634E5336FFD8FFD7263D14A5ECA0F835" box="[1042,1179,1987,2011]" pageId="16" pageNumber="319">P. Grootaert</collectorName>
).
<collectorName id="634E5336FFD8FFD7268214A5ED29F835" box="[1197,1298,1987,2011]" pageId="16" pageNumber="319">GenBank</collectorName>
accession code: PP893312
</materialsCitation>
</paragraph>
<caption id="9AC46668FFD9FFD6225E105DEA6FFC83" ID-DOI="http://doi.org/10.5281/zenodo.14683098" ID-Zenodo-Dep="14683098" httpUri="https://zenodo.org/record/14683098/files/figure.png" pageId="17" pageNumber="320" startId="17.[113,149,827,849]" targetBox="[113,749,189,811]" targetPageId="17" targetType="figure">
<paragraph id="CE0436E0FFD9FFD6225E105DEA6FFC83" blockId="17.[113,748,827,877]" pageId="17" pageNumber="320">
Fig. 17.
<taxonomicName id="09BB4D63FFD9FFD622E5105DE984FCBE" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[202,447,827,848]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="320" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD9FFD622E5105DE984FCBE" box="[202,447,827,848]" italics="true" pageId="17" pageNumber="320">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD9FFD623E0105DEA70FCBF" box="[463,587,827,849]" pageId="17" pageNumber="320" rank="species">new species</taxonomicNameLabel>
, female habitus (ZRCBDP008400). Photograph by Kristy Chang.
</paragraph>
</caption>
<paragraph id="CE0436E0FFD9FFD6225E10C3E912FA13" blockId="17.[113,755,933,1533]" pageId="17" pageNumber="320">
<materialsCitation id="7ED33CBDFFD9FFD6225E10C3E91EFA13" collectingDate="2016-03-16" collectingDateMax="2019-04-02" collectingDateMin="2016-03-16" collectionCode="ZRCBDP, PU" country="Singapore" county="GenBank" location="GenBank" municipality="GenBank" pageId="17" pageNumber="320" specimenCode="ZRCBDP0081888, PU01, ZRCBDP0084331, ZRCBDP0084003, ZRCBDP0085270, PU04, ZRCBDP0085236, ZRCBDP0085235, ZRCBDP0085269, ZRCBDP0085280, PU07, ZRCBDP0083285, PU10, ZRCBDP0257071, PU19, ZRCBDP0256595, ZRCBDP0317777, PU24" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFD9FFD6225E10C3E8D0FC53" bold="true" box="[113,235,933,957]" pageId="17" pageNumber="320">
<typeStatus id="11008842FFD9FFD6225E10C3E8DDFC53" box="[113,230,933,957]" pageId="17" pageNumber="320" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFD9FFD622DA10C3E9B5FC53" box="[245,398,933,957]" name="Singapore" pageId="17" pageNumber="320">SINGAPORE</collectingCountry>
, female,
<specimenCode id="9E1D9E9BFFD9FFD623DA10C3EA86FC53" box="[501,701,933,957]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0081888</specimenCode>
_12_
<specimenCode id="9E1D9E9BFFD9FFD6225E10A2E894FC32" box="[113,175,964,988]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_
<date id="BA051020FFD9FFD6229010A2E97AFC32" box="[191,321,964,988]" pageId="17" pageNumber="320" value="2016-08-10">
<collectingDate id="AA41E9C8FFD9FFD6229010A2E97AFC32" box="[191,321,964,988]" pageId="17" pageNumber="320" value="2016-08-10">10Aug2016</collectingDate>
</date>
;
<location id="CB64603BFFD9FFD6236010A3E98EFC33" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6236010A3E98EFC33" box="[335,437,965,989]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893313; female
<specimenCode id="9E1D9E9BFFD9FFD622E51082E99BFC12" box="[202,416,996,1020]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0084331</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD623811082E9D6FC12" box="[430,493,996,1020]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_
<date id="BA051020FFD9FFD623D31082EA4DFC12" box="[508,630,996,1020]" pageId="17" pageNumber="320" value="2016-06-01">
<collectingDate id="AA41E9C8FFD9FFD623D31082EA4DFC12" box="[508,630,996,1020]" pageId="17" pageNumber="320" value="2016-06-01">01Jun2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD620A81083EACBFC13" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD620A81083EACBFC13" box="[647,752,997,1021]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893314; female
<specimenCode id="9E1D9E9BFFD9FFD6203B1763EAD8FBF3" box="[532,739,1029,1053]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0084003</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E1743E889FBD3" box="[113,178,1061,1085]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU01</specimenCode>
_07
<location id="CB64603BFFD9FFD622F21743E937FBD3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD622F21743E937FBD3" box="[221,268,1061,1085]" country="Singapore" county="GenBank" municipality="GenBank" name="Apr" pageId="17" pageNumber="320">Apr</location>
201 (imaged),
<location id="CB64603BFFD9FFD623911743EA11FBD3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD623911743EA11FBD3" box="[446,554,1061,1085]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893315; female
<specimenCode id="9E1D9E9BFFD9FFD623631722EA21FBB2" box="[332,538,1092,1116]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085270</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620071722EA5CFBB2" box="[552,615,1092,1116]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD6205A1722EAD6FBB2" box="[629,749,1092,1116]" pageId="17" pageNumber="320" value="2016-06-08">
<collectingDate id="AA41E9C8FFD9FFD6205A1722EAD6FBB2" box="[629,749,1092,1116]" pageId="17" pageNumber="320" value="2016-06-08">08Jun2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD6225D1703E8D6FB93" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6225D1703E8D6FB93" box="[114,237,1125,1149]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893316; female
<specimenCode id="9E1D9E9BFFD9FFD6225A17E2E953FB72" box="[117,360,1156,1180]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085236</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6235717E2E9F8FB72" box="[376,451,1156,1180]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD623FC17E2EA5DFB72" box="[467,614,1156,1180]" pageId="17" pageNumber="320" value="2016-04-07">
<collectingDate id="AA41E9C8FFD9FFD623FC17E2EA5DFB72" box="[467,614,1156,1180]" pageId="17" pageNumber="320" value="2016-04-07">07Apr2016</collectingDate>
</date>
,
<location id="CB64603BFFD9FFD6205317E3EACAFB73" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6205317E3EACAFB73" box="[636,753,1157,1181]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893317; female
<specimenCode id="9E1D9E9BFFD9FFD6203917C2EADDFB52" box="[534,742,1188,1212]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085235</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E17A2E884FB32" box="[113,191,1220,1244]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD622FE17A2E950FB32" box="[209,363,1220,1244]" pageId="17" pageNumber="320" value="2016-04-07">
<collectingDate id="AA41E9C8FFD9FFD622FE17A2E950FB32" box="[209,363,1220,1244]" pageId="17" pageNumber="320" value="2016-04-07">07Apr2016</collectingDate>
</date>
;
<location id="CB64603BFFD9FFD623AC17A3E9C5FB33" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD623AC17A3E9C5FB33" box="[387,510,1221,1245]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893317; female
<specimenCode id="9E1D9E9BFFD9FFD6237E1782EA1AFB12" box="[337,545,1252,1276]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085269</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620001782EA55FB12" box="[559,622,1252,1276]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU04</specimenCode>
_
<date id="BA051020FFD9FFD620521782EAC8FB12" box="[637,755,1252,1276]" pageId="17" pageNumber="320" value="2016-06-01">
<collectingDate id="AA41E9C8FFD9FFD620521782EAC8FB12" box="[637,755,1252,1276]" pageId="17" pageNumber="320" value="2016-06-01">01Jun2016</collectingDate>
</date>
(Ma9036);
<location id="CB64603BFFD9FFD622DC1663E960FAF3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD622DC1663E960FAF3" box="[243,347,1285,1309]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893319; female
<specimenCode id="9E1D9E9BFFD9FFD6225D1642E976FAD2" box="[114,333,1316,1340]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0085280</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD623731642E9A4FAD2" box="[348,415,1316,1340]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU07</specimenCode>
_
<date id="BA051020FFD9FFD623821642EA08FAD2" box="[429,563,1316,1340]" pageId="17" pageNumber="320" value="2016-03-16">
<collectingDate id="AA41E9C8FFD9FFD623821642EA08FAD2" box="[429,563,1316,1340]" pageId="17" pageNumber="320" value="2016-03-16">16Mar2016</collectingDate>
</date>
(mitogenome);
<specimenCode id="9E1D9E9BFFD9FFD6225C1622E969FAB2" box="[115,338,1348,1372]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0083285</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6234E1622E998FAB2" box="[353,419,1348,1372]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU10</specimenCode>
,
<location id="CB64603BFFD9FFD6239B1623EA18FAB3" LSID="urn:lsid:plazi:treatment:461287F6FFD8FFD5219D1604EDF9F8DB:CB64603BFFD9FFD6239B1623EA18FAB3" box="[436,547,1349,1373]" country="Singapore" county="GenBank" municipality="GenBank" name="GenBank" pageId="17" pageNumber="320">GenBank</location>
accession code: PP893320;
<specimenCode id="9E1D9E9BFFD9FFD623231602E9CDFA92" box="[268,502,1380,1404]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0257071</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD620281602EA77FA92" box="[519,588,1380,1404]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU19</specimenCode>
_
<date id="BA051020FFD9FFD620731602EAD1FA92" box="[604,746,1380,1404]" pageId="17" pageNumber="320" value="2018-04-26">
<collectingDate id="AA41E9C8FFD9FFD620731602EAD1FA92" box="[604,746,1380,1404]" pageId="17" pageNumber="320" value="2018-04-26">26Apr2018</collectingDate>
</date>
;
<collectingMunicipality id="2E60AC9AFFD9FFD6225B16E3E8E1FA73" box="[116,218,1413,1437]" pageId="17" pageNumber="320">GenBank</collectingMunicipality>
accession code: PP893321;
<specimenCode id="9E1D9E9BFFD9FFD6203216E2EADEFA72" box="[541,741,1412,1436]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0256595</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6225E16C3E894FA53" box="[113,175,1445,1469]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU19</specimenCode>
_
<date id="BA051020FFD9FFD6229216C3E978FA53" box="[189,323,1445,1469]" pageId="17" pageNumber="320" value="2018-05-03">
<collectingDate id="AA41E9C8FFD9FFD6229216C3E978FA53" box="[189,323,1445,1469]" pageId="17" pageNumber="320" value="2018-05-03">03May2018</collectingDate>
</date>
;
<collectingCounty id="27654E6CFFD9FFD6237E16C3E98CFA53" box="[337,439,1445,1469]" pageId="17" pageNumber="320">GenBank</collectingCounty>
accession code: PP893322;
<specimenCode id="9E1D9E9BFFD9FFD6225D16A3E901FA33" box="[114,314,1477,1501]" collectionCode="ZRCBDP" pageId="17" pageNumber="320">ZRCBDP0317777</specimenCode>
_
<specimenCode id="9E1D9E9BFFD9FFD6236716A3E9BEFA33" box="[328,389,1477,1501]" collectionCode="PU" country="USA" name="Princeton University" pageId="17" pageNumber="320">PU24</specimenCode>
_
<date id="BA051020FFD9FFD623BD16A3EA37FA33" box="[402,524,1477,1501]" pageId="17" pageNumber="320" value="2019-04-02">
<collectingDate id="AA41E9C8FFD9FFD623BD16A3EA37FA33" box="[402,524,1477,1501]" pageId="17" pageNumber="320" value="2019-04-02">02Apr2019</collectingDate>
</date>
; GenBank accession code: PP893323
</materialsCitation>
.
</paragraph>
<caption id="9AC46668FFD9FFD6210217D3EBF0FAED" ID-DOI="http://doi.org/10.5281/zenodo.14683100" ID-Zenodo-Dep="14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="17" pageNumber="320" startId="17.[813,848,1205,1227]" targetBox="[816,1451,194,1187]" targetPageId="17" targetType="figure">
<paragraph id="CE0436E0FFD9FFD6210217D3EBF0FAED" blockId="17.[813,1453,1205,1284]" pageId="17" pageNumber="320">
Fig. 18.
<taxonomicName id="09BB4D63FFD9FFD621AB17D3EC4FFB24" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[900,1140,1205,1226]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="320" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFD9FFD621AB17D3EC4FFB24" box="[900,1140,1205,1226]" italics="true" pageId="17" pageNumber="320">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFD9FFD626AC17D3ECC5FB25" box="[1155,1278,1205,1227]" pageId="17" pageNumber="320" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFD9FFD6225E1543E8E1F9B2" blockId="17.[113,752,1572,1628]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E1543E889F9D3" bold="true" box="[113,178,1573,1597]" pageId="17" pageNumber="320">Male.</emphasis>
Length: body
<quantity id="09439B05FFD9FFD6237A1543E982F9D2" box="[341,441,1573,1597]" metricMagnitude="-3" metricUnit="m" metricValue="1.79" pageId="17" pageNumber="320" unit="mm" value="1.79">1.79 mm</quantity>
; wing
<quantity id="09439B05FFD9FFD6202B1542EA53F9D2" box="[516,616,1572,1596]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="17" pageNumber="320" unit="mm" value="1.66">1.66 mm</quantity>
; wing width
<quantity id="09439B05FFD9FFD6225E1522E8EDF9B2" box="[113,214,1604,1628]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="320" unit="mm" value="0.55">0.55 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFD9FFD6225E15E3E90FF893" blockId="17.[113,753,1669,1917]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E15E3E88EF973" bold="true" box="[113,181,1669,1693]" pageId="17" pageNumber="320">Head.</emphasis>
Frons and face shiny metallic green. Face triangular, wide below antenna, narrowing down to clypeus, there a little wider than pedicel; indented near middle with a furrow. Clypeus narrow. Hairs and bristles on head brownish. A pair of strong retroclinate ocellar bristles, a pair of strong proclinate vertical bristles (
<figureCitation id="56802A65FFD9FFD623801443EA38F8D2" box="[431,515,1828,1853]" captionStart="Fig" captionStartId="16.[838,875,1049,1071]" captionTargetBox="[836,1472,189,1034]" captionTargetId="figure-264@16.[836,1472,189,1034]" captionTargetPageId="16" captionText="Fig. 16. Protomedetera uncinata, new species, male habitus (ZRCBDP0085280). Photograph by Kristy Chang." figureDoi="http://doi.org/10.5281/zenodo.14683094" httpUri="https://zenodo.org/record/14683094/files/figure.png" pageId="17" pageNumber="320">Fig. 16</figureCitation>
) and a pair of short postvertical bristles pointing inward. Postocular bristles uniseriate, brown.
</paragraph>
<paragraph id="CE0436E0FFD9FFD6225E14C2ED96F9FD" blockId="17.[113,753,1956,2109]" lastBlockId="17.[812,1453,1339,1555]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD6225E14C2E8DDF852" bold="true" box="[113,230,1956,1980]" pageId="17" pageNumber="320">Antennae.</emphasis>
Scape yellow, short, with an interior projection. Pedicel yellow, cup-like, large, with postpedicel sunken in it; in the apical crown of bristles 3 long apical bristles longer than and bending over the postpedicel; below the long ventral apical bristle a second a long finer bristle; with some additional short, scattered bristles, most at the inner side. Postpedicel with apex dark brown, spheroid, with short pubescence and a subapical long arista (shifted toward outside). All bristles on postpedicel brown. Arista 3 times as long as all antennal segments together; black, naked, basal portion of arista short. Palpus brown, small, circular, with a long brown subapical bristle, and a second bristle beneath.
</paragraph>
<paragraph id="CE0436E0FFD9FFD62102155DED90F8BD" blockId="17.[811,1452,1595,1876]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD62102155DEBB3F9BD" bold="true" box="[813,904,1595,1619]" pageId="17" pageNumber="320">Thorax.</emphasis>
Brownish metallic (pollinosity probably absent, not visible in the ethanol conserved specimen). Basal half of mesonotum uniformly covered with short bristles, the rows of 4 serial acrostichals indistinctly separated from dc. Apical half of mesonotum flat before scutellum, lacking bristles in central area. Hairs and bristles on thorax brownish black; 2 long prescutellar dorsocentral bristles, anterior dorsocentrals short followed by 2 long prescutellar dorsocentrals. A pair of very long crossing scutellars with a short bristle at outside.
</paragraph>
<paragraph id="CE0436E0FFD9FFD62102141DED57F83D" blockId="17.[813,1452,1915,2003]" pageId="17" pageNumber="320">
<emphasis id="FCCFEAF2FFD9FFD62102141DEB52F87D" bold="true" box="[813,873,1915,1939]" pageId="17" pageNumber="320">Legs.</emphasis>
Mainly brown but trochanters, tip of all femora, all tibiae and tarsomeres yellow except tarsomere 5 of all legs brownish. Hairs and bristles on legs brownish black.
</paragraph>
<paragraph id="CE0436E0FFD9FFD52102149AEA84FF1B" blockId="17.[813,1451,2043,2099]" lastBlockId="18.[136,775,189,245]" lastPageId="18" lastPageNumber="321" pageId="17" pageNumber="320">Fore legs. Coxa densely covered with short pale bristles, long apical bristles; one bristle in middle longer than the others. Fore femur with short pale ventrals and no distinct posterior preapical bristles. Tibia with short apicals.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7127BEAA2FE3B" blockId="18.[135,776,285,469]" pageId="18" pageNumber="321">Mid legs. Coxa with short anterior bristles and a long pale brownish exterior bristle as long as coxa is long. Femur with a fine preapical anterior bristle. Tibia with a short anterodorsal and a longer posterodorsal near base; a stronger anterodorsal at ⅓ of base and a small anterodorsal in preapical crown; ventral preapical twice as long as width of tibia.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7129BE921FD5B" blockId="18.[136,776,509,693]" pageId="18" pageNumber="321">Hind legs. Coxa with a short brownish exterior bristle near middle. Femur in apical half with a row of short brown anteroventral bristles; preapical anterior bristles indistinct. Hind femur with indistinct preapical bristles. Hind tibia lacking distinct bristles, apart from the short bristles in apical crown.</paragraph>
<paragraph id="CE0436E0FFDAFFD522A711BBE919FCDB" blockId="18.[136,775,733,821]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A711BBE8E8FD1B" bold="true" box="[136,211,733,757]" pageId="18" pageNumber="321">Wings.</emphasis>
Hyaline, tinged yellowish grey. Veins yellowish brown. Squama white, with a single short brown bristle. Haltere white.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7103BE96CFC9B" blockId="18.[136,343,861,885]" box="[136,343,861,885]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7103BE8C5FC9B" bold="true" box="[136,254,861,885]" pageId="18" pageNumber="321">Abdomen.</emphasis>
Brown.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A710FBE90FFB1B" blockId="18.[135,777,925,1269]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A710FBE968FC5B" bold="true" box="[136,339,925,949]" pageId="18" pageNumber="321">Male terminalia.</emphasis>
<figureCitation id="56802A65FFDAFFD5237110FBE98FFC5B" box="[350,436,925,949]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18</figureCitation>
. Foramen situated at base of epandrium. Surstylus brown, wide truncate composed of a single element supposed to be the dorsal surstylus. Apex surstylus rounded with 3 short and 3 long bristles on the ventral side, and 2 strong subapical bristles on the lateral side (
<figureCitation id="56802A65FFDAFFD522EB175BE91DFBBB" box="[196,294,1085,1109]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
). Cercus short oval, with a short and a very long apical bristle (
<figureCitation id="56802A65FFDAFFD52379173BE98EFB9B" box="[342,437,1117,1141]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
), other bristles short (
<figureCitation id="56802A65FFDAFFD520B3173BEAC7FB9B" box="[668,764,1117,1141]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18C</figureCitation>
). Hypandrium bifurcate (
<figureCitation id="56802A65FFDAFFD523BF171BE9C2FB7B" box="[400,505,1149,1173]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18A</figureCitation>
), the fork bent dorsally (
<figureCitation id="56802A65FFDAFFD522BF17FBE8CBFB5B" box="[144,240,1181,1205]" captionStart="Fig" captionStartId="17.[813,848,1205,1227]" captionTargetBox="[816,1451,194,1187]" captionTargetId="figure-17@17.[798,1469,176,1197]" captionTargetPageId="17" captionText="Fig. 18. Protomedetera uncinata, new species, male terminalia. A, Ventral; B, Lateral; C, Dorsal. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683100" httpUri="https://zenodo.org/record/14683100/files/figure.png" pageId="18" pageNumber="321">Fig. 18B</figureCitation>
). Phallus hidden below hypandrium. Apicoventral epandrial lobe lacking, but 3 almost equally long epandrial bristles present.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7167BE973FA9B" blockId="18.[136,775,1309,1397]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7167BE8D9FADB" bold="true" box="[136,226,1309,1333]" pageId="18" pageNumber="321">Female.</emphasis>
Length: body
<quantity id="09439B05FFDAFFD523A9167BEA3EFADB" box="[390,517,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="1.775" metricValueMax="2.0" metricValueMin="1.55" pageId="18" pageNumber="321" unit="mm" value="1.775" valueMax="2.0" valueMin="1.55">1.552 mm</quantity>
; wing
<quantity id="09439B05FFDAFFD5207E167BEADFFADB" box="[593,740,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="1.92" metricValueMax="2.2" metricValueMin="1.64" pageId="18" pageNumber="321" unit="mm" value="1.92" valueMax="2.2" valueMin="1.64">1.642.2 mm</quantity>
. In most aspects, resembles male as there are no secondary sexual characters.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFDAFFD522A716FBEDF9F8DB" pageId="18" pageNumber="321" type="discussion">
<paragraph id="CE0436E0FFDAFFD522A716FBEA73F9DB" blockId="18.[136,775,1437,1589]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A716FBE936FA5B" bold="true" box="[136,269,1437,1461]" pageId="18" pageNumber="321">Comments.</emphasis>
Variability is noted in the darkening of the postpedicel from brown with a paler base to almost black apex.
<taxonomicName id="09BB4D63FFDAFFD522E716BBE9F0FA1B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[200,459,1501,1525]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFDAFFD522E716BBE9F0FA1B" box="[200,459,1501,1525]" italics="true" pageId="18" pageNumber="321">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFDAFFD523F616B8EA60FA1B" box="[473,603,1501,1525]" pageId="18" pageNumber="321" rank="species">new species</taxonomicNameLabel>
, differs from all other known
<taxonomicName id="09BB4D63FFDAFFD52336169BE982F9FB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[281,441,1533,1557]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD52336169BE982F9FB" box="[281,441,1533,1557]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
by the strong forked tip of the hypandrium. The forks are bent dorsally.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7153BEACAF8BB" blockId="18.[135,775,1629,1877]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7153BE9A5F99B" bold="true" box="[136,414,1629,1653]" pageId="18" pageNumber="321">National distribution.</emphasis>
Thanks to the barcoding,
<specimenCount id="D8BDFD69FFDAFFD520C4153BE8C2F97B" count="13" pageId="18" pageNumber="321" type="generic">13 specimens</specimenCount>
were identified.
<taxonomicName id="09BB4D63FFDAFFD52393151BEAF3F97B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[444,712,1661,1685]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="uncinata" status="sp. nov.">
<emphasis id="FCCFEAF2FFDAFFD52393151BEAF3F97B" box="[444,712,1661,1685]" italics="true" pageId="18" pageNumber="321">Protomedetera uncinata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFDAFFD520F51518E8E3F95B" pageId="18" pageNumber="321" rank="species">new species</taxonomicNameLabel>
, is the most abundant
<taxonomicName id="09BB4D63FFDAFFD523F815FBEAF5F95B" box="[471,718,1693,1717]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD523F815FBEA4CF95B" box="[471,631,1693,1717]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
species
</taxonomicName>
ever found in
<collectingCountry id="B6AC7670FFDAFFD522C015DBE964F93B" box="[239,351,1725,1749]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
; it is exclusive to Pulau Ubin. All the records are from back mangroves, except for a single record from a coastal forest (PU26). This site is very close to a back mangrove, so sample contamination could have occurred during flooding of the area by a storm and a spring tide.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A7141BEA43F85B" blockId="18.[136,775,1917,1973]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD522A7141BE9FDF87B" bold="true" box="[136,454,1917,1941]" pageId="18" pageNumber="321">
Other
<taxonomicName id="09BB4D63FFDAFFD522FF141BE9FDF87B" box="[208,454,1917,1941]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFDAFFD522FF141BE955F87B" bold="true" box="[208,366,1917,1941]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
species.
</taxonomicName>
</emphasis>
Two more unidentified species of
<taxonomicName id="09BB4D63FFDAFFD5228814FBE97CF85B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[167,327,1949,1973]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5228814FBE97CF85B" box="[167,327,1949,1973]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
were found on Pulau Ubin.
</paragraph>
<paragraph id="CE0436E0FFDAFFD522A714BBED84FEFB" blockId="18.[136,776,2013,2101]" lastBlockId="18.[835,1474,189,277]" pageId="18" pageNumber="321">
1. A female of
<taxonomicName id="09BB4D63FFDAFFD5230014BBE9F4F81B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[303,463,2013,2037]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5230014BBE9F4F81B" box="[303,463,2013,2037]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
with quadriserial acrostichal bristles, 4 long dorsocentrals, and yellow hind metatarsus was found in station PU01 on
<date id="BA051020FFDAFFD523C31B7BEAA7F7DB" box="[492,668,2077,2101]" pageId="18" pageNumber="321" value="2019-03-19">19 March 2019</date>
(labelled sp4). It seems to be different from the other three species reported above. Unfortunately, no barcode is available yet to give an indication if it might belong to another species.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C125BEB82FCFB" blockId="18.[834,1475,317,789]" pageId="18" pageNumber="321">
2. A female, provisionally labelled as sp5 (PU01, ZRCBDP0083912, image on BOS: https://singapore. biodiversity.online/species/A-Arth-Hexa-Diptera-002292) has the proboscis as long as head is high, which is unusually long for
<taxonomicName id="09BB4D63FFDAFFD521B512DBEC02FE3B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[922,1081,445,469]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD521B512DBEC02FE3B" box="[922,1081,445,469]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. It has 2 long dorsocentrals, but since the specimen was severely damaged, some dorsocentrals may have been lost. The basal tarsomere of the hind tarsus is slender and longer than the following tarsomere, which is also unusual for
<taxonomicName id="09BB4D63FFDAFFD5263C115BEC89FDBB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1043,1202,573,597]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5263C115BEC89FDBB" box="[1043,1202,573,597]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. Since the specimen was destroyed during extraction for sequencing, no nomenclatorial actions can be taken. The mitogenome was successful and the species clusters at the outside of the other
<taxonomicName id="09BB4D63FFDAFFD5273411FBED81FD5B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1307,1466,669,693]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5273411FBED81FD5B" box="[1307,1466,669,693]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. Following barcode is available (courtesy of Darren Yeo): ZRCBDP0083912 sp5 PU01, GenBank accession code: PP893324.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C105BEC48FC5B" blockId="18.[835,1476,829,949]" pageId="18" pageNumber="321">
<materialsCitation id="7ED33CBDFFDAFFD5216C105BEC54FC5B" collectingDate="2018-03-22" collectionCode="PU, ZRCBDP, PP" pageId="18" pageNumber="321" specimenCode="PU12, ZRCBDP0281845, PP893325" specimenCount="1">
3. A female belonging to the
<taxonomicName id="09BB4D63FFDAFFD526AC1058ECD2FCBB" box="[1155,1257,829,853]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD526AC1058ECD2FCBB" box="[1155,1257,829,853]" italics="true" pageId="18" pageNumber="321">P. glabra</emphasis>
</taxonomicName>
group was found in a strip of secondary forest in front of Endut Senit Campsite (
<specimenCode id="9E1D9E9BFFDAFFD52163101BEBB4FC7B" box="[844,911,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="18" pageNumber="321">PU12</specimenCode>
,
<specimenCode id="9E1D9E9BFFDAFFD521B0101BEC54FC7B" box="[927,1135,893,917]" collectionCode="ZRCBDP" pageId="18" pageNumber="321">ZRCBDP0281845</specimenCode>
_
<specimenCode id="9E1D9E9BFFDAFFD52651101BEC80FC7B" box="[1150,1211,893,917]" collectionCode="PU" country="USA" name="Princeton University" pageId="18" pageNumber="321">PU12</specimenCode>
_
<date id="BA051020FFDAFFD526E5101BED70FC7B" box="[1226,1355,893,917]" pageId="18" pageNumber="321" value="2018-03-22">
<collectingDate id="AA41E9C8FFDAFFD526E5101BED70FC7B" box="[1226,1355,893,917]" pageId="18" pageNumber="321" value="2018-03-22">22Mar2018</collectingDate>
</date>
, GenBank accession code:
<specimenCode id="9E1D9E9BFFDAFFD521DA10FBEC53FC5B" box="[1013,1128,925,949]" collectionCode="PP" pageId="18" pageNumber="321">PP893325</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C10B8EDF8FA9B" blockId="18.[834,1475,989,1397]" pageId="18" pageNumber="321">
As already mentioned above,
<taxonomicName id="09BB4D63FFDAFFD5268110B8ED24FC1B" box="[1198,1311,989,1013]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD5268110B8ED24FC1B" box="[1198,1311,989,1013]" italics="true" pageId="18" pageNumber="321">P. glabra</emphasis>
</taxonomicName>
has different morphology from the type species of
<taxonomicName id="09BB4D63FFDAFFD5273C109BEDF9FBFB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1299,1474,1021,1045]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5273C109BEDF9FBFB" box="[1299,1474,1021,1045]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
and should probably be attributed to another genus. The postpedicel does not bear long apical bristles and the position of the male terminalia on the abdomen is different.
<taxonomicName id="09BB4D63FFDAFFD5216B171BECD1FB7B" authority="Tang et al., 2018" authorityName="Tang" authorityYear="2018" box="[836,1258,1149,1174]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="FCCFEAF2FFDAFFD5216B171BEC0AFB7B" box="[836,1073,1149,1173]" italics="true" pageId="18" pageNumber="321">Protomedetera glabra</emphasis>
<bibRefCitation id="AA2A4B11FFDAFFD526181718ECD1FB7B" author="Tang C &amp; Grootaert P &amp; Yang D" box="[1079,1258,1149,1174]" pageId="18" pageNumber="321" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
</taxonomicName>
was found in
<collectingCountry id="B6AC7670FFDAFFD527AE1718EBF0FB5B" name="Papua New Guinea" pageId="18" pageNumber="321">Papua New Guinea</collectingCountry>
(
<typeStatus id="11008842FFDAFFD521F517FBEC00FB5B" box="[986,1083,1181,1205]" pageId="18" pageNumber="321" type="holotype">holotype</typeStatus>
) and in
<collectingCountry id="B6AC7670FFDAFFD526B917FBED3EFB5B" box="[1174,1285,1181,1205]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
, but although the male terminalia of the specimens from
<collectingCountry id="B6AC7670FFDAFFD526DF17D8EDF9FB3B" box="[1264,1474,1213,1237]" name="Papua New Guinea" pageId="18" pageNumber="321">Papua New Guinea</collectingCountry>
and those from
<collectingCountry id="B6AC7670FFDAFFD521D817BBEC5EFB1B" box="[1015,1125,1245,1269]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
are similar, they are distinctly different and hence the taxonomy of this species group needs revision. The species was generally found in anthropogenic habitats, like Clementi Woods (park), Evolution Garden (Botanic Gardens), NUS (university park), and Sime Forest.
</paragraph>
<paragraph id="CE0436E0FFDAFFD5216C16FBEDF9F8DB" blockId="18.[834,1476,1437,1846]" pageId="18" pageNumber="321">
<emphasis id="FCCFEAF2FFDAFFD5216C16FBED7CFA5B" bold="true" box="[835,1351,1437,1461]" pageId="18" pageNumber="321">
Comments on
<taxonomicName id="09BB4D63FFDAFFD521C116FBECABFA5B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1006,1168,1437,1461]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD521C116FBECABFA5B" bold="true" box="[1006,1168,1437,1461]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
in Pulau Ubin.
</emphasis>
The genus
<taxonomicName id="09BB4D63FFDAFFD5216C16DBEBD8FA3B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[835,995,1469,1493]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5216C16DBEBD8FA3B" box="[835,995,1469,1493]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
is very rare in
<collectingCountry id="B6AC7670FFDAFFD526AC16DBECD4FA3B" box="[1155,1263,1469,1493]" name="Singapore" pageId="18" pageNumber="321">Singapore</collectingCountry>
and although many sites and various biotopes were surveyed during the last twenty years for prolonged periods, only two species were recorded in the past (
<bibRefCitation id="AA2A4B11FFDAFFD5261C157BECC3F9DB" author="Tang C &amp; Grootaert P &amp; Yang D" box="[1075,1272,1565,1589]" pageId="18" pageNumber="321" pagination="137 - 151" refId="ref15356" refString="Tang C, Grootaert P &amp; Yang D (2018) Protomedetera, a new genus from the Oriental and Australasian realms (Diptera, Dolichopodidae, Medeterinae). Zookeys, 743: 137 - 151." type="journal article" year="2018">Tang et al., 2018</bibRefCitation>
). With six species now, Pulau Ubin seems to be a hot spot for the genus
<taxonomicName id="09BB4D63FFDAFFD5216C153BEBD8F99B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[835,995,1629,1653]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5216C153BEBD8F99B" box="[835,995,1629,1653]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
. It is unclear what the reason might be. On Pulau Ubin, all species were found in mangrove areas or very close to mangroves.
<taxonomicName id="09BB4D63FFDAFFD5260A15FBECFEF95B" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1061,1221,1693,1717]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5260A15FBECFEF95B" box="[1061,1221,1693,1717]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
is a tree trunk dweller, but its precise biology is unknown. The presence of lots of dead decaying wood present at the surveyed sites might favour the presence of
<taxonomicName id="09BB4D63FFDAFFD5266F159BECE5F8FB" authorityName="Tang, Yang &amp; Grootaert" authorityYear="2018" box="[1088,1246,1789,1813]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD5266F159BECE5F8FB" box="[1088,1246,1789,1813]" italics="true" pageId="18" pageNumber="321">Protomedetera</emphasis>
</taxonomicName>
, where it might feed on wood inhabiting insects like the related genus
<taxonomicName id="09BB4D63FFDAFFD52779147BED87F8DB" authorityName="Fischer von Waldheim" authorityYear="1819" box="[1366,1468,1821,1845]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFDAFFD52779147BED87F8DB" box="[1366,1468,1821,1845]" italics="true" pageId="18" pageNumber="321">Medetera</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,104 @@
<document id="74415B4E4C5CC07CCEA75191DDF06778" ID-DOI="10.15298/rusentj.33.4.01" ID-ISSN="0132-8069" ID-Zenodo-Dep="14683166" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737172112549" checkinUser="felipe" docAuthor="Shcherbakov, Dmitry E. &amp; Shmakov, Alexei S." docDate="2024" docId="9B5FBC4801419E63FF48173EFC571EF0" docLanguage="en" docName="RussEntomolJ.33.4.415-418.pdf" docOrigin="Russian Entomological Journal 33 (4)" docSource="http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.4:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="4" docTitle="Iotacypha vishniakovae Shcherbakov et Shmakov 2024, sp.n." docType="treatment" docVersion="2" lastPageNumber="417" masterDocId="6766C43001439E61FFF51265FFD31A00" masterDocTitle="Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber" masterLastPageNumber="418" masterPageNumber="415" pageNumber="417" updateTime="1737175928815" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="AD63985FE66BD90D0596268251986F4F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B92F5BF6ED766C854706B2F423466F92">
<mods:title id="C04EE0E824872ABC8CE5053014EAAD3A">Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber</mods:title>
</mods:titleInfo>
<mods:name id="64EB87EC106EB5DD7FD8968CE55940B5" type="personal">
<mods:role id="6FEF8CC445FCEB5846D780E93B123F3E">
<mods:roleTerm id="3D6EF141F6915033280BAE5B5D3E0A06">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F1447348D416ACC34AA5AA5F0B9D3E73">Shcherbakov, Dmitry E.</mods:namePart>
</mods:name>
<mods:name id="D4410975AEC1942174978632636336D7" type="personal">
<mods:role id="3957900F60360BF352531441C43BC755">
<mods:roleTerm id="F1A70652B0EAC953B20198C415E7E6EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A567FA53E5D67FE41ECDEFE4A43F40E3">Shmakov, Alexei S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5EFA87FB8DD990787449142204813ED0">text</mods:typeOfResource>
<mods:relatedItem id="34BA9DE6F766FCC154F0E1C81C33F850" type="host">
<mods:titleInfo id="453E11177056E5F4EA55EE966F069902">
<mods:title id="292F66D312F30460F88D4F0B4E853A2A">Russian Entomological Journal</mods:title>
</mods:titleInfo>
<mods:part id="94E9221D6CA1765819CB2CE46378E163">
<mods:date id="237DD099BBAFFCF7EA5D24BD4E43520F">2024</mods:date>
<mods:detail id="2BDA08CDDB3D3D9A08617AF0C8B1A37A" type="volume">
<mods:number id="10240A18353FE7180DEF5A14C8ED19E0">33</mods:number>
</mods:detail>
<mods:detail id="AE89B4C63C08AA15179E77C6A29E94CE" type="issue">
<mods:number id="7A191F19FDE7CBF4FC80AE8B085C0181">4</mods:number>
</mods:detail>
<mods:extent id="496B22405EB8BC4B181456091D0CA446" unit="page">
<mods:start id="1820D87A64877CDC69CC64DC79BAB80C">415</mods:start>
<mods:end id="CF852BF1CD98DC6DD90CB1C12A11D783">418</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="A9E04403D76C7D0E9FDBFBE72690DF9C">
<mods:url id="EB39D50CB7A95EA339A2BC94C1007F44">http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc</mods:url>
</mods:location>
<mods:classification id="79A114DAC8F22EBEB6AF2678CCE87490">journal article</mods:classification>
<mods:identifier id="56435C6CD70FEF1D0059215B0AF58BD3" type="DOI">10.15298/rusentj.33.4.01</mods:identifier>
<mods:identifier id="17CB4D9E47019C676CFB82F28387D150" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="4DECA9DF10D55EE5FE26402109F60A6B" type="Zenodo-Dep">14683166</mods:identifier>
</mods:mods>
<treatment id="9B5FBC4801419E63FF48173EFC571EF0" LSID="urn:lsid:plazi:treatment:9B5FBC4801419E63FF48173EFC571EF0" httpUri="http://treatment.plazi.org/id/9B5FBC4801419E63FF48173EFC571EF0" lastPageNumber="417" pageId="2" pageNumber="417">
<subSubSection id="5BEC5ED501419E63FF48173EFCC11F72" box="[189,786,1371,1394]" pageId="2" pageNumber="417" type="nomenclature">
<paragraph id="13490D5E01419E63FF48173EFCC11F72" blockId="2.[189,786,1371,1424]" box="[189,786,1371,1394]" pageId="2" pageNumber="417">
<heading id="4801BA3201419E63FF48173EFCC11F72" box="[189,786,1371,1394]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="417" reason="4">
<taxonomicName id="D4F676DD01419E63FF48173EFD011F72" authority="Shcherbakov et Shmakov" authorityName="Shcherbakov et Shmakov" authorityYear="2024" box="[189,722,1371,1394]" class="Insecta" family="Lophioneuridae" genus="Iotacypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thysanoptera" pageId="2" pageNumber="417" phylum="Arthropoda" rank="species" species="vishniakovae" status="sp. nov.">
<emphasis id="2182D14C01419E63FF48173EFE691F72" box="[189,442,1371,1394]" italics="true" pageId="2" pageNumber="417">Iotacypha vishniakovae</emphasis>
Shcherbakov et Shmakov
</taxonomicName>
,
<emphasis id="2182D14C01419E63FD28173EFCC11F72" bold="true" box="[733,786,1371,1394]" pageId="2" pageNumber="417">
<taxonomicNameLabel id="3AB16C3701419E63FD28173EFCC11F72" box="[733,786,1371,1394]" pageId="2" pageNumber="417" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5BEC5ED501419E63FE43171CFDCA1F90" box="[438,537,1401,1424]" pageId="2" pageNumber="417" type="description">
<paragraph id="13490D5E01419E63FE43171CFDCA1F90" blockId="2.[189,786,1371,1424]" box="[438,537,1401,1424]" pageId="2" pageNumber="417">
<figureCitation id="8BCD11DB01419E63FE43171CFDC61F90" box="[438,533,1401,1424]" captionStart="Figs 19" captionStartId="1.[188,226,1842,1861]" captionTargetBox="[280,1329,302,1799]" captionTargetId="figure-6@1.[280,1329,302,1799]" captionTargetPageId="1" captionText="Figs 19. Iotacypha vishniakovae Shcherbakov et Shmakov, sp.n., female holotype PIN 5608/174 (photographs by Dmitry Vorontsov, except 78; 56 in polarized light): 1 — habitus, dorsal (a — apical cell; d — discal cell; pa — preapical cell; arrow — precostal carina); 2 — habitus, ventral; 3 — head and thorax, dorsal; 4 — apex of abdomen, ventral; 5 — anterior part of body, left lateral (arrow, left ocellus); 6 — distal part of abdomen, right lateral; 7 — right antenna, dorsal; 8 — left antenna, ventral; 9 — right fore tarsus, ventral. Scale bars: 12 — 0.2 mm; 39 — 0.1 mm. Рис. 19. Iotacypha vishniakovae Shcherbakov et Shmakov, sp.n., голотип ПИН 5608/174, самка (фото ДмитриЯ Воронцова, кроме 78; 56 в полЯриЗованном свете): 1 — общий вид сверху (a — апикальнаЯ, d — дискальнаЯ, pa — преапикальные Ячейки; стрелка — прекостальный киль); 2 — общий вид сниЗу; 3 — голова и грудь сверху; 4 — вершина брюшка сниЗу; 5 — переднЯЯ часть тела слева (стрелка левый глаЗок); 6 — дистальнаЯ часть брюшка справа; 7 — правый усик сверху; 8 — левый усик сниЗу; 9 — праваЯ переднЯЯ лапка сниЗу. Длина масштабных линеек: 12 — 0,2 мм; 39 — 0,1 мм." figureDoi="http://doi.org/10.5281/zenodo.14683168" httpUri="https://zenodo.org/record/14683168/files/figure.png" pageId="2" pageNumber="417">Figs 19</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5BEC5ED501419E63FF1617D3FE041C06" pageId="2" pageNumber="417" type="materials_examined">
<paragraph id="13490D5E01419E63FF1617D3FE041C06" blockId="2.[189,787,1462,2076]" pageId="2" pageNumber="417">
<materialsCitation id="A39E070301419E63FF1617D3FE041C06" collectionCode="PIN" country="Myanmar" location="Burmese" municipality="Hukawng Valley" pageId="2" pageNumber="417" specimenCode="PIN 5608" specimenCount="1" stateProvince="Kachin" typeStatus="holotype">
MATERIAL.
<typeStatus id="CC4DB3FC01419E63FE9D17D3FE121FCB" box="[360,449,1462,1483]" pageId="2" pageNumber="417" type="holotype">Holotype</typeStatus>
<specimenCode id="4350A52501419E63FE3317D3FDF71FCB" box="[454,548,1462,1483]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="417">PIN 5608</specimenCode>
/174, female —
<location id="16295B8501419E63FD4817D3FCC11FCB" LSID="urn:lsid:plazi:treatment:9B5FBC4801419E63FF48173EFC571EF0:16295B8501419E63FD4817D3FCC11FCB" box="[701,786,1462,1483]" country="Myanmar" municipality="Hukawng Valley" name="Burmese" pageId="2" pageNumber="417" stateProvince="Kachin">Burmese</location>
amber,
<collectingMunicipality id="F32D972401419E63FEF017B1FE7B1FE9" box="[261,424,1492,1513]" pageId="2" pageNumber="417">Hukawng Valley</collectingMunicipality>
,
<collectingRegion id="D132C3BC01419E63FE4617B1FDE11FE9" box="[435,562,1492,1513]" country="Myanmar" name="Kachin" pageId="2" pageNumber="417">Kachin State</collectingRegion>
,
<collectingCountry id="6BE14DCE01419E63FDC817B1FD4E1FE9" box="[573,669,1492,1513]" name="Myanmar" pageId="2" pageNumber="417">Myanmar</collectingCountry>
; mid-Cretaceous (AlbianCenomanian).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5BEC5ED501419E63FF16146AFAFA1E98" pageId="2" pageNumber="417" type="description">
<paragraph id="13490D5E01419E63FF16146AFBE3182F" blockId="2.[189,787,1462,2076]" lastBlockId="2.[824,1422,303,1264]" pageId="2" pageNumber="417">
DESCRIPTION. Very small, robust lophioneurids; female body length
<quantity id="D40EA0BB01419E63FE991448FE111C42" box="[364,450,1581,1602]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="417" unit="mm" value="0.65">0.65 mm</quantity>
(as preserved). Subbrachypterous. Forewings
<quantity id="D40EA0BB01419E63FEDF142FFEA51C5F" box="[298,374,1610,1631]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="417" unit="mm" value="0.7">0.7 mm</quantity>
long, rather broad (1.9:1), widest beyond 2/3 wing length, broadly rounded apically, steeply tectiform in repose (their apical margins held vertically and meeting along midline), deeply convex, strongly sclerotized (especially proximally); wing base pale; membrane translucent, covered with stub-like microtrichia and fine dark hexagonal areolation, areoles increase and somewhat fade towards apex; veins dark, raised, beset with strong curved setae (including distal C). Costal margin sinuate about midlength (at least in anterior view); costal vein submarginal, with well-developed areolate precostal carina reaching RP2. Sc absent. RA strongly sigmoidal, almost as long as wing width, distant from C distally, leaving short R+M stem about its midlength (short RP+M stem developed). Arculus (basal
<emphasis id="2182D14C01419E63FE6515A9FE131DE1" box="[400,448,1996,2017]" italics="true" pageId="2" pageNumber="417">m-cu</emphasis>
crossvein) between R+M and CuA closing broad basal cell. RP and M both forked before 2/3 wing length, RP1 and M2 transverse, oblique RP2 and M1 fused for a short distance, forming petiole of apical cell enclosing wing apex. RP1 apex not far from RA; discal cell longer than each of two preapical cells; apical and anterior preapical cells transverse, posterior preapical cell elongated. Clavus short and broad, with two anal veins. Hind wings narrow, broadly rounded at apex; veins dark; R+M+CuA stem dividing distal to 1/2 wing length, R and M separating distal to 1/3 wing length; R, M and CuA subtransverse; claval furrow distinct; clavus with areolation near base.
</paragraph>
<paragraph id="13490D5E01419E63FCAB105DFAFA1E98" blockId="2.[824,1422,303,1264]" pageId="2" pageNumber="417">Antennae 7-segmented, about 1/2 as long as forewing; scape and pedicel stout; flagellum moderately slender; first and last flagellomeres shortest, terminal flagellomere with two very long curved sense cones (much surpassing its apex) and three unequal apical setae. Eyes protruding, rounded; ommatidia few in number, loosely arranged. Cranium rounded; lateral ocelli near eyes; median ocellus near base of clypeus. Mouth cone wide, directed ventrally and slightly anteriorly; maxillary palps moderately long, stout, probably 3-segmented, 2nd segment long and massive, 3rd segment minute; labial palps short, unsegmented. Pronotum transverse, saddle-shaped; mesoscutum subtriangular. Legs rather short, hind tibia no longer than 1/4 forewing length. Hind coxae largest. Tibiae with rows of setae. Tarsi 2-segmented, basitarsus about 1/2 distitarsus length; right fore tarsus deformed or malformed, longer than any other tarsus and appearing 3-segmented; two slender curved claws with sub-basal tooth. Female abdomen as long as head + thorax, widest and more sclerotized proximally. Last two sternites with small rounded median projection on posterior margin. Ovipositor rudimentary, concealed by last sternite.</paragraph>
</subSubSection>
<subSubSection id="5BEC5ED501419E63FCAB16C4FC571EF0" pageId="2" pageNumber="417" type="etymology">
<paragraph id="13490D5E01419E63FCAB16C4FC571EF0" blockId="2.[824,1422,303,1264]" pageId="2" pageNumber="417">
ETYMOLOGY.After paleoentomologist
<taxonomicName id="D4F676DD01419E63FB1716C4FC501ED3" authority="N. Vishniakova" authorityName="N. Vishniakova" class="Magnoliopsida" family="Heliotropiaceae" genus="Valentina" kingdom="Plantae" order="Boraginales" pageId="2" pageNumber="417" phylum="Tracheophyta" rank="genus">Valentina N. Vishniakova</taxonomicName>
for her great contribution to our knowledge of lophioneurids.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,124 @@
<document id="C04D782051625A9BBAE72439DCB704B6" ID-DOI="10.26107/RBZ-2024-0032" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682797" ID-ZooBank="D7A0B570-07AC-47F5-9B56-88EBDB277EC0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171734098" checkinUser="felipe" docAuthor="Inkhavilay, Khamla, Sutcharit, &amp; Chirasak" docDate="2024" docId="9F6A8E412302FFD2FF615EBBA933F7E0" docLanguage="en" docName="RafflesBZool.72.438-449.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Krobylos Panha &amp; Burch 2002" docType="treatment" docVersion="2" lastPageNumber="446" masterDocId="6353F639230AFFDAFFB55B66AB49FFCF" masterDocTitle="A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos" masterLastPageNumber="449" masterPageNumber="438" pageNumber="446" updateTime="1737175246880" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8624FBEFD8441C8C4FAC9C9F504F4AEB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9EFE6F7E8836323BCF67EB120B6E21CC">
<mods:title id="898417986C473B73D751266E446D6442">A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos</mods:title>
</mods:titleInfo>
<mods:name id="6376FDBFE94233A28382E77E4E3C35C8" type="personal">
<mods:role id="2B38C3690A7EEFCD60FAF9C8026F98A7">
<mods:roleTerm id="3FC025AA02DBE35BB4F9D476AF463FEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9FA1834698C9E2979D2B0B7E9421AF1A">Inkhavilay, Khamla</mods:namePart>
</mods:name>
<mods:name id="F74C3565757DE1F2CF300B640C1C45AB" type="personal">
<mods:role id="C5CD9177789BC159298E9C4F03D8A902">
<mods:roleTerm id="BE8CF90442FB0B36DB12DB353C971B50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15DA7264246C2EE5A12E92552C3CB282">Sutcharit,</mods:namePart>
</mods:name>
<mods:name id="955AE3B5797426BA80DE1C5CFED50E4D" type="personal">
<mods:role id="2C8C2F734F85A9E0691B47D35BB320E3">
<mods:roleTerm id="31032093D72902A360EC02B7154AD9F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9164A0CC51462093E92F06BD85207EE">Chirasak</mods:namePart>
</mods:name>
<mods:typeOfResource id="CB0CBC1C441F11AE686C6E5143DE8FBA">text</mods:typeOfResource>
<mods:relatedItem id="521AB7C583A0354FCB89F5BB916D2F57" type="host">
<mods:titleInfo id="69AD574B3E9BE586070A3C1319245C0C">
<mods:title id="B6DA78AAD1C84D98C6BBEC558BB6712C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="76F8155497691E546A99574CB0A6508F">
<mods:date id="C1A198AC495D186464210221D4F0CA3F">2024</mods:date>
<mods:detail id="BBBFB50A1B04A89D25A18D2CDB3F9CFB" type="pubDate">
<mods:number id="F057AD3D15CC911F1AE207260C38A59B">2024-11-04</mods:number>
</mods:detail>
<mods:detail id="132606E204876259E5EC63165A8FDF6B" type="volume">
<mods:number id="28591FFCE2DB174205B671ACCBDF2C00">72</mods:number>
</mods:detail>
<mods:extent id="6BEA309E57738057200370DA1C80753A" unit="page">
<mods:start id="5A71D79D38BD493662BFFED281DAA058">438</mods:start>
<mods:end id="EC606F9B7D534CE0D49DFFD8A3E2F560">449</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6C140FE042799CFCF7D0D07683CA123C">journal article</mods:classification>
<mods:identifier id="4E224AC5C42718A2D509F0020B163D51" type="DOI">10.26107/RBZ-2024-0032</mods:identifier>
<mods:identifier id="1AC871BB3BA5AF94603229916B8EEF72" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C67665289AAA76D92B8123C14DD0B967" type="Zenodo-Dep">14682797</mods:identifier>
<mods:identifier id="CEC6BC9E0276842E88D329ED09713D7C" type="ZooBank">D7A0B570-07AC-47F5-9B56-88EBDB277EC0</mods:identifier>
</mods:mods>
<treatment id="9F6A8E412302FFD2FF615EBBA933F7E0" ID-DOI="http://doi.org/10.5281/zenodo.14682809" ID-Zenodo-Dep="14682809" LSID="urn:lsid:plazi:treatment:9F6A8E412302FFD2FF615EBBA933F7E0" httpUri="http://treatment.plazi.org/id/9F6A8E412302FFD2FF615EBBA933F7E0" lastPageNumber="446" pageId="8" pageNumber="446">
<subSubSection id="5FD96CDC2302FFD2FF615EBBA9C2FA3A" box="[212,651,1501,1525]" pageId="8" pageNumber="446" type="nomenclature">
<paragraph id="177C3F572302FFD2FF615EBBA9C2FA3A" blockId="8.[212,651,1501,1525]" box="[212,651,1501,1525]" pageId="8" pageNumber="446">
<heading id="4C34883B2302FFD2FF615EBBA9C2FA3A" bold="true" box="[212,651,1501,1525]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="446" reason="2">
<emphasis id="25B7E3452302FFD2FF615EBBA9C2FA3A" bold="true" box="[212,651,1501,1525]" pageId="8" pageNumber="446">
Genus
<taxonomicName id="D0C344D42302FFD2FE905EBBA9C2FA3A" ID-CoL="7P6LY" authority="Panha &amp; Burch, 2002" authorityName="Panha &amp; Burch" authorityYear="2002" box="[293,651,1501,1525]" class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FE905EBBAACEFA3A" bold="true" box="[293,391,1501,1525]" italics="true" pageId="8" pageNumber="446">Krobylos</emphasis>
Panha &amp; Burch, 2002
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2302FFD2FFC45D7AAAFDF980" pageId="8" pageNumber="446" type="reference_group">
<paragraph id="177C3F572302FFD2FFC45D7AAAFDF980" blockId="8.[113,752,1564,1615]" pageId="8" pageNumber="446">
<treatmentCitationGroup id="37D318792302FFD2FFC45D7AAAFDF980" pageId="8" pageNumber="446">
<treatmentCitation id="966219462302FFD2FFC45D7AAA96F9FE" author="Panha S &amp; Burch JB" box="[113,479,1564,1586]" page="127" pageId="8" pageNumber="446" year="2002">
<taxonomicName id="D0C344D42302FFD2FFC45D7AAA96F9FE" ID-CoL="7P6LY" authority="Panha &amp; Burch, 2002 b: 127" authorityName="Panha &amp; Burch" authorityPageNumber="127" authorityYear="2002" box="[113,479,1564,1586]" class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FFC45D7AAB80F9FE" box="[113,201,1564,1585]" italics="true" pageId="8" pageNumber="446">Krobylos</emphasis>
<bibRefCitation id="735242A62302FFD2FF7A5D7AAA96F9FE" author="Panha S &amp; Burch JB" box="[207,479,1564,1586]" pageId="8" pageNumber="446" pagination="113 - 134" refId="ref7772" refString="Panha S &amp; Burch JB (2002 b) New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana, 10 [1999] (24): 113 - 134. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2002">Panha &amp; Burch, 2002b: 127</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="966219462302FFD2FE5D5D7AA9A5F9FE" author="Panha S &amp; Burch JB" box="[488,748,1564,1586]" page="104" pageId="8" pageNumber="446" year="2005">
<bibRefCitation id="735242A62302FFD2FE5D5D7AA9A5F9FE" author="Panha S &amp; Burch JB" box="[488,748,1564,1586]" pageId="8" pageNumber="446" pagination="2004 - 2005" refId="ref7868" refString="Panha S &amp; Burch JB (2005) An introduction to the microsnails of Thailand. Malacological Review, 37 / 38 [2004 - 2005]: 1 - 155. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2005">Panha &amp; Burch, 2005: 104</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="966219462302FFD2FF235D5FAAF8F980" author="Pall-Gergely B &amp; Hunyadi A &amp; Jochum A &amp; Asami T" box="[150,433,1593,1615]" httpUri="http://treatment.plazi.org/id/1326C88CCDA1F2DF880EC204E90FE263" page="52" pageId="8" pageNumber="446" year="2015">
<bibRefCitation id="735242A62302FFD2FF235D5FAAF8F980" author="Pall-Gergely B &amp; Hunyadi A &amp; Jochum A &amp; Asami T" box="[150,433,1593,1615]" pageId="8" pageNumber="446" pagination="31 - 62" refId="ref7468" refString="Pall-Gergely B, Hunyadi A, Jochum A &amp; Asami T (2015) Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys, 523: 31 - 62." type="journal article" year="2015">Páll-Gergely et al., 2015: 52</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2302FFD2FFC45D11AA06F960" pageId="8" pageNumber="446" type="type_taxon">
<paragraph id="177C3F572302FFD2FFC45D11AA06F960" blockId="8.[113,752,1655,1711]" pageId="8" pageNumber="446">
<emphasis id="25B7E3452302FFD2FFC45D11AA40F940" bold="true" box="[113,265,1655,1679]" pageId="8" pageNumber="446">
<typeStatus id="C87881F52302FFD2FFC45D11ABE2F940" box="[113,171,1655,1679]" pageId="8" pageNumber="446">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="D0C344D42302FFD2FEA75D11A98EF940" ID-CoL="99JQX" authority="Panha &amp; Burch, 2002" authorityName="Panha &amp; Burch" authorityYear="2002" box="[274,711,1655,1679]" class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="pomjuk">
<emphasis id="25B7E3452302FFD2FEA75D11AA85F940" box="[274,460,1655,1679]" italics="true" pageId="8" pageNumber="446">Krobylos pomjuk</emphasis>
Panha &amp; Burch, 2002
</taxonomicName>
, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2302FFD2FFC65DB1A993F840" pageId="8" pageNumber="446" type="diagnosis">
<paragraph id="177C3F572302FFD2FFC65DB1A993F840" blockId="8.[111,752,1751,1935]" pageId="8" pageNumber="446">
<emphasis id="25B7E3452302FFD2FFC65DB1ABAEF920" bold="true" box="[115,231,1751,1775]" pageId="8" pageNumber="446">Diagnosis.</emphasis>
Shell depressed to elevated conic, brownish to light brown, whorl more or less angular, and suture deep. Shell surface with dense and irregular growth lines, and very weak spiral striae may be present. Aperture subvertical; peristome simple (not expanded) to weakly expanded; apertural dentition absent. Umbilicus narrow and rimate.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2302FFD2FFC45CD1A933F7E0" pageId="8" pageNumber="446" type="discussion">
<paragraph id="177C3F572302FFD2FFC45CD1A933F7E0" blockId="8.[112,752,1975,2095]" pageId="8" pageNumber="446">
<emphasis id="25B7E3452302FFD2FFC45CD1AB96F800" bold="true" box="[113,223,1975,1999]" pageId="8" pageNumber="446">Remarks.</emphasis>
Currently,
<taxonomicName id="D0C344D42302FFD2FEE95CD1AAF7F800" authorityName="Panha &amp; Burch" authorityYear="2002" box="[348,446,1975,1999]" class="Gastropoda" family="Gastrocoptidae" genus="Krobylos" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FEE95CD1AAF7F800" box="[348,446,1975,1999]" italics="true" pageId="8" pageNumber="446">Krobylos</emphasis>
</taxonomicName>
consists of nine species, six of which are known exclusively in
<collectingCountry id="6FD47FC72302FFD2FE565CB1A90CF820" box="[483,581,2007,2031]" name="Thailand" pageId="8" pageNumber="446">Thailand</collectingCountry>
, two from
<collectingCountry id="6FD47FC72302FFD2FD035CB1A9A5F821" box="[694,748,2007,2030]" name="Laos" pageId="8" pageNumber="446">Laos</collectingCountry>
, and only one from southern
<collectingCountry id="6FD47FC72302FFD2FE1D5C91AAA1F7C0" box="[424,488,2039,2063]" name="China" pageId="8" pageNumber="446">China</collectingCountry>
(
<bibRefCitation id="735242A62302FFD2FE4D5C91A9A2F7C0" author="Panha S &amp; Burch JB" box="[504,747,2039,2063]" pageId="8" pageNumber="446" pagination="2004 - 2005" refId="ref7868" refString="Panha S &amp; Burch JB (2005) An introduction to the microsnails of Thailand. Malacological Review, 37 / 38 [2004 - 2005]: 1 - 155. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2005">Panha &amp; Burch, 2005</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FFC65371AACBF7E0" author="Pall-Gergely B &amp; Hunyadi A &amp; Jochum A &amp; Asami T" box="[115,386,2071,2095]" pageId="8" pageNumber="446" pagination="31 - 62" refId="ref7468" refString="Pall-Gergely B, Hunyadi A, Jochum A &amp; Asami T (2015) Seven new hypselostomatid species from China, including some of the world's smallest land snails (Gastropoda, Pulmonata, Orthurethra). ZooKeys, 523: 31 - 62." type="journal article" year="2015">Páll-Gergely et al., 2015</bibRefCitation>
; MolluscaBase, 2024).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,333 @@
<document id="FA1987F1A196CA878ED4849E7D425C39" ID-DOI="10.26107/RBZ-2024-0032" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682797" ID-ZooBank="D7A0B570-07AC-47F5-9B56-88EBDB277EC0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171734098" checkinUser="felipe" docAuthor="Inkhavilay, Khamla, Sutcharit, &amp; Chirasak" docDate="2024" docId="9F6A8E412309FFDCFEB45C3DABB5FC5A" docLanguage="en" docName="RafflesBZool.72.438-449.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Boysidia houaphanica Inkhavilay, Sutcharit &amp; Chirasak, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="444" masterDocId="6353F639230AFFDAFFB55B66AB49FFCF" masterDocTitle="A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos" masterLastPageNumber="449" masterPageNumber="438" pageNumber="441" updateTime="1737175246880" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="504B989842389BFF9B7F310005E0876C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A4E8F9A6108869DC08E0A8B9249D1DBF">
<mods:title id="65331D7201A5033E4D8013EBF5FF7222">A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos</mods:title>
</mods:titleInfo>
<mods:name id="83A86F0AC92B84F77B1703BCFF5EFF3B" type="personal">
<mods:role id="A7191077E30CD8CA6CA1C7713A916043">
<mods:roleTerm id="8A56949B3C1312A4924F9294AB09F8A5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A260834F22FEB46D0F9286D9614BFADF">Inkhavilay, Khamla</mods:namePart>
</mods:name>
<mods:name id="ED8E3AEFC677F4261A1B733DB8F4A6A5" type="personal">
<mods:role id="E7A3BF9D6840C84CE93A0A9811F22784">
<mods:roleTerm id="DA2C6B5C4EE4362726D02239B395CDFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A3C78FAABF70FEDE125B311B2929B77">Sutcharit,</mods:namePart>
</mods:name>
<mods:name id="327CEABAF431EAB56FA958BB95B239BF" type="personal">
<mods:role id="F85AB2A5A331F67D49BC149F57463C36">
<mods:roleTerm id="ACF3CE46E76EF3E38168751A4D7C70BB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="31BE2137D1490BCE5BD1CF4C12FB7B03">Chirasak</mods:namePart>
</mods:name>
<mods:typeOfResource id="4BBDC71556ECEEA4703924D772DEB581">text</mods:typeOfResource>
<mods:relatedItem id="DDE00AFCF606C72A238AE19F1938B290" type="host">
<mods:titleInfo id="E38DF4477319AA949CE6C31866421CB5">
<mods:title id="4CF91C906C20C907A4113B5AE845ADA8">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="051D1D8B99D22CD1AF56C6E20A005AEB">
<mods:date id="883B4909FA414059159C2E9D33B97ECE">2024</mods:date>
<mods:detail id="F39550E230F0B48864EEFFE39F00E0C1" type="pubDate">
<mods:number id="33CC3978CF3C169260D974EFCFCD94B4">2024-11-04</mods:number>
</mods:detail>
<mods:detail id="DCFF8073EB363FFD6CE15F9D602C0791" type="volume">
<mods:number id="75ABEFE43A0BA2ED502631A5B75CF199">72</mods:number>
</mods:detail>
<mods:extent id="7EC0BEBDFA6CA999EED3A8EA897E294D" unit="page">
<mods:start id="1D9A6CC32FB1385D314537F2DE0B066F">438</mods:start>
<mods:end id="4E3DCD6ECC977AB3FDEDD70D06F25598">449</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7ACB1F56324621A34B56A912D334A79B">journal article</mods:classification>
<mods:identifier id="4EFFB556BA0EE68655D1F5BAF780BC93" type="DOI">10.26107/RBZ-2024-0032</mods:identifier>
<mods:identifier id="DF293A9DFF58681B475028D53A098FA0" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="6A835C387FA27E099A75AF87316A2B63" type="Zenodo-Dep">14682797</mods:identifier>
<mods:identifier id="422182AC8E25D68A576D5D76CAE3F8D8" type="ZooBank">D7A0B570-07AC-47F5-9B56-88EBDB277EC0</mods:identifier>
</mods:mods>
<treatment id="9F6A8E412309FFDCFEB45C3DABB5FC5A" LSID="urn:lsid:plazi:treatment:9F6A8E412309FFDCFEB45C3DABB5FC5A" httpUri="http://treatment.plazi.org/id/9F6A8E412309FFDCFEB45C3DABB5FC5A" lastPageId="6" lastPageNumber="444" pageId="3" pageNumber="441">
<subSubSection id="5FD96CDC2309FFD9FEB45C3DA9C7F8BC" box="[257,654,1883,1907]" pageId="3" pageNumber="441" type="nomenclature">
<paragraph id="177C3F572309FFD9FEB45C3DA9C7F8BC" blockId="3.[257,654,1883,1939]" box="[257,654,1883,1907]" pageId="3" pageNumber="441">
<heading id="4C34883B2309FFD9FEB45C3DA9C7F8BC" bold="true" box="[257,654,1883,1907]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="441" reason="2">
<emphasis id="25B7E3452309FFD9FEB45C3DA9C7F8BC" bold="true" box="[257,654,1883,1907]" pageId="3" pageNumber="441">
<taxonomicName id="D0C344D42309FFD9FEB45C3DAAB0F8BC" ID-CoL="CPTDR" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[257,505,1883,1907]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="441" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E3452309FFD9FEB45C3DAAB0F8BC" bold="true" box="[257,505,1883,1907]" italics="true" pageId="3" pageNumber="441">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E2309FFD9FDBC5C3DA9C7F8BC" box="[521,654,1883,1907]" pageId="3" pageNumber="441" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2309FFD9FE305C1DA943F85C" box="[389,522,1915,1939]" pageId="3" pageNumber="441" type="description">
<paragraph id="177C3F572309FFD9FE305C1DA943F85C" blockId="3.[257,654,1883,1939]" box="[389,522,1915,1939]" pageId="3" pageNumber="441">
(
<figureCitation id="8FF823D22309FFD9FE385C1DAAAFF85C" box="[397,486,1915,1939]" captionStart="Fig" captionStartId="4.[113,148,1330,1352]" captionTargetBox="[111,1450,189,1314]" captionTargetId="figure-10@4.[111,1450,189,1314]" captionTargetPageId="4" captionText="Fig. 2. A, Boysidia pahpetensis Saurin, 1953, syntype MNHN-IM-2000-33880 from Pah Hia. B, Boysidia houaphanica, new species, paratype CUMZ 14421 from Samtay, Houaphanh, Laos. Abbreviations: b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (B)." figureDoi="http://doi.org/10.5281/zenodo.14682801" httpUri="https://zenodo.org/record/14682801/files/figure.png" pageId="3" pageNumber="441">Figs 2B</figureCitation>
,
<figureCitation id="8FF823D22309FFD9FE465C1DA949F85C" box="[499,512,1915,1939]" captionStart="Fig" captionStartId="5.[136,171,1902,1924]" captionTargetBox="[136,1474,293,1887]" captionTargetId="figure-8@5.[136,1474,293,1887]" captionTargetPageId="5" captionText="Fig. 3. Boysidia houaphanica, new species. AC, holotype CUMZ 14420 from Samtay, Houaphanh, Laos; A, shell and enlarged apertures of different view angles. B, protoconch and C, inner wall of peristome around basal plica showing small nodules. D, paratype CUMZ 14421 from the type locality. Abbreviations: b: basal plica, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica." figureDoi="http://doi.org/10.5281/zenodo.14682805" httpUri="https://zenodo.org/record/14682805/files/figure.png" pageId="3" pageNumber="441">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2309FFD9FF3D5CDDAF23F95C" pageId="3" pageNumber="441" type="materials_examined">
<paragraph id="177C3F572309FFD9FF3D5CDDAF23F95C" blockId="3.[136,777,1979,2099]" lastBlockId="3.[835,1474,1563,1683]" pageId="3" pageNumber="441">
<emphasis id="25B7E3452309FFD9FF3D5CDDAA27F81C" bold="true" box="[136,366,1979,2003]" pageId="3" pageNumber="441">Material examined.</emphasis>
<materialsCitation id="A7AB350A2309FFD9FECD5CDDAFB1F9FC" collectingDate="2022-05" collectionCode="CUMZ" collectorName="K. Inkhavilay &amp; Inkhavilay, A &amp; C. Oulaiseng" country="Laos" latitude="20.014555" location="Ban Lao" longLatPrecision="1" longitude="104.682495" municipality="Samtay District" pageId="3" pageNumber="441" specimenCode="CUMZ 14420" specimenCount="1" stateProvince="Houaphan" typeStatus="holotype">
<typeStatus id="C87881F52309FFD9FECD5CDDAA94F81C" box="[376,477,1979,2003]" pageId="3" pageNumber="441" type="holotype">Holotype</typeStatus>
<specimenCode id="4765972C2309FFD9FE525CDDA9CDF81C" box="[487,644,1979,2003]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="3" pageNumber="441">CUMZ 14420</specimenCode>
(height
<quantity id="D03B92B22309FFD9FD505CDDABFDF83C" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="441" unit="mm" value="3.2">3.2 mm</quantity>
, width
<quantity id="D03B92B22309FFD9FEBD5CBDAA16F83C" box="[264,351,2011,2035]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="3" pageNumber="441" unit="mm" value="2.8">2.8 mm</quantity>
, 6 whorls;
<figureCitation id="8FF823D22309FFD9FE695CBDA918F83C" box="[476,593,2011,2035]" captionStart="Fig" captionStartId="5.[136,171,1902,1924]" captionTargetBox="[136,1474,293,1887]" captionTargetId="figure-8@5.[136,1474,293,1887]" captionTargetPageId="5" captionText="Fig. 3. Boysidia houaphanica, new species. AC, holotype CUMZ 14420 from Samtay, Houaphanh, Laos; A, shell and enlarged apertures of different view angles. B, protoconch and C, inner wall of peristome around basal plica showing small nodules. D, paratype CUMZ 14421 from the type locality. Abbreviations: b: basal plica, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica." figureDoi="http://doi.org/10.5281/zenodo.14682805" httpUri="https://zenodo.org/record/14682805/files/figure.png" pageId="3" pageNumber="441">Fig. 3AC</figureCitation>
) from limestone outcrop at
<location id="121C698C2309FFD9FF485C9AAA15F7DC" LSID="urn:lsid:plazi:treatment:9F6A8E412309FFDCFEB45C3DABB5FC5A:121C698C2309FFD9FF485C9AAA15F7DC" box="[253,348,2044,2067]" country="Laos" latitude="20.014555" longLatPrecision="1" longitude="104.682495" municipality="Samtay District" name="Ban Lao" pageId="3" pageNumber="441" stateProvince="Houaphan">Ban Lao</location>
,
<collectingMunicipality id="F718A52D2309FFD9FEDD5C9DA95AF7DC" box="[360,531,2043,2067]" pageId="3" pageNumber="441">Samtay District</collectingMunicipality>
,
<collectingRegion id="D507F1B52309FFD9FDAB5C9DA84AF7DC" box="[542,771,2043,2067]" country="Laos" name="Houaphan" pageId="3" pageNumber="441">Houaphanh Province</collectingRegion>
,
<collectingCountry id="6FD47FC72309FFD9FF3C537AABF4F7FC" box="[137,189,2076,2099]" name="Laos" pageId="3" pageNumber="441">Laos</collectingCountry>
(
<geoCoordinate id="72F759902309FFD9FF7F537CAA28F7FC" box="[202,353,2074,2099]" degrees="20" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="441" precision="1" seconds="52.4" value="20.014555">20°0052.4″N</geoCoordinate>
,
<geoCoordinate id="72F759902309FFD9FEDF537CA945F7FC" box="[362,524,2074,2099]" degrees="104" direction="east" minutes="40" orientation="longitude" pageId="3" pageNumber="441" precision="1" seconds="57.0" value="104.682495">104°4057.0″E</geoCoordinate>
), coll.
<collectorName id="BA365A812309FFD9FDE4537AA9ACF7FC" box="[593,741,2075,2099]" pageId="3" pageNumber="441">K. Inkhavilay</collectorName>
, A. Inthavong and
<collectorName id="BA365A812309FFD9FC5D5D7DAF31F9FC" box="[1000,1144,1563,1587]" pageId="3" pageNumber="441">C. Oulaiseng</collectorName>
,
<date id="637D19972309FFD9FB305D7AAFB1F9FC" box="[1157,1272,1563,1588]" pageId="3" pageNumber="441" value="2022-05">
<collectingDate id="7339E07F2309FFD9FB305D7AAFB1F9FC" box="[1157,1272,1563,1588]" pageId="3" pageNumber="441" value="2022-05">May 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="A7AB350A2309FFD9FAB15D7AAFCBF99C" collectionCode="CUMZ" country="Laos" pageId="3" pageNumber="441" specimenCode="CUMZ 14421" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
<typeStatus id="C87881F52309FFD9FAB15D7AAE25F9FB" box="[1284,1388,1564,1588]" pageId="3" pageNumber="441" type="paratype">Paratypes</typeStatus>
<specimenCode id="4765972C2309FFD9FAC05D7DA8CFF99C" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="3" pageNumber="441">CUMZ 14421</specimenCode>
(6 shells;
<figureCitation id="8FF823D22309FFD9FC405D5DAF02F99C" box="[1013,1099,1595,1619]" captionStart="Fig" captionStartId="4.[113,148,1330,1352]" captionTargetBox="[111,1450,189,1314]" captionTargetId="figure-10@4.[111,1450,189,1314]" captionTargetPageId="4" captionText="Fig. 2. A, Boysidia pahpetensis Saurin, 1953, syntype MNHN-IM-2000-33880 from Pah Hia. B, Boysidia houaphanica, new species, paratype CUMZ 14421 from Samtay, Houaphanh, Laos. Abbreviations: b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (B)." figureDoi="http://doi.org/10.5281/zenodo.14682801" httpUri="https://zenodo.org/record/14682801/files/figure.png" pageId="3" pageNumber="441">Figs 2B</figureCitation>
,
<figureCitation id="8FF823D22309FFD9FBE25D5DAF30F99C" box="[1111,1145,1595,1619]" captionStart="Fig" captionStartId="5.[136,171,1902,1924]" captionTargetBox="[136,1474,293,1887]" captionTargetId="figure-8@5.[136,1474,293,1887]" captionTargetPageId="5" captionText="Fig. 3. Boysidia houaphanica, new species. AC, holotype CUMZ 14420 from Samtay, Houaphanh, Laos; A, shell and enlarged apertures of different view angles. B, protoconch and C, inner wall of peristome around basal plica showing small nodules. D, paratype CUMZ 14421 from the type locality. Abbreviations: b: basal plica, c: columellar lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica." figureDoi="http://doi.org/10.5281/zenodo.14682805" httpUri="https://zenodo.org/record/14682805/files/figure.png" pageId="3" pageNumber="441">3D</figureCitation>
)
</materialsCitation>
,
<materialsCitation id="A7AB350A2309FFD9FB3B5D5DAE8BF99C" box="[1166,1474,1595,1619]" collectionCode="CUMZ" country="Laos" pageId="3" pageNumber="441" specimenCode="CUMZ 14422" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
<specimenCode id="4765972C2309FFD9FB3B5D5DAE6CF99C" box="[1166,1317,1595,1619]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="3" pageNumber="441">CUMZ 14422</specimenCode>
(3 shells), and
</materialsCitation>
<materialsCitation id="A7AB350A2309FFD9FCF15D3AAF2FF95C" collectionCode="NHMUK" country="Laos" pageId="3" pageNumber="441" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
<collectionCode id="71D2A7922309FFD9FCF15D3AA8E0F9BC" box="[836,937,1628,1651]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="3" pageNumber="441" type="Museum">NHMUK</collectionCode>
(2 shells) were all collected from the type locality together with the
<typeStatus id="C87881F52309FFD9FBB05D1DAF2FF95C" box="[1029,1126,1659,1683]" pageId="3" pageNumber="441" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2309FFD9FCF05DDDAFA5F89C" pageId="3" pageNumber="441" type="diagnosis">
<paragraph id="177C3F572309FFD9FCF05DDDAFA5F89C" blockId="3.[834,1475,1723,1875]" pageId="3" pageNumber="441">
<emphasis id="25B7E3452309FFD9FCF05DDDA8FFF91C" bold="true" box="[837,950,1723,1747]" pageId="3" pageNumber="441">Diagnosis.</emphasis>
Shell minute, heliciform, without tuba. Protoconch sculpture with weakly malleated pits; teleoconch with irregular growth lines. Apertural dentition consisting of five lamellae (parietal, upper and lower palatals, basal and columellar), and small palatal tubercle.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC2309FFDEFCF65C1DA99CF81F" lastPageId="4" lastPageNumber="442" pageId="3" pageNumber="441" type="description">
<paragraph id="177C3F572309FFDEFCF65C1DA99CF81F" blockId="3.[835,1475,1915,2099]" lastBlockId="4.[112,754,1464,2000]" lastPageId="4" lastPageNumber="442" pageId="3" pageNumber="441">
<emphasis id="25B7E3452309FFD9FCF65C1DA89BF85C" bold="true" box="[835,978,1915,1939]" pageId="3" pageNumber="441">Description.</emphasis>
Shell elongate conical, yellowish brown to chestnut-brown, with 5½ to 6¾ convex whorls. Shell height
<quantity id="D03B92B22309FFD9FCF65CDDA883F81C" box="[835,970,1979,2003]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="3" pageNumber="441" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.13.5 mm</quantity>
, shell width
<quantity id="D03B92B22309FFD9FBEE5CDDAFABF81C" box="[1115,1250,1979,2003]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="3" pageNumber="441" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.52.8 mm</quantity>
. Apex rather round; protoconch about 1½ whorls with weak malleated pits. Teleoconch generally smooth with scattered, prominent and irregular growth lines; suture wide, deeply impressed. Last whorl large, rounded, slightly descending near aperture. Peristome attached to preceding whorl; inner surface including plicae and lamellae covered with scattered small nodules; lip slightly thickened, whitish, broadly expanded. Aperture sub-rectangular, with six apertural teeth. Parietal lamella (p) strongly developed, elevated, sinuous deeply inside aperture, consisting of two parts separated at concrescent point: outer part elevated, strong, curved, reaching peristome margin; inner part thickened, elevated, stronger than outer part. Palatal tubercle (pt) strongly developed with triangular shape. Upper palatal plica (upl) thin, flake shaped, long inside aperture, elevated to nearly touch parietal lamella, forming an almost separate chamber at upper junction of peristome; lower (lpl) palatal strong, elevated. Basal plica (b) prominent, small and low. Columellar lamella (c) large, strong, distinct horizontally. Umbilicus perforate, about one-fourth of shell width, rounded, shallow, not revealing preceding whorl.
</paragraph>
</subSubSection>
<caption id="43BC6FDF230EFFDEFFC45E54AFD0FA4F" ID-DOI="http://doi.org/10.5281/zenodo.14682801" ID-Zenodo-Dep="14682801" httpUri="https://zenodo.org/record/14682801/files/figure.png" pageId="4" pageNumber="442" startId="4.[113,148,1330,1352]" targetBox="[111,1450,189,1314]" targetPageId="4" targetType="figure">
<paragraph id="177C3F57230EFFDEFFC45E54AFD0FA4F" blockId="4.[113,1452,1330,1409]" pageId="4" pageNumber="442">
Fig. 2. A,
<taxonomicName id="D0C344D4230EFFDEFF695E54A97CFA87" authority="Saurin, 1953" authorityName="Saurin" authorityYear="1953" box="[220,565,1330,1352]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="pahpetensis">
<emphasis id="25B7E345230EFFDEFF695E54AAE2FA88" box="[220,427,1330,1351]" italics="true" pageId="4" pageNumber="442">Boysidia pahpetensis</emphasis>
<bibRefCitation id="735242A6230EFFDEFE005E54A97CFA87" author="Saurin E" box="[437,565,1330,1352]" pageId="4" pageNumber="442" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
</taxonomicName>
, syntype MNHN-IM-2000-33880 from Pah Hia. B,
<taxonomicName id="D0C344D4230EFFDEFBF65E54AE57FA88" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[1091,1310,1330,1351]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230EFFDEFBF65E54AE57FA88" box="[1091,1310,1330,1351]" italics="true" pageId="4" pageNumber="442">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230EFFDEFA9B5E54AEEEFA87" box="[1326,1447,1330,1352]" pageId="4" pageNumber="442" rank="species">new species</taxonomicNameLabel>
, paratype CUMZ 14421 from Samtay, Houaphanh, Laos. Abbreviations:
<emphasis id="25B7E345230EFFDEFC9B5E29A80AFAAB" bold="true" box="[814,835,1359,1380]" pageId="4" pageNumber="442">b:</emphasis>
basal plica,
<emphasis id="25B7E345230EFFDEFC0B5E29A898FAAB" bold="true" box="[958,977,1359,1380]" pageId="4" pageNumber="442">c:</emphasis>
columellar lamella,
<emphasis id="25B7E345230EFFDEFB225E28AFFAFAAB" bold="true" box="[1175,1203,1358,1380]" pageId="4" pageNumber="442">ip:</emphasis>
infraparietal lamella,
<emphasis id="25B7E345230EFFDEFA3C5E29AEE5FAAB" bold="true" box="[1417,1452,1359,1380]" pageId="4" pageNumber="442">lpl:</emphasis>
lower palatal plica,
<emphasis id="25B7E345230EFFDEFE805E0DAA03FA4F" bold="true" box="[309,330,1387,1408]" pageId="4" pageNumber="442">p:</emphasis>
parietal lamella,
<emphasis id="25B7E345230EFFDEFE435E0DA95DFA4F" bold="true" box="[502,532,1387,1408]" pageId="4" pageNumber="442">pt:</emphasis>
palatal tubercle,
<emphasis id="25B7E345230EFFDEFD0B5E0DA9A1FA4F" bold="true" box="[702,744,1387,1408]" pageId="4" pageNumber="442">upl:</emphasis>
upper palatal plica. Photo: M. Caballer (B).
</paragraph>
</caption>
<subSubSection id="5FD96CDC230EFFDCFFC45C9EA968FE5A" lastPageId="6" lastPageNumber="444" pageId="4" pageNumber="442" type="diagnosis">
<paragraph id="177C3F57230EFFDEFFC45C9EA9A6F7FF" blockId="4.[112,752,2040,2096]" pageId="4" pageNumber="442">
<emphasis id="25B7E345230EFFDEFFC45C9EAA22F7DF" bold="true" box="[113,363,2040,2064]" pageId="4" pageNumber="442">Differential diagnosis.</emphasis>
<taxonomicName id="D0C344D4230EFFDEFEDB5C9EA906F7DF" authorityName="Saurin" authorityYear="1953" box="[366,591,2040,2064]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="pahpetensis">
<emphasis id="25B7E345230EFFDEFEDB5C9EA906F7DF" box="[366,591,2040,2064]" italics="true" pageId="4" pageNumber="442">Boysidia pahpetensis</emphasis>
</taxonomicName>
(
<figureCitation id="8FF823D2230EFFDEFDE85C9EA9F9F7DE" box="[605,688,2040,2065]" captionStart="Fig" captionStartId="4.[113,148,1330,1352]" captionTargetBox="[111,1450,189,1314]" captionTargetId="figure-10@4.[111,1450,189,1314]" captionTargetPageId="4" captionText="Fig. 2. A, Boysidia pahpetensis Saurin, 1953, syntype MNHN-IM-2000-33880 from Pah Hia. B, Boysidia houaphanica, new species, paratype CUMZ 14421 from Samtay, Houaphanh, Laos. Abbreviations: b: basal plica, c: columellar lamella, ip: infraparietal lamella, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (B)." figureDoi="http://doi.org/10.5281/zenodo.14682801" httpUri="https://zenodo.org/record/14682801/files/figure.png" pageId="4" pageNumber="442">Fig. 2A</figureCitation>
) from
<collectingCountry id="6FD47FC7230EFFDEFFC5537FABEDF7FF" box="[112,164,2073,2096]" name="Laos" pageId="4" pageNumber="442">Laos</collectingCountry>
differs from the new species in having a conic shell,
</paragraph>
<paragraph id="177C3F57230EFFDEFC985EDEA8C5F95F" blockId="4.[813,1454,1464,1680]" pageId="4" pageNumber="442">
four whorls, rounded aperture, small infraparietal lamella, and a concrescent point situated close to the peristome margin (
<bibRefCitation id="735242A6230EFFDEFC3D5E9FAF5EF9DE" author="Ancey CF" box="[904,1047,1529,1553]" pageId="4" pageNumber="442" pagination="373 - 374" refId="ref6651" refString="Ancey CF (1881) Description de mollusques terrestres nouveaux. Le Naturaliste, 3 (47): 373 - 374." type="journal article" year="1881">Ancey, 1881</bibRefCitation>
;
<bibRefCitation id="735242A6230EFFDEFB915E9EAE57F9DE" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[1060,1310,1528,1553]" pageId="4" pageNumber="442" pagination="1 - 166" refId="ref6956" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
). In contrast, the new species possesses an elongate conical shape, five to six whorls, a sub-rectangular aperture, without infraparietal lamella, and a concrescent point situated deeper inside the aperture.
</paragraph>
<paragraph id="177C3F57230EFFDEFC985DDEAE52F7DF" blockId="4.[811,1452,1720,2064]" pageId="4" pageNumber="442">
<taxonomicName id="D0C344D4230EFFDEFC985DDEAE58F91E" authority="Bavay &amp; Dautzenberg, 1912" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[813,1297,1720,1745]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="robusta">
<emphasis id="25B7E345230EFFDEFC985DDEA8A9F91F" box="[813,992,1720,1744]" italics="true" pageId="4" pageNumber="442">Boysidia robusta</emphasis>
<bibRefCitation id="735242A6230EFFDEFC505DDFAE58F91E" author="Bavay A &amp; Dautzenberg P" box="[997,1297,1720,1745]" pageId="4" pageNumber="442" pagination="1 - 54" refId="ref6674" refString="Bavay A &amp; Dautzenberg P (1912) Description de coquilles nouvelles de l Indo-Chine. Journal de Conchyliologie, 60 (1): 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="6FD47FC7230EFFDEFAE55DDEAEE5F91F" box="[1360,1452,1720,1744]" name="Vietnam" pageId="4" pageNumber="442">Vietnam</collectingCountry>
and
<taxonomicName id="D0C344D4230EFFDEFCEB5DBEAE24F93E" authority="Panha &amp; Burch, 2002" authorityName="Panha &amp; Burch" authorityYear="2002" box="[862,1389,1752,1777]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="chiangmaiensis">
<emphasis id="25B7E345230EFFDEFCEB5DBEAF26F93F" box="[862,1135,1752,1776]" italics="true" pageId="4" pageNumber="442">Boysidia chiangmaiensis</emphasis>
Panha &amp; Burch, 2002
</taxonomicName>
from
<collectingCountry id="6FD47FC7230EFFDEFC985D9EA8DBF8DF" box="[813,914,1784,1808]" name="Thailand" pageId="4" pageNumber="442">Thailand</collectingCountry>
differ from the new species in having ovate conic and large shells (height
<quantity id="D03B92B2230EFFDEFBC75C7EAF84F8FE" box="[1138,1229,1816,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="4" pageNumber="442" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
) with basal plica. In addition,
<taxonomicName id="D0C344D4230EFFDEFC215C5EAF05F89F" authorityName="Bavay &amp; Dautzenberg" authorityYear="1912" box="[916,1100,1848,1872]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="robusta">
<emphasis id="25B7E345230EFFDEFC215C5EAF05F89F" box="[916,1100,1848,1872]" italics="true" pageId="4" pageNumber="442">Boysidia robusta</emphasis>
</taxonomicName>
also has an ovate aperture, and with small infraparietal lamella and lower basal plica, while
<taxonomicName id="D0C344D4230EFFDEFC985C1EAF0BF85F" authorityName="Panha &amp; Burch" authorityYear="2002" box="[813,1090,1912,1936]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="chiangmaiensis">
<emphasis id="25B7E345230EFFDEFC985C1EAF0BF85F" box="[813,1090,1912,1936]" italics="true" pageId="4" pageNumber="442">Boysidia chiangmaiensis</emphasis>
</taxonomicName>
has a rounded aperture (
<bibRefCitation id="735242A6230EFFDEFAD15C1FAF6BF87E" author="Bavay A &amp; Dautzenberg P" pageId="4" pageNumber="442" pagination="1 - 54" refId="ref6674" refString="Bavay A &amp; Dautzenberg P (1912) Description de coquilles nouvelles de l Indo-Chine. Journal de Conchyliologie, 60 (1): 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
;
<bibRefCitation id="735242A6230EFFDEFB845CFEAE7FF87E" author="Panha S &amp; Burch JB" box="[1073,1334,1944,1969]" pageId="4" pageNumber="442" pagination="117 - 122" refId="ref7719" refString="Panha S &amp; Burch JB (2002 a) First records and new species of Boysidia and Sinoennea from Thailand. Malacological Review, 31 / 32 [1998 - 1999] (2): 117 - 122. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2002">Panha &amp; Burch, 2002a</bibRefCitation>
,
<bibRefCitation id="735242A6230EFFDEFAF15CFEAE36F87F" author="Panha S &amp; Burch JB" box="[1348,1407,1944,1968]" pageId="4" pageNumber="442" pagination="2004 - 2005" refId="ref7868" refString="Panha S &amp; Burch JB (2005) An introduction to the microsnails of Thailand. Malacological Review, 37 / 38 [2004 - 2005]: 1 - 155. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2005">2005</bibRefCitation>
). In comparison,
<taxonomicName id="D0C344D4230EFFDEFC025CDEAFEEF81F" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[951,1191,1976,2000]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="442" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230EFFDEFC025CDEAFEEF81F" box="[951,1191,1976,2000]" italics="true" pageId="4" pageNumber="442">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230EFFDEFB035CDFAE73F81F" box="[1206,1338,1976,2000]" pageId="4" pageNumber="442" rank="species">new species</taxonomicNameLabel>
, possesses an elongate conical, smaller shell (height
<quantity id="D03B92B2230EFFDEFAA15CBEAEE9F83E" box="[1300,1440,2008,2033]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.5" metricValueMin="3.1" pageId="4" pageNumber="442" unit="mm" value="3.3" valueMax="3.5" valueMin="3.1">3.13.5 mm</quantity>
), subrectangular aperture, and with basal plica.
</paragraph>
<caption id="43BC6FDF230FFFDFFF3D5C08AA99F814" ID-DOI="http://doi.org/10.5281/zenodo.14682805" ID-Zenodo-Dep="14682805" httpUri="https://zenodo.org/record/14682805/files/figure.png" pageId="5" pageNumber="443" startId="5.[136,171,1902,1924]" targetBox="[136,1474,293,1887]" targetPageId="5" targetType="figure">
<paragraph id="177C3F57230FFFDFFF3D5C08AA99F814" blockId="5.[136,1474,1902,2011]" pageId="5" pageNumber="443">
Fig. 3.
<taxonomicName id="D0C344D4230FFFDFFF785C08AAECF84C" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[205,421,1902,1923]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="443" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230FFFDFFF785C08AAECF84C" box="[205,421,1902,1923]" italics="true" pageId="5" pageNumber="443">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230FFFDFFE045C09A961F84B" box="[433,552,1902,1924]" pageId="5" pageNumber="443" rank="species">new species</taxonomicNameLabel>
. AC, holotype CUMZ 14420 from Samtay, Houaphanh, Laos; A, shell and enlarged apertures of different view angles. B, protoconch and C, inner wall of peristome around basal plica showing small nodules. D, paratype CUMZ 14421 from the type locality. Abbreviations:
<emphasis id="25B7E345230FFFDFFDF15CCEA910F872" bold="true" box="[580,601,1960,1981]" pageId="5" pageNumber="443">b:</emphasis>
basal plica,
<emphasis id="25B7E345230FFFDFFD635CCEA9A0F872" bold="true" box="[726,745,1960,1981]" pageId="5" pageNumber="443">c:</emphasis>
columellar lamella,
<emphasis id="25B7E345230FFFDFFC075CCEA89CF872" bold="true" box="[946,981,1960,1981]" pageId="5" pageNumber="443">lpl:</emphasis>
lower palatal plica,
<emphasis id="25B7E345230FFFDFFB2B5CCEAFFAF872" bold="true" box="[1182,1203,1960,1981]" pageId="5" pageNumber="443">p:</emphasis>
parietal lamella,
<emphasis id="25B7E345230FFFDFFAEA5CCEAE35F872" bold="true" box="[1375,1404,1960,1981]" pageId="5" pageNumber="443">pt:</emphasis>
palatal tubercle,
<emphasis id="25B7E345230FFFDFFF565CA3AA45F815" bold="true" box="[227,268,1989,2010]" pageId="5" pageNumber="443">upl:</emphasis>
upper palatal plica.
</paragraph>
</caption>
<paragraph id="177C3F57230CFFDCFFC45BD8A968FE5A" blockId="6.[113,752,189,405]" pageId="6" pageNumber="444">
For further comparison,
<taxonomicName id="D0C344D4230CFFDCFE305BDBABFCFF3A" authority="Panha &amp; Burch, 2002 b" authorityName="Panha &amp; Burch" authorityYear="2002" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="tholos">
<emphasis id="25B7E345230CFFDCFE305BDBA978FF1A" box="[389,561,189,213]" italics="true" pageId="6" pageNumber="444">Boysidia tholos</emphasis>
<bibRefCitation id="735242A6230CFFDCFD8E5BDBABFCFF3A" author="Panha S &amp; Burch JB" pageId="6" pageNumber="444" pagination="113 - 134" refId="ref7772" refString="Panha S &amp; Burch JB (2002 b) New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana, 10 [1999] (24): 113 - 134. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2002">Panha &amp; Burch, 2002b</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="6FD47FC7230CFFDCFF485BBBAA14FF3A" box="[253,349,221,245]" name="Thailand" pageId="6" pageNumber="444">Thailand</collectingCountry>
can be distinguished from this new species by having a dome-shaped shell, five whorls, rounded aperture, and infrapalatal lamella. In contrast,
<taxonomicName id="D0C344D4230CFFDCFD3A5A7BABB5FE9A" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230CFFDCFD3A5A7BABB5FE9A" italics="true" pageId="6" pageNumber="444">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230CFFDCFEBF5A58AAC3FE9A" box="[266,394,317,341]" pageId="6" pageNumber="444" rank="species">new species</taxonomicNameLabel>
has an elongated conic shell, 56 whorls, subrectangular aperture, and without infrapalatal lamella (
<bibRefCitation id="735242A6230CFFDCFF655A1BAA87FE5A" author="Panha S &amp; Burch JB" box="[208,462,381,405]" pageId="6" pageNumber="444" pagination="113 - 134" refId="ref7772" refString="Panha S &amp; Burch JB (2002 b) New taxa of Pupillidae (Pulmonata: Stylommatophora) from Thailand. Walkerana, 10 [1999] (24): 113 - 134. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2002">Panha &amp; Burch, 2002b</bibRefCitation>
,
<bibRefCitation id="735242A6230CFFDCFE6E5A1BA95CFE5A" author="Panha S &amp; Burch JB" box="[475,533,381,405]" pageId="6" pageNumber="444" pagination="2004 - 2005" refId="ref7868" refString="Panha S &amp; Burch JB (2005) An introduction to the microsnails of Thailand. Malacological Review, 37 / 38 [2004 - 2005]: 1 - 155. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2005">2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFFC45ADBAAC4FDDA" pageId="6" pageNumber="444" type="etymology">
<paragraph id="177C3F57230CFFDCFFC45ADBAAC4FDDA" blockId="6.[113,752,445,533]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFFC45ADBABBFFE1A" bold="true" box="[113,246,445,469]" pageId="6" pageNumber="444">Etymology.</emphasis>
The species name
<taxonomicName id="D0C344D4230CFFDCFE605ADBA924FE1A" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[469,621,445,469]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="houaphanica">
<emphasis id="25B7E345230CFFDCFE6B5ADBA924FE1A" box="[478,621,445,469]" italics="true" pageId="6" pageNumber="444">houaphanica</emphasis>
</taxonomicName>
is derived from the
<typeStatus id="C87881F5230CFFDCFF665AB8AA48FE39" box="[211,257,478,502]" pageId="6" pageNumber="444">type</typeStatus>
locality,
<collectingRegion id="D507F1B5230CFFDCFED05ABBA900FE3A" box="[357,585,477,501]" country="Laos" name="Houaphan" pageId="6" pageNumber="444">Houaphanh Province</collectingRegion>
, where the
<typeStatus id="C87881F5230CFFDCFD775AB8A9B9FE39" box="[706,752,478,502]" pageId="6" pageNumber="444">type</typeStatus>
specimens were collected.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFFC4595BAA30FD7A" pageId="6" pageNumber="444" type="distribution">
<paragraph id="177C3F57230CFFDCFFC4595BAA30FD7A" blockId="6.[113,753,573,693]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFFC4595BAA43FD9A" bold="true" box="[113,266,573,597]" pageId="6" pageNumber="444">Distribution.</emphasis>
This species,
<taxonomicName id="D0C344D4230CFFDCFE01595BA9F8FD9A" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[436,689,573,597]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230CFFDCFE01595BA9F8FD9A" box="[436,689,573,597]" italics="true" pageId="6" pageNumber="444">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230CFFDCFD765958AB88FDBA" pageId="6" pageNumber="444" rank="species">new species</taxonomicNameLabel>
, is known only from the
<typeStatus id="C87881F5230CFFDCFE785938AAB2FDB9" box="[461,507,606,630]" pageId="6" pageNumber="444">type</typeStatus>
locality in
<collectingRegion id="D507F1B5230CFFDCFDC0593BAB9DFD5A" country="Laos" name="Houaphan" pageId="6" pageNumber="444">Houaphanh Province</collectingRegion>
,
<collectingCountry id="6FD47FC7230CFFDCFF545918AA51FD5A" box="[225,280,638,661]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
. The snails were found under leaf litter in limestone rock crevices.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFFC659BBABB5FC5A" pageId="6" pageNumber="444" type="discussion">
<paragraph id="177C3F57230CFFDCFFC659BBABB5FC5A" blockId="6.[112,752,733,917]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFFC659BBABAEFD3A" bold="true" box="[115,231,733,757]" pageId="6" pageNumber="444">Remarks.</emphasis>
This new species occurs sympatrically with
<taxonomicName id="D0C344D4230CFFDCFFC4599BAAD2FCDA" authorityName="Xaisomboun" baseAuthorityName="Saurin" baseAuthorityYear="1953" box="[113,411,765,789]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E345230CFFDCFFC4599BAAD2FCDA" box="[113,411,765,789]" italics="true" pageId="6" pageNumber="444">Bensonella novemdentata</emphasis>
</taxonomicName>
. The occurrence of multiple hypselostomatid species in the same locality is not uncommon (e.g.,
<bibRefCitation id="735242A6230CFFDCFF1B585BAAEEFC9A" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[174,423,829,853]" pageId="6" pageNumber="444" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay et al., 2016</bibRefCitation>
;
<bibRefCitation id="735242A6230CFFDCFE06585BA9F9FC9A" author="Vermeulen JJ &amp; Luu HT &amp; Keum T &amp; Anker K" box="[435,688,829,853]" pageId="6" pageNumber="444" pagination="7 - 41" refId="ref8868" refString="Vermeulen JJ, Luu HT, Keum T &amp; Anker K (2019) New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam). Folia Malacologica, 27: 7 - 41." type="journal article" year="2019">Vermeulen et al., 2019</bibRefCitation>
;
<bibRefCitation id="735242A6230CFFDCFD09585BAA3AFCBA" author="Pall-Gergely B &amp; White TS" pageId="6" pageNumber="444" pagination="2011 - 2029" refId="ref7633" refString="Pall-Gergely B &amp; White TS (2022) Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History, 56 (45 - 48): 2011 - 2029." type="journal article" year="2022">Páll-Gergely &amp; White, 2022</bibRefCitation>
;
<bibRefCitation id="735242A6230CFFDCFECA583BA92AFCBA" author="Sutcharit C &amp; Ngor PB &amp; Pall-Gergely B &amp; Jeratthitikul E &amp; Siriwut W &amp; Srisonchai R &amp; Ng TH &amp; Jirapatrasilp P &amp; Panha S" box="[383,611,861,885]" pageId="6" pageNumber="444" pagination="1287 - 1303" refId="ref8467" refString="Sutcharit C, Ngor PB, Pall-Gergely B, Jeratthitikul E, Siriwut W, Srisonchai R, Ng TH, Jirapatrasilp P &amp; Panha S (2023) Notes on the hypselostomatid snails (Gastropoda: Heterobranchia) from limestone hills in Western Cambodia with a new record and a new species. Journal of Natural History, 57 (25 - 28): 1287 - 1303." type="journal article" year="2023">Sutcharit et al., 2023</bibRefCitation>
;
<bibRefCitation id="735242A6230CFFDCFDDB583BABB8FC5A" author="Jirapatrasilp P &amp; Panha S &amp; Sutcharit C &amp; Tongkerd P" pageId="6" pageNumber="444" pagination="87 - 98" refId="ref7082" refString="Jirapatrasilp P, Panha S, Sutcharit C &amp; Tongkerd P (2024) Four new species of micro land snail in the genus Bensonella Pilsbry &amp; Vanatta, 1900 (Stylommatophora: Hypselostomatidae) from Thailand. Archiv fur Molluskenkunde, 153 (1): 87 - 98." type="journal article" year="2024">Jirapatrasilp et al., 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,353 @@
<document id="70326D162DAC690700E351DDEB377E55" ID-DOI="10.26107/RBZ-2024-0032" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682797" ID-ZooBank="D7A0B570-07AC-47F5-9B56-88EBDB277EC0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171734098" checkinUser="felipe" docAuthor="Inkhavilay, Khamla, Sutcharit, &amp; Chirasak" docDate="2024" docId="9F6A8E41230CFFD2FF045C05AAB4FA7A" docLanguage="en" docName="RafflesBZool.72.438-449.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Bensonella novemdentata Inkhavilay &amp; Sutcharit &amp; Chirasak 2024, new combination" docType="treatment" docVersion="2" lastPageNumber="446" masterDocId="6353F639230AFFDAFFB55B66AB49FFCF" masterDocTitle="A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos" masterLastPageNumber="449" masterPageNumber="438" pageNumber="444" updateTime="1737175246880" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="39CDB3E389EF6DEF7C9039A1A3683C80" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5FA4D2D8E0409F282F61827506477ADC">
<mods:title id="735C18470D779C2D14538CEE7EBA9043">A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos</mods:title>
</mods:titleInfo>
<mods:name id="A9481158D96766493648757C54B2611C" type="personal">
<mods:role id="2379DD941A311B48FAE20459F33F703E">
<mods:roleTerm id="166399EA4DDF44A69018E457C3D369DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AE323E74D158BD628A39ADE2DD66F9A5">Inkhavilay, Khamla</mods:namePart>
</mods:name>
<mods:name id="B30FB082E511A0B70B3ADDDB2372F1AA" type="personal">
<mods:role id="A9E2B8A5A10DEC5B3F79623B360E4382">
<mods:roleTerm id="965802C5D30481089773008AC19011DC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="92424C7D82D3F46FE8F27CA1517C2C88">Sutcharit,</mods:namePart>
</mods:name>
<mods:name id="25856C41829237D98DC91047C7BE9D7A" type="personal">
<mods:role id="A935EA1400D26F15A9207698FE5A8971">
<mods:roleTerm id="079E78CA8D03737EF7D88857E7148B40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DA41BD175EF76727F39D2884B1D5DCD9">Chirasak</mods:namePart>
</mods:name>
<mods:typeOfResource id="3E1F75F9FBF26C8A9B9D0ADE1FAE159E">text</mods:typeOfResource>
<mods:relatedItem id="B5C2483184E20F45E906B0352DCD019F" type="host">
<mods:titleInfo id="17E824D112842AB5D0B8CEABB0C819B9">
<mods:title id="D8CB812A85AE2387B6D53E454E0F70CC">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="F3555C881AFC198D641C6C747E677E74">
<mods:date id="56C45E9BE28A4F6AE02E4B6BE9B50BFC">2024</mods:date>
<mods:detail id="04942F70F59865E80F38AC76799AAC7C" type="pubDate">
<mods:number id="F9AF8850E5213108949DF6A5F587954D">2024-11-04</mods:number>
</mods:detail>
<mods:detail id="952C630865CEDD1D47923572D8516056" type="volume">
<mods:number id="6E301B49AEDFD28C79D28D4B547D3DCD">72</mods:number>
</mods:detail>
<mods:extent id="71B34C355AD446F76BD83E1329BD5F74" unit="page">
<mods:start id="B039B25CDA07B2F28D457E0A0351F373">438</mods:start>
<mods:end id="7AC16092B07D737F9360A1B89454D94D">449</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4D7B2485C997B70602D537BC8C235798">journal article</mods:classification>
<mods:identifier id="8E0F0DF2256F75748521F9A2CA5D7D75" type="DOI">10.26107/RBZ-2024-0032</mods:identifier>
<mods:identifier id="527BA9D305089BAB4558B527EF3686F1" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="3CD0A8FE83A7C2979B219B8282148FC2" type="Zenodo-Dep">14682797</mods:identifier>
<mods:identifier id="F76657F63CE1C2607E24B6A26DB81C17" type="ZooBank">D7A0B570-07AC-47F5-9B56-88EBDB277EC0</mods:identifier>
</mods:mods>
<treatment id="9F6A8E41230CFFD2FF045C05AAB4FA7A" ID-DOI="http://doi.org/10.5281/zenodo.14682803" ID-Zenodo-Dep="14682803" LSID="urn:lsid:plazi:treatment:9F6A8E41230CFFD2FF045C05AAB4FA7A" httpUri="http://treatment.plazi.org/id/9F6A8E41230CFFD2FF045C05AAB4FA7A" lastPageId="8" lastPageNumber="446" pageId="6" pageNumber="444">
<subSubSection id="5FD96CDC230CFFDCFF045C05AAB3F854" pageId="6" pageNumber="444" type="nomenclature">
<paragraph id="177C3F57230CFFDCFF045C05AAB3F854" blockId="6.[177,689,1891,1980]" pageId="6" pageNumber="444">
<heading id="4C34883B230CFFDCFF045C05AAB3F854" bold="true" centered="true" fontSize="10" level="2" pageId="6" pageNumber="444" reason="2">
<emphasis id="25B7E345230CFFDCFF045C05AAB3F854" bold="true" pageId="6" pageNumber="444">
<taxonomicName id="D0C344D4230CFFDCFF045C05A932F8B4" ID-CoL="CPT5Q" authority="(Saurin, 1953)" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" baseAuthorityName="Saurin" baseAuthorityYear="1953" box="[177,635,1891,1915]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="novemdentata" status="comb. nov.">
<emphasis id="25B7E345230CFFDCFF045C05AA87F8B4" bold="true" box="[177,462,1891,1915]" italics="true" pageId="6" pageNumber="444">Bensonella novemdentata</emphasis>
(
<bibRefCitation id="735242A6230CFFDCFE6B5C05A93AF8B4" author="Saurin E" box="[478,627,1891,1915]" pageId="6" pageNumber="444" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="3E845E3E230CFFDCFD365C02AAB3F854" pageId="6" pageNumber="444" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFE3D5CC2AA93F873" box="[392,474,1956,1980]" pageId="6" pageNumber="444" type="description">
<paragraph id="177C3F57230CFFDCFE3D5CC2AA93F873" blockId="6.[177,689,1891,1980]" box="[392,474,1956,1980]" pageId="6" pageNumber="444">
(
<figureCitation id="8FF823D2230CFFDCFE255CC2AA9BF873" box="[400,466,1956,1980]" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="6" pageNumber="444">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFFC45C84AF0FFEC2" pageId="6" pageNumber="444" type="reference_group">
<paragraph id="177C3F57230CFFDCFFC45C84AF0FFEC2" blockId="6.[113,752,2018,2098]" lastBlockId="6.[849,1452,189,269]" pageId="6" pageNumber="444">
<treatmentCitationGroup id="37D31879230CFFDCFFC45C84AF0FFEC2" pageId="6" pageNumber="444">
<treatmentCitation id="96621946230CFFDCFFC45C84A959F837" author="Saurin E" box="[113,528,2018,2040]" page="115" pageId="6" pageNumber="444" year="1953">
<taxonomicName id="D0C344D4230CFFDCFFC45C84A959F837" ID-CoL="7QD4K" authority="Saurin, 1953: 115" authorityName="Saurin" authorityPageNumber="115" authorityYear="1953" box="[113,528,2018,2040]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E345230CFFDCFFC45C84AA1CF838" box="[113,341,2018,2039]" italics="true" pageId="6" pageNumber="444">Boysidia novemdentata</emphasis>
<bibRefCitation id="735242A6230CFFDCFEE85C84A959F837" author="Saurin E" box="[349,528,2018,2040]" pageId="6" pageNumber="444" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953: 115</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 116, fig. 1, and pl. 4, fig. 4ac.
<typeStatus id="C87881F5230CFFDCFF4C5366AA63F7DA" box="[249,298,2048,2069]" pageId="6" pageNumber="444">Type</typeStatus>
locality: environs du village méo de Pah Hia, à 100 kilomètres au Sud de Xieng-Khouang, chef-lieu de la province du Tran Ninh,
<collectingCountry id="6FD47FC7230CFFDCFB875BD8AF29FF1C" box="[1074,1120,190,211]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
[probably refers to Ban Namthong, Longchaeng District, Xaisomboun Province,
<collectingCountry id="6FD47FC7230CFFDCFAB45BBDAE7CFF3F" box="[1281,1333,219,240]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
].
<treatmentCitation id="96621946230CFFDCFAF65BBCA8A0FEC2" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" httpUri="http://treatment.plazi.org/id/C12AA132AABEB5977B6FF52156ECCCF4" page="59" pageId="6" pageNumber="444" year="2019">
<bibRefCitation id="735242A6230CFFDCFAF65BBCA8A0FEC2" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" pageId="6" pageNumber="444" pagination="1 - 166" refId="ref6956" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019: 59</bibRefCitation>
</treatmentCitation>
, fig. 26b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFC985A53A820FE22" pageId="6" pageNumber="444" type="materials_examined">
<paragraph id="177C3F57230CFFDCFC985A53A820FE22" blockId="6.[813,1452,309,493]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFC985A53AF5AFE82" bold="true" box="[813,1043,309,333]" pageId="6" pageNumber="444">Material examined.</emphasis>
<materialsCitation id="A7AB350A230CFFDCFBA85A53A82FFE22" collectingDate="2022-05" collectionCode="CUMZ" collectorName="K. Inkhavilay &amp; Inkhavilay, A &amp; C. Oulaiseng" country="Laos" latitude="20.014555" location="Ban Lao" longLatPrecision="1" longitude="104.682495" municipality="Samtay District" pageId="6" pageNumber="444" specimenCode="CUMZ 14426" specimenCount="1" stateProvince="Houaphan" typeStatus="syntype">
<typeStatus id="C87881F5230CFFDCFBA85A53AF3FFE82" box="[1053,1142,309,333]" pageId="6" pageNumber="444" type="syntype">Syntype</typeStatus>
MNHN-IM-2000-33881 (1 shell;
<figureCitation id="8FF823D2230CFFDCFCC55A33A88CFEA2" box="[880,965,341,365]" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="6" pageNumber="444">Fig. 4A</figureCitation>
) from Pah Hia.
<specimenCode id="4765972C230CFFDCFBC95A33AE5CFEA2" box="[1148,1301,341,365]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="6" pageNumber="444">CUMZ 14426</specimenCode>
(1 shell;
<figureCitation id="8FF823D2230CFFDCFA345A33A804FE42" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="6" pageNumber="444">Fig. 4B</figureCitation>
) from limestone outcrop at
<location id="121C698C230CFFDCFB3C5A10AFA2FE42" LSID="urn:lsid:plazi:treatment:9F6A8E41230CFFD2FF045C05AAB4FA7A:121C698C230CFFDCFB3C5A10AFA2FE42" box="[1161,1259,374,397]" country="Laos" latitude="20.014555" longLatPrecision="1" longitude="104.682495" municipality="Samtay District" name="Ban Lao" pageId="6" pageNumber="444" stateProvince="Houaphan">Ban Lao</location>
,
<collectingMunicipality id="F718A52D230CFFDCFB4C5A13AEEEFE42" box="[1273,1447,373,397]" pageId="6" pageNumber="444">Samtay District</collectingMunicipality>
,
<collectingRegion id="D507F1B5230CFFDCFC985AF3AF59FE62" box="[813,1040,405,429]" country="Laos" name="Houaphan" pageId="6" pageNumber="444">Houaphanh Province</collectingRegion>
,
<collectingCountry id="6FD47FC7230CFFDCFBAE5AF0AF06FE62" box="[1051,1103,406,429]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
(
<geoCoordinate id="72F75990230CFFDCFBE85AF2AFBAFE62" box="[1117,1267,404,429]" degrees="20" direction="north" minutes="00" orientation="latitude" pageId="6" pageNumber="444" precision="1" seconds="52.4" value="20.014555">20°0052.4″N</geoCoordinate>
,
<geoCoordinate id="72F75990230CFFDCFB4A5AF2AEE9FE62" box="[1279,1440,404,429]" degrees="104" direction="east" minutes="40" orientation="longitude" pageId="6" pageNumber="444" precision="1" seconds="57.0" value="104.682495">104°4057.0″E</geoCoordinate>
), coll.
<collectorName id="BA365A81230CFFDCFCD05AD0A8B7FE02" box="[869,1022,437,461]" pageId="6" pageNumber="444">K. Inkhavilay</collectorName>
, A. Inthavong, and
<collectorName id="BA365A81230CFFDCFB695AD3AE24FE02" box="[1244,1389,437,461]" pageId="6" pageNumber="444">C. Oulaiseng</collectorName>
,
<date id="637D1997230CFFDCFACF5AD0A82FFE22" pageId="6" pageNumber="444" value="2022-05">
<collectingDate id="7339E07F230CFFDCFACF5AD0A82FFE22" pageId="6" pageNumber="444" value="2022-05">May 2022</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFC985973AECEFD42" pageId="6" pageNumber="444" type="diagnosis">
<paragraph id="177C3F57230CFFDCFC985973AECEFD42" blockId="6.[812,1452,533,653]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFC985973A8E8FDE2" bold="true" box="[813,929,533,557]" pageId="6" pageNumber="444">Diagnosis.</emphasis>
Shell elongate-conic, yellowish brown in colour. Cervical crest situated far from peristome. Aperture with eight teeth (i.e., parietal, infraparietal, angular, columellar, basal, and upper, inter and lower palatals), not hooked.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFC9859D3AE54FAC2" pageId="6" pageNumber="444" type="description">
<paragraph id="177C3F57230CFFDCFC9859D3AE54FAC2" blockId="6.[811,1453,693,1293]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFC9859D3A8F7FD02" bold="true" box="[813,958,693,717]" pageId="6" pageNumber="444">Description.</emphasis>
Shell conical, yellowish brown to reddish brown with 5¼ to 6 widely convex whorls. Apex rather blunt; protoconch about 1½ whorls with wrinkles, very weak spiral ridges. Teleoconch with strong, irregular growth lines and conspicuous; suture impressed and deep. Last whorl large, rounded. Peristome thickened, slightly expanded; constriction weak; lip reddish brown. Cervical crest rather sharp, situated close to peristome. Aperture subquadrate with many apertural dentitions. Parietal lamella (p) strongly developed, rectangular. Infraparietal lamella (ip) small and blunt. Angular lamella (a) slightly higher than parietal lamella, sinuous: outer part short, low, reaching peristome; inner part strongly elevated. Palatal tubercle (pt) strongly developed with triangular shape; one upper palatal plica (upl) with strong folding; one inter- (itpl) and one lower (lpl) palatal relatively large, nearly equal in size, strong plica. Basal plica (b) small, low, tubercle-like. Columellar lamella (c) large, strong, distinct horizontally. Umbilicus narrowly perforate, about one-fifth of shell width, rounded, deep.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFDCFC985E53A8FFFA22" pageId="6" pageNumber="444" type="distribution">
<paragraph id="177C3F57230CFFDCFC985E53A8FFFA22" blockId="6.[812,1453,1333,1517]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFC985E53A8F6FA82" bold="true" box="[813,959,1333,1357]" pageId="6" pageNumber="444">Distribution.</emphasis>
This species was previously known only from the
<typeStatus id="C87881F5230CFFDCFCE15E30A8CBFAA1" box="[852,898,1366,1390]" pageId="6" pageNumber="444">type</typeStatus>
locality (
<bibRefCitation id="735242A6230CFFDCFC5C5E33AF3FFAA2" author="Saurin E" box="[1001,1142,1365,1389]" pageId="6" pageNumber="444" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
;
<bibRefCitation id="735242A6230CFFDCFB375E33AE3FFAA2" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" box="[1154,1398,1365,1389]" pageId="6" pageNumber="444" pagination="1 - 166" refId="ref6956" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
) and is newly recorded from
<collectingRegion id="D507F1B5230CFFDCFB8A5E13AE6FFA42" box="[1087,1318,1397,1421]" country="Laos" name="Houaphan" pageId="6" pageNumber="444">Houaphanh Province</collectingRegion>
in northern
<collectingCountry id="6FD47FC7230CFFDCFC985EF0A82DFA62" box="[813,868,1430,1453]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
. The snails live on limestone karsts and can be found on the limestone wall together with
<taxonomicName id="D0C344D4230CFFDCFB005ED3AEECFA02" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" box="[1205,1445,1461,1485]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="houaphanica" status="sp. nov.">
<emphasis id="25B7E345230CFFDCFB005ED3AEECFA02" box="[1205,1445,1461,1485]" italics="true" pageId="6" pageNumber="444">Boysidia houaphanica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3E845E3E230CFFDCFC985EB0A8FBFA22" box="[813,946,1493,1517]" pageId="6" pageNumber="444" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="5FD96CDC230CFFD2FC985D73AAB4FA7A" lastPageId="8" lastPageNumber="446" pageId="6" pageNumber="444" type="discussion">
<paragraph id="177C3F57230CFFDCFC985D73AF04F8C1" blockId="6.[813,1452,1557,1806]" pageId="6" pageNumber="444">
<emphasis id="25B7E345230CFFDCFC985D73A8EBF9E2" bold="true" box="[813,930,1557,1581]" pageId="6" pageNumber="444">Remarks.</emphasis>
The recently collected specimens are from
<collectingRegion id="D507F1B5230CFFDCFC985D53AF5EF982" box="[813,1047,1589,1613]" country="Laos" name="Houaphan" pageId="6" pageNumber="444">Houaphanh Province</collectingRegion>
, which is about
<quantity id="D03B92B2230CFFDCFB605D53AE63F982" box="[1237,1322,1589,1613]" metricMagnitude="5" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="444" unit="km" value="200.0">200 km</quantity>
east of the “type locality, which is stated to be Pahai”. However, the morphology of our specimens deviates slightly from that of the
<typeStatus id="C87881F5230CFFDCFC1B5DF0AF4AF961" box="[942,1027,1686,1710]" pageId="6" pageNumber="444" type="syntype">syntype</typeStatus>
(
<figureCitation id="8FF823D2230CFFDCFBA25DF3AF38F962" box="[1047,1137,1685,1709]" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="6" pageNumber="444">Fig. 4A</figureCitation>
) in having a more rounded whorl and deeper suture (
<figureCitation id="8FF823D2230CFFDCFBF35DD3AFD4F902" box="[1094,1181,1717,1741]" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="6" pageNumber="444">Fig. 4B</figureCitation>
). Otherwise, the number and arrangement of the apertural dentitions are practically identical with the
<typeStatus id="C87881F5230CFFDCFC475D90AF01F8C1" box="[1010,1096,1782,1806]" pageId="6" pageNumber="444" type="syntype">syntype</typeStatus>
.
</paragraph>
<paragraph id="177C3F57230CFFD2FC985C53AAF8FE1A" blockId="6.[811,1455,1845,2093]" lastBlockId="8.[112,752,189,469]" lastPageId="8" lastPageNumber="446" pageId="6" pageNumber="444">
Among the morphologically similar species recorded from
<collectingCountry id="6FD47FC7230CFFDCFC985C30A82CF8A2" box="[813,869,1878,1901]" name="Laos" pageId="6" pageNumber="444">Laos</collectingCountry>
,
<taxonomicName id="D0C344D4230CFFDCFCC15C33AFDAF8A2" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" baseAuthorityName="Saurin" baseAuthorityYear="1953" box="[884,1171,1877,1901]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E345230CFFDCFCC15C33AFDAF8A2" box="[884,1171,1877,1901]" italics="true" pageId="6" pageNumber="444">Bensonella novemdentata</emphasis>
</taxonomicName>
differs from
<taxonomicName id="D0C344D4230CFFDCFA845C33AFA3F842" authority="(Bavay &amp; Dautzenberg, 1912)" baseAuthorityName="Bavay &amp; Dautzenberg" baseAuthorityYear="1912" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="paviei">
<emphasis id="25B7E345230CFFDCFA845C33A83DF842" italics="true" pageId="6" pageNumber="444">Bensonella paviei</emphasis>
(
<bibRefCitation id="735242A6230CFFDCFC3D5C13AFABF842" author="Bavay A &amp; Dautzenberg P" box="[904,1250,1909,1933]" pageId="6" pageNumber="444" pagination="1 - 54" refId="ref6674" refString="Bavay A &amp; Dautzenberg P (1912) Description de coquilles nouvelles de l Indo-Chine. Journal de Conchyliologie, 60 (1): 1 - 54." type="journal article" year="1912">Bavay &amp; Dautzenberg, 1912</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D0C344D4230CFFDCFA9E5C13A8F7F862" baseAuthorityName="Inkhavilay &amp; Panha" baseAuthorityYear="2016" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="444" phylum="Mollusca" rank="species" species="anguloobtusa">
<emphasis id="25B7E345230CFFDCFA9E5C13A8F7F862" italics="true" pageId="6" pageNumber="444">Bensonella anguloobtusa</emphasis>
</taxonomicName>
(
<bibRefCitation id="735242A6230CFFDCFC7A5CF3AFBAF862" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[975,1267,1941,1965]" pageId="6" pageNumber="444" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">
Inkhavilay &amp; Panha,
<quantity id="D03B92B2230CFFDCFB0B5CF3AFBAF862" box="[1214,1267,1941,1965]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="6" pageNumber="444" unit="in" value="2016.0">2016</quantity>
</bibRefCitation>
in
<bibRefCitation id="735242A6230CFFDCFAA95CF3A8C4F802" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" pageId="6" pageNumber="444" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay et al., 2016</bibRefCitation>
) by having a prominent cervical crest and palatal plicae forming a short tubercle and situated near the aperture. In comparison, the latter two species have very weak to no cervical crest and palatal plicae extending far inside the aperture;
<taxonomicName id="D0C344D42302FFD2FEBF5BDBA96CFF1A" baseAuthorityName="Inkhavilay &amp; Panha" baseAuthorityYear="2016" box="[266,549,189,213]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="anguloobtusa">
<emphasis id="25B7E3452302FFD2FEBF5BDBA96CFF1A" box="[266,549,189,213]" italics="true" pageId="8" pageNumber="446">Bensonella anguloobtusa</emphasis>
</taxonomicName>
further possesses a subcolumellar plica without lower parietal lamella. In addition,
<taxonomicName id="D0C344D42302FFD2FF695B9BAAB2FEDA" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" baseAuthorityName="Saurin" baseAuthorityYear="1953" box="[220,507,253,277]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E3452302FFD2FF695B9BAAB2FEDA" box="[220,507,253,277]" italics="true" pageId="8" pageNumber="446">Bensonella novemdentata</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="D0C344D42302FFD2FF1B5A7BA982FEFA" authority="(Inkhavilay &amp; Panha, 2016)" baseAuthorityName="Inkhavilay &amp; Panha" baseAuthorityYear="2016" box="[174,715,285,309]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="paralella">
<emphasis id="25B7E3452302FFD2FF1B5A7BAAD9FEFA" box="[174,400,285,309]" italics="true" pageId="8" pageNumber="446">Bensonella paralella</emphasis>
(
<bibRefCitation id="735242A62302FFD2FE155A7BA98DFEFA" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[416,708,285,309]" pageId="8" pageNumber="446" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay &amp; Panha, 2016</bibRefCitation>
)
</taxonomicName>
by having a subrectangular aperture and prominent interpalatal and cervical crest, while
<taxonomicName id="D0C344D42302FFD2FE345A3BA928FEBA" baseAuthorityName="Inkhavilay &amp; Panha" baseAuthorityYear="2016" box="[385,609,349,373]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="paralella">
<emphasis id="25B7E3452302FFD2FE345A3BA928FEBA" box="[385,609,349,373]" italics="true" pageId="8" pageNumber="446">Bensonella paralella</emphasis>
</taxonomicName>
possesses an ovate-shaped aperture, and without interpalatal plica and cervical crest (
<bibRefCitation id="735242A62302FFD2FEBD5AFBAABEFE7A" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[264,503,413,437]" pageId="8" pageNumber="446" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay et al., 2016</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FDB45AFBABE0FE1A" author="Pall-Gergely B &amp; White TS" pageId="8" pageNumber="446" pagination="2011 - 2029" refId="ref7633" refString="Pall-Gergely B &amp; White TS (2022) Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History, 56 (45 - 48): 2011 - 2029." type="journal article" year="2022">Páll-Gergely &amp; White, 2022</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FF035ADBAAECFE1A" author="Tongkerd P &amp; Lwin N &amp; Pall-Gergely B &amp; Chanabun R &amp; Pholyotha A &amp; Prasankok P &amp; Seesamut T &amp; Siriwut W &amp; Srisonchai R &amp; Sutcharit C &amp; Panha S" box="[182,421,445,469]" pageId="8" pageNumber="446" pagination="157 - 197" refId="ref8756" refString="Tongkerd P, Lwin N, Pall-Gergely B, Chanabun R, Pholyotha A, Prasankok P, Seesamut T, Siriwut W, Srisonchai R, Sutcharit C &amp; Panha S (2024) Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species. ZooKeys, 1195: 157 - 197." type="journal article" year="2024">Tongkerd et al., 2024</bibRefCitation>
).
</paragraph>
<caption id="43BC6FDF230DFFDDFF3D5CABA97CF7F7" ID-DOI="http://doi.org/10.5281/zenodo.14682807" ID-Zenodo-Dep="14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="7" pageNumber="445" startId="7.[136,171,1997,2019]" targetBox="[136,1474,189,1981]" targetPageId="7" targetType="figure">
<paragraph id="177C3F57230DFFDDFF3D5CABA97CF7F7" blockId="7.[136,1477,1997,2105]" pageId="7" pageNumber="445">
Fig. 4.
<taxonomicName id="D0C344D4230DFFDDFF655CABA914F82C" authority="(Saurin, 1953)" authorityName="Inkhavilay &amp; Sutcharit &amp; Chirasak" authorityYear="2024" baseAuthorityName="Saurin" baseAuthorityYear="1953" box="[208,605,1997,2019]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="445" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E345230DFFDDFF655CABAA80F82D" box="[208,457,1997,2018]" italics="true" pageId="7" pageNumber="445">Bensonella novemdentata</emphasis>
(
<bibRefCitation id="735242A6230DFFDDFE605CABA91FF82C" author="Saurin E" box="[469,598,1997,2019]" pageId="7" pageNumber="445" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
)
</taxonomicName>
. A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations:
<emphasis id="25B7E345230DFFDDFBEF5C8CAF27F830" bold="true" box="[1114,1134,2026,2047]" pageId="7" pageNumber="445">a:</emphasis>
angular lamella,
<emphasis id="25B7E345230DFFDDFAA95C8CAE78F830" bold="true" box="[1308,1329,2026,2047]" pageId="7" pageNumber="445">b:</emphasis>
basal plica,
<emphasis id="25B7E345230DFFDDFA055C8CAE8AF830" bold="true" box="[1456,1475,2026,2047]" pageId="7" pageNumber="445">c:</emphasis>
columellar lamella,
<emphasis id="25B7E345230DFFDDFEF85360AA20F7D3" bold="true" box="[333,361,2054,2076]" pageId="7" pageNumber="445">ip:</emphasis>
infraparietal lamella,
<emphasis id="25B7E345230DFFDDFDF75360A924F7D3" bold="true" box="[578,621,2054,2076]" pageId="7" pageNumber="445">itpl:</emphasis>
interpalatal plica,
<emphasis id="25B7E345230DFFDDFC905361A801F7D3" bold="true" box="[805,840,2055,2076]" pageId="7" pageNumber="445">lpl:</emphasis>
lower palatal plica,
<emphasis id="25B7E345230DFFDDFBA75361AF6EF7D3" bold="true" box="[1042,1063,2055,2076]" pageId="7" pageNumber="445">p:</emphasis>
parietal lamella,
<emphasis id="25B7E345230DFFDDFB665361AFB9F7D3" bold="true" box="[1235,1264,2055,2076]" pageId="7" pageNumber="445">pt:</emphasis>
palatal tubercle,
<emphasis id="25B7E345230DFFDDFA2E5361AE8CF7D3" bold="true" box="[1435,1477,2055,2076]" pageId="7" pageNumber="445">upl:</emphasis>
upper palatal plica. Photo: M. Caballer (A).
</paragraph>
</caption>
<paragraph id="177C3F572302FFD2FFC45A9BA927FC5A" blockId="8.[111,752,509,917]" pageId="8" pageNumber="446">
The two species,
<taxonomicName id="D0C344D42302FFD2FE8B5A9BAA63FDFA" authority="(Panha &amp; Tongkerd, 2003)" baseAuthorityName="Panha &amp; Tongkerd" baseAuthorityYear="2003" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="wangviangensis">
<emphasis id="25B7E3452302FFD2FE8B5A9BA93EFDDA" box="[318,631,509,533]" italics="true" pageId="8" pageNumber="446">Bensonella wangviangensis</emphasis>
(
<bibRefCitation id="735242A62302FFD2FD3F5A9BAA6AFDFA" author="Panha S &amp; Tongkerd P &amp; Sutcharit C &amp; Tumpeesuwan S &amp; Vongsombath C" pageId="8" pageNumber="446" pagination="123 - 128" refId="ref7955" refString="Panha S, Tongkerd P, Sutcharit C, Tumpeesuwan S &amp; Vongsombath C (2003) A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana, 13 [2002] (29 / 30): 123 - 128. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2003">Panha &amp; Tongkerd, 2003</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D0C344D42302FFD2FED3597BABFAFD9A" authority="(Maassen, 2008)" baseAuthorityName="Maassen" baseAuthorityYear="2008" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="gittenbergeri">
<emphasis id="25B7E3452302FFD2FED3597BA93EFDFA" box="[358,631,541,565]" italics="true" pageId="8" pageNumber="446">Bensonella gittenbergeri</emphasis>
(
<bibRefCitation id="735242A62302FFD2FD3D597BABE5FD9A" author="Maassen WJM" pageId="8" pageNumber="446" pagination="233 - 240" refId="ref7262" refString="Maassen WJM (2008) Remarks on a small collection of terrestrial molluscs from north-west Laos, with descriptions of three new species (Mollusca: Pulmonata: Streptaxidae, Vertiginidae). Basteria, 72: 233 - 240." type="journal article" year="2008">Maassen, 2008</bibRefCitation>
)
</taxonomicName>
, previously known from
<collectingCountry id="6FD47FC72302FFD2FE635958A947FD9A" box="[470,526,574,597]" name="Laos" pageId="8" pageNumber="446">Laos</collectingCountry>
, clearly differ from this species in both shell shape and apertural dentition. For comparison,
<taxonomicName id="D0C344D42302FFD2FE87591BA92CFD5A" baseAuthorityName="Panha &amp; Tongkerd" baseAuthorityYear="2003" box="[306,613,637,661]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="wangviangensis">
<emphasis id="25B7E3452302FFD2FE87591BA92CFD5A" box="[306,613,637,661]" italics="true" pageId="8" pageNumber="446">Bensonella wangviangensis</emphasis>
</taxonomicName>
has a small shell (height ≤
<quantity id="D03B92B22302FFD2FEAE59FBAA14FD7A" box="[283,349,669,693]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="446" unit="mm" value="2.0">2 mm</quantity>
) with 4 whorls, without basal plica, the parietal wall at the posterior corner elevated, and an aperture elliptical with prominent posterior corner balloonshaped, formed by angular lamella and upper palatal plica. Meanwhile,
<taxonomicName id="D0C344D42302FFD2FF46587BAABFFCFA" baseAuthorityName="Maassen" baseAuthorityYear="2008" box="[243,502,797,821]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="gittenbergeri">
<emphasis id="25B7E3452302FFD2FF46587BAABFFCFA" box="[243,502,797,821]" italics="true" pageId="8" pageNumber="446">Bensonella gittenbergeri</emphasis>
</taxonomicName>
has a large shell (height
<quantity id="D03B92B22302FFD2FFDA585BAB83FC9A" box="[111,202,829,853]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="8" pageNumber="446" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
), ovate conic shape, inconspicuous palatal tubercle, and lower palatal long, reaching expanded lip (
<bibRefCitation id="735242A62302FFD2FDDC583BABE3FC5A" author="Panha S &amp; Tongkerd P &amp; Sutcharit C &amp; Tumpeesuwan S &amp; Vongsombath C" pageId="8" pageNumber="446" pagination="123 - 128" refId="ref7955" refString="Panha S, Tongkerd P, Sutcharit C, Tumpeesuwan S &amp; Vongsombath C (2003) A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana, 13 [2002] (29 / 30): 123 - 128. [Date of publication follows Jirapatrasilp et al. (2023)]" type="journal article" year="2003">Panha et al., 2003</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FF025818AA15FC5A" author="Maassen WJM" box="[183,348,893,917]" pageId="8" pageNumber="446" pagination="233 - 240" refId="ref7262" refString="Maassen WJM (2008) Remarks on a small collection of terrestrial molluscs from north-west Laos, with descriptions of three new species (Mollusca: Pulmonata: Streptaxidae, Vertiginidae). Basteria, 72: 233 - 240." type="journal article" year="2008">Maassen, 2008</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FEDC581BA92AFC5A" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[361,611,893,917]" pageId="8" pageNumber="446" pagination="213 - 232" refId="ref7007" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay et al., 2016</bibRefCitation>
).
</paragraph>
<paragraph id="177C3F572302FFD2FFC458DBAAB4FA7A" blockId="8.[111,752,957,1462]" pageId="8" pageNumber="446">
Regardless of their phylogenetic relationships, the traditional diagnostic characters set by their respective type species,
<taxonomicName id="D0C344D42302FFD2FFC4589BAB99FBDA" authorityName=", Pilsbry" authorityYear="1917" box="[113,208,1021,1045]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FFC4589BAB99FBDA" box="[113,208,1021,1045]" italics="true" pageId="8" pageNumber="446">Boysidia</emphasis>
</taxonomicName>
and
<taxonomicName id="D0C344D42302FFD2FEBE589BAACAFBDA" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" box="[267,387,1021,1045]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FEBE589BAACAFBDA" box="[267,387,1021,1045]" italics="true" pageId="8" pageNumber="446">Bensonella</emphasis>
</taxonomicName>
, share a peristome being adnate to the last whorl and lacking the free portion of the last whorl (tuba). In contrast,
<taxonomicName id="D0C344D42302FFD2FE3F5F5BAAAEFB9A" authorityName=", Pilsbry" authorityYear="1917" box="[394,487,1085,1109]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FE3F5F5BAAAEFB9A" box="[394,487,1085,1109]" italics="true" pageId="8" pageNumber="446">Boysidia</emphasis>
</taxonomicName>
has one parietal lamella (without angular lamella), while
<taxonomicName id="D0C344D42302FFD2FE515F3BA914FBBA" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" box="[484,605,1117,1141]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FE515F3BA914FBBA" box="[484,605,1117,1141]" italics="true" pageId="8" pageNumber="446">Bensonella</emphasis>
</taxonomicName>
has separate parietal and angular lamellae. Based on that criterion of generic classification, the species
<taxonomicName id="D0C344D42302FFD2FE5E5FFBA9AEFB7A" authorityName="Saurin" authorityYear="1953" box="[491,743,1181,1205]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="species" species="novemdentata">
<emphasis id="25B7E3452302FFD2FE5E5FFBA9AEFB7A" box="[491,743,1181,1205]" italics="true" pageId="8" pageNumber="446">Boysidia novemdentata</emphasis>
</taxonomicName>
is placed under the
<taxonomicName id="D0C344D42302FFD2FEEF5FDBAAF5FB1A" authorityName=", Pilsbry" authorityYear="1917" box="[346,444,1213,1237]" class="Gastropoda" family="Gastrocoptidae" genus="Boysidia" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FEEF5FDBAAF5FB1A" box="[346,444,1213,1237]" italics="true" pageId="8" pageNumber="446">Boysidia</emphasis>
</taxonomicName>
(
<bibRefCitation id="735242A62302FFD2FE7A5FDBA921FB1A" author="Saurin E" box="[463,616,1213,1237]" pageId="8" pageNumber="446" pagination="113 - 120" refId="ref8343" refString="Saurin E (1953) Coquilles nouvelles de lIndochine. Journal de Conchyliologie, 93: 113 - 120." type="journal article" year="1953">Saurin, 1953</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FDC25FDBABBAFB3A" author="Inkhavilay K &amp; Sutcharit C &amp; Bantaowong U &amp; Chanabun R &amp; Siriwut W &amp; Srisonchai R &amp; Pholyotha A &amp; Jirapatrasilp P &amp; Panha S" pageId="8" pageNumber="446" pagination="1 - 166" refId="ref6956" refString="Inkhavilay K, Sutcharit C, Bantaowong U, Chanabun R, Siriwut W, Srisonchai R, Pholyotha A, Jirapatrasilp P &amp; Panha S (2019) Annotated checklist of the terrestrial molluscs from Laos (Mollusca, Gastropoda). ZooKeys, 834: 1 - 166." type="journal article" year="2019">Inkhavilay et al., 2019</bibRefCitation>
; MolluscaBase, 2024). However, the
<typeStatus id="C87881F52302FFD2FD2E5FB8A9A6FB39" box="[667,751,1246,1270]" pageId="8" pageNumber="446" type="syntype">syntype</typeStatus>
(
<figureCitation id="8FF823D22302FFD2FFCD5F9BAB86FADA" box="[120,207,1277,1301]" captionStart="Fig" captionStartId="7.[136,171,1997,2019]" captionTargetBox="[136,1474,189,1981]" captionTargetId="figure-7@7.[136,1474,189,1981]" captionTargetPageId="7" captionText="Fig. 4. Bensonella novemdentata (Saurin, 1953). A, syntype MNHN-IM-2000-33881 from Pah Hia. B, shell and enlarged apertures show different view angles of specimen CUMZ 14426 from Sam Tai, Houaphanh, Laos. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, upl: upper palatal plica. Photo: M. Caballer (A)." figureDoi="http://doi.org/10.5281/zenodo.14682807" httpUri="https://zenodo.org/record/14682807/files/figure.png" pageId="8" pageNumber="446">Fig. 4A</figureCitation>
) has well-separated parietal and angular lamellae and peristome adnate to the preceding whorl; these are the unique characteristics of
<taxonomicName id="D0C344D42302FFD2FE3A5E5BA941FA9A" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" box="[399,520,1341,1365]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FE3A5E5BA941FA9A" box="[399,520,1341,1365]" italics="true" pageId="8" pageNumber="446">Bensonella</emphasis>
</taxonomicName>
(
<bibRefCitation id="735242A62302FFD2FDAF5E5BABE0FABA" author="Pilsbry HA &amp; Vanatta EG" pageId="8" pageNumber="446" pagination="582 - 611" refId="ref8179" refString="Pilsbry HA &amp; Vanatta EG (1900) A partial revision of the Pupae of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, 52: 582 - 611." type="journal article" year="1900">Pilsbry &amp; Vanatta, 1900</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FF075E3BAA08FABA" author="Pilsbry HA" box="[178,321,1373,1397]" pageId="8" pageNumber="446" refId="ref8014" refString="Pilsbry HA (1917, 1918 [1916 - 1918]) Pupillidae (Gastrocoptinae). Manual of Conchology. Second Series, Volume 24. The Academy of Natural Sciences of Philadelphia, Philadelphia. 380 pp. [pp. 1 - 112, pl. 1 - 13 (1916); pp. 113 - 256, pl. 14 - 38 (1917); pp. 257 - 380, pl. 39 - 49 (1918)]" type="book" year="1917">Pilsbry, 1917</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FEFF5E3BAAB1FABA" author="Schileyko AA" box="[330,504,1373,1397]" pageId="8" pageNumber="446" pagination="129 - 261" refId="ref8363" refString="Schileyko AA (1998) Treatise on Recent terrestrial pulmonate molluscs. Part 2. Gastrocoptidae, Hypselostomatidae, Vertiginidae, Truncatellinidae, Pachnodidae, Enidae, Sagdidae. Ruthenica, Supplement 2: 129 - 261." type="journal article" year="1998">Schileyko, 1998</bibRefCitation>
;
<bibRefCitation id="735242A62302FFD2FDB75E3BABE2FA5A" author="Pall-Gergely B &amp; White TS" pageId="8" pageNumber="446" pagination="2011 - 2029" refId="ref7633" refString="Pall-Gergely B &amp; White TS (2022) Solving the mystery of the misunderstood Bensonella plicidens (Benson, 1849) (Gastropoda: Stylommatophora: Hypselostomatidae). Journal of Natural History, 56 (45 - 48): 2011 - 2029." type="journal article" year="2022">Páll-Gergely &amp; White, 2022</bibRefCitation>
). Therefore, we have transferred this species into a more appropriate genus,
<taxonomicName id="D0C344D42302FFD2FE375EFBAABFFA7A" authorityName="Pilsbry &amp; Vanatta" authorityYear="1900" box="[386,502,1437,1461]" class="Gastropoda" family="Gastrocoptidae" genus="Bensonella" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="446" phylum="Mollusca" rank="genus">
<emphasis id="25B7E3452302FFD2FE375EFBAABFFA7A" box="[386,502,1437,1461]" italics="true" pageId="8" pageNumber="446">Bensonella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,246 @@
<document id="A9D051FF3C2E49FD76438D30B55C1163" ID-DOI="10.26107/RBZ-2024-0034" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682900" ID-ZooBank="10B1E24B-518C-4A85-A7C2-400C1D6D5EE7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171819659" checkinUser="felipe" docAuthor="Ng, Peter K. L., Wowor, &amp; Daisy" docDate="2024" docId="C06087FDF159FFC6FEFDF90BFBA6F7D9" docLanguage="en" docName="RafflesBZool.72.469-478.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Geosesarma nigripes Ng, Wowor &amp; Daisy, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="473" masterDocId="3C59FF85F15DFFC2FFF4FFB6FFDAFFCC" masterDocTitle="Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia" masterLastPageNumber="478" masterPageNumber="469" pageNumber="473" updateTime="1737175501798" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="B8B786E670D2DA89ABC4211A33B05D82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="841907344E01793FB989FB102FE04DF6">
<mods:title id="7C0A60ACFD7DBE3F6B0CE2019AE070E6">Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="2729E10EC44478343E8CF39C3FF9965A" type="personal">
<mods:role id="03B0C173FC93E97586B3E418C6A8187E">
<mods:roleTerm id="935B4DF30624E651DBA3DF30D1201C0E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="683627EA5AC5D187C82F8AB2ED2C4517">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="FF918753DE9A4442CB1295374A58F078" type="personal">
<mods:role id="D7A5AE0854A9C735175B2E00CDB8DEE2">
<mods:roleTerm id="73C695D2998C6295FA73BD0128AD6239">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F20B40ABE761F318717134840D6D8A9C">Wowor,</mods:namePart>
</mods:name>
<mods:name id="6CD8F894EF4922364290600D794C8474" type="personal">
<mods:role id="A1BD3AD906249857E30AA65C51F95542">
<mods:roleTerm id="3E7E00B5FB507FA58CB2CEA740296927">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C44C95A6718EB6A829FD7B80C770EF45">Daisy</mods:namePart>
</mods:name>
<mods:typeOfResource id="287831E1BB6E21F72A6337F4183AD782">text</mods:typeOfResource>
<mods:relatedItem id="392C1DBF4DC9648EE44325DAFE2A6045" type="host">
<mods:titleInfo id="53F2390DD436E69BE4AC965F9A478740">
<mods:title id="2B125F97D48A7ADE70C618464DC45C4E">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="2E194A5B97C122DB87B6D16A574E2A09">
<mods:date id="A9043B3BBFE36A87E7284B752A28A398">2024</mods:date>
<mods:detail id="D4E7F71C8BCD942BEFF235E2737DCBD4" type="pubDate">
<mods:number id="E41950C64AE855A1CB7701B417B515D0">2024-11-27</mods:number>
</mods:detail>
<mods:detail id="8B20CCF7EC2E3835DDA8A57B2EBA5352" type="volume">
<mods:number id="A6FAB67BE524C4BA62E24A962B6D61DF">72</mods:number>
</mods:detail>
<mods:extent id="68844F38E68A85AD14001EC7E7C42B53" unit="page">
<mods:start id="DC06A985F5ED916ECE55078444467B36">469</mods:start>
<mods:end id="BB371B3C8C75BEE8887B7E9F66E95E3B">478</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="37362A564C7DDF3C501072316A843DA5">journal article</mods:classification>
<mods:identifier id="CF58490AB8DB5734A7E59BE27D4DAC76" type="DOI">10.26107/RBZ-2024-0034</mods:identifier>
<mods:identifier id="58B2B5B319EC3BA6612F65F0022BE8AB" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="11D4590765BDA3FBF5B71EA045C266E9" type="Zenodo-Dep">14682900</mods:identifier>
<mods:identifier id="FD178B5591F63E8F7C29ECEEBD865B53" type="ZooBank">10B1E24B-518C-4A85-A7C2-400C1D6D5EE7</mods:identifier>
</mods:mods>
<treatment id="C06087FDF159FFC6FEFDF90BFBA6F7D9" ID-DOI="http://doi.org/10.5281/zenodo.14682908" ID-Zenodo-Dep="14682908" LSID="urn:lsid:plazi:treatment:C06087FDF159FFC6FEFDF90BFBA6F7D9" httpUri="http://treatment.plazi.org/id/C06087FDF159FFC6FEFDF90BFBA6F7D9" lastPageNumber="473" pageId="4" pageNumber="473">
<subSubSection id="00D36560F159FFC6FEFDF90BFD5CF919" box="[265,646,1725,1749]" pageId="4" pageNumber="473" type="nomenclature">
<paragraph id="487636EBF159FFC6FEFDF90BFD5CF919" blockId="4.[265,646,1725,1781]" box="[265,646,1725,1749]" pageId="4" pageNumber="473">
<heading id="133E8187F159FFC6FEFDF90BFD5CF919" bold="true" box="[265,646,1725,1749]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="473" reason="2">
<emphasis id="7ABDEAF9F159FFC6FEFDF90BFD5CF919" bold="true" box="[265,646,1725,1749]" pageId="4" pageNumber="473">
<taxonomicName id="8FC94D68F159FFC6FEFDF90BFE2BF919" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[265,497,1725,1749]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="7ABDEAF9F159FFC6FEFDF90BFE2BF919" bold="true" box="[265,497,1725,1749]" italics="true" pageId="4" pageNumber="473">Geosesarma nigripes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F159FFC6FDF5F90BFD5CF919" box="[513,646,1725,1749]" pageId="4" pageNumber="473" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00D36560F159FFC6FEDDF96BFDBCF939" box="[297,614,1757,1781]" pageId="4" pageNumber="473" type="description">
<paragraph id="487636EBF159FFC6FEDDF96BFDBCF939" blockId="4.[265,646,1725,1781]" box="[297,614,1757,1781]" pageId="4" pageNumber="473">
(
<figureCitation id="D0F22A6EF159FFC6FEC5F96BFE75F939" box="[305,431,1757,1781]" captionStart="Fig" captionStartId="1.[113,148,2013,2035]" captionTargetBox="[114,1450,240,1997]" captionTargetId="figure-50@1.[114,1450,240,1997]" captionTargetPageId="1" captionText="Fig. 1. Live colouration of males. AD, Geosesarma riani, new species; EH, G. nigripes, new species. All specimens not preserved. Photographs: A, E, F, Achmad Rian Dietra; B, Oliver Mengedoht; C, D, G, H, Chris Lukhaup." figureDoi="http://doi.org/10.5281/zenodo.14682904" httpUri="https://zenodo.org/record/14682904/files/figure.png" pageId="4" pageNumber="473">Figs. 1EH</figureCitation>
,
<figureCitation id="D0F22A6EF159FFC6FE49F96BFE16F939" box="[445,460,1757,1781]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">3</figureCitation>
,
<figureCitation id="D0F22A6EF159FFC6FE2EF96BFDC3F939" box="[474,537,1757,1781]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="4" pageNumber="473">4DF</figureCitation>
,
<figureCitation id="D0F22A6EF159FFC6FDD2F96BFD84F939" box="[550,606,1757,1781]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">5JR</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="00D36560F159FFC6FF7CF8ABFA19FE79" pageId="4" pageNumber="473" type="materials_examined">
<paragraph id="487636EBF159FFC6FF7CF8ABFA19FE79" blockId="4.[136,775,1821,2069]" lastBlockId="4.[835,1475,189,437]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FF7CF8ABFEB8F8F9" bold="true" box="[136,354,1821,1845]" pageId="4" pageNumber="473">Material examined.</emphasis>
<materialsCitation id="F8A13CB6F159FFC6FE90F8ABFF1BF879" collectingDate="2022-06" collectionCode="MZB" collectorName="Achmad Sofyan" country="Indonesia" location="Adimulya village" municipality="Wanareja District" pageId="4" pageNumber="473" specimenCount="1" stateProvince="Jawa Tengah" typeStatus="holotype">
<typeStatus id="97728849F159FFC6FE90F8ABFE10F8F9" box="[356,458,1821,1845]" pageId="4" pageNumber="473" type="holotype">Holotype</typeStatus>
: male (12.3 ×
<quantity id="8F319B0EF159FFC6FDAEF8ABFD6DF8F9" box="[602,695,1821,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="473" unit="mm" value="11.3">11.3 mm</quantity>
) (
<collectionCode id="2ED8AE2EF159FFC6FD3FF8ABFCDCF8F9" box="[715,774,1821,1845]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="473" type="Museum">MZB</collectionCode>
Cru 5720), Cikukun hamlet,
<location id="4D166030F159FFC6FE3AF88BFD4BF899" LSID="urn:lsid:plazi:treatment:C06087FDF159FFC6FEFDF90BFBA6F7D9:4D166030F159FFC6FE3AF88BFD4BF899" box="[462,657,1853,1877]" country="Indonesia" municipality="Wanareja District" name="Adimulya village" pageId="4" pageNumber="473" stateProvince="Jawa Tengah">Adimulya village</location>
,
<collectingMunicipality id="A812AC91F159FFC6FD6BF88BFF01F8B9" pageId="4" pageNumber="473">Wanareja District</collectingMunicipality>
, Cilacap Regency,
<collectingRegion id="8A0DF809F159FFC6FE48F8EBFD93F8B9" box="[444,585,1885,1909]" country="Indonesia" name="Jawa Tengah" pageId="4" pageNumber="473">Central Java</collectingRegion>
,
<collectingCountry id="30DE767BF159FFC6FDAEF8EBFD1FF8B9" box="[602,709,1885,1909]" name="Indonesia" pageId="4" pageNumber="473">Indonesia</collectingCountry>
, coll.
<collectorName id="E53C533DF159FFC6FF7CF8CBFEEFF859" box="[136,309,1917,1941]" pageId="4" pageNumber="473">Achmad Sofyan</collectorName>
, transferred from Achmad Rian Dietra,
<date id="3C77102BF159FFC6FD22F8C8FF1BF879" pageId="4" pageNumber="473" value="2022-06">
<collectingDate id="2C33E9C3F159FFC6FD22F8C8FF1BF879" pageId="4" pageNumber="473" value="2022-06">June 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="F8A13CB6F159FFC6FF3EF828FEAEF819" collectingDate="2022-06" collectionCode="ZRC" collectorName="Achmad Sofyan" country="Indonesia" location="Adimulya village" municipality="Wanareja District" pageId="4" pageNumber="473" specimenCode="ZRC 2023.0321" specimenCount="1" specimenCount-male="1" stateProvince="Jawa Tengah" typeStatus="paratype">
<typeStatus id="97728849F159FFC6FF3EF828FEF6F87A" box="[202,300,1950,1974]" pageId="4" pageNumber="473" type="paratype">Paratype</typeStatus>
:
<specimenCount id="5ECFFD62F159FFC6FEC1F82BFEA6F879" box="[309,380,1949,1973]" count="1" pageId="4" pageNumber="473" type="male">1 male</specimenCount>
(12.7 ×
<quantity id="8F319B0EF159FFC6FE21F82BFDEFF879" box="[469,565,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="4" pageNumber="473" unit="mm" value="11.9">11.9 mm</quantity>
) (
<specimenCode id="186F9E90F159FFC6FDBEF82BFD21F879" box="[586,763,1949,1973]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="473">ZRC 2023.0321</specimenCode>
), same data as holotype
</materialsCitation>
;
<specimenCount id="5ECFFD62F159FFC6FE8AF80BFE1CF819" box="[382,454,1981,2005]" count="1" pageId="4" pageNumber="473" type="male">1 male</specimenCount>
(13.3 ×
<quantity id="8F319B0EF159FFC6FDEBF80BFD5BF819" box="[543,641,1981,2005]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="473" unit="mm" value="11.8">11.8 mm</quantity>
),
<materialsCitation id="F8A13CB6F159FFC6FD67F80BFD31F7D9" collectionCode="MZB" country="Indonesia" pageId="4" pageNumber="473" specimenCount="10" specimenCount-female="10" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="5ECFFD62F159FFC6FD67F80BFCDDF819" box="[659,775,1981,2005]" count="10" pageId="4" pageNumber="473" type="female">10 females</specimenCount>
(10.8 ×
<quantity id="8F319B0EF159FFC6FF15F86BFEEEF839" box="[225,308,2013,2037]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="4" pageNumber="473" unit="mm" value="9.9">9.9 mm</quantity>
14.5 ×
<quantity id="8F319B0EF159FFC6FE50F86BFDD3F839" box="[420,521,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="4" pageNumber="473" unit="mm" value="13.1">13.1 mm</quantity>
), 12 ovigerous females (11.8 × 11.0 mm 12.8 ×
<quantity id="8F319B0EF159FFC6FE43F84BFDC1F7D9" box="[439,539,2045,2069]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="4" pageNumber="473" unit="mm" value="12.4">12.4 mm</quantity>
) (
<collectionCode id="2ED8AE2EF159FFC6FDC7F84BFDB4F7D9" box="[563,622,2045,2069]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="473" type="Museum">MZB</collectionCode>
Cru 5721)
</materialsCitation>
,
<materialsCitation id="F8A13CB6F159FFC6FD0DF84BFAC9FED9" collectingDate="2023-07" collectionCode="ZRC" country="Indonesia" pageId="4" pageNumber="473" specimenCode="ZRC 2023.0562" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="5ECFFD62F159FFC6FD0DF84BFC4DFF19" box="[761,919,189,2069]" count="2" pageId="4" pageNumber="473" type="female">2 females</specimenCount>
(13.9 ×
<quantity id="8F319B0EF159FFC6FC09FF0BFBB8FF19" box="[1021,1122,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="473" unit="mm" value="12.8">12.8 mm</quantity>
, 12.7 ×
<quantity id="8F319B0EF159FFC6FB37FF0BFAF1FF19" box="[1219,1323,189,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="4" pageNumber="473" unit="mm" value="11.8">11.8 mm</quantity>
), 1 ovigerous female (11.8 ×
<quantity id="8F319B0EF159FFC6FC1AFF6BFB8BFF39" box="[1006,1105,221,245]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="473" unit="mm" value="11.3">11.3 mm</quantity>
) (
<specimenCode id="186F9E90F159FFC6FB9DFF6BFAC4FF39" box="[1129,1310,221,245]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="473">ZRC 2023.0562</specimenCode>
), same locality and collector as
<typeStatus id="97728849F159FFC6FC0DFF4BFB83FED9" box="[1017,1113,253,277]" pageId="4" pageNumber="473" type="holotype">holotype</typeStatus>
, early
<date id="3C77102BF159FFC6FB50FF4BFAC9FED9" box="[1188,1299,253,277]" pageId="4" pageNumber="473" value="2023-07">
<collectingDate id="2C33E9C3F159FFC6FB50FF4BFAC9FED9" box="[1188,1299,253,277]" pageId="4" pageNumber="473" value="2023-07">July 2023</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="5ECFFD62F159FFC6FAEBFF4BFA5BFED9" box="[1311,1409,253,277]" count="34" pageId="4" pageNumber="473" type="male">34 males</specimenCount>
(11.5 ×
<quantity id="8F319B0EF159FFC6FCAFFEABFC74FEF9" box="[859,942,285,309]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="4" pageNumber="473" unit="mm" value="9.9">9.9 mm</quantity>
13.8 ×
<quantity id="8F319B0EF159FFC6FBE9FEABFB58FEF9" box="[1053,1154,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="4" pageNumber="473" unit="mm" value="12.6">12.6 mm</quantity>
),
<materialsCitation id="F8A13CB6F159FFC6FB63FEABFC21FEB9" collectionCode="MZB" country="Indonesia" pageId="4" pageNumber="473" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="5ECFFD62F159FFC6FB63FEABFADAFEF9" box="[1175,1280,285,309]" count="2" pageId="4" pageNumber="473" type="female">2 females</specimenCount>
(11.5 ×
<quantity id="8F319B0EF159FFC6FA95FEABFA18FEF9" box="[1377,1474,285,309]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="4" pageNumber="473" unit="mm" value="10.3">10.3 mm</quantity>
13.3 ×
<quantity id="8F319B0EF159FFC6FC5EFE8BFBD5FE99" box="[938,1039,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="4" pageNumber="473" unit="mm" value="11.4">11.4 mm</quantity>
), 1 ovigerous female (12.5 ×
<quantity id="8F319B0EF159FFC6FAACFE8BFA61FE99" box="[1368,1467,317,341]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="4" pageNumber="473" unit="mm" value="11.1">11.1 mm</quantity>
) (
<collectionCode id="2ED8AE2EF159FFC6FCBFFEEBFC5CFEB9" box="[843,902,349,373]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="4" pageNumber="473" type="Museum">MZB</collectionCode>
Cru 5732)
</materialsCitation>
,
<specimenCount id="5ECFFD62F159FFC6FBF1FEEBFBBEFEB9" box="[1029,1124,349,373]" count="20" pageId="4" pageNumber="473" type="male">20 males</specimenCount>
(10.7 × 9.6 15.1 ×
<quantity id="8F319B0EF159FFC6FAB5FEEBFA79FEB9" box="[1345,1443,349,373]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="4" pageNumber="473" unit="mm" value="13.8">13.8 mm</quantity>
),
<materialsCitation id="F8A13CB6F159FFC6FA41FEEBFA65FE79" collectingDate="2023-11-28" collectionCode="ZRC" country="Indonesia" pageId="4" pageNumber="473" specimenCode="ZRC 2023.0567" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Tengah" typeStatus="paratype">
<specimenCount id="5ECFFD62F159FFC6FA41FEEBFC4FFE59" count="2" pageId="4" pageNumber="473" type="female">2 females</specimenCount>
(12.3 ×
<quantity id="8F319B0EF159FFC6FC1EFECBFB9CFE59" box="[1002,1094,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="4" pageNumber="473" unit="mm" value="11.1">11.1 mm</quantity>
, 15.0 ×
<quantity id="8F319B0EF159FFC6FB6CFECBFB2CFE59" box="[1176,1270,381,405]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="4" pageNumber="473" unit="mm" value="12.6">12.6 mm</quantity>
) (
<specimenCode id="186F9E90F159FFC6FAFDFECBFA6DFE59" box="[1289,1463,381,405]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="473">ZRC 2023.0567</specimenCode>
), same locality and collector as
<typeStatus id="97728849F159FFC6FB70FE2BFB3EFE79" box="[1156,1252,413,437]" pageId="4" pageNumber="473" type="holotype">holotype</typeStatus>
,
<date id="3C77102BF159FFC6FB1AFE2BFA65FE79" box="[1262,1471,413,437]" pageId="4" pageNumber="473" value="2023-11-28">
<collectingDate id="2C33E9C3F159FFC6FB1AFE2BFA65FE79" box="[1262,1471,413,437]" pageId="4" pageNumber="473" value="2023-11-28">28 November 2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00D36560F159FFC6FCB7FE6BFB85F859" pageId="4" pageNumber="473" type="diagnosis">
<paragraph id="487636EBF159FFC6FCB7FE6BFBE3F9F9" blockId="4.[835,1475,477,1589]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FCB7FE6BFC62FE39" bold="true" box="[835,952,477,501]" pageId="4" pageNumber="473">Diagnosis.</emphasis>
Carapace subsquarish, slightly trapezoidal with posterior part wider than anterior part, lateral margins gently diverging posteriorly (
<figureCitation id="D0F22A6EF159FFC6FB7BFDABFAD4FDF9" box="[1167,1294,541,565]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3A, B</figureCitation>
); dorsal surface with distinct regions, anterior regions with numerous small rounded granules (
<figureCitation id="D0F22A6EF159FFC6FBF8FDEBFB58FDB9" box="[1036,1154,605,629]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3A, B</figureCitation>
); front deflexed, frontal lobes very broad with almost truncate margins, separated by broad median concavity; postfrontal cristae sharp (
<figureCitation id="D0F22A6EF159FFC6FACEFD2BFA6FFD79" box="[1338,1461,669,693]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3AC</figureCitation>
); external orbital tooth triangular, directed obliquely outwards, tip extending beyond anterior part of lateral carapace margin, first epibranchial tooth triangular, low, visible, separated from margin by shallow notch, second epibranchial tooth small but visible (
<figureCitation id="D0F22A6EF159FFC6FBE1FC8BFBB5FC99" box="[1045,1135,829,853]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3B</figureCitation>
). Ischium of third maxilliped subtrapezoidal, merus ovate, wide; exopod relatively slender, with long flagellum that extends beyond merus width (
<figureCitation id="D0F22A6EF159FFC6FA63FCCBFC84FC79" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">Fig. 5J</figureCitation>
). Outer surface of palm of adult male chelae with low granules and rugae; inner surface finely granulated but without distinct transverse comb-like ridge; dorsal margin of dactylus with 5 or 6 evenly spaced tubercles, each with chitinous tip (
<figureCitation id="D0F22A6EF159FFC6FC28FBABFB88FBF9" box="[988,1106,1053,1077]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3DF</figureCitation>
). Ambulatory legs with relatively stouter meri (fourth leg length-to-width ratio ca. 3.0), with sharp subdistal spine on dorsal margin; ventral margins of propodus and dactylus of adult first ambulatory leg with scattered stiff setae (
<figureCitation id="D0F22A6EF159FFC6FBE3FB2BFB53FB79" box="[1047,1161,1181,1205]" captionStart="Fig" captionStartId="3.[113,148,2015,2037]" captionTargetBox="[114,1452,209,2001]" captionTargetId="figure-13@3.[113,1453,209,2001]" captionTargetPageId="3" captionText="Fig. 3. Geosesarma nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="4" pageNumber="473">Fig. 3A, G</figureCitation>
). Male pleon wide, triangular; especially somite 6 relatively wide; telson subtriangular with rounded tip, slightly shorter than broad (length-to-width ratio ca. 0.8); somite 6 with convex lateral margins (
<figureCitation id="D0F22A6EF159FFC6FAAEFB4BFA64FAD9" box="[1370,1470,1277,1301]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="4" pageNumber="473">Figs. 4D</figureCitation>
,
<figureCitation id="D0F22A6EF159FFC6FCB7FAABFCB2FAF9" box="[835,872,1309,1333]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">5K</figureCitation>
). G1 relatively slender; subdistal part of outer margin of stem before chitinous distal part rounded, outer margin of junction between stem and chitinous distal part concave, long; chitinous distal part bent about 45º along longitudinal axis, elongate, tapering, distalmost part conical in lateral view, subspatuliform in mesial view (
<figureCitation id="D0F22A6EF159FFC6FB2CFA0BFA97FA19" box="[1240,1357,1469,1493]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">Fig. 5LQ</figureCitation>
). Vulva on anterior part of somite 6, with raised vulvar cover, directed mesially (
<figureCitation id="D0F22A6EF159FFC6FC5BFA4BFBD8F9D9" box="[943,1026,1533,1557]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="4" pageNumber="473">Fig. 4F</figureCitation>
). Preserved egg size 1.8 ×
<quantity id="8F319B0EF159FFC6FADAFA4BFA5BF9D9" box="[1326,1409,1533,1557]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="473" unit="mm" value="1.6">1.6 mm</quantity>
1.6 ×
<quantity id="8F319B0EF159FFC6FCAFF9ABFC74F9F9" box="[859,942,1565,1589]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="473" unit="mm" value="1.5">1.5 mm</quantity>
in diameter.
</paragraph>
<paragraph id="487636EBF159FFC6FCB7F9EBFA5CF979" blockId="4.[835,1474,1629,1717]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FCB7F9EBFBDDF9B9" bold="true" box="[835,1031,1629,1653]" pageId="4" pageNumber="473">Notes on habitat.</emphasis>
<taxonomicName id="8FC94D68F159FFC6FBFAF9E8FB28F9B9" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[1038,1266,1630,1653]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="7ABDEAF9F159FFC6FBFAF9E8FB28F9B9" box="[1038,1266,1630,1653]" italics="true" pageId="4" pageNumber="473">Geosesarma nigripes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F159FFC6FAF4F9E8FA5EF9B9" box="[1280,1412,1629,1653]" pageId="4" pageNumber="473" rank="species">new species</taxonomicNameLabel>
, lives among wet bushes with stony substrates along streams in lowlands in
<collectingRegion id="8A0DF809F159FFC6FC3EF92BFB62F979" box="[970,1208,1693,1717]" country="Indonesia" name="Jawa Tengah" pageId="4" pageNumber="473">Central Java Province</collectingRegion>
at about
<quantity id="8F319B0EF159FFC6FAD4F92BFA83F979" box="[1312,1369,1693,1717]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="473" unit="m" value="40.0">40 m</quantity>
asl.
</paragraph>
<paragraph id="487636EBF159FFC6FCB7F96BFB85F859" blockId="4.[835,1474,1757,1941]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FCB7F96BFC7EF939" bold="true" box="[835,932,1757,1781]" pageId="4" pageNumber="473">Colour.</emphasis>
Carapace pale cream to pale yellow, with posterolateral regions dark purple to almost black; frontal surfaces of cephalothorax dark brown to almost black; H-shaped grooves dark brown; eyes iridescent orange to yellow; chelipeds orange; ambulatory legs dark purple to almost black (
<figureCitation id="D0F22A6EF159FFC6FC29F8CBFB89F859" box="[989,1107,1917,1941]" captionStart="Fig" captionStartId="1.[113,148,2013,2035]" captionTargetBox="[114,1450,240,1997]" captionTargetId="figure-50@1.[114,1450,240,1997]" captionTargetPageId="1" captionText="Fig. 1. Live colouration of males. AD, Geosesarma riani, new species; EH, G. nigripes, new species. All specimens not preserved. Photographs: A, E, F, Achmad Rian Dietra; B, Oliver Mengedoht; C, D, G, H, Chris Lukhaup." figureDoi="http://doi.org/10.5281/zenodo.14682904" httpUri="https://zenodo.org/record/14682904/files/figure.png" pageId="4" pageNumber="473">Fig. 1EH</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00D36560F159FFC6FCB7F80BFBA6F7D9" pageId="4" pageNumber="473" type="etymology">
<paragraph id="487636EBF159FFC6FCB7F80BFBA6F7D9" blockId="4.[835,1474,1981,2069]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FCB7F80BFC1FF819" bold="true" box="[835,965,1981,2005]" pageId="4" pageNumber="473">Etymology.</emphasis>
The species is named after its black ambulatory legs; from the Latin “nigra” for black and “pes” for legs. The name is used as a noun.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,83 @@
<document id="399D4E3076E3666D46FDA236F0D57E69" ID-DOI="10.26107/RBZ-2024-0034" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682900" ID-ZooBank="10B1E24B-518C-4A85-A7C2-400C1D6D5EE7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171819659" checkinUser="felipe" docAuthor="Ng, Peter K. L., Wowor, &amp; Daisy" docDate="2024" docId="C06087FDF15DFFC2FC35FB4FFA47FABE" docLanguage="en" docName="RafflesBZool.72.469-478.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Geosesarma De Man 1892" docType="treatment" docVersion="2" lastPageNumber="469" masterDocId="3C59FF85F15DFFC2FFF4FFB6FFDAFFCC" masterDocTitle="Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia" masterLastPageNumber="478" masterPageNumber="469" pageNumber="469" updateTime="1737175501798" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="45BE0E9FEE51977F69B09E399C64AA17" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A161D99CF6C7203357A8119EFED0B528">
<mods:title id="72AB3007878C8D20DC691FF66F0ECD50">Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="20897A1540EDCDC88A15FAB67A289FEF" type="personal">
<mods:role id="01804F11DCEE5F2A58D5546EEEB8E82A">
<mods:roleTerm id="891FB5AB5D3AAF8223EE4C30F5154D83">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D960E83782B443443D2135EB74498E7">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="0FE411D22B932D8BDE4B40BE0EE7B2E2" type="personal">
<mods:role id="3882DF0FD50F7402C1397DF27CA4C1F2">
<mods:roleTerm id="697B3BF591111345FF7FFEBDE1BFEC21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE4FC651EAE7C728131CFBB411E0B361">Wowor,</mods:namePart>
</mods:name>
<mods:name id="D11C0A69196F672FFA7E3C38583DA051" type="personal">
<mods:role id="83E525F71D3ACD79A29295E478ABF373">
<mods:roleTerm id="5BF3DE01164F4E17F23529CC8A4160E1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="24A8D40610D89432F6E2122D46926F1C">Daisy</mods:namePart>
</mods:name>
<mods:typeOfResource id="2E3FD54A5956528305DB25770DBEED88">text</mods:typeOfResource>
<mods:relatedItem id="C44F1545223F94F0202C6DC8ECB91B91" type="host">
<mods:titleInfo id="67188269E614A96D8B2FF64768A985B6">
<mods:title id="E9E24AD27A4F86625A8699F6AF3051D4">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="D7F4149A38C64D389BAD91E810F5B081">
<mods:date id="68FA63D0FE5507CAEE8B9AEB58DB6D4B">2024</mods:date>
<mods:detail id="DCD7265BFEEF07EC99BBAD21A7A22420" type="pubDate">
<mods:number id="4E306444986D6034945E6A13F8ED7653">2024-11-27</mods:number>
</mods:detail>
<mods:detail id="EA41A1325EBBC5B892C2982C5BE6B5DD" type="volume">
<mods:number id="03BB920F9531CA7EDE203CC136FAE1E2">72</mods:number>
</mods:detail>
<mods:extent id="A5C4681E2A40C8B7A34227F6A3677B9B" unit="page">
<mods:start id="234B190C13C2071D6EDC10400A97AF22">469</mods:start>
<mods:end id="9E59B9A69B8D96F7C1D6AD42B48587D0">478</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AA0E3758C5212CA702E89C49E3966C16">journal article</mods:classification>
<mods:identifier id="F9627A92768EBBC596303AAF6FC0E910" type="DOI">10.26107/RBZ-2024-0034</mods:identifier>
<mods:identifier id="83F442B977E1F0238EB70CD98FCFEA40" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A965DA44ADC7B5424101BAA5A52DD615" type="Zenodo-Dep">14682900</mods:identifier>
<mods:identifier id="A31D18272D27DE19F91F24716B81AC4F" type="ZooBank">10B1E24B-518C-4A85-A7C2-400C1D6D5EE7</mods:identifier>
</mods:mods>
<treatment id="C06087FDF15DFFC2FC35FB4FFA47FABE" ID-DOI="http://doi.org/10.5281/zenodo.14682902" ID-Zenodo-Dep="14682902" LSID="urn:lsid:plazi:treatment:C06087FDF15DFFC2FC35FB4FFA47FABE" httpUri="http://treatment.plazi.org/id/C06087FDF15DFFC2FC35FB4FFA47FABE" lastPageNumber="469" pageId="0" pageNumber="469">
<subSubSection id="00D36560F15DFFC2FC35FB4FFA9EFADD" box="[961,1348,1273,1298]" pageId="0" pageNumber="469" type="nomenclature">
<paragraph id="487636EBF15DFFC2FC35FB4FFA9EFADD" blockId="0.[961,1348,1273,1298]" box="[961,1348,1273,1298]" pageId="0" pageNumber="469">
<heading id="133E8187F15DFFC2FC35FB4FFA9EFADD" bold="true" box="[961,1348,1273,1298]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="469" reason="2">
<emphasis id="7ABDEAF9F15DFFC2FC35FB4FFA9EFADD" bold="true" box="[961,1348,1273,1298]" pageId="0" pageNumber="469">
Genus
<taxonomicName id="8FC94D68F15DFFC2FBE7FB4CFA9EFADD" ID-CoL="7P3H8" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" box="[1043,1348,1273,1298]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="7ABDEAF9F15DFFC2FBE7FB4CFB40FADE" bold="true" box="[1043,1178,1274,1298]" italics="true" pageId="0" pageNumber="469">Geosesarma</emphasis>
<bibRefCitation id="2C584B1AF15DFFC2FB56FB4CFA9EFADD" author="De Man JG" box="[1186,1348,1273,1298]" pageId="0" pageNumber="469" pagination="265 - 527" refId="ref4608" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Volume 2. Pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00D36560F15DFFC2FCB7FA8CFA47FABE" pageId="0" pageNumber="469" type="type_taxon">
<paragraph id="487636EBF15DFFC2FCB7FA8CFA47FABE" blockId="0.[835,1474,1337,1394]" pageId="0" pageNumber="469">
<emphasis id="7ABDEAF9F15DFFC2FCB7FA8CFC06FA9D" bold="true" box="[835,988,1337,1362]" pageId="0" pageNumber="469">
<typeStatus id="97728849F15DFFC2FCB7FA8CFCA7FA9E" box="[835,893,1338,1362]" pageId="0" pageNumber="469">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="8FC94D68F15DFFC2FC11FA8CFCA1FABE" ID-CoL="84HBY" authority="De Man, 1892" authorityName="De Man" authorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="469" phylum="Arthropoda" rank="species" species="nodulifera" subGenus="Geosesarma">
<emphasis id="7ABDEAF9F15DFFC2FC11FA8CFB9BFA9D" box="[997,1089,1338,1361]" italics="true" pageId="0" pageNumber="469">Sesarma</emphasis>
(
<emphasis id="7ABDEAF9F15DFFC2FBA7FA8CFB02FA9D" box="[1107,1240,1338,1361]" italics="true" pageId="0" pageNumber="469">Geosesarma</emphasis>
)
<emphasis id="7ABDEAF9F15DFFC2FB1EFA8CFA83FA9E" box="[1258,1369,1338,1362]" italics="true" pageId="0" pageNumber="469">nodulifera</emphasis>
<bibRefCitation id="2C584B1AF15DFFC2FA96FA8CFCA1FABE" author="De Man JG" pageId="0" pageNumber="469" pagination="265 - 527" refId="ref4608" refString="De Man JG (1892) Decapoden des indischen Archipels. In: Weber M (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Volume 2. Pp. 265 - 527, pls. 15 - 29." type="book chapter" year="1892">De Man, 1892</bibRefCitation>
</taxonomicName>
, subsequent designation by
<bibRefCitation id="2C584B1AF15DFFC2FB5AFAECFA43FABE" author="Serene R &amp; Soh CL" box="[1198,1433,1370,1394]" pageId="0" pageNumber="469" pagination="387 - 416" refId="ref5157" refString="Serene R &amp; Soh CL (1970) New Indo-Pacific Genera allied to Sesarma Say 1817 (Brachyura, Decapoda, Crustacea). Treubia, 27 (4): 387 - 416, pls. 1 - 8." type="journal article" year="1970">Serène &amp; Soh (1970)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,265 @@
<document id="197A09D7B339616A6B541A134168B9D2" ID-DOI="10.26107/RBZ-2024-0034" ID-ISSN="2345-7600" ID-Zenodo-Dep="14682900" ID-ZooBank="10B1E24B-518C-4A85-A7C2-400C1D6D5EE7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171819659" checkinUser="felipe" docAuthor="Ng, Peter K. L., Wowor, &amp; Daisy" docDate="2024" docId="C06087FDF15DFFC6FC22FA2CFE11F959" docLanguage="en" docName="RafflesBZool.72.469-478.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Geosesarma riani Ng, Wowor &amp; Daisy, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="473" masterDocId="3C59FF85F15DFFC2FFF4FFB6FFDAFFCC" masterDocTitle="Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia" masterLastPageNumber="478" masterPageNumber="469" pageNumber="469" updateTime="1737175501798" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="5BBAD07BF7EB4F9D4F3FB231A3608A25" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B5EA64C8F75F846FC8C41C48406ABE42">
<mods:title id="70F6011CCB4A95B38BFDF0BD63CB2D2F">Two new species of vampire crabs (Crustacea: Decapoda: Brachyura: Sesarmidae: Geosesarma De Man, 1892) from Java, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="58E18C0C464535E434F4F80BD04CFD75" type="personal">
<mods:role id="CAFCEC898E4BEB6059DF7D917998D347">
<mods:roleTerm id="DFB08604A540E2118014B31D052F568D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83AAD6D819950D73FD482733FB1E96EB">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:name id="CD3D8441BAE383190536B381DE7BE481" type="personal">
<mods:role id="7984249450523959C9D93B019D086F22">
<mods:roleTerm id="B8FCFDA5ECD10FD27C4275C8B55554AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="16EEDC872C4F5A408738DD50A9E6B744">Wowor,</mods:namePart>
</mods:name>
<mods:name id="33D4A6AC7EBE01F3CD581B48C28B59ED" type="personal">
<mods:role id="50C12C981C98A099213C79D424B2468B">
<mods:roleTerm id="6B1FB86BBE9B2360A6E1DAA2C19B7FAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65D140F0A096891D88F5805C6C5CF834">Daisy</mods:namePart>
</mods:name>
<mods:typeOfResource id="4121CD59E8157B37ED34E38473588948">text</mods:typeOfResource>
<mods:relatedItem id="79564DD909CB9CD8D24D9845A98811C0" type="host">
<mods:titleInfo id="BFA91265990D531C7030BE46E44E1FCF">
<mods:title id="EF13543EE68B96C5975E6664C44ADD61">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1DC1051815E95DA440E49269F517569D">
<mods:date id="E6902E3AF184DA20F4CC9D0A91368223">2024</mods:date>
<mods:detail id="5E0A6A43941E285B7FBB867DECA4FBC7" type="pubDate">
<mods:number id="57039BD40EE445343BC2E5BE1EABA756">2024-11-27</mods:number>
</mods:detail>
<mods:detail id="6FC44FF916D86788A5BB33D6F9EBAF5A" type="volume">
<mods:number id="9127B00C2FF7F860AC33D6FA7AA5EA4E">72</mods:number>
</mods:detail>
<mods:extent id="62CE4FEE057D6F88A55A339B0C5CFA44" unit="page">
<mods:start id="E8FEF119ED683C38E328999BA41788F0">469</mods:start>
<mods:end id="DFFFA38E191CDFB1AC1FC4AF3499112D">478</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="580B0BE7965087CE024CA95B77651317">journal article</mods:classification>
<mods:identifier id="AC067C10CC8467E26D815230E27F4431" type="DOI">10.26107/RBZ-2024-0034</mods:identifier>
<mods:identifier id="2C97FA5794B4C676F0DFA31EE81D494F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C06B48F1621136BA6F2FF3134ACFCC30" type="Zenodo-Dep">14682900</mods:identifier>
<mods:identifier id="49A27E2133DDBCB9049F8E2C65CC4AFF" type="ZooBank">10B1E24B-518C-4A85-A7C2-400C1D6D5EE7</mods:identifier>
</mods:mods>
<treatment id="C06087FDF15DFFC6FC22FA2CFE11F959" LSID="urn:lsid:plazi:treatment:C06087FDF15DFFC6FC22FA2CFE11F959" httpUri="http://treatment.plazi.org/id/C06087FDF15DFFC6FC22FA2CFE11F959" lastPageId="4" lastPageNumber="473" pageId="0" pageNumber="469">
<subSubSection id="00D36560F15DFFC2FC22FA2CFAEAFA7D" box="[982,1328,1433,1458]" pageId="0" pageNumber="469" type="nomenclature">
<paragraph id="487636EBF15DFFC2FC22FA2CFAEAFA7D" blockId="0.[982,1328,1433,1490]" box="[982,1328,1433,1458]" pageId="0" pageNumber="469">
<heading id="133E8187F15DFFC2FC22FA2CFAEAFA7D" bold="true" box="[982,1328,1433,1458]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="469" reason="2">
<emphasis id="7ABDEAF9F15DFFC2FC22FA2CFAEAFA7D" bold="true" box="[982,1328,1433,1458]" pageId="0" pageNumber="469">
<taxonomicName id="8FC94D68F15DFFC2FC22FA2CFB41FA7E" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[982,1179,1434,1458]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="469" phylum="Arthropoda" rank="species" species="riani" status="sp. nov.">
<emphasis id="7ABDEAF9F15DFFC2FC22FA2CFB41FA7E" bold="true" box="[982,1179,1434,1458]" italics="true" pageId="0" pageNumber="469">Geosesarma riani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F15DFFC2FB5EFA2CFAEAFA7D" box="[1194,1328,1433,1458]" pageId="0" pageNumber="469" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00D36560F15DFFC2FC16FA0CFAFEFA1D" box="[994,1316,1465,1490]" pageId="0" pageNumber="469" type="description">
<paragraph id="487636EBF15DFFC2FC16FA0CFAFEFA1D" blockId="0.[982,1328,1433,1490]" box="[994,1316,1465,1490]" pageId="0" pageNumber="469">
(
<figureCitation id="D0F22A6EF15DFFC2FC1EFA0CFBB1FA1E" box="[1002,1131,1466,1490]" captionStart="Fig" captionStartId="1.[113,148,2013,2035]" captionTargetBox="[114,1450,240,1997]" captionTargetId="figure-50@1.[114,1450,240,1997]" captionTargetPageId="1" captionText="Fig. 1. Live colouration of males. AD, Geosesarma riani, new species; EH, G. nigripes, new species. All specimens not preserved. Photographs: A, E, F, Achmad Rian Dietra; B, Oliver Mengedoht; C, D, G, H, Chris Lukhaup." figureDoi="http://doi.org/10.5281/zenodo.14682904" httpUri="https://zenodo.org/record/14682904/files/figure.png" pageId="0" pageNumber="469">Figs. 1AD</figureCitation>
,
<figureCitation id="D0F22A6EF15DFFC2FB8CFA0CFB5DFA1E" box="[1144,1159,1466,1490]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="0" pageNumber="469">2</figureCitation>
,
<figureCitation id="D0F22A6EF15DFFC2FB61FA0CFB0DFA1E" box="[1173,1239,1466,1490]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="0" pageNumber="469">4AC</figureCitation>
,
<figureCitation id="D0F22A6EF15DFFC2FB10FA0FFAC6FA1D" box="[1252,1308,1465,1489]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="0" pageNumber="469">5AI</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="00D36560F15DFFC2FCB7FA4FFC29F81E" pageId="0" pageNumber="469" type="materials_examined">
<paragraph id="487636EBF15DFFC2FCB7FA4FFC29F81E" blockId="0.[834,1476,1529,2002]" pageId="0" pageNumber="469">
<emphasis id="7ABDEAF9F15DFFC2FCB7FA4FFBF5F9DD" bold="true" box="[835,1071,1529,1553]" pageId="0" pageNumber="469">Material examined.</emphasis>
<materialsCitation id="F8A13CB6F15DFFC2FBCDFA4CFCA1F97E" collectingDate="2022-06" collectionCode="MZB" collectorName="Achmad Sofyan" country="Indonesia" county="Banjar Patroman" location="Purwaharja village" municipality="Purwaharja District" pageId="0" pageNumber="469" specimenCount="1" stateProvince="Jawa Barat" typeStatus="holotype">
<typeStatus id="97728849F15DFFC2FBCDFA4CFB7FF9DE" box="[1081,1189,1530,1554]" pageId="0" pageNumber="469" type="holotype">Holotype</typeStatus>
: male (12.8 ×
<quantity id="8F319B0EF15DFFC2FAA7FA4CFA61F9DE" box="[1363,1467,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="0" pageNumber="469" unit="mm" value="11.8">11.8 mm</quantity>
) (
<collectionCode id="2ED8AE2EF15DFFC2FCBFF9ACFC5CF9FE" box="[843,902,1562,1586]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="469" type="Museum">MZB</collectionCode>
Cru 5717), Pulo Majeti hamlet,
<location id="4D166030F15DFFC2FB04F9ACFA64F9FE" LSID="urn:lsid:plazi:treatment:C06087FDF15DFFC6FC22FA2CFE11F959:4D166030F15DFFC2FB04F9ACFA64F9FE" box="[1264,1470,1562,1586]" country="Indonesia" county="Banjar Patroman" municipality="Purwaharja District" name="Purwaharja village" pageId="0" pageNumber="469" stateProvince="Jawa Barat">Purwaharja village</location>
,
<collectingMunicipality id="A812AC91F15DFFC2FCB7F98CFBC0F99E" box="[835,1050,1594,1618]" pageId="0" pageNumber="469">Purwaharja District</collectingMunicipality>
, Kota Banjar, ca.
<quantity id="8F319B0EF15DFFC2FB13F98CFAEFF99E" box="[1255,1333,1594,1618]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="0" pageNumber="469" unit="km" value="2.6">2.6 km</quantity>
northeast of
<collectingCounty id="A1174E67F15DFFC2FCB7F9ECFC21F9BD" box="[835,1019,1626,1650]" pageId="0" pageNumber="469">Banjar Patroman</collectingCounty>
(Pataruman),
<collectingRegion id="8A0DF809F15DFFC2FB6EF9ECFAD7F9BD" box="[1178,1293,1626,1649]" country="Indonesia" name="Jawa Barat" pageId="0" pageNumber="469">West Java</collectingRegion>
,
<collectingCountry id="30DE767BF15DFFC2FAEFF9ECFA5CF9BE" box="[1307,1414,1626,1650]" name="Indonesia" pageId="0" pageNumber="469">Indonesia</collectingCountry>
, coll.
<collectorName id="E53C533DF15DFFC2FCB7F9CCFC2BF95E" box="[835,1009,1658,1682]" pageId="0" pageNumber="469">Achmad Sofyan</collectorName>
, transferred from Achmad Rian Dietra,
<date id="3C77102BF15DFFC2FA65F9CCFCA1F97E" pageId="0" pageNumber="469" value="2022-06">
<collectingDate id="2C33E9C3F15DFFC2FA65F9CCFCA1F97E" pageId="0" pageNumber="469" value="2022-06">June 2022</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="F8A13CB6F15DFFC2FC70F92CFAB8F91D" collectionCode="MZB" country="Indonesia" pageId="0" pageNumber="469" specimenCount="4" specimenCount-male="4" stateProvince="Jawa Barat" typeStatus="paratype">
<typeStatus id="97728849F15DFFC2FC70F92CFC2AF97E" box="[900,1008,1690,1714]" pageId="0" pageNumber="469" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="5ECFFD62F15DFFC2FC0EF92CFB91F97E" box="[1018,1099,1690,1714]" count="4" pageId="0" pageNumber="469" type="male">4 males</specimenCount>
(11.1 ×
<quantity id="8F319B0EF15DFFC2FB51F92CFADCF97D" box="[1189,1286,1690,1714]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="0" pageNumber="469" unit="mm" value="10.7">10.7 mm</quantity>
, 10.9 ×
<quantity id="8F319B0EF15DFFC2FAAFF92FFA66F97D" box="[1371,1468,1689,1713]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="0" pageNumber="469" unit="mm" value="10.5">10.5 mm</quantity>
, 10.3 ×
<quantity id="8F319B0EF15DFFC2FC65F90CFC2EF91D" box="[913,1012,1722,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="0" pageNumber="469" unit="mm" value="10.1">10.1 mm</quantity>
, 9.4 ×
<quantity id="8F319B0EF15DFFC2FBB6F90CFB4CF91E" box="[1090,1174,1722,1746]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="0" pageNumber="469" unit="mm" value="8.7">8.7 mm</quantity>
) (
<collectionCode id="2ED8AE2EF15DFFC2FB59F90CFB32F91E" box="[1197,1256,1722,1746]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="469" type="Museum">MZB</collectionCode>
Cru 5718)
</materialsCitation>
,
<materialsCitation id="F8A13CB6F15DFFC2FA9AF90FFC19F8FE" collectingDate="2022-06" collectionCode="ZRC" collectorName="Achmad Sofyan" country="Indonesia" county="Banjar Patroman" location="Purwaharja village" municipality="Purwaharja District" pageId="0" pageNumber="469" specimenCode="ZRC 2023.0320" specimenCount="5" specimenCount-male="5" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="5ECFFD62F15DFFC2FA9AF90FFA18F91E" box="[1390,1474,1721,1746]" count="5" pageId="0" pageNumber="469" type="male">5 males</specimenCount>
(11.7 ×
<quantity id="8F319B0EF15DFFC2FC6AF96FFBD9F93D" box="[926,1027,1753,1777]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="0" pageNumber="469" unit="mm" value="11.5">11.5 mm</quantity>
, 11.7 ×
<quantity id="8F319B0EF15DFFC2FB95F96CFB1FF93D" box="[1121,1221,1754,1778]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="0" pageNumber="469" unit="mm" value="11.2">11.2 mm</quantity>
, 11.5 ×
<quantity id="8F319B0EF15DFFC2FAD7F96CFA5DF93D" box="[1315,1415,1754,1778]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="0" pageNumber="469" unit="mm" value="10.7">10.7 mm</quantity>
, 11.2 ×
<quantity id="8F319B0EF15DFFC2FCA8F94CFC69F8DD" box="[860,947,1786,1810]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="0" pageNumber="469" unit="mm" value="9.9">9.9 mm</quantity>
, 11.1 ×
<quantity id="8F319B0EF15DFFC2FBE5F94CFBAEF8DE" box="[1041,1140,1786,1810]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="0" pageNumber="469" unit="mm" value="10.4">10.4 mm</quantity>
) (
<specimenCode id="186F9E90F15DFFC2FB78F94CFA9BF8DE" box="[1164,1345,1786,1810]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="469">ZRC 2023.0320</specimenCode>
), same data as holotype
</materialsCitation>
;
<specimenCount id="5ECFFD62F15DFFC2FC24F8ACFBC0F8FE" box="[976,1050,1818,1842]" count="1" pageId="0" pageNumber="469" type="male">1 male</specimenCount>
(15.3 ×
<quantity id="8F319B0EF15DFFC2FB8EF8ACFB05F8FE" box="[1146,1247,1818,1842]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="0" pageNumber="469" unit="mm" value="13.8">13.8 mm</quantity>
),
<materialsCitation id="F8A13CB6F15DFFC2FB00F8ACFA91F8BD" collectionCode="MZB" country="Indonesia" pageId="0" pageNumber="469" specimenCount="12" specimenCount-female="12" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="5ECFFD62F15DFFC2FB00F8ACFAB0F8FE" box="[1268,1386,1818,1842]" count="12" pageId="0" pageNumber="469" type="female">12 females</specimenCount>
(11.5 ×
<quantity id="8F319B0EF15DFFC2FCB7F88CFC79F89D" box="[835,931,1850,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="0" pageNumber="469" unit="mm" value="10.6">10.6 mm</quantity>
13.4 ×
<quantity id="8F319B0EF15DFFC2FBFAF88CFBABF89E" box="[1038,1137,1850,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="0" pageNumber="469" unit="mm" value="12.1">12.1 mm</quantity>
), 14 ovigerous females (11.5 ×
<quantity id="8F319B0EF15DFFC2FCB7F8EFFC7FF8BD" box="[835,933,1881,1905]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="0" pageNumber="469" unit="mm" value="10.5">10.5 mm</quantity>
12.6 × 12.0 mm) (
<collectionCode id="2ED8AE2EF15DFFC2FB67F8ECFB14F8BE" box="[1171,1230,1882,1906]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="469" type="Museum">MZB</collectionCode>
Cru 5719)
</materialsCitation>
,
<materialsCitation id="F8A13CB6F15DFFC2FAACF8ECFC35F81E" collectingDate="2023-07" collectionCode="ZRC" collectorName="Achmad Sofyan" country="Indonesia" county="Banjar Patroman" location="Purwaharja village" municipality="Purwaharja District" pageId="0" pageNumber="469" specimenCode="ZRC 2023.0561" specimenCount="2" specimenCount-female="2" stateProvince="Jawa Barat" typeStatus="paratype">
<specimenCount id="5ECFFD62F15DFFC2FAACF8ECFA18F8BE" box="[1368,1474,1882,1906]" count="2" pageId="0" pageNumber="469" type="female">2 females</specimenCount>
(13.9 × 12.0 mm, 12.0 ×
<quantity id="8F319B0EF15DFFC2FBBCF8CCFB73F85E" box="[1096,1193,1914,1938]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="0" pageNumber="469" unit="mm" value="10.7">10.7 mm</quantity>
), 1 ovigerous female (11.2 ×
<quantity id="8F319B0EF15DFFC2FCAFF82CFC65F87E" box="[859,959,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="0" pageNumber="469" unit="mm" value="10.2">10.2 mm</quantity>
) (
<specimenCode id="186F9E90F15DFFC2FC23F82CFB57F87D" box="[983,1165,1945,1970]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="469">ZRC 2023.0561</specimenCode>
), same locality as holotype, early
<date id="3C77102BF15DFFC2FC75F80CFC35F81E" box="[897,1007,1978,2002]" pageId="0" pageNumber="469" value="2023-07">
<collectingDate id="2C33E9C3F15DFFC2FC75F80CFC35F81E" box="[897,1007,1978,2002]" pageId="0" pageNumber="469" value="2023-07">July 2023</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00D36560F15DFFC6FCB7F84FFCDDFA39" lastPageId="4" lastPageNumber="473" pageId="0" pageNumber="469" type="diagnosis">
<paragraph id="487636EBF15DFFC6FCB7F84FFEE6FB79" blockId="0.[835,1474,2041,2098]" lastBlockId="4.[136,775,189,1205]" lastPageId="4" lastPageNumber="473" pageId="0" pageNumber="469">
<emphasis id="7ABDEAF9F15DFFC2FCB7F84FFC65F7DD" bold="true" box="[835,959,2041,2065]" pageId="0" pageNumber="469">Diagnosis.</emphasis>
Carapace squarish, wider than long, lateral margins almost parallel (
<figureCitation id="D0F22A6EF15DFFC2FBAEF7ACFB0FF7FE" box="[1114,1237,2074,2098]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="0" pageNumber="469">Fig. 2A, B</figureCitation>
); dorsal surface with distinct regions, anterior regions with numerous small rounded granules (
<figureCitation id="D0F22A6EF159FFC6FEA5FF6BFE1CFF39" box="[337,454,221,245]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="4" pageNumber="473">Fig. 2A, B</figureCitation>
); front deflexed, frontal lobes very broad with almost truncate margins, separated by broad median concavity; postfrontal cristae sharp (
<figureCitation id="D0F22A6EF159FFC6FD8BFEABFD2CFEF9" box="[639,758,285,310]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="4" pageNumber="473">Fig. 2AC</figureCitation>
); external orbital tooth acutely triangular, directed obliquely outwards, tip slightly extending beyond anterior part of lateral carapace margin, first epibranchial tooth triangular, low, visible, separated from margin by low, wide concavity, second epibranchial tooth small but visible (
<figureCitation id="D0F22A6EF159FFC6FDBAFE0BFD47FE19" box="[590,669,445,469]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="4" pageNumber="473">Fig. 2B</figureCitation>
). Ischium of third maxilliped subtrapezoidal, merus ovate, wide; exopod relatively slender, with long flagellum that extends beyond merus width (
<figureCitation id="D0F22A6EF159FFC6FEE8FDABFEA9FDF9" box="[284,371,541,565]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">Fig. 5A</figureCitation>
). Outer surface of palm of adult male chelae granulate; inner surface distinctly granulated but without distinct transverse comb-like ridge; dorsal margin of dactylus with 5 or 6 evenly spaced tubercle, each with chitinous tip (
<figureCitation id="D0F22A6EF159FFC6FED5FD2BFE4CFD79" box="[289,406,669,693]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="4" pageNumber="473">Fig. 2DF</figureCitation>
). Ambulatory legs with relatively more slender meri (fourth leg length-to-width ratio ca. 3.2), with sharp subdistal spine on dorsal margin; ventral margins of propodus and dactylus of adult first ambulatory leg with scattered stiff setae (
<figureCitation id="D0F22A6EF159FFC6FE11FCABFDB8FCF9" box="[485,610,797,822]" captionStart="Fig" captionStartId="2.[135,170,2016,2038]" captionTargetBox="[135,1474,208,2001]" captionTargetId="figure-8@2.[134,1475,208,2001]" captionTargetPageId="2" captionText="Fig. 2. Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="4" pageNumber="473">Fig. 2A, G</figureCitation>
). Male pleon, wide, triangular; somite 6 wide; telson subtriangular with rounded tip, slightly shorter than broad (length-to-width ratio ca. 0.9); somite 6 with convex lateral margins (
<figureCitation id="D0F22A6EF159FFC6FD6BFCCBFCD8FC59" box="[671,770,893,917]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="4" pageNumber="473">Figs. 4A</figureCitation>
,
<figureCitation id="D0F22A6EF159FFC6FF7CFC2BFF73FC79" box="[136,169,925,949]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">5B</figureCitation>
). G1 relatively slender; subdistal part of outer margin of stem before chitinous distal part rounded, outer margin of junction between stem and chitinous distal part concave, short; chitinous distal part bent about 45º along longitudinal axis, elongate, tapering, distalmost part flared in lateral view, subspatuliform in mesial view (
<figureCitation id="D0F22A6EF159FFC6FDE8FB8BFD4BFB99" box="[540,657,1085,1109]" captionStart="Fig" captionStartId="6.[135,170,1916,1938]" captionTargetBox="[134,1473,252,1889]" captionTargetId="figure-127@6.[120,1486,249,1904]" captionTargetPageId="6" captionText="Fig. 5. AI, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; JR, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, J, left third maxilliped; B, K, pleon; C, L, left G1 (ventral view); D, M, left G1 (dorsal view); E, N, distal part of left G1 (ventral view); F, O, distal part of left G1 (dorsal view); G, P, distal part of left G1 (mesial view); H, Q, distal part of left G1 (outer lateral view); I, R, left G2. Scales: A, C, D, I, J, L, M, R = 0.5 mm; B, K = 1.0 mm; EH, NQ = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14682914" httpUri="https://zenodo.org/record/14682914/files/figure.png" pageId="4" pageNumber="473">Fig. 5CH</figureCitation>
). Vulva on anterior part of somite 6, with raised vulvar cover, directed mesially (
<figureCitation id="D0F22A6EF159FFC6FF06FBCBFE9DFB59" box="[242,327,1149,1173]" captionStart="Fig" captionStartId="5.[113,148,1614,1636]" captionTargetBox="[113,1451,189,1599]" captionTargetId="figure-13@5.[112,1453,189,1599]" captionTargetPageId="5" captionText="Fig. 4. A, Geosesarma riani, new species, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java; B, C, G. riani, new species, paratype female (13.2 × 11.8 mm) (MZB Cru 5719), Central Java; D, G. nigripes, new species, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java; E, F, G. nigripes, new species, paratype female (15.0 × 13.3 mm) (MZB Cru 5721), Central Java. A, D, male thoracic sternum and pleon; B, E, female pleon; C, F, sternopleonal cavity showing vulvae." figureDoi="http://doi.org/10.5281/zenodo.14682912" httpUri="https://zenodo.org/record/14682912/files/figure.png" pageId="4" pageNumber="473">Fig. 4C</figureCitation>
). Preserved egg size 1.6 × 1.3 1.5 ×
<quantity id="8F319B0EF159FFC6FD11FBCBFF6BFB79" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="473" unit="mm" value="1.5">1.5 mm</quantity>
in diameter.
</paragraph>
<caption id="1CB66663F15CFFC3FF85F86BFBC0F7C3" ID-DOI="http://doi.org/10.5281/zenodo.14682904" ID-Zenodo-Dep="14682904" httpUri="https://zenodo.org/record/14682904/files/figure.png" pageId="1" pageNumber="470" startId="1.[113,148,2013,2035]" targetBox="[114,1450,240,1997]" targetPageId="1" targetType="figure">
<paragraph id="487636EBF15CFFC3FF85F86BFBC0F7C3" blockId="1.[113,1452,2013,2063]" pageId="1" pageNumber="470">
Fig. 1. Live colouration of males. AD,
<taxonomicName id="8FC94D68F15CFFC3FDFDF86BFD62F83E" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[521,696,2013,2034]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="riani" status="sp. nov.">
<emphasis id="7ABDEAF9F15CFFC3FDFDF86BFD62F83E" box="[521,696,2013,2034]" italics="true" pageId="1" pageNumber="470">Geosesarma riani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F15CFFC3FD33F86BFC9AF83F" box="[711,832,2013,2035]" pageId="1" pageNumber="470" rank="species">new species</taxonomicNameLabel>
; EH,
<taxonomicName id="8FC94D68F15CFFC3FC7CF86BFC22F83E" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[904,1016,2013,2034]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="470" phylum="Arthropoda" rank="species" species="nigripes" status="new species">
<emphasis id="7ABDEAF9F15CFFC3FC7CF86BFC22F83E" box="[904,1016,2013,2034]" italics="true" pageId="1" pageNumber="470">G. nigripes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F15CFFC3FBF2F86BFBA5F83F" box="[1030,1151,2013,2035]" pageId="1" pageNumber="470" rank="species">new species</taxonomicNameLabel>
. All specimens not preserved. Photographs: A, E, F, Achmad Rian Dietra; B, Oliver Mengedoht; C, D, G, H, Chris Lukhaup.
</paragraph>
</caption>
<caption id="1CB66663F15FFFC0FF73F856FD75F7FC" ID-DOI="http://doi.org/10.5281/zenodo.14682906" ID-Zenodo-Dep="14682906" httpUri="https://zenodo.org/record/14682906/files/figure.png" pageId="2" pageNumber="471" startId="2.[135,170,2016,2038]" targetBox="[135,1474,208,2001]" targetPageId="2" targetType="figure">
<paragraph id="487636EBF15FFFC0FF73F856FD75F7FC" blockId="2.[135,1475,2016,2096]" pageId="2" pageNumber="471">
Fig. 2.
<taxonomicName id="8FC94D68F15FFFC0FF3AF857FEA1F83A" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[206,379,2017,2038]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="471" phylum="Arthropoda" rank="species" species="riani" status="sp. nov.">
<emphasis id="7ABDEAF9F15FFFC0FF3AF857FEA1F83A" box="[206,379,2017,2038]" italics="true" pageId="2" pageNumber="471">Geosesarma riani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F15FFFC0FE73F857FE24F83A" box="[391,510,2016,2038]" pageId="2" pageNumber="471" rank="species">new species</taxonomicNameLabel>
, holotype male (12.8 × 11.8 mm) (MZB Cru 5717), West Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg.
</paragraph>
</caption>
<caption id="1CB66663F15EFFC1FF85F869FD0BF7E3" ID-DOI="http://doi.org/10.5281/zenodo.14682910" ID-Zenodo-Dep="14682910" httpUri="https://zenodo.org/record/14682910/files/figure.png" pageId="3" pageNumber="472" startId="3.[113,148,2015,2037]" targetBox="[114,1452,209,2001]" targetPageId="3" targetType="figure">
<paragraph id="487636EBF15EFFC1FF85F869FD0BF7E3" blockId="3.[113,1454,2015,2095]" pageId="3" pageNumber="472">
Fig. 3.
<taxonomicName id="8FC94D68F15EFFC1FF4DF856FE5CF839" authorityName="Ng &amp; Wowor &amp; Daisy" authorityYear="2024" box="[185,390,2016,2037]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="472" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
<emphasis id="7ABDEAF9F15EFFC1FF4DF856FE5CF839" box="[185,390,2016,2037]" italics="true" pageId="3" pageNumber="472">Geosesarma nigripes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="618E5782F15EFFC1FE60F856FDD1F839" box="[404,523,2015,2037]" pageId="3" pageNumber="472" rank="species">new species</taxonomicNameLabel>
, holotype male (12.3 × 11.3 mm) (MZB Cru 5720), Central Java. A, overall dorsal habitus; B, dorsal view of carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, dorsal view of right cheliped; F, submarginal view of dactylus of right chela; G, left fourth ambulatory leg.
</paragraph>
</caption>
<paragraph id="487636EBF159FFC6FF7CFB6BFEEBFAF9" blockId="4.[136,775,1245,1333]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FF7CFB6BFE81FB39" bold="true" box="[136,347,1245,1269]" pageId="4" pageNumber="473">Notes on habitat.</emphasis>
This species inhabits wet areas in a forest, ground which is covered with stones and rocks, at about
<quantity id="8F319B0EF159FFC6FF38FAABFEDEFAF9" box="[204,260,1309,1333]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="473" unit="m" value="60.0">60 m</quantity>
asl.
</paragraph>
<paragraph id="487636EBF159FFC6FF7CFAEBFCDDFA39" blockId="4.[136,775,1373,1525]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FF7CFAEBFF05FAB9" bold="true" box="[136,223,1373,1397]" pageId="4" pageNumber="473">Colour.</emphasis>
Carapace grey to greyish-blue, with posterolateral regions dark brown, H-shaped grooves dark brown; frontal surfaces of cephalothorax orange to dull red; eyes iridescent yellow; chelipeds orange; ambulatory legs pale orange to orangish-red, with numerous small dark spots (
<figureCitation id="D0F22A6EF159FFC6FD77FA6BFD21FA39" box="[643,763,1501,1525]" captionStart="Fig" captionStartId="1.[113,148,2013,2035]" captionTargetBox="[114,1450,240,1997]" captionTargetId="figure-50@1.[114,1450,240,1997]" captionTargetPageId="1" captionText="Fig. 1. Live colouration of males. AD, Geosesarma riani, new species; EH, G. nigripes, new species. All specimens not preserved. Photographs: A, E, F, Achmad Rian Dietra; B, Oliver Mengedoht; C, D, G, H, Chris Lukhaup." figureDoi="http://doi.org/10.5281/zenodo.14682904" httpUri="https://zenodo.org/record/14682904/files/figure.png" pageId="4" pageNumber="473">Fig. 1AD</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="00D36560F159FFC6FF7CF9ABFE11F959" pageId="4" pageNumber="473" type="etymology">
<paragraph id="487636EBF159FFC6FF7CF9ABFE11F959" blockId="4.[136,776,1565,1685]" pageId="4" pageNumber="473">
<emphasis id="7ABDEAF9F159FFC6FF7CF9ABFED0F9F9" bold="true" box="[136,266,1565,1589]" pageId="4" pageNumber="473">Etymology.</emphasis>
The species is named after Achmad Rian Dietra or Rian, who has great interest in all ornamental Indonesian crustaceans especially
<taxonomicName id="8FC94D68F159FFC6FE8AF9E8FDE3F9B9" box="[382,569,1630,1653]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="7ABDEAF9F159FFC6FE8AF9E8FDD9F9B9" box="[382,515,1630,1653]" italics="true" pageId="4" pageNumber="473">Geosesarma</emphasis>
spp.
</taxonomicName>
, and has helped us obtain material for this study.
</paragraph>
</subSubSection>
</treatment>
</document>