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<document id="51F23C24320743CAB19BBDC6CBA03634" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFCAFFC3231F151BECD3FD3B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Chrysotus dot Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="307" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="305" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7F1882515B6A256A0F53899AE9A1698B">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
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<mods:namePart id="4256AEA0B39E5A952C49B103E8EBBCE1">Grootaert, Patrick</mods:namePart>
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<mods:namePart id="1FF7EC112FF455FE2711006276071789">Velde,</mods:namePart>
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<mods:namePart id="12E723E3B491A55E53A2E1A8433ECD39">Isabella Van de</mods:namePart>
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<mods:title id="F13997BA03B25F159900C1CE80EF60A0">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="736A669FF045E85A3E63CD15576CE74C">2024</mods:date>
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<taxonomicName id="09BB4D63FFCAFFC5231F151BE9F1F97B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[304,458,1661,1685]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCAFFC5231F151BE9F1F97B" bold="true" box="[304,458,1661,1685]" italics="true" pageId="2" pageNumber="305">Chrysotus dot</emphasis>
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,
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(
<figureCitation id="56802A65FFCAFFC523BC15FBE9DBF95B" box="[403,480,1693,1717]" captionStart="Fig" captionStartId="2.[835,873,843,865]" captionTargetBox="[835,1474,189,828]" captionTargetId="figure-671@2.[835,1474,189,828]" captionTargetPageId="2" captionText="Fig. 2. Chrysotus dot, new species, habitus paratype male (ZRCBDP0083923). Photograph by Miteshravin S/O Ravindran." figureDoi="http://doi.org/10.5281/zenodo.14683060" httpUri="https://zenodo.org/record/14683060/files/figure.png" pageId="2" pageNumber="305">Figs. 2</figureCitation>
,
<figureCitation id="56802A65FFCAFFC523C115FBE9C0F95B" box="[494,507,1693,1717]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="2" pageNumber="305">3</figureCitation>
)
</paragraph>
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<subSubSection id="86A1656BFFCAFFC522A715BBEC41FAE3" pageId="2" pageNumber="305" type="materials_examined">
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<emphasis id="FCCFEAF2FFCAFFC522A715BBE902F91B" bold="true" box="[136,313,1757,1781]" pageId="2" pageNumber="305">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFCAFFC5236A15BBE97BF89B" collectingDate="2019-03-19" collectionCode="OBS, LKCNHM" collectorName="P. Grootaert &amp; GenBank" country="Singapore" county="Tock Kew" location="Pulau Ubin" municipality="Sungei" pageId="2" pageNumber="305" specimenCode="PU22, ZRCBDP0317760" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFCAFFC5236A15BBE997F91B" box="[325,428,1757,1781]" pageId="2" pageNumber="305" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="B6AC7670FFCAFFC523D215BBEA4BF91B" box="[509,624,1757,1781]" name="Singapore" pageId="2" pageNumber="305">Singapore</collectingCountry>
,
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,
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[= River]
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basin (
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),
<date id="BA051020FFCAFFC52076159BEB38F8FB" box="[601,771,1789,1813]" pageId="2" pageNumber="305" value="2019-03-19">
<collectingDate id="AA41E9C8FFCAFFC52076159BEB38F8FB" box="[601,771,1789,1813]" pageId="2" pageNumber="305" value="2019-03-19">19 March 2019</collectingDate>
</date>
,
<collectingMethod id="17FA4EF7FFCAFFC522A5147BE936F8DB" box="[138,269,1821,1845]" pageId="2" pageNumber="305">Malaise trap</collectingMethod>
(
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) in back mangroves (
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, leg.
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; in ethanol,
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).
<collectorName id="634E5336FFCAFFC52016145BEAA6F8BB" box="[569,669,1853,1877]" pageId="2" pageNumber="305">GenBank</collectorName>
accession code: PP893289.
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</paragraph>
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<materialsCitation id="7ED33CBDFFCAFFC522A714FBE929F83B" collectionCode="OBS" country="Singapore" location="Pulau Ubin" municipality="Sungei Tock Kew" pageId="2" pageNumber="305" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFCAFFC522A714FBE932F85B" bold="true" box="[136,265,1949,1973]" pageId="2" pageNumber="305">
<typeStatus id="11008842FFCAFFC522A714FBE93FF85B" box="[136,260,1949,1973]" pageId="2" pageNumber="305" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCountry id="B6AC7670FFCAFFC5233A14FBE9B7F85B" box="[277,396,1949,1973]" name="Singapore" pageId="2" pageNumber="305">Singapore</collectingCountry>
,
<location id="CB64603BFFCAFFC523B414FBEA18F85B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCAFFC523B414FBEA18F85B" box="[411,547,1949,1973]" country="Singapore" municipality="Sungei Tock Kew" name="Pulau Ubin" pageId="2" pageNumber="305">Pulau Ubin</location>
,
<collectingMunicipality id="2E60AC9AFFCAFFC5201C14FBEB3DF85B" box="[563,774,1949,1973]" pageId="2" pageNumber="305">Sungei Tock Kew</collectingMunicipality>
basin (
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)
</materialsCitation>
,
<materialsCitation id="7ED33CBDFFCAFFC5230E14DBE93FF81B" collectingDate="2019-03-19" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="31" specimenCount-female="22" specimenCount-male="9" typeStatus="paratype">
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,
<specimenCount id="D8BDFD69FFCAFFC523A814DBEA39F83B" box="[391,514,1981,2005]" count="22" pageId="2" pageNumber="305" type="female">22 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5203F14DBEA74F83B" box="[528,591,1981,2005]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5207214DBEB3DF83B" box="[605,774,1981,2005]" pageId="2" pageNumber="305" value="2019-03-19">
<collectingDate id="AA41E9C8FFCAFFC5207214DBEB3DF83B" box="[605,774,1981,2005]" pageId="2" pageNumber="305" value="2019-03-19">19 March 2019</collectingDate>
</date>
(Ma10640)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5233F14BBEB38F81B" box="[272,771,2013,2037]" collectingDate="2019-03-26" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5233F14BBE940F81B" box="[272,379,2013,2037]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC523A814BBE9FDF81B" box="[391,454,2013,2037]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC523FD14BBEA43F81B" box="[466,632,2013,2037]" pageId="2" pageNumber="305" value="2019-03-26">
<collectingDate id="AA41E9C8FFCAFFC523FD14BBEA43F81B" box="[466,632,2013,2037]" pageId="2" pageNumber="305" value="2019-03-26">26 March 2019</collectingDate>
</date>
(MA10955)
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;
<materialsCitation id="7ED33CBDFFCAFFC522A7149BEAD1F7FB" box="[136,746,2045,2069]" collectingDate="2019-04-02" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="15" specimenCount-female="9" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC522A7149BE8DEF7FB" box="[136,229,2045,2069]" count="6" pageId="2" pageNumber="305" type="male">6 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC522DB149BE95CF7FB" box="[244,359,2045,2069]" count="9" pageId="2" pageNumber="305" type="female">9 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC52359149BE983F7FB" box="[374,440,2045,2069]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC523E8149BEA66F7FB" box="[455,605,2045,2069]" pageId="2" pageNumber="305" value="2019-04-02">
<collectingDate id="AA41E9C8FFCAFFC523E8149BEA66F7FB" box="[455,605,2045,2069]" pageId="2" pageNumber="305" value="2019-04-02">2 April 2019</collectingDate>
</date>
(Ma10718)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC520D6149BEAD0F7DB" collectingDate="2019-04-16" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="8" specimenCount-female="2" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC520D6149BE8F6F7DB" count="6" pageId="2" pageNumber="305" type="male">6 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC522F21B7BE96BF7DB" box="[221,336,2077,2101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC523701B7BE99AF7DB" box="[351,417,2077,2101]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5239E1B7BEA6DF7DB" box="[433,598,2077,2101]" pageId="2" pageNumber="305" value="2019-04-16">
<collectingDate id="AA41E9C8FFCAFFC5239E1B7BEA6DF7DB" box="[433,598,2077,2101]" pageId="2" pageNumber="305" value="2019-04-16">16 April 2019</collectingDate>
</date>
(MA10986)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC520D51B7BED28FC23" box="[762,1299,949,2101]" collectingDate="2019-04-24" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC520D51B7BEBA1FC23" box="[762,922,949,2101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5218A10D3EBDEFC23" box="[933,997,949,973]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC521DF10D3ECBCFC23" box="[1008,1159,949,973]" pageId="2" pageNumber="305" value="2019-04-24">
<collectingDate id="AA41E9C8FFCAFFC521DF10D3ECBCFC23" box="[1008,1159,949,973]" pageId="2" pageNumber="305" value="2019-04-24">24 April 2019</collectingDate>
</date>
(MA10972)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5273110D3EC57FC03" collectingDate="2019-05-15" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5273110D3ED4FFC23" box="[1310,1396,949,973]" count="3" pageId="2" pageNumber="305" type="male">3 males</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5275010D3ED84FC23" box="[1407,1471,949,973]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5216C10B3EBE0FC03" box="[835,987,981,1006]" pageId="2" pageNumber="305" value="2019-05-15">
<collectingDate id="AA41E9C8FFCAFFC5216C10B3EBE0FC03" box="[835,987,981,1006]" pageId="2" pageNumber="305" value="2019-05-15">15 May 2019</collectingDate>
</date>
(MA11506)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5265510B3EC7FFBE3" collectingDate="2019-05-28" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5265510B3ECF1FC03" box="[1146,1226,981,1005]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526F610B3ED72FC03" box="[1241,1353,981,1005]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5277810B3EDA3FC03" box="[1367,1432,981,1005]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5278810B3EB8EFBE3" pageId="2" pageNumber="305" value="2019-05-28">
<collectingDate id="AA41E9C8FFCAFFC5278810B3EB8EFBE3" pageId="2" pageNumber="305" value="2019-05-28">28 May 2019</collectingDate>
</date>
(MA11189)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5267D1093EC29FBC3" collectingDate="2019-06-03" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5267D1093EC9BFBE3" box="[1106,1184,1013,1037]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526801093ED26FBE3" box="[1199,1309,1013,1037]" count="1" pageId="2" pageNumber="305" type="female">1 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC527041093ED50FBE3" box="[1323,1387,1013,1037]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC527561093EB40FBC3" pageId="2" pageNumber="305" value="2019-06-03">
<collectingDate id="AA41E9C8FFCAFFC527561093EB40FBC3" pageId="2" pageNumber="305" value="2019-06-03">3 June 2019</collectingDate>
</date>
(MA10982)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5260D1773EC3DFBA3" collectingDate="2019-06-11" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="9" specimenCount-female="4" specimenCount-male="5" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5260D1773ECBBFBC3" box="[1058,1152,1045,1069]" count="5" pageId="2" pageNumber="305" type="male">5 males</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526BF1773ED3EFBC3" box="[1168,1285,1045,1069]" count="4" pageId="2" pageNumber="305" type="female">4 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC5273B1773ED63FBC3" box="[1300,1368,1045,1069]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC527481773EB42FBA3" pageId="2" pageNumber="305" value="2019-06-11">
<collectingDate id="AA41E9C8FFCAFFC527481773EB42FBA3" pageId="2" pageNumber="305" value="2019-06-11">11 June 2019</collectingDate>
</date>
(MA11180)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC5263D1753EBFCFB83" collectingDate="2019-06-19" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC5263D1753EC65FBA3" box="[1042,1118,1077,1101]" count="1" pageId="2" pageNumber="305" type="male">1 male</specimenCount>
,
<specimenCount id="D8BDFD69FFCAFFC526441753ECE3FBA3" box="[1131,1240,1077,1101]" count="2" pageId="2" pageNumber="305" type="female">2 females</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC526CB1753ED18FBA3" box="[1252,1315,1077,1101]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC5271F1753EDF8FBA3" box="[1328,1475,1077,1101]" pageId="2" pageNumber="305" value="2019-06-19">
<collectingDate id="AA41E9C8FFCAFFC5271F1753EDF8FBA3" box="[1328,1475,1077,1101]" pageId="2" pageNumber="305" value="2019-06-19">19 June 2019</collectingDate>
</date>
(MA10976)
</materialsCitation>
;
<materialsCitation id="7ED33CBDFFCAFFC521FA1733ED84FB83" box="[981,1471,1109,1133]" collectingDate="2019-06-25" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC521FA1733EC03FB83" box="[981,1080,1109,1133]" count="1" pageId="2" pageNumber="305" type="female">1 female</specimenCount>
,
<specimenCode id="9E1D9E9BFFCAFFC526691733ECBDFB83" box="[1094,1158,1109,1133]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
,
<date id="BA051020FFCAFFC526BB1733ED17FB83" box="[1172,1324,1109,1133]" pageId="2" pageNumber="305" value="2019-06-25">
<collectingDate id="AA41E9C8FFCAFFC526BB1733ED17FB83" box="[1172,1324,1109,1133]" pageId="2" pageNumber="305" value="2019-06-25">25 June 2019</collectingDate>
</date>
, (MA10967)
</materialsCitation>
. Barcodes of
<materialsCitation id="7ED33CBDFFCAFFC521E11713ECD8FB63" box="[974,1251,1141,1165]" collectionCode="PU" country="Singapore" pageId="2" pageNumber="305" specimenCode="PU22" specimenCount="58" typeStatus="paratype">
<specimenCount id="D8BDFD69FFCAFFC521E11713EC5BFB63" box="[974,1120,1141,1165]" count="58" pageId="2" pageNumber="305" type="generic">58 specimens</specimenCount>
from
<specimenCode id="9E1D9E9BFFCAFFC526891713ECD8FB63" box="[1190,1251,1141,1165]" collectionCode="PU" country="USA" name="Princeton University" pageId="2" pageNumber="305">PU22</specimenCode>
</materialsCitation>
are available.
</paragraph>
<paragraph id="CE0436E0FFCAFFC5216C17D3EC41FAE3" blockId="2.[835,1474,1205,1293]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C17D3EC53FB23" bold="true" box="[835,1128,1205,1229]" pageId="2" pageNumber="305">Other material examined</emphasis>
. 2,357 barcodes of
<taxonomicName id="09BB4D63FFCAFFC5276C17D0EDBDFB23" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1347,1414,1205,1229]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot" status="new species">
<emphasis id="FCCFEAF2FFCAFFC5276C17D0EDBDFB23" box="[1347,1414,1205,1229]" italics="true" pageId="2" pageNumber="305">C. dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCAFFC527B917D0EBADFB03" pageId="2" pageNumber="305" rank="species">new species</taxonomicNameLabel>
, are available from Pulau Ubin, Pulau [=
<collectingCountry id="B6AC7670FFCAFFC5275D17B3ED80FB03" box="[1394,1467,1237,1261]" name="Iceland" pageId="2" pageNumber="305">Island</collectingCountry>
] Semakau, and Coney Island.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC5216C1653ED2DFA63" pageId="2" pageNumber="305" type="etymology">
<paragraph id="CE0436E0FFCAFFC5216C1653ED2DFA63" blockId="2.[834,1474,1333,1421]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C1653EBFEFAA3" bold="true" box="[835,965,1333,1357]" pageId="2" pageNumber="305">Etymology.</emphasis>
The species is named after the brown spot or dot that is present on the fore femora of the males and females. The English word
<taxonomicName id="09BB4D63FFCAFFC5263F1613EC7BFA63" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[1040,1088,1397,1421]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="305" phylum="Arthropoda" rank="species" species="dot">“dot</taxonomicName>
” is used as a noun.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCAFFC3216C16D3E91BFDDB" lastPageId="4" lastPageNumber="307" pageId="2" pageNumber="305" type="diagnosis">
<paragraph id="CE0436E0FFCAFFC5216C16D3EC85F823" blockId="2.[834,1476,1461,1997]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C16D3EB83FA23" bold="true" box="[835,952,1461,1485]" pageId="2" pageNumber="305">Diagnosis.</emphasis>
A small species (
<quantity id="09439B05FFCAFFC5265716D3ECC0FA23" box="[1144,1275,1461,1485]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
) with a black spot located dorsally on the middle of the fore femur in both sexes. Four dorsocentrals, 5 uniserial acrostichals. Haltere yellowish. Legs yellow, with apical 2 tarsomeres of all legs brown. Male: Antenna yellowish brown, pedicel contrastingly darker brown than the yellowish brown postpedicel. Frons very wide, eyes touching near middle of face, with at least 4 rows of enlarged eye facets on each side of face. Palpus yellow, large, oval, hiding proboscis and shorter in females, lacking black bristling. Apical 2 tarsomeres of all legs brown, slightly dorsoventrally flattened. Pulvilli present on all legs, though minute and narrow. Sternite 8 lacking macrosetae. Cerci yellow, ventral surstylus dark brown. Tip of aedeagus widened with complex structure. Female: palpus brown, not hiding the proboscis, covered with black bristles and a distinct apical. Legs coloured like in males. Apical tarsomeres of all legs brownish but not widened.
</paragraph>
<paragraph id="CE0436E0FFCAFFC5216C1493EDBBF7E3" blockId="2.[835,1408,2037,2061]" box="[835,1408,2037,2061]" pageId="2" pageNumber="305">
<emphasis id="FCCFEAF2FFCAFFC5216C1493EBBFF7E3" bold="true" box="[835,900,2037,2061]" pageId="2" pageNumber="305">Male.</emphasis>
Length: body
<quantity id="09439B05FFCAFFC526091493EC97F7E3" box="[1062,1196,2037,2061]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" metricValueMax="1.3" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.2" valueMax="1.3" valueMin="1.1">1.11.3 mm</quantity>
; wing
<quantity id="09439B05FFCAFFC526D91493ED47F7E3" box="[1270,1404,2037,2061]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="305" unit="mm" value="1.15" valueMax="1.2" valueMin="1.1">1.11.2 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCBFFC423621314ECEBFF64" blockId="3.[333,1232,114,138]" box="[333,1232,114,138]" pageId="3" pageNumber="306">Grootaert &amp; Van de Velde: New long-legged flies from mangroves on Pulau Ubin</paragraph>
<caption id="9AC46668FFCBFFC4225E1776EA7EFBAC" ID-DOI="http://doi.org/10.5281/zenodo.14683064" ID-Zenodo-Dep="14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306" startId="3.[113,148,1040,1062]" targetBox="[95,1465,167,1091]" targetPageId="3" targetType="figure">
<paragraph id="CE0436E0FFCBFFC4225E1776EA7EFBAC" blockId="3.[113,1453,1040,1091]" pageId="3" pageNumber="306">
Fig. 3.
<taxonomicName id="09BB4D63FFCBFFC422971776E904FBCB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[184,319,1040,1061]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="306" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCBFFC422971776E904FBCB" box="[184,319,1040,1061]" italics="true" pageId="3" pageNumber="306">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCBFFC423641776E9F9FBC8" box="[331,450,1040,1062]" pageId="3" pageNumber="306" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFCBFFC4225E171CE9D4FADC" blockId="3.[113,753,1146,1330]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E171CE88EFB7C" bold="true" box="[113,181,1146,1170]" pageId="3" pageNumber="306">Head.</emphasis>
Frons very wide, metallic green. Eyes touching near middle of face, with at least 4 rows of enlarged eye facets at each side of face. All bristles black. A pair of long retroclinate, diverging ocellar bristles, a pair of proclinate vertical bristles as long as ocellars. Postoculars, short, fine, uniserial above, not distinct below.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E163FE9D1F9FC" blockId="3.[113,754,1369,1554]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E163FE8DCFA9F" bold="true" box="[113,231,1369,1393]" pageId="3" pageNumber="306">Antennae.</emphasis>
Yellowish brown, pedicel contrastingly darker brown than postpedicel which is more yellowish. Pedicel with dorsal apical bristle nearly as long as pedicel is high. Postpedicel kidney-shaped, arista apical near middle, set with minute hairs. Palpus yellow, very large, oval, hiding proboscis, lacking distinct bristles.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E155CE9B0F95C" blockId="3.[112,753,1594,1714]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E155CE8EAF9BC" bold="true" box="[113,209,1594,1618]" pageId="3" pageNumber="306">Thorax.</emphasis>
Shiny metallic green. All bristles black. Four dorsocentrals almost equally long. Five uniserial acrostichals. A pair of widely separated marginal scutellars 2.5 times as long as scutellum is long.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E15BCEA28F8DC" blockId="3.[112,752,1754,1842]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4225E15BCE895F91C" bold="true" box="[113,174,1754,1778]" pageId="3" pageNumber="306">Legs.</emphasis>
Yellow, fore femur with a blackish brown spot located dorsally in middle. Mid coxa brown, hind coxae paler brown. Apical 2 tarsomeres of all legs brown.
</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E143CEACBF85C" blockId="3.[112,752,1882,1970]" pageId="3" pageNumber="306">Fore legs. Coxa with indistinct pale apical bristles. Femur and tibia lacking distinct bristles. Length femur, tibia, tarsomeres (in mm): 0.33: 0.34: 0.12: 0.047: 0.039: 0.032: 0.032.</paragraph>
<paragraph id="CE0436E0FFCBFFC4225E14BCEBEEFB1C" blockId="3.[113,753,2010,2066]" lastBlockId="3.[813,1452,1146,1266]" pageId="3" pageNumber="306">Mid legs. Coxa lacking an exterior bristle, but a long bristle on middle of margin between anterior and dorsal part of the coxa. Femur with a brown posterior preapical bristle. Tibia with a short anterodorsal in basal quarter. Length femur, tibia, tarsomeres (in mm): 0.39: 0.39: 0.19: 0.09: 0.06: 0.032: 0.032.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102167CED23FA3C" blockId="3.[812,1452,1306,1490]" pageId="3" pageNumber="306">Hind legs. A brown exterior bristle near base of coxa, longer than coxa is high and a slightly shorter apical bristle. Trochanter with a long dorsal bristle longer than trochanter. Femur with 3 preapical anteroventral bristles, middle bristle longest, longer than width of femur. Tibia lacking distinct bristles except for a few short apical bristles.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102169CEC71F9DC" blockId="3.[813,1452,1530,1586]" pageId="3" pageNumber="306">Length femur, tibia, tarsomeres (in mm): 0.39: 0.39: 0.17: 0.08: 0.05: 0.32: 0.05.</paragraph>
<paragraph id="CE0436E0FFCBFFC42102153FEB8EF95C" blockId="3.[813,1452,1625,1714]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC42102153FEB43F99F" bold="true" box="[813,888,1625,1649]" pageId="3" pageNumber="306">Wings.</emphasis>
Brownish tinge with pale brown veins. Squama brown with 2 pale brown bristles. Haltere yellowish (not clear white).
</paragraph>
<paragraph id="CE0436E0FFCBFFC4210215BFEBCEF8DC" blockId="3.[812,1452,1753,1842]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC4210215BFEB98F91F" bold="true" box="[813,931,1753,1777]" pageId="3" pageNumber="306">Abdomen.</emphasis>
Brown, indistinctly shiny. All bristles black and short. Sternite 8 lacking macrosetae though a row of bristles is present (
<figureCitation id="56802A65FFCBFFC4218B147CEBDCF8DC" box="[932,999,1818,1842]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFCBFFC42102143CEDB3F7FC" blockId="3.[813,1453,1881,2066]" pageId="3" pageNumber="306">
<emphasis id="FCCFEAF2FFCBFFC42102143CEBD5F89F" bold="true" box="[813,1006,1881,1906]" pageId="3" pageNumber="306">Male terminalia.</emphasis>
<figureCitation id="56802A65FFCBFFC421D7143CEC07F89C" box="[1016,1084,1882,1906]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3</figureCitation>
. Cercus yellow, surstylus brown. Epandrial lobe with 3 bristles, the apical bristle as long as lobe is wide, other 2 bristles minute. Ventral surstylus with a strong subapical bristle and a smaller bristle below. Apex of aedeagus widened with a complex structure (
<figureCitation id="56802A65FFCBFFC4270414BCED9AF81C" box="[1323,1441,2010,2034]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3AC</figureCitation>
). Postgonite short, densely set with short hairs (
<figureCitation id="56802A65FFCBFFC42709149CED47F7FC" box="[1318,1404,2042,2066]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="3" pageNumber="306">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A713DBEAC3FF3B" blockId="4.[136,760,189,213]" box="[136,760,189,213]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A713DBE8D9FF3B" bold="true" box="[136,226,189,213]" pageId="4" pageNumber="307">Female.</emphasis>
Length: body
<quantity id="09439B05FFCCFFC323AB13DBEA31FF3B" box="[388,522,189,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="4" pageNumber="307" unit="mm" value="1.25" valueMax="1.3" valueMin="1.2">1.21.3 mm</quantity>
; wing
<quantity id="09439B05FFCCFFC3207B13DBEACFFF3B" box="[596,756,189,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.105" metricValueMax="1.11" metricValueMin="1.1" pageId="4" pageNumber="307" unit="mm" value="1.105" valueMax="1.11" valueMin="1.1">1.101.11 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7139BE91BFDDB" blockId="4.[135,777,253,565]" pageId="4" pageNumber="307">Similar to males in colouration of legs. Palpus brown and much smaller than in males, with longer black bristles and a distinct apical. Proboscis yellowish, not hidden by palpus, long, about half as long as eye is high. Face as wide as postpedicel, apically protruding and pointed. Postpedicel reniform as in males, but more uniformly brown. Eye facets not enlarged and eyes not as wide as in males so that the row of lower postoculars is visible. Postoculars black above, longer and yellow below. Marginal bristles on tergites longer than in males.</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC322A7113BED82FF1B" pageId="4" pageNumber="307" type="discussion">
<paragraph id="CE0436E0FFCCFFC322A7113BEB3DFC3B" blockId="4.[135,776,605,981]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7113BE937FD9B" bold="true" box="[136,268,605,629]" pageId="4" pageNumber="307">Comments.</emphasis>
This small new species is easily recognised by the yellow legs with a black spot located dorsally in the middle of the fore femora and this is in both sexes. The postpedicel is yellowish brown and somewhat kidney-shaped with an apical arista. The male has a very large oval yellowish palpus, while the female has a smaller brown palpus. It is the most common
<taxonomicName id="09BB4D63FFCCFFC3237B107BEA28FCDB" box="[340,531,797,821]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3237B107BE9FBFCDB" box="[340,448,797,821]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
in back mangroves. It hardly visits front mangroves and was never observed in non-mangrove biotopes where many species of the entirely black
<taxonomicName id="09BB4D63FFCCFFC322E7101BE9BCFC7B" box="[200,391,893,917]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322E7101BE90FFC7B" box="[200,308,893,917]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
are dominant. The genetic variation measured by the number of haplotypes is low, especially in comparison with the terrestrial black
<taxonomicName id="09BB4D63FFCCFFC3201010DBEB3DFC3B" box="[575,774,957,981]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFCCFFC3201010DBEA90FC3B" box="[575,683,957,981]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7109BEAB9FB1B" blockId="4.[136,776,1021,1270]" pageId="4" pageNumber="307">
The tip of the aedeagus in
<taxonomicName id="09BB4D63FFCCFFC32380109BEA78FBFB" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[431,579,1021,1045]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCCFFC32380109BEA78FBFB" box="[431,579,1021,1045]" italics="true" pageId="4" pageNumber="307">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC3207D1098EAECFBFB" box="[594,727,1021,1045]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, has a complex structure with folds and a spine-like projection (
<figureCitation id="56802A65FFCCFFC322BF175BE936FBBB" box="[144,269,1085,1109]" captionStart="Fig" captionStartId="3.[113,148,1040,1062]" captionTargetBox="[95,1465,167,1091]" captionTargetId="figure-16@3.[95,1465,167,1042]" captionTargetPageId="3" captionText="Fig. 3. Chrysotus dot, new species, male terminalia. A, Hypopygium lateral; B, hypopygium ventral; C, Detail postgonite and tip phallus. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683064" httpUri="https://zenodo.org/record/14683064/files/figure.png" pageId="4" pageNumber="307">Fig. 3AC</figureCitation>
), similar to species of the
<taxonomicName id="09BB4D63FFCCFFC3206B1758EB3CFBBB" authorityName="Wei &amp; Yang" authorityYear="2007" box="[580,775,1085,1109]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="leigongshanus">
<emphasis id="FCCFEAF2FFCCFFC3206B1758EB3CFBBB" box="[580,775,1085,1109]" italics="true" pageId="4" pageNumber="307">C. leigongshanus</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="AA2A4B11FFCCFFC32307173BE9A6FB9B" author="Wei L" box="[296,413,1117,1141]" pageId="4" pageNumber="307" pagination="1 - 158" refId="ref15479" refString="Wei L (2018) A review of the genus Chrysotus Meigen (Diptera: Dolichopodidae) from China with definition of the leigongshanus species group. Oriental Insects, 52 (1): 1 - 158." type="journal article" year="2018">Wei, 2018</bibRefCitation>
). However, this species group has black femora, no pulvilli on the hind tarsus and sternite 8 with a pair of strong setae. In
<taxonomicName id="09BB4D63FFCCFFC323F817FBEA55FB5B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[471,622,1181,1205]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="sp. nov.">
<emphasis id="FCCFEAF2FFCCFFC323F817FBEA55FB5B" box="[471,622,1181,1205]" italics="true" pageId="4" pageNumber="307">Chrysotus dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC3205117F8EB38FB5B" box="[638,771,1181,1205]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, the femora are yellow, there are small pulvilli present on all legs, and sternite 8 does not bear strong setae.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7167BEA32FA3B" blockId="4.[135,777,1309,1493]" pageId="4" pageNumber="307">
Several other, but less common and yet to be described
<taxonomicName id="09BB4D63FFCCFFC322A7165BE97EFABB" box="[136,325,1341,1365]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7165BE8C8FABB" box="[136,243,1341,1365]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
species
</taxonomicName>
with yellow legs occur in back mangroves. They do not bear the typical black spot on the fore femora and have a brown triangular or trapezoid postpedicel with a short ventral protuberance. One of these species,
<taxonomicName id="09BB4D63FFCCFFC3205016F8EAC4FA5B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[639,767,1437,1461]" class="Insecta" family="Dolichopodidae" genus="Protomedetera" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="ubinensis" status="new species">
<emphasis id="FCCFEAF2FFCCFFC3205016F8EAC4FA5B" box="[639,767,1437,1461]" italics="true" pageId="4" pageNumber="307">C. ubinensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC322A716D8E936FA3B" box="[136,269,1469,1493]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, is described hereafter.
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A7169BE93DF91B" blockId="4.[135,777,1533,1781]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC322A7169BE9BBF9FB" bold="true" box="[136,384,1533,1557]" pageId="4" pageNumber="307">National distribution.</emphasis>
The new species is one of the most common
<taxonomicName id="09BB4D63FFCCFFC322C2157BE962F9DB" authorityName="Meigen" authorityYear="1824" box="[237,345,1565,1589]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFCCFFC322C2157BE962F9DB" box="[237,345,1565,1589]" italics="true" pageId="4" pageNumber="307">Chrysotus</emphasis>
</taxonomicName>
and dolichopodid species, in
<collectingCountry id="B6AC7670FFCCFFC320B4157BEB3CF9DB" box="[667,775,1565,1589]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
in the back mangroves of Pulau Ubin. It was found all over the island in all 26 back mangrove stations that were sampled with Malaise traps (
<figureCitation id="56802A65FFCCFFC3234F151BE99BF97B" box="[352,416,1661,1685]" captionStart="Fig" captionStartId="1.[113,148,1433,1455]" captionTargetBox="[113,1451,189,1417]" captionTargetId="figure-15@1.[113,1451,189,1417]" captionTargetPageId="1" captionText="Fig. 1. Distribution of the sampling stations with Malaise traps on Pulau Ubin. A, all sites from 2005 to 2019; B, detailed position of traps in the OBS area in 2019." figureDoi="http://doi.org/10.5281/zenodo.14683058" httpUri="https://zenodo.org/record/14683058/files/figure.png" pageId="4" pageNumber="307">Fig. 1</figureCitation>
). In total, 2,784 specimens were collected in the back mangroves and 2,357 specimens were barcoded, showing very little variation in haplotypes (less than 0.7%).
</paragraph>
<paragraph id="CE0436E0FFCCFFC322A71478ED82FF1B" blockId="4.[135,776,1821,2101]" lastBlockId="4.[834,1473,189,246]" pageId="4" pageNumber="307">
<materialsCitation id="7ED33CBDFFCCFFC322A71478ED8EFF1B" collectionCode="SMO" county="Mandai" location="It" municipality="Lim Chu Kang" pageId="4" pageNumber="307" specimenCode="SMN3" specimenCount="484">
In the front mangroves of the replanted mangroves of Pulau Semakau, only a
<collectingMethod id="17FA4EF7FFCCFFC323BD145BE96DF89B" pageId="4" pageNumber="307">single specimen was found in the three Malaise traps</collectingMethod>
that were operational for two full years (
<specimenCode id="9E1D9E9BFFCCFFC322BE141BE8E6F87B" box="[145,221,1917,1941]" collectionCode="SMN" country="China" lsid="urn:lsid:biocol.org:col:14058" name="Simao District National Medical and Pharmaceutical Institute" pageId="4" pageNumber="307" type="Herbarium">SMN3</specimenCode>
), while
<specimenCount id="D8BDFD69FFCCFFC32301141BE9F2F87B" box="[302,457,1917,1941]" count="239" pageId="4" pageNumber="307" type="generic">239 specimens</specimenCount>
were found in the adjacent old mangroves (
<collectionCode id="A8AAAE25FFCCFFC3232714FBE97DF85B" box="[264,326,1949,1973]" pageId="4" pageNumber="307">SMO</collectionCode>
), thus underlining that
<taxonomicName id="09BB4D63FFCCFFC3201A14F8EA4EF85B" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[565,629,1949,1973]" class="Insecta" family="Dolichopodidae" genus="Chrysotus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="307" phylum="Arthropoda" rank="species" species="dot" status="new species">
<emphasis id="FCCFEAF2FFCCFFC3201A14F8EA4EF85B" box="[565,629,1949,1973]" italics="true" pageId="4" pageNumber="307">C. dot</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFCCFFC320AD14F8EB38F85B" box="[642,771,1949,1973]" pageId="4" pageNumber="307" rank="species">new species</taxonomicNameLabel>
, is really a back mangrove species.
<location id="CB64603BFFCCFFC3202514D8EA21F83B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3202514D8EA21F83B" box="[522,538,1982,2005]" county="Mandai" municipality="Lim Chu Kang" name="It" pageId="4" pageNumber="307">It</location>
was further found in the back mangroves of
<location id="CB64603BFFCCFFC323BC14BBEA1DF81B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC323BC14BBEA1DF81B" box="[403,550,2013,2037]" county="Mandai" municipality="Lim Chu Kang" name="Coney Island" pageId="4" pageNumber="307">Coney Island</location>
(
<specimenCount id="D8BDFD69FFCCFFC3201614BBEAEBF81B" box="[569,720,2013,2037]" count="17" pageId="4" pageNumber="307" type="generic">17 specimens</specimenCount>
in 5 sampled stations),
<location id="CB64603BFFCCFFC32360149BE9CAF7FB" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC32360149BE9CAF7FB" box="[335,497,2045,2069]" county="Mandai" municipality="Lim Chu Kang" name="Berlayer Creek" pageId="4" pageNumber="307">Berlayer Creek</location>
(
<specimenCount id="D8BDFD69FFCCFFC3202D149BEA42F7FB" box="[514,633,2045,2069]" count="1" pageId="4" pageNumber="307" type="generic">1 specimen</specimenCount>
in 2 sampled stations), and
<location id="CB64603BFFCCFFC323371B7BEA64F7DB" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC323371B7BEA64F7DB" box="[280,607,2077,2101]" county="Mandai" municipality="Lim Chu Kang" name="Sungei Buloh Wetland Reserve" pageId="4" pageNumber="307">Sungei Buloh Wetland Reserve</location>
(
<specimenCount id="D8BDFD69FFCCFFC320441B7BEB3CF7DB" box="[619,775,2077,2101]" count="227" pageId="4" pageNumber="307" type="generic">227 specimens</specimenCount>
in 14 sampled stations).
<location id="CB64603BFFCCFFC3267A13DBECE0FF3B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3267A13DBECE0FF3B" box="[1109,1243,189,213]" county="Mandai" municipality="Lim Chu Kang" name="Remarkably" pageId="4" pageNumber="307">Remarkably</location>
, it was not found in the mangroves at
<location id="CB64603BFFCCFFC3262913BBEC4BFF1B" LSID="urn:lsid:plazi:treatment:461287F6FFCAFFC3231F151BECD3FD3B:CB64603BFFCCFFC3262913BBEC4BFF1B" box="[1030,1136,221,245]" county="Mandai" municipality="Lim Chu Kang" name="Sarimbun" pageId="4" pageNumber="307">Sarimbun</location>
,
<collectingMunicipality id="2E60AC9AFFCCFFC3265213BBED18FF18" box="[1149,1315,221,246]" pageId="4" pageNumber="307">Lim Chu Kang</collectingMunicipality>
, and
<collectingCounty id="27654E6CFFCCFFC3274F13BBED8EFF1B" box="[1376,1461,221,245]" pageId="4" pageNumber="307">Mandai</collectingCounty>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFCCFFC3216C127BECD3FD3B" pageId="4" pageNumber="307" type="distribution">
<paragraph id="CE0436E0FFCCFFC3216C127BEDFFFDFB" blockId="4.[834,1476,285,533]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C127BECA6FEDB" bold="true" box="[835,1181,285,309]" pageId="4" pageNumber="307">Distribution in Southeast Asia.</emphasis>
The new species was found in all three mangrove sites in
<collectingCountry id="B6AC7670FFCCFFC3265E125BEC82FEBB" box="[1137,1209,317,341]" name="Brunei" pageId="4" pageNumber="307">Brunei</collectingCountry>
we investigated (
<collectingRegion id="0C7FF802FFCCFFC32742125BED86FEBB" box="[1389,1469,317,341]" country="Brunei" name="Tutong" pageId="4" pageNumber="307">Tutong</collectingRegion>
, Berambang, and Labu). They differ by 2.3% in haplotype structure from the
<collectingCountry id="B6AC7670FFCCFFC3260E121BECA8FE7B" box="[1057,1171,381,405]" name="Singapore" pageId="4" pageNumber="307">Singapore</collectingCountry>
populations and a deeper morphological study is needed to show if they represent the same species. Only a single female was found in a mangrove site in southern
<collectingCountry id="B6AC7670FFCCFFC321DF12BBEC74FE1B" box="[1008,1103,477,501]" name="Thailand" pageId="4" pageNumber="307">Thailand</collectingCountry>
(
<collectingRegion id="0C7FF802FFCCFFC3267012BBECD9FE1B" box="[1119,1250,477,501]" country="Thailand" name="Surat Thani" pageId="4" pageNumber="307">Surat Thani</collectingRegion>
, leg. A. Samoh) and genetically it falls entirely in the population of Pulau Ubin.
</paragraph>
<paragraph id="CE0436E0FFCCFFC3216C115BECD3FD3B" blockId="4.[835,1476,573,725]" pageId="4" pageNumber="307">
<emphasis id="FCCFEAF2FFCCFFC3216C115BEBFAFDBB" bold="true" box="[835,961,573,597]" pageId="4" pageNumber="307">Phenology.</emphasis>
The new species is active throughout the year, but with its peak activity at the start of the rainy season in April and the lowest activity in the dryer periods of the year from end June to end September, with a second peak of activity from October to December.
</paragraph>
</subSubSection>
</treatment>
</document>