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<document id="395F3E52AD765CA180ADE98A24B78DEF" ID-DOI="10.26107/RBZ-2024-0025" ID-ISSN="2345-7600" ID-Zenodo-Dep="14683054" ID-ZooBank="079CC651-8D4D-4B95-B022-388B2FA51FE2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737171912946" checkinUser="felipe" docAuthor="Grootaert, Patrick, Velde, &amp; Isabella Van de" docDate="2024" docId="461287F6FFC4FFC9230E1083E97DFE1B" docLanguage="en" docName="RafflesBZool.72.303-323.pdf" docOrigin="Raffles Bulletin of Zoology 72" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Medetera stigma Grootaert, Velde &amp; de, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="316" masterDocId="BA2BFF8EFFC8FFC7222F1366E83BFFEE" masterDocTitle="New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)" masterLastPageNumber="323" masterPageNumber="303" pageNumber="315" updateTime="1737175760734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DE39DC881575B2978694F3FD8D993658">New long-legged flies from mangroves on Pulau Ubin (Singapore) (Insecta: Diptera: Dolichopodidae)</mods:title>
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<mods:namePart id="7B13122A3568942E26F097ADC3E63C43">Grootaert, Patrick</mods:namePart>
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<mods:namePart id="BF3E16D3F22748AF77B03ABD8C6AF5C9">Velde,</mods:namePart>
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<mods:namePart id="8D2461E3A55C0B07EE9B6E2D38598037">Isabella Van de</mods:namePart>
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<mods:date id="60BBEC73559D5378D652CA91692CA4A4">2024</mods:date>
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<mods:number id="4138E91349F9BA10E6C569279DD216E5">2024-09-19</mods:number>
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<treatment id="461287F6FFC4FFC9230E1083E97DFE1B" LSID="urn:lsid:plazi:treatment:461287F6FFC4FFC9230E1083E97DFE1B" httpUri="http://treatment.plazi.org/id/461287F6FFC4FFC9230E1083E97DFE1B" lastPageId="14" lastPageNumber="316" pageId="12" pageNumber="315">
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<paragraph id="CE0436E0FFC4FFCB230E1083EA54FC10" blockId="12.[289,623,997,1054]" box="[289,623,997,1022]" pageId="12" pageNumber="315">
<heading id="954C818CFFC4FFCB230E1083EA54FC10" bold="true" box="[289,623,997,1022]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="315" reason="2">
<emphasis id="FCCFEAF2FFC4FFCB230E1083EA54FC10" bold="true" box="[289,623,997,1022]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB230E1083E9DAFC10" bold="true" box="[289,481,997,1022]" italics="true" pageId="12" pageNumber="315">
<taxonomicName id="09BB4D63FFC4FFCB230E1083E9E0FC10" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[289,475,997,1022]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">Medetera stigma</taxonomicName>
,
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<taxonomicNameLabel id="E7FC5789FFC4FFCB23C61080EA54FC10" box="[489,623,998,1022]" pageId="12" pageNumber="315" rank="species">new species</taxonomicNameLabel>
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<paragraph id="CE0436E0FFC4FFCB23511760EA29FBF0" blockId="12.[289,623,997,1054]" box="[382,530,1030,1054]" pageId="12" pageNumber="315">
(
<figureCitation id="56802A65FFC4FFCB23AA1760E9DAFBF0" box="[389,481,1030,1054]" captionStart="Fig" captionStartId="12.[136,172,764,786]" captionTargetBox="[136,774,189,749]" captionTargetId="figure-8@12.[136,774,189,749]" captionTargetPageId="12" captionText="Fig. 10. Medetera stigma, new species, holotype male habitus, dorsal (ZRCBDP0317875). Photograph by Isabella Van de Velde." figureDoi="http://doi.org/10.5281/zenodo.14683080" httpUri="https://zenodo.org/record/14683080/files/figure.png" pageId="12" pageNumber="315">Figs. 10</figureCitation>
,
<figureCitation id="56802A65FFC4FFCB23C11760EA32FBF0" box="[494,521,1030,1054]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCB22A71720E8C6FA70" pageId="12" pageNumber="315" type="materials_examined">
<paragraph id="CE0436E0FFC4FFCB22A71720EA93FB30" blockId="12.[134,775,1094,1246]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71720E917FBB0" bold="true" box="[136,300,1094,1118]" pageId="12" pageNumber="315">Type material.</emphasis>
<materialsCitation id="7ED33CBDFFC4FFCB231E1720EA93FB30" collectingDate="2019-03-19" collectionCode="LKCNHM" latitude="1.4181389" longLatPrecision="1" longitude="103.93514" pageId="12" pageNumber="315" specimenCode="PU23, ZRCBDP0317875" specimenCount="1" typeStatus="holotype">
<typeStatus id="11008842FFC4FFCB231E1720E9A8FBB0" box="[305,403,1094,1118]" pageId="12" pageNumber="315" type="holotype">Holotype</typeStatus>
male. Pulau Ubin, Sungei [= River] Sempit,
<collectingMethod id="17FA4EF7FFC4FFCB22F31700E966FB93" box="[220,349,1126,1150]" pageId="12" pageNumber="315">Malaise trap</collectingMethod>
along creek in back mangrove,
<date id="BA051020FFC4FFCB208E1700E885FB70" pageId="12" pageNumber="315" value="2019-03-19">
<collectingDate id="AA41E9C8FFC4FFCB208E1700E885FB70" pageId="12" pageNumber="315" value="2019-03-19">19 March 2019</collectingDate>
</date>
,
<specimenCode id="9E1D9E9BFFC4FFCB22E417E0E931FB70" box="[203,266,1158,1182]" collectionCode="PU" country="USA" name="Princeton University" pageId="12" pageNumber="315">PU23</specimenCode>
,
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<geoCoordinate id="AB8F5027FFC4FFCB238A17E2EA7FFB73" box="[421,580,1156,1181]" degrees="103" direction="east" minutes="56" orientation="longitude" pageId="12" pageNumber="315" precision="1" seconds="06.5" value="103.93514">103°5606.5″E</geoCoordinate>
, (in 70% ethanol,
<specimenCode id="9E1D9E9BFFC4FFCB22A717C0E963FB50" box="[136,344,1190,1214]" collectionCode="ZRCBDP" pageId="12" pageNumber="315">ZRCBDP0317875</specimenCode>
_Jayanthi_JPP930_
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_
<date id="BA051020FFC4FFCB20AC17C0EB38FB50" box="[643,771,1190,1214]" pageId="12" pageNumber="315" value="2019-03-19">
<collectingDate id="AA41E9C8FFC4FFCB20AC17C0EB38FB50" box="[643,771,1190,1214]" pageId="12" pageNumber="315" value="2019-03-19">19Mar2019</collectingDate>
</date>
;
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). GenBank accession code: PP893304.
</materialsCitation>
</paragraph>
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<materialsCitation id="7ED33CBDFFC4FFCB22A71660E8C1FA70" collectingDate="2018-03-29" collectionCode="LKCNHM" latitude="1.4123555" longLatPrecision="1" longitude="103.96643" pageId="12" pageNumber="315" specimenCode="PU17, ZRCBDP0288845" specimenCount="1" typeStatus="paratype">
<emphasis id="FCCFEAF2FFC4FFCB22A71660E8C0FAF0" bold="true" box="[136,251,1286,1310]" pageId="12" pageNumber="315">
<typeStatus id="11008842FFC4FFCB22A71660E8CDFAF0" box="[136,246,1286,1310]" pageId="12" pageNumber="315" type="paratype">Paratype</typeStatus>
.
</emphasis>
Pulau Ubin, male, Sungei [= River] Maman, creek,
<date id="BA051020FFC4FFCB22E31640E957FAD0" box="[204,364,1318,1342]" pageId="12" pageNumber="315" value="2018-03-29">
<collectingDate id="AA41E9C8FFC4FFCB22E31640E957FAD0" box="[204,364,1318,1342]" pageId="12" pageNumber="315" value="2018-03-29">29 March 2018</collectingDate>
</date>
,
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,
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<geoCoordinate id="AB8F5027FFC4FFCB207F1642EAC0FAD3" box="[592,763,1316,1341]" degrees="103" direction="east" minutes="57" orientation="longitude" pageId="12" pageNumber="315" precision="1" seconds="59.17" value="103.96643">103°5759.17″E</geoCoordinate>
), front mangrove (70% ethanol)
<specimenCode id="9E1D9E9BFFC4FFCB20301620EB3BFAB0" box="[543,768,1350,1374]" collectionCode="ZRCBDP" pageId="12" pageNumber="315">ZRCBDP0288845</specimenCode>
,
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_
<date id="BA051020FFC4FFCB22FD1600E96AFA90" box="[210,337,1382,1406]" pageId="12" pageNumber="315" value="2018-03-29">
<collectingDate id="AA41E9C8FFC4FFCB22FD1600E96AFA90" box="[210,337,1382,1406]" pageId="12" pageNumber="315" value="2018-03-29">29Mar2018</collectingDate>
</date>
<collectionCode id="A8AAAE25FFC4FFCB23741600E9EEFA90" box="[347,469,1382,1406]" pageId="12" pageNumber="315">LKCNHM</collectionCode>
). GenBank accession code: PP893305
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCB22A716A0E9E5F9F3" pageId="12" pageNumber="315" type="etymology">
<paragraph id="CE0436E0FFC4FFCB22A716A0E9E5F9F3" blockId="12.[136,776,1478,1566]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A716A0E931FA30" bold="true" box="[136,266,1478,1502]" pageId="12" pageNumber="315">Etymology.</emphasis>
The new species is named
<taxonomicName id="09BB4D63FFC4FFCB201716A1EA45FA30" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[568,638,1479,1502]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma">
<emphasis id="FCCFEAF2FFC4FFCB201716A1EA45FA30" box="[568,638,1479,1502]" italics="true" pageId="12" pageNumber="315">stigma</emphasis>
</taxonomicName>
, referring to the presence of a brown spot at the base of the costa. The name
<taxonomicName id="09BB4D63FFC4FFCB22E51561E92AF9F0" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[202,273,1543,1566]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="315" phylum="Arthropoda" rank="species" species="stigma">
<emphasis id="FCCFEAF2FFC4FFCB22E51561E92AF9F0" box="[202,273,1543,1566]" italics="true" pageId="12" pageNumber="315">stigma</emphasis>
</taxonomicName>
is used as a noun.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC4FFCA22A71520E97FF89D" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315" type="diagnosis">
<paragraph id="CE0436E0FFC4FFCB22A71520EA2AF870" blockId="12.[135,777,1606,1950]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71520E8CDF9B0" bold="true" box="[136,246,1606,1630]" pageId="12" pageNumber="315">Diagnosis</emphasis>
. A small-sized species (about
<quantity id="09439B05FFC4FFCB20681520EA9BF9B0" box="[583,672,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="315" unit="mm" value="2.5">2.5 mm</quantity>
). Base of wings with a brownish streak at both sides of R
<subScript id="523F34A5FFC4FFCB20A01514EAACF96E" attach="left" box="[655,663,1650,1664]" fontSize="6" pageId="12" pageNumber="315">1</subScript>
. Face and palpus metallic green. Antenna yellow, postpedicel brownish on apex. Scutum without yellow colouration. Acrostichals biserial; 9 dorsocentrals, anterior 7 short, posterior 2 bristles long. Fore legs entirely black. Mid legs with trochanter and basal half of femur yellow, apical half femur, tibia and tarsus brown. Hind tibia with a black posterodorsal tooth beneath the preapical comb of bristles. Male terminalia with base of aedeagus not recurved, bent at a right angle. Tip hypandrium lacking denticles. Cercus undivided.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB22A714A0EA5CF833" blockId="12.[136,615,1990,2014]" box="[136,615,1990,2014]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A714A0E8F2F830" bold="true" box="[136,201,1990,2014]" pageId="12" pageNumber="315">Male.</emphasis>
Length: body
<quantity id="09439B05FFC4FFCB234414A0E9F9F833" box="[363,450,1990,2014]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="315" unit="mm" value="2.6">2.6 mm</quantity>
; wing
<quantity id="09439B05FFC4FFCB202314A0EA58F833" box="[524,611,1990,2014]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="12" pageNumber="315" unit="mm" value="2.1">2.1 mm</quantity>
.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB22A71B60ECCBFD1B" blockId="12.[135,775,2054,2110]" lastBlockId="12.[834,1474,477,757]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB22A71B60E8F7F7F0" bold="true" box="[136,204,2054,2078]" pageId="12" pageNumber="315">Head.</emphasis>
Face wide, with a pruinosity, but shiny metallic green with some bluish reflection at base. Frons very short (shorter
<emphasis id="FCCFEAF2FFC4FFCB216C12BBEBACFE1B" bold="true" box="[835,919,477,501]" pageId="12" pageNumber="315">Thorax</emphasis>
. Brownish black as base colour with a green metallic green shine, covered with a fine pruinosity. All bristles black. Acrostichals biserial, about 8 pairs; 9 dorsocentrals, anterior 7 bristles short, posterior 2 much longer, prescutellar bristle longest. Apical pair of scutellar bristles twice as long as bristles outside. A long white propleural bristle. Squama white with apical border brown, set with a row of pale brown bristles a little longer than squama is wide. Knob of haltere white, but with a brownish cap.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C107BECB8FC3B" blockId="12.[834,1475,797,981]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C107BEBBBFCDB" bold="true" box="[835,896,797,821]" pageId="12" pageNumber="315">Legs.</emphasis>
Fore legs entirely black. Mid legs with tip of coxa pale brown, trochanter, basal half of femur yellow; apical half femur, tibia and tarsus brown. Hind legs with coxa brown, trochanter yellow, basal half femur yellow, but apical half yellowish brown, basal half tibia pale brown, apical half brown, tarsus entirely brown.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C1098ED97FBBB" blockId="12.[834,1474,1021,1109]" pageId="12" pageNumber="315">Fore legs. Coxa with apical bristles yellow, shorter than coxa is wide; anterior bristles short, black. Femur with a posterior row of short brown bristles. No distinct bristles on tibia.</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C171BECB5FADB" blockId="12.[834,1474,1149,1333]" pageId="12" pageNumber="315">Mid legs. Coxa with whitish bristles; exterior bristle near apex coxa. Mid femur swollen on basal half with pale bristles, brown in apical half, half as long as femur is wide. Tibia with a short anterodorsal and posterodorsal bristle at basal quarter, as long as tibia is wide and a preapical ventral a little longer than tibia is wide.</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C163BEDA1F95B" blockId="12.[834,1476,1373,1717]" pageId="12" pageNumber="315">
Hind legs. Coxa near middle with a pale brown bristle, shorter than coxa is long. Femur somewhat dorsoventrally bent in basal third; with 5 long black dorsal bristles near base; 4 anterior bristles on apical third and 3 anteroventral bristles as long as femur is wide; basal half of femur with short ventral whitish bristles. Tibia with a black posterodorsal tooth-like projection composed of 4 closely set bristles, beneath the preapical comb of pale bristles; apical quarter more densely set with short ventral bristles than the basal ¾ of tibia. Basal tarsomere much shorter than second tarsomere. Length ratio tarsomeres
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mm: 0.16: 0.36: 0.22: 0.08: 0.95.
</paragraph>
<paragraph id="CE0436E0FFC4FFCB216C15BBEDF8F8DB" blockId="12.[835,1475,1757,1845]" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C15BBEB8CF91B" bold="true" box="[835,951,1757,1781]" pageId="12" pageNumber="315">Abdomen</emphasis>
. Brown in ground-colour with a faint green metallic shine. Tergites with all bristles brown, marginal bristles a little longer. Sternites with minute white bristles.
</paragraph>
<paragraph id="CE0436E0FFC4FFCA216C143BE8D0F8DD" blockId="12.[834,1475,1885,2101]" lastBlockId="13.[113,753,1723,1843]" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315">
<emphasis id="FCCFEAF2FFC4FFCB216C143BEC2EF89B" bold="true" box="[835,1045,1885,1909]" pageId="12" pageNumber="315">Male terminalia.</emphasis>
<figureCitation id="56802A65FFC4FFCB260E143BEC41F89B" box="[1057,1146,1885,1909]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11</figureCitation>
. Entirely brown. A pair of apicoventral epandrial bristles of equal length inserted on globular papilla. Basal epandrial bristle minute (
<figureCitation id="56802A65FFC4FFCB271714FBED8DF85B" box="[1336,1462,1949,1973]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11B, D</figureCitation>
). Tip hypandrium much widened before tip, lacking denticles (
<figureCitation id="56802A65FFC4FFCB216414BBEB8AF81B" box="[843,945,2013,2037]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11G</figureCitation>
). Base of aedeagus not recurved, bent at a right angle (
<figureCitation id="56802A65FFC4FFCB21A5149BEBD0F7FB" box="[906,1003,2045,2069]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="12" pageNumber="315">Fig. 11G</figureCitation>
). Dorsal and ventral surstylus well separated. Dorsal surstylus with an apical slit, bearing a lateral (
<figureCitation id="56802A65FFC4FFCA27B81B7BE886F93D" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" lastPageId="13" lastPageNumber="316" pageId="12" pageNumber="315">Fig. 11CE</figureCitation>
) and reflexed appendage at the outside. Postgonite pointed in lateral view (
<figureCitation id="56802A65FFC5FFCA235515BDE9DBF91D" box="[378,480,1755,1779]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11G</figureCitation>
). Cercus undivided (
<figureCitation id="56802A65FFC5FFCA20EA15BDE8F9F8FD" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11F, G</figureCitation>
), with a digitiform ventral appendage near middle (
<figureCitation id="56802A65FFC5FFCA2256147DE8E5F8DD" box="[121,222,1819,1843]" captionStart="Fig" captionStartId="13.[116,151,1588,1610]" captionTargetBox="[113,1457,190,1575]" captionTargetId="figure-16@13.[113,1475,167,1592]" captionTargetPageId="13" captionText="Fig. 11. Medetera stigma, new species, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert." figureDoi="http://doi.org/10.5281/zenodo.14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316">Fig. 11G</figureCitation>
).
</paragraph>
<caption id="9AC46668FFC5FFCA225B1552E991F96D" ID-DOI="http://doi.org/10.5281/zenodo.14683082" ID-Zenodo-Dep="14683082" httpUri="https://zenodo.org/record/14683082/files/figure.png" pageId="13" pageNumber="316" startId="13.[116,151,1588,1610]" targetBox="[113,1457,190,1575]" targetPageId="13" targetType="figure">
<paragraph id="CE0436E0FFC5FFCA225B1552E991F96D" blockId="13.[116,1454,1588,1668]" pageId="13" pageNumber="316">
Fig. 11.
<taxonomicName id="09BB4D63FFC5FFCA22E41552E94AF9A4" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[203,369,1588,1610]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">
<emphasis id="FCCFEAF2FFC5FFCA22E41552E94AF9A4" box="[203,369,1588,1610]" italics="true" pageId="13" pageNumber="316">Medetera stigma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC5FFCA23AE1553E9C1F9A4" box="[385,506,1588,1610]" pageId="13" pageNumber="316" rank="species">new species</taxonomicNameLabel>
, male terminalia. A, Ventral view; B, Hypandrium, epandrial bristles, ventral; C, Apex dorsal surstylus, right side; D, Ventral surstylus; E, Apex dorsal surstylus, left side; F, Cerci dorsal; G, Hypopygium lateral. Scale bar = 0.1 mm. Drawings by P. Grootaert.
</paragraph>
</caption>
<paragraph id="CE0436E0FFC5FFCA225E143DE97FF89D" blockId="13.[113,324,1883,1907]" box="[113,324,1883,1907]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA225E143DE8F0F89D" bold="true" box="[113,203,1883,1907]" pageId="13" pageNumber="316">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="86A1656BFFC5FFC9225E14FDE97DFE1B" lastPageId="14" lastPageNumber="317" pageId="13" pageNumber="316" type="discussion">
<paragraph id="CE0436E0FFC5FFCA225E14FDE8FDF7DD" blockId="13.[113,754,1947,2099]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA225E14FDE8C4F85D" bold="true" box="[113,255,1947,1971]" pageId="13" pageNumber="316">Comments.</emphasis>
<taxonomicName id="09BB4D63FFC5FFCA232314FDE9E2F85D" authorityName="Grootaert &amp; Velde &amp; Isabella Van de" authorityYear="2024" box="[268,473,1947,1971]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="stigma" status="sp. nov.">
<emphasis id="FCCFEAF2FFC5FFCA232314FDE9E2F85D" box="[268,473,1947,1971]" italics="true" pageId="13" pageNumber="316">Medetera stigma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E7FC5789FFC5FFCA23C214FAEABFF85D" box="[493,644,1947,1971]" pageId="13" pageNumber="316" rank="species">new species</taxonomicNameLabel>
, is quite characteristic in having at the base of the wings a brownish streak between the costa and R
<subScript id="523F34A5FFC5FFCA23911481E9FDF81B" attach="left" box="[446,454,2023,2037]" fontSize="6" pageId="13" pageNumber="316">1</subScript>
and below R
<subScript id="523F34A5FFC5FFCA207B1481EA67F81B" attach="left" box="[596,604,2023,2037]" fontSize="6" pageId="13" pageNumber="316">1</subScript>
. As far as we know, this characteristic is not observed in other
<taxonomicName id="09BB4D63FFC5FFCA20A3149DE8FDF7DD" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="FCCFEAF2FFC5FFCA20A3149DEAC9F7FD" box="[652,754,2043,2067]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="CE0436E0FFC5FFCA210215DDED97F8FD" blockId="13.[813,1452,1723,1812]" pageId="13" pageNumber="316">
<materialsCitation id="7ED33CBDFFC5FFCA210215DDED93F8FD" collectionCode="PU" pageId="13" pageNumber="316" specimenCode="PU23, PU17" specimenCount="2" typeStatus="holotype">
Only
<specimenCount id="D8BDFD69FFC5FFCA214415DAEC30F93D" box="[875,1035,1723,1747]" count="2" pageId="13" pageNumber="316" type="generic">two specimens</specimenCount>
are known: the
<typeStatus id="11008842FFC5FFCA26EE15DDED25F93D" box="[1217,1310,1723,1747]" pageId="13" pageNumber="316" type="holotype">holotype</typeStatus>
from a back mangrove near the mouth of Sungei Sempit (
<specimenCode id="9E1D9E9BFFC5FFCA273015BDED65F91D" box="[1311,1374,1755,1779]" collectionCode="PU" country="USA" name="Princeton University" pageId="13" pageNumber="316">PU23</specimenCode>
) and a
<typeStatus id="11008842FFC5FFCA2102159AEBBCF8FA" box="[813,903,1788,1812]" pageId="13" pageNumber="316" type="paratype">paratype</typeStatus>
from a front mangrove along Sungei Maman (
<specimenCode id="9E1D9E9BFFC5FFCA274F159DED9AF8FD" box="[1376,1441,1787,1811]" collectionCode="PU" country="USA" name="Princeton University" pageId="13" pageNumber="316">PU17</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CE0436E0FFC5FFCA2102145AEB8DF85D" blockId="13.[812,1451,1851,1971]" pageId="13" pageNumber="316">
None of the species handled by
<bibRefCitation id="AA2A4B11FFC5FFCA2652145DED2BF8BD" author="Bickel D" box="[1149,1296,1851,1875]" pageId="13" pageNumber="316" pagination="195 - 259" refId="ref14423" refString="Bickel D (1987) A revision of the Oriental and Australasian Medetera (Diptera: Dolichopodidae). Records of the Australian Museum, 39: 195 - 259." type="journal article" year="1987">Bickel (1987)</bibRefCitation>
in his revision of the
<collectingRegion id="0C7FF802FFC5FFCA215A143DEBF7F89D" box="[885,972,1883,1907]" country="Egypt" name="Ash Sharqiyah" pageId="13" pageNumber="316">Oriental</collectingRegion>
and Australasian
<taxonomicName id="09BB4D63FFC5FFCA26BC143DECC2F89D" authorityName="Fischer von Waldheim" authorityYear="1819" box="[1171,1273,1883,1907]" class="Insecta" family="Dolichopodidae" genus="Medetera" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="FCCFEAF2FFC5FFCA26BC143DECC2F89D" box="[1171,1273,1883,1907]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
</taxonomicName>
and by
<bibRefCitation id="AA2A4B11FFC5FFCA277A143AEBA1F87D" author="Yang D &amp; Zhang LL &amp; Wang MQ &amp; Zhu YJ" pageId="13" pageNumber="316" refId="ref15650" refString="Yang D, Zhang LL, Wang MQ &amp; Zhu YJ (2011) Fauna Sinica Insecta, Vol. 53. Diptera Dolichopodidae. Science Press, Beijing, 1912 pp. [in Chinese, with English summary]" type="book" year="2011">Yang et al. (2011)</bibRefCitation>
in the Fauna sinica, correspond to the present new species.
</paragraph>
<paragraph id="CE0436E0FFC5FFC9210214BDE97DFE1B" blockId="13.[813,1452,2011,2099]" lastBlockId="14.[136,776,189,501]" lastPageId="14" lastPageNumber="317" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA210214BDEB96F81D" bold="true" box="[813,941,2011,2035]" pageId="13" pageNumber="316">Bionomics.</emphasis>
Adults of most
<taxonomicName id="09BB4D63FFC5FFCA264514BDED17F81D" box="[1130,1324,2011,2035]" pageId="13" pageNumber="316">
<emphasis id="FCCFEAF2FFC5FFCA264514BDECE9F81D" box="[1130,1234,2011,2035]" italics="true" pageId="13" pageNumber="316">Medetera</emphasis>
species
</taxonomicName>
are known to dwell on tree trunks where they hunt for small insects. The larvae of most species are predacious and live inside the burrows of bark beetles where they feed on the larvae of these beetles. Having this
<typeStatus id="11008842FFC6FFC923FF13B8E9C5FF18" box="[464,510,222,246]" pageId="14" pageNumber="317">type</typeStatus>
of biology that on first sight is independent of the saline mangrove conditions, it is remarkable that a number of these
<taxonomicName id="09BB4D63FFC6FFC9200B127BEAE6FEDB" box="[548,733,285,309]" pageId="14" pageNumber="317">
<emphasis id="FCCFEAF2FFC6FFC9200B127BEAB1FEDB" box="[548,650,285,309]" italics="true" pageId="14" pageNumber="317">Medetera</emphasis>
species
</taxonomicName>
are exclusive for back mangroves and occur in the terrestrial forests behind the mangroves. At present, four of the eleven species actually known from
<collectingCountry id="B6AC7670FFC6FFC923FC121BEA7AFE7B" box="[467,577,381,405]" name="Singapore" pageId="14" pageNumber="317">Singapore</collectingCountry>
are exclusive for mangroves. Some species are dominant in mangroves and rarely occur in other biotopes, while some are exclusive for terrestrial forests.
</paragraph>
</subSubSection>
</treatment>
</document>