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<document id="FF0A990B8A3BCAC0888F3741BEC8AC6A" ID-CLB-Dataset="302074" ID-DOI="10.3897/phytokeys.246.129476" ID-GBIF-Dataset="b381880a-e0bb-4b02-ad0f-2f90a0b6c995" ID-publisher-id="129476" URI-arpha="7631F16E-4CD8-5A76-9F28-45E5F398CE20" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1725298343909" checkinUser="pensoft" docAuthor="Meerts, Pierre J. &amp; Paton, Alan J." docDate="2024" docId="0AFC2799833954CDA12EFA64A538A1D5" docLanguage="en" docName="PhytoKeys 246: 71-178" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/129476/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Coleus esculentus var. kolweziensis Meerts &amp; A. J. Paton A 2024, var. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="178" masterDocId="7631F16E4CD85A769F2845E5F398CE20" masterDocTitle="The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species" masterLastPageNumber="178" masterPageNumber="71" pageNumber="71" updateTime="1725322260938" updateUser="ExternalLinkService">
<mods:mods id="7B26305073C1620DE0BB582469618431" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="751F568BCFA682BA29C4378305C40ADB">
<mods:title id="1BC799F5F1CE468555267DDEFF64841C">The genus Coleus (Lamiaceae) in Central Africa (Democratic Republic of the Congo, Rwanda, Burundi), with the description of 15 new species</mods:title>
</mods:titleInfo>
<mods:name id="AC46FA6B334A82C3AF01BD340351BC39" type="personal">
<mods:role id="9650D61225D40C00BFAD0D0E76C9F3D4">
<mods:roleTerm id="77CB72588A231A630133785CF8351215">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="976D3F91D14EC5E6C65DA720A60358C8">Meerts, Pierre J.</mods:namePart>
<mods:nameIdentifier id="340AA6DEE9EAC4650323B3232313FD29" type="ORCID">0000-0003-4215-027X</mods:nameIdentifier>
<mods:affiliation id="3495CB28D9A0EF1000B90422D4A3D777">Meise Botanic Garden, Nieuwelaan 38, BE- 1860 Meise, Belgium &amp; Université Libre de Bruxelles, Av. F. D. Roosevelt 50 CP 244, BE- 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles, Service général de lEnseignement supérieur et de la Recherche scientifique, Brussels, Belgium</mods:affiliation>
</mods:name>
<mods:name id="832727532C5E97D06980653BA1F2D293" type="personal">
<mods:role id="A36F5D039266629C8393428F9206E42C">
<mods:roleTerm id="E36AD8871ED829D7E2C7235000491D50">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DB5B27C978877034554601B4C4F9D266">Paton, Alan J.</mods:namePart>
<mods:nameIdentifier id="D0390359F72105B7501817949A87F0D8" type="ORCID">0000-0002-6052-6675</mods:nameIdentifier>
<mods:affiliation id="F22A11F4170CD0CBAB5A139E3D6F57FC">The Herbarium, Royal Botanic Gardens, Kew, Richmond TW 9 3 AB, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5EBB6014E6821F8F0FE1705F402A7201">text</mods:typeOfResource>
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<mods:titleInfo id="8722FBA4E42DE7279B7AF1DDBD518B07">
<mods:title id="BDE115C8796BC4B3E2236E874C9A90F6">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="8B8E139F07D3DFB0F8A297D634D7FFD1">
<mods:date id="C8ADEC21223EDD3D9BE6A918C048EFB4">2024</mods:date>
<mods:detail id="2DA246220D8CE171BFD7748673F1576A" type="pubDate">
<mods:number id="C580A2374B702FD81923F285F4E8BCAC">2024-09-02</mods:number>
</mods:detail>
<mods:detail id="A3B3B282DC046A72B8A0398EA84E4FA6" type="volume">
<mods:number id="0C811D2EFB0DE8B2A65ACF2F0DF3D7E9">246</mods:number>
</mods:detail>
<mods:extent id="7E9DE7D025E1A3104CB117F321C0D003" unit="page">
<mods:start id="D25BB2E7075F5B296E30E912A476AC4D">71</mods:start>
<mods:end id="F13FBE347F77FB8F52F6E435085E90BA">178</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="7EE667A65C1EA5B2D4D152C6E2E0A8A8">journal article</mods:classification>
<mods:identifier id="2A98781681BC8797F382AB79303DEB0B" type="CLB-Dataset">302074</mods:identifier>
<mods:identifier id="46F51A0631675E33A4AACA186ACF5470" type="DOI">10.3897/phytokeys.246.129476</mods:identifier>
<mods:identifier id="B38AC823D37F7CABA56326C5739687AD" type="GBIF-Dataset">b381880a-e0bb-4b02-ad0f-2f90a0b6c995</mods:identifier>
</mods:mods>
<treatment id="0AFC2799833954CDA12EFA64A538A1D5" ID-DOI="http://doi.org/10.5281/zenodo.13629793" ID-GBIF-Taxon="240061482" ID-Zenodo-Dep="13629793" ID-arpha="0AFC2799-8339-54CD-A12E-FA64A538A1D5" ID-ipni="urn:lsid:ipni.org:names:77347692-1" LSID="urn:lsid:plazi:treatment:0AFC2799833954CDA12EFA64A538A1D5" LSID-IPNI="urn:lsid:ipni.org:names:77347692-1" httpUri="http://treatment.plazi.org/id/0AFC2799833954CDA12EFA64A538A1D5">
<subSubSection id="2A8025E0D38C348D78D3818141C2544E" type="nomenclature">
<paragraph id="FB8023D7D67A22FDAD0FE2E50481768B">
<taxonomicName id="F5DBD1015F5749AAF229E338FE0571E7" authority="Meerts &amp; A. J. Paton" authorityName="Meerts &amp; A. J. Paton A" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="variety" species="esculentus" status="var. nov." variety="kolweziensis">
<emphasis id="EC270C6CC9F7B5C0F3B633B798CF27CC" italics="true">Coleus esculentus var. kolweziensis</emphasis>
Meerts &amp; A. J. Paton
</taxonomicName>
<taxonomicNameLabel id="1B42DC6E2CB6FBFA344A48E22314B268" rank="variety">var. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="8EAA28E04BBB48E484510A9A4151CA57" type="description">
<paragraph id="9804DE43F3F76B191757A927FA13705D">
<figureCitation id="AB02B34A0049D757E3CB9D2F4E56A8D7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Coleus esculentus var. kolweziensis Meerts &amp; A. J. Paton A habit B detail of inflorescence and bracts C detail of calyx A M. Schaijes 3354 B, C Breyne 1197. Drawn by Hilde Orye. Scale bars: 1 cm (A); 5 mm (B); 2 mm (C)." figureDoi="10.3897/phytokeys.246.129476.figure4" httpUri="https://binary.pensoft.net/fig/1126711">Fig. 4 A C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EABGK" type="material">
<paragraph id="EF1BCEA551D3C0CE8F8C53881BFD018E">
<heading id="41D5482DCA7A5A890E74238860F60D67" reason="title">Type.</heading>
</paragraph>
<paragraph id="874D931301132CBE5B4B4D02D8A71CA1">
<materialsCitation id="3F6C1B5002A97EE6F24D3198DD6F255A" ID-GBIF-Occurrence="4927580310" collectingDate="1987-03-01" collectionCode="BR" collectorName="M. Schaijes" country="DR. Congo" elevation="1435" latitude="-10.508056" location="Piste Nzilo-Kyamasumba" longLatPrecision="21" longitude="25.436666" specimenCode="BR 0000021718298" specimenCount="1" stateProvince="Haut-Katanga" typeStatus="holotype">
<collectingCountry id="76D64B5943C0111A0FF02908CF56D8D0">DR. Congo</collectingCountry>
,
<collectingRegion id="C3851599635369F24C4F0224EA99DB5D">Haut-Katanga</collectingRegion>
,
<location id="D8CD3ACD15C7F0F98988A639EBA75709" LSID="urn:lsid:plazi:treatment:0AFC2799833954CDA12EFA64A538A1D5:D8CD3ACD15C7F0F98988A639EBA75709" country="DR. Congo" latitude="-10.508056" longLatPrecision="21" longitude="25.436666" name="Piste Nzilo-Kyamasumba" stateProvince="Haut-Katanga">Piste Nzilo-Kyamasumba</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E2BGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[25.436667,-10.508056]}">
<geoCoordinate id="DBF5006A6DFB7523280590DC1C8E2AEA" degrees="10" direction="south" minutes="30" orientation="latitude" precision="15" seconds="29" value="-10.508056">10 ° 30 ' 29 &quot; S</geoCoordinate>
,
<geoCoordinate id="3488DB0ADC5FD3F3648671A258266ACA" degrees="25" direction="east" minutes="26" orientation="longitude" precision="15" seconds="12" value="25.436666">25 ° 26 ' 12 &quot; E</geoCoordinate>
</named-content>
,
<quantity id="FE32C6CBD3B8DD026E444B0852020727" metricMagnitude="3" metricUnit="m" metricValue="1.435" unit="m" value="1435.0">
<elevation id="9DE89A2359FB2602B533BDA5C5B05B37" metricMagnitude="3" metricUnit="m" metricValue="1.435" unit="m" value="1435.0">1435 m</elevation>
</quantity>
elev.,
<date id="3841093947F514DBE92D4C0D512F2E68" value="1987-03-01">
<collectingDate id="01CC628FD59D59A99FE9A09B41F73D7F" value="1987-03-01">1 Mar 1987</collectingDate>
</date>
,
<emphasis id="F4D122BDAC38E77A553D8B6AE04BF3DA" italics="true">
<collectorName id="A94C740154FD5B52A902FC5287157033">M. Schaijes</collectorName>
3354
</emphasis>
(
<collectionCode id="8B0231D270CC17C78EF0A66CF8036B91" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
[
<specimenCode id="14B737BB51EAED83C583E69696FD8DC9">BR 0000021718298</specimenCode>
], photos)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCGK" type="diagnosis">
<paragraph id="211291A00E8A2BA566C86056E80EB604">
<heading id="4D3A77758667CA6B522879F3D833D28E" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="EFAC9D1A155DEEED1628C7B252FFB891">
Differs from the type in the following combination of traits: thyrses borne on pairs of ascending branches, at the 25 upper nodes of the stem, forming a panicle; bracts broadly ovate-elliptic, ca. 5 ×
<quantity id="510E6B3CB14471307743A17A173F55DE" metricMagnitude="-3" metricUnit="m" metricValue="3.0" unit="mm" value="3.0">3 mm</quantity>
, 5 - veined; rhizome creeping; stem leafy at flowering; leaves narrowly obovate, &lt;
<quantity id="F64551E578A55ED3F9FEEA42302401E0" metricMagnitude="-2" metricUnit="m" metricValue="1.5" unit="mm" value="15.0">15 mm</quantity>
wide, pubescent on veins beneath, very shortly pubescent above.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EHCGK" type="etymology">
<paragraph id="7856F71A9BD36BAAAB87EDA1E4FAA8BE">
<heading id="4EBF987B91043CE5BE964046F5DDE527" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="2B9F248BDBD99C2AFB6E545066CE0CFF">
All collections of this variety originate from the region of Kolwezi in western Upper
<collectingRegion id="0A9DD6A736E73F29D9CF78DC4EEAB178" country="Republic of the Congo" name="Katanga">Katanga</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETCGK" type="distribution">
<paragraph id="63C291EC0A2365D160F360FFDB5360A4">
<heading id="0591E444E95D733093EBBB8CA6500A5A" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="70542D132827153076A19777D9B5A40B">
Endemic to SE
<collectingCountry id="B0BC50CBBF2B8CD19F2CE6DE7B715A5E">DR. Congo</collectingCountry>
(
<collectingRegion id="572B7026935A27F300CA9370DC8243CB">Haut-Katanga</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0ELDGK" type="habitat">
<paragraph id="3458D0142DCFAC0CB8F951DCEA0D7781">
<heading id="862A863DF81DF025479C18E5AA06A8B2" reason="title">Habitat and ecology.</heading>
</paragraph>
<paragraph id="8795026E96C9F8251E58E82E7E033AA0">
Steppic savannah, dry woodland, rocky slopes;
<quantity id="036F8497BF1C42248BAECE9F65237B48" metricMagnitude="3" metricUnit="m" metricValue="1.3875" metricValueMax="1.475" metricValueMin="1.3" unit="m" value="1387.5" valueMax="1475.0" valueMin="1300.0">13001475 m</quantity>
elev.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQDGK" type="Additional specimens">
<paragraph id="8110BF9ED6BC12AB1A35031984EFE0DD">
<heading id="0144DE78C34638F0F6A1900B84939AEA" reason="title">Additional specimens.</heading>
</paragraph>
<paragraph id="D0D167AE11B573AC6C7DC40E02C1101F">
<collectingCountry id="F94F44D9C91CB4B400B0EBADE9D99D1E">DR. Congo</collectingCountry>
,
<emphasis id="82DD390F5F043076E13964FBE84DC355" bold="true" italics="true">
<collectingRegion id="361C219E258863E5A38D3CA5278B0789">Haut-Katanga</collectingRegion>
</emphasis>
, Env. de Sakabinda, sources de la Kengere,
<date id="FF9400827AB1F8BDD92C55FC8FDFD373" value="1971-04-09">9 Apr 1971</date>
,
<emphasis id="C0FBD8346EC06B41A27BE850B25591D2" italics="true">S. Lisowski</emphasis>
23304 (
<collectionCode id="EBF2517C0361620C2E9B959DDE2DCF82" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15257" lsid="urn:lsid:biocol.org:col:15257" name="Adam Mickiewicz University" type="Herbarium">POZG</collectionCode>
); Env. Kolwezi, près du village Muilu,
<date id="158EB4EE91CAF6150CE5D5D29FCAED0A" value="1971-04-08">8 Apr 1971</date>
,
<emphasis id="21BF864378ACC2130A8DE4AA6D419E5A" italics="true">S. Lisowski</emphasis>
23703 (
<collectionCode id="0EE7EC83EA1F93605E00FDE8FF3AC23E" country="Poland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15257" lsid="urn:lsid:biocol.org:col:15257" name="Adam Mickiewicz University" type="Herbarium">POZG</collectionCode>
); without locality, 1937,
<emphasis id="34A21D4D007461AB5D3BE91DDE8B432C" italics="true">Salésiens 1264</emphasis>
(
<collectionCode id="D72A296C3253E48E677C480E38C98C95" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
);
<quantity id="B5BA52153AD176F3782B5655B70EA901" metricMagnitude="4" metricUnit="m" metricValue="1.9" unit="km" value="19.0">19 km</quantity>
SSE of Kolwezi,
<date id="32CA38AD14E9FE01D2F08EDFFC22FF3F" value="1983-03-13">13 Mar 1983</date>
,
<emphasis id="48FDEFAB5327B70191E2AA1EAC779072" italics="true">
<collectionCode id="9DC96EB8FA74C3F586AE77384731C942" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Schaijes 1873
</emphasis>
(
<collectionCode id="D7C6579771C9973B17F2A1D2A1FFDAD2" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
); Piste Kolwezi-Musokantanda (Plateau de la Manika),
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKEGK" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[25.266389,-10.811667]}">
<geoCoordinate id="109DD97D305B299DCA0BBA86909BE1F7" degrees="10" direction="south" minutes="48" orientation="latitude" precision="15" seconds="42" value="-10.8116665">10 ° 48 ' 42 &quot; S</geoCoordinate>
,
<geoCoordinate id="FEAC642D9C537885486EC77C63983447" degrees="25" direction="east" minutes="15" orientation="longitude" precision="15" seconds="59" value="25.266388">25 ° 15 ' 59 &quot; E</geoCoordinate>
</named-content>
,
<date id="EC0E572F9C079D6367BBA042677481D4" value="1989-02-19">19 Feb 1989</date>
,
<emphasis id="64E3724753AA5D963E0CC3E17B9C21DF" italics="true">
<collectionCode id="45DA3CB835AEBF899F5D8D7D5D923495" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" type="Herbarium">M</collectionCode>
. Schaijes 4305
</emphasis>
(
<collectionCode id="39F20101E324B0D3F93058A1A8583EF4" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection">BR</collectionCode>
);
</paragraph>
<caption id="6321369F10AE160E7283F9B794987F78" ID-DOI="10.3897/phytokeys.246.129476.figure4" ID-arpha="49A290D4-FFFB-50B2-A44D-F33FCE7F2677" httpUri="https://binary.pensoft.net/fig/1126711" startId="F4">
<paragraph id="7856ABB16C245050DC40B8111C9E224A">
<label id="7D2DC3194BD6BC38A4AB13859CD0CC13">Figure 4.</label>
</paragraph>
<paragraph id="13541A117A0265BBF0F28CC01ECDC1B6">
<taxonomicName id="B9D1CD91C22822B85C14D1B862AAAF5C" authority="Meerts &amp; A. J. Paton A" authorityName="Meerts &amp; A. J. Paton A" authorityYear="2024" class="Magnoliopsida" family="Lamiaceae" genus="Coleus" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="variety" species="esculentus" variety="kolweziensis">
<emphasis id="A2FC1F168254815471773C9486188A78" italics="true">Coleus esculentus var. kolweziensis</emphasis>
Meerts &amp; A. J. Paton
<emphasis id="84333CF127C86919ECC473286C571222" bold="true">A</emphasis>
</taxonomicName>
habit
<emphasis id="B4BFC7EFDB6DF6DC4A9F471118F169E7" bold="true">B</emphasis>
detail of inflorescence and bracts
<emphasis id="121C0C6D4E450E2D8DE3065D4EDFC1F4" bold="true">C</emphasis>
detail of calyx
<emphasis id="B71ABD927AED8E7CD9226AE6CAF327E7" bold="true">A</emphasis>
<emphasis id="28C2123C943C5DD54F3704224D1033E0" italics="true">M. Schaijes 3354</emphasis>
<emphasis id="142C5BE2225D57038191972147A55282" bold="true">B, C</emphasis>
<emphasis id="F474D4D42616AAF5A91187B5D8329C21" italics="true">Breyne 1197</emphasis>
. Drawn by Hilde Orye. Scale bars: 1 cm (
<emphasis id="C8BB1939FBC9C38C6C637ED93609FACB" bold="true">A</emphasis>
); 5 mm (
<emphasis id="4AAD95C40B7C0E9237F4EA500CC26A3F" bold="true">B</emphasis>
); 2 mm (
<emphasis id="712FF831A3318602B3928299B257CEC0" bold="true">C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDGGK" type="notes">
<paragraph id="D6D8DAFFBAAA0F152701FF6C1444A2A6">
<heading id="E0ADFD223859E12B9BF83A374130B197" reason="title">Note.</heading>
</paragraph>
<paragraph id="B1EC3C8E3D3FF106EFD31D69C2E14AF2">This new variety has a restricted distribution range in the region of Kolwezi.</paragraph>
</subSubSection>
</treatment>
</document>