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<document id="EEEE7BDF28A9320C9D44CAF5D97A2E1B" ID-DOI="10.11646/zootaxa.5543.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386385" ID-ZooBank="8A21E43A-5605-4238-97B2-81CCA432A503" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1733918705237" checkinUser="plazi" docAuthor="Omelko, Mikhail M. &amp; Fomichev, Alexander A." docDate="2024" docId="B20419093767FF9C69A0FE3D01EC99E0" docLanguage="en" docName="zootaxa.5543.3.5.pdf" docOrigin="Zootaxa 5543 (3)" docSource="https://doi.org/10.11646/zootaxa.5543.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bowie catopuma Omelko &amp; Fomichev, 2024, sp. nov." docType="treatment" docVersion="1" lastPageNumber="388" masterDocId="4E3D61713763FF996937FF91026F9F4A" masterDocTitle="New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia" masterLastPageNumber="403" masterPageNumber="383" pageNumber="387" updateTime="1734117937186" updateUser="guilherme" zenodo-license-document="CLOSED">
<mods:mods id="24D672DB2B3D12C32891C8EDBAD91C6B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DF89019691BF7FE81393F345017A095B">
<mods:title id="0AA3FBAE55A88D845AC5B507FCDDEF85">New and poorly known species of Bowie Jäger, 2022 (Araneae: Ctenidae) from Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name id="04D709582D1387027F267318616FEE29" type="personal">
<mods:role id="9DB93A6B9487EA5C9C1654B5C2AFE512">
<mods:roleTerm id="7F4A70EA10C96E77A7B21894186905B0">Author</mods:roleTerm>
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<mods:namePart id="8C6083E3DFA2F471165D60D21C2E5267">Omelko, Mikhail M.</mods:namePart>
<mods:nameIdentifier id="04B2E26DA4AD39AACB61B0249A232586" type="ORCID">0000-0002-1556-6248</mods:nameIdentifier>
<mods:affiliation id="E360EDCD7D6141B7E89A75E65288CE99">Federal Scientific Center of East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok 690022, Russia.</mods:affiliation>
<mods:nameIdentifier id="D94659124DDC78797F9BD1FC939AB21E" type="email">omelkom@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="786F9BA606357E195ECD2CB445B41CD3" type="personal">
<mods:role id="A274EBB2165567A3355F5A02BDCF2909">
<mods:roleTerm id="0FA54445449600691116C2CE4CC2A9D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D139B29FA1B1418BC95EFA392BED0CD8">Fomichev, Alexander A.</mods:namePart>
<mods:nameIdentifier id="04F4852162B4C62B431138081A1B895E" type="ORCID">0000-0001-9268-622X</mods:nameIdentifier>
<mods:affiliation id="8CF6FDA500057C4898E46C93B5DCBF87">Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia. &amp; Tomsk State University, Lenina Pr., 36, Tomsk, RF- 634050, Russia. &amp; Western Caspian University, 31 Istiglaliyyat Street, Baku, Azerbaijan</mods:affiliation>
<mods:nameIdentifier id="82A3D85316C033301C7C05F0649C20F5" type="email">a.fomichov@mail.ru</mods:nameIdentifier>
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<mods:typeOfResource id="47D34F3D2FDAF20ED47B093D602E1882">text</mods:typeOfResource>
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<mods:title id="71E50C029DCB92472608DCE803D909A7">Zootaxa</mods:title>
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<mods:part id="9E2B3D39447F6EC21A7DE654AD02B36B">
<mods:date id="543F1F7DA8F4C19DADD5BBFD67307892">2024</mods:date>
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<mods:number id="F89BDA2E99763CC3FE911459A9613BE7">2024-12-04</mods:number>
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<mods:number id="FFB16BA93093553E10992A1EB197C83A">5543</mods:number>
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<mods:number id="28CB14D0E0D728EEDEDC40D540E46E0C">3</mods:number>
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<mods:identifier id="0E07BBA20BC068904663F363F1514E03" type="ISSN">1175-5326</mods:identifier>
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<treatment id="B20419093767FF9C69A0FE3D01EC99E0" LSID="urn:lsid:plazi:treatment:B20419093767FF9C69A0FE3D01EC99E0" httpUri="http://treatment.plazi.org/id/B20419093767FF9C69A0FE3D01EC99E0" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387">
<subSubSection id="72B7FB943767FF9D69A0FE3D03AA9E8C" box="[151,453,428,454]" pageId="4" pageNumber="387" type="nomenclature">
<paragraph id="3A12A81F3767FF9D69A0FE3D03AA9E8C" blockId="4.[151,453,428,490]" box="[151,453,428,454]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0FE3D03AA9E8C" bold="true" box="[151,453,428,454]" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D69A0FE3D030F9E8C" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[151,352,428,454]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D69A0FE3D030F9E8C" bold="true" box="[151,352,428,454]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763767FF9D685FFE3D03AA9E8C" box="[360,453,428,454]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69A0FE4103CA9EA0" box="[151,421,464,490]" pageId="4" pageNumber="387" type="description">
<paragraph id="3A12A81F3767FF9D69A0FE4103CA9EA0" blockId="4.[151,453,428,490]" box="[151,421,464,490]" pageId="4" pageNumber="387">
(
<figureCitation id="A296B49A3767FF9D69A8FE41036B9EA0" box="[159,260,464,490]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6827FE4103259EA0" box="[272,330,464,490]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">9–12</figureCitation>
,
<figureCitation id="A296B49A3767FF9D6861FE4103F29EA0" box="[342,413,464,490]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="4" pageNumber="387">68–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69A0FD8906F49D1C" pageId="4" pageNumber="387" type="materials_examined">
<paragraph id="3A12A81F3767FF9D69A0FD8906F49D1C" blockId="4.[151,1436,535,671]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0FD8903299D78" bold="true" box="[151,326,536,562]" pageId="4" pageNumber="387">Type material.</emphasis>
<materialsCitation id="8AC5A2423767FF9D687AFD8900049D1C" collectingDate="1994-08-30" collectionCode="ZMMU" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1020" latitude="4.4666667" location="Lucia Mt." longLatPrecision="1306" longitude="117.933334" municipality="Borneo" pageId="4" pageNumber="387" specimenCount="1" specimenCount-female="1" stateProvince="Sabah State" typeStatus="holotype">
<emphasis id="08D9740D3767FF9D687AFD8900939D78" bold="true" box="[333,764,535,562]" pageId="4" pageNumber="387">
<collectingCountry id="42BAE88F3767FF9D687AFD8903889D78" box="[333,487,536,562]" name="Malaysia" pageId="4" pageNumber="387">MALAYSIA</collectingCountry>
:
<emphasis id="08D9740D3767FF9D68C3FD8600119D78" bold="true" box="[500,638,535,562]" italics="true" pageId="4" pageNumber="387">
<collectingRegion id="F86966FD3767FF9D68C3FD8600119D78" box="[500,638,535,562]" country="Malaysia" name="Sabah" pageId="4" pageNumber="387">
<taxonomicName id="FDADD39C3767FF9D68C3FD8600529D78" authorityName="Li &amp; Yao" authorityYear="2022" box="[500,573,535,562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="sabah">Sabah</taxonomicName>
State
</collectingRegion>
</emphasis>
:
<typeStatus id="E51616BD3767FF9D6BB8FD8900939D78" box="[655,764,536,562]" pageId="4" pageNumber="387" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="2CAB63963767FF9D6A3CFD88014F9D79" box="[779,800,537,563]" count="1" pageId="4" pageNumber="387" type="female">♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A18FD8901E39D78" box="[815,908,536,562]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="387" type="Museum">ZMMU</collectionCode>
),
<collectingMunicipality id="DA7632653767FF9D6A97FD88019A9D79" box="[928,1013,537,563]" pageId="4" pageNumber="387">Borneo</collectingMunicipality>
,
<location id="3F72FEC43767FF9D6D36FD89061F9D79" LSID="urn:lsid:plazi:treatment:B20419093767FF9C69A0FE3D01EC99E0:3F72FEC43767FF9D6D36FD89061F9D79" box="[1025,1136,536,563]" country="Malaysia" latitude="4.4666667" longLatPrecision="1306" longitude="117.933334" municipality="Borneo" name="Lucia Mt." pageId="4" pageNumber="387" stateProvince="Sabah State">Lucia Mt.</location>
[
<geoCoordinate id="5F99CED83767FF9D6D48FD8906BA9D78" box="[1151,1237,536,562]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="4" pageNumber="387" precision="925" value="4.4666667">4°28’N</geoCoordinate>
,
<geoCoordinate id="5F99CED83767FF9D6DD5FD89073E9D79" box="[1250,1361,536,563]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="4" pageNumber="387" precision="925" value="117.933334">117°56’E</geoCoordinate>
],
<quantity id="FD5505FA3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">
<elevation id="B1804F2C3767FF9D6C53FD8902C09D1D" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="4" pageNumber="387" unit="m" value="1020.0">1020 m</elevation>
</quantity>
,
<date id="4E138EDF3767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">
<collectingDate id="5E5777373767FF9D698CFDAD03009D1C" box="[187,367,572,598]" pageId="4" pageNumber="387" value="1994-08-30">30 August 1994</collectingDate>
</date>
,
<collectorName id="9758CDC93767FF9D684EFDAD00549D1C" box="[377,571,572,598]" pageId="4" pageNumber="387">A.M. Emelyanov</collectorName>
leg.
</materialsCitation>
<materialsCitation id="8AC5A2423767FF9D6B45FDAC06F89D1C" box="[626,1175,572,599]" collectingDate="1994-08-30" collectionCode="IBSS" collectorName="A. M. Emelyanov" country="Malaysia" elevation="1020" latitude="4.4666667" location="Lucia Mt." longLatPrecision="1306" longitude="117.933334" municipality="Borneo" pageId="4" pageNumber="387" specimenCount="2" specimenCount-female="2" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="E51616BD3767FF9D6B45FDAC008C9D1D" box="[626,739,573,599]" pageId="4" pageNumber="387" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2CAB63963767FF9D6BDAFDAD017F9D1C" box="[749,784,572,598]" count="2" pageId="4" pageNumber="387" type="female">2♀</specimenCount>
(
<collectionCode id="5CBC30DA3767FF9D6A28FDAD01329D1C" box="[799,861,572,598]" collectionName="IBSS" pageId="4" pageNumber="387">IBSS</collectionCode>
) together with the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9D69F0FDF100059DD4" pageId="4" pageNumber="387" type="etymology">
<paragraph id="3A12A81F3767FF9D69F0FDF100059DD4" blockId="4.[151,1436,535,671]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0FDF103219D30" bold="true" box="[199,334,608,634]" pageId="4" pageNumber="387">Etymology.</emphasis>
The specific name is derived from the generic name of small wild cat from Southeast Asia,
<emphasis id="08D9740D3767FF9D69A0FD1503639DD4" box="[151,268,644,670]" italics="true" pageId="4" pageNumber="387">Catopuma</emphasis>
Severtzov; noun in apposition.
</paragraph>
</subSubSection>
<caption id="6ED2F8973767FF9D69A0F9310172984A" ID-DOI="http://doi.org/10.5281/zenodo.14386395" ID-Zenodo-Dep="14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387" startId="4.[151,264,1696,1720]" targetBox="[151,1435,694,1672]" targetPageId="4" targetType="figure">
<paragraph id="3A12A81F3767FF9D69A0F9310172984A" blockId="4.[151,1436,1696,1792]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69A0F931032999F2" bold="true" box="[151,326,1696,1720]" pageId="4" pageNumber="387">FIGURES 9–12.</emphasis>
Epigyne of
<taxonomicName id="FDADD39C3767FF9D688AF930000A99F2" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[445,613,1697,1720]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D688AF930000A99F2" box="[445,613,1697,1720]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6B5DF93100D699F2" bold="true" box="[618,697,1696,1720]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6B5DF93100D699F2" box="[618,697,1696,1720]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations:
<emphasis id="08D9740D3767FF9D6BFBF95400819996" box="[716,750,1733,1756]" italics="true" pageId="4" pageNumber="387">FD</emphasis>
—fertilization duct,
<emphasis id="08D9740D3767FF9D6A8BF95401B79996" box="[956,984,1733,1756]" italics="true" pageId="4" pageNumber="387">LT</emphasis>
—lateral tooth,
<emphasis id="08D9740D3767FF9D6D40F95506DC9991" box="[1143,1203,1732,1755]" italics="true" pageId="4" pageNumber="387">MC—</emphasis>
anterior constriction of median plate,
<emphasis id="08D9740D3767FF9D6811F9780324984A" box="[294,331,1769,1792]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
—median plate of epigyne,
<emphasis id="08D9740D3767FF9D6B52F979001199B5" box="[613,638,1768,1791]" italics="true" pageId="4" pageNumber="387">Sp</emphasis>
—spermatheca.
</paragraph>
</caption>
<subSubSection id="72B7FB943767FF9D69F0F8D6073198E3" pageId="4" pageNumber="387" type="diagnosis">
<paragraph id="3A12A81F3767FF9D69F0F8D6073198E3" blockId="4.[151,1437,1863,2034]" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F8D6032D982B" bold="true" box="[199,322,1863,1889]" pageId="4" pageNumber="387">Diagnosis.</emphasis>
By shape of the median plate (
<emphasis id="08D9740D3767FF9D6BA1F8D900D1982B" box="[662,702,1864,1889]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
) widest anteriorly females of
<taxonomicName id="FDADD39C3767FF9D6D3EF8D906AD982B" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1033,1218,1864,1889]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3767FF9D6D3EF8D906AD982B" box="[1033,1218,1864,1889]" italics="true" pageId="4" pageNumber="387">Bowie catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3767FF9D6DFEF8D9074E9828" bold="true" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387">
<taxonomicNameLabel id="13EAC9763767FF9D6DFEF8D9074E9828" box="[1225,1313,1864,1890]" pageId="4" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="FDADD39C3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="387" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3767FF9D6826F8FD03FB98CC" box="[273,404,1900,1926]" italics="true" pageId="4" pageNumber="387">B. neukoeln</emphasis>
</taxonomicName>
, but may be easily distinguished by 1)
<emphasis id="08D9740D3767FF9D6A74F8FD010498CF" box="[835,875,1900,1925]" italics="true" pageId="4" pageNumber="387">MP</emphasis>
with long anterior constriction (
<emphasis id="08D9740D3767FF9D6DE6F8FD069498CC" box="[1233,1275,1900,1926]" italics="true" pageId="4" pageNumber="387">MC</emphasis>
) (vs. short; cf.
<figureCitation id="A296B49A3767FF9D69A0F80102B698E0" box="[151,217,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 9</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6838F801033D98E0" box="[271,338,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 5</figureCitation>
); 2) vulval folds (
<emphasis id="08D9740D3767FF9D6B2BF801005198E3" box="[540,574,1936,1961]" italics="true" pageId="4" pageNumber="387">VF</emphasis>
) almost touching each other (vs. widely spaced; cf.
<figureCitation id="A296B49A3767FF9D6DB1F80106B998E0" box="[1158,1238,1936,1962]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="4" pageNumber="387">Fig. 10</figureCitation>
and
<figureCitation id="A296B49A3767FF9D6C3BF801072098E3" box="[1292,1359,1935,1962]" captionStart="FIGURES 5–8" captionStartId="3.[151,264,1935,1959]" captionTargetBox="[151,1435,829,1910]" captionTargetId="figure-182@3.[151,1435,829,1910]" captionTargetPageId="3" captionText="FIGURES 5–8. Epigyne of Bowie neukoeln Jäger. 5—intact epigyne, ventral; 6—ditto, macerated, ventral; 7—internal genitalia, dorsal; 8—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca, VF—vulval folds." figureDoi="http://doi.org/10.5281/zenodo.14386393" httpUri="https://zenodo.org/record/14386393/files/figure.png" pageId="4" pageNumber="387">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943767FF9C69F0F822073999CC" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387" type="description">
<paragraph id="3A12A81F3767FF9C69F0F82206FB9E0B" blockId="4.[151,1437,1863,2034]" lastBlockId="5.[151,1437,151,537]" lastPageId="5" lastPageNumber="388" pageId="4" pageNumber="387">
<emphasis id="08D9740D3767FF9D69F0F82203369887" bold="true" box="[199,345,1971,1997]" pageId="4" pageNumber="387">Description.</emphasis>
<emphasis id="08D9740D3767FF9D6856F82503D29884" box="[353,445,1972,1998]" italics="true" pageId="4" pageNumber="387">Female.</emphasis>
(
<figureCitation id="A296B49A3767FF9D68FAF825005A9884" box="[461,565,1972,1998]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1214,1238]" captionTargetBox="[183,1407,180,1182]" captionTargetId="figure-30@2.[177,1411,179,1188]" captionTargetPageId="2" captionText="FIGURES 1–4. Female habitus of Bowie neukoeln Jäger (1–2) and B. catopuma sp. nov. (3–4). 1, 3—dorsal; 2, 4—ventral. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.14386389" httpUri="https://zenodo.org/record/14386389/files/figure.png" pageId="4" pageNumber="387">Figs 3–4</figureCitation>
). Total length 11.76. Prosoma 5.79 long, 4.28 wide. Opisthosoma 5.37 long, 3.44 wide. Prosoma reddish brown, with brown median band. Median band widened and with projections near ocular area. Lateral bands absent. Fovea thin, black. Clypeus brown. Chelicerae dark brown, with 3 promarginal and 4 retromarginal teeth, denticles absent. Sternum light brown without pattern. Labium dark brown. Endites dark brown with yellowish distal edge. Dorsal part of opisthosoma light brown with yellowish cardiac mark and series of poorly visible gray spots. Lateral sides of opisthosoma light brown with blurry spots. Ventral part of opisthosoma light brown with yellow spots forming two poorly visible V-marks. Spinnerets light brown.
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FEDA07109EC3" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">Eye diameters: AME 0.22, ALE 0.18, PME 0.31, PLE 0.32; interdistances: AME–AME 0.19, AME–ALE 0.38, PME–PME 0.26, PME–PLE 0.42, AME–PME 0.11, ALE–PLE 0.17. Clypeus height at AME 0.14, at ALE 0.51.</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FE0207629D53" blockId="5.[151,1437,151,537]" pageId="5" pageNumber="388">
For palp and legs measurements see
<tableCitation id="772F9DA43766FF9C6B5AFE0200AC9EE7" box="[621,707,403,429]" captionStart="TABLE 3" captionStartId="5.[151,239,620,644]" captionTargetPageId="5" captionText="TABLE 3. Palp and legs measurements of female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 3</tableCitation>
. Palp coloration: femora, patellae, tibiae light brown; tarsi light brown with darker tip. Leg coloration: femora I–IV light brown with poorly visible spots. Patellae I–IV light brown. Tibiae I–II light brown; III–IV light brown with blurred brown semi-rings. Metatarsi I–II light brown; III–IV light brown with blurred brown semi-rings. Tarsi I–IV light brown. For palp and legs spination see
<tableCitation id="772F9DA43766FF9C6D84FE6E07669D53" box="[1203,1289,511,537]" captionStart="TABLE 4" captionStartId="5.[151,239,971,995]" captionTargetPageId="5" captionText="TABLE 4. Palp and legs spination for the female of Bowie catopuma sp. nov." pageId="5" pageNumber="388">Table 4</tableCitation>
.
</paragraph>
<caption id="6ED2F8973766FF9C69A0FDFD01B29DCF" box="[151,989,620,645]" pageId="5" pageNumber="388" startId="5.[151,239,620,644]" targetBox="[163,1370,661,886]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FDFD01B29DCF" blockId="5.[151,989,620,645]" box="[151,989,620,645]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FDFD03659DCE" bold="true" box="[151,266,620,644]" pageId="5" pageNumber="388">TABLE 3.</emphasis>
Palp and legs measurements of female of
<taxonomicName id="FDADD39C3766FF9C6BF9FDFC01EF9DCE" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[718,896,621,644]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF9FDFC01EF9DCE" box="[718,896,621,644]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB0FDFC01B29DCF" box="[903,989,621,645]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C684EFD0407359C3C" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FD0407359C3C" box="[163,1370,661,886]" gridcols="7" gridrows="6" pageId="5" pageNumber="388">
<tr id="849DAA5D376600666994FD0407359DE7" box="[163,1370,661,685]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666858FD0403F49DE7" box="[367,411,661,685]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666B19FD0400359DE7" box="[558,602,661,685]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666BDBFD0401779DE7" box="[748,792,661,685]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666A9CFD0401B89DE7" box="[939,983,661,685]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
<th id="C74CC321376600666D5EFD0406FA9DE7" box="[1129,1173,661,685]" gridcol="5" gridrow="0" pageId="5" pageNumber="388">Ta</th>
<th id="C74CC321376600666C16FD0407359DE7" box="[1313,1370,661,685]" gridcol="6" gridrow="0" pageId="5" pageNumber="388">Total</th>
</tr>
<tr id="849DAA5D376600666994FD2C07359D9F" box="[163,1370,701,725]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD2C02BF9D9F" box="[163,208,701,725]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666858FD2C03F49D9F" box="[367,411,701,725]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">2.00</td>
<td id="C74CC321376600666B19FD2C00359D9F" box="[558,602,701,725]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1.06</td>
<td id="C74CC321376600666BDBFD2C01779D9F" box="[748,792,701,725]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1.19</td>
<td id="C74CC321376600666A9CFD2C01B89D9F" box="[939,983,701,725]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
<td id="C74CC321376600666D5EFD2C06FA9D9F" box="[1129,1173,701,725]" gridcol="5" gridrow="1" pageId="5" pageNumber="388">1.35</td>
<td id="C74CC321376600666C16FD2C07359D9F" box="[1313,1370,701,725]" gridcol="6" gridrow="1" pageId="5" pageNumber="388">5.6</td>
</tr>
<tr id="849DAA5D376600666994FD7407359DB7" box="[163,1370,741,765]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FD7402BF9DB7" box="[163,208,741,765]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666858FD7403F49DB7" box="[367,411,741,765]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">4.08</td>
<td id="C74CC321376600666B19FD7400359DB7" box="[558,602,741,765]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">2.17</td>
<td id="C74CC321376600666BDBFD7401779DB7" box="[748,792,741,765]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">3.77</td>
<td id="C74CC321376600666A9CFD7401B89DB7" box="[939,983,741,765]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">3.10</td>
<td id="C74CC321376600666D5EFD7406FA9DB7" box="[1129,1173,741,765]" gridcol="5" gridrow="2" pageId="5" pageNumber="388">1.00</td>
<td id="C74CC321376600666C16FD7407359DB7" box="[1313,1370,741,765]" gridcol="6" gridrow="2" pageId="5" pageNumber="388">14.12</td>
</tr>
<tr id="849DAA5D376600666994FC9C07359C6C" box="[163,1370,781,806]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FC9C02BF9C6C" box="[163,208,781,806]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666858FC9C03F49C6C" box="[367,411,781,806]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3.94</td>
<td id="C74CC321376600666B19FC9C00359C6C" box="[558,602,781,806]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">2.14</td>
<td id="C74CC321376600666BDBFC9C01779C6C" box="[748,792,781,806]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">3.32</td>
<td id="C74CC321376600666A9CFC9C01B89C6C" box="[939,983,781,806]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2.92</td>
<td id="C74CC321376600666D5EFC9C06FA9C6C" box="[1129,1173,781,806]" gridcol="5" gridrow="3" pageId="5" pageNumber="388">1.05</td>
<td id="C74CC321376600666C16FC9C07359C6C" box="[1313,1370,781,806]" gridcol="6" gridrow="3" pageId="5" pageNumber="388">13.37</td>
</tr>
<tr id="849DAA5D376600666994FCA407359C04" box="[163,1370,821,846]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCA402BF9C04" box="[163,208,821,846]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666858FCA403F49C04" box="[367,411,821,846]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3.31</td>
<td id="C74CC321376600666B19FCA400359C04" box="[558,602,821,846]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1.71</td>
<td id="C74CC321376600666BDBFCA401779C04" box="[748,792,821,846]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">2.45</td>
<td id="C74CC321376600666A9CFCA401B89C04" box="[939,983,821,846]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">2.91</td>
<td id="C74CC321376600666D5EFCA406FA9C04" box="[1129,1173,821,846]" gridcol="5" gridrow="4" pageId="5" pageNumber="388">1.09</td>
<td id="C74CC321376600666C16FCA407359C04" box="[1313,1370,821,846]" gridcol="6" gridrow="4" pageId="5" pageNumber="388">11.47</td>
</tr>
<tr id="849DAA5D376600666994FCCF07359C3C" box="[163,1370,862,886]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FCCF02BF9C3C" box="[163,208,862,886]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666858FCCF03F49C3C" box="[367,411,862,886]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">4.61</td>
<td id="C74CC321376600666B19FCCF00359C3C" box="[558,602,862,886]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1.77</td>
<td id="C74CC321376600666BDBFCCF01779C3C" box="[748,792,862,886]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">4.00</td>
<td id="C74CC321376600666A9CFCCF01B89C3C" box="[939,983,862,886]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">5.34</td>
<td id="C74CC321376600666D5EFCCF06FA9C3C" box="[1129,1173,862,886]" gridcol="5" gridrow="5" pageId="5" pageNumber="388">1.52</td>
<td id="C74CC321376600666C16FCCF07359C3C" box="[1313,1370,862,886]" gridcol="6" gridrow="5" pageId="5" pageNumber="388">17.24</td>
</tr>
</table>
</paragraph>
<caption id="6ED2F8973766FF9C69A0FC5A01B99CA9" box="[151,982,971,995]" pageId="5" pageNumber="388" startId="5.[151,239,971,995]" targetBox="[163,1366,1011,1237]" targetIsTable="true" targetPageId="5" targetType="table">
<paragraph id="3A12A81F3766FF9C69A0FC5A01B99CA9" blockId="5.[151,982,971,995]" box="[151,982,971,995]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69A0FC5A03659CA9" bold="true" box="[151,266,971,995]" pageId="5" pageNumber="388">TABLE 4.</emphasis>
Palp and legs spination for the female of
<taxonomicName id="FDADD39C3766FF9C6BF0FC5D01159CA9" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[711,890,972,995]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6BF0FC5D01159CA9" box="[711,890,972,995]" italics="true" pageId="5" pageNumber="388">Bowie catopuma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="13EAC9763766FF9C6AB7FC5A01B99CA9" box="[896,982,971,995]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3A12A81F3766FF9C68B8FC6507399B9F" pageId="5" pageNumber="388">
<table id="48AD5ABF376600666994FC6207399B9F" box="[163,1366,1011,1237]" gridcols="5" gridrows="6" pageId="5" pageNumber="388">
<tr id="849DAA5D376600666994FC6207399B41" box="[163,1366,1011,1035]" gridrow="0" pageId="5" pageNumber="388" rowspan-0="1">
<th id="C74CC321376600666854FC6203BA9B41" box="[355,469,1011,1035]" gridcol="1" gridrow="0" pageId="5" pageNumber="388">Fe</th>
<th id="C74CC321376600666BA3FC6200809B41" box="[660,751,1011,1035]" gridcol="2" gridrow="0" pageId="5" pageNumber="388">Pa</th>
<th id="C74CC321376600666ABEFC62062A9B41" box="[905,1093,1011,1035]" gridcol="3" gridrow="0" pageId="5" pageNumber="388">Ti</th>
<th id="C74CC321376600666DF5FC6207399B41" box="[1218,1366,1011,1035]" gridcol="4" gridrow="0" pageId="5" pageNumber="388">Mt</th>
</tr>
<tr id="849DAA5D376600666994FB8D07399B7E" box="[163,1366,1052,1076]" gridrow="1" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB8D02BF9B7E" box="[163,208,1052,1076]" gridcol="0" gridrow="1" pageId="5" pageNumber="388">Palp</th>
<td id="C74CC321376600666854FB8D03BA9B7E" box="[355,469,1052,1076]" gridcol="1" gridrow="1" pageId="5" pageNumber="388">3d 1p 1r</td>
<td id="C74CC321376600666BA3FB8D00809B7E" box="[660,751,1052,1076]" gridcol="2" gridrow="1" pageId="5" pageNumber="388">1(0)p</td>
<td id="C74CC321376600666ABEFB8D062A9B7E" box="[905,1093,1052,1076]" gridcol="3" gridrow="1" pageId="5" pageNumber="388">1d 2p 1r</td>
<td id="C74CC321376600666DF5FB8D07399B7E" box="[1218,1366,1052,1076]" gridcol="4" gridrow="1" pageId="5" pageNumber="388">-</td>
</tr>
<tr id="849DAA5D376600666994FBD507399B16" box="[163,1366,1092,1116]" gridrow="2" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBD502BF9B16" box="[163,208,1092,1116]" gridcol="0" gridrow="2" pageId="5" pageNumber="388">I</th>
<td id="C74CC321376600666854FBD503BA9B16" box="[355,469,1092,1116]" gridcol="1" gridrow="2" pageId="5" pageNumber="388">3d 3p 3(2)r</td>
<td id="C74CC321376600666BA3FBD500809B16" box="[660,751,1092,1116]" gridcol="2" gridrow="2" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBD5062A9B16" box="[905,1093,1092,1116]" gridcol="3" gridrow="2" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBD507399B16" box="[1218,1366,1092,1116]" gridcol="4" gridrow="2" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FBFD07399BCE" box="[163,1366,1132,1156]" gridrow="3" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FBFD02BF9BCE" box="[163,208,1132,1156]" gridcol="0" gridrow="3" pageId="5" pageNumber="388">II</th>
<td id="C74CC321376600666854FBFD03BA9BCE" box="[355,469,1132,1156]" gridcol="1" gridrow="3" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FBFD00809BCE" box="[660,751,1132,1156]" gridcol="2" gridrow="3" pageId="5" pageNumber="388">spineless</td>
<td id="C74CC321376600666ABEFBFD062A9BCE" box="[905,1093,1132,1156]" gridcol="3" gridrow="3" pageId="5" pageNumber="388">2-2-2-2-2v</td>
<td id="C74CC321376600666DF5FBFD07399BCE" box="[1218,1366,1132,1156]" gridcol="4" gridrow="3" pageId="5" pageNumber="388">2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB0507399BE7" box="[163,1366,1172,1197]" gridrow="4" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB0502BF9BE7" box="[163,208,1172,1197]" gridcol="0" gridrow="4" pageId="5" pageNumber="388">III</th>
<td id="C74CC321376600666854FB0503BA9BE7" box="[355,469,1172,1197]" gridcol="1" gridrow="4" pageId="5" pageNumber="388">3d 4p 4r</td>
<td id="C74CC321376600666BA3FB0500809BE7" box="[660,751,1172,1197]" gridcol="2" gridrow="4" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB05062A9BE7" box="[905,1093,1172,1197]" gridcol="3" gridrow="4" pageId="5" pageNumber="388">3d 2p 2r 2-2v</td>
<td id="C74CC321376600666DF5FB0507399BE7" box="[1218,1366,1172,1197]" gridcol="4" gridrow="4" pageId="5" pageNumber="388">4p 5r 2-2-2v</td>
</tr>
<tr id="849DAA5D376600666994FB2C07399B9F" box="[163,1366,1213,1237]" gridrow="5" pageId="5" pageNumber="388">
<th id="C74CC321376600666994FB2C02BF9B9F" box="[163,208,1213,1237]" gridcol="0" gridrow="5" pageId="5" pageNumber="388">IV</th>
<td id="C74CC321376600666854FB2C03BA9B9F" box="[355,469,1213,1237]" gridcol="1" gridrow="5" pageId="5" pageNumber="388">3d 4p 2r</td>
<td id="C74CC321376600666BA3FB2C00809B9F" box="[660,751,1213,1237]" gridcol="2" gridrow="5" pageId="5" pageNumber="388">1p 1r</td>
<td id="C74CC321376600666ABEFB2C062A9B9F" box="[905,1093,1213,1237]" gridcol="3" gridrow="5" pageId="5" pageNumber="388">3(2)d 2p 2r 2-2-2v</td>
<td id="C74CC321376600666DF5FB2C07399B9F" box="[1218,1366,1213,1237]" gridcol="4" gridrow="5" pageId="5" pageNumber="388">4p 4r 2-1-2-2v</td>
</tr>
</table>
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FAB903E59AE4" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
Female genitalia as shown in
<figureCitation id="A296B49A3766FF9C6B2BFAB900FE9A08" box="[540,657,1320,1346]" captionStart="FIGURES 9–12" captionStartId="4.[151,264,1696,1720]" captionTargetBox="[151,1435,694,1672]" captionTargetId="figure-189@4.[151,1435,694,1672]" captionTargetPageId="4" captionText="FIGURES 9–12. Epigyne of Bowie catopuma sp. nov. 9—intact, ventral; 10—macerated, ventral; 11—internal genitalia, dorsal; 12—ditto, posterior. Scale bar: 0.2 mm. Abbreviations: FD—fertilization duct, LT—lateral tooth, MC—anterior constriction of median plate, MP—median plate of epigyne, Sp—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.14386395" httpUri="https://zenodo.org/record/14386395/files/figure.png" pageId="5" pageNumber="388">Figs 9–12</figureCitation>
. Epigyne 1.5 times wider than long. Median plate (
<emphasis id="08D9740D3766FF9C6DE9FAB807699A08" box="[1246,1286,1321,1346]" italics="true" pageId="5" pageNumber="388">MP</emphasis>
) trapezoidal, strongly sclerotized anteriorly. Lateral teeth (
<emphasis id="08D9740D3766FF9C6B87FADC00A09A2C" box="[688,719,1357,1382]" italics="true" pageId="5" pageNumber="388">LT</emphasis>
) well developed, located posterior of median plate, slightly extending beyond epigastric furrow, with rounded tips. Spermathecae (
<emphasis id="08D9740D3766FF9C6A8BFAE001B79AC1" box="[956,984,1393,1419]" italics="true" pageId="5" pageNumber="388">Sp</emphasis>
) large, piriform, separated by a third of spermatheca’s length.
</paragraph>
<paragraph id="3A12A81F3766FF9C69F0FA29073999CC" blockId="5.[151,1436,1320,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0FA29037E9A98" bold="true" box="[199,273,1464,1490]" pageId="5" pageNumber="388">Notes.</emphasis>
Since males
<taxonomicName id="FDADD39C3766FF9C689DFA2800579A98" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[426,568,1465,1490]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C689DFA2800579A98" box="[426,568,1465,1490]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6B76FA2900F39A98" bold="true" box="[577,668,1464,1490]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6B76FA2900F39A98" box="[577,668,1464,1490]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are unknown, it is difficult to place it in a species group. There are species with a similar shape of median plate and lateral teeth situated at the posterior margin of the epigyne in
<emphasis id="08D9740D3766FF9C69A0F99102919950" box="[151,254,1536,1562]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
and
<taxonomicName id="FDADD39C3766FF9C680EF99003B49950" authorityName="Jager" authorityYear="2022" box="[313,475,1537,1562]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C680EF99003B49950" box="[313,475,1537,1562]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
species groups. Because the new species have spermathecae consisting of only one compartment, while those in
<emphasis id="08D9740D3766FF9C68E8F9B500299974" box="[479,582,1572,1598]" italics="true" pageId="5" pageNumber="388">chinagirl</emphasis>
-group have particularly structured spermathecae, we place
<taxonomicName id="FDADD39C3766FF9C6DD2F9B4071C9974" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[1253,1395,1573,1598]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C6DD2F9B4071C9974" box="[1253,1395,1573,1598]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6C4CF9B502A79928" bold="true" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6C4CF9B502A79928" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the
<taxonomicName id="FDADD39C3766FF9C6820F9D803D69928" authorityName="Jager" authorityYear="2022" box="[279,441,1609,1634]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">
<emphasis id="08D9740D3766FF9C6820F9D803D69928" box="[279,441,1609,1634]" italics="true" pageId="5" pageNumber="388">scarymonsters</emphasis>
</taxonomicName>
-species group. Thus, with the description of the new species, this group now consists of five species:
<taxonomicName id="FDADD39C3766FF9C681EF9FC038C99CC" authorityName="Omelko &amp; Fomichev" authorityYear="2023" box="[297,483,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="binturong">
<emphasis id="08D9740D3766FF9C681EF9FC038C99CC" box="[297,483,1644,1670]" italics="true" pageId="5" pageNumber="388">Bowie binturong</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C68C5F9FC001099CC" authorityName="Omelko &amp; Fomichev" authorityYear="2024" box="[498,639,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="catopuma" status="sp. nov.">
<emphasis id="08D9740D3766FF9C68C5F9FC001099CC" box="[498,639,1645,1670]" italics="true" pageId="5" pageNumber="388">B. catopuma</emphasis>
</taxonomicName>
<emphasis id="08D9740D3766FF9C6BB1F9FD008F99CC" bold="true" box="[646,736,1644,1670]" pageId="5" pageNumber="388">
<taxonomicNameLabel id="13EAC9763766FF9C6BB1F9FD008F99CC" box="[646,736,1644,1670]" pageId="5" pageNumber="388" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="FDADD39C3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="neukoeln">
<emphasis id="08D9740D3766FF9C6BD9F9FC011D99CC" box="[750,882,1644,1670]" italics="true" pageId="5" pageNumber="388">B. neukoeln</emphasis>
</taxonomicName>
,
<taxonomicName id="FDADD39C3766FF9C6AB7F9FC063B99CD" authority="Li &amp; Yao" authorityName="Li &amp; Yao" authorityYear="2022" box="[896,1108,1644,1671]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="sabah">
<emphasis id="08D9740D3766FF9C6AB7F9FC018D99CC" box="[896,994,1644,1670]" italics="true" pageId="5" pageNumber="388">B. sabah</emphasis>
Li &amp; Yao
</taxonomicName>
, and
<emphasis id="08D9740D3766FF9C6DB9F9FC073999CC" box="[1166,1366,1645,1670]" italics="true" pageId="5" pageNumber="388">
<taxonomicName id="FDADD39C3766FF9C6DB9F9FC073D99CC" authorityName="Jager" authorityYear="2022" box="[1166,1362,1645,1670]" class="Arachnida" family="Ctenidae" genus="Bowie" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="388" phylum="Arthropoda" rank="species" species="scarymonsters">B. scarymonsters</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="72B7FB943766FF9C69F0F90101EC99E0" box="[199,899,1680,1707]" pageId="5" pageNumber="388" type="distribution">
<paragraph id="3A12A81F3766FF9C69F0F90101EC99E0" blockId="5.[151,1436,1320,1707]" box="[199,899,1680,1707]" pageId="5" pageNumber="388">
<emphasis id="08D9740D3766FF9C69F0F901030E99E0" bold="true" box="[199,353,1680,1706]" pageId="5" pageNumber="388">Distribution.</emphasis>
<typeStatus id="E51616BD3766FF9C6850F90003F099E1" box="[359,415,1681,1707]" pageId="5" pageNumber="388">Type</typeStatus>
locality only, Borneo:
<collectingRegion id="F86966FD3766FF9C6B97F901008B99E0" box="[672,740,1680,1706]" country="Malaysia" name="Sabah" pageId="5" pageNumber="388">Sabah</collectingRegion>
(
<figureCitation id="A296B49A3766FF9C6BC4F901011899E0" box="[755,887,1680,1706]" captionStart="FIGURES 68–69" captionStartId="19.[151,264,533,557]" captionTargetBox="[151,1435,181,504]" captionTargetId="figure-19@19.[151,1435,181,504]" captionTargetPageId="19" captionText="FIGURES 68–69. Collection localities of the studied Bowie species. Circle—Bowie catopuma sp. nov.; star—Bowie musang sp. nov.; diamond—Bowie neukoeln Jäger; triangle—Bowie tangalunga sp. nov.; inverted triangle—Bowie shakeit Jäger; square—Bowie hosei (F.O. Pickard-Cambridge). The frame on Fig. 69 refers to the content of Fig. 68." figureDoi="http://doi.org/10.5281/zenodo.14386431" httpUri="https://zenodo.org/record/14386431/files/figure.png" pageId="5" pageNumber="388">Figs 68–69</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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