225 lines
15 KiB
XML
225 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.46.32823" ID-GBIF-Dataset="e546c2a6-4668-4253-b2e2-b462f23dab8c" ID-PMC="PMC6379322" ID-Pensoft-Pub="1314-4049-46-1" ID-PubMed="30787668" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-4049-46-1" ModsDocOrigin="MycoKeys 46" ModsDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" checkinTime="1555333436509" checkinUser="pensoft" docAuthor="Cripps, Cathy L., Eberhardt, Ursula, Schuetz, Nicole, Beker, Henry J., Vera S. Evenson, & Horak, Egon" docDate="2019" docId="D31F8AFF540C7A1114F3A457E7DA1FFC" docLanguage="en" docName="MycoKeys 46: 1-54" docOrigin="MycoKeys 46" docSource="http://dx.doi.org/10.3897/mycokeys.46.32823" docTitle="Hebeloma spetsbergense Beker & U. Eberh., Fungi Europaei 14 (Lomazzo): 180 2016" docType="treatment" docVersion="4" lastPageNumber="41" masterDocId="AC18AA047D78FFBFD06BFFFDFF948577" masterDocTitle="The genus Hebeloma in the Rocky Mountain Alpine Zone" masterLastPageNumber="54" masterPageNumber="1" pageNumber="39" updateTime="1668136151891" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Hebeloma in the Rocky Mountain Alpine Zone</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cripps, Cathy L.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Eberhardt, Ursula</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schuetz, Nicole</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beker, Henry J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vera S. Evenson,</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Horak, Egon</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>MycoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>46</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/mycokeys.46.32823</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.46.32823</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-4049-46-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="156202101" LSID="urn:lsid:plazi:treatment:D31F8AFF540C7A1114F3A457E7DA1FFC" httpUri="http://treatment.plazi.org/id/D31F8AFF540C7A1114F3A457E7DA1FFC" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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<subSubSection pageId="38" pageNumber="39" type="nomenclature">
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<paragraph pageId="38" pageNumber="39">
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16.
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<taxonomicName authority="Beker & U. Eberh., Fungi Europaei 14 (Lomazzo): 180 (2016)" class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma spetsbergense" order="Agaricales" pageId="38" pageNumber="39" phylum="Basidiomycota" rank="species" species="spetsbergense">Hebeloma spetsbergense Beker & U. Eberh., Fungi Europaei 14 (Lomazzo): 180 (2016)</taxonomicName>
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Figures 6A, 22
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</paragraph>
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</subSubSection>
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<subSubSection pageId="39" pageNumber="40" type="etymology">
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<paragraph pageId="39" pageNumber="40">
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<pageBreakToken pageId="39" pageNumber="40" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="39" pageNumber="40">Originally found in Svalbard.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
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<paragraph pageId="39" pageNumber="40">Description.</paragraph>
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<paragraph pageId="39" pageNumber="40">
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Cortina present. Pileus 10-25 mm in diameter, shallow convex, almost applanate with indistinct umbo or not, smooth, tacky to dry, brown in center, outwards paler brown or more cinnamon, with white edge, not hygrophanous; margin turned down in young specimens, entire. Lamellae attached, adnexed, medium close, L = 26-30, pale cream to milk coffee, to brown; edges indistinct fimbriate. Stipe long and thin, 20-40
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<normalizedToken originalValue="×">x</normalizedToken>
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2-3 mm, equal, cream at apex to dark brown at base, fibrils at apex, and below silky-smooth with longitundinal fibrils. Context cream and brown to black in lower part. Odor raphanoid. Exsiccata: pileus brown, darker brown in center; lamellae reddish brown; stipe thin, cream but darkening at base.
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</paragraph>
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<paragraph lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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Basidiospores yellow brown, amygdaliform, without a large snout, apiculate, not guttulate, finely verrucose (O1, O2), distinctly and sometimes strongly dextri
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<pageBreakToken pageId="40" pageNumber="41" start="start">noid</pageBreakToken>
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(D2, D3), no loosening perispore observed (P0), 11-14
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<normalizedToken originalValue="×">x</normalizedToken>
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7-8.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, on average 12.5
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<normalizedToken originalValue="×">x</normalizedToken>
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7.6
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, Q = 1.65. Basidia 28-35
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<normalizedToken originalValue="×">x</normalizedToken>
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8-10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, clavate, mostly four-spored. Cheilocystidia lageniform, with long cylindrical neck, 30-80
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<normalizedToken originalValue="×">x</normalizedToken>
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4-7
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at apex, 4-5.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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in middle, and 7-10.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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at base. Pleurocystidia absent. Epicutis thickness 30-35
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, with no encrusted hyphae recorded.
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</paragraph>
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<caption pageId="40" pageNumber="41">
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<paragraph pageId="40" pageNumber="41">
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Figure 22.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma spetsbergense" order="Agaricales" pageId="40" pageNumber="41" phylum="Basidiomycota" rank="species" species="spetsbergense">Hebeloma spetsbergense</taxonomicName>
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, DBG-F-027678 and BR5020184126599 (HJB 11982, from Svalbard). Scale bar for basidia and cheilocystidia 5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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, for spores 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. Drawing G. Walther, reproduced from
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<bibRefCitation pageId="40" pageNumber="41">Beker et al. (2016)</bibRefCitation>
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.
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</paragraph>
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</caption>
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<caption pageId="40" pageNumber="41">
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<paragraph pageId="40" pageNumber="41">
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Figure 23. Micro-morphological features (basidiospores, basidia, cheilocystidia) of
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma" order="Agaricales" pageId="40" pageNumber="41" phylum="Basidiomycota" rank="genus">Hebeloma</taxonomicName>
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species found in the Rocky Mountain alpine zone. 1
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<taxonomicName lsidName="H. vaccinum" pageId="40" pageNumber="41" rank="species" species="vaccinum">H. vaccinum</taxonomicName>
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(holotype, Herb. PC) 2
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<taxonomicName lsidName="H. aurantioumbrinum" pageId="40" pageNumber="41" rank="species" species="aurantioumbrinum">H. aurantioumbrinum</taxonomicName>
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ZT12730 3
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<taxonomicName lsidName="H. subconcolor" pageId="40" pageNumber="41" rank="species" species="subconcolor">H. subconcolor</taxonomicName>
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ZT 13776 4
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<taxonomicName lsidName="H. hiemale" pageId="40" pageNumber="41" rank="species" species="hiemale">H. hiemale</taxonomicName>
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ZT9828 5
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<taxonomicName lsidName="H. avellaneum" pageId="40" pageNumber="41" rank="species" species="avellaneum">H. avellaneum</taxonomicName>
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DBG-F-019533 6
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<taxonomicName lsidName="H. velutipes" pageId="40" pageNumber="41" rank="species" species="velutipes">H. velutipes</taxonomicName>
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ZT6100 7
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<taxonomicName lsidName="H. alpinum" pageId="40" pageNumber="41" rank="species" species="alpinum">H. alpinum</taxonomicName>
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CLC2875 8
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<taxonomicName lsidName="H. marginatulum" pageId="40" pageNumber="41" rank="species" species="marginatulum">H. marginatulum</taxonomicName>
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ZT9002 9
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<taxonomicName lsidName="H. alpinicola" pageId="40" pageNumber="41" rank="species" species="alpinicola">H. alpinicola</taxonomicName>
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ZT13763 10
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<taxonomicName lsidName="H. dunense" pageId="40" pageNumber="41" rank="species" species="dunense">H. dunense</taxonomicName>
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ZT9001 11
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<taxonomicName lsidName="H. mesophaeum" pageId="40" pageNumber="41" rank="species" species="mesophaeum">H. mesophaeum</taxonomicName>
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ZT8082 12
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<taxonomicName lsidName="H. excedens" pageId="40" pageNumber="41" rank="species" species="excedens">H. excedens</taxonomicName>
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ZT7475 13
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<taxonomicName lsidName="H. oreophilum" pageId="40" pageNumber="41" rank="species" species="oreophilum">H. oreophilum</taxonomicName>
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ZT12733 14
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<taxonomicName lsidName="H. hygrophilum" pageId="40" pageNumber="41" rank="species" species="hygrophilum">H. hygrophilum</taxonomicName>
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CLC1462 15
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<taxonomicName lsidName="H. nigellum" pageId="40" pageNumber="41" rank="species" species="nigellum">H. nigellum</taxonomicName>
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ZT6425 16
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<taxonomicName lsidName="H. spetsbergense" pageId="40" pageNumber="41" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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micro in Fig. 22. Both two and four-spored basidiospores shown for 2, 5, 7, 8, 10, 12, 13, 15. Scale bar: 10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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. All drawings by E. Horak.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="rocky mountain ecology">
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<paragraph pageId="40" pageNumber="41">Rocky Mountain ecology.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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In alpine habitats in Colorado, in moss near
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix" order="Aplousobranchia" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">Salix</taxonomicName>
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species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="materials_examined">
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<paragraph pageId="40" pageNumber="41">Rocky Mountain specimens examined.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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U.S.A. COLORADO: San Juan County, San Juan Mountains, Mineral Basin, 31 July 2002, CLC1879 (MONT), C. Cripps. Clear Creek County, Denver Mountain Park, Summit Lake, 3911 m, in
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<taxonomicName class="Magnoliopsida" family="Salicaceae" genus="Salix" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Salix arctica" order="Malpighiales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="species" species="arctica">Salix arctica</taxonomicName>
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and
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<taxonomicName lsidName="S. glauca" pageId="40" pageNumber="41" rank="species" species="glauca">S. glauca</taxonomicName>
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, 20 Aug 2013, DBG-F-027678, L. Gillman.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="40" pageNumber="41" type="discussion">
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<paragraph pageId="40" pageNumber="41">Discussion.</paragraph>
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<paragraph pageId="40" pageNumber="41">
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According to
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<bibRefCitation pageId="40" pageNumber="41">Beker et al. (2018)</bibRefCitation>
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,
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<taxonomicName lsidName="H. spetsbergense" pageId="40" pageNumber="41" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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cannot be distinguished from similar species by ITS. The two RM collections (Fig. 6A) differ by 4 [0] bp, the variation of
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<taxonomicName lsidName="H. spetsbergense" pageId="40" pageNumber="41" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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within the sample (7 sequences) is 0-5 [0-2] bp.
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<taxonomicName class="Agaricomycetes" family="Hymenogastraceae" genus="Hebeloma" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Hebeloma nigellum" order="Agaricales" pageId="40" pageNumber="41" phylum="Basidiomycota" rank="species" species="nigellum">Hebeloma nigellum</taxonomicName>
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is the most similar species occurring in the same habitat and, within this sample, differs in its ITS by 1-8 [0-3] bp from
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<taxonomicName lsidName="H. spetsbergense" pageId="40" pageNumber="41" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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. Morphologically
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<taxonomicName lsidName="H. spetsbergense" pageId="40" pageNumber="41" rank="species" species="spetsbergense">H. spetsbergense</taxonomicName>
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is similar to
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<taxonomicName lsidName="H. hygrophilum" pageId="40" pageNumber="41" rank="species" species="hygrophilum">H. hygrophilum</taxonomicName>
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and
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<taxonomicName lsidName="H. nigellum" pageId="40" pageNumber="41" rank="species" species="nigellum">H. nigellum</taxonomicName>
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, but its spores appear to be larger. Previously this species was only known from Svalbard (
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<bibRefCitation pageId="40" pageNumber="41">Beker et al. 2016</bibRefCitation>
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,
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<bibRefCitation pageId="40" pageNumber="41">2018</bibRefCitation>
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), and we report it here from North America for the first time. In Svalbard, it was found with
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<taxonomicName class="Ascidiacea" family="Polycitoridae" genus="Salix" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Salix Polaris" order="Aplousobranchia" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="Polaris">Salix Polaris</taxonomicName>
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near sea level at a latitude of 78°N. In Colorado, it is reported at elevations of 3700-3800 m and latitudes from 36-38°N, and there is a distance between localities of 6500 km, greatly extending its disjunct range. It remains to be seen, if it also occurs in other arctic and alpine habitats.
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</paragraph>
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<paragraph pageId="40" pageNumber="41">
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With the persistent presence of a cortina and the lageniform or ventricose cheilocystidia, this taxon clearly belongs in
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<taxonomicName infraspecific-rank="sect." lsidName="H." pageId="40" pageNumber="41" rank="section" section="Hebeloma">H. sect. Hebeloma</taxonomicName>
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. The rather strongly dextrinoid amygdaloid spores, less than 14
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<normalizedToken originalValue="µm">µm</normalizedToken>
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long but more than 7.5
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<normalizedToken originalValue="µm">µm</normalizedToken>
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wide, distinguish this taxon from the other alpine-arctic species of this section.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |