271 lines
30 KiB
XML
271 lines
30 KiB
XML
<document id="C5AA46558D4340461328525B4D4485E2" ID-DOI="10.11646/zootaxa.3955.3.1" ID-GBIF-Dataset="d7702a61-10c6-4f20-bcce-47ce105dfb5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="234363" ID-ZooBank="0A556FED-C620-4D5D-94BD-CE44F49C33CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461050407125" checkinUser="plazi" docAuthor="Simonsen, Thomas J." docDate="2015" docId="925A87B4FFAAFFFFFF206418B6C58566" docLanguage="en" docName="zt03955p328.pdf" docOrigin="Zootaxa 3955 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elhamma viettei Simonsen, 2015, sp. n." docType="treatment" docVersion="8" lastPageNumber="320" masterDocId="6E63FFCCFFBBFFECFFB76153B626813E" masterDocTitle="Elhamma Walker (Lepidoptera: Hepialidae) revisited: adult morphology, assessment of recently proposed synonyms and descriptions of two species" masterLastPageNumber="328" masterPageNumber="301" pageNumber="318" updateTime="1698578654870" updateUser="plazi">
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<mods:title id="14C9766227DFF22EC50FD9C78F7D6FFA">Elhamma Walker (Lepidoptera: Hepialidae) revisited: adult morphology, assessment of recently proposed synonyms and descriptions of two species</mods:title>
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<mods:name id="836EE296434ED76973AA19A34CDD8C15" type="personal">
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<mods:namePart id="9B0827B947017623FC7295B439806C19">Simonsen, Thomas J.</mods:namePart>
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<mods:title id="D40D9BA391FE017A48A6E0876722B988">Zootaxa</mods:title>
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<mods:part id="B8D65FA467E1518A08DBEE863989F3AF">
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<mods:date id="D9C8B3FD31BD1E41BE4F26B8A5C652ED">2015</mods:date>
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<mods:detail id="0374CFD1840037D41601BD504A9C1155" type="volume">
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<mods:number id="B9C7084A8A34E9D8259110128EC271FE">3955</mods:number>
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<mods:number id="CDF7D3D9A74E48A102DE5969FBE9A3D2">3</mods:number>
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<mods:classification id="96386A8A31538F44382C9CDC77C7B21F">journal article</mods:classification>
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<mods:identifier id="2EF8F630DCFB867BECF3431944E8B3F1" type="DOI">10.11646/zootaxa.3955.3.1</mods:identifier>
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<mods:identifier id="EF4DDFDE47B55298D712307E075C63AE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="925A87B4FFAAFFFFFF206418B6C58566" ID-DOI="http://doi.org/10.5281/zenodo.6102415" ID-GBIF-Taxon="119611773" ID-Zenodo-Dep="6102415" LSID="urn:lsid:plazi:treatment:925A87B4FFAAFFFFFF206418B6C58566" httpUri="http://treatment.plazi.org/id/925A87B4FFAAFFFFFF206418B6C58566" lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318">
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<subSubSection id="52E96529FFAAFFFDFF206418B7B184B8" pageId="17" pageNumber="318" type="nomenclature">
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<paragraph id="1A4C36A2FFAAFFFDFF206418B78B845B" blockId="17.[151,429,1355,1414]" box="[151,429,1355,1381]" pageId="17" pageNumber="318">
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<heading id="410481CEFFAAFFFDFF206418B78B845B" bold="true" box="[151,429,1355,1381]" fontSize="11" level="1" pageId="17" pageNumber="318" reason="1">
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<emphasis id="2887EAB0FFAAFFFDFF206418B78B845B" bold="true" box="[151,429,1355,1381]" pageId="17" pageNumber="318">
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<taxonomicName id="DDF34D21FFAAFFFDFF206418B77A845B" box="[151,348,1355,1381]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="viettei" status="sp. nov.">
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<emphasis id="2887EAB0FFAAFFFDFF206418B77A845B" bold="true" box="[151,348,1355,1381]" italics="true" pageId="17" pageNumber="318">Elhamma viettei</emphasis>
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</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="33B457CBFFAAFFFDFEDC6418B78B845B" box="[363,429,1355,1381]" pageId="17" pageNumber="318" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
<paragraph id="1A4C36A2FFAAFFFDFF20643EB7B184B8" blockId="17.[151,429,1355,1414]" box="[151,407,1389,1414]" pageId="17" pageNumber="318">
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||
(
|
||
<figureCitation id="82C82A27FFAAFFFDFF28643EB73484B8" box="[159,274,1389,1414]" captionStart="FIGURES 1 – 12" captionStartId="1.[151,264,1850,1873]" captionTargetBox="[153,1433,479,1793]" captionTargetId="figure@1.[153,1433,479,1794]" captionTargetPageId="1" captionText="FIGURES 1 – 12. Exemplars of all Elhamma species: 1 – 4, E. australasiae; 1 — female, 2 – 4 — males (2 is E. inconcluso Holotype); 5, E. diakonoffi male (Holotype); 6, E. roebkei male (Holotype); 7 – 8, E. grehani males (7 is Holotype, 8 is paratype RMNH INS 910279); 9 – 10, E. viettei males (9 is Holotype, 10 is paratype RMNH INS 910276); 11 – 12, E. toxopeusi males (11 is Holotype, 12 is the non-type RMNH INS 910272)." httpUri="https://zenodo.org/record/234364/files/figure.png" pageId="17" pageNumber="318">Figs 9–10</figureCitation>
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,
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<figureCitation id="82C82A27FFAAFFFDFEAA643EB74084B8" box="[285,358,1389,1414]" captionStart="FIGURES 42 – 46. E" captionStartId="18.[151,264,1529,1552]" captionTargetBox="[174,1415,209,1501]" captionTargetId="figure@18.[164,1423,193,1508]" captionTargetPageId="18" captionText="FIGURES 42 – 46. E. viettei, male wings and genitalia morphology. 42: Wings; 43: Genitalia, ' classic' slide preparation; 44: Genitalia, ethanol preparation, posterior view; 45: Genitalia, ethanol preparation, lateral view with valva removed—note ' in vivo ' position of S 8; 46: Sternum 8, posterior view." httpUri="https://zenodo.org/record/234374/files/figure.png" pageId="17" pageNumber="318">42–46</figureCitation>
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,
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<figureCitation id="82C82A27FFAAFFFDFEC6643DB7A884B8" box="[369,398,1390,1414]" captionStart="FIGURE 77" captionStartId="25.[151,250,1174,1196]" captionTargetBox="[153,1433,430,1152]" captionTargetId="figure@25.[153,1433,430,1153]" captionTargetPageId="25" captionText="FIGURE 77. Distribution of New Guinea Elhamma species based on specimen records. Filled circle, E. viettei; open circle, E. grehani; filled square, E. toxopeusi; open square, E. diakonoffi; crossed square, E. roebkei." httpUri="https://zenodo.org/record/234380/files/figure.png" pageId="17" pageNumber="318">77</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="52E96529FFAAFFFDFF2064E5B52584D4" pageId="17" pageNumber="318" type="materials_examined">
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||
<paragraph id="1A4C36A2FFAAFFFDFF2064E5B43684F5" blockId="17.[151,771,1461,1514]" box="[151,528,1461,1483]" pageId="17" pageNumber="318">
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||
<typeStatus id="C5488800FFAAFFFDFF2064E5B6EF84F5" box="[151,201,1462,1483]" pageId="17" pageNumber="318">Type</typeStatus>
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||
data:
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<typeStatus id="C5488800FFAAFFFDFEB064E6B74384F5" box="[263,357,1461,1483]" pageId="17" pageNumber="318" type="holotype">Holotype</typeStatus>
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Male, Naturalis.
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</paragraph>
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<paragraph id="1A4C36A2FFAAFFFDFF206486B52584D4" blockId="17.[151,771,1461,1514]" box="[151,771,1492,1514]" pageId="17" pageNumber="318">
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||
<typeStatus id="C5488800FFAAFFFDFF206486B6EF84D4" box="[151,201,1493,1514]" pageId="17" pageNumber="318">Type</typeStatus>
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locality: Walmak, Kecamatan Nipsan,
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<collectingCountry id="62E47632FFAAFFFDFDE26486B4B384D4" box="[597,661,1493,1514]" name="Papua New Guinea" pageId="17" pageNumber="318">Papua</collectingCountry>
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||
,
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<collectingCountry id="62E47632FFAAFFFDFD176487B52584D4" box="[672,771,1492,1514]" name="Indonesia" pageId="17" pageNumber="318">Indonesia</collectingCountry>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="52E96529FFAAFFFDFF206744B403876D" pageId="17" pageNumber="318" type="etymology">
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||
<paragraph id="1A4C36A2FFAAFFFDFF206744B403876D" blockId="17.[151,1437,1559,1980]" pageId="17" pageNumber="318">
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||
<emphasis id="2887EAB0FFAAFFFDFF206744B738870E" bold="true" box="[151,286,1559,1584]" pageId="17" pageNumber="318">Etymology.</emphasis>
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The species is named in honour of the late French
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<taxonomicName id="DDF34D21FFAAFFFDFCDD6744B5C5870E" box="[874,995,1559,1584]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
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expert Pierre Viette (
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<date id="6E4D1062FFAAFFFDFB6D6744B37B8711" box="[1242,1373,1559,1583]" pageId="17" pageNumber="318" value="1921" valueMax="2011">1921–2011</date>
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). The name is a noun in the genitive case.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="52E96529FFAAFFFDFF70670CB4CB8636" pageId="17" pageNumber="318" type="materials_examined">
|
||
<paragraph id="1A4C36A2FFAAFFFDFF70670CB20087A2" blockId="17.[151,1437,1559,1980]" pageId="17" pageNumber="318">
|
||
<emphasis id="2887EAB0FFAAFFFDFF70670CB7958746" bold="true" box="[199,435,1631,1656]" pageId="17" pageNumber="318">Material examined.</emphasis>
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||
<typeStatus id="C5488800FFAAFFFDFE0A670CB4778749" box="[445,593,1631,1655]" pageId="17" pageNumber="318" type="holotype">HOLOTYPE</typeStatus>
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male, (
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||
<collectionCode id="7CE2AE67FFAAFFFDFD1D670CB5268749" LSID="urn:lsid:biocol.org:col:34992" box="[682,768,1631,1655]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="318">RMNH</collectionCode>
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||
<collectionCode id="7CE2AE67FFAAFFFDFCBE670CB5108749" box="[777,822,1631,1655]" pageId="17" pageNumber="318">INS</collectionCode>
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||
910275—dissected, FW:
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||
<quantity id="DD0B9B47FFAAFFFDFBD4670CB2E78749" box="[1123,1217,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="17" pageNumber="318" unit="mm" value="23.9">23.9mm</quantity>
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||
)
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||
<collectingCountry id="62E47632FFAAFFFDFB64670CB3658746" box="[1235,1347,1631,1656]" name="Indonesia" pageId="17" pageNumber="318">Indonesia</collectingCountry>
|
||
,
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||
<collectingCountry id="62E47632FFAAFFFDFAE66733B3BE8749" box="[1361,1432,1632,1655]" name="Papua New Guinea" pageId="17" pageNumber="318">Papua</collectingCountry>
|
||
, Kecamatan Nipsan, Walmak,
|
||
<quantity id="DD0B9B47FFAAFFFDFE5367D7B41287A2" box="[484,564,1668,1692]" metricMagnitude="3" metricUnit="m" metricValue="1.71" pageId="17" pageNumber="318" unit="m" value="1710.0">1710m</quantity>
|
||
, 4˚07’S–138˚38’E,
|
||
<date id="6E4D1062FFAAFFFDFCA667D0B5A587A2" box="[785,899,1667,1692]" pageId="17" pageNumber="318" value="1931-02-01" valueMax="1931-02-16" valueMin="1931-02-01">31/1–16/2</date>
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||
2005, at light.
|
||
</paragraph>
|
||
<paragraph id="1A4C36A2FFAAFFFDFF7067FBB4CB8636" blockId="17.[151,1437,1559,1980]" pageId="17" pageNumber="318">
|
||
<typeStatus id="C5488800FFAAFFFDFF7067FBB70F87FE" box="[199,297,1704,1728]" pageId="17" pageNumber="318" type="paratype">Paratype</typeStatus>
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||
male, (
|
||
<collectionCode id="7CE2AE67FFAAFFFDFECB67F4B7F48781" LSID="urn:lsid:biocol.org:col:34992" box="[380,466,1703,1727]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="318">RMNH</collectionCode>
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<collectionCode id="7CE2AE67FFAAFFFDFE6E67F4B4208781" box="[473,518,1703,1727]" pageId="17" pageNumber="318">INS</collectionCode>
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||
910276—dissected, FW:
|
||
<quantity id="DD0B9B47FFAAFFFDFC9C67F4B5AF87FE" box="[811,905,1703,1728]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" pageId="17" pageNumber="318" unit="mm" value="23.5">23.5mm</quantity>
|
||
)
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||
<collectingCountry id="62E47632FFAAFFFDFC2F67F4B22E87FE" box="[920,1032,1703,1728]" name="Indonesia" pageId="17" pageNumber="318">Indonesia</collectingCountry>
|
||
,
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||
<collectingCountry id="62E47632FFAAFFFDFBA467FBB27C8781" box="[1043,1114,1704,1727]" name="Papua New Guinea" pageId="17" pageNumber="318">Papua</collectingCountry>
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||
, Biak Utara, Sansundi,
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<date id="6E4D1062FFAAFFFDFAD467F4B6F487DA" pageId="17" pageNumber="318" value="2006-06-11">11/6/ 2006</date>
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||
, leg. Rinto H. Mambrasar.
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||
<typeStatus id="C5488800FFAAFFFDFDBF679FB44C87DA" box="[520,618,1740,1764]" pageId="17" pageNumber="318" type="paratype">Paratype</typeStatus>
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||
male, (
|
||
<collectionCode id="7CE2AE67FFAAFFFDFD77679FB53087DA" LSID="urn:lsid:biocol.org:col:34992" box="[704,790,1740,1764]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="318">RMNH</collectionCode>
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||
<collectionCode id="7CE2AE67FFAAFFFDFCA9679FB56D87DA" box="[798,843,1740,1764]" pageId="17" pageNumber="318">INS</collectionCode>
|
||
910277, FW:
|
||
<quantity id="DD0B9B47FFAAFFFDFC466798B26987DA" box="[1009,1103,1739,1764]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="17" pageNumber="318" unit="mm" value="25.8">25.8mm</quantity>
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||
) Bivak 39, Sterren Gebergte, Nieuw
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||
<collectingCountry id="62E47632FFAAFFFDFF5F67BCB71A8636" box="[232,316,1775,1800]" name="Guinea-Bissau" pageId="17" pageNumber="318">Guinea</collectingCountry>
|
||
,
|
||
<quantity id="DD0B9B47FFAAFFFDFEF067BCB7BE8639" box="[327,408,1775,1799]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="318" unit="m" value="1300.0">1300m</quantity>
|
||
, Ned. Exp. 1959,
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||
<date id="6E4D1062FFAAFFFDFDD667BCB4CF8636" box="[609,745,1775,1800]" pageId="17" pageNumber="318" value="1959-06-30">30-VI-1959</date>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52E96529FFAAFFFDFF706640B7FD8682" pageId="17" pageNumber="318" type="distribution">
|
||
<paragraph id="1A4C36A2FFAAFFFDFF706640B7FD8682" blockId="17.[151,1437,1559,1980]" pageId="17" pageNumber="318">
|
||
<emphasis id="2887EAB0FFAAFFFDFF706640B7F68612" bold="true" box="[199,464,1811,1836]" pageId="17" pageNumber="318">
|
||
Distribution (
|
||
<figureCitation id="82C82A27FFAAFFFDFEDD6640B7E48612" box="[362,450,1811,1836]" captionStart="FIGURE 77" captionStartId="25.[151,250,1174,1196]" captionTargetBox="[153,1433,430,1152]" captionTargetId="figure@25.[153,1433,430,1153]" captionTargetPageId="25" captionText="FIGURE 77. Distribution of New Guinea Elhamma species based on specimen records. Filled circle, E. viettei; open circle, E. grehani; filled square, E. toxopeusi; open square, E. diakonoffi; crossed square, E. roebkei." httpUri="https://zenodo.org/record/234380/files/figure.png" pageId="17" pageNumber="318">Fig. 77</figureCitation>
|
||
):
|
||
</emphasis>
|
||
The species is apparently widespread as the three specimens are from three different and quite distant localities. The
|
||
<typeStatus id="C5488800FFAAFFFDFDBF6664B44C866E" box="[520,618,1847,1872]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
|
||
from Walmak in the Jayawijaya Mountains (
|
||
<geoCoordinate id="7FC75065FFAAFFFDFBC36664B2FA8671" box="[1140,1244,1847,1871]" direction="east" orientation="longitude" pageId="17" pageNumber="318" precision="925" value="139.63333">139°38'E</geoCoordinate>
|
||
<geoCoordinate id="7FC75065FFAAFFFDFB526664B3098671" box="[1253,1327,1847,1871]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="925" value="-4.6833334">4°41'S</geoCoordinate>
|
||
) and one
|
||
<typeStatus id="C5488800FFAAFFFDFF20660FB6D1864A" box="[151,247,1884,1908]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
|
||
from the Star Mountains (Bivak 39,
|
||
<geoCoordinate id="7FC75065FFAAFFFDFD166608B4EA864A" box="[673,716,1883,1908]" direction="east" orientation="longitude" pageId="17" pageNumber="318" precision="925" value="140.76666">140</geoCoordinate>
|
||
°46'E
|
||
<geoCoordinate id="7FC75065FFAAFFFDFCA66608B57A864A" box="[785,860,1883,1908]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="925" value="-4.85">4°51'S</geoCoordinate>
|
||
) are both from the central highlands. Whereas the last
|
||
<typeStatus id="C5488800FFAAFFFDFF7066D3B70186A6" box="[199,295,1920,1944]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
|
||
is from the island of Biak (
|
||
<geoCoordinate id="7FC75065FFAAFFFDFDDA662CB4F086A6" box="[621,726,1919,1944]" direction="east" orientation="longitude" pageId="17" pageNumber="318" precision="925" value="135.81667">135°49'E</geoCoordinate>
|
||
<geoCoordinate id="7FC75065FFAAFFFDFD68662CB50F86A9" box="[735,809,1919,1943]" direction="south" orientation="latitude" pageId="17" pageNumber="318" precision="925" value="-0.68333334">0°41'S</geoCoordinate>
|
||
) to the north east of the New
|
||
<collectingCountry id="62E47632FFAAFFFDFB3C662CB2FA86A6" box="[1163,1244,1919,1944]" name="Guinea-Bissau" pageId="17" pageNumber="318">Guinea</collectingCountry>
|
||
mainland (exact localities from de Vos 2013).
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||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="4E8C662AFFA9FFFEFF2064AAB4BC8772" httpUri="https://zenodo.org/record/234374/files/figure.png" pageId="18" pageNumber="319" targetBox="[174,1415,209,1501]" targetPageId="18">
|
||
<paragraph id="1A4C36A2FFA9FFFEFF2064AAB4BC8772" blockId="18.[151,1436,1529,1613]" pageId="18" pageNumber="319">
|
||
<emphasis id="2887EAB0FFA9FFFEFF2064AAB7708730" bold="true" box="[151,342,1529,1552]" pageId="18" pageNumber="319">FIGURES 42–46.</emphasis>
|
||
<emphasis id="2887EAB0FFA9FFFEFEE964A9B79C8731" box="[350,442,1530,1551]" italics="true" pageId="18" pageNumber="319">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA9FFFEFECB64A9B79C8731" box="[380,442,1530,1551]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="viettei">viettei</taxonomicName>
|
||
</emphasis>
|
||
, male wings and genitalia morphology. 42: Wings; 43: Genitalia, 'classic' slide preparation; 44: Genitalia, ethanol preparation, posterior view; 45: Genitalia, ethanol preparation, lateral view with valva removed—note '
|
||
<emphasis id="2887EAB0FFA9FFFEFA3F674AB6E78773" italics="true" pageId="18" pageNumber="319">in vivo</emphasis>
|
||
' position of S8; 46: Sternum 8, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="52E96529FFA9FFFEFF70672AB74B87E4" pageId="18" pageNumber="319" type="diagnosis">
|
||
<paragraph id="1A4C36A2FFA9FFFEFF70672AB74B87E4" blockId="18.[151,1437,1657,2006]" pageId="18" pageNumber="319">
|
||
<emphasis id="2887EAB0FFA9FFFEFF70672AB76787AC" bold="true" box="[199,321,1657,1682]" pageId="18" pageNumber="319">Diagnosis.</emphasis>
|
||
Most similar to
|
||
<emphasis id="2887EAB0FFA9FFFEFDB26729B4BF87AC" box="[517,665,1657,1682]" italics="true" pageId="18" pageNumber="319">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA9FFFEFD9F672AB4BF87AC" box="[552,665,1657,1682]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="diakonoffi">diakonoffi</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="2887EAB0FFA9FFFEFD626729B56C87AC" box="[725,842,1657,1682]" italics="true" pageId="18" pageNumber="319">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA9FFFEFD40672AB56C87AC" box="[759,842,1657,1682]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="roepkei">roepkei</taxonomicName>
|
||
</emphasis>
|
||
, but can be distinguished from both species by the much darker overall habitus including dark brown forewings, dorsal thorax, head, antennae and almost black basal antenna scale tufts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52E96529FFA9FFFFFF7067B6B78582D3" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319" type="description">
|
||
<paragraph id="1A4C36A2FFA9FFFFFF7067B6B4FE80EE" blockId="18.[151,1437,1657,2006]" lastBlockId="19.[151,1437,151,1112]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
|
||
<emphasis id="2887EAB0FFA9FFFEFF7067B6B77F87C0" bold="true" box="[199,345,1765,1790]" pageId="18" pageNumber="319">Description.</emphasis>
|
||
Medium size (FW: HT
|
||
<quantity id="DD0B9B47FFA9FFFEFDDC67B5B4ED87C0" box="[619,715,1766,1790]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="18" pageNumber="319" unit="mm" value="23.9">23.9mm</quantity>
|
||
; PT
|
||
<quantity id="DD0B9B47FFA9FFFEFCB767B6B58687C0" box="[768,928,1765,1790]" metricMagnitude="-2" metricUnit="m" metricValue="2.465" metricValueMax="2.58" metricValueMin="2.35" pageId="18" pageNumber="319" unit="mm" value="24.65" valueMax="25.8" valueMin="23.5">23.5–25.8mm</quantity>
|
||
). Head with dark golden-brown, short, semirough vestiture dorsally, and protruding, rough blackish-brown vestiture frontally; antennae short, with scape scaled, otherwise naked, length more than half the width of the thorax, not serrate, each flagellomere keel-shaped; labial palpus short and pointed; with blackish-brown antennal scale tufts, and a broad circle of blackish-brown scales around the base of the antenna; eyes as described for genus; palps short with dark golden-brown vestiture. Wing pattern as in species diagnosis; wing vestiure type-2 bi-layered; both scale
|
||
<typeStatus id="C5488800FFA9FFFEFBEF66C9B2B5868C" box="[1112,1171,1946,1970]" pageId="18" pageNumber="319">types</typeStatus>
|
||
droplet shaped with a rounded apical margin, but cover scales twice the size as ground scales; both
|
||
<typeStatus id="C5488800FFA9FFFEFC4D66EDB21386E8" box="[1018,1077,1982,2006]" pageId="18" pageNumber="319">types</typeStatus>
|
||
with stout primary ridges, large windows and cross-ribs on abwing surface; secondary ridges on cover scales only; adwing surface with primary ridges, windows and cross-ribs, but windows more numerous on cover scales. FW overall dark olive brown with a mottled pattern of paler dark-beige scales; costal region and remaining margins with a row of dark spots (along latter margins restricted to the ‘cilia’); cubital cells basally with a large, diffuse dark spot of blackish-brown scales. HW uniform pale yellow-beige; margin darker, with ‘cilia’ at the veins being black. Wing venetion generally similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFEBB601EB790805B" box="[268,438,332,357]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFE9B601FB790805B" box="[300,438,332,357]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="australasiae">australasiae</taxonomicName>
|
||
</emphasis>
|
||
, but HW with M3-CuA1 and CuA1-CuA2 cross veins present. Thorax ventrally with a mixture of dark brown and yellow beige scales; legs yellow-golden; tarsal hooks short and stout with a clear basal point; arolium melanised and U-shaped. Abdomen generally yellow beige except of T6-8 which are dark grey. S3- 6 without dark spots; postrior edge of S3-6 smooth.
|
||
</paragraph>
|
||
<paragraph id="1A4C36A2FFA8FFFFFF70608FB3BB822B" blockId="19.[151,1437,151,1112]" pageId="19" pageNumber="320">
|
||
<emphasis id="2887EAB0FFA8FFFFFF70608FB71380CB" bold="true" box="[199,309,476,501]" pageId="19" pageNumber="320">Genitalia</emphasis>
|
||
: Sternum 8 (
|
||
<figureCitation id="82C82A27FFA8FFFFFE7B608FB46980CB" box="[460,591,476,501]" captionStart="FIGURES 42 – 46. E" captionStartId="18.[151,264,1529,1552]" captionTargetBox="[174,1415,209,1501]" captionTargetId="figure@18.[164,1423,193,1508]" captionTargetPageId="18" captionText="FIGURES 42 – 46. E. viettei, male wings and genitalia morphology. 42: Wings; 43: Genitalia, ' classic' slide preparation; 44: Genitalia, ethanol preparation, posterior view; 45: Genitalia, ethanol preparation, lateral view with valva removed—note ' in vivo ' position of S 8; 46: Sternum 8, posterior view." httpUri="https://zenodo.org/record/234374/files/figure.png" pageId="19" pageNumber="320">Figs 45–46</figureCitation>
|
||
) larger than in
|
||
<emphasis id="2887EAB0FFA8FFFFFCB6608EB55E80CB" box="[769,888,476,501]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFC95608FB55E80CB" box="[802,888,476,501]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
, lateral and anterior margins U-shape, posterior margin with a strongly sclerotised trapezoid central projection, margin more sclerotised laterally than in
|
||
<emphasis id="2887EAB0FFA8FFFFFAA86352B3B28326" box="[1311,1428,511,536]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFA8960ACB3B28326" box="[1342,1428,511,536]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
; central hook present and similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFDA86376B4B58303" box="[543,659,548,573]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFD8A6377B4B58303" box="[573,659,548,573]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
. Tergal lobe clearly bilobed and setose. Pseudoteguminal lobes broad and rounded, not very high, similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFDD7631AB4FF835E" box="[608,729,583,608]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFD346314B4FF835E" box="[643,729,583,608]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
; pseudoteguminal arms strongly sclerotised, both dorsal and ventral arms present; dorsal arms short and broad with a pointed tip, curved outwards (pointed straight up in unmounted specimens); ventral arms overall similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFCAC63C2B5B28396" box="[795,916,655,680]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFC8963DCB5B28396" box="[830,916,655,680]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
, long and slender, with a well-defined outer margin, arms synclerotised ventrally terminating in two small, sclerotised tips. Intermediate plate small and narrow, synclerotised with dorso-basal corner of the pseudoteguminal lobe. Valva slightly upwards curved, relatively broad; sacculus short; a strong, sclerotised, inwards-downwards curved tooth present disto-dorsally on sacculus.
|
||
</paragraph>
|
||
<paragraph id="1A4C36A2FFA8FFFFFF70624CB20282F6" blockId="19.[151,1437,151,1112]" pageId="19" pageNumber="320">
|
||
Trulleum overall similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFDBC6272B4A58206" box="[523,643,799,824]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFD99624CB4A58206" box="[558,643,799,824]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
: bilobed at base, attached basally to the base of juxta by a narrow, membranous connection; attached high on the ventral pseudoteguminal arms by a narrow membranous connection. Juxta similar to
|
||
<emphasis id="2887EAB0FFA8FFFFFEFE623AB79882BE" box="[329,446,871,896]" italics="true" pageId="19" pageNumber="320">
|
||
E.
|
||
<taxonomicName id="DDF34D21FFA8FFFFFEDF6234B79882BE" box="[360,446,871,896]" class="Insecta" family="Hepialidae" genus="Elhamma" kingdom="Animalia" order="Lepidoptera" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="grehani">grehani</taxonomicName>
|
||
</emphasis>
|
||
; deeply cup shaped with a basal ridge. Phallus very long, approximately
|
||
<date id="6E4D1062FFA8FFFFFB5B6234B32F82BE" box="[1260,1289,871,896]" pageId="19" pageNumber="320">6x</date>
|
||
height of the genitalia. Vinculum and saccus broad U-shape with a flat dorsal cross-ridge and a U-shaped sulcus separating vinculum proper and the apodemal vinculum (sensu Nielsen & Kristensen 1989).
|
||
</paragraph>
|
||
<paragraph id="1A4C36A2FFA8FFFFFF706287B78582D3" blockId="19.[151,1437,151,1112]" box="[199,419,980,1005]" pageId="19" pageNumber="320">
|
||
<emphasis id="2887EAB0FFA8FFFFFF706287B73882D3" bold="true" box="[199,286,980,1005]" pageId="19" pageNumber="320">Female</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52E96529FFA8FFFFFF7062A4B6C58566" pageId="19" pageNumber="320" type="discussion">
|
||
<paragraph id="1A4C36A2FFA8FFFFFF7062A4B6C58566" blockId="19.[151,1437,151,1112]" pageId="19" pageNumber="320">
|
||
<emphasis id="2887EAB0FFA8FFFFFF7062A4B71D852E" bold="true" box="[199,315,1015,1040]" pageId="19" pageNumber="320">Remarks.</emphasis>
|
||
Females are unknown, and nothing is known about the biology of the species. But an additional label on the
|
||
<typeStatus id="C5488800FFA8FFFFFE96654FB7AF850B" box="[289,393,1052,1077]" pageId="19" pageNumber="320" type="holotype">Holotype</typeStatus>
|
||
reads: “cultivated area/disturbed montane forest UNCEN-ZMA Expedition
|
||
<collectingCountry id="62E47632FFA8FFFFFB54654EB301850A" box="[1251,1319,1053,1076]" name="Papua New Guinea" pageId="19" pageNumber="320">Papua</collectingCountry>
|
||
<collectingCountry id="62E47632FFA8FFFFFA98654FB3BA850B" box="[1327,1436,1052,1077]" name="Indonesia" pageId="19" pageNumber="320">Indonesia</collectingCountry>
|
||
2005”.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |