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<document id="59A319B5393A638C447385414FC55AC4" ID-CLB-Dataset="26928" ID-DOI="10.11646/zootaxa.4596.1.1" ID-GBIF-Dataset="cb288575-93a8-4e59-86fd-ee6bae784b6a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2659091" ID-ZooBank="B88EA161-04D3-4D50-B3FD-E98A60DA3740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556872200693" checkinUser="plazi" docAuthor="A, Contreras-Félix Gerardo &amp; Oscar, Francke B." docDate="2019" docId="022787FEFFA91952FF163931FA32FA3A" docLanguage="en" docName="zootaxa.4596.1.1.pdf" docOrigin="Zootaxa 4596 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Vaejovis mexicanus C. L. Koch 1836" docType="key" docVersion="12" lastPageNumber="18" masterDocId="FE1EFF86FFA41943FF813C15FFA7FF9B" masterDocTitle="Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)" masterLastPageNumber="100" masterPageNumber="1" pageNumber="14" updateTime="1698725788431" updateUser="ExternalLinkService">
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<mods:title id="8E0DC594A4A87A521B831B879F516CFA">Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)</mods:title>
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<mods:namePart id="99D4C82777DACCBDABADDC56A14BF7BD">A, Contreras-Félix Gerardo</mods:namePart>
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<mods:namePart id="82BA69E2B3276A13E81F8535896A7915">Oscar, Francke B.</mods:namePart>
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<treatment id="022787FEFFA91952FF163931FA32FA3A" ID-GBIF-Taxon="156941558" LSID="urn:lsid:plazi:treatment:022787FEFFA91952FF163931FA32FA3A" httpUri="http://treatment.plazi.org/id/022787FEFFA91952FF163931FA32FA3A" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
<subSubSection id="C2946563FFA9194EFF163931FC50FAA5" box="[151,1015,1316,1342]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8A3136E8FFA9194EFF163931FC50FAA5" blockId="13.[151,1015,1316,1342]" box="[151,1015,1316,1342]" pageId="13" pageNumber="14">
<heading id="D1798184FFA9194EFF163931FC50FAA5" bold="true" box="[151,1015,1316,1342]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
<emphasis id="B8FAEAFAFFA9194EFF163931FC50FAA5" bold="true" box="[151,1015,1316,1342]" pageId="13" pageNumber="14">
Key to identification of males of the species in the “
<taxonomicName id="4D8E4D6BFFA9194EFC943931FC30FAA5" ID-CoL="7F8PV" authorityName="C. L. Koch" authorityYear="1836" box="[789,919,1316,1342]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B8FAEAFAFFA9194EFC943931FC30FAA5" bold="true" box="[789,919,1316,1342]" italics="true" pageId="13" pageNumber="14">mexicanus</emphasis>
</taxonomicName>
” group
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C2946563FFA91952FF16397FFA32FA3A" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14" type="key">
<keyStep id="317A2E4DFFA9194EFF16397FFA3CFA4C" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF16397FFA3CFA06" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF16397FFA3CFA06" pageId="13" pageNumber="14">
1 Lateral inframedian carinae usually with two setae at least on the first three segment of the metasoma; three
<emphasis id="B8FAEAFAFFA9194EFB7B397FFA85FA1B" box="[1274,1314,1386,1408]" italics="true" pageId="13" pageNumber="14">pi/ri</emphasis>
setae on the third and fourth telotarsi................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF1639B0FA3CFA4C" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF1639B0FA3CFA4C" pageId="13" pageNumber="14">
1 Lateral inframedian carinae with one or no seta on the first three segments of the metasoma; two or less
<emphasis id="B8FAEAFAFFA9194EFB4C39B1FB51FA21" box="[1229,1270,1444,1466]" italics="true" pageId="13" pageNumber="14">pi/ri</emphasis>
setae on the third and fourth telotarsi.................................................................................... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF1639F5FA3BF989" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF1639F5FA3BFA6E" blockId="13.[151,1436,1386,2033]" box="[151,1436,1504,1525]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF1639F5FA3BFA6E" box="[151,1436,1504,1525]" pageId="13" pageNumber="14">2(1) Hemispermatophore lamella short and compact; less than two times longer than wide (L/W = 1.76) (Fig: 6-a)........... 3</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF1639E8FA3BF989" blockId="13.[151,1436,1386,2033]" box="[151,1436,1533,1554]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF1639E8FA3BF989" box="[151,1436,1533,1554]" pageId="13" pageNumber="14">2 Hemispermatophore lamella long and slender; more than three times longer than wide (L/W = 3.8) (Fig: 6-b)........... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF163A0FFA3BF9D6" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF163A0FFA3BF9AB" blockId="13.[151,1436,1386,2033]" box="[151,1436,1562,1584]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163A0FFA3BF9AB" box="[151,1436,1562,1584]" pageId="13" pageNumber="14">
3(2) Vesicle thin, twice longer than wide...........................................................
<taxonomicName id="4D8E4D6BFFA9194EFB7E3A0EFAF8F9AB" ID-CoL="7F928" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[1279,1375,1562,1584]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tapalpa" status="sp. nov.">
<emphasis id="B8FAEAFAFFA9194EFB7E3A0EFAF8F9AB" box="[1279,1375,1562,1584]" italics="true" pageId="13" pageNumber="14">V. tapalpa</emphasis>
</taxonomicName>
<emphasis id="B8FAEAFAFFA9194EFAE73A0FFA3BF9AB" bold="true" box="[1382,1436,1562,1584]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A3C95781FFA9194EFAE73A0FFA3BF9AB" box="[1382,1436,1562,1584]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF163A2DFA3BF9D6" blockId="13.[151,1436,1386,2033]" box="[151,1436,1591,1613]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163A2DFA3BF9D6" box="[151,1436,1591,1613]" pageId="13" pageNumber="14">
3 Vesicle globose, usually as long as wide...........................................................
<taxonomicName id="4D8E4D6BFFA9194EFAA23A2DFA3BF9D6" ID-CoL="7F8Q6" authorityName="Sissom" authorityYear="1989" box="[1315,1436,1591,1613]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B8FAEAFAFFA9194EFAA23A2DFA3BF9D6" box="[1315,1436,1591,1613]" italics="true" pageId="13" pageNumber="14">V. monticola</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF163A40FA3CF91C" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF163A40FA3CF9F1" blockId="13.[151,1436,1386,2033]" box="[151,1435,1620,1642]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163A40FA3CF9F1" box="[151,1435,1620,1642]" pageId="13" pageNumber="14">
4(2) Caudal gland absent (Fig: 7-a).................................................................
<taxonomicName id="4D8E4D6BFFA9194EFA9A3A40FA3CF9F1" ID-CoL="7F8N2" authorityName="Contreras-Felix &amp; Francke" authorityYear="2014" box="[1307,1435,1620,1642]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="coalcoman">
<emphasis id="B8FAEAFAFFA9194EFA9A3A40FA3CF9F1" box="[1307,1435,1620,1642]" italics="true" pageId="13" pageNumber="14">V. coalcoman</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF163A67FA3CF91C" blockId="13.[151,1436,1386,2033]" box="[151,1435,1650,1671]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163A67FA3CF91C" box="[151,1435,1650,1671]" pageId="13" pageNumber="14">4 Caudal gland present (Fig: 7-b)......................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF163A85FA3CF959" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF163A85FA3BF93E" blockId="13.[151,1436,1386,2033]" box="[151,1436,1679,1701]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163A85FA3BF93E" box="[151,1436,1679,1701]" pageId="13" pageNumber="14">
5(4) Pedipalp chela manus with more than four granular carinae (Fig: 7-c)...........................
<taxonomicName id="4D8E4D6BFFA9194EFB4E3A85FAF8F93E" ID-CoL="7F8PP" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[1231,1375,1679,1701]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="santibagnezi" status="sp. nov.">
<emphasis id="B8FAEAFAFFA9194EFB4E3A85FAF8F93E" box="[1231,1375,1679,1701]" italics="true" pageId="13" pageNumber="14">V. santibagnezi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C95781FFA9194EFAE73A9AFA3BF93E" box="[1382,1436,1679,1701]" pageId="13" pageNumber="14" rank="species">
<emphasis id="B8FAEAFAFFA9194EFAE73A9AFA3BF93E" bold="true" box="[1382,1436,1679,1701]" pageId="13" pageNumber="14">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF163AB8FA3CF959" blockId="13.[151,1436,1386,2033]" box="[151,1435,1709,1730]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163AB8FA3CF959" box="[151,1435,1709,1730]" pageId="13" pageNumber="14">5 Pedipalp chela manus with three or fewer granular carinae (Fig: 7-d)............................................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF163ADFFA3BF8A3" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF163ADFFA3BF966" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163ADFFA3BF966" pageId="13" pageNumber="14">
6(5) Pedipalp chela ventral accessory carina granular; hemispermatophore hooks bent 90° at the tip; pectinal tooth counts 1415........................................................................................
<taxonomicName id="4D8E4D6BFFA9194EFB6E3AFDFAF8F966" ID-CoL="7F8MR" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[1263,1375,1767,1789]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ceboruco" status="sp. nov.">
<emphasis id="B8FAEAFAFFA9194EFB6E3AFDFAF8F966" box="[1263,1375,1767,1789]" italics="true" pageId="13" pageNumber="14">V. ceboruco</emphasis>
</taxonomicName>
<emphasis id="B8FAEAFAFFA9194EFAE73AF2FA3BF966" bold="true" box="[1382,1436,1767,1789]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A3C95781FFA9194EFAE73AF2FA3BF966" box="[1382,1436,1767,1789]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF163B10FA3BF8A3" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163B10FA3BF8A3" pageId="13" pageNumber="14">
6´Pedipalp chela ventral accessory carina smooth or absent: hemispermatophore hooks straight; pectinal tooth counts 1112............................................................................................
<taxonomicName id="4D8E4D6BFFA9194EFA963B36FAF8F8A3" ID-CoL="7F8Q8" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[1303,1375,1826,1848]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="talpa" status="sp. nov.">
<emphasis id="B8FAEAFAFFA9194EFA963B36FAF8F8A3" box="[1303,1375,1826,1848]" italics="true" pageId="13" pageNumber="14">V. talpa</emphasis>
</taxonomicName>
<emphasis id="B8FAEAFAFFA9194EFAE73B37FA3BF8A3" bold="true" box="[1382,1436,1826,1848]" pageId="13" pageNumber="14">
<taxonomicNameLabel id="A3C95781FFA9194EFAE73B37FA3BF8A3" box="[1382,1436,1826,1848]" pageId="13" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFA9194EFF163B55FA3BF82D" pageId="13" pageNumber="14">
<paragraph id="8A3136E8FFA9194EFF163B55FA3BF8EE" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163B55FA3BF8EE" pageId="13" pageNumber="14">7(1) Caudal gland evident, covering about one third of the dorsal surface of the vesicle (Fig: 7-e); hemispermatophore short, with an apical crest and a well-sclerotized median lobe with small granules on the edge.................................. 8</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194EFF163B94FA3BF82D" blockId="13.[151,1436,1386,2033]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163B94FA3BF82D" pageId="13" pageNumber="14">7 Caudal gland small or absent (Fig: 7-f); hemispermatophore without an apical crest and the median lobe weak and translucent................................................................................................... 15</keyLead>
</paragraph>
</keyStep>
<paragraph id="8A3136E8FFA9194EFF163BABFA3CF84F" blockId="13.[151,1436,1386,2033]" box="[151,1435,1982,2004]" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194EFF163BABFA3CF84F" box="[151,1435,1982,2004]" pageId="13" pageNumber="14">
<keyStep id="317A2E4DFFA9194EFF163BABFA3CF84F" box="[151,1435,1982,2004]" pageId="13" pageNumber="14">
8(7) Mesosomal sternite V with light-colored glandular area not differentiated (Fig: 7-g)......................
<taxonomicName id="4D8E4D6BFFA9194EFA903BAAFA3CF84F" ID-CoL="7F8QG" authorityName="Contreras-Felix, Francke &amp; Bryson" authorityYear="2015" box="[1297,1435,1982,2004]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="tenamaztlei">
<emphasis id="B8FAEAFAFFA9194EFA903BAAFA3CF84F" box="[1297,1435,1982,2004]" italics="true" pageId="13" pageNumber="14">V. tenamaztlei</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFA9194DFF163BC9FA3CFA84" blockId="13.[151,1436,1386,2033]" lastBlockId="14.[151,1437,1290,1986]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<keyLead id="317F95DDFFA9194DFF163BC9FA3CFA84" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
8 Mesosomal sternite V with distinct, whitish glandular area (Fig: 7-h)........................................... 9
<keyStep id="317A2E4DFFAA194DFF16391FFA3CFA84" box="[151,1435,1290,1311]" pageId="14" pageNumber="15">9(8) Mesosomal sternite V with the glandular area reaching the anterior margin of the segment.......................... 10</keyStep>
</keyLead>
</paragraph>
<caption id="DEF16660FFAA194DFF163864FDD2FB78" httpUri="https://zenodo.org/record/2659105/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,250,1137,1159]" targetBox="[484,1101,193,1100]" targetPageId="14">
<paragraph id="8A3136E8FFAA194DFF163864FDD2FB78" blockId="14.[151,1436,1137,1251]" pageId="14" pageNumber="15">
<emphasis id="B8FAEAFAFFAA194DFF163864FE86FB1C" bold="true" box="[151,289,1137,1159]" pageId="14" pageNumber="15">FIGURE 6.</emphasis>
Characters diagnosing species from the “
<taxonomicName id="4D8E4D6BFFAA194DFC9F3867FC2EFB1C" ID-CoL="7F8PV" authorityName="C. L. Koch" authorityYear="1836" box="[798,905,1138,1159]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B8FAEAFAFFAA194DFC9F3867FC2EFB1C" box="[798,905,1138,1159]" italics="true" pageId="14" pageNumber="15">mexicanus</emphasis>
</taxonomicName>
” group.
<emphasis id="B8FAEAFAFFAA194DFC7E3864FB82FB1C" bold="true" box="[1023,1061,1137,1159]" pageId="14" pageNumber="15">a—</emphasis>
Hemispermatophore
<taxonomicName id="4D8E4D6BFFAA194DFA883867FA32FB1C" ID-CoL="7F8Q6" authorityName="Sissom" authorityYear="1989" box="[1289,1429,1137,1159]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B8FAEAFAFFAA194DFA883867FA32FB1C" box="[1289,1429,1137,1159]" italics="true" pageId="14" pageNumber="15">V. monticola</emphasis>
</taxonomicName>
.
<emphasis id="B8FAEAFAFFAA194DFF163885FF19FB3D" bold="true" box="[151,190,1168,1190]" pageId="14" pageNumber="15">b—</emphasis>
Hemispermatophore of
<taxonomicName id="4D8E4D6BFFAA194DFE313884FD80FB3D" ID-CoL="7F8MR" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[432,551,1168,1190]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ceboruco" status="sp. nov.">
<emphasis id="B8FAEAFAFFAA194DFE313884FD80FB3D" box="[432,551,1168,1190]" italics="true" pageId="14" pageNumber="15">V. ceboruco</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A3C95781FFAA194DFDAF3884FDDAFB3D" box="[558,637,1169,1190]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
red line, representing the longitude of the lamella; black line, representing the comparative longitude of the trunk; red arrow, signaling the point where each hemispermatophore are wider; Blue arrow, signaling the median lobe and its development.
</paragraph>
</caption>
<paragraph id="8A3136E8FFAA194DFF16393DFA3CFAA6" blockId="14.[151,1437,1290,1986]" box="[151,1435,1320,1341]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF16393DFA3CFAA6" box="[151,1435,1320,1341]" pageId="14" pageNumber="15">9 Mesosomal sternite V with the glandular area marginal, only present on the posterior half of the segment.............. 12</keyLead>
</paragraph>
<keyStep id="317A2E4DFFAA194DFF163950FA3CFAEC" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF163950FA3CFAC1" blockId="14.[151,1437,1290,1986]" box="[151,1435,1349,1370]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163950FA3CFAC1" box="[151,1435,1349,1370]" pageId="14" pageNumber="15">
10(9) Intercarinal spaces on metasomal segments feebly granulose..........................................
<taxonomicName id="4D8E4D6BFFAA194DFAA63950FA3CFAC1" ID-CoL="7F927" authorityName="Graham &amp; Bryson" authorityYear="2010" box="[1319,1435,1349,1370]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B8FAEAFAFFAA194DFAA63950FA3CFAC1" box="[1319,1435,1349,1370]" italics="true" pageId="14" pageNumber="15">V. montanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163977FA3CFAEC" blockId="14.[151,1437,1290,1986]" box="[151,1435,1378,1399]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163977FA3CFAEC" box="[151,1435,1378,1399]" pageId="14" pageNumber="15">10 Intercarinal spaces on metasoma segments distinctly granulose................................................ 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF163995FA3CFA29" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF163995FA3CFA0E" blockId="14.[151,1437,1290,1986]" box="[151,1435,1408,1429]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163995FA3CFA0E" box="[151,1435,1408,1429]" pageId="14" pageNumber="15">
11(10) Caudal gland with a strong and noticeable depression..................................................
<taxonomicName id="4D8E4D6BFFAA194DFABE3995FA3CFA0E" ID-CoL="7F8PY" box="[1343,1435,1408,1429]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sierrae">
<emphasis id="B8FAEAFAFFAA194DFABE3995FA3CFA0E" box="[1343,1435,1408,1429]" italics="true" pageId="14" pageNumber="15">V. sierrae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163988FA3CFA29" blockId="14.[151,1437,1290,1986]" box="[151,1435,1437,1458]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163988FA3CFA29" box="[151,1435,1437,1458]" pageId="14" pageNumber="15">
11 Caudal gland flat or slightly depressed.............................................................
<taxonomicName id="4D8E4D6BFFAA194DFAB63988FA3CFA29" ID-CoL="7F8PG" box="[1335,1435,1437,1458]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mcwesti">
<emphasis id="B8FAEAFAFFAA194DFAB63988FA3CFA29" box="[1335,1435,1437,1458]" italics="true" pageId="14" pageNumber="15">V. mcwesti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF1639AFFA3CFA76" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF1639AFFA3CFA4B" blockId="14.[151,1437,1290,1986]" box="[151,1435,1466,1488]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF1639AFFA3CFA4B" box="[151,1435,1466,1488]" pageId="14" pageNumber="15">
12(9) Caudal gland well-developed, almost as long as the vesicle (Fig: 7-c).....................................
<taxonomicName id="4D8E4D6BFFAA194DFAC439AEFA3CFA4B" ID-CoL="7F8MX" authorityName="Pocock" authorityYear="1902" box="[1349,1435,1466,1488]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dugesi">
<emphasis id="B8FAEAFAFFAA194DFAC439AEFA3CFA4B" box="[1349,1435,1466,1488]" italics="true" pageId="14" pageNumber="15">V. dugesi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF1639CDFA3CFA76" blockId="14.[151,1437,1290,1986]" box="[151,1435,1496,1517]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF1639CDFA3CFA76" box="[151,1435,1496,1517]" pageId="14" pageNumber="15">12 Caudal gland poorly developed or absent (Fig: 7-d).......................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF1639E0FA3CF9F9" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF1639E0FA3CF9B3" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF1639E0FA3CF9B3" pageId="14" pageNumber="15">
13(12) Pedipalp chela with the dorsal marginal carina absent; fixed finger almost as long as the pedipalp chela manus (CM L/FF 0.98± 0.02); pectinal tooth count higher than 15 (usually 1819).............................................
<taxonomicName id="4D8E4D6BFFAA194DFAB23A06FA3CF9B3" ID-CoL="7F929" authorityName="Sissom" authorityYear="1990" box="[1331,1435,1555,1576]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sprousei">
<emphasis id="B8FAEAFAFFAA194DFAB23A06FA3CF9B3" box="[1331,1435,1555,1576]" italics="true" pageId="14" pageNumber="15">V. sprousei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163A25FA3CF9F9" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163A25FA3CF9F9" pageId="14" pageNumber="15">13 Pedipalp chela with the dorsal marginal carina present; fixed finger shorter than the pedipalp chela manus (CM L/FF Ẽ 1); pectinal tooth counts lower than 15......................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF163A7FFA3BF943" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF163A7FFA3CF906" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163A7FFA3CF906" pageId="14" pageNumber="15">
14(13) Vesicle globular (L/W= 1.62); pedipalp patella more than three times longer than wide; hemispermatophore lamella bent 180° at the tip....................................................................................
<taxonomicName id="4D8E4D6BFFAA194DFAA63A9DFA3CF906" ID-CoL="7F8QF" authorityName="Hendrixson &amp; Sissom" authorityYear="2001" box="[1319,1435,1671,1693]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tesselatus">
<emphasis id="B8FAEAFAFFAA194DFAA63A9DFA3CF906" box="[1319,1435,1671,1693]" italics="true" pageId="14" pageNumber="15">V. tesselatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163AB0FA3BF943" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163AB0FA3BF943" pageId="14" pageNumber="15">
14 Vesicle thin (L/W= 2.08); pedipalp patella less than three times longer than wide; hemispermatophore lamella tip straight...............................................................................................
<taxonomicName id="4D8E4D6BFFAA194DFAAC3AD6FA3BF943" ID-CoL="7F922" authorityName="Sissom" authorityYear="1989" box="[1325,1436,1731,1752]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="rossmani">
<emphasis id="B8FAEAFAFFAA194DFAAC3AD6FA3BF943" box="[1325,1436,1731,1752]" italics="true" pageId="14" pageNumber="15">V. rossmani</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF163AF5FA3CF889" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF163AF5FA3FF96E" blockId="14.[151,1437,1290,1986]" box="[151,1432,1759,1781]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163AF5FA3FF96E" box="[151,1432,1759,1781]" pageId="14" pageNumber="15">
15(7) One telotarsal ventral distal spinule...............................................................
<taxonomicName id="4D8E4D6BFFAA194DFABB3AF5FA3FF96E" ID-CoL="7F8N3" authorityName="Sissom" authorityYear="1989" box="[1338,1432,1759,1781]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chiapas">
<emphasis id="B8FAEAFAFFAA194DFABB3AF5FA3FF96E" box="[1338,1432,1759,1781]" italics="true" pageId="14" pageNumber="15">V. chiapas</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163AE8FA3CF889" blockId="14.[151,1437,1290,1986]" box="[151,1435,1789,1810]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163AE8FA3CF889" box="[151,1435,1789,1810]" pageId="14" pageNumber="15">15 Two or more telotarsal ventral distal spinules.............................................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAA194DFF163B0FFA3CF81C" pageId="14" pageNumber="15">
<paragraph id="8A3136E8FFAA194DFF163B0FFA3BF8D6" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163B0FFA3BF8D6" pageId="14" pageNumber="15">16(15) Vesicle rounded and flattened dorsoventrally, less than one and a half times longer than wide and at most one and a half times wider than deep (Fig: 8-a); pedipalp chela fingers with basal notch (Fig: 8-b).................................... 17</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194DFF163B40FA3CF81C" blockId="14.[151,1437,1290,1986]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163B40FA3CF81C" pageId="14" pageNumber="15">16 Vesicle thin or globular, more than one and a half times longer than wide and at least one and a half times wider than deep (Fig: 8-c); pedipalp chela fingers straight basally (Fig: 8-d)...................................................... 20</keyLead>
</paragraph>
</keyStep>
<paragraph id="8A3136E8FFAA194DFF163B85FA3BF83E" blockId="14.[151,1437,1290,1986]" box="[151,1436,1935,1957]" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194DFF163B85FA3BF83E" box="[151,1436,1935,1957]" pageId="14" pageNumber="15">
<keyStep id="317A2E4DFFAA194DFF163B85FA3BF83E" box="[151,1436,1935,1957]" pageId="14" pageNumber="15">
17(16) Caudal gland absent.....................................................................
<taxonomicName id="4D8E4D6BFFAA194DFB703B85FA3BF83E" ID-CoL="7F925" authorityName="Hendrixson &amp; Sissom" authorityYear="2001" box="[1265,1436,1935,1957]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nigrofemoratus">
<emphasis id="B8FAEAFAFFAA194DFB703B85FA3BF83E" box="[1265,1436,1935,1957]" italics="true" pageId="14" pageNumber="15">V. nigrofemoratus</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAA194CFF163BB8FA3CFAF7" blockId="14.[151,1437,1290,1986]" lastBlockId="15.[151,1436,1366,2033]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<keyLead id="317F95DDFFAA194CFF163BB8FA3CFAF7" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
17 Caudal gland vestigial................................................................................ 18
<keyStep id="317A2E4DFFAB194CFF163943FA3CFAF7" box="[151,1435,1366,1388]" pageId="15" pageNumber="16">
18(17) Metasomal intercarinal spaces smooth; metasomal carinae granular.....................................
<taxonomicName id="4D8E4D6BFFAB194CFAB43942FA3CFAF7" ID-CoL="7F8N7" authorityName="Sissom" authorityYear="1989" box="[1333,1435,1366,1388]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="franckei">
<emphasis id="B8FAEAFAFFAB194CFAB43942FA3CFAF7" box="[1333,1435,1366,1388]" italics="true" pageId="15" pageNumber="16">V. franckei</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
<caption id="DEF16660FFAB194CFF163874FF42FAB4" httpUri="https://zenodo.org/record/2659107/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1121,1143]" targetBox="[322,1264,193,1100]" targetPageId="15">
<paragraph id="8A3136E8FFAB194CFF163874FF42FAB4" blockId="15.[151,1437,1121,1327]" pageId="15" pageNumber="16">
<emphasis id="B8FAEAFAFFAB194CFF163874FEB1FBEC" bold="true" box="[151,278,1121,1143]" pageId="15" pageNumber="16">FIGURE 7.</emphasis>
Characters to identification species with males:
<emphasis id="B8FAEAFAFFAB194CFC973874FC9CFBEC" bold="true" box="[790,827,1121,1143]" pageId="15" pageNumber="16">a—</emphasis>
Telson showing the vesicular gland, highlighted in red (
<taxonomicName id="4D8E4D6BFFAB194CFA083877FF7EFB0D" ID-CoL="7F8MX" authorityName="Pocock" authorityYear="1902" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dugesi">
<emphasis id="B8FAEAFAFFAB194CFA083877FF7EFB0D" italics="true" pageId="15" pageNumber="16">V. dugesi</emphasis>
</taxonomicName>
);
<emphasis id="B8FAEAFAFFAB194CFF763895FEB9FB0D" bold="true" box="[247,286,1152,1174]" pageId="15" pageNumber="16">b—</emphasis>
Telson showing an absent vesicular gland (
<taxonomicName id="4D8E4D6BFFAB194CFC863894FC24FB0D" ID-CoL="7F8QB" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[775,899,1153,1174]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="zapoteca">
<emphasis id="B8FAEAFAFFAB194CFC863894FC24FB0D" box="[775,899,1153,1174]" italics="true" pageId="15" pageNumber="16">V. zapoteca</emphasis>
</taxonomicName>
);
<emphasis id="B8FAEAFAFFAB194CFC203895FC62FB0D" bold="true" box="[929,965,1152,1174]" pageId="15" pageNumber="16">c—</emphasis>
Pedipalp chela manus, showing the strong development of carinae (
<taxonomicName id="4D8E4D6BFFAB194CFE1338B5FD8DFB2F" ID-CoL="7F8PP" authorityName="A &amp; Oscar" authorityYear="2019" box="[402,554,1182,1205]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="santibagnezi">
<emphasis id="B8FAEAFAFFAB194CFE1338B5FD8DFB2F" box="[402,554,1182,1205]" italics="true" pageId="15" pageNumber="16">V. santibagnezi</emphasis>
</taxonomicName>
);
<emphasis id="B8FAEAFAFFAB194CFDC1388BFDC0FB2F" bold="true" box="[576,615,1182,1204]" pageId="15" pageNumber="16">d—</emphasis>
Pedipalp chela manus, showing the poor development of carinae (
<taxonomicName id="4D8E4D6BFFAB194CFB7E38B5FA2AFB2F" ID-CoL="7F8QG" authorityName="Contreras-Felix, Francke &amp; Bryson" authorityYear="2015" box="[1279,1421,1182,1205]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="tenamaztlei">
<emphasis id="B8FAEAFAFFAB194CFB7E38B5FA2AFB2F" box="[1279,1421,1182,1205]" italics="true" pageId="15" pageNumber="16">V. tenamaztlei</emphasis>
</taxonomicName>
);
<emphasis id="B8FAEAFAFFAB194CFF1638A8FF1CFB48" bold="true" box="[151,187,1213,1235]" pageId="15" pageNumber="16">e—</emphasis>
Telson with well-developed vesicular gland (highlighted in red);
<emphasis id="B8FAEAFAFFAB194CFCEB38A8FC2CFB48" bold="true" box="[874,907,1213,1235]" pageId="15" pageNumber="16">f—</emphasis>
Telson with vesicular gland moderately developed (Highlighted in red);
<emphasis id="B8FAEAFAFFAB194CFEFB38C9FE07FB69" bold="true" box="[378,416,1244,1266]" pageId="15" pageNumber="16">g—</emphasis>
Mesosomal sternite V (
<taxonomicName id="4D8E4D6BFFAB194CFD1938C8FCB8FB69" ID-CoL="7F8PJ" authorityName=", Sissom" authorityYear="1989" box="[664,799,1244,1266]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8FAEAFAFFAB194CFD1938C8FCB8FB69" box="[664,799,1244,1266]" italics="true" pageId="15" pageNumber="16">V. maculosus</emphasis>
</taxonomicName>
), highlighted in red the small white patch at the end of the segment;
<emphasis id="B8FAEAFAFFAB194CFF7938EEFEB8FA8A" bold="true" box="[248,287,1275,1297]" pageId="15" pageNumber="16">h—</emphasis>
Mesosomal sternite V (
<taxonomicName id="4D8E4D6BFFAB194CFD8D38E9FD20FA8A" ID-CoL="7F927" authorityName="Graham &amp; Bryson" authorityYear="2010" box="[524,647,1276,1297]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="montanus">
<emphasis id="B8FAEAFAFFAB194CFD8D38E9FD20FA8A" box="[524,647,1276,1297]" italics="true" pageId="15" pageNumber="16">V. montanus</emphasis>
</taxonomicName>
) highlighted in red the white patch from the posterior edge, reaching the front margin.
</paragraph>
</caption>
<paragraph id="8A3136E8FFAB194CFF163961FA3CFA3D" blockId="15.[151,1436,1366,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163961FA3CFA3D" pageId="15" pageNumber="16">18 Metasomal intercarinal spaces granulated or if they are smooth then the metasomal carinae have all their granules basally fused................................................................................................... 19</keyLead>
</paragraph>
<keyStep id="317A2E4DFFAB194CFF1639BBFA3CF987" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF1639BBFA3CFA7A" blockId="15.[151,1436,1366,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF1639BBFA3CFA7A" pageId="15" pageNumber="16">
19(18) Metasomal intercarinal spaces granular; metasomal carinae usually composed by a line of granules and serrated or crenulated..........................................................................................
<taxonomicName id="4D8E4D6BFFAB194CFA9D39D9FA3CFA7A" ID-CoL="7F8N5" authorityName="Pocock" authorityYear="1898" box="[1308,1435,1483,1505]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B8FAEAFAFFAB194CFA9D39D9FA3CFA7A" box="[1308,1435,1483,1505]" italics="true" pageId="15" pageNumber="16">V. granulatus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF1639FCFA3CF987" blockId="15.[151,1436,1366,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF1639FCFA3CF987" pageId="15" pageNumber="16">
19 Metasomal intercarinal spaces smooth; metasomal carinae composed by a raised ridge of the cuticle, and all the granules of the carinae basally fused (Fig: 8-e)...................................................................
<taxonomicName id="4D8E4D6BFFAB194CFAAE3A12FA3CF987" ID-CoL="7F8QB" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[1327,1435,1543,1564]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="zapoteca">
<emphasis id="B8FAEAFAFFAB194CFAAE3A12FA3CF987" box="[1327,1435,1543,1564]" italics="true" pageId="15" pageNumber="16">V. zapoteca</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163A31FA3BF90A" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163A31FA3CF9CD" blockId="15.[151,1436,1366,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163A31FA3CF9CD" pageId="15" pageNumber="16">
20(16) Anterior margin of the carapace distinctly concave (Fig: 8-f); metasomal carinae on segments IV and V densely hirsute (usually more than 5 setae on each carina), ventral carinae on IV and V with 5 or more setae.....................
<taxonomicName id="4D8E4D6BFFAB194CFAA83A54FA3CF9CD" ID-CoL="7F8PQ" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[1321,1435,1600,1622]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="prendinii">
<emphasis id="B8FAEAFAFFAB194CFAA83A54FA3CF9CD" box="[1321,1435,1600,1622]" italics="true" pageId="15" pageNumber="16">V. prendinii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163A4BFA3BF90A" blockId="15.[151,1436,1366,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163A4BFA3BF90A" pageId="15" pageNumber="16">20 Anterior margin of the carapace straight or very slightly concave (Fig: 8-g); metasomal carinae on segments IV and V sparsely hirsute (usually less than 5 setae on each carina), ventral carinae on IV and V with less than 5 setae.................. 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163A8CFA3BF950" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163A8CFA3BF934" blockId="15.[151,1436,1366,2033]" box="[151,1436,1688,1711]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163A8CFA3BF934" box="[151,1436,1688,1711]" pageId="15" pageNumber="16">
21(20) Vesicle ventrodistally with an aggregation of granules resembling a subaculear tubercle (Fig: 9-a)......
<taxonomicName id="4D8E4D6BFFAB194CFB613A8CFAC7F935" ID-CoL="7F8PT" authority="A &amp; Oscar, 2019" authorityName="A &amp; Oscar" authorityYear="2019" box="[1248,1376,1688,1710]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="nanchititla" status="sp. nov.">
<emphasis id="B8FAEAFAFFAB194CFB613A8CFAC7F935" box="[1248,1376,1688,1710]" italics="true" pageId="15" pageNumber="16">V. nanchititla</emphasis>
</taxonomicName>
<emphasis id="B8FAEAFAFFAB194CFAE73A8CFA3BF934" bold="true" box="[1382,1436,1689,1711]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A3C95781FFAB194CFAE73A8CFA3BF934" box="[1382,1436,1689,1711]" pageId="15" pageNumber="16" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163AA3FA3BF950" blockId="15.[151,1436,1366,2033]" box="[151,1436,1718,1739]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163AA3FA3BF950" box="[151,1436,1718,1739]" pageId="15" pageNumber="16">21 Vesicle ventrodistally smooth, without an aggregation of granules (Fig: 9-b).................................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163AC1FA3CF89D" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163AC1FA3BF972" blockId="15.[151,1436,1366,2033]" box="[151,1436,1748,1769]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163AC1FA3BF972" box="[151,1436,1748,1769]" pageId="15" pageNumber="16">22(21) Metasomal segment I distinctly wider than long (L/W &lt;0.7), appearing rectangular in dorsal view................... 23</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163AE4FA3CF89D" blockId="15.[151,1436,1366,2033]" box="[151,1435,1777,1798]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163AE4FA3CF89D" box="[151,1435,1777,1798]" pageId="15" pageNumber="16">22 Metasomal segment I nearly as long as wide, (L/W&gt; 0.7) appearing almost square in dorsal view.................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163B1BFA3CF8DA" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163B1BFA31F8BF" blockId="15.[151,1436,1366,2033]" box="[151,1430,1806,1828]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163B1BFA31F8BF" box="[151,1430,1806,1828]" pageId="15" pageNumber="16">
23(22) Pedipalp chela with dorsal marginal carinae smooth, vestigial, present as an irregular pigmented line............
<taxonomicName id="4D8E4D6BFFAB194CFABF3B1AFA31F8BF" ID-CoL="7F8N9" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[1342,1430,1806,1828]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="dzahui">
<emphasis id="B8FAEAFAFFAB194CFABF3B1AFA31F8BF" box="[1342,1430,1806,1828]" italics="true" pageId="15" pageNumber="16">V. dzahui</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163B39FA3CF8DA" blockId="15.[151,1436,1366,2033]" box="[151,1435,1836,1857]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163B39FA3CF8DA" box="[151,1435,1836,1857]" pageId="15" pageNumber="16">23 Pedipalp chela with dorsal marginal carina distinct, granular................................................... 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163B5CFA3CF8E7" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163B5CFA3CF8C5" blockId="15.[151,1436,1366,2033]" box="[151,1435,1864,1886]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163B5CFA3CF8C5" box="[151,1435,1864,1886]" pageId="15" pageNumber="16">
24(23) Pedipalp femur more than 3 times longer than wide, as wide as deep......................................
<taxonomicName id="4D8E4D6BFFAB194CFAB43B5CFA3CF8C5" ID-CoL="7F8Q2" authorityName="Pocock" authorityYear="1898" box="[1333,1435,1864,1886]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B8FAEAFAFFAB194CFAB43B5CFA3CF8C5" box="[1333,1435,1864,1886]" italics="true" pageId="15" pageNumber="16">V. pusillus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163B73FA3CF8E7" blockId="15.[151,1436,1366,2033]" box="[151,1435,1894,1916]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163B73FA3CF8E7" box="[151,1435,1894,1916]" pageId="15" pageNumber="16">
24 Pedipalp femur less than 3 times longer than wide, and almost one and a half times wider than deep............
<taxonomicName id="4D8E4D6BFFAB194CFAAF3B72FA3CF8E7" ID-CoL="7F92F" authorityName="Zarate-Galvez &amp; Francke" authorityYear="2009" box="[1326,1435,1895,1916]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="trespicos">
<emphasis id="B8FAEAFAFFAB194CFAAF3B72FA3CF8E7" box="[1326,1435,1895,1916]" italics="true" pageId="15" pageNumber="16">V. trespicos</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163B91FA3CF82D" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163B91FA3CF802" blockId="15.[151,1436,1366,2033]" box="[151,1435,1924,1945]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163B91FA3CF802" box="[151,1435,1924,1945]" pageId="15" pageNumber="16">25(22) Caudal gland circular................................................................................. 26</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163BB4FA3CF82D" blockId="15.[151,1436,1366,2033]" box="[151,1435,1953,1974]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163BB4FA3CF82D" box="[151,1435,1953,1974]" pageId="15" pageNumber="16">25 Caudal gland linear.................................................................................. 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="317A2E4DFFAB194CFF163BABFA3CF86A" pageId="15" pageNumber="16">
<paragraph id="8A3136E8FFAB194CFF163BABFA3EF84F" blockId="15.[151,1436,1366,2033]" box="[151,1433,1982,2004]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163BABFA3EF84F" box="[151,1433,1982,2004]" pageId="15" pageNumber="16">
26(25) Pectinal tooth counts 20 or more (usually 2022); vesicle elongated (L/W ḵ1.7).............................
<taxonomicName id="4D8E4D6BFFAB194CFACB3BAAFA3EF84F" ID-CoL="7F8PX" authorityName=", Pocock" authorityYear="1902" box="[1354,1433,1982,2004]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="B8FAEAFAFFAB194CFACB3BAAFA3EF84F" box="[1354,1433,1982,2004]" italics="true" pageId="15" pageNumber="16">V. smithi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8A3136E8FFAB194CFF163BC9FA3CF86A" blockId="15.[151,1436,1366,2033]" box="[151,1435,2012,2033]" pageId="15" pageNumber="16">
<keyLead id="317F95DDFFAB194CFF163BC9FA3CF86A" box="[151,1435,2012,2033]" pageId="15" pageNumber="16">
26 Pectinal tooth counts less than 20; vesicle globose (L/W
<geoCoordinate id="EFBA502FFFAB194CFC803BC9FC92F86A" box="[769,821,2012,2033]" direction="east" orientation="longitude" pageId="15" pageNumber="16" precision="5555" value="1.7">Ẽ 1.7</geoCoordinate>
)........................................
<taxonomicName id="4D8E4D6BFFAB194CFA9E3BC9FA3CF86A" ID-CoL="7F8PV" authorityName="C. L. Koch" authorityYear="1836" box="[1311,1435,2012,2033]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B8FAEAFAFFAB194CFA9E3BC9FA3CF86A" box="[1311,1435,2012,2033]" italics="true" pageId="15" pageNumber="16">V. mexicanus</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<caption id="DEF16660FFB41953FF163B50FB4CF84D" httpUri="https://zenodo.org/record/2659109/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1861,1883]" targetBox="[493,1093,193,1835]" targetPageId="16">
<paragraph id="8A3136E8FFB41953FF163B50FB4CF84D" blockId="16.[151,1436,1861,2006]" pageId="16" pageNumber="17">
<emphasis id="B8FAEAFAFFB41953FF163B50FEBFF8C0" bold="true" box="[151,280,1861,1883]" pageId="16" pageNumber="17">FIGURE 8.</emphasis>
Characters to differentiate males of species within the “
<taxonomicName id="4D8E4D6BFFB41953FC073B52FC56F8C7" ID-CoL="7F8PV" authorityName="C. L. Koch" authorityYear="1836" box="[902,1009,1863,1884]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B8FAEAFAFFB41953FC073B52FC56F8C7" box="[902,1009,1863,1884]" italics="true" pageId="16" pageNumber="17">mexicanus</emphasis>
</taxonomicName>
” group:
<emphasis id="B8FAEAFAFFB41953FBD73B50FBDCF8C0" bold="true" box="[1110,1147,1861,1883]" pageId="16" pageNumber="17">a—</emphasis>
Vesicle, showing a distinct dorsoventrally flattened shape (in relation on two parallel red lines);
<emphasis id="B8FAEAFAFFB41953FCDD3B71FC24F8E1" bold="true" box="[860,899,1892,1914]" pageId="16" pageNumber="17">b—</emphasis>
Pedipalp chela, red arrow signaling the notch on the fingers;
<emphasis id="B8FAEAFAFFB41953FF6A3B96FEA8F802" bold="true" box="[235,271,1923,1945]" pageId="16" pageNumber="17">c—</emphasis>
Vesicle, showing an oval shape (in relation to two parallel red lines);
<emphasis id="B8FAEAFAFFB41953FC503B96FC5FF802" bold="true" box="[977,1016,1923,1945]" pageId="16" pageNumber="17">d—</emphasis>
Pedipalp chela fingers straight, red arrow signaling the lack of a notch;
<emphasis id="B8FAEAFAFFB41953FE4C3BB4FE56F82C" bold="true" box="[461,497,1953,1975]" pageId="16" pageNumber="17">e—</emphasis>
Metasomal carinae showing the merging of the granules (Red arrow signaling each carina;
<emphasis id="B8FAEAFAFFB41953FF163BD5FF1EF84D" bold="true" box="[151,185,1984,2006]" pageId="16" pageNumber="17">f—</emphasis>
Anterior margin of the carapace concave;
<emphasis id="B8FAEAFAFFB41953FDE33BD5FD2FF84D" bold="true" box="[610,648,1984,2006]" pageId="16" pageNumber="17">g—</emphasis>
Anterior margin of the carapace straight or slightly concave.
</paragraph>
</caption>
<caption id="DEF16660FFB51952FF16386BFBDCFB28" httpUri="https://zenodo.org/record/2659111/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1150,1172]" targetBox="[508,1075,198,1127]" targetPageId="17">
<paragraph id="8A3136E8FFB51952FF16386BFBDCFB28" blockId="17.[151,1435,1150,1203]" pageId="17" pageNumber="18">
<emphasis id="B8FAEAFAFFB51952FF16386BFEB3FB0F" bold="true" box="[151,276,1150,1172]" pageId="17" pageNumber="18">FIGURE 9.</emphasis>
Characters to differentiate males of species within the “
<taxonomicName id="4D8E4D6BFFB51952FCDF3895FC6EFB0E" ID-CoL="7F8PV" authorityName="C. L. Koch" authorityYear="1836" box="[862,969,1152,1173]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="mexicanus">
<emphasis id="B8FAEAFAFFB51952FCDF3895FC6EFB0E" box="[862,969,1152,1173]" italics="true" pageId="17" pageNumber="18">mexicanus</emphasis>
</taxonomicName>
” group:
<emphasis id="B8FAEAFAFFB51952FBA6386BFB94FB0F" bold="true" box="[1063,1075,1150,1172]" pageId="17" pageNumber="18">a</emphasis>
—Vesicle smooth, with aggregated granules resembling a subaculear tubercle;
<emphasis id="B8FAEAFAFFB51952FDCD3888FDD4FB28" bold="true" box="[588,627,1181,1203]" pageId="17" pageNumber="18">b—</emphasis>
Vesicle granular, but without a subaculear tubercle.
</paragraph>
</caption>
<paragraph id="8A3136E8FFB51952FF1638ECFA32FA3A" blockId="17.[151,1436,1272,1441]" pageId="17" pageNumber="18">
<keyLead id="317F95DDFFB51952FF1638ECFA32FA3A" pageId="17" pageNumber="18">
<keyStep id="317A2E4DFFB51952FF1638ECFA32FA3A" pageId="17" pageNumber="18">
27(25) Pedipalp chela smooth, without granular carinae...................................................
<taxonomicName id="4D8E4D6BFFB51952FA9E38ECFA3CFA95" ID-CoL="7F8PJ" authorityName=", Sissom" authorityYear="1989" box="[1311,1435,1272,1294]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="maculosus">
<emphasis id="B8FAEAFAFFB51952FA9E38ECFA3CFA95" box="[1311,1435,1272,1294]" italics="true" pageId="17" pageNumber="18">V. maculosus</emphasis>
</taxonomicName>
27 Pedipalp chela with at least one granular carina............................................................. 28 28(27) Pedipalp femur two times wider than deep; vesicle more than two times longer than wide....................
<taxonomicName id="4D8E4D6BFFB51952FABB3921FA3BFAD2" ID-CoL="7F926" authorityName="Miranda-Lopez, Ponce-Saavedra and Francke" authorityYear="2012" box="[1338,1436,1331,1353]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="morelia">
<emphasis id="B8FAEAFAFFB51952FABB3921FA3BFAD2" box="[1338,1436,1331,1353]" italics="true" pageId="17" pageNumber="18">V. morelia</emphasis>
</taxonomicName>
28 Pedipalp femur never more than one and a half times wider than deep; vesicle less than two times longer than wide...... 29 29(28) Sternite VII with 1112 setae; chela with dorsal carinae moderate to strong, granulose......................
<taxonomicName id="4D8E4D6BFFB51952FABB397BFA3EFA18" ID-CoL="7F8MM" authorityName="Santibanez-Lopez &amp; Francke" authorityYear="2010" box="[1338,1433,1389,1411]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="darwini">
<emphasis id="B8FAEAFAFFB51952FABB397BFA3EFA18" box="[1338,1433,1389,1411]" italics="true" pageId="17" pageNumber="18">V. darwini</emphasis>
</taxonomicName>
29 Sternite VII with 15 or more setae; chela with dorsal carina weak, granulose..............................
<taxonomicName id="4D8E4D6BFFB51952FAB83999FA32FA3A" ID-CoL="7F8PZ" authorityName="Sissom" authorityYear="1989" box="[1337,1429,1420,1441]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="B8FAEAFAFFB51952FAB83999FA32FA3A" box="[1337,1429,1420,1441]" italics="true" pageId="17" pageNumber="18">V. setosus</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
</document>