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<mods:title id="F977E49A205E7026680F2BEC8BC1B302">The genus Hoplandrothrips and its relatives (Thysanoptera: Phlaeothripidae) from Southeast Asia and Taiwan</mods:title>
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<mods:namePart id="8472071FC8ED1FE3317120E13441B8A9">Okajima, Shûji</mods:namePart>
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<mods:affiliation id="D9B6EC3E7EAE60CDD8E19C261C966D08">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan.</mods:affiliation>
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<mods:namePart id="01A03264B4286CD01DB5735805A8C07B">Masumoto, Masami</mods:namePart>
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<mods:affiliation id="8463A52B7D2442B1769985E18B7F8965">Laboratory of Entomology, Tokyo University of Agriculture, 1737 Funako, Atsugi, Kanagawa, 243 - 0034 Japan.</mods:affiliation>
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<subSubSection id="C361DAA6850EC334D9B2A647AA16766C" box="[151,608,1812,1839]" pageId="6" pageNumber="28" type="nomenclature">
<paragraph id="8BC4892D850EC334D9B2A647AA16766C" blockId="6.[151,608,1812,1875]" box="[151,608,1812,1839]" pageId="6" pageNumber="28">
<heading id="D08C3E41850EC334D9B2A647AA16766C" bold="true" box="[151,608,1812,1839]" fontSize="11" level="1" pageId="6" pageNumber="28" reason="1">
<taxonomicName id="4C7BF2AE850EC334D9B2A647AA16766C" ID-CoL="38DHT" authority="Okajima" authorityName="Okajima" box="[151,608,1812,1839]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="28" phylum="Arthropoda" rank="species" species="claricornis">
<emphasis id="B90F553F850EC334D9B2A647AA16766C" bold="true" box="[151,608,1812,1839]" pageId="6" pageNumber="28">
<emphasis id="B90F553F850EC334D9B2A647A99F766D" bold="true" box="[151,489,1812,1839]" italics="true" pageId="6" pageNumber="28">Ecacanthothrips claricornis</emphasis>
Okajima
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361DAA6850EC334D9B2A66BA9297611" box="[151,351,1848,1875]" pageId="6" pageNumber="28" type="description">
<paragraph id="8BC4892D850EC334D9B2A66BA9297611" blockId="6.[151,608,1812,1875]" box="[151,351,1848,1875]" pageId="6" pageNumber="28">
(
<figureCitation id="134095A8850EC334D9BAA66BA9727611" box="[159,260,1848,1875]" captionStart="FIGURES 114" captionStartId="37.[151,264,1867,1891]" captionTargetBox="[158,1430,181,1844]" captionTargetId="figure-32@37.[158,1430,181,1844]" captionTargetPageId="37" captionText="FIGURES 114. Ecacanthothrips species. E. andrei 12: (1) female; (2) large male. E. brevicornis sp. n. 34: (3) female; (4) male. E. claricornis 56: (5) female; (6) large male. E. coniger 78: (7) female; (8) large male. E. leai 911: (9) female; (10) large male; (11) small male. E. moundi 1213: (12) fmale; (13) large male. E. nigellus: (14) female." figureDoi="http://doi.org/10.5281/zenodo.13211345" httpUri="https://zenodo.org/record/13211345/files/figure.png" pageId="6" pageNumber="28">Figs 56</figureCitation>
,
<figureCitation id="134095A8850EC334D835A66BA9217611" box="[272,343,1848,1875]" captionStart="FIGURES 7583" captionStartId="43.[151,264,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-32@43.[173,1414,181,1919]" captionTargetPageId="43" captionText="FIGURES 7583. Ecacanthothrips claricornis. (75) fore leg, female; (76) head and prothorax, female; (77) head and prothorax, large male; (78) antenna, female; (79) antennal segments IIIV, female; (80) metanotum and pelta, female; (81) prosternum and mesopresternum, female; (82) hind femur, female; (83) tube, female." figureDoi="http://doi.org/10.5281/zenodo.13211363" httpUri="https://zenodo.org/record/13211363/files/figure.png" pageId="6" pageNumber="28">7583</figureCitation>
)
</paragraph>
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<subSubSection id="C361DAA6850EC335D9B2A62AA9197093" lastPageId="7" lastPageNumber="29" pageId="6" pageNumber="28" type="reference_group">
<paragraph id="8BC4892D850EC334D9B2A62AAAD376D3" blockId="6.[151,677,1913,1937]" box="[151,677,1913,1937]" pageId="6" pageNumber="28">
<treatmentCitationGroup id="AB6BAE03850EC334D9B2A62AAAD376D3" box="[151,677,1913,1937]" pageId="6" pageNumber="28">
<treatmentCitation id="0ADAAF3C850EC334D9B2A62AAAD776D3" author="Okajima, S." box="[151,673,1913,1937]" page="59" pageId="6" pageNumber="28" year="1983">
<taxonomicName id="4C7BF2AE850EC334D9B2A62AAAD776D3" ID-CoL="38DHT" authority="Okajima, 1983: 57 - 59" authorityName="Okajima" authorityPageNumber="57 - 59" authorityYear="1983" box="[151,673,1913,1937]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="28" phylum="Arthropoda" rank="species" species="claricornis">
<emphasis id="B90F553F850EC334D9B2A62AA9C276D3" box="[151,436,1913,1937]" italics="true" pageId="6" pageNumber="28">Ecacanthothrips claricornis</emphasis>
<bibRefCitation id="EFEAF4DC850EC334D89FA62AAAD776D3" author="Okajima, S." box="[442,673,1913,1937]" pageId="6" pageNumber="28" pagination="56 - 65" refId="ref50428" refString="Okajima, S. (1983) Four new species of Ecacanthothrips from the Oriental region (Thysanoptera, Phlaeothripidae). Kontyu, 51 (1), 56 - 65." type="journal article" year="1983">Okajima, 1983: 5759</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC4892D850EC335D9B2A6EFA9197093" blockId="6.[151,1437,1979,2042]" lastBlockId="7.[151,1437,151,645]" lastPageId="7" lastPageNumber="29" pageId="6" pageNumber="28">
<taxonomicName id="4C7BF2AE850EC334D9B2A6EFA9437697" ID-CoL="38DHT" box="[151,309,1979,2005]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="28" phylum="Arthropoda" rank="species" species="claricornis">
<emphasis id="B90F553F850EC334D9B2A6EFA9437697" box="[151,309,1979,2005]" italics="true" pageId="6" pageNumber="28">E. claricornis</emphasis>
</taxonomicName>
was described based on
<specimenCount id="9D7D42A4850EC334DB78A6E8AAAD7697" box="[605,731,1979,2005]" count="17" pageId="6" pageNumber="28" type="female">17 females</specimenCount>
and
<specimenCount id="9D7D42A4850EC334DA3CA6E8ABF37697" box="[793,901,1979,2005]" count="6" pageId="6" pageNumber="28" type="male">six males</specimenCount>
collected from Mindanao Is., the
<collectingCountry id="F36CC9BD850EC334DC32A6E8ADEE7697" box="[1303,1432,1979,2005]" name="Philippines" pageId="6" pageNumber="28">Philippines</collectingCountry>
, together with non-paratypic
<specimenCount id="9D7D42A4850EC334D8F3A68CAA1A76BB" box="[470,620,2015,2041]" count="3" pageId="6" pageNumber="28" type="female">three females</specimenCount>
and
<specimenCount id="9D7D42A4850EC334DB86A68CAB6076BB" box="[675,790,2015,2041]" count="5" pageId="6" pageNumber="28" type="male">five males</specimenCount>
from Luzon Is., and is included in the
<taxonomicName id="4C7BF2AE850EC334DDEEA68CAD6F76BB" ID-CoL="6MTC6" authorityName="Ashmead" authorityYear="1905" box="[1227,1305,2015,2041]" class="Insecta" family="Phlaeothripidae" genus="Idolothrips" kingdom="Animalia" order="Thysanoptera" pageId="6" pageNumber="28" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B90F553F850EC334DDEEA68CAD6F76BB" box="[1227,1305,2015,2041]" italics="true" pageId="6" pageNumber="28">tibialis</emphasis>
</taxonomicName>
-group. It is newly recorded here from Central and
<collectingRegion id="49BF47CF850FC335DB66A1C4AA8571F3" box="[579,755,151,177]" country="Indonesia" name="Sulawesi Selatan" pageId="7" pageNumber="29">South Sulawesi</collectingRegion>
,
<collectingCountry id="F36CC9BD850FC335DBD8A1C4AB1B71F3" box="[765,877,151,177]" name="Indonesia" pageId="7" pageNumber="29">Indonesia</collectingCountry>
, based on a long series of females and males listed below. This species also shows remarkable size related structural variations associated with allometric growth as in
<taxonomicName id="4C7BF2AE850FC335D9B2A18CA89371BB" ID-CoL="6MTC6" authorityName="Ashmead" authorityYear="1905" box="[151,229,223,249]" class="Insecta" family="Phlaeothripidae" genus="Idolothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B90F553F850FC335D9B2A18CA89371BB" box="[151,229,223,249]" italics="true" pageId="7" pageNumber="29">tibialis</emphasis>
</taxonomicName>
. Although the specimens from Mindanao, the
<typeStatus id="54C0378F850FC335DBCBA18CAB6871BB" box="[750,798,223,249]" pageId="7" pageNumber="29">type</typeStatus>
series, have the femora fully brown, the specimens from both Luzon and
<collectingRegion id="49BF47CF850FC335D876A050A9CE705F" box="[339,440,259,285]" country="Indonesia" name="Sulawesi Utara" pageId="7" pageNumber="29">Sulawesi</collectingRegion>
have the femora largely brown but with apices clear yellow. Moreover, the size related variation of
<taxonomicName id="4C7BF2AE850FC335D80DA074A9D57003" ID-CoL="38DHT" box="[296,419,295,321]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="claricornis">
<emphasis id="B90F553F850FC335D80DA074A9D57003" box="[296,419,295,321]" italics="true" pageId="7" pageNumber="29">claricornis</emphasis>
</taxonomicName>
is more similar to
<taxonomicName id="4C7BF2AE850FC335DBAFA074AAA47003" ID-CoL="38DHQ" box="[650,722,295,321]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B90F553F850FC335DBAFA074AAA47003" box="[650,722,295,321]" italics="true" pageId="7" pageNumber="29">andrei</emphasis>
</taxonomicName>
rather than
<taxonomicName id="4C7BF2AE850FC335DA40A074ABC57003" ID-CoL="6MTC6" authorityName="Ashmead" authorityYear="1905" box="[869,947,295,321]" class="Insecta" family="Phlaeothripidae" genus="Idolothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="B90F553F850FC335DA40A074ABC57003" box="[869,947,295,321]" italics="true" pageId="7" pageNumber="29">tibialis</emphasis>
</taxonomicName>
, and the prothoracic anteromarginal setae elongate, the fore tibia with no sub-apical inner tubercle even in large males, and
<taxonomicName id="4C7BF2AE850FC335DD05A018ACED7027" ID-CoL="38DHT" box="[1056,1179,331,357]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="claricornis">
<emphasis id="B90F553F850FC335DD05A018ACED7027" box="[1056,1179,331,357]" italics="true" pageId="7" pageNumber="29">claricornis</emphasis>
</taxonomicName>
has only one stout seta with blunt or dilated apex on each mid and hind femur in both sexes (
<figureCitation id="134095A8850FC335DA9DA03CAC7870CB" box="[952,1038,367,393]" captionStart="FIGURES 7583" captionStartId="43.[151,264,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-32@43.[173,1414,181,1919]" captionTargetPageId="43" captionText="FIGURES 7583. Ecacanthothrips claricornis. (75) fore leg, female; (76) head and prothorax, female; (77) head and prothorax, large male; (78) antenna, female; (79) antennal segments IIIV, female; (80) metanotum and pelta, female; (81) prosternum and mesopresternum, female; (82) hind femur, female; (83) tube, female." figureDoi="http://doi.org/10.5281/zenodo.13211363" httpUri="https://zenodo.org/record/13211363/files/figure.png" pageId="7" pageNumber="29">Fig. 82</figureCitation>
). In contrast to
<taxonomicName id="4C7BF2AE850FC335DDE6A03CAD7D70CB" ID-CoL="38DHQ" box="[1219,1291,367,393]" class="Insecta" family="Phlaeothripidae" genus="Ecacanthothrips" kingdom="Animalia" order="Thysanoptera" pageId="7" pageNumber="29" phylum="Arthropoda" rank="species" species="andrei">
<emphasis id="B90F553F850FC335DDE6A03CAD7D70CB" box="[1219,1291,367,393]" italics="true" pageId="7" pageNumber="29">andrei</emphasis>
</taxonomicName>
, this species has somewhat enlarged antennal segment III (
<figureCitation id="134095A8850FC335DBBEA0C0AB4370EF" box="[667,821,403,429]" captionStart="FIGURES 7583" captionStartId="43.[151,264,1943,1967]" captionTargetBox="[173,1414,181,1919]" captionTargetId="figure-32@43.[173,1414,181,1919]" captionTargetPageId="43" captionText="FIGURES 7583. Ecacanthothrips claricornis. (75) fore leg, female; (76) head and prothorax, female; (77) head and prothorax, large male; (78) antenna, female; (79) antennal segments IIIV, female; (80) metanotum and pelta, female; (81) prosternum and mesopresternum, female; (82) hind femur, female; (83) tube, female." figureDoi="http://doi.org/10.5281/zenodo.13211363" httpUri="https://zenodo.org/record/13211363/files/figure.png" pageId="7" pageNumber="29">Figs 78 &amp; 79</figureCitation>
), that is distinctly wider than segment IV, with usually 1015 sense cones.
</paragraph>
</subSubSection>
<subSubSection id="C361DAA6850FC335D9E2A088AB6673C7" pageId="7" pageNumber="29" type="description">
<paragraph id="8BC4892D850FC335D9E2A088AB6673C7" blockId="7.[151,1437,151,645]" pageId="7" pageNumber="29">
<emphasis id="B90F553F850FC335D9E2A088A9CF70B7" box="[199,441,475,501]" italics="true" pageId="7" pageNumber="29">Specimens examined.</emphasis>
<emphasis id="B90F553F850FC335D8E5A088A99970B7" bold="true" box="[448,495,475,501]" pageId="7" pageNumber="29">The</emphasis>
<materialsCitation id="3B138370850FC335D8D2A089ADEA70B7" box="[503,1436,474,501]" collectorName="Mindanao Is. &amp; Luzon Is." country="Philippines" location="Mindanao Is" pageId="7" pageNumber="29" specimenCount="1" specimenCount-female="3" specimenCount-male="5" typeStatus="holotype">
<emphasis id="B90F553F850FC335D8D2A089AA0B70B6" bold="true" box="[503,637,474,500]" pageId="7" pageNumber="29">
<collectingCountry id="F36CC9BD850FC335D8D2A089AA0B70B6" box="[503,637,474,500]" name="Philippines" pageId="7" pageNumber="29">Philippines</collectingCountry>
</emphasis>
, holotype female and paratype females and males from
<location id="8EA4DFF6850FC335DDDAA088ADE470B7" LSID="urn:lsid:plazi:treatment:03D2383B850EC335D9B2A647AB6673C7:8EA4DFF6850FC335DDDAA088ADE470B7" box="[1279,1426,475,501]" country="Philippines" name="Mindanao Is" pageId="7" pageNumber="29">Mindanao Is</location>
.;
</materialsCitation>
<materialsCitation id="3B138370850FC335D9B2A0ACABA6735B" box="[151,976,510,537]" collectorName="Mindanao Is. &amp; Luzon Is." country="Philippines" location="Luzon Is." pageId="7" pageNumber="29" specimenCount="8" specimenCount-female="3" specimenCount-male="5" typeStatus="holotype">
<specimenCount id="9D7D42A4850FC335D9B2A0ACA975735B" box="[151,259,511,537]" count="3" pageId="7" pageNumber="29" type="female">3 females</specimenCount>
and
<specimenCount id="9D7D42A4850FC335D81FA0ADA9E2735B" box="[314,404,510,537]" count="5" pageId="7" pageNumber="29" type="male">5 males</specimenCount>
, from
<location id="8EA4DFF6850FC335D8F8A0ACAA31735B" LSID="urn:lsid:plazi:treatment:03D2383B850EC335D9B2A647AB6673C7:8EA4DFF6850FC335D8F8A0ACAA31735B" box="[477,583,511,537]" country="Philippines" name="Luzon Is." pageId="7" pageNumber="29">Luzon Is.</location>
(data described in
<bibRefCitation id="EFEAF4DC850FC335DA38A0ACABB1735B" author="Okajima, S." box="[797,967,511,537]" pageId="7" pageNumber="29" pagination="56 - 65" refId="ref50428" refString="Okajima, S. (1983) Four new species of Ecacanthothrips from the Oriental region (Thysanoptera, Phlaeothripidae). Kontyu, 51 (1), 56 - 65." type="journal article" year="1983">Okajima, 1983</bibRefCitation>
)
</materialsCitation>
.
<materialsCitation id="3B138370850FC335DAFEA0ADAA7F7323" collectingDate="1984-07-31" collectingDateMax="1984-08-03" collectingDateMin="1984-07-31" country="Indonesia" elevation="900" location="Malino" municipality="South" pageId="7" pageNumber="29" specimenCount="14" specimenCount-female="6" specimenCount-male="8" stateProvince="South Celebes" typeStatus="holotype">
<emphasis id="B90F553F850FC335DAFEA0ADAC26735A" bold="true" box="[987,1104,510,536]" pageId="7" pageNumber="29">
<collectingCountry id="F36CC9BD850FC335DAFEA0ADAC26735A" box="[987,1104,510,536]" name="Indonesia" pageId="7" pageNumber="29">Indonesia</collectingCountry>
</emphasis>
,
<collectingRegion id="49BF47CF850FC335DD7BA0ACAD76735B" box="[1118,1280,511,537]" pageId="7" pageNumber="29">South Celebes</collectingRegion>
(= Sulawesi),
<location id="8EA4DFF6850FC335D9B2A370A89D737F" LSID="urn:lsid:plazi:treatment:03D2383B850EC335D9B2A647AB6673C7:8EA4DFF6850FC335D9B2A370A89D737F" box="[151,235,547,573]" country="Indonesia" municipality="South" name="Malino" pageId="7" pageNumber="29" stateProvince="South Celebes">Malino</location>
, alt. about
<quantity id="4C8324C8850FC335D842A370A9DF737F" box="[359,425,547,573]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="29" unit="m" value="900.0">
<elevation id="00566E1E850FC335D842A370A9DF737F" box="[359,425,547,573]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="29" unit="m" value="900.0">900m</elevation>
</quantity>
,
<specimenCount id="9D7D42A4850FC335D890A370AA57737F" box="[437,545,547,573]" count="4" pageId="7" pageNumber="29" type="female">4 females</specimenCount>
and
<specimenCount id="9D7D42A4850FC335DB7CA371AAC4737F" box="[601,690,546,573]" count="5" pageId="7" pageNumber="29" type="male">5 males</specimenCount>
, on dead leaves and branches,
<date id="FFC5AFED850FC335DD28A370ACF9737F" box="[1037,1167,547,573]" pageId="7" pageNumber="29" value="1984-07-31">
<collectingDate id="EF815605850FC335DD28A370ACF9737F" box="[1037,1167,547,573]" pageId="7" pageNumber="29" value="1984-07-31">31.vii.1984</collectingDate>
</date>
,
<specimenCount id="9D7D42A4850FC335DDBEA370AD71737F" box="[1179,1287,547,573]" count="2" pageId="7" pageNumber="29" type="female">2 females</specimenCount>
and
<specimenCount id="9D7D42A4850FC335DC1BA370ADEE737F" box="[1342,1432,547,573]" count="3" pageId="7" pageNumber="29" type="male">3 males</specimenCount>
, on dead Palmae,
<date id="FFC5AFED850FC335D873A314A9A47323" box="[342,466,583,609]" pageId="7" pageNumber="29" value="1984-08-03">
<collectingDate id="EF815605850FC335D873A314A9A47323" box="[342,466,583,609]" pageId="7" pageNumber="29" value="1984-08-03">3.viii.1984</collectingDate>
</date>
, SO;
</materialsCitation>
<materialsCitation id="3B138370850FC335DB2AA314AB7A73C7" collectingDate="1984-08-08" collectingDateMax="1984-08-14" collectingDateMin="1984-08-08" country="Indonesia" elevation="1000" location="Pedamaran" municipality="South" pageId="7" pageNumber="29" specimenCount="107" specimenCount-female="62" specimenCount-male="45" stateProvince="Central Celebes" typeStatus="holotype">
<collectingRegion id="49BF47CF850FC335DB2AA314AAB57323" box="[527,707,583,609]" pageId="7" pageNumber="29">Central Celebes</collectingRegion>
,
<locationDeviation id="9A76900F850FC335DBEBA314AB047323" box="[718,882,583,609]" pageId="7" pageNumber="29">near Rantepao</locationDeviation>
,
<location id="8EA4DFF6850FC335DA58A314AB8A7323" LSID="urn:lsid:plazi:treatment:03D2383B850EC335D9B2A647AB6673C7:8EA4DFF6850FC335DA58A314AB8A7323" box="[893,1020,583,609]" country="Indonesia" municipality="South" name="Pedamaran" pageId="7" pageNumber="29" stateProvince="Central Celebes">Pedamaran</location>
, alt. about
<quantity id="4C8324C8850FC335DD50A314ACB37323" box="[1141,1221,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="29" unit="m" value="1000.0">
<elevation id="00566E1E850FC335DD50A314ACB37323" box="[1141,1221,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="29" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<specimenCount id="9D7D42A4850FC335DDF4A314AD3C7323" box="[1233,1354,583,609]" count="62" pageId="7" pageNumber="29" type="female">62 females</specimenCount>
and
<specimenCount id="9D7D42A4850FC335DCA5A315A8AD73C7" count="45" pageId="7" pageNumber="29" type="male">45 males</specimenCount>
, on dead leaves and branches,
<date id="FFC5AFED850FC335DB16A338AAAF73C7" box="[563,729,619,645]" pageId="7" pageNumber="29" value="1984-08-08" valueMax="1984-08-14" valueMin="1984-08-08">
<collectingDate id="EF815605850FC335DB16A338AAAF73C7" box="[563,729,619,645]" pageId="7" pageNumber="29" value="1984-08-08" valueMax="1984-08-14" valueMin="1984-08-08">814.viii.1984</collectingDate>
</date>
, SO
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>