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<document id="2AAF7A74043100D32012CE4F322673C5" ID-CLB-Dataset="24286" ID-DOI="10.11646/zootaxa.4728.2.2" ID-GBIF-Dataset="a1c40b73-0c11-4863-a188-412c33523e02" ID-ISSN="1175-5326" ID-Zenodo-Dep="3621117" ID-ZooBank="AE4F8089-7D2E-4ABA-AA09-CD504705788A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579677041620" checkinUser="plazi" docAuthor="Wagner, Rüdiger &amp; Withers, Phil" docDate="2020" docId="784D878B601A21038FF1034CFE508821" docLanguage="en" docName="zootaxa.4728.2.2.pdf" docOrigin="Zootaxa 4728 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonnoiriella arcuata Jezek 1997" docType="treatment" docVersion="7" lastPageNumber="205" masterDocId="8474FFF3600C21158F66006DFFB28F20" masterDocTitle="The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)" masterLastPageNumber="210" masterPageNumber="183" pageNumber="205" updateTime="1698778572713" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F36C87F6976BB4F3876521BFB447FE9E">The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae)</mods:title>
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<treatment id="784D878B601A21038FF1034CFE508821" ID-DOI="http://doi.org/10.5281/zenodo.5918615" ID-GBIF-Taxon="161827862" ID-Zenodo-Dep="5918615" LSID="urn:lsid:plazi:treatment:784D878B601A21038FF1034CFE508821" httpUri="http://treatment.plazi.org/id/784D878B601A21038FF1034CFE508821" lastPageNumber="205" pageId="22" pageNumber="205">
<subSubSection id="B8FE6516601A21038FF1034CFD958C1B" box="[151,551,801,828]" pageId="22" pageNumber="205" type="nomenclature">
<paragraph id="F05B369D601A21038FF1034CFD958C1B" blockId="22.[151,551,801,863]" box="[151,551,801,828]" pageId="22" pageNumber="205">
<heading id="AB1381F1601A21038FF1034CFD958C1B" bold="true" box="[151,551,801,828]" fontSize="11" level="1" pageId="22" pageNumber="205" reason="1">
<taxonomicName id="37E44D1E601A21038FF1034CFD958C1B" ID-CoL="7CHTC" authority="Jezek, 1997" authorityName="Jezek" authorityYear="1997" box="[151,551,801,828]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="205" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="C290EA8F601A21038FF1034CFD958C1B" bold="true" box="[151,551,801,828]" pageId="22" pageNumber="205">
<emphasis id="C290EA8F601A21038FF1034CFE228C1B" bold="true" box="[151,400,801,828]" italics="true" pageId="22" pageNumber="205">Tonnoiriella arcuata</emphasis>
<bibRefCitation id="94754B6C601A21038EFE034CFD958C1B" author="Jezek, J." box="[408,551,801,827]" pageId="22" pageNumber="205" pagination="105 - 122" refId="ref20050" refString="Jezek, J. (1997) New and interesting taxa of moth flies (Diptera, Psychodidae) from different moist biotopes of the Palaearctic Region. Casopis Narodniho muzea Rada prirodovedna, 166 (1 - 4), 105 - 122." type="journal article" year="1997">Ježek, 1997</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="B8FE6516601A21038FF10328FE958C7F" box="[151,295,837,863]" pageId="22" pageNumber="205" type="description">
<paragraph id="F05B369D601A21038FF10328FE958C7F" blockId="22.[151,551,801,863]" box="[151,295,837,863]" pageId="22" pageNumber="205">
(
<figureCitation id="68DF2A18601A21038FF90328FEAD8C7F" box="[159,287,837,863]" captionStart="FIGURES 4349" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 4349: 4345 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 4849 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="22" pageNumber="205">Figs 48, 49</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601A21038FF103E4FB348C81" box="[151,1158,904,929]" pageId="22" pageNumber="205" type="reference_group">
<paragraph id="F05B369D601A21038FF103E4FB348C81" blockId="22.[151,1158,904,929]" box="[151,1158,904,929]" pageId="22" pageNumber="205">
<taxonomicName id="37E44D1E601A21038FF103E4FE5B8C81" ID-CoL="7CHTC" authority="Jezek, 1997" authorityName="Jezek" authorityYear="1997" box="[151,489,904,929]" class="Insecta" family="Psychodidae" genus="Tonnoiriella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="205" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="C290EA8F601A21038FF103E4FEDA8C81" box="[151,360,905,929]" italics="true" pageId="22" pageNumber="205">Tonnoiriella arcuata</emphasis>
<bibRefCitation id="94754B6C601A21038E0803E5FE5B8C81" author="Jezek, J." box="[366,489,904,929]" pageId="22" pageNumber="205" pagination="105 - 122" refId="ref20050" refString="Jezek, J. (1997) New and interesting taxa of moth flies (Diptera, Psychodidae) from different moist biotopes of the Palaearctic Region. Casopis Narodniho muzea Rada prirodovedna, 166 (1 - 4), 105 - 122." type="journal article" year="1997">Ježek, 1997</bibRefCitation>
</taxonomicName>
: Časopis Národniho Muzea v Praze Řada přirodovedna, 166: 1 16.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601A21038FF103A2FEED8BBE" pageId="22" pageNumber="205" type="materials_examined">
<paragraph id="F05B369D601A21038FF103A2FEED8BBE" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
<emphasis id="C290EA8F601A21038FF103A2FF4D8CCA" bold="true" box="[151,255,975,1002]" pageId="22" pageNumber="205">Material</emphasis>
:
<specimenCount id="E6E2FD14601A21038E6903BDFE808CCA" box="[271,306,976,1002]" pageId="22" pageNumber="205" type="male">1♂</specimenCount>
<typeStatus id="2F5F883F601A21038E5F03A2FE2C8CC9" box="[313,414,975,1001]" pageId="22" pageNumber="205" type="holotype">holotype</typeStatus>
, slide mounted: head, mothparts with 2 palpi, 2 antennae, thorax cut in half, 2 wings, legs, abdomen part, 1 stylus, aedeagus, epandrium with epandrial processes; left side white label: Caucasus, Pskhu,
<date id="845A105D601A21038FF1047AFEBA8B11" box="[151,264,1047,1073]" pageId="22" pageNumber="205" value="1988-06-27">27.6.1988</date>
Ježek leg.; bowl shaped basin (village Pskhu). Swamp slopes nr. Stream Betaga,
<quantity id="371C9B78601A21038BE90475FB598B12" box="[1167,1259,1048,1074]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="22" pageNumber="205" unit="m" value="2.0">2.000 m</quantity>
a.s.l., mountain forest,
<taxonomicName id="37E44D1E601A21038F820451FE968B76" box="[228,292,1084,1110]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="C290EA8F601A21038F820451FE968B76" box="[228,292,1084,1110]" italics="true" pageId="22" pageNumber="205">Alnus</emphasis>
</taxonomicName>
,
<taxonomicName id="37E44D1E601A21038E570451FE388B76" authorityName="C.Linnaeus" authorityYear="1753" box="[305,394,1084,1110]" class="Magnoliopsida" family="Betulaceae" genus="Corylus" kingdom="Plantae" order="Fagales" pageId="22" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="C290EA8F601A21038E570451FE388B76" box="[305,394,1084,1110]" italics="true" pageId="22" pageNumber="205">Corylus</emphasis>
</taxonomicName>
,
<taxonomicName id="37E44D1E601A21038EF00451FDBA8B76" box="[406,520,1084,1110]" class="Magnoliopsida" family="Adoxaceae" genus="Sambucus" kingdom="Plantae" order="Dipsacales" pageId="22" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="C290EA8F601A21038EF00451FDBA8B76" box="[406,520,1084,1110]" italics="true" pageId="22" pageNumber="205">Sambucus</emphasis>
</taxonomicName>
,
<taxonomicName id="37E44D1E601A21038D730451FC3D8B75" authority=", Musci Hepaticae. Right" authorityName="Musci Hepaticae. Right" box="[533,911,1083,1110]" class="Magnoliopsida" family="Rosaceae" genus="Fragaria" kingdom="Plantae" order="Rosales" pageId="22" pageNumber="205" phylum="Tracheophyta" rank="genus">
<emphasis id="C290EA8F601A21038D730451FDCE8B75" box="[533,636,1084,1109]" italics="true" pageId="22" pageNumber="205">Fragaria</emphasis>
, Musci Hepaticae. Right
</taxonomicName>
side red label:
<taxonomicName id="37E44D1E601A21038B500451FA2A8B76" authority="Jezek 1999" authorityName="Jezek" authorityYear="1999" box="[1078,1432,1083,1110]" class="Insecta" family="Limoniidae" genus="Tonnoirella" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="205" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="C290EA8F601A21038B500451FAA18B75" box="[1078,1299,1084,1110]" italics="true" pageId="22" pageNumber="205">Tonnoirella arcuata</emphasis>
<bibRefCitation id="94754B6C601A21038A7F0456FA2A8B76" author="Jezek, J." box="[1305,1432,1083,1110]" pageId="22" pageNumber="205" pagination="7 - 18" refId="ref20095" refString="Jezek, J. (1999) Rare and new Palaearctic Tonnoiriella - species (Diptera, Psychodidae). Casopis Narodniho muzea Rada prirodovedna, 168 (1 - 4), 7 - 18." type="journal article" year="1999">Ježek 1999</bibRefCitation>
</taxonomicName>
,
<typeStatus id="2F5F883F601A21038FF10432FF4B8B59" box="[151,249,1119,1145]" pageId="22" pageNumber="205" type="holotype">holotype</typeStatus>
Cat. No. 33663.
<typeStatus id="2F5F883F601A21038ED0040DFD958B5A" box="[438,551,1120,1146]" pageId="22" pageNumber="205" type="paratype">Paratypes</typeStatus>
: Cat. Nos. 33666, 33667, 33669; collected with
<typeStatus id="2F5F883F601A21038B580432FB128B59" box="[1086,1184,1119,1145]" pageId="22" pageNumber="205" type="holotype">holotype</typeStatus>
(all in Natural History Museum
<collectingRegion id="3220F87F601A21038E6604E9FEE18BBE" box="[256,339,1156,1182]" country="Czech Republic" name="Praha" pageId="22" pageNumber="205">Prague</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601A21038FA104CAFE6B89B6" pageId="22" pageNumber="205" type="description">
<paragraph id="F05B369D601A21038FA104CAFC9E8A55" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
<emphasis id="C290EA8F601A21038FA104CAFEE38BE2" bold="true" box="[199,337,1191,1218]" pageId="22" pageNumber="205">Description</emphasis>
: Head with eyebridge of 3 facet rows; distance between eyes 5.5 facet diameters; one row of postocular bristles. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042-0.042-0.036-0.036-0.036- 0.036-0.034-0.034-0.03-0.028-0.022-0.026-0.026-0.018-0.02-
<quantity id="371C9B78601A21038C280482FC0A8A2A" box="[846,952,1263,1290]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="22" pageNumber="205" unit="mm" value="0.03">0.03 mm</quantity>
; relative length 21-21-18-18-18-18-17-
<date id="845A105D601A21038A11049DFF4C8A0E" pageId="22" pageNumber="205" value="2017-11-15" valueMax="2017-11-14" valueMin="2017-11-15">17- 15-14-11</date>
-13-13-9-10-15; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 511; one antenna with two flagellomeres fused so that the flagellum appears 13-segmented. Palpus segments absolute length: 0.048-0.068-
<quantity id="371C9B78601A21038E400536FE578A56" box="[294,485,1371,1398]" metricMagnitude="-4" metricUnit="m" metricValue="1.09" metricValueMax="1.34" metricValueMin="0.84" pageId="22" pageNumber="205" unit="mm" value="0.109" valueMax="0.134" valueMin="0.084">0.084-0.134 mm</quantity>
; relative length 24-34-42-67.
</paragraph>
<paragraph id="F05B369D601A21038FA10512FC708ABA" blockId="22.[151,1437,975,1793]" box="[199,962,1407,1434]" pageId="22" pageNumber="205">
Wing 2.95 times longer than wide; length
<quantity id="371C9B78601A21038DF90512FCB48ABA" box="[671,774,1407,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="22" pageNumber="205" unit="mm" value="1.67">1.67 mm</quantity>
. width
<quantity id="371C9B78601A21038C310512FC0C8ABA" box="[855,958,1407,1434]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="22" pageNumber="205" unit="mm" value="0.57">0.57 mm</quantity>
.
</paragraph>
<paragraph id="F05B369D601A21038FA105CEFB488925" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
Male terminalia with thin hypandrium with 2 small lateral angular projections. Gonocoxites (probably artificially compressed) are approximately
<date id="845A105D601A21038D5805A5FDE88AC2" box="[574,602,1480,1506]" pageId="22" pageNumber="205">2x</date>
longer than wide, with a long seta in the distal fourth, condyles pentagonal. Gonostyli 1.3 times longer than gonocoxites, slightly bent. Parameres small, not clearly discernable.
</paragraph>
<paragraph id="F05B369D601A21038FA10662FA30896D" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
Aedeagus (
<figureCitation id="68DF2A18601A21038E200662FE25890A" box="[326,407,1551,1578]" captionStart="FIGURES 4349" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 4349: 4345 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 4849 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="22" pageNumber="205">Fig. 48</figureCitation>
) with short basiphallus sclerite; ventral arm of basiphallus bilobed, laterally connected to a U-shaped ventral distiphallus sclerite. Dorsal lobe of basiphallus broad straight, no distiphallus sclerite attached.
</paragraph>
<paragraph id="F05B369D601A21038FA1063AFE6B89B6" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
Epandrium (
<figureCitation id="68DF2A18601A21038E34063AFE108952" box="[338,418,1623,1650]" captionStart="FIGURES 4349" captionStartId="20.[151,264,1910,1935]" captionTargetBox="[231,1376,218,1841]" captionTargetId="figure@20.[191,1397,181,1884]" captionTargetPageId="20" captionText="FIGURES 4349: 4345 T. nigricauda 43 hypandrium, gonopods, aedeagus, 44 aedeagus ventral view; 45 variability of the ventral distiphallus sclerite in England, Poland, North- and South Germany (left to right?); 46 T. goncalvesi genitalia ventral view; 47 T. andradei hypandrium, gonopods, aedeagus; 4849 T. arcuata (holotype) 48 hypandrium, gonopods, aedeagus, 49 epandrium and epandrial processes, oblique view." figureDoi="http://doi.org/10.5281/zenodo.3621129" httpUri="https://zenodo.org/record/3621129/files/figure.png" pageId="22" pageNumber="205">Fig. 49</figureCitation>
) not clearly detectable; epandrial processes elongate, almost straight with 8 tenacula, apical ones shorter than basal ones.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601A21038FA106F2FDE78999" box="[199,597,1695,1722]" pageId="22" pageNumber="205" type="distribution">
<paragraph id="F05B369D601A21038FA106F2FDE78999" blockId="22.[151,1437,975,1793]" box="[199,597,1695,1722]" pageId="22" pageNumber="205">
<emphasis id="C290EA8F601A21038FA106F2FEEB899A" bold="true" box="[199,345,1695,1722]" pageId="22" pageNumber="205">Distribution</emphasis>
: Caucasus mountains.
</paragraph>
</subSubSection>
<subSubSection id="B8FE6516601A21038FA106A9FE508821" pageId="22" pageNumber="205" type="discussion">
<paragraph id="F05B369D601A21038FA106A9FE508821" blockId="22.[151,1437,975,1793]" pageId="22" pageNumber="205">
<emphasis id="C290EA8F601A21038FA106A9FE8689FE" bold="true" box="[199,308,1732,1758]" pageId="22" pageNumber="205">Remarks</emphasis>
: A species, easy to identify by the the double-headed hypandrium, and the distiphallus sclerite, almost semicircular and evenly bent.
</paragraph>
</subSubSection>
</treatment>
</document>