The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae) Author Wagner, Rüdiger Author Withers, Phil text Zootaxa 2020 2020-01-22 4728 2 183 210 journal article 24286 10.11646/zootaxa.4728.2.2 a1c40b73-0c11-4863-a188-412c33523e02 1175-5326 3621117 AE4F8089-7D2E-4ABA-AA09-CD504705788A Tonnoiriella arcuata Ježek, 1997 ( Figs 48, 49 ) Tonnoiriella arcuata Ježek, 1997 : Časopis Národniho Muzea v Praze Řada přirodovedna, 166: 1 16. Material : 1♂ holotype , slide mounted: head, mothparts with 2 palpi, 2 antennae, thorax cut in half, 2 wings, legs, abdomen part, 1 stylus, aedeagus, epandrium with epandrial processes; left side white label: Caucasus, Pskhu, 27.6.1988 Ježek leg.; bowl shaped basin (village Pskhu). Swamp slopes nr. Stream Betaga, 2.000 m a.s.l., mountain forest, Alnus , Corylus , Sambucus , Fragaria , Musci Hepaticae. Right side red label: Tonnoirella arcuata Ježek 1999 , holotype Cat. No. 33663. Paratypes : Cat. Nos. 33666, 33667, 33669; collected with holotype (all in Natural History Museum Prague ). Description : Head with eyebridge of 3 facet rows; distance between eyes 5.5 facet diameters; one row of postocular bristles. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042-0.042-0.036-0.036-0.036- 0.036-0.034-0.034-0.03-0.028-0.022-0.026-0.026-0.018-0.02- 0.03 mm ; relative length 21-21-18-18-18-18-17- 17- 15-14-11 -13-13-9-10-15; terminal flagellomere with apiculus. Pairs of digitate ascoids on flagellomeres 5–11; one antenna with two flagellomeres fused so that the flagellum appears 13-segmented. Palpus segments absolute length: 0.048-0.068- 0.084-0.134 mm ; relative length 24-34-42-67. Wing 2.95 times longer than wide; length 1.67 mm . width 0.57 mm . Male terminalia with thin hypandrium with 2 small lateral angular projections. Gonocoxites (probably artificially compressed) are approximately 2x longer than wide, with a long seta in the distal fourth, condyles pentagonal. Gonostyli 1.3 times longer than gonocoxites, slightly bent. Parameres small, not clearly discernable. Aedeagus ( Fig. 48 ) with short basiphallus sclerite; ventral arm of basiphallus bilobed, laterally connected to a U-shaped ventral distiphallus sclerite. Dorsal lobe of basiphallus broad straight, no distiphallus sclerite attached. Epandrium ( Fig. 49 ) not clearly detectable; epandrial processes elongate, almost straight with 8 tenacula, apical ones shorter than basal ones. Distribution : Caucasus mountains. Remarks : A species, easy to identify by the the double-headed hypandrium, and the distiphallus sclerite, almost semicircular and evenly bent.