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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1159.99218" ID-Pensoft-Pub="1313-2970-1159-87" ID-Pensoft-UUID="ECCECB868071523DB7628E304850BD62" ID-ZooBank="CD97DD120549412F976508D8DE21605B" ModsDocID="1313-2970-1159-87" checkinTime="1682458941538" checkinUser="pensoft" docAuthor="Okada, Ryohei, Jaitrong, Weeyawat &amp; Wewalka, Guenther" docDate="2023" docId="B70D2C12B4DC50A0B553DCC9BD80B24A" docLanguage="en" docName="ZooKeys 1159: 87-119" docOrigin="ZooKeys 1159" docPubDate="2023-04-25" docSource="http://dx.doi.org/10.3897/zookeys.1159.99218" docTitle="Microdytes schwendingeri Wewalka 1997" docType="treatment" docVersion="1" id="ECCECB868071523DB7628E304850BD62" lastPageNumber="87" masterDocId="ECCECB868071523DB7628E304850BD62" masterDocTitle="A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)" masterLastPageNumber="119" masterPageNumber="87" pageNumber="87" updateTime="1682458941538" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Okada, Ryohei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8488-0660</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand &amp; Coleopterological Society of Japan, National Museum of Nature and Science, Tsukuba, Japan</mods:affiliation>
<mods:nameIdentifier type="email">wasserinsekt@kub.biglobe.ne.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaitrong, Weeyawat</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1362-0754</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wewalka, Guenther</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2994-8001</mods:nameIdentifier>
<mods:affiliation>Starkfriedgasse 16 / 1 / 3, A - 1190 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1159</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>119</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1159.99218</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1159.99218</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1159-87</mods:identifier>
<mods:identifier type="ZooBank">CD97DD120549412F976508D8DE21605B</mods:identifier>
<mods:identifier type="Pensoft-UUID">ECCECB868071523DB7628E304850BD62</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:B70D2C12B4DC50A0B553DCC9BD80B24A" httpUri="http://treatment.plazi.org/id/B70D2C12B4DC50A0B553DCC9BD80B24A" lastPageNumber="87" pageId="0" pageNumber="87">
<subSubSection pageId="0" pageNumber="87" type="nomenclature">
<paragraph pageId="0" pageNumber="87">
<taxonomicName ID-CoL="42WF4" LSID="B70D2C12-B4DC-50A0-B553-DCC9BD80B24A" authority="Wewalka, 1997" authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes schwendingeri" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="schwendingeri">Microdytes schwendingeri Wewalka, 1997</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">Figs 34</figureCitation>
<figureCitation captionStart="Figures 3840" captionStartId="F8" captionText="Figures 38 - 40. Diagnostic characters of Microdytes species. 38 Impression of lateral side (M. maximiliani sp. nov.) 39 ventral surface A with setae (M. heineri) B with very coarse punctures (M. schoedli) C with coarse punctures (M. maculatus) D with fine punctures (M. balkei) E with very fine punctures (M. jeenthongi sp. nov.) 40 clypeus bead A not bordered (M. ubonensis sp. nov.) B finely bordered (M. schwendingeri) C produced (M. dimorphus)." figureDoi="10.3897/zookeys.1159.99218.figures38-40" httpUri="https://binary.pensoft.net/fig/842228" pageId="0" pageNumber="87">, 40B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="reference_group">
<paragraph pageId="0" pageNumber="87">
<taxonomicName authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes schwendingeri" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="schwendingeri">Microdytes schwendingeri</taxonomicName>
Wewalka, 1997: 36;
<bibRefCitation author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="87" pagination="21 - 39" refId="B24" refString="Wewalka, G, 2011. New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946. Koleopterologische Rundschau 81: 21 - 39" title="New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946." volume="81" year="2011">Wewalka 2011</bibRefCitation>
: 37;
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Etudes Entomologiques Motschulsky" pageId="0" pageNumber="87" publicationUrl="http://www.norrent.se" refId="B13" refString="Nilsson, AN, Hajek, J, 2023. A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2023. http://www.norrent.se" title="A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2023." url="http://www.norrent.se" year="2023">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2023
</bibRefCitation>
: 212.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="type locality">
<paragraph pageId="0" pageNumber="87">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand, Sakon Nakhon Province, Phu Pan NP.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="materials_examined">
<paragraph pageId="0" pageNumber="87">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="2020-12-27" collectorName="R. Okada" country="Thailand" elevation="200" location="Phai Lom St." municipality="Bang Phaeng District" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nakhon Phanom Province">
<collectingCountry name="Thailand">Thailand</collectingCountry>
:
<collectingRegion country="Thailand" name="Nakhon Phanom">Nakhon Phanom Province</collectingRegion>
.
<specimenCount type="male">1♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
,
<collectingMunicipality>Bang Phaeng District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:B70D2C12B4DC50A0B553DCC9BD80B24A:F70CD91BF819D9D831228C3EDBCBFF4C" country="Thailand" municipality="Bang Phaeng District" name="Phai Lom St." stateProvince="Nakhon Phanom Province">Phai Lom St.</location>
203 (alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</elevation>
</quantity>
),
<collectingDate value="2020-12-27">27.XII.2020</collectingDate>
,
<collectorName>R. Okada</collectorName>
leg. (CRO, THNHM) (Fig.
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">34</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="2011-05-31" collectingDateMax="2011-06-06" collectingDateMin="2011-05-31" country="Laos" elevation="325" latitude="17.0" location="Phou Xang He" longLatPrecision="1280" longitude="105.63333" specimenCount="1" specimenCount-female="1" stateProvince="Savannakhet Province">
<collectingCountry name="Laos">Laos</collectingCountry>
:
<collectingRegion country="Laos" name="Savannakhet">Savannakhet Province</collectingRegion>
,
<specimenCount type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:B70D2C12B4DC50A0B553DCC9BD80B24A:A7A2CB1F115587D047D950A1194E3BB7" country="Laos" latitude="17.0" longLatPrecision="1280" longitude="105.63333" name="Phou Xang He" stateProvince="Savannakhet Province">Phou Xang He</location>
NBCA, ca.
<locationDeviation location="Ban Pa Phaknau">
<quantity metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
SW Ban Pa Phaknau
</locationDeviation>
,
<geoCoordinate degrees="17" direction="north" minutes="00" orientation="latitude" precision="925" value="17.0">17°00'N</geoCoordinate>
,
<geoCoordinate degrees="105" direction="east" minutes="38" orientation="longitude" precision="925" value="105.63333">105°38'E</geoCoordinate>
, alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" unit="m" value="325.0" valueMax="400.0" valueMin="250.0">250-400 m</elevation>
</quantity>
,
<collectingDate value="2011-05-31" valueMax="2011-06-06" valueMin="2011-05-31">31.V.-6.VI. 2011</collectingDate>
, NHMB Basel
</materialsCitation>
,
<materialsCitation collectingDate="2010-05-10" collectingDateMax="2010-05-12" collectingDateMin="2010-05-10" collectorName="Expedition, M, Brancucci, M, Geiser, D, Hauk, Z, Kraus, A, E. Vongphachan, J. Hajek" country="Laos" elevation="415" latitude="15.135" location="Bolavens Plateau" longLatPrecision="128" longitude="106.64667" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Champasak Province">
<collectingCountry name="Laos">Laos</collectingCountry>
2011
<collectorName>Expedition, M</collectorName>
.
<collectorName>Brancucci, M</collectorName>
.
<collectorName>Geiser, D</collectorName>
.
<collectorName>Hauk, Z</collectorName>
.
<collectorName>Kraus, A</collectorName>
. Phantala &amp;
<collectorName>E. Vongphachan</collectorName>
leg. (NMB);
<collectingRegion country="Laos" name="Champasak">Champasak Province</collectingRegion>
.
<specimenCount type="male">2♂♂</specimenCount>
,
<specimenCount type="female">1♀</specimenCount>
,
<location LSID="urn:lsid:plazi:treatment:B70D2C12B4DC50A0B553DCC9BD80B24A:C4E7E6D40F3845638ECEAF53AFF8C916" country="Laos" latitude="15.135" longLatPrecision="128" longitude="106.64667" name="Bolavens Plateau" stateProvince="Champasak Province">Bolavens Plateau</location>
, waterfall ca.
<locationDeviation location="Tad Katamtok">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
E Tad Katamtok
</locationDeviation>
,
<geoCoordinate degrees="15" direction="north" minutes="08.1" orientation="latitude" precision="92" value="15.135">15°08.1'N</geoCoordinate>
,
<geoCoordinate degrees="106" direction="east" minutes="38.8" orientation="longitude" precision="92" value="106.64667">106°38.8'E</geoCoordinate>
, alt.
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.15" unit="m" value="415.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.15" unit="m" value="415.0">415 m</elevation>
</quantity>
,
<collectingDate value="2010-05-10" valueMax="2010-05-12" valueMin="2010-05-10">10.-12.V.2010</collectingDate>
,
<collectorName>
J.
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
</collectorName>
leg. (CGW, NMP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="comments">
<paragraph pageId="0" pageNumber="87">Comments.</paragraph>
<paragraph pageId="0" pageNumber="87">Some specimens from Laos have less distinct elytral markings but no other differences from typical specimens have been observed.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand: Nakhon Phanom (first record) and Sakon Nakhon provinces; Laos (first record): Savannakhet and Champasak provinces.</paragraph>
</subSubSection>
</treatment>
</document>