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<document id="23BC9BFB0AF04CCE60DAA94DFC84F641" ID-CLB-Dataset="169751" ID-DOI="10.11646/zootaxa.5195.5.4" ID-GBIF-Dataset="38cca3f8-185a-4feb-a0e5-f7e51475d54e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7223747" ID-ZooBank="9238817E-3772-4CDF-9B50-045F456050E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666163934792" checkinUser="plazi" docAuthor="Luo, Jiu-Yang, Wang, Yan-Hui, Chen, Ping-Ping &amp; Xie, Qiang" docDate="2022" docId="C8089363FFCCCB43FF7345B0FA446141" docLanguage="en" docName="zootaxa.5195.5.4.pdf" docOrigin="Zootaxa 5195 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Omaniidae Cobben 1970" docType="key" docVersion="6" lastPageNumber="456" masterDocId="3431EB1BFFCBCB44FFE4477FFFD86747" masterDocTitle="First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha)" masterLastPageNumber="467" masterPageNumber="449" pageNumber="456" updateTime="1699408882582" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="43FCA604CF1D0EA19617A0FEC40F5613">First record of the intertidal dwarf bug family Omaniidae from China, with description of a new species (Heteroptera: Leptopodomorpha)</mods:title>
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<mods:namePart id="58D935920C46422D72F425135F99A696">Luo, Jiu-Yang</mods:namePart>
<mods:affiliation id="BCAB5ED8C78C5FD83BC5CEB9C951757B">State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China. &amp; School of Life Sciences, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China.</mods:affiliation>
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<mods:namePart id="8FE1EE046B765686A259D3A89B3BC11A">Wang, Yan-Hui</mods:namePart>
<mods:affiliation id="5DA4A3D060DDC0C44C043930E48D9C5A">State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China. &amp; School of Life Sciences, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China.</mods:affiliation>
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<mods:namePart id="FF18C87B01F2B75D8A96FBC8FCBF4388">Chen, Ping-Ping</mods:namePart>
<mods:affiliation id="4F3F5E5454DEF80A2B6B437C89689F8D">Netherlands Centre of Biodiversity Naturalis, 2300 RA, Leiden, The Netherlands</mods:affiliation>
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<mods:roleTerm id="AE01A448C18DFC5FC8DE67A012DDB6DC">Author</mods:roleTerm>
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<mods:namePart id="772CA3ED65BE9F71D6C352A97BC05344">Xie, Qiang</mods:namePart>
<mods:affiliation id="825F133C0EA36946C63747D7E87DE6C3">State Key Laboratory of Biocontrol, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China. &amp; School of Life Sciences, Sun Yat-sen University, 135 Xingangxi Road, Guangzhou 510275, Guangdong, China.</mods:affiliation>
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<mods:title id="C07E6F2BD79DC05B78AD7E8660D00189">Zootaxa</mods:title>
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<mods:date id="499A9949EC58AF091AC85892696F5E94">2022</mods:date>
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<mods:number id="F344B1B00366E239AEC9021F44C44693">2022-10-14</mods:number>
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<treatment id="C8089363FFCCCB43FF7345B0FA446141" ID-DOI="http://doi.org/10.5281/zenodo.7223757" ID-GBIF-Taxon="203118055" ID-Zenodo-Dep="7223757" LSID="urn:lsid:plazi:treatment:C8089363FFCCCB43FF7345B0FA446141" httpUri="http://treatment.plazi.org/id/C8089363FFCCCB43FF7345B0FA446141" lastPageNumber="456" pageId="7" pageNumber="456">
<subSubSection id="08BB71FEFFCCCB43FF7345B0FE2C644A" pageId="7" pageNumber="456" type="nomenclature">
<paragraph id="401E2275FFCCCB43FF7345B0FD0265AE" blockId="7.[151,730,719,781]" box="[151,730,719,745]" pageId="7" pageNumber="456">
<heading id="1B569519FFCCCB43FF7345B0FD0265AE" bold="true" box="[151,730,719,745]" fontSize="11" level="1" pageId="7" pageNumber="456" reason="1">
<emphasis id="72D5FE67FFCCCB43FF7345B0FD0265AE" bold="true" box="[151,730,719,745]" pageId="7" pageNumber="456">
Key to the species of
<taxonomicName id="87A159F6FFCCCB43FE7945B0FD0265AE" ID-CoL="DKV" authority="Cobben, 1970" authorityName="Cobben" authorityYear="1970" box="[413,730,719,745]" class="Insecta" family="Omaniidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="family">
Omaniidae
<bibRefCitation id="24305F84FFCCCB43FDCB45B0FD0265AE" author="Cobben, R. H." box="[559,730,719,745]" pageId="7" pageNumber="456" pagination="61 - 90" refId="ref9614" refString="Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90." type="journal article" year="1970">Cobben, 1970</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="401E2275FFCCCB43FF73458CFE2C644A" blockId="7.[151,730,719,781]" box="[151,500,755,781]" pageId="7" pageNumber="456">
Modified from
<bibRefCitation id="24305F84FFCCCB43FEA0458CFE28644A" author="Cobben, R. H." box="[324,496,755,781]" pageId="7" pageNumber="456" pagination="61 - 90" refId="ref9614" refString="Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90." type="journal article" year="1970">Cobben (1970)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="08BB71FEFFCCCB43FF73444EFA446141" pageId="7" pageNumber="456" type="key">
<keyStep id="FB553AD0FFCCCB43FF73444EFA4464D9" pageId="7" pageNumber="456">
<paragraph id="401E2275FFCCCB43FF73444EFA446423" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
<keyLead id="FB508140FFCCCB43FF73444EFA446423" pageId="7" pageNumber="456">
1. Body longer than 2.0 mm; forewings with well-developed membrane or flap-like leaving most of abdominal dorsum exposed; eye without trichobothria in adults.............................. Families
<taxonomicName id="87A159F6FFCCCB43FC104431FBA46423" ID-CoL="63R" authorityName="Lethierry &amp; Severin" authorityYear="1896" box="[1012,1148,846,868]" class="Insecta" family="Aepophilidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="family">
<emphasis id="72D5FE67FFCCCB43FC104431FBA46423" bold="true" box="[1012,1148,846,868]" pageId="7" pageNumber="456">Aepophilidae</emphasis>
</taxonomicName>
,
<taxonomicName id="87A159F6FFCCCB43FB6C4431FACD6423" ID-CoL="BY3" authorityName="Brulle" authorityYear="1836" box="[1160,1301,846,868]" class="Insecta" family="Leptopodidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="family">
<emphasis id="72D5FE67FFCCCB43FB6C4431FACD6423" bold="true" box="[1160,1301,846,868]" pageId="7" pageNumber="456">Leptopodidae</emphasis>
</taxonomicName>
and
<taxonomicName id="87A159F6FFCCCB43FAA04431FA446423" ID-CoL="FWD" authorityName="Amyot &amp; Serville" authorityYear="1843" box="[1348,1436,846,868]" class="Insecta" family="Saldidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="family">
<emphasis id="72D5FE67FFCCCB43FAA04431FA446423" bold="true" box="[1348,1436,846,868]" pageId="7" pageNumber="456">Saldidae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="401E2275FFCCCB43FF734413FA4464D9" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
<keyLead id="FB508140FFCCCB43FF734413FA4464D9" pageId="7" pageNumber="456">
– Body shorter than 2.0 mm; forewings elytra-like without membrane (
<figureCitation id="D89A3EF0FFCCCB43FC9A4413FC1864C6" box="[894,960,876,898]" captionStart="FIGURE 1" captionStartId="2.[151,250,1810,1834]" captionTargetBox="[151,1435,376,1787]" captionTargetId="figure-38@2.[151,1435,376,1787]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Corallocoris xishaensis sp. nov.: A–C, male holotype; D–F, female paratype. A, dorsal view; B, lateral view; C, ventral view; D, dorsal view; E, lateral view; F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223749" httpUri="https://zenodo.org/record/7223749/files/figure.png" pageId="7" pageNumber="456">Figs. 1</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FC294413FC3564C6" box="[973,1005,876,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">2D</figureCitation>
); eye with 3 trichobothria in adults (
<figureCitation id="D89A3EF0FFCCCB43FAA94413FF2A64D9" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Fig. 2B, E</figureCitation>
)...............................................................................
<taxonomicName id="87A159F6FFCCCB43FB2C44F6FA5364D8" ID-CoL="DKV" authority="Cobben" authorityName="Cobben" authorityYear="1970" box="[1224,1419,905,927]" class="Insecta" family="Omaniidae" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="family">
<emphasis id="72D5FE67FFCCCB43FB2C44F6FAE164D8" bold="true" box="[1224,1337,905,927]" pageId="7" pageNumber="456">Omaniidae</emphasis>
Cobben
</taxonomicName>
2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="FB553AD0FFCCCB43FF7344D8FA446376" pageId="7" pageNumber="456">
<paragraph id="401E2275FFCCCB43FF7344D8FA4564B0" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
<keyLead id="FB508140FFCCCB43FF7344D8FA4564B0" pageId="7" pageNumber="456">
2. Collar of head sharply projecting medially into the vertex region (
<figureCitation id="D89A3EF0FFCCCB43FC8744D9FC6964FB" box="[867,945,934,956]" captionStart="FIGURE 8" captionStartId="12.[151,250,1853,1877]" captionTargetBox="[162,1435,192,1829]" captionTargetId="figure-20@12.[151,1435,181,1829]" captionTargetPageId="12" captionText="FIGURE 8.A–B, Habitus of Omania coleoptrata, paratypes, female:A, dorsal view; B, lateral view C–D, habitus of Corallocoris nauruensis, paratypes, male: C, dorsal view; D, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7223769" httpUri="https://zenodo.org/record/7223769/files/figure.png" pageId="7" pageNumber="456">Fig. 8A</figureCitation>
, also see
<bibRefCitation id="24305F84FFCCCB43FBEB44D9FB4064FB" author="Horvath, G." box="[1039,1176,934,956]" pageId="7" pageNumber="456" pagination="598 - 599" refId="ref10048" refString="Horvath, G. (1915) Novum genus peculliare Acanthiidarum. Annales Musei Nationalis Hungarici, 13, 598 - 599." type="journal article" year="1915">Horváth 1915</bibRefCitation>
: fig. 1;
<bibRefCitation id="24305F84FFCCCB43FB0744D9FAB164FB" author="Cobben, R. H." box="[1251,1385,934,956]" pageId="7" pageNumber="456" pagination="61 - 90" refId="ref9614" refString="Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90." type="journal article" year="1970">Cobben 1970</bibRefCitation>
: figs. 1A, 6A); ocelli not circular (see
<bibRefCitation id="24305F84FFCCCB43FDFC44BBFD45649E" author="Cobben, R. H." box="[536,669,964,986]" pageId="7" pageNumber="456" pagination="61 - 90" refId="ref9614" refString="Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90." type="journal article" year="1970">Cobben 1970</bibRefCitation>
: fig. 6A); forewing without sutures (
<figureCitation id="D89A3EF0FFCCCB43FC1E44BBFBBB649E" box="[1018,1123,964,986]" captionStart="FIGURE 8" captionStartId="12.[151,250,1853,1877]" captionTargetBox="[162,1435,192,1829]" captionTargetId="figure-20@12.[151,1435,181,1829]" captionTargetPageId="12" captionText="FIGURE 8.A–B, Habitus of Omania coleoptrata, paratypes, female:A, dorsal view; B, lateral view C–D, habitus of Corallocoris nauruensis, paratypes, male: C, dorsal view; D, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7223769" httpUri="https://zenodo.org/record/7223769/files/figure.png" pageId="7" pageNumber="456">Fig. 8A–B</figureCitation>
, also see
<bibRefCitation id="24305F84FFCCCB43FB5944BBFA9A649E" author="Cobben, R. H." box="[1213,1346,964,986]" pageId="7" pageNumber="456" pagination="61 - 90" refId="ref9614" refString="Cobben, R. H. (1970) Morphology and taxonomy of intertidal dwarf bugs (Heteroptera: Omaniidae Fam. Nov.). TiJdschrift voor Entomologie, 113, 61 - 90." type="journal article" year="1970">Cobben 1970</bibRefCitation>
: fig. 1A). Femora entirely stramineous (
<collectingCountry id="38B662E5FFCCCB43FDE5449EFDE764B1" box="[513,575,993,1014]" name="Egypt" pageId="7" pageNumber="456">Egypt</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FDAC449EFD1064B0" box="[584,712,993,1015]" name="Saudi Arabia" pageId="7" pageNumber="456">Saudi Arabia</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FD36449EFCC864B1" box="[722,784,993,1014]" name="Oman" pageId="7" pageNumber="456">Oman</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FCFD449EFCB464B0" box="[793,876,993,1015]" name="Pakistan" pageId="7" pageNumber="456">Pakistan</collectingCountry>
).............
<taxonomicName id="87A159F6FFCCCB43FBEE449EFB6A64B0" ID-CoL="67WL" authority="Horvath" authorityName="Horvath" authorityYear="1915" box="[1034,1202,993,1015]" class="Insecta" family="Omaniidae" genus="Omania" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="72D5FE67FFCCCB43FBEE449EFB8264B1" bold="true" box="[1034,1114,993,1014]" italics="true" pageId="7" pageNumber="456">Omania</emphasis>
Horváth
</taxonomicName>
:
<taxonomicName id="87A159F6FFCCCB43FB58449EFA4564B0" ID-CoL="74JSQ" authority="Horvath" authorityName="Horvath" authorityYear="1915" box="[1212,1437,993,1015]" class="Insecta" family="Omaniidae" genus="Omania" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="coleoptrata">
<emphasis id="72D5FE67FFCCCB43FB58449EFA9064B0" bold="true" box="[1212,1352,993,1015]" italics="true" pageId="7" pageNumber="456">O. coleoptrata</emphasis>
Horváth
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="401E2275FFCCCB43FF734480FA446376" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
<keyLead id="FB508140FFCCCB43FF734480FA446376" pageId="7" pageNumber="456">
– Collar of head with broadly arcuate anterior margin (
<figureCitation id="D89A3EF0FFCCCB43FD104481FC936353" box="[756,843,1022,1044]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2B</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FCBC4480FCA16353" box="[856,889,1023,1044]" captionStart="FIGURE 4" captionStartId="5.[151,250,1867,1891]" captionTargetBox="[242,1345,181,1844]" captionTargetId="figure-17@5.[242,1345,181,1844]" captionTargetPageId="5" captionText="FIGURE 4. Scanning electron micrographs of morphological structures of Corallocoris xishaensis sp. nov., female non-type specimen:A, head and part of pronotum in dorsal view; B, microtrichia on collar of head; C, fovea of pronotum; D, microtrichia and setae on pronotum; E, meso- and metathorax in ventral view; F, structure on mesosternum; G, hind coxa (sa = setose area); H, setose area." figureDoi="http://doi.org/10.5281/zenodo.7223759" httpUri="https://zenodo.org/record/7223759/files/figure.png" pageId="7" pageNumber="456">4A</figureCitation>
); ocelli circular (
<figureCitation id="D89A3EF0FFCCCB43FBC04481FBA46353" box="[1060,1148,1022,1044]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2B</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FB6C4480FB7E6353" box="[1160,1190,1023,1044]" captionStart="FIGURE 3" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[242,1345,181,1844]" captionTargetId="figure-20@4.[242,1345,181,1844]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron micrographs of morphological structures of Corallocoris xishaensis sp. nov., female non-type specimen: A, body; B, vertex of head in dorsal view; C, base of forewing in dorsal view; D, forewing in dorsal view; E, scutellum in dorsal view; F, lower surface of forewing; G, base of lower surface of forewing; H, microtrichia on lower surface of forewing. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223755" httpUri="https://zenodo.org/record/7223755/files/figure.png" pageId="7" pageNumber="456">3B</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FB574481FB096353" box="[1203,1233,1022,1044]" captionStart="FIGURE 6" captionStartId="7.[151,250,612,636]" captionTargetBox="[151,1435,181,589]" captionTargetId="figure-17@7.[151,1435,181,589]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Corallocoris xishaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7223763" httpUri="https://zenodo.org/record/7223763/files/figure.png" pageId="7" pageNumber="456">6E</figureCitation>
); forewing with two longitudinal sutures (
<figureCitation id="D89A3EF0FFCCCB43FE494363FDDB6376" box="[429,515,1052,1074]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2D</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FDE94363FD926375" box="[525,586,1052,1074]" captionStart="FIGURE 6" captionStartId="7.[151,250,612,636]" captionTargetBox="[151,1435,181,589]" captionTargetId="figure-17@7.[151,1435,181,589]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Corallocoris xishaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7223763" httpUri="https://zenodo.org/record/7223763/files/figure.png" pageId="7" pageNumber="456">6A–B</figureCitation>
). Femora with dark area and light apex.....................
<taxonomicName id="87A159F6FFCCCB43FB5B4364FA536375" ID-CoL="3TX3" authority="Cobben" authorityName="Cobben" authorityYear="1970" box="[1215,1419,1051,1074]" class="Insecta" family="Omaniidae" genus="Corallocoris" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="genus">
<emphasis id="72D5FE67FFCCCB43FB5B4364FAE26376" bold="true" box="[1215,1338,1051,1073]" italics="true" pageId="7" pageNumber="456">Corallocoris</emphasis>
Cobben
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3
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3. Cephalic trichobothria pair
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front of pair 1 (see
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: Plate fig. 1A); cephalic trichobothria pair 3 not arising from light spots (see
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: Plate fig. 1A) (
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).........................................
<taxonomicName id="87A159F6FFCCCB43FB3C4329FA44632B" authority="(Miyamoto)" baseAuthorityName="Miyamoto" box="[1240,1436,1110,1132]" class="Insecta" family="Omaniidae" genus="Omania" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="satoi">
<emphasis id="72D5FE67FFCCCB43FB3C4329FAFB632C" bold="true" box="[1240,1315,1110,1131]" italics="true" pageId="7" pageNumber="456">C. satoi</emphasis>
(Miyamoto)
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– Cephalic trichobothria pairs 1 and
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a straight line (
<figureCitation id="D89A3EF0FFCCCB43FD1E430BFCB463CE" box="[762,876,1140,1162]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2B, E</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FC9C430BFC4E63CE" box="[888,918,1140,1161]" captionStart="FIGURE 3" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[242,1345,181,1844]" captionTargetId="figure-20@4.[242,1345,181,1844]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron micrographs of morphological structures of Corallocoris xishaensis sp. nov., female non-type specimen: A, body; B, vertex of head in dorsal view; C, base of forewing in dorsal view; D, forewing in dorsal view; E, scutellum in dorsal view; F, lower surface of forewing; G, base of lower surface of forewing; H, microtrichia on lower surface of forewing. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223755" httpUri="https://zenodo.org/record/7223755/files/figure.png" pageId="7" pageNumber="456">3B</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FC45430BFC1A63CD" box="[929,962,1140,1162]" captionStart="FIGURE 6" captionStartId="7.[151,250,612,636]" captionTargetBox="[151,1435,181,589]" captionTargetId="figure-17@7.[151,1435,181,589]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Corallocoris xishaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7223763" httpUri="https://zenodo.org/record/7223763/files/figure.png" pageId="7" pageNumber="456">6D</figureCitation>
); cephalic trichobothria pair 3 arising from light elevated spots (
<figureCitation id="D89A3EF0FFCCCB43FE9243EEFE3E63E0" box="[374,486,1169,1191]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2B, E</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FE0B43EEFDD563E1" box="[495,525,1169,1190]" captionStart="FIGURE 3" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[242,1345,181,1844]" captionTargetId="figure-20@4.[242,1345,181,1844]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron micrographs of morphological structures of Corallocoris xishaensis sp. nov., female non-type specimen: A, body; B, vertex of head in dorsal view; C, base of forewing in dorsal view; D, forewing in dorsal view; E, scutellum in dorsal view; F, lower surface of forewing; G, base of lower surface of forewing; H, microtrichia on lower surface of forewing. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223755" httpUri="https://zenodo.org/record/7223755/files/figure.png" pageId="7" pageNumber="456">3B</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FDF343EEFD8863E0" box="[535,592,1169,1191]" captionStart="FIGURE 6" captionStartId="7.[151,250,612,636]" captionTargetBox="[151,1435,181,589]" captionTargetId="figure-17@7.[151,1435,181,589]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Corallocoris xishaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7223763" httpUri="https://zenodo.org/record/7223763/files/figure.png" pageId="7" pageNumber="456">6D–E</figureCitation>
).................................................................... 4
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<paragraph id="401E2275FFCCCB43FF7343D0FA4463A5" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
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4. Pronotum and scutellum with light areas (
<figureCitation id="D89A3EF0FFCCCB43FD9943D1FD136383" box="[637,715,1198,1220]" captionStart="FIGURE 8" captionStartId="12.[151,250,1853,1877]" captionTargetBox="[162,1435,192,1829]" captionTargetId="figure-20@12.[151,1435,181,1829]" captionTargetPageId="12" captionText="FIGURE 8.A–B, Habitus of Omania coleoptrata, paratypes, female:A, dorsal view; B, lateral view C–D, habitus of Corallocoris nauruensis, paratypes, male: C, dorsal view; D, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7223769" httpUri="https://zenodo.org/record/7223769/files/figure.png" pageId="7" pageNumber="456">Fig. 8C</figureCitation>
); median fovea of pronotal dome dividing the dome posteriorly (
<figureCitation id="D89A3EF0FFCCCB43FAAE43D1FA4C6383" box="[1354,1428,1198,1220]" captionStart="FIGURE 8" captionStartId="12.[151,250,1853,1877]" captionTargetBox="[162,1435,192,1829]" captionTargetId="figure-20@12.[151,1435,181,1829]" captionTargetPageId="12" captionText="FIGURE 8.A–B, Habitus of Omania coleoptrata, paratypes, female:A, dorsal view; B, lateral view C–D, habitus of Corallocoris nauruensis, paratypes, male: C, dorsal view; D, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7223769" httpUri="https://zenodo.org/record/7223769/files/figure.png" pageId="7" pageNumber="456">Fig. 8C</figureCitation>
) (
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)................................................................
<taxonomicName id="87A159F6FFCCCB43FBD243B3FA4463A5" authority="(Herring &amp; Chapman)" baseAuthorityName="Herring &amp; Chapman" box="[1078,1436,1228,1250]" class="Insecta" family="Omaniidae" genus="Omania" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="nauruensis">
<emphasis id="72D5FE67FFCCCB43FBD243B3FB1963A6" bold="true" box="[1078,1217,1228,1249]" italics="true" pageId="7" pageNumber="456">C. nauruensis</emphasis>
(Herring &amp; Chapman)
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</paragraph>
<paragraph id="401E2275FFCCCB43FF734395FA44625B" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
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– Pronotum and scutellum concolorous (
<figureCitation id="D89A3EF0FFCCCB43FDB34396FD1363B8" box="[599,715,1257,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1810,1834]" captionTargetBox="[151,1435,376,1787]" captionTargetId="figure-38@2.[151,1435,376,1787]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Corallocoris xishaensis sp. nov.: A–C, male holotype; D–F, female paratype. A, dorsal view; B, lateral view; C, ventral view; D, dorsal view; E, lateral view; F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223749" httpUri="https://zenodo.org/record/7223749/files/figure.png" pageId="7" pageNumber="456">Figs. 1A, D</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FD324396FD2F63B8" box="[726,759,1257,1279]" captionStart="FIGURE 6" captionStartId="7.[151,250,612,636]" captionTargetBox="[151,1435,181,589]" captionTargetId="figure-17@7.[151,1435,181,589]" captionTargetPageId="7" captionText="FIGURE 6. Habitat of Corallocoris xishaensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7223763" httpUri="https://zenodo.org/record/7223763/files/figure.png" pageId="7" pageNumber="456">6A</figureCitation>
); median fovea of pronotal dome present as a simple pit (
<figureCitation id="D89A3EF0FFCCCB43FAC04396FA4063B8" box="[1316,1432,1257,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1810,1834]" captionTargetBox="[151,1435,376,1787]" captionTargetId="figure-38@2.[151,1435,376,1787]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Corallocoris xishaensis sp. nov.: A–C, male holotype; D–F, female paratype. A, dorsal view; B, lateral view; C, ventral view; D, dorsal view; E, lateral view; F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223749" httpUri="https://zenodo.org/record/7223749/files/figure.png" pageId="7" pageNumber="456">Figs. 1A, D</figureCitation>
,
<figureCitation id="D89A3EF0FFCCCB43FF064278FEC5625B" box="[226,285,1287,1308]" captionStart="FIGURE 3" captionStartId="4.[151,250,1867,1891]" captionTargetBox="[242,1345,181,1844]" captionTargetId="figure-20@4.[242,1345,181,1844]" captionTargetPageId="4" captionText="FIGURE 3. Scanning electron micrographs of morphological structures of Corallocoris xishaensis sp. nov., female non-type specimen: A, body; B, vertex of head in dorsal view; C, base of forewing in dorsal view; D, forewing in dorsal view; E, scutellum in dorsal view; F, lower surface of forewing; G, base of lower surface of forewing; H, microtrichia on lower surface of forewing. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223755" httpUri="https://zenodo.org/record/7223755/files/figure.png" pageId="7" pageNumber="456">3A, D</figureCitation>
).............................................................................................. 5
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5. Elevated spots on frons distinctly larger than ocelli, contact with eyes; base of hypocostal lamina clearly notched along ventral side (
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).......................................................................
<taxonomicName id="87A159F6FFCCCB43FB3D423EFA456210" ID-CoL="5ZZNC" authority="Cobben" authorityName="Cobben" authorityYear="1987" box="[1241,1437,1345,1367]" class="Insecta" family="Omaniidae" genus="Corallocoris" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="aldabrae">
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<emphasis id="72D5FE67FFCCCB43FB3D423EFB316211" bold="true" box="[1241,1257,1345,1366]" italics="true" pageId="7" pageNumber="456">C</emphasis>
.
<emphasis id="72D5FE67FFCCCB43FB11423EFA946210" bold="true" box="[1269,1356,1345,1367]" italics="true" pageId="7" pageNumber="456">aldabrae</emphasis>
</emphasis>
Cobben
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<paragraph id="401E2275FFCCCB43FF734220FA4462D5" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
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– Elevated spots on frons not distinctly larger than ocelli, and not contact with eyes (
<figureCitation id="D89A3EF0FFCCCB43FBF24221FB516233" box="[1046,1161,1374,1396]" captionStart="FIGURE 2" captionStartId="3.[151,250,1867,1891]" captionTargetBox="[172,1416,181,1844]" captionTargetId="figure-17@3.[172,1416,181,1844]" captionTargetPageId="3" captionText="FIGURE 2. Morphological structures of Corallocoris xishaensis sp. nov.: A, G, K, male paratype; B–F, H–J, female paratype. A, meso-and metathorax in dorsal view; B, head and prothorax in dorsal view; C, abdomen in dorsal view; D, right forewing in dorsal view; E, head and prothorax in frontal view; F, abdomen in lateral view; G, abdomen in lateral view; H, foreleg; I, middle leg; J, hind leg; K, abdomen in dorsal view. See Materials and methods for structure abbreviations." figureDoi="http://doi.org/10.5281/zenodo.7223753" httpUri="https://zenodo.org/record/7223753/files/figure.png" pageId="7" pageNumber="456">Figs. 2B, E</figureCitation>
;
<figureCitation id="D89A3EF0FFCCCB43FB714220FB0B6233" box="[1173,1235,1375,1396]" captionStart="FIGURE 7" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1755]" captionTargetId="figure-17@11.[151,1435,181,1755]" captionTargetPageId="11" captionText="FIGURE 7. Corallocoris marksae, non-type specimen, female: A–C, habitus; A, dorsal view; B, lateral view; C, ventral view; D, head in frontal view; E, head and pronotum in dorsal view; F, light fascia on right forewing; G, labels." figureDoi="http://doi.org/10.5281/zenodo.7223767" httpUri="https://zenodo.org/record/7223767/files/figure.png" pageId="7" pageNumber="456">7D, E</figureCitation>
); base of hypocostal lamina clearly without notched along ventral side............................................................ 6
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<keyLead id="FB508140FFCCCB43FF7342E6FA44628B" pageId="7" pageNumber="456">
6. Forewing with distinct grey transverse fascia and grey apical margin (
<figureCitation id="D89A3EF0FFCCCB43FC9B42E6FBD262E9" box="[895,1034,1433,1455]" captionStart="FIGURE 7" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1755]" captionTargetId="figure-17@11.[151,1435,181,1755]" captionTargetPageId="11" captionText="FIGURE 7. Corallocoris marksae, non-type specimen, female: A–C, habitus; A, dorsal view; B, lateral view; C, ventral view; D, head in frontal view; E, head and pronotum in dorsal view; F, light fascia on right forewing; G, labels." figureDoi="http://doi.org/10.5281/zenodo.7223767" httpUri="https://zenodo.org/record/7223767/files/figure.png" pageId="7" pageNumber="456">Figs. 7A–B, F</figureCitation>
); entire trochanters and femora except for apices dark-brown (
<figureCitation id="D89A3EF0FFCCCB43FE4542C9FDDE628B" box="[417,518,1462,1484]" captionStart="FIGURE 7" captionStartId="11.[151,250,1779,1803]" captionTargetBox="[151,1435,181,1755]" captionTargetId="figure-17@11.[151,1435,181,1755]" captionTargetPageId="11" captionText="FIGURE 7. Corallocoris marksae, non-type specimen, female: A–C, habitus; A, dorsal view; B, lateral view; C, ventral view; D, head in frontal view; E, head and pronotum in dorsal view; F, light fascia on right forewing; G, labels." figureDoi="http://doi.org/10.5281/zenodo.7223767" httpUri="https://zenodo.org/record/7223767/files/figure.png" pageId="7" pageNumber="456">Fig. 7B–C</figureCitation>
) (
<collectingCountry id="38B662E5FFCCCB43FDFE42C9FDAE628B" box="[538,630,1462,1484]" name="Australia" pageId="7" pageNumber="456">Australia</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FD6442C9FD37628B" box="[640,751,1462,1484]" name="Philippines" pageId="7" pageNumber="456">Philippines</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FD1D42C8FCE5628B" box="[761,829,1463,1484]" name="Samoa" pageId="7" pageNumber="456">Samoa</collectingCountry>
;
<collectingCountry id="38B662E5FFCCCB43FCA342C9FC72628B" box="[839,938,1462,1484]" name="Singapore" pageId="7" pageNumber="456">Singapore</collectingCountry>
)................... ..
<taxonomicName id="87A159F6FFCCCB43FB4A42C9FA44628B" authority="(Woodward)" baseAuthorityName="Woodward" box="[1198,1436,1462,1484]" class="Insecta" family="Omaniidae" genus="Omania" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="marksae">
<emphasis id="72D5FE67FFCCCB43FB4A42C9FAC6628B" bold="true" box="[1198,1310,1462,1484]" italics="true" pageId="7" pageNumber="456">C. marksae</emphasis>
(Woodward)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="401E2275FFCCCB43FF7342ABFA446141" blockId="7.[151,1437,817,1543]" pageId="7" pageNumber="456">
<keyLead id="FB508140FFCCCB43FF7342ABFA446141" pageId="7" pageNumber="456">
– Forewing nearly concolorous, with apical margin paler (
<figureCitation id="D89A3EF0FFCCCB43FCE542ABFC7C62AE" box="[769,932,1492,1514]" captionStart="FIGURE 1" captionStartId="2.[151,250,1810,1834]" captionTargetBox="[151,1435,376,1787]" captionTargetId="figure-38@2.[151,1435,376,1787]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Corallocoris xishaensis sp. nov.: A–C, male holotype; D–F, female paratype. A, dorsal view; B, lateral view; C, ventral view; D, dorsal view; E, lateral view; F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223749" httpUri="https://zenodo.org/record/7223749/files/figure.png" pageId="7" pageNumber="456">Fig. 1A–B, D–E</figureCitation>
); entire trochanters and femora except for subapices whitish-yellow (
<figureCitation id="D89A3EF0FFCCCB43FE9B428EFDC26141" box="[383,538,1521,1543]" captionStart="FIGURE 1" captionStartId="2.[151,250,1810,1834]" captionTargetBox="[151,1435,376,1787]" captionTargetId="figure-38@2.[151,1435,376,1787]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Corallocoris xishaensis sp. nov.: A–C, male holotype; D–F, female paratype. A, dorsal view; B, lateral view; C, ventral view; D, dorsal view; E, lateral view; F, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7223749" httpUri="https://zenodo.org/record/7223749/files/figure.png" pageId="7" pageNumber="456">Fig. 1B–C, E–F</figureCitation>
) (
<collectingCountry id="38B662E5FFCCCB43FDC9428EFDB36140" box="[557,619,1521,1543]" name="China" pageId="7" pageNumber="456">China</collectingCountry>
: Xisha Islands).....................................
<emphasis id="72D5FE67FFCCCB43FB23428EFA446141" bold="true" box="[1223,1436,1521,1543]" pageId="7" pageNumber="456">
<taxonomicName id="87A159F6FFCCCB43FB23428EFA926140" authority="Luo &amp; Wang &amp; Chen &amp; Xie, 2022" authorityName="Luo &amp; Wang &amp; Chen &amp; Xie" authorityYear="2022" box="[1223,1354,1521,1543]" class="Insecta" family="Omaniidae" genus="Corallocoris" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="456" phylum="Arthropoda" rank="species" species="xishaensis" status="sp. nov.">
<emphasis id="72D5FE67FFCCCB43FB23428EFA926140" bold="true" box="[1223,1354,1521,1543]" italics="true" pageId="7" pageNumber="456">C. xishaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="69E6431CFFCCCB43FAB4428EFA446141" box="[1360,1436,1521,1542]" pageId="7" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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