208 lines
23 KiB
XML
208 lines
23 KiB
XML
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<mods:title id="7EC4807652B50BF5C0698A9CEA5B7D2C">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="3429DF1921BF45263AD2E6FE529DAE1B">Lackner, Tomas</mods:namePart>
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<mods:namePart id="78FA063D0401F998ADC58F2937BCD51B">Leschen, Richard A. B.</mods:namePart>
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<treatment id="BE685745BE55312A3B4EC3E981B8FFAB" ID-GBIF-Taxon="132648719" LSID="urn:lsid:plazi:treatment:BE685745BE55312A3B4EC3E981B8FFAB" httpUri="http://treatment.plazi.org/id/BE685745BE55312A3B4EC3E981B8FFAB" lastPageId="39" lastPageNumber="40" pageId="34" pageNumber="35">
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<subSubSection id="161BD8E8FEF9B2B71D25E0775B8C5E82" pageId="34" pageNumber="35" type="nomenclature">
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<paragraph id="C7C01514F037781586B45E5572813AF7" pageId="34" pageNumber="35">
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<taxonomicName id="6371CED1A4E611FE3265B2717B80F48C" ID-CoL="7VHT5" authority="Marseul, 1862" authorityName="Marseul" authorityYear="1862" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae (Marseul, 1862)</taxonomicName>
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Figs 161, 162-173, 174-182, 756
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</paragraph>
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</subSubSection>
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<subSubSection id="1584A92E43AAAACC5BD8BDD8896A429A" pageId="35" pageNumber="36" type="reference_group">
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<paragraph id="EA8C05A0377D4884842932CEFA1FBB00" pageId="35" pageNumber="36">
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<taxonomicName id="F9A707A5AD02CC7492CDFBC69233897B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus sinae" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sinae">
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<pageBreakToken id="60FF339E161CF5F6981268BC08AFE290" pageId="35" pageNumber="36" start="start">Saprinus</pageBreakToken>
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sinae
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</taxonomicName>
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Marseul, 1862: 496.
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</paragraph>
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<paragraph id="8FD54A0B5A03D27855C7F5AD30C4F781" pageId="35" pageNumber="36">
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<taxonomicName id="083876B8807E4AB224334474D74A59D3" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus vernulus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="vernulus">Saprinus vernulus</taxonomicName>
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Blackburn, 1903: 108 - syn. n.
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</paragraph>
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</subSubSection>
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<subSubSection id="E4EE38E20FE45C95927554D07E1F8D9F" pageId="35" pageNumber="36" type="type locality">
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<paragraph id="C0D57A1BE89DEA969F8E3759A2DD0ABF" pageId="35" pageNumber="36">Type locality.</paragraph>
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<paragraph id="CB361918257EC66E8B87F638FAC9E75B" pageId="35" pageNumber="36">China: Shanghai.</paragraph>
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<caption id="8D96B433584F7A7701993E602118BF68" pageId="35" pageNumber="36">
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<paragraph id="CDCEDFBF3A272A685A7FEAEED1552D36" pageId="35" pageNumber="36">
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Figure 161.
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<taxonomicName id="5523A04774C38443AB218E4C9F6A9560" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae</taxonomicName>
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(Marseul, 1862) habitus, dorsal view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="68154B7DB9062D44E672FF8FFEFB827C" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="type material">
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<paragraph id="2CAE45C1F8888130D92FA641F5DD0374" pageId="35" pageNumber="36">Type material examined.</paragraph>
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<paragraph id="55A3A3ECC06D666C656982E7CF544D33" pageId="35" pageNumber="36">
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<taxonomicName id="D547AADB7C362134344F947C7C9248C4" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus sinae" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sinae">Saprinus sinae</taxonomicName>
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Marseul, 1862: Lectotype, present designation: ♂, mounted on tip of mounting card, with genitalia extracted, dismembered and placed in a separate plastic slide under the specimen, right antennal club, right protarsus and right hind-leg missing, with the following labels: tiny, dark blue rectangular label, followed by: "
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<taxonomicName id="95956C03994B8B1AFD757E9BEF27EF3C" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="1B2A4A09541A7F146C0416845FEDFDCF" lsidName="Sinae" pageId="35" pageNumber="36" rank="species" species="Sinae">Sinae</taxonomicName>
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m. / Shanghai / Dej. 63" (round, yellow label, written); followed by:
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<normalizedToken id="5FE78696019297D4ECA31FBA01EAF710" originalValue="“Shanghai”">"Shanghai"</normalizedToken>
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(written); followed by: "52 (158a)
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<taxonomicName id="7EE8DF7F16FAEB1A128D1C60C3F289E1" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
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/
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<taxonomicName id="DB6C34AB4DC17D5A23B20FF0AEB9BADF" lsidName="Sinae" pageId="35" pageNumber="36" rank="species" species="Sinae">Sinae</taxonomicName>
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m.60 / Shanghai" (written); followed by: "MUSEUM PARIS / COLL. / DE MARSEUL 1890" (printed); followed by:
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<normalizedToken id="823EF539F348193D357009E7205D06B3" originalValue="“TYPE”">"TYPE"</normalizedToken>
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(red-printed label); followed by: "
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<taxonomicName id="2437BB3903D599604CFA518F44E0D38E" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus sinae" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sinae">Saprinus sinae</taxonomicName>
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/ Marseul, 1862 / LECTOTYPE 2014 / des. T. Lackner" (red label, written) (MNHN). This species was described from unknown number of specimens and the lectotype designation fixes the identity of the species.
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</paragraph>
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<caption id="A0B9801D50BB7B4AAA0B05334A432E15" pageId="35" pageNumber="36">
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<paragraph id="EE6C5A466B6986A1CB602506C2DDC038" pageId="35" pageNumber="36">
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Figures 162-173. 162
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<taxonomicName id="673AB68AF69FD31FA9D15A00E841BC76" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae</taxonomicName>
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(Marseul, 1862) head, dorsal view 163 antennal club, dorsal view 164 ditto, ventral view 165 mentum, ventral view 166 propygidium + pygidium 167 prosternum 168 mesoventrite 169 lateral disc of metaventrite + metepisternum 170 protibia, dorsal view 171 ditto, ventral view 172 mesotibia, ventral view 173 metatibia, ventral view.
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</paragraph>
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</caption>
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<paragraph id="1D8363645B431CED64DFE708F83AD741" pageId="36" pageNumber="37">
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<taxonomicName id="8BEBA70AAB06039AEFA3049222F5454D" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus vernulus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vernulus">
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<pageBreakToken id="EE5AEB5FE08C9327B7E5CF562BE5137F" pageId="36" pageNumber="37" start="start">Saprinus</pageBreakToken>
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vernulus
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</taxonomicName>
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Blackburn, 1903: Lectotype, present designantion: ♀, mounted on a card with
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<normalizedToken id="0162706F1CE8D85A2BBCBE70A9C9101C" originalValue="“♀”">"♀"</normalizedToken>
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(printed), followed by:
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<normalizedToken id="EE143D2F5C104770F17B3C295E1D3772" originalValue="“7249”">"7249"</normalizedToken>
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(mount card with a red number, printed); followed by: "N.S. Wales" (printed); followed by: "K. 270214" (printed); followed by: "
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<taxonomicName id="50D2A61705F59C3F9749A2AE593B305B" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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/
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<taxonomicName id="03C6D3034D8A54D5994CFBC083BC49F1" lsidName="vernulus" pageId="36" pageNumber="37" rank="species" species="vernulus">vernulus</taxonomicName>
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/ Cotype" (written); followed by: "
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<taxonomicName id="CCFF761A3ABB311B3D042D48A71C2CCC" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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/
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<taxonomicName id="B60015ABB49F8ED3EDBF2E1202BCC786" lsidName="vernulus" pageId="36" pageNumber="37" rank="species" species="vernulus">vernulus</taxonomicName>
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Bl." (written); followed by:
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<normalizedToken id="8B7664726DBDCA880E0651A83D14CB04" originalValue="“SYNTYPE”">"SYNTYPE"</normalizedToken>
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(yellow label, written); followed by: "
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<taxonomicName id="B368358F8174719E74A78867C3725F53" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus vernulus" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="vernulus">Saprinus vernulus</taxonomicName>
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/ Blackburn, 1903 / LECTOTYPE des. / T. Lackner 2016" (red label, written) (AMS). This specimen has been described based on unknown number of specimens and the lectotype designation fixes the identity of the species.
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</paragraph>
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<caption id="6C9FA8394DCBE3D955A1B7E9A6319C91" pageId="36" pageNumber="37">
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<paragraph id="30B6E3904CD3F35577BBD7F88436DE8A" pageId="36" pageNumber="37">
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Figures 174-182. 174
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<taxonomicName id="8ADD5DD9B5E11C7407214A9FC8BAA859" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae</taxonomicName>
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(Marseul, 1862) male terminalia: 8th sternite + 8th tergite, ventral view 175 ditto, dorsal view 176 ditto, lateral view 177 male terminalia: 9th + 10th tergites, dorsal view 178 ditto, lateral view 179 male terminalia: spiculum gastrale, ventral view 180 ditto, lateral view 181 male terminalia: aedeagus, dorsal view 182 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="FF048532650A781787DDD93FAE64DFD7" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="materials_examined">
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<paragraph id="F7AF28C869E52941FE5AB3CE70F76231" pageId="36" pageNumber="37">Additional material examined.</paragraph>
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<paragraph id="6E0C8A27D9D80B6B2AE67D63E24D4BEC" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
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AUSTRALIA. New South Wales: 2 specs., re-mounted on a single pin, with the same labels as the lectotype, (and certainly from the same collection - D. Smith pers. comm.) except the original mount with red number
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<normalizedToken id="C8ED34F02D90812AEF449793708F93EF" originalValue="“7248”">"7248"</normalizedToken>
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instead of
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<normalizedToken id="CAB7B638461506E3305D5B521F45BF90" originalValue="“7249”">"7249"</normalizedToken>
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and with
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<normalizedToken id="3C54DA5C1963F5B726E0BC787B36C564" originalValue="“K270229”">"K270229"</normalizedToken>
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instead of "K. 270214" of the lecto
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<pageBreakToken id="EC9E302362B52CB69C8B191B9806A47A" pageId="37" pageNumber="38" start="start">type</pageBreakToken>
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(AMS). 2 ♂♂, without further data (MAMU); 9 specs., New South Wales, without further data (SAMA); 3 specs., Sydney, Lea (SAMA). South Australia: 3 specs., Adelaide, Griffith (SAMA).
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</paragraph>
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</subSubSection>
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<subSubSection id="44AA2421A18112A2B691B80925076999" pageId="37" pageNumber="38" type="biology_ecology">
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<paragraph id="4B664C382B66B056603D652A6C19B394" pageId="37" pageNumber="38">Biology.</paragraph>
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<paragraph id="7C8AC34CEE3DB7256274CBEE1AB3B856" pageId="37" pageNumber="38">A typical littoral psammophile found on beaches.</paragraph>
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</subSubSection>
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<subSubSection id="A345B83519CB5A5DB440E23C9CDBE632" pageId="38" pageNumber="39" type="distribution">
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||
<paragraph id="EA959CE6029ADF5737405EF91B388F76" pageId="38" pageNumber="39">
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<pageBreakToken id="F607AE95AF7492CAA4F12617543DDFC5" pageId="38" pageNumber="39" start="start">Distribution</pageBreakToken>
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.
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</paragraph>
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<paragraph id="E3D41FF10288FC67ADE6132761801431" pageId="38" pageNumber="39">
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Described from Shanghai (China), distributed along the coasts of China, Indonesia, Russian Far East, Thailand, Vietnam, Cambodia, Myanmar, Japan, Korea, Malaysia, Australia (
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<bibRefCitation id="59696E51AF2D69B28BDCCB6EBA322F15" pageId="38" pageNumber="39">Mazur 2011</bibRefCitation>
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||
; present paper). Reported also from Afghanistan by
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<bibRefCitation id="FB4EAAEA6D50292587B7808103BBA0BD" author="Therond, J" journalOrPublisher="Revue Francaise d'Entomologie" pageId="218" pageNumber="219" pagination="62 - 65" title="Contribution a la connaissance de la faune de l'Afghanistan. 54. Coleoptera - Histeridae." volume="29" year="1969">
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<normalizedToken id="95D142C606C6CD540CE4158FA75927DA" originalValue="Thérond">Therond</normalizedToken>
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||
(1969)
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</bibRefCitation>
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together with the preceding species. From the Australopacific Region found only in Australia (New South Wales and South Australia; Fig. 756).
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</paragraph>
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</subSubSection>
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<subSubSection id="F176C10F1B800E93345FCC5D817A9DCB" pageId="39" pageNumber="40" type="remarks">
|
||
<paragraph id="58ED0271C09F19D15F86100F34A61C5E" pageId="39" pageNumber="40">
|
||
<pageBreakToken id="C3EF825C9E3B58A4F94CFDD3FD5BB108" pageId="39" pageNumber="40" start="start">Remarks</pageBreakToken>
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||
.
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||
</paragraph>
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||
<paragraph id="295F0905C4835CEF73CD37C64DC3C477" pageId="39" pageNumber="40">
|
||
<bibRefCitation id="E4A7A7D3D8EDE0295B8B2C094068DAF5" author="Blackburn, T" journalOrPublisher="Transactions and proceedings of the Royal Society of South Australia" pageId="213" pageNumber="214" pagination="91 - 182" title="Further notes on Australian Coleoptera, with descriptions of new genera and species." volume="27" year="1903">Blackburn (1903</bibRefCitation>
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||
: 108) mentioned 'distinctly green colour in combination with elytra having a very strong external subhumeral
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<normalizedToken id="D9140BC24D86BBBDC3FF3C1E2D16E513" originalValue="stria’">stria'</normalizedToken>
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||
as the main distinguishing characters between
|
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<taxonomicName id="A6F837AF006F6D1529C9977E22A6E69B" lsidName="H. vernulus" pageId="39" pageNumber="40" rank="species" species="vernulus">H. vernulus</taxonomicName>
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and
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<taxonomicName id="1FE2304815555CC290D395AC1A5EDD1B" lsidName="H. sinae" pageId="39" pageNumber="40" rank="species" species="sinae">H. sinae</taxonomicName>
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. Since the external morphological characters of many
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<taxonomicName id="B93D437A9AE35D284DE5B52042CFC379" lsidName="" pageId="39" pageNumber="40" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
|
||
species are known to be intra-specifically rather variable, we assign the characters used by Blackburn for the separation of
|
||
<taxonomicName id="F29D483EEC8404C1EC5AA5F32D8341EB" lsidName="H. vernulus" pageId="39" pageNumber="40" rank="species" species="vernulus">H. vernulus</taxonomicName>
|
||
from
|
||
<taxonomicName id="A7AC2F985810051E2772ED2CE5EE8FE7" lsidName="H. sinae" pageId="39" pageNumber="40" rank="species" species="sinae">H. sinae</taxonomicName>
|
||
as intra-specific variation and synonymize the two species. A single syntype of
|
||
<taxonomicName id="A03084F65C3DB3523E917BC7C607949F" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus vernulus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vernulus">Saprinus vernulus</taxonomicName>
|
||
Blackburn, 1903 (currently
|
||
<taxonomicName id="73273DD0E4F0910D5B831AE22FB6394F" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) vernulus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vernulus" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) vernulus</taxonomicName>
|
||
) housed at ANIC has been inspected and we can herewith conclude that it is conspecific with the species
|
||
<taxonomicName id="5B25DF09035B4AD516C157B691802599" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae</taxonomicName>
|
||
(Marseul, 1862); thus
|
||
<taxonomicName id="713EB694FB89794B99BAF2A4959E9593" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) vernulus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="vernulus" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) vernulus</taxonomicName>
|
||
Blackburn, 1903 =
|
||
<taxonomicName id="9AE99439B08D9660AB2D0EF03552F562" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) sinae" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinae" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) sinae</taxonomicName>
|
||
(Marseul, 1862) syn. n.
|
||
<taxonomicName id="82CD1FDDC390F1A4F3157742FC7D0295" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) sinae" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinae" subGenus="H.">Hypocaccus (H.) sinae</taxonomicName>
|
||
is above included in the key to the Australopacific species of
|
||
<taxonomicName id="B505EFC03412EA8A5EE93BE771DD0E61" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
|
||
that separates it from the related
|
||
<taxonomicName id="E851C454D266CE09091A315E762DB440" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) brasiliensis" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="H.">Hypocaccus (H.) brasiliensis</taxonomicName>
|
||
. We chose not to fully re-describe
|
||
<taxonomicName id="BCA873BB222F5DAAE1FA6C195E18A781" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) sinae" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="sinae" subGenus="H.">Hypocaccus (H.) sinae</taxonomicName>
|
||
here, leaving its re-description to the revision of the genus
|
||
<taxonomicName id="02CB955B041436C107C58C99E845B6A7" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
|
||
. For the sake of the better species recognition, however, we decided to depict it here, including its male terminalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |