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<document id="1963227BEB8429A978382D87BD20E51E" ID-DOI="10.11646/zootaxa.4007.2.9" ID-GBIF-Dataset="55ee7eb4-65b4-4fbc-80eb-ee9e32e477dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="236500" ID-ZooBank="60ABECAF-9687-4172-A309-D2222DFEC473" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461218491837" checkinUser="plazi" docAuthor="Kim, Su-Hwan, Kim, Hyeong-Su &amp; Park, Jong-Young" docDate="2015" docId="492087BFFFE37969A7E8F8EDFEA9C59A" docLanguage="en" docName="zt04007p275.pdf" docOrigin="Zootaxa 4007 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liobagrus hyeongsanensis Kim, Kim &amp; Park, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="270" masterDocId="B519FFC7FFE3796AA77FFFC3FFE7C350" masterDocTitle="A new species of torrent catfish, Liobagrus hyeongsanensis (Teleostei: Siluriformes: Amblycipitidae), from Korea" masterLastPageNumber="275" masterPageNumber="267" pageNumber="267" updateTime="1698603186691" updateUser="plazi">
<mods:mods id="C627C48BCC839EA52516EB55ADA5DB2C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EFC816EF8C5F4D86D208685BDC452961">A new species of torrent catfish, Liobagrus hyeongsanensis (Teleostei: Siluriformes: Amblycipitidae), from Korea</mods:title>
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<mods:namePart id="65206C484E0E7D059BD71B86F02A23C1">Kim, Su-Hwan</mods:namePart>
</mods:name>
<mods:name id="ED63D7DA99801CA3E0CB3E240212E4A7" type="personal">
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<mods:namePart id="29F19AA3A0649F5B9167DD209512F6FA">Kim, Hyeong-Su</mods:namePart>
</mods:name>
<mods:name id="24C7CF817B611CA7EE86113630F28EF7" type="personal">
<mods:role id="D79D3A5D5E30D6F58D99B9867925FD27">
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</mods:role>
<mods:namePart id="F0BFBA593B6FBBD11DC165162F9534F7">Park, Jong-Young</mods:namePart>
</mods:name>
<mods:typeOfResource id="6D5C592C01899D0B6A10A6A0288963CF">text</mods:typeOfResource>
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<mods:title id="83A577C9D24167A45B95A4C66959D306">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3D2028FF24D779C4D75E87C4384E3A7D">
<mods:date id="200957E65778B425C5838C3186AA4041">2015</mods:date>
<mods:detail id="75FE93BAFB5FF28C00E1E593646DA182" type="volume">
<mods:number id="D0BB987C4E6FB4F73E4C3C3E6A30D5EA">4007</mods:number>
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<mods:number id="CFC728FCCB10BFA171B81A10838DD06A">1</mods:number>
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<mods:start id="7381B94C508482C2080FB87EA62C6EEF">267</mods:start>
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<mods:classification id="5E9DE5D6DBF9AEE4F5061BFE64DF3187">journal article</mods:classification>
<mods:identifier id="9627ABCDC04DA807A1BC3705D13E413D" type="DOI">10.11646/zootaxa.4007.2.9</mods:identifier>
<mods:identifier id="3B4BC57D7582ADC98CD01ABDC0BF09F8" type="GBIF-Dataset">55ee7eb4-65b4-4fbc-80eb-ee9e32e477dc</mods:identifier>
<mods:identifier id="1CD1598440E955821837E38C151AB69D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A63E1A6FF40A1142547729D4A878B861" type="Zenodo-Dep">236500</mods:identifier>
<mods:identifier id="0DE5BD99C580415801A4466BB3E4E2C3" type="ZooBank">60ABECAF-9687-4172-A309-D2222DFEC473</mods:identifier>
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<treatment id="492087BFFFE37969A7E8F8EDFEA9C59A" ID-DOI="http://doi.org/10.5281/zenodo.5621746" ID-GBIF-Taxon="119631238" ID-Zenodo-Dep="5621746" LSID="urn:lsid:plazi:treatment:492087BFFFE37969A7E8F8EDFEA9C59A" httpUri="http://treatment.plazi.org/id/492087BFFFE37969A7E8F8EDFEA9C59A" lastPageId="3" lastPageNumber="270" pageId="0" pageNumber="267">
<subSubSection id="89936522FFE3796AA7E8F8EDFF0CC43A" pageId="0" pageNumber="267" type="nomenclature">
<paragraph id="C13636A9FFE3796AA7E8F8EDFDDAC419" blockId="0.[151,573,1838,1899]" box="[151,573,1838,1865]" pageId="0" pageNumber="267">
<heading id="9A7E81C5FFE3796AA7E8F8EDFDDAC419" bold="true" box="[151,573,1838,1865]" fontSize="11" level="1" pageId="0" pageNumber="267" reason="1">
<emphasis id="F3FDEABBFFE3796AA7E8F8EDFDDAC419" bold="true" box="[151,573,1838,1865]" pageId="0" pageNumber="267">
<taxonomicName id="06894D2AFFE3796AA7E8F8EDFE3FC418" ID-CoL="6QHDZ" box="[151,472,1838,1864]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="0" pageNumber="267" phylum="Chordata" rank="species" species="hyeongsanensis" status="sp. nov.">
<emphasis id="F3FDEABBFFE3796AA7E8F8EDFE3FC418" bold="true" box="[151,472,1838,1864]" italics="true" pageId="0" pageNumber="267">Liobagrus hyeongsanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8CE57C0FFE3796AA69FF8ECFDDAC419" box="[480,573,1839,1865]" pageId="0" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C13636A9FFE3796AA7E8F891FF0CC43A" blockId="0.[151,573,1838,1899]" box="[151,235,1874,1899]" pageId="0" pageNumber="267">
(
<figureCitation id="59B22A2CFFE3796AA7E0F891FF07C43A" box="[159,224,1874,1899]" captionStart="FIGURE 1" captionStartId="1.[151,250,1288,1310]" captionTargetBox="[340,1246,177,1265]" captionTargetId="figure@1.[340,1247,177,1266]" captionTargetPageId="1" captionText="FIGURE 1. Liobagrus hyeongsanensis, sp. nov., CNUC 38547, holotype, 84.1 mm SL, Hyeongsan River, South Korea." httpUri="https://zenodo.org/record/236501/files/figure.png" pageId="0" pageNumber="267">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89936522FFE3796BA7E8F859FD02C6FB" lastPageId="1" lastPageNumber="268" pageId="0" pageNumber="267" type="materials_examined">
<paragraph id="C13636A9FFE3796AA7E8F859FB65C486" blockId="0.[151,1436,1946,2006]" pageId="0" pageNumber="267">
<emphasis id="F3FDEABBFFE3796AA7E8F859FEADC4E3" bold="true" box="[151,330,1946,1971]" pageId="0" pageNumber="267">
<typeStatus id="1E32880BFFE3796AA7E8F859FF35C4E3" box="[151,210,1946,1971]" pageId="0" pageNumber="267">Type</typeStatus>
material.
</emphasis>
<typeStatus id="1E32880BFFE3796AA629F859FE25C4E3" box="[342,450,1946,1971]" pageId="0" pageNumber="267" type="holotype">Holotype</typeStatus>
:
<collectionCode id="A798AE6CFFE3796AA6ACF859FDC6C4E2" box="[467,545,1946,1970]" pageId="0" pageNumber="267">CNUC</collectionCode>
38547,
<quantity id="06719B4CFFE3796AA5F9F859FD17C4E2" box="[646,752,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="8.41" pageId="0" pageNumber="267" unit="mm" value="84.1">84.1 mm</quantity>
standard length (SL); Hyeongsan River, Yangbuk-myeon, Gyeongju-si,
<collectingCountry id="B99E7639FFE3796AA650F87EFE58C486" box="[303,447,1981,2006]" name="South Korea" pageId="0" pageNumber="267">South Korea</collectingCountry>
,
<geoCoordinate id="A4BD506EFFE3796AA6B5F87EFDA9C486" box="[458,590,1981,2006]" direction="north" orientation="latitude" pageId="0" pageNumber="267" precision="15" value="35.832775">35°49'58&quot;N</geoCoordinate>
<geoCoordinate id="A4BD506EFFE3796AA52AF87DFD01C486" box="[597,742,1982,2006]" direction="east" orientation="longitude" pageId="0" pageNumber="267" precision="15" value="129.41138">129°24'41&quot;E</geoCoordinate>
; J.
<collectingCountry id="B99E7639FFE3796AA47DF87CFCFFC486" box="[770,792,1983,2006]" name="Italy" pageId="0" pageNumber="267">Y</collectingCountry>
. Park &amp; S.H. Kim,
<date id="B5371069FFE3796AA48CF87DFB99C486" box="[1011,1150,1982,2006]" pageId="0" pageNumber="267" value="2013-06-02">2 June 2013</date>
.
</paragraph>
<caption id="95F66621FFE2796BA7E8FACBFABCC64E" box="[151,1371,1288,1310]" httpUri="https://zenodo.org/record/236501/files/figure.png" pageId="1" pageNumber="268" targetBox="[340,1246,177,1265]" targetPageId="1">
<paragraph id="C13636A9FFE2796BA7E8FACBFABCC64E" blockId="1.[151,1371,1288,1310]" box="[151,1371,1288,1310]" pageId="1" pageNumber="268">
<emphasis id="F3FDEABBFFE2796BA7E8FACBFEF4C64D" bold="true" box="[151,275,1288,1310]" pageId="1" pageNumber="268">FIGURE 1.</emphasis>
<emphasis id="F3FDEABBFFE2796BA666FACBFDCAC64E" box="[281,557,1288,1310]" italics="true" pageId="1" pageNumber="268">
<taxonomicName id="06894D2AFFE2796BA666FACBFDCFC64E" box="[281,552,1288,1310]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="hyeongsanensis" status="sp. nov.">Liobagrus hyeongsanensis</taxonomicName>
,
</emphasis>
<emphasis id="F3FDEABBFFE2796BA54CFACBFD6DC64D" bold="true" box="[563,650,1288,1309]" pageId="1" pageNumber="268">
<taxonomicNameLabel id="E8CE57C0FFE2796BA54CFACBFD62C64D" box="[563,645,1288,1309]" pageId="1" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
CNUC 38547, holotype, 84.1 mm SL, Hyeongsan River, South Korea.
</paragraph>
</caption>
<paragraph id="C13636A9FFE2796BA7B8FA88FBABC633" blockId="1.[151,1436,1354,2027]" box="[199,1100,1354,1379]" pageId="1" pageNumber="268">
<typeStatus id="1E32880BFFE2796BA7B8FA88FEDFC632" box="[199,312,1355,1378]" pageId="1" pageNumber="268" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="A798AE6CFFE2796BA63DFA88FE77C633" box="[322,400,1355,1379]" pageId="1" pageNumber="268">CNUC</collectionCode>
38548–38567 (20),
<quantity id="06719B4CFFE2796BA50AFA89FCFFC632" box="[629,792,1354,1379]" metricMagnitude="-2" metricUnit="m" metricValue="7.06" metricValueMax="8.41" metricValueMin="5.71" pageId="1" pageNumber="268" unit="mm" value="70.6" valueMax="84.1" valueMin="57.1">57.1–84.1 mm</quantity>
SL; same data as
<typeStatus id="1E32880BFFE2796BA49DFA89FBAFC633" box="[994,1096,1354,1379]" pageId="1" pageNumber="268" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="C13636A9FFE2796BA7B8FAACFD02C6FB" blockId="1.[151,1436,1354,2027]" pageId="1" pageNumber="268">
Nontypes:
<collectionCode id="A798AE6CFFE2796BA63CFAACFE76C6D7" box="[323,401,1391,1415]" pageId="1" pageNumber="268">CNUC</collectionCode>
38830–38854 (25),
<quantity id="06719B4CFFE2796BA503FAADFCC6C6D6" box="[636,801,1390,1415]" metricMagnitude="-2" metricUnit="m" metricValue="5.4799999999999995" metricValueMax="6.779999999999999" metricValueMin="4.18" pageId="1" pageNumber="268" unit="mm" value="54.8" valueMax="67.8" valueMin="41.8">41.8–67.8 mm</quantity>
SL; Bukcheon River, Hwangnyong-dong, Gyeongju-si,
<collectingCountry id="B99E7639FFE2796BA7E8FA51FEC0C6FA" box="[151,295,1426,1451]" name="South Korea" pageId="1" pageNumber="268">South Korea</collectingCountry>
; J.
<collectingCountry id="B99E7639FFE2796BA63DFA50FEBFC6FA" box="[322,344,1427,1450]" name="Italy" pageId="1" pageNumber="268">Y</collectingCountry>
. Park &amp; S.H. Kim,
<date id="B5371069FFE2796BA54CFA50FD05C6FB" box="[563,738,1426,1451]" pageId="1" pageNumber="268" value="2010-03-28">28 March 2010</date>
.
</paragraph>
</subSubSection>
<subSubSection id="89936522FFE2796BA7B8FA75FC7FC5F7" pageId="1" pageNumber="268" type="diagnosis">
<paragraph id="C13636A9FFE2796BA7B8FA75FC7FC5F7" blockId="1.[151,1436,1354,2027]" pageId="1" pageNumber="268">
<emphasis id="F3FDEABBFFE2796BA7B8FA75FEDDC69F" bold="true" box="[199,314,1462,1487]" pageId="1" pageNumber="268">Diagnosis</emphasis>
.
<taxonomicName id="06894D2AFFE2796BA638FA75FD96C69E" box="[327,625,1462,1486]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="268" phylum="Chordata" rank="species" species="hyeongsanensis">
<emphasis id="F3FDEABBFFE2796BA638FA75FD96C69E" box="[327,625,1462,1486]" italics="true" pageId="1" pageNumber="268">Liobagrus hyeongsanensis</emphasis>
</taxonomicName>
is distinguished from all its congeners by a small size (a maximum of
<quantity id="06719B4CFFE2796BA2FFFA74FF24C6A2" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="268" unit="mm" value="90.0">90 mm</quantity>
SL, vs.
<quantity id="06719B4CFFE2796BA652FA19FE2CC6A3" box="[301,459,1498,1523]" metricMagnitude="-1" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="1" pageNumber="268" unit="mm" value="165.0" valueMax="180.0" valueMin="150.0">150–180 mm</quantity>
); a smaller number of eggs per gravid female (60–110 vs.130–210); and a smaller number of serrations on the pectoral fin (2–3, mostly 3, vs. 3–6 or 0–3, smaller or vestigial); a slenderer body width at pectoral-fin base (15.5–17.9 % SL, vs. 17.1–21.9%); a relatively short pectoral spine (3.7–6.5 % SL, vs. 6.8– 13.1%). The lower jaw is shorter than the upper jaw (vs. longer or equal), and the body and fins are entirely brownish yellow without any other markings (vs. a vertical broad yellowish crescent-shaped band on the caudal fin or black or whitish yellow outer margins of the dorsal and anal fins).
</paragraph>
</subSubSection>
<subSubSection id="89936522FFE27969A7B8F971FE34C3A8" lastPageId="3" lastPageNumber="270" pageId="1" pageNumber="268" type="description">
<paragraph id="C13636A9FFE2796BA7B8F971FBD2C42F" blockId="1.[151,1436,1354,2027]" pageId="1" pageNumber="268">
<emphasis id="F3FDEABBFFE2796BA7B8F971FEBEC59B" bold="true" box="[199,345,1714,1739]" pageId="1" pageNumber="268">Description.</emphasis>
Meristic and morphometric data are provided in
<tableCitation id="8C0B0312FFE2796BA4F0F971FC01C59B" box="[911,998,1714,1739]" captionStart="TABLE 1" captionStartId="2.[151,239,254,277]" captionTargetBox="[159,1411,378,648]" captionTargetPageId="2" captionText="TABLE 1. Morphometric and meristic data of Liobagrus hyeongsanensis (CNUC 38547 – 38567) and L. mediadiposalis (CNUC 37821 – 37838). H, holotype." httpUri="http://table.plazi.org/id/95F66621FFE17968A7E8FF3DFDF9C265" pageId="1" pageNumber="268" tableUuid="95F66621FFE17968A7E8FF3DFDF9C265">Table 1</tableCitation>
. Asterisks indicate meristic values for
<typeStatus id="1E32880BFFE2796BA7E8F915FF1AC5BF" box="[151,253,1750,1775]" pageId="1" pageNumber="268" type="holotype">holotype</typeStatus>
. Maximun size not exceeding
<quantity id="06719B4CFFE2796BA530F914FD79C5BE" box="[591,670,1751,1775]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="268" unit="mm" value="90.0">90 mm</quantity>
standard length. Body thin, compressed, and round; head depressed, caudal peduncle vertically depressed; dorsal and ventral profiles of body straight. Predorsal profile slightly sloping ventral from dorsal fin to occiput. Head depressed, broad. Eye small, dorsolateral, subcutaneous, ovoid. Snout rounded in dorsal view. Anterior nostril tubular, rim with fleshy flap forming short tube; posterior nostril porelike, rim posteriorly confluent with base of nasal barbel. Lateral line absent or vestigial.
</paragraph>
<paragraph id="C13636A9FFE27968A7B8F849FB67C385" blockId="1.[151,1436,1354,2027]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="269" pageId="1" pageNumber="268">Mouth terminal; lips thickened, papillate, premaxillary and mandibular toothpads curved, teeth small and ciliform or setiform, with upper jaw slightly longer than lower jaw. Four pairs of barbels: maxillary barbel longest, reaching base of pectoral spine; nasal barbel short, not reaching posterior margin of preoperculum; lateral mandibular barbel long, reaching posterior margin of pectoral-fin base; medial mandibular barbel shortest of the four pairs of barbels, about half length of outer mental barbel, reaching to gill membrane.</paragraph>
<caption id="95F66621FFE17968A7E8FF3DFDF9C265" ID-Table-UUID="95F66621FFE17968A7E8FF3DFDF9C265" httpUri="http://table.plazi.org/id/95F66621FFE17968A7E8FF3DFDF9C265" pageId="2" pageNumber="269" targetBox="[159,1411,378,648]" targetIsTable="true" targetPageId="2">
<paragraph id="C13636A9FFE17968A7E8FF3DFDF9C265" blockId="2.[151,1436,253,309]" pageId="2" pageNumber="269">
<emphasis id="F3FDEABBFFE17968A7E8FF3DFEEDC245" bold="true" box="[151,266,254,277]" pageId="2" pageNumber="269">TABLE 1.</emphasis>
Morphometric and meristic data of
<taxonomicName id="06894D2AFFE17968A5F0FF3EFC4CC245" box="[655,939,253,277]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="hyeongsanensis">
<emphasis id="F3FDEABBFFE17968A5F0FF3EFC4CC245" box="[655,939,253,277]" italics="true" pageId="2" pageNumber="269">Liobagrus hyeongsanensis</emphasis>
</taxonomicName>
(CNUC 38547–38567) and
<taxonomicName id="06894D2AFFE17968A3A1FF3DFA7BC245" box="[1246,1436,253,277]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="mediadiposalis">
<emphasis id="F3FDEABBFFE17968A3A1FF3DFA7BC245" box="[1246,1436,253,277]" italics="true" pageId="2" pageNumber="269">L. mediadiposalis</emphasis>
</taxonomicName>
(CNUC 37821–37838). H, holotype.
</paragraph>
</caption>
<paragraph id="C13636A9FFE17968A530FE91FB71C486" blockId="2.[591,1260,337,359]" lastBlockId="2.[151,1437,1693,2006]" pageId="2" pageNumber="269">
<emphasis id="F3FDEABBFFE17968A530FE91FB0BC237" box="[591,1260,337,359]" italics="true" pageId="2" pageNumber="269">
<taxonomicName id="06894D2AFFE17968A530FE91FCEEC237" box="[591,777,337,359]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="hyeongsanensis">L. hyeongsanensis</taxonomicName>
<taxonomicName id="06894D2AFFE17968A347FE91FB0BC237" authority="Length" authorityName="Length" box="[1080,1260,337,359]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="269" phylum="Chordata" rank="species" species="mediadiposalis">L. mediadiposalis</taxonomicName>
</emphasis>
Length from occiput to dorsal-fin origin 9.4 20 9.3–12.9 10.3 0.8 18 10.3–13.3 11.7 0.8 Dorsal fin with II, 6 rays; origin closer to snout tip than to anal-fin origin, its posterior margin convex. Dorsalfin spine a little shorter than pectoral-fin spine. Adipose fin base long, as long as or slightly exceeding anal-fin base length, confluent with caudal fin, its posterior margin convex. Pectoral fin with I, 7 rays, its origin anterior to vertical from posterior margin of operculum, partially covered by opercular membrane. Pectoral-fin spine sharp, short, with 2 (5), 3 (16) serrations on posterior edge. Pelvic fin with i, 5 rays; short, its origin located midway between dorsal and adipose fins, its tip not reaching base of anal fin. Anal fin with 15 (2), 16 (5), 17 (11), or 18 (3) rays, its distal margin rounded; anal fin short, origin slightly posterior to vertical through adipose-fin origin, posterior tip of anal fin not reaching beyond posterior margin of adipose fin. Caudal fin subtruncate, rounded. Vertebral column 38 (1), 39 (10), 40 (9), or 41 (1) post-Weberian elements. Gill rakers 6–8.
</paragraph>
<paragraph id="C13636A9FFE17968A530FEB8FA64C1D8" pageId="2" pageNumber="269">
<table id="B389C409FFE18695A7E0FEB9FA64C1D8" box="[159,1411,378,648]" gridcols="8" gridrows="7" pageId="2" pageNumber="269">
<tr id="7FB934EBFFE18695A7E0FEB9FA64C2C0" box="[159,1411,378,400]" gridrow="0" pageId="2" pageNumber="269" rowspan-0="1">
<th id="3C685D97FFE18695A530FEB9FD54C2C0" box="[591,691,378,400]" gridcol="1" gridrow="0" pageId="2" pageNumber="269">H n</th>
<th id="3C685D97FFE18695A59CFEB9FCAFC2C0" box="[739,840,378,400]" gridcol="2" gridrow="0" pageId="2" pageNumber="269">range</th>
<th id="3C685D97FFE18695A409FEB9FC4AC2C0" box="[886,941,378,400]" gridcol="3" gridrow="0" pageId="2" pageNumber="269">mean</th>
<th id="3C685D97FFE18695A4A7FEB9FC1FC2C0" box="[984,1016,378,400]" gridcol="4" gridrow="0" pageId="2" pageNumber="269">SD</th>
<th id="3C685D97FFE18695A347FEB9FBB5C2C0" box="[1080,1106,378,400]" gridcol="5" gridrow="0" pageId="2" pageNumber="269">n</th>
<th id="3C685D97FFE18695A302FEB9FAD9C2C0" box="[1149,1342,378,400]" gridcol="6" gridrow="0" pageId="2" pageNumber="269">range mean</th>
<th id="3C685D97FFE18695A21CFEB9FA64C2C0" box="[1379,1411,378,400]" gridcol="7" gridrow="0" pageId="2" pageNumber="269">SD</th>
</tr>
<tr id="7FB934EBFFE18695A7E0FE67FA64C2EA" box="[159,1411,420,442]" gridrow="1" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FE67FE2CC2EA" box="[159,459,420,442]" gridcol="0" gridrow="1" pageId="2" pageNumber="269">Standard length (mm)</th>
<td id="3C685D97FFE18695A530FE67FD54C2EA" box="[591,691,420,442]" gridcol="1" gridrow="1" pageId="2" pageNumber="269">84.1 20</td>
<td id="3C685D97FFE18695A59CFE67FCAFC2EA" box="[739,840,420,442]" gridcol="2" gridrow="1" pageId="2" pageNumber="269">57.1–84.1</td>
<td id="3C685D97FFE18695A409FE67FC4AC2EA" box="[886,941,420,442]" gridcol="3" gridrow="1" pageId="2" pageNumber="269">68.9</td>
<td id="3C685D97FFE18695A4A7FE67FC1FC2EA" box="[984,1016,420,442]" gridcol="4" gridrow="1" pageId="2" pageNumber="269">6.0</td>
<td id="3C685D97FFE18695A347FE67FBB5C2EA" box="[1080,1106,420,442]" gridcol="5" gridrow="1" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FE67FAD9C2EA" box="[1149,1342,420,442]" gridcol="6" gridrow="1" pageId="2" pageNumber="269">73.5–100.3 85.4</td>
<td id="3C685D97FFE18695A21CFE67FA64C2EA" box="[1379,1411,420,442]" gridcol="7" gridrow="1" pageId="2" pageNumber="269">7.7</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FE0EFA64C2B3" box="[159,1411,461,483]" gridrow="2" pageId="2" pageNumber="269" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="3C685D97FFE18695A7E0FE0EFE2CC2B3" box="[159,459,461,483]" gridcol="0" gridrow="2" pageId="2" pageNumber="269">In percents of standard length</th>
</tr>
<tr id="7FB934EBFFE18695A7E0FE35FA64C15C" box="[159,1411,502,524]" gridrow="3" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FE35FE2CC15C" box="[159,459,502,524]" gridcol="0" gridrow="3" pageId="2" pageNumber="269">Preocciput length</th>
<td id="3C685D97FFE18695A530FE35FD54C15C" box="[591,691,502,524]" gridcol="1" gridrow="3" pageId="2" pageNumber="269">19.1 20</td>
<td id="3C685D97FFE18695A59CFE35FCAFC15C" box="[739,840,502,524]" gridcol="2" gridrow="3" pageId="2" pageNumber="269">18.3–22.3</td>
<td id="3C685D97FFE18695A409FE35FC4AC15C" box="[886,941,502,524]" gridcol="3" gridrow="3" pageId="2" pageNumber="269">19.6</td>
<td id="3C685D97FFE18695A4A7FE35FC1FC15C" box="[984,1016,502,524]" gridcol="4" gridrow="3" pageId="2" pageNumber="269">1.1</td>
<td id="3C685D97FFE18695A347FE35FBB5C15C" box="[1080,1106,502,524]" gridcol="5" gridrow="3" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FE35FAD9C15C" box="[1149,1342,502,524]" gridcol="6" gridrow="3" pageId="2" pageNumber="269">17.9–20.3 19.2</td>
<td id="3C685D97FFE18695A21CFE35FA64C15C" box="[1379,1411,502,524]" gridcol="7" gridrow="3" pageId="2" pageNumber="269">0.6</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FDE3FA64C166" box="[159,1411,544,566]" gridrow="4" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FDE3FE2CC166" box="[159,459,544,566]" gridcol="0" gridrow="4" pageId="2" pageNumber="269">Predorsal length</th>
<td id="3C685D97FFE18695A530FDE3FD54C166" box="[591,691,544,566]" gridcol="1" gridrow="4" pageId="2" pageNumber="269">28.2 20</td>
<td id="3C685D97FFE18695A59CFDE3FCAFC166" box="[739,840,544,566]" gridcol="2" gridrow="4" pageId="2" pageNumber="269">27.8–31.3</td>
<td id="3C685D97FFE18695A409FDE3FC4AC166" box="[886,941,544,566]" gridcol="3" gridrow="4" pageId="2" pageNumber="269">29.4</td>
<td id="3C685D97FFE18695A4A7FDE3FC1FC166" box="[984,1016,544,566]" gridcol="4" gridrow="4" pageId="2" pageNumber="269">1.1</td>
<td id="3C685D97FFE18695A347FDE3FBB5C166" box="[1080,1106,544,566]" gridcol="5" gridrow="4" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FDE3FAD9C166" box="[1149,1342,544,566]" gridcol="6" gridrow="4" pageId="2" pageNumber="269">28.4–31.4 30.3</td>
<td id="3C685D97FFE18695A21CFDE3FA64C166" box="[1379,1411,544,566]" gridcol="7" gridrow="4" pageId="2" pageNumber="269">0.8</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FD8AFA64C10F" box="[159,1411,585,607]" gridrow="5" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FD8AFE2CC10F" box="[159,459,585,607]" gridcol="0" gridrow="5" pageId="2" pageNumber="269">Prepectoral length</th>
<td id="3C685D97FFE18695A530FD8AFD54C10F" box="[591,691,585,607]" gridcol="1" gridrow="5" pageId="2" pageNumber="269">18.7 20</td>
<td id="3C685D97FFE18695A59CFD8AFCAFC10F" box="[739,840,585,607]" gridcol="2" gridrow="5" pageId="2" pageNumber="269">18.7–21.4</td>
<td id="3C685D97FFE18695A409FD8AFC4AC10F" box="[886,941,585,607]" gridcol="3" gridrow="5" pageId="2" pageNumber="269">19.9</td>
<td id="3C685D97FFE18695A4A7FD8AFC1FC10F" box="[984,1016,585,607]" gridcol="4" gridrow="5" pageId="2" pageNumber="269">0.7</td>
<td id="3C685D97FFE18695A347FD8AFBB5C10F" box="[1080,1106,585,607]" gridcol="5" gridrow="5" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FD8AFAD9C10F" box="[1149,1342,585,607]" gridcol="6" gridrow="5" pageId="2" pageNumber="269">20.6–23.3 22.0</td>
<td id="3C685D97FFE18695A21CFD8AFA64C10F" box="[1379,1411,585,607]" gridcol="7" gridrow="5" pageId="2" pageNumber="269">0.8</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FDB1FA64C1D8" box="[159,1411,626,648]" gridrow="6" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FDB1FE2CC1D8" box="[159,459,626,648]" gridcol="0" gridrow="6" pageId="2" pageNumber="269">Prepelvic length</th>
<td id="3C685D97FFE18695A530FDB1FD54C1D8" box="[591,691,626,648]" gridcol="1" gridrow="6" pageId="2" pageNumber="269">45.8 20</td>
<td id="3C685D97FFE18695A59CFDB1FCAFC1D8" box="[739,840,626,648]" gridcol="2" gridrow="6" pageId="2" pageNumber="269">45.4–51.4</td>
<td id="3C685D97FFE18695A409FDB1FC4AC1D8" box="[886,941,626,648]" gridcol="3" gridrow="6" pageId="2" pageNumber="269">47.4</td>
<td id="3C685D97FFE18695A4A7FDB1FC1FC1D8" box="[984,1016,626,648]" gridcol="4" gridrow="6" pageId="2" pageNumber="269">1.6</td>
<td id="3C685D97FFE18695A347FDB1FBB5C1D8" box="[1080,1106,626,648]" gridcol="5" gridrow="6" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FDB1FAD9C1D8" box="[1149,1342,626,648]" gridcol="6" gridrow="6" pageId="2" pageNumber="269">44.2–49.0 46.0</td>
<td id="3C685D97FFE18695A21CFDB1FA64C1D8" box="[1379,1411,626,648]" gridcol="7" gridrow="6" pageId="2" pageNumber="269">1.2</td>
</tr>
</table>
</paragraph>
<paragraph id="C13636A9FFE17968A7E0FD06FA64C03B" pageId="2" pageNumber="269">
<table id="B389C409FFE18695A7E0FD06FA64C0DA" box="[159,1411,709,906]" gridcols="10" gridrows="3" pageId="2" pageNumber="269">
<tr id="7FB934EBFFE18695A7E0FD06FA64C1AA" box="[159,1411,709,762]" gridrow="0" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FD06FDD7C1AA" box="[159,560,709,762]" gridcol="0" gridrow="0" pageId="2" pageNumber="269">Length from pectoral-fin origin to dorsal-fin origin</th>
<th id="3C685D97FFE18695A530FD06FD9CC1AA" box="[591,635,709,762]" gridcol="1" gridrow="0" pageId="2" pageNumber="269">15.2</th>
<th id="3C685D97FFE18695A5E6FD06FD54C1AA" box="[665,691,709,762]" gridcol="2" gridrow="0" pageId="2" pageNumber="269">20</th>
<th id="3C685D97FFE18695A59CFD06FCAFC1AA" box="[739,840,709,762]" gridcol="3" gridrow="0" pageId="2" pageNumber="269">13.8–17.7</th>
<th id="3C685D97FFE18695A409FD06FC45C1AA" box="[886,930,709,762]" gridcol="4" gridrow="0" pageId="2" pageNumber="269">15.7</th>
<th id="3C685D97FFE18695A4A7FD06FC1FC1AA" box="[984,1016,709,762]" gridcol="5" gridrow="0" pageId="2" pageNumber="269">1.0</th>
<th id="3C685D97FFE18695A347FD06FBB5C1AA" box="[1080,1106,709,762]" gridcol="6" gridrow="0" pageId="2" pageNumber="269">18</th>
<th id="3C685D97FFE18695A302FD06FB04C1AA" box="[1149,1251,709,762]" gridcol="7" gridrow="0" pageId="2" pageNumber="269">14.8–23.9</th>
<th id="3C685D97FFE18695A278FD06FAD4C1AA" box="[1287,1331,709,762]" gridcol="8" gridrow="0" pageId="2" pageNumber="269">16.9</th>
<th id="3C685D97FFE18695A21CFD06FA64C1AA" box="[1379,1411,709,762]" gridcol="9" gridrow="0" pageId="2" pageNumber="269">1.9</th>
</tr>
<tr id="7FB934EBFFE18695A7E0FCCEFA64C012" box="[159,1411,781,834]" gridrow="1" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FCCEFDD7C012" box="[159,560,781,834]" gridcol="0" gridrow="1" pageId="2" pageNumber="269">Length from pectoral-fin origin to pelvic-fin origin</th>
<td id="3C685D97FFE18695A530FCCEFD9CC012" box="[591,635,781,834]" gridcol="1" gridrow="1" pageId="2" pageNumber="269">28.3</td>
<td id="3C685D97FFE18695A5E6FCCEFD54C012" box="[665,691,781,834]" gridcol="2" gridrow="1" pageId="2" pageNumber="269">20</td>
<td id="3C685D97FFE18695A59CFCCEFCAFC012" box="[739,840,781,834]" gridcol="3" gridrow="1" pageId="2" pageNumber="269">27.5–34.4</td>
<td id="3C685D97FFE18695A409FCCEFC45C012" box="[886,930,781,834]" gridcol="4" gridrow="1" pageId="2" pageNumber="269">30.2</td>
<td id="3C685D97FFE18695A4A7FCCEFC1FC012" box="[984,1016,781,834]" gridcol="5" gridrow="1" pageId="2" pageNumber="269">2.3</td>
<td id="3C685D97FFE18695A347FCCEFBB5C012" box="[1080,1106,781,834]" gridcol="6" gridrow="1" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FCCEFB04C012" box="[1149,1251,781,834]" gridcol="7" gridrow="1" pageId="2" pageNumber="269">25.4–28.5</td>
<td id="3C685D97FFE18695A278FCCEFAD4C012" box="[1287,1331,781,834]" gridcol="8" gridrow="1" pageId="2" pageNumber="269">27.0</td>
<td id="3C685D97FFE18695A21CFCCEFA64C012" box="[1379,1411,781,834]" gridcol="9" gridrow="1" pageId="2" pageNumber="269">0.9</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FC96FA64C0DA" box="[159,1411,853,906]" gridrow="2" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FC96FDD7C0DA" box="[159,560,853,906]" gridcol="0" gridrow="2" pageId="2" pageNumber="269">Length from dorsal-fin origin to pelvic- fin origin</th>
<td id="3C685D97FFE18695A530FC96FD9CC0DA" box="[591,635,853,906]" gridcol="1" gridrow="2" pageId="2" pageNumber="269">24.7</td>
<td id="3C685D97FFE18695A5E6FC96FD54C0DA" box="[665,691,853,906]" gridcol="2" gridrow="2" pageId="2" pageNumber="269">20</td>
<td id="3C685D97FFE18695A59CFC96FCAFC0DA" box="[739,840,853,906]" gridcol="3" gridrow="2" pageId="2" pageNumber="269">24.6–31.9</td>
<td id="3C685D97FFE18695A409FC96FC45C0DA" box="[886,930,853,906]" gridcol="4" gridrow="2" pageId="2" pageNumber="269">27.6</td>
<td id="3C685D97FFE18695A4A7FC96FC1FC0DA" box="[984,1016,853,906]" gridcol="5" gridrow="2" pageId="2" pageNumber="269">2.3</td>
<td id="3C685D97FFE18695A347FC96FBB5C0DA" box="[1080,1106,853,906]" gridcol="6" gridrow="2" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FC96FB04C0DA" box="[1149,1251,853,906]" gridcol="7" gridrow="2" pageId="2" pageNumber="269">23.0–25.7</td>
<td id="3C685D97FFE18695A278FC96FAD4C0DA" box="[1287,1331,853,906]" gridcol="8" gridrow="2" pageId="2" pageNumber="269">24.7</td>
<td id="3C685D97FFE18695A21CFC96FA64C0DA" box="[1379,1411,853,906]" gridcol="9" gridrow="2" pageId="2" pageNumber="269">1.0</td>
</tr>
</table>
</paragraph>
<paragraph id="C13636A9FFE17968A7E0FC5EFA64C50C" pageId="2" pageNumber="269">
<table id="B389C409FFE18695A7E0FC5EFA64C50C" box="[159,1411,925,1628]" gridcols="7" gridrows="16" pageId="2" pageNumber="269">
<tr id="7FB934EBFFE18695A7E0FC5EFA64C082" box="[159,1411,925,978]" gridrow="0" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FC5EFD54C082" box="[159,691,925,978]" gridcol="0" gridrow="0" pageId="2" pageNumber="269">Length from pelvic-fin origin to adipose- 22.2 20 fin origin</th>
<th id="3C685D97FFE18695A59DFC5EFCAFC082" box="[738,840,925,978]" gridcol="1" gridrow="0" pageId="2" pageNumber="269">15.9–26.0</th>
<th id="3C685D97FFE18695A409FC5EFC45C082" box="[886,930,925,978]" gridcol="2" gridrow="0" pageId="2" pageNumber="269">20.0</th>
<th id="3C685D97FFE18695A4A7FC5EFC1FC082" box="[984,1016,925,978]" gridcol="3" gridrow="0" pageId="2" pageNumber="269">2.3</th>
<th id="3C685D97FFE18695A347FC5EFBB5C082" box="[1080,1106,925,978]" gridcol="4" gridrow="0" pageId="2" pageNumber="269">18</th>
<th id="3C685D97FFE18695A302FC5EFAD4C082" box="[1149,1331,925,978]" gridcol="5" gridrow="0" pageId="2" pageNumber="269">18.1–23.5 20.4</th>
<th id="3C685D97FFE18695A21CFC5EFA64C082" box="[1379,1411,925,978]" gridcol="6" gridrow="0" pageId="2" pageNumber="269">1.4</th>
</tr>
<tr id="7FB934EBFFE18695A7E0FC26FA64C74A" box="[159,1411,997,1050]" gridrow="1" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FC26FD54C74A" box="[159,691,997,1050]" gridcol="0" gridrow="1" pageId="2" pageNumber="269">Length from pelvic-fin origin to anal-fin 16.6 20 origin</th>
<td id="3C685D97FFE18695A59DFC26FCAFC74A" box="[738,840,997,1050]" gridcol="1" gridrow="1" pageId="2" pageNumber="269">12.1–16.6</td>
<td id="3C685D97FFE18695A409FC26FC45C74A" box="[886,930,997,1050]" gridcol="2" gridrow="1" pageId="2" pageNumber="269">14.1</td>
<td id="3C685D97FFE18695A4A7FC26FC1FC74A" box="[984,1016,997,1050]" gridcol="3" gridrow="1" pageId="2" pageNumber="269">1.2</td>
<td id="3C685D97FFE18695A347FC26FBB5C74A" box="[1080,1106,997,1050]" gridcol="4" gridrow="1" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FC26FAD4C74A" box="[1149,1331,997,1050]" gridcol="5" gridrow="1" pageId="2" pageNumber="269">15.3–20.9 17.9</td>
<td id="3C685D97FFE18695A21CFC26FA64C74A" box="[1379,1411,997,1050]" gridcol="6" gridrow="1" pageId="2" pageNumber="269">1.5</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FBEEFA64C713" box="[159,1411,1069,1091]" gridrow="2" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FBEEFD54C713" box="[159,691,1069,1091]" gridcol="0" gridrow="2" pageId="2" pageNumber="269">Anal-fin base length 16.3 20</th>
<td id="3C685D97FFE18695A59DFBEEFCAFC713" box="[738,840,1069,1091]" gridcol="1" gridrow="2" pageId="2" pageNumber="269">15.5–28.1</td>
<td id="3C685D97FFE18695A409FBEEFC45C713" box="[886,930,1069,1091]" gridcol="2" gridrow="2" pageId="2" pageNumber="269">17.9</td>
<td id="3C685D97FFE18695A4A7FBEEFC1FC713" box="[984,1016,1069,1091]" gridcol="3" gridrow="2" pageId="2" pageNumber="269">2.6</td>
<td id="3C685D97FFE18695A347FBEEFBB5C713" box="[1080,1106,1069,1091]" gridcol="4" gridrow="2" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FBEEFAD4C713" box="[1149,1331,1069,1091]" gridcol="5" gridrow="2" pageId="2" pageNumber="269">14.7–18.6 16.8</td>
<td id="3C685D97FFE18695A21CFBEEFA64C713" box="[1379,1411,1069,1091]" gridcol="6" gridrow="2" pageId="2" pageNumber="269">1.1</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FB95FA64C73C" box="[159,1411,1110,1132]" gridrow="3" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FB95FD54C73C" box="[159,691,1110,1132]" gridcol="0" gridrow="3" pageId="2" pageNumber="269">Dorsal-fin base length 8.3 20</th>
<td id="3C685D97FFE18695A59DFB95FCAFC73C" box="[738,840,1110,1132]" gridcol="1" gridrow="3" pageId="2" pageNumber="269">7.8–10.9</td>
<td id="3C685D97FFE18695A409FB95FC45C73C" box="[886,930,1110,1132]" gridcol="2" gridrow="3" pageId="2" pageNumber="269">9.0</td>
<td id="3C685D97FFE18695A4A7FB95FC1FC73C" box="[984,1016,1110,1132]" gridcol="3" gridrow="3" pageId="2" pageNumber="269">0.8</td>
<td id="3C685D97FFE18695A347FB95FBB5C73C" box="[1080,1106,1110,1132]" gridcol="4" gridrow="3" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FB95FAD4C73C" box="[1149,1331,1110,1132]" gridcol="5" gridrow="3" pageId="2" pageNumber="269">7.7–10.4 9.6</td>
<td id="3C685D97FFE18695A21CFB95FA64C73C" box="[1379,1411,1110,1132]" gridcol="6" gridrow="3" pageId="2" pageNumber="269">0.6</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FB43FA64C7C6" box="[159,1411,1152,1174]" gridrow="4" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FB43FD54C7C6" box="[159,691,1152,1174]" gridcol="0" gridrow="4" pageId="2" pageNumber="269">Occiput to pectoral-fin origin 11.0 20</th>
<td id="3C685D97FFE18695A59DFB43FCAFC7C6" box="[738,840,1152,1174]" gridcol="1" gridrow="4" pageId="2" pageNumber="269">10.4–13.3</td>
<td id="3C685D97FFE18695A409FB43FC45C7C6" box="[886,930,1152,1174]" gridcol="2" gridrow="4" pageId="2" pageNumber="269">11.6</td>
<td id="3C685D97FFE18695A4A7FB43FC1FC7C6" box="[984,1016,1152,1174]" gridcol="3" gridrow="4" pageId="2" pageNumber="269">0.8</td>
<td id="3C685D97FFE18695A347FB43FBB5C7C6" box="[1080,1106,1152,1174]" gridcol="4" gridrow="4" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FB43FAD4C7C6" box="[1149,1331,1152,1174]" gridcol="5" gridrow="4" pageId="2" pageNumber="269">11.9–13.4 12.5</td>
<td id="3C685D97FFE18695A21CFB43FA64C7C6" box="[1379,1411,1152,1174]" gridcol="6" gridrow="4" pageId="2" pageNumber="269">0.5</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FB6AFA64C7EF" box="[159,1411,1193,1215]" gridrow="5" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FB6AFD54C7EF" box="[159,691,1193,1215]" gridcol="0" gridrow="5" pageId="2" pageNumber="269">Interorbital width 7.4 20</th>
<td id="3C685D97FFE18695A59DFB6AFCAFC7EF" box="[738,840,1193,1215]" gridcol="1" gridrow="5" pageId="2" pageNumber="269">5.6–7.8</td>
<td id="3C685D97FFE18695A409FB6AFC45C7EF" box="[886,930,1193,1215]" gridcol="2" gridrow="5" pageId="2" pageNumber="269">6.6</td>
<td id="3C685D97FFE18695A4A7FB6AFC1FC7EF" box="[984,1016,1193,1215]" gridcol="3" gridrow="5" pageId="2" pageNumber="269">0.5</td>
<td id="3C685D97FFE18695A347FB6AFBB5C7EF" box="[1080,1106,1193,1215]" gridcol="4" gridrow="5" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FB6AFAD4C7EF" box="[1149,1331,1193,1215]" gridcol="5" gridrow="5" pageId="2" pageNumber="269">6.2–7.5 6.7</td>
<td id="3C685D97FFE18695A21CFB6AFA64C7EF" box="[1379,1411,1193,1215]" gridcol="6" gridrow="5" pageId="2" pageNumber="269">0.3</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FB11FA64C7B8" box="[159,1411,1234,1256]" gridrow="6" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FB11FD54C7B8" box="[159,691,1234,1256]" gridcol="0" gridrow="6" pageId="2" pageNumber="269">Body width at pectoral-fin origin 16.9 20</th>
<td id="3C685D97FFE18695A59DFB11FCAFC7B8" box="[738,840,1234,1256]" gridcol="1" gridrow="6" pageId="2" pageNumber="269">15.5–17.9</td>
<td id="3C685D97FFE18695A409FB11FC45C7B8" box="[886,930,1234,1256]" gridcol="2" gridrow="6" pageId="2" pageNumber="269">16.5</td>
<td id="3C685D97FFE18695A4A7FB11FC1FC7B8" box="[984,1016,1234,1256]" gridcol="3" gridrow="6" pageId="2" pageNumber="269">0.6</td>
<td id="3C685D97FFE18695A347FB11FBB5C7B8" box="[1080,1106,1234,1256]" gridcol="4" gridrow="6" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FB11FAD4C7B8" box="[1149,1331,1234,1256]" gridcol="5" gridrow="6" pageId="2" pageNumber="269">17.1–20.0 18.5</td>
<td id="3C685D97FFE18695A21CFB11FA64C7B8" box="[1379,1411,1234,1256]" gridcol="6" gridrow="6" pageId="2" pageNumber="269">0.7</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FB3FFA64C642" box="[159,1411,1276,1298]" gridrow="7" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FB3FFD54C642" box="[159,691,1276,1298]" gridcol="0" gridrow="7" pageId="2" pageNumber="269">Body width under dorsal-fin origin 13.6 20</th>
<td id="3C685D97FFE18695A59DFB3FFCAFC642" box="[738,840,1276,1298]" gridcol="1" gridrow="7" pageId="2" pageNumber="269">13.4–19.3</td>
<td id="3C685D97FFE18695A409FB3FFC45C642" box="[886,930,1276,1298]" gridcol="2" gridrow="7" pageId="2" pageNumber="269">15.5</td>
<td id="3C685D97FFE18695A4A7FB3FFC1FC642" box="[984,1016,1276,1298]" gridcol="3" gridrow="7" pageId="2" pageNumber="269">1.6</td>
<td id="3C685D97FFE18695A347FB3FFBB5C642" box="[1080,1106,1276,1298]" gridcol="4" gridrow="7" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FB3FFAD4C642" box="[1149,1331,1276,1298]" gridcol="5" gridrow="7" pageId="2" pageNumber="269">13.0–16.5 14.9</td>
<td id="3C685D97FFE18695A21CFB3FFA64C642" box="[1379,1411,1276,1298]" gridcol="6" gridrow="7" pageId="2" pageNumber="269">0.9</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FAE6FA64C66B" box="[159,1411,1317,1339]" gridrow="8" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FAE6FD54C66B" box="[159,691,1317,1339]" gridcol="0" gridrow="8" pageId="2" pageNumber="269">Head width 19.9 20</th>
<td id="3C685D97FFE18695A59DFAE6FCAFC66B" box="[738,840,1317,1339]" gridcol="1" gridrow="8" pageId="2" pageNumber="269">16.4–19.9</td>
<td id="3C685D97FFE18695A409FAE6FC45C66B" box="[886,930,1317,1339]" gridcol="2" gridrow="8" pageId="2" pageNumber="269">18.0</td>
<td id="3C685D97FFE18695A4A7FAE6FC1FC66B" box="[984,1016,1317,1339]" gridcol="3" gridrow="8" pageId="2" pageNumber="269">0.9</td>
<td id="3C685D97FFE18695A347FAE6FBB5C66B" box="[1080,1106,1317,1339]" gridcol="4" gridrow="8" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FAE6FAD4C66B" box="[1149,1331,1317,1339]" gridcol="5" gridrow="8" pageId="2" pageNumber="269">17.5–19.6 18.5</td>
<td id="3C685D97FFE18695A21CFAE6FA64C66B" box="[1379,1411,1317,1339]" gridcol="6" gridrow="8" pageId="2" pageNumber="269">0.6</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FA8DFA64C634" box="[159,1411,1358,1380]" gridrow="9" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FA8DFD54C634" box="[159,691,1358,1380]" gridcol="0" gridrow="9" pageId="2" pageNumber="269">Pectoral spine length 3.7 20</th>
<td id="3C685D97FFE18695A59DFA8DFCAFC634" box="[738,840,1358,1380]" gridcol="1" gridrow="9" pageId="2" pageNumber="269">3.7–6.5</td>
<td id="3C685D97FFE18695A409FA8DFC45C634" box="[886,930,1358,1380]" gridcol="2" gridrow="9" pageId="2" pageNumber="269">5.2</td>
<td id="3C685D97FFE18695A4A7FA8DFC1FC634" box="[984,1016,1358,1380]" gridcol="3" gridrow="9" pageId="2" pageNumber="269">0.8</td>
<td id="3C685D97FFE18695A347FA8DFBB5C634" box="[1080,1106,1358,1380]" gridcol="4" gridrow="9" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FA8DFAD4C634" box="[1149,1331,1358,1380]" gridcol="5" gridrow="9" pageId="2" pageNumber="269">9.0–12.2 10.3</td>
<td id="3C685D97FFE18695A21CFA8DFA64C634" box="[1379,1411,1358,1380]" gridcol="6" gridrow="9" pageId="2" pageNumber="269">1.0</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FABBFA64C6DE" box="[159,1411,1400,1422]" gridrow="10" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FABBFD54C6DE" box="[159,691,1400,1422]" gridcol="0" gridrow="10" pageId="2" pageNumber="269">Dorsal spine length 3.3 20</th>
<td id="3C685D97FFE18695A59DFABBFCAFC6DE" box="[738,840,1400,1422]" gridcol="1" gridrow="10" pageId="2" pageNumber="269">3.2–6.8</td>
<td id="3C685D97FFE18695A409FABBFC45C6DE" box="[886,930,1400,1422]" gridcol="2" gridrow="10" pageId="2" pageNumber="269">4.7</td>
<td id="3C685D97FFE18695A4A7FABBFC1FC6DE" box="[984,1016,1400,1422]" gridcol="3" gridrow="10" pageId="2" pageNumber="269">0.9</td>
<td id="3C685D97FFE18695A347FABBFBB5C6DE" box="[1080,1106,1400,1422]" gridcol="4" gridrow="10" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FABBFAD4C6DE" box="[1149,1331,1400,1422]" gridcol="5" gridrow="10" pageId="2" pageNumber="269">5.5–8.9 7.4</td>
<td id="3C685D97FFE18695A21CFABBFA64C6DE" box="[1379,1411,1400,1422]" gridcol="6" gridrow="10" pageId="2" pageNumber="269">1.0</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FA62FA64C6E7" box="[159,1411,1441,1463]" gridrow="11" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FA62FD54C6E7" box="[159,691,1441,1463]" gridcol="0" gridrow="11" pageId="2" pageNumber="269">Caudal-fin length 17.3 20</th>
<td id="3C685D97FFE18695A59DFA62FCAFC6E7" box="[738,840,1441,1463]" gridcol="1" gridrow="11" pageId="2" pageNumber="269">14.9–18.2</td>
<td id="3C685D97FFE18695A409FA62FC45C6E7" box="[886,930,1441,1463]" gridcol="2" gridrow="11" pageId="2" pageNumber="269">17.0</td>
<td id="3C685D97FFE18695A4A7FA62FC1FC6E7" box="[984,1016,1441,1463]" gridcol="3" gridrow="11" pageId="2" pageNumber="269">0.8</td>
<td id="3C685D97FFE18695A347FA62FBB5C6E7" box="[1080,1106,1441,1463]" gridcol="4" gridrow="11" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FA62FAD4C6E7" box="[1149,1331,1441,1463]" gridcol="5" gridrow="11" pageId="2" pageNumber="269">15.6–20.6 18.8</td>
<td id="3C685D97FFE18695A21CFA62FA64C6E7" box="[1379,1411,1441,1463]" gridcol="6" gridrow="11" pageId="2" pageNumber="269">1.4</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FA09FA64C6B0" box="[159,1411,1482,1504]" gridrow="12" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FA09FD54C6B0" box="[159,691,1482,1504]" gridcol="0" gridrow="12" pageId="2" pageNumber="269">Maxillary-barbel length 20.1 20</th>
<td id="3C685D97FFE18695A59DFA09FCAFC6B0" box="[738,840,1482,1504]" gridcol="1" gridrow="12" pageId="2" pageNumber="269">18.2–22.6</td>
<td id="3C685D97FFE18695A409FA09FC45C6B0" box="[886,930,1482,1504]" gridcol="2" gridrow="12" pageId="2" pageNumber="269">20.9</td>
<td id="3C685D97FFE18695A4A7FA09FC1FC6B0" box="[984,1016,1482,1504]" gridcol="3" gridrow="12" pageId="2" pageNumber="269">1.1</td>
<td id="3C685D97FFE18695A347FA09FBB5C6B0" box="[1080,1106,1482,1504]" gridcol="4" gridrow="12" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FA09FAD4C6B0" box="[1149,1331,1482,1504]" gridcol="5" gridrow="12" pageId="2" pageNumber="269">17.5–24.5 21.0</td>
<td id="3C685D97FFE18695A21CFA09FA64C6B0" box="[1379,1411,1482,1504]" gridcol="6" gridrow="12" pageId="2" pageNumber="269">1.6</td>
</tr>
<tr id="7FB934EBFFE18695A7E0FA37FA64C55A" box="[159,1411,1524,1546]" gridrow="13" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0FA37FD54C55A" box="[159,691,1524,1546]" gridcol="0" gridrow="13" pageId="2" pageNumber="269">Nasal-barbel length 12.3 20</th>
<td id="3C685D97FFE18695A59DFA37FCAFC55A" box="[738,840,1524,1546]" gridcol="1" gridrow="13" pageId="2" pageNumber="269">9.4–14.5</td>
<td id="3C685D97FFE18695A409FA37FC45C55A" box="[886,930,1524,1546]" gridcol="2" gridrow="13" pageId="2" pageNumber="269">12.3</td>
<td id="3C685D97FFE18695A4A7FA37FC1FC55A" box="[984,1016,1524,1546]" gridcol="3" gridrow="13" pageId="2" pageNumber="269">1.3</td>
<td id="3C685D97FFE18695A347FA37FBB5C55A" box="[1080,1106,1524,1546]" gridcol="4" gridrow="13" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302FA37FAD4C55A" box="[1149,1331,1524,1546]" gridcol="5" gridrow="13" pageId="2" pageNumber="269">10.1–15.6 12.7</td>
<td id="3C685D97FFE18695A21CFA37FA64C55A" box="[1379,1411,1524,1546]" gridcol="6" gridrow="13" pageId="2" pageNumber="269">1.3</td>
</tr>
<tr id="7FB934EBFFE18695A7E0F9DEFA64C563" box="[159,1411,1565,1587]" gridrow="14" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0F9DEFD54C563" box="[159,691,1565,1587]" gridcol="0" gridrow="14" pageId="2" pageNumber="269">Outer-mental barbel length 16.6 20</th>
<td id="3C685D97FFE18695A59DF9DEFCAFC563" box="[738,840,1565,1587]" gridcol="1" gridrow="14" pageId="2" pageNumber="269">13.4–18.1</td>
<td id="3C685D97FFE18695A409F9DEFC45C563" box="[886,930,1565,1587]" gridcol="2" gridrow="14" pageId="2" pageNumber="269">16.0</td>
<td id="3C685D97FFE18695A4A7F9DEFC1FC563" box="[984,1016,1565,1587]" gridcol="3" gridrow="14" pageId="2" pageNumber="269">1.2</td>
<td id="3C685D97FFE18695A347F9DEFBB5C563" box="[1080,1106,1565,1587]" gridcol="4" gridrow="14" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302F9DEFAD4C563" box="[1149,1331,1565,1587]" gridcol="5" gridrow="14" pageId="2" pageNumber="269">12.4–18.6 15.2</td>
<td id="3C685D97FFE18695A21CF9DEFA64C563" box="[1379,1411,1565,1587]" gridcol="6" gridrow="14" pageId="2" pageNumber="269">1.7</td>
</tr>
<tr id="7FB934EBFFE18695A7E0F985FA64C50C" box="[159,1411,1606,1628]" gridrow="15" pageId="2" pageNumber="269">
<th id="3C685D97FFE18695A7E0F985FD54C50C" box="[159,691,1606,1628]" gridcol="0" gridrow="15" pageId="2" pageNumber="269">Inner-mental barbel length 7.2 20</th>
<td id="3C685D97FFE18695A59DF985FCAFC50C" box="[738,840,1606,1628]" gridcol="1" gridrow="15" pageId="2" pageNumber="269">6.5–10.6</td>
<td id="3C685D97FFE18695A409F985FC45C50C" box="[886,930,1606,1628]" gridcol="2" gridrow="15" pageId="2" pageNumber="269">8.9</td>
<td id="3C685D97FFE18695A4A7F985FC1FC50C" box="[984,1016,1606,1628]" gridcol="3" gridrow="15" pageId="2" pageNumber="269">1.2</td>
<td id="3C685D97FFE18695A347F985FBB5C50C" box="[1080,1106,1606,1628]" gridcol="4" gridrow="15" pageId="2" pageNumber="269">18</td>
<td id="3C685D97FFE18695A302F985FAD4C50C" box="[1149,1331,1606,1628]" gridcol="5" gridrow="15" pageId="2" pageNumber="269">7.4–9.1 8.3</td>
<td id="3C685D97FFE18695A21CF985FA64C50C" box="[1379,1411,1606,1628]" gridcol="6" gridrow="15" pageId="2" pageNumber="269">0.5</td>
</tr>
</table>
</paragraph>
<paragraph id="C13636A9FFE07969A7B8FF54FE34C3A8" blockId="3.[151,1436,151,248]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7B8FF54FEAFC3E0" bold="true" box="[199,328,151,176]" pageId="3" pageNumber="270">Coloration</emphasis>
. See
<figureCitation id="59B22A2CFFE07969A6F6FF54FE0EC3E0" box="[393,489,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1288,1310]" captionTargetBox="[340,1246,177,1265]" captionTargetId="figure@1.[340,1247,177,1266]" captionTargetPageId="1" captionText="FIGURE 1. Liobagrus hyeongsanensis, sp. nov., CNUC 38547, holotype, 84.1 mm SL, Hyeongsan River, South Korea." httpUri="https://zenodo.org/record/236501/files/figure.png" pageId="3" pageNumber="270">Figure 1</figureCitation>
for general appearance. In life (
<figureCitation id="59B22A2CFFE07969A425FF54FC78C3E0" box="[858,927,151,176]" captionStart="FIGURE 1" captionStartId="1.[151,250,1288,1310]" captionTargetBox="[340,1246,177,1265]" captionTargetId="figure@1.[340,1247,177,1266]" captionTargetPageId="1" captionText="FIGURE 1. Liobagrus hyeongsanensis, sp. nov., CNUC 38547, holotype, 84.1 mm SL, Hyeongsan River, South Korea." httpUri="https://zenodo.org/record/236501/files/figure.png" pageId="3" pageNumber="270">Fig. 1</figureCitation>
): body generally brownish yellow, fading to light yellow on ventral surface, without distinct markings. All barbels pale gray. All fins with similar color to body, without any other markings.
</paragraph>
</subSubSection>
<caption id="95F66621FFE07969A7E8FB2BFD00C64D" httpUri="https://zenodo.org/record/236502/files/figure.png" pageId="3" pageNumber="270" targetBox="[340,1246,302,1234]" targetPageId="3">
<paragraph id="C13636A9FFE07969A7E8FB2BFD00C64D" blockId="3.[151,1435,1256,1309]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7E8FB2BFEF2C7AD" bold="true" box="[151,277,1256,1278]" pageId="3" pageNumber="270">FIGURE 2.</emphasis>
<taxonomicName id="06894D2AFFE07969A662FB2BFDE3C7AE" box="[285,516,1256,1278]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="270" phylum="Chordata" rank="species" species="somjinensis">
<emphasis id="F3FDEABBFFE07969A662FB2BFDE3C7AE" box="[285,516,1256,1278]" italics="true" pageId="3" pageNumber="270">Liobagrus somjinensis</emphasis>
</taxonomicName>
(A), CNUC 39019, 87.7 mm SL, Somjin River, South Korea;
<taxonomicName id="06894D2AFFE07969A3EAFB2AFAACC7AE" box="[1173,1355,1256,1278]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="270" phylum="Chordata" rank="species" species="mediadiposalis">
<emphasis id="F3FDEABBFFE07969A3EAFB2AFAACC7AE" box="[1173,1355,1256,1278]" italics="true" pageId="3" pageNumber="270">L. mediadiposalis</emphasis>
</taxonomicName>
(B, C), CNUC 39020, 95.1 mm SL, Nakdong River, South Korea.
</paragraph>
</caption>
<subSubSection id="89936522FFE07969A7B8FA8AFE87C69D" pageId="3" pageNumber="270" type="distribution">
<paragraph id="C13636A9FFE07969A7B8FA8AFBC1C6D6" blockId="3.[151,1436,1353,1738]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7B8FA8AFE87C632" bold="true" box="[199,352,1353,1378]" pageId="3" pageNumber="270">Distribution.</emphasis>
<taxonomicName id="06894D2AFFE07969A60CFA89FD4EC632" box="[371,681,1354,1378]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="270" phylum="Chordata" rank="species" species="hyeongsanensis">
<emphasis id="F3FDEABBFFE07969A60CFA89FD4EC632" box="[371,681,1354,1378]" italics="true" pageId="3" pageNumber="270">Liobagrus hyeongsanensis</emphasis>
</taxonomicName>
is restricted to the rivers flowing to the eastern coast of Gyeongsangbuk-do,
<collectingCountry id="B99E7639FFE07969A6FFFAAEFDF6C6D5" box="[384,529,1389,1414]" name="South Korea" pageId="3" pageNumber="270">South Korea</collectingCountry>
: Upper Hyeongsan and Taehwa Rivers (
<figureCitation id="59B22A2CFFE07969A4ACFAAEFBFEC6D6" box="[979,1049,1389,1414]" captionStart="FIGURE 4" captionStartId="5.[151,250,1837,1859]" captionTargetBox="[156,1421,816,1802]" captionTargetId="figure@5.[151,1435,807,1815]" captionTargetPageId="5" captionText="FIGURE 4. South Korea, showing distributions of Liobagrus hyeongsanensis, L. somjinensis, L. mediadiposalis, L. obesus, and L. andersoni. Dense black lines: Baekdudaegan mountain range, showing three biogeographical ranges of ichthyofaunal endemisms in the Korean Peninsula (modified from Kim 1997)." httpUri="https://zenodo.org/record/236504/files/figure.png" pageId="3" pageNumber="270">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="C13636A9FFE07969A7B8FA52FE87C69D" blockId="3.[151,1436,1353,1738]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7B8FA52FE51C6FA" bold="true" box="[199,438,1425,1450]" pageId="3" pageNumber="270">Sexual dimorphism.</emphasis>
The adductor mandibulae in males is greatly expended during the spawning season, from late April to June.
</paragraph>
</subSubSection>
<subSubSection id="89936522FFE07969A7B8FA1AFE91C546" pageId="3" pageNumber="270" type="etymology">
<paragraph id="C13636A9FFE07969A7B8FA1AFE91C546" blockId="3.[151,1436,1353,1738]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7B8FA1AFEA9C6A2" bold="true" box="[199,334,1497,1522]" pageId="3" pageNumber="270">Etymology.</emphasis>
Named after the Hyeongsan River, the
<typeStatus id="1E32880BFFE07969A477FA19FCDFC6A2" box="[776,824,1498,1522]" pageId="3" pageNumber="270">type</typeStatus>
locality. We propose the Korean name Donbang-Jagasari for this species.
</paragraph>
</subSubSection>
<subSubSection id="89936522FFE07969A7B8F9E2FEA9C59A" pageId="3" pageNumber="270" type="biology_ecology">
<paragraph id="C13636A9FFE07969A7B8F9E2FEA9C59A" blockId="3.[151,1436,1353,1738]" pageId="3" pageNumber="270">
<emphasis id="F3FDEABBFFE07969A7B8F9E2FE5FC56A" bold="true" box="[199,440,1569,1594]" pageId="3" pageNumber="270">Biology and habitat.</emphasis>
<taxonomicName id="06894D2AFFE07969A6C0F9E1FD0FC56A" box="[447,744,1570,1594]" class="Actinopterygii" family="Amblycipitidae" genus="Liobagrus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="270" phylum="Chordata" rank="species" species="hyeongsanensis">
<emphasis id="F3FDEABBFFE07969A6C0F9E1FD0FC56A" box="[447,744,1570,1594]" italics="true" pageId="3" pageNumber="270">Liobagrus hyeongsanensis</emphasis>
</taxonomicName>
is small, nocturnal and benthic. It inhabits the bottom stratum of running waters with moderately fast currents, in upper streams and valley streams. It is generally seen in shallow waters (
<quantity id="06719B4CFFE07969A790F9A9FE84C5D1" box="[239,355,1642,1666]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="3" pageNumber="270" unit="m" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 m</quantity>
deep) with large or small stony or pebbly substrates. The spawning season is from late April to June. The adult females do not exceed 90.0 mm SL and lay a small number of eggs (60–110), 2.8–3.3 (3.1±0.2) mm in diameter.
</paragraph>
</subSubSection>
</treatment>
</document>
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