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<document id="9A7E4F5C828A7428577F395A47823270" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFFEBA01FC7BFD2DFC8AF8D8" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Danarma silus" docType="treatment" docVersion="10" lastPageNumber="923" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="922" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="36562A72C19B3511DAD42AAA7F26FA1C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D9B1E689ADC02F0EC0E9019B7F309C75">
<mods:title id="46DD7711489E8408E9C949776FBD5199">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:name id="BEAEEA7E483A66956078F881FEB9F9A6" type="personal">
<mods:role id="0B28F493A4FB7DEDB455327427E76A1C">
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<mods:namePart id="525F9286AC1C590F7C2ADA2A559B096B">Schubart, Christoph D.</mods:namePart>
<mods:affiliation id="34B8E02293BD9BDBFEE2837E7782DC58">Zoology &amp; Evolution, University of Regensburg, 93040 Regensburg, Germany</mods:affiliation>
<mods:nameIdentifier id="648F23C8D41738E8D7385B9E1029E0EB" type="email">christoph.schubart@ur.de</mods:nameIdentifier>
</mods:name>
<mods:name id="0C84C54CE280C327F4D7DCFED51C9603" type="personal">
<mods:role id="EE069E17FF2EDB9A2A6C3752B96F1732">
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</mods:role>
<mods:namePart id="A7329CA16805EC159647817DD7049C7E">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="6066FC05EE808D7F19DED9BD5D34D9B7">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
<mods:nameIdentifier id="0622EEA0438669E47245588CC853CB12" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="B7695FC9A867C776EF43EC64CAF7D791">text</mods:typeOfResource>
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<mods:titleInfo id="62844DB55841A114DB818E6600FC04CC">
<mods:title id="559A01BDB50120E3DFC722FC9C4EBC8B">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="49BC264337B36DDFF032DA424224B110">
<mods:date id="6DD0850C9D3F693B5D7558869F58E3B3">2020</mods:date>
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<mods:number id="712A617DA870344752399622FDBF609A">2020-12-23</mods:number>
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<mods:detail id="E52F2F12E82E1AC8DBB3AD07203ABFC1" type="volume">
<mods:number id="CD50DFEA812298D55898D62457292B16">68</mods:number>
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<mods:start id="F798491597656F7BB5EEC9AB21C858D9">891</mods:start>
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<mods:classification id="CD88C4640699ADE1E9A10503771EED04">journal article</mods:classification>
<mods:identifier id="B84207BE36353134135E58C3306C2ABD" type="CLB-Dataset">55667</mods:identifier>
<mods:identifier id="66F68A90D8FC5FA9B3263C42E5BAFBA5" type="DOI">10.26107/RBZ-2020-0097</mods:identifier>
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<mods:identifier id="E6B275657DE0EDFBEBE3292F6CF1E216" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="16D3358087A3A6BBD1DEF366077B953D" type="ZooBank">815E4670-B063-4FD8-B31E-3AD89B3A7942</mods:identifier>
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<treatment id="49152B56FFFEBA01FC7BFD2DFC8AF8D8" ID-DOI="http://doi.org/10.5281/zenodo.6524881" ID-GBIF-Taxon="195197176" ID-Zenodo-Dep="6524881" LSID="urn:lsid:plazi:treatment:49152B56FFFEBA01FC7BFD2DFC8AF8D8" httpUri="http://treatment.plazi.org/id/49152B56FFFEBA01FC7BFD2DFC8AF8D8" lastPageId="32" lastPageNumber="923" pageId="31" pageNumber="922">
<subSubSection id="89A6C9CBFFFEBA3EFC7BFD2DFABFFD04" box="[934,1330,669,693]" pageId="31" pageNumber="922" type="nomenclature">
<paragraph id="C1039A40FFFEBA3EFC7BFD2DFABFFD04" blockId="31.[934,1330,669,725]" box="[934,1330,669,693]" pageId="31" pageNumber="922">
<heading id="9A4B2D2CFFFEBA3EFC7BFD2DFABFFD04" bold="true" box="[934,1330,669,693]" centered="true" fontSize="10" level="2" pageId="31" pageNumber="922" reason="2">
<taxonomicName id="06BCE1C3FFFEBA3EFC7BFD2DFABFFD04" ID-CoL="84GST" authority="(Davie &amp; Ng, 2013)" baseAuthorityName="Davie &amp; Ng" baseAuthorityYear="2013" box="[934,1330,669,693]" class="Malacostraca" family="Sesarmidae" genus="Danarma" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="922" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="F3C84652FFFEBA3EFC7BFD2DFABFFD04" bold="true" box="[934,1330,669,693]" pageId="31" pageNumber="922">
<emphasis id="F3C84652FFFEBA3EFC7BFD2DFBC4FD04" bold="true" box="[934,1097,669,693]" italics="true" pageId="31" pageNumber="922">Danarma silus</emphasis>
(
<bibRefCitation id="A52DE7B1FFFEBA3EFB87FD2EFAA6FD04" author="Davie &amp; Ng" box="[1114,1323,669,693]" firstAuthor="Davie" pageId="31" pageNumber="922" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFEBA3EFBEEFD0EFB29FD64" box="[1075,1188,701,725]" pageId="31" pageNumber="922" type="description">
<paragraph id="C1039A40FFFEBA3EFBEEFD0EFB29FD64" blockId="31.[934,1330,669,725]" box="[1075,1188,701,725]" pageId="31" pageNumber="922">
(
<figureCitation id="598786C5FFFEBA3EFBE6FD0EFB10FD64" box="[1083,1181,701,725]" captionStart="Fig" captionStartId="26.[136,171,1910,1932]" captionTargetBox="[136,1474,199,1895]" captionTargetId="figure-8@26.[136,1474,199,1895]" captionTargetPageId="26" captionText="Fig. 13. Overall habitus. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Oahu, Hawaii; B, Danarma silus, holotype male (16.3 × 12.8 mm) (ZRC 2012.0787), Guam; C, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; D, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM), Greta Beach, Christmas Island; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; I, Platychirarma buettikoferi, lectotype male (14.1 × 11.6 mm) (RMNH D 148), Liberia; J, Platychirarma buettikoferi, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351323" httpUri="https://zenodo.org/record/5351323/files/figure.png" pageId="31" pageNumber="922">Fig. 13B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFEBA3EFCF0FD4FFB96FCD6" pageId="31" pageNumber="922" type="reference_group">
<paragraph id="C1039A40FFFEBA3EFCF0FD4FFBAEFC9F" blockId="31.[813,1451,764,871]" pageId="31" pageNumber="922">
<treatmentCitationGroup id="E1ACBD6EFFFEBA3EFCF0FD4FFBAEFC9F" pageId="31" pageNumber="922">
<taxonomicName id="06BCE1C3FFFEBA3EFCF0FD4FFBA8FCA0" ID-CoL="7S8NZ" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[813,1061,764,785]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="922" phylum="Arthropoda" rank="species" species="obtusifrons">
<emphasis id="F3C84652FFFEBA3EFCF0FD4FFBA8FCA0" box="[813,1061,764,785]" italics="true" pageId="31" pageNumber="922">Chiromantes obtusifrons</emphasis>
</taxonomicName>
–
<treatmentCitation id="401DBC51FFFEBA3EFB98FD4FFAB3FCA3" author="Paulay G &amp; Kropp R &amp; Ng PKL &amp; Eldredge LG" box="[1093,1342,764,786]" page="508" pageId="31" pageNumber="922" year="2003">
<bibRefCitation id="A52DE7B1FFFEBA3EFB98FD4FFAB3FCA3" author="Paulay" box="[1093,1342,764,786]" etAl="et al." firstAuthor="Paulay" page="508" pageId="31" pageNumber="922" pagination="456 - 513" refId="ref74719" refString="Paulay G, Kropp R, Ng PKL &amp; Eldredge LG (2003) The crustaceans and pycnogonids of the Mariana Islands. Micronesica, 35 - 36: 456 - 513." type="journal article" year="2003">Paulay et al., 2003: 508</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFFEBA3EFA91FD4FFC74FC9F" author="Paulay G &amp; Starmer J" page="2" pageId="31" pageNumber="922" year="2011">
<bibRefCitation id="A52DE7B1FFFEBA3EFA91FD4FFC74FC9F" author="Paulay &amp; Starmer" firstAuthor="Paulay" page="2" pageId="31" pageNumber="922" pagination="19916" refId="ref74752" refString="Paulay G &amp; Starmer J (2011) Evolution, insular restriction, and extinction of oceanic land crabs, exemplified by the loss of an endemic Geograpsus in the Hawaiian Islands. PLoS ONE, 6 (5): e 19916. doi: 10.1371 / journal. pone. 0019916" type="journal article" year="2011">Paulay &amp; Starmer, 2011: 2</bibRefCitation>
</treatmentCitation>
, 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFFEBA3EFCF0FC86FB96FCD6" blockId="31.[813,1451,764,871]" pageId="31" pageNumber="922">
<treatmentCitationGroup id="E1ACBD6EFFFEBA3EFCF0FC86FB96FCD6" pageId="31" pageNumber="922">
<treatmentCitation id="401DBC51FFFEBA3EFCF0FC86FB37FCFA" author="Davie PJF &amp; Ng PKL" box="[813,1210,821,843]" page="14" pageId="31" pageNumber="922" year="2013">
<taxonomicName id="06BCE1C3FFFEBA3EFCF0FC86FB37FCFA" ID-CoL="7TVP9" authority="Davie &amp; Ng, 2013: 14" authorityName="Davie &amp; Ng" authorityPageNumber="14" authorityYear="2013" box="[813,1210,821,843]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="31" pageNumber="922" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="F3C84652FFFEBA3EFCF0FC86FC56FCFB" box="[813,987,821,842]" italics="true" pageId="31" pageNumber="922">Chiromantes silus</emphasis>
<bibRefCitation id="A52DE7B1FFFEBA3EFC3CFC86FB37FCFA" author="Davie &amp; Ng" box="[993,1210,821,843]" firstAuthor="Davie" page="14" pageId="31" pageNumber="922" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; figs. 3A–C, 4B, 5C, 6C, 7C, 8B, 9C, 13A–E.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFEBA3EFCF0FC23FB42FAB8" pageId="31" pageNumber="922" type="materials_examined">
<paragraph id="C1039A40FFFEBA3EFCF0FC23FB42FAB8" blockId="31.[813,1452,912,1289]" pageId="31" pageNumber="922">
<emphasis id="F3C84652FFFEBA3EFCF0FC23FB8EFC19" bold="true" box="[813,1027,912,936]" pageId="31" pageNumber="922">Material examined</emphasis>
.
<materialsCitation id="71D4901DFFFEBA3EFBD2FC23FCE4FBB8" ID-GBIF-Occurrence="3763237482" collectedFrom="under limestone rocks" collectingDate="1997-10-05" collectionCode="ZRC" collectorName="G. Paulay" country="Guam" elevation="15" location="Tangulsson" pageId="31" pageNumber="922" specimenCode="ZRC 2012.0787" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="1E0724E2FFFEBA3EFBD2FC23FBFFFC19" box="[1039,1138,912,936]" pageId="31" pageNumber="922" type="holotype">Holotype</typeStatus>
<specimenCount id="D7BA51C9FFFEBA3EFBA5FC23FB21FC19" box="[1144,1196,912,936]" pageId="31" pageNumber="922" type="male">male</specimenCount>
(16.3 ×
<quantity id="064437A5FFFEBA3EFADEFC22FAEFFC18" box="[1283,1378,913,937]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="922" unit="mm" value="12.8">12.8 mm</quantity>
) (
<specimenCode id="911A323BFFFEBA3EFAABFC22FC2EFC78" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="922">ZRC 2012.0787</specimenCode>
),
<location id="C463CC9BFFFEBA3EFC6EFC03FBBCFC79" LSID="urn:lsid:plazi:treatment:49152B56FFFEBA01FC7BFD2DFC8AF8D8:C463CC9BFFFEBA3EFC6EFC03FBBCFC79" box="[947,1073,944,968]" country="Guam" name="Tangulsson" pageId="31" pageNumber="922">Tangulsson</location>
,
<collectingCountry id="B9ABDAD0FFFEBA3EFBE7FC02FBF3FC78" box="[1082,1150,945,969]" name="Guam" pageId="31" pageNumber="922">Guam</collectingCountry>
,
<quantity id="064437A5FFFEBA3EFB5AFC02FB68FC79" box="[1159,1253,945,969]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="31" pageNumber="922" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">
<elevation id="4A917D73FFFEBA3EFB5AFC02FB68FC79" box="[1159,1253,945,969]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="31" pageNumber="922" unit="m" value="15.0" valueMax="20.0" valueMin="10.0">10–20 m</elevation>
</quantity>
from shore,
<quantity id="064437A5FFFEBA3EFAB4FC03FA21FC79" box="[1385,1452,944,968]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="31" pageNumber="922" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">
<elevation id="4A917D73FFFEBA3EFAB4FC03FA21FC79" box="[1385,1452,944,968]" metricMagnitude="0" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="31" pageNumber="922" unit="m" value="6.5" valueMax="8.0" valueMin="5.0">5–8 m</elevation>
</quantity>
elevation,
<collectedFrom id="72A0D9BDFFFEBA3EFC41FC63FB07FC59" box="[924,1162,976,1000]" pageId="31" pageNumber="922">under limestone rocks</collectedFrom>
, coll.
<collectorName id="6C49FF96FFFEBA3EFB16FC62FABBFC59" box="[1227,1334,976,1001]" pageId="31" pageNumber="922">G. Paulay</collectorName>
,
<date id="B502BC80FFFEBA3EFA9CFC63FCEBFBB8" pageId="31" pageNumber="922" value="1997-10-05">
<collectingDate id="A5464568FFFEBA3EFA9CFC63FCEBFBB8" pageId="31" pageNumber="922" value="1997-10-05">5 October 1997</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="71D4901DFFFEBA3EFCADFC42FA1DFB99" ID-GBIF-Occurrence="3763237331" collectingDate="1997-10-05" collectionCode="ZRC" collectorName="G. Paulay" country="Guam" elevation="15" location="Tangulsson" pageId="31" pageNumber="922" specimenCode="ZRC 2012.0788" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="1E0724E2FFFEBA3EFCADFC42FC55FBB8" box="[880,984,1009,1033]" pageId="31" pageNumber="922" type="paratype">Paratypes</typeStatus>
–
<specimenCount id="D7BA51C9FFFEBA3EFC29FC42FBB1FBB9" box="[1012,1084,1008,1033]" pageId="31" pageNumber="922" type="male">1 male</specimenCount>
(15.1 ×
<quantity id="064437A5FFFEBA3EFB44FC42FB71FBB8" box="[1177,1276,1009,1033]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" pageId="31" pageNumber="922" unit="mm" value="11.3">11.3 mm</quantity>
),
<specimenCount id="D7BA51C9FFFEBA3EFAD2FC42FAE0FBB9" box="[1295,1389,1008,1033]" pageId="31" pageNumber="922" type="female">1 female</specimenCount>
(14.9 ×
<quantity id="064437A5FFFEBA3EFC9BFBA2FC21FB98" box="[838,940,1041,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="31" pageNumber="922" unit="mm" value="11.2">11.2 mm</quantity>
) (
<specimenCode id="911A323BFFFEBA3EFC19FBA2FBF0FB98" box="[964,1149,1041,1065]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="922">ZRC 2012.0788</specimenCode>
), same data as holotype
</materialsCitation>
;
<materialsCitation id="71D4901DFFFEBA3EFA43FBA2FB5FFBD8" ID-GBIF-Occurrence="3763237360" collectingDate="2001-08-03" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Guam" location="Hapato Beach" pageId="31" pageNumber="922" specimenCode="ZRC 2001.0742" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="D7BA51C9FFFEBA3EFA43FBA2FCECFBF9" pageId="31" pageNumber="922" type="male">1 male</specimenCount>
(15.7 ×
<quantity id="064437A5FFFEBA3EFC18FB82FBA5FBF8" box="[965,1064,1073,1097]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="31" pageNumber="922" unit="mm" value="11.7">11.7 mm</quantity>
) (
<specimenCode id="911A323BFFFEBA3EFB9CFB82FB7BFBF8" box="[1089,1270,1073,1097]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="922">ZRC 2001.0742</specimenCode>
),
<location id="C463CC9BFFFEBA3EFAD6FB82FA2AFBF9" LSID="urn:lsid:plazi:treatment:49152B56FFFEBA01FC7BFD2DFC8AF8D8:C463CC9BFFFEBA3EFAD6FB82FA2AFBF9" box="[1291,1447,1072,1096]" country="Guam" name="Hapato Beach" pageId="31" pageNumber="922">Hapato Beach</location>
,
<collectingCountry id="B9ABDAD0FFFEBA3EFCF0FBE2FCFCFBD8" box="[813,881,1105,1129]" name="Guam" pageId="31" pageNumber="922">Guam</collectingCountry>
, coll.
<collectorName id="6C49FF96FFFEBA3EFC6FFBE2FBAEFBD8" box="[946,1059,1105,1129]" pageId="31" pageNumber="922">P.K.L. Ng</collectorName>
,
<date id="B502BC80FFFEBA3EFBF2FBE2FB5FFBD8" box="[1071,1234,1105,1129]" pageId="31" pageNumber="922" value="2001-08-03">
<collectingDate id="A5464568FFFEBA3EFBF2FBE2FB5FFBD8" box="[1071,1234,1105,1129]" pageId="31" pageNumber="922" value="2001-08-03">3 August 2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFFEBA3EFB00FBE2FAB6FB18" ID-GBIF-Occurrence="3763237523" collectingDate="2001-08-04" collectionCode="ZRC" collectorName="P. K. L. Ng" country="Guam" location="Hapato Beach" pageId="31" pageNumber="922" specimenCode="ZRC 2001.0743" specimenCount="1" typeStatus="paratype">
1 ovigerous female (15.2 ×
<quantity id="064437A5FFFEBA3EFC59FBC3FC6AFB38" box="[900,999,1136,1161]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="31" pageNumber="922" unit="mm" value="11.6">11.6 mm</quantity>
) (with zoea 1) (
<specimenCode id="911A323BFFFEBA3EFB4AFBC2FAC6FB38" box="[1175,1355,1137,1161]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="922">ZRC 2001.0743</specimenCode>
),
<location id="C463CC9BFFFEBA3EFA83FBC2FCFEFB19" LSID="urn:lsid:plazi:treatment:49152B56FFFEBA01FC7BFD2DFC8AF8D8:C463CC9BFFFEBA3EFA83FBC2FCFEFB19" country="Guam" name="Hapato Beach" pageId="31" pageNumber="922">Hapato Beach</location>
,
<collectingCountry id="B9ABDAD0FFFEBA3EFC5CFB22FC4BFB18" box="[897,966,1169,1193]" name="Guam" pageId="31" pageNumber="922">Guam</collectingCountry>
, coll.
<collectorName id="6C49FF96FFFEBA3EFBD3FB22FB0EFB18" box="[1038,1155,1169,1193]" pageId="31" pageNumber="922">P.K.L. Ng</collectorName>
,
<date id="B502BC80FFFEBA3EFB4FFB22FAB6FB18" box="[1170,1339,1169,1193]" pageId="31" pageNumber="922" value="2001-08-04">
<collectingDate id="A5464568FFFEBA3EFB4FFB22FAB6FB18" box="[1170,1339,1169,1193]" pageId="31" pageNumber="922" value="2001-08-04">4 August 2001</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="71D4901DFFFEBA3EFA97FB22FB46FAB8" ID-GBIF-Occurrence="3763237384" collectingDate="2000-04-15" collectingDateMax="2000-04-18" collectingDateMin="2000-04-15" collectionCode="ZRC" collectorName="P. K. L. Ng &amp; C. - H. Wang" country="Guam" county="Marine Laboratory" location="Pago Bay" municipality="University" pageId="31" pageNumber="922" specimenCode="ZRC 2000.0573" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="D7BA51C9FFFEBA3EFA97FB22FA27FB19" box="[1354,1450,1168,1193]" pageId="31" pageNumber="922" type="female">1 female</specimenCount>
(15.2 ×
<quantity id="064437A5FFFEBA3EFC58FB02FC6AFB78" box="[901,999,1201,1225]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="31" pageNumber="922" unit="mm" value="11.2">11.2 mm</quantity>
) (
<specimenCode id="911A323BFFFEBA3EFC22FB02FB3EFB79" box="[1023,1203,1200,1225]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="31" pageNumber="922">ZRC 2000.0573</specimenCode>
),
<location id="C463CC9BFFFEBA3EFB1AFB02FABFFB78" LSID="urn:lsid:plazi:treatment:49152B56FFFEBA01FC7BFD2DFC8AF8D8:C463CC9BFFFEBA3EFB1AFB02FABFFB78" box="[1223,1330,1201,1225]" country="Guam" county="Marine Laboratory" municipality="University" name="Pago Bay" pageId="31" pageNumber="922">Pago Bay</location>
, outside of
<collectingMunicipality id="2167003AFFFEBA3EFCF0FB63FC13FB59" box="[813,926,1232,1256]" pageId="31" pageNumber="922">University</collectingMunicipality>
of
<collectingCountry id="B9ABDAD0FFFEBA3EFC1CFB62FB8FFB58" box="[961,1026,1233,1257]" name="Guam" pageId="31" pageNumber="922">Guam</collectingCountry>
<collectingCounty id="2862E2CCFFFEBA3EFBD4FB63FB5BFB59" box="[1033,1238,1232,1256]" pageId="31" pageNumber="922">Marine Laboratory</collectingCounty>
,
<collectingCountry id="B9ABDAD0FFFEBA3EFB3CFB62FAA8FB58" box="[1249,1317,1233,1257]" name="Guam" pageId="31" pageNumber="922">Guam</collectingCountry>
, coll.
<collectorName id="6C49FF96FFFEBA3EFAB8FB62FCC3FAB8" pageId="31" pageNumber="922">P.K.L. Ng</collectorName>
&amp;
<collectorName id="6C49FF96FFFEBA3EFCAEFB42FC74FAB8" box="[883,1017,1265,1289]" pageId="31" pageNumber="922">C.-H. Wang</collectorName>
,
<date id="B502BC80FFFEBA3EFBDBFB43FB46FAB8" box="[1030,1227,1264,1289]" pageId="31" pageNumber="922" value="2000-04-15" valueMax="2000-04-18" valueMin="2000-04-15">
<collectingDate id="A5464568FFFEBA3EFBDBFB43FB46FAB8" box="[1030,1227,1264,1289]" pageId="31" pageNumber="922" value="2000-04-15" valueMax="2000-04-18" valueMin="2000-04-15">15–18 April 2000</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFFEBA01FCF0FA83FD9FF998" lastPageId="32" lastPageNumber="923" pageId="31" pageNumber="922" type="diagnosis">
<paragraph id="C1039A40FFFEBA3EFCF0FA83FBBFF879" blockId="31.[812,1452,1328,1993]" pageId="31" pageNumber="922">
<emphasis id="F3C84652FFFEBA3EFCF0FA83FC11FAF9" bold="true" box="[813,924,1328,1352]" pageId="31" pageNumber="922">Diagnosis</emphasis>
. Carapace transversely subovate, ca. 1.3 times broader than long; dorsal carapace, lateral branchial regions markedly swollen; external orbital tooth weakly oblique, forming rounded obtuse angle posteriorly marking widest point of carapace, especially in larger specimens; front ca. 0.65 times carapace width, margin straight or slightly concave in dorsal view, appearing smooth but microscopically granular, with pair of low lateral swellings behind margin; supraorbital margin entire, semicircular; dorsal surface of cheliped carpus conspicuously granular; ambulatory legs relatively short; third ambulatory merus ca. 2.3 times longer than wide; fourth ambulatory merus similar, 2.4 times longer; third ambulatory propodus ca. 2.6 times longer than wide; fourth ambulatory merus ca. 2.3 times; male pleon moderately broad, somite 6 with lateral margins broadly divergent with margins relatively evenly convex, somite 3 width 3.2 times basal width of telson; G1 relatively slender, weakly tapering to broadly convex subdistal shoulder; distally slender, strongly bent to 45° angle; distal chitinous process long, with dorsal margin concave, apically pointed. (After
<bibRefCitation id="A52DE7B1FFFEBA3EFCF0F803FBABF879" author="Davie &amp; Ng" box="[813,1062,1968,1993]" firstAuthor="Davie" page="16" pageId="31" pageNumber="922" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013: 16</bibRefCitation>
).
</paragraph>
<paragraph id="C1039A40FFFEBA01FCF0F843FD9FF998" blockId="31.[813,1451,2032,2089]" lastBlockId="32.[136,776,1425,1577]" lastPageId="32" lastPageNumber="923" pageId="31" pageNumber="922">
<emphasis id="F3C84652FFFEBA3EFCF0F843FCF0F7B9" bold="true" box="[813,893,2032,2056]" pageId="31" pageNumber="922">Colour</emphasis>
. “Background colour of carapace typically maroon, with coarse paler orange to yellow-green blotching. Legs speckled with some indication of transverse banding. Legs and chelipeds dorsally reddish brown to dark orange, becoming paler orange in frontal and ventral view. Ocular peduncles speckled similar to carapace; corneas darker.” (
<bibRefCitation id="A52DE7B1FFC1BA01FF4DF9A2FE04F998" author="Davie &amp; Ng" box="[144,393,1553,1577]" firstAuthor="Davie" page="17" pageId="32" pageNumber="923" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013: 17</bibRefCitation>
, fig. 3A–C).
</paragraph>
</subSubSection>
<caption id="95C3CAC8FFC1BA01FF55FA94FB78FAE8" ID-DOI="http://doi.org/10.5281/zenodo.5351329" ID-Zenodo-Dep="5351329" httpUri="https://zenodo.org/record/5351329/files/figure.png" pageId="32" pageNumber="923" startId="32.[136,171,1319,1341]" targetBox="[136,1474,189,1303]" targetPageId="32">
<paragraph id="C1039A40FFC1BA01FF55FA94FB78FAE8" blockId="32.[136,1475,1318,1369]" pageId="32" pageNumber="923">
Fig. 16. Chela of species of
<taxonomicName id="06BCE1C3FFC1BA01FE79FA94FD0EFA8D" baseAuthorityName="Brito Capello" baseAuthorityYear="1864" box="[420,643,1319,1340]" class="Malacostraca" family="Sesarmidae" genus="Trapezarma" kingdom="Animalia" order="Decapoda" pageId="32" pageNumber="923" phylum="Arthropoda" rank="species" species="angolense">
<emphasis id="F3C84652FFC1BA01FE79FA94FD0EFA8D" box="[420,643,1319,1340]" italics="true" pageId="32" pageNumber="923">Trapezarma angolense</emphasis>
</taxonomicName>
. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; C–F, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view.
</paragraph>
</caption>
<subSubSection id="89A6C9CBFFC1BA01FF55F9E2FE04F938" pageId="32" pageNumber="923" type="discussion">
<paragraph id="C1039A40FFC1BA01FF55F9E2FE04F938" blockId="32.[136,775,1617,1673]" pageId="32" pageNumber="923">
<emphasis id="F3C84652FFC1BA01FF55F9E2FF7DF9D8" bold="true" box="[136,240,1617,1641]" pageId="32" pageNumber="923">Remarks</emphasis>
. The taxonomy of this species has been treated by
<bibRefCitation id="A52DE7B1FFC1BA01FF76F9C2FE08F938" author="Davie &amp; Ng" box="[171,389,1649,1673]" firstAuthor="Davie" pageId="32" pageNumber="923" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng (2013)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFC1BA01FF55F903FF75F898" pageId="32" pageNumber="923" type="biology_ecology">
<paragraph id="C1039A40FFC1BA01FF55F903FF75F898" blockId="32.[136,775,1712,1833]" pageId="32" pageNumber="923">
<emphasis id="F3C84652FFC1BA01FF55F903FF50F979" bold="true" box="[136,221,1712,1736]" pageId="32" pageNumber="923">Biology</emphasis>
. Lives under rocks and in karstic crevices in sparsely vegetated coastal platforms, up to ca.
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inland and ca.
<quantity id="064437A5FFC1BA01FF55F942FF4DF8B9" box="[136,192,1777,1801]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="923" unit="m" value="10.0">10 m</quantity>
elevation (
<bibRefCitation id="A52DE7B1FFC1BA01FEE4F942FDE3F8B8" author="Paulay &amp; Starmer" box="[313,622,1777,1801]" firstAuthor="Paulay" page="11" pageId="32" pageNumber="923" pagination="19916" refId="ref74752" refString="Paulay G &amp; Starmer J (2011) Evolution, insular restriction, and extinction of oceanic land crabs, exemplified by the loss of an endemic Geograpsus in the Hawaiian Islands. PLoS ONE, 6 (5): e 19916. doi: 10.1371 / journal. pone. 0019916" type="journal article" year="2011">Paulay &amp; Starmer, 2011: 11</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFC1BA01FDA6F942FF67F898" author="Davie &amp; Ng" firstAuthor="Davie" page="17" pageId="32" pageNumber="923" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013: 17</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFC1BA01FF55F8E3FC8AF8D8" box="[136,775,1872,1897]" pageId="32" pageNumber="923" type="distribution">
<paragraph id="C1039A40FFC1BA01FF55F8E3FC8AF8D8" blockId="32.[136,775,1872,1897]" box="[136,775,1872,1897]" pageId="32" pageNumber="923">
<emphasis id="F3C84652FFC1BA01FF55F8E3FE9EF8D9" bold="true" box="[136,275,1872,1896]" pageId="32" pageNumber="923">Distribution</emphasis>
. Known only from
<collectingCountry id="B9ABDAD0FFC1BA01FE39F8E2FDA8F8D8" box="[484,549,1873,1897]" name="Guam" pageId="32" pageNumber="923">Guam</collectingCountry>
(
<bibRefCitation id="A52DE7B1FFC1BA01FDEFF8E2FD71F8D8" author="Davie &amp; Ng" box="[562,764,1873,1897]" firstAuthor="Davie" pageId="32" pageNumber="923" pagination="1 - 25" refId="ref68942" refString="Davie PJF &amp; Ng PKL (2013) A review of Chiromantes obtusifrons (Dana, 1851) (Decapoda: Brachyura: Sesarmidae), with descriptions of four new sibling-species from Christmas Island, Guam and Taiwan. Zootaxa, 3609 (1): 1 - 25." type="journal article" year="2013">Davie &amp; Ng, 2013</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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