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<document id="1807B0FB348CFA665045BE60CC3642D5" ID-CLB-Dataset="55667" ID-DOI="10.26107/RBZ-2020-0097" ID-GBIF-Dataset="6cb155ce-8b9f-48ce-8e6c-b0dc00f8b1cc" ID-ISSN="2345-7600" ID-Zenodo-Dep="5351295" ID-ZooBank="815E4670-B063-4FD8-B31E-3AD89B3A7942" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1630205841501" checkinUser="diego" docAuthor="Schubart, Christoph D. &amp; Ng, Peter K. L." docDate="2020" docId="49152B56FFECBA2CFEE4FD8EFB96FAC4" docLanguage="en" docName="RafflesBZool.68.891-994.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Orisarma Schubart &amp; Ng 2020, new genus" docType="treatment" docUuid="65AE06A7-7D6B-4E62-8A26-223DE518AB3B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="904" masterDocId="B52C532EFFE1BA21FFDDFFB3FF8DFFB1" masterDocTitle="Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species" masterLastPageNumber="994" masterPageNumber="891" pageNumber="904" updateTime="1699203722016" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="547DEC4B03FCE62E88259E14C6A8FBE7">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<mods:namePart id="A9458057885080B76BEDB4294395F549">Schubart, Christoph D.</mods:namePart>
<mods:affiliation id="9624B9126D29D6D7156DF94E342D5817">Zoology &amp; Evolution, University of Regensburg, 93040 Regensburg, Germany</mods:affiliation>
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<mods:namePart id="93E117B95949468FFA2CC9B712D85EAD">Ng, Peter K. L.</mods:namePart>
<mods:affiliation id="DB357377273469478530F26768C23EA2">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore</mods:affiliation>
<mods:nameIdentifier id="641B6AA2736C01296B7A45E66421CA53" type="email">peterng@nus.edu.sg</mods:nameIdentifier>
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<treatment id="49152B56FFECBA2CFEE4FD8EFB96FAC4" ID-DOI="http://doi.org/10.5281/zenodo.6525277" ID-GBIF-Taxon="195197154" ID-Zenodo-Dep="6525277" LSID="urn:lsid:zoobank.org:act:65AE06A7-7D6B-4E62-8A26-223DE518AB3B" httpUri="http://treatment.plazi.org/id/49152B56FFECBA2CFEE4FD8EFB96FAC4" lastPageNumber="904" pageId="13" pageNumber="904">
<subSubSection id="89A6C9CBFFECBA2CFEE4FD8EFDA4FDE4" box="[313,553,573,597]" pageId="13" pageNumber="904" type="nomenclature">
<paragraph id="C1039A40FFECBA2CFEE4FD8EFDA4FDE4" blockId="13.[313,553,573,597]" box="[313,553,573,597]" pageId="13" pageNumber="904">
<heading id="9A4B2D2CFFECBA2CFEE4FD8EFDA4FDE4" bold="true" box="[313,553,573,597]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="904" reason="2">
<emphasis id="F3C84652FFECBA2CFEE4FD8EFDA4FDE4" bold="true" box="[313,553,573,597]" pageId="13" pageNumber="904">
<taxonomicName id="06BCE1C3FFECBA2CFEE4FD8EFE2FFDE4" ID-CoL="84GNY" authority="Schubart &amp; Ng, 2020" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[313,418,573,597]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F3C84652FFECBA2CFEE4FD8EFE2FFDE4" bold="true" box="[313,418,573,597]" italics="true" pageId="13" pageNumber="904">Orisarma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="E8FBFB29FFECBA2CFE6CFD8EFDA4FDE4" box="[433,553,573,597]" pageId="13" pageNumber="904" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="89A6C9CBFFECBA2CFFACFDCEFD9BFD04" pageId="13" pageNumber="904" type="type_taxon">
<paragraph id="C1039A40FFECBA2CFFACFDCEFD9BFD04" blockId="13.[113,752,637,693]" pageId="13" pageNumber="904">
<emphasis id="F3C84652FFECBA2CFFACFDCEFE8EFD24" bold="true" box="[113,259,637,661]" pageId="13" pageNumber="904">
<typeStatus id="1E0724E2FFECBA2CFFACFDCEFF26FD24" box="[113,171,637,661]" pageId="13" pageNumber="904">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="06BCE1C3FFECBA2CFEC9FDCDFD61FD24" ID-CoL="7Z299" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" box="[276,748,637,661]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis id="F3C84652FFECBA2CFEC9FDCDFE5DFD24" box="[276,464,637,661]" italics="true" pageId="13" pageNumber="904">Sesarma dehaani</emphasis>
H. 
<bibRefCitation id="A52DE7B1FFECBA2CFE20FDCEFD61FD24" author="Milne Edwards" box="[509,748,637,661]" firstAuthor="Milne Edwards" pageId="13" pageNumber="904" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
, by present designation. Gender neuter.
</paragraph>
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<subSubSection id="89A6C9CBFFECBA2CFFACFD6EFD7CF964" pageId="13" pageNumber="904" type="diagnosis">
<paragraph id="C1039A40FFECBA2CFFACFD6EFD7CF964" blockId="13.[113,753,733,1749]" pageId="13" pageNumber="904">
<emphasis id="F3C84652FFECBA2CFFACFD6EFF53FD44" bold="true" box="[113,222,733,757]" pageId="13" pageNumber="904">Diagnosis</emphasis>
. Carapace squarish to transversely subrectangular; frontal margin distinctly bilobed, gently deflexed, as wide as or slightly wider than posterior carapace margin; lateral margins of carapace entire, gently lobulated or with one epibranchial tooth, posterolateral part gently convex to subparallel; regions of carapace distinct to prominently demarcated; postfrontal and epigastric crests separated by relatively deep or distinct grooves, margin usually relatively rounded, regions clearly separated; basal articles of antenna and antennules clearly separated by septum formed by extension of front; dorsal margin of palm without any longitudinal pectinated ridge; in adult males, inner surface with prominent submedian transverse swelling, highest point with transverse ridge of granules, outer surface of palm and pollex convex, outer surface of palm usually with short longitudinal median smooth ridge (sometimes very low, obscured by granules); dorsal margin of chelipedal dactylus smooth, if small granules present, scattered, never in distinct row or of regular shape; inner distal margin of merus of cheliped not lamelliform; inner surfaces of first to third ambulatory coxae with only scattered short setae among them, not arranged into dense tufts; male thoracic sternites 2–4 relatively narrow in adults, suture between sternites 3 and 4 shallow, sometimes appearing medially interrupted; male sternopleonal cavity reaching two-thirds length of sternite 4 to just before anterior margin of sternite 2; male thoracic sternite 5 smooth, without depression on anterior part; G1 relatively slender, long or short, stout, chitinous part relatively short. Vulva on anterior part of sternite 6, anterior edge presses against sternite 5; anterior sternal vulvar cover very low, posterior sternal vulvar cover low, rim-like; opening projecting, directed anteriorly to obliquely anteriorly.
</paragraph>
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<subSubSection id="89A6C9CBFFECBA2CFFACF94EFDC3F8E4" pageId="13" pageNumber="904" type="etymology">
<paragraph id="C1039A40FFECBA2CFFACF94EFDC3F8E4" blockId="13.[113,754,1789,1878]" pageId="13" pageNumber="904">
<emphasis id="F3C84652FFECBA2CFFACF94EFF7AF8A4" bold="true" box="[113,247,1789,1813]" pageId="13" pageNumber="904">Etymology</emphasis>
. The name is derived from the arbritary combination of the term for 
<collectingCountry id="B9ABDAD0FFECBA2CFE71F8ADFD9BF884" box="[428,534,1821,1845]" name="China" pageId="13" pageNumber="904">East Asia</collectingCountry>
, “Oriental”, and the genus name 
<taxonomicName id="06BCE1C3FFECBA2CFF26F88DFEDAF8E4" authorityName="Say" authorityYear="1817" box="[251,343,1854,1877]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFF26F88DFEDAF8E4" box="[251,343,1854,1877]" italics="true" pageId="13" pageNumber="904">Sesarma</emphasis>
</taxonomicName>
. The gender is neuter.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFECBA2CFFACF8CEFF6AF784" pageId="13" pageNumber="904" type="discussion">
<paragraph id="C1039A40FFECBA2CFFACF8CEFF6AF784" blockId="13.[113,752,1917,2101]" pageId="13" pageNumber="904">
<emphasis id="F3C84652FFECBA2CFFACF8CEFEBAF824" bold="true" box="[113,311,1917,1941]" pageId="13" pageNumber="904">Included species</emphasis>
. 
<taxonomicName id="06BCE1C3FFECBA2CFE95F8CDFF2AF804" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis id="F3C84652FFECBA2CFE95F8CDFD86F824" box="[328,523,1917,1941]" italics="true" pageId="13" pageNumber="904">Sesarma dehaani</emphasis>
H. 
<bibRefCitation id="A52DE7B1FFECBA2CFDE1F8CEFF2AF804" author="Milne Edwards" firstAuthor="Milne Edwards" pageId="13" pageNumber="904" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
(= 
<taxonomicName id="06BCE1C3FFECBA2CFF14F82DFDAEF804" authority="Rathbun, 1897" authorityName="Rathbun" authorityYear="1897" box="[201,547,1949,1973]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="hanseni">
<emphasis id="F3C84652FFECBA2CFF14F82DFEF3F804" box="[201,382,1949,1973]" italics="true" pageId="13" pageNumber="904">Sesarma hanseni</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFE5EF82EFDAEF804" author="Rathbun" box="[387,547,1949,1973]" firstAuthor="Rathbun" pageId="13" pageNumber="904" pagination="89 - 92" refId="ref75039" refString="Rathbun MJ (1897) Synopsis of the American Sesarmae, with description of a new species. Proceedings of the Biological Society of Washington, 11: 89 - 92." type="journal article" year="1897">Rathbun, 1897</bibRefCitation>
</taxonomicName>
); 
<taxonomicName id="06BCE1C3FFECBA2CFDE8F82DFE9EF864" authority="De Man, 1887" authorityName="De Man" authorityYear="1887" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="neglecta">
<emphasis id="F3C84652FFECBA2CFDE8F82DFD7DF804" box="[565,752,1949,1973]" italics="true" pageId="13" pageNumber="904">Sesarma neglecta</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFFACF80DFE9EF864" author="De Man" box="[113,275,1981,2005]" firstAuthor="De Man" pageId="13" pageNumber="904" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="06BCE1C3FFECBA2CFEFFF80EFD61F864" authority="Komai &amp; Ng, 2013" authorityName="Komai &amp; Ng" authorityYear="2013" box="[290,748,1981,2006]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="magnus">
<emphasis id="F3C84652FFECBA2CFEFFF80EFD85F864" box="[290,520,1981,2005]" italics="true" pageId="13" pageNumber="904">Chiromantes magnus</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFDCFF80EFD61F864" author="Komai &amp; Ng" box="[530,748,1981,2006]" firstAuthor="Komai" pageId="13" pageNumber="904" pagination="539 - 551" refId="ref71033" refString="Komai T &amp; Ng PKL (2013) A new species of sesarmid crab of the genus Chiromantes (Crustacea: Decapoda: Brachyura) from the Ogasawara Islands, Japan. Zootaxa, 3681 (5): 539 - 551." type="journal article" year="2013">Komai &amp; Ng, 2013</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="06BCE1C3FFECBA2CFFACF86DFD07F844" authority="De Haan, 1835" authorityName="De Haan" authorityYear="1835" box="[113,650,2013,2037]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="intermedius" subGenus="Pachysoma">
<emphasis id="F3C84652FFECBA2CFFACF86DFF46F844" box="[113,203,2014,2037]" italics="true" pageId="13" pageNumber="904">Grapsus</emphasis>
(
<emphasis id="F3C84652FFECBA2CFF06F86EFEDBF844" box="[219,342,2013,2037]" italics="true" pageId="13" pageNumber="904">Pachysoma</emphasis>
) 
<emphasis id="F3C84652FFECBA2CFEB9F86EFE6FF844" box="[356,482,2013,2037]" italics="true" pageId="13" pageNumber="904">intermedius</emphasis>
De Haan, 1835
</taxonomicName>
; 
<taxonomicName id="06BCE1C3FFECBA2CFD49F86DFE54F7A4" authority="H. Milne Edwards, 1853" authorityName="H. Milne Edwards" authorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="F3C84652FFECBA2CFD49F86DFF48F7A4" italics="true" pageId="13" pageNumber="904">Sesarma sinensis</emphasis>
H. 
<bibRefCitation id="A52DE7B1FFECBA2CFF32F84EFE54F7A4" author="Milne Edwards" box="[239,473,2045,2069]" firstAuthor="Milne Edwards" pageId="13" pageNumber="904" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
</taxonomicName>
; 
<taxonomicName id="06BCE1C3FFECBA2CFE38F84EFF6EF784" authority="Soh, 1978" authorityName="Soh" authorityYear="1978" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="patshuni">
<emphasis id="F3C84652FFECBA2CFE38F84EFD7DF7A4" box="[485,752,2045,2069]" italics="true" pageId="13" pageNumber="904">Pseudosesarma patshuni</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFFACF7AEFF6EF784" author="Soh" box="[113,227,2077,2101]" firstAuthor="Soh" pageId="13" pageNumber="904" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFECBA2CFCF0FF0EFB96FAC4" pageId="13" pageNumber="904" type="discussion">
<paragraph id="C1039A40FFECBA2CFCF0FF0EFC37FC04" blockId="13.[812,1453,189,949]" pageId="13" pageNumber="904">
<emphasis id="F3C84652FFECBA2CFCF0FF0EFC1BFF64" bold="true" box="[813,918,189,213]" pageId="13" pageNumber="904">Remarks</emphasis>
. The major differences between 
<taxonomicName id="06BCE1C3FFECBA2CFACDFF0DFAF7FF64" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1296,1402,190,213]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFACDFF0DFAF7FF64" box="[1296,1402,190,213]" italics="true" pageId="13" pageNumber="904">Orisarma</emphasis>
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFECBA2CFCF0FF6EFC39FF44" authorityName="Gistel" authorityYear="1848" box="[813,948,221,245]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="F3C84652FFECBA2CFCF0FF6EFC39FF44" box="[813,948,221,245]" italics="true" pageId="13" pageNumber="904">Chiromantes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E8FBFB29FFECBA2CFC65FF6DFC62FF44" box="[952,1007,222,245]" pageId="13" pageNumber="904" sensu="stricto">s. str.</taxonomicNameLabel>
have been discussed under the latter genus. In addition to 
<taxonomicName id="06BCE1C3FFECBA2CFC65FF4EFA21FEA4" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[952,1452,253,277]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis id="F3C84652FFECBA2CFC65FF4EFB15FEA4" box="[952,1176,253,277]" italics="true" pageId="13" pageNumber="904">Chiromantes dehaani</emphasis>
(H. 
<bibRefCitation id="A52DE7B1FFECBA2CFB18FF4EFA29FEA4" author="Milne Edwards" box="[1221,1444,253,277]" firstAuthor="Milne Edwards" pageId="13" pageNumber="904" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFECBA2CFC84FEADFB1DFE84" authority="(De Man, 1887)" baseAuthorityName="De Man" baseAuthorityYear="1887" box="[857,1168,285,309]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="neglectus">
<emphasis id="F3C84652FFECBA2CFC84FEADFC50FE84" box="[857,989,285,309]" italics="true" pageId="13" pageNumber="904">C. neglectus</emphasis>
(
<bibRefCitation id="A52DE7B1FFECBA2CFC37FEAEFB05FE84" author="De Man" box="[1002,1160,285,309]" firstAuthor="De Man" pageId="13" pageNumber="904" pagination="639 - 722" refId="ref72089" refString="Man JG De (1887) Ubersicht der Indopacifischen Arten der Gattung Sesarma Say nebst einer Kritik der von W. Hess und E. Nauck in den Jahren 1865 und 1880 beschrieben Decapoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 2: 639 - 722." type="journal article" year="1887">De Man, 1887</bibRefCitation>
)
</taxonomicName>
, two species of 
<taxonomicName id="06BCE1C3FFECBA2CFAE5FEADFA21FE84" authorityName="Serene &amp; Soh" authorityYear="1970" box="[1336,1452,286,309]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFAE5FEADFA21FE84" box="[1336,1452,286,309]" italics="true" pageId="13" pageNumber="904">Sesarmops</emphasis>
</taxonomicName>
and one species of 
<taxonomicName id="06BCE1C3FFECBA2CFBDEFE8EFB27FEE4" authorityName=", Serene &amp; Soh" authorityYear="1970" box="[1027,1194,317,341]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFBDEFE8EFB27FEE4" box="[1027,1194,317,341]" italics="true" pageId="13" pageNumber="904">Pseudosesarma</emphasis>
</taxonomicName>
also need to be placed in 
<taxonomicName id="06BCE1C3FFECBA2CFC96FEEDFC3EFEC4" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[843,947,350,373]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFC96FEEDFC3EFEC4" box="[843,947,350,373]" italics="true" pageId="13" pageNumber="904">Orisarma</emphasis>
</taxonomicName>
. As 
<bibRefCitation id="A52DE7B1FFECBA2CFC37FEEEFB5BFEC4" author="Serene &amp; Soh" box="[1002,1238,349,373]" firstAuthor="Serene" pageId="13" pageNumber="904" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh (1970)</bibRefCitation>
had suspected (see Introduction), 
<taxonomicName id="06BCE1C3FFECBA2CFC1FFECDFAE5FE24" authority="(De Haan, 1835)" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[962,1384,381,405]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFECBA2CFC1FFECDFB38FE24" box="[962,1205,381,405]" italics="true" pageId="13" pageNumber="904">Sesarmops intermedius</emphasis>
(De Haan, 1835)
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFECBA2CFA4AFECDFB2EFE04" authority="(H. Milne Edwards, 1853)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="F3C84652FFECBA2CFA4AFECDFC0CFE04" italics="true" pageId="13" pageNumber="904">S. sinensis</emphasis>
(H. 
<bibRefCitation id="A52DE7B1FFECBA2CFC6FFE2EFB11FE04" author="Milne Edwards" box="[946,1180,413,437]" firstAuthor="Milne Edwards" pageId="13" pageNumber="904" pagination="163 - 228" refId="ref72839" refString="Milne Edwards H (1853) Memoires sur la famille des Ocypodiens, suite. Annales de Sciences Naturelle, Zoology, Paris, Serie 3 (Zoologie) 20: 163 - 228, pls. 6 - 11. [A continuation of H. Milne Edwards, 1852, and reprinted with it in the undated Melanges Carcinologiques pp. 129 - 196]" type="journal article" year="1853">Milne Edwards, 1853</bibRefCitation>
)
</taxonomicName>
are more closely related to each other and to species of 
<taxonomicName id="06BCE1C3FFECBA2CFB59FE0DFB67FE64" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1156,1258,446,469]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFB59FE0DFB67FE64" box="[1156,1258,446,469]" italics="true" pageId="13" pageNumber="904">Orisarma</emphasis>
</taxonomicName>
. Surprisingly, we also find that all the morphological characters of the small species, 
<taxonomicName id="06BCE1C3FFECBA2CFC52FE4EFAA4FDA4" authority="Soh, 1978" authorityName="Soh" authorityYear="1978" box="[911,1321,509,533]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="patshuni">
<emphasis id="F3C84652FFECBA2CFC52FE4EFB25FDA4" box="[911,1192,509,533]" italics="true" pageId="13" pageNumber="904">Pseudosesarma patshuni</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFB6FFE4EFAA4FDA4" author="Soh" box="[1202,1321,509,533]" firstAuthor="Soh" pageId="13" pageNumber="904" pagination="9 - 22" refId="ref77463" refString="Soh CL (1978) On a collection of sesarmine crabs (Decapoda, Brachyura, Grapsidae) from Hong Kong. Memoirs of the Hong Kong Natural History Society, 13: 9 - 22." type="journal article" year="1978">Soh, 1978</bibRefCitation>
</taxonomicName>
, also agree with 
<taxonomicName id="06BCE1C3FFECBA2CFCB7FDADFBF8FD84" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[874,1141,541,565]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="intermedius">
<emphasis id="F3C84652FFECBA2CFCB7FDADFBF8FD84" box="[874,1141,541,565]" italics="true" pageId="13" pageNumber="904">Sesarmops intermedius</emphasis>
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFECBA2CFB69FDADFAA3FD84" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1853" box="[1204,1326,542,565]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="F3C84652FFECBA2CFB69FDADFAA3FD84" box="[1204,1326,542,565]" italics="true" pageId="13" pageNumber="904">S. sinensis</emphasis>
</taxonomicName>
, including the form of the thoracic sternum, male chela with a low median longitudinal ridge, relatively wide male pleons, proportionately long ambulatory legs, structure of their G1s (
<figureCitation id="598786C5FFECBA2CFCB4FD2EFC62FD04" box="[873,1007,669,693]" captionStart="Fig" captionStartId="7.[113,148,1626,1648]" captionTargetBox="[113,1449,189,1611]" captionTargetId="figure-14@7.[113,1449,189,1611]" captionTargetPageId="7" captionText="Fig. 1. Overall habitus. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. dehaani, male (36.4 × 34.4 mm) (ZRC 2011.1027), Kumejima Island, Japan; E, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; F, O. magnum, paratype male (50.6 × 45.6 mm) (ZRC 2013.0173), Ogasawara Island, Japan; G, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; H, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351297" httpUri="https://zenodo.org/record/5351297/files/figure.png" pageId="13" pageNumber="904">Figs. 1G, H</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFC21FD2EFB97FD04" box="[1020,1050,669,693]" captionStart="Fig" captionStartId="10.[136,171,1435,1457]" captionTargetBox="[136,1476,189,1419]" captionTargetId="figure-7@10.[136,1476,189,1419]" captionTargetPageId="10" captionText="Fig. 3. Frontal view of cephalothorax. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; C, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; D, O. intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; E, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; F, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351301" httpUri="https://zenodo.org/record/5351301/files/figure.png" pageId="13" pageNumber="904">3F</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFBF5FD2EFBEEFD04" box="[1064,1123,669,693]" captionStart="Fig" captionStartId="14.[135,170,1987,2009]" captionTargetBox="[136,1476,189,1972]" captionTargetId="figure-7@14.[136,1476,189,1972]" captionTargetPageId="14" captionText="Fig. 5. Chela. A, B, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; C, C. haematocheir, male (18.6 × 20.7 mm) (RUMF-ZC-544), Japan; D–F, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; G, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong; I, O. patshuni, male (20.4 × 18.2 mm) (ZRC 2012.0032), Hong Kong. A, D, G, H, I, outer view; B, C, F, dorso-lateral view; E, inner view." figureDoi="http://doi.org/10.5281/zenodo.5351305" httpUri="https://zenodo.org/record/5351305/files/figure.png" pageId="13" pageNumber="904">5H, I</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFBACFD2EFB1EFD04" box="[1137,1171,669,693]" captionStart="Fig" captionStartId="20.[135,170,1977,1999]" captionTargetBox="[195,1417,189,1962]" captionTargetId="figure-7@20.[195,1417,189,1962]" captionTargetPageId="20" captionText="Fig. 9. Male anterior thoracic sternum and pleon. A, Chiromantes haematocheir, lectotype male (33.7 × 29.7 mm) (RMNH-D160), Japan; B, C. ryukyuanus, holotype male (33.1 × 29.6 mm) (RUMF-ZC-539), Okinawa Island, Japan; C, Orisarma dehaani, lectotype male (39.5 × 35.7 mm) (RMNH-D157), Japan; D, O. neglectum, neotype male (35.6 × 31.8 mm) (ZRC 1998.310), Shanghai, China; E, O. intermedium, lectotype male (23.0 × 19.9 mm) (RMNH-D165), Japan; F, O. sinense, lectotype male (19.0 × 16.6 mm) (MNHN-BP3635a), China; G, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; H, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351313" httpUri="https://zenodo.org/record/5351313/files/figure.png" pageId="13" pageNumber="904">9H</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFB7CFD2EFB7CFD04" box="[1185,1265,669,693]" captionStart="Fig" captionStartId="25.[113,148,1906,1928]" captionTargetBox="[116,1451,292,1890]" captionTargetId="figure-15@25.[99,1466,286,1902]" captionTargetPageId="25" captionText="Fig. 12. A–E, Orisarma intermedium, male (27.1 × 25.2 mm) (ZRC 1970.2.23.6), Japan; F–J, O. sinense, male (29.6 × 25.4 mm) (ZRC 1998.1204), Shanghai, China; K–O, O. patshuni, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong. A, F, K, male pleons; B, G, L, left G1 (ventral view, denuded); C, H, M, left G1 (dorsal view, denuded); D, I, N, left distal part of G1 (ventral view, denuded); E, J, O, left distal part of G1 (dorsal view, denuded). Scales: A, F = 5.0 mm; B–E, G–J, K–M = 1.0 mm; N, O = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351321" httpUri="https://zenodo.org/record/5351321/files/figure.png" pageId="13" pageNumber="904">12K–O</figureCitation>
) and even in the structure of the vulvae (
<figureCitation id="598786C5FFECBA2CFB9CFD0EFB21FD64" box="[1089,1196,701,725]" captionStart="Fig" captionStartId="72.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1882]" captionTargetId="figure-8@72.[136,1474,211,1882]" captionTargetPageId="72" captionText="Fig. 43. Vulva. A, Chiromantes haematocheir, female (27.0 × 23.7 mm) (ZRC 2002.0225), Japan; B, Orisarma dehaani, female (20.4 × 17.8 mm) (ZRC 2012.0057), Japan; C, Orisarma intermedium, female (31.8 × 28.4 mm) (ZRC 2013.0140), Japan; D, Orisarma patshuni, female (20.5 × 18.1 mm) (ZRC 2014.0817), Hong Kong; E, Danarma obtusifrons, female (15.7 × 11.6 mm) (UF-FLMNH 14837), Hawaii; F, Danarma eurymerus, paratype ovigerous female (18.0 × 13.4 mm) (ZRC 2012.0956), Taiwan; G, Cristarma eulimene, female (15.6 × 12.8 mm) (ZRC 1968.1.22.3), Mozambique; H, Cristarma ortmanni, female (13.1 × 10.2 mm) (ZRC 2000.1783), Kenya; I, Trapezarma angolense, female (18.2 × 14.3 mm) (ZRC 2015.0297), Cameroon; J, Platychirarma buettikoferi, ovigerous female (11.3 × 9.1 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351373" httpUri="https://zenodo.org/record/5351373/files/figure.png" pageId="13" pageNumber="904">Fig. 43D</figureCitation>
). The closest genus to 
<taxonomicName id="06BCE1C3FFECBA2CFCF0FD6DFC18FD44" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[813,917,734,757]" class="Malacostraca" family="Sesarmidae" genus="Orisarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFCF0FD6DFC18FD44" box="[813,917,734,757]" italics="true" pageId="13" pageNumber="904">Orisarma</emphasis>
</taxonomicName>
may be 
<taxonomicName id="06BCE1C3FFECBA2CFC28FD6DFBD3FD44" authority="Schubart &amp; Ng, 2020" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1013,1118,734,757]" class="Malacostraca" family="Sesarmidae" genus="Manarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F3C84652FFECBA2CFC28FD6DFBD3FD44" box="[1013,1118,734,757]" italics="true" pageId="13" pageNumber="904">Manarma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="E8FBFB29FFECBA2CFBB0FD6DFB6EFD47" box="[1133,1251,734,758]" pageId="13" pageNumber="904" rank="genus">new genus</taxonomicNameLabel>
, which consists of two species, 
<taxonomicName id="06BCE1C3FFECBA2CFC68FD4DFB6AFCA4" authority="(De Man, 1892)" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[949,1255,765,789]" class="Malacostraca" family="Sesarmidae" genus="Pseudosesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="moeschii">
<emphasis id="F3C84652FFECBA2CFC68FD4DFBBAFCA4" box="[949,1079,765,789]" italics="true" pageId="13" pageNumber="904">M. moeschii</emphasis>
(
<bibRefCitation id="A52DE7B1FFECBA2CFB99FD4EFB52FCA4" author="De Man" box="[1092,1247,765,789]" firstAuthor="De Man" pageId="13" pageNumber="904" pagination="225 - 264" refId="ref72297" refString="Man JG De (1892) Carcinological studies in the Leyden Museum, No. 6. Notes Leyden Museum, 14 (36): 225 - 264, pls. 7 - 10." type="journal article" year="1892">De Man, 1892</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="06BCE1C3FFECBA2CFAC4FD4DFC6CFC84" authority="(Tweedie, 1940)" baseAuthorityName="Tweedie" baseAuthorityYear="1940" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="johorensis">
<emphasis id="F3C84652FFECBA2CFAC4FD4DFA26FCA4" box="[1305,1451,765,789]" italics="true" pageId="13" pageNumber="904">M. johorensis</emphasis>
(
<bibRefCitation id="A52DE7B1FFECBA2CFCE9FCAEFC57FC84" author="Tweedie" box="[820,986,797,821]" firstAuthor="Tweedie" pageId="13" pageNumber="904" pagination="88 - 113" refId="ref78385" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. 24." type="journal article" year="1940">Tweedie, 1940</bibRefCitation>
)
</taxonomicName>
. These two species, as already highlighted by 
<bibRefCitation id="A52DE7B1FFECBA2CFC8CFC8EFBCCFCE4" author="Serene &amp; Soh" box="[849,1089,829,853]" firstAuthor="Serene" pageId="13" pageNumber="904" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh (1970)</bibRefCitation>
, are characterised by their shiny carapaces, narrow thoracic sterna, narrow male pleons, and form of their G1s (cf. 
<figureCitation id="598786C5FFECBA2CFBFCFCCEFB22FC24" box="[1057,1199,893,917]" captionStart="Fig" captionStartId="44.[136,172,1637,1659]" captionTargetBox="[136,1474,189,1621]" captionTargetId="figure-7@44.[136,1474,189,1621]" captionTargetPageId="44" captionText="Fig. 24. Overall habitus. A, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; B, Miersarma granosimanum, male (22.5 × 19.2 mm) (ZRC 1965.7.29.164), Sedili River, Johor, Malaysia; C, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; D, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; E, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; F, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; G, Bresedium laevimanum, male (18.1 × 16.4 mm) (ZRC 1972.3.7.25), Sarawak, Borneo; H, Bresedium laevimanum, male (21.2 × 18.2 mm) (ZRC 1965.7.29.121), Johor, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.5351343" httpUri="https://zenodo.org/record/5351343/files/figure.png" pageId="13" pageNumber="904">Figs. 24C, D</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFB61FCCEFA80FC24" box="[1212,1293,893,917]" captionStart="Fig" captionStartId="47.[113,149,901,923]" captionTargetBox="[113,1451,189,886]" captionTargetId="figure-530@47.[113,1451,189,886]" captionTargetPageId="47" captionText="Fig. 26. Frontal view of cephalothorax. A, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; B, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; C, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; D, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia." figureDoi="http://doi.org/10.5281/zenodo.5351345" httpUri="https://zenodo.org/record/5351345/files/figure.png" pageId="13" pageNumber="904">26A, B</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFAC7FCCEFAE7FC24" box="[1306,1386,893,917]" captionStart="Fig" captionStartId="51.[113,148,1446,1468]" captionTargetBox="[113,1451,189,1430]" captionTargetId="figure-15@51.[113,1451,189,1430]" captionTargetPageId="51" captionText="Fig. 29. Outer view of chela. A, Miersarma granosimanum, lectotype male (16.9 × 14.8 mm) (NHM 1880.6), Borneo; B, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; C, Manarma johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; D, Bresedium laevimanum, lectotype male (20.0 × 17.6 mm) (MNHG), Borneo; E, Bresedium laevimanum, male (26.3 × 23.7 mm) (NHM) (lectotype of Sesarma sediliensis), Johor, Malaysia; F, Bresedium laevimanum, male (18.1 × 16.4 mm) (ZRC 1972.3.7.25), Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5351349" httpUri="https://zenodo.org/record/5351349/files/figure.png" pageId="13" pageNumber="904">29B, C</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFAAAFCCEFC0EFC04" captionStart="Fig" captionStartId="56.[136,171,1595,1617]" captionTargetBox="[136,1474,189,1580]" captionTargetId="figure-7@56.[136,1474,189,1580]" captionTargetPageId="56" captionText="Fig. 33. A–E, male anterior thoracic sternum and pleon; F–H, left ambulatory leg. A, D, H, Manarma moeschii, male (18.2 × 16.3 mm) (ZRC 2000.1926), Thailand; B, E, M. johorense, male (14.6 × 12.6 mm) (ZRC 1971.9.24.14), Seletar River, Singapore; C, F, Orisarma intermedium, male (34.7 × 31.6 mm) (ZRC 2001.0034), Pingtung, Taiwan; G, O. patshuni (, male (14.2 × 13.0 mm) (ZRC 1998.345), Hong Kong." figureDoi="http://doi.org/10.5281/zenodo.5351357" httpUri="https://zenodo.org/record/5351357/files/figure.png" pageId="13" pageNumber="904">33A, B, D, E</figureCitation>
, 
<figureCitation id="598786C5FFECBA2CFC4DFC2EFC23FC04" box="[912,942,925,949]" captionStart="Fig" captionStartId="90.[136,171,1875,1897]" captionTargetBox="[136,1474,324,1860]" captionTargetId="figure-7@90.[136,1474,324,1860]" captionTargetPageId="90" captionText="Fig. 58. Colour in life. Manarma moeschii. A, B, male, Thailand, specimen not preserved [photograph: Oliver Mengedoht]; C–H, male (19.7 × 17.1 mm) (ZRC 2017.0170), Thailand [photograph: Paul Ng]." figureDoi="http://doi.org/10.5281/zenodo.5351403" httpUri="https://zenodo.org/record/5351403/files/figure.png" pageId="13" pageNumber="904">58</figureCitation>
).
</paragraph>
<paragraph id="C1039A40FFECBA2CFCF0FC6EFB96FAC4" blockId="13.[813,1452,989,1397]" pageId="13" pageNumber="904">
The remaining species of 
<taxonomicName id="06BCE1C3FFECBA2CFB90FC6DFA26FC44" authority="Serene &amp; Soh, 1970" authorityName="Serene &amp; Soh" authorityYear="1970" box="[1101,1451,989,1013]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFB90FC6DFB4CFC44" box="[1101,1217,990,1013]" italics="true" pageId="13" pageNumber="904">Sesarmops</emphasis>
<bibRefCitation id="A52DE7B1FFECBA2CFB16FC6EFA26FC44" author="Serene &amp; Soh" box="[1227,1451,989,1013]" firstAuthor="Serene" pageId="13" pageNumber="904" pagination="387 - 416" refId="ref76461" refString="Serene R &amp; Soh CL (1970) New IndoPacific genera allied to Sesarma Say 1877 (Brachyura, Decapoda, Crustacea). Treubia, 27: 387 - 416." type="journal article" year="1970">Serène &amp; Soh, 1970</bibRefCitation>
</taxonomicName>
(
<typeStatus id="1E0724E2FFECBA2CFCE9FC4EFCEEFBA4" box="[820,867,1021,1045]" pageId="13" pageNumber="904">type</typeStatus>
species 
<taxonomicName id="06BCE1C3FFECBA2CFC1FFC4DFA12FBA4" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[962,1439,1021,1045]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="impressa">
<emphasis id="F3C84652FFECBA2CFC1FFC4DFB0BFBA4" box="[962,1158,1022,1045]" italics="true" pageId="13" pageNumber="904">Sesarma impressa</emphasis>
H. Milne Edwards, 1837
</taxonomicName>
), have more trapezoidal carapaces and a relatively stout G1 with a very long chitinous distal process. One species still in 
<taxonomicName id="06BCE1C3FFECBA2CFC95FBEDFC31FBC4" authorityName="Serene &amp; Soh" authorityYear="1970" box="[840,956,1118,1141]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFC95FBEDFC31FBC4" box="[840,956,1118,1141]" italics="true" pageId="13" pageNumber="904">Sesarmops</emphasis>
</taxonomicName>
, 
<taxonomicName id="06BCE1C3FFECBA2CFC15FBEDFB53FBC4" authority="(De Man, 1902)" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[968,1246,1117,1141]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="weberi">
<emphasis id="F3C84652FFECBA2CFC15FBEDFBA6FBC4" box="[968,1067,1117,1141]" italics="true" pageId="13" pageNumber="904">S. weberi</emphasis>
(
<bibRefCitation id="A52DE7B1FFECBA2CFBE4FBEEFB5BFBC4" author="De Man" box="[1081,1238,1117,1141]" firstAuthor="De Man" pageId="13" pageNumber="904" pagination="467 - 929" refId="ref72431" refString="Man JG De (1902) Die von Herrn Professor Kukenthal im Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: Kukenthal W (ed.) Ergebnisse einer Zoologischen Forschungsreise in den Molukken und Borneo, in Auftrage der Senckenberg. Naturforsch. Gesellschaft ausgefuhrt von Dr. Willy Kukenthal. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25 (3): 467 - 929, pls. 19 - 27." type="journal article" year="1902">De Man, 1902</bibRefCitation>
)
</taxonomicName>
, does not belong to this genus on account of its short G1 and a male chelipedal dactylar finger that has a row of stridulatory granules on the dorsal margin (see 
<bibRefCitation id="A52DE7B1FFECBA2CFBFEFB0DFA84FB64" author="Ng" box="[1059,1289,1213,1238]" etAl="et al." firstAuthor="Ng" page="224" pageId="13" pageNumber="904" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 224</bibRefCitation>
; 
<bibRefCitation id="A52DE7B1FFECBA2CFAC8FB0DFCE5FB44" author="Naruse &amp; Ng" firstAuthor="Naruse" pageId="13" pageNumber="904" pagination="445 - 532" refId="ref73455" refString="Naruse T &amp; Ng PKL (2020) Revision of the sesarmid crab genera Labuanium Serene and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003, and Namlacium Serene and Soh, 1970 (Crustacea: Decapoda: Brachyura: Grapsoidea), with descriptions of four new genera and three new species. Journal of Natural History, 54 (7 - 8): 445 - 532." type="journal article" year="2020">Naruse &amp; Ng, 2020</bibRefCitation>
). 
<bibRefCitation id="A52DE7B1FFECBA2CFCA0FB6DFBB3FB44" author="Ng" box="[893,1086,1245,1270]" etAl="et al." firstAuthor="Ng" pageId="13" pageNumber="904" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al. (2008a)</bibRefCitation>
suggested the species was close to 
<taxonomicName id="06BCE1C3FFECBA2CFC94FB4EFA0FFAA4" authority="(H. Milne Edwards, 1837)" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[841,1410,1277,1301]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="species" species="trapezoideum">
<emphasis id="F3C84652FFECBA2CFC94FB4EFBD1FAA4" box="[841,1116,1277,1301]" italics="true" pageId="13" pageNumber="904">Labuanium trapezoideum</emphasis>
(H. Milne Edwards, 1837)
</taxonomicName>
but 
<bibRefCitation id="A52DE7B1FFECBA2CFCF0FAADFB96FA84" author="Naruse &amp; Ng" box="[813,1051,1309,1334]" firstAuthor="Naruse" pageId="13" pageNumber="904" pagination="445 - 532" refId="ref73455" refString="Naruse T &amp; Ng PKL (2020) Revision of the sesarmid crab genera Labuanium Serene and Soh, 1970, Scandarma Schubart, Liu and Cuesta, 2003, and Namlacium Serene and Soh, 1970 (Crustacea: Decapoda: Brachyura: Grapsoidea), with descriptions of four new genera and three new species. Journal of Natural History, 54 (7 - 8): 445 - 532." type="journal article" year="2020">Naruse &amp; Ng (2020)</bibRefCitation>
showed otherwise. The systematic position of this species should be dealt with when 
<taxonomicName id="06BCE1C3FFECBA2CFAEAFA8DFA27FAE4" authorityName="Serene &amp; Soh" authorityYear="1970" box="[1335,1450,1342,1365]" class="Malacostraca" family="Sesarmidae" genus="Sesarmops" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="904" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFECBA2CFAEAFA8DFA27FAE4" box="[1335,1450,1342,1365]" italics="true" pageId="13" pageNumber="904">Sesarmops</emphasis>
</taxonomicName>
is completely revised.
</paragraph>
</subSubSection>
</treatment>
</document>