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<mods:title id="197BAE8783775EFE01AF0F2545CBAEF9">Revision of the intertidal and semiterrestrial crab genera Chiromantes Gistel, 1848, and Pseudosesarma Serène &amp; Soh, 1970 (Crustacea: Brachyura: Sesarmidae), using morphology and molecular phylogenetics, with the establishment of nine new genera and two new species</mods:title>
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<emphasis id="F3C84652FFCCBA0CFF68FE6DFE3FFE44" bold="true" box="[181,434,477,501]" italics="true" pageId="45" pageNumber="936">Trapezarma angolense</emphasis>
(
<bibRefCitation id="A52DE7B1FFCCBA0CFE1FFE6EFD28FE44" author="Brito Capello" box="[450,677,477,501]" firstAuthor="Brito Capello" pageId="45" pageNumber="936" refId="ref67965" refString="Brito Capello F de (1864) Descripcao de tres especies novas de crustaceos da Africa Occidental e observacoes acerca do Penoeus [sic] bocagei. Johnson. Especie nova dos mares de Portugal. Typographia da Academia, Lisboa. 11 + [1 unnumbered page of errata] pp., 1 unnumbered plate with 10 figures." type="book" year="1864">Brito Capello, 1864</bibRefCitation>
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<paragraph id="C1039A40FFCCBA0CFF04FE4EFD04FDA4" blockId="45.[181,684,477,533]" box="[217,649,509,533]" pageId="45" pageNumber="936">
(
<figureCitation id="598786C5FFCCBA0CFF3CFE4EFEDDFDA4" box="[225,336,509,533]" captionStart="Fig" captionStartId="26.[136,171,1910,1932]" captionTargetBox="[136,1474,199,1895]" captionTargetId="figure-8@26.[136,1474,199,1895]" captionTargetPageId="26" captionText="Fig. 13. Overall habitus. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Oahu, Hawaii; B, Danarma silus, holotype male (16.3 × 12.8 mm) (ZRC 2012.0787), Guam; C, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; D, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM), Greta Beach, Christmas Island; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia; I, Platychirarma buettikoferi, lectotype male (14.1 × 11.6 mm) (RMNH D 148), Liberia; J, Platychirarma buettikoferi, male (12.6 × 10.4 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351323" httpUri="https://zenodo.org/record/5351323/files/figure.png" pageId="45" pageNumber="936">Figs. 13G</figureCitation>
,
<figureCitation id="598786C5FFCCBA0CFE80FE4EFE00FDA4" box="[349,397,509,533]" captionStart="Fig" captionStartId="28.[136,171,1850,1872]" captionTargetBox="[136,1474,287,1835]" captionTargetId="figure-7@28.[136,1474,287,1835]" captionTargetPageId="28" captionText="Fig. 14. Frontal view of cephalothorax. A, Danarma obtusifrons, male (17.7 × 13.2 mm) (UF-FLMNH 14837), Hawaii; B, Danarma garfunkel, holotype male (17.0 × 13.0 mm) (QM, ex ZRC 2009.0822), Christmas Island; C, D, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; E, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; F, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; G, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; H, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia." figureDoi="http://doi.org/10.5281/zenodo.5351325" httpUri="https://zenodo.org/record/5351325/files/figure.png" pageId="45" pageNumber="936">14G</figureCitation>
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<geoCoordinate id="A488FC87FFCCBA0CFE47FE4EFE7CFDA4" box="[410,497,509,533]" degrees="16, 18" direction="east" orientation="longitude" pageId="45" pageNumber="936" precision="555" value="16.18">
<figureCitation id="598786C5FFCCBA0CFE47FE4EFE3AFDA4" box="[410,439,509,533]" captionStart="Fig" captionStartId="32.[136,171,1319,1341]" captionTargetBox="[136,1474,189,1303]" captionTargetId="figure-8@32.[136,1474,189,1303]" captionTargetPageId="32" captionText="Fig. 16. Chela of species of Trapezarma angolense. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; CF, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5351329" httpUri="https://zenodo.org/record/5351329/files/figure.png" pageId="45" pageNumber="936">16</figureCitation>
,
<figureCitation id="598786C5FFCCBA0CFE19FE4EFE7CFDA4" box="[452,497,509,533]" captionStart="Fig" captionStartId="34.[136,171,1517,1539]" captionTargetBox="[136,1474,189,1501]" captionTargetId="figure-8@34.[136,1474,189,1501]" captionTargetPageId="34" captionText="Fig. 18. Male anterior thoracic sternum and pleon. A, Danarma obtusifrons, male (19.7 × 14.7 mm) (ZRC 2002.0220), Oahu, Hawaii; B, Danarma eurymerus, holotype male (18.7 × 14.8 mm) (NMNS-7028-002), Lanyu Island, Taiwan; C, Cristarma eulimene, male (22.2 × 17.2 mm) (ZRC 1968.1.22.2), Inhaca Island, Mozambique; D, Cristarma ortmanni, male (20.0 × 15.5 mm) (ZRC 1968.1.22.1), Inhaca Island, Mozambique; E, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; F, Platychirarma buettikoferi, paralectotype male (10.0 × 8.7 mm) (SMF-ZMG 636), Fisherman Lake, Liberia." figureDoi="http://doi.org/10.5281/zenodo.5351333" httpUri="https://zenodo.org/record/5351333/files/figure.png" pageId="45" pageNumber="936">18E</figureCitation>
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<figureCitation id="598786C5FFCCBA0CFE23FE4EFDC2FDA4" box="[510,591,509,533]" captionStart="Fig" captionStartId="40.[135,171,1614,1636]" captionTargetBox="[135,1474,190,1599]" captionTargetId="figure-8@40.[120,1494,180,1605]" captionTargetPageId="40" captionText="Fig. 21. AG, Trapezarma angolense, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; HL, Platychirarma buettikoferi, 1 male (10.7 × 8.8 mm) (SMF 25980), West Nimbe Cameroon. A, H, male pleon; B, I, left G1 (ventral view, denuded); C, left G1 (dorso-lateral view) (denuded); D, J, left G1 (dorsal view, denuded); E, K, left distal part of G1 (ventral view, denuded); G, L, left distal part of G1 (dorsal view, denuded); F, left distal part of G1 (dorso-lateral view) (denuded). Scales: A = 5.0 mm; BD, H = 2.0 mm; EG, I, J = 1.0 mm; K, L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5351339" httpUri="https://zenodo.org/record/5351339/files/figure.png" pageId="45" pageNumber="936">21AG</figureCitation>
,
<figureCitation id="598786C5FFCCBA0CFD81FE4EFD0CFDA4" box="[604,641,509,533]" captionStart="Fig" captionStartId="72.[136,171,1898,1920]" captionTargetBox="[136,1474,211,1882]" captionTargetId="figure-8@72.[136,1474,211,1882]" captionTargetPageId="72" captionText="Fig. 43. Vulva. A, Chiromantes haematocheir, female (27.0 × 23.7 mm) (ZRC 2002.0225), Japan; B, Orisarma dehaani, female (20.4 × 17.8 mm) (ZRC 2012.0057), Japan; C, Orisarma intermedium, female (31.8 × 28.4 mm) (ZRC 2013.0140), Japan; D, Orisarma patshuni, female (20.5 × 18.1 mm) (ZRC 2014.0817), Hong Kong; E, Danarma obtusifrons, female (15.7 × 11.6 mm) (UF-FLMNH 14837), Hawaii; F, Danarma eurymerus, paratype ovigerous female (18.0 × 13.4 mm) (ZRC 2012.0956), Taiwan; G, Cristarma eulimene, female (15.6 × 12.8 mm) (ZRC 1968.1.22.3), Mozambique; H, Cristarma ortmanni, female (13.1 × 10.2 mm) (ZRC 2000.1783), Kenya; I, Trapezarma angolense, female (18.2 × 14.3 mm) (ZRC 2015.0297), Cameroon; J, Platychirarma buettikoferi, ovigerous female (11.3 × 9.1 mm) (ZRC 2015.0298), Cameroon." figureDoi="http://doi.org/10.5281/zenodo.5351373" httpUri="https://zenodo.org/record/5351373/files/figure.png" pageId="45" pageNumber="936">43I</figureCitation>
)
</paragraph>
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<emphasis id="F3C84652FFCCBA0CFFACFD8FFEBFFDE0" box="[113,306,572,593]" italics="true" pageId="45" pageNumber="936">Sesarma angolensis</emphasis>
<bibRefCitation id="A52DE7B1FFCCBA0CFEE5FD8FFD9EFDE0" author="Brito Capello" box="[312,531,572,594]" firstAuthor="Brito Capello" page="4" pageId="45" pageNumber="936" refId="ref67965" refString="Brito Capello F de (1864) Descripcao de tres especies novas de crustaceos da Africa Occidental e observacoes acerca do Penoeus [sic] bocagei. Johnson. Especie nova dos mares de Portugal. Typographia da Academia, Lisboa. 11 + [1 unnumbered page of errata] pp., 1 unnumbered plate with 10 figures." type="book" year="1864">Brito Capello, 1864: 4</bibRefCitation>
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, fig. 2;
<bibRefCitation id="A52DE7B1FFCCBA0CFD83FD8FFF30FDDE" author="De Man" firstAuthor="De Man" page="161" pageId="45" pageNumber="936" pagination="150 - 169" refId="ref72057" refString="Man JG De (1883) Carcinological Studies in the Leyden Museum. No. 3. Notes from the Leyden Museum, 5 (3): 150 - 169." type="journal article" year="1883">
De
<treatmentCitation id="401DBC51FFCCBA0CFD5CFD8FFF30FDDE" author="Man JG De" page="161" pageId="45" pageNumber="936" year="1883">Man, 1883: 161</treatmentCitation>
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;
<treatmentCitation id="401DBC51FFCCBA0CFF16FDEAFE2EFDDF" author="Buttikofer J" box="[203,419,601,623]" page="487" pageId="45" pageNumber="936" year="1890">
<bibRefCitation id="A52DE7B1FFCCBA0CFF16FDEAFE2EFDDF" author="Buttikofer" box="[203,419,601,623]" firstAuthor="Buttikofer" page="487" pageId="45" pageNumber="936" refId="ref68064" refString="Buttikofer J (1890) Reisebilder aus Liberia: Resultate geographischer, naturwissenschaftlicher und ethnographischer Untersuchungen wahrend der Jahre 1879 - 1882 und 1886 - 1887, 2. Brill, Leiden, viii + 510 pp." type="book" year="1890">Büttikofer, 1890: 487</bibRefCitation>
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;
<treatmentCitation id="401DBC51FFCCBA0CFE6DFDEAFDF6FDDE" author="Johnston H" box="[432,635,601,623]" page="861" pageId="45" pageNumber="936" year="1906">
<bibRefCitation id="A52DE7B1FFCCBA0CFE6DFDEAFDF6FDDE" author="Johnston" box="[432,635,601,623]" firstAuthor="Johnston" page="861" pageId="45" pageNumber="936" pagination="860 - 883" refId="ref70766" refString="Johnston H (1906) List of Invertebrate Animals of Liberia Founded on the Collections of Buttikofer, Reynolds, Whicker, H. H. Johnston, A. White, etc. In: Johnston H (ed.) Liberia with an appendix on the flora of Libera by Dr. O. Stapf. Volume 2. Hutchinson &amp; Co., London. Pp. 860 - 883." type="book chapter" year="1906">Johnston, 1906: 861</bibRefCitation>
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;
<treatmentCitation id="401DBC51FFCCBA0CFD5AFDEAFF7CFD3A" author="Longhurst AR" page="88" pageId="45" pageNumber="936" year="1958">
<bibRefCitation id="A52DE7B1FFCCBA0CFD5AFDEAFF7CFD3A" author="Longhurst" firstAuthor="Longhurst" page="88" pageId="45" pageNumber="936" refId="ref71936" refString="Longhurst AR (1958) An Ecological Survey of the West African Marine Benthos. Colonial Office, Fishery Publications, Issue 11. H. M. Stationery Office, London, 102 pp., 11 figs." type="book" year="1958">Longhurst, 1958: 88</bibRefCitation>
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;
<treatmentCitation id="401DBC51FFCCBA0CFF21FDC6FE13FD3A" author="Gauld DT" box="[252,414,629,651]" page="71" pageId="45" pageNumber="936" year="1960">
<bibRefCitation id="A52DE7B1FFCCBA0CFF21FDC6FE13FD3A" author="Gauld" box="[252,414,629,651]" firstAuthor="Gauld" page="71" pageId="45" pageNumber="936" pagination="68 - 72" refId="ref69374" refString="Gauld DT (1960) Brachyura: An annotated check-list of the Crustacea of Ghana, IV. Journal of the West African Science Association, 6 (1): 68 - 72." type="journal article" year="1960">Gauld, 1960: 71</bibRefCitation>
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.
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<emphasis id="F3C84652FFCCBA0CFFACFD20FF49FD19" box="[113,196,659,680]" italics="true" pageId="45" pageNumber="936">Sesarma</emphasis>
(
<emphasis id="F3C84652FFCCBA0CFF0EFD21FEDEFD16" box="[211,339,658,679]" italics="true" pageId="45" pageNumber="936">Holometopus</emphasis>
)
<emphasis id="F3C84652FFCCBA0CFEBCFD21FE45FD16" box="[353,456,658,679]" italics="true" pageId="45" pageNumber="936">angolensis</emphasis>
</taxonomicName>
<bibRefCitation id="A52DE7B1FFCCBA0CFE3CFD20FD19FD19" author="De Man" box="[481,660,658,680]" firstAuthor="De Man" page="59" pageId="45" pageNumber="936" pagination="31 - 65" refId="ref72390" refString="Man JG De (1900) Note sur une petite collection de Crustaces decapodes provenant de la cote d'Angola (Afrique occidentale). Memoires de la Societe Zoologique de France, 13: 31 - 65, 2 pls." type="journal article" year="1900">
De
<treatmentCitation id="401DBC51FFCCBA0CFDD9FD20FD19FD19" author="Man JG De" box="[516,660,658,680]" page="59" pageId="45" pageNumber="936" year="1900">Man, 1900: 59</treatmentCitation>
</bibRefCitation>
, pl. 2 fig. 11;
<treatmentCitation id="401DBC51FFCCBA0CFF66FD1CFEEFFD75" author="Tesch JJ" box="[187,354,687,709]" page="130" pageId="45" pageNumber="936" year="1917">
<bibRefCitation id="A52DE7B1FFCCBA0CFF66FD1CFEEFFD75" author="Tesch" box="[187,354,687,709]" firstAuthor="Tesch" page="130" pageId="45" pageNumber="936" pagination="127 - 260" refId="ref78063" refString="Tesch JJ (1917) Synopsis of the genera Sesarma, Metasesarma, Sarmatium and Clistocoeloma, with a key to the determination of the IndoPacific species. Zoologische Mededelingen, 3: 127 - 260, pls. 15 - 17." type="journal article" year="1917">Tesch, 1917: 130</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFCCBA0CFEB6FD1CFDBCFD75" author="Dartevelle E" box="[363,561,687,709]" page="48" pageId="45" pageNumber="936" year="1950">
<bibRefCitation id="A52DE7B1FFCCBA0CFEB6FD1CFDBCFD75" author="Dartevelle" box="[363,561,687,709]" firstAuthor="Dartevelle" page="48" pageId="45" pageNumber="936" pagination="1 - 58" refId="ref68782" refString="Dartevelle E (1950) La cote et l'estuaire du Congo. Memoires de l'Institut Royal Colonial Belge, Section de Sciences Naturelles et Medicales, 19 (2): 1 - 58, figs. 1 - 8." type="journal article" year="1950">Dartevelle, 1950: 48</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFCCBA0CFDE7FD1CFD61FD74" author="Monod T" box="[570,748,687,709]" page="445" pageId="45" pageNumber="936" year="1956">
<bibRefCitation id="A52DE7B1FFCCBA0CFDE7FD1CFD61FD74" author="Monod" box="[570,748,687,709]" firstAuthor="Monod" page="445" pageId="45" pageNumber="936" pagination="1 - 674" refId="ref72973" refString="Monod T (1956) Hippidea et Brachyura ouest-africains. Memoires de I'Institut Francais d'Afrique Noire, 45: 1 - 674, 884 figs." type="journal article" year="1956">Monod, 1956: 445</bibRefCitation>
</treatmentCitation>
, fig. 605;
<treatmentCitation id="401DBC51FFCCBA0CFF2CFD7FFE49FD50" author="Rossignol M" box="[241,452,716,738]" page="120" pageId="45" pageNumber="936" year="1962">
<bibRefCitation id="A52DE7B1FFCCBA0CFF2CFD7FFE49FD50" author="Rossignol" box="[241,452,716,738]" firstAuthor="Rossignol" page="120" pageId="45" pageNumber="936" pagination="111 - 138" refId="ref75407" refString="Rossignol M (1962) Catalogue de Crustaces Decapodes Brachyoures, Anomoures et Macroures littoraux en collection au centre d'oceanographie de Pointe-Noire. Travaux du Centre Oceanographique de Pointe-Noire, 2: 111 - 138, pls. 1 - 4." type="journal article" year="1962">Rossignol, 1962: 120</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFCCBA0CFFACFD5AFEE9FCE5" blockId="45.[113,753,572,968]" pageId="45" pageNumber="936">
<treatmentCitationGroup id="E1ACBD6EFFCCBA0CFFACFD5AFEE9FCE5" pageId="45" pageNumber="936">
<taxonomicName id="06BCE1C3FFCCBA0CFFACFD5AFE5EFD4F" ID-CoL="9DBPG" authorityName="Brito Capello" authorityYear="1864" box="[113,467,745,766]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="species" species="angolense" subGenus="Holometopus">
<emphasis id="F3C84652FFCCBA0CFFACFD5AFF4BFD4F" box="[113,198,745,766]" italics="true" pageId="45" pageNumber="936">Sesarma</emphasis>
(
<emphasis id="F3C84652FFCCBA0CFF05FD5AFED1FD4F" box="[216,348,745,766]" italics="true" pageId="45" pageNumber="936">Holometopus</emphasis>
)
<emphasis id="F3C84652FFCCBA0CFEB2FD5AFE5EFD4F" box="[367,467,745,766]" italics="true" pageId="45" pageNumber="936">angolense</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFCCBA0CFE2FFD5AFD32FD4E" author="Rathbun MJ" box="[498,703,745,767]" page="451" pageId="45" pageNumber="936" year="1921">
<bibRefCitation id="A52DE7B1FFCCBA0CFE2FFD5AFD32FD4E" author="Rathbun" box="[498,703,745,767]" firstAuthor="Rathbun" page="451" pageId="45" pageNumber="936" pagination="379 - 484" refId="ref75176" refString="Rathbun MJ (1921) The Brachyuran crabs collected by the American Museum Congo Expedition, 1909 - 1915. In: Scientific results of the American Museum Congo Expedition. General Invertebrate Zoology, No. 6. Bulletin of the American Museum of Natural History, 43: 379 - 484, figs. 1 - 22, pls. 15 - 44." type="journal article" year="1921">Rathbun, 1921: 451</bibRefCitation>
</treatmentCitation>
, pls. 43, 45 fig. 1;
<treatmentCitation id="401DBC51FFCCBA0CFEC1FCB6FE68FCAA" author="Dartevelle E" box="[284,485,773,795]" page="50" pageId="45" pageNumber="936" year="1950">
<bibRefCitation id="A52DE7B1FFCCBA0CFEC1FCB6FE68FCAA" author="Dartevelle" box="[284,485,773,795]" firstAuthor="Dartevelle" page="50" pageId="45" pageNumber="936" pagination="1 - 58" refId="ref68782" refString="Dartevelle E (1950) La cote et l'estuaire du Congo. Memoires de l'Institut Royal Colonial Belge, Section de Sciences Naturelles et Medicales, 19 (2): 1 - 58, figs. 1 - 8." type="journal article" year="1950">Dartevelle, 1950: 50</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFCCBA0CFE2DFCB5FD2DFCAA" author="Capart A" box="[496,672,773,795]" page="191" pageId="45" pageNumber="936" year="1951">
<bibRefCitation id="A52DE7B1FFCCBA0CFE2DFCB5FD2DFCAA" author="Capart" box="[496,672,773,795]" firstAuthor="Capart" page="191" pageId="45" pageNumber="936" pagination="11 - 205" refId="ref68246" refString="Capart A (1951) Crustaces Decapodes Brachyures. In: Expedition Oceanographique Belge dans les Eaux Cotieres Africaines de I'Atlantique Sud (1948 - 1949), Resultats Scientifiques, 3 (1): 11 - 205, figs. 1 - 80, pls. 1 - 3." type="journal article" year="1951">Capart, 1951: 191</bibRefCitation>
</treatmentCitation>
, fig. 76;
<treatmentCitation id="401DBC51FFCCBA0CFF4AFC91FEEFFC86" author="Rossignol M" box="[151,354,802,824]" page="92" pageId="45" pageNumber="936" year="1957">
<bibRefCitation id="A52DE7B1FFCCBA0CFF4AFC91FEEFFC86" author="Rossignol" box="[151,354,802,824]" firstAuthor="Rossignol" page="92" pageId="45" pageNumber="936" pagination="1 - 3" refId="ref75333" refString="Rossignol M (1957) Crustaces Decapodes marins de la region de Pointe-Noire. In: Collignon J, Rossignol M &amp; Roux C (eds.) Mollusques, Crustaces, Poissons Marins des cotes d'A. E. F. en collection au Centre d'Oceanographie de d'Institut d'Etudes Centrafricaines de Pointe-Noire. Paris. Pp. 71 - 136, figs. 1 - 20, pls. 1 - 3." type="book chapter" year="1957">Rossignol, 1957: 92</bibRefCitation>
</treatmentCitation>
, 122 [key];
<treatmentCitation id="401DBC51FFCCBA0CFE3CFC91FD15FC86" author="Jordan HD" box="[481,664,802,824]" page="198" pageId="45" pageNumber="936" year="1957">
<bibRefCitation id="A52DE7B1FFCCBA0CFE3CFC91FD15FC86" author="Jordan" box="[481,664,802,824]" firstAuthor="Jordan" page="198" pageId="45" pageNumber="936" pagination="197 - 206" refId="ref70840" refString="Jordan HD (1957) Crabs as pests of rice on tidal swamps. Empire Journal of Experimental Agriculture, 25 (99): 197 - 206." type="journal article" year="1957">
<collectingCountry id="B9ABDAD0FFCCBA0CFE3CFC91FDA9FC89" box="[481,548,802,824]" name="Jordan" pageId="45" pageNumber="936">Jordan</collectingCountry>
, 1957: 198
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="401DBC51FFCCBA0CFD7BFC90FF71FCE4" author="Humes AG" page="186" pageId="45" pageNumber="936" year="1957">
<bibRefCitation id="A52DE7B1FFCCBA0CFD7BFC90FF71FCE4" author="Humes" firstAuthor="Humes" page="186" pageId="45" pageNumber="936" pagination="180 - 194" refId="ref70589" refString="Humes AG (1957) The genus Cancrincola (Copepoda, Harpacticoida) on the West Coast of Africa. Bulletin de l'Institut Francais d'Afrique Noire, (A) 19 (1): 180 - 194, pls. 1 - 3." type="journal article" year="1957">Humes, 1957: 186</bibRefCitation>
</treatmentCitation>
, 187, 189.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFCCBA0CFFACFCEFFD22FCC0" blockId="45.[113,753,572,968]" box="[113,687,860,882]" pageId="45" pageNumber="936">
<treatmentCitationGroup id="E1ACBD6EFFCCBA0CFFACFCEFFD22FCC0" box="[113,687,860,882]" pageId="45" pageNumber="936">
<taxonomicName id="06BCE1C3FFCCBA0CFFACFCEFFE46FCC0" ID-CoL="84HBP" authorityName="Brito Capello" authorityYear="1864" box="[113,459,860,882]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="species" species="angolensis" subGenus="Parasesarma">
<emphasis id="F3C84652FFCCBA0CFFACFCEFFF49FCC0" box="[113,196,860,881]" italics="true" pageId="45" pageNumber="936">Sesarma</emphasis>
(
<emphasis id="F3C84652FFCCBA0CFF09FCEEFED9FCC3" box="[212,340,861,882]" italics="true" pageId="45" pageNumber="936">Parasesarma</emphasis>
)
<emphasis id="F3C84652FFCCBA0CFEB9FCEFFE46FCC0" box="[356,459,860,881]" italics="true" pageId="45" pageNumber="936">angolensis</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFCCBA0CFE3BFCEFFD26FCC0" author="Rathbun MJ" box="[486,683,860,882]" page="280" pageId="45" pageNumber="936" year="1900">
<bibRefCitation id="A52DE7B1FFCCBA0CFE3BFCEFFD26FCC0" author="Rathbun" box="[486,683,860,882]" firstAuthor="Rathbun" page="280" pageId="45" pageNumber="936" pagination="271 - 316" refId="ref75071" refString="Rathbun MJ (1900) The decapod crustaceans of West Africa. Proceedings of the United States National Museum, 22 (1199): 271 - 316." type="journal article" year="1900">Rathbun, 1900: 280</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFCCBA0CFFACFCCAFF4FFC1A" blockId="45.[113,753,572,968]" pageId="45" pageNumber="936">
<treatmentCitationGroup id="E1ACBD6EFFCCBA0CFFACFCCAFF4FFC1A" pageId="45" pageNumber="936">
<taxonomicName id="06BCE1C3FFCCBA0CFFACFCCAFE4FFC3F" ID-CoL="9DBPG" authorityName="Brito Capello" authorityYear="1864" box="[113,450,889,910]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="species" species="angolense" subGenus="Chiromantes">
<emphasis id="F3C84652FFCCBA0CFFACFCCAFF49FC3F" box="[113,196,889,910]" italics="true" pageId="45" pageNumber="936">Sesarma</emphasis>
(
<emphasis id="F3C84652FFCCBA0CFF09FCCAFEDDFC3F" box="[212,336,889,910]" italics="true" pageId="45" pageNumber="936">Chiromantes</emphasis>
)
<emphasis id="F3C84652FFCCBA0CFEBCFCCAFE4FFC3F" box="[353,450,889,910]" italics="true" pageId="45" pageNumber="936">angolense</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFCCBA0CFE02FCCAFF33FC1A" author="Manning RB &amp; Holthuis LB" page="243" pageId="45" pageNumber="936" year="1981">
<bibRefCitation id="A52DE7B1FFCCBA0CFE02FCCAFF33FC1A" author="Manning &amp; Holthuis" firstAuthor="Manning" page="243" pageId="45" pageNumber="936" pagination="1 - 379" refId="ref72504" refString="Manning RB &amp; Holthuis LB (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306: i - xii, 1 - 379, figs. 1 - 88, table 1." type="journal article" year="1981">Manning &amp; Holthuis, 1981: 243</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C1039A40FFCCBA0CFFACFC01FDD2FC76" blockId="45.[113,753,572,968]" box="[113,607,946,968]" pageId="45" pageNumber="936">
<treatmentCitationGroup id="E1ACBD6EFFCCBA0CFFACFC01FDD2FC76" box="[113,607,946,968]" pageId="45" pageNumber="936">
<taxonomicName id="06BCE1C3FFCCBA0CFFACFC01FEE6FC76" ID-CoL="7NSNF" authorityName="Gistel" authorityYear="1848" box="[113,363,946,967]" class="Malacostraca" family="Sesarmidae" genus="Chiromantes" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="genus">
<emphasis id="F3C84652FFCCBA0CFFA1FC01FF75FC76" box="[124,248,946,967]" italics="true" pageId="45" pageNumber="936">Chiromantes</emphasis>
<emphasis id="F3C84652FFCCBA0CFED7FC01FEE6FC76" box="[266,363,946,967]" italics="true" pageId="45" pageNumber="936">angolense</emphasis>
</taxonomicName>
<treatmentCitation id="401DBC51FFCCBA0CFE5AFC00FDD6FC76" author="Ng PKL &amp; Guinot D &amp; Davie PJF" box="[391,603,946,968]" page="220" pageId="45" pageNumber="936" year="2008">
<bibRefCitation id="A52DE7B1FFCCBA0CFE5AFC00FDD6FC76" author="Ng" box="[391,603,946,968]" etAl="et al." firstAuthor="Ng" page="220" pageId="45" pageNumber="936" pagination="1 - 286" refId="ref73754" refString="Ng PKL, Guinot D &amp; Davie PJF (2008 a) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008" yearSuffix="a">Ng et al., 2008a: 220</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFCCBA0CFFACFC43FEDAFA38" pageId="45" pageNumber="936" type="materials_examined">
<paragraph id="C1039A40FFCCBA0CFFACFC43FEDAFA38" blockId="45.[113,753,1008,1417]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFFACFC43FEC9FBB9" bold="true" box="[113,324,1008,1032]" pageId="45" pageNumber="936">Material examined</emphasis>
.
<materialsCitation id="71D4901DFFCCBA0CFE8DFC42FD7CFBF9" ID-GBIF-Occurrence="3763237370" collectionCode="SMF-ZMG" collectorName="M. P. Kammerman" country="Angola" location="Benguella" municipality="Benguella" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635" specimenCount="1" specimenCount-male="1" typeStatus="neotype">
<collectingCountry id="B9ABDAD0FFCCBA0CFE8DFC42FE4DFBB8" box="[336,448,1009,1033]" name="Angola" pageId="45" pageNumber="936">ANGOLA</collectingCountry>
<typeStatus id="1E0724E2FFCCBA0CFE0AFC42FDBDFBB8" box="[471,560,1009,1033]" pageId="45" pageNumber="936" type="neotype">Neotype</typeStatus>
(here designated):
<specimenCount id="D7BA51C9FFCCBA0CFFACFBA3FF28FB99" box="[113,165,1040,1064]" pageId="45" pageNumber="936" type="male">male</specimenCount>
(39.3 ×
<quantity id="064437A5FFCCBA0CFEDCFBA2FEECFB98" box="[257,353,1041,1065]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="45" pageNumber="936" unit="mm" value="32.7">32.7 mm</quantity>
) (
<specimenCode id="911A323BFFCCBA0CFEAAFBA2FDA9FB99" box="[375,548,1040,1065]" collectionCode="SMF-ZMG" pageId="45" pageNumber="936">SMF-ZMG 635</specimenCode>
a),
<collectingMunicipality id="2167003AFFCCBA0CFD99FBA3FD3EFB99" box="[580,691,1040,1064]" country="Angola" county="Nimbe" municipality="West Nimbe" name="Benguella" pageId="45" pageNumber="936">Benguella</collectingMunicipality>
,
<locationDeviation id="D0B18362FFCCBA0CFD63FBA3FE63FBF9" pageId="45" pageNumber="936">river mouth of Catumbella near Benguella</locationDeviation>
, coll.
<collectorName id="6C49FF96FFCCBA0CFDF4FB82FD61FBF9" box="[553,748,1073,1096]" pageId="45" pageNumber="936">M.P. Kammerman</collectorName>
;
</materialsCitation>
<materialsCitation id="71D4901DFFCCBA0CFFACFBE2FED2FB38" ID-GBIF-Occurrence="3763237571" accessionNumber="SMF 25982" collectingDate="1984-01-07" collectingDateMax="2015-06-21" collectingDateMin="1984-01-07" collectionCode="RMNH, SMF" collectorName="M. P. Kammerman" country="Angola" county="Nimbe" location="Benguella" municipality="West Nimbe" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635, D 1200, ZRC 2015.0297, D 1937" specimenCount="9" specimenCount-female="5" specimenCount-juvenile="1" specimenCount-male="3" typeStatus="neotype">
1 ovigerous female (39.6 ×
<quantity id="064437A5FFCCBA0CFE74FBE2FD80FBD8" box="[425,525,1105,1129]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="45" pageNumber="936" unit="mm" value="33.8">33.8 mm</quantity>
) (
<specimenCode id="911A323BFFCCBA0CFDFBFBE2FD5BFBD9" box="[550,726,1104,1129]" collectionCode="SMF-ZMG" pageId="45" pageNumber="936">SMF-ZMG 635</specimenCode>
b), same data as neotype;
</materialsCitation>
<materialsCitation id="71D4901DFFCCBA0CFEBAFBC2FE5FFB18" ID-GBIF-Occurrence="3763237382" accessionNumber="SMF 25982" collectingDate="1984-01-07" collectingDateMax="2015-06-21" collectingDateMin="1984-01-07" collectionCode="RMNH, SMF" collectorName="M. P. Kammerman " country="Angola" county="Nimbe" location="Benguella" municipality="West Nimbe" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635, D 1200, ZRC 2015.0297, D 1937" specimenCount="9" specimenCount-female="5" specimenCount-juvenile="1" specimenCount-male="3" typeStatus="neotype">
<specimenCount id="D7BA51C9FFCCBA0CFEBAFBC2FE3FFB39" box="[359,434,1136,1161]" pageId="45" pageNumber="936" type="male">1 male</specimenCount>
,
<specimenCount id="D7BA51C9FFCCBA0CFE62FBC2FDAAFB39" box="[447,551,1136,1161]" pageId="45" pageNumber="936" type="female">2 females</specimenCount>
(
<collectionCode id="A7AD0285FFCCBA0CFDEBFBC2FD05FB38" box="[566,648,1137,1161]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="45" pageNumber="936" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFCCBA0CFD4DFBC2FD69FB38" box="[656,740,1137,1161]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="45" pageNumber="936" type="Museum">D 1200</specimenCode>
),
<collectorName id="6C49FF96FFCCBA0CFFACFB23FF36FB19" box="[113,187,1168,1192]" pageId="45" pageNumber="936">Lobito</collectorName>
, coll.
<collectorName id="6C49FF96FFCCBA0CFF24FB22FE01FB19" box="[249,396,1169,1192]" pageId="45" pageNumber="936">P. Kamerman</collectorName>
, 1899.
</materialsCitation>
<materialsCitation id="71D4901DFFCCBA0CFE0AFB22FEE4FAB8" ID-GBIF-Occurrence="3763237408" accessionNumber="SMF 25982" collectingDate="1984-01-07" collectingDateMax="2015-06-21" collectingDateMin="1984-01-07" collectionCode="RMNH, SMF" collectorName="C.H. Otto" country="Angola" county="Nimbe" location="Benguella" municipality="West Nimbe" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635, D 1200, ZRC 2015.0297, D 1937" specimenCount="9" specimenCount-female="5" specimenCount-juvenile="1" specimenCount-male="3" typeStatus="neotype">
<collectingCountry id="B9ABDAD0FFCCBA0CFE0AFB22FDFDFB18" box="[471,624,1169,1193]" name="Cameroon" pageId="45" pageNumber="936">CAMEROON</collectingCountry>
<specimenCount id="D7BA51C9FFCCBA0CFD55FB22FD7DFB19" box="[648,752,1168,1193]" pageId="45" pageNumber="936" type="juvenile">1 juvenile</specimenCount>
male (
<collectionCode id="A7AD0285FFCCBA0CFF69FB02FF66FB78" box="[180,235,1201,1225]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="45" pageNumber="936">SMF</collectionCode>
25982),
<collectingMunicipality id="2167003AFFCCBA0CFE90FB02FE59FB79" box="[333,468,1200,1224]" pageId="45" pageNumber="936">West Nimbe</collectingMunicipality>
(Limbe or Victoria),
<collectorName id="6C49FF96FFCCBA0CFD63FB03FD7DFB79" box="[702,752,1200,1224]" pageId="45" pageNumber="936">Mile</collectorName>
6, north of
<collectingCounty id="2862E2CCFFCCBA0CFF25FB63FECBFB59" box="[248,326,1232,1256]" pageId="45" pageNumber="936">Nimbe</collectingCounty>
, between pieces of basalt, coll.
<collectorName id="6C49FF96FFCCBA0CFD61FB62FF28FAB8" pageId="45" pageNumber="936">C.H. Otto</collectorName>
,
<date id="B502BC80FFCCBA0CFF69FB42FEE8FAB8" box="[180,357,1265,1289]" pageId="45" pageNumber="936" value="1984-01-07">
<collectingDate id="A5464568FFCCBA0CFF69FB42FEE8FAB8" box="[180,357,1265,1289]" pageId="45" pageNumber="936" value="1984-01-07">7 January 1984</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="71D4901DFFCCBA0CFEAEFB42FF3CFAD9" ID-GBIF-Occurrence="3763237505" accessionNumber="SMF 25982" collectingDate="1984-01-07" collectingDateMax="2015-06-21" collectingDateMin="1984-01-07" collectionCode="RMNH, SMF" collectorName="P.A. Mvogo-Ndongo" country="Angola" county="Nimbe" location="Benguella" municipality="West Nimbe" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635, D 1200, ZRC 2015.0297, D 1937" specimenCount="9" specimenCount-female="5" specimenCount-juvenile="1" specimenCount-male="3" typeStatus="neotype">
<specimenCount id="D7BA51C9FFCCBA0CFEAEFB42FE47FAB9" box="[371,458,1264,1289]" pageId="45" pageNumber="936" type="male">2 males</specimenCount>
(35.2 ×
<quantity id="064437A5FFCCBA0CFDEFFB43FD14FAB9" box="[562,665,1264,1288]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="45" pageNumber="936" unit="mm" value="28.6">28.6 mm</quantity>
, 21.2 ×
<quantity id="064437A5FFCCBA0CFFAFFAA3FF5AFA98" box="[114,215,1296,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="45" pageNumber="936" unit="mm" value="16.3">16.3 mm</quantity>
),
<specimenCount id="D7BA51C9FFCCBA0CFF33FAA2FEC3FA99" box="[238,334,1296,1321]" pageId="45" pageNumber="936" type="female">1 female</specimenCount>
(18.2 ×
<quantity id="064437A5FFCCBA0CFE6EFAA2FD9BFA98" box="[435,534,1297,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.43" pageId="45" pageNumber="936" unit="mm" value="14.3">14.3 mm</quantity>
) (
<specimenCode id="911A323BFFCCBA0CFDF2FAA2FD68FA99" box="[559,741,1296,1321]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="45" pageNumber="936">ZRC 2015.0297</specimenCode>
),
<collectorName id="6C49FF96FFCCBA0CFFACFA83FF61FAF9" box="[113,236,1328,1352]" pageId="45" pageNumber="936">Lokoundjié</collectorName>
mangrove, coll.
<collectorName id="6C49FF96FFCCBA0CFE7AFA82FD02FAF9" box="[423,655,1328,1353]" pageId="45" pageNumber="936">P.A. Mvogo-Ndongo</collectorName>
,
<date id="B502BC80FFCCBA0CFD41FA82FF21FAD9" pageId="45" pageNumber="936" value="2015-06-21">
<collectingDate id="A5464568FFCCBA0CFD41FA82FF21FAD9" pageId="45" pageNumber="936" value="2015-06-21">21 June 2015</collectingDate>
</date>
.
</materialsCitation>
<materialsCitation id="71D4901DFFCCBA0CFF61FAE2FEDAFA38" ID-GBIF-Occurrence="3763237400" accessionNumber="SMF 25982" collectingDate="1984-01-07" collectingDateMax="2015-06-21" collectingDateMin="1984-01-07" collectionCode="RMNH, SMF" collectorName="J. Demery" country="Angola" county="Nimbe" location="Benguella" municipality="West Nimbe" pageId="45" pageNumber="936" specimenCode="SMF-ZMG 635, D 1200, ZRC 2015.0297, D 1937" specimenCount="9" specimenCount-female="5" specimenCount-juvenile="1" specimenCount-male="3" typeStatus="neotype">
<collectingCountry id="B9ABDAD0FFCCBA0CFF61FAE2FEA4FAD8" box="[188,297,1361,1385]" name="Liberia" pageId="45" pageNumber="936">LIBERIA</collectingCountry>
<specimenCount id="D7BA51C9FFCCBA0CFE90FAE2FE30FAD9" box="[333,445,1360,1385]" pageId="45" pageNumber="936" type="female">2 females</specimenCount>
(
<collectionCode id="A7AD0285FFCCBA0CFE0DFAE2FDA9FAD8" box="[464,548,1361,1385]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="45" pageNumber="936" type="Museum">RMNH</collectionCode>
<specimenCode id="911A323BFFCCBA0CFDF2FAE2FD07FAD8" box="[559,650,1361,1385]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="45" pageNumber="936" type="Museum">D 1937</specimenCode>
), coll.
<collectorName id="6C49FF96FFCCBA0CFD03FAE2FF46FA38" pageId="45" pageNumber="936">J. Demery</collectorName>
, 18901897.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFCCBA0CFFACFA03FAFFFF44" pageId="45" pageNumber="936" type="diagnosis">
<paragraph id="C1039A40FFCCBA0CFFACFA03FE76F899" blockId="45.[113,753,1456,1832]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFFACFA03FF52FA79" bold="true" box="[113,223,1456,1480]" pageId="45" pageNumber="936">Diagnosis</emphasis>
. Carapace distinctly trapezoidal; frontal margin entire; lateral margins of carapace strongly converging to posterior carapace margin; cheliped merus with inner margin serrated but not expanded; adult male chela stout, outer surface gently convex, in large males, pollex may be expanded, shear-like, more flattened; male thoracic sternites 24 narrow in adults, suture between sternites 3 and 4 distinct; male sternopleonal cavity reaching three-quarters length of sternite 4 to just before anterior margin of sternite 2; male thoracic sternite 5 smooth, without depression on anterior part; male pleon elongate, narrow, telson linguiform; G1 relatively stout, chitinous part short.
</paragraph>
<paragraph id="C1039A40FFCCBA0CFFACF8E3FAFFFF44" blockId="45.[113,753,1872,2088]" lastBlockId="45.[812,1452,189,245]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFFACF8E3FF4FF8D9" bold="true" box="[113,194,1872,1896]" pageId="45" pageNumber="936">Colour</emphasis>
. The colour was described in detail by
<bibRefCitation id="A52DE7B1FFCCBA0CFD58F8E3FF62F838" author="Rossignol" firstAuthor="Rossignol" page="92" pageId="45" pageNumber="936" pagination="1 - 3" refId="ref75333" refString="Rossignol M (1957) Crustaces Decapodes marins de la region de Pointe-Noire. In: Collignon J, Rossignol M &amp; Roux C (eds.) Mollusques, Crustaces, Poissons Marins des cotes d'A. E. F. en collection au Centre d'Oceanographie de d'Institut d'Etudes Centrafricaines de Pointe-Noire. Paris. Pp. 71 - 136, figs. 1 - 20, pls. 1 - 3." type="book chapter" year="1957">Rossignol (1957: 92)</bibRefCitation>
: carapace blue-green with a yellow margin; ambulatory legs of the same colour as the carapace with a red dot to the joints, end dactylus yellow-orange dactyls; pincers colour of yellow leather with a red spot at the joint of the chela and fingers; eyes with light blue or gray-blue peduncle; cornea brown with a red dot; pleon dirty yellow with a blue median transverse line spanning somite 1; thoracic articles yellow and blue (translated from the French).
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFCCBA0CFCF0FEAEFA8BFBE4" pageId="45" pageNumber="936" type="discussion">
<paragraph id="C1039A40FFCCBA0CFCF0FEAEFB86FD24" blockId="45.[812,1452,285,661]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFCF0FEAEFC1AFE84" bold="true" box="[813,919,285,309]" pageId="45" pageNumber="936">Remarks</emphasis>
. In naming this species, Brito Capello did not indicate how many specimens he had. All he stated was that the species was from “mares de
<collectingCountry id="B9ABDAD0FFCCBA0CFB42FEEEFB7CFEC4" box="[1183,1265,349,373]" name="Angola" pageId="45" pageNumber="936">Angola</collectingCountry>
” (
<bibRefCitation id="A52DE7B1FFCCBA0CFAD1FEEEFC0EFE24" author="Brito Capello" firstAuthor="Brito Capello" page="4" pageId="45" pageNumber="936" refId="ref67965" refString="Brito Capello F de (1864) Descripcao de tres especies novas de crustaceos da Africa Occidental e observacoes acerca do Penoeus [sic] bocagei. Johnson. Especie nova dos mares de Portugal. Typographia da Academia, Lisboa. 11 + [1 unnumbered page of errata] pp., 1 unnumbered plate with 10 figures." type="book" year="1864">Brito Capello, 1864: 4</bibRefCitation>
). Unfortunately, all of Brito Capellos material in the Lisbon Museum were lost in a fire in 1978, and the types of this species are no longer extant. While the identity of this species is currently not in doubt, because it is here designated as the type species for
<taxonomicName id="06BCE1C3FFCCBA0CFB7FFE4DFAAEFDA4" authority="Schubart &amp; Ng, 2020" authorityName="Schubart &amp; Ng" authorityYear="2020" box="[1186,1315,510,533]" class="Malacostraca" family="Sesarmidae" genus="Trapezarma" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="F3C84652FFCCBA0CFB7FFE4DFAAEFDA4" box="[1186,1315,510,533]" italics="true" pageId="45" pageNumber="936">Trapezarma</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E8FBFB29FFCCBA0CFAEFFE4DFA2AFDA7" box="[1330,1447,510,534]" pageId="45" pageNumber="936" rank="genus">new genus</taxonomicNameLabel>
, it is important to ensure a stable taxonomy. As such, we designate a large male specimen in the Senckenberg Museum (SMF-ZMG 635a) as the
<typeStatus id="1E0724E2FFCCBA0CFB92FDEDFB28FDC7" box="[1103,1189,606,630]" pageId="45" pageNumber="936" type="neotype">neotype</typeStatus>
of
<taxonomicName id="06BCE1C3FFCCBA0CFB0DFDEDFB8AFD24" authority="Brito Capello, 1864" authorityName="Brito Capello" authorityYear="1864" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="45" pageNumber="936" phylum="Arthropoda" rank="species" species="angolensis">
<emphasis id="F3C84652FFCCBA0CFB0DFDEDFA21FDC4" box="[1232,1452,605,629]" italics="true" pageId="45" pageNumber="936">Sesarma angolensis</emphasis>
<bibRefCitation id="A52DE7B1FFCCBA0CFCF0FDCEFB8AFD24" author="Brito Capello" box="[813,1031,637,661]" firstAuthor="Brito Capello" pageId="45" pageNumber="936" refId="ref67965" refString="Brito Capello F de (1864) Descripcao de tres especies novas de crustaceos da Africa Occidental e observacoes acerca do Penoeus [sic] bocagei. Johnson. Especie nova dos mares de Portugal. Typographia da Academia, Lisboa. 11 + [1 unnumbered page of errata] pp., 1 unnumbered plate with 10 figures." type="book" year="1864">Brito Capello, 1864</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C1039A40FFCCBA0CFCF0FD0EFA8BFBE4" blockId="45.[813,1452,701,1110]" pageId="45" pageNumber="936">
There is considerable variation in the form of the male chela. The largest male from
<collectingCountry id="B9ABDAD0FFCCBA0CFBA3FD6EFB43FD44" box="[1150,1230,733,757]" name="Angola" pageId="45" pageNumber="936">Angola</collectingCountry>
(the
<typeStatus id="1E0724E2FFCCBA0CFAD0FD6DFAEEFD47" box="[1293,1379,734,758]" pageId="45" pageNumber="936" type="neotype">neotype</typeStatus>
male) has a more typical chela in which the pollex is relatively slender and the dactylus gently curved (
<figureCitation id="598786C5FFCCBA0CFB31FCAEFAD9FC84" box="[1260,1364,797,821]" captionStart="Fig" captionStartId="32.[136,171,1319,1341]" captionTargetBox="[136,1474,189,1303]" captionTargetId="figure-8@32.[136,1474,189,1303]" captionTargetPageId="32" captionText="Fig. 16. Chela of species of Trapezarma angolense. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; CF, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5351329" httpUri="https://zenodo.org/record/5351329/files/figure.png" pageId="45" pageNumber="936">Fig. 16A</figureCitation>
).
<location id="C463CC9BFFCCBA0CFAB7FCADFA0DFC84" LSID="urn:lsid:plazi:treatment:49152B56FFCCBA0CFF68FE6DFBE9FA84:C463CC9BFFCCBA0CFAB7FCADFA0DFC84" box="[1386,1408,798,821]" country="Angola" county="The" municipality="Other" name="In" pageId="45" pageNumber="936">In</location>
the smaller but clearly adult male from
<collectingCountry id="B9ABDAD0FFCCBA0CFB62FC8EFABCFCE4" box="[1215,1329,829,853]" name="Cameroon" pageId="45" pageNumber="936">Cameroon</collectingCountry>
, the pollex is strongly expanded, very broad, the outer surface more flattened, and the dactylus is strongly curved (
<figureCitation id="598786C5FFCCBA0CFA9DFCCEFCCEFC04" captionStart="Fig" captionStartId="32.[136,171,1319,1341]" captionTargetBox="[136,1474,189,1303]" captionTargetId="figure-8@32.[136,1474,189,1303]" captionTargetPageId="32" captionText="Fig. 16. Chela of species of Trapezarma angolense. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; CF, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5351329" httpUri="https://zenodo.org/record/5351329/files/figure.png" pageId="45" pageNumber="936">Fig. 16C, D</figureCitation>
).
<location id="C463CC9BFFCCBA0CFC84FC2EFC0FFC04" LSID="urn:lsid:plazi:treatment:49152B56FFCCBA0CFF68FE6DFBE9FA84:C463CC9BFFCCBA0CFC84FC2EFC0FFC04" box="[857,898,925,949]" country="Angola" county="The" municipality="Other" name="The" pageId="45" pageNumber="936">The</location>
specimen figured by
<bibRefCitation id="A52DE7B1FFCCBA0CFBAFFC2EFADCFC04" author="Brito Capello" box="[1138,1361,925,949]" firstAuthor="Brito Capello" pageId="45" pageNumber="936" refId="ref67965" refString="Brito Capello F de (1864) Descripcao de tres especies novas de crustaceos da Africa Occidental e observacoes acerca do Penoeus [sic] bocagei. Johnson. Especie nova dos mares de Portugal. Typographia da Academia, Lisboa. 11 + [1 unnumbered page of errata] pp., 1 unnumbered plate with 10 figures." type="book" year="1864">Brito Capello (1864</bibRefCitation>
: fig. 2c) shows a chela which is more intermediate in form.
<collectingMunicipality id="2167003AFFCCBA0CFAB3FC0EFA26FC64" box="[1390,1451,957,981]" pageId="45" pageNumber="936">Other</collectingMunicipality>
than this, the specimens from
<collectingCountry id="B9ABDAD0FFCCBA0CFBB9FC6EFB5EFC44" box="[1124,1235,989,1013]" name="Cameroon" pageId="45" pageNumber="936">Cameroon</collectingCountry>
and
<collectingCountry id="B9ABDAD0FFCCBA0CFADEFC6EFADCFC44" box="[1283,1361,989,1013]" name="Angola" pageId="45" pageNumber="936">Angola</collectingCountry>
are very similar and we have no reason to suspect they are different species.
<collectingCounty id="2862E2CCFFCCBA0CFC54FBAEFC3FFB84" box="[905,946,1053,1077]" pageId="45" pageNumber="936">The</collectingCounty>
characteristic male chela (
<figureCitation id="598786C5FFCCBA0CFB07FBAEFAD2FB84" box="[1242,1375,1053,1077]" captionStart="Fig" captionStartId="32.[136,171,1319,1341]" captionTargetBox="[136,1474,189,1303]" captionTargetId="figure-8@32.[136,1474,189,1303]" captionTargetPageId="32" captionText="Fig. 16. Chela of species of Trapezarma angolense. A, B, neotype male (39.3 × 32.7 mm) (SMF-ZMG 635), Benguella, Angola; CF, male (35.2 × 28.6 mm) (ZRC 2015.0297), Cameroon. A, C, outer view; D, inner view; B, E, F, dorso-lateral view." figureDoi="http://doi.org/10.5281/zenodo.5351329" httpUri="https://zenodo.org/record/5351329/files/figure.png" pageId="45" pageNumber="936">Fig. 16C, D</figureCitation>
) noted above may be a feature of dominant males.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFCCBA0CFCF0FBCEFACFFB64" pageId="45" pageNumber="936" type="biology_ecology">
<paragraph id="C1039A40FFCCBA0CFCF0FBCEFACFFB64" blockId="45.[813,1453,1149,1237]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFCF0FBCEFC0EFB24" bold="true" box="[813,899,1149,1173]" pageId="45" pageNumber="936">Biology</emphasis>
. The best account of the biology of this species is given in
<bibRefCitation id="A52DE7B1FFCCBA0CFC48FB2EFB34FB04" author="Rathbun" box="[917,1209,1181,1205]" firstAuthor="Rathbun" page="451 - 452" pageId="45" pageNumber="936" pagination="379 - 484" refId="ref75176" refString="Rathbun MJ (1921) The Brachyuran crabs collected by the American Museum Congo Expedition, 1909 - 1915. In: Scientific results of the American Museum Congo Expedition. General Invertebrate Zoology, No. 6. Bulletin of the American Museum of Natural History, 43: 379 - 484, figs. 1 - 22, pls. 15 - 44." type="journal article" year="1921">Rathbun (1921: 451452)</bibRefCitation>
, who observes that it prefers the more freshwater sections of the river.
</paragraph>
</subSubSection>
<subSubSection id="89A6C9CBFFCCBA0CFCF0FB4EFBE9FA84" pageId="45" pageNumber="936" type="distribution">
<paragraph id="C1039A40FFCCBA0CFCF0FB4EFBE9FA84" blockId="45.[813,1450,1277,1333]" pageId="45" pageNumber="936">
<emphasis id="F3C84652FFCCBA0CFCF0FB4EFC4EFAA4" bold="true" box="[813,963,1277,1301]" pageId="45" pageNumber="936">Distribution</emphasis>
.
<collectingCountry id="B9ABDAD0FFCCBA0CFC0AFB4EFBE2FAA4" box="[983,1135,1277,1301]" name="Sierra Leone" pageId="45" pageNumber="936">Sierra Leone</collectingCountry>
to
<collectingCountry id="B9ABDAD0FFCCBA0CFB41FB4EFB7DFAA4" box="[1180,1264,1277,1301]" name="Angola" pageId="45" pageNumber="936">Angola</collectingCountry>
(
<bibRefCitation id="A52DE7B1FFCCBA0CFAD9FB4EFA28FAA4" author="Monod" box="[1284,1445,1277,1301]" firstAuthor="Monod" pageId="45" pageNumber="936" pagination="1 - 674" refId="ref72973" refString="Monod T (1956) Hippidea et Brachyura ouest-africains. Memoires de I'Institut Francais d'Afrique Noire, 45: 1 - 674, 884 figs." type="journal article" year="1956">Monod, 1956</bibRefCitation>
;
<bibRefCitation id="A52DE7B1FFCCBA0CFCF3FAAEFBD5FA84" author="Manning &amp; Holthuis" box="[814,1112,1309,1333]" firstAuthor="Manning" pageId="45" pageNumber="936" pagination="1 - 379" refId="ref72504" refString="Manning RB &amp; Holthuis LB (1981) West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, 306: i - xii, 1 - 379, figs. 1 - 88, table 1." type="journal article" year="1981">Manning &amp; Holthuis, 1981</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>