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<document ID-DOI="10.11646/phytotaxa.594.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="7864827" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="jonas" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1682329282318" checkinUser="plazi" docAuthor="Meerts, Pierre" docDate="2023" docId="49146779FF9F691BFF07FB33FA64FC42" docLanguage="en" docName="phytotaxa.594.1.1.pdf" docOrigin="Phytotaxa 594 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.594.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.5:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="5" docTitle="Clerodendrum Linnaeus 1753" docType="key" docVersion="2" lastPageNumber="5" masterDocId="B52D1F01FF9E691FFF8EFFE3FFCEFFE3" masterDocTitle="The genus Clerodendrum (Lamiaceae) in the flora of Central Africa (D. R. Congo, Rwanda, Burundi)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="2" updateTime="1682452634697" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>The genus Clerodendrum (Lamiaceae) in the flora of Central Africa (D. R. Congo, Rwanda, Burundi)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Meerts, Pierre</mods:namePart>
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<mods:affiliation>Meise Botanic Garden, Nieuwelaan 38, 1860 Meise, Belgium &amp; Herbarium et bibliothèque de botanique africaine, Université Libre de Bruxelles, Avenue F. D. Roosevelt 50, CP 244, 1050 Brussels, Belgium &amp; Fédération Wallonie-Bruxelles. Service général de l’Enseignement supérieur et de la Recherche scientifique, Brussels, Belgium pierre. meerts @ plantentuinmeise. be; https: // orcid. org / 0000 - 0003 - 4215 - 027 X</mods:affiliation>
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<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-24</mods:number>
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<mods:number>594</mods:number>
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<subSubSection box="[137,642,1232,1259]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph blockId="1.[136,707,1232,1294]" box="[137,642,1232,1259]" pageId="1" pageNumber="2">
<taxonomicName ID-CoL="8VWBC" ID-ENA="13432" authority="Linnaeus (1753: 637)" authorityName="Linnaeus" authorityPageNumber="637" authorityYear="1753" box="[137,642,1232,1259]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="genus">
<emphasis bold="true" box="[137,394,1232,1258]" pageId="1" pageNumber="2">CLERODENDRUM</emphasis>
<bibRefCitation author="Linnaeus, C." box="[401,642,1232,1259]" pageId="1" pageNumber="2" pagination="561 - 1200" refId="ref39967" refString="Linnaeus, C. (1753) Species plantarum, vol. 2. Laurentii Salvii, Holmiae, pp. 561 - 1200." type="book chapter" year="1753">Linnaeus (1753: 637)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection box="[136,707,1272,1294]" pageId="1" pageNumber="2" type="type_taxon">
<paragraph blockId="1.[136,707,1232,1294]" box="[136,707,1272,1294]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[136,186,1272,1293]" pageId="1" pageNumber="2">
<typeStatus box="[136,186,1272,1293]" pageId="1" pageNumber="2">Type</typeStatus>
</emphasis>
:— 
<taxonomicName authority="Linnaeus (1753: 637)" authorityName="Linnaeus" authorityPageNumber="637" authorityYear="1753" box="[217,703,1272,1294]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="infortunatum">
<emphasis box="[217,487,1272,1293]" italics="true" pageId="1" pageNumber="2">Clerodendrum infortunatum</emphasis>
<bibRefCitation author="Linnaeus, C." box="[493,703,1272,1294]" pageId="1" pageNumber="2" pagination="561 - 1200" refId="ref39967" refString="Linnaeus, C. (1753) Species plantarum, vol. 2. Laurentii Salvii, Holmiae, pp. 561 - 1200." type="book chapter" year="1753">Linnaeus (1753: 637)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="description">
<paragraph blockId="1.[136,1452,1340,2050]" pageId="1" pageNumber="2">Trees, shrubs, lianas, or more rarely perennial herbs; stem quadrangular. Leaves opposite or 3–4-whorled, usually petiolate, simple; lamina entire, undulate, repand, dentate or crenate; petiole articulated, sometimes the lower part persisting as a woody spine. Inflorescence of axillary cymes, generally grouped in racemose or corymbose thyrses; bracts often foliaceous, bracteoles small, mostly not exceeding calyx. Flower: calyx usually regular, campanulate to tubular or cup-shaped, ± deeply 5-lobed or truncate, sometimes 5-angled, usually persistent and accrescent in fruit; corolla symmetrical in bud or, if asymmetrical, usually expanding abruptly on upper side due to resupination, actinomorphic; tube ± long, narrowly cylindrical or funnel-shaped; limb of 5 equal or subequal spreading or reflexed lobes, larger than the others. Stamens 4, inserted on the corolla tube, usually long exserted; anthers versatile. Carpels 2, each incompletely divided into 2 loculi; locules 1-ovulate. Style terminal, stigma exserted, 2-fid, the lobes equal, short. Fruit drupaceous, obovoid or globose, generally 4-lobed, often appearing 1-lobed by abortion; endocarp separated into four 1-seeded pyrenes.</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="reference_group">
<paragraph blockId="1.[136,1452,1340,2050]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[189,319,1736,1762]" pageId="1" pageNumber="2">References</emphasis>
:— 
<bibRefCitation author="Baker, E. G." box="[352,506,1736,1762]" pageId="1" pageNumber="2" pagination="292 - 314" refId="ref37331" refString="Baker, E. G. (1900) Clerodendrum. In: Thiselton-Dyer, W. T. (Ed.) Flora of Tropical Africa 5. Lovell Reeve &amp; Co, London, pp. 292 - 314." type="book chapter" year="1900">Baker (1900)</bibRefCitation>
; 
<bibRefCitation author="Briquet, J." box="[517,685,1736,1763]" pageId="1" pageNumber="2" pagination="173 - 177" refId="ref37511" refString="Briquet, J. (1895) Viticoideae. Clerodendreae. In: Engler, A. &amp; Prantl, K. (Eds.) Die naturlichen Pflanzenfamilien IV (3 a). W. Engelmann, Leipzig, pp. 173 - 177." type="book chapter" year="1895">Briquet (1895)</bibRefCitation>
; 
<bibRefCitation author="Fernandes, R." box="[695,885,1736,1762]" pageId="1" pageNumber="2" pagination="5 - 109" refId="ref38766" refString="Fernandes, R. (1998) Notes sur les Verbenaceae. XI. Remarques sur quelques taxa africains du genre Clerodendrum L. Memorias da Sociedade Broteriana 30: 5 - 109." type="journal article" year="1998">Fernandes (1998</bibRefCitation>
, 
<bibRefCitation author="Fernandes, R." box="[895,956,1736,1763]" pageId="1" pageNumber="2" pagination="6 - 61" refId="ref38800" refString="Fernandes, R. (2005) Verbenaceae. In: Pope, G. V. &amp; Martins, E. S. (Eds.) Flora Zambesiaca. 8 (7). Royal Botanic Gardens, Kew, pp 6 - 61." type="book chapter" year="2005">2005</bibRefCitation>
); 
<bibRefCitation author="Harley, R. M. &amp; Atkins, S. &amp; Budantsev, A. &amp; Cantino, P. D. &amp; Conn, B. &amp; Grayer, R. J. &amp; Harley, M. M. &amp; Kok, R. de &amp; Krestovskaja, T. &amp; Morales, A. &amp; Paton, A. J. &amp; Ryding, O. &amp; Upson, T." box="[974,1195,1736,1762]" pageId="1" pageNumber="2" pagination="167 - 275" refId="ref39106" refString="Harley, R. M., Atkins, S., Budantsev, A., Cantino, P. D., Conn, B., Grayer, R. J., Harley, M. M., Kok, R. de, Krestovskaja, T., Morales, A., Paton, A. J., Ryding, O. &amp; Upson, T. (2004) Labiatae. In: Kadereit, J. W. (Ed.) The families and genera of vascular plants (Lamiales), VII. Springer Verlag, Berlin, pp. 167 - 275. https: // doi. org / 10.1007 / 978 - 3 - 642 - 18617 - 2 _ 11" type="book chapter" year="2004">
Harley 
<emphasis box="[1056,1112,1736,1762]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
(2004)
</bibRefCitation>
; 
<bibRefCitation author="Huber, H." box="[1206,1360,1736,1762]" pageId="1" pageNumber="2" pagination="439 - 445" refId="ref39459" refString="Huber, H. (1963) Clerodendrum. In: Hepper, F. N. (Ed.) Flora of West Tropical Africa, 2 nd Ed., vol. 2. Crown Agents for Oversea Governments and Administrations, London, pp. 439 - 445." type="book chapter" year="1963">Huber (1963)</bibRefCitation>
; 
<bibRefCitation author="Lebrun, J. - P. &amp; Stork, A." pageId="1" pageNumber="2" refId="ref39728" refString="Lebrun, J. - P. &amp; Stork, A. (1997) Enumeration des plantes a fleurs d'Afrique. IV. Conservatoire et Jardin botaniques de la ville de Geneve, Geneve, 712 pp." type="book" year="1997">Lebrun &amp; Stork (1997)</bibRefCitation>
; 
<bibRefCitation author="Li, B. &amp; Cantino, P. D. &amp; Olmstead, R. G. &amp; Bramley, G. L. C. &amp; Xiang, C. - L. &amp; Ma, Z. - H. &amp; Tan, Y. - H. &amp; Zhang, D. - X." box="[322,494,1772,1798]" pageId="1" pageNumber="2" pagination="34343" refId="ref39853" refString="Li, B., Cantino, P. D., Olmstead, R. G., Bramley, G. L. C., Xiang, C. - L., Ma, Z. - H., Tan, Y. - H. &amp; Zhang, D. - X. (2016) A large-scale chloroplast phylogeny of the Lamiaceae sheds new light on its subfamilial classification. Scientific Reports 6: 34343. https: // doi. org / 10.1038 / srep 34343" type="journal article" year="2016">
Li 
<emphasis box="[354,410,1772,1798]" italics="true" pageId="1" pageNumber="2">et al.</emphasis>
(2016)
</bibRefCitation>
; 
<bibRefCitation author="Moldenke, H. N." box="[506,716,1772,1799]" pageId="1" pageNumber="2" pagination="334 - 365" refId="ref40388" refString="Moldenke, H. N. (1985 a) Notes on the genus Clerodendrum (Verbenaceae). IV. Phytologia 57 (5): 334 - 365." type="journal article" year="1985">Moldenke (1985a)</bibRefCitation>
; 
<bibRefCitation author="Pollard, B." box="[728,894,1772,1798]" pageId="1" pageNumber="2" pagination="17 - 40" refId="ref41070" refString="Pollard, B. (2022) Clerodendrum. In: Sosef, M. S. M., Florence, J., Bourobou Bourobou, H. P., Bissiengou, P. (eds.) Flore du Gabon, vol. 58. Labiatae. Margraf Publishers, Weikersheim, pp. 17 - 40." type="book chapter" year="2022">Pollard (2022)</bibRefCitation>
; 
<bibRefCitation author="Robyns, W." box="[905,1075,1772,1798]" pageId="1" pageNumber="2" refId="ref41247" refString="Robyns, W. (1947) Flore des Spermatophytes du Parc national Albert, vol. II. Sympetales. Institut des Parcs Nationaux du Congo belge, Bruxelles, 627 pp." type="book" year="1947">Robyns (1947)</bibRefCitation>
; 
<bibRefCitation author="Thomas, B." box="[1087,1261,1772,1798]" pageId="1" pageNumber="2" pagination="1 - 106" refId="ref41701" refString="Thomas, B. (1936) Die Gattung Clerodendrum inAfrika. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 68: 1 - 106." type="journal article" year="1936">Thomas (1936)</bibRefCitation>
; 
<bibRefCitation author="Malaise, P." box="[1273,1447,1772,1799]" pageId="1" pageNumber="2" pagination="267 - 291" refId="ref40061" refString="Malaise, P. (1985) Verbenaceae. In: Troupin, G. (Ed.) Flore du Rwanda, Spermatophytes. 3. Musee royal de l'Afrique centrale, Tervueren, pp. 267 - 291." type="book chapter" year="1985">Malaise (1985)</bibRefCitation>
; 
<bibRefCitation author="Verdcourt, B." box="[136,331,1808,1834]" pageId="1" pageNumber="2" pagination="84 - 144" refId="ref41825" refString="Verdcourt, B. (1992) Clerodendrum. In: Polhill, R. M. (Ed.) Flora of tropical East Africa. Verbenaceae. Balkema. Rotterdam, pp. 84 - 144." type="book chapter" year="1992">Verdcourt (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="2" type="discussion">
<paragraph blockId="1.[136,1452,1340,2050]" pageId="1" pageNumber="2">
<emphasis bold="true" box="[189,245,1844,1870]" pageId="1" pageNumber="2">Note</emphasis>
:—Two species have been collected in cultivation in only one occasion and are not dealt with in the following account: 
<taxonomicName authority="Thunberg (1784: 256)" authorityName="Thunberg" authorityPageNumber="256" authorityYear="1784" box="[241,821,1880,1907]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="trichotomum">
<emphasis box="[241,557,1880,1906]" italics="true" pageId="1" pageNumber="2">Clerodendrum trichotomum</emphasis>
<bibRefCitation author="Thunberg, C. P." box="[567,821,1880,1907]" pageId="1" pageNumber="2" refId="ref41747" refString="Thunberg, C. P. (1784) Flora Japonica. Lipsiae, 419 pp." type="book" year="1784">Thunberg (1784: 256)</bibRefCitation>
</taxonomicName>
(D.R. 
<collectingCountry box="[904,1007,1880,1906]" name="Democratic Republic of the Congo" pageId="1" pageNumber="2">CONGO</collectingCountry>
. Eala, 
<date box="[1088,1240,1880,1907]" pageId="1" pageNumber="2" value="1932-04-05">5 April 1932</date>
, 
<emphasis italics="true" pageId="1" pageNumber="2">Corbisier-Baland 1361</emphasis>
(BR!)); 
<taxonomicName authority="(Osbeck 1757: 215) Mabberley (1989: 707)" authorityName="Mabberley" authorityPageNumber="707" authorityYear="1989" baseAuthorityName="Osbeck" baseAuthorityPageNumber="215" baseAuthorityYear="1757" box="[289,1055,1916,1943]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="2" phylum="Tracheophyta" rank="species" species="chinense">
<emphasis box="[289,557,1916,1942]" italics="true" pageId="1" pageNumber="2">Clerodendrum chinense</emphasis>
(
<bibRefCitation author="Osbeck, P." box="[571,778,1916,1943]" pageId="1" pageNumber="2" refId="ref40962" refString="Osbeck, P. (1757) Dagbok ofwer en ostindsk resa. Stockholm, 376 pp." type="book" year="1757">Osbeck 1757: 215</bibRefCitation>
) 
<bibRefCitation author="Mabberley, D. J." box="[794,1055,1916,1942]" pageId="1" pageNumber="2" refId="ref40026" refString="Mabberley, D. J. (1989) The plant-book. A portable dictionary of the higher plants. [Reprint with corrections]. Cambridge, Cambridge University Press, 707 pp." type="book" year="1989">Mabberley (1989: 707)</bibRefCitation>
</taxonomicName>
(D.R. 
<collectingCountry box="[1131,1234,1916,1942]" name="Democratic Republic of the Congo" pageId="1" pageNumber="2">CONGO</collectingCountry>
. Kinshasa, Gombe, 
<date box="[136,316,1952,1978]" pageId="1" pageNumber="2" value="2001-08-27">27 August 2001</date>
, 
<emphasis box="[327,485,1952,1978]" italics="true" pageId="1" pageNumber="2">Pauwels 8540</emphasis>
(BR!)).
</paragraph>
<paragraph blockId="1.[136,1452,1340,2050]" pageId="1" pageNumber="2">
The species present in our territory belong in the following sections of the genus, as defined by 
<bibRefCitation author="Thomas, B." box="[1272,1447,1988,2014]" pageId="1" pageNumber="2" pagination="1 - 106" refId="ref41701" refString="Thomas, B. (1936) Die Gattung Clerodendrum inAfrika. Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 68: 1 - 106." type="journal article" year="1936">Thomas (1936)</bibRefCitation>
, validated and amended by 
<bibRefCitation author="Verdcourt, B." box="[436,631,2024,2050]" pageId="1" pageNumber="2" pagination="84 - 144" refId="ref41825" refString="Verdcourt, B. (1992) Clerodendrum. In: Polhill, R. M. (Ed.) Flora of tropical East Africa. Verbenaceae. Balkema. Rotterdam, pp. 84 - 144." type="book chapter" year="1992">Verdcourt (1992)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="taxon_list">
<paragraph blockId="2.[136,793,158,219]" box="[136,371,158,184]" pageId="2" pageNumber="3">Sect. Capitata Verdc.</paragraph>
<paragraph blockId="2.[136,793,158,219]" box="[136,793,198,219]" pageId="2" pageNumber="3">
<taxonomicName box="[136,249,198,219]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buchneri">
<emphasis box="[136,152,198,219]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,249,198,219]" italics="true" pageId="2" pageNumber="3">buchneri</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Schumacher" baseAuthorityName="Willdenow" baseAuthorityYear="1800" box="[261,385,198,219]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="capitatum">
<emphasis box="[261,277,198,219]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[289,385,198,219]" italics="true" pageId="2" pageNumber="3">capitatum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="De Wildeman" authorityYear="1909" box="[397,528,198,219]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="excavatum">
<emphasis box="[397,413,198,219]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[425,528,198,219]" italics="true" pageId="2" pageNumber="3">excavatum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="S. Moore" authorityYear="1919" box="[540,681,198,219]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="frutectorum">
<emphasis box="[540,556,198,219]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[568,681,198,219]" italics="true" pageId="2" pageNumber="3">frutectorum</emphasis>
</taxonomicName>
, 
<taxonomicName box="[693,786,198,219]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="poggei">
<emphasis box="[693,709,198,219]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[721,786,198,219]" italics="true" pageId="2" pageNumber="3">poggei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,719,266,327]" box="[136,523,266,293]" pageId="2" pageNumber="3">Sect. Eurycalyx Thomas ex Verdc.</paragraph>
<paragraph blockId="2.[136,719,266,327]" box="[136,719,306,327]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Gurke. Thomas" authorityYear="1936" box="[136,268,306,327]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="bipindense">
<emphasis box="[136,152,306,327]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,268,306,327]" italics="true" pageId="2" pageNumber="3">bipindense</emphasis>
</taxonomicName>
, 
<taxonomicName box="[280,437,306,327]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="melanocrater">
<emphasis box="[280,296,306,327]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[308,437,306,327]" italics="true" pageId="2" pageNumber="3">melanocrater</emphasis>
</taxonomicName>
, 
<taxonomicName box="[449,596,306,327]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="thyrsoideum">
<emphasis box="[449,465,306,327]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[477,596,306,327]" italics="true" pageId="2" pageNumber="3">thyrsoideum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Palisot de Beauvois" authorityYear="1805" box="[608,713,306,327]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="volubile">
<emphasis box="[608,624,306,327]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[636,713,306,327]" italics="true" pageId="2" pageNumber="3">volubile</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,1062,374,435]" box="[136,542,374,401]" pageId="2" pageNumber="3">
Sect. 
<taxonomicName authority="Thomas ex Verdc." box="[198,542,374,401]" class="Magnoliopsida" family="Euphorbiaceae" genus="Neoboutonia" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="macrocalyx">Macrocalyx Thomas ex Verdc.</taxonomicName>
</paragraph>
<paragraph blockId="2.[136,1062,374,435]" box="[136,1062,414,435]" pageId="2" pageNumber="3">
<taxonomicName box="[136,231,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="baumii">
<emphasis box="[136,152,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,231,414,435]" italics="true" pageId="2" pageNumber="3">baumii</emphasis>
</taxonomicName>
, 
<taxonomicName box="[243,358,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buettneri">
<emphasis box="[243,259,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[271,358,414,435]" italics="true" pageId="2" pageNumber="3">buettneri</emphasis>
</taxonomicName>
, 
<taxonomicName box="[369,465,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fuscum">
<emphasis box="[369,385,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[397,465,414,435]" italics="true" pageId="2" pageNumber="3">fuscum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="De Wildeman" authorityYear="1909" box="[477,592,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="pynaertii">
<emphasis box="[477,493,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[505,592,414,435]" italics="true" pageId="2" pageNumber="3">pynaertii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Oliver" authorityYear="1875" box="[604,763,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="rotundifolium">
<emphasis box="[604,620,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[632,763,414,435]" italics="true" pageId="2" pageNumber="3">rotundifolium</emphasis>
</taxonomicName>
<taxonomicName box="[769,908,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="thomsoniae">
<emphasis box="[769,785,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[797,908,414,435]" italics="true" pageId="2" pageNumber="3">thomsoniae</emphasis>
</taxonomicName>
, 
<taxonomicName box="[920,1056,414,435]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis box="[920,936,414,435]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[948,1056,414,435]" italics="true" pageId="2" pageNumber="3">welwitschii</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,809,482,543]" box="[136,537,482,509]" pageId="2" pageNumber="3">Sect. Microcalyx Thomas ex Verdc.</paragraph>
<paragraph blockId="2.[136,809,482,543]" box="[136,809,522,543]" pageId="2" pageNumber="3">
<taxonomicName box="[136,277,522,543]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="formicarum">
<emphasis box="[136,152,522,543]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,277,522,543]" italics="true" pageId="2" pageNumber="3">formicarum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Good" authorityYear="1930" box="[289,492,522,543]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="inaequipetiolatum">
<emphasis box="[289,305,522,543]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[317,492,522,543]" italics="true" pageId="2" pageNumber="3">inaequipetiolatum</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Oliver" authorityYear="1887" box="[504,628,522,543]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="johnstonii">
<emphasis box="[504,520,522,543]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[532,628,522,543]" italics="true" pageId="2" pageNumber="3">johnstonii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Baker" authorityYear="1895" box="[640,803,522,543]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="polycephalum">
<emphasis box="[640,656,522,543]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[668,803,522,543]" italics="true" pageId="2" pageNumber="3">polycephalum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,566,590,651]" box="[136,517,590,617]" pageId="2" pageNumber="3">Sect. Oxycalyx Thomas ex Verdc.</paragraph>
<paragraph blockId="2.[136,566,590,651]" box="[136,566,630,651]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Walpers" authorityYear="1845" baseAuthorityName="Roxburgh" baseAuthorityYear="1832" box="[136,272,630,651]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buchananii">
<emphasis box="[136,152,630,651]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,272,630,651]" italics="true" pageId="2" pageNumber="3">buchananii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Don" authorityYear="1824" box="[284,407,630,651]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="splendens">
<emphasis box="[284,300,630,651]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[312,407,630,651]" italics="true" pageId="2" pageNumber="3">splendens</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Poiret in de Lamarck" authorityYear="1804" box="[419,560,630,651]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="umbellatum">
<emphasis box="[419,435,630,651]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[447,560,630,651]" italics="true" pageId="2" pageNumber="3">umbellatum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,890,698,759]" box="[136,562,698,725]" pageId="2" pageNumber="3">Sect. Siphonocalyx Thomas ex Verdc.</paragraph>
<paragraph blockId="2.[136,890,698,759]" box="[136,890,738,759]" pageId="2" pageNumber="3">
<taxonomicName authorityName="Moldenke" authorityYear="1953" box="[136,240,738,759]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="dewittei">
<emphasis box="[136,152,738,759]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[164,240,738,759]" italics="true" pageId="2" pageNumber="3">dewittei</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Thomas" authorityYear="1936" box="[252,403,738,759]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fasciculatum">
<emphasis box="[252,268,738,759]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[280,403,738,759]" italics="true" pageId="2" pageNumber="3">fasciculatum</emphasis>
</taxonomicName>
, 
<taxonomicName box="[415,580,738,759]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="schweinfurthii">
<emphasis box="[415,431,738,759]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[443,580,738,759]" italics="true" pageId="2" pageNumber="3">schweinfurthii</emphasis>
</taxonomicName>
, 
<taxonomicName authorityName="Henriq. Henriques" authorityYear="1892" box="[592,707,738,759]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="silvanum">
<emphasis box="[592,608,738,759]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[620,707,738,759]" italics="true" pageId="2" pageNumber="3">silvanum</emphasis>
</taxonomicName>
, 
<taxonomicName box="[719,884,738,759]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="tanganyikense">
<emphasis box="[719,735,738,759]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[747,884,738,759]" italics="true" pageId="2" pageNumber="3">tanganyikense</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="2.[136,1451,806,869]" pageId="2" pageNumber="3">
All the species in this account belong in the African clade except 
<taxonomicName authorityName="Walpers" authorityYear="1845" baseAuthorityName="Roxburgh" baseAuthorityYear="1832" box="[868,1027,806,833]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buchananii">
<emphasis box="[868,887,807,833]" italics="true" pageId="2" pageNumber="3">C</emphasis>
. 
<emphasis box="[901,1027,806,832]" italics="true" pageId="2" pageNumber="3">buchananii</emphasis>
</taxonomicName>
, an introduced species which belongs in the Asian clade as defined by 
<bibRefCitation author="Steane, D. A. &amp; Scotland, R. W. &amp; Mabberley, D. J." box="[499,721,842,868]" pageId="2" pageNumber="3" pagination="229 - 243" refId="ref41574" refString="Steane, D. A., Scotland, R. W. &amp; Mabberley, D. J. (1997) Phylogenetic Relationships of Clerodendrum s. l. (Lamiaceae) Inferred from Chloroplast DNA. Systematic Botany 22 (2): 229 - 243. https: // doi. org / 10.2307 / 2419455" type="journal article" year="1997">
Steane 
<emphasis box="[581,637,842,868]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
(1997)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[136,1159,914,941]" pageId="2" pageNumber="3" type="multiple">
<paragraph blockId="2.[136,1159,914,941]" box="[136,1159,914,941]" pageId="2" pageNumber="3">
<heading bold="true" box="[136,1159,914,941]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis bold="true" box="[136,801,914,941]" pageId="2" pageNumber="3">
Key to the species of 
<taxonomicName ID-CoL="8VWBC" ID-ENA="13432" authorityName="Linnaeus" authorityYear="1753" box="[386,554,914,941]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="genus">
<emphasis bold="true" box="[386,554,914,941]" italics="true" pageId="2" pageNumber="3">Clerodendrum</emphasis>
</taxonomicName>
in Central Africa (D
</emphasis>
. 
<emphasis bold="true" box="[808,828,914,940]" pageId="2" pageNumber="3">R</emphasis>
. 
<emphasis bold="true" box="[842,920,914,940]" pageId="2" pageNumber="3">
<collectingCountry box="[842,920,914,940]" name="Democratic Republic of the Congo" pageId="2" pageNumber="3">Congo</collectingCountry>
</emphasis>
, 
<emphasis bold="true" box="[934,1034,914,940]" pageId="2" pageNumber="3">
<collectingCountry box="[934,1034,914,940]" name="Rwanda" pageId="2" pageNumber="3">Rwanda</collectingCountry>
</emphasis>
, 
<emphasis bold="true" box="[1048,1159,914,940]" pageId="2" pageNumber="3">
<collectingCountry box="[1048,1151,914,940]" name="Burundi" pageId="2" pageNumber="3">Burundi</collectingCountry>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="key">
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,983,1033]" box="[136,1451,983,1005]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,983,1005]" pageId="2" pageNumber="3">1. Corolla wholly orange to red.................................................................................................................................................. Group 1</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,983,1033]" box="[136,1451,1011,1033]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1011,1033]" pageId="2" pageNumber="3">- Corolla white, cream, yellowish, greenish, occasionally red-spotted in throat or with one red lobe.................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1452,1069,1149]" pageId="2" pageNumber="3">
<keyLead pageId="2" pageNumber="3">
2. Inflorescence congested, capitate, with axis hidden by flowers; calyx membranous, petaloid, whitish, yellowish, purplish or mauve, 
<quantity box="[211,314,1098,1120]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.6" metricValueMin="1.0" pageId="2" pageNumber="3" unit="mm" value="18.0" valueMax="26.0" valueMin="10.0">10–26 mm</quantity>
long....................................................................................................................................................................... Group 2
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1452,1069,1149]" box="[136,1451,1127,1149]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1127,1149]" pageId="2" pageNumber="3">- Inflorescence looser, branched, or relatively compact but, in this case, calyx greenish, not petaloid, 3–10(–12) mm long .............3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1184,1235]" box="[136,1451,1184,1206]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1184,1206]" pageId="2" pageNumber="3">3. Inflorescence congested, hemispheric to globose, with branches inconspicuous at anthesis ................................................ Group 3</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1184,1235]" box="[136,1451,1213,1235]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1213,1235]" pageId="2" pageNumber="3">- Inflorescence of loose cymes, or elongate thyrses, with branches conspicuous at anthesis ..............................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1270,1321]" box="[136,1451,1270,1293]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1270,1293]" pageId="2" pageNumber="3">
4. Calyx lobes at anthesis 
<quantity box="[437,534,1270,1292]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="2" pageNumber="3" unit="mm" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 mm</quantity>
long, &lt;50(–60)% of total calyx length............................................................................ Group 4
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1270,1321]" box="[136,1451,1299,1321]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1299,1321]" pageId="2" pageNumber="3">
- Calyx lobes at anthesis (3–) 
<quantity box="[476,568,1299,1321]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="2" pageNumber="3" unit="mm" value="15.0" valueMax="25.0" valueMin="5.0">5–25 mm</quantity>
long,&gt; 50% of total calyx length............................................................................... Group 5
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection box="[136,513,1357,1379]" pageId="2" pageNumber="3" type="multiple">
<paragraph blockId="2.[136,513,1357,1379]" box="[136,513,1357,1379]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[136,222,1357,1379]" pageId="2" pageNumber="3">Group 1</emphasis>
: Corolla wholly orange or red
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="3" type="key">
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1414,1465]" box="[136,1451,1414,1437]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1414,1437]" pageId="2" pageNumber="3">
1. Calyx whitish to yellowish, pentagonal, gibbose at base, 
<quantity box="[757,860,1414,1436]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="2" pageNumber="3" unit="mm" value="18.5" valueMax="25.0" valueMin="12.0">12–25 mm</quantity>
long........................................................................................2
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1414,1465]" box="[136,1451,1443,1465]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1443,1465]" pageId="2" pageNumber="3">
- Calyx greenish or reddish, neither pentagonal nor gibbose, 
<quantity box="[779,870,1444,1465]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 mm</quantity>
long ......................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1501,1552]" box="[136,1451,1501,1523]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1501,1523]" pageId="2" pageNumber="3">
2. Stem and inflorescence branches densely pubescent, with brown hairs; leaves conspicuously pubescent ..... 
<taxonomicName box="[1238,1451,1501,1522]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="fuscum">
<emphasis box="[1238,1451,1501,1522]" italics="true" pageId="2" pageNumber="3">Clerodendrum fuscum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1501,1552]" box="[136,1451,1530,1552]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1530,1552]" pageId="2" pageNumber="3">
- Stem and inflorescence branches subglabrous or puberulous; leaves glabrous ........................................ 
<taxonomicName box="[1195,1451,1530,1551]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="thomsoniae">
<emphasis box="[1195,1451,1530,1551]" italics="true" pageId="2" pageNumber="3">Clerodendrum thomsoniae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1452,1587,1667]" box="[136,1451,1587,1609]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1587,1609]" pageId="2" pageNumber="3">
3. Petiole 
<quantity box="[283,369,1587,1609]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="2.5" metricValueMin="0.2" pageId="2" pageNumber="3" unit="cm" value="13.5" valueMax="25.0" valueMin="2.0">2–25 cm</quantity>
; lamina deeply cordate at base; lower surface of lamina tomentose .............................. 
<taxonomicName authorityName="Walpers" authorityYear="1845" baseAuthorityName="Roxburgh" baseAuthorityYear="1832" box="[1198,1451,1587,1608]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buchananii">
<emphasis box="[1198,1451,1587,1608]" italics="true" pageId="2" pageNumber="3">Clerodendrum buchananii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1452,1587,1667]" pageId="2" pageNumber="3">
<keyLead pageId="2" pageNumber="3">
- Petiole 0.3–2(–4) cm; lamina rounded to slightly cordate at base; lower surface of lamina glabrous or shortly pubescent on veins (rarely on whole surface).............................................................................................................................. 
<taxonomicName authorityName="Don" authorityYear="1824" box="[1211,1451,1645,1666]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="splendens">
<emphasis box="[1211,1451,1645,1666]" italics="true" pageId="2" pageNumber="3">Clerodendrum splendens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection box="[136,809,1702,1725]" pageId="2" pageNumber="3" type="multiple">
<paragraph blockId="2.[136,809,1702,1725]" box="[136,809,1702,1725]" pageId="2" pageNumber="3">
<emphasis bold="true" box="[136,222,1702,1724]" pageId="2" pageNumber="3">Group 2</emphasis>
: Inflorescence capitate, unbranched; calyx petaloid&gt; 
<quantity box="[742,809,1703,1724]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="mm" value="10.0">10 mm</quantity>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="key">
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1760,1811]" box="[136,1451,1760,1782]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1760,1782]" pageId="2" pageNumber="3">
1. Inflorescences on old defoliated shoots often near ground level; leaves glabrous..................................... 
<taxonomicName authorityName="De Wildeman" authorityYear="1909" box="[1203,1451,1760,1782]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="excavatum">
<emphasis box="[1203,1451,1760,1782]" italics="true" pageId="2" pageNumber="3">Clerodendrum excavatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1760,1811]" box="[136,1451,1789,1811]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1789,1811]" pageId="2" pageNumber="3">- Inflorescences on leafy twigs; leaves pubescent at least on lower surface ........................................................................................2</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1846,1926]" pageId="2" pageNumber="3">
<keyLead pageId="2" pageNumber="3">
2. Suffrutex (0.10–) 
<quantity box="[378,468,1846,1868]" metricMagnitude="-1" metricUnit="m" metricValue="5.75" metricValueMax="10.0" metricValueMin="1.5" pageId="2" pageNumber="3" unit="m" value="0.575" valueMax="1.0" valueMin="0.15">0.15–1 m</quantity>
in height, with shoots renewed every year; corolla tube 
<quantity box="[1007,1099,1846,1868]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="7.5" metricValueMin="5.0" pageId="2" pageNumber="3" unit="cm" value="6.25" valueMax="7.5" valueMin="5.0">5–7.5 cm</quantity>
; leaf lamina &lt;11 × 
<quantity box="[1290,1355,1846,1868]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="3" unit="cm" value="5.5">5.5 cm</quantity>
............... ....................................................................................................................................................................... 
<taxonomicName box="[1220,1450,1875,1896]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="buchneri">
<emphasis box="[1220,1450,1875,1896]" italics="true" pageId="2" pageNumber="3">Clerodendrum buchneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1846,1926]" box="[136,1451,1904,1926]" pageId="2" pageNumber="3">
<keyLead box="[136,1451,1904,1926]" pageId="2" pageNumber="3">
- Shrub or liana 
<quantity box="[353,416,1904,1926]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="2" pageNumber="3" unit="m" value="3.5" valueMax="6.0" valueMin="1.0">1–6 m</quantity>
, with persisting shoots; corolla tube 
<quantity box="[767,853,1904,1926]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.9" metricValueMin="0.6" pageId="2" pageNumber="3" unit="cm" value="12.5" valueMax="19.0" valueMin="6.0">6–19 cm</quantity>
; lamina of variable size, often&gt; 11 × 
<quantity box="[1217,1282,1904,1926]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="2" pageNumber="3" unit="cm" value="5.5">5.5 cm</quantity>
..........................3
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="2" pageNumber="3">
<paragraph blockId="2.[136,1451,1962,2070]" pageId="2" pageNumber="3">
<keyLead pageId="2" pageNumber="3">
3. Shrub with unbranched or sparingly branched shoots; petiole base not persistent; leaves mostly grouped at shoot apex; lamina 9–48 × 
<quantity box="[230,316,1990,2012]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="4.5" metricValueMin="0.6" pageId="2" pageNumber="3" unit="cm" value="25.5" valueMax="45.0" valueMin="6.0">6–45 cm</quantity>
, mostly cordate at base; petiole mostly&gt; 
<quantity box="[714,761,1991,2013]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="3" unit="cm" value="6.0">6 cm</quantity>
................................................................................................................4
</keyLead>
</paragraph>
<paragraph blockId="2.[136,1451,1962,2070]" pageId="2" pageNumber="3">
<keyLead pageId="2" pageNumber="3">
- Shrub or liana with branched shoots; petiole base persistent, spiny; leaves not grouped near shoot tip; lamina &lt;17 × 
<quantity box="[1310,1370,2020,2041]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="3" unit="cm" value="11.0">11 cm</quantity>
, cuneate or rounded at base; petiole &lt;
<quantity box="[491,538,2048,2070]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="3" unit="cm" value="6.0">6 cm</quantity>
............................................................................................................... 
<taxonomicName authorityName="Schumacher" baseAuthorityName="Willdenow" baseAuthorityYear="1800" box="[1210,1451,2048,2070]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="capitatum">
<emphasis box="[1210,1451,2048,2070]" italics="true" pageId="2" pageNumber="3">Clerodendrum capitatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1452,159,296]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
4. Inflorescence ovoid to subcylindric, 
<quantity box="[562,645,159,181]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="3" pageNumber="4" unit="cm" value="12.5" valueMax="20.0" valueMin="5.0">5–20 cm</quantity>
long (corolla excluded), markedly longer than wide; peduncle hollow, up to 
<quantity box="[1384,1451,159,181]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="10.0">10 mm</quantity>
thick; stem and petiole papillate, lacking long patent hairs; calyx purplish-violaceous ................................... 
<taxonomicName box="[1240,1450,188,209]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="poggei">
<emphasis box="[1240,1450,188,209]" italics="true" pageId="3" pageNumber="4">Clerodendrum poggei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,159,296]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
- Inflorescence hemispheric to shortly ovoid, 
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in diam. (corolla excluded); peduncle not exceeding 
<quantity box="[1230,1286,216,238]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
in diam., mostly solid; stem and petiole with mixed indumentum of papilliform and long patent hairs; calyx mostly greenish, occasionally with a purplish tinge ............................................................................................................................................. 
<taxonomicName authorityName="S. Moore" authorityYear="1919" box="[1192,1450,274,295]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="frutectorum">
<emphasis box="[1192,1450,274,295]" italics="true" pageId="3" pageNumber="4">Clerodendrum frutectorum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection box="[136,973,332,354]" pageId="3" pageNumber="4" type="multiple">
<paragraph blockId="3.[136,973,332,354]" box="[136,973,332,354]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[136,222,332,354]" pageId="3" pageNumber="4">Group 3</emphasis>
: Inflorescence congested, hemispheric; calyx green at anthesis, &lt;
<quantity box="[857,924,332,353]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="10.0">10 mm</quantity>
long
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="key">
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,389,440]" box="[136,1451,389,411]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,389,411]" pageId="3" pageNumber="4">
1. Calyx 
<quantity box="[274,353,389,411]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="4" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
long; peduncle papillate, pubescent, or tomentose ....................................................................................................2
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,389,440]" box="[136,1451,418,440]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,418,440]" pageId="3" pageNumber="4">
- Calyx 
<quantity box="[274,353,418,440]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="3" pageNumber="4" unit="mm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 mm</quantity>
long; peduncle glabrous to puberulous......................................................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1452,476,584]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
2. Corolla tube 
<quantity box="[335,438,476,498]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="3" pageNumber="4" unit="mm" value="30.0" valueMax="35.0" valueMin="25.0">25–35 mm</quantity>
long; petiole and young twigs glabrous or with patent stiff hairs; peduncle 
<quantity box="[1156,1241,476,497]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="2.2" metricValueMin="0.4" pageId="3" pageNumber="4" unit="cm" value="13.0" valueMax="22.0" valueMin="4.0">4–22 cm</quantity>
, papillate, puberulous or shortly pubescent............................................................................................................................... 
<taxonomicName box="[1168,1450,505,526]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="schweinfurthii">
<emphasis box="[1168,1450,505,526]" italics="true" pageId="3" pageNumber="4">Clerodendrum schweinfurthii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,476,584]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
- Corolla tube 
<quantity box="[341,447,533,555]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
long; petiole and young twigs beige to fulvous tomentose; peduncle 
<quantity box="[1139,1234,533,555]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="3" pageNumber="4" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.5–4 cm</quantity>
, densely pubescent to tomentose................................................................................................................................................. 
<taxonomicName authorityName="Thomas" authorityYear="1936" box="[1183,1451,562,583]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="fasciculatum">
<emphasis box="[1183,1451,562,583]" italics="true" pageId="3" pageNumber="4">Clerodendrum fasciculatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,620,728]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
3. Calyx narrowly tubular, 
<quantity box="[451,532,620,641]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="3" pageNumber="4" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
wide near throat, lobes not conspicuously veined; corolla tube 7--13(--18) mm long; stamens exserted 5--10(--15) mm; petiole glabrous; liana up to 
<quantity box="[735,784,649,670]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="m" value="10.0">10 m</quantity>
................................................. 
<taxonomicName authorityName="Fernandes" authorityYear="1998" baseAuthorityName="Exell" baseAuthorityYear="1930" box="[1080,1450,649,671]" class="Magnoliopsida" family="Lamiaceae" form="caulanthum" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="form" species="silvanum">
<emphasis box="[1080,1312,649,670]" italics="true" pageId="3" pageNumber="4">Clerodendrum silvanum</emphasis>
f. 
<emphasis box="[1337,1450,649,670]" italics="true" pageId="3" pageNumber="4">caulanthum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,620,728]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
- Calyx funnel-shaped, ca. 
<quantity box="[459,516,677,699]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
wide near throat, lobes conspicuously 3-veined; corolla tube 
<quantity box="[1118,1222,677,699]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="3" pageNumber="4" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15–22 mm</quantity>
long; stamens exserted 
<quantity box="[211,317,706,728]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="4" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
; petiole pubescent; suffrutex or small liana &lt;
<quantity box="[755,792,706,728]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="m" value="2.0">2 m</quantity>
in height ......................................................... 
<taxonomicName authorityName="Moldenke" authorityYear="1953" box="[1230,1451,706,727]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="dewittei">
<emphasis box="[1230,1451,706,727]" italics="true" pageId="3" pageNumber="4">Clerodendrum dewittei</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection box="[136,1200,764,786]" pageId="3" pageNumber="4" type="multiple">
<paragraph blockId="3.[136,1200,764,786]" box="[136,1200,764,786]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[136,222,764,786]" pageId="3" pageNumber="4">Group 4</emphasis>
: Inflorescence conspicuously branched; calyx lobes 
<quantity box="[728,825,764,786]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="3.0" metricValueMin="0.5" pageId="3" pageNumber="4" unit="mm" value="1.75" valueMax="3.0" valueMin="0.5">0.5–3 mm</quantity>
long, &lt;50(–60)% of total calyx length
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="4" type="key">
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,821,872]" box="[136,1451,821,843]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,821,843]" pageId="3" pageNumber="4">1. Calyx shortly campanulate to cupular, with depth equal to or smaller than diameter; calyx lobes spreading...................................2</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,821,872]" box="[136,1451,850,872]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,850,872]" pageId="3" pageNumber="4">- Calyx tubular to narrowly funnel-shaped, with depth markedly longer than diameter; calyx lobes erect.........................................5</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,908,959]" box="[136,1451,908,930]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,908,930]" pageId="3" pageNumber="4">
2. Inflorescence on old defoliated shoots, often near the ground................................................................... 
<taxonomicName authorityName="Gurke. Thomas" authorityYear="1936" box="[1202,1451,908,929]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="bipindense">
<emphasis box="[1202,1451,908,929]" italics="true" pageId="3" pageNumber="4">Clerodendrum bipindense</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,908,959]" box="[136,1451,937,959]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,937,959]" pageId="3" pageNumber="4">- Inflorescence on leafy shoots .............................................................................................................................................................3</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,994,1074]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
3. Whole plant turning black in herbarium; corolla dark-spotted after anthesis (in the field); lamina base rounded to obtusely cuneate ................................................................................................................................................... 
<taxonomicName box="[1176,1450,1023,1044]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="melanocrater">
<emphasis box="[1176,1450,1023,1044]" italics="true" pageId="3" pageNumber="4">Clerodendrum melanocrater</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,994,1074]" box="[136,1451,1052,1074]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,1052,1074]" pageId="3" pageNumber="4">- Whole plant green to brownish in herbarium; corolla not dark-spotted after anthesis; lamina base cuneate ....................................4</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1452,1109,1218]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
4. Flower bud and corolla tube greyish-tomentellous; inflorescence main axis pubescent; inflorescence paniculiform; rachis 
<quantity box="[1363,1448,1109,1131]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" metricValueMax="4.0" metricValueMin="0.6" pageId="3" pageNumber="4" unit="cm" value="23.0" valueMax="40.0" valueMin="6.0">6–40 cm</quantity>
, with internodes 
<quantity box="[365,456,1138,1160]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="6.5" metricValueMin="0.5" pageId="3" pageNumber="4" unit="mm" value="35.0" valueMax="65.0" valueMin="5.0">5–65 mm</quantity>
......................................................................................................................... 
<taxonomicName box="[1187,1451,1138,1159]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="thyrsoideum">
<emphasis box="[1187,1451,1138,1159]" italics="true" pageId="3" pageNumber="4">Clerodendrum thyrsoideum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,1109,1218]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
- Flower bud and corolla tube subglabrous; main axis of inflorescence subglabrous or puberulent only in the upper part; inflorescence corymbiform; rachis 1.5–6(–12) cm, with internodes &lt;
<quantity box="[741,808,1196,1217]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="mm" value="10.0">10 mm</quantity>
...................................................................... 
<taxonomicName authorityName="Palisot de Beauvois" authorityYear="1805" box="[1228,1450,1196,1217]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="volubile">
<emphasis box="[1228,1450,1196,1217]" italics="true" pageId="3" pageNumber="4">Clerodendrum volubile</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1452,1253,1391]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
5. Leaves 3–4-whorled; lamina conspicuously 3-veined from base, glabrous; inflorescence umbelliform, rachis 1–2(–7) cm, inflorescence branches almost horizontally spreading, glabrous to puberulous; stem often conspicuously striate or sulcate ............ ................................................................................................................................................................... 
<taxonomicName box="[1192,1450,1311,1332]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="formicarum">
<emphasis box="[1192,1450,1311,1332]" italics="true" pageId="3" pageNumber="4">Clerodendrum formicarum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,1253,1391]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">- Leaves opposite, or, more rarely, 3-whorled, but, in this case, pubescent; lamina 1-nerved at base; inflorescence thyrsiform or, rarely, umbellate; stem cylindric or slightly striate ............................................................................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,1425,1477]" box="[136,1451,1425,1448]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,1425,1448]" pageId="3" pageNumber="4">
6. Calyx (3–) 
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long, lobed ca. ¼ in length; corolla tube 
<quantity box="[806,897,1426,1448]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.0" metricValueMin="0.7" pageId="3" pageNumber="4" unit="mm" value="13.5" valueMax="20.0" valueMin="7.0">7–20 mm</quantity>
..........................................................................................7
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,1425,1477]" box="[136,1451,1455,1477]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,1455,1477]" pageId="3" pageNumber="4">
- Calyx 
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, lobed 40–50% in length; corolla tube 4–10(–11) mm ...........................................................................................8
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</paragraph>
</keyStep>
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<paragraph blockId="3.[136,1452,1513,1592]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
7. Stem (at least near apex) and leaves (at least the petiole), more or less pubescent; upper surface of lamina dull, with reticulum not prominulent............................................................................................................................................ 
<taxonomicName box="[1168,1450,1541,1562]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="tanganyikense">
<emphasis box="[1168,1450,1541,1562]" italics="true" pageId="3" pageNumber="4">Clerodendrum tanganyikense</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,1513,1592]" box="[136,1451,1570,1592]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,1570,1592]" pageId="3" pageNumber="4">
- Stem and leaves wholly glabrous; upper surface of lamina shiny, with reticulum prominulent................... 
<taxonomicName authorityName="Henriq. Henriques" authorityYear="1892" box="[1219,1451,1570,1591]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="silvanum">
<emphasis box="[1219,1451,1570,1591]" italics="true" pageId="3" pageNumber="4">Clerodendrum silvanum</emphasis>
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</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1452,1628,1736]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
8. Calyx glabrous to puberulent; first-order inflorescence axes branched only near tip, spreading at a very open angle, forming an umbel; leaves often 3-whorled .............................................................................................................. 
<taxonomicName authorityName="Baker" authorityYear="1895" box="[1171,1451,1657,1678]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="polycephalum">
<emphasis box="[1171,1451,1657,1678]" italics="true" pageId="3" pageNumber="4">Clerodendrum polycephalum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="3.[136,1452,1628,1736]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">- Calyx pubescent, either hirsute or with dense indumentum of curly hairs; first order inflorescence axes more ramified, forming a corymbose to hemispheric inflorescence; leaves opposite.................................................................................................................9</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,1772,1909]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
9. Young twigs, inflorescence branches and calyx covered with short curly rusty or fulvous hairs; inflorescence 4–14 × 
<quantity box="[1360,1447,1772,1793]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.9" metricValueMin="0.4" pageId="3" pageNumber="4" unit="cm" value="11.5" valueMax="19.0" valueMin="4.0">4–19 cm</quantity>
; petiole 
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; lower surface of lamina most often tomentose, with curly hairs, sometimes only on veins ...................................... ..................................................................................................................................................................... 
<taxonomicName authorityName="Oliver" authorityYear="1887" box="[1210,1451,1829,1850]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="johnstonii">
<emphasis box="[1210,1451,1829,1850]" italics="true" pageId="3" pageNumber="4">Clerodendrum johnstonii</emphasis>
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</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,1772,1909]" pageId="3" pageNumber="4">
<keyLead pageId="3" pageNumber="4">
- Young twigs, inflorescence branches and calyx with straight or undulate, not curly, pale fulvous hairs; inflorescence 3–4 × 
<quantity metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" pageId="3" pageNumber="4" unit="cm" value="5.75" valueMax="8.0" valueMin="3.5">3.5–8 cm</quantity>
; petiole &lt;
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; lamina hispid on lower surface of veins...................................................... 
<taxonomicName authorityName="Good" authorityYear="1930" box="[1131,1451,1887,1908]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="inaequipetiolatum">
<emphasis box="[1131,1451,1887,1908]" italics="true" pageId="3" pageNumber="4">Clerodendrum inaequipetiolatum</emphasis>
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</keyLead>
</paragraph>
</keyStep>
</subSubSection>
<subSubSection box="[136,1085,1944,1967]" pageId="3" pageNumber="4" type="multiple">
<paragraph blockId="3.[136,1085,1944,1967]" box="[136,1085,1944,1967]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[136,222,1944,1966]" pageId="3" pageNumber="4">Group 5</emphasis>
: Inflorescence branched; calyx divided over ca. 50% of its length, with lobes&gt; (3–) 
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</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="key">
<key lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<keyStep pageId="3" pageNumber="4">
<paragraph blockId="3.[136,1451,2002,2053]" box="[136,1451,2002,2024]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,2002,2024]" pageId="3" pageNumber="4">1. Liana or shrub; shoots perennial, branched,&gt; 1 m long ....................................................................................................................2</keyLead>
</paragraph>
<paragraph blockId="3.[136,1451,2002,2053]" box="[136,1451,2031,2053]" pageId="3" pageNumber="4">
<keyLead box="[136,1451,2031,2053]" pageId="3" pageNumber="4">- Suffrutex; shoots annual, unbranched, up to 1.2 m long ....................................................................................................................7</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1452,159,239]" box="[136,1451,159,181]" pageId="4" pageNumber="5">
<keyLead box="[136,1451,159,181]" pageId="4" pageNumber="5">
2. Inflorescence terminal, rather dense, with branches &lt;2.5(–4) cm, not subtended by leafy bracts........... 
<taxonomicName box="[1198,1451,159,180]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="welwitschii">
<emphasis box="[1198,1451,159,180]" italics="true" pageId="4" pageNumber="5">Clerodendrum welwitschii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,159,239]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
- Inflorescence in axillary cymes subtended by foliaceous bracts, branches&gt; 
<quantity box="[935,985,188,209]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="cm" value="3.0">3 cm</quantity>
, occasionally the uppermost ones grouped into a loose corymb or panicle......................................................................................................................................................................3
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1451,274,325]" box="[136,1451,274,296]" pageId="4" pageNumber="5">
<keyLead box="[136,1451,274,296]" pageId="4" pageNumber="5">
3. Shoot and inflorescence with long patent, brownish-violaceous articulated hairs 
<quantity box="[981,1060,274,296]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="4" pageNumber="5" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long ................. 
<taxonomicName box="[1219,1451,274,295]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="buettneri">
<emphasis box="[1219,1451,274,295]" italics="true" pageId="4" pageNumber="5">Clerodendrum buettneri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1451,274,325]" box="[136,1451,303,325]" pageId="4" pageNumber="5">
<keyLead box="[136,1451,303,325]" pageId="4" pageNumber="5">- Shoot and inflorescence glabrous or shortly pubescent, with hairs not remarkable ..........................................................................4</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1451,361,469]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
4. Corolla tube 
<quantity box="[342,427,361,383]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="4" pageNumber="5" unit="cm" value="8.5" valueMax="11.0" valueMin="6.0">6–11 cm</quantity>
long; calyx densely velvety-pubescent; leaf lamina tomentose on lower surface; pedicel 
<quantity box="[1312,1393,361,382]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1--4 cm</quantity>
; stem hollow..................................................................................................................................................... 
<taxonomicName authorityName="Oliver" authorityYear="1875" box="[1175,1451,389,410]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="rotundifolium">
<emphasis box="[1175,1451,389,410]" italics="true" pageId="4" pageNumber="5">Clerodendrum rotundifolium</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1451,361,469]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
- Corolla tube 
<quantity box="[339,415,418,440]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="cm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 cm</quantity>
; calyx glabrous to shortly pubescent; leaf lamina glabrous or slightly pubescent on lower surface; pedicel 0.4–1.8(–3.0) cm; stem solid ..............................................................................................................................................................5
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1452,504,613]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
5. Calyx cream to yellow; stem and inflorescence branches densely pubescent, with brown hairs; inflorescence branches subtended by foliaceous, subsessile, cordate, clasping bracts ........................................................................................... 
<taxonomicName box="[1238,1451,533,554]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="fuscum">
<emphasis box="[1238,1451,533,554]" italics="true" pageId="4" pageNumber="5">Clerodendrum fuscum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,504,613]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">- Calyx whitish to whitish-green, occasionally purple-tinged; stem and inflorescence axes puberulous to shortly pubescent; inflorescence branches subtended by ovate, petiolate, not clasping, foliaceous bracts .....................................................................6</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1452,648,786]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
6. Inflorescence comprised of small 3–5(10)-flowered axillary cymes on a peduncle 
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, in the 3–10 upper nodes of the stem, forming a very loose narrow thyrse (9–32 × 
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); calyx gibbose at base, pentagonal, accrescent to 
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during anthesis...... ....................................................................................................................................................................... 
<taxonomicName authorityName="De Wildeman" authorityYear="1909" box="[1219,1451,706,728]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="pynaertii">
<emphasis box="[1219,1451,706,728]" italics="true" pageId="4" pageNumber="5">Clerodendrum pynaertii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,648,786]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
- Inflorescence comprised of multiflorous corymbiform cymes, peduncle 
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, in the 2–3 upper nodes; calyx funnel-shaped or campanulate, neither gibbose nor pentagonal, &lt;
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long................................................................... 
<taxonomicName authorityName="Poiret in de Lamarck" authorityYear="1804" box="[1192,1450,764,785]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="umbellatum">
<emphasis box="[1192,1450,764,785]" italics="true" pageId="4" pageNumber="5">Clerodendrum umbellatum</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="4" pageNumber="5">
<paragraph blockId="4.[136,1452,821,929]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
7. Calyx gibbose, truncate at base, 
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long at anthesis, accrescent up to 
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long; inflorescence branches 
<quantity box="[1372,1447,822,843]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
; corolla tube ca. 
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long............................................................................................................................... 
<taxonomicName box="[1239,1451,850,871]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="baumii">
<emphasis box="[1239,1451,850,871]" italics="true" pageId="4" pageNumber="5">Clerodendrum baumii</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="4.[136,1452,821,929]" pageId="4" pageNumber="5">
<keyLead pageId="4" pageNumber="5">
- Calyx funnel-shaped, neither gibbose nor truncate, 
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long; inflorescence branches 
<quantity box="[1081,1155,879,901]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="9.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="cm" value="5.0" valueMax="9.0" valueMin="1.0">1–9 cm</quantity>
; corolla tube ca. 
<quantity box="[1318,1383,879,901]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="5" unit="cm" value="1.5">1.5 cm</quantity>
long... ................................................................................................................................................................... 
<taxonomicName authorityName="Poiret in de Lamarck" authorityYear="1804" box="[1192,1450,908,929]" class="Magnoliopsida" family="Lamiaceae" genus="Clerodendrum" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="5" phylum="Tracheophyta" rank="species" species="umbellatum">
<emphasis box="[1192,1450,908,929]" italics="true" pageId="4" pageNumber="5">Clerodendrum umbellatum</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
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