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<document id="4F823C3B24C7537A9F3426BB79ECA37D" ID-CLB-Dataset="8612" ID-DOI="10.11646/zootaxa.4851.2.10" ID-GBIF-Dataset="af1d4768-dd24-484d-8ad6-a7fe1d300e12" ID-ISSN="1175-5326" ID-Zenodo-Dep="4476997" ID-ZooBank="43C5F7B3-1C03-4738-84CC-FF17B4FE0889" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609432312793" checkinUser="plazi" docAuthor="Ghaderi, Reza, Hesar, Abas Mokaram, Karegar, Akbar &amp; Pereira, Tiago José" docDate="2020" docId="924E87E396464250FF4C3FCD1829F70F" docLanguage="en" docName="zootaxa.4851.2.10.pdf" docOrigin="Zootaxa 4851 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diphtherophora de Man 1880" docType="treatment" docVersion="8" lastPageNumber="382" masterDocId="6E77FF9B964E425BFFDB3A071B28F64E" masterDocTitle="Molecular phylogeny of Diphtherophora de Man, 1880 (Nematoda: Diphtherophoridae), with description of a new species" masterLastPageNumber="385" masterPageNumber="371" pageNumber="379" updateTime="1698850660153" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D0094E6ECC72A0749D7452A871393D97">Molecular phylogeny of Diphtherophora de Man, 1880 (Nematoda: Diphtherophoridae), with description of a new species</mods:title>
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<mods:namePart id="D672D467FE9C8BFB39A281A728647471">Ghaderi, Reza</mods:namePart>
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<mods:namePart id="DBB2FDFB3FA9A2D1E8A94C95809C54BC">Hesar, Abas Mokaram</mods:namePart>
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<mods:affiliation id="0CEE68E627CC8AC5ED2F495B21BDF219">abasmokaram @ gmail. com; https: // orcid. org / 0000 - 0002 - 0244 - 1366</mods:affiliation>
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<mods:namePart id="05D3863890BE5434F700AF6E872E10B8">Karegar, Akbar</mods:namePart>
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<mods:affiliation id="C16749B3E2FE9B8F958CEA00CC422607">karegar @ shirazu. ac. ir; https: // orcid. org / 0000 - 0002 - 2041 - 816 X</mods:affiliation>
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<mods:namePart id="AE8F712E83CD913FE9E80EBCD4CFE2B0">Pereira, Tiago José</mods:namePart>
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<mods:affiliation id="F8884400A37452391A817A766B4EC645">Department of Nematology, University of California, Riverside, 900 University Avenue, Riverside, CA 92521, USA &amp; Present address: Department of Marine Sciences, University of Georgia, 325 Sanford Drive, Athens, GA 30602, USA. Tiago. Pereira @ uga. edu; https: // orcid. org / 0000 - 0002 - 6424 - 2848</mods:affiliation>
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<paragraph id="1A5836F596464253FF4C3FCD1969F3AB" blockId="8.[151,577,1482,1509]" box="[151,577,1482,1509]" pageId="8" pageNumber="379">
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<taxonomicName id="DDE74D7696464253FF4C3FCD1A73F3AB" authorityName="de Man" authorityYear="1880" box="[151,347,1482,1509]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796464253FF4C3FCD1A73F3AB" bold="true" box="[151,347,1482,1509]" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
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species phylogeny
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<paragraph id="1A5836F596464253FF4C3C141F86F14B" blockId="8.[151,1437,1555,2014]" pageId="8" pageNumber="379">
In total, 26 DNA sequences, specifically covering the D2–D3 domains of the 28S rRNA gene, were used in our molecular phylogenetic analyses: 18 
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<emphasis id="2893EAE796464253FDC33C3019E5F01F" box="[536,717,1591,1617]" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
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, two 
<taxonomicName id="DDE74D7696464253FCD43C3018E4F01F" authorityName="de Man" authorityYear="1880" box="[783,972,1591,1617]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="8" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796464253FCD43C3018E4F01F" box="[783,972,1591,1617]" italics="true" pageId="8" pageNumber="379">Tylolaimophorus</emphasis>
</taxonomicName>
, one representative of 
<taxonomicName id="DDE74D7696464253FB0A3C301EB0F01F" authorityName="Micoletzky" authorityYear="1922" box="[1233,1432,1591,1617]" class="Adenophorea" family="Prismatolaimidae" kingdom="Animalia" order="Enoplida" pageId="8" pageNumber="379" phylum="Nematoda" rank="family">Prismatolaimidae</taxonomicName>
, six representatives of 
<taxonomicName id="DDE74D7696464253FE4B3C5C1919F03B" authorityName="Thorne" authorityYear="1935" box="[400,561,1627,1653]" class="Adenophorea" family="Trichodoridae" kingdom="Animalia" order="Triplonchida" pageId="8" pageNumber="379" phylum="Nematoda" rank="family">Trichodoridae</taxonomicName>
, two representatives of 
<taxonomicName id="DDE74D7696464253FCE53C5C189DF03B" authorityName="de Man" authorityYear="1876" box="[830,949,1627,1653]" class="Adenophorea" family="Tripylidae" kingdom="Animalia" order="Enoplida" pageId="8" pageNumber="379" phylum="Nematoda" rank="family">Tripylidae</taxonomicName>
, and three representatives of 
<taxonomicName id="DDE74D7696464253FAD93C5C1BF9F0D7" authorityName="Zhao" authorityYear="2011" class="Adenophorea" family="Trischistomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="8" pageNumber="379" phylum="Nematoda" rank="family">Trischistomatidae</taxonomicName>
. Our final alignment was 785 bp long and contained 326 conserved sites, 455 variable sites, 415 parsimonyinformative sites, and 40 singletons. Phylogenetic analyses based on both inference methods (i.e., ML and BI) were highly congruent with respect to the monophyly of the genus 
<taxonomicName id="DDE74D7696464253FC803CC01F38F0AF" authorityName="de Man" authorityYear="1880" box="[859,1040,1735,1761]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796464253FC803CC01F38F0AF" box="[859,1040,1735,1761]" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
</taxonomicName>
, the relationships among 
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<emphasis id="2893EAE796464253FAE23CC01BDAF14B" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
</taxonomicName>
species and outgroups, and the phylogenetic placement of 
<taxonomicName id="DDE74D7696464253FC5C3CEB1F2EF14B" authority="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira, 2020" authorityName="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira" authorityYear="2020" box="[903,1030,1771,1797]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="379" phylum="Tracheophyta" rank="species" species="eldarica" status="sp. nov.">
<emphasis id="2893EAE796464253FC5C3CEB1F2EF14B" box="[903,1030,1771,1797]" italics="true" pageId="8" pageNumber="379">D. eldarica</emphasis>
</taxonomicName>
<emphasis id="2893EAE796464253FBD63CEC1F64F14B" bold="true" box="[1037,1100,1771,1797]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="33A0579C96464253FBD63CEC1F64F14B" box="[1037,1100,1771,1797]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="82DC2A7096464253FB873CEC1F89F14B" box="[1116,1185,1771,1797]" captionStart="FIGURE 4" captionStartId="9.[151,250,1809,1833]" captionTargetBox="[151,1436,705,1779]" captionTargetId="figure-254@9.[151,1436,705,1785]" captionTargetPageId="9" captionText="FIGURE 4. Molecular phylogeny of Diphtherophora species based on the 28S rRNA gene. The 50% majority rule consensus tree from the Bayesian analysis is presented. Six clades (A–F) are identified among Diphtherophora species. Branch support (only above 50%) is shown on branches as ML/BI. For simplicity, the phylogenetic tree is rooted to the branch leading to all Tripyla species. Sequences produced in this study are highlighted in gray (and in bold for the new species)." figureDoi="http://doi.org/10.5281/zenodo.4477005" httpUri="https://zenodo.org/record/4477005/files/figure.png" pageId="8" pageNumber="379">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="1A5836F596464252FF1C3D08194FF70C" blockId="8.[151,1437,1555,2014]" lastBlockId="9.[151,1437,151,681]" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">
Phylogenetic trees showed the existence of six strongly supported clades (A–F) within the 
<taxonomicName id="DDE74D7696464253FB3C3D081EB4F167" authorityName="de Man" authorityYear="1880" box="[1255,1436,1807,1833]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796464253FB3C3D081EB4F167" box="[1255,1436,1807,1833]" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
</taxonomicName>
group: clade A is represented by 
<taxonomicName id="DDE74D7696464253FDDA3D3319B7F103" authorityName="Ivanova" authorityYear="1958" box="[513,671,1843,1869]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="species" species="caudata">
<emphasis id="2893EAE796464253FDDA3D331934F103" box="[513,540,1844,1869]" italics="true" pageId="8" pageNumber="379">D.</emphasis>
cf. 
<emphasis id="2893EAE796464253FD9E3D3419B7F103" box="[581,671,1843,1869]" italics="true" pageId="8" pageNumber="379">caudata</emphasis>
</taxonomicName>
from Arasbaran (Northwestern 
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) and 
<taxonomicName id="DDE74D7696464253FBB63D331FC0F103" authorityName="Ivanova" authorityYear="1958" box="[1133,1256,1843,1869]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="species" species="caudata">
<emphasis id="2893EAE796464253FBB63D331FC0F103" box="[1133,1256,1843,1869]" italics="true" pageId="8" pageNumber="379">D. caudata</emphasis>
</taxonomicName>
from Maragheh (Northwestern 
<collectingCountry id="62F0766596464253FEE43D501A58F13F" box="[319,368,1879,1905]" name="Iran" pageId="8" pageNumber="379">Iran</collectingCountry>
), the latter species recovered in this study; clade B included sequences representing 
<taxonomicName id="DDE74D7696464253FACD3D5F1EB4F13F" authorityName="Thorne" authorityYear="1939" box="[1302,1436,1879,1905]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="species" species="obesa">
<emphasis id="2893EAE796464253FACD3D5F1E1AF13F" box="[1302,1330,1880,1905]" italics="true" pageId="8" pageNumber="379">D.</emphasis>
cf. 
<emphasis id="2893EAE796464253FA803D501EB4F13F" box="[1371,1436,1879,1905]" italics="true" pageId="8" pageNumber="379">obesa</emphasis>
</taxonomicName>
and 
<taxonomicName id="DDE74D7696464253FF1D3D7B1A18F1DB" authorityName="Ivanova" authorityYear="1980" box="[198,304,1916,1941]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="species" species="tenera">
<emphasis id="2893EAE796464253FF1D3D7B1A18F1DB" box="[198,304,1916,1941]" italics="true" pageId="8" pageNumber="379">D. tenera</emphasis>
</taxonomicName>
from different regions of Northern and Southern 
<collectingCountry id="62F0766596464253FC8C3D7C18AFF1DB" box="[855,903,1915,1941]" name="Iran" pageId="8" pageNumber="379">Iran</collectingCountry>
, the latter species, however, being paraphyletic; clade C was comprised of 
<taxonomicName id="DDE74D7696464253FE653DA71973F1F7" authorityName="Goodey" authorityYear="1951" baseAuthorityName="Van Der Linde" baseAuthorityYear="1938" box="[446,603,1951,1977]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="species" species="perplexans">
<emphasis id="2893EAE796464253FE653DA71973F1F7" box="[446,603,1951,1977]" italics="true" pageId="8" pageNumber="379">D. perplexans</emphasis>
</taxonomicName>
from different sites in Northwestern (Takab and Urmia) and Southern 
<collectingCountry id="62F0766596464253FAB43D981EB4F1F7" box="[1391,1436,1951,1977]" name="Iran" pageId="8" pageNumber="379">Iran</collectingCountry>
(Shiraz), the latter population recovered in this study, and a specimen characterized only as 
<taxonomicName id="DDE74D7696464253FB5D3DC41E13F193" authorityName="de Man" authorityYear="1880" box="[1158,1339,1987,2013]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="379" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796464253FB5D3DC41E13F193" box="[1158,1339,1987,2013]" italics="true" pageId="8" pageNumber="379">Diphtherophora</emphasis>
</taxonomicName>
sp. from CA, 
<collectingCountry id="62F0766596474252FF173A901A20F6FF" box="[204,264,151,177]" name="United States of America" pageId="9" pageNumber="380">USA</collectingCountry>
; clade D was solely represented by the new species 
<taxonomicName id="DDE74D7696474252FC8A3A9018F8F6FF" authority="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira, 2020" authorityName="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira" authorityYear="2020" box="[849,976,151,177]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="380" phylum="Tracheophyta" rank="species" species="eldarica" status="sp. nov.">
<emphasis id="2893EAE796474252FC8A3A9018F8F6FF" box="[849,976,151,177]" italics="true" pageId="9" pageNumber="380">D. eldarica</emphasis>
</taxonomicName>
<emphasis id="2893EAE796474252FC033A901F3EF6FF" bold="true" box="[984,1046,151,177]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="33A0579C96474252FC033A901F3EF6FF" box="[984,1046,151,177]" pageId="9" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; clade E harbored uncharacterized 
<taxonomicName id="DDE74D7696474252FF4C3ABC1A64F69B" authorityName="de Man" authorityYear="1880" box="[151,332,187,213]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FF4C3ABC1A64F69B" box="[151,332,187,213]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
species from different sites in Northern and Northwestern 
<collectingCountry id="62F0766596474252FC393ABC1F27F69B" box="[994,1039,187,213]" name="Iran" pageId="9" pageNumber="380">Iran</collectingCountry>
including Arasbaran and Ali Abad; clade F is solely represented by sequences of 
<taxonomicName id="DDE74D7696474252FD7B3AD81835F6B6" authorityName="Ghaderi, Kashi, Mirbabaei Karani &amp; Karegar" authorityYear="2017" box="[672,797,223,248]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="species" species="geraerti">
<emphasis id="2893EAE796474252FD7B3AD81835F6B6" box="[672,797,223,248]" italics="true" pageId="9" pageNumber="380">D. geraerti</emphasis>
</taxonomicName>
from Shiraz (Southern 
<collectingCountry id="62F0766596474252FBF03AD81F75F6B7" box="[1067,1117,223,249]" name="Iran" pageId="9" pageNumber="380">Iran</collectingCountry>
). Our phylogenetic analyses strongly supported (ML = 97, BI = 1.0) the new species 
<taxonomicName id="DDE74D7696474252FCC23B0418B1F753" authority="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira, 2020" authorityName="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira" authorityYear="2020" box="[793,921,259,285]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="380" phylum="Tracheophyta" rank="species" species="eldarica" status="sp. nov.">
<emphasis id="2893EAE796474252FCC23B0418B1F753" box="[793,921,259,285]" italics="true" pageId="9" pageNumber="380">D. eldarica</emphasis>
</taxonomicName>
<emphasis id="2893EAE796474252FC793B0418CAF753" bold="true" box="[930,994,259,285]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="33A0579C96474252FC793B0418CAF753" box="[930,994,259,285]" pageId="9" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
as a sister taxon of clade D, that is, 
<taxonomicName id="DDE74D7696474252FA5A3B041A3AF70F" authorityName="Goodey" authorityYear="1951" baseAuthorityName="Van Der Linde" baseAuthorityYear="1938" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="species" species="perplexans">
<emphasis id="2893EAE796474252FA5A3B041A3AF70F" italics="true" pageId="9" pageNumber="380">D. perplexans</emphasis>
</taxonomicName>
+ 
<taxonomicName id="DDE74D7696474252FEEB3B201ACDF70F" authorityName="de Man" authorityYear="1880" box="[304,485,295,321]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FEEB3B201ACDF70F" box="[304,485,295,321]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
sp. (
<figureCitation id="82DC2A7096474252FDC03B201973F70C" box="[539,603,295,322]" captionStart="FIGURE 4" captionStartId="9.[151,250,1809,1833]" captionTargetBox="[151,1436,705,1779]" captionTargetId="figure-254@9.[151,1436,705,1785]" captionTargetPageId="9" captionText="FIGURE 4. Molecular phylogeny of Diphtherophora species based on the 28S rRNA gene. The 50% majority rule consensus tree from the Bayesian analysis is presented. Six clades (A–F) are identified among Diphtherophora species. Branch support (only above 50%) is shown on branches as ML/BI. For simplicity, the phylogenetic tree is rooted to the branch leading to all Tripyla species. Sequences produced in this study are highlighted in gray (and in bold for the new species)." figureDoi="http://doi.org/10.5281/zenodo.4477005" httpUri="https://zenodo.org/record/4477005/files/figure.png" pageId="9" pageNumber="380">Fig.4</figureCitation>
).
</paragraph>
<paragraph id="1A5836F596474252FF1C3B4C19C3F4E7" blockId="9.[151,1437,151,681]" pageId="9" pageNumber="380">
Genetic divergence (p-distance %) within 
<taxonomicName id="DDE74D7696474252FD7D3B4C1873F72B" authorityName="de Man" authorityYear="1880" box="[678,859,331,357]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FD7D3B4C1873F72B" box="[678,859,331,357]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
clades was relatively low, ranging from 0% (clade A, among 
<taxonomicName id="DDE74D7696474252FED53B681AA3F7C7" authorityName="Ivanova" authorityYear="1958" box="[270,395,367,393]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="species" species="caudata">
<emphasis id="2893EAE796474252FED53B681AA3F7C7" box="[270,395,367,393]" italics="true" pageId="9" pageNumber="380">D. caudata</emphasis>
</taxonomicName>
sequences) to 5.2% (variation within clade E, represented by 
<taxonomicName id="DDE74D7696474252FB9C3B681FD4F7C7" authorityName="de Man" authorityYear="1880" box="[1095,1276,367,393]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FB9C3B681FD4F7C7" box="[1095,1276,367,393]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
sp. sequences from 
<collectingCountry id="62F0766596474252FF0F3B941A2DF7E3" box="[212,261,403,429]" name="Iran" pageId="9" pageNumber="380">Iran</collectingCountry>
; 
<tableCitation id="5765034E96474252FECB3B941A4CF7E3" box="[272,356,403,429]" captionStart="TABLE 3" captionStartId="10.[151,239,702,728]" captionTargetId="graphics-47@10.[151,2037,731,1449]" captionTargetPageId="10" captionText="TABLE 3. Genetic divergence as estimated by p–distance (%) within (diagonal in bold) and between clades." httpUri="http://table.plazi.org/id/4E98667D96444251FF4C38B91E37F496" pageId="9" pageNumber="380" tableUuid="4E98667D96444251FF4C38B91E37F496">Table 3</tableCitation>
). On the other hand, genetic divergence between 
<taxonomicName id="DDE74D7696474252FC563B941F6AF7E3" authorityName="de Man" authorityYear="1880" box="[909,1090,403,429]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FC563B941F6AF7E3" box="[909,1090,403,429]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
clades was much higher, ranging from 8.2% (clade A vs. B) to as high as 23.7% (clade C vs. F). For comparison, genetic divergence between any of the 
<taxonomicName id="DDE74D7696474252FF053BDC1ABBF7BB" authorityName="de Man" authorityYear="1880" box="[222,403,475,501]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FF053BDC1ABBF7BB" box="[222,403,475,501]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
clades and 
<taxonomicName id="DDE74D7696474252FDCD3BDC19FBF7BB" authorityName="de Man" authorityYear="1880" box="[534,723,475,501]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FDCD3BDC19FBF7BB" box="[534,723,475,501]" italics="true" pageId="9" pageNumber="380">Tylolaimophorus</emphasis>
</taxonomicName>
(also considered a representative of 
<taxonomicName id="DDE74D7696474252FBB73BDC1E6AF7B8" authorityName="Micoletzky" authorityYear="1922" box="[1132,1346,475,502]" class="Adenophorea" family="Diphtherophoridae" kingdom="Animalia" order="Triplonchida" pageId="9" pageNumber="380" phylum="Nematoda" rank="family">Diphtherophoridae</taxonomicName>
) ranged from 25.6% (
<taxonomicName id="DDE74D7696474252FEF43BF81AC4F457" authorityName="de Man" authorityYear="1880" box="[303,492,511,537]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FEF43BF81AC4F457" box="[303,492,511,537]" italics="true" pageId="9" pageNumber="380">Tylolaimophorus</emphasis>
</taxonomicName>
vs. clade A) to 32.7% (
<taxonomicName id="DDE74D7696474252FD203BF81890F457" authorityName="de Man" authorityYear="1880" box="[763,952,511,537]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FD203BF81890F457" box="[763,952,511,537]" italics="true" pageId="9" pageNumber="380">Tylolaimophorus</emphasis>
</taxonomicName>
vs. clade F). Similarly, genetic divergence between 
<taxonomicName id="DDE74D7696474252FF2038241A98F473" authorityName="de Man" authorityYear="1880" box="[251,432,547,573]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FF2038241A98F473" box="[251,432,547,573]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
clades and trichodorid genera were in the same range (24.1% to 30.1%) as those observed for comparisons with 
<taxonomicName id="DDE74D7696474252FE433840197DF42F" authorityName="de Man" authorityYear="1880" box="[408,597,583,609]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FE433840197DF42F" box="[408,597,583,609]" italics="true" pageId="9" pageNumber="380">Tylolaimophorus</emphasis>
</taxonomicName>
. This pattern is also consistent when comparing the genetic divergence between 
<taxonomicName id="DDE74D7696474252FF26386C1A9AF4CB" authorityName="de Man" authorityYear="1880" box="[253,434,619,645]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FF26386C1A9AF4CB" box="[253,434,619,645]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
clades and other outgroup taxa including genera 
<taxonomicName id="DDE74D7696474252FC3E386C1E17F4CB" authority="de Man, 1880" authorityName="de Man" authorityYear="1880" box="[997,1343,619,645]" class="Adenophorea" family="Prismatolaimidae" genus="Prismatolaimus" kingdom="Animalia" order="Enoplida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FC3E386C1FBEF4CB" box="[997,1174,619,645]" italics="true" pageId="9" pageNumber="380">Prismatolaimus</emphasis>
<bibRefCitation id="7E764B0496474252FB45386C1E17F4CB" author="de Man, J. G." box="[1182,1343,619,645]" pageId="9" pageNumber="380" pagination="1 - 104" refId="ref7257" refString="de Man, J. G. (1880) Die Einheimischen, frei in der reinen Erde und im sussen Wasser lebende Nematoden [Free-living soil and freshwater nematodes]. Monographisch Bearbeitet. Tijdschrift der Nederlandsche dierk undige Vereniging, 5, 1 - 104. [in German]" type="journal article" year="1880">de Man, 1880</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="DDE74D7696474252FA90386C1A1CF4E7" authority="Bastian, 1865" authorityName="Bastian" authorityYear="1865" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FA90386C1EB4F4CB" box="[1355,1436,619,645]" italics="true" pageId="9" pageNumber="380">Tripyla</emphasis>
<bibRefCitation id="7E764B0496474252FF4C38881A1CF4E7" author="Bastian, H. C." box="[151,308,654,681]" pageId="9" pageNumber="380" pagination="73 - 184" refId="ref6880" refString="Bastian, H. C. (1865) Monograph on the Anguillulidae. Transactions of the Linnean Society of London, 25, 73 - 184. https: // doi. org / 10.1111 / j. 1096 - 3642.1865. tb 00179. x" type="journal article" year="1865">Bastian, 1865</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="DDE74D7696474252FEB438881950F4E7" authority="Brzeski, 1963" authorityName="Brzeski" authorityYear="1963" box="[367,632,655,681]" class="Adenophorea" family="Trischistomatidae" genus="Tripylina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FEB438881AFEF4E7" box="[367,470,655,681]" italics="true" pageId="9" pageNumber="380">Tripylina</emphasis>
<bibRefCitation id="7E764B0496474252FE0738881950F4E7" author="Brzeski, M. W." box="[476,632,655,681]" pageId="9" pageNumber="380" pagination="295 - 308" refId="ref6973" refString="Brzeski, M. W. (1963) Nematode genera of the family Tripylidae (Nematoda, Enoplida). Acta Zoologica Cracoviensia, 8, 295 - 308." type="journal article" year="1963">Brzeski, 1963</bibRefCitation>
</taxonomicName>
(
<tableCitation id="5765034E96474252FD5D388819F5F4E7" box="[646,733,655,681]" captionStart="TABLE 3" captionStartId="10.[151,239,702,728]" captionTargetId="graphics-47@10.[151,2037,731,1449]" captionTargetPageId="10" captionText="TABLE 3. Genetic divergence as estimated by p–distance (%) within (diagonal in bold) and between clades." httpUri="http://table.plazi.org/id/4E98667D96444251FF4C38B91E37F496" pageId="9" pageNumber="380" tableUuid="4E98667D96444251FF4C38B91E37F496">Table 3</tableCitation>
).
</paragraph>
<caption id="4E98667D96474252FF4C3D161FFCF1DB" ID-DOI="http://doi.org/10.5281/zenodo.4477005" ID-Zenodo-Dep="4477005" httpUri="https://zenodo.org/record/4477005/files/figure.png" pageId="9" pageNumber="380" startId="9.[151,250,1809,1833]" targetBox="[151,1436,705,1779]" targetPageId="9">
<paragraph id="1A5836F596474252FF4C3D161FFCF1DB" blockId="9.[151,1437,1809,1941]" pageId="9" pageNumber="380">
<emphasis id="2893EAE796474252FF4C3D161A3BF167" bold="true" box="[151,275,1809,1833]" pageId="9" pageNumber="380">FIGURE 4.</emphasis>
Molecular phylogeny of 
<taxonomicName id="DDE74D7696474252FDCC3D161993F167" authorityName="de Man" authorityYear="1880" box="[535,699,1809,1833]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FDCC3D161993F167" box="[535,699,1809,1833]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
species based on the 28S rRNA gene. The 50% majority rule consensus tree from the Bayesian analysis is presented. Six clades (A–F) are identified among 
<taxonomicName id="DDE74D7696474252FC263D321F89F103" authorityName="de Man" authorityYear="1880" box="[1021,1185,1845,1869]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FC263D321F89F103" box="[1021,1185,1845,1869]" italics="true" pageId="9" pageNumber="380">Diphtherophora</emphasis>
</taxonomicName>
species. Branch support (only above 50%) is shown on branches as ML/BI. For simplicity, the phylogenetic tree is rooted to the branch leading to all 
<taxonomicName id="DDE74D7696474252FF4C3D7A1BC8F1DB" authorityName="Bastian" authorityYear="1865" box="[151,224,1917,1941]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="9" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796474252FF4C3D7A1BC8F1DB" box="[151,224,1917,1941]" italics="true" pageId="9" pageNumber="380">Tripyla</emphasis>
</taxonomicName>
species. Sequences produced in this study are highlighted in gray (and in bold for the new species).
</paragraph>
</caption>
<caption id="4E98667D96444251FF4C38B91E37F496" ID-Table-UUID="4E98667D96444251FF4C38B91E37F496" box="[151,1311,702,728]" httpUri="http://table.plazi.org/id/4E98667D96444251FF4C38B91E37F496" pageId="10" pageNumber="381" targetBox="[166,2008,745,1441]" targetIsTable="true" targetPageId="10">
<paragraph id="1A5836F596444251FF4C38B91E37F496" blockId="10.[151,1311,702,728]" box="[151,1311,702,728]" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF4C38B91A2BF496" bold="true" box="[151,259,702,728]" pageId="10" pageNumber="381">TABLE 3</emphasis>
. Genetic divergence as estimated by p–distance (%) within (diagonal in bold) and between clades.
</paragraph>
</caption>
<paragraph id="1A5836F596444251FF7D38EC1C80F3EE" pageId="10" pageNumber="381">
<table id="68E7C4559644BDA4FF7D38EE1CF0F3EF" box="[166,2008,745,1441]" gridcols="15" gridrows="15" pageId="10" pageNumber="381">
<tr id="A4D734B79644BDA4FF7D38EE1CF0F54A" box="[166,2008,745,772]" gridrow="0" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-6="1" rowspan-7="1">
<th id="E7065DCB9644BDA4FF7D38EE1BDEF54A" box="[166,246,745,772]" gridcol="0" gridrow="0" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D38EC1BCDF54A" bold="true" box="[166,229,747,772]" pageId="10" pageNumber="381">Clade</emphasis>
<emphasis id="2893EAE796444251FF3E38EE1BC4F4B9" bold="true" box="[229,236,745,759]" pageId="10" pageNumber="381">1</emphasis>
</th>
<th id="E7065DCB9644BDA4FEF038EE1986F54A" box="[299,686,745,772]" gridcol="1" gridrow="0" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF038EC1A52F54A" bold="true" box="[299,378,747,772]" pageId="10" pageNumber="381">Species</emphasis>
</th>
<th id="E7065DCB9644BDA4FCDF38EE1F05F54A" box="[772,1069,745,772]" colspan="4" colspanRight="3" gridcol="2" gridrow="0" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FCDF38EC18DDF54A" bold="true" box="[772,1013,747,772]" italics="true" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FCDF38EC1885F54A" authorityName="de Man" authorityYear="1880" box="[772,941,747,772]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">Diphtherophora</taxonomicName>
clades
</emphasis>
</th>
<th id="E7065DCB9644BDA4FADE38EE1EB5F54A" box="[1285,1437,745,772]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FADE38EC1E51F54A" bold="true" box="[1285,1401,747,772]" pageId="10" pageNumber="381">Outgroups</emphasis>
</th>
</tr>
<tr id="A4D734B79644BDA4FF7D391D1CF0F57D" box="[166,2008,794,819]" gridrow="1" pageId="10" pageNumber="381" rowspan-0="1" rowspan-1="1">
<td id="E7065DCB9644BDA4FCDF391D1818F57D" box="[772,816,794,819]" gridcol="2" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FCDF391D183EF57D" bold="true" box="[772,790,794,819]" pageId="10" pageNumber="381">A</emphasis>
</td>
<td id="E7065DCB9644BDA4FC80391D18FEF57D" box="[859,982,794,819]" colspan="2" colspanRight="1" gridcol="3" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FC80391D188AF57D" bold="true" box="[859,930,794,819]" pageId="10" pageNumber="381">B C</emphasis>
</td>
<td id="E7065DCB9644BDA4FBDA391D1F05F57D" box="[1025,1069,794,819]" gridcol="5" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FBDA391D1F3BF57D" bold="true" box="[1025,1043,794,819]" pageId="10" pageNumber="381">D</emphasis>
</td>
<td id="E7065DCB9644BDA4FB83391D1FACF57D" box="[1112,1156,794,819]" gridcol="6" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FB83391D1F41F57D" bold="true" box="[1112,1129,794,819]" pageId="10" pageNumber="381">E</emphasis>
</td>
<td id="E7065DCB9644BDA4FB74391D1FF3F57D" box="[1199,1243,794,819]" gridcol="7" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FB74391D1F96F57D" bold="true" box="[1199,1214,794,819]" pageId="10" pageNumber="381">F</emphasis>
</td>
<td id="E7065DCB9644BDA4FADE391D1E13F57D" box="[1285,1339,794,819]" gridcol="8" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FADE391D1E13F57D" bold="true" box="[1285,1339,794,819]" pageId="10" pageNumber="381">Tylo.</emphasis>
</td>
<td id="E7065DCB9644BDA4FAB1391D1EB5F57D" box="[1386,1437,794,819]" gridcol="9" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FAB1391D1EB5F57D" bold="true" box="[1386,1437,794,819]" pageId="10" pageNumber="381">Tric.</emphasis>
</td>
<td id="E7065DCB9644BDA4FA1D391D1D28F57D" box="[1478,1536,794,819]" gridcol="10" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FA1D391D1D28F57D" bold="true" box="[1478,1536,794,819]" pageId="10" pageNumber="381">Para.</emphasis>
</td>
<td id="E7065DCB9644BDA4F9FD391D1D43F57D" box="[1574,1643,794,819]" gridcol="11" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F9FD391D1D43F57D" bold="true" box="[1574,1643,794,819]" pageId="10" pageNumber="381">Mono.</emphasis>
</td>
<td id="E7065DCB9644BDA4F943391D1DF7F57D" box="[1688,1759,794,819]" gridcol="12" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F943391D1DF7F57D" bold="true" box="[1688,1759,794,819]" pageId="10" pageNumber="381">Prism.</emphasis>
</td>
<td id="E7065DCB9644BDA4F8D2391D1C71F57D" box="[1801,1881,794,819]" gridcol="13" gridrow="1" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F8D2391D1C71F57D" bold="true" box="[1801,1881,794,819]" pageId="10" pageNumber="381">Tripyli.</emphasis>
</td>
<td id="E7065DCB9644BDA4F853391D1CF0F57D" box="[1928,2008,794,819]" gridcol="14" gridrow="1" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251F853391D1CF0F57D" authorityName="Bastian" authorityYear="1865" box="[1928,2008,794,819]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251F853391D1CF0F57D" bold="true" box="[1928,2008,794,819]" pageId="10" pageNumber="381">Tripyla</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D394D1CF0F52D" box="[166,2008,842,867]" gridrow="2" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D394D1BDEF52D" box="[166,246,842,867]" gridcol="0" gridrow="2" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF71394D1B94F52D" bold="true" box="[170,188,842,867]" pageId="10" pageNumber="381">A</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF0394D1986F52D" box="[299,686,842,867]" gridcol="1" gridrow="2" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF0394D1A94F52D" authorityName="Ivanova" authorityYear="1958" box="[299,444,842,867]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="caudata">
<emphasis id="2893EAE796444251FEF0394D1A6CF52D" box="[299,324,842,867]" italics="true" pageId="10" pageNumber="381">D.</emphasis>
cf. 
<emphasis id="2893EAE796444251FEB1394D1A94F52D" box="[362,444,842,867]" italics="true" pageId="10" pageNumber="381">caudata</emphasis>
</taxonomicName>
</td>
<td id="E7065DCB9644BDA4FCDF394D1818F52D" box="[772,816,842,867]" gridcol="2" gridrow="2" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FCDF394D180CF52D" bold="true" box="[772,804,842,867]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D397E1CF0F5DD" box="[166,2008,889,915]" gridrow="3" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D397E1BDEF5DD" box="[166,246,889,915]" gridcol="0" gridrow="3" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF77397E1B95F5DC" bold="true" box="[172,189,889,914]" pageId="10" pageNumber="381">B</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF0397E1986F5DD" box="[299,686,889,915]" gridcol="1" gridrow="3" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF0397D1A8DF5DD" authorityName="Thorne" authorityYear="1939" box="[299,421,890,915]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="obesa">
<emphasis id="2893EAE796444251FEF0397D1A6CF5DD" box="[299,324,890,915]" italics="true" pageId="10" pageNumber="381">D.</emphasis>
cf. 
<emphasis id="2893EAE796444251FEB1397D1A8DF5DD" box="[362,421,890,915]" italics="true" pageId="10" pageNumber="381">obesa</emphasis>
</taxonomicName>
<emphasis id="2893EAE796444251FE77397D190BF5DD" box="[428,547,890,915]" italics="true" pageId="10" pageNumber="381">
+ 
<taxonomicName id="DDE74D7696444251FE18397D190BF5DD" authorityName="Ivanova" authorityYear="1980" box="[451,547,890,915]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="tenera">D. tenera</taxonomicName>
</emphasis>
</td>
<td id="E7065DCB9644BDA4FCDF397E1818F5DD" box="[772,816,889,915]" gridcol="2" gridrow="3" pageId="10" pageNumber="381">8.2</td>
<td id="E7065DCB9644BDA4FC80397E18AFF5DD" box="[859,903,889,915]" gridcol="3" gridrow="3" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FC80397E1853F5DC" bold="true" box="[859,891,889,914]" pageId="10" pageNumber="381">3.1</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D39AE1CF0F58D" box="[166,2008,937,963]" gridrow="4" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D39AE1BDEF58D" box="[166,246,937,963]" gridcol="0" gridrow="4" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7739AE1B96F58C" bold="true" box="[172,190,937,962]" pageId="10" pageNumber="381">C</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF039AE1986F58D" box="[299,686,937,963]" gridcol="1" gridrow="4" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF039AD1953F58D" box="[299,635,938,963]" italics="true" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF039AD1A91F58D" authorityName="Goodey" authorityYear="1951" baseAuthorityName="Van Der Linde" baseAuthorityYear="1938" box="[299,441,938,963]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="perplexans">D. perplexans</taxonomicName>
+ 
<taxonomicName id="DDE74D7696444251FE0C39AD1953F58D" authorityName="de Man" authorityYear="1880" box="[471,635,938,963]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">Diphtherophora</taxonomicName>
</emphasis>
sp.
</td>
<td id="E7065DCB9644BDA4FCDF39AE1818F58D" box="[772,816,937,963]" gridcol="2" gridrow="4" pageId="10" pageNumber="381">16.5</td>
<td id="E7065DCB9644BDA4FC8039AE18AFF58D" box="[859,903,937,963]" gridcol="3" gridrow="4" pageId="10" pageNumber="381">17.7</td>
<td id="E7065DCB9644BDA4FC7139AE18FEF58D" box="[938,982,937,963]" gridcol="4" gridrow="4" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FC7139AE18E2F58C" bold="true" box="[938,970,937,962]" pageId="10" pageNumber="381">0.2</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D39DF1CF0F5BC" box="[166,2008,984,1010]" gridrow="5" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D39DF1BDEF5BC" box="[166,246,984,1010]" gridcol="0" gridrow="5" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7739DE1B96F5BC" bold="true" box="[172,190,985,1010]" pageId="10" pageNumber="381">D</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF039DF1986F5BC" box="[299,686,984,1010]" gridcol="1" gridrow="5" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF039DE1AF5F5BC" bold="true" box="[299,477,985,1010]" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF039DE1AB6F5BC" authority="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira, 2020" authorityName="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira" authorityYear="2020" box="[299,414,985,1010]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="381" phylum="Tracheophyta" rank="species" species="eldarica" status="sp. nov.">
<emphasis id="2893EAE796444251FEF039DE1AB6F5BC" bold="true" box="[299,414,985,1010]" italics="true" pageId="10" pageNumber="381">D. eldarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33A0579C96444251FE7E39DE1AF5F5BC" box="[421,477,985,1010]" pageId="10" pageNumber="381" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
<emphasis id="2893EAE796444251FE0539DF1ACDF5A8" bold="true" box="[478,485,984,998]" pageId="10" pageNumber="381">2</emphasis>
</td>
<td id="E7065DCB9644BDA4FCDF39DF1818F5BC" box="[772,816,984,1010]" gridcol="2" gridrow="5" pageId="10" pageNumber="381">14.2</td>
<td id="E7065DCB9644BDA4FC8039DF18AFF5BC" box="[859,903,984,1010]" gridcol="3" gridrow="5" pageId="10" pageNumber="381">13.5</td>
<td id="E7065DCB9644BDA4FC7139DF18FEF5BC" box="[938,982,984,1010]" gridcol="4" gridrow="5" pageId="10" pageNumber="381">17.5</td>
<td id="E7065DCB9644BDA4FBDA39DF1F05F5BC" box="[1025,1069,984,1010]" gridcol="5" gridrow="5" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FBDA39DE1F09F5BC" bold="true" box="[1025,1057,985,1010]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3E0E1CF0F26C" box="[166,2008,1033,1058]" gridrow="6" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D3E0E1BDEF26C" box="[166,246,1033,1058]" gridcol="0" gridrow="6" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF773E0E1B95F26C" bold="true" box="[172,189,1033,1058]" pageId="10" pageNumber="381">E</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03E0E1986F26C" box="[299,686,1033,1058]" gridcol="1" gridrow="6" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03E0E1AE7F26C" authorityName="de Man" authorityYear="1880" box="[299,463,1033,1058]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FEF03E0E1AE7F26C" box="[299,463,1033,1058]" italics="true" pageId="10" pageNumber="381">Diphtherophora</emphasis>
</taxonomicName>
sp.
</td>
<td id="E7065DCB9644BDA4FCDF3E0E1818F26C" box="[772,816,1033,1058]" gridcol="2" gridrow="6" pageId="10" pageNumber="381">17.0</td>
<td id="E7065DCB9644BDA4FC803E0E18AFF26C" box="[859,903,1033,1058]" gridcol="3" gridrow="6" pageId="10" pageNumber="381">16.8</td>
<td id="E7065DCB9644BDA4FC713E0E18FEF26C" box="[938,982,1033,1058]" gridcol="4" gridrow="6" pageId="10" pageNumber="381">20.2</td>
<td id="E7065DCB9644BDA4FBDA3E0E1F05F26C" box="[1025,1069,1033,1058]" gridcol="5" gridrow="6" pageId="10" pageNumber="381">19.5</td>
<td id="E7065DCB9644BDA4FB833E0E1FACF26C" box="[1112,1156,1033,1058]" gridcol="6" gridrow="6" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FB833E0E1F50F26C" bold="true" box="[1112,1144,1033,1058]" pageId="10" pageNumber="381">5.2</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3E3E1CF0F21C" box="[166,2008,1081,1106]" gridrow="7" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-8="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D3E3E1BDEF21C" box="[166,246,1081,1106]" gridcol="0" gridrow="7" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF773E3E1B93F21C" bold="true" box="[172,187,1081,1106]" pageId="10" pageNumber="381">F</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03E3E1986F21C" box="[299,686,1081,1106]" gridcol="1" gridrow="7" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03E3E1AB4F21C" authorityName="Ghaderi, Kashi, Mirbabaei Karani &amp; Karegar" authorityYear="2017" box="[299,412,1081,1106]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="geraerti">
<emphasis id="2893EAE796444251FEF03E3E1AB4F21C" box="[299,412,1081,1106]" italics="true" pageId="10" pageNumber="381">D. geraerti</emphasis>
</taxonomicName>
</td>
<td id="E7065DCB9644BDA4FCDF3E3E1818F21C" box="[772,816,1081,1106]" gridcol="2" gridrow="7" pageId="10" pageNumber="381">18.4</td>
<td id="E7065DCB9644BDA4FC803E3E18AFF21C" box="[859,903,1081,1106]" gridcol="3" gridrow="7" pageId="10" pageNumber="381">17.9</td>
<td id="E7065DCB9644BDA4FC713E3E18FEF21C" box="[938,982,1081,1106]" gridcol="4" gridrow="7" pageId="10" pageNumber="381">23.7</td>
<td id="E7065DCB9644BDA4FBDA3E3E1F05F21C" box="[1025,1069,1081,1106]" gridcol="5" gridrow="7" pageId="10" pageNumber="381">19.1</td>
<td id="E7065DCB9644BDA4FB833E3E1FACF21C" box="[1112,1156,1081,1106]" gridcol="6" gridrow="7" pageId="10" pageNumber="381">21.5</td>
<td id="E7065DCB9644BDA4FB743E3E1FF3F21C" box="[1199,1243,1081,1106]" gridcol="7" gridrow="7" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FB743E3E1FE7F21C" bold="true" box="[1199,1231,1081,1106]" pageId="10" pageNumber="381">4.6</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3E6E1CF0F2CC" box="[166,2008,1129,1154]" gridrow="8" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1" rowspan-9="1">
<th id="E7065DCB9644BDA4FF7D3E6E1BDEF2CC" box="[166,246,1129,1154]" gridcol="0" gridrow="8" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3E6E1BF3F2CC" bold="true" box="[166,219,1129,1154]" pageId="10" pageNumber="381">Tylo.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03E6E1986F2CC" box="[299,686,1129,1154]" gridcol="1" gridrow="8" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03E6E1AA9F2CC" box="[299,385,1129,1154]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="minor">
<emphasis id="2893EAE796444251FEF03E6E1AA9F2CC" box="[299,385,1129,1154]" italics="true" pageId="10" pageNumber="381">T. minor</emphasis>
</taxonomicName>
</td>
<td id="E7065DCB9644BDA4FCDF3E6E1818F2CC" box="[772,816,1129,1154]" gridcol="2" gridrow="8" pageId="10" pageNumber="381">25.8</td>
<td id="E7065DCB9644BDA4FC803E6E18AFF2CC" box="[859,903,1129,1154]" gridcol="3" gridrow="8" pageId="10" pageNumber="381">28.4</td>
<td id="E7065DCB9644BDA4FC713E6E18FEF2CC" box="[938,982,1129,1154]" gridcol="4" gridrow="8" pageId="10" pageNumber="381">30.2</td>
<td id="E7065DCB9644BDA4FBDA3E6E1F05F2CC" box="[1025,1069,1129,1154]" gridcol="5" gridrow="8" pageId="10" pageNumber="381">32.3</td>
<td id="E7065DCB9644BDA4FB833E6E1FACF2CC" box="[1112,1156,1129,1154]" gridcol="6" gridrow="8" pageId="10" pageNumber="381">28.9</td>
<td id="E7065DCB9644BDA4FB743E6E1FF3F2CC" box="[1199,1243,1129,1154]" gridcol="7" gridrow="8" pageId="10" pageNumber="381">32.7</td>
<td id="E7065DCB9644BDA4FADE3E6E1E13F2CC" box="[1285,1339,1129,1154]" gridcol="8" gridrow="8" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FADE3E6E1E0DF2CC" bold="true" box="[1285,1317,1129,1154]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3E9F1CF0F2FC" box="[166,2008,1176,1202]" gridrow="9" pageId="10" pageNumber="381" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1">
<th id="E7065DCB9644BDA4FF7D3E9F1BDEF2FC" box="[166,246,1176,1202]" gridcol="0" gridrow="9" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3E9F1BF0F2FF" bold="true" box="[166,216,1176,1201]" pageId="10" pageNumber="381">Tric.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03E9F1986F2FC" box="[299,686,1176,1202]" gridcol="1" gridrow="9" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF03E9E1907F2FC" box="[299,559,1177,1202]" italics="true" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03E9E1ABFF2FC" box="[299,407,1177,1202]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="iranicus">T. iranicus</taxonomicName>
+ 
<taxonomicName id="DDE74D7696444251FE6E3E9E1907F2FC" box="[437,559,1177,1202]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="variabilis">T. variabilis</taxonomicName>
</emphasis>
</td>
<td id="E7065DCB9644BDA4FCDF3E9F1818F2FC" box="[772,816,1176,1202]" gridcol="2" gridrow="9" pageId="10" pageNumber="381">24.1</td>
<td id="E7065DCB9644BDA4FC803E9F18AFF2FC" box="[859,903,1176,1202]" gridcol="3" gridrow="9" pageId="10" pageNumber="381">24.3</td>
<td id="E7065DCB9644BDA4FC713E9F18FEF2FC" box="[938,982,1176,1202]" gridcol="4" gridrow="9" pageId="10" pageNumber="381">27.4</td>
<td id="E7065DCB9644BDA4FBDA3E9F1F05F2FC" box="[1025,1069,1176,1202]" gridcol="5" gridrow="9" pageId="10" pageNumber="381">27.5</td>
<td id="E7065DCB9644BDA4FB833E9F1FACF2FC" box="[1112,1156,1176,1202]" gridcol="6" gridrow="9" pageId="10" pageNumber="381">26.8</td>
<td id="E7065DCB9644BDA4FB743E9F1FF3F2FC" box="[1199,1243,1176,1202]" gridcol="7" gridrow="9" pageId="10" pageNumber="381">28.8</td>
<td id="E7065DCB9644BDA4FADE3E9F1E13F2FC" box="[1285,1339,1176,1202]" gridcol="8" gridrow="9" pageId="10" pageNumber="381">31.0</td>
<td id="E7065DCB9644BDA4FAB13E9F1EB5F2FC" box="[1386,1437,1176,1202]" gridcol="9" gridrow="9" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FAB13E9F1EA2F2FF" bold="true" box="[1386,1418,1176,1201]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3ECF1CF0F2AC" box="[166,2008,1224,1250]" gridrow="10" pageId="10" pageNumber="381" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-14="1">
<th id="E7065DCB9644BDA4FF7D3ECF1BDEF2AC" box="[166,246,1224,1250]" gridcol="0" gridrow="10" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3ECF1BF7F2AF" bold="true" box="[166,223,1224,1249]" pageId="10" pageNumber="381">Para.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03ECF1986F2AC" box="[299,686,1224,1250]" gridcol="1" gridrow="10" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03ECE1A98F2AC" box="[299,432,1225,1250]" class="Adenophorea" family="Prismatolaimidae" genus="Prismatolaimus" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="hispanicus">
<emphasis id="2893EAE796444251FEF03ECE1A98F2AC" box="[299,432,1225,1250]" italics="true" pageId="10" pageNumber="381">P. hispanicus</emphasis>
</taxonomicName>
</td>
<td id="E7065DCB9644BDA4FCDF3ECF1818F2AC" box="[772,816,1224,1250]" gridcol="2" gridrow="10" pageId="10" pageNumber="381">25.2</td>
<td id="E7065DCB9644BDA4FC803ECF18AFF2AC" box="[859,903,1224,1250]" gridcol="3" gridrow="10" pageId="10" pageNumber="381">25.3</td>
<td id="E7065DCB9644BDA4FC713ECF18FEF2AC" box="[938,982,1224,1250]" gridcol="4" gridrow="10" pageId="10" pageNumber="381">27.3</td>
<td id="E7065DCB9644BDA4FBDA3ECF1F05F2AC" box="[1025,1069,1224,1250]" gridcol="5" gridrow="10" pageId="10" pageNumber="381">27.7</td>
<td id="E7065DCB9644BDA4FB833ECF1FACF2AC" box="[1112,1156,1224,1250]" gridcol="6" gridrow="10" pageId="10" pageNumber="381">28.5</td>
<td id="E7065DCB9644BDA4FB743ECF1FF3F2AC" box="[1199,1243,1224,1250]" gridcol="7" gridrow="10" pageId="10" pageNumber="381">30.1</td>
<td id="E7065DCB9644BDA4FADE3ECF1E13F2AC" box="[1285,1339,1224,1250]" gridcol="8" gridrow="10" pageId="10" pageNumber="381">31.3</td>
<td id="E7065DCB9644BDA4FAB13ECF1EB5F2AC" box="[1386,1437,1224,1250]" gridcol="9" gridrow="10" pageId="10" pageNumber="381">20.4</td>
<td id="E7065DCB9644BDA4FA1D3ECF1D28F2AC" box="[1478,1536,1224,1250]" gridcol="10" gridrow="10" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FA1D3ECF1ECEF2AF" bold="true" box="[1478,1510,1224,1249]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3EFF1CF0F35F" box="[166,2008,1272,1297]" gridrow="11" pageId="10" pageNumber="381" rowspan-12="1" rowspan-13="1" rowspan-14="1">
<th id="E7065DCB9644BDA4FF7D3EFF1BDEF35F" box="[166,246,1272,1297]" gridcol="0" gridrow="11" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3EFF1BC3F35F" bold="true" box="[166,235,1272,1297]" pageId="10" pageNumber="381">Mono.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03EFF1986F35F" box="[299,686,1272,1297]" gridcol="1" gridrow="11" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03EFF195CF35F" authority="+ Monotricodorus" authorityName="Monotricodorus" box="[299,628,1272,1297]" pageId="10" pageNumber="381" rank="species" species="vangunfyi">
<emphasis id="2893EAE796444251FEF03EFF195CF35F" box="[299,628,1272,1297]" italics="true" pageId="10" pageNumber="381">M. vangunfyi + Monotricodorus</emphasis>
</taxonomicName>
sp.
</td>
<td id="E7065DCB9644BDA4FCDF3EFF1818F35F" box="[772,816,1272,1297]" gridcol="2" gridrow="11" pageId="10" pageNumber="381">26.4</td>
<td id="E7065DCB9644BDA4FC803EFF18AFF35F" box="[859,903,1272,1297]" gridcol="3" gridrow="11" pageId="10" pageNumber="381">26.1</td>
<td id="E7065DCB9644BDA4FC713EFF18FEF35F" box="[938,982,1272,1297]" gridcol="4" gridrow="11" pageId="10" pageNumber="381">29.1</td>
<td id="E7065DCB9644BDA4FBDA3EFF1F05F35F" box="[1025,1069,1272,1297]" gridcol="5" gridrow="11" pageId="10" pageNumber="381">29.6</td>
<td id="E7065DCB9644BDA4FB833EFF1FACF35F" box="[1112,1156,1272,1297]" gridcol="6" gridrow="11" pageId="10" pageNumber="381">28.9</td>
<td id="E7065DCB9644BDA4FB743EFF1FF3F35F" box="[1199,1243,1272,1297]" gridcol="7" gridrow="11" pageId="10" pageNumber="381">29.7</td>
<td id="E7065DCB9644BDA4FADE3EFF1E13F35F" box="[1285,1339,1272,1297]" gridcol="8" gridrow="11" pageId="10" pageNumber="381">30.1</td>
<td id="E7065DCB9644BDA4FAB13EFF1EB5F35F" box="[1386,1437,1272,1297]" gridcol="9" gridrow="11" pageId="10" pageNumber="381">22.4</td>
<td id="E7065DCB9644BDA4FA1D3EFF1D28F35F" box="[1478,1536,1272,1297]" gridcol="10" gridrow="11" pageId="10" pageNumber="381">21.9</td>
<td id="E7065DCB9644BDA4F9FD3EFF1D43F35F" box="[1574,1643,1272,1297]" gridcol="11" gridrow="11" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F9F53EFF1D72F35F" bold="true" box="[1582,1626,1272,1297]" pageId="10" pageNumber="381">10.3</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3F201CF0F30F" box="[166,2008,1319,1345]" gridrow="12" pageId="10" pageNumber="381" rowspan-13="1" rowspan-14="1">
<th id="E7065DCB9644BDA4FF7D3F201BDEF30F" box="[166,246,1319,1345]" gridcol="0" gridrow="12" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3F2F1BC4F30F" bold="true" box="[166,236,1320,1345]" pageId="10" pageNumber="381">Prism.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03F201986F30F" box="[299,686,1319,1345]" gridcol="1" gridrow="12" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03F2F1A98F30F" box="[299,432,1320,1345]" class="Adenophorea" family="Prismatolaimidae" genus="Prismatolaimus" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="dolichurus">
<emphasis id="2893EAE796444251FEF03F2F1A98F30F" box="[299,432,1320,1345]" italics="true" pageId="10" pageNumber="381">P. dolichurus</emphasis>
</taxonomicName>
1
</td>
<td id="E7065DCB9644BDA4FCDF3F201818F30F" box="[772,816,1319,1345]" gridcol="2" gridrow="12" pageId="10" pageNumber="381">27.9</td>
<td id="E7065DCB9644BDA4FC803F2018AFF30F" box="[859,903,1319,1345]" gridcol="3" gridrow="12" pageId="10" pageNumber="381">27.9</td>
<td id="E7065DCB9644BDA4FC713F2018FEF30F" box="[938,982,1319,1345]" gridcol="4" gridrow="12" pageId="10" pageNumber="381">30.2</td>
<td id="E7065DCB9644BDA4FBDA3F201F05F30F" box="[1025,1069,1319,1345]" gridcol="5" gridrow="12" pageId="10" pageNumber="381">30.3</td>
<td id="E7065DCB9644BDA4FB833F201FACF30F" box="[1112,1156,1319,1345]" gridcol="6" gridrow="12" pageId="10" pageNumber="381">26.6</td>
<td id="E7065DCB9644BDA4FB743F201FF3F30F" box="[1199,1243,1319,1345]" gridcol="7" gridrow="12" pageId="10" pageNumber="381">29.4</td>
<td id="E7065DCB9644BDA4FADE3F201E13F30F" box="[1285,1339,1319,1345]" gridcol="8" gridrow="12" pageId="10" pageNumber="381">28.3</td>
<td id="E7065DCB9644BDA4FAB13F201EB5F30F" box="[1386,1437,1319,1345]" gridcol="9" gridrow="12" pageId="10" pageNumber="381">29.7</td>
<td id="E7065DCB9644BDA4FA1D3F201D28F30F" box="[1478,1536,1319,1345]" gridcol="10" gridrow="12" pageId="10" pageNumber="381">30.7</td>
<td id="E7065DCB9644BDA4F9FD3F201D43F30F" box="[1574,1643,1319,1345]" gridcol="11" gridrow="12" pageId="10" pageNumber="381">30.4</td>
<td id="E7065DCB9644BDA4F9433F201DF7F30F" box="[1688,1759,1319,1345]" gridcol="12" gridrow="12" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F9433F2F1D90F30F" bold="true" box="[1688,1720,1320,1345]" pageId="10" pageNumber="381">0.0</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3F5F1CF0F33F" box="[166,2008,1368,1393]" gridrow="13" pageId="10" pageNumber="381" rowspan-14="1">
<th id="E7065DCB9644BDA4FF7D3F5F1BDEF33F" box="[166,246,1368,1393]" gridcol="0" gridrow="13" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FF7D3F5F1BDEF33F" bold="true" box="[166,246,1368,1393]" pageId="10" pageNumber="381">Tripyli.</emphasis>
</th>
<td id="E7065DCB9644BDA4FEF03F5F1986F33F" box="[299,686,1368,1393]" gridcol="1" gridrow="13" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF03F5F1986F33F" box="[299,686,1368,1393]" italics="true" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03F5F1AA1F33F" box="[299,393,1368,1393]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="tamaki">T. tamaki</taxonomicName>
+ 
<taxonomicName id="DDE74D7696444251FE7C3F5F1926F33F" box="[423,526,1368,1393]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="tearoha">T. tearoha</taxonomicName>
+ 
<taxonomicName id="DDE74D7696444251FDF03F5F1986F33F" box="[555,686,1368,1393]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="manurewa">T. manurewa</taxonomicName>
</emphasis>
</td>
<td id="E7065DCB9644BDA4FCDF3F5F1818F33F" box="[772,816,1368,1393]" gridcol="2" gridrow="13" pageId="10" pageNumber="381">28.6</td>
<td id="E7065DCB9644BDA4FC803F5F18AFF33F" box="[859,903,1368,1393]" gridcol="3" gridrow="13" pageId="10" pageNumber="381">27.1</td>
<td id="E7065DCB9644BDA4FC713F5F18FEF33F" box="[938,982,1368,1393]" gridcol="4" gridrow="13" pageId="10" pageNumber="381">30.1</td>
<td id="E7065DCB9644BDA4FBDA3F5F1F05F33F" box="[1025,1069,1368,1393]" gridcol="5" gridrow="13" pageId="10" pageNumber="381">31.8</td>
<td id="E7065DCB9644BDA4FB833F5F1FACF33F" box="[1112,1156,1368,1393]" gridcol="6" gridrow="13" pageId="10" pageNumber="381">28.9</td>
<td id="E7065DCB9644BDA4FB743F5F1FF3F33F" box="[1199,1243,1368,1393]" gridcol="7" gridrow="13" pageId="10" pageNumber="381">32.7</td>
<td id="E7065DCB9644BDA4FADE3F5F1E13F33F" box="[1285,1339,1368,1393]" gridcol="8" gridrow="13" pageId="10" pageNumber="381">30.6</td>
<td id="E7065DCB9644BDA4FAB13F5F1EB5F33F" box="[1386,1437,1368,1393]" gridcol="9" gridrow="13" pageId="10" pageNumber="381">29.5</td>
<td id="E7065DCB9644BDA4FA1D3F5F1D28F33F" box="[1478,1536,1368,1393]" gridcol="10" gridrow="13" pageId="10" pageNumber="381">31.5</td>
<td id="E7065DCB9644BDA4F9FD3F5F1D43F33F" box="[1574,1643,1368,1393]" gridcol="11" gridrow="13" pageId="10" pageNumber="381">29.7</td>
<td id="E7065DCB9644BDA4F9433F5F1DF7F33F" box="[1688,1759,1368,1393]" gridcol="12" gridrow="13" pageId="10" pageNumber="381">28.0</td>
<td id="E7065DCB9644BDA4F8D23F5F1C71F33F" box="[1801,1881,1368,1393]" gridcol="13" gridrow="13" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251F8D23F5F1C01F33F" bold="true" box="[1801,1833,1368,1393]" pageId="10" pageNumber="381">3.8</emphasis>
</td>
</tr>
<tr id="A4D734B79644BDA4FF7D3F801CF0F3EF" box="[166,2008,1415,1441]" gridrow="14" pageId="10" pageNumber="381">
<th id="E7065DCB9644BDA4FF7D3F801BDEF3EF" box="[166,246,1415,1441]" gridcol="0" gridrow="14" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FF7D3F801BDEF3EE" authorityName="Bastian" authorityYear="1865" box="[166,246,1415,1440]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="380" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FF7D3F801BDEF3EE" bold="true" box="[166,246,1415,1440]" pageId="10" pageNumber="381">Tripyla</emphasis>
</taxonomicName>
</th>
<td id="E7065DCB9644BDA4FEF03F801986F3EF" box="[299,686,1415,1441]" gridcol="1" gridrow="14" pageId="10" pageNumber="381">
<emphasis id="2893EAE796444251FEF03F8F191CF3EF" box="[299,564,1416,1441]" italics="true" pageId="10" pageNumber="381">
<taxonomicName id="DDE74D7696444251FEF03F8F1A9AF3EF" authorityName="de Man" authorityYear="1880" box="[299,434,1416,1441]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="filicaudata">T. filicaudata</taxonomicName>
+ 
<taxonomicName id="DDE74D7696444251FE0B3F8F191CF3EF" box="[464,564,1416,1441]" class="Adenophorea" family="Tripylidae" genus="Tripyla" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="species" species="bioblitz">T. bioblitz</taxonomicName>
</emphasis>
</td>
<td id="E7065DCB9644BDA4FCDF3F801818F3EF" box="[772,816,1415,1441]" gridcol="2" gridrow="14" pageId="10" pageNumber="381">21.9</td>
<td id="E7065DCB9644BDA4FC803F8018AFF3EF" box="[859,903,1415,1441]" gridcol="3" gridrow="14" pageId="10" pageNumber="381">22.5</td>
<td id="E7065DCB9644BDA4FC713F8018FEF3EF" box="[938,982,1415,1441]" gridcol="4" gridrow="14" pageId="10" pageNumber="381">24.8</td>
<td id="E7065DCB9644BDA4FBDA3F801F05F3EF" box="[1025,1069,1415,1441]" gridcol="5" gridrow="14" pageId="10" pageNumber="381">25.6</td>
<td id="E7065DCB9644BDA4FB833F801FACF3EF" box="[1112,1156,1415,1441]" gridcol="6" gridrow="14" pageId="10" pageNumber="381">23.8</td>
<td id="E7065DCB9644BDA4FB743F801FF3F3EF" box="[1199,1243,1415,1441]" gridcol="7" gridrow="14" pageId="10" pageNumber="381">26.5</td>
<td id="E7065DCB9644BDA4FADE3F801E13F3EF" box="[1285,1339,1415,1441]" gridcol="8" gridrow="14" pageId="10" pageNumber="381">26.0</td>
<td id="E7065DCB9644BDA4FAB13F801EB5F3EF" box="[1386,1437,1415,1441]" gridcol="9" gridrow="14" pageId="10" pageNumber="381">24.7</td>
<td id="E7065DCB9644BDA4FA1D3F801D28F3EF" box="[1478,1536,1415,1441]" gridcol="10" gridrow="14" pageId="10" pageNumber="381">24.6</td>
<td id="E7065DCB9644BDA4F9FD3F801D43F3EF" box="[1574,1643,1415,1441]" gridcol="11" gridrow="14" pageId="10" pageNumber="381">28.1</td>
<td id="E7065DCB9644BDA4F9433F801DF7F3EF" box="[1688,1759,1415,1441]" gridcol="12" gridrow="14" pageId="10" pageNumber="381">25.0</td>
<td id="E7065DCB9644BDA4F8D23F801C71F3EF" box="[1801,1881,1415,1441]" gridcol="13" gridrow="14" pageId="10" pageNumber="381">25.3</td>
<td id="E7065DCB9644BDA4F8533F801CF0F3EF" box="[1928,2008,1415,1441]" gridcol="14" gridrow="14" pageId="10" pageNumber="381">3.4</td>
</tr>
</table>
</paragraph>
<tableNote id="E701377B96444251FF4C3FA819BEF3A0" pageId="10" pageNumber="381" targetBox="[166,2008,745,1441]" targetPageId="10">
<paragraph id="1A5836F596444251FF4C3FA81CA2F384" blockId="10.[151,1930,1455,1518]" box="[151,1930,1455,1482]" pageId="10" pageNumber="381">
<superScript id="ED929BBD96444251FF4C3FA81BB7F3F0" attach="right" box="[151,159,1455,1470]" fontSize="6" pageId="10" pageNumber="381">1</superScript>
Taxon abbreviations: 
<taxonomicName id="DDE74D7696444251FE5D3FB719A4F384" authority="(Tylo.)" baseAuthorityName="Tylo." box="[390,652,1456,1482]" class="Secernentea" family="Diplogasteridae" genus="Tylolaimophorus" kingdom="Animalia" order="Diplogasterida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FE5D3FB71912F384" box="[390,570,1456,1482]" italics="true" pageId="10" pageNumber="381">Tylolaimophorus</emphasis>
(Tylo.)
</taxonomicName>
, 
<taxonomicName id="DDE74D7696444251FD4C3FB7184EF384" authority="(Tric.)" baseAuthorityName="Tric." box="[663,870,1456,1482]" class="Adenophorea" family="Trichodoridae" genus="Trichodorus" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FD4C3FB71831F384" box="[663,793,1456,1482]" italics="true" pageId="10" pageNumber="381">Trichodorus</emphasis>
(Tric.)
</taxonomicName>
, 
<taxonomicName id="DDE74D7696444251FCAA3FB71F5CF384" authority="(Para.)" baseAuthorityName="Para." box="[881,1140,1456,1482]" class="Adenophorea" family="Trichodoridae" genus="Paratrichodorus" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FCAA3FB71F0BF384" box="[881,1059,1456,1482]" italics="true" pageId="10" pageNumber="381">Paratrichodorus</emphasis>
(Para.)
</taxonomicName>
, 
<taxonomicName id="DDE74D7696444251FBA43FB71EB3F384" authority="(Mono.)" baseAuthorityName="Mono." box="[1151,1435,1456,1482]" class="Adenophorea" family="Trichodoridae" genus="Monotrichodorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Triplonchida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FBA43FB71E12F384" box="[1151,1338,1456,1482]" italics="true" pageId="10" pageNumber="381">Monotrichodorus</emphasis>
(Mono.)
</taxonomicName>
, 
<taxonomicName id="DDE74D7696444251FA7E3FB71D86F384" authority="(Prism.)" baseAuthorityName="Prism." box="[1445,1710,1456,1482]" class="Adenophorea" family="Prismatolaimidae" genus="Prismatolaimus" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251FA7E3FB71D66F384" box="[1445,1614,1456,1482]" italics="true" pageId="10" pageNumber="381">Prismatolaimus</emphasis>
(Prism.)
</taxonomicName>
, 
<taxonomicName id="DDE74D7696444251F9623FB71CAFF384" authority="(Tripyli.)" baseAuthorityName="Tripyli." box="[1721,1927,1456,1482]" class="Adenophorea" family="Trischistomatidae" genus="Tripylina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Enoplida" pageId="10" pageNumber="381" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796444251F9623FB71C33F384" box="[1721,1819,1456,1482]" italics="true" pageId="10" pageNumber="381">Tripylina</emphasis>
(Tripyli.)
</taxonomicName>
.
</paragraph>
<paragraph id="1A5836F596444251FF4C3FD419BEF3A0" blockId="10.[151,1930,1455,1518]" box="[151,662,1491,1518]" pageId="10" pageNumber="381">
<superScript id="ED929BBD96444251FF4C3FD41BB7F3AC" attach="right" box="[151,159,1491,1506]" fontSize="6" pageId="10" pageNumber="381">2</superScript>
Taxon represented by only one DNA sequence.
</paragraph>
</tableNote>
<paragraph id="1A5836F596454250FF1C3A901829F70F" blockId="11.[151,1437,150,321]" pageId="11" pageNumber="382">
With respect to the new species 
<taxonomicName id="DDE74D7696454250FDF03A901982F6FF" authority="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira, 2020" authorityName="Ghaderi &amp; Hesar &amp; Karegar &amp; Pereira" authorityYear="2020" box="[555,682,151,177]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="382" phylum="Tracheophyta" rank="species" species="eldarica" status="sp. nov.">
<emphasis id="2893EAE796454250FDF03A901982F6FF" box="[555,682,151,177]" italics="true" pageId="11" pageNumber="382">D. eldarica</emphasis>
</taxonomicName>
<emphasis id="2893EAE796454250FD6A3A9019C7F6FF" bold="true" box="[689,751,151,177]" pageId="11" pageNumber="382">
<taxonomicNameLabel id="33A0579C96454250FD6A3A9019C7F6FF" box="[689,751,151,177]" pageId="11" pageNumber="382" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, genetic divergence ranged from 13.5% (between clade B, 
<taxonomicName id="DDE74D7696454250FA5A3A901BF7F69A" authorityName="Ivanova" authorityYear="1980" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="11" pageNumber="382" phylum="Nematoda" rank="species" species="tenera">
<emphasis id="2893EAE796454250FA5A3A901BF7F69A" italics="true" pageId="11" pageNumber="382">D. tenera</emphasis>
</taxonomicName>
+ 
<taxonomicName id="DDE74D7696454250FF263ABC1AACF69B" authorityName="Thorne" authorityYear="1939" box="[253,388,187,213]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="11" pageNumber="382" phylum="Nematoda" rank="species" species="obesa">
<emphasis id="2893EAE796454250FF263ABC1A30F69A" box="[253,280,187,212]" italics="true" pageId="11" pageNumber="382">D.</emphasis>
cf. 
<emphasis id="2893EAE796454250FE983ABC1AACF69B" box="[323,388,187,213]" italics="true" pageId="11" pageNumber="382">obesa</emphasis>
</taxonomicName>
) to 19.5% (between clade E, 
<taxonomicName id="DDE74D7696454250FD0A3ABC18AEF69B" authorityName="de Man" authorityYear="1880" box="[721,902,187,213]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="11" pageNumber="382" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796454250FD0A3ABC18AEF69B" box="[721,902,187,213]" italics="true" pageId="11" pageNumber="382">Diphtherophora</emphasis>
</taxonomicName>
sp.). Although all six clades were strongly supported in our analyses, with ML branch support always greater than 98% and BPP always with maximum support, some relationships among 
<taxonomicName id="DDE74D7696454250FE1F3B041951F753" authorityName="de Man" authorityYear="1880" box="[452,633,259,285]" class="Adenophorea" family="Diphterophoridae" genus="Diphtherophora" kingdom="Animalia" order="Dorylaimida" pageId="11" pageNumber="382" phylum="Nematoda" rank="genus">
<emphasis id="2893EAE796454250FE1F3B041951F753" box="[452,633,259,285]" italics="true" pageId="11" pageNumber="382">Diphtherophora</emphasis>
</taxonomicName>
clades (
<emphasis id="2893EAE796454250FD033B0419D2F752" box="[728,762,259,284]" italics="true" pageId="11" pageNumber="382">i.e.</emphasis>
some of the deeper nodes) were either weakly supported or not recovered by one of the inference methods (
<figureCitation id="82DC2A7096454250FD743B2019DCF70F" box="[687,756,295,321]" captionStart="FIGURE 4" captionStartId="9.[151,250,1809,1833]" captionTargetBox="[151,1436,705,1779]" captionTargetId="figure-254@9.[151,1436,705,1785]" captionTargetPageId="9" captionText="FIGURE 4. Molecular phylogeny of Diphtherophora species based on the 28S rRNA gene. The 50% majority rule consensus tree from the Bayesian analysis is presented. Six clades (A–F) are identified among Diphtherophora species. Branch support (only above 50%) is shown on branches as ML/BI. For simplicity, the phylogenetic tree is rooted to the branch leading to all Tripyla species. Sequences produced in this study are highlighted in gray (and in bold for the new species)." figureDoi="http://doi.org/10.5281/zenodo.4477005" httpUri="https://zenodo.org/record/4477005/files/figure.png" pageId="11" pageNumber="382">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>