147 lines
18 KiB
XML
147 lines
18 KiB
XML
<document id="01F5153AA6884DE20A6EFDB0F9B916D2" ID-DOI="10.11646/zootaxa.5543.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14386776" ID-ZooBank="E34AF3EF-4D03-4C08-8E11-C9514D42021B" IM.bibliography_approvedBy="guilherme" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1733919186538" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2024" docId="03C83A1CFF81C35DFF772F2653B14438" docLanguage="en" docName="zootaxa.5543.4.1.pdf" docOrigin="Zootaxa 5543 (4)" docSource="https://doi.org/10.11646/zootaxa.5543.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pillsburiaster micropeltus Mah 2024, nov. comb." docType="treatment" docVersion="2" lastPageNumber="479" masterDocId="FFF14264FF9DC341FFE02E495341417D" masterDocTitle="New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific" masterLastPageNumber="500" masterPageNumber="451" pageNumber="479" updateTime="1734380713031" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BE209F7C2952B7822B42BD91C7A841C6">New genera and species of deep-sea Goniasteridae (Asteroidea) from the North Pacific</mods:title>
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<mods:namePart id="57786282CB78863CAC3BE9680E46136A">Mah, Christopher L.</mods:namePart>
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<mods:title id="158102AD8C8DF70FB2833F9047D8615B">Zootaxa</mods:title>
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<mods:part id="6B8C8899853A964EBC1AA2138C09E406">
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<mods:date id="CBF91A6E62E94BCEA1FC91764020C88F">2024</mods:date>
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<mods:number id="E81DC3304F698AD85E04DE3EFB64A1AB">2024-12-05</mods:number>
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<treatment id="03C83A1CFF81C35DFF772F2653B14438" ID-DOI="http://doi.org/10.5281/zenodo.14503284" ID-Zenodo-Dep="14503284" LSID="urn:lsid:plazi:treatment:03C83A1CFF81C35DFF772F2653B14438" httpUri="http://treatment.plazi.org/id/03C83A1CFF81C35DFF772F2653B14438" lastPageNumber="479" pageId="28" pageNumber="479">
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<subSubSection id="C37BD881FF81C35DFF772F26504840F4" box="[151,777,367,394]" pageId="28" pageNumber="479" type="nomenclature">
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<paragraph id="8BDE8B0AFF81C35DFF772F26504840F4" blockId="28.[151,777,367,429]" box="[151,777,367,394]" pageId="28" pageNumber="479">
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<heading id="D0963C66FF81C35DFF772F26504840F4" bold="true" box="[151,777,367,394]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
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<emphasis id="B9155718FF81C35DFF772F26504840F4" bold="true" box="[151,777,367,394]" pageId="28" pageNumber="479">
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<taxonomicName id="4C61F089FF81C35DFF772F26513A40F4" authority="(Fisher 1906)" authorityName="Mah" authorityYear="2024" baseAuthorityName="Fisher" baseAuthorityYear="1906" box="[151,635,367,394]" class="Asteroidea" family="Goniasteridae" genus="Pillsburiaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="micropeltus" status="comb. nov.">
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<emphasis id="B9155718FF81C35DFF772F26529140F7" bold="true" box="[151,464,367,394]" italics="true" pageId="28" pageNumber="479">Pillsburiaster micropeltus</emphasis>
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(
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<bibRefCitation id="EFF0F6FBFF81C35DFE3E2F26513240F4" author="Fisher, W. K." box="[478,627,367,393]" pageId="28" pageNumber="479" pagination="987 - 1130" refId="ref21055" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher 1906</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A226EA63FF81C35DFD632F26504840F4" box="[643,777,367,393]" pageId="28" pageNumber="479" rank="species">nov. comb.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BD881FF81C35DFF772FDA520940D0" box="[151,328,403,429]" pageId="28" pageNumber="479" type="description">
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<paragraph id="8BDE8B0AFF81C35DFF772FDA520940D0" blockId="28.[151,777,367,429]" box="[151,328,403,429]" pageId="28" pageNumber="479">
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<figureCitation id="135A978FFF81C35DFF772FDA520940D0" box="[151,328,403,429]" captionStart="FIGURE 9" captionStartId="29.[151,250,1679,1703]" captionTargetBox="[151,1436,181,1655]" captionTargetId="figure-15@29.[151,1436,181,1655]" captionTargetPageId="29" captionText="FIGURE 9. Pillsburiaster micropeltus nov. comb. Holotype USNM 21164. A. Abactinal. B. Closeup of coarse abactinal granules. C. Armtip. D. Actinal. E. Adambulacral plates, furrow and subambulacral spination. Scale bar=5.0 mm: A, D; Scale bar=2.0 mm: B; Scale bar=3.0 mm: C; Scale bar=2.0 mm: E." figureDoi="http://doi.org/10.5281/zenodo.14386808" httpUri="https://zenodo.org/record/14386808/files/figure.png" pageId="28" pageNumber="479">FIGURE 9A–D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BD881FF81C35DFF772F9151F14350" pageId="28" pageNumber="479" type="reference_group">
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<paragraph id="8BDE8B0AFF81C35DFF772F9151474092" blockId="28.[151,790,471,557]" box="[151,518,471,496]" pageId="28" pageNumber="479">
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<treatmentCitationGroup id="AB71AC24FF81C35DFF772F9151474092" box="[151,518,471,496]" pageId="28" pageNumber="479">
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<treatmentCitation id="0AC0AD1BFF81C35DFF772F9151434092" author="Fisher, W. K." box="[151,514,471,496]" page="1006" pageId="28" pageNumber="479" year="1906">
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<taxonomicName id="4C61F089FF81C35DFF772F9151434092" ID-CoL="57KCB" authority="Fisher 1906: 1006" authorityName="Fisher" authorityPageNumber="1006" authorityYear="1906" box="[151,514,471,496]" class="Asteroidea" family="Goniasteridae" genus="Tosia" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="micropelta">
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<emphasis id="B9155718FF81C35DFF772F915201408D" box="[151,320,472,496]" italics="true" pageId="28" pageNumber="479">Tosia micropelta</emphasis>
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<bibRefCitation id="EFF0F6FBFF81C35DFEA72F9151434092" author="Fisher, W. K." box="[327,514,471,496]" pageId="28" pageNumber="479" pagination="987 - 1130" refId="ref21055" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher 1906: 1006</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BDE8B0AFF81C35DFF772FBF50574373" blockId="28.[151,790,471,557]" box="[151,790,502,526]" pageId="28" pageNumber="479">
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<treatmentCitationGroup id="AB71AC24FF81C35DFF772FBF50574373" box="[151,790,502,526]" pageId="28" pageNumber="479">
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<treatmentCitation id="0AC0AD1BFF81C35DFF772FBF51654373" author="Fisher, W. K." box="[151,548,502,526]" page="205" pageId="28" pageNumber="479" year="1911">
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<taxonomicName id="4C61F089FF81C35DFF772FBF51654373" ID-CoL="76DC8" authority="Fisher 1911: 205" authorityName="Fisher" authorityPageNumber="205" authorityYear="1911" box="[151,548,502,526]" class="Asteroidea" family="Goniasteridae" genus="Peltaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="micropeltus">
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<emphasis id="B9155718FF81C35DFF772FBF52314373" box="[151,368,502,526]" italics="true" pageId="28" pageNumber="479">Peltaster micropeltus</emphasis>
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<bibRefCitation id="EFF0F6FBFF81C35DFE962FBF51654373" author="Fisher, W. K." box="[374,548,502,526]" pageId="28" pageNumber="479" pagination="1 - 420" refId="ref21085" refString="Fisher, W. K. (1911) Asteroidea of the North Pacific and adjacent waters. 1. Phanerozonia and Spinulosida. Bulletin of the US National Museum, 76, i - xiii + 1 - 420. https: // doi. org / 10.5479 / si. 03629236.76. i" type="journal article" year="1911">Fisher 1911: 205</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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; A.M.
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<treatmentCitation id="0AC0AD1BFF81C35DFD8A2FBF50534373" author="Clark, A. M." box="[618,786,502,526]" page="269" pageId="28" pageNumber="479" year="1993">
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<bibRefCitation id="EFF0F6FBFF81C35DFD8A2FBF50534373" author="Clark, A. M." box="[618,786,502,526]" pageId="28" pageNumber="479" pagination="187 - 366" refId="ref20685" refString="Clark, A. M. (1993) An index of names of recent Asteroidea Part 2: Valvatida. E chinoderm Studies, 4, 187 - 366. https: // doi. org / 10.1201 / 9781003072553 - 4" type="journal article" year="1993">Clark 1993: 269</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BDE8B0AFF81C35DFF772C5C51F14350" blockId="28.[151,790,471,557]" box="[151,688,533,557]" pageId="28" pageNumber="479">
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<taxonomicName id="4C61F089FF81C35DFF772C5C514B4350" ID-CoL="6VKK3" authority="McKnight 1973" authorityName="McKnight" authorityYear="1973" box="[151,522,533,557]" class="Asteroidea" family="Goniasteridae" genus="Pillsburiaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="maini" status="syn. nov.">
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<emphasis id="B9155718FF81C35DFF772C5C52224350" box="[151,355,533,557]" italics="true" pageId="28" pageNumber="479">Pillsburiaster maini</emphasis>
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<bibRefCitation id="EFF0F6FBFF81C35DFE892C5C514B4350" author="McKnight, D. G." box="[361,522,533,557]" pageId="28" pageNumber="479" pagination="171 - 195" refId="ref22755" refString="McKnight, D. G. (1973) Additions to the asteroid fauna of New Zealand: Family Goniasteridae. NZOI Records, 1 (13), 171 - 195." type="journal article" year="1973">McKnight 1973</bibRefCitation>
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</taxonomicName>
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<emphasis id="B9155718FF81C35DFDF02C5C51F14350" bold="true" box="[528,688,533,557]" pageId="28" pageNumber="479">
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<taxonomicNameLabel id="A226EA63FF81C35DFDF02C5C51F14350" box="[528,688,533,557]" pageId="28" pageNumber="479">new synonymy</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37BD881FF81C35DFF772C12567A457C" pageId="28" pageNumber="479" type="diagnosis">
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<paragraph id="8BDE8B0AFF81C35DFF772C1252514308" blockId="28.[151,272,603,629]" box="[151,272,603,629]" pageId="28" pageNumber="479">
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<emphasis id="B9155718FF81C35DFF772C1252514308" bold="true" box="[151,272,603,629]" pageId="28" pageNumber="479">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BDE8B0AFF81C35DFF772CEA577F43C0" blockId="28.[151,1437,675,1026]" box="[151,1086,675,702]" pageId="28" pageNumber="479">Body stellate (R/r=1.93), interradial arcs weakly curved, arms triangular, tips blunt.</paragraph>
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<paragraph id="8BDE8B0AFF81C35DFF272C8E567A457C" blockId="28.[151,1437,675,1026]" pageId="28" pageNumber="479">
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<emphasis id="B9155718FF81C35DFF272C8E57964278" bold="true" pageId="28" pageNumber="479">Abactinal plate round to irregularly polygonal, each covered by coarse, round granules, 1 to 4, mostly 2 to 3. Pedicellariae, small upright, spoon-shaped valves similar in size to adjacent granules.</emphasis>
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Superomarginals and inferomarginals 26 from armtip to armtip (13 per side) each with a distinct naked bald spot on the abactinal and lateral surface covered by granules. Spatulate pedicellariae present on proximal plates. Inferomarginals covered by granules with a similar bald spot on the actinal face-smaller in size than the corresponding superomarginal plate. Terminal plate large with a thimble-shaped tubercle. Actinal plates irregular, quadrate to hexagonal, covered by round granules. Weakly developed fasciolar grooves. Furrow spines, blunt tipped, 5 to 6 with subambulacrals in two series, set off from furrow spines, each composed of cylindrical spinelets, 3 to 4 per row, close-set. Mouth plates with 10 furrow spinelets, surface with granules similar to subambulacral spinelets on adambulacral plates.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37BD881FF81C35DFF772A1A53B14438" pageId="28" pageNumber="479" type="discussion">
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<paragraph id="8BDE8B0AFF81C35DFF772A1A52524510" blockId="28.[151,275,1107,1133]" box="[151,275,1107,1133]" pageId="28" pageNumber="479">
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<heading id="D0963C66FF81C35DFF772A1A52524510" bold="true" box="[151,275,1107,1133]" fontSize="11" level="1" pageId="28" pageNumber="479" reason="1">
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<emphasis id="B9155718FF81C35DFF772A1A52524510" bold="true" box="[151,275,1107,1133]" pageId="28" pageNumber="479">Comment</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDE8B0AFF81C35DFF772AD253B14438" blockId="28.[151,1437,1179,1349]" pageId="28" pageNumber="479">
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USNM 21164, the
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<typeStatus id="54DA35A8FF81C35DFE9A2AD2529D45C8" box="[378,476,1179,1205]" pageId="28" pageNumber="479" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C61F089FF81C35DFDEB2AD5502145C8" authority="Fisher 1906" authorityName="Fisher" authorityYear="1906" box="[523,864,1179,1205]" class="Asteroidea" family="Goniasteridae" genus="Tosia" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="micropelta">
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<emphasis id="B9155718FF81C35DFDEB2AD5518B45C8" box="[523,714,1179,1205]" italics="true" pageId="28" pageNumber="479">Tosia micropelta</emphasis>
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<bibRefCitation id="EFF0F6FBFF81C35DFD362AD2502145C8" author="Fisher, W. K." box="[726,864,1179,1205]" pageId="28" pageNumber="479" pagination="987 - 1130" refId="ref21055" refString="Fisher, W. K. (1906) The starfishes of the Hawaiian islands. Bulletin of the United States Fish Commission, 23, 987 - 1130." type="journal article" year="1906">Fisher 1906</bibRefCitation>
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</taxonomicName>
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possesses the coarse, round, clustered, abactinal granulation diagnostic of
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<taxonomicName id="4C61F089FF81C35DFE5E2AF651BF45A4" authority="Halpern 1970" authorityName="Halpern" authorityYear="1970" box="[446,766,1215,1241]" class="Asteroidea" family="Goniasteridae" genus="Pillsburiaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="genus">
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<emphasis id="B9155718FF81C35DFE5E2AF6511945A4" box="[446,600,1215,1241]" italics="true" pageId="28" pageNumber="479">Pillsburiaster</emphasis>
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Halpern 1970
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</taxonomicName>
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rather than
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<taxonomicName id="4C61F089FF81C35DFC6C2AF657C645A4" authority="Verrill 1899" authorityName="Verrill" authorityYear="1899" box="[908,1159,1215,1241]" class="Asteroidea" family="Goniasteridae" genus="Peltaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="genus">
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<emphasis id="B9155718FF81C35DFC6C2AF650B045A4" box="[908,1009,1215,1241]" italics="true" pageId="28" pageNumber="479">Peltaster</emphasis>
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<bibRefCitation id="EFF0F6FBFF81C35DFC1B2AF657C645A4" author="Verrill, A. E." box="[1019,1159,1215,1241]" pageId="28" pageNumber="479" pagination="145 - 234" refId="ref23351" refString="Verrill, A. E. (1899) Revision of certain genera and species of starfishes, with descriptions of new forms. Transactions of the Connecticut Academy of Arts and Sciences, 10 (1), 145 - 234. https: // doi. org / 10.5962 / bhl. part. 7031" type="journal article" year="1899">Verrill 1899</bibRefCitation>
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</taxonomicName>
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, which is defined based on the presence of secondary plates in addition to weakly fasciolar, tabulate plates; all of which are absent. The weakly stellate to stellate body shape (
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<collectionCode id="ED7013CFFF81C35DFDA12B4E5114445C" box="[577,597,1287,1313]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="28" pageNumber="479">R</collectionCode>
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/r=1.93) and its depth,
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<quantity id="4C9926EFFF81C35DFCB22B4E50B3445F" box="[850,1010,1287,1314]" metricMagnitude="3" metricUnit="m" metricValue="1.0825" metricValueMax="1.593" metricValueMin="0.572" pageId="28" pageNumber="479" unit="m" value="1082.5" valueMax="1593.0" valueMin="572.0">572 to 1593 m</quantity>
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are also consistent with
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<taxonomicName id="4C61F089FF81C35DFAE22B4E53B14438" class="Asteroidea" family="Goniasteridae" genus="Pillsburiaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="479" phylum="Echinodermata" rank="species" species="undetermined">
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<emphasis id="B9155718FF81C35DFAE22B4E56DD445C" box="[1282,1436,1287,1313]" italics="true" pageId="28" pageNumber="479">Pillsburiaster</emphasis>
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species.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |