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<mods:title id="BA6CF8B762AF2E9D0C5517C36D6CD365">Taxonomic revision of the “ mexicanus ” group of the genus Vaejovis C. L. Koch, 1836 (Scorpiones: Vaejovidae)</mods:title>
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<emphasis id="B8FAEAFAFF801967FF163F00FEF5FCB4" bold="true" box="[151,338,789,815]" italics="true" pageId="36" pageNumber="37">Vaejovis dzahui</emphasis>
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<figureCitation id="12B52A6DFF801967FF163F22FEA5FCCB" box="[151,258,823,848]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="36" pageNumber="37">Figures 4</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FE8F3F22FE92FCCB" box="[270,309,823,848]" captionStart="FIGURE 13" captionStartId="22.[151,250,989,1011]" captionTargetBox="[151,1435,193,968]" captionTargetId="figure@22.[151,1435,193,968]" captionTargetPageId="22" captionText="FIGURE 13. Comparison in shapes on carapace in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659119/files/figure.png" pageId="36" pageNumber="37">13f</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FEC03F2DFECBFCCB" box="[321,364,824,848]" captionStart="FIGURE 14" captionStartId="23.[151,250,1558,1580]" captionTargetBox="[340,1236,345,1537]" captionTargetId="figure@23.[340,1247,337,1537]" captionTargetPageId="23" captionText="FIGURE 14. Comparison of the different shapes in the caudal gland on species within the “mexicanus” group (each caudal gland, if present, is highlighted in red): a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov." httpUri="https://zenodo.org/record/2659121/files/figure.png" pageId="36" pageNumber="37">14e</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FEF63F22FE38FCCB" box="[375,415,823,848]" captionStart="FIGURE 15" captionStartId="24.[151,250,1398,1420]" captionTargetBox="[340,1247,195,1374]" captionTargetId="figure@24.[340,1247,195,1374]" captionTargetPageId="24" captionText="FIGURE 15. Comparison in shapes and trichobothrial patterns in the pedipalp femur, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659123/files/figure.png" pageId="36" pageNumber="37">15f</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FE2A3F22FE74FCCB" box="[427,467,823,848]" captionStart="FIGURE 16" captionStartId="25.[151,250,1480,1502]" captionTargetBox="[340,1247,258,1458]" captionTargetId="figure@25.[340,1247,258,1458]" captionTargetPageId="25" captionText="FIGURE 16. Comparison in shapes and trichobothrial patterns in the pedipalp patella, in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659125/files/figure.png" pageId="36" pageNumber="37">16f</figureCitation>
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<figureCitation id="12B52A6DFF801967FE5F3F22FDA1FCCB" box="[478,518,823,848]" captionStart="FIGURE 17" captionStartId="26.[151,250,1585,1607]" captionTargetBox="[335,1252,335,1567]" captionTargetId="figure@26.[335,1252,335,1567]" captionTargetPageId="26" captionText="FIGURE 17. Comparison in shapes and trichobothrial patterns in the pedipalp chela of males in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659127/files/figure.png" pageId="36" pageNumber="37">17f</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FD933F22FD9EFCCB" box="[530,569,823,848]" captionStart="FIGURE 18" captionStartId="27.[151,250,1747,1769]" captionTargetBox="[340,1247,589,1726]" captionTargetId="figure@27.[340,1247,589,1726]" captionTargetPageId="27" captionText="FIGURE 18. Comparison in the different shapes and trichobothrial patterns in the pedipalp chela of females, that selected species within “mexicanus” group present: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659129/files/figure.png" pageId="36" pageNumber="37">18f</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FDC43F22FDCAFCCB" box="[581,621,823,848]" captionStart="FIGURE 19" captionStartId="29.[151,250,1805,1827]" captionTargetBox="[319,1255,877,1784]" captionTargetId="figure@29.[319,1267,877,1784]" captionTargetPageId="29" captionText="FIGURE 19. Comparison on the shapes in the hemispermatophore in species within “mexicanus” group: a—V. ceboruco sp. nov.; b—V. chiapas; c—V. coalcoman; d—V. darwini; e—V. dugesi; f—V. dzahui; g—V. franckei; h—V. granulatus; i—V. maculosus; j—V. mcwesti; k—V. mexicanus; l—V. montanus; m—V. monticola; n—V. morelia; o—V. nanchititla sp. nov. Scale bar = 1 mm." httpUri="https://zenodo.org/record/2659131/files/figure.png" pageId="36" pageNumber="37">19f</figureCitation>
, 
<figureCitation id="12B52A6DFF801967FDF93F2DFD33FCCB" box="[632,660,824,848]" captionStart="FIGURE 24" captionStartId="37.[151,250,1268,1290]" captionTargetBox="[340,1244,195,1247]" captionTargetId="figure@37.[340,1247,195,1247]" captionTargetPageId="37" captionText="FIGURE 24. Habitus of Vaejovis dzahui holotype (♂): a—dorsal view; b—ventral view. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2659141/files/figure.png" pageId="36" pageNumber="37">24</figureCitation>
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<emphasis id="B8FAEAFAFF801967FF163F94FE91FC0D" box="[151,310,896,918]" italics="true" pageId="36" pageNumber="37">Vaejovis dzahui</emphasis>
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–559, 566–567, Fig: 30–36, 52.
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<emphasis id="B8FAEAFAFF801967FF163FD7FEF7FC40" bold="true" box="[151,336,962,987]" pageId="36" pageNumber="37">Type material.</emphasis>
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male. 
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: 
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, 
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road to 
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(
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). 
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. 
<collectorName id="277B533EFF801967FB243FF3FB44FC64" box="[1189,1251,998,1023]" pageId="36" pageNumber="37">Cols.</collectorName>
: 
<collectorName id="277B533EFF801967FB733FF2FAD2FC64" box="[1266,1397,998,1023]" pageId="36" pageNumber="37">O. Francke</collectorName>
, 
<collectorName id="277B533EFF801967FA043FF2FF52FBB8" pageId="36" pageNumber="37">G. Villegas</collectorName>
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and 
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(
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). 
<collectorName id="277B533EFF801967FD77381FFCC8FBB8" box="[758,879,1034,1059]" pageId="36" pageNumber="37">Examined.</collectorName>
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</paragraph>
<paragraph id="8A3136E8FF801967FF46383AFE21FBF0" blockId="36.[151,1437,962,2031]" pageId="36" pageNumber="37">
<materialsCitation id="3AE63CB5FF801967FF46383AFBAAFBDC" ID-GBIF-Occurrence="2237797860" box="[199,1037,1070,1095]" collectingDate="2006-06-28" collectionCode="CNAN-T" collectorName="Cols. &amp; O. Francke &amp; G. Villegas &amp; H. Montano &amp; A. Valdez &amp; Examined." country="Mexico" elevation="2290" latitude="17.7" location="San Cristobal Suchixtlahuaca-Santiago Tepujan" longLatPrecision="5555" longitude="-97.38333" municipality="Distrito Coixtlahuaca" pageId="36" pageNumber="37" specimenCode="CNAN-T0387" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Oaxaca" typeStatus="paratype">
<typeStatus id="5535884AFF801967FF46383AFE9EFBDC" box="[199,313,1071,1095]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9C88FD61FF801967FEC7383BFED1FBDD" box="[326,374,1070,1095]" pageId="36" pageNumber="37" type="male">5 ♂</specimenCount>
, 
<specimenCount id="9C88FD61FF801967FE07383AFE14FBDD" box="[390,435,1071,1095]" pageId="36" pageNumber="37" type="female">2 ♀</specimenCount>
, same data as holotype (
<specimenCode id="DA289E93FF801967FD58383AFC25FBDC" box="[729,898,1071,1095]" collectionCode="CNAN-T" pageId="36" pageNumber="37">CNAN-T0387</specimenCode>
). Examined
</materialsCitation>
. 
<materialsCitation id="3AE63CB5FF801967FB9A383AFE25FBF0" ID-GBIF-Occurrence="2237797757" collectingDate="2006-06-28" collectionCode="AMNH" collectorName="Cols. &amp; O. Francke &amp; G. Villegas &amp; H. Montano &amp; A. Valdez &amp; Examined." country="Mexico" elevation="2290" latitude="17.7" location="San Cristobal Suchixtlahuaca-Santiago Tepujan" longLatPrecision="5555" longitude="-97.38333" municipality="Distrito Coixtlahuaca" pageId="36" pageNumber="37" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Oaxaca" typeStatus="paratype">
<specimenCount id="9C88FD61FF801967FB9A383AFBECFBDD" box="[1051,1099,1071,1095]" pageId="36" pageNumber="37" type="male">4 ♂</specimenCount>
, 
<specimenCount id="9C88FD61FF801967FBDA383AFB2EFBDD" box="[1115,1161,1071,1095]" pageId="36" pageNumber="37" type="female">2 ♀</specimenCount>
, same data as holotype (
<collectionCode id="EC9FAE2DFF801967FF1E3846FF5CFBF0" box="[159,251,1107,1131]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="36" pageNumber="37">AMNH</collectionCode>
). Examined
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C2946563FF801967FF463863FB5DF818" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8A3136E8FF801967FF463863FEC2F88C" blockId="36.[151,1437,962,2031]" pageId="36" pageNumber="37">
<emphasis id="B8FAEAFAFF801967FF463863FEE6FB14" bold="true" box="[199,321,1142,1167]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Adults 
<quantity id="4D769B0DFF801967FE273862FD84FB15" box="[422,547,1143,1167]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="36" pageNumber="37" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 mm</quantity>
long (Fig: 24-a, b). Carapace (Fig: 13-f), slightly shorter than metasomal segment V in males, longer in females (Carapace L/MS-V 0.94±0.1 [1.09±0.03]); longer than femur (Carapace L/ Femur L 1.37±0.1 [1.4±0.02]); anterior margin of carapace slightly concave. Mesosomal tergite VII with two pairs of weak to inconspicuous carinae. Pectinal teeth 13–14 [12–13]; sternite V on males with a whitish patch almost absent, present only on posterior margin; sternite VII with carinae obsolete, only noticed as a small elevation of the cuticle, with 10 to 15 setae. Segment I considerably wider than long, rectangular in dorsal view (MSI L/ 
<geoCoordinate id="EFBA502FFF801967FA03393EFF60FAFC" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-0.61">W 0.61</geoCoordinate>
±0.08 [0.61±0.01]); ventral submedian carinae on metasomal segment I obsolete, on II weak and crenulated; intercarinal spaces on metasoma smooth to shagreened; lateral supramedian carinae on segment IV absent; segment V longer than wide (MS-V L/ 
<geoCoordinate id="EFBA502FFF801967FDEA3982FD64FA34" box="[619,707,1430,1455]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-1.84">W 1.84</geoCoordinate>
±0.15 [1.75±0.17]), nearly as deep as wide (W/D 1.16 ±0.09 [1.1±0.1]). Vesicle slender (V L/ 
<geoCoordinate id="EFBA502FFF801967FDA439AEFDD9FA48" box="[549,638,1467,1491]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-1.87">W 1.87</geoCoordinate>
±0.14 [1.81±0.3]) (Fig: 15-d), wider than deep (W/D 1.39± 0.15 [1.38±0.19]). Ventral surface of vesicle smooth, with one dark, median longitudinal band; dorsal surface flat, medially with a small, whitish spot on distal one-third of vesicle (Fig: 14-f). Pedipalp femur (Fig: 15-f) less than three times longer than wide (L/ 
<geoCoordinate id="EFBA502FFF801967FD883A32FDF8F9A4" box="[521,607,1575,1599]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-2.77">W 2.77</geoCoordinate>
±0.2 [2.81±0.2]), slightly wider than deep (W/D 1.29±0.2 [1.07± 0.07]); anterior face of pedipalp femur flat, with some scattered large granules on basal fourth. Pedipalp patella (Fig: 16-f) less than three times longer than wide (L/ 
<geoCoordinate id="EFBA502FFF801967FD2C3A7AFCACF91C" box="[685,779,1646,1671]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-2.58">W 2.58</geoCoordinate>
±0.25 [2.56±0.05]), wider than deep (W/D 1.15±0.01 [1.19±0.06]); dorsointernal carina of patella inconspicuous. Pedipalp chela manus (Fig: 17, 18-f) less than two times longer than wide (L/ 
<geoCoordinate id="EFBA502FFF801967FE5A3AA2FD93F954" box="[475,564,1719,1743]" direction="west" orientation="longitude" pageId="36" pageNumber="37" precision="555" value="-1.72">W 1.72</geoCoordinate>
±0.08 [1.94±0.06]); fixed finger shorter than manus (CM L/FF 1.39±0.1 [1.17±0.12]), with all the carinae positions pigmented but the cuticle not raised from the surface. Dentate margins of fingers straight.
</paragraph>
<paragraph id="8A3136E8FF801967FF463B37FABAF8C5" blockId="36.[151,1437,962,2031]" pageId="36" pageNumber="37">
Hemispermatophore (Fig: 19-f): lamelliform (TL 2.5; LL 1.4; LW 
<quantity id="4D769B0DFF801967FC4E3B36FB8AF8A0" box="[975,1069,1827,1851]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="36" pageNumber="37" unit="mm" value="0.3">0.3 mm</quantity>
); lamella long, with no crest or decoration; two hooks located basally, lobe-shaped. Capsular region with an inner median lobe present.
</paragraph>
<paragraph id="8A3136E8FF801967FF463B7FFB5DF818" blockId="36.[151,1437,962,2031]" box="[199,1274,1898,1923]" pageId="36" pageNumber="37">Full morphometric variation is given in tables 13 and 14. Setae variation is given in tables 9–12.</paragraph>
</subSubSection>
<subSubSection id="C2946563FF801967FF463B9BFC5FF874" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8A3136E8FF801967FF463B9BFB74F83C" blockId="36.[151,1437,962,2031]" box="[199,1235,1934,1959]" pageId="36" pageNumber="37">
<emphasis id="B8FAEAFAFF801967FF463B9BFEC7F83C" bold="true" box="[199,352,1934,1959]" pageId="36" pageNumber="37">Distribution.</emphasis>
<collectingRegion id="484AF80AFF801967FEE63B9AFE19F83C" box="[359,446,1935,1959]" country="Mexico" name="Oaxaca" pageId="36" pageNumber="37">Oaxaca</collectingRegion>
, 
<collectingCountry id="F2997678FF801967FE483B9BFD85F83C" box="[457,546,1934,1959]" name="Mexico" pageId="36" pageNumber="37">México</collectingCountry>
. This species is only known from the 
<typeStatus id="5535884AFF801967FC483B9AFC5EF83C" box="[969,1017,1935,1959]" pageId="36" pageNumber="37">type</typeStatus>
locality (
<figureCitation id="12B52A6DFF801967FBE23B9BFB61F83C" box="[1123,1222,1934,1959]" captionStart="FIGURE 4" captionStartId="12.[151,250,1698,1720]" captionTargetBox="[162,1430,957,1668]" captionTargetId="figure@12.[150,1436,949,1676]" captionTargetPageId="12" captionText="FIGURE 4. Known distribution of all the species of Vaejovis, included in the “mexicanus” group." httpUri="https://zenodo.org/record/2659101/files/figure.png" pageId="36" pageNumber="37">Figure 4</figureCitation>
).
</paragraph>
<paragraph id="8A3136E8FF801967FF463BA7FC5FF874" blockId="36.[151,1437,962,2031]" pageId="36" pageNumber="37">
<emphasis id="B8FAEAFAFF801967FF463BA7FE2FF850" bold="true" box="[199,392,1970,1995]" pageId="36" pageNumber="37">Natural history.</emphasis>
This species inhabits in mixed oak-pine forest. Adult males and females have been found during the rainy season and were collected with black lights during the night.
</paragraph>
</subSubSection>
</treatment>
</document>