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<mods:title id="6E5796D8F6A0209F7793C91249D75B9B">Barbatula leoparda (Actinopterygii, Nemacheilidae), a new endemic species of stone loach of French Catalonia</mods:title>
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<heading id="D0B9F26AFF9AFFF6FC21FA0DFB10B462" bold="true" box="[994,1273,1523,1548]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="171" reason="2">
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<taxonomicName id="4C4E3E85FF9AFFF6FC21FA0DFB51B465" authorityName="Gauliard &amp; Dettai &amp; Persat &amp; Keith &amp; Denys" authorityYear="2019" box="[994,1208,1523,1547]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="171" phylum="Chordata" rank="species" species="leoparda" status="sp. nov.">
<emphasis id="B93A9914FF9AFFF6FC21FA0DFB51B465" bold="true" box="[994,1208,1523,1547]" italics="true" pageId="2" pageNumber="171">Barbatula leoparda</emphasis>
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<paragraph id="8BF14506FF9AFFF6FBD7F9EAFB21B442" blockId="2.[1044,1224,1556,1580]" box="[1044,1224,1556,1580]" pageId="2" pageNumber="171">
(
<figureCitation id="13755983FF9AFFF6FBDFF9EAFB9EB442" box="[1052,1143,1556,1580]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="3.[132,200,917,939]" captionStartId-1="3.[132,200,1572,1594]" captionStartId-2="4.[132,195,714,736]" captionTargetBox-0="[549,1455,215,1293]" captionTargetBox-1="[549,1456,1331,1860]" captionTargetBox-2="[132,775,215,697]" captionTargetId-0="figure-17@3.[549,1455,215,1293]" captionTargetId-1="figure-174@3.[549,1456,1331,1860]" captionTargetId-2="figure-352@4.[132,775,215,697]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. – Specimens of Barbatula spp. in lateral view. A: Holotype of Barbatula leoparda MNHN 2018-0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.; B: Paratype of Barbatula leoparda n. sp. MNHN 2016-0351, 46.5 mm SL ♀, Tech River at Céret, 9 Sep. 2016, Denys, Hautecoeur and ONEMA coll; C: Barbatula barbatula MNHN 2013-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; D: Holotype of Barbatula quignardi MGAB77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll." captionText-1="Figure 2. – Specimens of Barbatula spp. in ventral view. A: Holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefi- ach, 7 Sep. 2016, Denys and ONEMA coll.; B: Barbatula barbatula MNHN 2013-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; C: Holotype of Barbatula quignardi MGAB77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll." captionText-2="Figure 3. – Mouth of holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.Abbreviations: medial indentation of upper lip (mi), mental lobe of lower lip (ml). Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.12711144" figureDoi-1="http://doi.org/10.5281/zenodo.12711150" figureDoi-2="http://doi.org/10.5281/zenodo.12711152" httpUri-0="https://zenodo.org/record/12711144/files/figure.png" httpUri-1="https://zenodo.org/record/12711150/files/figure.png" httpUri-2="https://zenodo.org/record/12711152/files/figure.png" pageId="2" pageNumber="171">Figs 1-3</figureCitation>
; 
<tableCitation id="C6CC70BDFF9AFFF6FB42F9EAFB56B442" box="[1153,1215,1556,1580]" captionStart="Table I" captionStartId="4.[132,185,1100,1122]" captionText="Table I. – Morphometry of Barbatula leoparda, B. barbatula and B. quignardi for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). Values in parentheses: mean. Values of holotype are included in range. Bold mean values highlight morphometric differences (see Fig. 4)." pageId="2" pageNumber="171">Tab. I</tableCitation>
)
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<emphasis id="B93A9914FF9AFFF6FCEEF9ABFBEDB403" bold="true" box="[813,1028,1621,1645]" pageId="2" pageNumber="171">Material examined</emphasis>
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. –
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MNHN 2018-0228, 
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SL 
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at 
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, Denys and ONEMA coll.;
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. –
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MNHN 2016-0347, 4, 
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SL 
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SL 
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SL 
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SL 
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, collected with holotype; MNHN 2010-0997, 1, 
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: Dept. Pyrénées-Orientales: 
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at 
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, Denys and ONEMA coll.; MNHN erage of 218, minimum coverage 164)), genseq-1 complete 12S rDNA (GenBank accession MK518372, 950 bp, average coverage of 305, minimum coverage 93))
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.
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<caption id="DF31158EFF9BFFF7FF47FC6BFE8EB77D" ID-DOI="http://doi.org/10.5281/zenodo.12711144" ID-Zenodo-Dep="12711144" httpUri="https://zenodo.org/record/12711144/files/figure.png" pageId="3" pageNumber="172" startId="3.[132,200,917,939]" targetBox="[549,1455,215,1293]" targetPageId="3" targetType="figure">
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Figure 1. – Specimens of 
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<emphasis id="B93A9914FF9BFFF7FE56FC63FE17B1C5" box="[405,510,925,939]" italics="true" pageId="3" pageNumber="172">Barbatula</emphasis>
spp.
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in lateral view. 
<emphasis id="B93A9914FF9BFFF7FE9DFC53FE86B1AC" bold="true" box="[350,367,941,962]" pageId="3" pageNumber="172">A</emphasis>
: Holotype of 
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<emphasis id="B93A9914FF9BFFF7FF47FC33FED6B1B5" box="[132,319,973,987]" italics="true" pageId="3" pageNumber="172">Barbatula leoparda</emphasis>
MNHN 2018
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-0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.; 
<emphasis id="B93A9914FF9BFFF7FF47FBF3FF7DB64C" bold="true" box="[132,148,1037,1058]" pageId="3" pageNumber="172">B</emphasis>
: Paratype of 
<taxonomicName id="4C4E3E85FF9BFFF7FEDDFBEBFE0CB64D" authorityName="Gauliard &amp; Dettai &amp; Persat &amp; Keith &amp; Denys" authorityYear="2019" box="[286,485,1045,1059]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="leoparda" status="sp. nov.">
<emphasis id="B93A9914FF9BFFF7FEDDFBEBFE0CB64D" box="[286,485,1045,1059]" italics="true" pageId="3" pageNumber="172">Barbatula leoparda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209246FFF9BFFF7FE2FFBF3FF4BB655" pageId="3" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
MNHN 2016-0351, 46.5 mm SL ♀, Tech River at Céret, 9 Sep. 2016, Denys, Hautecoeur and ONEMA coll; 
<emphasis id="B93A9914FF9BFFF7FF79FB93FF22B6ED" bold="true" box="[186,203,1133,1155]" pageId="3" pageNumber="172">C</emphasis>
: 
<taxonomicName id="4C4E3E85FF9BFFF7FF19FB8BFF52B6F5" authority="MNHN 2013" authorityName="MNHN" authorityYear="2013" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9BFFF7FF19FB8BFE45B6ED" box="[218,428,1141,1155]" italics="true" pageId="3" pageNumber="172">Barbatula barbatula</emphasis>
MNHN 2013
</taxonomicName>
-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; 
<emphasis id="B93A9914FF9BFFF7FF47FB33FF7CB68C" bold="true" box="[132,149,1229,1250]" pageId="3" pageNumber="172">D</emphasis>
: Holotype of 
<taxonomicName id="4C4E3E85FF9BFFF7FEE9FB2BFF3DB694" authority="MGAB" authorityName="MGAB" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9BFFF7FEE9FB2BFE16B68D" box="[298,511,1237,1251]" italics="true" pageId="3" pageNumber="172">Barbatula quignardi</emphasis>
MGAB
</taxonomicName>
77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll.
</paragraph>
</caption>
<caption id="DF31158EFF9BFFF7FF47F9DAFE8EB52C" ID-DOI="http://doi.org/10.5281/zenodo.12711150" ID-Zenodo-Dep="12711150" httpUri="https://zenodo.org/record/12711150/files/figure.png" pageId="3" pageNumber="172" startId="3.[132,200,1572,1594]" targetBox="[549,1456,1331,1860]" targetPageId="3" targetType="figure">
<paragraph id="8BF14506FF9BFFF7FF47F9DAFE8EB52C" blockId="3.[132,512,1572,1858]" pageId="3" pageNumber="172">
Figure 2. – Specimens of 
<taxonomicName id="4C4E3E85FF9BFFF7FE56F9D2FF59B43F" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B93A9914FF9BFFF7FE56F9D2FE17B454" box="[405,510,1580,1594]" italics="true" pageId="3" pageNumber="172">Barbatula</emphasis>
spp.
</taxonomicName>
in ventral view. 
<emphasis id="B93A9914FF9BFFF7FEA2F9C5FE9BB43E" bold="true" box="[353,370,1595,1616]" pageId="3" pageNumber="172">A</emphasis>
: Holotype of 
<taxonomicName id="4C4E3E85FF9BFFF7FF47F9A2FE1EB407" authority="MNHN 2018" authorityName="MNHN" authorityYear="2018" box="[132,503,1620,1642]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9BFFF7FF47F9A2FEB6B404" box="[132,351,1628,1642]" italics="true" pageId="3" pageNumber="172">Barbatula leoparda</emphasis>
MNHN 2018
</taxonomicName>
- 0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.; 
<emphasis id="B93A9914FF9BFFF7FF7CF962FF26B4DF" bold="true" box="[191,207,1692,1713]" pageId="3" pageNumber="172">B</emphasis>
: 
<taxonomicName id="4C4E3E85FF9BFFF7FF1EF95AFF52B4A4" authority="MNHN 2013" authorityName="MNHN" authorityYear="2013" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9BFFF7FF1EF95AFE44B4DC" box="[221,429,1700,1714]" italics="true" pageId="3" pageNumber="172">Barbatula barbatula</emphasis>
MNHN 2013
</taxonomicName>
-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; 
<emphasis id="B93A9914FF9BFFF7FF47F905FF7CB57F" bold="true" box="[132,149,1787,1809]" pageId="3" pageNumber="172">C</emphasis>
: Holotype of 
<taxonomicName id="4C4E3E85FF9BFFF7FEE9F8FAFF3DB547" authority="MGAB" authorityName="MGAB" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9BFFF7FEE9F8FAFE16B57C" box="[298,511,1796,1810]" italics="true" pageId="3" pageNumber="172">Barbatula quignardi</emphasis>
MGAB
</taxonomicName>
77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll.
</paragraph>
</caption>
<caption id="DF31158EFF9CFFF0FF47FD34FDB8B146" ID-DOI="http://doi.org/10.5281/zenodo.12711152" ID-Zenodo-Dep="12711152" httpUri="https://zenodo.org/record/12711152/files/figure.png" pageId="4" pageNumber="173" startId="4.[132,195,714,736]" targetBox="[132,775,215,697]" targetPageId="4" targetType="figure">
<paragraph id="8BF14506FF9CFFF0FF47FD34FDB8B146" blockId="4.[132,775,714,808]" pageId="4" pageNumber="173">
Figure 3. – Mouth of holotype of 
<taxonomicName id="4C4E3E85FF9CFFF0FE03FD2DFCE8B0B1" authority="MNHN 2018" authorityName="MNHN" authorityYear="2018" box="[448,769,714,737]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9CFFF0FE03FD2DFD95B08F" box="[448,636,723,737]" italics="true" pageId="4" pageNumber="173">Barbatula leoparda</emphasis>
MNHN 2018
</taxonomicName>
- 0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll. Abbreviations: medial indentation of upper lip (mi), mental lobe of lower lip (ml). Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BF14506FF9CFFF0FF47FCB4FD12B1EA" blockId="4.[132,774,842,900]" pageId="4" pageNumber="173">
<materialsCitation id="3B264F5BFF9CFFF0FF47FCB4FD1EB1EA" collectingDate="2016-09-09" latitude="42.496" location="Tech River" longLatPrecision="69" longitude="2.755" municipality="Ceret" pageId="4" pageNumber="173" specimenCount="1" specimenCount-female="1">
2016-0351, 1, 
<quantity id="4CB6E8E3FF9CFFF0FEDCFCB4FE94B10C" box="[287,381,842,866]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" pageId="4" pageNumber="173" unit="mm" value="46.5">46.5 mm</quantity>
SL 
<specimenCount id="9D488E8FFF9CFFF0FE6BFCB5FE55B10C" box="[424,444,843,866]" count="1" pageId="4" pageNumber="173" type="female">♀</specimenCount>
, 
<location id="8E9113DDFF9CFFF0FE0BFCB4FDD4B10C" LSID="urn:lsid:plazi:treatment:03E7F410FF9AFFF1FC21FA0DFE90B4BE:8E9113DDFF9CFFF0FE0BFCB4FDD4B10C" box="[456,573,842,866]" latitude="42.496" longLatPrecision="69" longitude="2.755" municipality="Ceret" name="Tech River" pageId="4" pageNumber="173">Tech River</location>
at 
<collectingMunicipality id="6B95DF7CFF9CFFF0FD9EFCB4FD72B10C" box="[605,667,842,866]" pageId="4" pageNumber="173">Céret</collectingMunicipality>
: 
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<geoCoordinate id="EE7A23C1FF9CFFF0FF47FC92FF33B1EA" box="[132,218,876,900]" degrees="2.755" direction="east" orientation="longitude" pageId="4" pageNumber="173" precision="55" value="2.755">E 2.755</geoCoordinate>
, 
<date id="FFF063C6FF9CFFF0FF20FC92FE8AB1EA" box="[227,355,876,900]" pageId="4" pageNumber="173" value="2016-09-09">
<collectingDate id="EFB49A2EFF9CFFF0FF20FC92FE8AB1EA" box="[227,355,876,900]" pageId="4" pageNumber="173" value="2016-09-09">9 Sep. 2016</collectingDate>
</date>
, Denys, Hautecoeur and ONEMA coll
</materialsCitation>
.
</paragraph>
<paragraph id="8BF14506FF9CFFF0FF47FC53FDE8B1AB" blockId="4.[132,775,941,1067]" box="[132,513,941,966]" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FF47FC53FDE8B1AB" bold="true" box="[132,513,941,966]" pageId="4" pageNumber="173">Material for molecular references</emphasis>
</paragraph>
<paragraph id="8BF14506FF9CFFF0FF69FC27FCEEB644" blockId="4.[132,775,941,1067]" pageId="4" pageNumber="173">
<materialsCitation id="3B264F5BFF9CFFF0FF69FC27FCEEB644" country="France" latitude="42.697" location="Tet River" longLatPrecision="68" longitude="2.663" municipality="Nefiach" pageId="4" pageNumber="173" specimenCount="1" typeStatus="holotype">
<emphasis id="B93A9914FF9CFFF0FF69FC27FECFB186" box="[170,294,985,1000]" italics="true" pageId="4" pageNumber="173">
<typeStatus id="54F5FBA4FF9CFFF0FF69FC27FEE6B186" box="[170,271,985,1000]" pageId="4" pageNumber="173" type="holotype">Holotype</typeStatus>
. –
</emphasis>
MNHN 2018-0228, 
<collectingCountry id="F3590596FF9CFFF0FDC5FC2EFDBAB189" box="[518,595,976,999]" name="France" pageId="4" pageNumber="173">France</collectingCountry>
: Dept. Pyrénées-Orientales: 
<location id="8E9113DDFF9CFFF0FF3DFC0FFE8BB667" LSID="urn:lsid:plazi:treatment:03E7F410FF9AFFF1FC21FA0DFE90B4BE:8E9113DDFF9CFFF0FF3DFC0FFE8BB667" box="[254,354,1009,1033]" country="France" latitude="42.697" longLatPrecision="68" longitude="2.663" municipality="Nefiach" name="Tet River" pageId="4" pageNumber="173">Têt River</location>
at 
<collectingMunicipality id="6B95DF7CFF9CFFF0FE42FC0FFE31B667" box="[385,472,1009,1033]" pageId="4" pageNumber="173">Nefiach</collectingMunicipality>
: 
<geoCoordinate id="EE7A23C1FF9CFFF0FE22FC0FFDAAB667" box="[481,579,1009,1033]" degrees="42.697" direction="north" orientation="latitude" pageId="4" pageNumber="173" precision="55" value="42.697">N 42.697</geoCoordinate>
<geoCoordinate id="EE7A23C1FF9CFFF0FD8AFC0FFD77B667" box="[585,670,1009,1033]" degrees="2.663" direction="east" orientation="longitude" pageId="4" pageNumber="173" precision="55" value="2.663">E 2.663</geoCoordinate>
, genseq-1 COI (GenBank accession MK518369, 650 bp, average cov-
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C354168DFF9CFFF1FCEEFE9EFE90B4BE" lastPageId="5" lastPageNumber="174" pageId="4" pageNumber="173" type="diagnosis">
<paragraph id="8BF14506FF9CFFF0FCEEFE9EFC72B316" blockId="4.[813,1456,352,1066]" box="[813,923,352,376]" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FCEEFE9EFC72B316" bold="true" box="[813,923,352,376]" pageId="4" pageNumber="173">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF14506FF9CFFF0FC91FE73FAAFB157" blockId="4.[813,1456,352,1066]" pageId="4" pageNumber="173">
<taxonomicName id="4C4E3E85FF9CFFF0FC91FE73FBCDB3F2" authorityName="MNHN" authorityYear="2018" box="[850,1060,397,412]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9CFFF0FC91FE73FBCDB3F2" box="[850,1060,397,412]" italics="true" pageId="4" pageNumber="173">Barbatula leoparda</emphasis>
</taxonomicName>
larger than 
<quantity id="4CB6E8E3FF9CFFF0FB60FE7DFB04B3F5" box="[1187,1261,387,411]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="173" unit="mm" value="47.0">47 mm</quantity>
SL is distinguishable from 
<taxonomicName id="4C4E3E85FF9CFFF0FC60FE51FBDDB3D0" authorityName="MNHN" authorityYear="2013" box="[931,1076,431,446]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9CFFF0FC60FE51FBDDB3D0" box="[931,1076,431,446]" italics="true" pageId="4" pageNumber="173">B. barbatula</emphasis>
</taxonomicName>
of the same size by the presence of dark-brown blotches on the belly and the jugular area (
<emphasis id="B93A9914FF9CFFF0FCF5FE0AFCBDB06D" box="[822,852,500,515]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
absence of pigmentation) (
<figureCitation id="13755983FF9CFFF0FB42FE15FB2CB06D" box="[1153,1221,491,515]" captionStart="Figure 2" captionStartId="3.[132,200,1572,1594]" captionTargetBox="[549,1456,1331,1860]" captionTargetId="figure-174@3.[549,1456,1331,1860]" captionTargetPageId="3" captionText="Figure 2. – Specimens of Barbatula spp. in ventral view. A: Holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefi- ach, 7 Sep. 2016, Denys and ONEMA coll.; B: Barbatula barbatula MNHN 2013-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; C: Holotype of Barbatula quignardi MGAB77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll." figureDoi="http://doi.org/10.5281/zenodo.12711150" httpUri="https://zenodo.org/record/12711150/files/figure.png" pageId="4" pageNumber="173">Fig. 2</figureCitation>
), a larger interorbital width (35.6 to 41.8% HL, 
<emphasis id="B93A9914FF9CFFF0FB8BFDE9FB8CB048" box="[1096,1125,535,550]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
23.0 to 38.1% HL; 
<figureCitation id="13755983FF9CFFF0FAF8FDF3FA64B04B" box="[1339,1421,525,549]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4A</figureCitation>
), a deeper caudal peduncle (12.1 to 13.0% SL, 
<emphasis id="B93A9914FF9CFFF0FAC1FDC7FAC9B026" box="[1282,1312,569,584]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
8.0 to 12.4% SL; 
<figureCitation id="13755983FF9CFFF0FC9FFDACFC5AB004" box="[860,947,594,618]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4C</figureCitation>
), and a fewer ratio length/depth of the caudal peduncle (1.30 to 1.48, 
<emphasis id="B93A9914FF9CFFF0FBE8FD80FBA0B0E3" box="[1067,1097,638,653]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
1.38 to 2.56; 
<figureCitation id="13755983FF9CFFF0FB1DFD8BFAD9B0E2" box="[1246,1328,629,653]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4D</figureCitation>
). Sub-adult specimens (
<quantity id="4CB6E8E3FF9CFFF0FC71FD69FBAAB0C1" box="[946,1091,663,687]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.7" metricValueMin="3.6" pageId="4" pageNumber="173" unit="mm" value="41.5" valueMax="47.0" valueMin="36.0">36 to 47 mm</quantity>
SL) are diagnosable by a larger interorbital width (35.5 to 40.5% HL, 
<emphasis id="B93A9914FF9CFFF0FB07FD3DFB0BB0BC" box="[1220,1250,707,722]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
28.7 to 38.4% HL; 
<figureCitation id="13755983FF9CFFF0FCEEFD22FC68B09A" box="[813,897,732,756]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4B</figureCitation>
), a fewer ratio length/depth of the caudal peduncle (1.41 to 1.45, 
<emphasis id="B93A9914FF9CFFF0FC03FCF6FC34B179" box="[960,989,776,791]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
1.45 to 2.15; 
<figureCitation id="13755983FF9CFFF0FBADFD00FB54B179" box="[1134,1213,766,791]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4E</figureCitation>
) and longer pelvic fins (15.8 to 16.5% SL, 
<emphasis id="B93A9914FF9CFFF0FC3FFCD4FBF0B157" box="[1020,1049,810,825]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
12.4 to 15.5% SL; 
<figureCitation id="13755983FF9CFFF0FB26FCDFFAD3B157" box="[1253,1338,801,825]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4G</figureCitation>
).
</paragraph>
<paragraph id="8BF14506FF9CFFF1FC91FCB3FEE1B1AF" blockId="4.[813,1456,352,1066]" lastBlockId="5.[132,776,937,1745]" lastPageId="5" lastPageNumber="174" pageId="4" pageNumber="173">
<taxonomicName id="4C4E3E85FF9CFFF0FC91FCB3FBCDB132" authorityName="MNHN" authorityYear="2018" box="[850,1060,845,860]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9CFFF0FC91FCB3FBCDB132" box="[850,1060,845,860]" italics="true" pageId="4" pageNumber="173">Barbatula leoparda</emphasis>
</taxonomicName>
larger than 
<quantity id="4CB6E8E3FF9CFFF0FB60FCBAFB04B135" box="[1187,1261,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="4" pageNumber="173" unit="mm" value="47.0">47 mm</quantity>
SL is distinguishable from 
<taxonomicName id="4C4E3E85FF9CFFF0FC60FC91FBDAB110" box="[931,1075,879,894]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9CFFF0FC60FC91FBDAB110" box="[931,1075,879,894]" italics="true" pageId="4" pageNumber="173">B. quignardi</emphasis>
</taxonomicName>
of the same size by the presence of dark-brown blotches on the belly and the jugular area (
<emphasis id="B93A9914FF9CFFF0FCF5FC4AFCBDB1AD" box="[822,852,948,963]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
absence of pigmentation) (
<figureCitation id="13755983FF9CFFF0FB42FC55FB2CB1AD" box="[1153,1221,939,963]" captionStart="Figure 2" captionStartId="3.[132,200,1572,1594]" captionTargetBox="[549,1456,1331,1860]" captionTargetId="figure-174@3.[549,1456,1331,1860]" captionTargetPageId="3" captionText="Figure 2. – Specimens of Barbatula spp. in ventral view. A: Holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefi- ach, 7 Sep. 2016, Denys and ONEMA coll.; B: Barbatula barbatula MNHN 2013-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; C: Holotype of Barbatula quignardi MGAB77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll." figureDoi="http://doi.org/10.5281/zenodo.12711150" httpUri="https://zenodo.org/record/12711150/files/figure.png" pageId="4" pageNumber="173">Fig. 2</figureCitation>
), a larger interorbital width (35.6 to 41.8% HL, 
<emphasis id="B93A9914FF9CFFF0FB97FC29FB9BB188" box="[1108,1138,983,998]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
21.6 to 36.0% HL; 
<figureCitation id="13755983FF9CFFF0FA92FC33FA4CB188" box="[1361,1445,973,998]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4A</figureCitation>
) and a shorter dorsal-fin length (17.1 to 20.1% SL, 
<emphasis id="B93A9914FF9CFFF0FA99FC07FA91B666" box="[1370,1400,1017,1032]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
18.9 to 23.9% HL; 
<figureCitation id="13755983FF9CFFF0FC05FBECFBFEB644" box="[966,1047,1042,1066]" captionStart="Figure 4" captionStartId="5.[132,197,736,758]" captionTargetBox="[133,1455,217,718]" captionTargetId="figure-354@5.[132,1455,215,719]" captionTargetPageId="5" captionText="Figure 4. – Boxplots highlighting morphometric differences between Barbatula leoparda (in grey) and the two other species Barbatula barbatula and Barbatula quignardi, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). only by a larger interorbital width (35.5 to 40.5% HL, vs. 21.4 to 35.6% HL; Fig. 4B)." figureDoi="http://doi.org/10.5281/zenodo.12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="4" pageNumber="173">Fig. 4F</figureCitation>
). Subadult specimens are diagnosable 
<emphasis id="B93A9914FF9DFFF1FF47FC57FEE1B1AF" bold="true" box="[132,264,937,961]" pageId="5" pageNumber="174">Description</emphasis>
</paragraph>
<caption id="DF31158EFF9CFFF0FF47FBB2FA4AB617" pageId="4" pageNumber="173" startId="4.[132,185,1100,1122]" targetType="table">
<paragraph id="8BF14506FF9CFFF0FF47FBB2FA4AB617" blockId="4.[132,1455,1100,1146]" pageId="4" pageNumber="173">
Table I. – Morphometry of 
<taxonomicName id="4C4E3E85FF9CFFF0FE44FBAAFDADB60C" authorityName="MNHN" authorityYear="2018" box="[391,580,1108,1122]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9CFFF0FE44FBAAFDADB60C" box="[391,580,1108,1122]" italics="true" pageId="4" pageNumber="173">Barbatula leoparda</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4E3E85FF9CFFF0FD8CFBAAFD21B60C" authorityName="MNHN" authorityYear="2013" box="[591,712,1108,1122]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9CFFF0FD8CFBAAFD21B60C" box="[591,712,1108,1122]" italics="true" pageId="4" pageNumber="173">B. barbatula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E3E85FF9CFFF0FD36FBAAFC84B60C" box="[757,877,1108,1122]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9CFFF0FD36FBAAFC84B60C" box="[757,877,1108,1122]" italics="true" pageId="4" pageNumber="173">B. quignardi</emphasis>
</taxonomicName>
for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL). Values in parentheses: mean. Values of holotype are included in range. Bold mean values highlight morphometric differences (see Fig. 4).
</paragraph>
</caption>
<paragraph id="8BF14506FF9CFFF0FE34FB6EFA4FB507" pageId="4" pageNumber="173">
<table id="F94EB7A6FF9C000BFF4DFB77FA4FB507" box="[142,1446,1161,1897]" gridcols="7" gridrows="24" pageId="4" pageNumber="173">
<tr id="357E4744FF9C000BFF4DFB77FA4FB6F2" box="[142,1446,1161,1180]" gridrow="0" pageId="4" pageNumber="173" rowspan-0="1">
<th id="76AF2E38FF9C000BFE5CFB77FCC1B6F2" box="[415,808,1161,1180]" colspan="2" colspanRight="1" gridcol="1" gridrow="0" pageId="4" pageNumber="173">
<taxonomicName id="4C4E3E85FF9CFFF0FE34FB6EFD76B6F2" authorityName="Gauliard &amp; Dettai &amp; Persat &amp; Keith &amp; Denys" authorityYear="2019" box="[503,671,1168,1180]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="leoparda" status="sp. nov.">
<emphasis id="B93A9914FF9CFFF0FE34FB6EFD76B6F2" box="[503,671,1168,1180]" italics="true" pageId="4" pageNumber="173">Barbatula leoparda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209246FFF9CFFF0FD66FB77FD3DB6F2" box="[677,724,1161,1180]" pageId="4" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</th>
<th id="76AF2E38FF9C000BFCFCFB77FB81B6F2" box="[831,1128,1161,1180]" colspan="2" colspanRight="1" gridcol="3" gridrow="0" pageId="4" pageNumber="173">
<taxonomicName id="4C4E3E85FF9CFFF0FCBFFB6EFBC2B6F2" authorityName="MNHN" authorityYear="2013" box="[892,1067,1168,1180]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9CFFF0FCBFFB6EFBC2B6F2" box="[892,1067,1168,1180]" italics="true" pageId="4" pageNumber="173">Barbatula barbatula</emphasis>
</taxonomicName>
</th>
<th id="76AF2E38FF9C000BFBBFFB77FA4FB6F2" box="[1148,1446,1161,1180]" colspan="2" colspanRight="1" gridcol="5" gridrow="0" pageId="4" pageNumber="173">
<taxonomicName id="4C4E3E85FF9CFFF0FB7AFB6EFA81B6F2" box="[1209,1384,1168,1180]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9CFFF0FB7AFB6EFA81B6F2" box="[1209,1384,1168,1180]" italics="true" pageId="4" pageNumber="173">Barbatula quignardi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="357E4744FF9C000BFF4DFB56FA4FB6D2" box="[142,1446,1192,1212]" gridrow="1" pageId="4" pageNumber="173" rowspan-0="1">
<td id="76AF2E38FF9C000BFE5CFB56FDC3B6D2" box="[415,554,1192,1212]" gridcol="1" gridrow="1" pageId="4" pageNumber="173">36-47 mm SL</td>
<td id="76AF2E38FF9C000BFDFDFB56FCC1B6D2" box="[574,808,1192,1212]" gridcol="2" gridrow="1" pageId="4" pageNumber="173">&gt; 47 mm SL Holoype</td>
<td id="76AF2E38FF9C000BFCFCFB56FC23B6D2" box="[831,970,1192,1212]" gridcol="3" gridrow="1" pageId="4" pageNumber="173">36-47 mm SL</td>
<td id="76AF2E38FF9C000BFC1DFB56FB81B6D2" box="[990,1128,1192,1212]" gridcol="4" gridrow="1" pageId="4" pageNumber="173">&gt; 47 mm SL</td>
<td id="76AF2E38FF9C000BFBBFFB56FAEEB6D2" box="[1148,1287,1192,1212]" gridcol="5" gridrow="1" pageId="4" pageNumber="173">36-47 mm SL</td>
<td id="76AF2E38FF9C000BFAD8FB56FA4FB6D2" box="[1307,1446,1192,1212]" gridcol="6" gridrow="1" pageId="4" pageNumber="173">&gt; 47 mm SL</td>
</tr>
<tr id="357E4744FF9C000BFF4DFB39FA4FB6B5" box="[142,1446,1223,1243]" gridrow="2" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFB39FE6DB6B5" box="[142,388,1223,1243]" gridcol="0" gridrow="2" pageId="4" pageNumber="173">N</th>
<td id="76AF2E38FF9C000BFE5CFB39FDC3B6B5" box="[415,554,1223,1243]" gridcol="1" gridrow="2" pageId="4" pageNumber="173">3</td>
<td id="76AF2E38FF9C000BFDFDFB39FCC1B6B5" box="[574,808,1223,1243]" gridcol="2" gridrow="2" pageId="4" pageNumber="173">3</td>
<td id="76AF2E38FF9C000BFCFCFB39FC23B6B5" box="[831,970,1223,1243]" gridcol="3" gridrow="2" pageId="4" pageNumber="173">8</td>
<td id="76AF2E38FF9C000BFC1DFB39FB81B6B5" box="[990,1128,1223,1243]" gridcol="4" gridrow="2" pageId="4" pageNumber="173">37</td>
<td id="76AF2E38FF9C000BFBBFFB39FAEEB6B5" box="[1148,1287,1223,1243]" gridcol="5" gridrow="2" pageId="4" pageNumber="173">15</td>
<td id="76AF2E38FF9C000BFAD8FB39FA4FB6B5" box="[1307,1446,1223,1243]" gridcol="6" gridrow="2" pageId="4" pageNumber="173">52</td>
</tr>
<tr id="357E4744FF9C000BFF4DFB18FA4FB695" box="[142,1446,1254,1275]" gridrow="3" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFB18FE6DB695" box="[142,388,1254,1275]" gridcol="0" gridrow="3" pageId="4" pageNumber="173">Standard length SL (mm)</th>
<td id="76AF2E38FF9C000BFE5CFB18FDC3B695" box="[415,554,1254,1275]" gridcol="1" gridrow="3" pageId="4" pageNumber="173">36.6-46.5 (42.6)</td>
<td id="76AF2E38FF9C000BFDFDFB18FCC1B695" box="[574,808,1254,1275]" gridcol="2" gridrow="3" pageId="4" pageNumber="173">48.2-62.6 (57.3) 62.6</td>
<td id="76AF2E38FF9C000BFCFCFB18FC23B695" box="[831,970,1254,1275]" gridcol="3" gridrow="3" pageId="4" pageNumber="173">37.8-46.9 (41.5)</td>
<td id="76AF2E38FF9C000BFC1DFB18FB81B695" box="[990,1128,1254,1275]" gridcol="4" gridrow="3" pageId="4" pageNumber="173">47.1-81.5 (63.6)</td>
<td id="76AF2E38FF9C000BFBBFFB18FAEEB695" box="[1148,1287,1254,1275]" gridcol="5" gridrow="3" pageId="4" pageNumber="173">36.4-46.8 (42.4)</td>
<td id="76AF2E38FF9C000BFAD8FB18FA4FB695" box="[1307,1446,1254,1275]" gridcol="6" gridrow="3" pageId="4" pageNumber="173">47.8-77.6 (59.8)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFAF3FA4FB777" box="[142,1446,1293,1305]" gridrow="4" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFAF3FA4FB777" box="[142,1446,1293,1305]" colspan="7" colspanRight="6" gridcol="0" gridrow="4" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FF4DFAF3FEFEB777" box="[142,279,1293,1305]" italics="true" pageId="4" pageNumber="173">In percent of SL</emphasis>
</th>
</tr>
<tr id="357E4744FF9C000BFF4DFADBFA4FB757" box="[142,1446,1317,1337]" gridrow="5" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFADBFE6DB757" box="[142,388,1317,1337]" gridcol="0" gridrow="5" pageId="4" pageNumber="173">Head length HL</th>
<td id="76AF2E38FF9C000BFE5CFADBFDC3B757" box="[415,554,1317,1337]" gridcol="1" gridrow="5" pageId="4" pageNumber="173">22.8-23.4 (23.1)</td>
<td id="76AF2E38FF9C000BFDFDFADBFCC1B757" box="[574,808,1317,1337]" gridcol="2" gridrow="5" pageId="4" pageNumber="173">22.1-23.4 (22.7) 22.1</td>
<td id="76AF2E38FF9C000BFCFCFADBFC23B757" box="[831,970,1317,1337]" gridcol="3" gridrow="5" pageId="4" pageNumber="173">21.7-25.0 (23.8)</td>
<td id="76AF2E38FF9C000BFC1DFADBFB81B757" box="[990,1128,1317,1337]" gridcol="4" gridrow="5" pageId="4" pageNumber="173">20.6-24.9 (22.7)</td>
<td id="76AF2E38FF9C000BFBBFFADBFAEEB757" box="[1148,1287,1317,1337]" gridcol="5" gridrow="5" pageId="4" pageNumber="173">22.4-26.7 (24.5)</td>
<td id="76AF2E38FF9C000BFAD8FADBFA4FB757" box="[1307,1446,1317,1337]" gridcol="6" gridrow="5" pageId="4" pageNumber="173">19.8-26.5 (23.2)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFABAFA4FB736" box="[142,1446,1348,1368]" gridrow="6" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFABAFE6DB736" box="[142,388,1348,1368]" gridcol="0" gridrow="6" pageId="4" pageNumber="173">Predorsal length</th>
<td id="76AF2E38FF9C000BFE5CFABAFDC3B736" box="[415,554,1348,1368]" gridcol="1" gridrow="6" pageId="4" pageNumber="173">50.4-53.7 (52.3)</td>
<td id="76AF2E38FF9C000BFDFDFABAFCC1B736" box="[574,808,1348,1368]" gridcol="2" gridrow="6" pageId="4" pageNumber="173">50.5-53.8 (52.2) 52.3</td>
<td id="76AF2E38FF9C000BFCFCFABAFC23B736" box="[831,970,1348,1368]" gridcol="3" gridrow="6" pageId="4" pageNumber="173">51.4-54.4 (53.3)</td>
<td id="76AF2E38FF9C000BFC1DFABAFB81B736" box="[990,1128,1348,1368]" gridcol="4" gridrow="6" pageId="4" pageNumber="173">50.6-55.8 (52.7)</td>
<td id="76AF2E38FF9C000BFBBFFABAFAEEB736" box="[1148,1287,1348,1368]" gridcol="5" gridrow="6" pageId="4" pageNumber="173">49.5-56.4 (52.9)</td>
<td id="76AF2E38FF9C000BFAD8FABAFA4FB736" box="[1307,1446,1348,1368]" gridcol="6" gridrow="6" pageId="4" pageNumber="173">48.7-56.0 (52.5)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA9DFA4FB719" box="[142,1446,1379,1399]" gridrow="7" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA9DFE6DB719" box="[142,388,1379,1399]" gridcol="0" gridrow="7" pageId="4" pageNumber="173">Postdorsal length</th>
<td id="76AF2E38FF9C000BFE5CFA9DFDC3B719" box="[415,554,1379,1399]" gridcol="1" gridrow="7" pageId="4" pageNumber="173">31.7-37.7 (35.6)</td>
<td id="76AF2E38FF9C000BFDFDFA9DFCC1B719" box="[574,808,1379,1399]" gridcol="2" gridrow="7" pageId="4" pageNumber="173">34.1-38.6 (35.6) 34.1</td>
<td id="76AF2E38FF9C000BFCFCFA9DFC23B719" box="[831,970,1379,1399]" gridcol="3" gridrow="7" pageId="4" pageNumber="173">33.0-38.1 (36.5)</td>
<td id="76AF2E38FF9C000BFC1DFA9DFB81B719" box="[990,1128,1379,1399]" gridcol="4" gridrow="7" pageId="4" pageNumber="173">33.9-42.8 (38.1)</td>
<td id="76AF2E38FF9C000BFBBFFA9DFAEEB719" box="[1148,1287,1379,1399]" gridcol="5" gridrow="7" pageId="4" pageNumber="173">32.8-39.4 (36.1)</td>
<td id="76AF2E38FF9C000BFAD8FA9DFA4FB719" box="[1307,1446,1379,1399]" gridcol="6" gridrow="7" pageId="4" pageNumber="173">32.7-39.6 (36.5)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA7CFA4FB7F9" box="[142,1446,1410,1431]" gridrow="8" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA7CFE6DB7F9" box="[142,388,1410,1431]" gridcol="0" gridrow="8" pageId="4" pageNumber="173">Prepectoral length</th>
<td id="76AF2E38FF9C000BFE5CFA7CFDC3B7F9" box="[415,554,1410,1431]" gridcol="1" gridrow="8" pageId="4" pageNumber="173">22.7-26.4 (24.4)</td>
<td id="76AF2E38FF9C000BFDFDFA7CFCC1B7F9" box="[574,808,1410,1431]" gridcol="2" gridrow="8" pageId="4" pageNumber="173">22.8-24.8 (23.6) 22.8</td>
<td id="76AF2E38FF9C000BFCFCFA7CFC23B7F9" box="[831,970,1410,1431]" gridcol="3" gridrow="8" pageId="4" pageNumber="173">23.7-28.0 (25.7)</td>
<td id="76AF2E38FF9C000BFC1DFA7CFB81B7F9" box="[990,1128,1410,1431]" gridcol="4" gridrow="8" pageId="4" pageNumber="173">19.7-27.6 (23.3)</td>
<td id="76AF2E38FF9C000BFBBFFA7CFAEEB7F9" box="[1148,1287,1410,1431]" gridcol="5" gridrow="8" pageId="4" pageNumber="173">21.1-29.1 (24.9)</td>
<td id="76AF2E38FF9C000BFAD8FA7CFA4FB7F9" box="[1307,1446,1410,1431]" gridcol="6" gridrow="8" pageId="4" pageNumber="173">20.4-26.4 (23.8)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA5FFA4FB7D8" box="[142,1446,1441,1462]" gridrow="9" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA5FFE6DB7D8" box="[142,388,1441,1462]" gridcol="0" gridrow="9" pageId="4" pageNumber="173">Prepelvic length</th>
<td id="76AF2E38FF9C000BFE5CFA5FFDC3B7D8" box="[415,554,1441,1462]" gridcol="1" gridrow="9" pageId="4" pageNumber="173">53.4-56.2 (55.2)</td>
<td id="76AF2E38FF9C000BFDFDFA5FFCC1B7D8" box="[574,808,1441,1462]" gridcol="2" gridrow="9" pageId="4" pageNumber="173">51.1-56.1 (53.7) 56.1</td>
<td id="76AF2E38FF9C000BFCFCFA5FFC23B7D8" box="[831,970,1441,1462]" gridcol="3" gridrow="9" pageId="4" pageNumber="173">51.9-58.0 (54.9)</td>
<td id="76AF2E38FF9C000BFC1DFA5FFB81B7D8" box="[990,1128,1441,1462]" gridcol="4" gridrow="9" pageId="4" pageNumber="173">49.1-61.6 (53.5)</td>
<td id="76AF2E38FF9C000BFBBFFA5FFAEEB7D8" box="[1148,1287,1441,1462]" gridcol="5" gridrow="9" pageId="4" pageNumber="173">51.9-57.8 (54.3)</td>
<td id="76AF2E38FF9C000BFAD8FA5FFA4FB7D8" box="[1307,1446,1441,1462]" gridcol="6" gridrow="9" pageId="4" pageNumber="173">50.9-56.7 (53.9)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA3FFA4FB7BB" box="[142,1446,1473,1493]" gridrow="10" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA3FFE6DB7BB" box="[142,388,1473,1493]" gridcol="0" gridrow="10" pageId="4" pageNumber="173">Preanal length</th>
<td id="76AF2E38FF9C000BFE5CFA3FFDC3B7BB" box="[415,554,1473,1493]" gridcol="1" gridrow="10" pageId="4" pageNumber="173">76.3-80.3 (78.5)</td>
<td id="76AF2E38FF9C000BFDFDFA3FFCC1B7BB" box="[574,808,1473,1493]" gridcol="2" gridrow="10" pageId="4" pageNumber="173">74.0-79.5 (76.7) 76.8</td>
<td id="76AF2E38FF9C000BFCFCFA3FFC23B7BB" box="[831,970,1473,1493]" gridcol="3" gridrow="10" pageId="4" pageNumber="173">74.1-79.6 (76.7)</td>
<td id="76AF2E38FF9C000BFC1DFA3FFB81B7BB" box="[990,1128,1473,1493]" gridcol="4" gridrow="10" pageId="4" pageNumber="173">74.1-80.1 (76.1)</td>
<td id="76AF2E38FF9C000BFBBFFA3FFAEEB7BB" box="[1148,1287,1473,1493]" gridcol="5" gridrow="10" pageId="4" pageNumber="173">75.4-83.1 (77.8)</td>
<td id="76AF2E38FF9C000BFAD8FA3FFA4FB7BB" box="[1307,1446,1473,1493]" gridcol="6" gridrow="10" pageId="4" pageNumber="173">74.7-79.6 (77.1)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA1EFA4FB79A" box="[142,1446,1504,1524]" gridrow="11" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA1EFE6DB79A" box="[142,388,1504,1524]" gridcol="0" gridrow="11" pageId="4" pageNumber="173">Caudal peduncle length CPL</th>
<td id="76AF2E38FF9C000BFE5CFA1EFDC3B79A" box="[415,554,1504,1524]" gridcol="1" gridrow="11" pageId="4" pageNumber="173">16.6-17.9 (17.2)</td>
<td id="76AF2E38FF9C000BFDFDFA1EFCC1B79A" box="[574,808,1504,1524]" gridcol="2" gridrow="11" pageId="4" pageNumber="173">17.0-17.8 (17.3) 17.0</td>
<td id="76AF2E38FF9C000BFCFCFA1EFC23B79A" box="[831,970,1504,1524]" gridcol="3" gridrow="11" pageId="4" pageNumber="173">15.0-20.2 (17.5)</td>
<td id="76AF2E38FF9C000BFC1DFA1EFB81B79A" box="[990,1128,1504,1524]" gridcol="4" gridrow="11" pageId="4" pageNumber="173">15.0-20.6 (17.7)</td>
<td id="76AF2E38FF9C000BFBBFFA1EFAEEB79A" box="[1148,1287,1504,1524]" gridcol="5" gridrow="11" pageId="4" pageNumber="173">13.1-20.4 (16.7)</td>
<td id="76AF2E38FF9C000BFAD8FA1EFA4FB79A" box="[1307,1446,1504,1524]" gridcol="6" gridrow="11" pageId="4" pageNumber="173">14.2-19.7 (17.3)</td>
</tr>
<tr id="357E4744FF9C000BFF4DFA01FA4FB47C" box="[142,1446,1535,1554]" gridrow="12" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DFA01FE6DB47C" box="[142,388,1535,1554]" gridcol="0" gridrow="12" pageId="4" pageNumber="173">Caudal peduncle depth CPD</th>
<td id="76AF2E38FF9C000BFE5CFA01FDC3B47C" box="[415,554,1535,1554]" gridcol="1" gridrow="12" pageId="4" pageNumber="173">11.7-12.3 (12.1)</td>
<td id="76AF2E38FF9C000BFDFDFA01FCC1B47C" box="[574,808,1535,1554]" gridcol="2" gridrow="12" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FDFDFA01FD21B47C" bold="true" box="[574,712,1535,1554]" pageId="4" pageNumber="173">12.1-13.0 (12.7)</emphasis>
12.9
</td>
<td id="76AF2E38FF9C000BFCFCFA01FC23B47C" box="[831,970,1535,1554]" gridcol="3" gridrow="12" pageId="4" pageNumber="173">9.3-12.0 (10.5)</td>
<td id="76AF2E38FF9C000BFC1DFA01FB81B47C" box="[990,1128,1535,1554]" gridcol="4" gridrow="12" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FC20FA01FB8AB47C" bold="true" box="[995,1123,1535,1554]" pageId="4" pageNumber="173">8.0-12.4 (10.8)</emphasis>
</td>
<td id="76AF2E38FF9C000BFBBFFA01FAEEB47C" box="[1148,1287,1535,1554]" gridcol="5" gridrow="12" pageId="4" pageNumber="173">9.5-13.2 (11.4)</td>
<td id="76AF2E38FF9C000BFAD8FA01FA4FB47C" box="[1307,1446,1535,1554]" gridcol="6" gridrow="12" pageId="4" pageNumber="173">10.0-14.0 (12.2)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF9E0FA4FB45D" box="[142,1446,1566,1587]" gridrow="13" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF9E0FE6DB45D" box="[142,388,1566,1587]" gridcol="0" gridrow="13" pageId="4" pageNumber="173">Body depth</th>
<td id="76AF2E38FF9C000BFE5CF9E0FDC3B45D" box="[415,554,1566,1587]" gridcol="1" gridrow="13" pageId="4" pageNumber="173">14.1-14.9 (14.4)</td>
<td id="76AF2E38FF9C000BFDFDF9E0FCC1B45D" box="[574,808,1566,1587]" gridcol="2" gridrow="13" pageId="4" pageNumber="173">13.4-15.8 (14.5) 14.2</td>
<td id="76AF2E38FF9C000BFCFCF9E0FC23B45D" box="[831,970,1566,1587]" gridcol="3" gridrow="13" pageId="4" pageNumber="173">13.8-15.8 (14.6)</td>
<td id="76AF2E38FF9C000BFC1DF9E0FB81B45D" box="[990,1128,1566,1587]" gridcol="4" gridrow="13" pageId="4" pageNumber="173">10.6-18.1 (15.0)</td>
<td id="76AF2E38FF9C000BFBBFF9E0FAEEB45D" box="[1148,1287,1566,1587]" gridcol="5" gridrow="13" pageId="4" pageNumber="173">13.2-16.7 (14.5)</td>
<td id="76AF2E38FF9C000BFAD8F9E0FA4FB45D" box="[1307,1446,1566,1587]" gridcol="6" gridrow="13" pageId="4" pageNumber="173">12.2-19.8 (15.1)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF9C3FA4FB43C" box="[142,1446,1597,1618]" gridrow="14" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF9C3FE6DB43C" box="[142,388,1597,1618]" gridcol="0" gridrow="14" pageId="4" pageNumber="173">Pectoral-fin length</th>
<td id="76AF2E38FF9C000BFE5CF9C3FDC3B43C" box="[415,554,1597,1618]" gridcol="1" gridrow="14" pageId="4" pageNumber="173">18.9-23.0 (20.5)</td>
<td id="76AF2E38FF9C000BFDFDF9C3FCC1B43C" box="[574,808,1597,1618]" gridcol="2" gridrow="14" pageId="4" pageNumber="173">16.0-21.2 (18.4) 16.0</td>
<td id="76AF2E38FF9C000BFCFCF9C3FC23B43C" box="[831,970,1597,1618]" gridcol="3" gridrow="14" pageId="4" pageNumber="173">17.0-23.1 (19.3)</td>
<td id="76AF2E38FF9C000BFC1DF9C3FB81B43C" box="[990,1128,1597,1618]" gridcol="4" gridrow="14" pageId="4" pageNumber="173">13.4-23.0 (17.9)</td>
<td id="76AF2E38FF9C000BFBBFF9C3FAEEB43C" box="[1148,1287,1597,1618]" gridcol="5" gridrow="14" pageId="4" pageNumber="173">17.9-26.0 (20.5)</td>
<td id="76AF2E38FF9C000BFAD8F9C3FA4FB43C" box="[1307,1446,1597,1618]" gridcol="6" gridrow="14" pageId="4" pageNumber="173">16.0-23.9 (19.9)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF9A2FA4FB41F" box="[142,1446,1628,1649]" gridrow="15" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF9A2FE6DB41F" box="[142,388,1628,1649]" gridcol="0" gridrow="15" pageId="4" pageNumber="173">Pelvic-fin length</th>
<td id="76AF2E38FF9C000BFE5CF9A2FDC3B41F" box="[415,554,1628,1649]" gridcol="1" gridrow="15" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FE5CF9A3FDC3B401" bold="true" box="[415,554,1628,1648]" pageId="4" pageNumber="173">15.8-16.5 (16.3)</emphasis>
</td>
<td id="76AF2E38FF9C000BFDFDF9A2FCC1B41F" box="[574,808,1628,1649]" gridcol="2" gridrow="15" pageId="4" pageNumber="173">13.5-16.0 (15.1) 13.5</td>
<td id="76AF2E38FF9C000BFCFCF9A2FC23B41F" box="[831,970,1628,1649]" gridcol="3" gridrow="15" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FCFCF9A3FC20B401" bold="true" box="[831,969,1628,1648]" pageId="4" pageNumber="173">12.4-15.5 (14.5)</emphasis>
</td>
<td id="76AF2E38FF9C000BFC1DF9A2FB81B41F" box="[990,1128,1628,1649]" gridcol="4" gridrow="15" pageId="4" pageNumber="173">11.1-16.7 (14.0)</td>
<td id="76AF2E38FF9C000BFBBFF9A2FAEEB41F" box="[1148,1287,1628,1649]" gridcol="5" gridrow="15" pageId="4" pageNumber="173">13.9-18.2 (15.8)</td>
<td id="76AF2E38FF9C000BFAD8F9A2FA4FB41F" box="[1307,1446,1628,1649]" gridcol="6" gridrow="15" pageId="4" pageNumber="173">12.7-18.5 (15.8)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF982FA4FB4FE" box="[142,1446,1660,1680]" gridrow="16" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF982FE6DB4FE" box="[142,388,1660,1680]" gridcol="0" gridrow="16" pageId="4" pageNumber="173">Dorsal-fin length</th>
<td id="76AF2E38FF9C000BFE5CF982FDC3B4FE" box="[415,554,1660,1680]" gridcol="1" gridrow="16" pageId="4" pageNumber="173">20.9-22.7 (21.8)</td>
<td id="76AF2E38FF9C000BFDFDF982FCC1B4FE" box="[574,808,1660,1680]" gridcol="2" gridrow="16" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FDFDF982FD21B4E1" bold="true" box="[574,712,1660,1679]" pageId="4" pageNumber="173">17.1-20.1 (19.0)</emphasis>
17.1
</td>
<td id="76AF2E38FF9C000BFCFCF982FC23B4FE" box="[831,970,1660,1680]" gridcol="3" gridrow="16" pageId="4" pageNumber="173">18.7-22.5 (20.2)</td>
<td id="76AF2E38FF9C000BFC1DF982FB81B4FE" box="[990,1128,1660,1680]" gridcol="4" gridrow="16" pageId="4" pageNumber="173">16.0-24.2 (19.2)</td>
<td id="76AF2E38FF9C000BFBBFF982FAEEB4FE" box="[1148,1287,1660,1680]" gridcol="5" gridrow="16" pageId="4" pageNumber="173">18.6-25.6 (21.9)</td>
<td id="76AF2E38FF9C000BFAD8F982FA4FB4FE" box="[1307,1446,1660,1680]" gridcol="6" gridrow="16" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FAD8F982FA4FB4E1" bold="true" box="[1307,1446,1660,1679]" pageId="4" pageNumber="173">18.9-23.9 (21.6)</emphasis>
</td>
</tr>
<tr id="357E4744FF9C000BFF4DF965FA4FB4C1" box="[142,1446,1691,1711]" gridrow="17" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF965FE6DB4C1" box="[142,388,1691,1711]" gridcol="0" gridrow="17" pageId="4" pageNumber="173">Anal-fin length</th>
<td id="76AF2E38FF9C000BFE5CF965FDC3B4C1" box="[415,554,1691,1711]" gridcol="1" gridrow="17" pageId="4" pageNumber="173">15.2-17.0 (15.9)</td>
<td id="76AF2E38FF9C000BFDFDF965FCC1B4C1" box="[574,808,1691,1711]" gridcol="2" gridrow="17" pageId="4" pageNumber="173">14.2-17.2 (15.4) 14.2</td>
<td id="76AF2E38FF9C000BFCFCF965FC23B4C1" box="[831,970,1691,1711]" gridcol="3" gridrow="17" pageId="4" pageNumber="173">14.0-17.0 (15.6)</td>
<td id="76AF2E38FF9C000BFC1DF965FB81B4C1" box="[990,1128,1691,1711]" gridcol="4" gridrow="17" pageId="4" pageNumber="173">10.0-17.9 (14.8)</td>
<td id="76AF2E38FF9C000BFBBFF965FAEEB4C1" box="[1148,1287,1691,1711]" gridcol="5" gridrow="17" pageId="4" pageNumber="173">13.9-19.0 (16.6)</td>
<td id="76AF2E38FF9C000BFAD8F965FA4FB4C1" box="[1307,1446,1691,1711]" gridcol="6" gridrow="17" pageId="4" pageNumber="173">13.4-19.4 (16.4)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF944FA4FB4A3" box="[142,1446,1722,1741]" gridrow="18" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF944FE6DB4A3" box="[142,388,1722,1741]" gridcol="0" gridrow="18" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FF4DF944FECBB4A3" bold="true" box="[142,290,1722,1741]" pageId="4" pageNumber="173">Ratio CPL/CPD</emphasis>
</th>
<td id="76AF2E38FF9C000BFE5CF944FDC3B4A3" box="[415,554,1722,1741]" gridcol="1" gridrow="18" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FE5CF944FDC3B4A3" bold="true" box="[415,554,1722,1741]" pageId="4" pageNumber="173">1.40-1.45 (1.43)</emphasis>
</td>
<td id="76AF2E38FF9C000BFDFDF944FCC1B4A3" box="[574,808,1722,1741]" gridcol="2" gridrow="18" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FDFDF944FD21B4A3" bold="true" box="[574,712,1722,1741]" pageId="4" pageNumber="173">1.30-1.48 (1.37)</emphasis>
1.32
</td>
<td id="76AF2E38FF9C000BFCFCF944FC23B4A3" box="[831,970,1722,1741]" gridcol="3" gridrow="18" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FCFCF944FC20B4A3" bold="true" box="[831,969,1722,1741]" pageId="4" pageNumber="173">1.45-2.15 (1.68)</emphasis>
</td>
<td id="76AF2E38FF9C000BFC1DF944FB81B4A3" box="[990,1128,1722,1741]" gridcol="4" gridrow="18" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FC1DF944FB81B4A3" bold="true" box="[990,1128,1722,1741]" pageId="4" pageNumber="173">1.38-2.56 (1.66)</emphasis>
</td>
<td id="76AF2E38FF9C000BFBBFF944FAEEB4A3" box="[1148,1287,1722,1741]" gridcol="5" gridrow="18" pageId="4" pageNumber="173">1.26-1.77 (1.47)</td>
<td id="76AF2E38FF9C000BFAD8F944FA4FB4A3" box="[1307,1446,1722,1741]" gridcol="6" gridrow="18" pageId="4" pageNumber="173">1.14-1.71 (1.42)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF91FFA4FB483" box="[142,1446,1761,1773]" gridrow="19" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF91FFA4FB483" box="[142,1446,1761,1773]" colspan="7" colspanRight="6" gridcol="0" gridrow="19" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FF4DF91FFEF2B483" box="[142,283,1761,1773]" italics="true" pageId="4" pageNumber="173">In percent of HL</emphasis>
</th>
</tr>
<tr id="357E4744FF9C000BFF4DF907FA4FB563" box="[142,1446,1785,1805]" gridrow="20" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF907FE6DB563" box="[142,388,1785,1805]" gridcol="0" gridrow="20" pageId="4" pageNumber="173">Snout length</th>
<td id="76AF2E38FF9C000BFE5CF907FDC3B563" box="[415,554,1785,1805]" gridcol="1" gridrow="20" pageId="4" pageNumber="173">41.4-48.6 (44.3)</td>
<td id="76AF2E38FF9C000BFDFDF907FCC1B563" box="[574,808,1785,1805]" gridcol="2" gridrow="20" pageId="4" pageNumber="173">39.5-48.3 (44.5) 45.8</td>
<td id="76AF2E38FF9C000BFCFCF907FC23B563" box="[831,970,1785,1805]" gridcol="3" gridrow="20" pageId="4" pageNumber="173">39.7-53.8 (45.2)</td>
<td id="76AF2E38FF9C000BFC1DF907FB81B563" box="[990,1128,1785,1805]" gridcol="4" gridrow="20" pageId="4" pageNumber="173">39.4-50.2 (44.3)</td>
<td id="76AF2E38FF9C000BFBBFF907FAEEB563" box="[1148,1287,1785,1805]" gridcol="5" gridrow="20" pageId="4" pageNumber="173">36.0-47.9 (43.1)</td>
<td id="76AF2E38FF9C000BFAD8F907FA4FB563" box="[1307,1446,1785,1805]" gridcol="6" gridrow="20" pageId="4" pageNumber="173">38.0-51.7 (44.0)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF8E6FA4FB542" box="[142,1446,1816,1836]" gridrow="21" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF8E6FE6DB542" box="[142,388,1816,1836]" gridcol="0" gridrow="21" pageId="4" pageNumber="173">Eye diameter</th>
<td id="76AF2E38FF9C000BFE5CF8E6FDC3B542" box="[415,554,1816,1836]" gridcol="1" gridrow="21" pageId="4" pageNumber="173">15.6-19.5 (17.8)</td>
<td id="76AF2E38FF9C000BFDFDF8E6FCC1B542" box="[574,808,1816,1836]" gridcol="2" gridrow="21" pageId="4" pageNumber="173">16.0-19.3 (17.8) 18.0</td>
<td id="76AF2E38FF9C000BFCFCF8E6FC23B542" box="[831,970,1816,1836]" gridcol="3" gridrow="21" pageId="4" pageNumber="173">15.9-24.6 (18.6)</td>
<td id="76AF2E38FF9C000BFC1DF8E6FB81B542" box="[990,1128,1816,1836]" gridcol="4" gridrow="21" pageId="4" pageNumber="173">12.8-23.9 (16.3)</td>
<td id="76AF2E38FF9C000BFBBFF8E6FAEEB542" box="[1148,1287,1816,1836]" gridcol="5" gridrow="21" pageId="4" pageNumber="173">14.9-21.9 (18.9)</td>
<td id="76AF2E38FF9C000BFAD8F8E6FA4FB542" box="[1307,1446,1816,1836]" gridcol="6" gridrow="21" pageId="4" pageNumber="173">13.5-21.0 (17.6)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF8C9FA4FB525" box="[142,1446,1847,1867]" gridrow="22" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF8C9FE6DB525" box="[142,388,1847,1867]" gridcol="0" gridrow="22" pageId="4" pageNumber="173">Postorbital length</th>
<td id="76AF2E38FF9C000BFE5CF8C9FDC3B525" box="[415,554,1847,1867]" gridcol="1" gridrow="22" pageId="4" pageNumber="173">46.0-51.6 (48.3)</td>
<td id="76AF2E38FF9C000BFDFDF8C9FCC1B525" box="[574,808,1847,1867]" gridcol="2" gridrow="22" pageId="4" pageNumber="173">45.7-47.0 (46.5) 46.9</td>
<td id="76AF2E38FF9C000BFCFCF8C9FC23B525" box="[831,970,1847,1867]" gridcol="3" gridrow="22" pageId="4" pageNumber="173">45.7-54.2 (47.8)</td>
<td id="76AF2E38FF9C000BFC1DF8C9FB81B525" box="[990,1128,1847,1867]" gridcol="4" gridrow="22" pageId="4" pageNumber="173">40.7-53.5 (46.9)</td>
<td id="76AF2E38FF9C000BFBBFF8C9FAEEB525" box="[1148,1287,1847,1867]" gridcol="5" gridrow="22" pageId="4" pageNumber="173">38.4-48.8 (42.2)</td>
<td id="76AF2E38FF9C000BFAD8F8C9FA4FB525" box="[1307,1446,1847,1867]" gridcol="6" gridrow="22" pageId="4" pageNumber="173">37.3-56.3 (43.9)</td>
</tr>
<tr id="357E4744FF9C000BFF4DF8A8FA4FB507" box="[142,1446,1878,1897]" gridrow="23" pageId="4" pageNumber="173">
<th id="76AF2E38FF9C000BFF4DF8A8FE6DB507" box="[142,388,1878,1897]" gridcol="0" gridrow="23" pageId="4" pageNumber="173">Interorbital width</th>
<td id="76AF2E38FF9C000BFE5CF8A8FDC3B507" box="[415,554,1878,1897]" gridcol="1" gridrow="23" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FE5CF8A8FDC3B507" bold="true" box="[415,554,1878,1897]" pageId="4" pageNumber="173">35.5-40.5 (37.6)</emphasis>
</td>
<td id="76AF2E38FF9C000BFDFDF8A8FCC1B507" box="[574,808,1878,1897]" gridcol="2" gridrow="23" pageId="4" pageNumber="173">
<emphasis id="B93A9914FF9CFFF0FDFDF8A8FD20B507" bold="true" box="[574,713,1878,1897]" pageId="4" pageNumber="173">35.6-41.8 (38.0)</emphasis>
41.8
</td>
<td id="76AF2E38FF9C000BFCFCF8A8FC23B507" box="[831,970,1878,1897]" gridcol="3" gridrow="23" pageId="4" pageNumber="173">28.7-38.4 (33.3)</td>
<td id="76AF2E38FF9C000BFC1DF8A8FB81B507" box="[990,1128,1878,1897]" gridcol="4" gridrow="23" pageId="4" pageNumber="173">23.0-38.1 (30.9)</td>
<td id="76AF2E38FF9C000BFBBFF8A8FAEEB507" box="[1148,1287,1878,1897]" gridcol="5" gridrow="23" pageId="4" pageNumber="173">21.4-35.6 (29.4)</td>
<td id="76AF2E38FF9C000BFAD8F8A8FA4FB507" box="[1307,1446,1878,1897]" gridcol="6" gridrow="23" pageId="4" pageNumber="173">21.6-36.0 (29.4)</td>
</tr>
</table>
</paragraph>
<caption id="DF31158EFF9DFFF1FF47FD1EFE46B1EF" ID-DOI="http://doi.org/10.5281/zenodo.12711154" ID-Zenodo-Dep="12711154" httpUri="https://zenodo.org/record/12711154/files/figure.png" pageId="5" pageNumber="174" startId="5.[132,197,736,758]" targetBox="[133,1455,217,718]" targetPageId="5" targetType="figure">
<paragraph id="8BF14506FF9DFFF1FF47FD1EFBDCB163" blockId="5.[132,1455,736,897]" pageId="5" pageNumber="174">
Figure 4. – Boxplots highlighting morphometric differences between 
<taxonomicName id="4C4E3E85FF9DFFF1FCF1FD16FC1DB098" authorityName="Gauliard &amp; Dettai &amp; Persat &amp; Keith &amp; Denys" authorityYear="2019" box="[818,1012,744,758]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="174" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9DFFF1FCF1FD16FC1DB098" box="[818,1012,744,758]" italics="true" pageId="5" pageNumber="174">Barbatula leoparda</emphasis>
</taxonomicName>
(in grey) and the two other species 
<taxonomicName id="4C4E3E85FF9DFFF1FA95FD16FF12B160" authorityName="MNHN" authorityYear="2013" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="174" phylum="Chordata" rank="species" species="barbatula">
<emphasis id="B93A9914FF9DFFF1FA95FD16FF12B160" italics="true" pageId="5" pageNumber="174">Barbatula barbatula</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4E3E85FF9DFFF1FEEBFCFEFE04B160" box="[296,493,768,782]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="174" phylum="Chordata" rank="species" species="quignardi">
<emphasis id="B93A9914FF9DFFF1FEEBFCFEFE04B160" box="[296,493,768,782]" italics="true" pageId="5" pageNumber="174">Barbatula quignardi</emphasis>
</taxonomicName>
, for the two sizes classes (36 to 47 mm SL and&gt; 47 mm SL).
</paragraph>
<paragraph id="8BF14506FF9DFFF1FF47FCB6FE46B1EF" blockId="5.[132,1455,736,897]" pageId="5" pageNumber="174">
only by a larger interorbital width (35.5 to 40.5% HL, 
<emphasis id="B93A9914FF9DFFF1FD2AFCAFFCE1B10E" box="[745,776,849,864]" italics="true" pageId="5" pageNumber="174">vs.</emphasis>
21.4 to 35.6% HL; Fig. 4B).
</paragraph>
</caption>
<paragraph id="8BF14506FF9DFFF1FF69FC34FE90B4BE" blockId="5.[132,776,937,1745]" pageId="5" pageNumber="174">
For general appearance, see figure 1; morphometric data are provided in table I. Medium sized species. The stone loaches belonging to 
<taxonomicName id="4C4E3E85FF9DFFF1FEA4FBEBFE0DB64A" authorityName="Gauliard &amp; Dettai &amp; Persat &amp; Keith &amp; Denys" authorityYear="2019" box="[359,484,1045,1060]" family="Nemacheilidae" genus="Barbatula" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="174" phylum="Chordata" rank="species" species="leoparda">
<emphasis id="B93A9914FF9DFFF1FEA4FBEBFE0DB64A" box="[359,484,1045,1060]" italics="true" pageId="5" pageNumber="174">B. leoparda</emphasis>
</taxonomicName>
present a marbled pigmentation on the body with brown-dark blotches on the belly and in the jugular area for specimens larger than 
<quantity id="4CB6E8E3FF9DFFF1FD55FBB2FD0BB60A" box="[662,738,1100,1124]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="174" unit="mm" value="47.0">47 mm</quantity>
SL (
<figureCitation id="13755983FF9DFFF1FF4FFB93FF27B6EB" box="[140,206,1133,1157]" captionStart="Figure 2" captionStartId="3.[132,200,1572,1594]" captionTargetBox="[549,1456,1331,1860]" captionTargetId="figure-174@3.[549,1456,1331,1860]" captionTargetPageId="3" captionText="Figure 2. – Specimens of Barbatula spp. in ventral view. A: Holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefi- ach, 7 Sep. 2016, Denys and ONEMA coll.; B: Barbatula barbatula MNHN 2013-0826, 72.9 mm SL ♂, stream Clauge (Rhône drainage) at La Loye, 1 Jul. 2013, Denys and ONEMA coll.; C: Holotype of Barbatula quignardi MGAB77, 52 mm SL ♂, Lez River, Jul. 1962, Băcescu coll." figureDoi="http://doi.org/10.5281/zenodo.12711150" httpUri="https://zenodo.org/record/12711150/files/figure.png" pageId="5" pageNumber="174">Fig. 2</figureCitation>
). The lateral profile is straight and the head is round- ed. The body is cylindrical in front of the dorsal-fin origin and the tail is laterally compressed. The body width is equal on all its length, except for the head. The anterior and posteri- or nostrils are close to each other without any space between them. The mouth bears 3 pairs of barbels, and the last one located at the corners does not reach the posterior part of the eye. The upper lip is slim with a medial incision not exceeding the half of the width of the upper lip (
<figureCitation id="13755983FF9DFFF1FDF8FA8CFD92B7E4" box="[571,635,1394,1418]" captionStart="Figure 3" captionStartId="4.[132,195,714,736]" captionTargetBox="[132,775,215,697]" captionTargetId="figure-352@4.[132,775,215,697]" captionTargetPageId="4" captionText="Figure 3. – Mouth of holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.Abbreviations: medial indentation of upper lip (mi), mental lobe of lower lip (ml). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12711152" httpUri="https://zenodo.org/record/12711152/files/figure.png" pageId="5" pageNumber="174">Fig. 3</figureCitation>
). The mental lobes on the lower lip are not well marked (
<figureCitation id="13755983FF9DFFF1FD92FA6DFD78B7C5" box="[593,657,1427,1451]" captionStart="Figure 3" captionStartId="4.[132,195,714,736]" captionTargetBox="[132,775,215,697]" captionTargetId="figure-352@4.[132,775,215,697]" captionTargetPageId="4" captionText="Figure 3. – Mouth of holotype of Barbatula leoparda MNHN 2018- 0228, 62.6 mm SL ♀, Têt River at Nefiach, 7 Sep. 2016, Denys and ONEMA coll.Abbreviations: medial indentation of upper lip (mi), mental lobe of lower lip (ml). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.12711152" httpUri="https://zenodo.org/record/12711152/files/figure.png" pageId="5" pageNumber="174">Fig. 3</figureCitation>
). This species has III simple and 7 to 8 branched dorsal fin rays (
<typeStatus id="54F5FBA4FF9DFFF1FD0EFA4DFF5BB782" pageId="5" pageNumber="174" type="holotype">holotype</typeStatus>
III7), III simple and 5 to 6 branched anal fin rays (
<typeStatus id="54F5FBA4FF9DFFF1FD0EFA2AFF5DB463" pageId="5" pageNumber="174" type="holotype">holotype</typeStatus>
III5), I simple and 9 to 11 branched pectoral fin rays (
<typeStatus id="54F5FBA4FF9DFFF1FF4FF9EBFF07B443" box="[140,238,1557,1581]" pageId="5" pageNumber="174" type="holotype">holotype</typeStatus>
I11), I simple and 6 to 7 branched pelvic fin rays (
<typeStatus id="54F5FBA4FF9DFFF1FF4FF9C8FF07B420" box="[140,238,1590,1614]" pageId="5" pageNumber="174" type="holotype">holotype</typeStatus>
I7), and 14 to 18 branched caudal-fin rays (
<typeStatus id="54F5FBA4FF9DFFF1FD08F9C8FF5BB401" pageId="5" pageNumber="174" type="holotype">holotype</typeStatus>
18). The lateral line is complete. Specimens larger than 
<quantity id="4CB6E8E3FF9DFFF1FF47F989FF27B4E1" box="[132,206,1655,1679]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="5" pageNumber="174" unit="mm" value="47.0">47 mm</quantity>
SL have a proportionally deeper caudal peduncle and a smaller dorsal fin than smaller specimens. Largest known specimen 
<quantity id="4CB6E8E3FF9DFFF1FF2CF946FEA4B4BE" box="[239,333,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="6.26" pageId="5" pageNumber="174" unit="mm" value="62.6">62.6 mm</quantity>
SL.
</paragraph>
</subSubSection>
</treatment>
</document>