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<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="A1610A2DB89353C2957F92D8AB247D0F" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Tschaidicancha joseochoai Benedetti &amp; Pinto-da-Rocha 2022, sp. nov." docType="treatment" docUuid="FC63BBD6-6531-46D8-95BC-1A217C7B9B0B" docUuidSource="ZooBank" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
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<mods:start>309</mods:start>
<mods:end>388</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:FC63BBD6-6531-46D8-95BC-1A217C7B9B0B" httpUri="http://treatment.plazi.org/id/A1610A2DB89353C2957F92D8AB247D0F" lastPageNumber="309" pageId="0" pageNumber="309">
<subSubSection pageId="0" pageNumber="309" type="nomenclature">
<paragraph pageId="0" pageNumber="309">
3.51.
<taxonomicName LSID="http://zoobank.org/FC63BBD6-6531-46D8-95BC-1A217C7B9B0B" authority="Benedetti &amp; Pinto-da-Rocha, 2022" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Tschaidicancha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tschaidicancha joseochoai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="joseochoai" status="sp. nov.">Tschaidicancha joseochoai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="309">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">Figs 6D</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">, 11I, J</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">, 14I, J</figureCitation>
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Live specimens of Tschaidicancha. A T. joseochoai sp. nov., male; B female; C T. chaplini sp. nov., male; D female; E T. weyrauchi Roewer, 1957, male; F: female." figureDoi="10.3897/asp.80.e73829.figure27" httpUri="https://binary.pensoft.net/fig/729901" pageId="0" pageNumber="309">, 27A, B</figureCitation>
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">, 29B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">Description.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 4,4; DSL: 4,2; CL: 1,7; FIVL: 7,8; ChL: 3,0.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Live specimens of Tschaidicancha. A T. joseochoai sp. nov., male; B female; C T. chaplini sp. nov., male; D female; E T. weyrauchi Roewer, 1957, male; F: female." figureDoi="10.3897/asp.80.e73829.figure27" httpUri="https://binary.pensoft.net/fig/729901" pageId="0" pageNumber="309">27A</figureCitation>
) Predominantly orange brown, with two white spots on the carapace, laterally posterior to the ocularium. Dark brown legs. Chelicerae orange brown (in a shade darker than body). Tubercles of the DS areas, free tergites and lateral margins whitish.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D</figureCitation>
) Gamma-P-type DSS, with concave posterior margin of DS. Anterior margin of the carapace with medium elevation, practically smooth, with few sparse granules. Ocularium with mild median depression; a pair of acuminate spines. Carapace with few granules, distributed mainly in the lateral region. Areas I-IV without granules, except elevation on area III. Area I undivided longitudinally; with a median pair of small tubercles. Area II with a median pair of tubercles and a right lateral tubercle. Area III with two spines, directed posteriorly located in elevations of the integument, very granular. Area IV with a median pair of tubercles. Posterior margin of DS smooth; with a left lateral tubercle. Free tergites I-III smooth. Free tergite I unarmed. Free tergite II with a pair of lateral tubercles. Free tergite III with three tubercles. Lateral margins of DS smooth; seven-nine tubercles in the posterior half, close to areas II-IV and posterior margin of DS.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D</figureCitation>
) Swollen. Segments I-II with few granules. Segment II with finger with one tooth. Segment III with 5-6 tiny teeth.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: With sparse granules dorsally in the femur. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal tubercle; a ventral row of four setiferous tubercles, the basal-most being smallest and the remaining three of equal size, distributed throughout the article, except at the apex; a proapical spine. Patella with a proapical tubercle. Tibia: prolateral iiIii, retrolateral IiIi. Tarsus: prolateral iIiii, retrolateral IiIi.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxae I-II with rows of four and six tubercles, respectively, those of the coxa I larger. Coxae III-IV with granules. Genital area with few granules. Free sternites with row of granules. Anal operculum granular.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">11I, J</figureCitation>
) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with 7-10 tubercles, distributed on the prodorsal side. Trochanters I-IV unarmed and with few granules. Femora I-III unarmed and with small granules. Femur IV with dense dorsal granulation; a prolateral row of 29 acuminate tubercles along apcal ⅔; a retrolateral row of 34 acuminate tubercles; a prodorsal row of five acuminate tubercles and three blunt tubercles along basal ⅓, smaller than the other tubercles present in the article; a ventral row of granules. Patellae I-III unarmed, with few granules. Patella IV granular; with three dorsoapical spiniform tubercles. Tibiae I-IV unarmed and densely granular. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) 9 / 16 / 14 / 17.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B Lumieria woodyalleni gen. et sp. nov.; C - D Metasarcus beni sp. nov.; E - F Metasarcus kurosawai sp. nov.; G - H Metasarcus limachii sp. nov.; I - J Tschaidicancha joseochoai sp. nov.; K - L Tschaidicancha weyrauchi Roewer, 1957; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure14" httpUri="https://binary.pensoft.net/fig/729888" pageId="0" pageNumber="309">14I, J</figureCitation>
) VP subrectangular, long; distal margin slightly convex, with distal-lateral projections; straight in lateral view. MS C1-C3(4) subdistal long (MS C1-C2 shorter than the others) and slightly curved; MS A1 short and straight, medially placed; MS B1 sub basal long and straight. Lateral sacs long and apically blunt; with long T3-like microsetae. Stylus expanded in the shape of a wedge and flattened laterally, with serrated edges; with large projection in the dorsal direction and with several tiny apical projections. Dorsal process subrectangular. -
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) DSW: 4.2; DSL: 4.1; CL: 1.5. FIVL: 8.7. ChL: 1.7. (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Live specimens of Tschaidicancha. A T. joseochoai sp. nov., male; B female; C T. chaplini sp. nov., male; D female; E T. weyrauchi Roewer, 1957, male; F: female." figureDoi="10.3897/asp.80.e73829.figure27" httpUri="https://binary.pensoft.net/fig/729901" pageId="0" pageNumber="309">27B</figureCitation>
) Chelicerae smaller than the male. Femur IV unarmed. Patella IV with acuminate tubercles as in the male, but less robust and smaller. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=1) 8, 15, 14, 17.
</paragraph>
<caption doi="10.3897/asp.80.e73829.figure30" httpUri="https://binary.pensoft.net/fig/729904" pageId="0" pageNumber="309" start="Figure 30" startId="F30">
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">Figure 30.</emphasis>
Distribution of
<taxonomicName authorityName="Kury &amp; Maury" authorityYear="1998" class="Arachnida" family="Metasarcidae" genus="Incasarcus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Incasarcus" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="309">Incasarcus</emphasis>
</taxonomicName>
in Cusco, Peru.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
It differs from other species of the genus by the following set of characteristics: gamma-P-type DSS; ocularium with a pair of acuminate tubercles; DS and scutal areas smooth; scutal area I undivided; free tergites I-III unarmed (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Habitus, dorsal of males of Lumieria gen. nov. and Tschaidicancha. A L. antonionii gen. et sp. nov.; B L. woodyalleni gen. et sp. nov.; C T. chaplini sp. nov.; D T. joseochoai sp. nov.; E T. scorsesei sp. nov.; F T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure6" httpUri="https://binary.pensoft.net/fig/729880" pageId="0" pageNumber="309">6D</figureCitation>
); male femur IV with a prolateral row of 29 acuminate tubercles and a retrolateral row of 34 acuminate tubercles (Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Male trochanter-patella / tibia IV of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha, dorsal and ventral view, respectively. A - B H. kubricki gen. et sp. nov.; C - D L. antonionii gen. et sp. nov.; E - F L. woodyalleni gen. et sp. nov.; G - H T. chaplini sp. nov.; I - J T. joseochoai sp. nov.; K - L T. scorsesei sp. nov.; M - N T. weyrauchi Roewer, 1957; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure11" httpUri="https://binary.pensoft.net/fig/729885" pageId="0" pageNumber="309">11I, J</figureCitation>
); body predominantly orange-brown with two white spots on the carapace; tubercles of the DS areas, free tergites and lateral margins whitish (Fig.
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Live specimens of Tschaidicancha. A T. joseochoai sp. nov., male; B female; C T. chaplini sp. nov., male; D female; E T. weyrauchi Roewer, 1957, male; F: female." figureDoi="10.3897/asp.80.e73829.figure27" httpUri="https://binary.pensoft.net/fig/729901" pageId="0" pageNumber="309">27A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="etymology">
<paragraph pageId="0" pageNumber="309">Derivatio nominis.</paragraph>
<paragraph pageId="0" pageNumber="309">
The specific epithet of masculine gender, in the genitive form, dedicated Dr.
<normalizedToken originalValue="José">Jose</normalizedToken>
Ochoa C. (MUBI), for his great help in fieldwork in Bolivia and Peru.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig.
<figureCitation captionStart="Figure 29" captionStartId="F29" captionText="Figure 29. Distribution of Huancabamba gen. nov., Lumieria gen. nov. and Tschaidicancha in Peru. A H. kubricki gen. et sp. nov., L. antonionii gen. et sp. nov., T. chaplini sp. nov., T. scorsesei sp. nov., T. weyrauchi; B L. woodyalleni gen. et sp. nov., T. joseochoai sp. nov." figureDoi="10.3897/asp.80.e73829.figure29" httpUri="https://binary.pensoft.net/fig/729903" pageId="0" pageNumber="309">29B</figureCitation>
) PERU. Pasco. Oxapampa. Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<materialsCitation collectingDate="2011-04-22" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" latitude="-10.545029" location="Parque Nacional Yanachaga-Chemillen" longLatPrecision="1" longitude="-75.35677" municipality="Oxapampa" specimenCount="♂" stateProvince="Pasco" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Holotype</typeStatus>
</emphasis>
<specimenCount></specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<collectingRegion country="Peru" name="Pasco">Pasco</collectingRegion>
,
<collectingMunicipality>Oxapampa</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:A1610A2DB89353C2957F92D8AB247D0F:1390B9249C541BD6984EB2EBAFB60BD6" country="PERU" latitude="-10.545029" longLatPrecision="1" longitude="-75.35677" municipality="Oxapampa" name="Parque Nacional Yanachaga-Chemillen" stateProvince="Pasco">
Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
</location>
<geoCoordinate degrees="10" direction="south" minutes="32" orientation="latitude" precision="1" seconds="42.1" value="-10.545029">10°3242.1″S</geoCoordinate>
<geoCoordinate degrees="75" direction="west" minutes="21" orientation="longitude" precision="1" seconds="24.4" value="-75.35677">75°2124.4″W</geoCoordinate>
,
<collectingDate value="2011-04-22">22/IV/2011</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
,
<collectorName>A. Benedetti</collectorName>
,
<collectorName>J. Ochoa</collectorName>
&amp;
<collectorName>D. Silva</collectorName>
leg. (MUBI)
</materialsCitation>
-
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount type="female">1 ♀</specimenCount>
,
<normalizedToken originalValue="ditto">'ditto'</normalizedToken>
(MZSP 36988).
<materialsCitation collectingDate="2011-04-09" collectingDateMax="2011-04-11" collectingDateMin="2011-04-09" collectorName="S. Cardonel, J. Grados" country="PERU" latitude="-10.545139" location="Parque Nacional Yanachaga-Chemillen" longLatPrecision="1" longitude="-75.35828" municipality="Oxapampa" specimenCount="1" specimenCount-male="1" stateProvince="Pasco" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount type="male">1 ♂</specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<collectingRegion country="Peru" name="Pasco">Pasco</collectingRegion>
,
<collectingMunicipality>Oxapampa</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:A1610A2DB89353C2957F92D8AB247D0F:E124E249A0EE429AB2C89359307F4E49" country="PERU" latitude="-10.545139" longLatPrecision="1" longitude="-75.35828" municipality="Oxapampa" name="Parque Nacional Yanachaga-Chemillen" stateProvince="Pasco">
Parque Nacional
<normalizedToken originalValue="Yanachaga-Chemillén">Yanachaga-Chemillen</normalizedToken>
</location>
<geoCoordinate degrees="10" direction="south" minutes="32" orientation="latitude" precision="1" seconds="42.5" value="-10.545139">10°3242.5″S</geoCoordinate>
<geoCoordinate degrees="75" direction="west" minutes="21" orientation="longitude" precision="1" seconds="29.8" value="-75.35828">75°2129.8″W</geoCoordinate>
,
<collectingDate value="2011-04-09" valueMax="2011-04-11" valueMin="2011-04-09">9-11/IV/2011</collectingDate>
,
<collectorName>S. Cardonel</collectorName>
&amp;
<collectorName>J. Grados</collectorName>
leg. (MUSM)
</materialsCitation>
;
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratypes</typeStatus>
</emphasis>
<specimenCount type="male">3 ♂</specimenCount>
,
<specimenCount type="female">3 ♀</specimenCount>
,
<normalizedToken originalValue="ditto">'ditto'</normalizedToken>
(MZSP 36984).
</paragraph>
</subSubSection>
</treatment>
</document>