Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species Author Benedetti, Alipio Rezende Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil & Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil alipiobenedetti@gmail.com Author Pinto-da-Rocha, Ricardo https://orcid.org/0000-0002-3959-2205 Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil text Arthropod Systematics & amp; Phylogeny 2022 2022-08-16 80 309 388 http://dx.doi.org/10.3897/asp.80.e73829 journal article http://dx.doi.org/10.3897/asp.80.e73829 1864-8312-80-309 D5C0468B99A14EF39237D9BC51A8BDA3 11907D1B6F665D96B5FD2F9751431366 3.51. Tschaidicancha joseochoai sp. nov. Figs 6D , 11I, J , 14I, J , 27A, B , 29B Description. MALE: Measurements ( n =1) DSW: 4,4; DSL: 4,2; CL: 1,7; FIVL: 7,8; ChL: 3,0. Coloration : (Fig. 27A ) Predominantly orange brown, with two white spots on the carapace, laterally posterior to the ocularium. Dark brown legs. Chelicerae orange brown (in a shade darker than body). Tubercles of the DS areas, free tergites and lateral margins whitish. Dorsum : (Fig. 6D ) Gamma-P-type DSS, with concave posterior margin of DS. Anterior margin of the carapace with medium elevation, practically smooth, with few sparse granules. Ocularium with mild median depression; a pair of acuminate spines. Carapace with few granules, distributed mainly in the lateral region. Areas I-IV without granules, except elevation on area III. Area I undivided longitudinally; with a median pair of small tubercles. Area II with a median pair of tubercles and a right lateral tubercle. Area III with two spines, directed posteriorly located in elevations of the integument, very granular. Area IV with a median pair of tubercles. Posterior margin of DS smooth; with a left lateral tubercle. Free tergites I-III smooth. Free tergite I unarmed. Free tergite II with a pair of lateral tubercles. Free tergite III with three tubercles. Lateral margins of DS smooth; seven-nine tubercles in the posterior half, close to areas II-IV and posterior margin of DS. Chelicerae : (Fig. 6D ) Swollen. Segments I-II with few granules. Segment II with finger with one tooth. Segment III with 5-6 tiny teeth. Pedipalpus : With sparse granules dorsally in the femur. Trochanter with a ventroapical setiferous tubercle. Femur with a ventrobasal tubercle; a ventral row of four setiferous tubercles, the basal-most being smallest and the remaining three of equal size, distributed throughout the article, except at the apex; a proapical spine. Patella with a proapical tubercle. Tibia: prolateral iiIii, retrolateral IiIi. Tarsus: prolateral iIiii, retrolateral IiIi. Venter : Coxae I-II with rows of four and six tubercles, respectively, those of the coxa I larger. Coxae III-IV with granules. Genital area with few granules. Free sternites with row of granules. Anal operculum granular. Legs : (Figs 6D , 11I, J ) Coxae I-II each one with a prolateral and a retrolateral apophysis. Coxa III with a prolateral apophysis. Coxa IV with 7-10 tubercles, distributed on the prodorsal side. Trochanters I-IV unarmed and with few granules. Femora I-III unarmed and with small granules. Femur IV with dense dorsal granulation; a prolateral row of 29 acuminate tubercles along apcal ⅔; a retrolateral row of 34 acuminate tubercles; a prodorsal row of five acuminate tubercles and three blunt tubercles along basal ⅓, smaller than the other tubercles present in the article; a ventral row of granules. Patellae I-III unarmed, with few granules. Patella IV granular; with three dorsoapical spiniform tubercles. Tibiae I-IV unarmed and densely granular. Tarsal segmentation: ( n =1) 9 / 16 / 14 / 17. Penis : (Fig. 14I, J ) VP subrectangular, long; distal margin slightly convex, with distal-lateral projections; straight in lateral view. MS C1-C3(4) subdistal long (MS C1-C2 shorter than the others) and slightly curved; MS A1 short and straight, medially placed; MS B1 sub basal long and straight. Lateral sacs long and apically blunt; with long T3-like microsetae. Stylus expanded in the shape of a wedge and flattened laterally, with serrated edges; with large projection in the dorsal direction and with several tiny apical projections. Dorsal process subrectangular. - FEMALE: Measurements ( n =1) DSW: 4.2; DSL: 4.1; CL: 1.5. FIVL: 8.7. ChL: 1.7. (Fig. 27B ) Chelicerae smaller than the male. Femur IV unarmed. Patella IV with acuminate tubercles as in the male, but less robust and smaller. Tarsal segmentation: ( n =1) 8, 15, 14, 17. Figure 30. Distribution of Incasarcus in Cusco, Peru. Diagnosis. It differs from other species of the genus by the following set of characteristics: gamma-P-type DSS; ocularium with a pair of acuminate tubercles; DS and scutal areas smooth; scutal area I undivided; free tergites I-III unarmed (Fig. 6D ); male femur IV with a prolateral row of 29 acuminate tubercles and a retrolateral row of 34 acuminate tubercles (Fig. 11I, J ); body predominantly orange-brown with two white spots on the carapace; tubercles of the DS areas, free tergites and lateral margins whitish (Fig. 27A ). Derivatio nominis. The specific epithet of masculine gender, in the genitive form, dedicated Dr. Jose Ochoa C. (MUBI), for his great help in fieldwork in Bolivia and Peru. Distribution. (Fig. 29B ) PERU. Pasco. Oxapampa. Parque Nacional Yanachaga-Chemillen . Material examined. Holotype ♂ , ' PERU , Pasco , Oxapampa , Parque Nacional Yanachaga-Chemillen 10°32′42.1″S 75°21′24.4″W , 22/IV/2011 , R. Pinto-da-Rocha , A. Benedetti , J. Ochoa & D. Silva leg. (MUBI) - Paratype 1 ♀ , 'ditto' (MZSP 36988). Paratype 1 ♂ , ' PERU , Pasco , Oxapampa , Parque Nacional Yanachaga-Chemillen 10°32′42.5″S 75°21′29.8″W , 9-11/IV/2011 , S. Cardonel & J. Grados leg. (MUSM) ; Paratypes 3 ♂ , 3 ♀ , 'ditto' (MZSP 36984).