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<document id="C5698901EB232BB30A12C2C4B9793484" ID-DOI="10.11646/zootaxa.3884.5.4" ID-GBIF-Dataset="77bac6b4-70c7-44b9-a1f4-8f8a0aa894cf" ID-ISSN="1175-5326" ID-Zenodo-Dep="224374" ID-ZooBank="D02F0C43-F20E-430B-A584-430D9FCF652F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460971347875" checkinUser="plazi" docAuthor="Ng, Heok Hee &amp; Kottelat, Maurice" docDate="2014" docId="F55F8F44FFA2FFACE68A0669FA6CF89F" docLanguage="en" docName="zt03884p444.pdf" docOrigin="Zootaxa 3884 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Clarias serniosus, sp. nov." docType="treatment" docVersion="11" lastPageNumber="440" masterDocId="0966F73CFFA3FFAFE61D042FF938FFF8" masterDocTitle="Clarias serniosus, a new walking catfish (Teleostei: Clariidae) from Laos" masterLastPageNumber="444" masterPageNumber="437" pageNumber="438" updateTime="1698558878701" updateUser="plazi">
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<mods:title id="5408BC6D089EF3A36D2EDAFE41F6AB90">Clarias serniosus, a new walking catfish (Teleostei: Clariidae) from Laos</mods:title>
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<mods:namePart id="C46D8D3FC7F5BD387B0F3D507226C19A">Ng, Heok Hee</mods:namePart>
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<mods:namePart id="076DA326792DDC08FEF2AFB52BF5E8A1">Kottelat, Maurice</mods:namePart>
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<mods:date id="8EA6CA6A2857689E75D4306A1C34F46C">2014</mods:date>
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<treatment id="F55F8F44FFA2FFACE68A0669FA6CF89F" ID-DOI="http://doi.org/10.5281/zenodo.5657609" ID-GBIF-Taxon="119593882" ID-Zenodo-Dep="5657609" LSID="urn:lsid:plazi:treatment:F55F8F44FFA2FFACE68A0669FA6CF89F" httpUri="http://treatment.plazi.org/id/F55F8F44FFA2FFACE68A0669FA6CF89F" lastPageId="3" lastPageNumber="440" pageId="1" pageNumber="438">
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<paragraph id="7D493E52FFA2FFAEE68A0669F8F5FD98" blockId="1.[151,461,582,642]" box="[151,461,582,608]" pageId="1" pageNumber="438">
<heading id="2601893EFFA2FFAEE68A0669F8F5FD98" bold="true" box="[151,461,582,608]" fontSize="11" level="1" pageId="1" pageNumber="438" reason="1">
<emphasis id="4F82E240FFA2FFAEE68A0669F8F5FD98" bold="true" box="[151,461,582,608]" pageId="1" pageNumber="438">
<taxonomicName id="BAF645D1FFA2FFAEE68A0669F850FD98" ID-CoL="5Z88H" authority="Ng &amp; Kottelat, 2014" box="[151,360,582,608]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="serniosus" status="sp. nov.">
<emphasis id="4F82E240FFA2FFAEE68A0669F850FD98" bold="true" box="[151,360,582,608]" italics="true" pageId="1" pageNumber="438">Clarias serniosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54B15F3BFFA2FFAEE7720669F8F5FD98" box="[367,461,582,608]" pageId="1" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="35EC6DD9FFA2FFAEE68A0646F9D3FD7A" box="[151,235,617,642]" pageId="1" pageNumber="438" type="description">
<paragraph id="7D493E52FFA2FFAEE68A0646F9D3FD7A" blockId="1.[151,461,582,642]" box="[151,235,617,642]" pageId="1" pageNumber="438">
(
<figureCitation id="E5CD22D7FFA2FFAEE6820646F9D8FD7A" box="[159,224,617,642]" captionStart="FIGURE 1" captionStartId="3.[151,250,1058,1080]" captionTargetBox="[151,1436,193,1036]" captionTargetId="figure@3.[151,1436,193,1037]" captionTargetPageId="3" captionText="FIGURE 1. Clarias serniosus, holotype, MHNG 2745.072, 141 mm SL; Laos: Bolavens Plateau, Xe Pian drainage. Dorsal and lateral views." httpUri="https://zenodo.org/record/224375/files/figure.png" pageId="1" pageNumber="438">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA2FFAEE68A069EFA23FCCE" pageId="1" pageNumber="438" type="materials_examined">
<paragraph id="7D493E52FFA2FFAEE68A069EFA00FCEA" blockId="1.[151,1437,689,2010]" pageId="1" pageNumber="438">
<emphasis id="4F82E240FFA2FFAEE68A069EF87FFD32" bold="true" box="[151,327,689,714]" pageId="1" pageNumber="438">
<typeStatus id="A24D80F0FFA2FFAEE68A069EF9EAFD32" box="[151,210,689,714]" pageId="1" pageNumber="438">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="CD9E340FFFA2FFAEE752069EFA00FCEA" ID-GBIF-Occurrence="3467425301" collectingDate="2013-01-17" collectionCode="MHNG" collectorName="M. Kottelat &amp; T. Phommavong" country="Laos" elevation="0" latitude="15.042778" location="Bolavens Plateau" longLatPrecision="21" longitude="106.36472" municipality="Champasak Province" pageId="1" pageNumber="438" specimenCode="MHNG 2745.072" specimenCount="1" typeStatus="holotype">
<typeStatus id="A24D80F0FFA2FFAEE752069EF884FD32" box="[335,444,689,714]" pageId="1" pageNumber="438" type="holotype">Holotype</typeStatus>
:
<specimenCode id="2D509629FFA2FFAEE7D5069DFBAAFD32" box="[456,658,689,714]" pageId="1" pageNumber="438">MHNG 2745.072</specimenCode>
, 141 mm SL;
<collectingCountry id="05E17EC2FFA2FFAEE525069DFA4BFD31" box="[824,883,690,713]" name="Laos" pageId="1" pageNumber="438">Laos</collectingCountry>
:
<collectingMunicipality id="9D2DA428FFA2FFAEE562069EFD4AFD32" box="[895,1138,689,714]" pageId="1" pageNumber="438">Champasak Province</collectingMunicipality>
,
<location id="78296889FFA2FFAEE263069EFC7CFD32" LSID="urn:lsid:plazi:treatment:F55F8F44FFA2FFACE68A0669FA6CF89F:78296889FFA2FFAEE263069EFC7CFD32" box="[1150,1348,689,714]" country="Laos" latitude="15.042778" longLatPrecision="21" longitude="106.36472" municipality="Champasak Province" name="Bolavens Plateau" pageId="1" pageNumber="438">Bolavens Plateau</location>
,
<location id="78296889FFA2FFAEE34F069DFB3DFD16" LSID="urn:lsid:plazi:treatment:F55F8F44FFA2FFACE68A0669FA6CF89F:78296889FFA2FFAEE34F069DFB3DFD16" country="Laos" latitude="15.042778" longLatPrecision="21" longitude="106.36472" municipality="Champasak Province" name="Houay Slarm near Ban Pha Nouandong" pageId="1" pageNumber="438">Houay Slarm near Ban Pha Nouandong</location>
, a tributary of
<location id="78296889FFA2FFAEE4B706F9FA15FD16" LSID="urn:lsid:plazi:treatment:F55F8F44FFA2FFACE68A0669FA6CF89F:78296889FFA2FFAEE4B706F9FA15FD16" box="[682,813,726,750]" country="Laos" latitude="15.042778" longLatPrecision="21" longitude="106.36472" municipality="Champasak Province" name="Houay Xoy" pageId="1" pageNumber="438">Houay Xoy</location>
, itself a tributary of
<location id="78296889FFA2FFAEE20E06F9FD48FD16" LSID="urn:lsid:plazi:treatment:F55F8F44FFA2FFACE68A0669FA6CF89F:78296889FFA2FFAEE20E06F9FD48FD16" box="[1043,1136,725,750]" country="Laos" latitude="15.042778" longLatPrecision="21" longitude="106.36472" municipality="Champasak Province" name="Xe Pian" pageId="1" pageNumber="438">Xe Pian</location>
,
<geoCoordinate id="18C25895FFA2FFAEE26606FAFDC7FD16" box="[1147,1279,725,750]" direction="north" orientation="latitude" pageId="1" pageNumber="438" precision="15" value="15.042778">15°02'34&quot;N</geoCoordinate>
<geoCoordinate id="18C25895FFA2FFAEE31B06FAFCAFFD16" box="[1286,1431,725,750]" direction="east" orientation="longitude" pageId="1" pageNumber="438" precision="15" value="106.36472">106°21'53&quot;E</geoCoordinate>
,
<elevation id="F6DBD961FFA2FFAEE68A06D6F839FCEA" box="[151,257,761,786]" pageId="1" pageNumber="438">985 masl</elevation>
;
<collectorName id="D0035B84FFA2FFAEE71106D5F8B4FCEA" box="[268,396,761,786]" pageId="1" pageNumber="438">M. Kottelat</collectorName>
&amp;
<collectorName id="D0035B84FFA2FFAEE7AD06D5FB55FCEA" box="[432,621,761,786]" pageId="1" pageNumber="438">T. Phommavong</collectorName>
,
<date id="09481892FFA2FFAEE46A06D5FA0CFCEA" box="[631,820,762,786]" pageId="1" pageNumber="438" value="2013-01-17">
<collectingDate id="190CE17AFFA2FFAEE46A06D5FA0CFCEA" box="[631,820,762,786]" pageId="1" pageNumber="438" value="2013-01-17">17 January 2013</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="7D493E52FFA2FFAEE68A0731FA23FCCE" blockId="1.[151,1437,689,2010]" box="[151,795,797,822]" pageId="1" pageNumber="438">
<materialsCitation id="CD9E340FFFA2FFAEE68A0731FA23FCCE" ID-GBIF-Occurrence="3467425302" box="[151,795,797,822]" pageId="1" pageNumber="438" typeStatus="paratype">
<typeStatus id="A24D80F0FFA2FFAEE68A0731F9C6FCCE" box="[151,254,798,822]" pageId="1" pageNumber="438" type="paratype">Paratype</typeStatus>
: CMK
<quantity id="BA0E93B7FFA2FFAEE7530731F8AFFCCE" box="[334,407,798,822]" metricMagnitude="4" metricUnit="m" metricValue="2.3382161" pageId="1" pageNumber="438" unit="mm" value="2.3382161E7">23382</quantity>
, 161 mm SL; data as for
<typeStatus id="A24D80F0FFA2FFAEE4AC0732FA2FFCCE" box="[689,791,797,822]" pageId="1" pageNumber="438" type="holotype">holotype</typeStatus>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA2FFAEE6DA076EFD7DFBAE" pageId="1" pageNumber="438" type="diagnosis">
<paragraph id="7D493E52FFA2FFAEE6DA076EFD7DFBAE" blockId="1.[151,1437,689,2010]" pageId="1" pageNumber="438">
<emphasis id="4F82E240FFA2FFAEE6DA076EF879FCA2" bold="true" box="[199,321,833,858]" pageId="1" pageNumber="438">Diagnosis.</emphasis>
<taxonomicName id="BAF645D1FFA2FFAEE755076EFB34FCA1" box="[328,524,833,857]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA2FFAEE755076EFB34FCA1" box="[328,524,833,857]" italics="true" pageId="1" pageNumber="438">Clarias serniosus</emphasis>
</taxonomicName>
is distinguished from all Southeast Asian congeners, except for
<taxonomicName id="BAF645D1FFA2FFAEE2C7076DFC41FCA1" box="[1242,1401,833,857]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="meladerma">
<emphasis id="4F82E240FFA2FFAEE2C7076DFC41FCA1" box="[1242,1401,833,857]" italics="true" pageId="1" pageNumber="438">C. meladerma</emphasis>
</taxonomicName>
, in having a dark gray head and body with black blotches (vs. black blotches absent). C
<emphasis id="4F82E240FFA2FFAEE223074AFDC8FC85" box="[1086,1264,869,893]" italics="true" pageId="1" pageNumber="438">
larias
<taxonomicName id="BAF645D1FFA2FFAEE29B0749FDC8FC85" box="[1158,1264,870,893]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="serniosus">serniosus</taxonomicName>
</emphasis>
differs from
<taxonomicName id="BAF645D1FFA2FFAEE39F0749F82DFC59" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="meladerma">
<emphasis id="4F82E240FFA2FFAEE39F0749F82DFC59" italics="true" pageId="1" pageNumber="438">C. meladerma</emphasis>
</taxonomicName>
in having a greater distance between the occipital process and the base of the first dorsal-fin ray (8.5% SL vs. 2.84.8), a smooth (vs. serrated) anterior edge of the pectoral spine, a longer pectoral fin (8.510.1% SL vs. 5.28.5) and a deeper caudal peduncle (8.48.6% SL vs. 5.37.3). The following combination of characters additionally diagnoses
<taxonomicName id="BAF645D1FFA2FFAEE78007D9FB12FBF5" box="[413,554,1014,1037]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="438" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA2FFAEE78007D9FB12FBF5" box="[413,554,1014,1037]" italics="true" pageId="1" pageNumber="438">C. serniosus</emphasis>
</taxonomicName>
from Southeast Asian congeners: occipital process length 1517% HL, head length 28.228.6% SL, head width 18.519.2% SL, head depth 13.013.7% SL, smooth anterior edge of pectoral spine, 67 dorsal-fin rays, body depth at anus 16.216.5% SL, and 57 total vertebrae.
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA2FFADE6DA004EFC6DF833" lastPageId="2" lastPageNumber="439" pageId="1" pageNumber="438" type="description">
<paragraph id="7D493E52FFA2FFAEE6DA004EFA58FA8E" blockId="1.[151,1437,689,2010]" pageId="1" pageNumber="438">
<emphasis id="4F82E240FFA2FFAEE6DA004EF861FB82" bold="true" box="[199,345,1121,1146]" pageId="1" pageNumber="438">Description.</emphasis>
Morphometric data in
<tableCitation id="30740BE9FFA2FFAEE47A004EFB86FB82" box="[615,702,1121,1146]" captionStart="TABLE 1" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1331,382,1758]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data for Clarias serniosus (n = 2)." httpUri="http://table.plazi.org/id/29896EDAFFA1FFADE68A0563FA36FE9B" pageId="1" pageNumber="438" tableUuid="29896EDAFFA1FFADE68A0563FA36FE9B">Table 1</tableCitation>
. Head depressed; dorsal profile slightly convex; ventral profile almost straight. Bony elements of dorsal surface of head covered with thick skin; bones and sutures not readily visible. Anterior pair of nostrils tubular, medial to maxillary barbel base. Posterior pair of nostrils bordered by nasal barbels anteriorly and fleshy membrane posteriorly; posteromedial to maxillary barbel base. Eye ovoid, horizontal axis longest, subcutaneous; located dorsolaterally on head. Anterior fontanel long, narrow (“knifeshaped” of
<bibRefCitation id="196743A3FFA2FFAEE70A013AF880FAD6" author="Teugels, G. G." box="[279,440,1301,1326]" firstAuthor="Teugels" journalOrPublisher="Annales du Musee Royal de l'Afrique Centrale (Zoologie)" pageId="1" pageNumber="438" pagination="1 - 199" part="247" refId="ref5228" refString="Teugels, G. G. (1986) A systematic revision of the African species of the genus Clarias (Pisces; Clariidae). Annales du Musee Royal de l'Afrique Centrale (Zoologie), 247, 1 - 199." title="A systematic revision of the African species of the genus Clarias (Pisces; Clariidae)" type="journal article" year="1986">Teugels, 1986</bibRefCitation>
); anterior tip reaching to line through anterior orbital margin. Occipital process rounded. Gill openings narrow, extending from dorsal-most point of pectoral-fin base to isthmus. Gill membranes free from isthmus but united to each other, with 8 (2) branchiostegal rays.
</paragraph>
<paragraph id="7D493E52FFA2FFAEE6DA01AEF803F942" blockId="1.[151,1437,689,2010]" pageId="1" pageNumber="438">Mouth subterminal, with fleshy, plicate lips. Oral teeth small and in irregular rows on all tooth-bearing surfaces. Premaxillary tooth band rectangular, with median notch on posterior edge. Dentary tooth band much narrower than premaxillary tooth band at symphysis, tapering laterally. Vomerine tooth band unpaired, continuous across midline, crescentic, smoothly arched along anterior and posterior margins. Premaxillary and dentary teeth viliform; vomerine teeth subgranular. Barbels in four pairs; long, slender, with thick fleshy bases. Maxillary barbel extending to middle of pectoral-fin base. Nasal barbel extending nearly to tip of supraoccipital process. Inner mandibular-barbel origin close to midline, extending to halfway between posterior orbital margin and base of pectoral spine. Outer mandibular barbel originating posterolateral of inner mandibular barbel, extending to base of pectoral spine.</paragraph>
<paragraph id="7D493E52FFA2FFAEE6DA02EAFB91F8B2" blockId="1.[151,1437,689,2010]" pageId="1" pageNumber="438">Body terete, becoming compressed towards caudal peduncle. Dorsal profile rising very gently from tip of snout to origin of dorsal fin, thereafter almost horizontal to end of caudal peduncle. Ventral profile slightly convex to middle of head, thereafter almost horizontal to end of caudal peduncle. Skin smooth. Lateral line complete, midlateral in position. Vertebrae 19+38=57 (2).</paragraph>
<paragraph id="7D493E52FFA2FFADE6DA037AFD83FEE5" blockId="1.[151,1437,689,2010]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="439" pageId="1" pageNumber="438">Dorsal fin with long base, spanning posterior three-quarters of body; with 67 (2) rays covered by thick layer of skin, without spine. Dorsal-fin margin straight, parallel to dorsal profile of body. Pectoral fin with small spine, sharply pointed at tip, and 8* (1) or 9 (1) rays. Anterior spine margin smooth; posterior spine margin smooth or with uneven asperities. Pectoral-fin margin straight anteriorly, convex posteriorly. Pelvic-fin origin at anterior third of body, with i,5 (2) rays and convex margin; tip of adpressed fin reaching base of first two or three anal-fin rays. Anus and urogenital openings located at vertical through middle of adpressed pelvic fin. Anal fin with long base, extending for posterior three-fifths of body, and 49* (1) or 54 (1) rays covered by thick layer of skin; margin straight and parallel to ventral edge of body. Caudal fin rounded, with i,7,6,i (2) principal rays.</paragraph>
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<paragraph id="7D493E52FFA1FFADE68A0563FA36FE9B" blockId="2.[151,782,331,355]" box="[151,782,331,355]" pageId="2" pageNumber="439">
<emphasis id="4F82E240FFA1FFADE68A0563F832FE9B" bold="true" box="[151,266,332,355]" pageId="2" pageNumber="439">TABLE 1.</emphasis>
Morphometric data for
<taxonomicName id="BAF645D1FFA1FFADE4170564FBFDFE9B" box="[522,709,331,355]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="439" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA1FFADE4170564FBFDFE9B" box="[522,709,331,355]" italics="true" pageId="2" pageNumber="439">Clarias serniosus</emphasis>
</taxonomicName>
(n=2).
</paragraph>
</caption>
<paragraph id="7D493E52FFA1FFADE5980551FDF5F926" pageId="2" pageNumber="439">
<table id="0FF6CCF2FFA10050E6820551FC0BF926" box="[159,1331,382,1758]" gridcols="3" gridrows="33" pageId="2" pageNumber="439">
<tr id="C3C63C10FFA10050E6820551FC0BFE4B" box="[159,1331,382,435]" gridrow="0" pageId="2" pageNumber="439" rowspan-0="1">
<th id="8017556CFFA10050E5980551FD00FE4B" box="[901,1080,382,435]" gridcol="1" gridrow="0" pageId="2" pageNumber="439">Holotype MHNG 2745.072</th>
<th id="8017556CFFA10050E2A90551FC0BFE4B" box="[1204,1331,382,435]" gridcol="2" gridrow="0" pageId="2" pageNumber="439">Paratype CMK 23382</th>
</tr>
<tr id="C3C63C10FFA10050E68205E9FC0BFE24" box="[159,1331,454,476]" gridrow="1" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68205E9FBA1FE24" box="[159,665,454,476]" gridcol="0" gridrow="1" pageId="2" pageNumber="439">Standard length</th>
<td id="8017556CFFA10050E59805E9FD00FE24" box="[901,1080,454,476]" gridcol="1" gridrow="1" pageId="2" pageNumber="439">141</td>
<td id="8017556CFFA10050E2A905E9FC0BFE24" box="[1204,1331,454,476]" gridcol="2" gridrow="1" pageId="2" pageNumber="439">161</td>
</tr>
<tr id="C3C63C10FFA10050E68205DFFC0BFDFE" box="[159,1331,496,518]" gridrow="2" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68205DFFC0BFDFE" box="[159,1331,496,518]" colspan="3" colspanRight="2" gridcol="0" gridrow="2" pageId="2" pageNumber="439">In % SL</th>
</tr>
<tr id="C3C63C10FFA10050E6820636FC0BFDD7" box="[159,1331,537,559]" gridrow="3" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820636FBA1FDD7" box="[159,665,537,559]" gridcol="0" gridrow="3" pageId="2" pageNumber="439">Predorsal length</th>
<td id="8017556CFFA10050E5980636FD00FDD7" box="[901,1080,537,559]" gridcol="1" gridrow="3" pageId="2" pageNumber="439">36.1</td>
<td id="8017556CFFA10050E2A90636FC0BFDD7" box="[1204,1331,537,559]" gridcol="2" gridrow="3" pageId="2" pageNumber="439">35.9</td>
</tr>
<tr id="C3C63C10FFA10050E682066DFC0BFDA0" box="[159,1331,578,600]" gridrow="4" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682066DFBA1FDA0" box="[159,665,578,600]" gridcol="0" gridrow="4" pageId="2" pageNumber="439">Preanal length</th>
<td id="8017556CFFA10050E598066DFD00FDA0" box="[901,1080,578,600]" gridcol="1" gridrow="4" pageId="2" pageNumber="439">51.4</td>
<td id="8017556CFFA10050E2A9066DFC0BFDA0" box="[1204,1331,578,600]" gridcol="2" gridrow="4" pageId="2" pageNumber="439">50.3</td>
</tr>
<tr id="C3C63C10FFA10050E6820643FC0BFD7A" box="[159,1331,620,642]" gridrow="5" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820643FBA1FD7A" box="[159,665,620,642]" gridcol="0" gridrow="5" pageId="2" pageNumber="439">Prepelvic length</th>
<td id="8017556CFFA10050E5980643FD00FD7A" box="[901,1080,620,642]" gridcol="1" gridrow="5" pageId="2" pageNumber="439">42.9</td>
<td id="8017556CFFA10050E2A90643FC0BFD7A" box="[1204,1331,620,642]" gridcol="2" gridrow="5" pageId="2" pageNumber="439">41.9</td>
</tr>
<tr id="C3C63C10FFA10050E68206BAFC0BFD53" box="[159,1331,661,683]" gridrow="6" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68206BAFBA1FD53" box="[159,665,661,683]" gridcol="0" gridrow="6" pageId="2" pageNumber="439">Prepectoral length</th>
<td id="8017556CFFA10050E59806BAFD00FD53" box="[901,1080,661,683]" gridcol="1" gridrow="6" pageId="2" pageNumber="439">21.9</td>
<td id="8017556CFFA10050E2A906BAFC0BFD53" box="[1204,1331,661,683]" gridcol="2" gridrow="6" pageId="2" pageNumber="439">20.3</td>
</tr>
<tr id="C3C63C10FFA10050E6820691FC0BFD2C" box="[159,1331,702,724]" gridrow="7" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820691FBA1FD2C" box="[159,665,702,724]" gridcol="0" gridrow="7" pageId="2" pageNumber="439">Length of dorsal-fin base</th>
<td id="8017556CFFA10050E5980691FD00FD2C" box="[901,1080,702,724]" gridcol="1" gridrow="7" pageId="2" pageNumber="439">64.4</td>
<td id="8017556CFFA10050E2A90691FC0BFD2C" box="[1204,1331,702,724]" gridcol="2" gridrow="7" pageId="2" pageNumber="439">66.8</td>
</tr>
<tr id="C3C63C10FFA10050E68206C7FC0BFD06" box="[159,1331,744,766]" gridrow="8" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68206C7FBA1FD06" box="[159,665,744,766]" gridcol="0" gridrow="8" pageId="2" pageNumber="439">Anal-fin length</th>
<td id="8017556CFFA10050E59806C7FD00FD06" box="[901,1080,744,766]" gridcol="1" gridrow="8" pageId="2" pageNumber="439">50.1</td>
<td id="8017556CFFA10050E2A906C7FC0BFD06" box="[1204,1331,744,766]" gridcol="2" gridrow="8" pageId="2" pageNumber="439">46.2</td>
</tr>
<tr id="C3C63C10FFA10050E682073EFC0BFCDF" box="[159,1331,785,807]" gridrow="9" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682073EFBA1FCDF" box="[159,665,785,807]" gridcol="0" gridrow="9" pageId="2" pageNumber="439">Pelvic-fin length</th>
<td id="8017556CFFA10050E598073EFD00FCDF" box="[901,1080,785,807]" gridcol="1" gridrow="9" pageId="2" pageNumber="439">10.7</td>
<td id="8017556CFFA10050E2A9073EFC0BFCDF" box="[1204,1331,785,807]" gridcol="2" gridrow="9" pageId="2" pageNumber="439">10.1</td>
</tr>
<tr id="C3C63C10FFA10050E6820715FC0BFCA8" box="[159,1331,826,848]" gridrow="10" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820715FBA1FCA8" box="[159,665,826,848]" gridcol="0" gridrow="10" pageId="2" pageNumber="439">Pectoral-fin length</th>
<td id="8017556CFFA10050E5980715FD00FCA8" box="[901,1080,826,848]" gridcol="1" gridrow="10" pageId="2" pageNumber="439">13.9</td>
<td id="8017556CFFA10050E2A90715FC0BFCA8" box="[1204,1331,826,848]" gridcol="2" gridrow="10" pageId="2" pageNumber="439">13.4</td>
</tr>
<tr id="C3C63C10FFA10050E682074BFC0BFC82" box="[159,1331,868,890]" gridrow="11" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682074BFBA1FC82" box="[159,665,868,890]" gridcol="0" gridrow="11" pageId="2" pageNumber="439">Pectoral-spine length</th>
<td id="8017556CFFA10050E598074BFD00FC82" box="[901,1080,868,890]" gridcol="1" gridrow="11" pageId="2" pageNumber="439">9.7</td>
<td id="8017556CFFA10050E2A9074BFC0BFC82" box="[1204,1331,868,890]" gridcol="2" gridrow="11" pageId="2" pageNumber="439">10.5</td>
</tr>
<tr id="C3C63C10FFA10050E68207A2FC0BFC5B" box="[159,1331,909,931]" gridrow="12" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68207A2FBA1FC5B" box="[159,665,909,931]" gridcol="0" gridrow="12" pageId="2" pageNumber="439">Caudal-fin length</th>
<td id="8017556CFFA10050E59807A2FD00FC5B" box="[901,1080,909,931]" gridcol="1" gridrow="12" pageId="2" pageNumber="439">14.9</td>
<td id="8017556CFFA10050E2A907A2FC0BFC5B" box="[1204,1331,909,931]" gridcol="2" gridrow="12" pageId="2" pageNumber="439">14.8</td>
</tr>
<tr id="C3C63C10FFA10050E6820799FC0BFC34" box="[159,1331,950,972]" gridrow="13" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820799FBA1FC34" box="[159,665,950,972]" gridcol="0" gridrow="13" pageId="2" pageNumber="439">Distance between occipital process and dorsal fin</th>
<td id="8017556CFFA10050E5980799FD00FC34" box="[901,1080,950,972]" gridcol="1" gridrow="13" pageId="2" pageNumber="439">8.5</td>
<td id="8017556CFFA10050E2A90799FC0BFC34" box="[1204,1331,950,972]" gridcol="2" gridrow="13" pageId="2" pageNumber="439">8.5</td>
</tr>
<tr id="C3C63C10FFA10050E68207CFFC0BFC0E" box="[159,1331,992,1014]" gridrow="14" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68207CFFBA1FC0E" box="[159,665,992,1014]" gridcol="0" gridrow="14" pageId="2" pageNumber="439">Caudal peduncle depth</th>
<td id="8017556CFFA10050E59807CFFD00FC0E" box="[901,1080,992,1014]" gridcol="1" gridrow="14" pageId="2" pageNumber="439">8.4</td>
<td id="8017556CFFA10050E2A907CFFC0BFC0E" box="[1204,1331,992,1014]" gridcol="2" gridrow="14" pageId="2" pageNumber="439">8.6</td>
</tr>
<tr id="C3C63C10FFA10050E6820026FC0BFBE7" box="[159,1331,1033,1055]" gridrow="15" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820026FBA1FBE7" box="[159,665,1033,1055]" gridcol="0" gridrow="15" pageId="2" pageNumber="439">Body depth at anus</th>
<td id="8017556CFFA10050E5980026FD00FBE7" box="[901,1080,1033,1055]" gridcol="1" gridrow="15" pageId="2" pageNumber="439">16.5</td>
<td id="8017556CFFA10050E2A90026FC0BFBE7" box="[1204,1331,1033,1055]" gridcol="2" gridrow="15" pageId="2" pageNumber="439">16.2</td>
</tr>
<tr id="C3C63C10FFA10050E682001DFC0BFBB0" box="[159,1331,1074,1096]" gridrow="16" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682001DFBA1FBB0" box="[159,665,1074,1096]" gridcol="0" gridrow="16" pageId="2" pageNumber="439">Head length</th>
<td id="8017556CFFA10050E598001DFD00FBB0" box="[901,1080,1074,1096]" gridcol="1" gridrow="16" pageId="2" pageNumber="439">28.6</td>
<td id="8017556CFFA10050E2A9001DFC0BFBB0" box="[1204,1331,1074,1096]" gridcol="2" gridrow="16" pageId="2" pageNumber="439">28.2</td>
</tr>
<tr id="C3C63C10FFA10050E6820073FC0BFB8A" box="[159,1331,1116,1138]" gridrow="17" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820073FBA1FB8A" box="[159,665,1116,1138]" gridcol="0" gridrow="17" pageId="2" pageNumber="439">Head width</th>
<td id="8017556CFFA10050E5980073FD00FB8A" box="[901,1080,1116,1138]" gridcol="1" gridrow="17" pageId="2" pageNumber="439">19.2</td>
<td id="8017556CFFA10050E2A90073FC0BFB8A" box="[1204,1331,1116,1138]" gridcol="2" gridrow="17" pageId="2" pageNumber="439">18.5</td>
</tr>
<tr id="C3C63C10FFA10050E68200AAFC0BFB63" box="[159,1331,1157,1179]" gridrow="18" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68200AAFBA1FB63" box="[159,665,1157,1179]" gridcol="0" gridrow="18" pageId="2" pageNumber="439">Head depth</th>
<td id="8017556CFFA10050E59800AAFD00FB63" box="[901,1080,1157,1179]" gridcol="1" gridrow="18" pageId="2" pageNumber="439">13.7</td>
<td id="8017556CFFA10050E2A900AAFC0BFB63" box="[1204,1331,1157,1179]" gridcol="2" gridrow="18" pageId="2" pageNumber="439">13.0</td>
</tr>
<tr id="C3C63C10FFA10050E6820081FC0BFB3C" box="[159,1331,1198,1220]" gridrow="19" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820081FC0BFB3C" box="[159,1331,1198,1220]" colspan="3" colspanRight="2" gridcol="0" gridrow="19" pageId="2" pageNumber="439">In % HL</th>
</tr>
<tr id="C3C63C10FFA10050E68200F7FC0BFB16" box="[159,1331,1240,1262]" gridrow="20" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68200F7FBA1FB16" box="[159,665,1240,1262]" gridcol="0" gridrow="20" pageId="2" pageNumber="439">Snout length</th>
<td id="8017556CFFA10050E59800F7FD00FB16" box="[901,1080,1240,1262]" gridcol="1" gridrow="20" pageId="2" pageNumber="439">30</td>
<td id="8017556CFFA10050E2A900F7FC0BFB16" box="[1204,1331,1240,1262]" gridcol="2" gridrow="20" pageId="2" pageNumber="439">30</td>
</tr>
<tr id="C3C63C10FFA10050E682012EFC0BFAEF" box="[159,1331,1281,1303]" gridrow="21" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682012EFBA1FAEF" box="[159,665,1281,1303]" gridcol="0" gridrow="21" pageId="2" pageNumber="439">Interorbital distance</th>
<td id="8017556CFFA10050E598012EFD00FAEF" box="[901,1080,1281,1303]" gridcol="1" gridrow="21" pageId="2" pageNumber="439">45</td>
<td id="8017556CFFA10050E2A9012EFC0BFAEF" box="[1204,1331,1281,1303]" gridcol="2" gridrow="21" pageId="2" pageNumber="439">45</td>
</tr>
<tr id="C3C63C10FFA10050E6820105FC0BFAB8" box="[159,1331,1322,1344]" gridrow="22" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820105FBA1FAB8" box="[159,665,1322,1344]" gridcol="0" gridrow="22" pageId="2" pageNumber="439">Eye diameter</th>
<td id="8017556CFFA10050E5980105FD00FAB8" box="[901,1080,1322,1344]" gridcol="1" gridrow="22" pageId="2" pageNumber="439">9</td>
<td id="8017556CFFA10050E2A90105FC0BFAB8" box="[1204,1331,1322,1344]" gridcol="2" gridrow="22" pageId="2" pageNumber="439">8</td>
</tr>
<tr id="C3C63C10FFA10050E682017BFC0BFA92" box="[159,1331,1364,1386]" gridrow="23" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682017BFBA1FA92" box="[159,665,1364,1386]" gridcol="0" gridrow="23" pageId="2" pageNumber="439">Nasal barbel length</th>
<td id="8017556CFFA10050E598017BFD00FA92" box="[901,1080,1364,1386]" gridcol="1" gridrow="23" pageId="2" pageNumber="439">59</td>
<td id="8017556CFFA10050E2A9017BFC0BFA92" box="[1204,1331,1364,1386]" gridcol="2" gridrow="23" pageId="2" pageNumber="439">51</td>
</tr>
<tr id="C3C63C10FFA10050E6820152FC0BFA6B" box="[159,1331,1405,1427]" gridrow="24" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820152FBA1FA6B" box="[159,665,1405,1427]" gridcol="0" gridrow="24" pageId="2" pageNumber="439">Maxillary barbel length</th>
<td id="8017556CFFA10050E5980152FD00FA6B" box="[901,1080,1405,1427]" gridcol="1" gridrow="24" pageId="2" pageNumber="439">78</td>
<td id="8017556CFFA10050E2A90152FC0BFA6B" box="[1204,1331,1405,1427]" gridcol="2" gridrow="24" pageId="2" pageNumber="439">77</td>
</tr>
<tr id="C3C63C10FFA10050E6820189FC0BFA44" box="[159,1331,1446,1468]" gridrow="25" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820189FBA1FA44" box="[159,665,1446,1468]" gridcol="0" gridrow="25" pageId="2" pageNumber="439">Inner mandibular barbel length</th>
<td id="8017556CFFA10050E5980189FD00FA44" box="[901,1080,1446,1468]" gridcol="1" gridrow="25" pageId="2" pageNumber="439">60</td>
<td id="8017556CFFA10050E2A90189FC0BFA44" box="[1204,1331,1446,1468]" gridcol="2" gridrow="25" pageId="2" pageNumber="439">46</td>
</tr>
<tr id="C3C63C10FFA10050E68201FFFC0BFA1E" box="[159,1331,1488,1510]" gridrow="26" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68201FFFBA1FA1E" box="[159,665,1488,1510]" gridcol="0" gridrow="26" pageId="2" pageNumber="439">Outer mandibular barbel length</th>
<td id="8017556CFFA10050E59801FFFD00FA1E" box="[901,1080,1488,1510]" gridcol="1" gridrow="26" pageId="2" pageNumber="439">73</td>
<td id="8017556CFFA10050E2A901FFFC0BFA1E" box="[1204,1331,1488,1510]" gridcol="2" gridrow="26" pageId="2" pageNumber="439">70</td>
</tr>
<tr id="C3C63C10FFA10050E68201D6FC0BF9F7" box="[159,1331,1529,1551]" gridrow="27" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68201D6FBA1F9F7" box="[159,665,1529,1551]" gridcol="0" gridrow="27" pageId="2" pageNumber="439">Front fontanel length</th>
<td id="8017556CFFA10050E59801D6FD00F9F7" box="[901,1080,1529,1551]" gridcol="1" gridrow="27" pageId="2" pageNumber="439">22</td>
<td id="8017556CFFA10050E2A901D6FC0BF9F7" box="[1204,1331,1529,1551]" gridcol="2" gridrow="27" pageId="2" pageNumber="439">21</td>
</tr>
<tr id="C3C63C10FFA10050E682020DFC0BF9C0" box="[159,1331,1570,1592]" gridrow="28" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682020DFBA1F9C0" box="[159,665,1570,1592]" gridcol="0" gridrow="28" pageId="2" pageNumber="439">Front fontanel width</th>
<td id="8017556CFFA10050E598020DFD00F9C0" box="[901,1080,1570,1592]" gridcol="1" gridrow="28" pageId="2" pageNumber="439">3</td>
<td id="8017556CFFA10050E2A9020DFC0BF9C0" box="[1204,1331,1570,1592]" gridcol="2" gridrow="28" pageId="2" pageNumber="439">5</td>
</tr>
<tr id="C3C63C10FFA10050E6820263FC0BF99A" box="[159,1331,1612,1634]" gridrow="29" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E6820263FBA1F99A" box="[159,665,1612,1634]" gridcol="0" gridrow="29" pageId="2" pageNumber="439">Occipital fontanel length</th>
<td id="8017556CFFA10050E5980263FD00F99A" box="[901,1080,1612,1634]" gridcol="1" gridrow="29" pageId="2" pageNumber="439">19</td>
<td id="8017556CFFA10050E2A90263FC0BF99A" box="[1204,1331,1612,1634]" gridcol="2" gridrow="29" pageId="2" pageNumber="439">13</td>
</tr>
<tr id="C3C63C10FFA10050E682025AFC0BF973" box="[159,1331,1653,1675]" gridrow="30" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E682025AFBA1F973" box="[159,665,1653,1675]" gridcol="0" gridrow="30" pageId="2" pageNumber="439">Occipital fontanel width</th>
<td id="8017556CFFA10050E598025AFD00F973" box="[901,1080,1653,1675]" gridcol="1" gridrow="30" pageId="2" pageNumber="439">6</td>
<td id="8017556CFFA10050E2A9025AFC0BF973" box="[1204,1331,1653,1675]" gridcol="2" gridrow="30" pageId="2" pageNumber="439">4</td>
</tr>
<tr id="C3C63C10FFA10050E68202B1FC0BF94C" box="[159,1331,1694,1716]" gridrow="31" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68202B1FBA1F94C" box="[159,665,1694,1716]" gridcol="0" gridrow="31" pageId="2" pageNumber="439">Occipital process length</th>
<td id="8017556CFFA10050E59802B1FD00F94C" box="[901,1080,1694,1716]" gridcol="1" gridrow="31" pageId="2" pageNumber="439">17</td>
<td id="8017556CFFA10050E2A902B1FC0BF94C" box="[1204,1331,1694,1716]" gridcol="2" gridrow="31" pageId="2" pageNumber="439">15</td>
</tr>
<tr id="C3C63C10FFA10050E68202E7FC0BF926" box="[159,1331,1736,1758]" gridrow="32" pageId="2" pageNumber="439">
<th id="8017556CFFA10050E68202E7FBA1F926" box="[159,665,1736,1758]" gridcol="0" gridrow="32" pageId="2" pageNumber="439">Occipital process width</th>
<td id="8017556CFFA10050E59802E7FD00F926" box="[901,1080,1736,1758]" gridcol="1" gridrow="32" pageId="2" pageNumber="439">28</td>
<td id="8017556CFFA10050E2A902E7FC0BF926" box="[1204,1331,1736,1758]" gridcol="2" gridrow="32" pageId="2" pageNumber="439">30</td>
</tr>
</table>
</paragraph>
<paragraph id="7D493E52FFA1FFADE6DA030DFC25F85F" blockId="2.[151,1436,1826,1995]" pageId="2" pageNumber="439">
<emphasis id="4F82E240FFA1FFADE6DA030DF877F8C3" bold="true" box="[199,335,1826,1851]" pageId="2" pageNumber="439">Coloration.</emphasis>
In 70% alcohol: dorsal and lateral surfaces of head and body dark gray, fading to pale gray on ventral surfaces. Dorsal and lateral surfaces of head and body with irregular, small black blotches randomly distributed throughout. Dorsal, anal and caudal fins dark gray. Pectoral-fin rays dark gray, with hyaline interradial membranes. Pelvic fins hyaline. Barbels and pectoral spines dark gray dorsally and light gray ventrally.
</paragraph>
<paragraph id="7D493E52FFA1FFADE6DA039CFC6DF833" blockId="2.[151,1436,1826,1995]" box="[199,1365,1970,1995]" pageId="2" pageNumber="439">In life: Body and fins dark gray, almost black. Blotches black, distinct but not conspicuously contrasted.</paragraph>
</subSubSection>
<caption id="29896EDAFFA0FFACE68A000DF827FBAF" httpUri="https://zenodo.org/record/224375/files/figure.png" pageId="3" pageNumber="440" targetBox="[151,1436,193,1036]" targetPageId="3">
<paragraph id="7D493E52FFA0FFACE68A000DF827FBAF" blockId="3.[151,1435,1058,1111]" pageId="3" pageNumber="440">
<emphasis id="4F82E240FFA0FFACE68A000DF82AFBCF" bold="true" box="[151,274,1058,1080]" pageId="3" pageNumber="440">FIGURE 1.</emphasis>
<taxonomicName id="BAF645D1FFA0FFACE705000DF8F1FBC0" box="[280,457,1058,1080]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE705000DF8F1FBC0" box="[280,457,1058,1080]" italics="true" pageId="3" pageNumber="440">Clarias serniosus</emphasis>
</taxonomicName>
, holotype, MHNG 2745.072, 141 mm SL; Laos: Bolavens Plateau, Xe Pian drainage. Dorsal and lateral views.
</paragraph>
</caption>
<subSubSection id="35EC6DD9FFA0FFACE6DA0051FC1BFB42" pageId="3" pageNumber="440" type="distribution">
<paragraph id="7D493E52FFA0FFACE6DA0051FC1BFB42" blockId="3.[151,1437,1150,1895]" pageId="3" pageNumber="440">
<emphasis id="4F82E240FFA0FFACE6DA0051F858FB6F" bold="true" box="[199,352,1150,1175]" pageId="3" pageNumber="440">Distribution.</emphasis>
<taxonomicName id="BAF645D1FFA0FFACE77A0051FB14FB6E" box="[359,556,1150,1174]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE77A0051FB14FB6E" box="[359,556,1150,1174]" italics="true" pageId="3" pageNumber="440">Clarias serniosus</emphasis>
</taxonomicName>
is known only from its
<typeStatus id="A24D80F0FFA0FFACE52A0050FA5FFB6F" box="[823,871,1151,1175]" pageId="3" pageNumber="440">type</typeStatus>
locality in the Xe Pian drainage (a tributary of the Xe Kong, itself a tributary of the Mekong) flowing from the Bolavens Plateau in southern
<collectingCountry id="05E17EC2FFA0FFACE296008CFDF9FB42" box="[1163,1217,1187,1210]" name="Laos" pageId="3" pageNumber="440">Laos</collectingCountry>
(
<figureCitation id="E5CD22D7FFA0FFACE2CC008DFC2EFB42" box="[1233,1302,1186,1211]" captionStart="FIGURE 2" captionStartId="4.[151,250,1960,1982]" captionTargetBox="[151,1436,193,1939]" captionTargetId="figure@4.[151,1436,193,1939]" captionTargetPageId="4" captionText="FIGURE 2. Map showing type locality of Clarias serniosus." httpUri="https://zenodo.org/record/224376/files/figure.png" pageId="3" pageNumber="440">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA0FFACE6DA00E9FD1BFA6B" pageId="3" pageNumber="440" type="biology_ecology">
<paragraph id="7D493E52FFA0FFACE6DA00E9FD1BFA6B" blockId="3.[151,1437,1150,1895]" pageId="3" pageNumber="440">
<emphasis id="4F82E240FFA0FFACE6DA00E9F8FAFB27" bold="true" box="[199,450,1222,1247]" pageId="3" pageNumber="440">Habitat and ecology.</emphasis>
The Bolavens Plateau is a large basalt plateau of southern
<collectingCountry id="05E17EC2FFA0FFACE26B00E8FD97FB26" box="[1142,1199,1223,1246]" name="Laos" pageId="3" pageNumber="440">Laos</collectingCountry>
, about 50 by
<quantity id="BA0E93B7FFA0FFACE34F00E8FCA4FB27" box="[1362,1436,1222,1247]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="440" unit="km" value="80.0">80 km</quantity>
(
<bibRefCitation id="196743A3FFA0FFACE68300C5F89BFAFA" author="Sanematsu, K. &amp; Moriyama, T. &amp; Sotouky, L. &amp; Watanabe, Y." box="[158,419,1258,1282]" etAl="et al." firstAuthor="Sanematsu" journalOrPublisher="Bulletin of the Geological Survey of Japan" pageId="3" pageNumber="440" pagination="105 - 129" part="62" refId="ref5074" refString="Sanematsu, K., Moriyama, T., Sotouky, L. &amp; Watanabe, Y. (2011) Laterization of basalts and sandstone associated with the enrichment of Al, Ga and Sc in the Bolaven Plateau, southern Laos. Bulletin of the Geological Survey of Japan, 62, 105 - 129." title="Laterization of basalts and sandstone associated with the enrichment of Al, Ga and Sc in the Bolaven Plateau, southern Laos" type="journal article" year="2011">
Sanematsu
<emphasis id="4F82E240FFA0FFACE73C00C4F862FAFA" box="[289,346,1258,1282]" italics="true" pageId="3" pageNumber="440">et al.</emphasis>
, 2011
</bibRefCitation>
), 1000-1300 masl, surrounded by the Mekong and Xe Kong floodplains (about 100 masl). It has very steep slopes, especially along its southern and eastern edges. It has a very wet climate with heavy rainfall in all seasons. Most rivers leave the plateau through high and dramatic series of waterfalls and gorges. The Houay Xoy, a tributary of the Xe Pian, itself tributary of the Xe Kong, is one of the few streams without high falls (based on 1:100,000 maps and air photographs; ground survey was not possible).
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA0FFACE6DA01B1FB77F8E7" pageId="3" pageNumber="440" type="discussion">
<paragraph id="7D493E52FFA0FFACE6DA01B1FB77F8E7" blockId="3.[151,1437,1150,1895]" pageId="3" pageNumber="440">
The
<typeStatus id="A24D80F0FFA0FFACE6E701B0F812FA4F" box="[250,298,1439,1463]" pageId="3" pageNumber="440">type</typeStatus>
series of
<taxonomicName id="BAF645D1FFA0FFACE78401B0FB1DFA4E" box="[409,549,1439,1462]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE78401B0FB1DFA4E" box="[409,549,1439,1462]" italics="true" pageId="3" pageNumber="440">C. serniosus</emphasis>
</taxonomicName>
was obtained in a small tributary of the Houay Xoy on which a small wooden weir had been constructed in an area recently logged and converted for agriculture. In the sampling area, the stream was about
<quantity id="BA0E93B7FFA0FFACE70801C8F867FA06" box="[277,351,1511,1535]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="3" pageNumber="440" unit="m" value="3.0" valueMax="4.0" valueMin="2.0">24 m</quantity>
wide, and less than
<quantity id="BA0E93B7FFA0FFACE44F01C8FBA2FA06" box="[594,666,1511,1535]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="440" unit="cm" value="70.0">70 cm</quantity>
deep (at the beginning of the low-water season). The bottom was almost exclusively soft sand, probably the result of erosion and sedimentation. About
<quantity id="BA0E93B7FFA0FFACE2660225FDFDF9DA" box="[1147,1221,1546,1570]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="440" unit="m" value="100.0">100 m</quantity>
of the stream was sampled with electricity. The two individuals of
<taxonomicName id="BAF645D1FFA0FFACE4D10200FA62F9BE" box="[716,858,1583,1606]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE4D10200FA62F9BE" box="[716,858,1583,1606]" italics="true" pageId="3" pageNumber="440">C. serniosus</emphasis>
</taxonomicName>
were obtained in a shaded area of very degraded forest. Only three other fish species were present:
<taxonomicName id="BAF645D1FFA0FFACE4FF027CFAA8F992" box="[738,912,1618,1642]" class="Actinopterygii" family="Cyprinidae" genus="Devario" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="gibber">
<emphasis id="4F82E240FFA0FFACE4FF027CFAA8F992" box="[738,912,1618,1642]" italics="true" pageId="3" pageNumber="440">Devario gibber</emphasis>
</taxonomicName>
,
<taxonomicName id="BAF645D1FFA0FFACE5BC027DFD56F992" box="[929,1134,1618,1642]" class="Actinopterygii" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="clatrata">
<emphasis id="4F82E240FFA0FFACE5BC027DFD56F992" box="[929,1134,1618,1642]" italics="true" pageId="3" pageNumber="440">Schistura clatrata</emphasis>
</taxonomicName>
and
<taxonomicName id="BAF645D1FFA0FFACE2B7027DFC67F992" box="[1194,1375,1618,1642]" class="Actinopterygii" family="Channidae" genus="Channa" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="gachua">
<emphasis id="4F82E240FFA0FFACE2B7027DFC67F992" box="[1194,1375,1618,1642]" italics="true" pageId="3" pageNumber="440">Channa gachua</emphasis>
</taxonomicName>
. The small number of species is a common situation of disturbed areas and these three species are common species, often observed in degraded habitats with running water, and they are known throughout the plateau as well as downstream.
<taxonomicName id="BAF645D1FFA0FFACE7330291F8CBF92E" box="[302,499,1726,1750]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE7330291F8CBF92E" box="[302,499,1726,1750]" italics="true" pageId="3" pageNumber="440">Clarias serniosus</emphasis>
</taxonomicName>
is likely to be present also downstream of the plateau in the Xe Kong drainage. On the day of capture, water temperature was estimated to be 1015°C (air temperature fluctuated between 6°C at night and about 25°C in the afternoon).
</paragraph>
</subSubSection>
<subSubSection id="35EC6DD9FFA0FFACE6DA0305FA6CF89F" pageId="3" pageNumber="440" type="etymology">
<paragraph id="7D493E52FFA0FFACE6DA0305FA6CF89F" blockId="3.[151,1437,1150,1895]" pageId="3" pageNumber="440">
<emphasis id="4F82E240FFA0FFACE6DA0305F876F8BB" bold="true" box="[199,334,1834,1859]" pageId="3" pageNumber="440">Etymology.</emphasis>
The specific epithet is the Latin adjective
<taxonomicName id="BAF645D1FFA0FFACE5360304FAADF8BA" box="[811,917,1835,1858]" class="Actinopterygii" family="Clariidae" genus="Clarias" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="440" phylum="Chordata" rank="species" species="serniosus">
<emphasis id="4F82E240FFA0FFACE5360304FAADF8BA" box="[811,917,1835,1858]" italics="true" pageId="3" pageNumber="440">serniosus</emphasis>
</taxonomicName>
, meaning covered with an eruption or scabby, in allusion to the color pattern of small, irregular dark patches.
</paragraph>
</subSubSection>
</treatment>
</document>