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<document id="CB6375EC1646EF2F451CFF5B4A383D1B" ID-CLB-Dataset="31436" ID-DOI="10.11646/zootaxa.4085.2.1" ID-GBIF-Dataset="ba498adf-4dec-4fa3-8213-9d4d8c3b9dd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052535" ID-ZooBank="E41CB99C-5177-47A7-A424-2453D27E48F0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510873310154" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2016" docId="F50D3F20FFB9D351EBE6F94FFB97FBFB" docLanguage="en" docName="zootaxa.4085.2.1.pdf" docOrigin="Zootaxa 4085 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Prionocyphon uncatus Zwick 2016, n. sp." docType="treatment" docVersion="9" lastPageNumber="174" masterDocId="09344758FFACD346EB71FFF6FFF9FFE9" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae" masterLastPageNumber="198" masterPageNumber="151" pageNumber="172" updateTime="1698499212072" updateUser="ExternalLinkService">
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<mods:title id="342A0DF7C4DE7C36B2B1D4F9B2CA2EE1">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<treatment id="F50D3F20FFB9D351EBE6F94FFB97FBFB" ID-DOI="http://doi.org/10.5281/zenodo.6076952" ID-GBIF-Taxon="137308974" ID-Zenodo-Dep="6076952" LSID="urn:lsid:plazi:treatment:F50D3F20FFB9D351EBE6F94FFB97FBFB" httpUri="http://treatment.plazi.org/id/F50D3F20FFB9D351EBE6F94FFB97FBFB" lastPageId="23" lastPageNumber="174" pageId="21" pageNumber="172">
<subSubSection id="35BEDDBDFFB9D353EBE6F94FFE73F91D" pageId="21" pageNumber="172" type="nomenclature">
<paragraph id="7D1B8E36FFB9D353EBE6F94FFE03F93A" blockId="21.[151,506,1721,1781]" box="[151,506,1721,1747]" pageId="21" pageNumber="172">
<heading id="2653395AFFB9D353EBE6F94FFE03F93A" bold="true" box="[151,506,1721,1747]" fontSize="11" level="1" pageId="21" pageNumber="172" reason="1">
<emphasis id="4FD05224FFB9D353EBE6F94FFE03F93A" bold="true" box="[151,506,1721,1747]" pageId="21" pageNumber="172">
<emphasis id="4FD05224FFB9D353EBE6F94FFE49F93A" bold="true" box="[151,432,1721,1747]" italics="true" pageId="21" pageNumber="172">
<taxonomicName id="BAA4F5B5FFB9D353EBE6F94FFE53F93A" authority="Zwick, 2016" authorityName="Zwick" authorityYear="2016" box="[151,426,1721,1747]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="172" phylum="Arthropoda" rank="species" species="uncatus" status="sp. nov.">Prionocyphon uncatus</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="54E3EF5FFFB9D353EAC9F94FFE03F93A" box="[440,506,1721,1747]" pageId="21" pageNumber="172" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="7D1B8E36FFB9D353EBE6F92AFE73F91D" blockId="21.[151,506,1721,1781]" box="[151,394,1756,1781]" pageId="21" pageNumber="172">
(Figs 4853,
<tableCitation id="3026BB8DFFB9D353EA5CF92AFE78F91D" box="[301,385,1756,1781]" captionStart="TABLE 2" captionStartId="17.[151,239,152,175]" captionText="TABLE 2. Character definitions and character matrix for the European type species, Prionocyphon serricornis, and most Australian Prionocyphon species. Asterisked names: not seen, information from Watts (2010 b). Not included: P. kurandaensis Watts, P. microflavidus Watts, and P. ocellatus Watts (not seen, insufficient information), and P. insolitus Watts, P. lamingtonensis Watts and P. macroflavidus Watts which are misplaced in Prionocyphon (Watts 2014)." pageId="21" pageNumber="172">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFFB9D353EBE6F8D2FD30F86D" pageId="21" pageNumber="172" type="materials_examined">
<paragraph id="7D1B8E36FFB9D353EBE6F8D2FB0AF8D5" blockId="21.[151,1436,1828,2033]" box="[151,1267,1828,1853]" pageId="21" pageNumber="172">
<emphasis id="4FD05224FFB9D353EBE6F8D2FD86F8D4" bold="true" box="[151,639,1828,1853]" pageId="21" pageNumber="172">
<typeStatus id="A21F3094FFB9D353EBE6F8D2FF2BF8D4" box="[151,210,1828,1853]" pageId="21" pageNumber="172">Type</typeStatus>
material.
<typeStatus id="A21F3094FFB9D353EA3DF8D2FE40F8D4" box="[332,441,1828,1853]" pageId="21" pageNumber="172" type="holotype">Holotype</typeStatus>
<specimenCount id="6BA245BFFFB9D353EACEF8D2FE2DF8D4" box="[447,468,1828,1853]" pageId="21" pageNumber="172" type="male"></specimenCount>
,
<specimenCount id="6BA245BFFFB9D353EA93F8D2FD86F8D4" box="[482,639,1828,1853]" pageId="21" pageNumber="172" type="female" typeStatus="paratype">1 paratype ♀</specimenCount>
</emphasis>
: Black Mtn, light trap,
<emphasis id="4FD05224FFB9D353E8F7F8D2FC46F8D4" bold="true" box="[902,959,1828,1853]" pageId="21" pageNumber="172">
<collectionCode id="1BB516F3FFB9D353E8F7F8D2FC46F8D4" box="[902,959,1828,1853]" pageId="21" pageNumber="172">ACT</collectionCode>
</emphasis>
,
<date id="091AA8F6FFB9D353E8BCF8D2FBDAF8D4" box="[973,1059,1828,1853]" pageId="21" pageNumber="172" value="69-01-13">13.1.69</date>
, S.Misko (
<collectionCode id="1BB516F3FFB9D353EFECF8D2FB11F8D5" box="[1181,1256,1828,1852]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="21" pageNumber="172">ANIC</collectionCode>
).
</paragraph>
<paragraph id="7D1B8E36FFB9D353EBB6F8B1FD30F86D" blockId="21.[151,1436,1828,2033]" pageId="21" pageNumber="172">
<emphasis id="4FD05224FFB9D353EBB6F8B1FE3FF889" bold="true" box="[199,454,1863,1888]" pageId="21" pageNumber="172">Additional paratypes</emphasis>
: 1♂: Black Mtn, ACT,
<date id="091AA8F6FFB9D353E9AAF8B1FCB5F889" box="[731,844,1863,1888]" pageId="21" pageNumber="172" value="1965-02-22">22.ii.1965</date>
T.F.B. Common; 1♂: ibidem,
<date id="091AA8F6FFB9D353EFDFF8B1FAEBF889" box="[1198,1298,1863,1888]" pageId="21" pageNumber="172" value="68-11-29">29.11.68</date>
, M.S.Upton (all ANIC). 1♂: Mittagong,
<emphasis id="4FD05224FFB9D353EAA2F89AFDDCF86C" bold="true" box="[467,549,1900,1925]" pageId="21" pageNumber="172">N.S.W.</emphasis>
Lea (SAMA).
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFFB9D351EBB6F879FDD2FCC3" lastPageId="23" lastPageNumber="174" pageId="21" pageNumber="172" type="description">
<paragraph id="7D1B8E36FFB9D350EBB6F879FEE8F808" blockId="21.[151,1436,1828,2033]" lastBlockId="22.[151,1435,1934,2017]" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172">
<emphasis id="4FD05224FFB9D353EBB6F879FEDFF841" bold="true" box="[199,294,1935,1960]" pageId="21" pageNumber="172">Habitus</emphasis>
. Stout, compact, rather convex, widest before midlength, narrowing a little caudally, almost dropshaped. BL
<quantity id="BA5C23D3FFB9D353EA59F842FE41F825" box="[296,440,1972,1997]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="21" pageNumber="172" unit="mm" value="2.35" valueMax="2.6" valueMin="2.1">2.12.6 mm</quantity>
, BL/BW ~1.55, HCW equals ~60% of BW. Dorsal side, base of antennae and legs uniformly brownish, head darker than rest, flagellum infuscate. Punctures on head and pronotum very fine, fine on
<emphasis id="4FD05224FFBAD350EBE6F878FEAFF84D" bold="true" box="[151,342,1934,1956]" pageId="22" pageNumber="173">FIGURES 4853.</emphasis>
<taxonomicName id="BAA4F5B5FFBAD350EA2CF878FDBAF84D" authority="Zwick, 2016" authorityName="Zwick" authorityYear="2016" box="[349,579,1934,1956]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="173" phylum="Arthropoda" rank="species" species="uncatus" status="sp. nov.">
<emphasis id="4FD05224FFBAD350EA2CF878FDBAF84D" box="[349,579,1934,1956]" italics="true" pageId="22" pageNumber="173">Prionocyphon uncatus</emphasis>
</taxonomicName>
<emphasis id="4FD05224FFBAD350E93BF878FD7DF84D" bold="true" box="[586,644,1934,1956]" pageId="22" pageNumber="173">
<taxonomicNameLabel id="54E3EF5FFFBAD350E93BF878FD7DF84D" box="[586,644,1934,1956]" pageId="22" pageNumber="173" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male (4852) and female (53). 48, T8 and S8; 49, T9; 50, S9; 51, penis and tegmen, superimposed; 52, apex of same, beginning eversion of endophallus; 53, bursal sclerite. 4850 to the same scale, 52 not to scale.
</paragraph>
<paragraph id="7D1B8E36FFBAD350EBE6FF61FE7BFF11" pageId="22" pageNumber="173">
<table id="0FA47C96FFBA2CB9EBE6FF61FA65FF11" box="[151,1436,151,248]" gridcols="6" gridrows="3" pageId="22" pageNumber="173">
<tr id="C3948C74FFBA2CB9EBE6FF61FA65FF59" box="[151,1436,151,176]" gridrow="0" pageId="22" pageNumber="173">
<th id="8045E508FFBA2CB9EBE6FF61FCEEFF59" box="[151,791,151,176]" gridcol="0" gridrow="0" pageId="22" pageNumber="173">elytra. Pilosity blond, semi-erect. Frontoclypeus with</th>
<th id="8045E508FFBA2CB9E86EFF61FC49FF59" box="[799,944,151,176]" gridcol="1" gridrow="0" pageId="22" pageNumber="173">lateral lobes</th>
<th id="8045E508FFBA2CB9E8CBFF61FB84FF59" box="[954,1149,151,176]" gridcol="2" gridrow="0" pageId="22" pageNumber="173">projecting either</th>
<th id="8045E508FFBA2CB9EFF7FF61FB45FF59" box="[1158,1212,151,176]" gridcol="3" gridrow="0" pageId="22" pageNumber="173">side</th>
<th id="8045E508FFBA2CB9EFB6FF61FAF6FF59" box="[1223,1295,151,176]" gridcol="4" gridrow="0" pageId="22" pageNumber="173">of the</th>
<th id="8045E508FFBA2CB9EE69FF61FA65FF59" box="[1304,1436,151,176]" gridcol="5" gridrow="0" pageId="22" pageNumber="173">rectangular</th>
</tr>
<tr id="C3948C74FFBA2CB9EBE6FF4AFA65FF3C" box="[151,1436,188,213]" gridrow="1" pageId="22" pageNumber="173">
<th id="8045E508FFBA2CB9EBE6FF4AFCEEFF3C" box="[151,791,188,213]" gridcol="0" gridrow="1" pageId="22" pageNumber="173">clypeolabrum. Front margin of pronotum curved, partly</th>
<td id="8045E508FFBA2CB9E86EFF4AFC49FF3C" box="[799,944,188,213]" gridcol="1" gridrow="1" pageId="22" pageNumber="173">covering the</td>
<td id="8045E508FFBA2CB9E8CBFF4AFB84FF3C" box="[954,1149,188,213]" gridcol="2" gridrow="1" pageId="22" pageNumber="173">occiput. Pronotal</td>
<td id="8045E508FFBA2CB9EFF7FF4AFB45FF3C" box="[1158,1212,188,213]" gridcol="3" gridrow="1" pageId="22" pageNumber="173">front</td>
<td id="8045E508FFBA2CB9EFB6FF4AFAF6FF3C" box="[1223,1295,188,213]" gridcol="4" gridrow="1" pageId="22" pageNumber="173">angles</td>
<td id="8045E508FFBA2CB9EE69FF4AFA65FF3C" box="[1304,1436,188,213]" gridcol="5" gridrow="1" pageId="22" pageNumber="173">rectangular,</td>
</tr>
<tr id="C3948C74FFBA2CB9EBE6FF29FA65FF11" box="[151,1436,223,248]" gridrow="2" pageId="22" pageNumber="173" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="8045E508FFBA2CB9EBE6FF29FCEEFF11" box="[151,791,223,248]" gridcol="0" gridrow="2" pageId="22" pageNumber="173">blunt, not projecting.</th>
</tr>
</table>
</paragraph>
<paragraph id="7D1B8E36FFBBD351EBB6FF61FDE3FEDF" blockId="23.[151,1437,151,1043]" pageId="23" pageNumber="174">
<emphasis id="4FD05224FFBBD351EBB6FF61FEF2FF59" bold="true" box="[199,267,151,176]" pageId="23" pageNumber="174">Male.</emphasis>
Antenna serrate near base, distal flagellar antennomeres triangular. Pilosity of abdominal sternites heterogenous. Plate of T8 wide, barely longer than apodemes, caudal edge truncate, with setae and microtrichia, S8 a shallow thin sclerite arch (Fig. 48). T9 has longer apodemes but a smaller plate than T8 which is relatively well defined, caudally truncate, bare (Fig. 49). S9 large, caudally bilobed and pilose, its anterior portion supported by a thin loop-shaped sclerite (Fig. 50).
</paragraph>
<paragraph id="7D1B8E36FFBBD351EBB6FEC8FC07FE52" blockId="23.[151,1437,151,1043]" pageId="23" pageNumber="174">The hairless tegmen (Fig. 51) with a deep keyhole-shaped basal notch surrounded by a strong sclerite. Styles long and slender. Parameres with convex convergent sides, ending in a strongly sclerotized dark hook with sharp tip. Transparent dorsal and ventral flanges surround the long parallel penis. Pala rounded in front and more than twice as long as the section occupied by parameroids and trigonium (Fig. 51).</paragraph>
<paragraph id="7D1B8E36FFBBD351EBB6FE32FD4CFD6A" blockId="23.[151,1437,151,1043]" pageId="23" pageNumber="174">Trigonium with wide base, diameter decreasing stepwise to a long narrow tip. Parameroids a little shorter than trigonium, with hooked apices overlying membranous cone through which ejaculatory duct can apparently be extruded. Orientation of parameroid apex changes with condition. At rest, tips directed mediad (Fig. 51). When penis apex spread open, parameroid tips turn outward (Fig. 52). At the same time, two slender bacula and two triangular sclerotized areas become exposed on spreading membranous penis apex. Armed endophallus apparently absent, visible parts of wall of wide ductus bare.</paragraph>
<paragraph id="7D1B8E36FFBBD351EBB6FD7AFDD2FCC3" blockId="23.[151,1437,151,1043]" pageId="23" pageNumber="174">
<emphasis id="4FD05224FFBBD351EBB6FD7AFEE7FD4C" bold="true" box="[199,286,652,677]" pageId="23" pageNumber="174">Female</emphasis>
(presumed). Antenna unmodified, distal flagellar antennomeres indistinctly conical, barely longer than apically wide. S6 with central porus. A prehensor was not recognized. Bursal sclerite consisting of a funnel-shaped sclerotized section of the gonoduct, the sides diverge cephalad and appear reticulate. Between their rear ends lies an elongate plate, between their divergent front ends are some small asymmetrical sclerites (Fig. 53) resembling female morphotype A (
<figureCitation id="E59F92B3FFBBD351EAEAFCE7FDE7FCC3" box="[411,542,785,810]" captionStart="FIGURES 45 47" captionStartId="20.[151,264,1125,1147]" captionTargetBox="[152,1436,377,1104]" captionTargetId="figure@20.[150,1436,373,1104]" captionTargetPageId="20" captionText="FIGURES 45 47. Prionocyphon spp., bursal sclerites, front end at top of page. Morphotypes A (45, 46) and B (47): 45, Acacia Plateau N. S. Wales H. Davidson (ANIC); 46, Tasmania A. Simson (SAMA); 47, Star Valley Lookout c. 5 km W of Paluma QLD 3. vii, 67 at light J. G. Brooks (ANIC). The scale lines are 100 µm." httpUri="https://zenodo.org/record/1052555/files/figure.png" pageId="23" pageNumber="174">Figs 45, 46</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="35BEDDBDFFBBD351EBB6FCC4FB97FBFB" pageId="23" pageNumber="174" type="etymology">
<paragraph id="7D1B8E36FFBBD351EBB6FCC4FB97FBFB" blockId="23.[151,1437,151,1043]" pageId="23" pageNumber="174">
<emphasis id="4FD05224FFBBD351EBB6FCC4FE3AFCA2" bold="true" box="[199,451,818,843]" pageId="23" pageNumber="174">Etymology and note</emphasis>
. The presumed female was taken together with several males, and there were no additional species in the sample. The Latin adjective
<taxonomicName id="BAA4F5B5FFBBD351E98EFCA3FCAFFC85" authorityName="Zwick" authorityYear="2016" box="[767,854,853,876]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="4FD05224FFBBD351E98EFCA3FCAFFC85" box="[767,854,853,876]" italics="true" pageId="23" pageNumber="174">uncatus</emphasis>
</taxonomicName>
means hooked, the same as
<taxonomicName id="BAA4F5B5FFBBD351EFD5FCA2FAFAFC85" box="[1188,1283,852,876]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4FD05224FFBBD351EFD5FCA2FAFAFC85" box="[1188,1283,852,876]" italics="true" pageId="23" pageNumber="174">hamatus</emphasis>
</taxonomicName>
. Both names describe the apex of the tegmen.
<taxonomicName id="BAA4F5B5FFBBD351E976FC83FCABFC64" authority="Watts" authorityName="Watts" box="[519,850,885,909]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4FD05224FFBBD351E976FC83FCF5FC64" box="[519,780,885,909]" italics="true" pageId="23" pageNumber="174">Prionocyphon hamatus</emphasis>
Watts
</taxonomicName>
and
<taxonomicName id="BAA4F5B5FFBBD351E8F9FC80FC03FC64" authority="Zwick, 2016" authorityName="Zwick" authorityYear="2016" box="[904,1018,886,909]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="uncatus" status="sp. nov.">
<emphasis id="4FD05224FFBBD351E8F9FC80FC03FC64" box="[904,1018,886,909]" italics="true" pageId="23" pageNumber="174">P. uncatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54E3EF5FFFBBD351EF73FC80FBCFFC64" box="[1026,1078,886,909]" pageId="23" pageNumber="174" rank="species">n.sp.</taxonomicNameLabel>
are related but different species with serrate antennae.
<taxonomicName id="BAA4F5B5FFBBD351EAE4FC60FD62FC47" box="[405,667,918,942]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="4FD05224FFBBD351EAE4FC60FD62FC47" box="[405,667,918,942]" italics="true" pageId="23" pageNumber="174">Prionocyphon hamatus</emphasis>
</taxonomicName>
differs mainly by the almost equal lengths of the pala and the distal portion of the penis, and the longer and more slender trigonium, the straight parameroids, and the long and thin paramere apices which were described as “resembling the hook on the apex of the penis in
<taxonomicName id="BAA4F5B5FFBBD351EFA3FC2CFAC5FC18" authorityName="Watts" authorityYear="2010" box="[1234,1340,986,1009]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="storeyi">
<emphasis id="4FD05224FFBBD351EFA3FC2CFAC5FC18" box="[1234,1340,986,1009]" italics="true" pageId="23" pageNumber="174">P. storeyi</emphasis>
</taxonomicName>
” (
<bibRefCitation id="1935F3C7FFBBD351EE2AFC2CFF18FBFA" author="Watts" pageId="23" pageNumber="174" refString="Watts, C. H. S. (2010 b). Revision of Australian Prionocyphon Redtenbacher (Scirtidae: Coleoptera). Transactions of the Royal Society of South Australia, 134, 53 - 88." type="journal article" year="2010" yearSuffix="b">Watts 2010b</bibRefCitation>
). They are apparently not strongly sclerotized while this is the case in
<taxonomicName id="BAA4F5B5FFBBD351E885FC0DFB9EFBFB" authorityName="Zwick" authorityYear="2016" box="[1012,1127,1019,1042]" class="Insecta" family="Scirtidae" genus="Prionocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="174" phylum="Arthropoda" rank="species" species="uncatus">
<emphasis id="4FD05224FFBBD351E885FC0DFB9EFBFB" box="[1012,1127,1019,1042]" italics="true" pageId="23" pageNumber="174">P. uncatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>