377 lines
52 KiB
XML
377 lines
52 KiB
XML
<document id="217539E89691AE50520F383F6ED8C5B3" ID-CLB-Dataset="31436" ID-DOI="10.11646/zootaxa.4085.2.1" ID-GBIF-Dataset="ba498adf-4dec-4fa3-8213-9d4d8c3b9dd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052535" ID-ZooBank="E41CB99C-5177-47A7-A424-2453D27E48F0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510873310154" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2016" docId="F50D3F20FFAFD340EBE6FF61FC9EFD19" docLanguage="en" docName="zootaxa.4085.2.1.pdf" docOrigin="Zootaxa 4085 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon chlorizans" docType="treatment" docVersion="10" lastPageNumber="157" masterDocId="09344758FFACD346EB71FFF6FFF9FFE9" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae" masterLastPageNumber="198" masterPageNumber="151" pageNumber="154" updateTime="1698499212072" updateUser="ExternalLinkService">
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<mods:title id="44608EF155302F5967A40467252AA152">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<mods:namePart id="35E23B731B7A26EC0489FCFCAAD6A421">Zwick, Peter</mods:namePart>
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<mods:title id="48D0D004472E60C28DF6FB3514DED525">Zootaxa</mods:title>
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<mods:date id="F6B37F0715A466028563AD978E44A1E5">2016</mods:date>
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<mods:number id="9AAF093E449646389F23126A69AAC399">2016-03-02</mods:number>
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<treatment id="F50D3F20FFAFD340EBE6FF61FC9EFD19" ID-DOI="http://doi.org/10.5281/zenodo.6076920" ID-GBIF-Taxon="137308960" ID-Zenodo-Dep="6076920" LSID="urn:lsid:plazi:treatment:F50D3F20FFAFD340EBE6FF61FC9EFD19" httpUri="http://treatment.plazi.org/id/F50D3F20FFAFD340EBE6FF61FC9EFD19" lastPageId="6" lastPageNumber="157" pageId="3" pageNumber="154">
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<subSubSection id="35BEDDBDFFAFD345EBE6FF61FEF2FF3A" pageId="3" pageNumber="154" type="nomenclature">
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<paragraph id="7D1B8E36FFAFD345EBE6FF61FD39FF5B" blockId="3.[151,704,151,211]" box="[151,704,151,178]" pageId="3" pageNumber="154">
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<heading id="2653395AFFAFD345EBE6FF61FD39FF5B" bold="true" box="[151,704,151,178]" fontSize="11" level="1" pageId="3" pageNumber="154" reason="1">
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<taxonomicName id="BAA4F5B5FFAFD345EBE6FF61FD39FF5B" authority="(Klausnitzer, 1973)" baseAuthorityName="Klausnitzer" baseAuthorityYear="1973" box="[151,704,151,178]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">
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<emphasis id="4FD05224FFAFD345EBE6FF61FD39FF5B" bold="true" box="[151,704,151,178]" pageId="3" pageNumber="154">
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<emphasis id="4FD05224FFAFD345EBE6FF61FE35FF58" bold="true" box="[151,460,151,177]" italics="true" pageId="3" pageNumber="154">Ypsiloncyphon chlorizans</emphasis>
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(
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||
<bibRefCitation id="1935F3C7FFAFD345EAADFF6EFD41FF5B" author="Klausnitzer" box="[476,696,152,178]" pageId="3" pageNumber="154" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon- Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Col., Helodidae). Folia entomologica Hungarica, 26, 97 - 110." type="journal article" year="1973">Klausnitzer, 1973</bibRefCitation>
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)
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||
</emphasis>
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||
</taxonomicName>
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||
</heading>
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</paragraph>
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<paragraph id="7D1B8E36FFAFD345EBE6FF4CFEF2FF3A" blockId="3.[151,704,151,211]" box="[151,267,186,211]" pageId="3" pageNumber="154">
|
||
(
|
||
<figureCitation id="E59F92B3FFAFD345EBEEFF4CFEFBFF3A" box="[159,258,186,211]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Figs 1, 2</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="35BEDDBDFFAFD345EBE6FEF4FD9AFDB6" pageId="3" pageNumber="154" type="materials_examined">
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<paragraph id="7D1B8E36FFAFD345EBE6FEF4FB2EFE8A" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
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<materialsCitation id="CDCC846BFFAFD345EBE6FEF4FEF3FE8A" ID-GBIF-Occurrence="1701200967" collectingDate="1961-11-21" collectionCode="ZMUC" collectorName="Noona Dan Exp. & Penis & Euparal & P. Zwick" country="Philippines" location="Tawi Tawi Lapid Lapid" municipality="Manalik Channel" pageId="3" pageNumber="154" specimenCount="3" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="4FD05224FFAFD345EBE6FEF4FE1DFEF2" bold="true" box="[151,484,258,283]" pageId="3" pageNumber="154">
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||
<typeStatus id="A21F3094FFAFD345EBE6FEF4FF2BFEF2" box="[151,210,258,283]" pageId="3" pageNumber="154">Type</typeStatus>
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||
material.
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||
<specimenCount id="6BA245BFFFAFD345EA21FEF4FE9CFEF2" box="[336,357,258,283]" pageId="3" pageNumber="154" type="male">♂</specimenCount>
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||
<typeStatus id="A21F3094FFAFD345EA1FFEF4FE26FEF2" box="[366,479,258,283]" pageId="3" pageNumber="154" type="holotype">Holotype</typeStatus>
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||
:
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||
</emphasis>
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||
<typeStatus id="A21F3094FFAFD345EA9CFEF4FD9BFEF2" box="[493,610,258,283]" pageId="3" pageNumber="154" type="holotype">Holotypus</typeStatus>
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||
<taxonomicName id="BAA4F5B5FFAFD345E91AFEF4FC33FEF2" authority="Klausnitzer" authorityName="Klausnitzer" authorityYear="1973" box="[619,970,258,283]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">Cyphon chlorizans Klausnitzer</taxonomicName>
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||
\
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||
<collectingCountry id="05B3CEA6FFAFD345E895FEF4FB9CFEF2" box="[996,1125,258,283]" name="Philippines" pageId="3" pageNumber="154">Philippines</collectingCountry>
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||
,
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||
<location id="787BD8EDFFAFD345EF03FEF4FA86FEF2" LSID="urn:lsid:plazi:treatment:F50D3F20FFAFD340EBE6FF61FC9EFD19:787BD8EDFFAFD345EF03FEF4FA86FEF2" box="[1138,1407,258,283]" country="Philippines" municipality="Manalik Channel" name="Tawi Tawi Lapid Lapid" pageId="3" pageNumber="154">Tawi Tawi Lapid Lapid</location>
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at
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<collectingMunicipality id="9D7F144CFFAFD345EBE6FED0FEA2FED6" box="[151,347,294,319]" pageId="3" pageNumber="154">Manalik Channel</collectingMunicipality>
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<date id="091AA8F6FFAFD345EA14FED1FE06FED6" box="[357,511,294,319]" pageId="3" pageNumber="154" value="1961-11-21">
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<collectingDate id="195E511EFFAFD345EA14FED1FE06FED6" box="[357,511,294,319]" pageId="3" pageNumber="154" value="1961-11-21">21.Nov. 1961</collectingDate>
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</date>
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||
<collectorName id="D051EBE0FFAFD345E978FED1FD32FED7" box="[521,715,295,318]" pageId="3" pageNumber="154">Noona Dan Exp.</collectorName>
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61-62 \
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||
<collectorName id="D051EBE0FFAFD345E840FED0FC97FED6" box="[817,878,294,319]" pageId="3" pageNumber="154">Penis</collectorName>
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mounted in
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<collectorName id="D051EBE0FFAFD345EF72FED0FBA3FED6" box="[1027,1114,294,319]" pageId="3" pageNumber="154">Euparal</collectorName>
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||
by
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||
<collectorName id="D051EBE0FFAFD345EFF8FED1FB17FED6" box="[1161,1262,294,319]" pageId="3" pageNumber="154">P. Zwick</collectorName>
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||
2015 \
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||
<collectionCode id="1BB516F3FFAFD345EE3AFED1FA65FED6" box="[1355,1436,295,319]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="154">ZMUC</collectionCode>
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0 0 518973
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="CDCC846BFFAFD345EA64FEBDFB2AFE8A" ID-GBIF-Occurrence="1701200969" box="[277,1235,330,355]" collectingDate="1961-11-21" collectionCode="ZMUC" collectorName="P. Zwick" country="Philippines" location="Tawi Tawi Lapid Lapid" municipality="Manalik Channel" pageId="3" pageNumber="154" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<specimenCount id="6BA245BFFFAFD345EA64FEBDFEC6FE8A" box="[277,319,330,355]" pageId="3" pageNumber="154" type="female">
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||
1
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||
<emphasis id="4FD05224FFAFD345EA5BFEBCFEC6FE8A" bold="true" box="[298,319,330,355]" pageId="3" pageNumber="154">♀</emphasis>
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||
</specimenCount>
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||
<typeStatus id="A21F3094FFAFD345EA34FEBCFE4DFE8A" box="[325,436,330,355]" pageId="3" pageNumber="154" type="paratype">paratype</typeStatus>
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||
: the same data, plus: Genitalia preparation by
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<collectorName id="D051EBE0FFAFD345E8CCFEBDFBDBFE8A" box="[957,1058,330,355]" pageId="3" pageNumber="154">P. Zwick</collectorName>
|
||
, 2015 (
|
||
<collectionCode id="1BB516F3FFAFD345EF02FEBDFB33FE8A" box="[1139,1226,331,355]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="154">ZMUC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="7D1B8E36FFAFD345EBE6FE99FCBEFE42" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
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Two additional
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<typeStatus id="A21F3094FFAFD345EA3DFE99FE4EFE6E" box="[332,439,367,391]" pageId="3" pageNumber="154" type="paratype">paratypes</typeStatus>
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taken at the same locality lack the abdomen (
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<collectionCode id="1BB516F3FFAFD345E8A1FE99FBDFFE6E" box="[976,1062,367,391]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="3" pageNumber="154">ZMUC</collectionCode>
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||
; collecting date
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<date id="091AA8F6FFAFD345EF94FE98FAB1FE6E" box="[1253,1352,366,391]" pageId="3" pageNumber="154" value="61-11-19">19.11.61</date>
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||
) or the genitalia slide is not available (BK; collecting date
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||
<date id="091AA8F6FFAFD345E9A4FE64FCC2FE42" box="[725,827,402,427]" pageId="3" pageNumber="154" value="61-11-21">21.11.61</date>
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||
).
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||
</paragraph>
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||
<paragraph id="7D1B8E36FFAFD345EBB6FE40FD9AFDB6" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
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||
<emphasis id="4FD05224FFAFD345EBB6FE40FE7BFE26" bold="true" box="[199,386,438,463]" pageId="3" pageNumber="154">Technical notes</emphasis>
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||
. The
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<typeStatus id="A21F3094FFAFD345EABAFE41FDFFFE26" box="[459,518,439,463]" pageId="3" pageNumber="154">types</typeStatus>
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sit on inverted minuten pins pushed into the mesothorax from below, slightly laterally, and the mesoventral process remains visible. The abdomen and male parts of the
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<typeStatus id="A21F3094FFAFD345EFA3FE2CFACDFE1A" box="[1234,1332,474,499]" pageId="3" pageNumber="154" type="holotype">holotype</typeStatus>
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are in a semitransparent film of glue on a card, only the penis needed to be removed, cleared of adherent tissue in KOH and mounted in Euparal. The detached empty female abdomen had been glued on a card. Some tissue next to it was cleared in KOH and mounted in Euparal.
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="35BEDDBDFFAFD340EBB6FD9CFC9EFD19" lastPageId="6" lastPageNumber="157" pageId="3" pageNumber="154" type="description">
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FD9CFCCBFD06" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
<emphasis id="4FD05224FFAFD345EBB6FD9CFE98FD6A" bold="true" box="[199,353,618,643]" pageId="3" pageNumber="154">Supplements</emphasis>
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||
to the original description of
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||
<emphasis id="4FD05224FFAFD345E9A2FD9DFC88FD6A" box="[723,881,619,643]" italics="true" pageId="3" pageNumber="154">
|
||
<taxonomicName id="BAA4F5B5FFAFD345E9A2FD9DFC95FD6A" authorityName="Klausnitzer" authorityYear="1973" box="[723,876,619,643]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">Y. chlorizans</taxonomicName>
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.
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</emphasis>
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||
Body surface dark brown, no greenish sheen. Maxillary palpus 3-segmented through fusion of segments 3 and 4, composite terminal segment large, with basal half thick, distal half slender, similar to
|
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<figureCitation id="E59F92B3FFAFD345E929FD44FD64FD22" box="[600,669,690,715]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 4</figureCitation>
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||
. Mesoventral process widening caudally and truncate, mesepimeron very narrow, as in Australian species (
|
||
<bibRefCitation id="1935F3C7FFAFD345E930FD20FCDFFD06" author="Zwick" box="[577,806,726,751]" pageId="3" pageNumber="154" refString="Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014">Zwick 2014: fig. 57</bibRefCitation>
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||
).
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FD0CFB96FCB2" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
<emphasis id="4FD05224FFAFD345EBB6FD0CFEFDFCFA" bold="true" box="[199,260,762,787]" pageId="3" pageNumber="154">Male</emphasis>
|
||
. Apodemes of T8 only half as long as those of T9. S8 rod-shaped and long, it must have extended foward into segment 4, similar to
|
||
<figureCitation id="E59F92B3FFAFD345EAB8FCE8FDF6FCDE" box="[457,527,798,823]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
|
||
. Caudally it supports rudimentary plate of sternite. Narrow forked tegmen and large S9 resembling intermediate between U and V as illustrated in original description.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FC90FDC1FC2E" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
Penis (
|
||
<figureCitation id="E59F92B3FFAFD345EA67FC90FEA0FC96" box="[278,345,870,895]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
|
||
) slender, in lateral view ventral face straight and flat at tip, except for minute low trigonium. Contour of middle third rises gradually posteriorly, then drops abruptly. Wrinkled cuticle of efferent duct (
|
||
<emphasis id="4FD05224FFAFD345EE07FC7DFF0CFC2F" italics="true" pageId="3" pageNumber="154">vas deferens</emphasis>
|
||
) seen in basal third of penis.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FC25FEB2FB2A" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
In dorsoventral view (
|
||
<figureCitation id="E59F92B3FFAFD345EAB1FC24FDF8FC02" box="[448,513,978,1003]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
|
||
) penis resembles a popsicle shaped like a sugar loaf. Parallel sclerites of the long pala end abruptly at a structure resembling the eye of a needle. Caudally from this point sclerites embedded in flat general surface, remaining sharply delimited for a short distance, distally turning into stripes which are more strongly pigmented than surroundings. Stripes converging, meeting and ending shortly before reaching nippleshaped trigonium. The raised little nipple stands at about 13% of total penis length from the regularly tongueshaped apex. A narrow medial seam on each of straight sclerites of pala widens, lies over needle-eye, and then turns to outside.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FB38FB45FAC6" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
<emphasis id="4FD05224FFAFD345EBB6FB38FEE7FB0E" bold="true" box="[199,286,1230,1255]" pageId="3" pageNumber="154">Female</emphasis>
|
||
. Ovipositor unmodified, bacula articulating with the long and slender gonocoxites. Gonostyli slender drums with apical sensilla. Edges of S8 converging and merging to a slender unpaired rod about half as long as apodemes of relatively wide T8 (
|
||
<figureCitation id="E59F92B3FFAFD345E978FAE0FDB4FAC6" box="[521,589,1302,1327]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 2</figureCitation>
|
||
). No vulvar sclerite and no dictyon on accessory gland.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FACCFC68FA9E" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
<emphasis id="4FD05224FFAFD345EBB6FACCFEE8FABA" bold="true" box="[199,273,1338,1363]" pageId="3" pageNumber="154">Notes.</emphasis>
|
||
Neither
|
||
<taxonomicName id="BAA4F5B5FFAFD345EA04FACDFDFFFABA" authorityName="Klausnitzer" authorityYear="1973" box="[373,518,1339,1363]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="4FD05224FFAFD345EA04FACDFDFFFABA" box="[373,518,1339,1363]" italics="true" pageId="3" pageNumber="154">Y. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
nor
|
||
<taxonomicName id="BAA4F5B5FFAFD345E94DFACDFD5EFABB" authorityName="Klausnitzer" authorityYear="1973" box="[572,679,1339,1362]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="micans">
|
||
<emphasis id="4FD05224FFAFD345E94DFACDFD5EFABB" box="[572,679,1339,1362]" italics="true" pageId="3" pageNumber="154">Y. micans</emphasis>
|
||
</taxonomicName>
|
||
exhibit the greenish sheen originally ascribed to them. Probably it resulted only from the kind of illumination used (
|
||
<bibRefCitation id="1935F3C7FFAFD345E9CEFAA8FC7CFA9E" author="Klausnitzer" box="[703,901,1374,1399]" pageId="3" pageNumber="154" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon- Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Col., Helodidae). Folia entomologica Hungarica, 26, 97 - 110." type="journal article" year="1973">Klausnitzer 1973</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6FA74FAB4F9A6" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
The
|
||
<typeStatus id="A21F3094FFAFD345EB8DFA75FED5FA72" box="[252,300,1411,1435]" pageId="3" pageNumber="154">type</typeStatus>
|
||
and the other Asian species examined (see below) as well as all Australian members agree in the structure of the mesoventrite. In a key to
|
||
<taxonomicName id="BAA4F5B5FFAFD345E912FA50FD43FA57" box="[611,698,1446,1470]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FD05224FFAFD345E912FA50FD43FA57" box="[611,698,1446,1470]" italics="true" pageId="3" pageNumber="154">Cyphon</emphasis>
|
||
</taxonomicName>
|
||
-like
|
||
<taxonomicName id="BAA4F5B5FFAFD345E984FA50FC1CFA56" authority="(Ruta 2009)" baseAuthorityName="Ruta" baseAuthorityYear="2009" box="[757,997,1446,1471]" class="Insecta" family="Scirtidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="family">
|
||
Scirtidae (
|
||
<bibRefCitation id="1935F3C7FFAFD345E817FA51FC27FA56" author="Ruta" box="[870,990,1447,1471]" pageId="3" pageNumber="154" refString="Ruta, R. (2009) Brachycyphon Fairmaire, 1896, a neglected genus of Afrotropical Scirtidae (Coleoptera). Annales Zoologici, 59 (4), 621 - 628. [Warszawa, 2009] http: // dx. doi. org / 10.3161 / 000345409 x 484982" type="journal article" year="2009">Ruta 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
the narrow mesepimeron distinguishes
|
||
<taxonomicName id="BAA4F5B5FFAFD345EBE6FA3DFEC0FA0A" authorityName="Klausnitzer" authorityYear="2009" box="[151,313,1483,1507]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FD05224FFAFD345EBE6FA3DFEC0FA0A" box="[151,313,1483,1507]" italics="true" pageId="3" pageNumber="154">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
. The mesoventral process of the illustrated
|
||
<taxonomicName id="BAA4F5B5FFAFD345E854FA3DFC3DFA0A" box="[805,964,1483,1507]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="jaloszynskii">
|
||
<emphasis id="4FD05224FFAFD345E854FA3DFC3DFA0A" box="[805,964,1483,1507]" italics="true" pageId="3" pageNumber="154">Y. jaloszynskii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1935F3C7FFAFD345E8A3FA3DFB5BFA0A" author="Ruta" box="[978,1186,1482,1507]" pageId="3" pageNumber="154" refString="Ruta, R. (2009) Brachycyphon Fairmaire, 1896, a neglected genus of Afrotropical Scirtidae (Coleoptera). Annales Zoologici, 59 (4), 621 - 628. [Warszawa, 2009] http: // dx. doi. org / 10.3161 / 000345409 x 484982" type="journal article" year="2009">Ruta 2009: fig. 19</bibRefCitation>
|
||
) widens, but less than in the other Asian species examined by
|
||
<bibRefCitation id="1935F3C7FFAFD345E92AFA19FD1AF9EE" author="Ruta" box="[603,739,1519,1543]" pageId="3" pageNumber="154" refString="Ruta, R. (2004). Five new species of Oriental Scirtidae (Coleoptera: Scirtoidea). Genus, 15, 363 - 379." type="journal article" year="2004">Ruta (2004)</bibRefCitation>
|
||
and myself. Klausnitzer (2009) calls the mesoventral process "breit abgestutzt und beborstet". The caudal widening of the mesoventral process may be a generic character not generally considered in the past but recently described in a number of Oriental species (
|
||
<bibRefCitation id="1935F3C7FFAFD345EF1DF9C0FAB8F9A6" author="Klausnitzer" box="[1132,1345,1590,1615]" pageId="3" pageNumber="154" refString="Klausnitzer, B. (2015 b) Neue Arten der Gattung Ypsiloncyphon Klausnitzer, 2009 (Coleoptera, Scirtidae) aus der Orientalischen Region (201. Beitrag zur Kenntnis der Scirtidae). Entomologische Blatter und Coleoptera, 111, 465 - 479." type="journal article" year="2015" yearSuffix="b">Klausnitzer 2015b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6F9ACFDA1F8EB" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
The maxillary palpus of
|
||
<taxonomicName id="BAA4F5B5FFAFD345EAADF9ADFD95F99A" authorityName="Klausnitzer" authorityYear="1973" box="[476,620,1627,1651]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="4FD05224FFAFD345EAADF9ADFD95F99A" box="[476,620,1627,1651]" italics="true" pageId="3" pageNumber="154">Y. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
and of the other Asian species I studied as well as those recently named by
|
||
<bibRefCitation id="1935F3C7FFAFD345EBCBF989FE44F97E" author="Klausnitzer" box="[186,445,1662,1687]" pageId="3" pageNumber="154" refString="Klausnitzer, B. (2015 b) Neue Arten der Gattung Ypsiloncyphon Klausnitzer, 2009 (Coleoptera, Scirtidae) aus der Orientalischen Region (201. Beitrag zur Kenntnis der Scirtidae). Entomologische Blatter und Coleoptera, 111, 465 - 479." type="journal article" year="2015" yearSuffix="b">B. Klausnitzer (2015b)</bibRefCitation>
|
||
is derived.
|
||
<bibRefCitation id="1935F3C7FFAFD345E933F989FD30F97E" author="Ruta" box="[578,713,1663,1687]" pageId="3" pageNumber="154" refString="Ruta, R. (2007) Scirtidae of India and Sri Lanka. Part 1. The chlorizans - group of Cyphon Paykull, 1799 (Insecta: Coleoptera). Genus, 17, 323 - 340." type="journal article" year="2007">Ruta (2007)</bibRefCitation>
|
||
did not mention the number of segments when he described the palpus as “fusiform, with broadened, globular proximal half, and narrow, tempered apex” which fits
|
||
<figureCitation id="E59F92B3FFAFD345EE5EF954FA8FF952" box="[1327,1398,1698,1723]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 4</figureCitation>
|
||
. In
|
||
<tableCitation id="3026BB8DFFAFD345EBE6F930FF10F936" box="[151,233,1734,1759]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Presumed phylogenetic relationships between the species of genus Ypsiloncyphon. Boxes enclose members of" pageId="3" pageNumber="154">Table 1</tableCitation>
|
||
the figure 3 stands before the name of species known to have a three-segmented palpus. Possibly even more Asian species are similar in this respect.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAFD345EBB6F8F9FCCCF85E" blockId="3.[151,1437,258,1975]" pageId="3" pageNumber="154">
|
||
S8 and S9 of
|
||
<taxonomicName id="BAA4F5B5FFAFD345EA2FF8F9FE17F8CF" authorityName="Klausnitzer" authorityYear="1973" box="[350,494,1806,1830]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="4FD05224FFAFD345EA2FF8F9FE17F8CF" box="[350,494,1806,1830]" italics="true" pageId="3" pageNumber="154">Y. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
were originally described as S9 and presumed T9, respectively. Their large size and forward prolongation into anterior sternites are also observed in other species of group 2 (
|
||
<figureCitation id="E59F92B3FFAFD345EF92F8C4FAD5F8A2" box="[1251,1324,1842,1867]" captionStart="FIGURES 1 – 7" captionStartId="5.[151,264,1800,1822]" captionTargetBox="[232,1360,206,1763]" captionTargetId="figure@5.[214,1372,195,1776]" captionTargetPageId="5" captionText="FIGURES 1 – 7. Ypsiloncyphon spp. Y. chlorizans (Klausnitzer): 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype. Y. javanicus Yoshitomi, male: 3, diagrammatic view of abdominal sternites 4 – 7, the invaginated S 8 shown in black (after a photograph); 4, maxillary palpus. Y. longus Zwick: 5, maxillary palpus. Y. katherinae Zwick, female (from: 16.34 S 135.41 E 14 km NW of Cape Crawford, NT. 6 Nov. 19 M. S. Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3 – 5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite." httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
|
||
) and are apomorphic. S9 varies within the
|
||
<taxonomicName id="BAA4F5B5FFAFD345E962F8A0FD8EF886" authorityName="Fleming" authorityYear="1821" box="[531,631,1878,1903]" class="Insecta" family="Scirtidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="subFamily" subFamily="Scirtinae">Scirtinae</taxonomicName>
|
||
but in the groundplan (
|
||
<bibRefCitation id="1935F3C7FFAFD345E80FF8A0FBECF886" author="Zwick" box="[894,1045,1878,1903]" pageId="3" pageNumber="154" refString="Zwick, P. (2015 c) To the knowledge of Sarabandus robustus (LeConte) (Col.: Scirtidae: Scirtinae), and on the groundplan of male marsh beetle genitalia. Linzer biologische Beitrage, 47 (2), 1439 - 1449." type="journal article" year="2015" yearSuffix="c">Zwick 2015c</bibRefCitation>
|
||
) S8 is a small structure. It tends to be strongly reduced, often consisting only of weak sclerites arranged like the letters V or Y. In a number of genera S8 is absent, also in all Australasian
|
||
<taxonomicName id="BAA4F5B5FFAFD345E940F868FD2AF85F" authorityName="Klausnitzer" authorityYear="2009" box="[561,723,1950,1974]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FD05224FFAFD345E940F868FD2AF85F" box="[561,723,1950,1974]" italics="true" pageId="3" pageNumber="154">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
-species.
|
||
</paragraph>
|
||
<caption id="29DBDEBEFFA8D342EBE6FF6EFA65FF46" box="[151,1436,151,175]" pageId="4" pageNumber="155">
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FF6EFA65FF46" blockId="4.[151,1436,151,360]" box="[151,1436,151,175]" pageId="4" pageNumber="155">
|
||
<emphasis id="4FD05224FFA8D342EBE6FF6EFEFBFF46" bold="true" box="[151,258,152,175]" pageId="4" pageNumber="155">TABLE 1</emphasis>
|
||
. Presumed phylogenetic relationships between the species of genus
|
||
<emphasis id="4FD05224FFA8D342E8A6FF61FB80FF46" box="[983,1145,151,175]" italics="true" pageId="4" pageNumber="155">
|
||
<taxonomicName id="BAA4F5B5FFA8D342E8A6FF61FB8CFF46" authorityName="Klausnitzer" authorityYear="2009" box="[983,1141,151,175]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">Ypsiloncyphon</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Boxes enclose members of
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FF4EFA79FF26" blockId="4.[151,1436,151,360]" box="[151,1408,184,207]" pageId="4" pageNumber="155">monophyletic units, compare the text. Probable apomorphies which are known only in a few species are identified by:</paragraph>
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FF2EFDC8FEE5" blockId="4.[151,1436,151,360]" pageId="4" pageNumber="155">
|
||
<emphasis id="4FD05224FFA8D342EBE6FF2EFF5DFF04" bold="true" box="[151,164,216,237]" italics="true" pageId="4" pageNumber="155">3</emphasis>
|
||
(preceding the name): maxillary palpus 3-segmented, terminal segment composite.: mesoventral process widens caudally.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FEE3FD77FEC2" blockId="4.[151,1436,151,360]" box="[151,654,277,299]" pageId="4" pageNumber="155">*: Female S8 with unpaired rod-shaped apodeme.</paragraph>
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FEC3FCE1FEA3" blockId="4.[151,1436,151,360]" box="[151,792,308,330]" pageId="4" pageNumber="155">#: Female S8 with apodemes connected to form an open frame.</paragraph>
|
||
<paragraph id="7D1B8E36FFA8D342EBE6FEA4FD20FE81" blockId="4.[151,1436,151,360]" box="[151,729,338,360]" pageId="4" pageNumber="155">Abbreviations: Klausn. = Klausnitzer, Yosh. = Yoshitomi</paragraph>
|
||
<caption id="29DBDEBEFFA9D343EBE6F8FEFB1FF83F" httpUri="https://zenodo.org/record/1052537/files/figure.png" pageId="5" pageNumber="156" targetBox="[232,1360,206,1763]" targetPageId="5">
|
||
<paragraph id="7D1B8E36FFA9D343EBE6F8FEFB1FF83F" blockId="5.[151,1437,1800,2006]" pageId="5" pageNumber="156">
|
||
<emphasis id="4FD05224FFA9D343EBE6F8FEFEC4F8F7" bold="true" box="[151,317,1800,1822]" pageId="5" pageNumber="156">FIGURES 1–7.</emphasis>
|
||
<taxonomicName id="BAA4F5B5FFA9D343EA37F8FEFE20F8F7" authorityName="Klausnitzer" authorityYear="2009" box="[326,473,1800,1822]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FD05224FFA9D343EA37F8FEFE20F8F7" box="[326,473,1800,1822]" italics="true" pageId="5" pageNumber="156">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<taxonomicName id="BAA4F5B5FFA9D343E965F8FCFCD5F8F7" authority="(Klausnitzer)" authorityName="Klausnitzer" box="[532,812,1800,1823]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="4FD05224FFA9D343E965F8FCFD63F8F7" box="[532,666,1800,1823]" italics="true" pageId="5" pageNumber="156">Y. chlorizans</emphasis>
|
||
(Klausnitzer)
|
||
</taxonomicName>
|
||
: 1, penis of the holotype with labels; 2, terminal section of ovipositor and segment 8 of the female paratype.
|
||
<taxonomicName id="BAA4F5B5FFA9D343E9F8F8DEFC8AF8D4" authority="Yoshitomi" authorityName="Yoshitomi" authorityYear="2015" box="[649,883,1831,1853]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="javanicus">
|
||
<emphasis id="4FD05224FFA9D343E9F8F8DEFCFBF8D4" box="[649,770,1832,1853]" italics="true" pageId="5" pageNumber="156">Y. javanicus</emphasis>
|
||
Yoshitomi
|
||
</taxonomicName>
|
||
, male: 3, diagrammatic view of abdominal sternites 4– 7, the invaginated S8 shown in black (after a photograph); 4, maxillary palpus.
|
||
<taxonomicName id="BAA4F5B5FFA9D343E880F8B1FB58F8B5" authority="Zwick" authorityName="Zwick" authorityYear="2014" box="[1009,1185,1862,1884]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="longus">
|
||
<emphasis id="4FD05224FFA9D343E880F8B1FBA8F8B5" box="[1009,1105,1862,1884]" italics="true" pageId="5" pageNumber="156">Y. longus</emphasis>
|
||
Zwick
|
||
</taxonomicName>
|
||
: 5, maxillary palpus.
|
||
<taxonomicName id="BAA4F5B5FFA9D343EEFBF8B1FEB4F893" authority="Zwick" authorityName="Zwick" authorityYear="2014" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="katherinae">
|
||
<emphasis id="4FD05224FFA9D343EEFBF8B1FEFAF893" italics="true" pageId="5" pageNumber="156">Y. katherinae</emphasis>
|
||
Zwick
|
||
</taxonomicName>
|
||
, female (from: 16.34S 135.41E 14km NW of Cape Crawford, NT. 6 Nov.19 M.S.Upton; ANIC): 6, terminal section of ovipositor and segment 8; 7, bursella with dictyon. Figures 3–5 are not to scale, scale bars of the other figures are 100 µm. Abbreviations: ba, baculum of ovipositor; gs, gonostylite; gx, gonocoxite; in, gonoduct leading into bursella; ne, needle eye; out, ductus leading out of bursella to accessory gland; tr, trigonium; vd, vas deferens; vs, vulvar sclerite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7D1B8E36FFAAD340EBB6FF61FE87FEA9" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="157">
|
||
Earlier descriptions of the penis of
|
||
<taxonomicName id="BAA4F5B5FFAAD340E903FF6FFCF1FF59" box="[626,776,152,176]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="chlorizans">
|
||
<emphasis id="4FD05224FFAAD340E903FF6FFCF1FF59" box="[626,776,152,176]" italics="true" pageId="6" pageNumber="157">Y. chlorizans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1935F3C7FFAAD340E86AFF61FC1FFF59" author="Klausnitzer" box="[795,998,151,176]" pageId="6" pageNumber="157" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon- Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Col., Helodidae). Folia entomologica Hungarica, 26, 97 - 110." type="journal article" year="1973">Klausnitzer 1973</bibRefCitation>
|
||
, 2009) are incorrect. The needle-eye described above is homologous to the axil-like opening observed in Australian species where the parameroids separate from the trigonium (
|
||
<bibRefCitation id="1935F3C7FFAAD340EA93FF29FD97FF11" author="Zwick" box="[482,622,223,248]" pageId="6" pageNumber="157" refString="Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014">Zwick 2014</bibRefCitation>
|
||
, e.g., figs 77, 101, 104). The plate to which the trigonium is attached is formed by the fused parameroids which is also suggested by the free apices observed in several species (
|
||
<bibRefCitation id="1935F3C7FFAAD340EE56FEF2FF2BFEA9" author="Yoshitomi" pageId="6" pageNumber="157" refString="Yoshitomi, H. (2015) New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae). Zootaxa, 3904 (4), 451 - 500. http: // dx. doi. org / 10.11646 / zootaxa. 3904.4.1" type="journal article" year="2015">Yoshitomi 2015</bibRefCitation>
|
||
, e.g., figs 4, 6).
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAAD340EBB6FEBBFCEFFE39" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="157">
|
||
In most
|
||
<taxonomicName id="BAA4F5B5FFAAD340EA53FEBAFE7FFE8C" authorityName="Fleming" authorityYear="1821" box="[290,390,332,357]" class="Insecta" family="Scirtidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="subFamily" subFamily="Scirtinae">Scirtinae</taxonomicName>
|
||
the lateral sclerites of female S8 are more or less embedded in the segment. In
|
||
<taxonomicName id="BAA4F5B5FFAAD340EF8BFEBAFA65FE8D" authorityName="Klausnitzer" authorityYear="2009" box="[1274,1436,332,356]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4FD05224FFAAD340EF8BFEBAFA65FE8D" box="[1274,1436,332,356]" italics="true" pageId="6" pageNumber="157">Ypsiloncyphon</emphasis>
|
||
</taxonomicName>
|
||
they form freely projecting apodemes which appears to be derived. Different conditions occur. I regard the unpaired rod to which the apodemes of female S8 merge as apomorphic. Figures by
|
||
<bibRefCitation id="1935F3C7FFAAD340EF39FE62FAF7FE44" author="Yoshitomi" box="[1096,1294,404,429]" pageId="6" pageNumber="157" refString="Yoshitomi, H. (2015) New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae). Zootaxa, 3904 (4), 451 - 500. http: // dx. doi. org / 10.11646 / zootaxa. 3904.4.1" type="journal article" year="2015">Yoshitomi (2015)</bibRefCitation>
|
||
show this in all known Asian females which are asterisked in
|
||
<tableCitation id="3026BB8DFFAAD340E9CDFE41FCE9FE39" box="[700,784,439,464]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Presumed phylogenetic relationships between the species of genus Ypsiloncyphon. Boxes enclose members of" pageId="6" pageNumber="157">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAAD340EBE6FE09FD33FDF1" blockId="6.[151,1436,511,752]" box="[151,714,511,536]" pageId="6" pageNumber="157">
|
||
<emphasis id="4FD05224FFAAD340EBE6FE09FD33FDF1" box="[151,714,511,536]" italics="true" pageId="6" pageNumber="157">
|
||
<emphasis id="4FD05224FFAAD340EBE6FE09FD3AFDF1" bold="true" box="[151,707,511,536]" italics="true" pageId="6" pageNumber="157">Additional Asian species studied for comparison</emphasis>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAAD340EBB6FDD5FDDAFD89" blockId="6.[151,1436,511,752]" pageId="6" pageNumber="157">
|
||
<materialsCitation id="CDCC846BFFAAD340EBB6FDD5FDDAFD89" ID-GBIF-Occurrence="1701200970" collectingDate="1994-05-15" collectionCode="NHMW" country="Vietnam" location="Pacholatko" municipality="Dembicki" pageId="6" pageNumber="157" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<taxonomicName id="BAA4F5B5FFAAD340EBB6FDD5FDAFFDD4" authority="Yoshitomi" authorityName="Yoshitomi" authorityYear="2015" box="[199,598,547,573]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="4FD05224FFAAD340EBB6FDD5FDAFFDD4" bold="true" box="[199,598,547,573]" pageId="6" pageNumber="157">
|
||
<emphasis id="4FD05224FFAAD340EBB6FDD5FE28FDD5" bold="true" box="[199,465,547,572]" italics="true" pageId="6" pageNumber="157">Ypsiloncyphon bicolor</emphasis>
|
||
Yoshitomi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="4FD05224FFAAD340E900FDD2FCE2FDD4" bold="true" box="[625,795,548,573]" pageId="6" pageNumber="157">
|
||
<specimenCount id="6BA245BFFFAAD340E900FDD2FD6DFDD4" box="[625,660,548,573]" pageId="6" pageNumber="157" type="female">2♀</specimenCount>
|
||
<typeStatus id="A21F3094FFAAD340E9D4FDD2FCE2FDD4" box="[677,795,548,573]" pageId="6" pageNumber="157" type="paratype">paratypes</typeStatus>
|
||
</emphasis>
|
||
: S.
|
||
<collectingCountry id="05B3CEA6FFAAD340E82BFDD2FC40FDD4" box="[858,953,548,573]" name="Vietnam" pageId="6" pageNumber="157">Vietnam</collectingCountry>
|
||
Nam Cat Tien Nat. Park 1.-
|
||
<date id="091AA8F6FFAAD340EE53FDD2FA65FDD4" box="[1314,1436,548,573]" pageId="6" pageNumber="157" value="1994-05-15">
|
||
<collectingDate id="195E511EFFAAD340EE53FDD2FA65FDD4" box="[1314,1436,548,573]" pageId="6" pageNumber="157" value="1994-05-15">15.5.1994</collectingDate>
|
||
</date>
|
||
<location id="787BD8EDFFAAD340EBE6FDB1FEEDFD89" LSID="urn:lsid:plazi:treatment:F50D3F20FFAFD340EBE6FF61FC9EFD19:787BD8EDFFAAD340EBE6FDB1FEEDFD89" box="[151,276,583,608]" country="Vietnam" municipality="Dembicki" name="Pacholatko" pageId="6" pageNumber="157">Pacholatko</location>
|
||
&
|
||
<collectingMunicipality id="9D7F144CFFAAD340EA46FDB1FE5CFD89" box="[311,421,583,608]" pageId="6" pageNumber="157">Dembicki</collectingMunicipality>
|
||
(
|
||
<collectionCode id="1BB516F3FFAAD340EAC5FDBEFDEFFD89" box="[436,534,584,608]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="6" pageNumber="157">NHMW</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAAD340EBB6FD9DFAD1FD41" blockId="6.[151,1436,511,752]" pageId="6" pageNumber="157">
|
||
<taxonomicName id="BAA4F5B5FFAAD340EBB6FD9DFD57FD6C" authority="Yoshitomi" authorityName="Yoshitomi" authorityYear="2015" box="[199,686,619,645]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="chiangmaiensis">
|
||
<emphasis id="4FD05224FFAAD340EBB6FD9DFD57FD6C" bold="true" box="[199,686,619,645]" pageId="6" pageNumber="157">
|
||
<emphasis id="4FD05224FFAAD340EBB6FD9DFDD6FD6D" bold="true" box="[199,559,619,644]" italics="true" pageId="6" pageNumber="157">Ypsiloncyphon chiangmaiensis</emphasis>
|
||
Yoshitomi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="4FD05224FFAAD340E9B4FD9AFC90FD6C" bold="true" box="[709,873,620,645]" pageId="6" pageNumber="157">7♂ paratypes</emphasis>
|
||
: N-THAILAND, 18º49’N 98º57’E Chiang Mai, Zoo (Licht); several dates between
|
||
<date id="091AA8F6FFAAD340E952FD66FCDCFD41" box="[547,805,655,680]" pageId="6" pageNumber="157" value="1989-05-01" valueMax="1989-06-01" valueMin="1989-05-01">1 Mai and 1 June 1989</date>
|
||
, leg. Chantaramongkol & Malicky (NHMW).
|
||
</paragraph>
|
||
<paragraph id="7D1B8E36FFAAD340EBB6FD45FC9EFD19" blockId="6.[151,1436,511,752]" pageId="6" pageNumber="157">
|
||
<taxonomicName id="BAA4F5B5FFAAD340EBB6FD45FD9AFD24" authority="Yoshitomi" authorityName="Yoshitomi" authorityYear="2015" box="[199,611,691,717]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="javanicus">
|
||
<emphasis id="4FD05224FFAAD340EBB6FD45FD9AFD24" bold="true" box="[199,611,691,717]" pageId="6" pageNumber="157">
|
||
<emphasis id="4FD05224FFAAD340EBB6FD45FE1FFD25" bold="true" box="[199,486,691,716]" italics="true" pageId="6" pageNumber="157">Ypsiloncyphon javanicus</emphasis>
|
||
Yoshitomi
|
||
</emphasis>
|
||
</taxonomicName>
|
||
: 1♂: JAVA: Cibodas Kibun Raya caa. 4k SW of Cipana
|
||
<date id="091AA8F6FFAAD340EF9AFD42FA65FD25" box="[1259,1436,692,716]" pageId="6" pageNumber="157" value="1991-10-14" valueMax="1991-10-18" valueMin="1991-10-14">14-18 Oct 1991</date>
|
||
C.Reid, D.Subasli \ Sweeping grassy stream ca.
|
||
<quantity id="BA5C23D3FFAAD340E9C2FD21FCFBFD19" box="[691,770,727,752]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="157" unit="m" value="1350.0">1350m</quantity>
|
||
(ANIC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |