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125 lines
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<document id="BD22A6FD93F780FE8D970488C63807E0" ID-CLB-Dataset="31436" ID-DOI="10.11646/zootaxa.4085.2.1" ID-GBIF-Dataset="ba498adf-4dec-4fa3-8213-9d4d8c3b9dd3" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052535" ID-ZooBank="E41CB99C-5177-47A7-A424-2453D27E48F0" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510873310154" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2016" docId="F50D3F20FFAED344EBE6FB35FD49F804" docLanguage="en" docName="zootaxa.4085.2.1.pdf" docOrigin="Zootaxa 4085 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ypsiloncyphon Klausnitzer 2009" docType="treatment" docVersion="9" lastPageNumber="153" masterDocId="09344758FFACD346EB71FFF6FFF9FFE9" masterDocTitle="Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae" masterLastPageNumber="198" masterPageNumber="151" pageNumber="153" updateTime="1698499212072" updateUser="ExternalLinkService">
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<mods:title id="BD296CE300AE9AE59DF12A06C514AC4C">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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<mods:namePart id="40451D9F181C997643FC0F3DADD89A89">Zwick, Peter</mods:namePart>
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<mods:date id="32E5A172280C41777E6B5033923D78EC">2016</mods:date>
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<mods:number id="968FB4D38D01BE93579FB6385614825C">2016-03-02</mods:number>
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<mods:number id="E77F5CEAD99134E02FAC888008EF01CA">4085</mods:number>
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<treatment id="F50D3F20FFAED344EBE6FB35FD49F804" ID-DOI="http://doi.org/10.5281/zenodo.6076916" ID-GBIF-Taxon="137308959" ID-Zenodo-Dep="6076916" LSID="urn:lsid:plazi:treatment:F50D3F20FFAED344EBE6FB35FD49F804" httpUri="http://treatment.plazi.org/id/F50D3F20FFAED344EBE6FB35FD49F804" lastPageNumber="153" pageId="2" pageNumber="153">
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<subSubSection id="35BEDDBDFFAED344EBE6FB35FF05FB17" pageId="2" pageNumber="153" type="nomenclature">
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<paragraph id="7D1B8E36FFAED344EBE6FB35FD77FB34" blockId="2.[151,654,1218,1278]" box="[151,654,1218,1245]" pageId="2" pageNumber="153">
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<heading id="2653395AFFAED344EBE6FB35FD77FB34" bold="true" box="[151,654,1218,1245]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
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<emphasis id="4FD05224FFAED344EBE6FB35FD77FB34" bold="true" box="[151,654,1218,1245]" pageId="2" pageNumber="153">
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Genus
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<taxonomicName id="BAA4F5B5FFAED344EB9FFB34FD77FB34" authority="Klausnitzer, 2009 a" authorityName="Klausnitzer" authorityYear="2009" box="[238,654,1218,1245]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344EB9FFB34FE66FB35" bold="true" box="[238,415,1218,1244]" italics="true" pageId="2" pageNumber="153">Ypsiloncyphon</emphasis>
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<bibRefCitation id="1935F3C7FFAED344EAD7FB35FD77FB34" author="Klausnitzer" box="[422,654,1219,1245]" pageId="2" pageNumber="153" refString="Klausnitzer, B. (2009 a) Beschreibung einer neuen Gattung der Scirtidae aus der Orientalischen Region (Coleoptera) (144. Beitrag zur Kenntnis der Scirtidae). Beitrage zur Entomologie, 59, 247 - 261." type="journal article" year="2009" yearSuffix="a">Klausnitzer, 2009a</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="7D1B8E36FFAED344EBE6FB13FF05FB17" blockId="2.[151,654,1218,1278]" box="[151,252,1253,1278]" pageId="2" pageNumber="153">
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(
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<tableCitation id="3026BB8DFFAED344EBEFFB13FF08FB17" box="[158,241,1253,1278]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Presumed phylogenetic relationships between the species of genus Ypsiloncyphon. Boxes enclose members of" pageId="2" pageNumber="153">Table 1</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="35BEDDBDFFAED344EBE6FADBFD49F804" pageId="2" pageNumber="153" type="materials_examined">
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<paragraph id="7D1B8E36FFAED344EBE6FADBFD53FAAA" blockId="2.[151,682,1325,1347]" box="[151,682,1325,1347]" pageId="2" pageNumber="153">
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<emphasis id="4FD05224FFAED344EBE6FADBFEDFFAAA" bold="true" box="[151,294,1325,1347]" pageId="2" pageNumber="153">Type species:</emphasis>
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<taxonomicName id="BAA4F5B5FFAED344EA5AFADBFD5EFAAA" authority="Klausnitzer, 1973" authorityName="Klausnitzer" authorityYear="1973" box="[299,679,1325,1347]" class="Insecta" family="Scirtidae" genus="Cyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="chlorizans">
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<emphasis id="4FD05224FFAED344EA5AFADBFE13FAAA" box="[299,490,1325,1347]" italics="true" pageId="2" pageNumber="153">Cyphon chlorizans</emphasis>
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<bibRefCitation id="1935F3C7FFAED344EA81FADBFD5EFAAA" author="Klausnitzer" box="[496,679,1325,1347]" pageId="2" pageNumber="153" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon- Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Col., Helodidae). Folia entomologica Hungarica, 26, 97 - 110." type="journal article" year="1973">Klausnitzer, 1973</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="7D1B8E36FFAED344EBE6FA99FECDF96D" blockId="2.[151,1437,1391,2029]" pageId="2" pageNumber="153">
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The Australasian species of the genus
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<taxonomicName id="BAA4F5B5FFAED344E935FA86FD1FFA61" authorityName="Klausnitzer" authorityYear="2009" box="[580,742,1392,1416]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344E935FA86FD1FFA61" box="[580,742,1392,1416]" italics="true" pageId="2" pageNumber="153">Ypsiloncyphon</emphasis>
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</taxonomicName>
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were recently revised (
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<bibRefCitation id="1935F3C7FFAED344E89DFA99FB8EFA61" author="Zwick" box="[1004,1143,1391,1416]" pageId="2" pageNumber="153" refString="Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014">Zwick 2014</bibRefCitation>
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) but their affinities within the large genus were not clarified. Presently,
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<taxonomicName id="BAA4F5B5FFAED344E9FCFA62FCD6FA45" authorityName="Klausnitzer" authorityYear="2009" box="[653,815,1428,1452]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344E9FCFA62FCD6FA45" box="[653,815,1428,1452]" italics="true" pageId="2" pageNumber="153">Ypsiloncyphon</emphasis>
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</taxonomicName>
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includes over 50 Asian and about a dozen Australasian species (
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<tableCitation id="3026BB8DFFAED344EB8BFA41FEAAFA39" box="[250,339,1463,1488]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Presumed phylogenetic relationships between the species of genus Ypsiloncyphon. Boxes enclose members of" pageId="2" pageNumber="153">Table 1</tableCitation>
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, which contains all author names). The genus was established for species with, among other characteristics, a Y-shaped tegmen and a male T9 with apodemes longer than those of T8. Other elements of the male terminalia can take very different forms which made me question the monophyly of the genus (
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<bibRefCitation id="1935F3C7FFAED344EE72FA09FA68F9F1" author="Zwick" box="[1283,1425,1535,1560]" pageId="2" pageNumber="153" refString="Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014">Zwick 2014</bibRefCitation>
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). Today, various species are known in other Australian marsh beetle genera that are only distantly related but also present the name-giving Y-shaped tegmen (e.g., several
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<taxonomicName id="BAA4F5B5FFAED344E87CF9BEFC51F989" authorityName="Zwick" authorityYear="2013" box="[781,936,1608,1632]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344E87CF9BEFC51F989" box="[781,936,1608,1632]" italics="true" pageId="2" pageNumber="153">Austrocyphon</emphasis>
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</taxonomicName>
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spp.,
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<taxonomicName id="BAA4F5B5FFAED344E89AF9BEFAE9F989" authority="(Calvariellum)" authorityName="Calvariellum" box="[1003,1296,1608,1632]" class="Insecta" family="Scirtidae" genus="Calvarium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344E89AF9BEFAE9F989" box="[1003,1296,1608,1632]" italics="true" pageId="2" pageNumber="153">Calvarium (Calvariellum)</emphasis>
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</taxonomicName>
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spp.:
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<bibRefCitation id="1935F3C7FFAED344EE25F9B1FF26F96C" author="Zwick" pageId="2" pageNumber="153" refString="Zwick, P. (2013 d) ERRATUM: Peter Zwick (2013) Australian Marsh Beetles (Coleoptera: Scirtidae) 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 001 - 074. Zootaxa, 3710 (6), 600. http: // dx. doi. org / 10.11646 / zootaxa. 3710.6.6" type="journal article" year="2013" yearSuffix="d">Zwick 2013d</bibRefCitation>
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,
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<bibRefCitation id="1935F3C7FFAED344EB9AF99AFED1F96D" author="Zwick" box="[235,296,1644,1668]" pageId="2" pageNumber="153" refString="Zwick, P. (2014) Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer. Zootaxa, 3846 (1), 1 - 41. http: // dx. doi. org / 10.11646 / zootaxa. 3846.1.1" type="journal article" year="2014">2014</bibRefCitation>
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).
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</paragraph>
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<paragraph id="7D1B8E36FFAED344EBB6F979FD51F8B4" blockId="2.[151,1437,1391,2029]" pageId="2" pageNumber="153">
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Presently, four informal species groups within
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<taxonomicName id="BAA4F5B5FFAED344E980F966FC6AF941" authorityName="Klausnitzer" authorityYear="2009" box="[753,915,1680,1704]" class="Insecta" family="Scirtidae" genus="Ypsiloncyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="4FD05224FFAED344E980F966FC6AF941" box="[753,915,1680,1704]" italics="true" pageId="2" pageNumber="153">Ypsiloncyphon</emphasis>
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</taxonomicName>
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are separated by diagnostic characters. The definitions of groups 2 and 3 (the latter contains the Australasian species) partly rely on apomorphic characters (
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<bibRefCitation id="1935F3C7FFAED344EBEFF92EFEEEF919" author="Ruta" box="[158,279,1752,1776]" pageId="2" pageNumber="153" refString="Ruta, R. (2007) Scirtidae of India and Sri Lanka. Part 1. The chlorizans - group of Cyphon Paykull, 1799 (Insecta: Coleoptera). Genus, 17, 323 - 340." type="journal article" year="2007">Ruta 2007</bibRefCitation>
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; Klausnitzer 2009) while the definition of group 1 admits a variety of character expressions and is less helpful. The two species in the recently proposed group 4 have asymmetrical penes with setae or spines (
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<bibRefCitation id="1935F3C7FFAED344EE56F90AFF2DF8D1" author="Yoshitomi" pageId="2" pageNumber="153" refString="Yoshitomi, H. (2015) New species of the genus Ypsiloncyphon (Insecta: Coleoptera: Scirtidae: Scirtinae). Zootaxa, 3904 (4), 451 - 500. http: // dx. doi. org / 10.11646 / zootaxa. 3904.4.1" type="journal article" year="2015">Yoshitomi 2015</bibRefCitation>
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). Unfortunately, many Asian species, also the
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<typeStatus id="A21F3094FFAED344E9A5F8D6FCFDF8D1" box="[724,772,1824,1848]" pageId="2" pageNumber="153">type</typeStatus>
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species, are incompletely known, with only the habitus and selected details of the male genitalia described.
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</paragraph>
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<paragraph id="7D1B8E36FFAED344EBB6F891FD49F804" blockId="2.[151,1437,1391,2029]" pageId="2" pageNumber="153">
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The description of the
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<typeStatus id="A21F3094FFAED344EA91F89EFDE9F869" box="[480,528,1896,1920]" pageId="2" pageNumber="153">type</typeStatus>
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species is supplemented and partly corrected. A new understanding of the groundplan of male terminalia of
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<taxonomicName id="BAA4F5B5FFAED344E963F87AFCDAF84C" authority="(Zwick 2015c)" authorityName="Fleming" authorityYear="1821" baseAuthorityName="Zwick" baseAuthorityYear="2015" box="[530,803,1932,1957]" class="Insecta" family="Scirtidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="subFamily" subFamily="Scirtinae">
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Scirtinae (
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<bibRefCitation id="1935F3C7FFAED344E9F5F87AFCE2F84C" author="Zwick" box="[644,795,1932,1957]" pageId="2" pageNumber="153" refString="Zwick, P. (2015 c) To the knowledge of Sarabandus robustus (LeConte) (Col.: Scirtidae: Scirtinae), and on the groundplan of male marsh beetle genitalia. Linzer biologische Beitrage, 47 (2), 1439 - 1449." type="journal article" year="2015" yearSuffix="c">Zwick 2015c</bibRefCitation>
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)
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</taxonomicName>
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is helpful in definitions of monophyla within the genus, in order to determine the relations of the Australasian fauna to the other species. A comprehensive analysis of the Asian fauna is not intended in the present study.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |