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<mods:title id="1EED1CB7918183C239B41B5C6DC6B871">Australian Marsh Beetles (Coleoptera: Scirtidae). 9. The relations of Australasian Ypsiloncyphon species to their Asian congeners, additions, mainly to Petrocyphon and Prionocyphon, and a key to Australian genera of Scirtinae</mods:title>
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Genus
<taxonomicName id="BAA4F5B5FF8FD365EB9FFB1FFDBBFAEA" authority="Zwick, 2013 c" authorityName="Zwick" authorityYear="2013" box="[238,578,1257,1283]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="4FD05224FF8FD365EB9FFB1FFE61FAEA" bold="true" box="[238,408,1257,1283]" italics="true" pageId="35" pageNumber="186">Austrocyphon</emphasis>
<bibRefCitation id="1935F3C7FF8FD365EAEFFB1FFDBBFAEA" author="Zwick" box="[414,578,1257,1283]" pageId="35" pageNumber="186" refString="Zwick, P. (2013 c) Australian Marsh Beetles (Coleoptera: Scirtidae). 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 3706.1.1" type="journal article" year="2013" yearSuffix="c">Zwick, 2013c</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<paragraph id="7D1B8E36FF8FD365EBE6FADDFBACFA3A" blockId="35.[151,1436,1322,1563]" pageId="35" pageNumber="186">
<taxonomicName id="BAA4F5B5FF8FD365EBE6FADDFECBFAAA" authorityName="Zwick" authorityYear="2013" box="[151,306,1323,1347]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="4FD05224FF8FD365EBE6FADDFECBFAAA" box="[151,306,1323,1347]" italics="true" pageId="35" pageNumber="186">Austrocyphon</emphasis>
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includes many species. Males can be recognized by their distinctive terminalia but females have neither prehensors nor bursal sclerites.
<taxonomicName id="BAA4F5B5FF8FD365E912FAB8FC53FA8E" authority="Zwick" authorityName="Zwick" authorityYear="2013" box="[611,938,1358,1383]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="papilio">
<emphasis id="4FD05224FF8FD365E912FAB8FCAEFA8F" box="[611,855,1358,1382]" italics="true" pageId="35" pageNumber="186">Austrocyphon papilio</emphasis>
Zwick
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from Western Australia is an exception. I described elongate structures with transversely folded (concertinated) front ends in the vulvar area of individual females of several species (e.g.,
<taxonomicName id="BAA4F5B5FF8FD365E97AFA61FD47FA46" authority="(Lea)" authorityName="Lea" box="[523,702,1430,1455]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="doctus">
<emphasis id="4FD05224FF8FD365E97AFA61FD8CFA47" box="[523,629,1430,1454]" italics="true" pageId="35" pageNumber="186">A. doctus</emphasis>
(Lea)
</taxonomicName>
,
<taxonomicName id="BAA4F5B5FF8FD365E9BAFA61FBADFA46" authority="Zwick: Zwick 2013 c" authorityName="Zwick: Zwick" authorityYear="2013" box="[715,1108,1430,1455]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="tribulator">
<emphasis id="4FD05224FF8FD365E9BAFA61FCA0FA47" box="[715,857,1430,1454]" italics="true" pageId="35" pageNumber="186">A. tribulator</emphasis>
<bibRefCitation id="1935F3C7FF8FD365E813FA60FBADFA46" author="Zwick" box="[866,1108,1430,1455]" pageId="35" pageNumber="186" refString="Zwick, P. (2013 c) Australian Marsh Beetles (Coleoptera: Scirtidae). 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 3706.1.1" type="journal article" year="2013" yearSuffix="c">Zwick: Zwick 2013c</bibRefCitation>
</taxonomicName>
). These objects are probably spermatophores. Meanwhile I have seen them also in females of several other genera.
</paragraph>
<paragraph id="7D1B8E36FF8FD365EBB6FA29FADBF9F2" blockId="35.[151,1436,1322,1563]" pageId="35" pageNumber="186">
However, a few species can be recognized by their dorsal colour pattern or habitus (e.g.,
<taxonomicName id="BAA4F5B5FF8FD365EFDAFA29FA65FA1E" authority="(Blackburn)" authorityName="Blackburn" box="[1195,1436,1502,1527]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="4FD05224FF8FD365EFDAFA29FAF4FA1F" box="[1195,1293,1503,1526]" italics="true" pageId="35" pageNumber="186">A. pictus</emphasis>
(Blackburn)
</taxonomicName>
(
<bibRefCitation id="1935F3C7FF8FD365EBEFF9F4FE74F9F2" author="Zwick" box="[158,397,1538,1563]" pageId="35" pageNumber="186" refString="Zwick, P. (2013 c) Australian Marsh Beetles (Coleoptera: Scirtidae). 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 3706.1.1" type="journal article" year="2013" yearSuffix="c">Zwick 2013c: fig. 17</bibRefCitation>
),
<taxonomicName id="BAA4F5B5FF8FD365EAD3F9F5FD63F9F2" authority="Zwick" authorityName="Zwick" authorityYear="2013" box="[418,666,1538,1563]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="leptophallus">
<emphasis id="4FD05224FF8FD365EAD3F9F5FDB5F9F2" box="[418,588,1539,1563]" italics="true" pageId="35" pageNumber="186">A. leptophallus</emphasis>
Zwick
</taxonomicName>
(
<figureCitation id="E59F92B3FF8FD365E9DBF9F4FD04F9F2" box="[682,765,1538,1563]" captionStart="FIGURES 87 89" captionStartId="37.[151,264,874,896]" captionTargetBox="[160,1428,197,844]" captionTargetId="figure@37.[151,1435,195,853]" captionTargetPageId="37" captionText="FIGURES 87 89. Austrocyphon spp., dorsal view. 87, A. curvispina Zwick (female from Brown Mt.) 88, A. curvispina (presumed; female from Melong); 89, A. leptophallus Zwick, male from The Millstream, near Ravenshoe. Not to scale." httpUri="https://zenodo.org/record/1052573/files/figure.png" pageId="35" pageNumber="186">Fig. 89</figureCitation>
; also
<bibRefCitation id="1935F3C7FF8FD365E84CF9F4FC2DF9F2" author="Zwick" box="[829,980,1538,1563]" pageId="35" pageNumber="186" refString="Zwick, P. (2013 c) Australian Marsh Beetles (Coleoptera: Scirtidae). 4. Two new genera, Austrocyphon and Tasmanocyphon. Zootaxa, 3706 (1), 1 - 74. http: // dx. doi. org / 10.11646 / zootaxa. 3706.1.1" type="journal article" year="2013" yearSuffix="c">Zwick 2013c</bibRefCitation>
: fig. 184), and
<taxonomicName id="BAA4F5B5FF8FD365EF0DF9F5FAEBF9F3" authorityName="Zwick" authorityYear="2013" box="[1148,1298,1539,1562]" class="Insecta" family="Scirtidae" genus="Austrocyphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="species" species="curvispina">
<emphasis id="4FD05224FF8FD365EF0DF9F5FAEBF9F3" box="[1148,1298,1539,1562]" italics="true" pageId="35" pageNumber="186">A. curvispina</emphasis>
</taxonomicName>
).
</paragraph>
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