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<document id="E540A0798A6CFA16A2536C505BF0FE73" ID-CLB-Dataset="306077" ID-DOI="10.1093/zoolinnean/zlad185" ID-GBIF-Dataset="22dfc63b-d0de-464c-9983-1d3fa8738bb3" ID-ISSN="0024-4082" ID-Zenodo-Dep="14284394" ID-ZooBank="60FED76-A053-43D1-993A-E89C17167796" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732844651186" checkinUser="plazi" docAuthor="García-Guerrero, Fernando, Miller, Jonathan P., Delicado, Diana, Novo, Marta &amp; Ramos, Marian A." docDate="2024" docId="03FC87D6BA3CFFA0DD92AA25FDFA657C" docLanguage="en" docName="zlad185.pdf" docOrigin="Zoological Journal of the Linnean Society 202" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.9:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="9" docTitle="Navarriella Boeters 2000" docType="treatment" docVersion="3" lastPageNumber="9" masterDocId="FFC5FFAEBA3BFFA9DE7BAC0AFF896418" masterDocTitle="A systematic review of the Iberian springsnail subgenus Alzoniella (Navarriella) (Caenogastropoda: Hydrobiidae), with the description of a new potentially relict subfamily" masterLastPageNumber="17" masterPageNumber="1" pageNumber="8" updateTime="1733418490196" updateUser="ExternalLinkService">
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<mods:title id="12224421E39E37F958A8E751E60F4AB2">A systematic review of the Iberian springsnail subgenus Alzoniella (Navarriella) (Caenogastropoda: Hydrobiidae), with the description of a new potentially relict subfamily</mods:title>
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<mods:namePart id="6B0EFF0778A74F925446E5E9F18BA3CA">García-Guerrero, Fernando</mods:namePart>
<mods:affiliation id="E960F35C28BBD37BC36ED8367F0C47DB">National Museum of Natural ºciences (MNCN-CºIC), José Gutiérrez Abascal 2, 28006 Madrid, ºpain</mods:affiliation>
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<mods:namePart id="E9779A205BA0CAC6B031AF2306623A95">Miller, Jonathan P.</mods:namePart>
<mods:affiliation id="9603D17E4E6968CC77C46BB232B9765D">National Museum of Natural ºciences (MNCN-CºIC), José Gutiérrez Abascal 2, 28006 Madrid, ºpain</mods:affiliation>
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<mods:namePart id="A5F1C3427DF5F6AFFE5044E9A0E5DB77">Delicado, Diana</mods:namePart>
<mods:affiliation id="0F20EB98C1B2F247427387A252878FE2">Justus Liebig University Giessen, Department of Animal Ecology % ºystematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany</mods:affiliation>
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<mods:namePart id="173E84C12C6DF089C8701C0A09B7F738">Novo, Marta</mods:namePart>
<mods:affiliation id="48FB313DEC22344739BE9728AE62BD32">Department of Biodiversity, Ecology and Evolution, Faculty of Biological ºciences, Complutense University of Madrid, 28040 Madrid, ºpain</mods:affiliation>
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<mods:namePart id="129AE17DE4AF7B186B286F2CB11358A2">Ramos, Marian A.</mods:namePart>
<mods:affiliation id="CD0655E3455A04E77AD4432A56F5234B">National Museum of Natural ºciences (MNCN-CºIC), José Gutiérrez Abascal 2, 28006 Madrid, ºpain</mods:affiliation>
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<mods:title id="78BB4470E2FABFD16C1D01B439B8951C">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="DA8DEA4155026B6ED336B7F5E69F522E">2024</mods:date>
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<treatment id="03FC87D6BA3CFFA0DD92AA25FDFA657C" LSID="urn:lsid:plazi:treatment:03FC87D6BA3CFFA0DD92AA25FDFA657C" httpUri="http://treatment.plazi.org/id/03FC87D6BA3CFFA0DD92AA25FDFA657C" lastPageId="9" lastPageNumber="9" pageId="7" pageNumber="8" scope_class="Gastropoda" scope_family="Hydrobiidae" scope_order="Littorinimorpha" scope_subClass="Caenogastropoda">
<subSubSection id="C34F654BBA3CFFAEDD92AA25FB7D6251" box="[1001,1268,1583,1610]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BEA36C0BA3CFFAEDD92AA25FB7D6251" blockId="7.[810,1304,1583,1677]" box="[1001,1268,1583,1610]" pageId="7" pageNumber="8">
<emphasis id="B921EAD2BA3CFFAEDD92AA25FB7D6251" bold="true" box="[1001,1268,1583,1610]" pageId="7" pageNumber="8">
<taxonomicName id="4C554D43BA3CFFAEDD92AA25FB216251" authority="Boeters" authorityName="Boeters" authorityYear="2000" box="[1001,1192,1583,1610]" class="Gastropoda" family="Hydrobiidae" genus="Navarriella" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA3CFFAEDD92AA25FBD46252" bold="true" box="[1001,1117,1583,1610]" italics="true" pageId="7" pageNumber="8">Navarriella</emphasis>
Boeters
</taxonomicName>
, 2000
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34F654BBA3CFFAEDD51AA5CFA916295" box="[810,1304,1622,1677]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8BEA36C0BA3CFFAEDD51AA5CFC056275" blockId="7.[810,1304,1583,1677]" box="[810,908,1622,1645]" pageId="7" pageNumber="8">
<emphasis id="B921EAD2BA3CFFAEDD51AA5CFC056275" box="[810,908,1622,1645]" italics="true" pageId="7" pageNumber="8">Synonyms</emphasis>
</paragraph>
<paragraph id="8BEA36C0BA3CFFAEDD51AA7FFA916295" blockId="7.[810,1304,1583,1677]" box="[810,1304,1653,1677]" pageId="7" pageNumber="8">
<taxonomicName id="4C554D43BA3CFFAEDD51AA7FFA9D6295" authority="Boeters, 2000: 160 - 161" authorityName="Boeters" authorityPageNumber="160 - 161" authorityYear="2000" box="[810,1300,1653,1677]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="subGenus" subGenus="Navarriella">
<emphasis id="B921EAD2BA3CFFAEDD51AA7FFC056295" box="[810,908,1653,1677]" italics="true" pageId="7" pageNumber="8">Alzoniella</emphasis>
(
<emphasis id="B921EAD2BA3CFFAEDDE6AA7FFB846295" box="[925,1037,1653,1677]" italics="true" pageId="7" pageNumber="8">Navarriella</emphasis>
)
<bibRefCitation id="EFC44B31BA3CFFAEDA66AA7FFA9D6295" author="Boeters" box="[1053,1300,1653,1677]" firstAuthor="Boeters" page="160 - 161" pageId="7" pageNumber="8" pagination="151 - 63" refId="ref12022" refString="Boeters HD. Ne genus Alzoniella Giusti % Bodon, 1984, in France West European Hydrobiidae, 91. Basteria 2000; 64: 151 - 63." type="book chapter" year="2000">Boeters, 2000: 160–161</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F654BBA3CFFAEDD51AAA5FC4462FD" pageId="7" pageNumber="8" type="type_taxon">
<paragraph id="8BEA36C0BA3CFFAEDD51AAA5FC4462FD" blockId="7.[810,1459,1710,1766]" pageId="7" pageNumber="8">
<emphasis id="B921EAD2BA3CFFAEDD51AAA5FC2E62DE" box="[810,935,1711,1734]" italics="true" pageId="7" pageNumber="8">Type species:</emphasis>
<taxonomicName id="4C554D43BA3CFFAEDDD5AAA4FA9A62DE" authority="Paladilhe, 1874" authorityName="Paladilhe" authorityYear="1874" box="[942,1299,1710,1734]" class="Gastropoda" family="Assimineidae" genus="Paludinella" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="elliptica">
<emphasis id="B921EAD2BA3CFFAEDDD5AAA4FBE662DE" box="[942,1135,1710,1734]" italics="true" pageId="7" pageNumber="8">Paludinella elliptica</emphasis>
<bibRefCitation id="EFC44B31BA3CFFAEDA0EAAA4FA9A62DE" author="Paladilhe" box="[1141,1299,1710,1734]" firstAuthor="Paladilhe" pageId="7" pageNumber="8" pagination="1 - 38" refId="ref14785" refString="Paladilhe A. Monographie du nouveau genre Peringia, suivie des descriptions d'especes nouvelles de Paludinees francaises. Annales des Sciences Naturelles, Series 6, Zoologie et Paleontologie 1874; 1: 1 - 38." type="journal article" year="1874">Paladilhe, 1874</bibRefCitation>
</taxonomicName>
. Designated by
<bibRefCitation id="EFC44B31BA3CFFAEDD51AAC4FC4162FE" author="Boeters" box="[810,968,1742,1766]" firstAuthor="Boeters" pageId="7" pageNumber="8" pagination="151 - 63" refId="ref12022" refString="Boeters HD. Ne genus Alzoniella Giusti % Bodon, 1984, in France West European Hydrobiidae, 91. Basteria 2000; 64: 151 - 63." type="book chapter" year="2000">Boeters (2000)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F654BBA3CFFA1DA3BAB06FF4063A8" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BEA36C0BA3CFFAEDA3BAB06FB14633D" blockId="7.[810,1459,1804,1991]" box="[1088,1181,1804,1829]" pageId="7" pageNumber="8">
<emphasis id="B921EAD2BA3CFFAEDA3BAB06FB14633D" box="[1088,1181,1804,1829]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BEA36C0BA3CFFA1DD51AB38FF4063A8" blockId="7.[810,1459,1804,1991]" lastBlockId="8.[129,777,1725,1968]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Shell cylindrical with a rounded apex; aperture obliquely ovate; periostracum pale yellow to whitish; protoconch low and dome shaped. Operculum corneous, orangish, thin, pliable, oval, paucispiral, with a submarginal nucleus. Two pairs of basal cusps on the central radular tooth. Ctenidium well developed.
<collectingRegion id="4991F822BA33FFA1DF7BAAB4FEB562CD" box="[256,316,1726,1749]" country="Turkey" name="Bursa" pageId="8" pageNumber="9">Bursa</collectingRegion>
copulatrix large, pyriform, pedunculated, and lying against the posterior section of the albumen gland; two seminal receptacles with a long duct; Sr2 smaller than Sr1 and arising at the renal oviduct loop. Penis unpigmented, strap-like; distal end of the penis gradually tapering; more than two penial lobes. Nervous system scarcely pigmented, moderately concentrated with cerebral ganglia roughly equal in size.
</paragraph>
</subSubSection>
<caption id="DF2A6648BA33FFA1DEFAAA2BFDB56295" ID-DOI="http://doi.org/10.5281/zenodo.14284410" ID-Zenodo-Dep="14284410" httpUri="https://zenodo.org/record/14284410/files/figure.png" pageId="8" pageNumber="9" startId="8.[129,194,1569,1593]" targetBox="[130,1473,146,1539]" targetPageId="8" targetType="figure">
<paragraph id="8BEA36C0BA33FFA1DEFAAA2BFA156221" blockId="8.[129,1467,1569,1678]" box="[129,1436,1569,1594]" pageId="8" pageNumber="9">
<emphasis id="B921EAD2BA33FFA1DEFAAA2BFF516222" bold="true" box="[129,216,1569,1594]" pageId="8" pageNumber="9">Figure 5.</emphasis>
Geometric Morphometric analyses for the shells of
<taxonomicName id="4C554D43BA33FFA1DCB9AA2BFC176222" authorityName="Boeters" authorityYear="2000" box="[706,926,1569,1594]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="subGenus" subGenus="Navarriella">
<emphasis id="B921EAD2BA33FFA1DCB9AA2BFC946221" box="[706,797,1569,1593]" italics="true" pageId="8" pageNumber="9">Alzoniella</emphasis>
(
<emphasis id="B921EAD2BA33FFA1DD57AA2BFC1A6221" box="[812,915,1569,1593]" italics="true" pageId="8" pageNumber="9">Navarriella</emphasis>
)
</taxonomicName>
species based on 20 coordinates (eight landmarks and
</paragraph>
<paragraph id="8BEA36C0BA33FFA1DEFAAA34FDB56295" blockId="8.[129,1467,1569,1678]" pageId="8" pageNumber="9">12 semilandmarks). A, Principal Component Analysis (PCA) for species. B, Linear Discriminant Analysis (LDA) for species. C, PCA for populations. D, Thin-plate spline (TPS) plot displaying expansion (in red) and contraction (in blue) of the shell shape. Label abbreviations are explained in Supporting Information, Table S1.</paragraph>
</caption>
<subSubSection id="C34F654BBA33FFA0DA28AAB7FDFA657C" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BEA36C0BA33FFA1DA28AAB7FB2E62CF" blockId="8.[825,1475,1725,1976]" box="[1107,1191,1725,1751]" pageId="8" pageNumber="9">
<emphasis id="B921EAD2BA33FFA1DA28AAB7FB2E62CF" box="[1107,1191,1725,1751]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
</paragraph>
<paragraph id="8BEA36C0BA33FFA0DD42AAEEFDFA657C" blockId="8.[825,1475,1725,1976]" lastBlockId="9.[113,763,144,356]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<taxonomicName id="4C554D43BA33FFA1DD42AAEEFC2062E4" authorityName="Boeters" authorityYear="2000" box="[825,937,1764,1788]" class="Gastropoda" family="Hydrobiidae" genus="Navarriella" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA33FFA1DD42AAEEFC2062E4" box="[825,937,1764,1788]" italics="true" pageId="8" pageNumber="9">Navarriella</emphasis>
</taxonomicName>
is a monospecific genus, belonging to an independent lineage separate from
<taxonomicName id="4C554D43BA33FFA1DA06AB09FB566303" box="[1149,1247,1795,1819]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA33FFA1DA06AB09FB566303" box="[1149,1247,1795,1819]" italics="true" pageId="8" pageNumber="9">Alzoniella</emphasis>
</taxonomicName>
, and it is unclassified within the subfamily
<taxonomicName id="4C554D43BA33FFA1DA6EAB28FBF76322" authorityName="Radoman" authorityYear="1973" box="[1045,1150,1826,1850]" higherTaxonomySource="Manual Input" pageId="8" pageNumber="9" rank="subFamily" subFamily="Islamiinae">Islamiinae</taxonomicName>
(
<figureCitation id="136E2A45BA33FFA1DAF4AB28FB536322" box="[1167,1242,1826,1850]" captionStart="Figure3" captionStartId="6.[129,191,1243,1267]" captionTargetBox="[133,1469,149,1210]" captionTargetId="figure-157@6.[129,1473,144,1215]" captionTargetPageId="6" captionText="Figure3.Phylogenetic relationships ofhydrobiid taxa incorporating thespecies of Alzoniella(Navarriella).A,Bayesian tree ofthe family Hydrobiidae basedon theconcatenated dataset (COI and 18S), including some selected species ofthe hydrobiidsubfamilies (highlighted with vertical bars). Brancheswith bootstrap support (BS) values fromthe maximumlikelihood (ML) analysis and Bayesian posterior probabilities(BPP) greater than 75% and 0.95,respectively,are indicatedwith asterisks.Branches only with BS&gt; 75% are denoted with a triangle.Branchesonly with BPP&gt; 0.95are shownwith a square.B, Bayesian phylogenetictree ofspecimens of Alzoniella (Navarriella) based onthe concatenated dataset(COI, 16S, H3, and 28S).BS and BPP areprovided abovebranches when greaterthan 75% and 0.95,respectively.The first vertical bar representsthe taxonomic species of eachpopulationassigned in previous studies asAlzoniella (Navarriella) elliptica (in red)and Alzoniella (Navarriella) pellitica (in blue).The remaining vertical barsrefer to thespecies delimitation methods:ABGD,distance-based automatic gap discovery;GMYC, single-thresholdgeneralized mixed Yule-coalescent;bPTP, Bayesian approach ofthe Poisson Tree Processes;mPTP,multirate PoissonTree Processes.ThebPTP bar is displayed with its Bayesian support value.Labelabbreviations areexplained in Supporting Information,Table S2. Scale bars:expected change per site." figureDoi="http://doi.org/10.5281/zenodo.14284404" httpUri="https://zenodo.org/record/14284404/files/figure.png" pageId="8" pageNumber="9">Fig. 3A</figureCitation>
). The
<emphasis id="B921EAD2BA33FFA1DB5BAB29FAC56322" box="[1312,1356,1827,1850]" italics="true" pageId="8" pageNumber="9">COI</emphasis>
average sequence divergence with the
<typeStatus id="54EE8862BA33FFA1DA1AAB48FB066342" box="[1121,1167,1858,1882]" pageId="8" pageNumber="9">type</typeStatus>
species
<taxonomicName id="4C554D43BA33FFA1DA9CAB48FAD96342" box="[1255,1360,1858,1882]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="finalina">
<emphasis id="B921EAD2BA33FFA1DA9CAB48FAD96342" box="[1255,1360,1858,1882]" italics="true" pageId="8" pageNumber="9">A. finalina</emphasis>
</taxonomicName>
is 18.54%. Morphologically,
<taxonomicName id="4C554D43BA33FFA1DD95AB6BFBD96361" box="[1006,1104,1889,1913]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA33FFA1DD95AB6BFBD96361" box="[1006,1104,1889,1913]" italics="true" pageId="8" pageNumber="9">Alzoniella</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C554D43BA33FFA1DAAFAB6BFACD6361" authorityName="Boeters" authorityYear="2000" box="[1236,1348,1889,1913]" class="Gastropoda" family="Hydrobiidae" genus="Navarriella" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA33FFA1DAAFAB6BFACD6361" box="[1236,1348,1889,1913]" italics="true" pageId="8" pageNumber="9">Navarriella</emphasis>
</taxonomicName>
by the presence of two sessile seminal receptacles and two penial lobes.
<taxonomicName id="4C554D43BA33FFA1DD42AB95FC2063AF" authorityName="Boeters" authorityYear="2000" box="[825,937,1951,1975]" class="Gastropoda" family="Hydrobiidae" genus="Navarriella" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA33FFA1DD42AB95FC2063AF" box="[825,937,1951,1975]" italics="true" pageId="8" pageNumber="9">Navarriella</emphasis>
</taxonomicName>
has cylindrical shells with a height of
<quantity id="4CAD9B25BA33FFA1DB3CAB95FA4B63AF" box="[1351,1474,1951,1975]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="8" pageNumber="9" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 mm</quantity>
(
<bibRefCitation id="EFC44B31BA32FFA0DE07AC9AFE8464B0" author="Boeters" box="[124,269,144,168]" firstAuthor="Boeters" pageId="9" pageNumber="10" pagination="151 - 63" refId="ref12022" refString="Boeters HD. Ne genus Alzoniella Giusti % Bodon, 1984, in France West European Hydrobiidae, 91. Basteria 2000; 64: 151 - 63." type="book chapter" year="2000">Boeters 2000</bibRefCitation>
,
<bibRefCitation id="EFC44B31BA32FFA0DF58AC9AFEDE64B0" author="Boeters" box="[291,343,144,168]" firstAuthor="Boeters" pageId="9" pageNumber="10" pagination="149 - 56" refId="ref12051" refString="Boeters HD. A contribution to the knowledge of Alzoniella Giusti % Bodon 1984 in France unknown Westeuropean Hydrobiidae, 12 (Prosobranchia: Hydrobiidae). Archiv fur Molluskenkunde 2001; 129: 149 - 56. hups: // doi. org / 10.1127 / arch. moll / 129 / 2001 / 149" type="journal article" year="2001">2001</bibRefCitation>
, Arconada, Bolán &amp; Boeters, 2007), whereas the shells of
<taxonomicName id="4C554D43BA32FFA0DF21ACA5FE3564DF" box="[346,444,175,199]" class="Gastropoda" family="Hydrobiidae" genus="Alzoniella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA32FFA0DF21ACA5FE3564DF" box="[346,444,175,199]" italics="true" pageId="9" pageNumber="10">Alzoniella</emphasis>
</taxonomicName>
are conic to cylindrical-conic with a height of
<quantity id="4CAD9B25BA32FFA0DF6AACC4FE0264FE" box="[273,395,206,230]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.5" metricValueMin="0.7" pageId="9" pageNumber="10" unit="mm" value="1.6" valueMax="2.5" valueMin="0.7">0.7–2.5 mm</quantity>
(
<bibRefCitation id="EFC44B31BA32FFA0DFE1ACC5FD0C64FF" author="Giusti and Bodon" box="[410,645,207,231]" firstAuthor="Giusti" pageId="9" pageNumber="10" pagination="157 - 81" refId="ref13407" refString="Giusti F, Bodon M. Notulae malacologicae, XXXI Nuove Hydrobiidae dell'Italia nord-occidentale (Gastropoda: Prosobanchia). Archiv fur Molluskenkunde 1984; 114: 157 - 81." type="journal article" year="1984">Giusti and Bodon 1984</bibRefCitation>
).
<taxonomicName id="4C554D43BA32FFA0DCE1ACC5FF77651E" authority="Boeters, 2003" authorityName="Boeters" authorityYear="2003" class="Gastropoda" family="Hydrobiidae" genus="Guadiella" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA32FFA0DCE1ACC5FD7364FF" box="[666,762,207,231]" italics="true" pageId="9" pageNumber="10">Guadiella</emphasis>
<bibRefCitation id="EFC44B31BA32FFA0DE0AACE4FF77651E" author="Boeters" box="[113,254,238,262]" firstAuthor="Boeters" pageId="9" pageNumber="10" pagination="1 - 41" refId="ref12107" refString="Boeters HD. ºupplementary notes on Moitessieriidae and Hydrobiidae from the Iberian Peninsula (Gastropoda, Caenogastropoda). Basteria 2003; 67: 1 - 41." type="journal article" year="2003">Boeters, 2003</bibRefCitation>
</taxonomicName>
differs from
<taxonomicName id="4C554D43BA32FFA0DF04ACE4FE66651E" authorityName="Boeters" authorityYear="2000" box="[383,495,238,262]" class="Gastropoda" family="Hydrobiidae" genus="Navarriella" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B921EAD2BA32FFA0DF04ACE4FE66651E" box="[383,495,238,262]" italics="true" pageId="9" pageNumber="10">Navarriella</emphasis>
</taxonomicName>
by having a single seminal receptacle, a slender penis without penial lobes, and narrow, cylindrical to slightly conical shells with a height of
<quantity id="4CAD9B25BA32FFA0DC1EAD27FD73655C" box="[613,762,301,325]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="9" pageNumber="10" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.40–1.70 mm</quantity>
(
<bibRefCitation id="EFC44B31BA32FFA0DE07AD46FE8A657C" author="Boeters" box="[124,259,332,356]" firstAuthor="Boeters" pageId="9" pageNumber="10" pagination="1 - 41" refId="ref12107" refString="Boeters HD. ºupplementary notes on Moitessieriidae and Hydrobiidae from the Iberian Peninsula (Gastropoda, Caenogastropoda). Basteria 2003; 67: 1 - 41." type="journal article" year="2003">Boeters 2003</bibRefCitation>
, Arconada, Bolán &amp; Boeters, 2007).
</paragraph>
</subSubSection>
</treatment>
</document>
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