364 lines
42 KiB
XML
364 lines
42 KiB
XML
<document id="29F127BCC930B1BA7937407FA9A2EBE5" ID-DOI="10.17348/jbrit.v15.i2.1206" ID-Zenodo-Dep="14111022" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1634280027544" checkinUser="donat" docAuthor="Fawcett, Susan & Smith, Alan R." docDate="2021" docId="03B787F6FFBF9B1C6200787DFF1FFA6C" docLanguage="en" docName="2021_ThelypteridaceaeFerns_OACopy3.pdf.imf" docOrigin="Fort Worth, Texas, USA: BRIT Press" docTitle="Reholttumia" docType="treatment" docVersion="3" lastPageNumber="77" masterDocId="FF8EFF8EFFE99B4B602B7D49FFA7FFBB" masterDocTitle="A Generic Classification of the Thelypteridaceae" pageNumber="76" updateTime="1731444927364" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="977B88EB139425E2D48B046B63679846">
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<mods:title id="AF68276F25A4FC2DF668FD65022CFA60">A Generic Classification of the Thelypteridaceae</mods:title>
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</mods:titleInfo>
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<mods:name id="5CC8F9E5462771C11817FDC4CDE9574B" type="personal">
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<mods:roleTerm id="433D5200FBF56D5C790F0C7C7452B843">Author</mods:roleTerm>
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<mods:namePart id="30D69A894D7A350CE67A6780C675A4A6">Fawcett, Susan</mods:namePart>
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</mods:name>
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<mods:name id="F749BE1CBFE2D56126D07552CB938DD6" type="personal">
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<mods:roleTerm id="1863C55073AD3495A4864CE0BCFF759A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E71EE46F4418E06C1B456ACA9A48AABF">Smith, Alan R.</mods:namePart>
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<mods:typeOfResource id="802C1FFABCBC59C42843063D4EE9BF68">text</mods:typeOfResource>
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<mods:dateIssued id="D1AE9E146F9E5F2220703239F449095E">2021</mods:dateIssued>
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<mods:publisher id="9C3A5281D1EC47C0E7E3BC21F4DBFB6B">BRIT Press</mods:publisher>
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<mods:placeTerm id="1C144ABA105069169F5CDE9DBABED18A">Fort Worth, Texas, USA</mods:placeTerm>
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<mods:classification id="C9C3B3F8D5A17A97B6FD313C5B781E9E">book</mods:classification>
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<mods:identifier id="95FE3A649E45272A731A62A6680888C2" type="DOI">10.17348/jbrit.v15.i2.1206</mods:identifier>
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<mods:identifier id="B4E5D591085D98C16A82DF8485666F80" type="Zenodo-Dep">14111022</mods:identifier>
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<treatment id="03B787F6FFBF9B1C6200787DFF1FFA6C" LSID="urn:lsid:plazi:treatment:03B787F6FFBF9B1C6200787DFF1FFA6C" httpUri="http://treatment.plazi.org/id/03B787F6FFBF9B1C6200787DFF1FFA6C" lastPageId="87" lastPageNumber="77" pageId="86" pageNumber="76">
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<subSubSection id="C304656BFFBF9B1D6200787DFD58FAF5" box="[555,767,1332,1358]" pageId="86" pageNumber="76" type="nomenclature">
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<paragraph id="8BA136E0FFBF9B1D6200787DFD58FAF5" blockId="86.[555,767,1332,1358]" box="[555,767,1332,1358]" pageId="86" pageNumber="76">
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<heading id="D0E9818CFFBF9B1D6200787DFD58FAF5" allCaps="true" bold="true" box="[555,767,1332,1358]" centered="true" fontSize="11" level="2" pageId="86" pageNumber="76" reason="0">
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<taxonomicName id="4C1E4D63FFBF9B1D6200787DFD58FAF5" ID-CoL="8RTZR" box="[555,767,1332,1358]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="genus" status="gen. nov.">
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<emphasis id="B96AEAF2FFBF9B1D6200787DFD58FAF5" bold="true" box="[555,767,1332,1358]" pageId="86" pageNumber="76">REHOLTTUMIA</emphasis>
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</taxonomicName>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C304656BFFBF9B1D604A7814FDC2FA2C" pageId="86" pageNumber="76" type="reference_group">
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<paragraph id="8BA136E0FFBF9B1D604A7814FB6FFACA" blockId="86.[97,1224,1372,1394]" box="[97,1224,1372,1394]" pageId="86" pageNumber="76">
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<taxonomicName id="4C1E4D63FFBF9B1D604A7814FE14FAC9" authority="S.E. Fawc. & A.R.Sm." authorityName="S. E. Fawc. & A. R. Sm." box="[97,435,1372,1394]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="genus" status="gen. nov.">
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<emphasis id="B96AEAF2FFBF9B1D604A7814FF78FAC9" bold="true" box="[97,223,1373,1394]" pageId="86" pageNumber="76">Reholttumia</emphasis>
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S.E. Fawc. & A.R.Sm.
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</taxonomicName>
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||
,
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||
<emphasis id="B96AEAF2FFBF9B1D61927815FDA8FACA" bold="true" box="[441,527,1372,1393]" pageId="86" pageNumber="76">
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||
<taxonomicNameLabel id="A2595789FFBF9B1D61927815FDA8FACA" box="[441,527,1372,1393]" pageId="86" pageNumber="76" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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||
—
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<typeStatus id="54A58842FFBF9B1D620A7829FDE7FACB" box="[545,576,1376,1392]" pageId="86" pageNumber="76">
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||
<collectionCode id="ED0FAE25FFBF9B1D620A7829FDE7FACB" box="[545,576,1376,1392]" pageId="86" pageNumber="76">TYPE</collectionCode>
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||
</typeStatus>
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||
:
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<taxonomicName id="4C1E4D63FFBF9B1D62637829FC6AFACA" authority="(C. Presl) S. E. Fawc. & A. R. Sm." authorityName="S. E. Fawc. & A. R. Sm." baseAuthorityName="C. Presl" box="[584,973,1376,1393]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="species" species="nitidula">
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<emphasis id="B96AEAF2FFBF9B1D62637829FD7AFACA" box="[584,733,1376,1393]" italics="true" pageId="86" pageNumber="76">Reholttumia nitidula</emphasis>
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(C.Presl) S.E. Fawc.& A.R.Sm.
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</taxonomicName>
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[=
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<taxonomicName id="4C1E4D63FFBF9B1D63CA7829FB63FACA" authority="C. Presl" authorityName="C. Presl" box="[993,1220,1376,1393]" class="Polypodiopsida" family="Thelypteridaceae" genus="Nephrodium" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="species" species="nitidulum">
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<emphasis id="B96AEAF2FFBF9B1D63CA7829FB22FACA" box="[993,1157,1376,1393]" italics="true" pageId="86" pageNumber="76">Nephrodium nitidulum</emphasis>
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C.Presl
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</taxonomicName>
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]
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</paragraph>
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<paragraph id="8BA136E0FFBF9B1D60AA78CCFDC2FA2C" blockId="86.[129,613,1413,1431]" box="[129,613,1413,1431]" pageId="86" pageNumber="76">
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For species synonymy, see
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<bibRefCitation id="EF8F4B11FFBF9B1D616178CFFE64FA2C" author="Holttum" box="[330,451,1414,1431]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="86" pageNumber="76" pagination="293 - 325" part="21" refId="ref109522" refString="HOLTTUM, R. E. 1973 a. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293 - 325." title="Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai" type="journal article" year="1973" yearSuffix="a">Holttum (1973a</bibRefCitation>
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,
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||
<bibRefCitation id="EF8F4B11FFBF9B1D61E278CFFE50FA2C" author="Holttum" box="[457,503,1414,1431]" firstAuthor="Holttum" journalOrPublisher="J. S. African Bot." pageId="86" pageNumber="76" pagination="123 - 168" part="40" refId="ref109577" refString="HOLTTUM, R. E. 1974 a. Thelypteridaceae of Africa and adjacent islands. J. S. African Bot. 40: 123 - 168." title="Thelypteridaceae of Africa and adjacent islands" type="journal article" year="1974" yearSuffix="a">1974a</bibRefCitation>
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,
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<bibRefCitation id="EF8F4B11FFBF9B1D61D578CFFD89FA2C" author="Holttum" box="[510,558,1414,1431]" firstAuthor="Holttum" journalOrPublisher="Allertonia" pageId="86" pageNumber="76" pagination="169 - 234" part="1" refId="ref109882" refString="HOLTTUM, R. E. 1977 b. The family Thelypteridaceae in the Pacific and Australasia. Allertonia 1: 169 - 234." title="The family Thelypteridaceae in the Pacific and Australasia" type="journal article" year="1977" yearSuffix="b">1977b</bibRefCitation>
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,
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<bibRefCitation id="EF8F4B11FFBF9B1D621F78CFFDFCFA2C" author="Holttum" box="[564,603,1414,1431]" firstAuthor="Holttum" journalOrPublisher="Flora Malesiana" pageId="86" pageNumber="76" pagination="334 - 560" part="1" refId="ref109955" refString="HOLTTUM, R. E. 1982. Thelypteridaceae. Flora Malesiana, Ser. II, 1 (5): 334 - 560." title="Thelypteridaceae" type="journal article" year="1982">1982</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C304656BFFBF9B1D604A78E5FF56F9FA" pageId="86" pageNumber="76" type="etymology">
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<paragraph id="8BA136E0FFBF9B1D604A78E5FF56F9FA" blockId="86.[97,1226,1452,1794]" pageId="86" pageNumber="76">
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<emphasis id="B96AEAF2FFBF9B1D604A78E5FF79FA7A" box="[97,222,1452,1473]" italics="true" pageId="86" pageNumber="76">Etymology.—</emphasis>
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The name honors Richard Eric Holttum (1895–1990), whose keen insights, perseverance, and fundamental contributions to our knowledge of
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<taxonomicName id="4C1E4D63FFBF9B1D61F87884FDDFFA59" baseAuthorityName="Holttum" baseAuthorityYear="1976" box="[467,632,1485,1506]" class="Polypodiopsida" family="Thelypteridaceae" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="family">Thelypteridaceae</taxonomicName>
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enabled the present work. Holttum’s focus was especially on species in the Old World tropics where the family is most diverse and least known. His careful examination of
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<typeStatus id="54A58842FFBF9B1D64B578A5FB6DF9BA" box="[1182,1226,1516,1537]" pageId="86" pageNumber="76">type</typeStatus>
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material in numerous
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<taxonomicName id="4C1E4D63FFBF9B1D61127B44FED6F999" box="[313,369,1549,1570]" genus="The" pageId="86" pageNumber="76" rank="species" species="major">major</taxonomicName>
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herbaria resolved many problems that had plagued the taxonomy of the group for more than a century.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C304656BFFBF9B1C60B97B04FE55FDCC" lastPageId="87" lastPageNumber="77" pageId="86" pageNumber="76" type="description">
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||
<paragraph id="8BA136E0FFBF9B1C60B97B04FE55FDCC" blockId="86.[97,1226,1452,1794]" lastBlockId="87.[121,1250,162,1495]" lastPageId="87" lastPageNumber="77" pageId="86" pageNumber="76">
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Plants terrestrial, medium-sized to large (to
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<quantity id="4CE69B05FFBF9B1D627C7B04FDD9F9DA" box="[599,638,1612,1634]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="76" unit="m" value="2.0">2 m</quantity>
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tall);
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||
<emphasis id="B96AEAF2FFBF9B1D62977B04FCB9F9D9" bold="true" box="[700,798,1613,1634]" pageId="86" pageNumber="76">rhizomes</emphasis>
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||
ascending, erect, short-creeping or rarely long-creeping, rhizome scales brown, often caducous;
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||
<emphasis id="B96AEAF2FFBF9B1D62527B24FD1AF939" bold="true" box="[633,701,1645,1666]" pageId="86" pageNumber="76">fronds</emphasis>
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||
monomorphic, erect to arching, rarely pendant, pinnate-pinnatifid;
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||
<emphasis id="B96AEAF2FFBF9B1D60D57BC4FE9DF919" bold="true" box="[254,314,1677,1698]" pageId="86" pageNumber="76">stipes</emphasis>
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||
stramineous, less often brown;
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||
<emphasis id="B96AEAF2FFBF9B1D625E7BC4FD4BF91A" bold="true" box="[629,748,1676,1698]" pageId="86" pageNumber="76">stipe scales</emphasis>
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||
thick, brown, broadly ovate to linear-lanceolate, glabrous or with marginal setae, apparently caducous, rarely extending to rachis (
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<taxonomicName id="4C1E4D63FFBF9B1D63527BE4FC5AF97A" box="[889,1021,1708,1730]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="86" pageNumber="76" phylum="Tracheophyta" rank="species" species="rodigasiana">
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<emphasis id="B96AEAF2FFBF9B1D63527BE4FC5AF97A" box="[889,1021,1708,1730]" italics="true" pageId="86" pageNumber="76">R. rodigasiana</emphasis>
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</taxonomicName>
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);
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<emphasis id="B96AEAF2FFBF9B1D643A7BE5FBF4F97A" bold="true" box="[1041,1107,1708,1729]" pageId="86" pageNumber="76">blades</emphasis>
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membranaceous to chartaceous, drying green, sometimes brown-olivaceous, but never reddish, blade apex gradually reduced, proximal pinnae not reduced, or gradually to subabruptly reduced to small, sometimes auriculate pinnae, the smallest typically at least
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<quantity id="4CE69B05FFBE9B1C61377DEBFEEAFF0C" box="[284,333,162,183]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="77" unit="cm" value="1.0">1 cm</quantity>
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long, rarely less than
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||
<quantity id="4CE69B05FFBE9B1C62087DEBFDF8FF0C" box="[547,607,162,183]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="87" pageNumber="77" unit="mm" value="5.0">5 mm</quantity>
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, proliferous buds absent;
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<emphasis id="B96AEAF2FFBE9B1C63737DEBFC39FF0C" bold="true" box="[856,926,162,183]" pageId="87" pageNumber="77">pinnae</emphasis>
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||
sessile to subsessile, pinna-bases truncate, margins usually incised 1/3–3/5 to costae, segment apices typically rounded, sometimes truncate (
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||
<taxonomicName id="4C1E4D63FFBE9B1C64BC7D8BFF38FF4C" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="truncata">
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||
<emphasis id="B96AEAF2FFBE9B1C64BC7D8BFF38FF4C" italics="true" pageId="87" pageNumber="77">R. truncata</emphasis>
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||
</taxonomicName>
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||
), rarely acute (
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||
<taxonomicName id="4C1E4D63FFBE9B1C61017DABFE27FF4C" box="[298,384,226,247]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="costata">
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||
<emphasis id="B96AEAF2FFBE9B1C61017DABFE27FF4C" box="[298,384,226,247]" italics="true" pageId="87" pageNumber="77">R.costata</emphasis>
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||
</taxonomicName>
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||
), proximal pinnae gradually tapered towards bases or with acroscopic auricles;
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||
<emphasis id="B96AEAF2FFBE9B1C64457DABFB04FF4C" bold="true" box="[1134,1187,226,247]" pageId="87" pageNumber="77">veins</emphasis>
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||
of one to three pairs anastomosing to form an excurrent veinlet running to sinus membrane, or ending below it (
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<taxonomicName id="4C1E4D63FFBE9B1C64587C4BFB74FEAD" box="[1139,1235,257,279]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="ecallosa">
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||
<emphasis id="B96AEAF2FFBE9B1C64587C4BFB74FEAD" box="[1139,1235,257,279]" italics="true" pageId="87" pageNumber="77">R. ecallosa</emphasis>
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||
</taxonomicName>
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||
), subsequent veins ending at margin above sinus;
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||
<emphasis id="B96AEAF2FFBE9B1C62727C68FD6BFE8D" bold="true" box="[601,716,289,310]" pageId="87" pageNumber="77">aerophores</emphasis>
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||
absent, or present as swelling or discoloration at pinna-base, never peg-like;
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||
<emphasis id="B96AEAF2FFBE9B1C614A7C0BFD8EFEEC" bold="true" box="[353,553,322,343]" pageId="87" pageNumber="77">indument abaxially</emphasis>
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||
of short, erect hairs sometimes present on axes, laminar tissue typically glabrous, but erect acicular hairs sometimes present (
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||
<taxonomicName id="4C1E4D63FFBE9B1C62AF7C2BFD42FECD" box="[644,741,353,375]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="ecallosa">
|
||
<emphasis id="B96AEAF2FFBE9B1C62AF7C2BFD42FECD" box="[644,741,353,375]" italics="true" pageId="87" pageNumber="77">R. ecallosa</emphasis>
|
||
</taxonomicName>
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||
);
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||
<emphasis id="B96AEAF2FFBE9B1C62D17C2BFC63FECC" bold="true" box="[762,964,354,375]" pageId="87" pageNumber="77">indument adaxially</emphasis>
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||
often restricted to antrorsely arching hyaline or reddish hairs along rachis and costae, but short-erect hairs sometimes present on axes;
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||
<emphasis id="B96AEAF2FFBE9B1C64A17CC8FB45FE2D" bold="true" box="[1162,1250,385,406]" pageId="87" pageNumber="77">pustules</emphasis>
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||
sometimes present on adaxial laminae, these large and irregular or minute, colorless, and glandular (
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||
<taxonomicName id="4C1E4D63FFBE9B1C646E7CEBFB02FE0D" box="[1093,1189,417,439]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="ecallosa">
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||
<emphasis id="B96AEAF2FFBE9B1C646E7CEBFB02FE0D" box="[1093,1189,417,439]" italics="true" pageId="87" pageNumber="77">R. ecallosa</emphasis>
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||
</taxonomicName>
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||
);
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||
<emphasis id="B96AEAF2FFBE9B1C64927CEBFB46FE0C" bold="true" box="[1209,1249,418,439]" pageId="87" pageNumber="77">sori</emphasis>
|
||
round, discrete, typically medial on veins, sometimes inframedial, almost always indusiate (except
|
||
<taxonomicName id="4C1E4D63FFBE9B1C64357C8BFBD1FE6C" box="[1054,1142,450,471]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="costata">
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||
<emphasis id="B96AEAF2FFBE9B1C64357C8BFBD1FE6C" box="[1054,1142,450,471]" italics="true" pageId="87" pageNumber="77">R. costata</emphasis>
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</taxonomicName>
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sometimes exindusiate), though shriveled indusia in mature sori may be obscure, indusia dark brown, glabrous or with acicular hairs;
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<emphasis id="B96AEAF2FFBE9B1C60987F4BFEBFFDAC" bold="true" box="[179,280,514,535]" pageId="87" pageNumber="77">sporangia</emphasis>
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glabrous, or with stipitate glands on capsules, these usually colorless, rarely bearing setulae (except
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<taxonomicName id="4C1E4D63FFBE9B1C64E67F4BFF65FD8C" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="oxyoura">
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||
<emphasis id="B96AEAF2FFBE9B1C64E67F4BFF65FD8C" italics="true" pageId="87" pageNumber="77">R. oxyoura</emphasis>
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</taxonomicName>
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);
|
||
<emphasis id="B96AEAF2FFBE9B1C60FE7F68FEB0FD8D" bold="true" box="[213,279,545,566]" pageId="87" pageNumber="77">spores</emphasis>
|
||
tan to brown, perine with tubercles or robust echinae (
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C63097F6BFC64FD8C" author="Patel" box="[802,963,546,567]" etAl="et al." firstAuthor="Patel" journalOrPublisher="Int. J. Pl. Sci." pageId="87" pageNumber="77" pagination="1016 - 1035" part="180" refId="ref112729" refString="PATEL, N., S. FAWCETT, M. SUNDUE, & J. BUDKE. 2019 a. Evolution of perine morphology in Thelypteridaceae. Int. J. Pl. Sci. 180: 1016 - 1035." title="Evolution of perine morphology in Thelypteridaceae" type="journal article" year="2019" yearSuffix="a">Patel et al. 2019a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C63E77F6BFB12FD8C" author="Tryon & Lugardon" box="[972,1205,546,567]" firstAuthor="Tryon" journalOrPublisher="Springer-Verlag, New York, U. S. A" pageId="87" pageNumber="77" refId="ref115218" refString="TRYON, A. F. & B. LUGARDON. 1991. Spores of Pteridophyta. Springer-Verlag, New York, U. S. A." title="Spores of Pteridophyta" type="book" year="1991">Tryon & Lugardon 1991</bibRefCitation>
|
||
);
|
||
<emphasis id="B96AEAF2FFBE9B1C64EE7F6BFB77FD8C" box="[1221,1232,546,567]" italics="true" pageId="87" pageNumber="77">x</emphasis>
|
||
= 36, based on five species counts, mostly diploid, but
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62417F0BFD76FDEC" box="[618,721,578,599]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="truncata">
|
||
<emphasis id="B96AEAF2FFBE9B1C62417F0BFD76FDEC" box="[618,721,578,599]" italics="true" pageId="87" pageNumber="77">R. truncata</emphasis>
|
||
</taxonomicName>
|
||
has both diploid and tetraploid cytotypes. No interspecific or intergeneric hybrids are known.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBE9B1C60827FC8FCEAFC6C" pageId="87" pageNumber="77" type="diagnosis">
|
||
<paragraph id="8BA136E0FFBE9B1C60827FC8FCEAFC6C" blockId="87.[121,1250,162,1495]" pageId="87" pageNumber="77">
|
||
<emphasis id="B96AEAF2FFBE9B1C60827FC8FE36FD2D" box="[169,401,641,662]" italics="true" pageId="87" pageNumber="77">
|
||
Diagnosis.—
|
||
<taxonomicName id="4C1E4D63FFBE9B1C610B7FC8FE36FD2D" box="[288,401,641,662]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">Reholttumia</taxonomicName>
|
||
</emphasis>
|
||
is distinguished from
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62447FCBFCA6FD2C" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[623,769,642,663]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C62447FCBFCA6FD2C" box="[623,769,642,663]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2595789FFBE9B1C632C7FCBFC84FD2C" box="[775,803,642,663]" pageId="87" pageNumber="77" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C632C7FCBFCBAFD2C" box="[775,797,642,663]" italics="true" pageId="87" pageNumber="77">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
by having inconspicuous or swollen (vs. peglike) aerophores; segment apices typically rounded or truncate (vs. broadly acute); sporangia sometimes bearing stipitate glands, but rarely setulose; proximal pinnae gradually or subabruptly (vs. abruptly) reduced, rarely reduced to less than
|
||
<quantity id="4CE69B05FFBE9B1C60867FABFF40FD4C" box="[173,231,738,759]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="87" pageNumber="77" unit="mm" value="5.0">5 mm</quantity>
|
||
long; and laminae drying green, or brown-olivaceous (vs. sometimes reddish). The pinna segments of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C60527E48FF4DFCAD" box="[121,234,769,790]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C60527E48FF4DFCAD" box="[121,234,769,790]" italics="true" pageId="87" pageNumber="77">Reholttumia</emphasis>
|
||
</taxonomicName>
|
||
typically have about 7 veins per cm (vs. about 11 veins per cm in
|
||
<taxonomicName id="4C1E4D63FFBE9B1C63537E4BFBACFCAC" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[888,1035,770,791]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C63537E4BFBACFCAC" box="[888,1035,770,791]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2595789FFBE9B1C64397E4BFB8BFCAC" box="[1042,1068,770,791]" pageId="87" pageNumber="77" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C64397E4BFB8EFCAC" box="[1042,1065,770,791]" italics="true" pageId="87" pageNumber="77">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
), but species with smaller proportions, and narrower pinnae (e.g.,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62637E6BFD6DFC8C" box="[584,714,802,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="macroptera">
|
||
<emphasis id="B96AEAF2FFBE9B1C62637E6BFD6DFC8C" box="[584,714,802,823]" italics="true" pageId="87" pageNumber="77">R. macroptera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62FD7E6BFCF8FC8D" box="[726,863,801,823]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="kerintjiensis">
|
||
<emphasis id="B96AEAF2FFBE9B1C62FD7E6BFCF8FC8D" box="[726,863,801,823]" italics="true" pageId="87" pageNumber="77">R. kerintjiensis</emphasis>
|
||
</taxonomicName>
|
||
) may have as many as 11 veins per cm. Another species,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C61367E0BFECCFCED" box="[285,363,833,855]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="laevis">
|
||
<emphasis id="B96AEAF2FFBE9B1C61367E0BFECCFCED" box="[285,363,833,855]" italics="true" pageId="87" pageNumber="77">R. laevis</emphasis>
|
||
</taxonomicName>
|
||
, is atypical in the genus in having oblique, asymmetric pinnae, small stature, and a creeping rhizome; it is also the earliest-diverging species in our sample.
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62E47E28FC8EFCCD" box="[719,809,865,886]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C62E47E28FC8EFCCD" box="[719,809,865,886]" italics="true" pageId="87" pageNumber="77">Christella</emphasis>
|
||
</taxonomicName>
|
||
differs by generally having abundant hairs on axes and laminae, and often by having characteristic pear-shaped orangish laminar and sporangial glands; it also lacks laminar pustules.
|
||
<taxonomicName id="4C1E4D63FFBE9B1C610E7EE8FE6EFC0D" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[293,457,929,950]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C610E7EE8FE6EFC0D" box="[293,457,929,950]" italics="true" pageId="87" pageNumber="77">Sphaerostephanos</emphasis>
|
||
</taxonomicName>
|
||
differs in sometimes having yellow, sessile spherical glands.
|
||
<taxonomicName id="4C1E4D63FFBE9B1C64267EE8FB25FC0D" box="[1037,1154,929,950]" class="Polypodiopsida" family="Thelypteridaceae" genus="Plesioneuron" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C64267EE8FB25FC0D" box="[1037,1154,929,950]" italics="true" pageId="87" pageNumber="77">Plesioneuron</emphasis>
|
||
</taxonomicName>
|
||
differs by having dark (vs. tan) spores, and pinnae usually incised nearly to the costae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBE9B1C60827EA8FC67FB2C" pageId="87" pageNumber="77" type="biology_ecology">
|
||
<paragraph id="8BA136E0FFBE9B1C60827EA8FC67FB2C" blockId="87.[121,1250,162,1495]" pageId="87" pageNumber="77">
|
||
<emphasis id="B96AEAF2FFBE9B1C60827EA8FE34FC4D" box="[169,403,993,1014]" italics="true" pageId="87" pageNumber="77">Biogeography and ecology</emphasis>
|
||
.—The 30 species of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C627E7EA8FD62FC4D" box="[597,709,993,1014]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C627E7EA8FD62FC4D" box="[597,709,993,1014]" italics="true" pageId="87" pageNumber="77">Reholttumia</emphasis>
|
||
</taxonomicName>
|
||
occur in forests and openings, often beside streams, generally at lower elevations, but some species reach about
|
||
<quantity id="4CE69B05FFBE9B1C62AE794BFD76FBAC" box="[645,721,1026,1047]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="87" pageNumber="77" unit="m" value="2300.0">2300 m</quantity>
|
||
. The greatest species diversity is in
|
||
<collectingCountry id="F3097670FFBE9B1C6434794BFBCCFBAC" box="[1055,1131,1026,1047]" name="Malaysia" pageId="87" pageNumber="77">Malesia</collectingCountry>
|
||
, but a dozen species occur in the Pacific and Australasia (
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C6231796BFD09FB8C" author="Holttum" box="[538,686,1058,1079]" firstAuthor="Holttum" journalOrPublisher="Allertonia" pageId="87" pageNumber="77" pagination="169 - 234" part="1" refId="ref109882" refString="HOLTTUM, R. E. 1977 b. The family Thelypteridaceae in the Pacific and Australasia. Allertonia 1: 169 - 234." title="The family Thelypteridaceae in the Pacific and Australasia" type="journal article" year="1977" yearSuffix="b">Holttum 1977b</bibRefCitation>
|
||
), with one species reaching the Hawaiian Islands (
|
||
<taxonomicName id="4C1E4D63FFBE9B1C64A0796BFF0BFBEC" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="hudsoniana">
|
||
<emphasis id="B96AEAF2FFBE9B1C64A0796BFF0BFBEC" italics="true" pageId="87" pageNumber="77">R. hudsoniana</emphasis>
|
||
</taxonomicName>
|
||
). The western range of the genus nearly corresponds to that of its most widespread and highly variable species,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C64E6790BFF6FFBCC" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="truncata">
|
||
<emphasis id="B96AEAF2FFBE9B1C64E6790BFF6FFBCC" italics="true" pageId="87" pageNumber="77">R. truncata</emphasis>
|
||
</taxonomicName>
|
||
, which occurs in
|
||
<collectingCountry id="F3097670FFBE9B1C6158792BFE6AFBCC" box="[371,461,1122,1143]" name="Australia" pageId="87" pageNumber="77">Australia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C61FD792BFDB3FBCC" box="[470,532,1122,1143]" name="China" pageId="87" pageNumber="77">China</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C6236792BFDF5FBCC" box="[541,594,1122,1143]" name="India" pageId="87" pageNumber="77">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C6271792BFD35FBCC" box="[602,658,1122,1143]" name="Japan" pageId="87" pageNumber="77">Japan</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C62B7792BFD6CFBCC" box="[668,715,1122,1143]" name="Laos" pageId="87" pageNumber="77">Laos</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C62FF792BFC8BFBCC" box="[724,812,1122,1143]" name="Malaysia" pageId="87" pageNumber="77">Malaysia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C631F792BFC27FBCC" box="[820,896,1122,1143]" name="Malaysia" pageId="87" pageNumber="77">Malesia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C63A1792BFC4EFBCC" box="[906,1001,1122,1143]" name="Myanmar" pageId="87" pageNumber="77">Myanmar</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C63D9792BFBF6FBCC" box="[1010,1105,1122,1143]" name="Sri Lanka" pageId="87" pageNumber="77">Sri Lanka</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3097670FFBE9B1C6471792BFB13FBCC" box="[1114,1204,1122,1143]" name="Thailand" pageId="87" pageNumber="77">Thailand</collectingCountry>
|
||
, and
|
||
<collectingCountry id="F3097670FFBE9B1C605279CBFF69FB2C" box="[121,206,1154,1175]" name="Vietnam" pageId="87" pageNumber="77">Vietnam</collectingCountry>
|
||
. The genus is represented on
|
||
<collectingCountry id="F3097670FFBE9B1C61C879CBFDF1FB2C" box="[483,598,1154,1175]" name="Madagascar" pageId="87" pageNumber="77">Madagascar</collectingCountry>
|
||
by at least one species,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C631979CBFC1DFB2C" box="[818,954,1154,1175]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="remotipinna">
|
||
<emphasis id="B96AEAF2FFBE9B1C631979CBFC1DFB2C" box="[818,954,1154,1175]" italics="true" pageId="87" pageNumber="77">R. remotipinna</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C304656BFFBE9B1C608279EBFF1FFA6C" pageId="87" pageNumber="77" type="discussion">
|
||
<paragraph id="8BA136E0FFBE9B1C608279EBFF1FFA6C" blockId="87.[121,1250,162,1495]" pageId="87" pageNumber="77">
|
||
<emphasis id="B96AEAF2FFBE9B1C608279EBFE49FB0D" box="[169,494,1185,1207]" italics="true" pageId="87" pageNumber="77">Taxonomic and phylogenetic studies</emphasis>
|
||
.—All constituent species of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62D479E8FCC8FB0D" box="[767,879,1185,1206]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C62D479E8FCC8FB0D" box="[767,879,1185,1206]" italics="true" pageId="87" pageNumber="77">Reholttumia</emphasis>
|
||
</taxonomicName>
|
||
were treated within
|
||
<taxonomicName id="4C1E4D63FFBE9B1C641879EBFB63FB0C" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[1075,1220,1186,1207]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C641879EBFB63FB0C" box="[1075,1220,1186,1207]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C6052798BFEAAFB6C" author="Holttum" box="[121,269,1218,1239]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="87" pageNumber="77" pagination="17 - 52" part="19" refId="ref109463" refString="HOLTTUM, R. E. 1971. Studies in the family Thelypteridaceae III. A new system of genera in the Old World. Blumea 19: 17 - 52." title="Studies in the family Thelypteridaceae III. A new system of genera in the Old World" type="journal article" year="1971">Holttum (1971</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C6133798BFEF2FB6C" author="Holttum" box="[280,341,1218,1239]" firstAuthor="Holttum" journalOrPublisher="Blumea" pageId="87" pageNumber="77" pagination="293 - 325" part="21" refId="ref109522" refString="HOLTTUM, R. E. 1973 a. Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai. Blumea 21: 293 - 325." title="Studies in the family Thelypteridaceae V. The genus Pneumatopteris Nakai" type="journal article" year="1973" yearSuffix="a">1973a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C614B798BFE3BFB6C" author="Holttum" box="[352,412,1218,1239]" firstAuthor="Holttum" journalOrPublisher="J. S. African Bot." pageId="87" pageNumber="77" pagination="123 - 168" part="40" refId="ref109577" refString="HOLTTUM, R. E. 1974 a. Thelypteridaceae of Africa and adjacent islands. J. S. African Bot. 40: 123 - 168." title="Thelypteridaceae of Africa and adjacent islands" type="journal article" year="1974" yearSuffix="a">1974a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C618D798BFE42FB6C" author="Holttum" box="[422,485,1218,1239]" firstAuthor="Holttum" journalOrPublisher="Allertonia" pageId="87" pageNumber="77" pagination="169 - 234" part="1" refId="ref109882" refString="HOLTTUM, R. E. 1977 b. The family Thelypteridaceae in the Pacific and Australasia. Allertonia 1: 169 - 234." title="The family Thelypteridaceae in the Pacific and Australasia" type="journal article" year="1977" yearSuffix="b">1977b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8F4B11FFBE9B1C61DB798BFD84FB6C" author="Holttum" box="[496,547,1218,1239]" firstAuthor="Holttum" journalOrPublisher="Flora Malesiana" pageId="87" pageNumber="77" pagination="334 - 560" part="1" refId="ref109955" refString="HOLTTUM, R. E. 1982. Thelypteridaceae. Flora Malesiana, Ser. II, 1 (5): 334 - 560." title="Thelypteridaceae" type="journal article" year="1982">1982</bibRefCitation>
|
||
). However, his taxonomic concept of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C63B7798BFB95FB6C" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[924,1074,1218,1239]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C63B7798BFB95FB6C" box="[924,1074,1218,1239]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
has proven to be extremely broad and heterogeneous, as it encompasses species from ten distinct lineages, which we treat herein within existing genera or in newly described genera (see
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62AB784BFCBDFAAC" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[640,794,1282,1303]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C62AB784BFCBDFAAC" box="[640,794,1282,1303]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
for further discussion). Because the
|
||
<typeStatus id="54A58842FFBE9B1C64B37848FB62FAAD" box="[1176,1221,1281,1302]" pageId="87" pageNumber="77">type</typeStatus>
|
||
of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C6052786BFEAEFA8C" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[121,265,1314,1335]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C6052786BFEAEFA8C" box="[121,265,1314,1335]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B96AEAF2FFBE9B1C613F786BFEC8FA8D" box="[276,367,1313,1335]" italics="true" pageId="87" pageNumber="77">
|
||
<taxonomicName id="4C1E4D63FFBE9B1C613F786BFECBFA8D" box="[276,364,1313,1335]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="species" species="callosa">P.callosa</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
is in a small clade that is sister to
|
||
<taxonomicName id="4C1E4D63FFBE9B1C62987868FCF0FA8D" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[691,855,1313,1334]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C62987868FCF0FA8D" box="[691,855,1313,1334]" italics="true" pageId="87" pageNumber="77">Sphaerostephanos</emphasis>
|
||
</taxonomicName>
|
||
(a genus of about 200 species;
|
||
<figureCitation id="13252A65FFBE9B1C6456786BFB14FA8C" box="[1149,1203,1314,1335]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="87" pageNumber="77">
|
||
<emphasis id="B96AEAF2FFBE9B1C6456786BFB14FA8C" bold="true" box="[1149,1203,1314,1335]" pageId="87" pageNumber="77">Fig. 1</emphasis>
|
||
</figureCitation>
|
||
), we recognize
|
||
<taxonomicName id="4C1E4D63FFBE9B1C60F5780BFED6FAEC" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[222,369,1346,1367]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C60F5780BFED6FAEC" box="[222,369,1346,1367]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
in a greatly restricted sense, and segregate
|
||
<taxonomicName id="4C1E4D63FFBE9B1C63357808FC28FAED" box="[798,911,1345,1366]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C63357808FC28FAED" box="[798,911,1345,1366]" italics="true" pageId="87" pageNumber="77">Reholttumia</emphasis>
|
||
</taxonomicName>
|
||
—a morphologically coherent and large clade that is sister to
|
||
<taxonomicName id="4C1E4D63FFBE9B1C615A782BFDA5FACC" baseAuthorityName="Holttum" baseAuthorityYear="1973" box="[369,514,1378,1399]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C615A782BFDA5FACC" box="[369,514,1378,1399]" italics="true" pageId="87" pageNumber="77">Pneumatopteris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2595789FFBE9B1C622D782BFD85FACD" box="[518,546,1377,1399]" pageId="87" pageNumber="77" sensu="stricto">
|
||
<emphasis id="B96AEAF2FFBE9B1C622D782BFDBBFACC" box="[518,540,1378,1399]" italics="true" pageId="87" pageNumber="77">s.s</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
plus
|
||
<taxonomicName id="4C1E4D63FFBE9B1C627F7828FD5EFACD" baseAuthorityName="Holttum" baseAuthorityYear="1982" box="[596,761,1377,1398]" class="Polypodiopsida" family="Thelypteridaceae" genus="Sphaerostephanos" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C627F7828FD5EFACD" box="[596,761,1377,1398]" italics="true" pageId="87" pageNumber="77">Sphaerostephanos</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B96AEAF2FFBE9B1C632E782BFCE6FACC" bold="true" box="[773,833,1378,1399]" pageId="87" pageNumber="77">
|
||
<figureCitation id="13252A65FFBE9B1C632E782BFC9BFACC" box="[773,828,1378,1399]" captionStart="FIG" captionStartId="21.[121,129,1674,1694]" captionTargetBox="[123,1242,160,1631]" captionTargetId="figure-8@21.[121,1246,150,1639]" captionTargetPageId="21" captionText="FIG. 1. A synoptical treeillustrating relationships among genera of Thelypteridaceae based on Fawcett et al.(in press)." figureDoi="http://doi.org/10.5281/zenodo.14111042" httpUri="https://zenodo.org/record/14111042/files/figure.png" pageId="87" pageNumber="77">Fig. 1</figureCitation>
|
||
;
|
||
</emphasis>
|
||
Fawcett et al. in press). This new genus represents the largest segregate of
|
||
<taxonomicName id="4C1E4D63FFBE9B1C618B78CBFD47FA2D" authority="sensu Holttum. In" authorityName="sensu Holttum. In" authorityYear="1982" box="[416,736,1409,1431]" class="Polypodiopsida" family="Thelypteridaceae" genus="Pneumatopteris" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C618B78CBFDCEFA2C" box="[416,617,1410,1431]" italics="true" pageId="87" pageNumber="77">Pneumatopteris sensu</emphasis>
|
||
Holttum. In
|
||
</taxonomicName>
|
||
our phylogenetic analyses,
|
||
<taxonomicName id="4C1E4D63FFBE9B1C63CD78C8FBF1FA2D" box="[998,1110,1409,1430]" class="Polypodiopsida" family="Thelypteridaceae" genus="Reholttumia" kingdom="Plantae" order="Polypodiales" pageId="87" pageNumber="77" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96AEAF2FFBE9B1C63CD78C8FBF1FA2D" box="[998,1110,1409,1430]" italics="true" pageId="87" pageNumber="77">Reholttumia</emphasis>
|
||
</taxonomicName>
|
||
is represented by 25 accessions, including 13 of 30 currently recognized species, and several undescribed species (Fawcett et al. in press).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |