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<document id="867FA32941F0A887E8DDB46C5E497FC4" ID-DOI="10.5070/P940454153" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913559" ID-ZooBank="58312615-0833-432E-BF5D-3DFFBF361AAA" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712169471054" checkinUser="felipe" docAuthor="Burrow, Carole Jan, Murphy, Michael &amp; Turner, Susan" docDate="2023" docId="B21CD55BFFC9FFF7584C8C56FEFA21CB" docLanguage="en" docName="PaleoBios.40.4.1-32.pdf" docOrigin="PaleoBios 40 (4)" docSource="http://dx.doi.org/10.5070/p940454153" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.3:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="3" docTitle="Lanarkia TRAQUAIR 1898" docType="treatment" docVersion="3" lastPageNumber="22" masterDocId="4E25AD23FFDEFFE0596C8867FF9B270C" masterDocTitle="Late Silurian to earliest Devonian vertebrate biostratigraphy of the Birch Creek II section, Roberts Mountains, Nevada, U. S. A." masterLastPageNumber="32" masterPageNumber="1" pageNumber="22" updateTime="1727476634305" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="AFF9EE8BC976EF35B190225B4568A96A">Late Silurian to earliest Devonian vertebrate biostratigraphy of the Birch Creek II section, Roberts Mountains, Nevada, U. S. A.</mods:title>
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<paragraph id="3A0A644DFFC9FFF7584C8C56FDEB2341" blockId="23.[259,653,1040,1101]" box="[288,624,1073,1101]" pageId="23" pageNumber="22">
<heading id="6142D321FFC9FFF7584C8C56FDEB2341" allCaps="true" box="[288,624,1073,1101]" centered="true" fontSize="10" level="2" pageId="23" pageNumber="22" reason="2">
<taxonomicName id="FDB51FCEFFC9FFF7584C8C56FDEB2341" authority="TRAQUAIR, 1898" authorityName="TRAQUAIR" authorityYear="1898" box="[288,624,1073,1101]" family="Lanarkiidae" genus="Lanarkia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Thelodontiformes" pageId="23" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="08C1B85FFFC9FFF7584C8C56FE382341" box="[288,419,1073,1101]" italics="true" pageId="23" pageNumber="22">LANARKIA</emphasis>
<bibRefCitation id="5E2419BCFFC9FFF758C58C54FDEB2341" author="Traquair, R. H." box="[425,624,1075,1101]" pageId="23" pageNumber="22" pagination="72 - 76" refId="ref25754" refString="Traquair, R. H. 1898. [Report on fossil fishes]. Summary of Progress, Geological Survey of Great Britain, 1897, p. 72 - 76." type="journal article" year="1898">TRAQUAIR, 1898</bibRefCitation>
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<emphasis id="08C1B85FFFC9FFF759FC8C1EFEA7239F" bold="true" box="[144,316,1145,1171]" pageId="23" pageNumber="22">
<typeStatus id="E50EDAEFFFC9FFF759FC8C1EFF48239F" box="[144,211,1145,1171]" pageId="23" pageNumber="22">Type</typeStatus>
species
</emphasis>
<taxonomicName id="FDB51FCEFFC9FFF758358C1FFD71239F" authority="Traquair, 1898" authorityName="Traquair" authorityYear="1898" box="[345,746,1144,1172]" family="Lanarkiidae" genus="Lanarkia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="22" phylum="Chordata" rank="species" species="horrida">
<emphasis id="08C1B85FFFC9FFF758358C1FFDB72398" box="[345,556,1144,1172]" italics="true" pageId="23" pageNumber="22">Lanarkia horrida</emphasis>
<bibRefCitation id="5E2419BCFFC9FFF75B5F8C1EFD71239F" author="Traquair, R. H." box="[563,746,1145,1171]" pageId="23" pageNumber="22" pagination="72 - 76" refId="ref25754" refString="Traquair, R. H. 1898. [Report on fossil fishes]. Summary of Progress, Geological Survey of Great Britain, 1897, p. 72 - 76." type="journal article" year="1898">Traquair, 1898</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="72AF37C6FFC9FFF7581F8CA7FD8623D6" box="[371,541,1214,1242]" pageId="23" pageNumber="22" type="taxon_list">
<paragraph id="3A0A644DFFC9FFF7581F8CA7FD8623D6" blockId="23.[239,673,1214,1277]" box="[371,541,1214,1242]" pageId="23" pageNumber="22">
?
<taxonomicName id="FDB51FCEFFC9FFF758138CA7FD8623D6" box="[383,541,1214,1242]" family="Lanarkiidae" genus="Lanarkia" kingdom="Animalia" order="Thelodontiformes" pageId="23" pageNumber="22" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="08C1B85FFFC9FFF758138CA7FE6F23D6" box="[383,500,1216,1242]" italics="true" pageId="23" pageNumber="22">LANARKIA</emphasis>
SP.
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<subSubSection id="72AF37C6FFC9FFF759838C84FD3A23F1" box="[239,673,1251,1277]" pageId="23" pageNumber="22" type="description">
<paragraph id="3A0A644DFFC9FFF759838C84FD3A23F1" blockId="23.[239,673,1214,1277]" box="[239,673,1251,1277]" pageId="23" pageNumber="22">
(
<figureCitation id="A28E78C8FFC9FFF7599A8C84FEC923F1" box="[246,338,1251,1277]" captionStart="Figure 7" captionStartId="22.[112,191,1658,1682]" captionTargetBox="[112,1520,192,1638]" captionTargetId="figure-254@22.[112,1520,192,1638]" captionTargetPageId="22" captionText="Figure 7. Thelodont scales from the uppermost Silurian of the Birch Creek II section, all level 430.5' except B, C, G, K, L, P, Q. A. Trimerolepis sp. cf. T. tricava. UCR 10746-43 in crown view. B. Trimerolepis sp. AMF97876 (Parkes 1995, fig. 22.19, 20) from level 503, base view. C.?Lanarkia sp. UCR 944-20 (level 527.25) in lateral view. D.?Turinia sp. UCR10746-19, showing taphonomic effects and ultrasculpture, in crown view. E-J. Nikolivia spp. E. UCR 10746-13 with cell pattern on surface in crown view; F, UCR 10746-20 in crown view; G, AMF97875 (Parkes 1995, fig. 22.17, 18) from level 402' with possible bite marks, in crown view; H, UCR 10746-27 in ventral basal view; I, UCR 10746-25 showing taphonomic effects in crown view; J, Nikolivia sp. cf. N. auriculata, UCR 10746-18 in crown view. K, L. Apalolepis sp. AMF97872 (level 503) (Parkes 2005, fig. 22.11, 12), crown and basal views. M-O. Talivalia sp. cf. T. elongata. M, UCR 10746-17 in crown view; N, UCR 10746-37 in crown view; O, UCR 10746-21 in lateral view. P-S. Barlowodus sp. cf. B. tridens. P,Q, AMF97874 (level 875) in crown and basal views (Parkes 2005, fig. 22.15, 16). R, S, UCR 944-21 (level 527.25') in laterocrown view, closeup in S showing striated ultrasculpture. Scale bars=0.2 mm; anterior to left or top, otherwise arrow points to anterior. See Table 1 for meterage." figureDoi="http://doi.org/10.5281/zenodo.10913573" httpUri="https://zenodo.org/record/10913573/files/figure.png" pageId="23" pageNumber="22">FIG. 7C</figureCitation>
;
<tableCitation id="773751F6FFC9FFF758318C84FE2423F1" box="[349,447,1251,1277]" captionStart="Table 1" captionStartId="5.[112,179,1869,1893]" captionTargetBox="[120,1534,204,1847]" captionTargetId="graphics-72@5.[112,1519,193,1849]" captionTargetPageId="5" captionText="Table 1. Distribution of vertebrate and agnathan remains in the Birch Creek Section BCII, northern Roberts Mts. (Přidolí- Lochkovian; bold line indicates probable S/D boundary). Vertebrates are to the left above the upper horizontal line and agnathans are to the right. RP indicates Ross Parke samples. Number of elements: x 1-10, xx 11-50, xxx 51-100, xxxx&gt;100." httpUri="http://table.plazi.org/id/6ECA34C5FFDBFFE5591C8F2AFA2520A9" pageId="23" pageNumber="22" tableUuid="6ECA34C5FFDBFFE5591C8F2AFA2520A9">TABLE 1</tableCitation>
; SUPPL. 1, FIG. 11)
</paragraph>
</subSubSection>
<subSubSection id="72AF37C6FFC9FFF759FC8D4EFEFA21CB" pageId="23" pageNumber="22" type="discussion">
<paragraph id="3A0A644DFFC9FFF759FC8D4EFEFA21CB" blockId="23.[112,801,1321,1735]" pageId="23" pageNumber="22">
<emphasis id="08C1B85FFFC9FFF759FC8D4EFDFD224F" bold="true" box="[144,614,1321,1347]" pageId="23" pageNumber="22">Referred specimens and Remarks</emphasis>
—At least two possible scales from BC II section: one from level 458.67 (
<quantity id="FD4DC9A8FFC9FFF759158D17FF7D2286" box="[121,230,1392,1418]" metricMagnitude="2" metricUnit="m" metricValue="1.398" pageId="23" pageNumber="22" unit="m" value="139.8">139.8 m</quantity>
), one from level 527.25 (
<quantity id="FD4DC9A8FFC9FFF75B428D17FD072286" box="[558,668,1392,1418]" metricMagnitude="2" metricUnit="m" metricValue="1.607" pageId="23" pageNumber="22" unit="m" value="160.7">160.7 m</quantity>
) (
<figureCitation id="A28E78C8FFC9FFF75BDA8D17FC882286" box="[694,787,1392,1418]" captionStart="Figure 7" captionStartId="22.[112,191,1658,1682]" captionTargetBox="[112,1520,192,1638]" captionTargetId="figure-254@22.[112,1520,192,1638]" captionTargetPageId="22" captionText="Figure 7. Thelodont scales from the uppermost Silurian of the Birch Creek II section, all level 430.5' except B, C, G, K, L, P, Q. A. Trimerolepis sp. cf. T. tricava. UCR 10746-43 in crown view. B. Trimerolepis sp. AMF97876 (Parkes 1995, fig. 22.19, 20) from level 503, base view. C.?Lanarkia sp. UCR 944-20 (level 527.25) in lateral view. D.?Turinia sp. UCR10746-19, showing taphonomic effects and ultrasculpture, in crown view. E-J. Nikolivia spp. E. UCR 10746-13 with cell pattern on surface in crown view; F, UCR 10746-20 in crown view; G, AMF97875 (Parkes 1995, fig. 22.17, 18) from level 402' with possible bite marks, in crown view; H, UCR 10746-27 in ventral basal view; I, UCR 10746-25 showing taphonomic effects in crown view; J, Nikolivia sp. cf. N. auriculata, UCR 10746-18 in crown view. K, L. Apalolepis sp. AMF97872 (level 503) (Parkes 2005, fig. 22.11, 12), crown and basal views. M-O. Talivalia sp. cf. T. elongata. M, UCR 10746-17 in crown view; N, UCR 10746-37 in crown view; O, UCR 10746-21 in lateral view. P-S. Barlowodus sp. cf. B. tridens. P,Q, AMF97874 (level 875) in crown and basal views (Parkes 2005, fig. 22.15, 16). R, S, UCR 944-21 (level 527.25') in laterocrown view, closeup in S showing striated ultrasculpture. Scale bars=0.2 mm; anterior to left or top, otherwise arrow points to anterior. See Table 1 for meterage." figureDoi="http://doi.org/10.5281/zenodo.10913573" httpUri="https://zenodo.org/record/10913573/files/figure.png" pageId="23" pageNumber="22">Fig. 7C</figureCitation>
).
<taxonomicName id="FDB51FCEFFC9FFF7591C8DF6FEA422A1" box="[112,319,1425,1453]" pageId="23" pageNumber="22">
<emphasis id="08C1B85FFFC9FFF7591C8DF6FF4422A1" box="[112,223,1425,1453]" italics="true" pageId="23" pageNumber="22">Lanarkia</emphasis>
species
</taxonomicName>
are known mainly from earlier Silurian rocks in
<collectingCountry id="42A224DDFFC9FFF759B28DD1FED022DC" box="[222,331,1462,1488]" name="United Kingdom" pageId="23" pageNumber="22">Scotland</collectingCountry>
, mainland Europe, and arctic
<collectingCountry id="42A224DDFFC9FFF75BA48DD1FC8022DC" box="[712,795,1462,1488]" name="Russia" pageId="23" pageNumber="22">Russia</collectingCountry>
, as well as in arctic
<collectingCountry id="42A224DDFFC9FFF7583A8DBEFE2922FF" box="[342,434,1497,1523]" name="Canada" pageId="23" pageNumber="22">Canada</collectingCountry>
(
<bibRefCitation id="5E2419BCFFC9FFF758AF8DBDFD0E22FF" author="Marss, T. &amp; S. Turner &amp; V. Karatajute-Talimaa" box="[451,661,1497,1524]" pageId="23" pageNumber="22" refId="ref24679" refString="Marss, T., S. Turner, and V. Karatajute-Talimaa. 2007. Agnatha II-Thelodonti. Volume 1 B. 143 pp. Verlag Dr Friedrich Pfeil, Munich." type="book" year="2007">Märss et al. 2007</bibRefCitation>
). The scale from 527.25 is most like those called
<taxonomicName id="FDB51FCEFFC9FFF75B518D9CFCBB211B" box="[573,800,1531,1559]" family="Thelodontidae" genus="Thelodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="22" phylum="Chordata" rank="species" species="traquairi">
<emphasis id="08C1B85FFFC9FFF75B518D9CFCBB211B" box="[573,800,1531,1559]" italics="true" pageId="23" pageNumber="22">Thelodus traquairi</emphasis>
</taxonomicName>
by
<bibRefCitation id="5E2419BCFFC9FFF759FA8E47FEA12136" author="Gross, W." box="[150,314,1568,1594]" pageId="23" pageNumber="22" pagination="1 - 67" refId="ref23396" refString="Gross, W. 1967. Uber Thelodontier-Schuppen. Palaeontographica Abt. A 127: 1 - 67." type="journal article" year="1967">Gross (1967)</bibRefCitation>
, originally “
<taxonomicName id="FDB51FCEFFC9FFF758BD8E79FD3A2136" box="[465,673,1566,1594]" family="Lanarkiidae" genus="Lanarkia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="22" phylum="Chordata" rank="species" species="costata">
<emphasis id="08C1B85FFFC9FFF758BD8E79FD3A2136" box="[465,673,1566,1594]" italics="true" pageId="23" pageNumber="22">Lanarkia costata</emphasis>
</taxonomicName>
” of
<bibRefCitation id="5E2419BCFFC9FFF75BB88E47FF2A2151" author="Gross, W." pageId="23" pageNumber="22" pagination="91 - 161" refId="ref23372" refString="Gross, W. 1947. Die Agnathen und Acanthodier des obersilurischen Beyrichienkalks. Palaeontographica Abt. A 96: 91 - 161." type="journal article" year="1947">Gross, 1947</bibRefCitation>
, from the north German Beyrichienkalk (Bey. 1, early Přidolí) and ones from
<collectingCountry id="42A224DDFFC9FFF758B48E01FDB4218C" box="[472,559,1638,1664]" name="Poland" pageId="23" pageNumber="22">Poland</collectingCountry>
(late Přidolí;
<bibRefCitation id="5E2419BCFFC9FFF75BB68E01FF2B21AF" author="Gross, W." pageId="23" pageNumber="22" pagination="91 - 161" refId="ref23372" refString="Gross, W. 1947. Die Agnathen und Acanthodier des obersilurischen Beyrichienkalks. Palaeontographica Abt. A 96: 91 - 161." type="journal article" year="1947">Gross 1947</bibRefCitation>
,
<bibRefCitation id="5E2419BCFFC9FFF759D68EEDFED221AF" author="Marss, T." box="[186,329,1673,1700]" pageId="23" pageNumber="22" pagination="17 - 41" refId="ref24364" refString="Marss, T. 1997. Vertebrates of the Pridoli and Silurian-Devonian boundary beds in Europe. Modern Geology 21: 17 - 41." type="journal article" year="1997">Märss 1997</bibRefCitation>
), which have been referred to
<taxonomicName id="FDB51FCEFFC9FFF75BDE8EEFFEC721CB" authority="(Marss et al. 2007)" baseAuthorityName="Marss" baseAuthorityYear="2007" family="Lanarkiidae" genus="Lanarkia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="23" pageNumber="22" phylum="Chordata" rank="genus">
<emphasis id="08C1B85FFFC9FFF75BDE8EEFFCBB21A8" box="[690,800,1672,1700]" italics="true" pageId="23" pageNumber="22">Lanarkia</emphasis>
(
<bibRefCitation id="5E2419BCFFC9FFF759178ECAFECB21CB" author="Marss, T. &amp; S. Turner &amp; V. Karatajute-Talimaa" box="[123,336,1708,1735]" pageId="23" pageNumber="22" refId="ref24679" refString="Marss, T., S. Turner, and V. Karatajute-Talimaa. 2007. Agnatha II-Thelodonti. Volume 1 B. 143 pp. Verlag Dr Friedrich Pfeil, Munich." type="book" year="2007">Märss et al. 2007</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
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