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<document id="FB188E7240C3109360BE8581E9F39CA6" ID-DOI="10.11646/zootaxa.5375.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10200036" ID-ZooBank="27D02F09-01B7-457A-8A99-D8644B7B6ADE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700728652862" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2023" docId="03AE8A2DFFA2CF62FF47EA01FD31FE64" docLanguage="en" docName="zootaxa.5375.3.1.pdf" docOrigin="Zootaxa 5375 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5375.3.1/52332" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penia alishanensis Arimoto 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="307" masterDocId="FF97F255FFA6CF64FFD0EB1BFFFFFFED" masterDocTitle="Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="335" masterPageNumber="301" pageNumber="305" updateTime="1700729251563" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="28B989676966768C984607B1FD64C7B5">Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan</mods:title>
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<mods:namePart id="6A3E70BAE1564D2FE53C06392504887E">Arimoto, Kôichi</mods:namePart>
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<mods:date id="356C17B95C71E819095FDCB890D483F9">2023</mods:date>
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<treatment id="03AE8A2DFFA2CF62FF47EA01FD31FE64" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA2CF62FF47EA01FD31FE64" httpUri="http://treatment.plazi.org/id/03AE8A2DFFA2CF62FF47EA01FD31FE64" lastPageId="6" lastPageNumber="307" pageId="4" pageNumber="305">
<subSubSection id="C31D68B0FFA2CF60FF47EA01FDA9FED8" box="[151,598,282,309]" pageId="4" pageNumber="305" type="nomenclature">
<paragraph id="8BB83B3BFFA2CF60FF47EA01FDA9FED8" blockId="4.[151,598,282,345]" box="[151,598,282,309]" pageId="4" pageNumber="305">
<heading id="D0F08C57FFA2CF60FF47EA01FDA9FED8" bold="true" box="[151,598,282,309]" fontSize="11" level="1" pageId="4" pageNumber="305" reason="1">
<emphasis id="B973E729FFA2CF60FF47EA01FDA9FED8" bold="true" box="[151,598,282,309]" pageId="4" pageNumber="305">
<taxonomicName id="4C0740B8FFA2CF60FF47EA01FE13FED9" authority="Arimoto" authorityName="Arimoto" authorityYear="2023" box="[151,492,282,309]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="alishanensis" status="sp. nov.">
<emphasis id="B973E729FFA2CF60FF47EA01FE84FED8" bold="true" box="[151,379,282,309]" italics="true" pageId="4" pageNumber="305">Penia alishanensis</emphasis>
Arimoto
</taxonomicName>
,
<taxonomicNameLabel id="A2405A52FFA2CF60FE29EA00FDA9FED8" box="[505,598,283,309]" pageId="4" pageNumber="305" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA2CF60FF47EA25FED0FEB4" box="[151,303,318,345]" pageId="4" pageNumber="305" type="description">
<paragraph id="8BB83B3BFFA2CF60FF47EA25FED0FEB4" blockId="4.[151,598,282,345]" box="[151,303,318,345]" pageId="4" pageNumber="305">
(
<figureCitation id="133C27BEFFA2CF60FF4FEA25FEF3FEB4" box="[159,268,318,345]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Figures 2</figureCitation>
,
<figureCitation id="133C27BEFFA2CF60FEC8EA24FED9FEB4" box="[280,294,319,345]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="4" pageNumber="305">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA2CF60FF47EA9EFC34FE73" box="[151,971,388,415]" pageId="4" pageNumber="305" type="etymology">
<paragraph id="8BB83B3BFFA2CF60FF47EA9EFC34FE73" blockId="4.[151,1437,388,2023]" box="[151,971,388,415]" pageId="4" pageNumber="305">
<emphasis id="B973E729FFA2CF60FF47EA9EFEE1FE72" bold="true" box="[151,286,389,415]" pageId="4" pageNumber="305">Etymology.</emphasis>
Specific epithet derived from Mt. Ali
<collectingRegion id="49C3F5D9FFA2CF60FD1BEA9FFCF9FE73" box="[715,774,388,414]" country="Myanmar" name="Shan" pageId="4" pageNumber="305">Shan</collectingRegion>
, the
<typeStatus id="54BC8599FFA2CF60FCEAEA9EFC95FE72" box="[826,874,389,415]" pageId="4" pageNumber="305">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA2CF60FF17EABCFC9EFC46" pageId="4" pageNumber="305" type="materials_examined">
<paragraph id="8BB83B3BFFA2CF60FF17EABCFDE8FE09" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
<emphasis id="B973E729FFA2CF60FF17EABCFE88FE2C" bold="true" box="[199,375,423,449]" pageId="4" pageNumber="305">Type material.</emphasis>
<materialsCitation id="3B6F3166FFA2CF60FE50EABCFDE8FE09" collectingDate="1977-08-18" collectionCode="NMNST" collectorName="Y. Watanabe" country="Taiwan" county="Male" location="Mt. Ali Shan" municipality="Alishan Township" pageId="4" pageNumber="305" specimenCode="PAA01" specimenCount="1" stateProvince="Chiayi" typeStatus="holotype">
<emphasis id="B973E729FFA2CF60FE50EABCFE0BFE2C" bold="true" box="[384,500,423,449]" pageId="4" pageNumber="305">
<typeStatus id="54BC8599FFA2CF60FE50EABCFE10FE2C" box="[384,495,423,449]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCounty id="62D943B7FFA2CF60FE2EEABCFDC5FE2C" box="[510,570,423,449]" pageId="4" pageNumber="305">Male</collectingCounty>
,
<collectingCountry id="F3107BABFFA2CF60FD97EABCFD63FE2C" box="[583,668,423,449]" name="Taiwan" pageId="4" pageNumber="305">Taiwan</collectingCountry>
,
<collectingRegion id="49C3F5D9FFA2CF60FD79EABCFCAEFE2C" box="[681,849,423,449]" country="Taiwan" name="Chiayi" pageId="4" pageNumber="305">Chiayi County</collectingRegion>
,
<collectingMunicipality id="6BDCA141FFA2CF60FC8CEABCFBD2FE2C" box="[860,1069,423,449]" pageId="4" pageNumber="305">Alishan Township</collectingMunicipality>
,
<location id="8ED86DE0FFA2CF60FBEAEAB3FB2DFE2C" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA2CF62FF47EA01FD31FE64:8ED86DE0FFA2CF60FBEAEAB3FB2DFE2C" box="[1082,1234,423,450]" country="Taiwan" county="Male" municipality="Alishan Township" name="Mt. Ali Shan" pageId="4" pageNumber="305" stateProvince="Chiayi">Mt. Ali Shan</location>
,
<date id="FFB91DFBFFA2CF60FB0FEABCFA87FE2C" box="[1247,1400,423,450]" pageId="4" pageNumber="305" value="1977-08-18">
<collectingDate id="EFFDE413FFA2CF60FB0FEABCFA87FE2C" box="[1247,1400,423,450]" pageId="4" pageNumber="305" value="1977-08-18">18 VIII 1977</collectingDate>
</date>
,
<collectorName id="26F25EEDFFA2CF60FA55EAB3FEFAFE09" pageId="4" pageNumber="305">Y. Watanabe</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA2CF60FE92EAD1FE56FE09" box="[322,425,458,484]" pageId="4" pageNumber="305">NMNST</collectionCode>
;
<specimenCode id="DBA19340FFA2CF60FE65EAD1FDF2FE09" box="[437,525,458,484]" collectionCode="PAA" pageId="4" pageNumber="305">PAA01</specimenCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8BB83B3BFFA2CF60FF17EAF6FC1FFCF2" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
<emphasis id="B973E729FFA2CF60FF17EAF6FE71FDEA" bold="true" box="[199,398,493,519]" pageId="4" pageNumber="305">Male. Diagnosis.</emphasis>
This species is characterized by the following features: eyes 0.45 x longer than interocular distance in dorsal view; antennae surpassing elytral half by antennomere XI; antennomeres
<collectionCode id="ED16A3FEFFA2CF60FB5EE90AFB55FDC6" box="[1166,1194,529,555]" pageId="4" pageNumber="305">III</collectionCode>
distinctly longer than
<collectionCode id="ED16A3FEFFA2CF60FF47E92FFF52FDA3" box="[151,173,564,590]" pageId="4" pageNumber="305">II</collectionCode>
;
<collectionCode id="ED16A3FEFFA2CF60FF69E92FFF29FDA3" box="[185,214,564,590]" pageId="4" pageNumber="305">IV</collectionCode>
1.4 x longer than
<collectionCode id="ED16A3FEFFA2CF60FE76E92FFE3AFDA3" box="[422,453,564,590]" pageId="4" pageNumber="305">III</collectionCode>
, 0.9 x longer than
<collectionCode id="ED16A3FEFFA2CF60FD49E92FFD54FDA3" box="[665,683,564,590]" pageId="4" pageNumber="305">II</collectionCode>
<collectionCode id="ED16A3FEFFA2CF60FD6AE92FFD29FDA3" box="[698,726,564,590]" pageId="4" pageNumber="305">III</collectionCode>
combined; apical maxillary palpomere 2.3 x longer than wide, shorter than eye maximum length; hypomeron with distinct mesial projection; anterior angle of hypomeron biangular, with outer angle right angle, with mesial angle rounded; scutellar shield 1.2 x longer than wide; mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view; posterior edge of mesosternal process 0.1 x wider than total length of mesosternum; elytra 4.5 x longer than wide; abdominal ventrite
<collectionCode id="ED16A3FEFFA2CF60FB68E9A4FB33FD34" box="[1208,1228,703,729]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="305" type="Museum">V</collectionCode>
curved triangular, rounded apically; phallobase 0.7 x longer than wide; apex of parameres beyond preapical expansions elongated fan-shaped; apex length 2.7 x width of parameres at expansions in ventral side.
</paragraph>
<paragraph id="8BB83B3BFFA2CF60FF17E833FC9EFC46" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
This species is similar to
<taxonomicName id="4C0740B8FFA2CF60FE37E833FD93FCAC" authorityName="Schimmel" authorityYear="1996" box="[487,620,808,834]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B973E729FFA2CF60FE37E833FE07FCAC" box="[487,504,808,833]" italics="true" pageId="4" pageNumber="305">P</emphasis>
.
<emphasis id="B973E729FFA2CF60FDD7E833FD93FCAC" box="[519,620,808,833]" italics="true" pageId="4" pageNumber="305">smetanai</emphasis>
</taxonomicName>
in features of the body shape, the eyes, the basal antennomeres, and the mesosternal process between mesocoxae. It is distinguished by the length-to-width ratios of the apical maxillary palpomere and scutellar shield, and the shapes of the hypomeron, abdominal ventrite
<collectionCode id="ED16A3FEFFA2CF60FB89E875FB92FC65" box="[1113,1133,878,904]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="305" type="Museum">V</collectionCode>
and apex of the parameres beyond the preapical expansions (see diagnosis in
<taxonomicName id="4C0740B8FFA2CF60FD1CE88AFCAEFC47" authorityName="Schimmel" authorityYear="1996" box="[716,849,913,939]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B973E729FFA2CF60FD1CE88AFD22FC47" box="[716,733,913,938]" italics="true" pageId="4" pageNumber="305">P</emphasis>
.
<emphasis id="B973E729FFA2CF60FD3CE88AFCAEFC47" box="[748,849,913,938]" italics="true" pageId="4" pageNumber="305">smetanai</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA2CF62FF17E8A8FE5DFE88" lastPageId="6" lastPageNumber="307" pageId="4" pageNumber="305" type="description">
<paragraph id="8BB83B3BFFA2CF60FF17E8A8FE56FC1D" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
<emphasis id="B973E729FFA2CF60FF17E8A8FE81FC20" bold="true" box="[199,382,947,973]" pageId="4" pageNumber="305">Measurements.</emphasis>
BL: around 7.3 (estimate value), BW: 2.31, MAE: 1.09, MBE: 0.58, OI: 188, PAW: 1.01, EL: 5.19, EW: 1.16, EI: 449.
</paragraph>
<paragraph id="8BB83B3BFFA2CF60FF17E8E2FCA7FB72" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
<emphasis id="B973E729FFA2CF60FF17E8E2FEA6FBFE" bold="true" box="[199,345,1017,1043]" pageId="4" pageNumber="305">Description.</emphasis>
Body elongated, widest behind elytral midlength (
<figureCitation id="133C27BEFFA2CF60FC5CE8E2FC19FBFE" box="[908,998,1017,1043]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2A</figureCitation>
); surface generally smooth; interspaces between punctures distinctly larger than fine puncture diameter (
<figureCitation id="133C27BEFFA2CF60FCA8EF07FC0CFBDB" box="[888,1011,1052,1078]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2A, B</figureCitation>
) but on elytra intermixed with larger and smaller punctures. Color. Body, antennae and legs orange. Head, median area of pronotum and elytral anterior half darker. External edges of mandible and scutellum, and anterior edge of elytra black. Metasternum and abdomen partly tinged with black. Body covered with long yellow setae.
</paragraph>
<paragraph id="8BB83B3BFFA2CF60FF17EFB3FBA5FA5B" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
Head. Frons slightly flatted medially (
<figureCitation id="133C27BEFFA2CF60FD5CEFB3FCEEFB2F" box="[652,785,1192,1218]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2C, D</figureCitation>
); frontal carina not complete (
<figureCitation id="133C27BEFFA2CF60FBAFEFB3FB22FB2F" box="[1151,1245,1192,1218]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2C</figureCitation>
); frontal margin hexagonal, almost straight apically in dorsal view (
<figureCitation id="133C27BEFFA2CF60FD19EFD0FCDDFB08" box="[713,802,1227,1253]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2D</figureCitation>
); frontoclypeal region protruding beyond base of labrum. Eyes protuberant, 0.45 x longer than interocular distance in dorsal view (
<figureCitation id="133C27BEFFA2CF60FC3CEFF5FBB5FAE5" box="[1004,1098,1262,1288]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2D</figureCitation>
). Antennae surpassing elytral half by antennomere XI; antennomeres longer than wide; II globular, shortest, 1.35 x longer than wide; III weakly serrated, 2.3 x longer than wide, 1.8 x longer than II; IV weakly serrated, 3.1 x longer than wide, 1.4 x longer than III, 0.9 x longer than IIIII combined (
<figureCitation id="133C27BEFFA2CF60FD96EE4DFD61FA9C" box="[582,670,1366,1393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2E</figureCitation>
); VXI filiform; V 3.35 x longer than wide, 1.1 x longer than IV; XI 8.85 x longer than wide, 1.1 x longer than X. Mandible bidentate. Apical maxillary palpomere semicircular (
<figureCitation id="133C27BEFFA2CF60FABFEE62FF47FA5A" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2F</figureCitation>
), 2.3 x longer than wide, shorter than eye maximum length; anterior edge rounded.
</paragraph>
<paragraph id="8BB83B3BFFA2CF60FF17EEA4FB34F80A" blockId="4.[151,1437,388,2023]" pageId="4" pageNumber="305">
Prothorax with back half broken and lost (
<figureCitation id="133C27BEFFA2CF60FD7BEEA4FCD4FA34" box="[683,811,1471,1497]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2A, G</figureCitation>
). Pronotum roundly widening anteriorly, with punctate lateral ridge; anterior edge concave; anterior angles nearly right angle. Hypomeron with distinct mesial projection (
<figureCitation id="133C27BEFFA2CF60FF4FED1EFF02F9F2" box="[159,253,1541,1567]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2H</figureCitation>
: arrow); anterior angle biangular, with outer angle right angle, with mesial angle rounded; mesial and posterior margins with impunctate ridge. Prosternum incurved ventrally in lateral view; anterior lobe distinctly protruding beyond prosternal ventral line in lateral view (
<figureCitation id="133C27BEFFA2CF60FCF3ED50FC80F988" box="[803,895,1611,1637]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2G</figureCitation>
); anterior edge broadly rounded in ventral view (
<figureCitation id="133C27BEFFA2CF60FF4FED75FF07F965" box="[159,248,1646,1672]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2B</figureCitation>
). Pronotosternal sutures not grooved, almost straight in ventral view (
<figureCitation id="133C27BEFFA2CF60FBD9ED75FB9EF965" box="[1033,1121,1646,1672]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2B</figureCitation>
), slightly opened anteriorly. Scutellar shield tongue-shaped (
<figureCitation id="133C27BEFFA2CF60FDD6ED8AFDA9F946" box="[518,598,1681,1707]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2I</figureCitation>
), 1.2 x longer than wide, widest anteriorly, weakly constricted anteriorly and then narrowed posterad, flat, inclined anterior-downwards, not visible in lateral view; anterior edge slightly broadly rounded; posterior edge rounded. Mesosternum: borders of mesosternal cavity straight anteriorly and then moderately curved in lateral view (
<figureCitation id="133C27BEFFA2CF60FDF6EDE1FD7EF8F9" box="[550,641,1786,1812]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2G</figureCitation>
); mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view (
<figureCitation id="133C27BEFFA2CF60FE4EEC06FE06F8DA" box="[414,505,1821,1847]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2G</figureCitation>
); posterior edge 0.1 x wider than total length of mesosternum, weakly emarginate (
<figureCitation id="133C27BEFFA2CF60FF4FEC5BFF10F8B7" box="[159,239,1856,1882]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2J</figureCitation>
). Mesepisternum reaching mesocoxal cavity (
<figureCitation id="133C27BEFFA2CF60FD36EC5BFCCAF8B7" box="[742,821,1856,1882]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2J</figureCitation>
). Metasternum sulcate medially and ahead of metacoxal cavities (
<figureCitation id="133C27BEFFA2CF60FF2CEC79FEAAF890" box="[252,341,1890,1917]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2B</figureCitation>
). Metacoxal plate narrowed toward outer side, becoming like a parallel-sided bar at its outer 1/
<quantity id="4CFF96DEFFA2CF60FAAEEC78FF52F872" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="305" unit="in" value="10.0">10 in</quantity>
ventral view (
<figureCitation id="133C27BEFFA2CF60FE9DEC9EFE5AF84D" box="[333,421,1925,1952]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2B</figureCitation>
). Elytron convex but flat in median area, but with outer margin narrowly depressed, widest behind midlength, 4.5 x longer than wide; apex rounded. Hind wings fully developed. Tibiae with paired spurs; relative tarsomere lengths: IV&lt;III II&lt;V&lt;I; tarsomeres III and IV with lobe ventrally (
<figureCitation id="133C27BEFFA2CF60FBB4ECD7FB41F80A" box="[1124,1214,1996,2023]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="4" pageNumber="305">Fig. 2K</figureCitation>
).
</paragraph>
<caption id="DF786BB3FFA3CF61FF47ED6DFC08F8F3" ID-DOI="http://doi.org/10.5281/zenodo.10200040" ID-Zenodo-Dep="10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="5" pageNumber="306" startId="5.[151,250,1654,1678]" targetBox="[151,1436,181,1630]" targetPageId="5" targetType="figure">
<paragraph id="8BB83B3BFFA3CF61FF47ED6DFC08F8F3" blockId="5.[151,1436,1654,1822]" pageId="5" pageNumber="306">
<emphasis id="B973E729FFA3CF61FF47ED6DFEEAF963" bold="true" box="[151,277,1654,1678]" pageId="5" pageNumber="306">FIGURE 2.</emphasis>
<taxonomicName id="4C0740B8FFA3CF61FECEED6CFE21F963" authorityName="Arimoto" authorityYear="2023" box="[286,478,1654,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="alishanensis">
<emphasis id="B973E729FFA3CF61FECEED6CFE21F963" box="[286,478,1654,1678]" italics="true" pageId="5" pageNumber="306">Penia alishanensis</emphasis>
</taxonomicName>
, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow: hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFA3CF62FF17EC57FE3EFEAC" blockId="5.[151,1437,1868,2039]" lastBlockId="6.[151,1437,151,393]" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
Abdomen. Ventrite V curved triangular, rounded apically (
<figureCitation id="133C27BEFFA3CF61FCA1EC57FC33F88A" box="[881,972,1868,1895]" captionStart="FIGURE 2" captionStartId="5.[151,250,1654,1678]" captionTargetBox="[151,1436,181,1630]" captionTargetId="figure-16@5.[151,1436,181,1630]" captionTargetPageId="5" captionText="FIGURE 2. Penia alishanensis, holotype, male (PAA01). A: habitus, dorsal side; B: habitus, ventral side; C: head, anterior view; D: head, dorsal view; E: basal antennomeres of right antenna; F: right maxillary palp, ventral side; G: head, prothorax and mesothorax, lateral view (arrow: the direction perpendicular to the mesosternal process between mesocoxae); H: right hypomeron (arrow:hypomeron with distinct mesial projection); I: scutellar shield; J: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (arrow of figure 2G); K: right mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200040" httpUri="https://zenodo.org/record/10200040/files/figure.png" pageId="5" pageNumber="306">Fig. 2B</figureCitation>
), 0.65 x longer than wide. Tergites and sternites VIIIX yellow. Tergite VIII 1.5 x longer than wide, trapezoidal, narrowed posterad; posterior margin weakly incurved (
<figureCitation id="133C27BEFFA3CF61FEB1EC8FFE44F842" box="[353,443,1940,1967]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="5" pageNumber="306">Fig. 3A</figureCitation>
). Sternite VIII posteriorly widely concave between two projections (
<figureCitation id="133C27BEFFA3CF61FB6DEC8FFAEAF842" box="[1213,1301,1940,1967]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="5" pageNumber="306">Fig. 3B</figureCitation>
). Tergite IX 1.35 x longer than wide; median notch 0.4 x total length of tergite IX (
<figureCitation id="133C27BEFFA3CF61FC44ECA3FC14F83E" box="[916,1003,1976,2003]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="5" pageNumber="306">Fig. 3C</figureCitation>
). Tergite X longer than wide, triangular apically (
<figureCitation id="133C27BEFFA3CF61FF2EECC7FEA9F81A" box="[254,342,2012,2039]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="5" pageNumber="306">Fig. 3C</figureCitation>
). Sternite IX 3.2 x longer than wide, constricted anterior 2/5 (
<figureCitation id="133C27BEFFA3CF61FC2EECC7FBA9F81A" box="[1022,1110,2012,2039]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="5" pageNumber="306">Fig. 3D</figureCitation>
), rounded apically. Aedeagus yellow (
<figureCitation id="133C27BEFFA0CF62FF22EB8CFE9BFF5C" box="[242,356,151,177]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="6" pageNumber="307">Fig. 3E, F</figureCitation>
). Phallobase 0.2 x total length of aedeagus, 0.7 x longer than wide. Median lobe exceeding apices of parameres by apical 1/20; basal struts 0.15 x total length of median lobe. Parameres slender, not fused ventrally (
<figureCitation id="133C27BEFFA0CF62FF4FEBC4FF08FF14" box="[159,247,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="6" pageNumber="307">Fig. 3F</figureCitation>
); preapical expansions protruding anterolaterad (
<figureCitation id="133C27BEFFA0CF62FCFAEBC4FC78FF14" box="[810,903,223,249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="6" pageNumber="307">Fig. 3G</figureCitation>
); apex beyond preapical expansions elongated fan-shaped (
<figureCitation id="133C27BEFFA0CF62FEF5EA18FE7DFEF0" box="[293,386,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1441,1465]" captionTargetBox="[151,1436,417,1415]" captionTargetId="figure-107@6.[151,1436,417,1418]" captionTargetPageId="6" captionText="FIGURE 3. Penia alishanensis, holotype, male (PAA01). A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal side; F: aedeagus, ventral side; G: apical part of right paramera, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200042" httpUri="https://zenodo.org/record/10200042/files/figure.png" pageId="6" pageNumber="307">Fig. 3G</figureCitation>
), with a seta dorsally, with three setae ventrally; apex length 2.7 x width of parameres at expansions in ventral side.
</paragraph>
<paragraph id="8BB83B3BFFA0CF62FF17EA50FE5DFE88" blockId="6.[151,1437,151,393]" box="[199,418,331,357]" pageId="6" pageNumber="307">
<emphasis id="B973E729FFA0CF62FF17EA50FEDAFE88" bold="true" box="[199,293,331,357]" pageId="6" pageNumber="307">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA0CF62FF17EA75FD31FE64" box="[199,718,366,393]" pageId="6" pageNumber="307" type="distribution">
<paragraph id="8BB83B3BFFA0CF62FF17EA75FD31FE64" blockId="6.[151,1437,151,393]" box="[199,718,366,393]" pageId="6" pageNumber="307">
<emphasis id="B973E729FFA0CF62FF17EA75FE9EFE65" bold="true" box="[199,353,366,392]" pageId="6" pageNumber="307">Distribution.</emphasis>
<collectingCountry id="F3107BABFFA0CF62FEB7EA74FE42FE64" box="[359,445,367,393]" name="Taiwan" pageId="6" pageNumber="307">Taiwan</collectingCountry>
:
<collectingRegion id="49C3F5D9FFA0CF62FE18EA74FD94FE64" box="[456,619,367,393]" country="Taiwan" name="Chiayi" pageId="6" pageNumber="307">Chiayi County</collectingRegion>
(
<figureCitation id="133C27BEFFA0CF62FDABEA74FD42FE64" box="[635,701,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,902,926]" captionTargetBox="[158,1429,188,872]" captionTargetId="figure-16@3.[151,1436,181,879]" captionTargetPageId="3" captionText="FIGURE 1. Map of Taiwan. A: location of Taiwan; B: the collection sites of the material used in this study." figureDoi="http://doi.org/10.5281/zenodo.10200038" httpUri="https://zenodo.org/record/10200038/files/figure.png" pageId="6" pageNumber="307">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>