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<document id="4F653F3F61EEFDB9313F75886068513E" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCAC7B435D0B83FE118B640420" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Ulandrena (Ulandrena) Warncke 1968" docType="treatment" docVersion="1" lastPageNumber="39" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="35" updateTime="1723562696088" updateUser="guilherme">
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<mods:title id="8C1B142DA47330DD34D0FEFDE5F8D4AE">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<treatment id="612B87FCAC7B435D0B83FE118B640420" LSID="urn:lsid:plazi:treatment:612B87FCAC7B435D0B83FE118B640420" httpUri="http://treatment.plazi.org/id/612B87FCAC7B435D0B83FE118B640420" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35">
<subSubSection id="A1986561AC7B43590B83FE118A0A07B5" box="[151,1135,474,501]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="E93D36EAAC7B43590B83FE118A0A07B5" blockId="34.[151,1135,474,501]" box="[151,1135,474,501]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B43590B83FE118A0A07B5" bold="true" box="[151,1135,474,501]" pageId="34" pageNumber="35">
3) Revision of the
<emphasis id="DBF6EAF8AC7B43590A6CFE108F8D07B4" bold="true" box="[376,488,475,501]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
-group (within subgenus
<taxonomicName id="2E824D69AC7B43590834FE118A0207B5" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[800,1127,474,501]" class="Insecta" family="Andrenidae" genus="Ulandrena" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC7B43590834FE118DC707B4" bold="true" box="[800,930,474,501]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
Warncke, 1968
</taxonomicName>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7B435D0B83FDE88B640420" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35" type="discussion">
<paragraph id="E93D36EAAC7B43590B83FDE88FED051C" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
The
<taxonomicName id="2E824D69AC7B43590BD3FDE88FF7047D" authorityName="Smith" authorityYear="1853" box="[199,402,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590BD3FDE88FF7047D" box="[199,402,547,573]" italics="true" pageId="34" pageNumber="35">Andrena concinna</emphasis>
</taxonomicName>
-group consists of a small number of very large-sized
<taxonomicName id="2E824D69AC7B435908CDFDE88A5D047C" baseAuthorityName="Popov" baseAuthorityYear="1960" box="[985,1080,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC7B435908CDFDE88A5D047C" box="[985,1080,547,573]" italics="true" pageId="34" pageNumber="35">Andrena</emphasis>
</taxonomicName>
within the subgenus
<taxonomicName id="2E824D69AC7B43590E09FDE88BF0047C" authorityName="Warncke" authorityYear="1968" box="[1309,1429,547,573]" pageId="34" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7B43590E09FDE88BF0047C" box="[1309,1429,547,573]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
</taxonomicName>
. These bees are instantly recognised within the subgenus by their huge body size, with specimens usually exceeding
<quantity id="2E7A9B0FAC7B43590B83FDA18E8F04C4" box="[151,234,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="mm" value="15.0">15 mm</quantity>
in length which makes them some of the largest
<taxonomicName id="2E824D69AC7B4359083BFDA08D8F04C4" box="[815,1002,619,645]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B4359083BFDA08DEB04C4" box="[815,910,619,645]" italics="true" pageId="34" pageNumber="35">Andrena</emphasis>
species
</taxonomicName>
globally. The
<taxonomicName id="2E824D69AC7B43590F81FDA08A9904C5" authorityName="Smith" authorityYear="1853" box="[1173,1276,619,644]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F81FDA08A9904C5" box="[1173,1276,619,644]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
-group can be recognised further because the hind tarsal claws of the females are bidentate, presenting an inner subapical tooth. This separates them from the other large-bodied
<taxonomicName id="2E824D69AC7B435909D9FD788D20048C" authorityName="Warncke" authorityYear="1968" box="[717,837,691,717]" pageId="34" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7B435909D9FD788D20048C" box="[717,837,691,717]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
</taxonomicName>
, specifically
<taxonomicName id="2E824D69AC7B435908F5FD788B72048C" authority="Popov, 1940" authorityName="Popov" authorityYear="1940" box="[993,1303,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="DBF6EAF8AC7B435908F5FD788A1E048D" box="[993,1147,691,717]" italics="true" pageId="34" pageNumber="35">A. armeniaca</emphasis>
Popov, 1940
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590E31FD788F5404B0" authority="Giraud, 1863" authorityName="Giraud" authorityYear="1863" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DBF6EAF8AC7B43590E31FD788BF8048C" box="[1317,1437,691,717]" italics="true" pageId="34" pageNumber="35">A. elegans</emphasis>
Giraud, 1863
</taxonomicName>
, and
<taxonomicName id="2E824D69AC7B43590A66FD1C8C8504B0" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" box="[370,736,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="DBF6EAF8AC7B43590A66FD1C8C4504B0" box="[370,544,727,753]" italics="true" pageId="34" pageNumber="35">A. fedtschenkoi</emphasis>
Morawitz, 1876
</taxonomicName>
which have the hind tarsal claws unidentate. The
<taxonomicName id="2E824D69AC7B43590E38FD1C8BF604B1" authorityName="Smith" authorityYear="1853" box="[1324,1427,727,752]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E38FD1C8BF604B1" box="[1324,1427,727,752]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
- group therefore nominally contains seven species:
<taxonomicName id="2E824D69AC7B435909D9FD308A620554" authority="Morawitz, 1877" authorityName="Morawitz" authorityYear="1877" box="[717,1031,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="DBF6EAF8AC7B435909D9FD308D2E0555" box="[717,843,763,789]" italics="true" pageId="34" pageNumber="35">A. carinata</emphasis>
<bibRefCitation id="8D134B1BAC7B43590845FD308A620554" author="Morawitz, F." box="[849,1031,763,789]" pageId="34" pageNumber="35" pagination="3 - 112" refId="ref89413" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14, 3 - 112." type="journal article" year="1877">Morawitz, 1877</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590F05FD308B4C0554" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[1041,1321,762,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F05FD308AF20555" box="[1041,1175,763,789]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F8AFD308B4C0554" author="Smith, F." box="[1182,1321,762,789]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590E20FD308F210578" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590E20FD308BF80555" box="[1332,1437,763,789]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
<bibRefCitation id="8D134B1BAC7B43590B83FCD48F210578" author="Warncke, K." box="[151,324,798,825]" pageId="34" pageNumber="35" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590A44FCD48CFA0578" authority="Grunwaldt, 2005" authorityName="Grunwaldt" authorityYear="2005" box="[336,671,798,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="heinrichi">
<emphasis id="DBF6EAF8AC7B43590A44FCD48FB00578" box="[336,469,799,825]" italics="true" pageId="34" pageNumber="35">A. heinrichi</emphasis>
<bibRefCitation id="8D134B1BAC7B43590ACAFCD48CFA0578" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[478,671,798,825]" pageId="34" pageNumber="35" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">Grünwaldt, 2005</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B435909BFFCD48DA80578" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[683,973,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7B435909BFFCD48D520578" box="[683,823,799,825]" italics="true" pageId="34" pageNumber="35">A. paradoxa</emphasis>
Friese, 1921
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B435908CEFCD48B6C0578" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[986,1289,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B435908CEFCD48A300578" box="[986,1109,799,825]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F48FCD48B6C0578" author="Warncke, K." box="[1116,1289,799,825]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="2E824D69AC7B43590E52FCD48FE1051C" authority="Radoszkowski, 1876" authorityName="Radoszkowski" authorityYear="1876" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7B43590E52FCD48BF90579" box="[1350,1436,799,825]" italics="true" pageId="34" pageNumber="35">A. tecta</emphasis>
Radoszkowski, 1876
</taxonomicName>
.
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FCAC8F7202A8" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
The oldest species name within this group is
<taxonomicName id="2E824D69AC7B435909C2FCAC8D3A05C1" authorityName="Smith" authorityYear="1853" box="[726,863,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B435909C2FCAC8D3A05C1" box="[726,863,871,897]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
which was described from
<collectingCountry id="9195767AAC7B43590FB4FCAC8A9905C0" box="[1184,1276,871,897]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
, and which is currently known only from Mediterranean
<collectingCountry id="9195767AAC7B4359096DFC408CB705E4" box="[633,722,907,933]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
and
<collectingCountry id="9195767AAC7B4359081EFC408D3805E4" box="[778,861,907,933]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
. The type series of
<taxonomicName id="2E824D69AC7B43590F2DFC408AA505E5" authorityName="Smith" authorityYear="1853" box="[1081,1216,907,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F2DFC408AA505E5" box="[1081,1216,907,933]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
could not be found in NHMUK which is listed as the type depository by
<bibRefCitation id="8D134B1BAC7B435909E6FC648A390588" author="Gusenleitner, F. &amp; Schwarz, M." box="[754,1116,943,969]" pageId="34" pageNumber="35" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
.
<bibRefCitation id="8D134B1BAC7B43590F73FC648B530588" author="Smith, F." box="[1127,1334,942,969]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith (1853: 106)</bibRefCitation>
lists that the type material comes from the collection of W.W. Saunders, but there is no entry in the type catalogue of the NHMUK. In contrast,
<bibRefCitation id="8D134B1BAC7B43590A83FC3C8C030250" author="Smith, F." box="[407,614,1014,1041]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith (1853: 107)</bibRefCitation>
does not list a collection for
<taxonomicName id="2E824D69AC7B435908A7FC3C8AB10250" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[947,1236,1014,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="DBF6EAF8AC7B435908A7FC3C8A5B0250" box="[947,1086,1015,1041]" italics="true" pageId="34" pageNumber="35">A. variabilis</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F53FC3C8AB10250" author="Smith, F." box="[1095,1236,1014,1041]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
, and this material can be found in the NHMUK type collection. This suggests that since Smith indicated a private collection for
<taxonomicName id="2E824D69AC7B43590E90FBD08E9B0219" authorityName="Smith" authorityYear="1853" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E90FBD08E9B0219" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, the material did not come from the NHMUK general collection and hence may be lost. However, it is not considered necessary to designate a
<typeStatus id="36398848AC7B4359097CFBA88CA4023C" box="[616,705,1123,1149]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
at this point, as the identity of this taxon is clear based on the description and also based on biogeography, since only one member of the
<taxonomicName id="2E824D69AC7B435908E6FB4C8A3C02E1" authorityName="Smith" authorityYear="1853" box="[1010,1113,1159,1184]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B435908E6FB4C8A3C02E1" box="[1010,1113,1159,1184]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
-group is present in
<collectingCountry id="9195767AAC7B43590E29FB4C8BFC02E0" box="[1341,1433,1159,1185]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
, i.e.
<taxonomicName id="2E824D69AC7B43590BD5FB608F220285" authorityName="Smith" authorityYear="1853" box="[193,327,1195,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590BD5FB608F220285" box="[193,327,1195,1221]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
. Additional searches should be made in the NHMUK and OUMNH collections before a
<typeStatus id="36398848AC7B43590E3EFB608BE60284" box="[1322,1411,1195,1221]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
is designated.
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FB388CF50380" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
Warncke took a very broad approach to this species complex. As females are almost morphologically inseparable, differing most obviously in the strength of the tergal hairbands, he treated taxa as subspecies of
<taxonomicName id="2E824D69AC7B43590E90FADC8E9B0315" authorityName="Smith" authorityYear="1853" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E90FADC8E9B0315" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, describing
<taxonomicName id="2E824D69AC7B43590A91FAF08C3F0315" authorityName="Warncke" authorityYear="1965" box="[389,602,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
<emphasis id="DBF6EAF8AC7B43590A91FAF08C3F0315" box="[389,602,1339,1365]" italics="true" pageId="34" pageNumber="35">A. concinna crecca</emphasis>
</taxonomicName>
from western
<collectingCountry id="9195767AAC7B435909ECFAF08D2D0314" box="[760,840,1339,1365]" name="Turkey" pageId="34" pageNumber="35">Turkey</collectingCountry>
(Brussa =
<collectingRegion id="2B46F808AC7B435908ABFAF08A610314" box="[959,1028,1339,1365]" country="Turkey" name="Bursa" pageId="34" pageNumber="35">Bursa</collectingRegion>
;
<figureCitation id="71B92A6FAC7B43590F1AFAF08A1F0314" box="[1038,1146,1339,1365]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="34" pageNumber="35">Figure 19</figureCitation>
) and
<taxonomicName id="2E824D69AC7B43590FA3FAF08BF90314" authorityName="Warncke" authorityYear="1973" box="[1207,1436,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="resoluta">
<emphasis id="DBF6EAF8AC7B43590FA3FAF08BF90314" box="[1207,1436,1339,1365]" italics="true" pageId="34" pageNumber="35">A. concinna resoluta</emphasis>
</taxonomicName>
from North
<collectingCountry id="9195767AAC7B43590A37FA948FFA0338" box="[291,415,1375,1401]" name="Macedonia" pageId="34" pageNumber="35">Macedonia</collectingCountry>
(Skopje) (
<bibRefCitation id="8D134B1BAC7B4359090FFA948CA10338" author="Warncke, K." box="[539,708,1374,1401]" pageId="34" pageNumber="35" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke 1965</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7B435909C7FA948D6A0338" author="Warncke, K." box="[723,783,1375,1401]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">1973</bibRefCitation>
). However, within this group, there are clear differences in the genital capsule, and
<bibRefCitation id="8D134B1BAC7B43590AD3FA488D4A03DC" author="Gusenleitner, F. &amp; Schwarz, M." box="[455,815,1411,1437]" pageId="34" pageNumber="35" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
recognised six of these taxa as species, with a seventh described soon after (
<bibRefCitation id="8D134B1BAC7B43590A93FA6C8CE10380" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[391,644,1446,1473]" pageId="34" pageNumber="35" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">
Grünwaldt
<emphasis id="DBF6EAF8AC7B4359091CFA6C8C5C0380" box="[520,569,1447,1473]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FA008CB50130" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
Both
<taxonomicName id="2E824D69AC7B43590A12FA008FE703A4" authorityName="Warncke" authorityYear="1973" box="[262,386,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B43590A12FA008FE703A4" box="[262,386,1483,1509]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7B43590AAFFA008C4303A5" authorityName="Warncke" authorityYear="1965" box="[443,550,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590AAFFA008C4303A5" box="[443,550,1483,1509]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
have been recognised as distinct species, but the differences between the two are obscure. Moreover, the distributional ranges are unclear;
<taxonomicName id="2E824D69AC7B4359085FFA248DA30048" authorityName="Warncke" authorityYear="1973" box="[843,966,1519,1545]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B4359085FFA248DA30048" box="[843,966,1519,1545]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
was considered a species of the southern Balkans around North
<collectingCountry id="9195767AAC7B43590AB4F9D88C79006C" box="[416,540,1555,1581]" name="Macedonia" pageId="34" pageNumber="35">Macedonia</collectingCountry>
and south-western
<collectingCountry id="9195767AAC7B43590816F9D88D07006C" box="[770,866,1555,1581]" name="Bulgaria" pageId="34" pageNumber="35">Bulgaria</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7B43590860F9D88A7A006C" author="Warncke, K." box="[884,1055,1555,1581]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke 1973</bibRefCitation>
), but specimens can be found in south-eastern
<collectingCountry id="9195767AAC7B43590A22F9FC8FF30010" box="[310,406,1591,1617]" name="Bulgaria" pageId="34" pageNumber="35">Bulgaria</collectingCountry>
close to the Turkish border (see below) and hence not far from Bursa, and specimens of
<taxonomicName id="2E824D69AC7B43590B83F9908F640035" authorityName="Warncke" authorityYear="1965" box="[151,257,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590B83F9908F640035" box="[151,257,1627,1653]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
have been reported from the East
<collectingRegion id="2B46F808AC7B4359099CF9908CBB0034" box="[648,734,1627,1653]" country="Greece" name="Voreio Aigaio" pageId="34" pageNumber="35">Aegean</collectingRegion>
Islands of
<collectingCountry id="9195767AAC7B43590849F9908DC90034" box="[861,940,1627,1653]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7B435908A8F9908ACD0034" author="Rasmont, P. &amp; Devalez, J. &amp; Pauly, A. &amp; Michez, D. &amp; Radchenko, V." box="[956,1192,1627,1653]" pageId="34" pageNumber="35" pagination="17 - 32" refId="ref90635" refString="Rasmont, P., Devalez, J., Pauly, A., Michez, D. &amp; Radchenko, V. (2017) Addition to the checklist of IUCN European wild bees (Hymenoptera: Apoidea). Annales de la Societe entomologique de France, New Series, 53, 17 - 32. https: // doi. org / 10.1080 / 00379271.2017.1307696" type="journal article" year="2017">
Rasmont
<emphasis id="DBF6EAF8AC7B43590F3DF9908A390034" box="[1065,1116,1627,1653]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2017
</bibRefCitation>
).
<bibRefCitation id="8D134B1BAC7B43590FAFF9908BF90034" author="Warncke, K." box="[1211,1436,1627,1653]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke (1973: 28)</bibRefCitation>
gives small differences between
<taxonomicName id="2E824D69AC7B4359091EF9B48CE000D8" authorityName="Warncke" authorityYear="1973" box="[522,645,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B4359091EF9B48CE000D8" box="[522,645,1663,1689]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
and what can be assumed is the nominate
<taxonomicName id="2E824D69AC7B43590F7AF9B48A9000D9" authorityName="Smith" authorityYear="1853" box="[1134,1269,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F7AF9B48A9000D9" box="[1134,1269,1663,1689]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, since he does not specify the comparison taxon until mentioning
<taxonomicName id="2E824D69AC7B435909CEF9688D3200FD" authorityName="Warncke" authorityYear="1965" box="[730,855,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
ssp.
<emphasis id="DBF6EAF8AC7B43590819F9688D3200FD" box="[781,855,1699,1724]" italics="true" pageId="34" pageNumber="35">crecca</emphasis>
</taxonomicName>
, where he argues that the gonocoxae of
<taxonomicName id="2E824D69AC7B43590E35F9688BF900FC" authorityName="Warncke" authorityYear="1973" box="[1313,1436,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B43590E35F9688BF900FC" box="[1313,1436,1699,1725]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
have a wide and sharply developed longitudinal ridge on their inner margin, whereas in
<taxonomicName id="2E824D69AC7B43590F75F90C8AAC00A1" authorityName="Warncke" authorityYear="1965" box="[1121,1225,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590F75F90C8AAC00A1" box="[1121,1225,1735,1761]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
the transition from the gonocoxae to the gonostyli is smooth [“
<emphasis id="DBF6EAF8AC7B43590986F9208D140169" italics="true" pageId="34" pageNumber="35">Gonocoxiten besitzen auf der Innenseite je eine scharf ausgebildete und breite, längsverlaufende Leiste (bei der östlichen Unterart</emphasis>
<taxonomicName id="2E824D69AC7B4359086FF8C48A000168" authority="Warncke" authorityName="Warncke" authorityYear="1965" box="[891,1125,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
<emphasis id="DBF6EAF8AC7B4359086FF8C48DC20169" box="[891,935,1807,1832]" italics="true" pageId="34" pageNumber="35">ssp.</emphasis>
crecca
<emphasis id="DBF6EAF8AC7B43590F10F8C48A000168" box="[1028,1125,1807,1833]" italics="true" pageId="34" pageNumber="35">Warncke</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC7B43590F7BF8C48CD70130" italics="true" pageId="34" pageNumber="35">ist der Übergang von den Gonocoxiten zu den Gonostylen gleichmässig, und der etwas höhere Penisaufsatz ist nach hinten zu noch starker verlängert, die Penisspitze nach unten gebogen)</emphasis>
”].
</paragraph>
<caption id="BDFD6662AC7A43580B83F85C8CCD01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209349" ID-Zenodo-Dep="13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1943,1967]" targetBox="[214,1374,181,1919]" targetPageId="35" targetType="figure">
<paragraph id="E93D36EAAC7A43580B83F85C8CCD01B6" blockId="35.[151,1437,1943,2040]" pageId="35" pageNumber="36">
<emphasis id="DBF6EAF8AC7A43580B83F85C8F4401EE" bold="true" box="[151,289,1943,1967]" pageId="35" pageNumber="36">FIGURE 19.</emphasis>
<taxonomicName id="2E824D69AC7A43580A3CF85C8C8801EE" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[296,749,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="crecca" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7A43580A3CF85C8F1B01EE" box="[296,382,1943,1967]" italics="true" pageId="35" pageNumber="36">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7A43580A9AF85C8F9E01EE" box="[398,507,1943,1967]" italics="true" pageId="35" pageNumber="36">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7A4358091EF8538C2801EE" box="[522,589,1944,1967]" italics="true" pageId="35" pageNumber="36">crecca</emphasis>
<bibRefCitation id="8D134B1BAC7A43580940F85C8C8801EE" author="Warncke, K." box="[596,749,1943,1967]" pageId="35" pageNumber="36" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC7A435809E0F8538D3401F1" bold="true" box="[756,849,1944,1968]" pageId="35" pageNumber="36">holotype</emphasis>
male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail.
</paragraph>
</caption>
<paragraph id="E93D36EAAC7D435F0BD3FF5C8A5307B4" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
It is not clear what Warncke meant by this, because examination of the respective type material shows that the
<typeStatus id="36398848AC7D435F0B83FF708E9C0694" box="[151,249,187,213]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC7D435F0A35FF708FEE0695" authorityName="Warncke" authorityYear="1965" box="[289,395,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F0A35FF708FEE0695" box="[289,395,187,213]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
does have a developed longitudinal ridge on the inner margins of each gonocoxite (
<figureCitation id="71B92A6FAC7D435F0E52FF708E8E06B8" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figures 19F; H</figureCitation>
). In the
<typeStatus id="36398848AC7D435F0A5EFF148FC906B8" box="[330,428,223,249]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
itself, it is not so strongly developed as in type material of
<taxonomicName id="2E824D69AC7D435F0F42FF148AB406B8" authorityName="Warncke" authorityYear="1973" box="[1110,1233,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F0F42FF148AB406B8" box="[1110,1233,223,249]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC7D435F0FF5FF148B0606B8" box="[1249,1379,223,249]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37">Figure 20F</figureCitation>
), but examination of material from across
<collectingCountry id="9195767AAC7D435F0923FEC88CE2075C" box="[567,647,259,285]" name="Turkey" pageId="36" pageNumber="37">Turkey</collectingCountry>
shows that this is a little variable, and many individuals can be found that have the internal ridges as strongly produced as
<taxonomicName id="2E824D69AC7D435F09F3FEEC8D040700" authorityName="Warncke" authorityYear="1973" box="[743,865,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F09F3FEEC8D040700" box="[743,865,295,321]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
. Unfortunately, the penis valves of the
<typeStatus id="36398848AC7D435F0E08FEEC8B1B0700" box="[1308,1406,295,321]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC7D435F0B83FE808F650725" authorityName="Warncke" authorityYear="1965" box="[151,256,331,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F0B83FE808F650725" box="[151,256,331,357]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
are not in a normal position (
<figureCitation id="71B92A6FAC7D435F095CFE808C920724" box="[584,759,331,357]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figures 19F–G</figureCitation>
), but the apex of the penis valves can be seen where they are straight (
<figureCitation id="71B92A6FAC7D435F0BE8FEA48FE607C8" box="[252,387,367,393]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figure 19G</figureCitation>
), whereas in
<taxonomicName id="2E824D69AC7D435F0908FEA48CFD07C8" authorityName="Warncke" authorityYear="1973" box="[540,664,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F0908FEA48CFD07C8" box="[540,664,367,393]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
the apex is slightly upturned (
<figureCitation id="71B92A6FAC7D435F08E2FEA48A1E07C8" box="[1014,1147,367,393]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37">Figure 20E</figureCitation>
). However, intermediates can be found between these two states, for example from Malko Tarnovo in south-eastern
<collectingCountry id="9195767AAC7D435F0F86FE588A9307EC" box="[1170,1270,403,429]" name="Bulgaria" pageId="36" pageNumber="37">Bulgaria</collectingCountry>
. In the context of examination of material from the southern Balkans to the Caucasus, these differences are considered to be both small and variable, and hence
<taxonomicName id="2E824D69AC7D435F0AFEFE108C0107B4" authorityName="Warncke" authorityYear="1973" box="[490,612,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta" status="syn. nov.">
<emphasis id="DBF6EAF8AC7D435F0AFEFE108C0107B4" box="[490,612,475,501]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC7D435F097FFE108CB607B4" bold="true" box="[619,723,475,501]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="C0C55783AC7D435F097FFE108CB607B4" box="[619,723,475,501]" pageId="36" pageNumber="37">syn. nov.</taxonomicNameLabel>
</emphasis>
is synonymised with
<taxonomicName id="2E824D69AC7D435F08D3FE108A5507B5" authorityName="Warncke" authorityYear="1965" box="[967,1072,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F08D3FE108A5507B5" box="[967,1072,475,501]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BDFD6662AC7D435F0B83F87C8AEB01B2" ID-DOI="http://doi.org/10.5281/zenodo.13209351" ID-Zenodo-Dep="13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1975,1999]" targetBox="[151,1436,510,1960]" targetPageId="36" targetType="figure">
<paragraph id="E93D36EAAC7D435F0B83F87C8AEB01B2" blockId="36.[151,1436,1975,2036]" pageId="36" pageNumber="37">
<emphasis id="DBF6EAF8AC7D435F0B83F87C8F45018E" bold="true" box="[151,288,1975,1999]" pageId="36" pageNumber="37">FIGURE 20.</emphasis>
<taxonomicName id="2E824D69AC7D435F0A31F87C8C930191" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[293,758,1975,2000]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7D435F0A31F87C8F1E018E" box="[293,379,1975,1999]" italics="true" pageId="36" pageNumber="37">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7D435F0A9EF87C8F92018E" box="[394,503,1975,1999]" italics="true" pageId="36" pageNumber="37">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7D435F0912F87C8C3D018E" box="[518,600,1975,1999]" italics="true" pageId="36" pageNumber="37">resoluta</emphasis>
<bibRefCitation id="8D134B1BAC7D435F0949F87C8C930191" author="Warncke, K." box="[605,758,1975,2000]" pageId="36" pageNumber="37" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC7D435F09E8F8738D380191" bold="true" box="[764,861,1976,2000]" pageId="36" pageNumber="37">paratype</emphasis>
male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<caption id="BDFD6662AC7C435E0B83F8BF8B7201B9" ID-DOI="http://doi.org/10.5281/zenodo.13209355" ID-Zenodo-Dep="13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,243,1908,1932]" targetBox="[214,1373,161,1792]" targetPageId="37" targetType="figure">
<paragraph id="E93D36EAAC7C435E0B83F8BF8B7201B9" blockId="37.[151,1437,1908,2040]" pageId="37" pageNumber="38">
<emphasis id="DBF6EAF8AC7C435E0B83F8BF8F7001CD" bold="true" box="[151,277,1908,1932]" pageId="37" pageNumber="38">FIGURE 21.</emphasis>
<taxonomicName id="2E824D69AC7C435E0A0DF8BF8CD001CD" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[281,693,1908,1932]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="concinna" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0A0DF8BF8F0C01CD" box="[281,361,1908,1932]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0A62F8BF8FBE01CD" box="[374,475,1908,1932]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0AFCF8BE8C5B01CD" box="[488,574,1909,1932]" italics="true" pageId="37" pageNumber="38">concinna</emphasis>
<bibRefCitation id="8D134B1BAC7C435E0957F8BF8CD001CD" author="Smith, F." box="[579,693,1908,1932]" pageId="37" pageNumber="38" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E0E58F8BF8F8701F1" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="paradoxa" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0E58F8BF8BF901CD" box="[1356,1436,1908,1932]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0B8BF8538F6101F1" box="[159,260,1944,1968]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0A05F8538F0E01F1" box="[273,363,1944,1968]" italics="true" pageId="37" pageNumber="38">paradoxa</emphasis>
Friese, 1921
</taxonomicName>
male. B. Genital capsule, lateral view; D. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E0F6DF8538F520195" authority="Grunwaldt, 2005" authorityName="Grunwaldt" authorityYear="2005" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="heinrichi" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0F6DF8538AAC01F1" box="[1145,1225,1944,1968]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0FC2F8538B5E01F1" box="[1238,1339,1944,1968]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0E53F8538BF901F1" box="[1351,1436,1944,1968]" italics="true" pageId="37" pageNumber="38">heinrichi</emphasis>
<bibRefCitation id="8D134B1BAC7C435E0B83F8778F520195" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[151,311,1980,2004]" pageId="37" pageNumber="38" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">Grünwaldt, 2005</bibRefCitation>
</taxonomicName>
male. E. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E09B3F8778A0A0195" authority="Radoszkowski, 1876" authorityName="Radoszkowski" authorityYear="1876" box="[679,1135,1980,2004]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="tecta" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E09B3F8778C920195" box="[679,759,1980,2004]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0811F8778D0F0195" box="[773,874,1980,2004]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0863F8768DC00195" box="[887,933,1981,2004]" italics="true" pageId="37" pageNumber="38">tecta</emphasis>
Radoszkowski, 1876
</taxonomicName>
male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view;
<taxonomicName id="2E824D69AC7C435E0913F82B8DCC01B9" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[519,937,2016,2040]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crecca" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0913F82B8C3201B9" box="[519,599,2016,2040]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0971F82B8CAF01B9" box="[613,714,2016,2040]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E09CCF82A8D7301B9" box="[728,790,2017,2040]" italics="true" pageId="37" pageNumber="38">crecca</emphasis>
<bibRefCitation id="8D134B1BAC7C435E080FF82B8DCC01B9" author="Warncke, K." box="[795,937,2016,2040]" pageId="37" pageNumber="38" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
male. H. Genital capsule, lateral view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC7F435D0BD3FF5C8CB507B4" blockId="38.[151,1437,151,609]" pageId="38" pageNumber="39">
Searches in the ZMHB were unable to find material of
<taxonomicName id="2E824D69AC7F435D0845FF5C8DCC06F1" authorityName="Radoszkowski" authorityYear="1876" box="[849,937,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0845FF5C8DCC06F1" box="[849,937,151,177]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
, the listed
<typeStatus id="36398848AC7F435D0F3AFF5C8A3B06F0" box="[1070,1118,151,177]" pageId="38" pageNumber="39">type</typeStatus>
repository (
<bibRefCitation id="8D134B1BAC7F435D0FFFFF5C8F580694" author="Gusenleitner, F. &amp; Schwarz, M." pageId="38" pageNumber="39" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
). In the relevant box, there is a pinned note indicating that
<typeStatus id="36398848AC7F435D08CBFF708A6A0694" box="[991,1039,187,213]" pageId="38" pageNumber="39">type</typeStatus>
material from both
<taxonomicName id="2E824D69AC7F435D0FE0FF708B2E0695" authorityName="Radoszkowski" authorityYear="1876" box="[1268,1355,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0FE0FF708B2E0695" box="[1268,1355,187,213]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7F435D0E90FF708F6706B8" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7F435D0E90FF708F6706B8" italics="true" pageId="38" pageNumber="39">A. paradoxa</emphasis>
</taxonomicName>
have been on loan to Robert Wilhelm Grünwaldt (1909–†2003) since 1977. Since Grünwaldt’s collection was donated to the ZSM before his death, the material is likely to be there. No taxonomic action is therefore taken regarding
<taxonomicName id="2E824D69AC7F435D0A1FFEEC8F070701" authorityName="Radoszkowski" authorityYear="1876" box="[267,354,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0A1FFEEC8F070701" box="[267,354,295,321]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
. Finally,
<taxonomicName id="2E824D69AC7F435D0ADBFEEC8C2B0701" authorityName="Morawitz" authorityYear="1877" box="[463,590,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="DBF6EAF8AC7F435D0ADBFEEC8C2B0701" box="[463,590,295,321]" italics="true" pageId="38" pageNumber="39">A. carinata</emphasis>
</taxonomicName>
was described in the male sex only from modern day
<collectingCountry id="9195767AAC7F435D0FAFFEEC8B780700" box="[1211,1309,295,321]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7F435D0E39FEEC8EB70724" author="Morawitz, F." pageId="38" pageNumber="39" pagination="3 - 112" refId="ref89413" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14, 3 - 112." type="journal article" year="1877">Morawitz 1877</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7F435D0BCAFE808FFA0724" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[222,415,331,357]" pageId="38" pageNumber="39" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">Astafurova 2021</bibRefCitation>
). Although it can confidently be placed into the group of large-bodied
<taxonomicName id="2E824D69AC7F435D0FDEFE808B270724" authorityName="Warncke" authorityYear="1968" box="[1226,1346,331,357]" pageId="38" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7F435D0FDEFE808B270724" box="[1226,1346,331,357]" italics="true" pageId="38" pageNumber="39">Ulandrena</emphasis>
</taxonomicName>
species based on its declared size (16.5 mm) and declared morphological similarity to
<taxonomicName id="2E824D69AC7F435D0F0BFEA48AA907C8" authorityName="Morawitz" authorityYear="1876" box="[1055,1228,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="DBF6EAF8AC7F435D0F0BFEA48AA907C8" box="[1055,1228,367,393]" italics="true" pageId="38" pageNumber="39">A. fedtschenkoi</emphasis>
</taxonomicName>
, it lacks a precise description or illustration of the genital capsule. As such, it could be a junior synonym of either
<taxonomicName id="2E824D69AC7F435D0FDDFE588B7A07ED" authorityName="Radoszkowski" authorityYear="1876" box="[1225,1311,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0FDDFE588B7A07ED" box="[1225,1311,403,429]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
or a senior synonym of
<taxonomicName id="2E824D69AC7F435D0A35FE7C8FEF0791" authorityName="Warncke" authorityYear="1965" box="[289,394,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7F435D0A35FE7C8FEF0791" box="[289,394,439,465]" italics="true" pageId="38" pageNumber="39">A. crecca</emphasis>
</taxonomicName>
since both taxa occur in the Caucasus, but as the
<typeStatus id="36398848AC7F435D08BFFE7C8DBE0790" box="[939,987,439,465]" pageId="38" pageNumber="39">type</typeStatus>
material is lost (
<bibRefCitation id="8D134B1BAC7F435D0F86FE7C8BF50790" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[1170,1424,439,465]" pageId="38" pageNumber="39" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC7F435D0E03FE7C8B220790" box="[1303,1351,439,465]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2021
</bibRefCitation>
), it is better to officially declare it a
<emphasis id="DBF6EAF8AC7F435D090EFE108CB507B4" bold="true" box="[538,720,474,501]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D090EFE108CAC07B4" bold="true" box="[538,713,474,501]" italics="true" pageId="38" pageNumber="39">nomen dubium</emphasis>
.
</emphasis>
</paragraph>
<paragraph id="E93D36EAAC7F435D0BD3FE348B640420" blockId="38.[151,1437,151,609]" pageId="38" pageNumber="39">
The
<taxonomicName id="2E824D69AC7F435D0BE8FE348F060459" authorityName="Smith" authorityYear="1853" box="[252,355,511,536]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D0BE8FE348F060459" box="[252,355,511,536]" italics="true" pageId="38" pageNumber="39">concinna</emphasis>
</taxonomicName>
-group is therefore reduced to five species,
<taxonomicName id="2E824D69AC7F435D084EFE348D860459" authorityName="Smith" authorityYear="1853" box="[858,995,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D084EFE348D860459" box="[858,995,511,537]" italics="true" pageId="38" pageNumber="39">A. concinna</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D08E0FE348A340458" box="[1012,1105,511,537]" name="Albania" pageId="38" pageNumber="39">Albania</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0F4BFE348AD60458" box="[1119,1203,511,537]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D0FDCFE348B560459" authorityName="Warncke" authorityYear="1965" box="[1224,1331,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7F435D0FDCFE348B560459" box="[1224,1331,511,537]" italics="true" pageId="38" pageNumber="39">A. crecca</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D0E50FE348BFD0458" box="[1348,1432,511,537]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0B83FDE88E81047C" box="[151,228,547,573]" name="Serbia" pageId="38" pageNumber="39">Serbia</collectingCountry>
*, North
<collectingCountry id="9195767AAC7F435D0A58FDE88FAE047C" box="[332,459,547,573]" name="Macedonia" pageId="38" pageNumber="39">Macedonia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0ACFFDE88C5B047C" box="[475,574,547,573]" name="Bulgaria" pageId="38" pageNumber="39">Bulgaria</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D095AFDE88CFA047C" box="[590,671,547,573]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D09B9FDE88D71047C" box="[685,788,547,573]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D083FFDE88DD1047C" authorityName="Grunwaldt" authorityYear="2005" box="[811,948,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="heinrichi">
<emphasis id="DBF6EAF8AC7F435D083FFDE88DD1047C" box="[811,948,547,573]" italics="true" pageId="38" pageNumber="39">A. heinrichi</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D08D3FDE88A79047C" box="[967,1052,547,573]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
: Samos and western
<collectingCountry id="9195767AAC7F435D0E0CFDE88B09047C" box="[1304,1388,547,573]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D0E90FDE88F670420" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7F435D0E90FDE88F670420" italics="true" pageId="38" pageNumber="39">A. paradoxa</emphasis>
</taxonomicName>
(southern
<collectingCountry id="9195767AAC7F435D0A6EFD8C8FAE0420" box="[378,459,583,609]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0AC2FD8C8C710420" box="[470,532,583,609]" name="Syria" pageId="38" pageNumber="39">Syria</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D090BFD8C8C040420" box="[543,609,583,609]" name="Israel" pageId="38" pageNumber="39">Israel</collectingCountry>
), and
<taxonomicName id="2E824D69AC7F435D09B0FD8C8C9C0421" authorityName="Radoszkowski" authorityYear="1876" box="[676,761,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D09B0FD8C8C9C0421" box="[676,761,583,609]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D081CFD8C8D3E0420" box="[776,859,583,609]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0872FD8C8DA60420" box="[870,963,583,609]" name="Georgia" pageId="38" pageNumber="39">Georgia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D08D8FD8C8A540420" box="[972,1073,583,609]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0F2EFD8C8ADD0420" box="[1082,1208,583,609]" name="Azerbaijan" pageId="38" pageNumber="39">Azerbaijan</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0FD7FD8C8A900420" box="[1219,1269,583,609]" name="Iran" pageId="38" pageNumber="39">Iran</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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