Add updates up until 2024-08-13 15:28:28

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ggserver 2024-08-13 15:34:37 +00:00
parent f2b33e0463
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27 changed files with 12368 additions and 45 deletions

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<mods:title id="017BF486AC13195397F95197A67A5221">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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;
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<collectingRegion id="2B46F808AC03432109ACFC3C8D700250" box="[696,789,1015,1041]" country="Turkey" name="Hakkari" pageId="90" pageNumber="91">Hakkâri</collectingRegion>
,
<location id="EC5D6031AC0343210836FC3C8D8E0250" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0343210836FC3C8D8E0250" box="[802,1003,1015,1041]" country="Turkey" name="Suvari Halil-Pass" pageId="90" pageNumber="91" stateProvince="Hakkari">Suvari Halil-Pass</location>
,
<quantity id="2E7A9B0FAC03432108ECFC3C8A370250" box="[1016,1106,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="90" pageNumber="91" unit="m" value="2800.0">
<elevation id="62AFD1D9AC03432108ECFC3C8A370250" box="[1016,1106,1015,1041]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="90" pageNumber="91" unit="m" value="2800.0">2800 m</elevation>
</quantity>
,
<date id="9D3C102AAC0343210F74FC3C8ABF0250" box="[1120,1242,1015,1041]" pageId="90" pageNumber="91" value="1984-06-14">
<collectingDate id="8D78E9C2AC0343210F74FC3C8ABF0250" box="[1120,1242,1015,1041]" pageId="90" pageNumber="91" value="1984-06-14">14.vi.1984</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0343210FFCFC3C8B680250" box="[1256,1293,1015,1041]" count="4" pageId="90" pageNumber="91" type="male">4♂</specimenCount>
,
<specimenCount id="FF84FD63AC0343210E0FFC3C8B5B0250" box="[1307,1342,1015,1041]" count="1" pageId="90" pageNumber="91" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0343210E95FC3C8E980274" pageId="90" pageNumber="91">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0343210A7BFBD08BF10274" box="[367,1428,1051,1079]" collectingDate="1989-05-20" collectorName="K. Warncke &amp; Nemrut Dagi" country="Turkey" elevation="2900" location="Suvari Halil-Pass" pageId="90" pageNumber="91" specimenCount="1" specimenCount-female="1" stateProvince="Hakkari" typeStatus="paratype">
<collectingRegion id="2B46F808AC0343210A7BFBD08FA90274" box="[367,460,1051,1077]" country="Turkey" name="Hakkari" pageId="90" pageNumber="91">Hakkâri</collectingRegion>
,
<location id="EC5D6031AC0343210ACFFBD08CC10274" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0343210ACFFBD08CC10274" box="[475,676,1051,1077]" country="Turkey" name="Suvari Halil-Pass" pageId="90" pageNumber="91" stateProvince="Hakkari">Suvari Halil-Pass</location>
,
<quantity id="2E7A9B0FAC03432109A6FBD08D690274" box="[690,780,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="90" pageNumber="91" unit="m" value="2900.0">
<elevation id="62AFD1D9AC03432109A6FBD08D690274" box="[690,780,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="90" pageNumber="91" unit="m" value="2900.0">2900 m</elevation>
</quantity>
,
<date id="9D3C102AAC034321080FFBD08DE90274" box="[795,908,1051,1077]" pageId="90" pageNumber="91" value="1989-05-20">
<collectingDate id="8D78E9C2AC034321080FFBD08DE90274" box="[795,908,1051,1077]" pageId="90" pageNumber="91" value="1989-05-20">20.v.1989</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC034321088EFBD08DD80274" box="[922,957,1051,1077]" count="1" pageId="90" pageNumber="91" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0343210F15FBD08AE90274" box="[1025,1164,1051,1077]" pageId="90" pageNumber="91">K. Warncke</collectorName>
, OÖLM;
<collectorName id="4477533CAC0343210FEAFBD08BF10274" box="[1278,1428,1051,1077]" pageId="90" pageNumber="91">Nemrut Dağı</collectorName>
</materialsCitation>
/
<materialsCitation id="59EA3CB7AC0343210B83FBF48D6E021A" box="[151,779,1086,1115]" collectingDate="1989-05-25" collectorName="K. Warncke" country="Turkey" elevation="2350" location="Bitlis" pageId="90" pageNumber="91" specimenCount="1" specimenCount-female="1" stateProvince="Bitlis" typeStatus="paratype">
<collectingRegion id="2B46F808AC0343210B83FBF48EB30218" box="[151,214,1087,1113]" country="Turkey" name="Bitlis" pageId="90" pageNumber="91">Bitlis</collectingRegion>
,
<quantity id="2E7A9B0FAC0343210BF6FBF58F5C0218" box="[226,313,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="90" pageNumber="91" unit="m" value="2350.0">
<elevation id="62AFD1D9AC0343210BF6FBF58F5C0218" box="[226,313,1086,1113]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="90" pageNumber="91" unit="m" value="2350.0">2350 m</elevation>
</quantity>
,
<date id="9D3C102AAC0343210A51FBF58FD30218" box="[325,438,1086,1113]" pageId="90" pageNumber="91" value="1989-05-25">
<collectingDate id="8D78E9C2AC0343210A51FBF58FD30218" box="[325,438,1086,1113]" pageId="90" pageNumber="91" value="1989-05-25">25.v.1989</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0343210AD5FBF48F810218" box="[449,484,1087,1113]" count="1" pageId="90" pageNumber="91" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0343210936FBF48CCF0218" box="[546,682,1087,1113]" pageId="90" pageNumber="91">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0343260BD3FBA98D9F0488" lastPageId="93" lastPageNumber="94" pageId="90" pageNumber="91" type="description">
<paragraph id="E93D36EAAC0343210BD3FBA98A720380" blockId="90.[151,1437,834,1905]" pageId="90" pageNumber="91">
<emphasis id="DBF6EAF8AC0343210BD3FBA98FA6023C" bold="true" box="[199,451,1122,1149]" pageId="90" pageNumber="91">Description: Female:</emphasis>
Body length: 7.5
<quantity id="2E7A9B0FAC034321098CFBA98CB2023C" box="[664,727,1122,1149]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="90" pageNumber="91" unit="mm" value="8.0">8 mm</quantity>
(
<figureCitation id="71B92A6FAC03432109F2FBA88D08023C" box="[742,877,1122,1149]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="90" pageNumber="91">Figure 59A</figureCitation>
).
<emphasis id="DBF6EAF8AC0343210895FBA88DA2023C" box="[897,967,1123,1149]" italics="true" pageId="90" pageNumber="91">Head:</emphasis>
Dark, 1.2 times longer than wide (
<figureCitation id="71B92A6FAC0343210E45FBA88EAC02E0" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="90" pageNumber="91">Figure 59B</figureCitation>
). Clypeus weakly domed, more or less flattened medially, densely punctate, punctures separated by ≤0.5 puncture diameters over majority of area, only very slightly sparser medially, underlying surface weakly shagreened, shining. Process of labrum rounded trapezoidal, 3 times wider than long, surface smooth and shining. Gena slightly exceeding width of compound eye; ocelloccipital distance slightly exceeding diameter of lateral ocellus. Foveae dorsally narrow, occupying ⅓ of space between compound eye and lateral ocellus, equalling width of flagellum, narrowing further ventrally at level of antennal insertions; foveae filled with dark brown hairs dorsally, becoming light brown ventrally. Face predominantly with pale whitish pubescence, with black hairs restricted to inner margins of compound eyes, gena with whitish hairs, vertex with longer golden-brown hairs. Antennae dark, A512 ventrally lightened by presence of grey scales; A3 exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="E93D36EAAC0343210BD3FA008AD10130" blockId="90.[151,1437,834,1905]" pageId="90" pageNumber="91">
<emphasis id="DBF6EAF8AC0343210BD3FA008F2D03A5" box="[199,328,1483,1508]" italics="true" pageId="90" pageNumber="91">Mesosoma:</emphasis>
Scutum anteriorly and laterally with microsculpture, weakly shining, medially becoming smooth and shining with very weak shagreenation; surface irregularly punctate, punctures separated by 0.52 puncture diameters, by up to 4 puncture diameters medially (
<figureCitation id="71B92A6FAC03432109FEF9D88D14006C" box="[746,881,1554,1581]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="90" pageNumber="91">Figure 59C</figureCitation>
). Pronotum rounded. Mesepisternum with dense granular microreticulation, dull. Dorsolateral parts of propodeum with granular microreticulation, weakly shining, sculpture overlain by sparse network of raised rugosity forming appearance of shallow punctures with raised rims. Propodeal triangle laterally with fine granular shagreen, shining, medially and basally covered with dense network of raised rugae (
<figureCitation id="71B92A6FAC0343210A4EF9688F8300FC" box="[346,486,1698,1725]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="90" pageNumber="91">Figure 59G</figureCitation>
). Mesepisternum laterally with moderate whitish to light brownish hairs, scutum and scutellum with short densely plumose golden-brown hairs, hairs thus appearing sub-squamous (
<figureCitation id="71B92A6FAC0343210E1FF90C8BF500A0" box="[1291,1424,1734,1761]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="90" pageNumber="91">Figure 59E</figureCitation>
). Propodeal corbicula incomplete, weakly formed, dorsal fringe composed of golden-brown plumose hairs, internal surface with scattered golden-brown simple hairs. Legs dark, black to light brown. Flocculus moderately developed, composed of brownish hairs; femoral and tibial scopae composed of golden-brown simple hairs. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark orange-brown, nervulus interstitial.
</paragraph>
<caption id="BDFD6662AC0243200B83F85C8A2901B6" ID-DOI="http://doi.org/10.5281/zenodo.13209431" ID-Zenodo-Dep="13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="91" pageNumber="92" startId="91.[151,250,1943,1967]" targetBox="[215,1373,181,1919]" targetPageId="91" targetType="figure">
<paragraph id="E93D36EAAC0243200B83F85C8A2901B6" blockId="91.[151,1436,1943,2040]" pageId="91" pageNumber="92">
<emphasis id="DBF6EAF8AC0243200B83F85C8F4701EE" bold="true" box="[151,290,1943,1967]" pageId="91" pageNumber="92">FIGURE 59.</emphasis>
<taxonomicName id="2E824D69AC0243200A3EF85C8C0B01EE" authorityName="WOOD" authorityYear="2024" box="[298,622,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="species" species="eosphora" status="sp. nov." subGenus="Euandrena">
<emphasis id="DBF6EAF8AC0243200A3EF85C8FE501EE" box="[298,384,1943,1967]" italics="true" pageId="91" pageNumber="92">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC0243200A84F85C8C6501EE" box="[400,512,1943,1967]" italics="true" pageId="91" pageNumber="92">Euandrena</emphasis>
)
<emphasis id="DBF6EAF8AC0243200904F85C8C0B01EE" box="[528,622,1943,1967]" italics="true" pageId="91" pageNumber="92">eosphora</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC0243200963F8538C8501F1" bold="true" box="[631,736,1944,1968]" pageId="91" pageNumber="92">
<taxonomicNameLabel id="C0C55783AC0243200963F8538C8501F1" box="[631,736,1944,1968]" pageId="91" pageNumber="92" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view.
<taxonomicName id="2E824D69AC0243200F65F8708FDF01B6" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="91" pageNumber="92" phylum="Arthropoda" rank="subSpecies" species="pileata" subGenus="Euandrena" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0243200F65F8708AA20192" box="[1137,1223,1979,2003]" italics="true" pageId="91" pageNumber="92">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC0243200FC3F8708B220192" box="[1239,1351,1979,2003]" italics="true" pageId="91" pageNumber="92">Euandrena</emphasis>
)
<emphasis id="DBF6EAF8AC0243200E42F8708BF90192" box="[1366,1436,1979,2003]" italics="true" pageId="91" pageNumber="92">pileata</emphasis>
ssp.
<emphasis id="DBF6EAF8AC0243200BD0F82B8F7E01B6" box="[196,283,2016,2039]" italics="true" pageId="91" pageNumber="92">canigica</emphasis>
Warncke, 1975
</taxonomicName>
female. D. Scutum, dorsal view; F. Mesepisternum, lateral view.
</paragraph>
</caption>
<caption id="BDFD6662AC0543270B83F9A58A7E00EB" ID-DOI="http://doi.org/10.5281/zenodo.13209435" ID-Zenodo-Dep="13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="92" pageNumber="93" startId="92.[151,250,1646,1670]" targetBox="[151,1436,181,1622]" targetPageId="92" targetType="figure">
<paragraph id="E93D36EAAC0543270B83F9A58A7E00EB" blockId="92.[151,1436,1646,1707]" pageId="92" pageNumber="93">
<emphasis id="DBF6EAF8AC0543270B83F9A58F7A00C7" bold="true" box="[151,287,1646,1670]" pageId="92" pageNumber="93">FIGURE 60.</emphasis>
<taxonomicName id="2E824D69AC0543270A30F9A58C0600C7" authorityName="WOOD" authorityYear="2024" box="[292,611,1646,1670]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="92" pageNumber="93" phylum="Arthropoda" rank="species" species="eosphora" status="sp. nov." subGenus="Euandrena">
<emphasis id="DBF6EAF8AC0543270A30F9A58F1F00C7" box="[292,378,1646,1670]" italics="true" pageId="92" pageNumber="93">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC0543270A9CF9A58F9D00C7" box="[392,504,1646,1670]" italics="true" pageId="92" pageNumber="93">Euandrena</emphasis>
)
<emphasis id="DBF6EAF8AC0543270911F9A58C0600C7" box="[517,611,1646,1670]" italics="true" pageId="92" pageNumber="93">eosphora</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC054327097DF9A58CAA00C7" bold="true" box="[617,719,1646,1670]" pageId="92" pageNumber="93">
<taxonomicNameLabel id="C0C55783AC054327097DF9A58CAA00C7" box="[617,719,1646,1670]" pageId="92" pageNumber="93" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC0543270BD3F9128F4701E7" blockId="92.[151,1437,1752,2031]" pageId="92" pageNumber="93">
<emphasis id="DBF6EAF8AC0543270BD3F9128F2000B3" box="[199,325,1753,1778]" italics="true" pageId="92" pageNumber="93">Metasoma:</emphasis>
Terga dark, margins of T24 distinctly depressed, lightened dark brown, apical rim of T14 narrowly lightened hyaline-yellow (
<figureCitation id="71B92A6FAC0543270AABF9378C220156" box="[447,583,1788,1815]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="92" pageNumber="93">Figure 59H</figureCitation>
). Tergal discs finely microreticulate, sparsely punctate, punctures separated by 13 puncture diameters, punctures not extending onto margins. Tergal discs medially with very scattered short hairs, almost hairless, laterally with erect light brownish hairs. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate rounded triangular, margins impunctate, medially with dense confluent punctures, surface dull.
</paragraph>
<paragraph id="E93D36EAAC0543260BD3F87B8DC60722" blockId="92.[151,1437,1752,2031]" lastBlockId="93.[151,1437,151,2035]" lastPageId="93" lastPageNumber="94" pageId="92" pageNumber="93">
<emphasis id="DBF6EAF8AC0543270BD3F87B8F68018B" bold="true" box="[199,269,1968,1994]" pageId="92" pageNumber="93">Male:</emphasis>
Body length:
<quantity id="2E7A9B0FAC0543270AB8F87A8F88018A" box="[428,493,1969,1995]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="92" pageNumber="93" unit="mm" value="7.0">7 mm</quantity>
(
<figureCitation id="71B92A6FAC0543270AEFF87B8CE4018B" box="[507,641,1968,1994]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="92" pageNumber="93">Figure 60A</figureCitation>
).
<emphasis id="DBF6EAF8AC0543270980F87B8CBF018B" box="[660,730,1968,1994]" italics="true" pageId="92" pageNumber="93">Head:</emphasis>
Dark, 1.2 times wider than long (
<figureCitation id="71B92A6FAC0543270F45F87B8AB0018B" box="[1105,1237,1968,1994]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="92" pageNumber="93">Figure 60B</figureCitation>
). Clypeus weakly domed, more or less flattened medially, densely punctate, punctures separated by ≤0.5 puncture diameters over majority of area, underlying surface shagreened, weakly shining. Process of labrum trapezoidal, 2 times wider than long, apical margin emarginate, surface smooth and shining. Gena equalling width of compound eye; ocelloccipital distance equalling diameter of lateral ocellus. Face medially with white hairs, black hairs restricted to inner margins of compound eyes, gena and vertex with long white hairs; facial hairs often exceeding length of scape.Antennae dark basally, A413 ventrally lightened by presence of grey scales; A3 exceeding A4, shorter than A45; A4 quadrate, slightly wider than long, A513 rectangular, clearly longer than wide.
</paragraph>
<paragraph id="E93D36EAAC0443260BD3FEA58D970457" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<emphasis id="DBF6EAF8AC0443260BD3FEA58F2D07C6" box="[199,328,366,391]" italics="true" pageId="93" pageNumber="94">Mesosoma:</emphasis>
Scutum and scutellum laterally shagreened, weakly shining, medially becoming smooth and shining, irregularly punctate with punctures of variable size, punctures separated by 0.53 puncture diameters (
<figureCitation id="71B92A6FAC0443260E18FE5A8BF507EA" box="[1292,1424,401,427]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60C</figureCitation>
). Pronotum rounded. Mesepisternum and propodeum structurally as in female (
<figureCitation id="71B92A6FAC0443260F19FE7F8AF0078F" box="[1037,1173,436,462]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60D</figureCitation>
). Mesosoma uniformly with long white hairs, longest exceeding length of scape. Legs dark, pubescence whitish. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange, nervulus interstitial.
</paragraph>
<paragraph id="E93D36EAAC0443260BD3FDEB8D9F0488" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<emphasis id="DBF6EAF8AC0443260BD3FDEB8F200478" box="[199,325,544,569]" italics="true" pageId="93" pageNumber="94">Metasoma:</emphasis>
Terga structurally as in female, terga laterally with erect scattered white hairs (
<figureCitation id="71B92A6FAC0443260FDEFDEB8B2A047B" box="[1226,1359,544,570]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60E</figureCitation>
). S67 with long white hairs overlying pygidial plate. S8 narrow, columnar, ventral surface covered with short white hairs. Genital capsule simple, gonocoxae apically produced into weak rounded bumps, gonostyli narrow, slightly broadened apically, spatulate, inner margins slightly raised (
<figureCitation id="71B92A6FAC0443260826FD408DD604E4" box="[818,947,651,677]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60F</figureCitation>
). Penis valves narrow, more or less parallel-sided, occupying ¼ of space between gonostyli, slightly constricted medially.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0443260BD3FD198F810125" pageId="93" pageNumber="94" type="diagnosis">
<paragraph id="E93D36EAAC0443260BD3FD198B2202FC" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<emphasis id="DBF6EAF8AC0443260BD3FD198F2704AD" bold="true" box="[199,322,722,748]" pageId="93" pageNumber="94">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC0443260A59FD198C4504AD" authorityName="WOOD" authorityYear="2024" box="[333,544,722,748]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC0443260A59FD198C4504AD" box="[333,544,722,748]" italics="true" pageId="93" pageNumber="94">Andrena eosphora</emphasis>
</taxonomicName>
can be placed in the subgenus
<taxonomicName id="2E824D69AC0443260888FD198A7204AD" authorityName="Hedicke" authorityYear="1933" box="[924,1047,722,748]" pageId="93" pageNumber="94" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC0443260888FD198A7204AD" box="[924,1047,722,748]" italics="true" pageId="93" pageNumber="94">Euandrena</emphasis>
</taxonomicName>
due to the narrow facial foveae, occupying slightly less than one third of the space between the compound eye and a lateral ocellus, the foveae narrowing further ventrally, pronotum without humeral angle, A3 long (clearly exceeding A4+5), and the simple hairs of the tibial scopae (hairs not plumose). Due to its pale facial pubescence with dark hairs limited to the inner margins of the compound eyes (
<figureCitation id="71B92A6FAC0443260AE1FCAA8C12053D" box="[501,631,865,892]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="93" pageNumber="94">Figure 59B</figureCitation>
), slightly domed and densely punctate clypeus (punctures predominantly separated by ≤0.5 puncture diameters), light brown and densely plumose hairs of the scutum and scutellum (
<figureCitation id="71B92A6FAC0443260E45FC4E8EAD0585" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="93" pageNumber="94">Figure 59E</figureCitation>
; thus appearing sub-squamous), propodeal triangle with raised rugae basally and medially, and terga with almost impunctate discs and clearly depressed marginal areas that are lightened brown (
<figureCitation id="71B92A6FAC0443260F94FC078B6205A6" box="[1152,1287,972,999]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="93" pageNumber="94">Figure 59H</figureCitation>
), it is closest to
<taxonomicName id="2E824D69AC0443260BA0FC3A8C54024A" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[180,561,1008,1035]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="allosa" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0443260BA0FC3A8F1B024B" box="[180,382,1008,1035]" italics="true" pageId="93" pageNumber="94">A. allosa canigica</emphasis>
Warncke, 1975
</taxonomicName>
. Given the work of
<bibRefCitation id="8D134B1BAC044326081BFC3A8DB4024B" author="Praz, C. &amp; Muller, A. &amp; Genoud, D." box="[783,977,1008,1035]" pageId="93" pageNumber="94" pagination="11 - 38" refId="ref90264" refString="Praz, C., Muller, A. and Genoud, D. (2019) Hidden diversity in European bees: Andrena amieti sp. n., a new Alpine bee species related to Andrena bicolor (Fabricius, 1775) (Hymenoptera, Apoidea, Andrenidae). Alpine Entomology, 3, 11 - 38. https: // doi. org / 10.3897 / alpento. 3.29675" type="journal article" year="2019">
Praz
<emphasis id="DBF6EAF8AC044326085CFC3A8D1C024B" box="[840,889,1008,1034]" italics="true" pageId="93" pageNumber="94">et al</emphasis>
. (2019)
</bibRefCitation>
who elevated
<taxonomicName id="2E824D69AC0443260F67FC3A8EAA026E" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="allosa" subSpecies="pileata">
<emphasis id="DBF6EAF8AC0443260F67FC3A8B49024B" box="[1139,1324,1008,1035]" italics="true" pageId="93" pageNumber="94">A. allosa pileata</emphasis>
Warncke, 1975
</taxonomicName>
(
<collectingCountry id="9195767AAC0443260BCAFBDF8F54026F" box="[222,305,1044,1070]" name="Greece" pageId="93" pageNumber="94">Greece</collectingCountry>
) to species status, the status of
<taxonomicName id="2E824D69AC0443260987FBDF8D3A026F" authorityName="Warncke" authorityYear="1975" box="[659,863,1044,1070]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="allosa" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0443260987FBDF8D3A026F" box="[659,863,1044,1070]" italics="true" pageId="93" pageNumber="94">A. allosa canigica</emphasis>
</taxonomicName>
must be dealt with, since it will not be conspecific with
<taxonomicName id="2E824D69AC0443260BC7FBF38F950213" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[211,496,1079,1106]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="allosa">
<emphasis id="DBF6EAF8AC0443260BC7FBF38F5F0213" box="[211,314,1080,1106]" italics="true" pageId="93" pageNumber="94">A. allosa</emphasis>
Warncke, 1975
</taxonomicName>
<emphasis id="DBF6EAF8AC0443260AEFFBF38C5C0210" box="[507,569,1080,1105]" italics="true" pageId="93" pageNumber="94">
<taxonomicNameLabel id="C0C55783AC0443260AEFFBF38C5C0210" box="[507,569,1080,1105]" pageId="93" pageNumber="94" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
for the same reasons given by Praz
<emphasis id="DBF6EAF8AC04432608FCFBF38A790213" box="[1000,1052,1080,1106]" italics="true" pageId="93" pageNumber="94">et al</emphasis>
. Given the strong morphological similarity between
<taxonomicName id="2E824D69AC0443260A63FB978F8C0234" authorityName="Warncke" authorityYear="1975" box="[375,489,1115,1142]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="pileata">
<emphasis id="DBF6EAF8AC0443260A63FB978F8C0234" box="[375,489,1115,1142]" italics="true" pageId="93" pageNumber="94">A. pileata</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC044326093CFB978C990234" authorityName="Warncke" authorityYear="1975" box="[552,764,1115,1142]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="allosa" subSpecies="canigica">
<emphasis id="DBF6EAF8AC044326093CFB978C990234" box="[552,764,1115,1142]" italics="true" pageId="93" pageNumber="94">A. allosa canigica</emphasis>
</taxonomicName>
(differences are predominantly related to hair colour; no sculptural differences could be detected based on examined material, see below), they are considered likely to be conspecific, and so
<taxonomicName id="2E824D69AC0443260A60FB688C2D02FC" authorityName="Warncke" authorityYear="1975" box="[372,584,1187,1213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="pileata" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0443260A60FB688C2D02FC" box="[372,584,1187,1213]" italics="true" pageId="93" pageNumber="94">A. pileata canigica</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC044326095BFB688CB502FC" bold="true" box="[591,720,1187,1213]" pageId="93" pageNumber="94">comb. nov.</emphasis>
is conservatively placed in combination with
<taxonomicName id="2E824D69AC0443260FC7FB688B2502FC" authorityName="Warncke" authorityYear="1975" box="[1235,1344,1187,1213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="pileata">
<emphasis id="DBF6EAF8AC0443260FC7FB688B2502FC" box="[1235,1344,1187,1213]" italics="true" pageId="93" pageNumber="94">A. pileata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E93D36EAAC0443260BD3FB0C8F3C0063" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<taxonomicName id="2E824D69AC0443260BD3FB0C8FFC02A0" authorityName="WOOD" authorityYear="2024" box="[199,409,1223,1249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC0443260BD3FB0C8FFC02A0" box="[199,409,1223,1249]" italics="true" pageId="93" pageNumber="94">Andrena eosphora</emphasis>
</taxonomicName>
is extremely similar to
<taxonomicName id="2E824D69AC04432609A7FB0C8DEB02A1" authorityName="Warncke" authorityYear="1975" box="[691,910,1223,1249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="pileata" subSpecies="canigica">
<emphasis id="DBF6EAF8AC04432609A7FB0C8DEB02A1" box="[691,910,1223,1249]" italics="true" pageId="93" pageNumber="94">A. pileata canigica</emphasis>
</taxonomicName>
, but the two species can be separated by the sculpture and punctation of the scutum which in
<taxonomicName id="2E824D69AC04432609F0FB208D140345" authorityName="WOOD" authorityYear="2024" box="[740,881,1258,1285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC04432609F0FB208D140345" box="[740,881,1258,1285]" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
is laterally microreticulate and dull, becoming smooth and shining over the majority of the centre of the disc as well as the centre of the scutellum, and irregularly punctate, laterally punctate by 0.52 puncture diameters, punctures becoming sparser medially, separated by up to 34 puncture diameters (
<figureCitation id="71B92A6FAC0443260AA5FA9E8C5D0331" box="[433,568,1365,1392]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="93" pageNumber="94">Figure 59C</figureCitation>
). In contrast,
<taxonomicName id="2E824D69AC04432609C1FA9D8DC9032E" authorityName="Warncke" authorityYear="1975" box="[725,940,1365,1392]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="pileata" subSpecies="canigica">
<emphasis id="DBF6EAF8AC04432609C1FA9D8DC9032E" box="[725,940,1365,1392]" italics="true" pageId="93" pageNumber="94">A. pileata canigica</emphasis>
</taxonomicName>
shows a uniformly shagreened and at most weakly shining scutum and scutellum, and punctation is also more regular, separated by 0.52 puncture diameters, typically by 2 puncture diameters medially, punctation not becoming as scattered as in
<taxonomicName id="2E824D69AC0443260F65FA568A9C03F6" authorityName="WOOD" authorityYear="2024" box="[1137,1273,1437,1463]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC0443260F65FA568A9C03F6" box="[1137,1273,1437,1463]" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC0443260E1DFA568BF503F9" box="[1289,1424,1437,1464]" captionStart="FIGURE 59" captionStartId="91.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@91.[215,1373,181,1919]" captionTargetPageId="91" captionText="FIGURE 59. Andrena (Euandrena) eosphora spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; E. Mesepisternum, lateral view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (Euandrena) pileata ssp. canigica Warncke, 1975 female. D. Scutum, dorsal view; F. Mesepisternum, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209431" httpUri="https://zenodo.org/record/13209431/files/figure.png" pageId="93" pageNumber="94">Figure 59D</figureCitation>
). Given that the two taxa can be found in sympatry in south-eastern
<collectingCountry id="9195767AAC0443260899FA0A8DB8039A" box="[909,989,1473,1499]" name="Turkey" pageId="93" pageNumber="94">Turkey</collectingCountry>
(specifically, the Suvari Halil-Pass and east of Uludere, see “Other material examined” below), without apparent introgression, these are considered to be stable characters.
</paragraph>
<paragraph id="E93D36EAAC0443260BD3F9E78F810125" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
Since the male of
<taxonomicName id="2E824D69AC0443260A85F9E78C000007" authorityName="Warncke" authorityYear="1975" box="[401,613,1580,1606]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="subSpecies" species="pileata" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0443260A85F9E78C000007" box="[401,613,1580,1606]" italics="true" pageId="93" pageNumber="94">A. pileata canigica</emphasis>
</taxonomicName>
(as well as the male of
<emphasis id="DBF6EAF8AC044326087BF9E78A780007" box="[879,1053,1580,1606]" italics="true" pageId="93" pageNumber="94">
<taxonomicName id="2E824D69AC044326087BF9E78DB90007" authorityName="Warncke" authorityYear="1975" box="[879,988,1580,1606]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="pileata">A. pileata</taxonomicName>
<taxonomicNameLabel id="C0C55783AC04432608F6F9E68A780007" box="[994,1053,1581,1606]" pageId="93" pageNumber="94" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
) is unknown, diagnosis against
<taxonomicName id="2E824D69AC0443260E90F9E78E9A002B" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC0443260E90F9E78E9A002B" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
is not possible. However, the male of
<taxonomicName id="2E824D69AC044326098AF99B8D46002B" authorityName="WOOD" authorityYear="2024" box="[670,803,1616,1642]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC044326098AF99B8D46002B" box="[670,803,1616,1642]" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
is similar to the male of
<taxonomicName id="2E824D69AC0443260F39F99B8AC6002B" authorityName="WOOD" authorityYear="2024" box="[1069,1187,1616,1642]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="ridicula" status="spec. nov.">
<emphasis id="DBF6EAF8AC0443260F39F99B8AC6002B" box="[1069,1187,1616,1642]" italics="true" pageId="93" pageNumber="94">A. ridicula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC0443260FBCF99B8B7D002B" box="[1192,1304,1616,1642]" pageId="93" pageNumber="94" rank="species">spec. nov.</taxonomicNameLabel>
, with which it can occur in sympatry, due to the predominantly pale facial pubescence with black hairs restricted to the inner margins of the compound eyes (
<figureCitation id="71B92A6FAC0443260AE8F95C8CE500F0" box="[508,640,1687,1713]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60B</figureCitation>
), sculptured and weakly punctate terga with depressed margins that are lightened brown (
<figureCitation id="71B92A6FAC0443260A4FF9708FBB0094" box="[347,478,1723,1749]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60E</figureCitation>
), and genital capsule with very narrow penis valves (
<figureCitation id="71B92A6FAC0443260F3DF9708ACF0094" box="[1065,1194,1723,1749]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60F</figureCitation>
). The two species can be separated by the length of the head which in
<taxonomicName id="2E824D69AC04432609BAF9148D5000B8" authorityName="WOOD" authorityYear="2024" box="[686,821,1759,1785]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC04432609BAF9148D5000B8" box="[686,821,1759,1785]" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
is relatively broad, 1.2 times wider than long, whereas
<taxonomicName id="2E824D69AC0443260B83F8C88F6B015D" authorityName="WOOD" authorityYear="2024" box="[151,270,1794,1821]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="ridicula">
<emphasis id="DBF6EAF8AC0443260B83F8C88F6B015D" box="[151,270,1794,1821]" italics="true" pageId="93" pageNumber="94">A. ridicula</emphasis>
</taxonomicName>
has the head comparatively elongate, only 1.05 times wider than long, and also by the weaker sculpturing of the scutum of
<taxonomicName id="2E824D69AC0443260A40F8ED8FBE0101" authorityName="WOOD" authorityYear="2024" box="[340,475,1830,1856]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="eosphora">
<emphasis id="DBF6EAF8AC0443260A40F8ED8FBE0101" box="[340,475,1830,1856]" italics="true" pageId="93" pageNumber="94">A. eosphora</emphasis>
</taxonomicName>
, medially smooth and shining (
<figureCitation id="71B92A6FAC044326082FF8ED8DA50101" box="[827,960,1830,1856]" captionStart="FIGURE 60" captionStartId="92.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1622]" captionTargetId="figure-28@92.[151,1436,181,1622]" captionTargetPageId="92" captionText="FIGURE 60. Andrena (Euandrena) eosphora spec. nov. male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209435" httpUri="https://zenodo.org/record/13209435/files/figure.png" pageId="93" pageNumber="94">Figure 60C</figureCitation>
), more or less consistently shagreened and weakly shining in
<taxonomicName id="2E824D69AC0443260A71F8818FB80125" authorityName="WOOD" authorityYear="2024" box="[357,477,1866,1892]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="93" pageNumber="94" phylum="Arthropoda" rank="species" species="ridicula">
<emphasis id="DBF6EAF8AC0443260A71F8818FB80125" box="[357,477,1866,1892]" italics="true" pageId="93" pageNumber="94">A. ridicula</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0443260BD3F8A58F7D01EA" pageId="93" pageNumber="94" type="discussion">
<paragraph id="E93D36EAAC0443260BD3F8A58F7D01EA" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<emphasis id="DBF6EAF8AC0443260BD3F8A58F5E01C9" bold="true" box="[199,315,1902,1928]" pageId="93" pageNumber="94">Remarks.</emphasis>
Specimens from the Suvari-Halil pass were collected from
<taxonomicName id="2E824D69AC04432608C5F8A68A3001C6" box="[977,1109,1901,1927]" class="Magnoliopsida" family="Ranunculaceae" genus="Ranunculus" kingdom="Plantae" order="Ranunculales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8AC04432608C5F8A68A3001C6" box="[977,1109,1901,1927]" italics="true" pageId="93" pageNumber="94">Ranunculus</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69AC0443260F77F8A68B7501C6" box="[1123,1296,1901,1927]" class="Magnoliopsida" family="Ranunculaceae" kingdom="Plantae" order="Ranunculales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="family">Ranunculaceae</taxonomicName>
) and
<taxonomicName id="2E824D69AC0443260E58F8A58BF901C9" box="[1356,1436,1902,1928]" class="Liliopsida" family="Iridaceae" genus="Crocus" kingdom="Plantae" order="Asparagales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8AC0443260E58F8A58BF901C9" box="[1356,1436,1902,1928]" italics="true" pageId="93" pageNumber="94">Crocus</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69AC0443260B8BF85A8F6901EA" box="[159,268,1937,1963]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="93" pageNumber="94" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0443260BD3F87E8F0A01B2" pageId="93" pageNumber="94" type="etymology">
<paragraph id="E93D36EAAC0443260BD3F87E8F0A01B2" blockId="93.[151,1437,151,2035]" pageId="93" pageNumber="94">
<emphasis id="DBF6EAF8AC0443260BD3F87E8F2B018E" bold="true" box="[199,334,1973,1999]" pageId="93" pageNumber="94">Etymology.</emphasis>
From the Greek
<emphasis id="DBF6EAF8AC0443260906F87E8CED018E" box="[530,648,1973,1999]" italics="true" pageId="93" pageNumber="94">eosophora</emphasis>
meaning dawn-bringer, in reference to its easterly distribution. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0743250BD3FF5D8A38045A" pageId="94" pageNumber="95" type="distribution">
<paragraph id="E93D36EAAC0743250BD3FF5D8DBC06F0" blockId="94.[151,1437,150,539]" box="[199,985,150,177]" pageId="94" pageNumber="95">
<emphasis id="DBF6EAF8AC0743250BD3FF5D8F0406F1" bold="true" box="[199,353,150,176]" pageId="94" pageNumber="95">Distribution.</emphasis>
South-eastern
<collectingCountry id="9195767AAC074325091CFF5C8C3D06F0" box="[520,600,151,177]" name="Turkey" pageId="94" pageNumber="95">Turkey</collectingCountry>
(provinces of
<collectingRegion id="2B46F808AC07432509EEFF5C8D5206F0" box="[762,823,151,177]" country="Turkey" name="Bitlis" pageId="94" pageNumber="95">Bitlis</collectingRegion>
and
<collectingRegion id="2B46F808AC0743250879FF5C8DA806F0" box="[877,973,151,177]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
).
</paragraph>
<paragraph id="E93D36EAAC0743250BD3FF718A38045A" blockId="94.[151,1437,150,539]" pageId="94" pageNumber="95">
<materialsCitation id="59EA3CB7AC0743250BD3FF718E9F06B8" collectingDate="1962-06-01" collectorName="M. Schwarz" country="Greece" elevation="1900" location="Chelmos" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Peloponnisos">
<emphasis id="DBF6EAF8AC0743250BD3FF718F970695" bold="true" box="[199,498,186,212]" pageId="94" pageNumber="95">
<typeStatus id="36398848AC0743250BD3FF718F970695" box="[199,498,186,212]" isOtherTypeStatus="true" pageId="94" pageNumber="95">Other material examined</typeStatus>
</emphasis>
(
<taxonomicName id="2E824D69AC0743250915FF708CD10694" authorityName="Warncke" authorityYear="1975" box="[513,692,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="pileata">
<emphasis id="DBF6EAF8AC0743250915FF708CD10694" box="[513,692,187,213]" italics="true" pageId="94" pageNumber="95">Andrena pileata</emphasis>
</taxonomicName>
)
<emphasis id="DBF6EAF8AC07432509D7FF718D260695" bold="true" box="[707,835,186,212]" pageId="94" pageNumber="95">
<collectingCountry id="9195767AAC07432509D7FF718D5B0695" box="[707,830,186,212]" name="Greece" pageId="94" pageNumber="95">GREECE</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC074325085CFF708DCB0694" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC074325085CFF708DCB0694" box="[840,942,187,213]" country="Greece" name="Chelmos" pageId="94" pageNumber="95" stateProvince="Peloponnisos">Chelmos</location>
,
<collectingRegion id="2B46F808AC07432508ADFF708A580694" box="[953,1085,187,213]" country="Greece" name="Peloponnisos" pageId="94" pageNumber="95">Peloponnes</collectingRegion>
,
<quantity id="2E7A9B0FAC0743250F53FF708AF90694" box="[1095,1180,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="94" pageNumber="95" unit="m" value="1900.0">
<elevation id="62AFD1D9AC0743250F53FF708AF90694" box="[1095,1180,187,213]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="94" pageNumber="95" unit="m" value="1900.0">1900 m</elevation>
</quantity>
,
<date id="9D3C102AAC0743250FB3FF708B760694" box="[1191,1299,187,213]" pageId="94" pageNumber="95" value="1962-06-01">
<collectingDate id="8D78E9C2AC0743250FB3FF708B760694" box="[1191,1299,187,213]" pageId="94" pageNumber="95" value="1962-06-01">1.vi.1962</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250E0AFF708B250694" box="[1310,1344,187,213]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250E68FF708E9F06B8" pageId="94" pageNumber="95">M. Schwarz</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0743250A13FF2B8B5B06B8" box="[263,1342,223,251]" collectingDate="1987-05-27" collectingDateMax="1987-05-29" collectingDateMin="1987-05-27" collectionCode="RMNH" collectorName="H. Teunissen" country="Greece" elevation="1700" location="Chelmos" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2">
OÖLM (
<emphasis id="DBF6EAF8AC0743250A7FFF148FB706B8" bold="true" box="[363,466,223,249]" pageId="94" pageNumber="95">
<typeStatus id="36398848AC0743250A7FFF148FB706B8" box="[363,466,223,249]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
</emphasis>
);
<location id="EC5D6031AC0743250AFFFF148C2A06B8" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250AFFFF148C2A06B8" box="[491,591,223,249]" country="Greece" name="Chelmos" pageId="94" pageNumber="95">Chelmos</location>
[Aroania],
<quantity id="2E7A9B0FAC07432509C7FF148D4E06B8" box="[723,811,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="94" pageNumber="95" unit="m" value="1700.0">
<elevation id="62AFD1D9AC07432509C7FF148D4E06B8" box="[723,811,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="94" pageNumber="95" unit="m" value="1700.0">1700 m</elevation>
</quantity>
,
<date id="9D3C102AAC074325082CFF148DB606B8" box="[824,979,223,249]" pageId="94" pageNumber="95" value="1987-05-27" valueMax="1987-05-29" valueMin="1987-05-27">
<collectingDate id="8D78E9C2AC074325082CFF148DB606B8" box="[824,979,223,249]" pageId="94" pageNumber="95" value="1987-05-27" valueMax="1987-05-29" valueMin="1987-05-27">2729.v.1987</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC07432508F4FF148A6106B8" box="[992,1028,223,249]" count="2" pageId="94" pageNumber="95" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250F50FF148ABF06B8" box="[1092,1242,223,249]" pageId="94" pageNumber="95">H. Teunissen</collectorName>
,
<collectionCode id="8F93AE2FAC0743250FF2FF148B5B06B8" box="[1254,1342,223,249]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="94" pageNumber="95" type="Museum">RMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250E5EFF148F320700" collectingDate="2024-04-09" collectionCode="TJWC" collectorName="T. J. Wood" country="Greece" elevation="1750" location="Mt. Parnassos" pageId="94" pageNumber="95" specimenCount="6" specimenCount-female="6" stateProvince="Sterea Ellada">
<collectingRegion id="2B46F808AC0743250E5EFF148E8C075C" country="Greece" name="Sterea Ellada" pageId="94" pageNumber="95">Central Greece</collectingRegion>
,
<location id="EC5D6031AC0743250BE3FEC88FFC075C" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250BE3FEC88FFC075C" box="[247,409,259,285]" country="Greece" name="Mt. Parnassos" pageId="94" pageNumber="95" stateProvince="Sterea Ellada">Mt. Parnassos</location>
, immediately NW of ski centre,
<quantity id="2E7A9B0FAC0743250806FEC98D0E075C" box="[786,875,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="94" pageNumber="95" unit="m" value="1750.0">
<elevation id="62AFD1D9AC0743250806FEC98D0E075C" box="[786,875,258,285]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="94" pageNumber="95" unit="m" value="1750.0">1750 m</elevation>
</quantity>
,
<date id="9D3C102AAC074325086DFEC88D86075C" box="[889,995,259,285]" pageId="94" pageNumber="95" value="2024-04-09">
<collectingDate id="8D78E9C2AC074325086DFEC88D86075C" box="[889,995,259,285]" pageId="94" pageNumber="95" value="2024-04-09">9.iv.2024</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC07432508E5FEC88A73075C" box="[1009,1046,259,285]" count="6" pageId="94" pageNumber="95" type="female">6♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250F43FEC88AA9075C" box="[1111,1228,259,285]" pageId="94" pageNumber="95">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC0743250FCDFEC88B43075C" box="[1241,1318,259,285]" pageId="94" pageNumber="95">TJWC</collectionCode>
; (
<taxonomicName id="2E824D69AC0743250E29FEC88F280701" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="pileata" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0743250E29FEC88F280701" italics="true" pageId="94" pageNumber="95">Andrena pileata canigica</emphasis>
</taxonomicName>
)
</materialsCitation>
<materialsCitation id="59EA3CB7AC0743250A4BFEEC8AE00700" box="[351,1157,294,321]" collectingDate="1959-05-15" collectorName="Schweiger" country="Turkey" location="Ulu Dag" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Bursa">
<emphasis id="DBF6EAF8AC0743250A4BFEEC8FBB0700" bold="true" box="[351,478,295,321]" pageId="94" pageNumber="95">
<collectingCountry id="9195767AAC0743250A4BFEEC8FBC0700" box="[351,473,295,321]" name="Turkey" pageId="94" pageNumber="95">TURKEY</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC0743250AF2FEEC8C2D0700" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250AF2FEEC8C2D0700" box="[486,584,295,321]" country="Turkey" name="Ulu Dag" pageId="94" pageNumber="95" stateProvince="Bursa">Ulu Dag</location>
[Uludağ] b.
<collectingRegion id="2B46F808AC07432509CFFEEC8D7A0700" box="[731,799,295,321]" country="Turkey" name="Bursa" pageId="94" pageNumber="95">Bursa</collectingRegion>
,
<date id="9D3C102AAC074325083FFEED8DF90700" box="[811,924,294,321]" pageId="94" pageNumber="95" value="1959-05-15">
<collectingDate id="8D78E9C2AC074325083FFEED8DF90700" box="[811,924,294,321]" pageId="94" pageNumber="95" value="1959-05-15">15.v.1959</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC07432508BDFEEC8DA90700" box="[937,972,295,321]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250F19FEEC8AE00700" box="[1037,1157,295,321]" pageId="94" pageNumber="95">Schweiger</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0743250F85FEE38DB10726" collectingDate="1994-04-28" collectionCode="OOLM" collectorName="K. Denes" country="Turkey" location="Bolu" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Bolu">
OÖLM (
<emphasis id="DBF6EAF8AC0743250FE1FEEC8B390700" bold="true" box="[1269,1372,295,321]" pageId="94" pageNumber="95">
<typeStatus id="36398848AC0743250FE1FEEC8B390700" box="[1269,1372,295,321]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
</emphasis>
of
<taxonomicName id="2E824D69AC0743250E90FEEC8F200725" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="allosa" subSpecies="canigica">
<emphasis id="DBF6EAF8AC0743250E90FEEC8F200725" italics="true" pageId="94" pageNumber="95">A. allosa canigica</emphasis>
</taxonomicName>
);
<collectingRegion id="2B46F808AC0743250A74FE808FF30724" box="[352,406,331,357]" country="Turkey" name="Bolu" pageId="94" pageNumber="95">Bolu</collectingRegion>
lake env.,
<date id="9D3C102AAC0743250902FE808CEB0724" box="[534,654,331,357]" pageId="94" pageNumber="95" value="1994-04-28">
<collectingDate id="8D78E9C2AC0743250902FE808CEB0724" box="[534,654,331,357]" pageId="94" pageNumber="95" value="1994-04-28">28.iv.1994</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250988FE808CDA0724" box="[668,703,331,357]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250817FE808D150724" box="[771,880,331,357]" pageId="94" pageNumber="95">K. Deneš</collectorName>
,
<collectionCode id="8F93AE2FAC074325086AFE878DB10726" box="[894,980,332,359]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC07432508F5FE808F8607CA" collectingDate="1977-06-06" collectionCode="OOLM" collectorName="K. Warncke" country="Turkey" location="E of Uludere" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Hakkari">
<collectingRegion id="2B46F808AC07432508F5FE808A5B0724" box="[993,1086,331,357]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
, pass
<location id="EC5D6031AC0743250F92FE808B7B0724" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250F92FE808B7B0724" box="[1158,1310,331,357]" country="Turkey" name="E of Uludere" pageId="94" pageNumber="95" stateProvince="Hakkari">E of Uludere</location>
,
<date id="9D3C102AAC0743250E3FFE808BFD0724" box="[1323,1432,331,357]" pageId="94" pageNumber="95" value="1977-06-06">
<collectingDate id="8D78E9C2AC0743250E3FFE808BFD0724" box="[1323,1432,331,357]" pageId="94" pageNumber="95" value="1977-06-06">6.vi.1977</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250B83FEA48EDF07C8" box="[151,186,367,393]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250BEDFEA48FE707C8" box="[249,386,367,393]" pageId="94" pageNumber="95">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC0743250A99FEBB8F8607CA" box="[397,483,368,395]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250AFBFEA48B3007CA" box="[495,1365,366,395]" collectingDate="1980-06-02" collectionCode="OOLM" collectorName="M. Schwarz" country="Turkey" elevation="2500" location="Suvari Halil-Pass" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" stateProvince="Hakkari">
<collectingRegion id="2B46F808AC0743250AFBFEA48C2907C8" box="[495,588,367,393]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
,
<location id="EC5D6031AC074325094CFEA48D7B07C8" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC074325094CFEA48D7B07C8" box="[600,798,367,393]" country="Turkey" name="Suvari Halil-Pass" pageId="94" pageNumber="95" stateProvince="Hakkari">Suvari Halil-Pass</location>
,
<quantity id="2E7A9B0FAC074325083EFEA58DE407C8" box="[810,897,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="94" pageNumber="95" unit="m" value="2500.0">
<elevation id="62AFD1D9AC074325083EFEA58DE407C8" box="[810,897,366,393]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="94" pageNumber="95" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="9D3C102AAC074325089AFEA48D9F07C8" box="[910,1018,367,393]" pageId="94" pageNumber="95" value="1980-06-02">
<collectingDate id="8D78E9C2AC074325089AFEA48D9F07C8" box="[910,1018,367,393]" pageId="94" pageNumber="95" value="1980-06-02">2.vi.1980</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250F12FEA48A4F07C8" box="[1030,1066,367,393]" count="2" pageId="94" pageNumber="95" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250F7CFEA48A9607C8" box="[1128,1267,367,393]" pageId="94" pageNumber="95">M. Schwarz</collectorName>
,
<collectionCode id="8F93AE2FAC0743250FEBFEBB8B3007CA" box="[1279,1365,368,395]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250E74FEA48DB507EE" collectingDate="1983-05-29" collectionCode="OOLM" collectorName="K. Warncke" country="Turkey" elevation="2100" location="20 km W of Sarikamis" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Kars">
<collectingRegion id="2B46F808AC0743250E74FEA48BFD07C8" box="[1376,1432,367,393]" country="Turkey" name="Kars" pageId="94" pageNumber="95">Kars</collectingRegion>
,
<location id="EC5D6031AC0743250B83FE588FFC07EC" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250B83FE588FFC07EC" box="[151,409,403,429]" country="Turkey" name="20 km W of Sarikamis" pageId="94" pageNumber="95" stateProvince="Kars">
<quantity id="2E7A9B0FAC0743250B83FE588EBA07EC" box="[151,223,403,429]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="94" pageNumber="95" unit="km" value="20.0">20 km</quantity>
W of Sarıkamış
</location>
,
<quantity id="2E7A9B0FAC0743250AB0FE588F9907EC" box="[420,508,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="94" pageNumber="95" unit="m" value="2100.0">
<elevation id="62AFD1D9AC0743250AB0FE588F9907EC" box="[420,508,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="94" pageNumber="95" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="9D3C102AAC074325091CFE588C1C07EC" box="[520,633,403,429]" pageId="94" pageNumber="95" value="1983-05-29">
<collectingDate id="8D78E9C2AC074325091CFE588C1C07EC" box="[520,633,403,429]" pageId="94" pageNumber="95" value="1983-05-29">29.v.1983</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250991FE588CCD07EC" box="[645,680,403,429]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC07432509F3FE588D0A07EC" box="[743,879,403,429]" pageId="94" pageNumber="95">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC074325086EFE5F8DB507EE" box="[890,976,404,431]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC07432508C8FE588CEB0792" collectingDate="1980-06-03" collectionCode="OOLM" collectorName="M. Schwarz" country="Turkey" elevation="1412" location="Hakkari" pageId="94" pageNumber="95" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Hakkari">
<collectingRegion id="2B46F808AC07432508C8FE588A5C07EC" box="[988,1081,403,429]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
, Tanin-Tanin-Pass,
<quantity id="2E7A9B0FAC0743250E02FE588BF207EC" box="[1302,1431,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.4115" metricValueMax="2.8" metricValueMin="0.023" pageId="94" pageNumber="95" unit="m" value="1411.5" valueMax="2800.0" valueMin="23.0">
<elevation id="62AFD1D9AC0743250E02FE588BF207EC" box="[1302,1431,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.4115" metricValueMax="2.8" metricValueMin="0.023" pageId="94" pageNumber="95" unit="m" value="1411.5" valueMax="2800.0" valueMin="23.0">232800 m</elevation>
</quantity>
,
<date id="9D3C102AAC0743250B83FE7C8F660790" box="[151,259,439,465]" pageId="94" pageNumber="95" value="1980-06-03">
<collectingDate id="8D78E9C2AC0743250B83FE7C8F660790" box="[151,259,439,465]" pageId="94" pageNumber="95" value="1980-06-03">3.vi.1980</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250A1BFE7C8F510790" box="[271,308,439,465]" count="1" pageId="94" pageNumber="95" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC0743250A55FE7C8F000790" box="[321,357,439,465]" count="4" pageId="94" pageNumber="95" type="female">4♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250AB6FE7C8C480790" box="[418,557,439,465]" pageId="94" pageNumber="95">M. Schwarz</collectorName>
,
<collectionCode id="8F93AE2FAC074325092CFE738CEB0792" box="[568,654,440,467]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC074325098EFE7C8E8807B6" collectingDate="1980-06-02" collectionCode="OOLM" collectorName="K. Warncke" country="Turkey" elevation="2500" location="Hakkari" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Hakkari">
<collectingRegion id="2B46F808AC074325098EFE7C8C920790" box="[666,759,439,465]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
, Tanin-Tanin-Pass,
<quantity id="2E7A9B0FAC07432508C7FE7D8A4F0790" box="[979,1066,438,465]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="94" pageNumber="95" unit="m" value="2500.0">
<elevation id="62AFD1D9AC07432508C7FE7D8A4F0790" box="[979,1066,438,465]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="94" pageNumber="95" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="9D3C102AAC0743250F23FE7C8AC60790" box="[1079,1187,439,465]" pageId="94" pageNumber="95" value="1980-06-02">
<collectingDate id="8D78E9C2AC0743250F23FE7C8AC60790" box="[1079,1187,439,465]" pageId="94" pageNumber="95" value="1980-06-02">2.vi.1980</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250FBBFE7C8AB70790" box="[1199,1234,439,465]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250E04FE7C8BFD0790" box="[1296,1432,439,465]" pageId="94" pageNumber="95">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC0743250B83FE178E8807B6" box="[151,237,476,503]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250BECFE108DED07B6" box="[248,904,474,503]" collectingDate="1977-06-05" collectionCode="OOLM" collectorName="K. Warncke" country="Turkey" location="Uludere" pageId="94" pageNumber="95" specimenCount="1" specimenCount-female="1" stateProvince="Hakkari">
<location id="EC5D6031AC0743250BECFE108F3007B4" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250BECFE108F3007B4" box="[248,341,475,501]" country="Turkey" name="Uludere" pageId="94" pageNumber="95" stateProvince="Hakkari">Uludere</location>
,
<collectingRegion id="2B46F808AC0743250A74FE108FD807B4" box="[352,445,475,501]" country="Turkey" name="Hakkari" pageId="94" pageNumber="95">Hakkâri</collectingRegion>
,
<date id="9D3C102AAC0743250ADCFE118C5607B4" box="[456,563,474,501]" pageId="94" pageNumber="95" value="1977-06-05">
<collectingDate id="8D78E9C2AC0743250ADCFE118C5607B4" box="[456,563,474,501]" pageId="94" pageNumber="95" value="1977-06-05">5.vi.1977</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC074325092BFE108C0707B4" box="[575,610,475,501]" count="1" pageId="94" pageNumber="95" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC07432509B4FE108D4207B4" box="[672,807,475,501]" pageId="94" pageNumber="95">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC0743250826FE178DED07B6" box="[818,904,476,503]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250886FE108F84045A" collectingDate="1977-05-31" collectionCode="OOLM" collectorName="K. Warncke" country="Turkey" location="Sarikamis" pageId="94" pageNumber="95" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kars">
W
<location id="EC5D6031AC07432508A7FE108A4607B4" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC07432508A7FE108A4607B4" box="[947,1059,475,501]" country="Turkey" name="Sarikamis" pageId="94" pageNumber="95" stateProvince="Kars">Sarikamis</location>
[Sarıkamış]/
<collectingRegion id="2B46F808AC0743250FA1FE108A8907B4" box="[1205,1260,475,501]" country="Turkey" name="Kars" pageId="94" pageNumber="95">Kars</collectingRegion>
,
<date id="9D3C102AAC0743250FE2FE108B0307B4" box="[1270,1382,475,501]" pageId="94" pageNumber="95" value="1977-05-31">
<collectingDate id="8D78E9C2AC0743250FE2FE108B0307B4" box="[1270,1382,475,501]" pageId="94" pageNumber="95" value="1977-05-31">31.v.1977</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250E66FE108BFD07B4" box="[1394,1432,475,501]" count="2" pageId="94" pageNumber="95" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC0743250B83FE348EDE0458" box="[151,187,511,537]" count="3" pageId="94" pageNumber="95" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250BECFE348FE50458" box="[248,384,511,537]" pageId="94" pageNumber="95">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC0743250A9FFDCB8F84045A" box="[395,481,512,539]" pageId="94" pageNumber="95">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0743250AFFFE348D9D0458" box="[491,1016,510,537]" collectingDate="1975-05-24" collectorName="K. Warncke" country="Turkey" location="Yenikoy" pageId="94" pageNumber="95" specimenCount="2" specimenCount-female="2" stateProvince="Kars">
<location id="EC5D6031AC0743250AFFFE348C290458" LSID="urn:lsid:plazi:treatment:612B87FCAC0343250B83FD1D8A38045A:EC5D6031AC0743250AFFFE348C290458" box="[491,588,511,537]" country="Turkey" name="Yenikoy" pageId="94" pageNumber="95" stateProvince="Kars">Yeniköy</location>
/
<collectingRegion id="2B46F808AC0743250940FE348CED0458" box="[596,648,511,537]" country="Turkey" name="Kars" pageId="94" pageNumber="95">Kars</collectingRegion>
,
<date id="9D3C102AAC0743250987FE358D610458" box="[659,772,510,537]" pageId="94" pageNumber="95" value="1975-05-24">
<collectingDate id="8D78E9C2AC0743250987FE358D610458" box="[659,772,510,537]" pageId="94" pageNumber="95" value="1975-05-24">24.v.1975</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0743250804FE348D510458" box="[784,820,511,537]" count="2" pageId="94" pageNumber="95" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC0743250865FE348D9D0458" box="[881,1016,511,537]" pageId="94" pageNumber="95">K. Warncke</collectorName>
</materialsCitation>
, OÖLM.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,415 @@
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<taxonomicName id="2E824D69AC0A43280B83FBF58C500218" authorityName="WOOD" authorityYear="2024" box="[151,565,1086,1113]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="83" pageNumber="84" phylum="Arthropoda" rank="species" species="karahisara" status="sp. nov." subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC0A43280B83FBF58F640218" bold="true" box="[151,257,1086,1113]" italics="true" pageId="83" pageNumber="84">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC0A43280A06FBF58FC50218" bold="true" box="[274,416,1086,1113]" italics="true" pageId="83" pageNumber="84">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC0A43280AA5FBF58C500218" bold="true" box="[433,565,1086,1113]" italics="true" pageId="83" pageNumber="84">karahisara</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="A1986561AC0A43280B83FBA88FC7023C" box="[151,418,1122,1149]" pageId="83" pageNumber="84" type="description">
<paragraph id="E93D36EAAC0A43280B83FBA88FC7023C" blockId="83.[151,692,1086,1149]" box="[151,418,1122,1149]" pageId="83" pageNumber="84">
<figureCitation id="71B92A6FAC0A43280B83FBA88F26023C" box="[151,323,1122,1149]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figures 55AF</figureCitation>
;
<figureCitation id="71B92A6FAC0A43280A5BFBA98FFB023C" box="[335,414,1122,1149]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="83" pageNumber="84">56AF</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0A43280B83FB618D0B0372" pageId="83" pageNumber="84" type="materials_examined">
<paragraph id="E93D36EAAC0A43280B83FB618B2A0286" blockId="83.[151,1437,1194,2013]" box="[151,1359,1194,1223]" pageId="83" pageNumber="84">
<materialsCitation id="59EA3CB7AC0A43280B83FB618B2A0286" box="[151,1359,1194,1223]" collectingDate="1972-05-23" collectorName="K. Warncke" country="Turkey" location="Urgup" pageId="83" pageNumber="84" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC0A43280B83FB618FA10284" bold="true" box="[151,452,1194,1221]" pageId="83" pageNumber="84">
<typeStatus id="36398848AC0A43280B83FB618F5F0285" box="[151,314,1194,1220]" pageId="83" pageNumber="84" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC0A43280A52FB608FA50284" box="[326,448,1195,1221]" name="Turkey" pageId="83" pageNumber="84">TURKEY</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC0A43280ADFFB678C770286" LSID="urn:lsid:plazi:treatment:612B87FCAC0A432E0B83FBF58D310145:EC5D6031AC0A43280ADFFB678C770286" box="[459,530,1196,1223]" country="Turkey" name="Urgup" pageId="83" pageNumber="84">Ürgüp</location>
[inferred 38.6419
<superScript id="1EF79BA2AC0A432809C9FB608C990284" attach="right" box="[733,764,1195,1221]" fontSize="6" pageId="83" pageNumber="84">oN</superScript>
, 34.8933
<superScript id="1EF79BA2AC0A43280877FB608DE50284" attach="right" box="[867,896,1195,1221]" fontSize="6" pageId="83" pageNumber="84">oE</superScript>
],
<date id="9D3C102AAC0A43280887FB608A610284" box="[915,1028,1195,1221]" pageId="83" pageNumber="84" value="1972-05-23">
<collectingDate id="8D78E9C2AC0A43280887FB608A610284" box="[915,1028,1195,1221]" pageId="83" pageNumber="84" value="1972-05-23">23.v.1972</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0A43280F1BFB608A430284" box="[1039,1062,1195,1221]" count="1" pageId="83" pageNumber="84" type="male"></specimenCount>
, leg.
<collectorName id="4477533CAC0A43280F77FB608A8F0284" box="[1123,1258,1195,1221]" pageId="83" pageNumber="84">K. Warncke</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC0A43280BD3FB058D0B0372" blockId="83.[151,1437,1194,2013]" pageId="83" pageNumber="84">
<materialsCitation id="59EA3CB7AC0A43280BD3FB058D0F0372" collectingDate="1975-05-10" collectingDateMax="1975-05-28" collectingDateMin="1975-05-10" collectionCode="TJWC" collectorName="K. Warncke &amp; Yesilhisar" country="Turkey" location="Altinekin" pageId="83" pageNumber="84" specimenCount="29" specimenCount-female="15" specimenCount-male="14" stateProvince="Konya" typeStatus="paratype">
<emphasis id="DBF6EAF8AC0A43280BD3FB058C6402A8" bold="true" box="[199,513,1230,1257]" pageId="83" pageNumber="84">
<typeStatus id="36398848AC0A43280BD3FB058F0A02A9" box="[199,367,1230,1256]" pageId="83" pageNumber="84" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="9195767AAC0A43280A97FB048F9802A8" box="[387,509,1231,1257]" name="Turkey" pageId="83" pageNumber="84">TURKEY</collectingCountry>
:
</emphasis>
same information as
<typeStatus id="36398848AC0A43280806FB048D1202A8" box="[786,887,1231,1257]" pageId="83" pageNumber="84" type="holotype">holotype</typeStatus>
,
<specimenCount id="FF84FD63AC0A4328089DFB048DD902A8" box="[905,956,1231,1257]" count="10" pageId="83" pageNumber="84" type="male">10♂</specimenCount>
,
<specimenCount id="FF84FD63AC0A432808C4FB048A6702A8" box="[976,1026,1231,1257]" count="13" pageId="83" pageNumber="84" type="female">13♀</specimenCount>
, leg.
<collectorName id="4477533CAC0A43280F5BFB048ABA02A8" box="[1103,1247,1231,1257]" pageId="83" pageNumber="84">K. Warncke</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC0A43280E5AFB1B8BFD02AA" box="[1358,1432,1232,1259]" pageId="83" pageNumber="84">TJWC</collectionCode>
;
<location id="EC5D6031AC0A43280B83FB388F61034C" LSID="urn:lsid:plazi:treatment:612B87FCAC0A432E0B83FBF58D310145:EC5D6031AC0A43280B83FB388F61034C" box="[151,260,1267,1293]" country="Turkey" name="Altinekin" pageId="83" pageNumber="84" stateProvince="Konya">Altinekin</location>
,
<collectingRegion id="2B46F808AC0A43280A1AFB388F3E034C" box="[270,347,1267,1293]" country="Turkey" name="Konya" pageId="83" pageNumber="84">Konya</collectingRegion>
,
<date id="9D3C102AAC0A43280A71FB398FB3034C" box="[357,470,1266,1293]" pageId="83" pageNumber="84" value="1975-05-28">
<collectingDate id="8D78E9C2AC0A43280A71FB398FB3034C" box="[357,470,1266,1293]" pageId="83" pageNumber="84" value="1975-05-28">28.v.1975</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0A43280AF4FB388C60034C" box="[480,517,1267,1293]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC0A43280904FB388C56034C" box="[528,563,1267,1293]" count="1" pageId="83" pageNumber="84" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0A4328097BFB388C90034C" box="[623,757,1267,1293]" pageId="83" pageNumber="84">K. Warncke</collectorName>
, OÖLM;
<collectorName id="4477533CAC0A4328084AFB388DAA034C" box="[862,975,1267,1293]" pageId="83" pageNumber="84">Yeşilhisar</collectorName>
,
<date id="9D3C102AAC0A432808CCFB398A2C034C" box="[984,1097,1266,1293]" pageId="83" pageNumber="84" value="1975-05-10">
<collectingDate id="8D78E9C2AC0A432808CCFB398A2C034C" box="[984,1097,1266,1293]" pageId="83" pageNumber="84" value="1975-05-10">10.v.1975</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0A43280F47FB388A1D034C" box="[1107,1144,1267,1293]" count="1" pageId="83" pageNumber="84" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0A43280FA6FB388B5D034C" box="[1202,1336,1267,1293]" pageId="83" pageNumber="84">K. Warncke</collectorName>
, OÖLM;
<collectorName id="4477533CAC0A43280B83FADC8F6C0370" box="[151,265,1303,1329]" pageId="83" pageNumber="84">Yeşilhisar</collectorName>
/
<collectingRegion id="2B46F808AC0A43280A05FADC8F020370" box="[273,359,1303,1329]" country="Turkey" name="Kayseri" pageId="83" pageNumber="84">Kayseri</collectingRegion>
,
<date id="9D3C102AAC0A43280A67FADD8F860370" box="[371,483,1302,1329]" pageId="83" pageNumber="84" value="1975-05-25">
<collectingDate id="8D78E9C2AC0A43280A67FADD8F860370" box="[371,483,1302,1329]" pageId="83" pageNumber="84" value="1975-05-25">25.v.1975</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0A43280AFBFADC8C710370" box="[495,532,1303,1329]" count="2" pageId="83" pageNumber="84" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC0A43280934FADC8C260370" box="[544,579,1303,1329]" count="1" pageId="83" pageNumber="84" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0A43280995FADC8D6C0370" box="[641,777,1303,1329]" pageId="83" pageNumber="84">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0A432E0BD3FAF18CEA04E9" lastPageId="85" lastPageNumber="86" pageId="83" pageNumber="84" type="description">
<paragraph id="E93D36EAAC0A43280BD3FAF18FF00034" blockId="83.[151,1437,1194,2013]" pageId="83" pageNumber="84">
<emphasis id="DBF6EAF8AC0A43280BD3FAF18FA60314" bold="true" box="[199,451,1338,1365]" pageId="83" pageNumber="84">Description: Female:</emphasis>
Body length: 4.5
<quantity id="2E7A9B0FAC0A43280983FAF18CB20314" box="[663,727,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="83" pageNumber="84" unit="mm" value="5.0">5 mm</quantity>
(
<figureCitation id="71B92A6FAC0A432809F1FAF08D080314" box="[741,877,1338,1365]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figure 55A</figureCitation>
).
<emphasis id="DBF6EAF8AC0A43280894FAF08DA30314" box="[896,966,1339,1365]" italics="true" pageId="83" pageNumber="84">Head:</emphasis>
Dark, 1.15 times wider than long (
<figureCitation id="71B92A6FAC0A43280E45FAF08EAF0338" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figure 55B</figureCitation>
). Clypeus weakly domed, irregularly punctate, punctures separated by 0.53 puncture diameters, underlying surface with strong uniform granular shagreen, weakly shining. Process of labrum narrowly trapezoidal, slightly wider than long, apical margin truncate, surface shining. Gena equalling width of compound eye; ocelloccipital distance ⅓ diameter of lateral ocellus. Foveae dorsally occupying ½ space between compound eye and lateral ocellus, ventrally narrowing, narrower than width of flagellum, separated from inner margin of compound eye by distance equal to its own ventral width; foveae filled with whitish hairs. Face, gena, vertex, and scape covered with short whitish hairs. Antennae basally dark, A412 ventrally lightened by presence of orange scales; A3 exceeding A4, subequal to A4+5.
</paragraph>
<paragraph id="E93D36EAAC0A43280BD3F9B48AF9019C" blockId="83.[151,1437,1194,2013]" pageId="83" pageNumber="84">
<emphasis id="DBF6EAF8AC0A43280BD3F9B48F2D00D9" box="[199,328,1663,1688]" italics="true" pageId="83" pageNumber="84">Mesosoma:</emphasis>
Scutum and scutellum with strong granular shagreen, weakly shining, irregularly punctate with small punctures, punctures separated by 0.54 puncture diameters (
<figureCitation id="71B92A6FAC0A43280869F9688A6400FC" box="[893,1025,1698,1725]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figure 55C</figureCitation>
). Pronotum rounded. Mesepisternum and dorsolateral parts of propodeum with fine granular shagreen, shining. Propodeal triangle covered with slightly coarser granular shagreen, joining and merging with short raised rugae basally, surface weakly shining; propodeal triangle thus defined by change in surface sculpture (
<figureCitation id="71B92A6FAC0A432809EDF8C48DE40168" box="[761,897,1806,1833]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figure 55D</figureCitation>
). Mesepisternum with moderately long whitish hairs, becoming shorter and sub-squamous on scutum and scutellum. Propodeal corbicula incomplete, dorsal fringe composed of long whitish plumose hairs, internal surface with scattered simple white hairs. Legs basally dark, tarsi and hind tibiae lightened orange, pubescence whitish to light brownish. Flocculus incomplete, composed of white plumose hairs; femoral and tibial scopae composed of white simple hairs. Hind tarsal claws without inner tooth. Wings hyaline, stigma and venation light orange, nervulus strongly antefurcal (
<figureCitation id="71B92A6FAC0A43280F18F8088AEA019C" box="[1036,1167,1986,2013]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="83" pageNumber="84">Figure 55E</figureCitation>
).
</paragraph>
<caption id="BDFD6662AC0D432F0B83F9A58A3800EB" ID-DOI="http://doi.org/10.5281/zenodo.13209423" ID-Zenodo-Dep="13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="84" pageNumber="85" startId="84.[151,250,1646,1670]" targetBox="[151,1436,181,1623]" targetPageId="84" targetType="figure">
<paragraph id="E93D36EAAC0D432F0B83F9A58A3800EB" blockId="84.[151,1436,1646,1707]" pageId="84" pageNumber="85">
<emphasis id="DBF6EAF8AC0D432F0B83F9A58F4500C6" bold="true" box="[151,288,1646,1671]" pageId="84" pageNumber="85">FIGURE 55.</emphasis>
<taxonomicName id="2E824D69AC0D432F0A33F9A58CE500C7" authorityName="WOOD" authorityYear="2024" box="[295,640,1646,1671]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="84" pageNumber="85" phylum="Arthropoda" rank="species" species="karahisara" status="sp. nov." subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC0D432F0A33F9A58F1800C7" box="[295,381,1646,1670]" italics="true" pageId="84" pageNumber="85">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC0D432F0A98F9A58C6400C7" box="[396,513,1646,1670]" italics="true" pageId="84" pageNumber="85">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC0D432F0905F9A58CE500C7" box="[529,640,1646,1670]" italics="true" pageId="84" pageNumber="85">karahisara</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC0D432F0993F9A48C8B00C6" bold="true" box="[647,750,1647,1671]" pageId="84" pageNumber="85">
<taxonomicNameLabel id="C0C55783AC0D432F0993F9A48C8B00C6" box="[647,750,1647,1671]" pageId="84" pageNumber="85" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC0D432F0BD3F9358A8B01E6" blockId="84.[151,1436,1789,1959]" pageId="84" pageNumber="85">
<emphasis id="DBF6EAF8AC0D432F0BD3F9358F200156" box="[199,325,1790,1815]" italics="true" pageId="84" pageNumber="85">Metasoma:</emphasis>
Tergal discs dark, tergal margins lightened hyaline-yellow (
<figureCitation id="71B92A6FAC0D432F08E7F9368A120159" box="[1011,1143,1789,1816]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="84" pageNumber="85">Figure 55F</figureCitation>
). Tergal discs with strong granular microreticulation, sculpture strongest basally, dull, becoming weaker apically, here weakly shining; terga impunctate. Terga laterally with sparse erect white hairs, T24 apically with weak hairbands, widely separated on T23, complete on T4, not obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate golden-white. Pygidial plate narrow, rounded triangular, surface flattened, obscurely punctate, dull.
</paragraph>
<paragraph id="E93D36EAAC0D432E0BD3F81E8C6C07C9" blockId="84.[199,1436,2005,2032]" lastBlockId="85.[151,1437,150,1797]" lastPageId="85" lastPageNumber="86" pageId="84" pageNumber="85">
<emphasis id="DBF6EAF8AC0D432F0BD3F81E8F6801AE" bold="true" box="[199,269,2005,2031]" pageId="84" pageNumber="85">Male:</emphasis>
Body length: 4.5
<quantity id="2E7A9B0FAC0D432F0AF7F81E8C4101AE" box="[483,548,2005,2032]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="84" pageNumber="85" unit="mm" value="5.0">5 mm</quantity>
(
<figureCitation id="71B92A6FAC0D432F0927F81E8CDE01B1" box="[563,699,2005,2032]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="84" pageNumber="85">Figure 56A</figureCitation>
).
<emphasis id="DBF6EAF8AC0D432F09C4F81E8D7001AE" box="[720,789,2005,2031]" italics="true" pageId="84" pageNumber="85">Head:</emphasis>
Dark, 1.15 times wider than long (
<figureCitation id="71B92A6FAC0D432F0FB2F81E8B4E01B1" box="[1190,1323,2005,2032]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="84" pageNumber="85">Figure 56B</figureCitation>
). Clypeus almost entirely yellow-marked, markings not extending to absolute edge of clypeus, medially with two small black dots; surface weakly domed, irregularly punctate, punctures separated by 0.53 puncture diameters, underlying surface shagreened, weakly shining. Process of labrum rounded trapezoidal, 2 times wider than long, anterior margin emarginate. Gena subequal to width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Face, gena, vertex, and scale with white hairs, some exceeding length of scape. Antennae basally dark, A413 ventrally lightened by presence of orange scales; A3 exceeding A4, shorter than A4+5, A4 subquadrate, A513 slightly longer than wide.
</paragraph>
<paragraph id="E93D36EAAC0C432E0BD3FE588D060791" blockId="85.[151,1437,150,1797]" pageId="85" pageNumber="86">
<emphasis id="DBF6EAF8AC0C432E0BD3FE588F2D07ED" box="[199,328,403,428]" italics="true" pageId="85" pageNumber="86">Mesosoma:</emphasis>
Mesosoma structurally as in female (
<figureCitation id="71B92A6FAC0C432E09E0FE598DC207EC" box="[756,935,402,429]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figures 56CD</figureCitation>
), covered with white hairs, hind tibiae dark, tarsi lightened orange, hind tarsal claws with strong inner tooth.
</paragraph>
<paragraph id="E93D36EAAC0C432E0BD3FE108CEA04E9" blockId="85.[151,1437,150,1797]" pageId="85" pageNumber="86">
<emphasis id="DBF6EAF8AC0C432E0BD3FE108F2007B5" box="[199,325,475,500]" italics="true" pageId="85" pageNumber="86">Metasoma:</emphasis>
Terga structurally as in female, though tergal margins slightly more distinctly depressed (
<figureCitation id="71B92A6FAC0C432E0E45FE118EAC0458" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56E</figureCitation>
). T67 with short whitish hairs overlying pygidial plate of T7. S8 short, apically broadened with truncate apex, ventrally covered by short whitish hairs. Genital capsule elongate, 1.5 times longer than wide, gonocoxae produced into long apically rounded teeth (
<figureCitation id="71B92A6FAC0C432E0919FD8D8CEA0420" box="[525,655,582,609]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56F</figureCitation>
). Gonostyli slightly broadened apically, spatulate, inner margins with clear bump on inner margins medially. Penis valves relatively narrow, basally slightly broadened, occupying ⅓ space between gonostyli, narrowing apically.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0C432E0BD3FD798F6100D9" pageId="85" pageNumber="86" type="diagnosis">
<paragraph id="E93D36EAAC0C432E0BD3FD798A910219" blockId="85.[151,1437,150,1797]" pageId="85" pageNumber="86">
<emphasis id="DBF6EAF8AC0C432E0BD3FD798F27048D" bold="true" box="[199,322,690,716]" pageId="85" pageNumber="86">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC0C432E0A58FD798C55048D" authorityName="WOOD" authorityYear="2024" box="[332,560,690,716]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0A58FD798C55048D" box="[332,560,690,716]" italics="true" pageId="85" pageNumber="86">Andrena karahisara</emphasis>
</taxonomicName>
can be recognised as an
<taxonomicName id="2E824D69AC0C432E084FFD798DB8048D" authorityName="Warncke" authorityYear="1968" box="[859,989,690,716]" pageId="85" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC0C432E084FFD798DB8048D" box="[859,989,690,716]" italics="true" pageId="85" pageNumber="86">Aciandrena</emphasis>
</taxonomicName>
due to the small body size, propodeal triangle not defined by lateral carinae (
<figureCitation id="71B92A6FAC0C432E0948FD1D8C8004B0" box="[604,741,726,753]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="85" pageNumber="86">Figure 55D</figureCitation>
), surface covered by fine granular shagreen, with weak and obscure rugae basally, facial foveae ventrally narrower than dorsally, supraclypeal and lower paraocular areas free of striations (
<figureCitation id="71B92A6FAC0C432E0A25FCD58FD20578" box="[305,439,798,825]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="85" pageNumber="86">Figure 55B</figureCitation>
), yellow male clypeus (
<figureCitation id="71B92A6FAC0C432E09D8FCD58D360578" box="[716,851,798,825]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56B</figureCitation>
), antefurcal nervulus (
<figureCitation id="71B92A6FAC0C432E0F42FCD58ABE0578" box="[1110,1243,798,825]" captionStart="FIGURE 55" captionStartId="84.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@84.[151,1436,181,1623]" captionTargetPageId="84" captionText="FIGURE 55. Andrena (Aciandrena) karahisara spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Forewing, dorsolateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209423" httpUri="https://zenodo.org/record/13209423/files/figure.png" pageId="85" pageNumber="86">Figure 55E</figureCitation>
), genital capsule with produced gonocoxal teeth (
<figureCitation id="71B92A6FAC0C432E0910FC898CE2051C" box="[516,647,834,861]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56F</figureCitation>
), and the female hind tibial claws which lack an inner tooth. Females are extremely difficult to recognised, because they fall into the group of species around
<taxonomicName id="2E824D69AC0C432E0F7EFCAC8AA905C1" authorityName="Morawitz" authorityYear="1877" box="[1130,1228,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0F7EFCAC8AA905C1" box="[1130,1228,871,897]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
for which females are almost identical (most clearly also including
<taxonomicName id="2E824D69AC0C432E09D3FC408D3505E5" authorityName="Morawitz" authorityYear="1886" box="[711,848,906,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E09D3FC408D3505E5" box="[711,848,906,933]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC0C432E089FFC408ADA05E4" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[907,1215,906,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lamiana">
<emphasis id="DBF6EAF8AC0C432E089FFC408A6D05E5" box="[907,1032,906,933]" italics="true" pageId="85" pageNumber="86">A. lamiana</emphasis>
<bibRefCitation id="8D134B1BAC0C432E0F04FC418ADA05E4" author="Warncke, K." box="[1040,1215,906,933]" pageId="85" pageNumber="86" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
). Indeed, European material of
<taxonomicName id="2E824D69AC0C432E0A0FFC648FC60589" authorityName="Morawitz" authorityYear="1886" box="[283,419,942,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E0A0FFC648FC60589" box="[283,419,942,969]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
was confused with
<taxonomicName id="2E824D69AC0C432E0992FC648C8F0589" authorityName="Morawitz" authorityYear="1877" box="[646,746,943,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0992FC648C8F0589" box="[646,746,943,969]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
for a long time (e.g.
<bibRefCitation id="8D134B1BAC0C432E08C9FC658AE80589" author="Stockhert, E." box="[989,1165,942,968]" pageId="85" pageNumber="86" pagination="897 - 986" refId="ref91171" refString="Stockhert, E. (1930) Andrena Fabr. In: Schmiedeknecht, O. (Ed.), Die Hymenopteren Nord-und Mitteleuropas. Gustav Fischer, Jena, pp. 897 - 986." type="book chapter" year="1930">Stöckhert 1930</bibRefCitation>
), whereas true
<taxonomicName id="2E824D69AC0C432E0E2DFC648BF90589" authorityName="Morawitz" authorityYear="1877" box="[1337,1436,943,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0E2DFC648BF90589" box="[1337,1436,943,969]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
is restricted to eastern
<collectingCountry id="9195767AAC0C432E0A80FC198F8005AD" box="[404,485,978,1004]" name="Turkey" pageId="85" pageNumber="86">Turkey</collectingCountry>
, the Caucasus, northern
<collectingCountry id="9195767AAC0C432E09EDFC188D4C05AC" box="[761,809,979,1005]" name="Iran" pageId="85" pageNumber="86">Iran</collectingCountry>
, and
<collectingCountry id="9195767AAC0C432E0870FC198D9A05AD" box="[868,1023,978,1004]" name="Turkmenistan" pageId="85" pageNumber="86">Turkmenistan</collectingCountry>
(
<bibRefCitation id="8D134B1BAC0C432E0F1AFC198B7105AC" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Cierzniak, T." box="[1038,1300,978,1005]" pageId="85" pageNumber="86" refId="ref89759" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Cierzniak, T. (2005) Andreninae of the Central and Eastern Palaearctic. Part 1. Polish Entomological Monographs II. Polish Entomological Society, Poznan, 235 pp." type="book" year="2005">
Osytshnjuk
<emphasis id="DBF6EAF8AC0C432E0F83FC188AAD05AD" box="[1175,1224,978,1004]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
. 2005
</bibRefCitation>
). No female diagnosis is therefore given at the present time until molecular revision allows the identification of consistent characters to permit morphological separation of females of
<emphasis id="DBF6EAF8AC0C432E083BFBD18AA80275" box="[815,1229,1050,1077]" italics="true" pageId="85" pageNumber="86">
<taxonomicName id="2E824D69AC0C432E083BFBD18DF20275" authorityName="Morawitz" authorityYear="1886" box="[815,919,1050,1076]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">aciculata</taxonomicName>
-
<taxonomicName id="2E824D69AC0C432E08B4FBD18A7E0275" authorityName="WOOD" authorityYear="2024" box="[928,1051,1050,1076]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">karahisara</taxonomicName>
-
<taxonomicName id="2E824D69AC0C432E0F31FBD18AE40275" authorityName="Warncke" authorityYear="1965" box="[1061,1153,1050,1076]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lamiana">lamiana</taxonomicName>
-
<taxonomicName id="2E824D69AC0C432E0F9EFBD08AA80275" authorityName="Morawitz" authorityYear="1877" box="[1162,1229,1051,1076]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">tenuis</taxonomicName>
</emphasis>
. Female
<typeStatus id="36398848AC0C432E0E25FBD08BF90274" box="[1329,1436,1051,1077]" pageId="85" pageNumber="86" type="paratype">paratypes</typeStatus>
were accepted only based on association with males that were concurrently active at sampling sites.
</paragraph>
<paragraph id="E93D36EAAC0C432E0BD3FBA98F6100D9" blockId="85.[151,1437,150,1797]" pageId="85" pageNumber="86">
Recognition is best made in males, specifically by using the male genital capsule. For
<taxonomicName id="2E824D69AC0C432E0F92FBA88B68023D" authorityName="Morawitz" authorityYear="1886" box="[1158,1293,1122,1149]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E0F92FBA88B68023D" box="[1158,1293,1122,1149]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
,
<taxonomicName id="2E824D69AC0C432E0E0FFBA88BF0023D" authorityName="Warncke" authorityYear="1965" box="[1307,1429,1122,1149]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lamiana">
<emphasis id="DBF6EAF8AC0C432E0E0FFBA88BF0023D" box="[1307,1429,1122,1149]" italics="true" pageId="85" pageNumber="86">A. lamiana</emphasis>
</taxonomicName>
, and
<taxonomicName id="2E824D69AC0C432E0BD3FB4C8F4C02E1" authorityName="Morawitz" authorityYear="1877" box="[199,297,1159,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0BD3FB4C8F4C02E1" box="[199,297,1159,1185]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC0C432E0A23FB4D8FD802E0" box="[311,445,1158,1185]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56H</figureCitation>
), the genital capsule is compact, more or less as long as broad, whereas in
<taxonomicName id="2E824D69AC0C432E0E16FB4C8BF902E1" authorityName="WOOD" authorityYear="2024" box="[1282,1436,1158,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0E16FB4C8BF902E1" box="[1282,1436,1158,1185]" italics="true" pageId="85" pageNumber="86">A. karahisara</emphasis>
</taxonomicName>
the genital capsule is clearly elongate, 1.5 times longer than wide, and with the gonostyli showing a clear bump or kink on their internal margin medially (
<figureCitation id="71B92A6FAC0C432E096DFB058C9902A8" box="[633,764,1230,1257]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56F</figureCitation>
). This elongate genital capsule with a bump or kink on the internal margin of the gonostyli is closer to
<taxonomicName id="2E824D69AC0C432E0968FB388DD7034C" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[636,946,1266,1293]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="pulicaria">
<emphasis id="DBF6EAF8AC0C432E0968FB388D64034D" box="[636,769,1266,1293]" italics="true" pageId="85" pageNumber="86">A. pulicaria</emphasis>
Warncke, 1975
</taxonomicName>
, but this species has a clearly more elongate head, only 1.05 times wider than long.
<taxonomicName id="2E824D69AC0C432E0944FADD8D570371" authorityName="WOOD" authorityYear="2024" box="[592,818,1302,1328]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0944FADD8D570371" box="[592,818,1302,1328]" italics="true" pageId="85" pageNumber="86">Andrena karahisara</emphasis>
</taxonomicName>
can also be separated from
<taxonomicName id="2E824D69AC0C432E0F79FADC8A8C0371" authorityName="Warncke" authorityYear="1965" box="[1133,1257,1302,1329]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lamiana">
<emphasis id="DBF6EAF8AC0C432E0F79FADC8A8C0371" box="[1133,1257,1302,1329]" italics="true" pageId="85" pageNumber="86">A. lamiana</emphasis>
</taxonomicName>
by the shape of S8 which slightly but distinctly broadens apically forming a truncate apical process (as in
<taxonomicName id="2E824D69AC0C432E0F93FAF08B6B0315" authorityName="Morawitz" authorityYear="1886" box="[1159,1294,1338,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E0F93FAF08B6B0315" box="[1159,1294,1338,1365]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
, and weakly so in
<taxonomicName id="2E824D69AC0C432E0BC5FA948F570339" authorityName="Morawitz" authorityYear="1877" box="[209,306,1375,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0BC5FA948F570339" box="[209,306,1375,1401]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
), whereas in
<taxonomicName id="2E824D69AC0C432E0AD1FA948C5A0339" authorityName="Warncke" authorityYear="1965" box="[453,575,1374,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lamiana">
<emphasis id="DBF6EAF8AC0C432E0AD1FA948C5A0339" box="[453,575,1374,1401]" italics="true" pageId="85" pageNumber="86">A. lamiana</emphasis>
</taxonomicName>
S8 is parallel sided and comes to a narrowly rounded point, the apical process therefore not presenting a broadened truncate apical process. Separation from
<taxonomicName id="2E824D69AC0C432E08EEFA488AE403DD" authorityName="Morawitz" authorityYear="1886" box="[1018,1153,1410,1437]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E08EEFA488AE403DD" box="[1018,1153,1410,1437]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
can also be provisionally made by the colouration of the clypeus, as males of
<taxonomicName id="2E824D69AC0C432E09FEFA6C8D170381" authorityName="Morawitz" authorityYear="1886" box="[746,882,1446,1473]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="aciculata">
<emphasis id="DBF6EAF8AC0C432E09FEFA6C8D170381" box="[746,882,1446,1473]" italics="true" pageId="85" pageNumber="86">A. aciculata</emphasis>
</taxonomicName>
tend to have the yellow facial markings reduced so that they do not cover the entire clypeus (
<figureCitation id="71B92A6FAC0C432E09B7FA018D4903A4" box="[675,812,1482,1509]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56G</figureCitation>
), where
<taxonomicName id="2E824D69AC0C432E0886FA008A4A03A5" authorityName="WOOD" authorityYear="2024" box="[914,1071,1482,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0886FA008A4A03A5" box="[914,1071,1482,1509]" italics="true" pageId="85" pageNumber="86">A. karahisara</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC0C432E0F78FA008AB403A5" authorityName="Morawitz" authorityYear="1877" box="[1132,1233,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E0F78FA008AB403A5" box="[1132,1233,1483,1509]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
have the clypeus entirely yellow marked (with the exception of two small black dots medially, see photographs in
<bibRefCitation id="8D134B1BAC0C432E0FCBFA258EB6006D" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." pageId="85" pageNumber="86" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC0C432E0E77FA248BF00049" box="[1379,1429,1518,1544]" italics="true" pageId="85" pageNumber="86">et al</emphasis>
. 2021
</bibRefCitation>
). Finally,
<taxonomicName id="2E824D69AC0C432E0A53F9D88F81006D" authorityName="WOOD" authorityYear="2024" box="[327,484,1554,1581]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0A53F9D88F81006D" box="[327,484,1554,1581]" italics="true" pageId="85" pageNumber="86">A. karahisara</emphasis>
</taxonomicName>
males may also be separable from
<taxonomicName id="2E824D69AC0C432E086BF9D88D81006D" authorityName="Morawitz" authorityYear="1877" box="[895,996,1555,1581]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E086BF9D88D81006D" box="[895,996,1555,1581]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
based on the width of the head, since
<taxonomicName id="2E824D69AC0C432E0B83F9FC8F560011" authorityName="WOOD" authorityYear="2024" box="[151,307,1590,1617]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="karahisara">
<emphasis id="DBF6EAF8AC0C432E0B83F9FC8F560011" box="[151,307,1590,1617]" italics="true" pageId="85" pageNumber="86">A. karahisara</emphasis>
</taxonomicName>
has the head 1.15 times wider than long (
<figureCitation id="71B92A6FAC0C432E0806F9FD8DFD0010" box="[786,920,1590,1617]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56B</figureCitation>
), in
<taxonomicName id="2E824D69AC0C432E08D8F9FC8A4A0011" authorityName="Morawitz" authorityYear="1877" box="[972,1071,1591,1617]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0C432E08D8F9FC8A4A0011" box="[972,1071,1591,1617]" italics="true" pageId="85" pageNumber="86">A. tenuis</emphasis>
</taxonomicName>
the head is 1.21.3 times wider than long (
<figureCitation id="71B92A6FAC0C432E0A05F9918FFD0034" box="[273,408,1626,1653]" captionStart="FIGURE 56" captionStartId="86.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@86.[214,1373,181,1919]" captionTargetPageId="86" captionText="FIGURE 56. Andrena (Aciandrena) karahisara spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; Andrena (Aciandrena) tenuis Morawitz, 1877 male. G. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209425" httpUri="https://zenodo.org/record/13209425/files/figure.png" pageId="85" pageNumber="86">Figure 56G</figureCitation>
). Additional study using molecular markers is necessary to confirm that this is a consistent character.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0C432E0BD3F9698CC200A1" pageId="85" pageNumber="86" type="etymology">
<paragraph id="E93D36EAAC0C432E0BD3F9698CC200A1" blockId="85.[151,1437,150,1797]" pageId="85" pageNumber="86">
<emphasis id="DBF6EAF8AC0C432E0BD3F9698F2B00FD" bold="true" box="[199,334,1698,1724]" pageId="85" pageNumber="86">Etymology.</emphasis>
From the Turkish “
<emphasis id="DBF6EAF8AC0C432E0926F9698CFA00FD" box="[562,671,1698,1724]" italics="true" pageId="85" pageNumber="86">karahisar</emphasis>
” meaning “black castle”. This name was selected by Warncke, but was never published. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC0C432E0BD3F9218D310145" box="[199,852,1770,1797]" pageId="85" pageNumber="86" type="distribution">
<paragraph id="E93D36EAAC0C432E0BD3F9218D310145" blockId="85.[151,1437,150,1797]" box="[199,852,1770,1797]" pageId="85" pageNumber="86">
<emphasis id="DBF6EAF8AC0C432E0BD3F9218F040145" bold="true" box="[199,353,1770,1796]" pageId="85" pageNumber="86">Distribution.</emphasis>
Central
<collectingCountry id="9195767AAC0C432E0AD4F9218C750145" box="[448,528,1770,1796]" name="Turkey" pageId="85" pageNumber="86">Turkey</collectingCountry>
(
<collectingRegion id="2B46F808AC0C432E090BF9218C1F0145" box="[543,634,1770,1796]" country="Turkey" name="Kayseri" pageId="85" pageNumber="86">Kayseri</collectingRegion>
,
<collectingRegion id="2B46F808AC0C432E0992F9208CB60144" box="[646,723,1771,1797]" country="Turkey" name="Konya" pageId="85" pageNumber="86">Konya</collectingRegion>
,
<collectingRegion id="2B46F808AC0C432E09CAF9218D2D0145" box="[734,840,1770,1796]" country="Turkey" name="Nevsehir" pageId="85" pageNumber="86">Nevşehir</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,383 @@
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<emphasis id="DBF6EAF8AC0E432C0B83FF5C8BF706F0" bold="true" box="[151,1426,151,178]" pageId="87" pageNumber="88">
Other material examined (Turkish
<taxonomicName id="2E824D69AC0E432C092CFF5C8CDA06F3" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[568,703,151,178]" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="106" phylum="Arthropoda" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC0E432C092CFF5C8CDA06F3" bold="true" box="[568,703,151,178]" italics="true" pageId="87" pageNumber="88">Aciandrena</emphasis>
</taxonomicName>
species, records supported by at least some male specimens).
</emphasis>
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<subSubSection id="A1986561AC0E432C0B83FF148AD40425" pageId="87" pageNumber="88" type="materials_examined">
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(
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<emphasis id="DBF6EAF8AC0E432C0BB5FF148F1406B8" box="[161,369,223,249]" italics="true" pageId="87" pageNumber="88">Andrena aciculata</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC0E432C0A90FF148BFD06B8" box="[388,1432,223,252]" collectingDate="1997-06-29" collectorName="Ma. Halada" country="Turkey" location="20 km NW Igdir" pageId="87" pageNumber="88" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Igdir">
<emphasis id="DBF6EAF8AC0E432C0A90FF148C6706B8" bold="true" box="[388,514,223,249]" pageId="87" pageNumber="88">
<collectingCountry id="9195767AAC0E432C0A90FF148F9B06B8" box="[388,510,223,249]" name="Turkey" pageId="87" pageNumber="88">TURKEY</collectingCountry>
:
</emphasis>
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NW
<collectingRegion id="2B46F808AC0E432C0981FF148CAE06B8" box="[661,715,223,249]" country="Turkey" name="Igdir" pageId="87" pageNumber="88">Igdir</collectingRegion>
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[
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],
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<collectingDate id="8D78E9C2AC0E432C083AFF148DCD06B8" box="[814,936,223,249]" pageId="87" pageNumber="88" value="1997-06-29">29.vi.1997</collectingDate>
</date>
,
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,
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, leg.
<collectorName id="4477533CAC0E432C0F5BFF2B8AB306B8" box="[1103,1238,223,250]" pageId="87" pageNumber="88">Ma. Halada</collectorName>
, OÖLM;
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</materialsCitation>
,
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SW
<collectingRegion id="2B46F808AC0E432C0A09FEC88F0B075C" box="[285,366,259,285]" country="Turkey" name="Burdur" pageId="87" pageNumber="88">Burdur</collectingRegion>
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,
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<elevation id="62AFD1D9AC0E432C0A68FEC88FA2075F" box="[380,455,259,286]" metricMagnitude="2" metricUnit="m" metricValue="9.4" pageId="87" pageNumber="88" unit="m" value="940.0">940 m</elevation>
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,
<date id="9D3C102AAC0E432C0AC1FEC88C2F075C" box="[469,586,259,285]" pageId="87" pageNumber="88" value="2006-07-07">
<collectingDate id="8D78E9C2AC0E432C0AC1FEC88C2F075C" box="[469,586,259,285]" pageId="87" pageNumber="88" value="2006-07-07">7.vii.2006</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0943FEC88CEF075C" box="[599,650,259,285]" count="12" pageId="87" pageNumber="88" type="male">12♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C09D8FECF8D5C075C" box="[716,825,259,286]" pageId="87" pageNumber="88">J. Halada</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC0E432C08B7FECE8D880761" box="[931,1005,261,288]" pageId="87" pageNumber="88">TJWC</collectionCode>
;
<collectingRegion id="2B46F808AC0E432C08EFFEC88A29075C" box="[1019,1100,259,285]" country="Turkey" name="Burdur" pageId="87" pageNumber="88">Burdur</collectingRegion>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0E432C0F4EFEC88D7E0703" collectingDate="2006-07-06" collectionCode="TJWC" collectorName="J. Halada" country="Turkey" elevation="1060" location="5 km NE Yesilova" pageId="87" pageNumber="88" specimenCount="2" specimenCount-male="2" stateProvince="Erzurum">
<location id="EC5D6031AC0E432C0F4EFEC88B54075C" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0F4EFEC88B54075C" box="[1114,1329,259,286]" country="Turkey" name="5 km NE Yesilova" pageId="87" pageNumber="88" stateProvince="Erzurum">
<quantity id="2E7A9B0FAC0E432C0F4EFEC88AF3075C" box="[1114,1174,259,286]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="87" pageNumber="88" unit="km" value="5.0">5 km</quantity>
NE Yesilova
</location>
,
<quantity id="2E7A9B0FAC0E432C0E2AFEC88BF2075F" box="[1342,1431,259,286]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="87" pageNumber="88" unit="m" value="1060.0">
<elevation id="62AFD1D9AC0E432C0E2AFEC88BF2075F" box="[1342,1431,259,286]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="87" pageNumber="88" unit="m" value="1060.0">1060 m</elevation>
</quantity>
,
<date id="9D3C102AAC0E432C0B83FEEC8F690700" box="[151,268,295,321]" pageId="87" pageNumber="88" value="2006-07-06">
<collectingDate id="8D78E9C2AC0E432C0B83FEEC8F690700" box="[151,268,295,321]" pageId="87" pageNumber="88" value="2006-07-06">6.vii.2006</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0A0EFEEC8F250700" box="[282,320,295,321]" count="2" pageId="87" pageNumber="88" type="male">2♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0A97FEE38F940700" box="[387,497,295,322]" pageId="87" pageNumber="88">J. Halada</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC0E432C0949FEE28CC20705" box="[605,679,297,324]" pageId="87" pageNumber="88">TJWC</collectionCode>
;
<collectingRegion id="2B46F808AC0E432C09A2FEE38D7E0703" box="[694,795,296,322]" country="Turkey" name="Erzurum" pageId="87" pageNumber="88">Erzurum</collectingRegion>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0E432C083EFEEC8B4D0703" box="[810,1320,295,324]" collectingDate="2001-07-01" collectorName="J. Straka" country="Turkey" location="Kars" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Kars">
<date id="9D3C102AAC0E432C083EFEEC8DF80700" box="[810,925,295,321]" pageId="87" pageNumber="88" value="2001-07-01">
<collectingDate id="8D78E9C2AC0E432C083EFEEC8DF80700" box="[810,925,295,321]" pageId="87" pageNumber="88" value="2001-07-01">1.vii.2001</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C08B9FEEC8DB40700" box="[941,977,295,321]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0F01FEE38A190700" box="[1045,1148,295,322]" pageId="87" pageNumber="88">J. Straka</collectorName>
, OÖLM;
<collectingRegion id="2B46F808AC0E432C0FE4FEE38B4D0703" box="[1264,1320,296,322]" country="Turkey" name="Kars" pageId="87" pageNumber="88">Kars</collectingRegion>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0E432C0E22FEEC8DA70724" collectingDate="1988-05-31" collectorName="K. Warncke" country="Turkey" elevation="1460" location="10 km E Karakurt" pageId="87" pageNumber="88" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kutahya">
<location id="EC5D6031AC0E432C0E22FEEC8E980724" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0E22FEEC8E980724" country="Turkey" name="10 km E Karakurt" pageId="87" pageNumber="88" stateProvince="Kutahya">
<quantity id="2E7A9B0FAC0E432C0E22FEEC8BE40700" box="[1334,1409,295,321]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="87" pageNumber="88" unit="km" value="10.0">10 km</quantity>
E Karakurt
</location>
,
<quantity id="2E7A9B0FAC0E432C0A13FE808F380727" box="[263,349,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="87" pageNumber="88" unit="m" value="1460.0">
<elevation id="62AFD1D9AC0E432C0A13FE808F380727" box="[263,349,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="87" pageNumber="88" unit="m" value="1460.0">1460 m</elevation>
</quantity>
,
<date id="9D3C102AAC0E432C0A73FE808FBD0724" box="[359,472,331,357]" pageId="87" pageNumber="88" value="1988-05-31">
<collectingDate id="8D78E9C2AC0E432C0A73FE808FBD0724" box="[359,472,331,357]" pageId="87" pageNumber="88" value="1988-05-31">31.v.1988</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0AF6FE808C6D0724" box="[482,520,331,357]" count="2" pageId="87" pageNumber="88" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC0E432C0906FE808C500724" box="[530,565,331,357]" count="1" pageId="87" pageNumber="88" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0964FE878C920724" box="[624,759,331,358]" pageId="87" pageNumber="88">K. Warncke</collectorName>
, OÖLM;
<collectingRegion id="2B46F808AC0E432C0874FE808DA70724" box="[864,962,331,357]" country="Turkey" name="Kutahya" pageId="87" pageNumber="88">Kutahya</collectingRegion>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0E432C08DFFE808C5507CB" collectingDate="2006-07-12" collectorName="J. Halada" country="Turkey" elevation="1110" location="28 km SSE Kutahya" pageId="87" pageNumber="88" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kutahya">
<location id="EC5D6031AC0E432C08DFFE808ACB0724" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C08DFFE808ACB0724" box="[971,1198,331,357]" country="Turkey" name="28 km SSE Kutahya" pageId="87" pageNumber="88" stateProvince="Kutahya">
<quantity id="2E7A9B0FAC0E432C08DFFE808A740724" box="[971,1041,331,357]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="87" pageNumber="88" unit="km" value="28.0">28 km</quantity>
SSE
<collectingRegion id="2B46F808AC0E432C0F58FE808ACB0724" box="[1100,1198,331,357]" country="Turkey" name="Kutahya" pageId="87" pageNumber="88">Kutahya</collectingRegion>
</location>
,
<quantity id="2E7A9B0FAC0E432C0FACFE808B6E0727" box="[1208,1291,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="87" pageNumber="88" unit="m" value="1110.0">
<elevation id="62AFD1D9AC0E432C0FACFE808B6E0727" box="[1208,1291,331,358]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="87" pageNumber="88" unit="m" value="1110.0">1110 m</elevation>
</quantity>
,
<date id="9D3C102AAC0E432C0E02FE808BFD0724" box="[1302,1432,331,357]" pageId="87" pageNumber="88" value="2006-07-12">
<collectingDate id="8D78E9C2AC0E432C0E02FE808BFD0724" box="[1302,1432,331,357]" pageId="87" pageNumber="88" value="2006-07-12">12.vii.2006</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0B83FEA48ED807C8" box="[151,189,367,393]" count="2" pageId="87" pageNumber="88" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC0E432C0BDCFEA48E8907C8" box="[200,236,367,393]" count="3" pageId="87" pageNumber="88" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0A3DFEBB8FF107C8" box="[297,404,367,394]" pageId="87" pageNumber="88">J. Halada</collectorName>
, OÖLM;
<collectingRegion id="2B46F808AC0E432C0914FEBB8C5507CB" box="[512,560,368,394]" country="Turkey" name="Van" pageId="87" pageNumber="88">Van</collectingRegion>
</materialsCitation>
:
<materialsCitation id="59EA3CB7AC0E432C092FFEA48B4A07CD" box="[571,1327,367,396]" collectingDate="1981-06-08" collectorName="K. Warncke" country="Turkey" elevation="2350" location="Baskale" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1">
<quantity id="2E7A9B0FAC0E432C092FFEA48CE707C8" box="[571,642,367,393]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="87" pageNumber="88" unit="km" value="40.0">40 km</quantity>
N
<location id="EC5D6031AC0E432C09B0FEA48C9A07C8" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C09B0FEA48C9A07C8" box="[676,767,367,393]" country="Turkey" name="Baskale" pageId="87" pageNumber="88">Başkale</location>
,
<quantity id="2E7A9B0FAC0E432C081EFEA48D0407CB" box="[778,865,367,394]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="87" pageNumber="88" unit="m" value="2350.0">
<elevation id="62AFD1D9AC0E432C081EFEA48D0407CB" box="[778,865,367,394]" metricMagnitude="3" metricUnit="m" metricValue="2.35" pageId="87" pageNumber="88" unit="m" value="2350.0">2350 m</elevation>
</quantity>
,
<date id="9D3C102AAC0E432C0879FEA48DBC07C8" box="[877,985,367,393]" pageId="87" pageNumber="88" value="1981-06-08">
<collectingDate id="8D78E9C2AC0E432C0879FEA48DBC07C8" box="[877,985,367,393]" pageId="87" pageNumber="88" value="1981-06-08">8.vi.1981</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C08F1FEA48A6C07C8" box="[997,1033,367,393]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0F52FEBB8AAB07C8" box="[1094,1230,367,394]" pageId="87" pageNumber="88">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
;
</paragraph>
<paragraph id="E93D36EAAC0E432C0BD3FE588A680401" blockId="87.[151,1437,223,612]" pageId="87" pageNumber="88">
(
<taxonomicName id="2E824D69AC0E432C0BC5FE588FF607EC" authorityName="Warncke" authorityYear="1965" box="[209,403,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="lamiana">
<emphasis id="DBF6EAF8AC0E432C0BC5FE588FF607EC" box="[209,403,403,429]" italics="true" pageId="87" pageNumber="88">Andrena lamiana</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC0E432C0AB7FE588B4B07F1" box="[419,1326,403,432]" collectingDate="2002-05-09" collectionCode="OOLM" collectorName="F. Kantner" country="Turkey" location="Hasankeyf" pageId="87" pageNumber="88" specimenCount="3" specimenCount-male="3" stateProvince="Mardin">
<emphasis id="DBF6EAF8AC0E432C0AB7FE588C4707EC" bold="true" box="[419,546,403,429]" pageId="87" pageNumber="88">
<collectingCountry id="9195767AAC0E432C0AB7FE588C7807EC" box="[419,541,403,429]" name="Turkey" pageId="87" pageNumber="88">TURKEY</collectingCountry>
:
</emphasis>
<locationDeviation id="F88F2FC8AC0E432C093DFE588C9707EC" box="[553,754,403,430]" location="Mardin" pageId="87" pageNumber="88">
<quantity id="2E7A9B0FAC0E432C093DFE588C1B07EC" box="[553,638,403,429]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="87" pageNumber="88" unit="km" value="110.0">110 km</quantity>
E
<collectingRegion id="2B46F808AC0E432C0989FE588C9707EC" box="[669,754,403,429]" country="Turkey" name="Mardin" pageId="87" pageNumber="88">Mardin</collectingRegion>
</locationDeviation>
,
<location id="EC5D6031AC0E432C09EAFE588D1C07EC" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C09EAFE588D1C07EC" box="[766,889,403,429]" country="Turkey" name="Hasankeyf" pageId="87" pageNumber="88" stateProvince="Mardin">Hasankeyf</location>
,
<date id="9D3C102AAC0E432C0890FE588D8207EC" box="[900,999,403,429]" pageId="87" pageNumber="88" value="2002-05-09">
<collectingDate id="8D78E9C2AC0E432C0890FE588D8207EC" box="[900,999,403,429]" pageId="87" pageNumber="88" value="2002-05-09">9.v.2002</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C08E7FE588A7D07EC" box="[1011,1048,403,429]" count="3" pageId="87" pageNumber="88" type="male">3♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0F41FE5F8AA907EF" box="[1109,1228,404,430]" pageId="87" pageNumber="88">F. Kantner</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C0FCCFE5E8B4B07F1" box="[1240,1326,405,432]" pageId="87" pageNumber="88">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0E2EFE588D4C0795" collectingDate="2002-05-10" collectionCode="OOLM" collectorName="F. Kantner" country="Turkey" location="Fevzipasa" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1">
<locationDeviation id="F88F2FC8AC0E432C0E2EFE588E970790" location="Islahiye" pageId="87" pageNumber="88">
<quantity id="2E7A9B0FAC0E432C0E2EFE588BE407EC" box="[1338,1409,403,429]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="87" pageNumber="88" unit="km" value="13.0">13 km</quantity>
N Islahiye
</locationDeviation>
,
<location id="EC5D6031AC0E432C0BE9FE7C8F150790" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0BE9FE7C8F150790" box="[253,368,439,465]" country="Turkey" name="Fevzipasa" pageId="87" pageNumber="88">Fevzipasa</location>
,
<date id="9D3C102AAC0E432C0A6DFE7C8F8F0790" box="[377,490,439,465]" pageId="87" pageNumber="88" value="2002-05-10">
<collectingDate id="8D78E9C2AC0E432C0A6DFE7C8F8F0790" box="[377,490,439,465]" pageId="87" pageNumber="88" value="2002-05-10">10.v.2002</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0AE0FE7C8C7C0790" box="[500,537,439,465]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0947FE738CAC0793" box="[595,713,440,466]" pageId="87" pageNumber="88">F. Kantner</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C09C7FE728D4C0795" box="[723,809,441,468]" pageId="87" pageNumber="88">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0827FE7C8FCB07B9" collectingDate="1998-05-22" collectionCode="OOLM" collectorName="Ma. Halada" country="Turkey" location="Porsuk Baraji" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Kutahya">
<locationDeviation id="F88F2FC8AC0E432C0827FE7C8D9F0790" box="[819,1018,439,466]" location="Kutahya" pageId="87" pageNumber="88">
<quantity id="2E7A9B0FAC0E432C0827FE7C8D1C0790" box="[819,889,439,465]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="87" pageNumber="88" unit="km" value="30.0">30 km</quantity>
N
<collectingRegion id="2B46F808AC0E432C088CFE7C8D9F0790" box="[920,1018,439,465]" country="Turkey" name="Kutahya" pageId="87" pageNumber="88">Kutahya</collectingRegion>
</locationDeviation>
,
<location id="EC5D6031AC0E432C0F10FE7C8AFE0790" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0F10FE7C8AFE0790" box="[1028,1179,439,465]" country="Turkey" name="Porsuk Baraji" pageId="87" pageNumber="88" stateProvince="Kutahya">Porsuk Baraji</location>
[Dam],
<date id="9D3C102AAC0E432C0FE3FE7C8B0D0790" box="[1271,1384,439,465]" pageId="87" pageNumber="88" value="1998-05-22">
<collectingDate id="8D78E9C2AC0E432C0FE3FE7C8B0D0790" box="[1271,1384,439,465]" pageId="87" pageNumber="88" value="1998-05-22">22.v.1998</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0E66FE7C8BF20790" box="[1394,1431,439,465]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0BD3FE178F2807B4" box="[199,333,475,502]" pageId="87" pageNumber="88">Ma. Halada</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C0A4CFE168FCB07B9" box="[344,430,477,504]" pageId="87" pageNumber="88">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0AAEFE108AFA07B9" box="[442,1183,475,504]" collectingDate="1997-06-18" collectionCode="OOLM" collectorName="Ma. Halada" country="Turkey" location="Cornelek" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1">
<locationDeviation id="F88F2FC8AC0E432C0AAEFE108C3407B7" box="[442,593,475,502]" location="Mut" pageId="87" pageNumber="88">
<quantity id="2E7A9B0FAC0E432C0AAEFE108C6407B4" box="[442,513,475,501]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="87" pageNumber="88" unit="km" value="40.0">40 km</quantity>
E Mut
</locationDeviation>
,
<location id="EC5D6031AC0E432C0948FE108CA307B4" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0948FE108CA307B4" box="[604,710,475,501]" country="Turkey" name="Cornelek" pageId="87" pageNumber="88">Cornelek</location>
,
<date id="9D3C102AAC0E432C09C5FE108D2E07B4" box="[721,843,475,501]" pageId="87" pageNumber="88" value="1997-06-18">
<collectingDate id="8D78E9C2AC0E432C09C5FE108D2E07B4" box="[721,843,475,501]" pageId="87" pageNumber="88" value="1997-06-18">18.vi.1997</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0843FE108D1E07B4" box="[855,891,475,501]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C08ACFE178A5B07B4" box="[952,1086,475,502]" pageId="87" pageNumber="88">Ma. Halada</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C0F5DFE168AFA07B9" box="[1097,1183,477,504]" pageId="87" pageNumber="88">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0FBFFE108F84045D" collectingDate="2002-05-02" collectionCode="OOLM" collectorName="F. Kantner" country="Turkey" location="Iskenderun" pageId="87" pageNumber="88" specimenCount="3" specimenCount-male="3">
<location id="EC5D6031AC0E432C0FBFFE108B4F07B4" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0FBFFE108B4F07B4" box="[1195,1322,475,501]" country="Turkey" name="Iskenderun" pageId="87" pageNumber="88">Iskenderun</location>
,
<date id="9D3C102AAC0E432C0E21FE108BFD07B4" box="[1333,1432,475,501]" pageId="87" pageNumber="88" value="2002-05-02">
<collectingDate id="8D78E9C2AC0E432C0E21FE108BFD07B4" box="[1333,1432,475,501]" pageId="87" pageNumber="88" value="2002-05-02">2.v.2002</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0B83FE348ED90458" box="[151,188,511,537]" count="3" pageId="87" pageNumber="88" type="male">3♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0A15FDCB8F19045B" box="[257,380,512,538]" pageId="87" pageNumber="88">F. Kantner</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C0A9FFDCA8F84045D" box="[395,481,513,540]" pageId="87" pageNumber="88">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0AE4FE348BFD045B" box="[496,1432,511,538]" collectingDate="1974-04-10" collectionCode="RMNH" collectorName="W. H. Gravestein &amp; S. J. van Ooststroom" country="Turkey" location="Kusadasi" pageId="87" pageNumber="88" specimenCount="8" specimenCount-male="8">
<location id="EC5D6031AC0E432C0AE4FE348C3C0458" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0AE4FE348C3C0458" box="[496,601,511,537]" country="Turkey" name="Kusadasi" pageId="87" pageNumber="88">Kuşadası</location>
,
<date id="9D3C102AAC0E432C097DFE348C870458" box="[617,738,511,537]" pageId="87" pageNumber="88" value="1974-04-10">
<collectingDate id="8D78E9C2AC0E432C097DFE348C870458" box="[617,738,511,537]" pageId="87" pageNumber="88" value="1974-04-10">10.iv.1974</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C09E5FE348D730458" box="[753,790,511,537]" count="8" pageId="87" pageNumber="88" type="male">8♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C084EFDCB8A7D0458" box="[858,1048,511,538]" pageId="87" pageNumber="88">W.H. Gravestein</collectorName>
&amp;
<collectorName id="4477533CAC0E432C0F51FE348B570458" box="[1093,1330,511,538]" pageId="87" pageNumber="88">S.J. van Ooststroom</collectorName>
,
<collectionCode id="8F93AE2FAC0E432C0E54FDCB8BFD045B" box="[1344,1432,512,538]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="87" pageNumber="88" type="Museum">RMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC0E432C0B83FDE88FFD047C" box="[151,408,547,574]" country="Turkey" elevation="2000" location="Muradiye" pageId="87" pageNumber="88" specimenCount="1">
<location id="EC5D6031AC0E432C0B83FDE88F61047C" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C0B83FDE88F61047C" box="[151,260,547,573]" country="Turkey" name="Muradiye" pageId="87" pageNumber="88">Muradiye</location>
env.,
<quantity id="2E7A9B0FAC0E432C0A53FDE88FFD047C" box="[327,408,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="87" pageNumber="88" unit="m" value="2000.0">
<elevation id="62AFD1D9AC0E432C0A53FDE88FFD047C" box="[327,408,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="87" pageNumber="88" unit="m" value="2000.0">2000m</elevation>
</quantity>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC0E432C0AB6FDE88DCD047C" box="[418,936,547,574]" collectingDate="2001-06-05" collectorName="K. Denes" country="Turkey" location="Van" pageId="87" pageNumber="88" specimenCount="1" specimenCount-male="1" stateProvince="Van">
<quantity id="2E7A9B0FAC0E432C0AB6FDE88F92047C" box="[418,503,547,573]" metricMagnitude="5" metricUnit="m" metricValue="1.2" pageId="87" pageNumber="88" unit="km" value="120.0">120 km</quantity>
NE
<collectingRegion id="2B46F808AC0E432C093EFDEF8C3C047F" box="[554,601,548,574]" country="Turkey" name="Van" pageId="87" pageNumber="88">Van</collectingRegion>
,
<date id="9D3C102AAC0E432C0970FDE88CB5047F" box="[612,720,547,574]" pageId="87" pageNumber="88" value="2001-06-05">
<collectingDate id="8D78E9C2AC0E432C0970FDE88CB5047F" box="[612,720,547,574]" pageId="87" pageNumber="88" value="2001-06-05">5.vi.2001</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C09CFFDE88D65047C" box="[731,768,547,573]" count="1" pageId="87" pageNumber="88" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C0829FDEF8DCD047C" box="[829,936,547,574]" pageId="87" pageNumber="88">K. Deneš</collectorName>
</materialsCitation>
, OÖLM.
</paragraph>
<paragraph id="E93D36EAAC0E432C0BD3FD8C8AD40425" blockId="87.[151,1437,223,612]" box="[199,1201,583,612]" pageId="87" pageNumber="88">
(
<taxonomicName id="2E824D69AC0E432C0BC5FD8C8F1F0420" authorityName="Morawitz" authorityYear="1877" box="[209,378,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="tenuis">
<emphasis id="DBF6EAF8AC0E432C0BC5FD8C8F1F0420" box="[209,378,583,609]" italics="true" pageId="87" pageNumber="88">Andrena tenuis</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC0E432C0A9EFD8C8AC80425" box="[394,1197,583,612]" collectingDate="1975-05-22" collectorName="K. Warncke" country="Turkey" location="Karakurt" municipality="Arastal" pageId="87" pageNumber="88" specimenCount="25" specimenCount-male="25">
<emphasis id="DBF6EAF8AC0E432C0A9EFD8C8C6D0420" bold="true" box="[394,520,583,609]" pageId="87" pageNumber="88">
<collectingCountry id="9195767AAC0E432C0A9EFD8C8C610420" box="[394,516,583,609]" name="Turkey" pageId="87" pageNumber="88">TURKEY</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC0E432C091BFD8C8C100420" LSID="urn:lsid:plazi:treatment:612B87FCAC0E432C0B83FF5C8AD40425:EC5D6031AC0E432C091BFD8C8C100420" box="[527,629,583,609]" country="Turkey" municipality="Arastal" name="Karakurt" pageId="87" pageNumber="88">Karakurt</location>
/
<collectingMunicipality id="0959AC90AC0E432C0969FD8C8CAB0420" box="[637,718,583,609]" pageId="87" pageNumber="88">Arastal</collectingMunicipality>
,
<date id="9D3C102AAC0E432C09CDFD8C8D2F0423" box="[729,842,583,610]" pageId="87" pageNumber="88" value="1975-05-22">
<collectingDate id="8D78E9C2AC0E432C09CDFD8C8D2F0423" box="[729,842,583,610]" pageId="87" pageNumber="88" value="1975-05-22">22.v.1975</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC0E432C0841FD8C8DEC0423" box="[853,905,583,610]" count="25" pageId="87" pageNumber="88" type="male">25♂</specimenCount>
, leg.
<collectorName id="4477533CAC0E432C08D1FD838A290420" box="[965,1100,583,610]" pageId="87" pageNumber="88">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,446 @@
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<treatment id="612B87FCAC1143310B83F8E68C880700" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700" httpUri="http://treatment.plazi.org/id/612B87FCAC1143310B83F8E68C880700" lastPageId="74" lastPageNumber="75" pageId="72" pageNumber="73">
<subSubSection id="A1986561AC1143330B83F8E68D3B0106" box="[151,862,1837,1864]" pageId="72" pageNumber="73" type="nomenclature">
<paragraph id="E93D36EAAC1143330B83F8E68D3B0106" blockId="72.[151,862,1837,1864]" box="[151,862,1837,1864]" pageId="72" pageNumber="73">
<heading id="B2758186AC1143330B83F8E68D3B0106" bold="true" box="[151,862,1837,1864]" fontSize="11" level="2" pageId="72" pageNumber="73" reason="4">
<emphasis id="DBF6EAF8AC1143330B83F8E68D3B0106" bold="true" box="[151,862,1837,1864]" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330B83F8E68C8D0106" ID-CoL="5FLQ9" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" baseAuthorityName="Warncke" baseAuthorityYear="1967" box="[151,744,1837,1864]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="donata" status="stat. nov." subGenus="Truncandrena">
<emphasis id="DBF6EAF8AC1143330B83F8E68F640109" bold="true" box="[151,257,1837,1864]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330A06F8E68FA40109" bold="true" box="[274,449,1837,1864]" italics="true" pageId="72" pageNumber="73">Truncandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330AC6F8E68C400109" bold="true" box="[466,549,1837,1864]" italics="true" pageId="72" pageNumber="73">donata</emphasis>
<bibRefCitation id="8D134B1BAC1143330938F8E68C8D0106" author="Warncke, K." box="[556,744,1837,1863]" pageId="72" pageNumber="73" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC11433309FBF8E68D3B0106" box="[751,862,1837,1863]" pageId="72" pageNumber="73">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330B83F8B98ADD01CB" box="[151,1208,1906,1932]" pageId="72" pageNumber="73" type="reference_group">
<paragraph id="E93D36EAAC1143330B83F8B98ADD01CB" blockId="72.[151,1208,1906,1932]" box="[151,1208,1906,1932]" pageId="72" pageNumber="73">
<treatmentCitationGroup id="C99211C4AC1143330B83F8B98ADD01CB" box="[151,1208,1906,1932]" pageId="72" pageNumber="73">
<treatmentCitation id="682310FBAC1143330B83F8B98D1501CB" author="Warncke, K." box="[151,880,1906,1930]" page="227" pageId="72" pageNumber="73" year="1967">
<taxonomicName id="2E824D69AC1143330B83F8B98D1501CB" ID-CoL="5FLQ9" authority="Warncke, 1967: 227" authorityName="Warncke" authorityPageNumber="227" authorityYear="1967" box="[151,880,1906,1930]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="medeninensis" subGenus="Truncandrena" subSpecies="donata">
<emphasis id="DBF6EAF8AC1143330B83F8B98E8801CB" box="[151,237,1906,1930]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330BE8F8B98FEE01CB" box="[252,395,1906,1930]" italics="true" pageId="72" pageNumber="73">Truncandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330A8EF8B98C4701CB" box="[410,546,1906,1930]" italics="true" pageId="72" pageNumber="73">medeninensis</emphasis>
<materialsCitation id="59EA3CB7AC114333093CF8B98D1501CB" box="[552,880,1906,1930]" collectionCode="OOLM" country="Spain" location="Spain" pageId="72" pageNumber="73" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
ssp.
<emphasis id="DBF6EAF8AC1143330942F8B98CF901CB" box="[598,668,1906,1930]" italics="true" pageId="72" pageNumber="73">donata</emphasis>
<bibRefCitation id="8D134B1BAC11433309B6F8B98D1501CB" author="Warncke, K." box="[674,880,1906,1930]" pageId="72" pageNumber="73" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967: 227</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC114333086FF8B98DEE01C8" box="[891,907,1906,1929]" count="1" pageId="72" pageNumber="73" type="female"></specimenCount>
<specimenCount id="FF84FD63AC114333089EF8B98DC501C8" box="[906,928,1906,1929]" count="1" pageId="72" pageNumber="73" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC11433308B9F8B98D8E01CB" box="[941,1003,1906,1930]" name="Spain" pageId="72" pageNumber="73">Spain</collectingCountry>
, OÖLM, examined]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143320B83F8738A2F0171" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73" type="discussion">
<paragraph id="E93D36EAAC1143320B83F8738DC80700" blockId="72.[151,1436,1976,2039]" lastBlockId="73.[151,1437,150,321]" lastPageId="73" lastPageNumber="74" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330B83F8738F6E0193" bold="true" box="[151,267,1976,2002]" pageId="72" pageNumber="73">Remarks.</emphasis>
<bibRefCitation id="8D134B1BAC1143330A06F8738FD10193" author="Wood, T. J." box="[274,436,1976,2002]" pageId="72" pageNumber="73" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood (2023b)</bibRefCitation>
presented genetic data on the subgenus
<taxonomicName id="2E824D69AC1143330860F8738AA40193" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[884,1217,1976,2002]" pageId="72" pageNumber="73" rank="subGenus" subGenus="Truncandrena">
<emphasis id="DBF6EAF8AC1143330860F8738A770193" box="[884,1042,1976,2002]" italics="true" pageId="72" pageNumber="73">Truncandrena</emphasis>
Warncke, 1968
</taxonomicName>
with a focus on the Western Mediterranean. This work elevated one member of the species complex around
<taxonomicName id="2E824D69AC1143320F81F8178EAA06F0" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" lastPageId="73" lastPageNumber="74" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC1143330F81F8178B2B01B7" box="[1173,1358,2012,2038]" italics="true" pageId="72" pageNumber="73">A. medeninensis</emphasis>
Pérez, 1895
</taxonomicName>
(
<taxonomicName id="2E824D69AC1043320BF5FF5C8C6D06F0" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" box="[225,520,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="abunda">
<emphasis id="DBF6EAF8AC1043320BF5FF5C8F3006F0" box="[225,341,151,177]" italics="true" pageId="73" pageNumber="74">A. abunda</emphasis>
<bibRefCitation id="8D134B1BAC1043320A49FF5C8C6D06F0" author="Warncke, K." box="[349,520,151,177]" pageId="73" pageNumber="74" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
) but was unable to conclude on the status of
<taxonomicName id="2E824D69AC1043320F1BFF5C8B3506F0" authorityName="Warncke" authorityYear="1967" box="[1039,1360,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="medeninensis" subSpecies="donata">
<emphasis id="DBF6EAF8AC1043320F1BFF5C8AA306F0" box="[1039,1222,151,177]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
ssp.
<emphasis id="DBF6EAF8AC1043320E16FF5C8B3506F0" box="[1282,1360,151,177]" italics="true" pageId="73" pageNumber="74">donata</emphasis>
</taxonomicName>
which was described from
<collectingCountry id="9195767AAC1043320A63FF708FDD0694" box="[375,440,187,213]" name="Georgia" pageId="73" pageNumber="74">Iberia</collectingCountry>
due to a lack of genetic data. Such data are now available (
<figureCitation id="71B92A6FAC1043320F50FF708AD10694" box="[1092,1204,187,213]" captionStart="FIGURE 49" captionStartId="73.[151,250,1383,1407]" captionTargetBox="[151,1434,346,1358]" captionTargetId="figure-104@73.[151,1435,345,1360]" captionTargetPageId="73" captionText="FIGURE 49. Phylogenetic tree (maximum likelihood) of Andrena subgenus Truncandrena Warncke, 1968 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209411" httpUri="https://zenodo.org/record/13209411/files/figure.png" pageId="73" pageNumber="74">Figure 49</figureCitation>
), and clearly support species status for
<taxonomicName id="2E824D69AC1043320A73FF148FB206B8" authorityName="WOOD" authorityYear="2024" baseAuthorityName="Warncke" baseAuthorityYear="1967" box="[359,471,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="donata" status="stat. nov.">
<emphasis id="DBF6EAF8AC1043320A73FF148FB206B8" box="[359,471,223,249]" italics="true" pageId="73" pageNumber="74">A. donata</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC1043320AF5FF148C2A06B8" bold="true" box="[481,591,223,249]" pageId="73" pageNumber="74">
<taxonomicNameLabel id="C0C55783AC1043320AF5FF148C2A06B8" box="[481,591,223,249]" pageId="73" pageNumber="74">stat. nov.</taxonomicNameLabel>
</emphasis>
<taxonomicName id="2E824D69AC104332094DFF148D7506B8" authorityName="WOOD" authorityYear="2024" baseAuthorityName="Warncke" baseAuthorityYear="1967" box="[601,784,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="donata">
<emphasis id="DBF6EAF8AC104332094DFF148D7506B8" box="[601,784,223,249]" italics="true" pageId="73" pageNumber="74">Andrena donata</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="2E824D69AC10433208B9FF148A4106B8" authorityName="Warncke" authorityYear="1974" box="[941,1060,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="abunda">
<emphasis id="DBF6EAF8AC10433208B9FF148A4106B8" box="[941,1060,223,249]" italics="true" pageId="73" pageNumber="74">A. abunda</emphasis>
</taxonomicName>
, forming a clearly differentiated clade with bootstrap support of 98. The two species are separated by 9.58% (range 9.449.73%), and
<taxonomicName id="2E824D69AC1043320E00FEC88BE7075C" authorityName="WOOD" authorityYear="2024" baseAuthorityName="Warncke" baseAuthorityYear="1967" box="[1300,1410,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="donata">
<emphasis id="DBF6EAF8AC1043320E00FEC88BE7075C" box="[1300,1410,259,285]" italics="true" pageId="73" pageNumber="74">A. donata</emphasis>
</taxonomicName>
is separated from
<emphasis id="DBF6EAF8AC1043320A51FEEC8C470700" box="[325,546,295,321]" italics="true" pageId="73" pageNumber="74">
<taxonomicName id="2E824D69AC1043320A51FEEC8F9E0700" authorityName="Perez" authorityYear="1895" box="[325,507,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">A. medeninensis</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1043320916FEEC8C470700" box="[514,546,295,321]" pageId="73" pageNumber="74" sensu="lato">s.l.</taxonomicNameLabel>
</emphasis>
by 10.76% (range 10.0512.18%).
</paragraph>
<caption id="BDFD6662AC1043320B83FAAC8AB303E2" ID-DOI="http://doi.org/10.5281/zenodo.13209411" ID-Zenodo-Dep="13209411" httpUri="https://zenodo.org/record/13209411/files/figure.png" pageId="73" pageNumber="74" startId="73.[151,250,1383,1407]" targetBox="[151,1434,346,1358]" targetPageId="73" targetType="figure">
<paragraph id="E93D36EAAC1043320B83FAAC8AB303E2" blockId="73.[151,1436,1383,1444]" pageId="73" pageNumber="74">
<emphasis id="DBF6EAF8AC1043320B83FAAC8F40033E" bold="true" box="[151,293,1383,1407]" pageId="73" pageNumber="74">FIGURE 49.</emphasis>
Phylogenetic tree (maximum likelihood) of
<taxonomicName id="2E824D69AC1043320812FAAC8D39033E" baseAuthorityName="Popov" baseAuthorityYear="1960" box="[774,860,1383,1407]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC1043320812FAAC8D39033E" box="[774,860,1383,1407]" italics="true" pageId="73" pageNumber="74">Andrena</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="2E824D69AC10433208C5FAAC8B6C033E" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[977,1289,1383,1407]" pageId="73" pageNumber="74" rank="subGenus" subGenus="Truncandrena">
<emphasis id="DBF6EAF8AC10433208C5FAAC8A05033E" box="[977,1120,1383,1407]" italics="true" pageId="73" pageNumber="74">Truncandrena</emphasis>
Warncke, 1968
</taxonomicName>
based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted).
</paragraph>
</caption>
<paragraph id="E93D36EAAC1043320BD3FA1A8A2F0171" blockId="73.[151,1437,1489,2022]" pageId="73" pageNumber="74">
Morphologically,
<taxonomicName id="2E824D69AC1043320A84FA198F9803AA" authorityName="WOOD" authorityYear="2024" baseAuthorityName="Warncke" baseAuthorityYear="1967" box="[400,509,1489,1516]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="donata">
<emphasis id="DBF6EAF8AC1043320A84FA198F9803AA" box="[400,509,1489,1516]" italics="true" pageId="73" pageNumber="74">A. donata</emphasis>
</taxonomicName>
can be separated from
<emphasis id="DBF6EAF8AC1043320817FA198D9903AA" box="[771,1020,1489,1516]" italics="true" pageId="73" pageNumber="74">
<taxonomicName id="2E824D69AC1043320817FA198DDF03AA" authorityName="Perez" authorityYear="1895" box="[771,954,1489,1516]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">A. medeninensis</taxonomicName>
<taxonomicNameLabel id="C0C55783AC10433208D5FA198D9903AA" box="[961,1020,1490,1515]" pageId="73" pageNumber="74" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
in the female sex due to the stronger tergal punctation, head with a mixture of light brown pubescence with abundant black hairs along the inner margins of the compound eyes (in
<taxonomicName id="2E824D69AC1043320AD4F9D18C1D0072" authorityName="Perez" authorityYear="1895" box="[448,632,1561,1588]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC1043320AD4F9D18C1D0072" box="[448,632,1561,1588]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1043320994F9D18CDB0075" box="[640,702,1562,1588]" pageId="73" pageNumber="74" sensu="stricto">
<emphasis id="DBF6EAF8AC1043320994F9D18CDD0072" box="[640,696,1562,1587]" italics="true" pageId="73" pageNumber="74">s. str</emphasis>
.
</taxonomicNameLabel>
with uniform orange-red pubescence), clypeus with longitudinal impunctate mid-line comparatively shorted and less strongly produced (in
<taxonomicName id="2E824D69AC10433208C2F9F58AE90016" authorityName="Perez" authorityYear="1895" box="[982,1164,1597,1624]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC10433208C2F9F58AE90016" box="[982,1164,1597,1624]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1043320F86F9F58AAA0019" box="[1170,1231,1598,1624]" pageId="73" pageNumber="74" sensu="stricto">
<emphasis id="DBF6EAF8AC1043320F86F9F58AAC0016" box="[1170,1225,1598,1623]" italics="true" pageId="73" pageNumber="74">s. str</emphasis>
.
</taxonomicNameLabel>
with this mid-line long and strongly produced), scutum with intermixed light brown and black hairs (in
<taxonomicName id="2E824D69AC1043320F70F9A98B7E003A" authorityName="Perez" authorityYear="1895" box="[1124,1307,1633,1660]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC1043320F70F9A98B7E003A" box="[1124,1307,1633,1660]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1043320E37F9A98B07003D" box="[1315,1378,1634,1660]" pageId="73" pageNumber="74" sensu="stricto">
<emphasis id="DBF6EAF8AC1043320E37F9A98B39003A" box="[1315,1372,1634,1659]" italics="true" pageId="73" pageNumber="74">s. str</emphasis>
.
</taxonomicNameLabel>
with uniform orange-red hairs), T12 with long loose light brown hairs (in
<taxonomicName id="2E824D69AC104332088BF94D8A3100DE" authorityName="Perez" authorityYear="1895" box="[927,1108,1669,1696]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC104332088BF94D8A3100DE" box="[927,1108,1669,1696]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1043320F4EF94D8AF300E1" box="[1114,1174,1670,1696]" pageId="73" pageNumber="74" sensu="stricto">
<emphasis id="DBF6EAF8AC1043320F4EF94D8AF500DE" box="[1114,1168,1670,1695]" italics="true" pageId="73" pageNumber="74">s. str</emphasis>
.
</taxonomicNameLabel>
with sparse short hairs, almost without hairs), and T24 apically with narrow light brown hairbands (terga without apical hairbands). Males can be separated using the same pubescence characters, as well as the comparatively narrower genitalia capsule with comparatively narrower penis valves. Further study is required to establish the status of
<taxonomicName id="2E824D69AC1043320F8FF9398F260171" authority="Warncke, 1980" authorityName="Warncke" authorityYear="1980" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="subSpecies" species="medeninensis" subSpecies="tiznita">
<emphasis id="DBF6EAF8AC1043320F8FF9398BF9014A" box="[1179,1436,1777,1804]" italics="true" pageId="73" pageNumber="74">A. medeninensis tiznita</emphasis>
<bibRefCitation id="8D134B1BAC1043320B83F8DE8F260171" author="Warncke, K." box="[151,323,1813,1840]" pageId="73" pageNumber="74" pagination="65 - 87" refId="ref91868" refString="Warncke, K. (1980) Zur Verbreitung der Bienengattung Andrena F. in Tunesien. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 65 - 87." type="journal article" year="1980">Warncke, 1980</bibRefCitation>
</taxonomicName>
, but for now it is retained as a dark western form of
<taxonomicName id="2E824D69AC1043320899F8DD8A26016E" authorityName="Perez" authorityYear="1895" box="[909,1091,1813,1840]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="medeninensis">
<emphasis id="DBF6EAF8AC1043320899F8DD8A26016E" box="[909,1091,1813,1840]" italics="true" pageId="73" pageNumber="74">A. medeninensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1043310BD3F8F28D2C075C" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74" type="materials_examined">
<paragraph id="E93D36EAAC1043310BD3F8F28D2C075C" blockId="73.[151,1437,1489,2022]" lastBlockId="74.[151,1436,150,321]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<emphasis id="DBF6EAF8AC1043320BD3F8F28FDD0112" bold="true" box="[199,440,1849,1875]" pageId="73" pageNumber="74">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC1043320AD2F8F28C9A0139" collectingDate="2019-03-21" collectionCode="TJWC" collectorName="C. Figueiredo" country="Portugal" county="Alto Alentejo" location="Cerros" municipality="Reguengos" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC1043320AD2F8F28C080112" bold="true" box="[454,621,1849,1875]" pageId="73" pageNumber="74">
<collectingCountry id="9195767AAC1043320AD2F8F28C0D0112" box="[454,616,1849,1875]" name="Portugal" pageId="73" pageNumber="74">PORTUGAL</collectingCountry>
:
</emphasis>
<collectingCounty id="005C4E66AC104332096DF8F28D7E0112" box="[633,795,1849,1875]" pageId="73" pageNumber="74">Alto Alentejo</collectingCounty>
,
<collectingMunicipality id="0959AC90AC1043320839F8F18DC90115" box="[813,940,1850,1876]" pageId="73" pageNumber="74">Reguengos</collectingMunicipality>
,
<location id="EC5D6031AC10433208ABF8F18A690115" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC10433208ABF8F18A690115" box="[959,1036,1850,1876]" country="Portugal" county="Alto Alentejo" municipality="Reguengos" name="Cerros" pageId="73" pageNumber="74">Cerros</location>
, near to
<location id="EC5D6031AC1043320F95F8F18E930139" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1043320F95F8F18E930139" country="Portugal" county="Alto Alentejo" municipality="Reguengos" name="Restaurante Herdade do Esporao" pageId="73" pageNumber="74">Restaurante Herdade do Esporão</location>
,
<date id="9D3C102AAC1043320A12F8968FE60136" box="[262,387,1885,1911]" pageId="73" pageNumber="74" value="2019-03-21">
<collectingDate id="8D78E9C2AC1043320A12F8968FE60136" box="[262,387,1885,1911]" pageId="73" pageNumber="74" value="2019-03-21">21.iii.2019</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1043320A87F8958FD30139" box="[403,438,1886,1912]" count="1" pageId="73" pageNumber="74" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1043320914F8958CC70136" box="[512,674,1885,1912]" pageId="73" pageNumber="74">C. Figueiredo</collectorName>
,
<collectionCode id="8F93AE2FAC10433209A6F8958C9A0139" box="[690,767,1886,1912]" pageId="73" pageNumber="74">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1043320804F8968B530139" box="[784,1334,1885,1912]" country="Spain" location="Hispania" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC1043320804F8968D0E0136" bold="true" box="[784,875,1885,1911]" pageId="73" pageNumber="74">
<collectingCountry id="9195767AAC1043320804F8968D030136" box="[784,870,1885,1911]" name="Spain" pageId="73" pageNumber="74">SPAIN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC1043320863F8968DBE0136" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1043320863F8968DBE0136" box="[887,987,1885,1911]" country="Spain" name="Hispania" pageId="73" pageNumber="74">Hispania</location>
[no further information],
<specimenCount id="FF84FD63AC1043320E07F8958B530139" box="[1299,1334,1886,1912]" count="1" pageId="73" pageNumber="74" type="female">1♀</specimenCount>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC1043320E5EF8948CDB01DD" collectingDate="1900-04-11" country="Spain" location="Chamartin" pageId="73" pageNumber="74" specimenCount="1" specimenCount-male="1" stateProvince="Madrid" typeStatus="holotype">
OÖLM (
<emphasis id="DBF6EAF8AC1043320BB5F8498F6D01DD" bold="true" box="[161,264,1922,1948]" pageId="73" pageNumber="74">
<typeStatus id="36398848AC1043320BB5F8498F6D01DD" box="[161,264,1922,1948]" pageId="73" pageNumber="74" type="holotype">holotype</typeStatus>
</emphasis>
);
<location id="EC5D6031AC1043320A35F84A8FF201DA" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1043320A35F84A8FF201DA" box="[289,407,1921,1947]" country="Spain" name="Chamartin" pageId="73" pageNumber="74" stateProvince="Madrid">Chamartín</location>
[
<collectingRegion id="2B46F808AC1043320AB2F84A8C6401DA" box="[422,513,1921,1947]" country="Spain" name="Madrid" pageId="73" pageNumber="74">Madrid</collectingRegion>
],
<date id="9D3C102AAC1043320901F84A8CE901DA" box="[533,652,1921,1947]" pageId="73" pageNumber="74" value="1900-04-11">
<collectingDate id="8D78E9C2AC1043320901F84A8CE901DA" box="[533,652,1921,1947]" pageId="73" pageNumber="74" value="1900-04-11">11.iv.1900</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC104332098DF8498CDB01DD" box="[665,702,1922,1948]" count="1" pageId="73" pageNumber="74" type="male">1♂</specimenCount>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC10433209D8F8488E8E01FE" collectingDate="1900-04-09" collectorName="J. M. Dusmet &amp; Alonso" country="Spain" location="Madrid" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1" stateProvince="Madrid" typeStatus="paratype">
OÖLM (
<emphasis id="DBF6EAF8AC1043320824F8498DFE01DD" bold="true" box="[816,923,1922,1948]" pageId="73" pageNumber="74">
<typeStatus id="36398848AC1043320824F8498DFE01DD" box="[816,923,1922,1948]" pageId="73" pageNumber="74" type="paratype">paratype</typeStatus>
</emphasis>
);
<collectingRegion id="2B46F808AC10433208A2F84A8A6E01DA" box="[950,1035,1921,1947]" country="Spain" name="Madrid" pageId="73" pageNumber="74">Madrid</collectingRegion>
,
<date id="9D3C102AAC1043320F0CF84A8AE701DA" box="[1048,1154,1921,1947]" pageId="73" pageNumber="74" value="1900-04-09">
<collectingDate id="8D78E9C2AC1043320F0CF84A8AE701DA" box="[1048,1154,1921,1947]" pageId="73" pageNumber="74" value="1900-04-09">9.iv.1900</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1043320F9BF8498AD701DD" box="[1167,1202,1922,1948]" count="1" pageId="73" pageNumber="74" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1043320FE0F8498BE001DD" box="[1268,1413,1922,1948]" pageId="73" pageNumber="74">J.M. Dusmet</collectorName>
y
<collectorName id="4477533CAC1043320B83F86E8E8E01FE" box="[151,235,1957,1983]" pageId="73" pageNumber="74">Alonso</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC1043320BE1F86C8A5101FE" box="[245,1076,1957,1986]" collectingDate="1907-04-10" collectorName="J. M. Dusmet &amp; Alonso" country="Spain" location="Villaverde" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
OÖLM (
<emphasis id="DBF6EAF8AC1043320A42F86D8FA40181" bold="true" box="[342,449,1958,1984]" pageId="73" pageNumber="74">
<typeStatus id="36398848AC1043320A42F86D8FA40181" box="[342,449,1958,1984]" pageId="73" pageNumber="74" type="paratype">paratype</typeStatus>
</emphasis>
);
<location id="EC5D6031AC1043320ACCF86E8C2A01FE" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1043320ACCF86E8C2A01FE" box="[472,591,1957,1983]" country="Spain" name="Villaverde" pageId="73" pageNumber="74">Villaverde</location>
,
<date id="9D3C102AAC104332094DF86E8CB701FE" box="[601,722,1957,1983]" pageId="73" pageNumber="74" value="1907-04-10">
<collectingDate id="8D78E9C2AC104332094DF86E8CB701FE" box="[601,722,1957,1983]" pageId="73" pageNumber="74" value="1907-04-10">10.iv.1907</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC10433209CFF86D8C9B0181" box="[731,766,1958,1984]" count="1" pageId="73" pageNumber="74" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC104332082EF86D8DAD0181" box="[826,968,1958,1984]" pageId="73" pageNumber="74">J.M. Dusmet</collectorName>
y
<collectorName id="4477533CAC10433208F4F86E8A5101FE" box="[992,1076,1957,1983]" pageId="73" pageNumber="74">Alonso</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC1043320F2AF86C8C1901A2" collectingDate="1920-04-03" collectorName="J. M. Dusmet &amp; Alonso" country="Spain" location="Villaverde" pageId="73" pageNumber="74" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
OÖLM (
<emphasis id="DBF6EAF8AC1043320F8BF86D8B6F0181" bold="true" box="[1183,1290,1958,1984]" pageId="73" pageNumber="74">
<typeStatus id="36398848AC1043320F8BF86D8B6F0181" box="[1183,1290,1958,1984]" pageId="73" pageNumber="74" type="paratype">paratype</typeStatus>
</emphasis>
);
<location id="EC5D6031AC1043320E35F86E8BFD01FE" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1043320E35F86E8BFD01FE" box="[1313,1432,1957,1983]" country="Spain" name="Villaverde" pageId="73" pageNumber="74">Villaverde</location>
,
<date id="9D3C102AAC1043320B83F8028F6701A2" box="[151,258,1993,2019]" pageId="73" pageNumber="74" value="1920-04-03">
<collectingDate id="8D78E9C2AC1043320B83F8028F6701A2" box="[151,258,1993,2019]" pageId="73" pageNumber="74" value="1920-04-03">3.iv.1920</collectingDate>
</date>
,
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, leg.
<collectorName id="4477533CAC1043320A62F8018C6D01A5" box="[374,520,1994,2020]" pageId="73" pageNumber="74">J.M. Dusmet</collectorName>
y
<collectorName id="4477533CAC104332093CF8028C1901A2" box="[552,636,1993,2019]" pageId="73" pageNumber="74">Alonso</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC104331099DF8008F2306F0" collectingDate="1988-02-19" collectionCode="FJOS" collectorName="F. J. Ortiz-Sanchez" country="Spain" lastPageId="74" lastPageNumber="75" location="Granada" municipality="Cacin" pageId="73" pageNumber="74" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
OÖLM (
<emphasis id="DBF6EAF8AC10433209FBF8018D3F01A5" bold="true" box="[751,858,1994,2020]" pageId="73" pageNumber="74">
<typeStatus id="36398848AC10433209FBF8018D3F01A5" box="[751,858,1994,2020]" pageId="73" pageNumber="74" type="paratype">paratype</typeStatus>
</emphasis>
);
<collectingMunicipality id="0959AC90AC1043320860F8018DD001A5" box="[884,949,1994,2020]" pageId="73" pageNumber="74">Cacín</collectingMunicipality>
(
<location id="EC5D6031AC10433208D3F8028A4901A2" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC10433208D3F8028A4901A2" box="[967,1068,1993,2019]" country="Spain" municipality="Cacin" name="Granada" pageId="73" pageNumber="74">Granada</location>
),
<date id="9D3C102AAC1043320F55F8028AD001A2" box="[1089,1205,1993,2019]" pageId="73" pageNumber="74" value="1988-02-19">
<collectingDate id="8D78E9C2AC1043320F55F8028AD001A2" box="[1089,1205,1993,2019]" pageId="73" pageNumber="74" value="1988-02-19">19.ii.1988</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1043320FD7F8018A8D01A5" box="[1219,1256,1994,2020]" count="1" pageId="73" pageNumber="74" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC1043310E3EF8018E9206F0" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">F.J. Ortiz-Sánchez</collectorName>
,
<collectionCode id="8F93AE2FAC1343310A10FF5C8F2306F0" box="[260,326,151,177]" pageId="74" pageNumber="75">FJOS</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1343310A46FF5C8A9906F0" box="[338,1276,150,177]" collectingDate="2005-03-25" collectionCode="FJOS" collectorName="F. J. Ortiz-Sanchez" country="Spain" elevation="700" location="Ciudad Real" municipality="Fuencaliente" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingMunicipality id="0959AC90AC1343310A46FF5C8F8406F0" box="[338,481,151,177]" pageId="74" pageNumber="75">Fuencaliente</collectingMunicipality>
(
<location id="EC5D6031AC1343310AE5FF5C8CE606F0" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1343310AE5FF5C8CE606F0" box="[497,643,151,177]" country="Spain" municipality="Fuencaliente" name="Ciudad Real" pageId="74" pageNumber="75">Ciudad Real</location>
),
<quantity id="2E7A9B0FAC134331098CFF5C8C8606F0" box="[664,739,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="700.0">
<elevation id="62AFD1D9AC134331098CFF5C8C8606F0" box="[664,739,151,177]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="74" pageNumber="75" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="9D3C102AAC13433109E4FF5D8D0906F0" box="[752,876,150,177]" pageId="74" pageNumber="75" value="2005-03-25">
<collectingDate id="8D78E9C2AC13433109E4FF5D8D0906F0" box="[752,876,150,177]" pageId="74" pageNumber="75" value="2005-03-25">25.iii.2005</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC134331086CFF5C8DF806F0" box="[888,925,151,177]" count="1" pageId="74" pageNumber="75" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC13433108C8FF5C8ACB06F0" box="[988,1198,151,177]" pageId="74" pageNumber="75">F.J. Ortiz-Sánchez</collectorName>
,
<collectionCode id="8F93AE2FAC1343310FAEFF5C8A9906F0" box="[1210,1276,151,177]" pageId="74" pageNumber="75">FJOS</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1343310E1DFF5C8D570694" collectingDate="1989-05-09" collectionCode="FJOS" collectorName="A. Jimenez" country="Spain" county="Loja" location="Granada" municipality="Charco" pageId="74" pageNumber="75" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
<collectingCounty id="005C4E66AC1343310E1DFF5C8B5A06F0" box="[1289,1343,151,177]" pageId="74" pageNumber="75">Loja</collectingCounty>
,
<collectingMunicipality id="0959AC90AC1343310E5FFF5C8BF906F0" box="[1355,1436,151,177]" pageId="74" pageNumber="75">Charco</collectingMunicipality>
del negro (
<location id="EC5D6031AC1343310A07FF708F1D0694" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1343310A07FF708F1D0694" box="[275,376,187,213]" country="Spain" county="Loja" municipality="Charco" name="Granada" pageId="74" pageNumber="75">Granada</location>
),
<date id="9D3C102AAC1343310A99FF708F950694" box="[397,496,187,213]" pageId="74" pageNumber="75" value="1989-05-09">
<collectingDate id="8D78E9C2AC1343310A99FF708F950694" box="[397,496,187,213]" pageId="74" pageNumber="75" value="1989-05-09">9.v.1989</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1343310AE9FF708C440694" box="[509,545,187,213]" count="6" pageId="74" pageNumber="75" type="female">6♀</specimenCount>
, leg.
<collectorName id="4477533CAC1343310974FF708C810694" box="[608,740,187,213]" pageId="74" pageNumber="75">A. Jiménez</collectorName>
,
<collectionCode id="8F93AE2FAC13433109E4FF708D570694" box="[752,818,187,213]" pageId="74" pageNumber="75">FJOS</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC134331082BFF708F5D06B8" collectingDate="2023-03-11" collectionCode="TJWC" collectorName="J. I. Pascal" country="Spain" location="Laguna de Ambroz" pageId="74" pageNumber="75" specimenCount="1" specimenCount-male="1" stateProvince="Madrid" typeStatus="paratype">
<collectingRegion id="2B46F808AC134331082BFF708DF10694" box="[831,916,187,213]" country="Spain" name="Madrid" pageId="74" pageNumber="75">Madrid</collectingRegion>
,
<location id="EC5D6031AC13433108B6FF708AE50694" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC13433108B6FF708AE50694" box="[930,1152,187,213]" country="Spain" name="Laguna de Ambroz" pageId="74" pageNumber="75" stateProvince="Madrid">Laguna de Ambroz</location>
,
<date id="9D3C102AAC1343310F98FF708B620694" box="[1164,1287,187,213]" pageId="74" pageNumber="75" value="2023-03-11">
<collectingDate id="8D78E9C2AC1343310F98FF708B620694" box="[1164,1287,187,213]" pageId="74" pageNumber="75" value="2023-03-11">11.iii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1343310E01FF708B5C0694" box="[1301,1337,187,213]" count="1" pageId="74" pageNumber="75" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC1343310E6EFF708E8406B8" pageId="74" pageNumber="75">J.I. Pascal</collectorName>
,
<collectionCode id="8F93AE2FAC1343310BFFFF148F5D06B8" box="[235,312,223,249]" pageId="74" pageNumber="75">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1343310A56FF148A8C06B8" box="[322,1257,223,251]" collectingDate="2023-03-12" collectionCode="TJWC" collectorName="El Pardo &amp; P. Alvarez Fidalgo" country="Spain" location="Mingorrubio" pageId="74" pageNumber="75" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Madrid" typeStatus="paratype">
<collectingRegion id="2B46F808AC1343310A56FF148FF206B8" box="[322,407,223,249]" country="Spain" name="Madrid" pageId="74" pageNumber="75">Madrid</collectingRegion>
,
<location id="EC5D6031AC1343310AB6FF148C5106B8" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1343310AB6FF148C5106B8" box="[418,564,223,249]" country="Spain" name="Mingorrubio" pageId="74" pageNumber="75" stateProvince="Madrid">Mingorrubio</location>
,
<collectorName id="4477533CAC134331092AFF148CC406B8" box="[574,673,223,249]" pageId="74" pageNumber="75">El Pardo</collectorName>
,
<date id="9D3C102AAC13433109B8FF148D4206B8" box="[684,807,223,249]" pageId="74" pageNumber="75" value="2023-03-12">
<collectingDate id="8D78E9C2AC13433109B8FF148D4206B8" box="[684,807,223,249]" pageId="74" pageNumber="75" value="2023-03-12">12.iii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1343310826FF148D3206B8" box="[818,855,223,249]" count="3" pageId="74" pageNumber="75" type="male">3♂</specimenCount>
,
<specimenCount id="FF84FD63AC1343310876FF148DE306B8" box="[866,902,223,249]" count="3" pageId="74" pageNumber="75" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC13433108D6FF148AF706B8" box="[962,1170,223,251]" pageId="74" pageNumber="75">P. Álvarez Fidalgo</collectorName>
,
<collectionCode id="8F93AE2FAC1343310F88FF148A8C06B8" box="[1180,1257,223,249]" pageId="74" pageNumber="75">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1343310FE7FF148D20075C" collectingDate="2023-03-16" collectionCode="TJWC" collectorName="P. Alvarez Fidalgo" country="Spain" location="Vaciamadrid" pageId="74" pageNumber="75" specimenCount="3" specimenCount-female="3" stateProvince="Madrid" typeStatus="paratype">
<collectingRegion id="2B46F808AC1343310FE7FF148B2D06B8" box="[1267,1352,223,249]" country="Spain" name="Madrid" pageId="74" pageNumber="75">Madrid</collectingRegion>
,
<location id="EC5D6031AC1343310E47FF148F4D075C" LSID="urn:lsid:plazi:treatment:612B87FCAC1143310B83F8E68C880700:EC5D6031AC1343310E47FF148F4D075C" country="Spain" name="Vaciamadrid" pageId="74" pageNumber="75" stateProvince="Madrid">Rivas-Vaciamadrid</location>
,
<date id="9D3C102AAC1343310A27FEC88FCB075C" box="[307,430,259,285]" pageId="74" pageNumber="75" value="2023-03-16">
<collectingDate id="8D78E9C2AC1343310A27FEC88FCB075C" box="[307,430,259,285]" pageId="74" pageNumber="75" value="2023-03-16">16.iii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1343310AAEFEC88FBB075C" box="[442,478,259,285]" count="3" pageId="74" pageNumber="75" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC134331090FFEC88C88075C" box="[539,749,259,287]" pageId="74" pageNumber="75">P. Álvarez Fidalgo</collectorName>
,
<collectionCode id="8F93AE2FAC13433109ECFEC88D20075C" box="[760,837,259,285]" pageId="74" pageNumber="75">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1343310BD3FEED8C880700" box="[199,749,294,321]" pageId="74" pageNumber="75" type="distribution">
<paragraph id="E93D36EAAC1343310BD3FEED8C880700" blockId="74.[151,1436,150,321]" box="[199,749,294,321]" pageId="74" pageNumber="75">
<emphasis id="DBF6EAF8AC1343310BD3FEED8F040701" bold="true" box="[199,353,294,320]" pageId="74" pageNumber="75">Distribution.</emphasis>
<collectingCountry id="9195767AAC1343310A73FEEC8FC20700" box="[359,423,295,321]" name="Spain" pageId="74" pageNumber="75">Spain</collectingCountry>
and
<collectingCountry id="9195767AAC1343310ACAFEEC8C580700" box="[478,573,295,321]" name="Portugal" pageId="74" pageNumber="75">Portugal</collectingCountry>
(
<bibRefCitation id="8D134B1BAC134331095FFEEC8C840700" author="Wood, T. J." box="[587,737,295,321]" pageId="74" pageNumber="75" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,411 @@
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<emphasis id="DBF6EAF8AC1143330B83FEA58D0307C8" bold="true" box="[151,870,366,393]" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330B83FEA58C9007C9" ID-CoL="DSMC" authority="Noskiewicz, 1939" authorityName="Noskiewicz" authorityYear="1939" box="[151,757,366,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" status="stat. nov." subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC1143330B83FEA58F6407C8" bold="true" box="[151,257,366,393]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330A06FEA58FD707C8" bold="true" box="[274,434,366,393]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330AD7FEA58C7D07C8" bold="true" box="[451,536,366,393]" italics="true" pageId="72" pageNumber="73">pillichi</emphasis>
<bibRefCitation id="8D134B1BAC114333090BFEA58C9007C9" author="Noskiewicz, J." box="[543,757,366,392]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz, 1939</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC11433309E8FEA48D0307C8" box="[764,870,367,393]" pageId="72" pageNumber="73">stat. rev.</taxonomicNameLabel>
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<paragraph id="E93D36EAAC1143330B83FE788D3E078A" blockId="72.[151,859,435,460]" box="[151,859,435,460]" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330B83FE788FBD078A" authorityName="Noskiewicz" authorityYear="1939" box="[151,472,435,459]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC1143330B83FE788E88078A" box="[151,237,435,459]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1143330BE8FE788FE4078A" box="[252,385,435,459]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1143330A84FE788FBD078A" box="[400,472,435,459]" italics="true" pageId="72" pageNumber="73">pillichi</emphasis>
</taxonomicName>
<materialsCitation id="59EA3CB7AC1143330ACAFE7F8D3E078A" box="[478,859,435,460]" collectionCode="UWCP" country="Hungary" location="Hungary" pageId="72" pageNumber="73" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="FF84FD63AC1143330ACAFE7F8F8B078A" box="[478,494,436,459]" count="1" pageId="72" pageNumber="73" type="female"></specimenCount>
<specimenCount id="FF84FD63AC1143330AF9FE7F8C66078A" box="[493,515,436,459]" count="1" pageId="72" pageNumber="73" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC1143330904FE7F8C09078D" box="[528,620,436,460]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
,
<collectionCode id="8F93AE2FAC1143330962FE788CDA078A" box="[630,703,435,459]" country="Poland" name="University of Wroclaw" pageId="72" pageNumber="73">UWCP</collectionCode>
, not examined]
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330B83FE328A85036A" pageId="72" pageNumber="73" type="discussion">
<paragraph id="E93D36EAAC1143330B83FE328DEF053A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330B83FE328F6E0452" bold="true" box="[151,267,505,531]" pageId="72" pageNumber="73">Remarks.</emphasis>
<taxonomicName id="2E824D69AC1143330A00FE328FA90452" authorityName="Noskiewicz" authorityYear="1939" box="[276,460,505,531]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330A00FE328FA90452" box="[276,460,505,531]" italics="true" pageId="72" pageNumber="73">Andrena pillichi</emphasis>
</taxonomicName>
was described from Simontornya in
<collectingCountry id="9195767AAC114333086DFE318DB90455" box="[889,988,506,532]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, collected by F. Pillich who sent much material from this region to other workers, leading to the descriptions of multiple
<taxonomicName id="2E824D69AC1143330F44FDD68B6E0476" box="[1104,1291,541,567]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330F44FDD68ACA0476" box="[1104,1199,541,567]" italics="true" pageId="72" pageNumber="73">Andrena</emphasis>
species
</taxonomicName>
(e.g.
<bibRefCitation id="8D134B1BAC1143330E5AFDD68EB7041A" author="Alfken, J. D." pageId="72" pageNumber="73" pagination="88 - 93" refId="ref87099" refString="Alfken, J. D. (1933) Beitrage zur Kenntnis palaarktischer Bienen (Hym. Apid.). 3. Beitrag. Mitteilungen der Deutschen Entomologischen Gesellschaft, 4, 88 - 93. https: // doi. org / 10.1002 / mmnd. 193319330105" type="journal article" year="1933">Alfken 1933</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1143330BF4FD8A8F7E041A" author="Alfken, J. D." box="[224,283,577,603]" pageId="72" pageNumber="73" pagination="377 - 383" refId="ref87148" refString="Alfken, J. D. (1936) Neue Andrena-Arten vom Wiener Becken, aus dem Burgenland und Ungarn. Veroffentlichungen aus dem Uberseemuseum Bremen, 1, 377 - 383." type="journal article" year="1936">1936</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1143330A3DFD8A8F97041A" author="Noskiewicz, J." box="[297,498,577,603]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz 1939</bibRefCitation>
). Noskiewicz gave detailed identification criteria to allow separation of
<taxonomicName id="2E824D69AC1143330E38FD898BF9041A" authorityName="Noskiewicz" authorityYear="1939" box="[1324,1436,577,604]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330E38FD898BF9041A" box="[1324,1436,577,604]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
from
<taxonomicName id="2E824D69AC1143330BC0FDAD8F05043E" authorityName="Perkins" authorityYear="1914" box="[212,352,613,640]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330BC0FDAD8F05043E" box="[212,352,613,640]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
; both species are bivoltine, but
<taxonomicName id="2E824D69AC11433309D6FDAD8D4A043E" authorityName="Noskiewicz" authorityYear="1939" box="[706,815,613,640]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC11433309D6FDAD8D4A043E" box="[706,815,613,640]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
is noticeably shinier on the clypeus, scutum, scutellum, and tergal discs. Detailed identification criteria are given by
<bibRefCitation id="8D134B1BAC1143330852FD428A7A04E2" author="Noskiewicz, J." box="[838,1055,649,675]" pageId="72" pageNumber="73" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz (1939)</bibRefCitation>
,
<bibRefCitation id="8D134B1BAC1143330F38FD428B6504E2" author="Osytshnjuk, A. Z." box="[1068,1280,649,675]" pageId="72" pageNumber="73" pagination="315 - 369" refId="ref89594" refString="Osytshnjuk, A. Z. (1978) The identification of the insects of the European part of the USSR. Vol. 3. Hymenoptera. Part one. Opred Faune SSSR, 119, 315 - 369. [in Russian]" type="journal article" year="1978">Osytshnjuk (1978)</bibRefCitation>
, and
<bibRefCitation id="8D134B1BAC1143330E29FD428FD90486" author="Schmid-Egger, C. &amp; Scheuchl, E." pageId="72" pageNumber="73" refId="ref90781" refString="Schmid-Egger, C. &amp; Scheuchl, E. (1997) Illustrierte Bestimmungstabellen der Wildbienen Deutschlands und Osterreichs und Berucksichtigung der Arten der Schweiz. Band III. Andrenidae. Velden, Eigenverlag, 180 pp." type="book" year="1997">Schmid-Egger &amp; Scheuchl (1997)</bibRefCitation>
.
<bibRefCitation id="8D134B1BAC1143330AD3FD668C190486" author="Warncke, K." box="[455,636,685,711]" pageId="72" pageNumber="73" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
considered
<taxonomicName id="2E824D69AC1143330812FD658D100486" authorityName="Noskiewicz" authorityYear="1939" box="[774,885,685,712]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330812FD658D100486" box="[774,885,685,712]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
to be a valid species, but
<bibRefCitation id="8D134B1BAC1143330F89FD668B3C0486" author="Dylewska, M." box="[1181,1369,685,711]" pageId="72" pageNumber="73" pagination="359 - 708" refId="ref88522" refString="Dylewska, M. (1987) Die Gattung Andrena Fabricius (Andrenidae, Apoidea) in Nord- und Mitteleuropa. Acta Zoologica Cracoviensia, 30, 359 - 708." type="journal article" year="1987">Dylewska (1987</bibRefCitation>
, as
<taxonomicName id="2E824D69AC1143330E90FD658EBA04AA" authorityName="Noskiewicz" authorityYear="1939" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pilichi">
<emphasis id="DBF6EAF8AC1143330E90FD658EBA04AA" italics="true" pageId="72" pageNumber="73">A. pilichi</emphasis>
</taxonomicName>
subsequent incorrect spelling) and
<bibRefCitation id="8D134B1BAC1143330979FD1A8D3B04AA" author="Schwarz, M. &amp; Gusenleitner, F. &amp; Westrich, P. &amp; Dathe, H. H." box="[621,862,721,748]" pageId="72" pageNumber="73" pagination="1 - 398" refId="ref90955" refString="Schwarz, M., Gusenleitner, F., Westrich, P. &amp; Dathe, H. H. (1996) Katalog der Bienen Osterreichs, Deutschlands und der Schweiz. (Hymenoptera, Apidae). Entomofauna, Supplement 8, 1 - 398." type="journal article" year="1996">
Schwarz
<emphasis id="DBF6EAF8AC11433309C0FD198D6304AA" box="[724,774,721,747]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
. (1996)
</bibRefCitation>
synonymised it with
<taxonomicName id="2E824D69AC1143330F44FD198AB804AA" authorityName="Perkins" authorityYear="1914" box="[1104,1245,721,748]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330F44FD198AB804AA" box="[1104,1245,721,748]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
. Dylewska (who revised the
<typeStatus id="36398848AC1143330A08FD3D8F290551" box="[284,332,758,784]" pageId="72" pageNumber="73">type</typeStatus>
series) argued that some of the distinguishing characters given by Noskiewicz were insufficient, such as the antennal ratio of the males, the identical genital capsule, and variation in the degree of polish on the clypeus and scutellum. Schwarz
<emphasis id="DBF6EAF8AC114333091CFCF58C5F0516" box="[520,570,829,855]" italics="true" pageId="72" pageNumber="73">et al</emphasis>
. also argued that, based on a study of these two species including specimens from the Warncke collection, they could not consider them distinct.
</paragraph>
<paragraph id="E93D36EAAC1143330BD3FC4E8A85036A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
Fieldwork in
<collectingCountry id="9195767AAC1143330A4BFC4E8FA105DE" box="[351,452,901,927]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
produced long series of
<emphasis id="DBF6EAF8AC11433309C8FC4D8D8105DE" box="[732,996,901,928]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC11433309C8FC4D8D0F05DE" authorityName="Perkins" authorityYear="1914" box="[732,874,901,928]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">A. alfkenella</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330866FC4D8D8105DE" box="[882,996,901,927]" pageId="72" pageNumber="73" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
specimens. From two sites around
<collectingRegion id="2B46F808AC1143330E60FC4C8BF905E3" box="[1396,1436,903,930]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
in north-eastern
<collectingCountry id="9195767AAC1143330A42FC628FDB0582" box="[342,446,937,963]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
, Ciurea (open fallow agricultural fields) and Ursoaia (open woody steppe grassland), seven sequences of
<taxonomicName id="2E824D69AC1143330A66FC058C3105A6" box="[370,596,973,1000]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330A66FC058C3105A6" box="[370,596,973,1000]" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
were generated which showed low average intraspecific variation of 0.17% (
<figureCitation id="71B92A6FAC1143330B8BFC3A8F77024A" box="[159,274,1009,1035]" captionStart="FIGURE 37" captionStartId="57.[151,250,1819,1843]" captionTargetBox="[151,1433,311,1794]" captionTargetId="figure-78@57.[151,1435,309,1796]" captionTargetPageId="57" captionText="FIGURE 37. Phylogenetic tree (maximum likelihood) of Andrena subgenus Micrandrena Ashmead, 1899 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209387" httpUri="https://zenodo.org/record/13209387/files/figure.png" pageId="72" pageNumber="73">Figure 37</figureCitation>
). At the Ciurea site, a single Romanian specimen of
<taxonomicName id="2E824D69AC1143330872FC398D91024A" authorityName="Perkins" authorityYear="1914" box="[870,1012,1009,1036]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC1143330872FC398D91024A" box="[870,1012,1009,1036]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC11433308E8FC398A5D024D" box="[1020,1080,1010,1036]" pageId="72" pageNumber="73" sensu="stricto">s. str.</taxonomicNameLabel>
was found in sympatry with
<taxonomicName id="2E824D69AC1143330E90FC398F3B026E" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330E90FC398F3B026E" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
. This specimen was separated from the
<taxonomicName id="2E824D69AC1143330827FBDD8A7E026E" box="[819,1051,1045,1072]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC1143330827FBDD8A7E026E" box="[819,1051,1045,1072]" italics="true" pageId="72" pageNumber="73">A. alfkenella pillichi</emphasis>
</taxonomicName>
sequences by an average genetic distance of 3.76% (range 3.653.95%). More broadly, when including
<taxonomicName id="2E824D69AC11433308B8FBF18A5C0212" authorityName="Perkins" authorityYear="1914" box="[940,1081,1081,1108]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC11433308B8FBF18A5C0212" box="[940,1081,1081,1108]" italics="true" pageId="72" pageNumber="73">A. alfkenella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330F54FBF18A1E0215" box="[1088,1147,1082,1108]" pageId="72" pageNumber="73" sensu="stricto">s. str.</taxonomicNameLabel>
sequences from
<collectingCountry id="9195767AAC1143330E22FBF28BFD0212" box="[1334,1432,1081,1107]" name="Portugal" pageId="72" pageNumber="73">Portugal</collectingCountry>
,
<collectingCountry id="9195767AAC1143330B83FB968EBF0236" box="[151,218,1117,1143]" name="Spain" pageId="72" pageNumber="73">Spain</collectingCountry>
,
<collectingCountry id="9195767AAC1143330BF7FB968F290236" box="[227,332,1117,1143]" name="Germany" pageId="72" pageNumber="73">Germany</collectingCountry>
, and
<collectingCountry id="9195767AAC1143330A90FB968FB60236" box="[388,467,1117,1143]" name="Greece" pageId="72" pageNumber="73">Greece</collectingCountry>
(
<collectingRegion id="2B46F808AC1143330AF4FB968C440236" box="[480,545,1117,1143]" country="Greece" name="Kriti" pageId="72" pageNumber="73">Crete</collectingRegion>
, see above), the two taxa were separated by an average genetic distance of 4.08% (range 3.654.56%). Since the two taxa can be found in direct sympatry with consistent genetic differentiation and since their morphology is consistent with the criteria identified by Noskiewicz (with the caveat that some morphological variation is typical in members of the subgenus
<taxonomicName id="2E824D69AC114333086EFB028A6802A2" authorityName="Ashmead" authorityYear="1899" box="[890,1037,1225,1251]" pageId="72" pageNumber="73" rank="subGenus" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC114333086EFB028A6802A2" box="[890,1037,1225,1251]" italics="true" pageId="72" pageNumber="73">Micrandrena</emphasis>
</taxonomicName>
, particularly bivoltine species),
<taxonomicName id="2E824D69AC1143330E90FB018E830346" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi" status="stat. rev.">
<emphasis id="DBF6EAF8AC1143330E90FB018E830346" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC1143330BFAFB268F360346" bold="true" box="[238,339,1261,1287]" pageId="72" pageNumber="73">
<taxonomicNameLabel id="C0C55783AC1143330BFAFB268F360346" box="[238,339,1261,1287]" pageId="72" pageNumber="73">stat. rev.</taxonomicNameLabel>
</emphasis>
is returned to species status. Revision of European material of
<emphasis id="DBF6EAF8AC1143330F0CFB258B7B0346" box="[1048,1310,1261,1288]" italics="true" pageId="72" pageNumber="73">
<taxonomicName id="2E824D69AC1143330F0CFB258AC30346" authorityName="Perkins" authorityYear="1914" box="[1048,1190,1261,1288]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="alfkenella">A. alfkenella</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1143330FB9FB258B7B0346" box="[1197,1310,1261,1287]" pageId="72" pageNumber="73" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
is required to establish the precise range limits, though
<taxonomicName id="2E824D69AC1143330990FAD98C97036A" authorityName="WOOD" authorityYear="2024" box="[644,754,1297,1324]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="72" pageNumber="73" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC1143330990FAD98C97036A" box="[644,754,1297,1324]" italics="true" pageId="72" pageNumber="73">A. pillichi</emphasis>
</taxonomicName>
appears to be a strongly Pannonian species.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330BD3FAFE8F57006B" pageId="72" pageNumber="73" type="materials_examined">
<paragraph id="E93D36EAAC1143330BD3FAFE8F57006B" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330BD3FAFE8FD4030E" bold="true" box="[199,433,1333,1359]" pageId="72" pageNumber="73">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC1143330AA3FAFE8BFD0313" box="[439,1432,1333,1362]" collectingDate="2009-07-22" collectorName="J. Halada" country="Austria" location="1 km E Poysdorf" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Niederoesterreich">
<emphasis id="DBF6EAF8AC1143330AA3FAFE8C5B030E" bold="true" box="[439,574,1333,1359]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330AA3FAFE8C5C030E" box="[439,569,1333,1359]" name="Austria" pageId="72" pageNumber="73">AUSTRIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC1143330951FAFE8D66030E" box="[581,771,1333,1359]" country="Austria" name="Niederoesterreich" pageId="72" pageNumber="73">Niederosterreich</collectingRegion>
,
<location id="EC5D6031AC114333081AFAFE8DAA030E" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC114333081AFAFE8DAA030E" box="[782,975,1333,1360]" country="Austria" name="1 km E Poysdorf" pageId="72" pageNumber="73" stateProvince="Niederoesterreich">
<quantity id="2E7A9B0FAC114333081AFAFE8D2D030E" box="[782,840,1333,1359]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="km" value="1.0">1 km</quantity>
E Poysdorf
</location>
,
<date id="9D3C102AAC11433308CFFAFE8A38030E" box="[987,1117,1333,1359]" pageId="72" pageNumber="73" value="2009-07-22">
<collectingDate id="8D78E9C2AC11433308CFFAFE8A38030E" box="[987,1117,1333,1359]" pageId="72" pageNumber="73" value="2009-07-22">22.vii.2009</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F7DFAFE8AE9030E" box="[1129,1164,1333,1359]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330FDFFAFD8B53030E" box="[1227,1334,1333,1360]" pageId="72" pageNumber="73">J. Halada</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FA928BFD0337" box="[151,1432,1369,1398]" collectingDate="2003-04-05" collectingDateMax="2003-04-10" collectingDateMin="2003-04-05" collectorName="Y. Budaschkin" country="Austria" county="Krim" location="Vodianja" municipality="Karadagh" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC1143330B83FA928F700332" bold="true" box="[151,277,1369,1395]" pageId="72" pageNumber="73">CRIMEA:</emphasis>
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,
<collectingMunicipality id="0959AC90AC1143330A79FA928FBF0332" box="[365,474,1369,1395]" pageId="72" pageNumber="73">Karadagh</collectingMunicipality>
[Kara Dag],
<location id="EC5D6031AC1143330963FA928CBF0332" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330963FA928CBF0332" box="[631,730,1369,1395]" country="Austria" county="Krim" municipality="Karadagh" name="Vodianja" pageId="72" pageNumber="73">Vodianja</location>
balka,
<location id="EC5D6031AC1143330827FA928D150332" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330827FA928D150332" box="[819,880,1369,1395]" country="Austria" county="Krim" municipality="Karadagh" name="Wald" pageId="72" pageNumber="73">Wald</location>
,
<date id="9D3C102AAC114333086BFA928A760335" box="[895,1043,1369,1396]" pageId="72" pageNumber="73" value="2003-04-05" valueMax="2003-04-10" valueMin="2003-04-05">
<collectingDate id="8D78E9C2AC114333086BFA928A760335" box="[895,1043,1369,1396]" pageId="72" pageNumber="73" value="2003-04-05" valueMax="2003-04-10" valueMin="2003-04-05">510.iv.2003</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F37FA928A200332" box="[1059,1093,1369,1395]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330F9EFA918B560332" box="[1162,1331,1369,1396]" pageId="72" pageNumber="73">Y. Budaschkin</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FAB68DCA03DB" box="[151,943,1405,1434]" collectingDate="1941-08-17" collectorName="V. Zavadil" country="Czech Republic" location="Cejc" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Jihomoravsky">
<emphasis id="DBF6EAF8AC1143330B83FAB68F4603D6" bold="true" box="[151,291,1405,1431]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330B83FAB68F7B03D6" box="[151,286,1405,1431]" name="Czech Republic" pageId="72" pageNumber="73">CZECHIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC1143330A3FFAB68FE803D6" box="[299,397,1405,1431]" country="Czech Republic" name="Jihomoravsky" pageId="72" pageNumber="73">Moravia</collectingRegion>
,
<location id="EC5D6031AC1143330A8DFAB48FAA03DB" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330A8DFAB48FAA03DB" box="[409,463,1407,1434]" country="Czech Republic" name="Cejc" pageId="72" pageNumber="73" stateProvince="Jihomoravsky">Čejč</location>
,
<date id="9D3C102AAC1143330AC8FAB68C0303D6" box="[476,614,1405,1431]" pageId="72" pageNumber="73" value="1941-08-17">
<collectingDate id="8D78E9C2AC1143330AC8FAB68C0303D6" box="[476,614,1405,1431]" pageId="72" pageNumber="73" value="1941-08-17">17.viii.1941</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330966FAB68CF003D6" box="[626,661,1405,1431]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC11433309C1FAB58D2803D6" box="[725,845,1405,1432]" pageId="72" pageNumber="73">V. Zavadil</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC11433308A8FAB68F1403FF" collectingDate="1941-08-12" collectorName="V. Zavadil" country="Czech Republic" location="Cejkovice" pageId="72" pageNumber="73" specimenCount="1" specimenCount-male="1" stateProvince="Jihomoravsky">
<collectingRegion id="2B46F808AC11433308A8FAB68A7B03D6" box="[956,1054,1405,1431]" country="Czech Republic" name="Jihomoravsky" pageId="72" pageNumber="73">Moravia</collectingRegion>
,
<location id="EC5D6031AC1143330F3EFAB48AFA03DB" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330F3EFAB48AFA03DB" box="[1066,1183,1407,1434]" country="Czech Republic" name="Cejkovice" pageId="72" pageNumber="73" stateProvince="Jihomoravsky">Čejkovice</location>
,
<date id="9D3C102AAC1143330FBEFAB68B5103D6" box="[1194,1332,1405,1431]" pageId="72" pageNumber="73" value="1941-08-12">
<collectingDate id="8D78E9C2AC1143330FBEFAB68B5103D6" box="[1194,1332,1405,1431]" pageId="72" pageNumber="73" value="1941-08-12">12.viii.1941</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330E55FAB68B0003D6" box="[1345,1381,1405,1431]" count="1" pageId="72" pageNumber="73" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC1143330B83FA698F6A03FA" box="[151,271,1441,1468]" pageId="72" pageNumber="73">V. Zavadil</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330A69FA6A8BFD03FA" box="[381,1432,1441,1470]" collectingDate="2023-08-23" collectionCode="TJWC" collectorName="T. J. Wood" country="Romania" location="1 km NNW Ciurea" pageId="72" pageNumber="73" specimenCount="4" specimenCount-female="4" stateProvince="Iasi">
<emphasis id="DBF6EAF8AC1143330A69FA6A8C7703FA" bold="true" box="[381,530,1441,1467]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330A69FA6A8C6803FA" box="[381,525,1441,1467]" name="Romania" pageId="72" pageNumber="73">ROMANIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC114333090EFA688C2003FF" box="[538,581,1443,1470]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<location id="EC5D6031AC1143330945FA6A8D4E03FA" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330945FA6A8D4E03FA" box="[593,811,1441,1468]" country="Romania" name="1 km NNW Ciurea" pageId="72" pageNumber="73" stateProvince="Iasi">
<quantity id="2E7A9B0FAC1143330945FA6A8CEE03FA" box="[593,651,1441,1467]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="72" pageNumber="73" unit="km" value="1.0">1 km</quantity>
NNW Ciurea
</location>
, fallow fields,
<date id="9D3C102AAC11433308C7FA6A8A3803FA" box="[979,1117,1441,1467]" pageId="72" pageNumber="73" value="2023-08-23">
<collectingDate id="8D78E9C2AC11433308C7FA6A8A3803FA" box="[979,1117,1441,1467]" pageId="72" pageNumber="73" value="2023-08-23">23.viii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330F7DFA6A8AEB03FA" box="[1129,1166,1441,1467]" count="4" pageId="72" pageNumber="73" type="female">4♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330FD8FA698B2503FA" box="[1228,1344,1441,1468]" pageId="72" pageNumber="73">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC1143330E5FFA6A8BFD03FA" box="[1355,1432,1441,1467]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330B83FA0C8A90039E" box="[151,1269,1477,1506]" collectingDate="2023-07-21" collectionCode="TJWC" collectorName="T. J. Wood" country="Romania" county="Ursoaia" elevation="500" location="Dumnezeu" municipality="Dealul" pageId="72" pageNumber="73" specimenCount="15" specimenCount-female="5" specimenCount-male="10" stateProvince="Iasi">
<collectingRegion id="2B46F808AC1143330B83FA0C8EA703A3" box="[151,194,1479,1506]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<collectingRegion id="2B46F808AC1143330BDBFA0C8E9F03A3" box="[207,250,1479,1506]" country="Romania" name="Iasi" pageId="72" pageNumber="73">Iași</collectingRegion>
,
<quantity id="2E7A9B0FAC1143330A12FA0E8F2B03A1" box="[262,334,1477,1504]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="72" pageNumber="73" unit="m" value="500.0">
<elevation id="62AFD1D9AC1143330A12FA0E8F2B03A1" box="[262,334,1477,1504]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="72" pageNumber="73" unit="m" value="500.0">500 m</elevation>
</quantity>
E of
<collectingCounty id="005C4E66AC1143330A9BFA0E8F8C039E" box="[399,489,1477,1503]" pageId="72" pageNumber="73">Ursoaia</collectingCounty>
,
<collectingMunicipality id="0959AC90AC1143330AE1FA0E8C25039E" box="[501,576,1477,1503]" pageId="72" pageNumber="73">Dealul</collectingMunicipality>
lui
<location id="EC5D6031AC114333097AFA0D8C8F03A1" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC114333097AFA0D8C8F03A1" box="[622,746,1478,1504]" country="Romania" county="Ursoaia" municipality="Dealul" name="Dumnezeu" pageId="72" pageNumber="73" stateProvince="Iasi">Dumnezeu</location>
,
<date id="9D3C102AAC11433309E2FA0E8D1D039E" box="[758,888,1477,1503]" pageId="72" pageNumber="73" value="2023-07-21">
<collectingDate id="8D78E9C2AC11433309E2FA0E8D1D039E" box="[758,888,1477,1503]" pageId="72" pageNumber="73" value="2023-07-21">21.vii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330891FA0E8DDD039E" box="[901,952,1477,1503]" count="10" pageId="72" pageNumber="73" type="male">10♂</specimenCount>
,
<specimenCount id="FF84FD63AC11433308D1FA0E8D8C03A1" box="[965,1001,1477,1504]" count="5" pageId="72" pageNumber="73" type="female">5♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330F3CFA0D8AF9039E" box="[1064,1180,1477,1504]" pageId="72" pageNumber="73">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC1143330FBCFA0E8A90039E" box="[1192,1269,1477,1503]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330E15FA0E8D260047" collectingDate="1972-07-28" collectionCode="TJWC" collectorName="M. Kocourek" country="Slovakia" location="Sturovo" pageId="72" pageNumber="73" specimenCount="9" specimenCount-female="9">
<emphasis id="DBF6EAF8AC1143330E15FA0E8BF9039E" bold="true" box="[1281,1436,1477,1503]" pageId="72" pageNumber="73">
<collectingCountry id="9195767AAC1143330E15FA0E8BF2039E" box="[1281,1431,1477,1503]" name="Slovakia" pageId="72" pageNumber="73">SLOVAKIA</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC1143330B83FA208E970047" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC1143330B83FA208E970047" box="[151,242,1515,1542]" country="Slovakia" name="Sturovo" pageId="72" pageNumber="73">Štúrovo</location>
,
<date id="9D3C102AAC1143330A14FA228FE70042" box="[256,386,1513,1539]" pageId="72" pageNumber="73" value="1972-07-28">
<collectingDate id="8D78E9C2AC1143330A14FA228FE70042" box="[256,386,1513,1539]" pageId="72" pageNumber="73" value="1972-07-28">28.vii.1972</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330A84FA228FD10042" box="[400,436,1513,1539]" count="9" pageId="72" pageNumber="73" type="female">9♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330AE1FA218CEA0042" box="[501,655,1513,1540]" pageId="72" pageNumber="73">M. Kocourek</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC11433309EDFA208D260047" box="[761,835,1515,1542]" pageId="72" pageNumber="73">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1143330844FA228F48006B" collectingDate="1987-07-21" collectorName="Z. Padr" country="Slovakia" location="Remetske Hamre" municipality="Vihorlat" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="0959AC90AC1143330844FA228DCA0042" box="[848,943,1513,1539]" pageId="72" pageNumber="73">Vihorlat</collectingMunicipality>
,
<location id="EC5D6031AC11433308A8FA228AE70042" LSID="urn:lsid:plazi:treatment:612B87FCAC1143330B83FEA58A60009A:EC5D6031AC11433308A8FA228AE70042" box="[956,1154,1513,1539]" country="Slovakia" municipality="Vihorlat" name="Remetske Hamre" pageId="72" pageNumber="73">Remetské Hámre</location>
,
<date id="9D3C102AAC1143330F9BFA228B740042" box="[1167,1297,1513,1539]" pageId="72" pageNumber="73" value="1987-07-21">
<collectingDate id="8D78E9C2AC1143330F9BFA228B740042" box="[1167,1297,1513,1539]" pageId="72" pageNumber="73" value="1987-07-21">21.vii.1987</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1143330E0BFA228B270042" box="[1311,1346,1513,1539]" count="1" pageId="72" pageNumber="73" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1143330E90FA218EA90066" pageId="72" pageNumber="73">Z. Pádr</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1143330BD3F9FA8A60009A" pageId="72" pageNumber="73" type="distribution">
<paragraph id="E93D36EAAC1143330BD3F9FA8A60009A" blockId="72.[151,1437,505,1756]" pageId="72" pageNumber="73">
<emphasis id="DBF6EAF8AC1143330BD3F9FA8F04000A" bold="true" box="[199,353,1585,1611]" pageId="72" pageNumber="73">Distribution.</emphasis>
Not completely clear due to synonymisation, but across Pannonian Europe, including eastern
<collectingCountry id="9195767AAC1143330B83F99E8E89002E" box="[151,236,1621,1647]" name="Austria" pageId="72" pageNumber="73">Austria</collectingCountry>
,
<collectingCountry id="9195767AAC1143330A15F99E8F3B002E" box="[257,350,1621,1647]" name="Czech Republic" pageId="72" pageNumber="73">Czechia</collectingCountry>
,
<collectingCountry id="9195767AAC1143330A66F99D8FB00031" box="[370,469,1622,1648]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, northern
<collectingCountry id="9195767AAC114333094CF99E8CC7002E" box="[600,674,1621,1647]" name="Serbia" pageId="72" pageNumber="73">Serbia</collectingCountry>
,
<collectingCountry id="9195767AAC11433309A3F99E8D79002E" box="[695,796,1621,1647]" name="Slovakia" pageId="72" pageNumber="73">Slovakia</collectingCountry>
,
<collectingCountry id="9195767AAC1143330824F99E8DFD002E" box="[816,920,1621,1647]" name="Romania" pageId="72" pageNumber="73">Romania</collectingCountry>
, and
<collectingCountry id="9195767AAC11433308F0F99E8A27002E" box="[996,1090,1621,1647]" name="Ukraine" pageId="72" pageNumber="73">Ukraine</collectingCountry>
. Warnckes distribution map (
<bibRefCitation id="8D134B1BAC1143330B8BF9B28F9100D2" author="Gusenleitner, F. &amp; Schwarz, M." box="[159,500,1657,1683]" pageId="72" pageNumber="73" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
) shows a predominantly Pannonian distribution focused on
<collectingCountry id="9195767AAC1143330F86F9B18A9000D5" box="[1170,1269,1658,1684]" name="Hungary" pageId="72" pageNumber="73">Hungary</collectingCountry>
, with scattered records across
<collectingCountry id="9195767AAC1143330A29F9568FFF00F6" box="[317,410,1693,1719]" name="Ukraine" pageId="72" pageNumber="73">Ukraine</collectingCountry>
, one in the Caucasus, and some around the northern part of the Adriatic Sea (
<collectingCountry id="9195767AAC1143330FEDF9568B4A00F6" box="[1273,1327,1693,1719]" name="Italy" pageId="72" pageNumber="73">Italy</collectingCountry>
,
<collectingCountry id="9195767AAC1143330E2DF9568BF400F6" box="[1337,1425,1693,1719]" name="Croatia" pageId="72" pageNumber="73">Croatia</collectingCountry>
). The records from the Caucasus and Adriatic region must be revised for clarity.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,176 @@
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<treatment id="612B87FCAC1243300B83FD1D8CA10284" LSID="urn:lsid:plazi:treatment:612B87FCAC1243300B83FD1D8CA10284" httpUri="http://treatment.plazi.org/id/612B87FCAC1243300B83FD1D8CA10284" lastPageNumber="76" pageId="75" pageNumber="76">
<subSubSection id="A1986561AC1243300B83FD1D8CA104B1" box="[151,708,726,753]" pageId="75" pageNumber="76" type="nomenclature">
<paragraph id="E93D36EAAC1243300B83FD1D8CA104B1" blockId="75.[151,708,726,753]" box="[151,708,726,753]" pageId="75" pageNumber="76">
<heading id="B2758186AC1243300B83FD1D8CA104B1" bold="true" box="[151,708,726,753]" fontSize="11" level="2" pageId="75" pageNumber="76" reason="4">
<taxonomicName id="2E824D69AC1243300B83FD1D8CA104B1" authority="Strand, 1921" authorityName="Strand" authorityYear="1921" box="[151,708,726,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="waschulzi" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC1243300B83FD1D8CA104B1" bold="true" box="[151,708,726,753]" pageId="75" pageNumber="76">
<emphasis id="DBF6EAF8AC1243300B83FD1D8F6404B0" bold="true" box="[151,257,726,753]" italics="true" pageId="75" pageNumber="76">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1243300A06FD1D8FF104B0" bold="true" box="[274,404,726,753]" italics="true" pageId="75" pageNumber="76">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1243300AB2FD1D8C7804B0" bold="true" box="[422,541,726,753]" italics="true" pageId="75" pageNumber="76">waschulzi</emphasis>
<bibRefCitation id="8D134B1BAC1243300930FD1D8CA104B1" author="Strand, E." box="[548,708,726,752]" pageId="75" pageNumber="76" pagination="266 - 304" refId="ref91362" refString="Strand, E. (1921) Apidologisches, insbesondere uber palaarktische Andrena - Arten, auf Grund von Material des Deutschen Entomologischen Museums. Archiv fur Naturgeschichte, 87 A, 266 - 304." type="journal article" year="1921">Strand, 1921</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1243300B83FCD48CAF05AC" pageId="75" pageNumber="76" type="discussion">
<paragraph id="E93D36EAAC1243300B83FCD48CAF05AC" blockId="75.[151,1437,799,1113]" pageId="75" pageNumber="76">
Examination of material in the DEI raised the issue that the name of
<taxonomicName id="2E824D69AC12433008B2FCD48B250578" authority="Strand, 1921" authorityName="Strand" authorityYear="1921" box="[934,1344,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="schulzi" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC12433008B2FCD48DDA0578" box="[934,959,799,825]" italics="true" pageId="75" pageNumber="76">A.</emphasis>
(
<emphasis id="DBF6EAF8AC12433008C4FCD48A2D0578" box="[976,1096,799,825]" italics="true" pageId="75" pageNumber="76">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1243300F4EFCD48ACD0578" box="[1114,1192,799,825]" italics="true" pageId="75" pageNumber="76">schulzi</emphasis>
<bibRefCitation id="8D134B1BAC1243300FA4FCD48B250578" author="Strand, E." box="[1200,1344,799,825]" pageId="75" pageNumber="76" pagination="266 - 304" refId="ref91362" refString="Strand, E. (1921) Apidologisches, insbesondere uber palaarktische Andrena - Arten, auf Grund von Material des Deutschen Entomologischen Museums. Archiv fur Naturgeschichte, 87 A, 266 - 304." type="journal article" year="1921">Strand, 1921</bibRefCitation>
</taxonomicName>
(and its variety
<taxonomicName id="2E824D69AC1243300BFAFC888F3B051C" authorityName="Strand" authorityYear="1921" box="[238,350,835,861]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="schulzi">
<emphasis id="DBF6EAF8AC1243300BFAFC888F3B051C" box="[238,350,835,861]" italics="true" pageId="75" pageNumber="76">A. schulzi</emphasis>
</taxonomicName>
ab.
<emphasis id="DBF6EAF8AC1243300A86FC888C76051C" box="[402,531,835,861]" italics="true" pageId="75" pageNumber="76">clypeopicta</emphasis>
<bibRefCitation id="8D134B1BAC1243300908FC888CCA051C" author="Strand, E." box="[540,687,835,861]" pageId="75" pageNumber="76" pagination="266 - 304" refId="ref91362" refString="Strand, E. (1921) Apidologisches, insbesondere uber palaarktische Andrena - Arten, auf Grund von Material des Deutschen Entomologischen Museums. Archiv fur Naturgeschichte, 87 A, 266 - 304." type="journal article" year="1921">Strand, 1921</bibRefCitation>
) has been consistently misspelt (e.g.
<bibRefCitation id="8D134B1BAC1243300F4AFC888B60051C" author="Warncke, K." box="[1118,1285,834,861]" pageId="75" pageNumber="76" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke 1965</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1243300E07FC888B2B051C" author="Warncke, K." box="[1299,1358,835,861]" pageId="75" pageNumber="76" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">1967</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1243300E48FC888BF2051C" author="Warncke, K." box="[1372,1431,835,861]" pageId="75" pageNumber="76" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">1973</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1243300B83FCAC8F9105C0" author="Gusenleitner, F. &amp; Schwarz, M." box="[151,500,871,897]" pageId="75" pageNumber="76" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
). In print (and also on the original
<typeStatus id="36398848AC1243300893FCAC8DD205C0" box="[903,951,871,897]" pageId="75" pageNumber="76">type</typeStatus>
labels;
<figureCitation id="71B92A6FAC1243300F06FCAC8AE605C0" box="[1042,1155,870,897]" captionStart="FIGURE 50" captionStartId="76.[151,250,1943,1967]" captionTargetBox="[214,1347,181,1919]" captionTargetId="figure-28@76.[214,1373,181,1919]" captionTargetPageId="76" captionText="FIGURE 50. Andrena (Ulandrena) waschulzi Strand, 1921 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; Andrena (Ulandrena) waschulzi ab. clypeopicta Strand, 1921 holotype male. E. Label details; F. Habitus, lateral view; G. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209413" httpUri="https://zenodo.org/record/13209413/files/figure.png" pageId="75" pageNumber="76">Figure 50</figureCitation>
), it appears as “
<taxonomicName id="2E824D69AC1243300E29FCAC8F4005E4" authority="W. A. Schulzi" authorityName="W. A. Schulzi" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC1243300E29FCAC8F4005E4" italics="true" pageId="75" pageNumber="76">Andrena W.A. Schulzi</emphasis>
</taxonomicName>
” (
<bibRefCitation id="8D134B1BAC1243300A55FC408C6905E4" author="Strand, E." box="[321,524,907,933]" pageId="75" pageNumber="76" pagination="266 - 304" refId="ref91362" refString="Strand, E. (1921) Apidologisches, insbesondere uber palaarktische Andrena - Arten, auf Grund von Material des Deutschen Entomologischen Museums. Archiv fur Naturgeschichte, 87 A, 266 - 304." type="journal article" year="1921">Strand 1921: 279</bibRefCitation>
). Unfortunately, it is clear that the name should therefore be
<taxonomicName id="2E824D69AC1243300FD2FC408BF905E4" authorityName="Strand" authorityYear="1921" box="[1222,1436,907,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="waschulzi">
<emphasis id="DBF6EAF8AC1243300FD2FC408BF905E4" box="[1222,1436,907,933]" italics="true" pageId="75" pageNumber="76">Andrena waschulzi</emphasis>
</taxonomicName>
following ICZN article 32.5.2.4.4 (
<bibRefCitation id="8D134B1BAC124330090BFC648CC30588" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[543,678,943,969]" pageId="75" pageNumber="76" refId="ref89116" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
); the logic employed in the comparative case of
<taxonomicName id="2E824D69AC1243300FD2FC648F4D05AC" authority="(Kohl, 1906)" baseAuthorityName="Kohl" baseAuthorityYear="1906" class="Insecta" family="Apidae" genus="Tetralonia" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="wfkirbyi">
<emphasis id="DBF6EAF8AC1243300FD2FC648BF90588" box="[1222,1436,943,969]" italics="true" pageId="75" pageNumber="76">Tetralonia wfkirbyi</emphasis>
(Kohl, 1906)
</taxonomicName>
is followed here (
<bibRefCitation id="8D134B1BAC1243300AE6FC188CDB05AC" author="Straka, J. &amp; Batelka, J. &amp; Pauly, A." box="[498,702,979,1005]" pageId="75" pageNumber="76" pagination="183 - 219" refId="ref91254" refString="Straka, J., Batelka, J. &amp; Pauly, A. (2017) Bees of the Socotra Archipelago (Hymenoptera: Anthophila), their biogeography and association with parasites. Acta Entomologica Musei Nationalis Pragae, 57, 183 - 219. https: // doi. org / 10.1515 / aemnp- 2017 - 0118" type="journal article" year="2017">
Straka
<emphasis id="DBF6EAF8AC1243300956FC188C1605AC" box="[578,627,979,1005]" italics="true" pageId="75" pageNumber="76">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1243300BD3FC3D8E920218" pageId="75" pageNumber="76" type="materials_examined">
<paragraph id="E93D36EAAC1243300BD3FC3D8E920218" blockId="75.[151,1437,799,1113]" pageId="75" pageNumber="76">
<emphasis id="DBF6EAF8AC1243300BD3FC3D8FD40251" bold="true" box="[199,433,1014,1040]" pageId="75" pageNumber="76">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC1243300AA3FC3D8E960218" collectingDate="1914-03" collectingDateMax="1914-04-31" collectingDateMin="1914-03" collectionCode="DEI" collectorName="Paganetti" country="Greece" location="Creta" pageId="75" pageNumber="76" specimenCount="2" specimenCount-male="2" stateProvince="Kriti" typeStatus="holotype">
<emphasis id="DBF6EAF8AC1243300AA3FC3D8C520251" bold="true" box="[439,567,1014,1040]" pageId="75" pageNumber="76">
<collectingCountry id="9195767AAC1243300AA3FC3D8C570251" box="[439,562,1014,1040]" name="Greece" pageId="75" pageNumber="76">GREECE</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC124330092AFC3C8C1B0250" box="[574,638,1015,1041]" country="Greece" name="Kriti" pageId="75" pageNumber="76">Creta</collectingRegion>
,
<date id="9D3C102AAC124330099CFC3C8D660250" box="[648,771,1015,1041]" pageId="75" pageNumber="76" value="1914-03" valueMax="1914-04-31" valueMin="1914-03-00">
<collectingDate id="8D78E9C2AC124330099CFC3C8D660250" box="[648,771,1015,1041]" pageId="75" pageNumber="76" value="1914-03" valueMax="1914-04-31" valueMin="1914-03-00">iiiiv.1914</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC124330081BFC3C8D560250" box="[783,819,1015,1041]" count="1" pageId="75" pageNumber="76" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC1243300864FC3C8DBE0250" box="[880,987,1015,1041]" pageId="75" pageNumber="76">Paganetti</collectorName>
,
<collectionCode id="8F93AE2FAC12433008F2FC3C8A700250" box="[998,1045,1015,1041]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="75" pageNumber="76" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<typeStatus id="36398848AC1243300F37FC3C8AE20250" box="[1059,1159,1015,1041]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC1243300FB8FC3C8B5C0250" authorityName="Strand" authorityYear="1921" box="[1196,1337,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="waschulzi">
<emphasis id="DBF6EAF8AC1243300FB8FC3C8B5C0250" box="[1196,1337,1015,1041]" italics="true" pageId="75" pageNumber="76">A. waschulzi</emphasis>
</taxonomicName>
;
<figureCitation id="71B92A6FAC1243300E5CFC3C8E8A0274" captionStart="FIGURE 50" captionStartId="76.[151,250,1943,1967]" captionTargetBox="[214,1347,181,1919]" captionTargetId="figure-28@76.[214,1373,181,1919]" captionTargetPageId="76" captionText="FIGURE 50. Andrena (Ulandrena) waschulzi Strand, 1921 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; Andrena (Ulandrena) waschulzi ab. clypeopicta Strand, 1921 holotype male. E. Label details; F. Habitus, lateral view; G. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209413" httpUri="https://zenodo.org/record/13209413/files/figure.png" pageId="75" pageNumber="76">Figures 50AD</figureCitation>
); Canea (
<collectingRegion id="2B46F808AC1243300A49FBD08FFA0274" box="[349,415,1051,1077]" country="Greece" name="Kriti" pageId="75" pageNumber="76">Creta</collectingRegion>
),
<date id="9D3C102AAC1243300AA1FBD08C550274" box="[437,560,1051,1077]" pageId="75" pageNumber="76" value="1914-03" valueMax="1914-04-31" valueMin="1914-03-00">
<collectingDate id="8D78E9C2AC1243300AA1FBD08C550274" box="[437,560,1051,1077]" pageId="75" pageNumber="76" value="1914-03" valueMax="1914-04-31" valueMin="1914-03-00">iiiiv.1914</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC124330092BFBD08C060274" box="[575,611,1051,1077]" count="1" pageId="75" pageNumber="76" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC12433009B2FBD08D740274" box="[678,785,1051,1077]" pageId="75" pageNumber="76">Paganetti</collectorName>
,
<collectionCode id="8F93AE2FAC124330080BFBD08D2B0274" box="[799,846,1051,1077]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="75" pageNumber="76" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<typeStatus id="36398848AC1243300874FBD08DA10274" box="[864,964,1051,1077]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC12433008FBFBD08A1A0274" authorityName="Strand" authorityYear="1921" box="[1007,1151,1051,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="waschulzi">
<emphasis id="DBF6EAF8AC12433008FBFBD08A1A0274" box="[1007,1151,1051,1077]" italics="true" pageId="75" pageNumber="76">A. waschulzi</emphasis>
</taxonomicName>
ab.
<emphasis id="DBF6EAF8AC1243300FA1FBD08B530274" box="[1205,1334,1051,1077]" italics="true" pageId="75" pageNumber="76">clypeopicta</emphasis>
;
<figureCitation id="71B92A6FAC1243300E5CFBD08E8E0218" captionStart="FIGURE 50" captionStartId="76.[151,250,1943,1967]" captionTargetBox="[214,1347,181,1919]" captionTargetId="figure-28@76.[214,1373,181,1919]" captionTargetPageId="76" captionText="FIGURE 50. Andrena (Ulandrena) waschulzi Strand, 1921 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; Andrena (Ulandrena) waschulzi ab. clypeopicta Strand, 1921 holotype male. E. Label details; F. Habitus, lateral view; G. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209413" httpUri="https://zenodo.org/record/13209413/files/figure.png" pageId="75" pageNumber="76">Figures 50EH</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1243300B83FB618CA10284" box="[151,708,1194,1221]" pageId="75" pageNumber="76" type="description">
<paragraph id="E93D36EAAC1243300B83FB618CA10284" blockId="75.[151,708,1194,1221]" box="[151,708,1194,1221]" pageId="75" pageNumber="76">
<emphasis id="DBF6EAF8AC1243300B83FB618CA10284" bold="true" box="[151,708,1194,1221]" pageId="75" pageNumber="76">10) Description of previously unknown sexes</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="04C65D928CB7BD9C10F2E17CF0BDEEE2">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<treatment id="612B87FCAC1943390B83F9E38CE5042E" LSID="urn:lsid:plazi:treatment:612B87FCAC1943390B83F9E38CE5042E" httpUri="http://treatment.plazi.org/id/612B87FCAC1943390B83F9E38CE5042E" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
<subSubSection id="A1986561AC19433B0B83F9E38CA60003" box="[151,707,1576,1603]" pageId="64" pageNumber="65" type="nomenclature">
<paragraph id="E93D36EAAC19433B0B83F9E38CA60003" blockId="64.[151,707,1576,1603]" box="[151,707,1576,1603]" pageId="64" pageNumber="65">
<heading id="B2758186AC19433B0B83F9E38CA60003" bold="true" box="[151,707,1576,1603]" fontSize="11" level="2" pageId="64" pageNumber="65" reason="4">
<taxonomicName id="2E824D69AC19433B0B83F9E38CA60003" ID-CoL="DS92" authority="Friese, 1922" authorityName="Friese" authorityYear="1922" box="[151,707,1576,1603]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC19433B0B83F9E38CA60003" bold="true" box="[151,707,1576,1603]" pageId="64" pageNumber="65">
<emphasis id="DBF6EAF8AC19433B0B83F9E38F640002" bold="true" box="[151,257,1576,1603]" italics="true" pageId="64" pageNumber="65">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC19433B0A06F9E38FD30002" bold="true" box="[274,438,1576,1603]" italics="true" pageId="64" pageNumber="65">incertae sedis</emphasis>
)
<emphasis id="DBF6EAF8AC19433B0AD3F9E38C430003" bold="true" box="[455,550,1576,1602]" italics="true" pageId="64" pageNumber="65">meripes</emphasis>
<bibRefCitation id="8D134B1BAC19433B0939F9E38CA60003" author="Friese, H." box="[557,707,1576,1602]" pageId="64" pageNumber="65" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese, 1922</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC19433B0B83F9A68A0600A0" pageId="64" pageNumber="65" type="reference_group">
<paragraph id="E93D36EAAC19433B0B83F9A68F9C00E2" blockId="64.[151,1436,1645,1763]" pageId="64" pageNumber="65">
<taxonomicName id="2E824D69AC19433B0B83F9A68FAF00C4" ID-CoL="DSDP" authority="Friese, 1914" authorityName="Friese" authorityYear="1914" box="[151,458,1645,1669]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="DBF6EAF8AC19433B0B83F9A68F2300C5" box="[151,326,1645,1669]" italics="true" pageId="64" pageNumber="65">Andrena nigripes</emphasis>
<materialsCitation id="59EA3CB7AC19433B0A58F9A68FAF00C4" box="[332,458,1645,1669]" collectionCode="ZMHB" country="Russia" location="Siberia" pageId="64" pageNumber="65" specimenCount="1" specimenCount-female="1" typeStatus="lectotype">
<bibRefCitation id="8D134B1BAC19433B0A58F9A68FAF00C4" author="Friese, H." box="[332,458,1645,1669]" pageId="64" pageNumber="65" pagination="218 - 233" refId="ref88699" refString="Friese, H. (1914) Neue Apiden der palaarktischen Region. Stettiner entomologische Zeitung, 75, 218 - 233. https: // doi. org / 10.1002 / mmnd. 191419140311" type="journal article" year="1914">Friese, 1914</bibRefCitation>
</materialsCitation>
</taxonomicName>
(nec.
<taxonomicName id="2E824D69AC19433B0913F9A68D1700C4" ID-CoL="DSDP" authority="Provancher, 1895" authorityName="Provancher" authorityYear="1895" box="[519,882,1645,1669]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="DBF6EAF8AC19433B0913F9A68CD300C5" box="[519,694,1645,1669]" italics="true" pageId="64" pageNumber="65">Andrena nigripes</emphasis>
Provancher, 1895
</taxonomicName>
): 225,
<specimenCount id="FF84FD63AC19433B08A1F9A68DAD00C5" box="[949,968,1645,1668]" count="1" pageId="64" pageNumber="65" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC19433B08C0F9A68A7B00C4" box="[980,1054,1645,1669]" name="Russia" pageId="64" pageNumber="65">Russia</collectingCountry>
:
<location id="EC5D6031AC19433B0F33F9A68A1400C4" LSID="urn:lsid:plazi:treatment:612B87FCAC1943390B83F9E38CE5042E:EC5D6031AC19433B0F33F9A68A1400C4" box="[1063,1137,1645,1669]" country="Russia" name="Siberia" pageId="64" pageNumber="65">Siberia</location>
, ZMHB,
<typeStatus id="36398848AC19433B0FC5F9A68B4B00C4" box="[1233,1326,1645,1669]" pageId="64" pageNumber="65" type="lectotype">lectotype</typeStatus>
by present designation] (
<figureCitation id="71B92A6FAC19433B0A46F9408F9700E2" box="[338,498,1675,1699]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@65.[214,1373,181,1919]" captionTargetPageId="65" captionText="FIGURE 43. Andrena (Tarsandrena) nigripes Friese, 1914 nec. Provancher = A. meripes Friese, 1922 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Process of labrum, ventral view; E. Head, dorsal view; F. Scutum, dorsolateral view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209399" httpUri="https://zenodo.org/record/13209399/files/figure.png" pageId="64" pageNumber="65">Figures 43AH</figureCitation>
)
</paragraph>
<paragraph id="E93D36EAAC19433B0B83F9618DE30083" blockId="64.[151,1436,1645,1763]" box="[151,902,1706,1730]" pageId="64" pageNumber="65">
<treatmentCitation id="682310FBAC19433B0B83F9618F980083" author="Friese, H." box="[151,509,1706,1730]" page="217" pageId="64" pageNumber="65" year="1922">
<taxonomicName id="2E824D69AC19433B0B83F9618F980083" ID-CoL="DS92" authority="Friese, 1922: 217" authorityName="Friese" authorityPageNumber="217" authorityYear="1922" box="[151,509,1706,1730]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="meripes" status="nom. nov.">
<emphasis id="DBF6EAF8AC19433B0B83F9618F210083" box="[151,324,1706,1730]" italics="true" pageId="64" pageNumber="65">Andrena meripes</emphasis>
<bibRefCitation id="8D134B1BAC19433B0A5EF9618F980083" author="Friese, H." box="[330,509,1706,1730]" pageId="64" pageNumber="65" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese, 1922: 217</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="DBF6EAF8AC19433B091CF9608C0F0083" box="[520,618,1707,1730]" italics="true" pageId="64" pageNumber="65">
<taxonomicNameLabel id="C0C55783AC19433B091CF9608C0F0083" box="[520,618,1707,1730]" pageId="64" pageNumber="65">nom. nov.</taxonomicNameLabel>
</emphasis>
for
<taxonomicName id="2E824D69AC19433B0987F9618DE30083" ID-CoL="DSDP" authority="Friese, 1914" authorityName="Friese" authorityYear="1914" box="[659,902,1706,1730]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="nigripes">
<emphasis id="DBF6EAF8AC19433B0987F9618D670083" box="[659,770,1706,1730]" italics="true" pageId="64" pageNumber="65">A. nigripes</emphasis>
<bibRefCitation id="8D134B1BAC19433B081DF9618DE30083" author="Friese, H." box="[777,902,1706,1730]" pageId="64" pageNumber="65" pagination="218 - 233" refId="ref88699" refString="Friese, H. (1914) Neue Apiden der palaarktischen Region. Stettiner entomologische Zeitung, 75, 218 - 233. https: // doi. org / 10.1002 / mmnd. 191419140311" type="journal article" year="1914">Friese, 1914</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="E93D36EAAC19433B0B83F9028A0600A0" blockId="64.[151,1436,1645,1763]" box="[151,1123,1737,1763]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="C99211C4AC19433B0B83F9028A0600A0" box="[151,1123,1737,1763]" pageId="64" pageNumber="65">
<treatmentCitation id="682310FBAC19433B0B83F9028CEA00A0" author="Wood, T. J." box="[151,655,1737,1761]" httpUri="http://treatment.plazi.org/id/03828781F80B7002D6C7F8908B2EA745" page="46" pageId="64" pageNumber="65" year="2023">
<taxonomicName id="2E824D69AC19433B0B83F9028CEA00A0" authority="Wood, 2023 a: 46" authorityName="Wood" authorityPageNumber="46" authorityYear="2023" box="[151,655,1737,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="palliata" subGenus="Tarsandrena">
<emphasis id="DBF6EAF8AC19433B0B83F9028E8800A0" box="[151,237,1737,1761]" italics="true" pageId="64" pageNumber="65">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC19433B0BE8F9028F1900A0" box="[252,380,1737,1761]" italics="true" pageId="64" pageNumber="65">Tarsandrena</emphasis>
)
<emphasis id="DBF6EAF8AC19433B0A9FF9028FBF00A0" box="[395,474,1737,1761]" italics="true" pageId="64" pageNumber="65">palliata</emphasis>
<materialsCitation id="59EA3CB7AC19433B0ACBF9028CEA00A0" box="[479,655,1737,1761]" collectionCode="OOLM" country="Mongolia" location="Mongolia" pageId="64" pageNumber="65" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC19433B0ACBF9028CEA00A0" author="Wood, T. J." box="[479,655,1737,1761]" pageId="64" pageNumber="65" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood, 2023a: 46</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC19433B098DF9028CCC00A1" box="[665,681,1737,1760]" count="1" pageId="64" pageNumber="65" type="female"></specimenCount>
<specimenCount id="FF84FD63AC19433B09BDF9028CDA00A1" box="[681,703,1737,1760]" count="1" pageId="64" pageNumber="65" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC19433B09DFF9028D5600A0" box="[715,819,1737,1761]" name="Mongolia" pageId="64" pageNumber="65">Mongolia</collectingCountry>
, OÖLM, examined]
<emphasis id="DBF6EAF8AC19433B0F12F9028A0600A0" bold="true" box="[1030,1123,1737,1761]" pageId="64" pageNumber="65">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1943390B83F8C48D03072D" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65" type="discussion">
<paragraph id="E93D36EAAC1943390B83F8C48D03072D" blockId="64.[151,1437,1807,1977]" lastBlockId="66.[151,1437,151,513]" lastPageId="66" lastPageNumber="67" pageId="64" pageNumber="65">
<emphasis id="DBF6EAF8AC19433B0B83F8C48F6E0168" bold="true" box="[151,267,1807,1833]" pageId="64" pageNumber="65">Remarks.</emphasis>
<taxonomicName id="2E824D69AC19433B0A07F8C48FB10168" authorityName="Friese" authorityYear="1922" box="[275,468,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC19433B0A07F8C48FB10168" box="[275,468,1807,1833]" italics="true" pageId="64" pageNumber="65">Andrena meripes</emphasis>
</taxonomicName>
has proved an obscure taxon; it was treated as a member of the subgenus
<taxonomicName id="2E824D69AC19433B0E35F8C48BF90168" authorityName="Hedicke" authorityYear="1933" box="[1313,1436,1807,1833]" pageId="64" pageNumber="65" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC19433B0E35F8C48BF90168" box="[1313,1436,1807,1833]" italics="true" pageId="64" pageNumber="65">Euandrena</emphasis>
</taxonomicName>
by
<bibRefCitation id="8D134B1BAC19433B0BA8F8F88C4C010C" author="Gusenleitner, F. &amp; Schwarz, M." box="[188,553,1843,1869]" pageId="64" pageNumber="65" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
, but not by
<bibRefCitation id="8D134B1BAC19433B09A1F8F88DAB010C" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[693,974,1843,1869]" pageId="64" pageNumber="65" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC19433B0829F8FF8D14010C" box="[829,881,1843,1869]" italics="true" pageId="64" pageNumber="65">et al</emphasis>
. (2008)
</bibRefCitation>
. Surprisingly, this species has long been overlooked as part of the Russian bee fauna (not listed until 2023,
<bibRefCitation id="8D134B1BAC19433B086EF89C8AFF0130" author="Proshchalykin M. Y. &amp; Astafurova Y. V. &amp; Sidorov D. A." box="[890,1178,1879,1905]" pageId="64" pageNumber="65" pagination="263 - 276" refId="ref90418" refString="Proshchalykin M. Y., Astafurova Y. V. &amp; Sidorov D. A. (2017) Family Andrenidae. In: Lelej, A. S., Proshchalykin, M. Y. &amp; Loktionov, V. M. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences, Supplement 6, 263 - 276." type="journal article" year="2017">
Proshchalykin
<emphasis id="DBF6EAF8AC19433B0F34F8938A340130" box="[1056,1105,1879,1905]" italics="true" pageId="64" pageNumber="65">et al</emphasis>
. 2017
</bibRefCitation>
; 2023), despite the fact that it was described from eastern Siberia (Irkutsk) and also eastern
<collectingCountry id="9195767AAC19433B086BF8B08A6701D4" box="[895,1026,1915,1941]" name="Kazakhstan" pageId="64" pageNumber="65">Kazakhstan</collectingCountry>
(Djarkent = Zharkent) (
<bibRefCitation id="8D134B1BAC19433B0E1CF8B08BF501D4" author="Friese, H." box="[1288,1424,1915,1941]" pageId="64" pageNumber="65" pagination="218 - 233" refId="ref88699" refString="Friese, H. (1914) Neue Apiden der palaarktischen Region. Stettiner entomologische Zeitung, 75, 218 - 233. https: // doi. org / 10.1002 / mmnd. 191419140311" type="journal article" year="1914">Friese 1914</bibRefCitation>
). Examination of material in the ZMHB and DEI revealed
<specimenCount id="FF84FD63AC19433B0803F8548DAE01F8" box="[791,971,1951,1977]" count="3" pageId="64" pageNumber="65" type="generic">three specimens</specimenCount>
from Irkutsk (
<figureCitation id="71B92A6FAC19433B0F79F8548ABB01F8" box="[1133,1246,1951,1977]" captionStart="FIGURE 43" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@65.[214,1373,181,1919]" captionTargetPageId="65" captionText="FIGURE 43. Andrena (Tarsandrena) nigripes Friese, 1914 nec. Provancher = A. meripes Friese, 1922 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Process of labrum, ventral view; E. Head, dorsal view; F. Scutum, dorsolateral view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209399" httpUri="https://zenodo.org/record/13209399/files/figure.png" pageId="64" pageNumber="65">Figure 43</figureCitation>
), and revealed that
<taxonomicName id="2E824D69AC1B43390BDFFF5C8F2006F1" authorityName="Friese" authorityYear="1922" box="[203,325,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC1B43390BDFFF5C8F2006F1" box="[203,325,151,177]" italics="true" pageId="66" pageNumber="67">A. meripes</emphasis>
</taxonomicName>
is the senior synonym of
<taxonomicName id="2E824D69AC1B43390960FF5C8C8806F0" authorityName="Wood" authorityYear="2023" box="[628,749,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="palliata" status="syn. nov.">
<emphasis id="DBF6EAF8AC1B43390960FF5C8C8806F0" box="[628,749,151,177]" italics="true" pageId="66" pageNumber="67">A. palliata</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC1B433909E2FF5C8D0506F0" bold="true" box="[758,864,151,177]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="C0C55783AC1B433909E2FF5C8D0506F0" box="[758,864,151,177]" pageId="66" pageNumber="67">syn. nov.</taxonomicNameLabel>
</emphasis>
<bibRefCitation id="8D134B1BAC1B4339087DFF5C8A6906F0" author="Wood, T. J." box="[873,1036,151,177]" pageId="66" pageNumber="67" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood (2023a)</bibRefCitation>
placed
<taxonomicName id="2E824D69AC1B43390F73FF5C8A8506F0" authorityName="Wood" authorityYear="2023" box="[1127,1248,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="palliata">
<emphasis id="DBF6EAF8AC1B43390F73FF5C8A8506F0" box="[1127,1248,151,177]" italics="true" pageId="66" pageNumber="67">A. palliata</emphasis>
</taxonomicName>
in the subgenus
<taxonomicName id="2E824D69AC1B43390B83FF778F900697" authority="Osytshnjuk, 1984" authorityName="Osytshnjuk" authorityYear="1984" box="[151,501,188,214]" pageId="66" pageNumber="67" rank="subGenus" subGenus="Tarsandrena">
<emphasis id="DBF6EAF8AC1B43390B83FF778F410697" box="[151,292,188,214]" italics="true" pageId="66" pageNumber="67">Tarsandrena</emphasis>
<bibRefCitation id="8D134B1BAC1B43390A38FF778F900697" author="Osytshnjuk, A. Z." box="[300,501,188,214]" pageId="66" pageNumber="67" pagination="3 - 15" refId="ref89639" refString="Osytshnjuk, A. Z. (1984) New and little known bees of the subgenus Carandrena War. (Hymenoptera, Andrenidae, Andrena F.) in the fauna of the USSR. Trudy Zoologicheskogo Instituta, Leningrad, 128, 3 - 15. [in Russian]" type="journal article" year="1984">Osytshnjuk, 1984</bibRefCitation>
</taxonomicName>
, but provisional data from ultra-conserved element analysis (S. Bossert, T.J. Wood, G. Pisanty,
<emphasis id="DBF6EAF8AC1B43390A0FFF2A8FBB06BA" box="[283,478,225,251]" italics="true" pageId="66" pageNumber="67">unpublished data</emphasis>
) strongly suggest that
<taxonomicName id="2E824D69AC1B433909F6FF298D3906BA" authorityName="Friese" authorityYear="1922" box="[738,860,226,252]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC1B433909F6FF298D3906BA" box="[738,860,226,252]" italics="true" pageId="66" pageNumber="67">A. meripes</emphasis>
</taxonomicName>
is part of an undescribed subgenus which requires additional characterisation.
<taxonomicName id="2E824D69AC1B43390AC2FECC8CFF0761" authorityName="Friese" authorityYear="1922" box="[470,666,263,289]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC1B43390AC2FECC8CFF0761" box="[470,666,263,289]" italics="true" pageId="66" pageNumber="67">Andrena meripes</emphasis>
</taxonomicName>
therefore has a broader distribution in eastern Siberia,
<collectingCountry id="9195767AAC1B43390E3CFECC8BFD0760" box="[1320,1432,263,289]" name="Mongolia" pageId="66" pageNumber="67">Mongolia</collectingCountry>
, western
<collectingCountry id="9195767AAC1B43390BE2FEE78F590707" box="[246,316,300,326]" name="China" pageId="66" pageNumber="67">China</collectingCountry>
, and eastern
<collectingCountry id="9195767AAC1B43390AC4FEE78C330707" box="[464,598,300,326]" name="Kazakhstan" pageId="66" pageNumber="67">Kazakhstan</collectingCountry>
. No specimens from Zharkent could be found in either the ZMHB or DEI collections, but the listing of Friese is considered to be accurate.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC18433A0B83F85C8DD501B6" ID-DOI="http://doi.org/10.5281/zenodo.13209399" ID-Zenodo-Dep="13209399" httpUri="https://zenodo.org/record/13209399/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="65" targetType="figure">
<paragraph id="E93D36EAAC18433A0B83F85C8DD501B6" blockId="65.[151,1437,1943,2040]" pageId="65" pageNumber="66">
<emphasis id="DBF6EAF8AC18433A0B83F85C8F4701EE" bold="true" box="[151,290,1943,1967]" pageId="65" pageNumber="66">FIGURE 43.</emphasis>
<taxonomicName id="2E824D69AC18433A0A3EF85C8C9801F1" authority="Friese, 1914" authorityName="Friese" authorityYear="1914" box="[298,765,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="nigripes" subGenus="Tarsandrena">
<emphasis id="DBF6EAF8AC18433A0A3EF85C8FE501EE" box="[298,384,1943,1967]" italics="true" pageId="65" pageNumber="66">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC18433A0A85F85C8C7401EE" box="[401,529,1943,1967]" italics="true" pageId="65" pageNumber="66">Tarsandrena</emphasis>
)
<emphasis id="DBF6EAF8AC18433A0936F8538C1001EE" box="[546,629,1944,1967]" italics="true" pageId="65" pageNumber="66">nigripes</emphasis>
<bibRefCitation id="8D134B1BAC18433A0969F85C8C9801F1" author="Friese, H." box="[637,765,1943,1968]" pageId="65" pageNumber="66" pagination="218 - 233" refId="ref88699" refString="Friese, H. (1914) Neue Apiden der palaarktischen Region. Stettiner entomologische Zeitung, 75, 218 - 233. https: // doi. org / 10.1002 / mmnd. 191419140311" type="journal article" year="1914">Friese, 1914</bibRefCitation>
</taxonomicName>
nec. Provancher =
<taxonomicName id="2E824D69AC18433A08DDF8538AA501F1" authority="Friese, 1922" authorityName="Friese" authorityYear="1922" box="[969,1216,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="meripes">
<emphasis id="DBF6EAF8AC18433A08DDF8538A5D01EE" box="[969,1080,1944,1967]" italics="true" pageId="65" pageNumber="66">A. meripes</emphasis>
<bibRefCitation id="8D134B1BAC18433A0F54F85C8AA501F1" author="Friese, H." box="[1088,1216,1943,1968]" pageId="65" pageNumber="66" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese, 1922</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC18433A0FDCF8538B4C01F1" bold="true" box="[1224,1321,1944,1968]" pageId="65" pageNumber="66">lectotype</emphasis>
female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Process of labrum, ventral view; E. Head, dorsal view; F. Scutum, dorsolateral view; G. Propodeum, dorsal view; H. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC1B43390BD3FEBD8F04079A" pageId="66" pageNumber="67" type="materials_examined">
<paragraph id="E93D36EAAC1B43390BD3FEBD8F04079A" blockId="66.[151,1437,151,513]" pageId="66" pageNumber="67">
<emphasis id="DBF6EAF8AC1B43390BD3FEBD8FD407D1" bold="true" box="[199,433,374,400]" pageId="66" pageNumber="67">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC1B43390AACFEBD8BFD07D0" box="[440,1432,374,401]" collectionCode="ZMHB" collectorName="Staudinger" country="Russia" location="Irkutsk" pageId="66" pageNumber="67" specimenCount="1" specimenCount-female="1" stateProvince="Irkutsk" typeStatus="lectotype">
<emphasis id="DBF6EAF8AC1B43390AACFEBD8C4D07D1" bold="true" box="[440,552,374,400]" pageId="66" pageNumber="67">
<collectingCountry id="9195767AAC1B43390AACFEBD8C4607D1" box="[440,547,374,400]" name="Russia" pageId="66" pageNumber="67">RUSSIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC1B4339093BFEBC8CE407D0" box="[559,641,375,401]" country="Russia" name="Irkutsk" pageId="66" pageNumber="67">Irkutsk</collectingRegion>
, 1897,
<specimenCount id="FF84FD63AC1B433909C7FEBC8C9307D0" box="[723,758,375,401]" count="1" pageId="66" pageNumber="67" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC1B43390820FEBC8DCA07D0" box="[820,943,375,401]" pageId="66" pageNumber="67">Staudinger</collectorName>
,
<collectionCode id="8F93AE2FAC1B433908AFFEBC8A6907D0" box="[955,1036,375,401]" pageId="66" pageNumber="67">ZMHB</collectionCode>
(
<emphasis id="DBF6EAF8AC1B43390F08FEBC8AE207D0" bold="true" box="[1052,1159,375,401]" pageId="66" pageNumber="67">
<typeStatus id="36398848AC1B43390F08FEBC8AE207D0" box="[1052,1159,375,401]" pageId="66" pageNumber="67" type="lectotype">lectotype</typeStatus>
</emphasis>
by present designation)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC1B43390B83FE578F39079A" collectionCode="DEI" collectorName="Staudinger" country="Russia" location="Irkutsk" pageId="66" pageNumber="67" specimenCount="2" specimenCount-female="2" stateProvince="Irkutsk" typeStatus="syntype">
<collectingRegion id="2B46F808AC1B43390B83FE578E8C07F7" box="[151,233,412,438]" country="Russia" name="Irkutsk" pageId="66" pageNumber="67">Irkutsk</collectingRegion>
, 1898,
<specimenCount id="FF84FD63AC1B43390A2AFE578F0607F7" box="[318,355,412,438]" count="2" pageId="66" pageNumber="67" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC1B43390AB0FE578C7A07F7" box="[420,543,412,438]" pageId="66" pageNumber="67">Staudinger</collectorName>
,
<collectionCode id="8F93AE2FAC1B43390938FE568C3E07F6" box="[556,603,413,439]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="66" pageNumber="67" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<emphasis id="DBF6EAF8AC1B43390979FE578CB107F7" bold="true" box="[621,724,412,438]" pageId="66" pageNumber="67">
<typeStatus id="36398848AC1B43390979FE578CB107F7" box="[621,724,412,438]" pageId="66" pageNumber="67" type="syntype">syntypes</typeStatus>
</emphasis>
) (see
<bibRefCitation id="8D134B1BAC1B4339080FFE578DCE07F7" author="Wood, T. J." box="[795,939,412,438]" pageId="66" pageNumber="67" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood 2023a</bibRefCitation>
for additional examined material under the name
<taxonomicName id="2E824D69AC1B43390BCFFE098F34079A" authorityName="Wood" authorityYear="2023" box="[219,337,449,476]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="palliata">
<emphasis id="DBF6EAF8AC1B43390BCFFE098F34079A" box="[219,337,449,476]" italics="true" pageId="66" pageNumber="67">A. palliata</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1B43390BD3FE2D8CE5042E" pageId="66" pageNumber="67" type="distribution">
<paragraph id="E93D36EAAC1B43390BD3FE2D8BEB0440" blockId="66.[151,1437,151,513]" box="[199,1422,486,513]" pageId="66" pageNumber="67">
<emphasis id="DBF6EAF8AC1B43390BD3FE2D8F040441" bold="true" box="[199,353,486,512]" pageId="66" pageNumber="67">Distribution.</emphasis>
<collectingCountry id="9195767AAC1B43390A73FE2C8F880440" box="[359,493,487,513]" name="Kazakhstan" pageId="66" pageNumber="67">Kazakhstan</collectingCountry>
,
<collectingCountry id="9195767AAC1B43390AECFE2C8C260440" box="[504,579,487,513]" name="Russia" pageId="66" pageNumber="67">Russia</collectingCountry>
(Eastern Siberia),
<collectingCountry id="9195767AAC1B43390801FE2C8DE00440" box="[789,901,487,513]" name="Mongolia" pageId="66" pageNumber="67">Mongolia</collectingCountry>
,
<collectingCountry id="9195767AAC1B43390884FE2C8DB60440" box="[912,979,487,513]" name="China" pageId="66" pageNumber="67">China</collectingCountry>
(
<collectingRegion id="2B46F808AC1B433908F5FE2C8A220440" box="[993,1095,487,513]" country="China" name="Xinjiang Uygur" pageId="66" pageNumber="67">Xinjiang</collectingRegion>
) (
<bibRefCitation id="8D134B1BAC1B43390F49FE2C8A800440" author="Friese, H." box="[1117,1253,487,513]" pageId="66" pageNumber="67" pagination="218 - 233" refId="ref88699" refString="Friese, H. (1914) Neue Apiden der palaarktischen Region. Stettiner entomologische Zeitung, 75, 218 - 233. https: // doi. org / 10.1002 / mmnd. 191419140311" type="journal article" year="1914">Friese 1914</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC1B43390FE4FE2C8BE60440" author="Wood, T. J." box="[1264,1411,487,513]" pageId="66" pageNumber="67" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood 2023a</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAAC1B43390B83FD9E8CE5042E" blockId="66.[151,640,597,623]" box="[151,640,597,623]" pageId="66" pageNumber="67">
<emphasis id="DBF6EAF8AC1B43390B83FD9E8CE5042E" bold="true" box="[151,640,597,623]" pageId="66" pageNumber="67">7) Taxa newly elevated to species status</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -1,47 +1,64 @@
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<mods:identifier id="D09EFE7A0E0F5E6A65068A8784299A04" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="79878F49BFF96BBEF9BB83B2E722A04B" type="Zenodo-Dep">13209307</mods:identifier>
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<treatment id="612B87FCAC1C43330B83FF148C70075C" ID-DOI="http://doi.org/10.5281/zenodo.13209485" ID-Zenodo-Dep="13209485" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C" httpUri="http://treatment.plazi.org/id/612B87FCAC1C43330B83FF148C70075C" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
<subSubSection id="A1986561AC1C433E0B83FF148B0D06B8" box="[151,1384,223,252]" pageId="69" pageNumber="70" type="nomenclature">
<treatment id="612B87FCAC1C43330B83FF5C8C70075C" ID-DOI="http://doi.org/10.5281/zenodo.13209485" ID-Zenodo-Dep="13209485" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C" httpUri="http://treatment.plazi.org/id/612B87FCAC1C43330B83FF5C8C70075C" lastPageId="72" lastPageNumber="73" pageId="69" pageNumber="70">
<subSubSection id="A1986561AC1C433E0B83FF5C8D0406F0" box="[151,865,151,178]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="E93D36EAAC1C433E0B83FF5C8D0406F0" blockId="69.[151,865,151,178]" box="[151,865,151,178]" pageId="69" pageNumber="70">
<heading id="B2758186AC1C433E0B83FF5C8D0406F0" bold="true" box="[151,865,151,178]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="4">
<emphasis id="DBF6EAF8AC1C433E0B83FF5C8D0406F0" bold="true" box="[151,865,151,178]" pageId="69" pageNumber="70">
<taxonomicName id="2E824D69AC1C433E0B83FF5C8C8E06F0" ID-CoL="7J9X5" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" baseAuthorityName="Warncke" baseAuthorityYear="1975" box="[151,747,151,178]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="pelagica" status="stat. nov." subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC1C433E0B83FF5C8F6406F3" bold="true" box="[151,257,151,178]" italics="true" pageId="69" pageNumber="70">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1C433E0A06FF5C8FD706F3" bold="true" box="[274,434,151,178]" italics="true" pageId="69" pageNumber="70">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1C433E0AD7FF5C8C4D06F3" bold="true" box="[451,552,151,178]" italics="true" pageId="69" pageNumber="70">pelagica</emphasis>
Warncke, 1975
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC1C433E09E6FF5C8D0406F0" box="[754,865,151,177]" pageId="69" pageNumber="70">stat. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1C433E0B83FF148B0D06B8" box="[151,1384,223,252]" pageId="69" pageNumber="70" type="reference_group">
<paragraph id="E93D36EAAC1C433E0B83FF148B0D06B8" blockId="69.[151,1384,223,252]" box="[151,1384,223,252]" pageId="69" pageNumber="70">
<heading id="B2758186AC1C433E0B83FF148B0D06B8" box="[151,1384,223,252]" fontSize="11" level="2" pageId="69" pageNumber="70" reason="5">
<treatmentCitationGroup id="C99211C4AC1C433E0B83FF148B0D06B8" box="[151,1384,223,252]" pageId="69" pageNumber="70">
@ -255,10 +272,7 @@ Dark, 1.35 times wider than long (
). Clypeus weakly domed, surface with weakly raised irregular latitudinal wrinkles, surface punctate between wrinkles, punctures separated by 0.51 puncture diameters; underlying surface shagreened, weakly shining. Process of labrum rounded rectangular, 2 times wider than long, surface shining. Gena subequal to width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Face, gena, vertex, and scape with long whitish hairs, none equalling length of scape. Antennae dark, A3 exceeding A4, shorter than A4+5; A4 extremely short, clearly 2 times wider than long, A5 quadrate, A613 progressively becoming slightly longer.
</paragraph>
<paragraph id="E93D36EAAC1C433E0BD3F9038D09019C" blockId="69.[151,1437,295,2013]" pageId="69" pageNumber="70">
<emphasis id="DBF6EAF8AC1C433E0BD3F9038F2D00A0" box="[199,328,1736,1761]" italics="true" pageId="69" pageNumber="70">
<taxonomicName id="2E824D69AC1C433E0BD3F9038F2700A0" box="[199,322,1736,1761]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
<emphasis id="DBF6EAF8AC1C433E0BD3F9038F2D00A0" box="[199,328,1736,1761]" italics="true" pageId="69" pageNumber="70">Mesosoma:</emphasis>
Scutum and scutellum uniformly microreticulate, weakly shining, surface irregularly punctate, punctures separated by 0.51 puncture diameters (
<figureCitation id="71B92A6FAC1C433E09C6F9208D3C0144" box="[722,857,1771,1797]" captionStart="FIGURE 48" captionStartId="71.[151,250,1700,1724]" captionTargetBox="[151,1436,237,1652]" captionTargetId="figure-39@71.[151,1436,237,1676]" captionTargetPageId="71" captionText="FIGURE 48. Andrena (Micrandrena) rugothorace ssp. pelagica Warncke, 1975 male.A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; Andrena (Micrandrena) rugothorace Warncke, 1965 male. F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209409" httpUri="https://zenodo.org/record/13209409/files/figure.png" pageId="69" pageNumber="70">Figure 48C</figureCitation>
). Pronotum rounded. Mesepisternum with granular microreticulation, weakly shining. Propodeum with granular microreticulation, sculpture overlain by sparse network of raised rugosity, surface weakly shining. Propodeal triangle lacking lateral carinae, internal surface with longitudinal raised rugae, covering entire surface of triangle, interspaces shining. Mesepisternum and propodeum with long white plumose hairs, hairs subequal to length of scape, hairs becoming shorter and light brown on scutum and scutellum. Legs dark, apical tarsal segments lightened orange-brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange-brown, nervulus interstitial.
@ -341,7 +355,7 @@ male. F. Genital capsule, dorsal view.
</emphasis>
<collectingRegion id="2B46F808AC1E433C0956F8F98CE4010D" box="[578,641,1842,1868]" country="Greece" name="Kriti" pageId="71" pageNumber="72">Crete</collectingRegion>
,
<location id="EC5D6031AC1E433C099AF8F88C95010C" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C099AF8F88C95010C" box="[654,752,1843,1869]" country="Greece" name="Knossos" pageId="71" pageNumber="72" stateProvince="Kriti">Knossos</location>
<location id="EC5D6031AC1E433C099AF8F88C95010C" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C099AF8F88C95010C" box="[654,752,1843,1869]" country="Greece" name="Knossos" pageId="71" pageNumber="72" stateProvince="Kriti">Knossos</location>
,
<date id="9D3C102AAC1E433C09E8F8F98D09010D" box="[764,876,1842,1868]" pageId="71" pageNumber="72" value="1963-05-21">
<collectingDate id="8D78E9C2AC1E433C09E8F8F98D09010D" box="[764,876,1842,1868]" pageId="71" pageNumber="72" value="1963-05-21">21.v.1963</collectingDate>
@ -362,7 +376,7 @@ male. F. Genital capsule, dorsal view.
,
<collectingMunicipality id="0959AC90AC1E433C0B83F89C8F4E0131" box="[151,299,1878,1905]" pageId="71" pageNumber="72">Dept. Chania</collectingMunicipality>
,
<location id="EC5D6031AC1E433C0A20F89D8FAE0131" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C0A20F89D8FAE0131" box="[308,459,1878,1904]" country="Greece" municipality="Dept. Chania" name="Omalos Plain" pageId="71" pageNumber="72" stateProvince="Kriti">Omalos Plain</location>
<location id="EC5D6031AC1E433C0A20F89D8FAE0131" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C0A20F89D8FAE0131" box="[308,459,1878,1904]" country="Greece" municipality="Dept. Chania" name="Omalos Plain" pageId="71" pageNumber="72" stateProvince="Kriti">Omalos Plain</location>
,
<date id="9D3C102AAC1E433C0AC0F89D8C520131" box="[468,567,1878,1904]" pageId="71" pageNumber="72" value="1973-05-04">
<collectingDate id="8D78E9C2AC1E433C0AC0F89D8C520131" box="[468,567,1878,1904]" pageId="71" pageNumber="72" value="1973-05-04">4.v.1973</collectingDate>
@ -386,7 +400,7 @@ male. F. Genital capsule, dorsal view.
,
<collectingMunicipality id="0959AC90AC1E433C0F7DF89C8B7B0131" box="[1129,1310,1878,1905]" pageId="71" pageNumber="72">Dept. Heraklion</collectingMunicipality>
,
<location id="EC5D6031AC1E433C0E30F89D8EA901D4" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C0E30F89D8EA901D4" country="Greece" municipality="Dept. Heraklion" name="Asterousia Mts." pageId="71" pageNumber="72" stateProvince="Kriti">Asterousia Mts.</location>
<location id="EC5D6031AC1E433C0E30F89D8EA901D4" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C0E30F89D8EA901D4" country="Greece" municipality="Dept. Heraklion" name="Asterousia Mts." pageId="71" pageNumber="72" stateProvince="Kriti">Asterousia Mts.</location>
, betw.
<collectorName id="4477533CAC1E433C0A30F8B18F1601D5" box="[292,371,1914,1940]" pageId="71" pageNumber="72">Loukia</collectorName>
&amp;
@ -408,7 +422,7 @@ male. F. Genital capsule, dorsal view.
,
<collectingMunicipality id="0959AC90AC1E433C0F90F8B08B2501D5" box="[1156,1344,1914,1941]" pageId="71" pageNumber="72">Dept. Heraklion</collectingMunicipality>
,
<location id="EC5D6031AC1E433C0E44F8B18BF901D5" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C0E44F8B18BF901D5" box="[1360,1436,1914,1940]" country="Greece" municipality="Dept. Heraklion" name="Gortys" pageId="71" pageNumber="72" stateProvince="Kriti">Gortys</location>
<location id="EC5D6031AC1E433C0E44F8B18BF901D5" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C0E44F8B18BF901D5" box="[1360,1436,1914,1940]" country="Greece" municipality="Dept. Heraklion" name="Gortys" pageId="71" pageNumber="72" stateProvince="Kriti">Gortys</location>
[Gortyn],
<date id="9D3C102AAC1E433C0A1FF8558FE101F9" box="[267,388,1950,1976]" pageId="71" pageNumber="72" value="1973-04-26">
<collectingDate id="8D78E9C2AC1E433C0A1FF8558FE101F9" box="[267,388,1950,1976]" pageId="71" pageNumber="72" value="1973-04-26">26.iv.1973</collectingDate>
@ -426,7 +440,7 @@ male. F. Genital capsule, dorsal view.
,
<collectingMunicipality id="0959AC90AC1E433C0871F8548A4401F9" box="[869,1057,1950,1977]" pageId="71" pageNumber="72">Dept. Heraklion</collectingMunicipality>
,
<location id="EC5D6031AC1E433C0F24F8558AF701F9" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C0F24F8558AF701F9" box="[1072,1170,1950,1976]" country="Greece" municipality="Dept. Heraklion" name="Phaestos" pageId="71" pageNumber="72" stateProvince="Kriti">Phaestos</location>
<location id="EC5D6031AC1E433C0F24F8558AF701F9" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C0F24F8558AF701F9" box="[1072,1170,1950,1976]" country="Greece" municipality="Dept. Heraklion" name="Phaestos" pageId="71" pageNumber="72" stateProvince="Kriti">Phaestos</location>
[Phaistos],
<date id="9D3C102AAC1E433C0E0BF8558BFD01F9" box="[1311,1432,1950,1976]" pageId="71" pageNumber="72" value="1973-04-26">
<collectingDate id="8D78E9C2AC1E433C0E0BF8558BFD01F9" box="[1311,1432,1950,1976]" pageId="71" pageNumber="72" value="1973-04-26">26.iv.1973</collectingDate>
@ -444,7 +458,7 @@ male. F. Genital capsule, dorsal view.
,
<collectingMunicipality id="0959AC90AC1E433C0977F8088D54019D" box="[611,817,1986,2013]" pageId="71" pageNumber="72">Dept. Rethymnon</collectingMunicipality>
,
<location id="EC5D6031AC1E433C0829F8098DF7019D" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1E433C0829F8098DF7019D" box="[829,914,1986,2012]" country="Greece" municipality="Dept. Rethymnon" name="Anogia" pageId="71" pageNumber="72" stateProvince="Kriti">Anogia</location>
<location id="EC5D6031AC1E433C0829F8098DF7019D" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1E433C0829F8098DF7019D" box="[829,914,1986,2012]" country="Greece" municipality="Dept. Rethymnon" name="Anogia" pageId="71" pageNumber="72" stateProvince="Kriti">Anogia</location>
,
<date id="9D3C102AAC1E433C08B5F8098A7C019D" box="[929,1049,1986,2012]" pageId="71" pageNumber="72" value="1973-04-27">
<collectingDate id="8D78E9C2AC1E433C08B5F8098A7C019D" box="[929,1049,1986,2012]" pageId="71" pageNumber="72" value="1973-04-27">27.iv.1973</collectingDate>
@ -460,7 +474,7 @@ male. F. Genital capsule, dorsal view.
<materialsCitation id="59EA3CB7AC1143330B83FF5C8D0806F0" box="[151,877,151,177]" collectingDate="1973-04-24" collectionCode="RMNH" collectorName="v. Ooststroom" country="Greece" location="Lassithi Plain" pageId="72" pageNumber="73" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC1143330B83FF5C8EB306F0" box="[151,214,151,177]" country="Greece" name="Kriti" pageId="72" pageNumber="73">Crete</collectingRegion>
,
<location id="EC5D6031AC1143330BF5FF5C8F1906F0" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1143330BF5FF5C8F1906F0" box="[225,380,151,177]" country="Greece" name="Lassithi Plain" pageId="72" pageNumber="73" stateProvince="Kriti">Lassithi Plain</location>
<location id="EC5D6031AC1143330BF5FF5C8F1906F0" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1143330BF5FF5C8F1906F0" box="[225,380,151,177]" country="Greece" name="Lassithi Plain" pageId="72" pageNumber="73" stateProvince="Kriti">Lassithi Plain</location>
,
<date id="9D3C102AAC1143330A9CFF5C8C6506F0" box="[392,512,151,177]" pageId="72" pageNumber="73" value="1973-04-24">
<collectingDate id="8D78E9C2AC1143330A9CFF5C8C6506F0" box="[392,512,151,177]" pageId="72" pageNumber="73" value="1973-04-24">24.iv.1973</collectingDate>
@ -476,7 +490,7 @@ male. F. Genital capsule, dorsal view.
<materialsCitation id="59EA3CB7AC1143330863FF5C8C070694" collectingDate="2023-06-01" collectorName="R. Le Divelec" country="Greece" elevation="1160" location="Plateau Katharo" pageId="72" pageNumber="73" specimenCount="2" specimenCount-female="2" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC1143330863FF5C8DD306F0" box="[887,950,151,177]" country="Greece" name="Kriti" pageId="72" pageNumber="73">Crete</collectingRegion>
,
<location id="EC5D6031AC11433308D6FF5C8A1006F0" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC11433308D6FF5C8A1006F0" box="[962,1141,151,177]" country="Greece" name="Plateau Katharo" pageId="72" pageNumber="73" stateProvince="Kriti">Plateau Katharo</location>
<location id="EC5D6031AC11433308D6FF5C8A1006F0" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC11433308D6FF5C8A1006F0" box="[962,1141,151,177]" country="Greece" name="Plateau Katharo" pageId="72" pageNumber="73" stateProvince="Kriti">Plateau Katharo</location>
[
<locationDeviation id="F88F2FC8AC1143330F96FF5C8B5406F0" box="[1154,1329,151,177]" location="Katharon" pageId="72" pageNumber="73">W of Katharon</locationDeviation>
],
@ -499,7 +513,7 @@ male. F. Genital capsule, dorsal view.
[
<collectingRegion id="2B46F808AC11433309BAFF708C970694" box="[686,754,187,213]" country="Greece" name="Kriti" pageId="72" pageNumber="73">Crete</collectingRegion>
],
<location id="EC5D6031AC1143330816FF708DF00694" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF148C70075C:EC5D6031AC1143330816FF708DF00694" box="[770,917,187,213]" country="Greece" name="Aghia Galini" pageId="72" pageNumber="73" stateProvince="Kriti">Aghia Galini</location>
<location id="EC5D6031AC1143330816FF708DF00694" LSID="urn:lsid:plazi:treatment:612B87FCAC1C43330B83FF5C8C70075C:EC5D6031AC1143330816FF708DF00694" box="[770,917,187,213]" country="Greece" name="Aghia Galini" pageId="72" pageNumber="73" stateProvince="Kriti">Aghia Galini</location>
,
<date id="9D3C102AAC114333088BFF708A7E0694" box="[927,1051,187,213]" pageId="72" pageNumber="73" value="1978-03-24">
<collectingDate id="8D78E9C2AC114333088BFF708A7E0694" box="[927,1051,187,213]" pageId="72" pageNumber="73" value="1978-03-24">24.iii.1978</collectingDate>

View file

@ -0,0 +1,787 @@
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<mods:title id="DAAEE92F1F17A9D28EA6B859896DDE03">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:number id="28B29B31C5BF52EDA4292A5EF8E71FA6">2024-07-22</mods:number>
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<mods:detail id="F9244FAFB245990B6C727F23AE9413D1" type="volume">
<mods:number id="A6E21100B6234AA532403CEADAFD7C04">5483</mods:number>
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<mods:number id="B0DADC3ED5851A1E7711BC23FC84FC2A">1</mods:number>
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<mods:extent id="EE4BAF6C599F3ADAD85BFE2BB4219BEE" unit="page">
<mods:start id="8ECE196055182422FA9A25D6857F3622">1</mods:start>
<mods:end id="BF84084FB87F474429B74194621A37C4">150</mods:end>
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<mods:location id="C255423767E3CD6460D49B64CF780855">
<mods:url id="742A142DE90F2587C550759742259CF4">http://dx.doi.org/10.11646/zootaxa.5483.1.1</mods:url>
</mods:location>
<mods:classification id="5D951B7B70F668C69C53917F7BFB693A">journal article</mods:classification>
<mods:identifier id="0955C9E5AC54F36291C6A48BDE3D1494" type="DOI">10.11646/zootaxa.5483.1.1</mods:identifier>
<mods:identifier id="AD355EA1729703804CFC3C78674DB118" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="6B3BEDE18D8AA8C8DC82378D83130F19" type="Zenodo-Dep">13209307</mods:identifier>
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</mods:mods>
<treatment id="612B87FCAC2243050B83F9698B270048" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048" httpUri="http://treatment.plazi.org/id/612B87FCAC2243050B83F9698B270048" lastPageId="126" lastPageNumber="127" pageId="123" pageNumber="124">
<subSubSection id="A1986561AC2243000B83F9698CD800FD" box="[151,701,1698,1725]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="E93D36EAAC2243000B83F9698CD800FD" blockId="123.[151,701,1698,1760]" box="[151,701,1698,1725]" pageId="123" pageNumber="124">
<heading id="B2758186AC2243000B83F9698CD800FD" bold="true" box="[151,701,1698,1725]" fontSize="11" level="2" pageId="123" pageNumber="124" reason="4">
<emphasis id="DBF6EAF8AC2243000B83F9698CD800FD" bold="true" box="[151,701,1698,1725]" pageId="123" pageNumber="124">
<taxonomicName id="2E824D69AC2243000B83F9698C5B00FD" authorityName="WOOD" authorityYear="2024" box="[151,574,1698,1725]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="concisura" status="sp. nov.">
<emphasis id="DBF6EAF8AC2243000B83F9698F6400FC" bold="true" box="[151,257,1698,1725]" italics="true" pageId="123" pageNumber="124">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2243000A06F9698FD300FC" bold="true" box="[274,438,1698,1725]" italics="true" pageId="123" pageNumber="124">incertae sedis</emphasis>
)
<emphasis id="DBF6EAF8AC2243000AD3F9698C5B00FD" bold="true" box="[455,574,1698,1724]" italics="true" pageId="123" pageNumber="124">concisura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC2243000951F9698CD800FD" box="[581,701,1698,1724]" pageId="123" pageNumber="124" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2243000B83F90D8F2900A1" box="[151,332,1734,1760]" pageId="123" pageNumber="124" type="description">
<paragraph id="E93D36EAAC2243000B83F90D8F2900A1" blockId="123.[151,701,1698,1760]" box="[151,332,1734,1760]" pageId="123" pageNumber="124">
<figureCitation id="71B92A6FAC2243000B83F90D8F2D00A1" box="[151,328,1734,1760]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="123" pageNumber="124">Figures 77AG</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2243000B83F8C58D9B0131" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="E93D36EAAC2243000B83F8C58FF0010E" blockId="123.[151,1437,1806,2013]" pageId="123" pageNumber="124">
<materialsCitation id="59EA3CB7AC2243000B83F8C58FF0010E" collectingDate="2006-09-04" collectorName="V. Gurko" country="Tajikistan" location="Tigrova Balka" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC2243000B83F8C58F9D0169" bold="true" box="[151,504,1806,1832]" pageId="123" pageNumber="124">
<typeStatus id="36398848AC2243000B83F8C58F5F0169" box="[151,314,1806,1832]" pageId="123" pageNumber="124" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC2243000A53F8C58F960169" box="[327,499,1806,1832]" name="Tajikistan" pageId="123" pageNumber="124">TAJIKISTAN</collectingCountry>
:
</emphasis>
S.,
<location id="EC5D6031AC2243000932F8C58CAD0169" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2243000932F8C58CAD0169" box="[550,712,1806,1832]" country="Tajikistan" name="Tigrova Balka" pageId="123" pageNumber="124">Tigrova Balka</location>
res., Pianj. Riv. [inferred 37.1975
<superScript id="1EF79BA2AC2243000F59F8C58A090168" attach="right" box="[1101,1132,1806,1833]" fontSize="6" pageId="123" pageNumber="124">oN</superScript>
, 68.5639
<superScript id="1EF79BA2AC2243000FC0F8C58A970168" attach="right" box="[1236,1266,1806,1833]" fontSize="6" pageId="123" pageNumber="124">oE</superScript>
],
<date id="9D3C102AAC2243000E13F8C58B160169" box="[1287,1395,1806,1832]" pageId="123" pageNumber="124" value="2006-09-04">
<collectingDate id="8D78E9C2AC2243000E13F8C58B160169" box="[1287,1395,1806,1832]" pageId="123" pageNumber="124" value="2006-09-04">4.ix.2006</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2243000E94F8C48BF00168" box="[1408,1429,1807,1833]" count="1" pageId="123" pageNumber="124" type="female"></specimenCount>
, leg.
<collectorName id="4477533CAC2243000BD3F8F88F4A010D" box="[199,303,1842,1869]" pageId="123" pageNumber="124">V. Gurko</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC2243000BD3F89D8D9B0131" blockId="123.[151,1437,1806,2013]" box="[199,1022,1878,1905]" pageId="123" pageNumber="124">
<materialsCitation id="59EA3CB7AC2243000BD3F89D8D9F0131" box="[199,1018,1878,1905]" collectionCode="TJWC" country="Tajikistan" location="TAJIKISTAN" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="DBF6EAF8AC2243000BD3F89D8C790131" bold="true" box="[199,540,1878,1904]" pageId="123" pageNumber="124">
<typeStatus id="36398848AC2243000BD3F89D8F050131" box="[199,352,1878,1904]" pageId="123" pageNumber="124" type="paratype">PARATYPE</typeStatus>
:
<collectingCountry id="9195767AAC2243000A7FF89D8C720131" box="[363,535,1878,1904]" name="Tajikistan" pageId="123" pageNumber="124">TAJIKISTAN</collectingCountry>
:
</emphasis>
same information as
<typeStatus id="36398848AC2243000819F89D8D170131" box="[781,882,1878,1904]" pageId="123" pageNumber="124" type="holotype">holotype</typeStatus>
,
<specimenCount id="FF84FD63AC2243000869F89D8DC50131" box="[893,928,1878,1904]" count="1" pageId="123" pageNumber="124" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FAC22430008B9F89D8D9F0131" box="[941,1018,1878,1904]" pageId="123" pageNumber="124">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2243070BD3F8B18FED0284" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124" type="description">
<paragraph id="E93D36EAAC2243070BD3F8B18F7107C8" blockId="123.[151,1437,1806,2013]" lastBlockId="124.[151,1437,151,2013]" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
<emphasis id="DBF6EAF8AC2243000BD3F8B18FA301D5" bold="true" box="[199,454,1914,1940]" pageId="123" pageNumber="124">Description: Female:</emphasis>
Body length:
<quantity id="2E7A9B0FAC2243000964F8B18CB401D4" box="[624,721,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="123" pageNumber="124" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
(74A).
<emphasis id="DBF6EAF8AC2243000824F8B18D1001D5" box="[816,885,1914,1940]" italics="true" pageId="123" pageNumber="124">Head:</emphasis>
Dark, 1.1 times wider than long (
<figureCitation id="71B92A6FAC2243000E1CF8B18BF501D5" box="[1288,1424,1914,1940]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="123" pageNumber="124">Figure 77B</figureCitation>
). Clypeus domed, polished and shining, irregularly punctate, punctures separated by 0.52 puncture diameters. Process of labrum rounded triangular, as long as wide (
<figureCitation id="71B92A6FAC2243000831F8098DCB019D" box="[805,942,1986,2012]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="123" pageNumber="124">Figure 77C</figureCitation>
). Gena equalling width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Frons slightly domed, surface polished and shining, densely punctate, puncture separated by 0.51 puncture diameters. Vertex posterolateral to lateral ocelli with shining, impunctate, roughly triangular spaces. Foveae dorsally broad, occupying ¾ space between compound eye and lateral ocellus, long, ventrally extending to below level of antennal insertions; foveae filled with light brown hairs (
<figureCitation id="71B92A6FAC2543070BCBFEEC8F0D0700" box="[223,360,295,321]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77D</figureCitation>
). Face, gena, scape with short white hairs, becoming light golden-brown on vertex. Antennae basally dark brown, A512 ventrally lightly lightened by presence of brownish scales; A3 exceeding A4, shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAAC2543070BD3FE588B24051C" blockId="124.[151,1437,151,2013]" pageId="124" pageNumber="125">
<emphasis id="DBF6EAF8AC2543070BD3FE588F2D07ED" box="[199,328,403,428]" italics="true" pageId="124" pageNumber="125">Mesosoma:</emphasis>
Scutum and scutellum smooth and shining, irregularly punctate, scutum with punctures typically separated by 0.52 puncture diameters, scutellum with punctures sparse medially, separated by up to 4 puncture diameters (
<figureCitation id="71B92A6FAC2543070A04FE108FF407B4" box="[272,401,475,501]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77E</figureCitation>
). Pronotum with weak humeral angle. Mesepisternum smooth and shining, punctate, punctures separated by 0.51 puncture diameters.Dorsolateral parts of propodeum with obscure weak shagreen, weakly shining; propodeal triangle laterally delineated by slightly impressed shining rim, surface weakly shagreened, medially with weak network of slightly raised rugae; propodeal triangle thus poorly defined (
<figureCitation id="71B92A6FAC2543070F08FD8C8AC50420" box="[1052,1184,583,609]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77F</figureCitation>
). Mesepisternum with moderately long plumose whitish to light brownish hairs, not equalling length of scape. Scutum, scutellum, and tegulae covered with very sort light brown squamous hairs, not covering and obscuring surface, hairs becoming sparse on discs of scutum and scutellum. Propodeal corbicula incomplete, dorsal fringe composed of white weakly plumose hairs, internal surface densely covered with abundant long white weakly plumose hairs that do not obscure the underlying surface. Legs brown, apical tarsal segments lightened orange, pubescence whitish to light brownish. Flocculus complete, composed of whitish plumose hairs; femoral and tibial scopae composed of simple white hairs. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange, nervulus interstitial.
</paragraph>
<paragraph id="E93D36EAAC2543070BD3FCAC8EBD02E0" blockId="124.[151,1437,151,2013]" pageId="124" pageNumber="125">
<emphasis id="DBF6EAF8AC2543070BD3FCAC8F2005C1" box="[199,325,871,896]" italics="true" pageId="124" pageNumber="125">Metasoma:</emphasis>
Tergal discs predominantly dark, marginal areas and parts of discs of T12 lightened reddish brown (
<figureCitation id="71B92A6FAC2543070BF9FC408F1605E4" box="[237,371,907,933]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77G</figureCitation>
). Disc of T1 medially with elongate punctures, punctures separated on average by 12 puncture diameters, laterally and on declivity with dense simple punctures, punctures separated by 0.51 puncture diameters. Discs of T24 densely and regularly punctate, punctures separated by 0.5 puncture diameters, interspaces polished and shining. Marginal areas of T24 strongly and clearly depressed, T14 with strong and dense apical whitish to yellowish hairbands, covering entire marginal area and extending onto following tergal disc, hairbands complete, obscuring underlying surface. Apical fringe and hairs flanking pygidial plate whitish to brownish. Pygidial plate laterally with raised margin, apically with deep broad emargination, medially slightly humped, surface dull (
<figureCitation id="71B92A6FAC2543070E45FBA88EA902E0" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77G</figureCitation>
).
</paragraph>
<paragraph id="E93D36EAAC2543070BD3FB608FED0284" blockId="124.[151,1437,151,2013]" box="[199,392,1195,1221]" pageId="124" pageNumber="125">
<emphasis id="DBF6EAF8AC2543070BD3FB608F6E0284" bold="true" box="[199,267,1195,1221]" pageId="124" pageNumber="125">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2543050BD3FB058A580554" lastPageId="126" lastPageNumber="127" pageId="124" pageNumber="125" type="diagnosis">
<paragraph id="E93D36EAAC2543070BD3FB058C210048" blockId="124.[151,1437,151,2013]" pageId="124" pageNumber="125">
<emphasis id="DBF6EAF8AC2543070BD3FB058F2702A9" bold="true" box="[199,322,1230,1256]" pageId="124" pageNumber="125">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC2543070A53FB048C7F02A9" authorityName="WOOD" authorityYear="2024" box="[327,538,1231,1257]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2543070A53FB048C7F02A9" box="[327,538,1231,1257]" italics="true" pageId="124" pageNumber="125">Andrena concisura</emphasis>
</taxonomicName>
presents an unusual morphology which is unique within the Central Asian fauna. Specifically, it presents a domed shining clypeus (
<figureCitation id="71B92A6FAC25430709C3FB388D38034C" box="[727,861,1267,1293]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77B</figureCitation>
), a triangular process of the labrum (
<figureCitation id="71B92A6FAC2543070E1DFB388BF5034C" box="[1289,1424,1267,1293]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77C</figureCitation>
), impunctate and shining triangular areas adjacent to the lateral ocelli (
<figureCitation id="71B92A6FAC25430708D1FADC8A350370" box="[965,1104,1303,1329]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77D</figureCitation>
), the pronotum with a weak humeral angle, the scutum, scutellum, and tegulae with short squamous hairs (
<figureCitation id="71B92A6FAC25430708E1FAF08A130314" box="[1013,1142,1339,1365]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77E</figureCitation>
), the mesepisternum shiny and punctate, the propodeal triangle very weakly defined with minimal sculpture (
<figureCitation id="71B92A6FAC2543070F55FA948AA60338" box="[1089,1219,1375,1401]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77F</figureCitation>
), tergal discs T24 very densely punctate (
<figureCitation id="71B92A6FAC2543070A8AFA488C4003DC" box="[414,549,1411,1437]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77G</figureCitation>
; contrasting T1 with sparse punctures), marginal areas strongly depressed and entirely covered with dense complete hairbands, and the pygidial plate with a clear and deep emargination medially (
<figureCitation id="71B92A6FAC2543070B8BFA008F4003A4" box="[159,293,1483,1509]" captionStart="FIGURE 77" captionStartId="125.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@125.[215,1373,181,1919]" captionTargetPageId="125" captionText="FIGURE 77. Andrena (incertae sedis) concisura spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view." figureDoi="http://doi.org/10.5281/zenodo.13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="124" pageNumber="125">Figure 77H</figureCitation>
). It lacks other distinctive characters such as a broadened hind tibial spur, plumose hairs, or spines on the posterior face of the hind femorae.
</paragraph>
<paragraph id="E93D36EAAC2543070BD3F9D88F7E019C" blockId="124.[151,1437,151,2013]" pageId="124" pageNumber="125">
Amongst Central Asian species it could perhaps be compared to
<taxonomicName id="2E824D69AC25430708D7F9D88EAA0010" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="carinifrons" subGenus="Carinandrena">
<emphasis id="DBF6EAF8AC25430708D7F9D88DB9006C" box="[963,988,1555,1581]" italics="true" pageId="124" pageNumber="125">A.</emphasis>
(
<emphasis id="DBF6EAF8AC25430708E4F9D88AEB006C" box="[1008,1166,1555,1581]" italics="true" pageId="124" pageNumber="125">Carinandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2543070FB6F9D88B78006C" box="[1186,1309,1555,1581]" italics="true" pageId="124" pageNumber="125">carinifrons</emphasis>
Morawitz, 1876
</taxonomicName>
(see
<bibRefCitation id="8D134B1BAC2543070A1FF9FC8C720010" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[267,535,1591,1617]" pageId="124" pageNumber="125" pagination="105 - 176" refId="ref87433" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. Zookeys, 1120, 105 - 176. https: // doi. org / 10.3897 / zookeys. 1120.90206" type="journal article" year="2022">
Astafurova
<emphasis id="DBF6EAF8AC2543070A84F9FC8FA60010" box="[400,451,1591,1617]" italics="true" pageId="124" pageNumber="125">et al</emphasis>
. 2022b
</bibRefCitation>
for illustrations) due to the more or less quadrate head, the domed and shining clypeus, and the shiny impunctate areas adjacent to the lateral ocelli, but there are numerous differences such as the triangular process of the labrum (in
<taxonomicName id="2E824D69AC2543070943F9B48C9700D8" authorityName="Morawitz" authorityYear="1876" box="[599,754,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="carinifrons">
<emphasis id="DBF6EAF8AC2543070943F9B48C9700D8" box="[599,754,1663,1689]" italics="true" pageId="124" pageNumber="125">A. carinifrons</emphasis>
</taxonomicName>
with the process of the labrum apically narrowly but clearly emarginate), the lack of a longitudinal carina on the frons (in
<taxonomicName id="2E824D69AC2543070853F9688D8400FC" authorityName="Morawitz" authorityYear="1876" box="[839,993,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="carinifrons">
<emphasis id="DBF6EAF8AC2543070853F9688D8400FC" box="[839,993,1699,1725]" italics="true" pageId="124" pageNumber="125">A. carinifrons</emphasis>
</taxonomicName>
with a conspicuous raised carina here), and terga with strong apical hairbands that cover and obscure the marginal areas (in
<taxonomicName id="2E824D69AC2543070F5BF90C8A8F00A0" authorityName="Morawitz" authorityYear="1876" box="[1103,1258,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="carinifrons">
<emphasis id="DBF6EAF8AC2543070F5BF90C8A8F00A0" box="[1103,1258,1735,1761]" italics="true" pageId="124" pageNumber="125">A. carinifrons</emphasis>
</taxonomicName>
with terga with hairbands predominantly arising from the tergal bases, marginal areas predominantly hairless, certainly medially). Two Central Asian species formerly placed in the polyphyletic subgenus
<taxonomicName id="2E824D69AC25430708C5F8C48B680168" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[977,1293,1807,1833]" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC25430708C5F8C48A390168" box="[977,1116,1807,1833]" italics="true" pageId="124" pageNumber="125">Poliandrena</emphasis>
Warncke, 1968
</taxonomicName>
(see
<bibRefCitation id="8D134B1BAC2543070E5EF8C48F7B010C" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." pageId="124" pageNumber="125" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC2543070B83F8F88EAD010C" box="[151,200,1843,1869]" italics="true" pageId="124" pageNumber="125">et al</emphasis>
. 2022b
</bibRefCitation>
) with squamous hairs on the scutum and scutellum (
<taxonomicName id="2E824D69AC254307087DF8F88AC8010C" authority="Morawitz, 1893" authorityName="Morawitz" authorityYear="1893" box="[873,1197,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="jakowlevi">
<emphasis id="DBF6EAF8AC254307087DF8F88D96010C" box="[873,1011,1843,1869]" italics="true" pageId="124" pageNumber="125">A. jakowlevi</emphasis>
<bibRefCitation id="8D134B1BAC25430708EEF8F88AC8010C" author="Morawitz, F." box="[1018,1197,1843,1869]" pageId="124" pageNumber="125" pagination="3 - 87" refId="ref89480" refString="Morawitz, F. (1893) Supplement zur Bienenfauna Turkestans. Horae Societatis Entomologicae Rossicae, 28, 3 - 87." type="journal article" year="1893">Morawitz, 1893</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2543070FF7F8F88F3A0130" authority="Osytshnjuk, 1993" authorityName="Osytshnjuk" authorityYear="1993" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="kryzhanovskii">
<emphasis id="DBF6EAF8AC2543070FF7F8F88BF9010C" box="[1251,1436,1843,1869]" italics="true" pageId="124" pageNumber="125">A. kryzhanovskii</emphasis>
Osytshnjuk, 1993
</taxonomicName>
) are strongly different, as these hairs are very short and dense, forming a clear patch of hairs as in several members of the subgenus
<taxonomicName id="2E824D69AC2543070955F8B08DF401D4" authority="Hedicke, 1933" authorityName="Hedicke" authorityYear="1933" box="[577,913,1915,1941]" class="Insecta" family="Andrenidae" genus="Lepidandrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC2543070955F8B08CB801D4" box="[577,733,1915,1941]" italics="true" pageId="124" pageNumber="125">Lepidandrena</emphasis>
Hedicke, 1933
</taxonomicName>
, whereas in
<taxonomicName id="2E824D69AC2543070F31F8B08AD001D5" authorityName="WOOD" authorityYear="2024" box="[1061,1205,1915,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2543070F31F8B08AD001D5" box="[1061,1205,1915,1941]" italics="true" pageId="124" pageNumber="125">A. concisura</emphasis>
</taxonomicName>
the squamous hairs are individually longer and also individualised, with each hair standing alone and distinctly separated from its neighbours.
</paragraph>
<caption id="BDFD6662AC2443060B83F85C8F8301B6" ID-DOI="http://doi.org/10.5281/zenodo.13209477" ID-Zenodo-Dep="13209477" httpUri="https://zenodo.org/record/13209477/files/figure.png" pageId="125" pageNumber="126" startId="125.[151,250,1943,1967]" targetBox="[215,1373,181,1919]" targetPageId="125" targetType="figure">
<paragraph id="E93D36EAAC2443060B83F85C8F8301B6" blockId="125.[151,1436,1943,2039]" pageId="125" pageNumber="126">
<emphasis id="DBF6EAF8AC2443060B83F85C8F4501F1" bold="true" box="[151,288,1943,1968]" pageId="125" pageNumber="126">FIGURE 77.</emphasis>
<taxonomicName id="2E824D69AC2443060A33F85C8CED01EE" authorityName="WOOD" authorityYear="2024" box="[295,648,1943,1967]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="concisura" status="sp. nov.">
<emphasis id="DBF6EAF8AC2443060A33F85C8F1801EE" box="[295,381,1943,1967]" italics="true" pageId="125" pageNumber="126">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8AC2443060930F8538CED01EE" box="[548,648,1944,1967]" italics="true" pageId="125" pageNumber="126">concisura</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC244306099BF8538C9301F1" bold="true" box="[655,758,1944,1968]" pageId="125" pageNumber="126">
<taxonomicNameLabel id="C0C55783AC244306099BF8538C9301F1" box="[655,758,1944,1968]" pageId="125" pageNumber="126" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Head, dorsal view; E. Scutum, dorsolateral view; F. Propodeum, dorsal view; G. Terga, dorsal view; H. Pygidial plate, posterior view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC2743050BD3FF5C8A580554" blockId="126.[151,1437,151,1545]" pageId="126" pageNumber="127">
<taxonomicName id="2E824D69AC2743050BD3FF5C8FF906F1" authorityName="WOOD" authorityYear="2024" box="[199,412,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2743050BD3FF5C8FF906F1" box="[199,412,151,177]" italics="true" pageId="126" pageNumber="127">Andrena concisura</emphasis>
</taxonomicName>
is more comparable to the two species that make up the
<taxonomicName id="2E824D69AC2743050F08FF5C8A1506F1" authorityName="Schmiedeknecht" authorityYear="1900" box="[1052,1136,151,176]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050F08FF5C8A1506F1" box="[1052,1136,151,176]" italics="true" pageId="126" pageNumber="127">arsinoe</emphasis>
</taxonomicName>
-group (
<taxonomicName id="2E824D69AC2743050FD8FF5C8B0B06F0" authorityName="Warncke" authorityYear="1968" box="[1228,1390,151,177]" pageId="126" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC2743050FD8FF5C8B0B06F0" box="[1228,1390,151,177]" italics="true" pageId="126" pageNumber="127">Graecandrena</emphasis>
</taxonomicName>
I in
<bibRefCitation id="8D134B1BAC2743050B83FF708F1B0694" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[151,382,187,213]" pageId="126" pageNumber="127" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC2743050BE7FF708F430694" box="[243,294,187,213]" italics="true" pageId="126" pageNumber="127">et al</emphasis>
. 2022b
</bibRefCitation>
) whose taxa were previously placed into the subgenera
<taxonomicName id="2E824D69AC2743050F17FF708AE00694" authorityName="Warncke" authorityYear="1968" box="[1027,1157,187,213]" pageId="126" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC2743050F17FF708AE00694" box="[1027,1157,187,213]" italics="true" pageId="126" pageNumber="127">Aciandrena</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2743050FABFF708B040694" authorityName="Warncke" authorityYear="1968" box="[1215,1377,187,213]" pageId="126" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC2743050FABFF708B040694" box="[1215,1377,187,213]" italics="true" pageId="126" pageNumber="127">Graecandrena</emphasis>
</taxonomicName>
. The two species
<taxonomicName id="2E824D69AC2743050A37FF148CC406B8" authority="Schmiedeknecht, 1900" authorityName="Schmiedeknecht" authorityYear="1900" box="[291,673,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050A37FF148FFD06B9" box="[291,408,223,249]" italics="true" pageId="126" pageNumber="127">A. arsinoe</emphasis>
Schmiedeknecht, 1900
</taxonomicName>
(
<collectingCountry id="9195767AAC27430509A6FF148C9D06B8" box="[690,760,223,249]" name="Libya" pageId="126" pageNumber="127">Libya</collectingCountry>
,
<collectingCountry id="9195767AAC2743050810FF148D2F06B8" box="[772,842,223,249]" name="Egypt" pageId="126" pageNumber="127">Egypt</collectingCountry>
,
<collectingCountry id="9195767AAC2743050842FF148DF206B8" box="[854,919,223,249]" name="Israel" pageId="126" pageNumber="127">Israel</collectingCountry>
,
<collectingCountry id="9195767AAC27430508B0FF148D9106B8" box="[932,1012,223,249]" name="Jordan" pageId="126" pageNumber="127">Jordan</collectingCountry>
*,
<collectingCountry id="9195767AAC2743050F18FF148AC706B8" box="[1036,1186,223,249]" name="Saudi Arabia" pageId="126" pageNumber="127">Saudi Arabia</collectingCountry>
,
<collectingCountry id="9195767AAC2743050FBBFF148A9C06B8" box="[1199,1273,223,249]" name="Oman" pageId="126" pageNumber="127">Oman</collectingCountry>
*,
<collectingCountry id="9195767AAC2743050E05FF148F64075C" name="United Arab Emirates" pageId="126" pageNumber="127">United Arab Emirates</collectingCountry>
*;
<bibRefCitation id="8D134B1BAC2743050A03FEC88C0B075C" author="Gusenleitner, F. &amp; Schwarz, M." box="[279,622,259,285]" pageId="126" pageNumber="127" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC274305096CFEC88D29075C" author="Pisanty, G. &amp; Scheuchl, E. &amp; Dorchin, N." box="[632,844,259,285]" pageId="126" pageNumber="127" pagination="151 - 188" refId="ref89992" refString="Pisanty, G., Scheuchl, E. &amp; Dorchin, N. (2018) Taxonomic review of the subgenus Andrena (Poecilandrena) (Hymenoptera: Andrenidae) in Israel and the Levant. Zootaxa, 4374 (2), 151 - 188. https: // doi. org / 10.11646 / zootaxa. 4374.2.1" type="journal article" year="2018">
Pisanty
<emphasis id="DBF6EAF8AC27430509C6FEC88D66075C" box="[722,771,259,285]" italics="true" pageId="126" pageNumber="127">et al</emphasis>
. 2018
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC2743050842FEC88A10075C" author="Ascher, J. S. &amp; Pickering, J." box="[854,1141,259,285]" pageId="126" pageNumber="127" refId="ref87181" refString="Ascher, J. S. &amp; Pickering, J. (2023) Discover Life bee species guide and world checklist (Hymenoptera: Apidae: Anthophila). Available from: https: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species &amp; flags = HAS (accessed 28 May 2024)" type="url" year="2023">Ascher &amp; Pickering 2023</bibRefCitation>
) and
<taxonomicName id="2E824D69AC2743050FA7FEC88EAA0700" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="amicula">
<emphasis id="DBF6EAF8AC2743050FA7FEC88B49075C" box="[1203,1324,259,285]" italics="true" pageId="126" pageNumber="127">A. amicula</emphasis>
<bibRefCitation id="8D134B1BAC2743050E26FEC88EAA0700" author="Warncke, K." pageId="126" pageNumber="127" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="9195767AAC2743050BC9FEEC8F500700" box="[221,309,295,321]" name="Algeria" pageId="126" pageNumber="127">Algeria</collectingCountry>
,
<collectingCountry id="9195767AAC2743050A54FEEC8FE30700" box="[320,390,295,321]" name="Egypt" pageId="126" pageNumber="127">Egypt</collectingCountry>
,
<collectingCountry id="9195767AAC2743050A84FEEC8F850700" box="[400,480,295,321]" name="Eritrea" pageId="126" pageNumber="127">Eritrea</collectingCountry>
;
<collectingCountry id="9195767AAC2743050AFEFEEC8CE50700" box="[490,640,295,321]" name="Saudi Arabia" pageId="126" pageNumber="127">Saudi Arabia</collectingCountry>
*,
<collectingCountry id="9195767AAC2743050982FEEC8C840700" box="[662,737,295,321]" name="Oman" pageId="126" pageNumber="127">Oman</collectingCountry>
*;
<bibRefCitation id="8D134B1BAC27430509E2FEEC8DFF0700" author="Warncke, K." box="[758,922,295,321]" pageId="126" pageNumber="127" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke 1967</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC27430508B7FEEC8AA50700" author="Ascher, J. S. &amp; Pickering, J." box="[931,1216,295,321]" pageId="126" pageNumber="127" refId="ref87181" refString="Ascher, J. S. &amp; Pickering, J. (2023) Discover Life bee species guide and world checklist (Hymenoptera: Apidae: Anthophila). Available from: https: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species &amp; flags = HAS (accessed 28 May 2024)" type="url" year="2023">Ascher &amp; Pickering 2023</bibRefCitation>
) present an unusual genital capsule that has the gonostyli strongly constricted medially with the apexes hyaline and finger-like. As the male of
<taxonomicName id="2E824D69AC2743050BE3FEA48FE307C9" authorityName="WOOD" authorityYear="2024" box="[247,390,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2743050BE3FEA48FE307C9" box="[247,390,367,393]" italics="true" pageId="126" pageNumber="127">A. concisura</emphasis>
</taxonomicName>
is unknown, comparison in this sex is currently not possible. However, females are similar to
<taxonomicName id="2E824D69AC2743050BA2FE588F4E07ED" authorityName="Schmiedeknecht" authorityYear="1900" box="[182,299,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050BA2FE588F4E07ED" box="[182,299,403,429]" italics="true" pageId="126" pageNumber="127">A. arsinoe</emphasis>
</taxonomicName>
due to the semi-squamous mesonotal hairs, the domed and shining clypeus (in
<taxonomicName id="2E824D69AC2743050FAFFE588B5307EC" authorityName="Warncke" authorityYear="1967" box="[1211,1334,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="amicula">
<emphasis id="DBF6EAF8AC2743050FAFFE588B5307EC" box="[1211,1334,403,429]" italics="true" pageId="126" pageNumber="127">A. amicula</emphasis>
</taxonomicName>
with the clypeus much shorter and densely punctate, thus lacking large shining interspaces&gt;1 puncture diameter), the weakly sculptured propodeal triangle, the strong tergal hairbands, the comparatively stronger punctures on T24 compared to T1, and the pygidial plate with a slight hint of a median emargination. The two species are easily separated, as
<taxonomicName id="2E824D69AC2743050E90FE348F60047D" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2743050E90FE348F60047D" italics="true" pageId="126" pageNumber="127">A. concisura</emphasis>
</taxonomicName>
presents dark terga that are very strongly and densely punctate, punctures almost confluent (in
<taxonomicName id="2E824D69AC2743050E3EFDE88BF9047D" authorityName="Schmiedeknecht" authorityYear="1900" box="[1322,1436,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050E3EFDE88BF9047D" box="[1322,1436,547,573]" italics="true" pageId="126" pageNumber="127">A. arsinoe</emphasis>
</taxonomicName>
the terga are extensively red-marked and weakly punctate, punctures shallow and separated by around 1 puncture diameter), the clypeus is more densely punctate with punctures separated by 0.52 puncture diameters (in
<taxonomicName id="2E824D69AC2743050E3EFDA08BF904C5" authorityName="Schmiedeknecht" authorityYear="1900" box="[1322,1436,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050E3EFDA08BF904C5" box="[1322,1436,619,645]" italics="true" pageId="126" pageNumber="127">A. arsinoe</emphasis>
</taxonomicName>
with clypeal punctures separated by&gt;2 puncture diameters), and the pygidial plate is apically clearly incised (in
<taxonomicName id="2E824D69AC2743050B83FD788F6E048D" authorityName="Schmiedeknecht" authorityYear="1900" box="[151,267,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050B83FD788F6E048D" box="[151,267,691,717]" italics="true" pageId="126" pageNumber="127">A. arsinoe</emphasis>
</taxonomicName>
with the apex of the pygidial plate rounded to presenting a hint of an emargination). No concrete action is taken until phylogenetic data are available that can support or oppose an association with the
<taxonomicName id="2E824D69AC2743050FCAFD1C8B5704B1" authorityName="Schmiedeknecht" authorityYear="1900" box="[1246,1330,727,752]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050FCAFD1C8B5704B1" box="[1246,1330,727,752]" italics="true" pageId="126" pageNumber="127">arsinoe</emphasis>
</taxonomicName>
-group of species, and the status of the monotypic subgenus
<taxonomicName id="2E824D69AC27430509DFFD308A5D0554" authority="Osytshnjuk, 1993" authorityName="Osytshnjuk" authorityYear="1993" box="[715,1080,763,789]" class="Insecta" family="Andrenidae" genus="Carinandrena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC27430509DFFD308D0C0554" box="[715,873,763,789]" italics="true" pageId="126" pageNumber="127">Carinandrena</emphasis>
Osytshnjuk, 1993
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2743050BD3FCD48DFC05C0" pageId="126" pageNumber="127" type="discussion">
<paragraph id="E93D36EAAC2743050BD3FCD48DFC05C0" blockId="126.[151,1437,151,1545]" pageId="126" pageNumber="127">
<emphasis id="DBF6EAF8AC2743050BD3FCD48F5E0578" bold="true" box="[199,315,799,825]" pageId="126" pageNumber="127">Remarks.</emphasis>
The observed flight period of early September is unusual, as very few univoltine Palaearctic
<taxonomicName id="2E824D69AC2743050E29FCD48BF90578" box="[1341,1436,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC2743050E29FCD48BF90578" box="[1341,1436,799,825]" italics="true" pageId="126" pageNumber="127">Andrena</emphasis>
</taxonomicName>
fly this late in the summer, with most examples coming from the subgenera
<taxonomicName id="2E824D69AC2743050F04FC888AA6051C" authorityName="Hedicke" authorityYear="1933" box="[1040,1219,835,861]" pageId="126" pageNumber="150" rank="subGenus" subGenus="Cnemidandrena">
<emphasis id="DBF6EAF8AC2743050F04FC888AA6051C" box="[1040,1219,835,861]" italics="true" pageId="126" pageNumber="127">Cnemidandrena</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2743050E14FC888FBD05C0" authority="Hirashima &amp; LaBerge, 1964" authorityName="Hirashima &amp; LaBerge" authorityYear="1964" class="Insecta" family="Andrenidae" genus="Habromelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC2743050E14FC888BF9051C" box="[1280,1436,835,861]" italics="true" pageId="126" pageNumber="127">Habromelissa</emphasis>
Hirashima &amp; LaBerge, 1964
</taxonomicName>
, to which
<taxonomicName id="2E824D69AC2743050958FCAC8CBC05C1" authorityName="WOOD" authorityYear="2024" box="[588,729,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="concisura">
<emphasis id="DBF6EAF8AC2743050958FCAC8CBC05C1" box="[588,729,871,897]" italics="true" pageId="126" pageNumber="127">A. concisura</emphasis>
</taxonomicName>
does not belong.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2743050BD3FC408D3F0588" pageId="126" pageNumber="127" type="etymology">
<paragraph id="E93D36EAAC2743050BD3FC408D3F0588" blockId="126.[151,1437,151,1545]" pageId="126" pageNumber="127">
<emphasis id="DBF6EAF8AC2743050BD3FC408F2B05E4" bold="true" box="[199,334,907,933]" pageId="126" pageNumber="127">Etymology.</emphasis>
Feminine singular nominative form of the Latin adjective
<emphasis id="DBF6EAF8AC27430508C2FC408A2A05E5" box="[982,1103,907,932]" italics="true" pageId="126" pageNumber="127">concisurus</emphasis>
meaning cut, incision, gap, in reference to the pygidial plate which is clearly incised apically.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2743050BD3FC198B270048" pageId="126" pageNumber="127" type="distribution">
<paragraph id="E93D36EAAC2743050BD3FC198FB805AC" blockId="126.[151,1437,151,1545]" box="[199,477,978,1005]" pageId="126" pageNumber="127">
<emphasis id="DBF6EAF8AC2743050BD3FC198F0405AD" bold="true" box="[199,353,978,1004]" pageId="126" pageNumber="127">Distribution.</emphasis>
<collectingCountry id="9195767AAC2743050A73FC188FBC05AC" box="[359,473,979,1005]" name="Tajikistan" pageId="126" pageNumber="127">Tajikistan</collectingCountry>
.
</paragraph>
<paragraph id="E93D36EAAC2743050BD3FC3D8DB10284" blockId="126.[151,1437,151,1545]" pageId="126" pageNumber="127">
<materialsCitation id="59EA3CB7AC2743050BD3FC3D8F3B0276" collectingDate="1989-03-31" collectorName="M. Schwarz" country="Algeria" location="Tamanrasset" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1" stateProvince="Tamanghasset">
<emphasis id="DBF6EAF8AC2743050BD3FC3D8F930251" bold="true" box="[199,502,1014,1040]" pageId="126" pageNumber="127">
<typeStatus id="36398848AC2743050BD3FC3D8F930251" box="[199,502,1014,1040]" isOtherTypeStatus="true" pageId="126" pageNumber="127">Other material examined</typeStatus>
</emphasis>
(
<taxonomicName id="2E824D69AC274305091CFC3C8CAC0250" authorityName="Warncke" authorityYear="1967" box="[520,713,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="amicula">
<emphasis id="DBF6EAF8AC274305091CFC3C8CAC0250" box="[520,713,1015,1041]" italics="true" pageId="126" pageNumber="127">Andrena amicula</emphasis>
</taxonomicName>
)
<emphasis id="DBF6EAF8AC27430509CEFC3D8D030251" bold="true" box="[730,870,1014,1040]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC27430509CEFC3D8D040251" box="[730,865,1014,1040]" name="Algeria" pageId="126" pageNumber="127">ALGERIA</collectingCountry>
:
</emphasis>
<quantity id="2E7A9B0FAC274305087AFC3C8DD30250" box="[878,950,1015,1041]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="126" pageNumber="127" unit="km" value="60.0">60 km</quantity>
E of
<collectingRegion id="2B46F808AC27430508E2FC3C8AE00250" box="[1014,1157,1015,1041]" country="Algeria" name="Tamanghasset" pageId="126" pageNumber="127">Tamanrasset</collectingRegion>
,
<date id="9D3C102AAC2743050F84FC3C8B680250" box="[1168,1293,1015,1041]" pageId="126" pageNumber="127" value="1989-03-31">
<collectingDate id="8D78E9C2AC2743050F84FC3C8B680250" box="[1168,1293,1015,1041]" pageId="126" pageNumber="127" value="1989-03-31">31.iii.1989</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050E0DFC3C8B590250" box="[1305,1340,1015,1041]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050E68FC3C8E9F0274" pageId="126" pageNumber="127">M. Schwarz</collectorName>
, OÖLM
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050A7FFBD18CF20218" collectingDate="1933-01-26" collectionCode="ZMHB" collectorName="H. Preisner" country="Egypt" determinerName="K. Warncke" location="Gabal Elba" pageId="126" pageNumber="127" specimenCount="2" specimenCount-male="2">
<emphasis id="DBF6EAF8AC2743050A7FFBD18FB60275" bold="true" box="[363,467,1050,1076]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050A7FFBD18FAB0275" box="[363,462,1050,1076]" name="Egypt" pageId="126" pageNumber="127">EGYPT</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC2743050AC8FBD08C3B0274" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050AC8FBD08C3B0274" box="[476,606,1051,1077]" country="Egypt" name="Gabal Elba" pageId="126" pageNumber="127">Gabal Elba</location>
,
<date id="9D3C102AAC274305097FFBD08CB20274" box="[619,727,1051,1077]" pageId="126" pageNumber="127" value="1933-01-26">
<collectingDate id="8D78E9C2AC274305097FFBD08CB20274" box="[619,727,1051,1077]" pageId="126" pageNumber="127" value="1933-01-26">26.i.1933</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC27430509F1FBD08D6F0274" box="[741,778,1051,1077]" count="2" pageId="126" pageNumber="127" type="male">2♂</specimenCount>
, det.
<determinerName id="1CE1DA2EAC274305085EFBD08DB10274" box="[842,980,1051,1077]" pageId="126" pageNumber="127">K. Warncke</determinerName>
, leg.
<collectorName id="4477533CAC2743050F07FBD08AF30274" box="[1043,1174,1051,1077]" pageId="126" pageNumber="127">H. Preisner</collectorName>
,
<collectionCode id="8F93AE2FAC2743050FB6FBD08A960274" box="[1186,1267,1051,1077]" pageId="126" pageNumber="127">ZMHB</collectionCode>
(
<emphasis id="DBF6EAF8AC2743050E12FBD08B190274" bold="true" box="[1286,1404,1051,1077]" pageId="126" pageNumber="127">
<typeStatus id="36398848AC2743050E12FBD08B190274" box="[1286,1404,1051,1077]" pageId="126" pageNumber="127" type="paratype">paratypes</typeStatus>
</emphasis>
of “
<taxonomicName id="2E824D69AC2743050BB0FBF48FCB0218" authority="&quot; Alfken" authorityName="Alfken" box="[164,430,1087,1113]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="elbae">
<emphasis id="DBF6EAF8AC2743050BB0FBF48F2C0218" box="[164,329,1087,1113]" italics="true" pageId="126" pageNumber="127">Andrena elbae</emphasis>
” Alfken
</taxonomicName>
, unpublished name)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC27430509B1FBF48FBD023C" collectingDate="1933-01-26" collectorName="H. Preisner" country="Egypt" determinerName="K. Warncke" location="Gabal Elba" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1">
<location id="EC5D6031AC27430509B1FBF48D410218" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC27430509B1FBF48D410218" box="[677,804,1087,1113]" country="Egypt" name="Gabal Elba" pageId="126" pageNumber="127">Gabal Elba</location>
[Jabal`Ilbah],
<date id="9D3C102AAC27430508C3FBF48A260218" box="[983,1091,1087,1113]" pageId="126" pageNumber="127" value="1933-01-26">
<collectingDate id="8D78E9C2AC27430508C3FBF48A260218" box="[983,1091,1087,1113]" pageId="126" pageNumber="127" value="1933-01-26">26.i.1933</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050F45FBF48A100218" box="[1105,1141,1087,1113]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
, det.
<determinerName id="1CE1DA2EAC2743050FA3FBF48B240218" box="[1207,1345,1087,1113]" pageId="126" pageNumber="127">K. Warncke</determinerName>
, leg.
<collectorName id="4477533CAC2743050E95FBF48E90023C" pageId="126" pageNumber="127">H. Preisner</collectorName>
, OÖLM (
<emphasis id="DBF6EAF8AC2743050A76FBA88FA8023C" bold="true" box="[354,461,1123,1149]" pageId="126" pageNumber="127">
<typeStatus id="36398848AC2743050A76FBA88FA8023C" box="[354,461,1123,1149]" pageId="126" pageNumber="127" type="paratype">paratype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050AF7FBA88BF3023C" box="[483,1430,1123,1151]" collectorName="O. Schmiedeknecht" country="Egypt" location="Aegyptien" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<location id="EC5D6031AC2743050AF7FBA88C3D023C" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050AF7FBA88C3D023C" box="[483,600,1123,1149]" country="Egypt" name="Aegyptien" pageId="126" pageNumber="127">Aegyptien</location>
[no further information],
<specimenCount id="FF84FD63AC274305086DFBA88DF9023C" box="[889,924,1123,1149]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC27430508CDFBA88AD2023C" box="[985,1207,1123,1149]" pageId="126" pageNumber="127">O. Schmiedeknecht</collectorName>
, OÖLM (
<emphasis id="DBF6EAF8AC2743050E30FBA88BEE023C" bold="true" box="[1316,1419,1123,1149]" pageId="126" pageNumber="127">
<typeStatus id="36398848AC2743050E30FBA88BEE023C" box="[1316,1419,1123,1149]" pageId="126" pageNumber="127" type="holotype">holotype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050B83FB4D8A3D02E0" box="[151,1112,1158,1185]" collectingDate="2022-03-07" collectionCode="CSE, TJWC" collectorName="C. Schmid-Egger" country="Oman" location="Al Fulaij" pageId="126" pageNumber="127" specimenCount="3" specimenCount-female="3">
<emphasis id="DBF6EAF8AC2743050B83FB4D8E9C02E1" bold="true" box="[151,249,1158,1184]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050B83FB4D8E9102E1" box="[151,244,1158,1184]" name="Oman" pageId="126" pageNumber="127">OMAN</collectingCountry>
:
</emphasis>
<locationDeviation id="F88F2FC8AC2743050A14FB4C8FCC02E0" box="[256,425,1159,1185]" location="Sur" pageId="126" pageNumber="127">
<quantity id="2E7A9B0FAC2743050A14FB4C8F2D02E0" box="[256,328,1159,1185]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="126" pageNumber="127" unit="km" value="10.0">10 km</quantity>
SW Sur
</locationDeviation>
,
<location id="EC5D6031AC2743050AA6FB4C8C7C02E0" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050AA6FB4C8C7C02E0" box="[434,537,1159,1185]" country="Oman" name="Al Fulaij" pageId="126" pageNumber="127">Al Fulaij</location>
,
<date id="9D3C102AAC2743050931FB4C8CF602E0" box="[549,659,1159,1185]" pageId="126" pageNumber="127" value="2022-03-07">
<collectingDate id="8D78E9C2AC2743050931FB4C8CF602E0" box="[549,659,1159,1185]" pageId="126" pageNumber="127" value="2022-03-07">7.iii.2022</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC274305098BFB4C8CA602E0" box="[671,707,1159,1185]" count="3" pageId="126" pageNumber="127" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050814FB4C8DA002E0" box="[768,965,1159,1185]" pageId="126" pageNumber="127">C. Schmid-Egger</collectorName>
,
<collectionCode id="8F93AE2FAC27430508C4FB4C8A6102E0" box="[976,1028,1159,1185]" pageId="126" pageNumber="127">CSE</collectionCode>
/
<collectionCode id="8F93AE2FAC2743050F19FB4C8A3D02E0" box="[1037,1112,1159,1185]" pageId="126" pageNumber="127">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050F77FB4D8DB40284" collectingDate="1983-01-23" collectionCode="NHMUK" collectorName="K. M. Guichard" country="Saudi Arabia" elevation="500" location="Al Ha'ir" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1" stateProvince="Ar Riyad">
<emphasis id="DBF6EAF8AC2743050F77FB4D8B5D02E1" bold="true" box="[1123,1336,1158,1184]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050F77FB4D8B5602E1" box="[1123,1331,1158,1184]" name="Saudi Arabia" pageId="126" pageNumber="127">SAUDI ARABIA</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC2743050E2AFB4C8BF902E0" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050E2AFB4C8BF902E0" box="[1342,1436,1159,1185]" country="Saudi Arabia" name="Al Ha'ir" pageId="126" pageNumber="127" stateProvince="Ar Riyad">Al Hair</location>
[Al Haeer],
<collectingRegion id="2B46F808AC2743050A0AFB608F170284" box="[286,370,1195,1221]" country="Saudi Arabia" name="Ar Riyad" pageId="126" pageNumber="127">Riyadh</collectingRegion>
,
<quantity id="2E7A9B0FAC2743050A69FB618FA30284" box="[381,454,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="126" pageNumber="127" unit="m" value="500.0">
<elevation id="62AFD1D9AC2743050A69FB618FA30284" box="[381,454,1194,1221]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="126" pageNumber="127" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="9D3C102AAC2743050AC5FB608C5B0284" box="[465,574,1195,1221]" pageId="126" pageNumber="127" value="1983-01-23">
<collectingDate id="8D78E9C2AC2743050AC5FB608C5B0284" box="[465,574,1195,1221]" pageId="126" pageNumber="127" value="1983-01-23">23.i.1983</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC274305095DFB608C0B0284" box="[585,622,1195,1221]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC27430509BEFB608D320284" box="[682,855,1195,1221]" pageId="126" pageNumber="127">K.M. Guichard</collectorName>
,
<collectionCode id="8F93AE2FAC2743050877FB608DB40284" box="[867,977,1195,1221]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="E93D36EAAC2743050BD3FB048B270048" blockId="126.[151,1437,151,1545]" pageId="126" pageNumber="127">
(
<taxonomicName id="2E824D69AC2743050BC5FB048FEE02A9" authorityName="Schmiedeknecht" authorityYear="1900" box="[209,395,1231,1257]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="arsinoe">
<emphasis id="DBF6EAF8AC2743050BC5FB048FEE02A9" box="[209,395,1231,1257]" italics="true" pageId="126" pageNumber="127">Andrena arsinoe</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC2743050A88FB058B3202AA" box="[412,1367,1230,1259]" collectingDate="1963-03-27" collectionCode="OOLM" collectorName="R. W. Hodges" country="Egypt" determinerName="K. Warncke" location="Dakhla Oasis" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC2743050A88FB058C6102A9" bold="true" box="[412,516,1230,1256]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050A88FB058F9A02A9" box="[412,511,1230,1256]" name="Egypt" pageId="126" pageNumber="127">EGYPT</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC274305091FFB048CC102A8" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC274305091FFB048CC102A8" box="[523,676,1231,1257]" country="Egypt" name="Dakhla Oasis" pageId="126" pageNumber="127">Dakhla Oasis</location>
,
<date id="9D3C102AAC27430509A4FB048D4E02A8" box="[688,811,1231,1257]" pageId="126" pageNumber="127" value="1963-03-27">
<collectingDate id="8D78E9C2AC27430509A4FB048D4E02A8" box="[688,811,1231,1257]" pageId="126" pageNumber="127" value="1963-03-27">27.iii.1963</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050823FB048D3F02A8" box="[823,858,1231,1257]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, det.
<determinerName id="1CE1DA2EAC274305088DFB048A4402A8" box="[921,1057,1231,1257]" pageId="126" pageNumber="127">K. Warncke</determinerName>
, leg.
<collectorName id="4477533CAC2743050F49FB048A9002A8" box="[1117,1269,1231,1257]" pageId="126" pageNumber="127">R.W. Hodges</collectorName>
,
<collectionCode id="8F93AE2FAC2743050E15FB1B8B3202AA" box="[1281,1367,1232,1259]" pageId="126" pageNumber="127">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050E76FB048D14034E" collectingDate="1932-02-12" collectionCode="OOLM" collectorName="H. Priesner" country="Egypt" determinerName="K. Warncke" location="Wadi Digla" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1">
<location id="EC5D6031AC2743050E76FB048EBD034C" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050E76FB048EBD034C" country="Egypt" name="Wadi Digla" pageId="126" pageNumber="127">Wadi Digla</location>
,
<date id="9D3C102AAC2743050BF0FB388F3D034C" box="[228,344,1267,1293]" pageId="126" pageNumber="127" value="1932-02-12">
<collectingDate id="8D78E9C2AC2743050BF0FB388F3D034C" box="[228,344,1267,1293]" pageId="126" pageNumber="127" value="1932-02-12">12.ii.1932</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050A71FB388FEC034C" box="[357,393,1267,1293]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
, det.
<determinerName id="1CE1DA2EAC2743050ADCFB388C34034C" box="[456,593,1267,1293]" pageId="126" pageNumber="127">K. Warncke</determinerName>
, leg.
<collectorName id="4477533CAC274305099AFB388D6A034C" box="[654,783,1267,1293]" pageId="126" pageNumber="127">H. Priesner</collectorName>
,
<collectionCode id="8F93AE2FAC274305080FFB3F8D14034E" box="[795,881,1268,1295]" pageId="126" pageNumber="127">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050869FB388E8E0370" collectingDate="1909-04-23" collectionCode="ZMHB" country="Egypt" determinerName="J. D. Alfken" location="Wadi Hof" pageId="126" pageNumber="127" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<location id="EC5D6031AC2743050869FB388D88034C" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050869FB388D88034C" box="[893,1005,1267,1293]" country="Egypt" name="Wadi Hof" pageId="126" pageNumber="127">Wadi Hof</location>
,
<date id="9D3C102AAC27430508EEFB388A17034C" box="[1018,1138,1267,1293]" pageId="126" pageNumber="127" value="1909-04-23">
<collectingDate id="8D78E9C2AC27430508EEFB388A17034C" box="[1018,1138,1267,1293]" pageId="126" pageNumber="127" value="1909-04-23">23.iv.1909</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050F6BFB388AC1034C" box="[1151,1188,1267,1293]" count="2" pageId="126" pageNumber="127" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC2743050FA5FB388AB6034C" box="[1201,1235,1267,1293]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, det.
<determinerName id="1CE1DA2EAC2743050E00FB388BFD034C" box="[1300,1432,1267,1293]" pageId="126" pageNumber="127">J.D. Alfken</determinerName>
,
<collectionCode id="8F93AE2FAC2743050B83FADC8E8E0370" box="[151,235,1303,1329]" pageId="126" pageNumber="127">ZMHB</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050BE3FADD8A5D0370" box="[247,1080,1302,1329]" collectingDate="1986-03-06" collectingDateMax="1986-03-19" collectingDateMin="1986-03-06" collectionCode="NHMUK" collectorName="K. M. Guichard" country="Jordan" location="Aqaba" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1" stateProvince="Al 'Aqabah">
<emphasis id="DBF6EAF8AC2743050BE3FADD8F100371" bold="true" box="[247,373,1302,1328]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050BE3FADD8F150371" box="[247,368,1302,1328]" name="Jordan" pageId="126" pageNumber="127">JORDAN</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC2743050A6FFADC8FA10370" box="[379,452,1303,1329]" country="Jordan" name="Al 'Aqabah" pageId="126" pageNumber="127">Aqaba</collectingRegion>
(s.l.),
<date id="9D3C102AAC2743050919FADC8CC00370" box="[525,677,1303,1329]" pageId="126" pageNumber="127" value="1986-03-06" valueMax="1986-03-19" valueMin="1986-03-06">
<collectingDate id="8D78E9C2AC2743050919FADC8CC00370" box="[525,677,1303,1329]" pageId="126" pageNumber="127" value="1986-03-06" valueMax="1986-03-19" valueMin="1986-03-06">619.iii.1986</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC27430509A4FADC8CB60370" box="[688,723,1303,1329]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050806FADC8DA50370" box="[786,960,1303,1329]" pageId="126" pageNumber="127">K.M. Guichard</collectorName>
,
<collectionCode id="8F93AE2FAC27430508DFFADC8A5D0370" box="[971,1080,1303,1329]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050F50FADD8C600314" collectingDate="1983-04-08" collectionCode="NHMUK" collectorName="I. L. Hamer" country="Oman" location="Hawrah" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC2743050F50FADD8AC30371" bold="true" box="[1092,1190,1302,1328]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050F50FADD8AC40371" box="[1092,1185,1302,1328]" name="Oman" pageId="126" pageNumber="127">OMAN</collectingCountry>
:
</emphasis>
J.
<location id="EC5D6031AC2743050FD2FADC8B440370" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050FD2FADC8B440370" box="[1222,1313,1303,1329]" country="Oman" name="Hawrah" pageId="126" pageNumber="127">Hawrah</location>
,
<date id="9D3C102AAC2743050E39FADC8BF20370" box="[1325,1431,1303,1329]" pageId="126" pageNumber="127" value="1983-04-08">
<collectingDate id="8D78E9C2AC2743050E39FADC8BF20370" box="[1325,1431,1303,1329]" pageId="126" pageNumber="127" value="1983-04-08">8.iv.1983</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050B83FAF08EDF0314" box="[151,186,1339,1365]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050A10FAF08FE20314" box="[260,391,1339,1365]" pageId="126" pageNumber="127">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC2743050A8CFAF08C600314" box="[408,517,1339,1365]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050903FAF08F610338" collectingDate="1987-01-16" collectionCode="NHMUK" collectorName="I. L. Hamer" country="United Arab Emirates" location="Abu Dhabi" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1" stateProvince="Abu Zaby">
<emphasis id="DBF6EAF8AC2743050903FAF08DED0315" bold="true" box="[535,904,1338,1365]" pageId="126" pageNumber="127">
<collectingCountry id="9195767AAC2743050903FAF08DE60315" box="[535,899,1338,1365]" name="United Arab Emirates" pageId="126" pageNumber="127">UNITED ARAB EMIRATES</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC2743050881FAF08A7C0314" box="[917,1049,1339,1365]" country="United Arab Emirates" name="Abu Zaby" pageId="126" pageNumber="127">Abu Dhabi</collectingRegion>
,
<date id="9D3C102AAC2743050F3EFAF08AF30314" box="[1066,1174,1339,1365]" pageId="126" pageNumber="127" value="1987-01-16">
<collectingDate id="8D78E9C2AC2743050F3EFAF08AF30314" box="[1066,1174,1339,1365]" pageId="126" pageNumber="127" value="1987-01-16">16.i.1987</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050FBCFAF08AAE0314" box="[1192,1227,1339,1365]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050E01FAF08BFD0314" box="[1301,1432,1339,1365]" pageId="126" pageNumber="127">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC2743050B83FA948F610338" box="[151,260,1375,1401]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050A1BFA948AF20338" box="[271,1175,1374,1401]" collectingDate="2015-01-15" collectionCode="CSE" collectorName="Saji &amp; v. Harten" country="United Arab Emirates" location="Al Wathba" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1" stateProvince="Abu Zaby">
<collectingRegion id="2B46F808AC2743050A1BFA948FE80338" box="[271,397,1375,1401]" country="United Arab Emirates" name="Abu Zaby" pageId="126" pageNumber="127">Abu Dhabi</collectingRegion>
,
<location id="EC5D6031AC2743050A8CFA948C760338" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050A8CFA948C760338" box="[408,531,1375,1401]" country="United Arab Emirates" name="Al Wathba" pageId="126" pageNumber="127" stateProvince="Abu Zaby">Al Wathba</location>
,
<collectingMethod id="30C34EFDAC274305090AFA948CD00338" box="[542,693,1375,1401]" pageId="126" pageNumber="127">Malaise Trap</collectingMethod>
,
<date id="9D3C102AAC27430509D4FA958D490338" box="[704,812,1374,1401]" pageId="126" pageNumber="127" value="2015-01-15">
<collectingDate id="8D78E9C2AC27430509D4FA958D490338" box="[704,812,1374,1401]" pageId="126" pageNumber="127" value="2015-01-15">15.i.2015</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC274305082CFA948D3E0338" box="[824,859,1375,1401]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC274305088EFA948DA30338" box="[922,966,1375,1401]" pageId="126" pageNumber="127">Saji</collectorName>
&amp;
<collectorName id="4477533CAC27430508FEFA948A310338" box="[1002,1108,1375,1401]" pageId="126" pageNumber="127">v. Harten</collectorName>
,
<collectionCode id="8F93AE2FAC2743050F74FA948AF20338" box="[1120,1175,1375,1401]" pageId="126" pageNumber="127">CSE</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050FB6FA948FA703DC" collectingDate="1984-03-10" collectionCode="NHMUK" collectorName="I. L. Hamer" country="United Arab Emirates" location="Liwa" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<location id="EC5D6031AC2743050FB6FA948ABB0338" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050FB6FA948ABB0338" box="[1186,1246,1375,1401]" country="United Arab Emirates" name="Liwa" pageId="126" pageNumber="127">Liwa</location>
,
<date id="9D3C102AAC2743050FFEFA948B020338" box="[1258,1383,1375,1401]" pageId="126" pageNumber="127" value="1984-03-10">
<collectingDate id="8D78E9C2AC2743050FFEFA948B020338" box="[1258,1383,1375,1401]" pageId="126" pageNumber="127" value="1984-03-10">10.iii.1984</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050E66FA948BF00338" box="[1394,1429,1375,1401]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050BDDFA488F2D03DC" box="[201,328,1411,1437]" pageId="126" pageNumber="127">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC2743050A41FA488FA703DC" box="[341,450,1411,1437]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050ADBFA488B3303DC" box="[463,1366,1411,1437]" collectingDate="2011-01-04" collectingDateMax="2011-01-13" collectingDateMin="2011-01-04" collectionCode="CSE" collectorName="C. Schmid-Egger" country="United Arab Emirates" location="Liwa" pageId="126" pageNumber="127" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<location id="EC5D6031AC2743050ADBFA488C6C03DC" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050ADBFA488C6C03DC" box="[463,521,1411,1437]" country="United Arab Emirates" name="Liwa" pageId="126" pageNumber="127">Liwa</location>
desert,
<locationDeviation id="F88F2FC8AC2743050970FA488D7603DC" box="[612,787,1411,1437]" location="Arada" pageId="126" pageNumber="127">
<quantity id="2E7A9B0FAC2743050970FA488CC803DC" box="[612,685,1411,1437]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="126" pageNumber="127" unit="km" value="12.0">12 km</quantity>
S Arada
</locationDeviation>
,
<date id="9D3C102AAC2743050834FA488DC203DC" box="[800,935,1411,1437]" pageId="126" pageNumber="127" value="2011-01-04" valueMax="2011-01-13" valueMin="2011-01-04">
<collectingDate id="8D78E9C2AC2743050834FA488DC203DC" box="[800,935,1411,1437]" pageId="126" pageNumber="127" value="2011-01-04" valueMax="2011-01-13" valueMin="2011-01-04">413.i.2011</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC27430508A1FA488DBC03DC" box="[949,985,1411,1437]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC27430508F3FA488A6F03DC" box="[999,1034,1411,1437]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050F58FA488B7703DC" box="[1100,1298,1411,1437]" pageId="126" pageNumber="127">C. Schmid-Egger</collectorName>
,
<collectionCode id="8F93AE2FAC2743050E0BFA488B3303DC" box="[1311,1366,1411,1437]" pageId="126" pageNumber="127">CSE</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050E77FA488DEF0380" collectingDate="2011-01-09" collectionCode="CSE" collectorName="C. Schmid-Egger" country="United Arab Emirates" location="Liwa Oasis" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1">
<location id="EC5D6031AC2743050E77FA488EBD0380" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050E77FA488EBD0380" country="United Arab Emirates" name="Liwa Oasis" pageId="126" pageNumber="127">Liwa Oasis</location>
,
<locationDeviation id="F88F2FC8AC2743050BF0FA6D8FCF0380" box="[228,426,1446,1473]" location="Mazairaa" pageId="126" pageNumber="127">
<quantity id="2E7A9B0FAC2743050BF0FA6D8F7B0380" box="[228,286,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="126" pageNumber="127" unit="km" value="5.0">5 km</quantity>
E Mazairaa
</locationDeviation>
,
<date id="9D3C102AAC2743050AA2FA6C8C760380" box="[438,531,1447,1473]" pageId="126" pageNumber="127" value="2011-01-09">
<collectingDate id="8D78E9C2AC2743050AA2FA6C8C760380" box="[438,531,1447,1473]" pageId="126" pageNumber="127" value="2011-01-09">9.i.2011</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC274305090BFA6C8C210380" box="[543,580,1447,1473]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC2743050996FA6C8D2D0380" box="[642,840,1447,1473]" pageId="126" pageNumber="127">C. Schmid-Egger</collectorName>
,
<collectionCode id="8F93AE2FAC2743050847FA6C8DEF0380" box="[851,906,1447,1473]" pageId="126" pageNumber="127">CSE</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050882FA6C8F1903A4" collectingDate="2011-01-13" collectionCode="CSE" collectorName="C. Schmid-Egger" country="United Arab Emirates" location="Arada" municipality="Liwa Oasis" pageId="126" pageNumber="127" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingMunicipality id="0959AC90AC2743050882FA6C8A7D0380" box="[918,1048,1447,1473]" pageId="126" pageNumber="127">Liwa Oasis</collectingMunicipality>
,
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,
<date id="9D3C102AAC2743050F62FA6C8A840380" box="[1142,1249,1447,1473]" pageId="126" pageNumber="127" value="2011-01-13">
<collectingDate id="8D78E9C2AC2743050F62FA6C8A840380" box="[1142,1249,1447,1473]" pageId="126" pageNumber="127" value="2011-01-13">13.i.2011</collectingDate>
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,
<specimenCount id="FF84FD63AC2743050E34FA6C8B260380" box="[1312,1347,1447,1473]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050E97FA6C8F5E03A4" pageId="126" pageNumber="127">C. Schmid-Egger</collectorName>
,
<collectionCode id="8F93AE2FAC2743050A51FA008F1903A4" box="[325,380,1483,1509]" pageId="126" pageNumber="127">CSE</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC2743050A92FA008A2203A6" box="[390,1095,1483,1511]" collectingDate="2017-03-12" collectionCode="OOLM" collectorName="Ma. Halada" country="United Arab Emirates" location="Mizerah" municipality="Liwa Oasis" pageId="126" pageNumber="127" specimenCount="1" specimenCount-male="1">
<collectingMunicipality id="0959AC90AC2743050A92FA008C6203A4" box="[390,519,1483,1509]" pageId="126" pageNumber="127">Liwa Oasis</collectingMunicipality>
,
<location id="EC5D6031AC2743050906FA008C1703A4" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050906FA008C1703A4" box="[530,626,1483,1509]" country="United Arab Emirates" municipality="Liwa Oasis" name="Mizerah" pageId="126" pageNumber="127">Mizerah</location>
,
<date id="9D3C102AAC2743050968FA008C9D03A4" box="[636,760,1483,1509]" pageId="126" pageNumber="127" value="2017-03-12">
<collectingDate id="8D78E9C2AC2743050968FA008C9D03A4" box="[636,760,1483,1509]" pageId="126" pageNumber="127" value="2017-03-12">12.iii.2017</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050817FA008D4203A4" box="[771,807,1483,1509]" count="1" pageId="126" pageNumber="127" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC2743050876FA008D8203A4" box="[866,999,1483,1509]" pageId="126" pageNumber="127">Ma. Halada</collectorName>
,
<collectionCode id="8F93AE2FAC27430508E5FA078A2203A6" box="[1009,1095,1484,1511]" pageId="126" pageNumber="127">OÖLM</collectionCode>
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;
<materialsCitation id="59EA3CB7AC2743050F46FA008C080048" collectingDate="1987-02-12" collectionCode="NHMUK" collectorName="I. L. Hamer" country="United Arab Emirates" location="Suweihan" municipality="Shweib" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="0959AC90AC2743050F46FA008AC003A4" box="[1106,1189,1483,1509]" pageId="126" pageNumber="127">Shweib</collectingMunicipality>
/
<location id="EC5D6031AC2743050FBAFA008B7903A4" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC2743050FBAFA008B7903A4" box="[1198,1308,1483,1509]" country="United Arab Emirates" municipality="Shweib" name="Suweihan" pageId="126" pageNumber="127">Suweihan</location>
[Sweihan],
<date id="9D3C102AAC2743050B83FA248F6E0048" box="[151,267,1519,1545]" pageId="126" pageNumber="127" value="1987-02-12">
<collectingDate id="8D78E9C2AC2743050B83FA248F6E0048" box="[151,267,1519,1545]" pageId="126" pageNumber="127" value="1987-02-12">12.ii.1987</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2743050A03FA248F5F0048" box="[279,314,1519,1545]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050A6CFA248F900048" box="[376,501,1519,1545]" pageId="126" pageNumber="127">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC2743050914FA248C080048" box="[512,621,1519,1545]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC274305096DFA248B5A0048" box="[633,1343,1519,1545]" collectingDate="1986-01-24" collectionCode="NHMUK" collectorName="I. L. Hamer" country="United Arab Emirates" location="Suweihan" pageId="126" pageNumber="127" specimenCount="1" specimenCount-female="1">
<location id="EC5D6031AC274305096DFA248C820048" LSID="urn:lsid:plazi:treatment:612B87FCAC2243050B83F9698B270048:EC5D6031AC274305096DFA248C820048" box="[633,743,1519,1545]" country="United Arab Emirates" name="Suweihan" pageId="126" pageNumber="127">Suweihan</location>
[Sweihan],
<date id="9D3C102AAC274305087BFA248DB90048" box="[879,988,1519,1545]" pageId="126" pageNumber="127" value="1986-01-24">
<collectingDate id="8D78E9C2AC274305087BFA248DB90048" box="[879,988,1519,1545]" pageId="126" pageNumber="127" value="1986-01-24">24.i.1986</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC27430508F3FA248A6F0048" box="[999,1034,1519,1545]" count="1" pageId="126" pageNumber="127" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC2743050F5CFA248AA30048" box="[1096,1222,1519,1545]" pageId="126" pageNumber="127">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC2743050FC5FA248B5A0048" box="[1233,1343,1519,1545]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="126" pageNumber="127" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,374 @@
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<emphasis id="DBF6EAF8AC2C430E0B83FF5C8CD006F0" bold="true" box="[151,693,151,178]" pageId="117" pageNumber="118">
<taxonomicName id="2E824D69AC2C430E0B83FF5C8C5306F0" authorityName="WOOD" authorityYear="2024" box="[151,566,151,178]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perpetua" status="sp. nov." subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2C430E0B83FF5C8F6406F3" bold="true" box="[151,257,151,178]" italics="true" pageId="117" pageNumber="118">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2C430E0A06FF5C8FDF06F3" bold="true" box="[274,442,151,178]" italics="true" pageId="117" pageNumber="118">Taeniandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2C430E0AD8FF5C8C5306F0" bold="true" box="[460,566,151,177]" italics="true" pageId="117" pageNumber="118">perpetua</emphasis>
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</emphasis>
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</paragraph>
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<subSubSection id="A1986561AC2C430E0B83FF708F200694" box="[151,325,187,213]" pageId="117" pageNumber="118" type="description">
<paragraph id="E93D36EAAC2C430E0B83FF708F200694" blockId="117.[151,693,151,213]" box="[151,325,187,213]" pageId="117" pageNumber="118">
<figureCitation id="71B92A6FAC2C430E0B83FF708F240694" box="[151,321,187,213]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figures 73AF</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2C430E0B83FEC88C8D0700" pageId="117" pageNumber="118" type="materials_examined">
<paragraph id="E93D36EAAC2C430E0B83FEC88C8D0700" blockId="117.[151,1437,259,2014]" pageId="117" pageNumber="118">
<materialsCitation id="59EA3CB7AC2C430E0B83FEC88C8D0700" collectingDate="2019-05-13" country="Iran" elevation="1741" location="Bam" pageId="117" pageNumber="118" specimenCount="1" specimenCount-male="1" stateProvince="Kerman" typeStatus="holotype">
<emphasis id="DBF6EAF8AC2C430E0B83FEC88FF2075C" bold="true" box="[151,407,259,285]" pageId="117" pageNumber="118">
<typeStatus id="36398848AC2C430E0B83FEC88F5F075C" box="[151,314,259,285]" pageId="117" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC2C430E0A52FEC88FF7075C" box="[326,402,259,285]" name="Iran" pageId="117" pageNumber="118">IRAN</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC2C430E0A8AFECF8F9E075F" box="[414,507,260,286]" country="Iran" name="Kerman" pageId="117" pageNumber="118">Kerman</collectingRegion>
,
<location id="EC5D6031AC2C430E0912FECF8C5B075F" LSID="urn:lsid:plazi:treatment:612B87FCAC2C430C0B83FF5C8C9407B4:EC5D6031AC2C430E0912FECF8C5B075F" box="[518,574,260,286]" country="Iran" name="Bam" pageId="117" pageNumber="118" stateProvince="Kerman">Bam</location>
,
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NW, 29.3969
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, 57.8503
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,
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<collectingDate id="8D78E9C2AC2C430E0F2EFEC88ACF075C" box="[1082,1194,259,285]" pageId="117" pageNumber="118" value="2019-05-13">13.v.2019</collectingDate>
</date>
,
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, leg. W-H. Liebig, OÖLM (BOLD accession number: WPATW976-23).
</materialsCitation>
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<subSubSection id="A1986561AC2C430E0BD3FE808B4A0314" pageId="117" pageNumber="118" type="description">
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<emphasis id="DBF6EAF8AC2C430E0BD3FE808FA70724" bold="true" box="[199,450,331,357]" pageId="117" pageNumber="118">Description: Female:</emphasis>
Unknown.
</paragraph>
<paragraph id="E93D36EAAC2C430E0BD3FEA48FC004C4" blockId="117.[151,1437,259,2014]" pageId="117" pageNumber="118">
<emphasis id="DBF6EAF8AC2C430E0BD3FEA48F6807C8" bold="true" box="[199,269,367,393]" pageId="117" pageNumber="118">Male:</emphasis>
Body length:
<quantity id="2E7A9B0FAC2C430E0ABFFEA48F8907CB" box="[427,492,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="117" pageNumber="118" unit="mm" value="9.0">9 mm</quantity>
(
<figureCitation id="71B92A6FAC2C430E0AEEFEA48CE507C8" box="[506,640,367,393]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73A</figureCitation>
).
<emphasis id="DBF6EAF8AC2C430E0986FEA48CBD07C8" box="[658,728,367,393]" italics="true" pageId="117" pageNumber="118">Head:</emphasis>
1.2 times wider than long (
<figureCitation id="71B92A6FAC2C430E0F1DFEA48AE807C8" box="[1033,1165,367,393]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73B</figureCitation>
). Clypeus flattened over majority of area, densely but shallowly punctate, punctures separated by 0.51 puncture diameter with exception of narrow impunctate longitudinal mid-line, underlying surface smooth and shining. Process of labrum trapezoidal, 2 times wider than long, anterior margin thickened, broadly emarginate, dorsal surface with weak latitudinal striations. Gena equalling width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Face, gena, vertex, and scape covered with whitish plumose hairs, none equalling length of scape. Antennae dark basally, A413 ventrally lightened by presence of grey scales; A3 0.8 times length of A4, A413 elongate rectangular, clearly longer than wide (
<figureCitation id="71B92A6FAC2C430E0A07FDA08FFD04C4" box="[275,408,619,645]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73C</figureCitation>
).
</paragraph>
<paragraph id="E93D36EAAC2C430E0BD3FD5B8F6A0250" blockId="117.[151,1437,259,2014]" pageId="117" pageNumber="118">
<emphasis id="DBF6EAF8AC2C430E0BD3FD5B8F2D04E8" box="[199,328,656,681]" italics="true" pageId="117" pageNumber="118">Mesosoma:</emphasis>
Scutum densely but somewhat irregularly punctate, punctures touching to separated by 1 puncture diameter, underlying surface with strong granular shagreen, weakly shining (
<figureCitation id="71B92A6FAC2C430E0F0EFD788AC1048C" box="[1050,1188,691,717]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73D</figureCitation>
). Scutellum similarly punctate, sculpture weaker, more clearly shining. Pronotum predominantly rounded, with slight hint of angle along dorsolateral margins. Mesepisternum and dorsolateral parts of propodeum with granular microreticulation, microreticulation overlain by network of fine raised rugosity, surface weakly shining overall; propodeal triangle laterally delineated by fine carinae, internal surface with thick granular microreticulation and irregular network of fine raised rugae, dull, thus additionally defined from remaining parts of propodeum by change in sculpture. Mesepisternum and propodeum with long whitish plumose hairs, none equalling length of scape, hairs becoming shorter and light whitish-brownish on scutum and scutellum. Legs dark, pubescence whitish to light brownish. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange-brown, nervulus strongly postfurcal.
</paragraph>
<paragraph id="E93D36EAAC2C430E0BD3FBD78B4A0314" blockId="117.[151,1437,259,2014]" pageId="117" pageNumber="118">
<emphasis id="DBF6EAF8AC2C430E0BD3FBD78F200274" box="[199,325,1052,1077]" italics="true" pageId="117" pageNumber="118">Metasoma:</emphasis>
Tergal discs dark, marginal areas lightened hyaline-brown (
<figureCitation id="71B92A6FAC2C430E0F07FBD08AF90274" box="[1043,1180,1051,1077]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73E</figureCitation>
). Tergal discs basally shagreened and weakly shining, sculpture becoming weaker apically and onto marginal areas, here more strongly shining; surface punctate, punctures separated by 12 puncture diameters.Tergal discs with scattered short hairs, T15 with dense complete apical hairbands of whitish plumose hairs, obscuring underlying surface. S8 columnar, ventral surface covered with dense short brown hairs. Genital capsule compact, gonocoxae with their inner margins gently diverging apically, forming 90
<superScript id="1EF79BA2AC2C430E0AF8FB048F91029C" attach="left" box="[492,500,1231,1245]" fontSize="6" pageId="117" pageNumber="118">o</superScript>
angle at their apexes (
<figureCitation id="71B92A6FAC2C430E09E4FB048D1702A8" box="[752,882,1231,1257]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73F</figureCitation>
). Gonostyli relatively narrow basally, broadening apically, more or less spatulate, outer corner forming rounded 90
<superScript id="1EF79BA2AC2C430E087AFB388D130340" attach="left" box="[878,886,1267,1281]" fontSize="6" pageId="117" pageNumber="118">o</superScript>
angle; outer margin basally at transition between gonostyli and gonocoxae with small but distinct impression, breaking outer profile. Penis valves basally slightly broadened, occupying ½ space between gonostyli, apically narrowing, with small oval opening medially.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2C430D0BD3FA948A3B0724" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118" type="diagnosis">
<paragraph id="E93D36EAAC2C430E0BD3FA948C2900A0" blockId="117.[151,1437,259,2014]" pageId="117" pageNumber="118">
<emphasis id="DBF6EAF8AC2C430E0BD3FA948F270338" bold="true" box="[199,322,1375,1401]" pageId="117" pageNumber="118">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC2C430E0A5CFA948C770338" authorityName="WOOD" authorityYear="2024" box="[328,530,1375,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2C430E0A5CFA948C770338" box="[328,530,1375,1401]" italics="true" pageId="117" pageNumber="118">Andrena perpetua</emphasis>
</taxonomicName>
can be placed in the subgenus
<taxonomicName id="2E824D69AC2C430E087DFA948A670338" authorityName="Hedicke" authorityYear="1933" box="[873,1026,1375,1401]" pageId="117" pageNumber="118" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2C430E087DFA948A670338" box="[873,1026,1375,1401]" italics="true" pageId="117" pageNumber="118">Taeniandrena</emphasis>
</taxonomicName>
due to the strongly flattened clypeus and typical genital capsule with broad flattened gonostyli and somewhat expanded penis valves. It can be placed into the
<taxonomicName id="2E824D69AC2C430E0BEEFA6C8F2C0380" authorityName="van der Vecht" authorityYear="1927" box="[250,329,1447,1473]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2C430E0BEEFA6C8F2C0380" box="[250,329,1447,1473]" italics="true" pageId="117" pageNumber="118">gelriae</emphasis>
</taxonomicName>
-group of species (the species genetically falling around
<taxonomicName id="2E824D69AC2C430E08CBFA6C8B230380" authority="van der Vecht, 1927" authorityName="van der Vecht" authorityYear="1927" box="[991,1350,1447,1474]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2C430E08CBFA6C8A340380" box="[991,1105,1447,1473]" italics="true" pageId="117" pageNumber="118">A. gelriae</emphasis>
van der Vecht, 1927
</taxonomicName>
and
<taxonomicName id="2E824D69AC2C430E0E90FA6C8FAF03A4" authority="Thomson, 1870" authorityName="Thomson" authorityYear="1870" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="DBF6EAF8AC2C430E0E90FA6C8F7503A4" italics="true" pageId="117" pageNumber="118">A. intermedia</emphasis>
Thomson,
<quantity id="2E7A9B0FAC2C430E0A86FA008FAF03A4" box="[402,458,1483,1509]" metricMagnitude="1" metricUnit="m" metricValue="4.7498" pageId="117" pageNumber="118" unit="in" value="1870.0">1870</quantity>
</taxonomicName>
in the analysis of
<bibRefCitation id="8D134B1BAC2C430E0989FA078DE903A4" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[669,908,1483,1510]" pageId="117" pageNumber="118" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC2C430E09C3FA078D6F03A4" box="[727,778,1483,1509]" italics="true" pageId="117" pageNumber="118">et al</emphasis>
. 2022: 381
</bibRefCitation>
) due to the long A4 that is clearly longer than A3 (
<figureCitation id="71B92A6FAC2C430E0BDCFA248F2C0048" box="[200,329,1519,1545]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73C</figureCitation>
) and the genital capsule with the penis valves slightly broadened basally (
<figureCitation id="71B92A6FAC2C430E0F96FA248B660048" box="[1154,1283,1519,1545]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73F</figureCitation>
). Genetically, based on the COI barcode (652 base pair sequence generated),
<taxonomicName id="2E824D69AC2C430E084AF9D88D87006C" authorityName="WOOD" authorityYear="2024" box="[862,994,1555,1581]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2C430E084AF9D88D87006C" box="[862,994,1555,1581]" italics="true" pageId="117" pageNumber="118">A. perpetua</emphasis>
</taxonomicName>
also falls into this group, and is placed basally within this clade (
<figureCitation id="71B92A6FAC2C430E0AA5F9FC8C440010" box="[433,545,1591,1617]" captionStart="FIGURE 74" captionStartId="119.[151,250,1494,1518]" captionTargetBox="[151,1433,527,1468]" captionTargetId="figure-158@119.[151,1435,525,1470]" captionTargetPageId="119" captionText="FIGURE 74. Phylogenetic tree (maximum likelihood) of Andrena subgenus Taeniandrena Hedicke, 1933 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209469" httpUri="https://zenodo.org/record/13209469/files/figure.png" pageId="117" pageNumber="118">Figure 74</figureCitation>
). The closest sequence (WPATW975-23) was separated by 1.38%, this sequence coming from a probably undescribed member of the
<taxonomicName id="2E824D69AC2C430E09FEF9908D5C0034" authorityName="van der Vecht" authorityYear="1927" box="[746,825,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2C430E09FEF9908D5C0034" box="[746,825,1627,1653]" italics="true" pageId="117" pageNumber="118">gelriae</emphasis>
</taxonomicName>
-group from central
<collectingCountry id="9195767AAC2C430E0F09F9908A050034" box="[1053,1120,1627,1653]" name="Spain" pageId="117" pageNumber="118">Spain</collectingCountry>
. Though not included in the tree since the sequence is short, the one available sequence for the most easterly distributed member of the
<taxonomicName id="2E824D69AC2C430E0E50F9B48BF600D8" authorityName="van der Vecht" authorityYear="1927" box="[1348,1427,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2C430E0E50F9B48BF600D8" box="[1348,1427,1663,1689]" italics="true" pageId="117" pageNumber="118">gelriae</emphasis>
</taxonomicName>
- group (
<taxonomicName id="2E824D69AC2C430E0BFEF9688C4600FC" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[234,547,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="DBF6EAF8AC2C430E0BFEF9688F0B00FC" box="[234,366,1699,1725]" italics="true" pageId="117" pageNumber="118">A. producta</emphasis>
<bibRefCitation id="8D134B1BAC2C430E0A62F9688C4600FC" author="Warncke, K." box="[374,547,1699,1725]" pageId="117" pageNumber="118" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
; HYMAA323-22; 292 base pair sequence) was separated from
<taxonomicName id="2E824D69AC2C430E0FE0F9688B1C00FC" authorityName="WOOD" authorityYear="2024" box="[1268,1401,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2C430E0FE0F9688B1C00FC" box="[1268,1401,1699,1725]" italics="true" pageId="117" pageNumber="118">A. perpetua</emphasis>
</taxonomicName>
by 3.08% over the 44% sequence overlap.
</paragraph>
<paragraph id="E93D36EAAC2C430D0BD3F9208A3B0724" blockId="117.[151,1437,259,2014]" lastBlockId="118.[151,1436,151,357]" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118">
This group is extremely challenging taxonomically, with many cryptic and endemic species in Western Europe (
<bibRefCitation id="8D134B1BAC2C430E0B8BF8C48F320168" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[159,343,1807,1834]" pageId="117" pageNumber="118" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC2C430E0BCEF8DB8F690168" box="[218,268,1807,1833]" italics="true" pageId="117" pageNumber="118">et al</emphasis>
. 2022
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC2C430E0A76F8C48F9D0168" author="Wood, T. J." box="[354,504,1807,1833]" pageId="117" pageNumber="118" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
). These taxa are referred to in
<figureCitation id="71B92A6FAC2C430E0846F8C48DDA0168" box="[850,959,1807,1833]" captionStart="FIGURE 74" captionStartId="119.[151,250,1494,1518]" captionTargetBox="[151,1433,527,1468]" captionTargetId="figure-158@119.[151,1435,525,1470]" captionTargetPageId="119" captionText="FIGURE 74. Phylogenetic tree (maximum likelihood) of Andrena subgenus Taeniandrena Hedicke, 1933 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209469" httpUri="https://zenodo.org/record/13209469/files/figure.png" pageId="117" pageNumber="118">Figure 74</figureCitation>
as the French/Iberian barcoding complex. Members of this group largely (with the exception of
<taxonomicName id="2E824D69AC2C430E0813F8F88A4D010C" authority="Wood, 2022" authorityName="Wood" authorityYear="2022" box="[775,1064,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="contracta">
<emphasis id="DBF6EAF8AC2C430E0813F8F88DF1010C" box="[775,916,1843,1869]" italics="true" pageId="117" pageNumber="118">A. contracta</emphasis>
Wood, 2022
</taxonomicName>
which is restricted to the Sierra Nevada mountain in southern
<collectingCountry id="9195767AAC2C430E0AF2F89C8C420130" box="[486,551,1879,1905]" name="Spain" pageId="117" pageNumber="118">Spain</collectingCountry>
) display a genital capsule close to
<taxonomicName id="2E824D69AC2C430E08B3F89C8A5A0130" authorityName="Thomson" authorityYear="1870" box="[935,1087,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="DBF6EAF8AC2C430E08B3F89C8A5A0130" box="[935,1087,1879,1905]" italics="true" pageId="117" pageNumber="118">A. intermedia</emphasis>
</taxonomicName>
, with strongly broadened penis valves which occupy the majority of the space between the gonostyli. Genetically and morphologically
<taxonomicName id="2E824D69AC2C430E0E0EF8B08BF801D4" authorityName="WOOD" authorityYear="2024" box="[1306,1437,1915,1941]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2C430E0E0EF8B08BF801D4" box="[1306,1437,1915,1941]" italics="true" pageId="117" pageNumber="118">A. perpetua</emphasis>
</taxonomicName>
falls outside of this group as the penis valves are comparatively much narrower, and it can also be recognised and separated from more widespread members of this group due to its genital capsule (
<figureCitation id="71B92A6FAC2C430E0F40F8088ABC019C" box="[1108,1241,1987,2013]" captionStart="FIGURE 73" captionStartId="118.[151,250,1843,1867]" captionTargetBox="[151,1436,381,1819]" captionTargetId="figure-138@118.[151,1436,381,1819]" captionTargetPageId="118" captionText="FIGURE 73. Andrena (Taeniandrena) perpetua spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="117" pageNumber="118">Figure 73F</figureCitation>
). The gonocoxae have their inner margins gently diverging apically (as in
<taxonomicName id="2E824D69AC2F430D0802FF5C8DE006F0" authorityName="van der Vecht" authorityYear="1927" box="[790,901,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2F430D0802FF5C8DE006F0" box="[790,901,151,177]" italics="true" pageId="118" pageNumber="119">A. gelriae</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2F430D08A9FF5C8A2506F0" authorityName="Warncke" authorityYear="1973" box="[957,1088,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="DBF6EAF8AC2F430D08A9FF5C8A2506F0" box="[957,1088,151,177]" italics="true" pageId="118" pageNumber="119">A. producta</emphasis>
</taxonomicName>
), but these margins form a 90
<superScript id="1EF79BA2AC2F430D0E80FF5C8BF906E4" attach="none" box="[1428,1436,151,165]" fontSize="6" pageId="118" pageNumber="119">o</superScript>
angle apically (acute angle in
<taxonomicName id="2E824D69AC2F430D0AF1FF708C360694" authorityName="van der Vecht" authorityYear="1927" box="[485,595,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2F430D0AF1FF708C360694" box="[485,595,187,213]" italics="true" pageId="118" pageNumber="119">A. gelriae</emphasis>
</taxonomicName>
, obtuse angle in
<taxonomicName id="2E824D69AC2F430D0805FF708DF60694" authorityName="Warncke" authorityYear="1973" box="[785,915,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="DBF6EAF8AC2F430D0805FF708DF60694" box="[785,915,187,213]" italics="true" pageId="118" pageNumber="119">A. producta</emphasis>
</taxonomicName>
), the penis valves themselves are only slightly broadened with very narrow valve medially (grossly broadened with a broad valve in
<taxonomicName id="2E824D69AC2F430D0F9FFF148B4206B8" authorityName="Thomson" authorityYear="1870" box="[1163,1319,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="DBF6EAF8AC2F430D0F9FFF148B4206B8" box="[1163,1319,223,249]" italics="true" pageId="118" pageNumber="119">A. intermedia</emphasis>
</taxonomicName>
and most members of the French/Iberian barcoding complex, moderately broadened in
<taxonomicName id="2E824D69AC2F430D08ECFEC88A00075C" authorityName="van der Vecht" authorityYear="1927" box="[1016,1125,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2F430D08ECFEC88A00075C" box="[1016,1125,259,285]" italics="true" pageId="118" pageNumber="119">A. gelriae</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2F430D0F8FFEC88B7B075C" authorityName="Warncke" authorityYear="1973" box="[1179,1310,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="DBF6EAF8AC2F430D0F8FFEC88B7B075C" box="[1179,1310,259,285]" italics="true" pageId="118" pageNumber="119">A. producta</emphasis>
</taxonomicName>
), and there is a distinct kink or impression on the outer face of the gonostyli basally, at the transition between the gonocoxae and the gonostyli (comparison species with the outer margin of the gonostyli straight).
</paragraph>
</subSubSection>
<caption id="BDFD6662AC2F430D0B83F8F88AE7012E" ID-DOI="http://doi.org/10.5281/zenodo.13209467" ID-Zenodo-Dep="13209467" httpUri="https://zenodo.org/record/13209467/files/figure.png" pageId="118" pageNumber="119" startId="118.[151,250,1843,1867]" targetBox="[151,1436,381,1819]" targetPageId="118" targetType="figure">
<paragraph id="E93D36EAAC2F430D0B83F8F88AE7012E" blockId="118.[151,1436,1843,1903]" pageId="118" pageNumber="119">
<emphasis id="DBF6EAF8AC2F430D0B83F8F88F44010A" bold="true" box="[151,289,1843,1867]" pageId="118" pageNumber="119">FIGURE 73.</emphasis>
<taxonomicName id="2E824D69AC2F430D0A3CF8F88CE7010A" authorityName="WOOD" authorityYear="2024" box="[296,642,1843,1867]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="perpetua" status="sp. nov." subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2F430D0A3CF8F88F1B010A" box="[296,382,1843,1867]" italics="true" pageId="118" pageNumber="119">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2F430D0A9AF8F88C7C010A" box="[398,537,1843,1867]" italics="true" pageId="118" pageNumber="119">Taeniandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2F430D093CF8FF8CE7010A" box="[552,642,1844,1867]" italics="true" pageId="118" pageNumber="119">perpetua</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC2F430D099DF8F88C94010A" bold="true" box="[649,753,1843,1867]" pageId="118" pageNumber="119">
<taxonomicNameLabel id="C0C55783AC2F430D099DF8F88C94010A" box="[649,753,1843,1867]" pageId="118" pageNumber="119" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Antennae detail, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC2F430C0BD3F8568D670724" lastPageId="119" lastPageNumber="120" pageId="118" pageNumber="119" type="discussion">
<paragraph id="E93D36EAAC2F430C0BD3F8568D670724" blockId="118.[151,1436,1949,2012]" lastBlockId="119.[151,1436,151,501]" lastPageId="119" lastPageNumber="120" pageId="118" pageNumber="119">
<emphasis id="DBF6EAF8AC2F430D0BD3F8568F5E01F6" bold="true" box="[199,315,1949,1975]" pageId="118" pageNumber="119">Remarks.</emphasis>
Previously, only four species of
<taxonomicName id="2E824D69AC2F430D09B6F8568D5E01F6" authorityName="Hedicke" authorityYear="1933" box="[674,827,1949,1975]" pageId="118" pageNumber="149" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2F430D09B6F8568D5E01F6" box="[674,827,1949,1975]" italics="true" pageId="118" pageNumber="119">Taeniandrena</emphasis>
</taxonomicName>
were confidently known from
<collectingCountry id="9195767AAC2F430D0F9BF8558AD901F9" box="[1167,1212,1950,1976]" name="Iran" pageId="118" pageNumber="119">Iran</collectingCountry>
(
<bibRefCitation id="8D134B1BAC2F430D0FDDF8568BF901F6" author="Wood, T. J. &amp; Monfared, A." box="[1225,1436,1949,1975]" pageId="118" pageNumber="119" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared</bibRefCitation>
2022;
<bibRefCitation id="8D134B1BAC2F430D0BCAF80A8F15019A" author="Wood, T. J." box="[222,368,1985,2011]" pageId="118" pageNumber="119" pagination="1 - 85" refId="ref92440" refString="Wood, T. J. (2023 c) Revisions to the Andrena fauna of north-western Africa with a focus on Morocco (Hymenoptera: Andrenidae). European Journal of Taxonomy, 916, 1 - 85. https: // doi. org / 10.5852 / ejt. 2023.916.2381" type="journal article" year="2023">Wood 2023c</bibRefCitation>
), none of which are found in the
<taxonomicName id="2E824D69AC2F430D09FDF80A8D5D019A" authorityName="van der Vecht" authorityYear="1927" box="[745,824,1985,2011]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2F430D09FDF80A8D5D019A" box="[745,824,1985,2011]" italics="true" pageId="118" pageNumber="119">gelriae</emphasis>
</taxonomicName>
-group.
<taxonomicName id="2E824D69AC2F430D0885F80A8A39019A" authorityName="WOOD" authorityYear="2024" box="[913,1116,1985,2011]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="118" pageNumber="119" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2F430D0885F80A8A39019A" box="[913,1116,1985,2011]" italics="true" pageId="118" pageNumber="119">Andrena perpetua</emphasis>
</taxonomicName>
represents the most easterly member of this group recognised to date (with
<taxonomicName id="2E824D69AC2E430C098FFF5C8D6D06F0" authorityName="van der Vecht" authorityYear="1927" box="[667,776,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2E430C098FFF5C8D6D06F0" box="[667,776,151,177]" italics="true" pageId="119" pageNumber="120">A. gelriae</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC2E430C0828FF5C8DDB06F0" authorityName="Warncke" authorityYear="1973" box="[828,958,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="producta">
<emphasis id="DBF6EAF8AC2E430C0828FF5C8DDB06F0" box="[828,958,151,177]" italics="true" pageId="119" pageNumber="120">A. producta</emphasis>
</taxonomicName>
previously reported from
<collectingCountry id="9195767AAC2E430C0FCBFF5C8B5506F0" box="[1247,1328,151,177]" name="Turkey" pageId="119" pageNumber="120">Turkey</collectingCountry>
,
<bibRefCitation id="8D134B1BAC2E430C0E2DFF5C8EBB0694" author="Warncke, K." pageId="119" pageNumber="120" pagination="81 - 116" refId="ref91798" refString="Warncke, K. (1975 a) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil A. Mitteilungen der Munchner Entomologischen Gesellschaft, 64, 81 - 116." type="journal article" year="1975">Warncke 1975a</bibRefCitation>
, although it is unclear if this is true
<taxonomicName id="2E824D69AC2E430C096AFF708C880694" authorityName="van der Vecht" authorityYear="1927" box="[638,749,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2E430C096AFF708C880694" box="[638,749,187,213]" italics="true" pageId="119" pageNumber="120">A. gelriae</emphasis>
</taxonomicName>
based on recent revisionary worth within this species group which has substantially altered our species concepts; e.g.
<bibRefCitation id="8D134B1BAC2E430C0809FF148D8406B8" author="Wood, T. J. &amp; Ghisbain, G. &amp; Michez, D. &amp; Praz, C." box="[797,993,223,249]" pageId="119" pageNumber="120" pagination="147 - 193" refId="ref92060" refString="Wood, T. J., Ghisbain, G., Michez, D. &amp; Praz, C. (2021) Revisions to the faunas of Andrena of the Iberian Peninsula and Morocco with the descriptions of four new species (Hymenoptera: Andrenidae). European Journal of Taxonomy, 758, 147 - 193. https: // doi. org / 10.5852 / ejt. 2021.758.1431" type="journal article" year="2021">
Wood
<emphasis id="DBF6EAF8AC2E430C0873FF148DFD06B8" box="[871,920,223,249]" italics="true" pageId="119" pageNumber="120">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC2E430C08F9FF148AC606B8" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[1005,1187,223,249]" pageId="119" pageNumber="120" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC2E430C0F33FF148A3D06B8" box="[1063,1112,223,249]" italics="true" pageId="119" pageNumber="120">et al</emphasis>
. 2022
</bibRefCitation>
). Based on patterns of endemism in
<collectingCountry id="9195767AAC2E430C0A3BFEC88F15075C" box="[303,368,259,285]" name="Georgia" pageId="119" pageNumber="120">Iberia</collectingCountry>
(
<bibRefCitation id="8D134B1BAC2E430C0A94FEC88C52075C" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[384,567,259,285]" pageId="119" pageNumber="120" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC2E430C0AAFFEC88F89075C" box="[443,492,259,285]" italics="true" pageId="119" pageNumber="120">et al</emphasis>
. 2022
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC2E430C0956FEC88CB3075C" author="Wood, T. J." box="[578,726,259,285]" pageId="119" pageNumber="120" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
), it is possible that
<taxonomicName id="2E824D69AC2E430C08A4FEC88A56075D" authorityName="WOOD" authorityYear="2024" box="[944,1075,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="perpetua">
<emphasis id="DBF6EAF8AC2E430C08A4FEC88A56075D" box="[944,1075,259,285]" italics="true" pageId="119" pageNumber="120">A. perpetua</emphasis>
</taxonomicName>
is narrowly endemic to specific mountain ranges in Central
<collectingCountry id="9195767AAC2E430C0AC8FEEC8C680700" box="[476,525,295,321]" name="Iran" pageId="119" pageNumber="120">Iran</collectingCountry>
. Additional sampling and genetic analysis of Iranian
<taxonomicName id="2E824D69AC2E430C0F6FFEEC8B710700" authorityName="Hedicke" authorityYear="1933" box="[1147,1300,295,321]" pageId="119" pageNumber="149" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2E430C0F6FFEEC8B710700" box="[1147,1300,295,321]" italics="true" pageId="119" pageNumber="120">Taeniandrena</emphasis>
</taxonomicName>
is likely to uncover additional undescribed members of this group.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2E430C0BD3FEA48C420790" pageId="119" pageNumber="120" type="etymology">
<paragraph id="E93D36EAAC2E430C0BD3FEA48C420790" blockId="119.[151,1436,151,501]" pageId="119" pageNumber="120">
<emphasis id="DBF6EAF8AC2E430C0BD3FEA48F2B07C8" bold="true" box="[199,334,367,393]" pageId="119" pageNumber="120">Etymology.</emphasis>
Nominative feminine singular form of the Latin adjective
<emphasis id="DBF6EAF8AC2E430C08D9FEA48A5E07C9" box="[973,1083,367,392]" italics="true" pageId="119" pageNumber="120">perpetuus</emphasis>
meaning perpetual, everlasting, uninterrupted, in reference to the seemingly never-ending number of cryptic
<taxonomicName id="2E824D69AC2E430C0F01FE588ACB07EC" authorityName="Hedicke" authorityYear="1933" box="[1045,1198,403,429]" pageId="119" pageNumber="149" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC2E430C0F01FE588ACB07EC" box="[1045,1198,403,429]" italics="true" pageId="119" pageNumber="120">Taeniandrena</emphasis>
</taxonomicName>
species, particularly within the
<taxonomicName id="2E824D69AC2E430C0A1BFE7C8F3B0790" authorityName="van der Vecht" authorityYear="1927" box="[271,350,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="gelriae">
<emphasis id="DBF6EAF8AC2E430C0A1BFE7C8F3B0790" box="[271,350,439,465]" italics="true" pageId="119" pageNumber="120">gelriae</emphasis>
</taxonomicName>
-group of species.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2E430C0BD3FE118C9407B4" box="[199,753,474,501]" pageId="119" pageNumber="120" type="distribution">
<paragraph id="E93D36EAAC2E430C0BD3FE118C9407B4" blockId="119.[151,1436,151,501]" box="[199,753,474,501]" pageId="119" pageNumber="120">
<emphasis id="DBF6EAF8AC2E430C0BD3FE118F0407B5" bold="true" box="[199,353,474,500]" pageId="119" pageNumber="120">Distribution.</emphasis>
Central
<collectingCountry id="9195767AAC2E430C0AD5FE108F8B07B4" box="[449,494,475,501]" name="Iran" pageId="119" pageNumber="120">Iran</collectingCountry>
(province of
<collectingRegion id="2B46F808AC2E430C0992FE108C8007B4" box="[646,741,475,501]" country="Iran" name="Kerman" pageId="119" pageNumber="120">Kerman</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,430 @@
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<subSubSection id="A1986561AC2E430C0B83F9AF8CF1003F" box="[151,660,1636,1663]" pageId="119" pageNumber="120" type="nomenclature">
<paragraph id="E93D36EAAC2E430C0B83F9AF8CF1003F" blockId="119.[151,660,1636,1699]" box="[151,660,1636,1663]" pageId="119" pageNumber="120">
<heading id="B2758186AC2E430C0B83F9AF8CF1003F" bold="true" box="[151,660,1636,1663]" fontSize="11" level="2" pageId="119" pageNumber="120" reason="4">
<emphasis id="DBF6EAF8AC2E430C0B83F9AF8CF1003F" bold="true" box="[151,660,1636,1663]" pageId="119" pageNumber="120">
<taxonomicName id="2E824D69AC2E430C0B83F9AF8C70003E" authorityName="WOOD" authorityYear="2024" box="[151,533,1636,1663]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="119" pageNumber="120" phylum="Arthropoda" rank="species" species="namaka" status="sp. nov." subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2E430C0B83F9AF8F64003E" bold="true" box="[151,257,1636,1663]" italics="true" pageId="119" pageNumber="120">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2E430C0A06F9AF8FC7003E" bold="true" box="[274,418,1636,1663]" italics="true" pageId="119" pageNumber="120">Troandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2E430C0AA7F9AF8C70003E" bold="true" box="[435,533,1636,1663]" italics="true" pageId="119" pageNumber="120">namaka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC2E430C0908F9AF8CF1003F" box="[540,660,1636,1662]" pageId="119" pageNumber="120" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2E430C0B83F9438F8B00E3" box="[151,494,1672,1699]" pageId="119" pageNumber="120" type="description">
<paragraph id="E93D36EAAC2E430C0B83F9438F8B00E3" blockId="119.[151,660,1636,1699]" box="[151,494,1672,1699]" pageId="119" pageNumber="120">
<figureCitation id="71B92A6FAC2E430C0B83F9438F2D00E2" box="[151,328,1672,1699]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="119" pageNumber="120">Figures 75AH</figureCitation>
;
<figureCitation id="71B92A6FAC2E430C0A47F9438F8F00E3" box="[339,490,1672,1698]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="119" pageNumber="120">76AC; EG</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2E430C0B83F91B8FAB0118" pageId="119" pageNumber="120" type="materials_examined">
<paragraph id="E93D36EAAC2E430C0B83F91B8DE30150" blockId="119.[151,1437,1744,2023]" pageId="119" pageNumber="120">
<materialsCitation id="59EA3CB7AC2E430C0B83F91B8DE30150" collectingDate="1956-02-10" collectorName="Ch. Lindemann &amp; Statsslg. Munchen" country="Pakistan" county="Kanatti-Kak" location="West Pakistan" municipality="Salt Range" pageId="119" pageNumber="120" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC2E430C0B83F91B8FBB00AB" bold="true" box="[151,478,1744,1770]" pageId="119" pageNumber="120">
<typeStatus id="36398848AC2E430C0B83F91B8F5F00AB" box="[151,314,1744,1770]" pageId="119" pageNumber="120" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC2E430C0A5CF91B8FBC00AB" box="[328,473,1744,1770]" name="Pakistan" pageId="119" pageNumber="120">PAKISTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC2E430C0AF3F91A8CEC00AB" LSID="urn:lsid:plazi:treatment:612B87FCAC2E43000B83F9AF8D3B0011:EC5D6031AC2E430C0AF3F91A8CEC00AB" box="[487,649,1744,1771]" country="Pakistan" county="Kanatti-Kak" municipality="Salt Range" name="West Pakistan" pageId="119" pageNumber="120">West Pakistan</location>
,
<collectingMunicipality id="0959AC90AC2E430C0982F91B8D7100AA" box="[662,788,1744,1771]" pageId="119" pageNumber="120">Salt Range</collectingMunicipality>
,
<collectingCounty id="005C4E66AC2E430C0836F91B8DC900AB" box="[802,940,1744,1770]" pageId="119" pageNumber="120">Kanatti-Kak</collectingCounty>
[inferred Kenhatti, 32.6826
<superScript id="1EF79BA2AC2E430C0FFFF91B8B6F00AA" attach="right" box="[1259,1290,1744,1771]" fontSize="6" pageId="119" pageNumber="120">oN</superScript>
, 72.2515
<superScript id="1EF79BA2AC2E430C0E67F91B8BF500AA" attach="right" box="[1395,1424,1744,1771]" fontSize="6" pageId="119" pageNumber="120">oE</superScript>
],
<date id="9D3C102AAC2E430C0B83F93F8F6E014E" box="[151,267,1780,1807]" pageId="119" pageNumber="120" value="1956-02-10">
<collectingDate id="8D78E9C2AC2E430C0B83F93F8F6E014E" box="[151,267,1780,1807]" pageId="119" pageNumber="120" value="1956-02-10">10.ii.1956</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2E430C0A03F93E8F49014E" box="[279,300,1781,1807]" count="1" pageId="119" pageNumber="120" type="female"></specimenCount>
, leg.
<collectorName id="4477533CAC2E430C0A7EF93F8C7E014F" box="[362,539,1780,1806]" pageId="119" pageNumber="120">Ch. Lindemann</collectorName>
, ex.
<collectorName id="4477533CAC2E430C095BF93F8D44014F" box="[591,801,1780,1806]" pageId="119" pageNumber="120">Statsslg. München</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC2E430C0BD3F8D38FAB0118" blockId="119.[151,1437,1744,2023]" pageId="119" pageNumber="120">
<materialsCitation id="59EA3CB7AC2E430C0BD3F8D38FAF0118" collectingDate="1956-02-14" collectorName="Ch. Lindemann &amp; Statsslg. Munchen" country="Pakistan" location="Salt Range" municipality="Kanatti-Kak" pageId="119" pageNumber="120" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="DBF6EAF8AC2E430C0BD3F8D38F000173" bold="true" box="[199,357,1816,1842]" pageId="119" pageNumber="120">
<typeStatus id="36398848AC2E430C0BD3F8D38F050173" box="[199,352,1816,1842]" pageId="119" pageNumber="120" type="paratype">PARATYPE</typeStatus>
:
</emphasis>
<collectingCountry id="9195767AAC2E430C0A7BF8D28C760173" box="[367,531,1816,1843]" name="Pakistan" pageId="119" pageNumber="120">West Pakistan</collectingCountry>
,
<location id="EC5D6031AC2E430C0936F8D38CC70172" LSID="urn:lsid:plazi:treatment:612B87FCAC2E43000B83F9AF8D3B0011:EC5D6031AC2E430C0936F8D38CC70172" box="[546,674,1816,1843]" country="Pakistan" municipality="Kanatti-Kak" name="Salt Range" pageId="119" pageNumber="120">Salt Range</location>
,
<collectingMunicipality id="0959AC90AC2E430C09A6F8D38D590173" box="[690,828,1816,1842]" pageId="119" pageNumber="120">Kanatti-Kak</collectingMunicipality>
[Kenhatti],
<date id="9D3C102AAC2E430C08DFF8D38A5A0172" box="[971,1087,1816,1843]" pageId="119" pageNumber="120" value="1956-02-14">
<collectingDate id="8D78E9C2AC2E430C08DFF8D38A5A0172" box="[971,1087,1816,1843]" pageId="119" pageNumber="120" value="1956-02-14">14.ii.1956</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC2E430C0F5BF8D38A160173" box="[1103,1139,1816,1842]" count="1" pageId="119" pageNumber="120" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC2E430C0FA3F8D38B090173" box="[1207,1388,1816,1842]" pageId="119" pageNumber="120">Ch. Lindemann</collectorName>
, ex.
<collectorName id="4477533CAC2E430C0B83F8F78F0C0117" box="[151,361,1852,1878]" pageId="119" pageNumber="120">Statsslg. München</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2E43030BD3F8AB8C0601B2" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120" type="description">
<paragraph id="E93D36EAAC2E43030BD3F8AB8CEC0725" blockId="119.[151,1437,1744,2023]" lastBlockId="120.[151,1437,150,2035]" lastPageId="120" lastPageNumber="121" pageId="119" pageNumber="120">
<emphasis id="DBF6EAF8AC2E430C0BD3F8AB8FA7013B" bold="true" box="[199,450,1888,1914]" pageId="119" pageNumber="120">Description: Female:</emphasis>
Body length:
<quantity id="2E7A9B0FAC2E430C0976F8AB8CC6013A" box="[610,675,1888,1915]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="119" pageNumber="120" unit="mm" value="8.0">8 mm</quantity>
(
<figureCitation id="71B92A6FAC2E430C09A6F8AB8D5C013A" box="[690,825,1888,1915]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="119" pageNumber="120">Figure 75A</figureCitation>
).
<emphasis id="DBF6EAF8AC2E430C085FF8AB8DF4013B" box="[843,913,1888,1914]" italics="true" pageId="119" pageNumber="120">Head:</emphasis>
Dark, 1.2 times wider than long (
<figureCitation id="71B92A6FAC2E430C0E1FF8AB8BEA013A" box="[1291,1423,1888,1915]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="119" pageNumber="120">Figure 75B</figureCitation>
). Clypeus domed, irregularly punctate, punctures in lateral ⅓ separated by 0.5 puncture diameters, medially with impunctate longitudinal mid-line, puncture adjacent to mid-line separated by 0.52 puncture diameters; underlying surface smooth and shining. Process of labrum broadly trapezoidal, 2 times wider than long, anterior margin deeply emarginate (
<figureCitation id="71B92A6FAC2143030A36FF5C8FC606F0" box="[290,419,150,177]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75C</figureCitation>
). Galea with anterior surface smooth and shining (
<figureCitation id="71B92A6FAC21430308CFFF5C8A0406F0" box="[987,1121,150,177]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75D</figureCitation>
). Gena subequal to width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus (
<figureCitation id="71B92A6FAC21430308C9FF718A3A0694" box="[989,1119,186,213]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75E</figureCitation>
). Foveae dorsally occupying ⅔ of space between compound eye and lateral ocellus, narrowing slightly ventrally at level of antennal insertions; foveae filled with light brownish hairs. Face, gena, and scape with short white hairs, becoming intermixed with dark brown hairs on vertex, no facial hairs equalling length of scape. Antennae basally dark, A412 ventrally with brown scales; A3 exceeding A4, shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAAC2143030BD3FEA58D7C0551" blockId="120.[151,1437,150,2035]" pageId="120" pageNumber="121">
<emphasis id="DBF6EAF8AC2143030BD3FEA58F2D07C6" box="[199,328,366,391]" italics="true" pageId="120" pageNumber="121">Mesosoma:</emphasis>
Scutum and scutellum deeply and strongly punctate, punctures separated by &lt;0.51 puncture diameter, underlying surface smooth and shining (
<figureCitation id="71B92A6FAC21430309C6FE5A8D3307ED" box="[722,854,401,428]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75F</figureCitation>
). Pronotum with weak humeral angle, surface with weak longitudinal striations. Mesepisternum smooth and shining, deeply and densely punctate, punctures separated by 1 puncture diameter. Dorsolateral parts of propodeum finely microreticulate, sculpture overlain by network of weakly raised rugosity. Propodeal triangle laterally defined by weak carinae, internal surface predominantly covered with network of more strongly raised rugae with exception of apical ¼ which lacks rugae, propodeal triangle thus defined by change in surface sculpture (
<figureCitation id="71B92A6FAC214303094BFD888C82041F" box="[607,743,579,606]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75G</figureCitation>
). Mesepisternum with moderately long finely plumose white hairs, hairs on scutum and scutellum shorter, light brown, slightly more strongly plumose, but not sub-squamous. Propodeal corbicula incomplete, dorsal fringe composed of white plumose hairs, internal surface with scattered white hairs. Legs dark brown, pubescence whitish. Flocculus moderately developed, composed of white plumose hairs; femoral and tibial scopae composed of white simple hairs. Hind tarsal claws with short inner tooth. Wings hyaline, stigma and venation orange, nervulus antefurcal.
</paragraph>
<paragraph id="E93D36EAAC2143030BD3FCD08B12024B" blockId="120.[151,1437,150,2035]" pageId="120" pageNumber="121">
<emphasis id="DBF6EAF8AC2143030BD3FCD08F200575" box="[199,325,795,820]" italics="true" pageId="120" pageNumber="121">Metasoma:</emphasis>
Discs of T12 orange-red, discs of T35 black, marginal area of all terga lightened hyaline-yellow to dark brown (
<figureCitation id="71B92A6FAC2143030A57FCF68FAC0519" box="[323,457,829,856]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="120" pageNumber="121">Figure 75H</figureCitation>
). T2 laterally with weak indication of dark brown oval marks. Tergal discs and margins strongly and densely punctate, punctures separated by 0.5 puncture diameters, underlying surface shagreened, weakly shining. Tergal discs with scattered extremely short sparse golden hairs, T24 apically with fine white hairbands, broadly interrupted on T2, more or less complete on T3, complete on T4, hairbands obscuring underlying surface. Apical fringe of T5 medially dark brown, laterally with scattered long white hairs, hairs flanking pygidial plate dark brown. Pygidial plate rounded triangular, medial triangular area elevated, weakly punctate basally, surface dull.
</paragraph>
<paragraph id="E93D36EAAC2143030BD3FBDF8FCB030D" blockId="120.[151,1437,150,2035]" pageId="120" pageNumber="121">
<emphasis id="DBF6EAF8AC2143030BD3FBDF8F68026F" bold="true" box="[199,269,1044,1070]" pageId="120" pageNumber="121">Male:</emphasis>
Body length: 6.5 mm (
<figureCitation id="71B92A6FAC214303091EFBDF8CEA026F" box="[522,655,1044,1070]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76A</figureCitation>
).
<emphasis id="DBF6EAF8AC21430309B4FBDF8C83026F" box="[672,742,1044,1070]" italics="true" pageId="120" pageNumber="121">Head:</emphasis>
Dark, 1.3 times wider than long (
<figureCitation id="71B92A6FAC2143030F41FBDF8ABC026F" box="[1109,1241,1044,1070]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76C</figureCitation>
). Clypeus weakly domed, densely punctate, punctures separated by ≤0.5 puncture diameters, interspaces shining. Process of labrum narrow, quadrate, apical margin weakly emarginate. Gena broad, 1.5 times broader than width of compound eye, surface smooth and shining, punctate, punctures separated by 12 puncture diameters, posterior margin rounded; ocelloccipital distance 1.5 times diameter of lateral ocellus. Face with intermixed black and white hairs, white hairs more or less restricted to scape and antennal insertions; gena with white hairs ventrally, hairs becoming black dorsally, vertex with intermixed black and white hairs; longest facial hairs exceeding length of scape. Antennae dark, A413 ventrally lightened by brown scales; A3 short, slightly under ½ length of A4, A413 rectangular, clearly longer than wide.
</paragraph>
<paragraph id="E93D36EAAC2143030BD3FA9D8D7C00CC" blockId="120.[151,1437,150,2035]" pageId="120" pageNumber="121">
<emphasis id="DBF6EAF8AC2143030BD3FA9D8F2D032E" box="[199,328,1366,1391]" italics="true" pageId="120" pageNumber="121">Mesosoma:</emphasis>
Scutum and scutellum deeply and irregularly punctate, punctures separated by 0.53 puncture diameters, underlying surface smooth and shining (
<figureCitation id="71B92A6FAC21430309D7FAB28D2003D2" box="[707,837,1401,1427]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76B</figureCitation>
). Pronotum with strong humeral angle. Mesepisternum with granular shagreen, weakly shining, irregularly punctate, punctures with unevenly raised rims, separated by 12 puncture diameters. Dorsolateral parts of propodeum microreticulate, sculpture overlain by network of weakly raised rugosity. Propodeal triangle laterally defined by weak carinae, internal surface entirely covered with network of more strongly raised rugae, propodeal triangle thus defined by change in surface sculpture (
<figureCitation id="71B92A6FAC2143030E1AF9C38BF50063" box="[1294,1424,1544,1570]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76E</figureCitation>
). Mesepisternum, scutum, and scutellum with intermixed long finely plumose black and white hairs, propodeum with white hairs only. Legs dark brown to dark orange, pubescence whitish. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange, nervulus antefurcal.
</paragraph>
<paragraph id="E93D36EAAC2143030BD3F9538C0601B2" blockId="120.[151,1437,150,2035]" pageId="120" pageNumber="121">
<emphasis id="DBF6EAF8AC2143030BD3F9538F2000F0" box="[199,325,1688,1713]" italics="true" pageId="120" pageNumber="121">Metasoma:</emphasis>
Terga variably coloured, T2 lightened orange-red, orange-red colouration spreading weakly to discs of T1 and T3, otherwise discs becoming brown, T45 progressively becoming darker, apex of T5 almost black (
<figureCitation id="71B92A6FAC2143030BF0F9148F0D00B8" box="[228,360,1759,1785]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76F</figureCitation>
). Terga punctate, punctures separated by 0.51 puncture diameters, underlying surface weakly shagreened, more or less shining. Tergal discs with scattered extremely short sparse whitish hairs, T25 apically with fine white hairbands, broadly interrupted on T2, more or less complete on T3, complete on T45, hairbands partially obscuring underlying surface. T67 with long dark brown hairs overlying pygidial plate of T7. S8 columnar, ventral surface with lateral fan of brown hairs. Genital capsule slightly oval-elongate, gonocoxae apically produced into weak rounded points, gonostyli slightly broadened apically, spatulate, inner margin with small but distinctly raised bump medially (
<figureCitation id="71B92A6FAC2143030A82F87E8C7E018E" box="[406,539,1973,1999]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="120" pageNumber="121">Figure 76G</figureCitation>
). Penis valves basally slightly broadened, occupying ½ space between gonostyli, slightly narrowing medially and apically.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC2043020B83F85C8F3D01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209471" ID-Zenodo-Dep="13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="121" pageNumber="122" startId="121.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="121" targetType="figure">
<paragraph id="E93D36EAAC2043020B83F85C8F3D01B6" blockId="121.[151,1437,1943,2040]" pageId="121" pageNumber="122">
<emphasis id="DBF6EAF8AC2043020B83F85C8F4401F1" bold="true" box="[151,289,1943,1968]" pageId="121" pageNumber="122">FIGURE 75.</emphasis>
<taxonomicName id="2E824D69AC2043020A3CF85C8C0601EE" authorityName="WOOD" authorityYear="2024" box="[296,611,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="namaka" status="sp. nov." subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2043020A3CF85C8F1B01EE" box="[296,382,1943,1967]" italics="true" pageId="121" pageNumber="122">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2043020A9AF85C8C6101EE" box="[398,516,1943,1967]" italics="true" pageId="121" pageNumber="122">Troandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2043020907F85C8C0601EE" box="[531,611,1943,1967]" italics="true" pageId="121" pageNumber="122">namaka</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC204302097FF8538CB601F1" bold="true" box="[619,723,1944,1968]" pageId="121" pageNumber="122">
<taxonomicNameLabel id="C0C55783AC204302097FF8538CB601F1" box="[619,723,1944,1968]" pageId="121" pageNumber="122" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view.
</paragraph>
</caption>
<caption id="BDFD6662AC2343010B83F85C8DF901B6" ID-DOI="http://doi.org/10.5281/zenodo.13209475" ID-Zenodo-Dep="13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="122" pageNumber="123" startId="122.[151,250,1943,1967]" targetBox="[214,1374,181,1919]" targetPageId="122" targetType="figure">
<paragraph id="E93D36EAAC2343010B83F85C8DF901B6" blockId="122.[151,1437,1943,2039]" pageId="122" pageNumber="123">
<emphasis id="DBF6EAF8AC2343010B83F85C8F7A01EE" bold="true" box="[151,287,1943,1967]" pageId="122" pageNumber="123">FIGURE 76.</emphasis>
<taxonomicName id="2E824D69AC2343010A31F85C8C3801EE" authorityName="WOOD" authorityYear="2024" box="[293,605,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="namaka" status="sp. nov." subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2343010A31F85C8F1E01EE" box="[293,379,1943,1967]" italics="true" pageId="122" pageNumber="123">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2343010A9DF85C8F9A01EE" box="[393,511,1943,1967]" italics="true" pageId="122" pageNumber="123">Troandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2343010919F85C8C3801EE" box="[525,605,1943,1967]" italics="true" pageId="122" pageNumber="123">namaka</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC2343010977F8538CAF01F1" bold="true" box="[611,714,1944,1968]" pageId="122" pageNumber="123">
<taxonomicNameLabel id="C0C55783AC2343010977F8538CAF01F1" box="[611,714,1944,1968]" pageId="122" pageNumber="123" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC2343010F41F8708F5501B6" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="saettana" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2343010F41F8708ACE0192" box="[1109,1195,1979,2003]" italics="true" pageId="122" pageNumber="123">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC2343010FAAF8708B510192" box="[1214,1332,1979,2003]" italics="true" pageId="122" pageNumber="123">Troandrena</emphasis>
)
<emphasis id="DBF6EAF8AC2343010E52F8778BF90192" box="[1350,1436,1980,2003]" italics="true" pageId="122" pageNumber="123">saettana</emphasis>
Warncke, 1975
</taxonomicName>
male. D. Face, frontal view; H. Genital capsule, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC2243000BD3FF5D8FFB03A5" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="E93D36EAAC2243000BD3FF5D8AC704C5" blockId="123.[151,1437,150,1616]" pageId="123" pageNumber="124">
<emphasis id="DBF6EAF8AC2243000BD3FF5D8F2706F1" bold="true" box="[199,322,150,176]" pageId="123" pageNumber="124">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC2243000A53FF5D8C6306F1" authorityName="WOOD" authorityYear="2024" box="[327,518,150,176]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000A53FF5D8C6306F1" box="[327,518,150,176]" italics="true" pageId="123" pageNumber="124">Andrena namaka</emphasis>
</taxonomicName>
can be placed into the subgenus
<taxonomicName id="2E824D69AC2243000865FF5D8D9606F1" authorityName="Warncke" authorityYear="1975" box="[881,1011,150,176]" pageId="123" pageNumber="124" rank="subGenus" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2243000865FF5D8D9606F1" box="[881,1011,150,176]" italics="true" pageId="123" pageNumber="124">Troandrena</emphasis>
</taxonomicName>
, though placement is not immediately straightforward as
<taxonomicName id="2E824D69AC2243000A7EFF708F870695" authorityName="WOOD" authorityYear="2024" box="[362,482,186,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000A7EFF708F870695" box="[362,482,186,213]" italics="true" pageId="123" pageNumber="124">A. namaka</emphasis>
</taxonomicName>
differs from some of the classical
<taxonomicName id="2E824D69AC224300087CFF718D8F0695" authorityName="Warncke" authorityYear="1975" box="[872,1002,186,212]" pageId="123" pageNumber="124" rank="subGenus" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC224300087CFF718D8F0695" box="[872,1002,186,212]" italics="true" pageId="123" pageNumber="124">Troandrena</emphasis>
</taxonomicName>
concepts (and the concept of Warncke is subject to change, as
<taxonomicName id="2E824D69AC2243000A88FF148CBA06B9" authority="Brulle, 1839" authorityName="Brulle" authorityYear="1839" box="[412,735,222,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="chalcogastra">
<emphasis id="DBF6EAF8AC2243000A88FF148C2806B9" box="[412,589,222,249]" italics="true" pageId="123" pageNumber="124">A. chalcogastra</emphasis>
Brullé, 1839
</taxonomicName>
will be removed from this subgenus based on genetic analysis; S. Bossert, T.J. Wood, G. Pisanty,
<emphasis id="DBF6EAF8AC224300093FFEC98C95075D" box="[555,752,258,284]" italics="true" pageId="123" pageNumber="124">unpublished data</emphasis>
). The subgenus has been previously recognised due to the strongly flattened clypeus with the surface shining and with a slightly raised impunctate mid-line, with relatively narrow facial foveae, and with the process of the labrum strongly emarginate medially. Accepting the removal of
<taxonomicName id="2E824D69AC2243000E90FE808F4C07C9" authorityName="Brulle" authorityYear="1839" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="chalcogastra">
<emphasis id="DBF6EAF8AC2243000E90FE808F4C07C9" italics="true" pageId="123" pageNumber="124">A. chalcogastra</emphasis>
</taxonomicName>
, subgenus
<taxonomicName id="2E824D69AC2243000AB1FEA58C4207C9" authorityName="Warncke" authorityYear="1975" box="[421,551,366,392]" pageId="123" pageNumber="124" rank="subGenus" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2243000AB1FEA58C4207C9" box="[421,551,366,392]" italics="true" pageId="123" pageNumber="124">Troandrena</emphasis>
</taxonomicName>
was thought to include two species (
<taxonomicName id="2E824D69AC22430008D4FEA48A8E07C8" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[960,1259,366,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="saettana">
<emphasis id="DBF6EAF8AC22430008D4FEA48A5B07C9" box="[960,1086,367,393]" italics="true" pageId="123" pageNumber="124">A. saettana</emphasis>
Warncke, 1975
</taxonomicName>
and
<taxonomicName id="2E824D69AC2243000E34FEA48F2707EC" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="troodica">
<emphasis id="DBF6EAF8AC2243000E34FEA48BF907C9" box="[1312,1436,366,393]" italics="true" pageId="123" pageNumber="124">A. troodica</emphasis>
Warncke, 1975
</taxonomicName>
) between eastern
<collectingCountry id="9195767AAC2243000904FE598C3A07ED" box="[528,607,402,428]" name="Greece" pageId="123" pageNumber="124">Greece</collectingCountry>
and western
<collectingCountry id="9195767AAC22430009ECFE598D3207ED" box="[760,855,402,428]" name="Pakistan" pageId="123" pageNumber="124">Pakistan</collectingCountry>
(see comments in
<bibRefCitation id="8D134B1BAC2243000F3AFE598AD107ED" author="Wood, T. J." box="[1070,1204,402,428]" pageId="123" pageNumber="124" pagination="167 - 188" refId="ref92492" refString="Wood, T. J. (2024) New Asian Andrena species, with notes on the subgenus Cnemidandrena (Hymenoptera: Andrenidae). Zootaxa, 5404 (1), 167 - 188. https: // doi. org / 10.11646 / zootaxa. 5404.1.11" type="journal article" year="2024">Wood 2024</bibRefCitation>
), and a third species from central
<collectingCountry id="9195767AAC2243000A32FE7C8F360790" box="[294,339,439,465]" name="Iran" pageId="123" pageNumber="124">Iran</collectingCountry>
(
<taxonomicName id="2E824D69AC2243000A77FE7C8CE70791" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[355,642,438,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000A77FE7C8F960791" box="[355,499,438,465]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
Wood, 2023
</taxonomicName>
) was recently described that displays a clypeus which is not completely flattened but instead is slightly more domed.
<taxonomicName id="2E824D69AC2243000980FE118D3107B5" authorityName="WOOD" authorityYear="2024" box="[660,852,474,500]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000980FE118D3107B5" box="[660,852,474,500]" italics="true" pageId="123" pageNumber="124">Andrena namaka</emphasis>
</taxonomicName>
displays this partially flattened and shining clypeus with an impunctate mid-line (
<figureCitation id="71B92A6FAC2243000AFDFE358C0B0458" box="[489,622,510,537]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75B</figureCitation>
), and is therefore closer to
<taxonomicName id="2E824D69AC22430008BCFE348A5E0459" authorityName="Wood" authorityYear="2023" box="[936,1083,510,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC22430008BCFE348A5E0459" box="[936,1083,510,537]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
, as well as showing a strongly emarginate process of the labrum. Placement in the
<taxonomicName id="2E824D69AC22430009E7FDE98D10047D" authorityName="Warncke" authorityYear="1975" box="[755,885,546,572]" pageId="123" pageNumber="124" rank="subGenus" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC22430009E7FDE98D10047D" box="[755,885,546,572]" italics="true" pageId="123" pageNumber="124">Troandrena</emphasis>
</taxonomicName>
can also be informed by the male morphology, specifically the gena that is broader than the compound eye, the pronotum with a very strong humeral angle, though the genital capsule is not typical for the two species for which
<taxonomicName id="2E824D69AC2243000847FDA18DB004C5" authorityName="Warncke" authorityYear="1975" box="[851,981,618,644]" pageId="123" pageNumber="124" rank="subGenus" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC2243000847FDA18DB004C5" box="[851,981,618,644]" italics="true" pageId="123" pageNumber="124">Troandrena</emphasis>
</taxonomicName>
males are known.
</paragraph>
<paragraph id="E93D36EAAC2243000BD3FD448A9102E1" blockId="123.[151,1437,150,1616]" pageId="123" pageNumber="124">
In the female sex,
<taxonomicName id="2E824D69AC2243000A8AFD448C7D04E9" authorityName="WOOD" authorityYear="2024" box="[414,536,654,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000A8AFD448C7D04E9" box="[414,536,654,681]" italics="true" pageId="123" pageNumber="124">A. namaka</emphasis>
</taxonomicName>
is specifically close to
<taxonomicName id="2E824D69AC224300083DFD448DD904E9" authorityName="Wood" authorityYear="2023" box="[809,956,654,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC224300083DFD448DD904E9" box="[809,956,654,681]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
due to the partially red-marked terga, the strongly and densely punctate terga with punctures separated by 0.5 puncture diameters (
<figureCitation id="71B92A6FAC2243000F8DFD798B47048C" box="[1177,1314,690,717]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75H</figureCitation>
), the short clypeus (not projecting ventrally beyond the lower margin of the malar areas), the relatively domed clypeus with a clear impunctate mid-line (
<figureCitation id="71B92A6FAC2243000AC8FD318C040554" box="[476,609,762,789]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75B</figureCitation>
), the scutum smooth and shining, with large and dense punctures (
<figureCitation id="71B92A6FAC2243000E45FD318EA30578" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75F</figureCitation>
), and by the relatively small body size (&lt;
<quantity id="2E7A9B0FAC224300099CFCD58CB30578" box="[648,726,798,825]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="123" pageNumber="124" unit="mm" value="10.0">10 mm</quantity>
in length).
<taxonomicName id="2E824D69AC2243000846FCD58A750579" authorityName="WOOD" authorityYear="2024" box="[850,1040,798,824]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000846FCD58A750579" box="[850,1040,798,824]" italics="true" pageId="123" pageNumber="124">Andrena namaka</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="2E824D69AC2243000E19FCD48BF90579" authorityName="Wood" authorityYear="2023" box="[1293,1436,798,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000E19FCD48BF90579" box="[1293,1436,798,825]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
due to its smaller body size of
<quantity id="2E7A9B0FAC2243000AFAFC898C4A051C" box="[494,559,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="123" pageNumber="124" unit="mm" value="8.0">8 mm</quantity>
(99.5 mm in
<taxonomicName id="2E824D69AC22430009C1FC888D03051D" authorityName="Wood" authorityYear="2023" box="[725,870,834,861]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC22430009C1FC888D03051D" box="[725,870,834,861]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
), the slightly narrower facial foveae, occupying ⅔ of the space between the compound eye and a lateral ocellus (
<figureCitation id="71B92A6FAC224300085BFCAD8DB705C0" box="[847,978,870,897]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75E</figureCitation>
), upper margin of foveae separated from a lateral ocellus by a distance greater than its diameter (in
<taxonomicName id="2E824D69AC2243000838FC408DDB05E5" authorityName="Wood" authorityYear="2023" box="[812,958,906,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000838FC408DDB05E5" box="[812,958,906,933]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
with the facial foveae occupying ¾ of the space between the compound eye and a lateral ocellus, upper margin of foveae separated from a lateral ocellus by a distance less than its diameter), clypeus adjacent to impunctate mid-line less densely punctate, punctures separated by 0.52 puncture diameters (in
<taxonomicName id="2E824D69AC2243000918FC3C8CFA0251" authorityName="Wood" authorityYear="2023" box="[524,671,1014,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000918FC3C8CFA0251" box="[524,671,1014,1041]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
with the clypeus densely punctate adjacent to the impunctate mid-line, punctures separated by 0.5 puncture diameters), the outer surface of the galea smooth and shining (
<figureCitation id="71B92A6FAC2243000E07FBD18BFD0274" box="[1299,1432,1050,1077]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75D</figureCitation>
; in
<taxonomicName id="2E824D69AC2243000BA1FBF48F220219" authorityName="Wood" authorityYear="2023" box="[181,327,1086,1113]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000BA1FBF48F220219" box="[181,327,1086,1113]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
with the surface of the galea showing clear fine microsculpture), terga with fine apical hairbands, interrupted on T23, complete only on T4 (
<figureCitation id="71B92A6FAC224300096BFBA98D60023C" box="[639,773,1122,1149]" captionStart="FIGURE 75" captionStartId="121.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@121.[214,1373,181,1919]" captionTargetPageId="121" captionText="FIGURE 75. Andrena (Troandrena) namaka spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Process of labrum, ventral view; D. Galea, lateral view; E. Head, dorsal view; F. Scutum, dorsal view; G. Propodeum, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209471" httpUri="https://zenodo.org/record/13209471/files/figure.png" pageId="123" pageNumber="124">Figure 75H</figureCitation>
; in
<taxonomicName id="2E824D69AC224300083AFBA88DA5023D" authorityName="Wood" authorityYear="2023" box="[814,960,1122,1149]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC224300083AFBA88DA5023D" box="[814,960,1122,1149]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
with complete apical hairbands on T24), and terminal fringe medially dark brown (in
<taxonomicName id="2E824D69AC224300099FFB4C8D7902E1" authorityName="Wood" authorityYear="2023" box="[651,796,1158,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC224300099FFB4C8D7902E1" box="[651,796,1158,1185]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
terminal fringe medially golden-whitish).
</paragraph>
<paragraph id="E93D36EAAC2243000BD3FB608FFB03A5" blockId="123.[151,1437,150,1616]" pageId="123" pageNumber="124">
In the male sex, the male of
<taxonomicName id="2E824D69AC2243000919FB608CFA0285" authorityName="Wood" authorityYear="2023" box="[525,671,1194,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="monfaredi">
<emphasis id="DBF6EAF8AC2243000919FB608CFA0285" box="[525,671,1194,1221]" italics="true" pageId="123" pageNumber="124">A. monfaredi</emphasis>
</taxonomicName>
is unknown. Separation from
<taxonomicName id="2E824D69AC22430008ECFB608A1D0285" authorityName="Warncke" authorityYear="1975" box="[1016,1144,1195,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="saettana">
<emphasis id="DBF6EAF8AC22430008ECFB608A1D0285" box="[1016,1144,1195,1221]" italics="true" pageId="123" pageNumber="124">A. saettana</emphasis>
</taxonomicName>
(which also displays red-marked terga, and which is present in western
<collectingCountry id="9195767AAC22430009B9FB058D6B02A9" box="[685,782,1230,1256]" name="Pakistan" pageId="123" pageNumber="124">Pakistan</collectingCountry>
) can be made by the genital capsule which in
<taxonomicName id="2E824D69AC2243000E37FB048BF802A9" authorityName="WOOD" authorityYear="2024" box="[1315,1437,1230,1257]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000E37FB048BF802A9" box="[1315,1437,1230,1257]" italics="true" pageId="123" pageNumber="124">A. namaka</emphasis>
</taxonomicName>
is simple and resembles that of a member of the
<taxonomicName id="2E824D69AC22430009C6FB398D28034D" authorityName="Hedicke" authorityYear="1933" box="[722,845,1266,1292]" pageId="123" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC22430009C6FB398D28034D" box="[722,845,1266,1292]" italics="true" pageId="123" pageNumber="124">Euandrena</emphasis>
</taxonomicName>
with the gonocoxae apically produced into weakly rounded points (
<figureCitation id="71B92A6FAC2243000A47FADD8FBE0371" box="[339,475,1302,1328]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="123" pageNumber="124">Figure 76G</figureCitation>
; in
<taxonomicName id="2E824D69AC224300091EFADC8CEE0371" authorityName="Warncke" authorityYear="1975" box="[522,651,1303,1329]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="saettana">
<emphasis id="DBF6EAF8AC224300091EFADC8CEE0371" box="[522,651,1303,1329]" italics="true" pageId="123" pageNumber="124">A. saettana</emphasis>
</taxonomicName>
the gonocoxae have their inner margins forming obtuse angles, not produced into rounded points,
<figureCitation id="71B92A6FAC2243000AFDFAF18C090315" box="[489,620,1338,1364]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="123" pageNumber="124">Figure 76H</figureCitation>
), by the clypeus which is extremely short, ventrally not projecting beyond the lower margin of the malar areas, covered in a mixture of black and grey hairs (
<figureCitation id="71B92A6FAC2243000F57FA958AAD0339" box="[1091,1224,1374,1400]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="123" pageNumber="124">Figure 76C</figureCitation>
; in
<taxonomicName id="2E824D69AC2243000FE7FA948B170339" authorityName="Warncke" authorityYear="1975" box="[1267,1394,1375,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="saettana">
<emphasis id="DBF6EAF8AC2243000FE7FA948B170339" box="[1267,1394,1375,1401]" italics="true" pageId="123" pageNumber="124">A. saettana</emphasis>
</taxonomicName>
the clypeus is long, ventrally clearly projecting beyond the lower margin of the malar areas, surface covered with pale hairs,
<figureCitation id="71B92A6FAC2243000BCAFA6D8F060381" box="[222,355,1446,1472]" captionStart="FIGURE 76" captionStartId="122.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-28@122.[214,1374,181,1919]" captionTargetPageId="122" captionText="FIGURE 76. Andrena (Troandrena) namaka spec. nov. male. A. Habitus, lateral view; B. Scutum, dorsal view; C. Face, frontal view; E. Propodeum, dorsal view; F. Terga, dorsal view; G. Genital capsule, dorsal view; Andrena (Troandrena) saettana Warncke, 1975 male. D. Face, frontal view; H. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209475" httpUri="https://zenodo.org/record/13209475/files/figure.png" pageId="123" pageNumber="124">Figure 76D</figureCitation>
), and finally
<taxonomicName id="2E824D69AC2243000AEAFA6C8C1D0381" authorityName="WOOD" authorityYear="2024" box="[510,632,1446,1473]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="namaka">
<emphasis id="DBF6EAF8AC2243000AEAFA6C8C1D0381" box="[510,632,1446,1473]" italics="true" pageId="123" pageNumber="124">A. namaka</emphasis>
</taxonomicName>
is also small (6.5 mm in length), whereas
<taxonomicName id="2E824D69AC2243000F72FA6C8A820381" authorityName="Warncke" authorityYear="1975" box="[1126,1255,1447,1473]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="saettana">
<emphasis id="DBF6EAF8AC2243000F72FA6C8A820381" box="[1126,1255,1447,1473]" italics="true" pageId="123" pageNumber="124">A. saettana</emphasis>
</taxonomicName>
males typically exceed
<quantity id="2E7A9B0FAC2243000BF8FA018F4803A4" box="[236,301,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="123" pageNumber="124" unit="mm" value="8.0">8 mm</quantity>
in length.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2243000BD3FA258A1B006D" pageId="123" pageNumber="124" type="etymology">
<paragraph id="E93D36EAAC2243000BD3FA258A1B006D" blockId="123.[151,1437,150,1616]" pageId="123" pageNumber="124">
<emphasis id="DBF6EAF8AC2243000BD3FA258F2B0049" bold="true" box="[199,334,1518,1544]" pageId="123" pageNumber="124">Etymology.</emphasis>
From the Urdu noun
<emphasis id="DBF6EAF8AC224300092FFA258CE30049" box="[571,646,1518,1544]" italics="true" pageId="123" pageNumber="124">namak</emphasis>
which means salt or rock salt, in reference to the Salt Range in northern
<collectingRegion id="2B46F808AC2243000B83F9D98E80006D" box="[151,229,1554,1580]" country="Pakistan" name="Punjab" pageId="123" pageNumber="124">Punjab</collectingRegion>
which is naturally known for its extensive salt deposits. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC2243000BD3F9FD8D3B0011" box="[199,862,1590,1616]" pageId="123" pageNumber="124" type="distribution">
<paragraph id="E93D36EAAC2243000BD3F9FD8D3B0011" blockId="123.[151,1437,150,1616]" box="[199,862,1590,1616]" pageId="123" pageNumber="124">
<emphasis id="DBF6EAF8AC2243000BD3F9FD8F040011" bold="true" box="[199,353,1590,1616]" pageId="123" pageNumber="124">Distribution.</emphasis>
Central
<collectingCountry id="9195767AAC2243000AD5F9FD8C450011" box="[449,544,1590,1616]" name="Pakistan" pageId="123" pageNumber="124">Pakistan</collectingCountry>
(northern
<collectingRegion id="2B46F808AC2243000982F9FD8D370011" box="[662,850,1590,1616]" country="Pakistan" name="Punjab" pageId="123" pageNumber="124">Punjab province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,423 @@
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<mods:title id="C03441610E75CC110B7D3DE3017543DC">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<paragraph id="E93D36EAAC3F431D0B83FB398CDC034C" blockId="102.[151,697,1266,1329]" box="[151,697,1266,1293]" pageId="102" pageNumber="103">
<heading id="B2758186AC3F431D0B83FB398CDC034C" bold="true" box="[151,697,1266,1293]" fontSize="11" level="2" pageId="102" pageNumber="103" reason="4">
<emphasis id="DBF6EAF8AC3F431D0B83FB398CDC034C" bold="true" box="[151,697,1266,1293]" pageId="102" pageNumber="103">
<taxonomicName id="2E824D69AC3F431D0B83FB398C5F034C" authorityName="WOOD" authorityYear="2024" box="[151,570,1266,1293]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="102" pageNumber="103" phylum="Arthropoda" rank="species" species="balticola" status="sp. nov." subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3F431D0B83FB398F64034C" bold="true" box="[151,257,1266,1293]" italics="true" pageId="102" pageNumber="103">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC3F431D0A06FB398FA5034C" bold="true" box="[274,448,1266,1293]" italics="true" pageId="102" pageNumber="103">Graecandrena</emphasis>
)
<emphasis id="DBF6EAF8AC3F431D0AC6FB398C5F034C" bold="true" box="[466,570,1266,1293]" italics="true" pageId="102" pageNumber="103">balticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC3F431D0955FB388CDC034C" box="[577,697,1267,1293]" pageId="102" pageNumber="103" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3F431D0B83FADC8F290370" box="[151,332,1302,1329]" pageId="102" pageNumber="103" type="description">
<paragraph id="E93D36EAAC3F431D0B83FADC8F290370" blockId="102.[151,697,1266,1329]" box="[151,332,1302,1329]" pageId="102" pageNumber="103">
<figureCitation id="71B92A6FAC3F431D0B83FADC8F2D0370" box="[151,328,1302,1329]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figures 65AH</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3F431D0B83FA958F5F03DC" pageId="102" pageNumber="103" type="materials_examined">
<paragraph id="E93D36EAAC3F431D0B83FA958F5F03DC" blockId="102.[151,1437,1374,2013]" pageId="102" pageNumber="103">
<materialsCitation id="59EA3CB7AC3F431D0B83FA958F5F03DC" collectingDate="2008-05-11" collectionCode="TUZ" collectorName="V. Soon" country="Estonia" location="Kalmistu" pageId="102" pageNumber="103" specimenCode="TUZ 051846" specimenCount="1" specimenCount-female="1" stateProvince="Tartumaa" typeStatus="holotype">
<emphasis id="DBF6EAF8AC3F431D0B83FA958FA90339" bold="true" box="[151,460,1374,1400]" pageId="102" pageNumber="103">
<typeStatus id="36398848AC3F431D0B83FA958F5F0339" box="[151,314,1374,1400]" pageId="102" pageNumber="103" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC3F431D0A51FA958FA20339" box="[325,455,1374,1400]" name="Estonia" pageId="102" pageNumber="103">ESTONIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC3F431D0AC6FA948C750338" box="[466,528,1375,1401]" country="Estonia" name="Tartumaa" pageId="102" pageNumber="103">Tartu</collectingRegion>
,
<location id="EC5D6031AC3F431D090EFA948CE40338" LSID="urn:lsid:plazi:treatment:612B87FCAC3F43120B83FB398FD104C6:EC5D6031AC3F431D090EFA948CE40338" box="[538,641,1375,1401]" country="Estonia" name="Kalmistu" pageId="102" pageNumber="103" stateProvince="Tartumaa">Kalmistu</location>
paljand, 58.3948
<superScript id="1EF79BA2AC3F431D0855FA948D050338" attach="right" box="[833,864,1375,1401]" fontSize="6" pageId="102" pageNumber="103">oN</superScript>
, 26.7098
<superScript id="1EF79BA2AC3F431D08D1FA948D850338" attach="right" box="[965,992,1375,1401]" fontSize="6" pageId="102" pageNumber="103">oE</superScript>
,
<date id="9D3C102AAC3F431D08FFFA948A3E0338" box="[1003,1115,1375,1401]" pageId="102" pageNumber="103" value="2008-05-11">
<collectingDate id="8D78E9C2AC3F431D08FFFA948A3E0338" box="[1003,1115,1375,1401]" pageId="102" pageNumber="103" value="2008-05-11">11.v.2008</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC3F431D0F72FA948A1E0338" box="[1126,1147,1375,1401]" count="1" pageId="102" pageNumber="103" type="female"></specimenCount>
, leg.
<collectorName id="4477533CAC3F431D0FA2FA948B750338" box="[1206,1296,1375,1401]" pageId="102" pageNumber="103">V. Soon</collectorName>
,
<collectionCode id="8F93AE2FAC3F431D0E0EFA948B350338" box="[1306,1360,1375,1401]" pageId="102" pageNumber="103">TUZ</collectionCode>
(code:
<specimenCode id="B9249E91AC3F431D0B83FA488F4B03DC" box="[151,302,1410,1437]" collectionCode="TUZ" pageId="102" pageNumber="103">TUZ 051846</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3F43130BD3FA6D8FEF047C" lastPageId="104" lastPageNumber="105" pageId="102" pageNumber="103" type="description">
<paragraph id="E93D36EAAC3F431D0BD3FA6D8F2B00A0" blockId="102.[151,1437,1374,2013]" pageId="102" pageNumber="103">
<emphasis id="DBF6EAF8AC3F431D0BD3FA6D8FA00380" bold="true" box="[199,453,1446,1473]" pageId="102" pageNumber="103">Description: Female:</emphasis>
Body length: 5.5 mm (
<figureCitation id="71B92A6FAC3F431D09C1FA6C8D3B0380" box="[725,862,1446,1473]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65A</figureCitation>
).
<emphasis id="DBF6EAF8AC3F431D0867FA6C8DDC0380" box="[883,953,1447,1473]" italics="true" pageId="102" pageNumber="103">Head:</emphasis>
Dark, 1.15 times wider than long (
<figureCitation id="71B92A6FAC3F431D0E45FA6C8EAF03A4" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65B</figureCitation>
). Clypeus strongly domed, irregularly and shallowly punctate, punctures separated by 13 puncture diameters, underlying surface with even granular shagreen, weakly shining (
<figureCitation id="71B92A6FAC3F431D08BFFA248A530048" box="[939,1078,1518,1545]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65C</figureCitation>
). Process of labrum narrowly trapezoidal, slightly longer than wide. Gena subequal to width of compound eye; ocelloccipital distance slightly less than ½ diameter of lateral ocellus. Foveae dorsally moderately broad, occupying ½ space between compound eye and lateral ocellus, ventrally only narrowly slightly at level of antennal insertions; foveae filled with light brown hairs (
<figureCitation id="71B92A6FAC3F431D0BC9F9B48F0600D8" box="[221,355,1662,1689]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65D</figureCitation>
). Face, gena, vertex, and scape with short whitish hairs. Antennae basally dark, A412 ventrally lightened by presence of brownish scales; A3 exceeding A4, shorter than A4+5, A4 and A5 subquadrate, slightly wider than long.
</paragraph>
<paragraph id="E93D36EAAC3F43130BD3F9208DE00700" blockId="102.[151,1437,1374,2013]" lastBlockId="104.[151,1437,151,2013]" lastPageId="104" lastPageNumber="105" pageId="102" pageNumber="103">
<emphasis id="DBF6EAF8AC3F431D0BD3F9208F2D0145" box="[199,328,1771,1796]" italics="true" pageId="102" pageNumber="103">Mesosoma:</emphasis>
Scutum and scutellum finely shagreened, weakly shining, with weak metallic green-copper reflections; surface finely and obscurely punctate, punctures separated by 2 puncture diameters, individual punctures difficult to discern against background sculpture (
<figureCitation id="71B92A6FAC3F431D09FDF8F88D17010C" box="[745,882,1842,1869]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65E</figureCitation>
). Pronotum rounded. Mesepisternum with fine granular shagreen, shining. Dorsolateral parts of propodeum with strong granular microreticulation, dull; propodeal triangle laterally without carinae, surface with strong granular microreticulation overlain almost entirely with dense network of irregular winding rugae, propodeal triangle thus defined by change in surface sculpture (
<figureCitation id="71B92A6FAC3F431D0E19F8548BF501F8" box="[1293,1424,1950,1977]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="102" pageNumber="103">Figure 65F</figureCitation>
). Mesepisternum almost hairless, dorsally with sparse densely plumose and sub-squamous light brownish hairs, becoming shorter on scutum and scutellum. Propodeal corbicula incomplete, dorsal fringe long, composed of plumose yellowish hairs, internal surface with occasional long yellowish simple hairs. Legs brownish, hind basitarsi lightened orange, pubescence light brown. Flocculus complete, composed of light brownish plumose hairs; femoral and tibial scopae composed of whitish simple hairs. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation orange-brown, nervulus strongly antefurcal (
<figureCitation id="71B92A6FAC31431309E6FEEC8D1D0700" box="[754,888,294,321]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="104" pageNumber="105">Figure 65G</figureCitation>
).
</paragraph>
<caption id="BDFD6662AC3E431C0B83F85C8F3D01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209447" ID-Zenodo-Dep="13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="103" pageNumber="104" startId="103.[151,250,1943,1967]" targetBox="[215,1373,181,1919]" targetPageId="103" targetType="figure">
<paragraph id="E93D36EAAC3E431C0B83F85C8F3D01B6" blockId="103.[151,1437,1943,2040]" pageId="103" pageNumber="104">
<emphasis id="DBF6EAF8AC3E431C0B83F85C8F4501EE" bold="true" box="[151,288,1943,1967]" pageId="103" pageNumber="104">FIGURE 65.</emphasis>
<taxonomicName id="2E824D69AC3E431C0A32F85C8CE201EE" authorityName="WOOD" authorityYear="2024" box="[294,647,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="balticola" status="sp. nov." subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3E431C0A32F85C8F1901EE" box="[294,380,1943,1967]" italics="true" pageId="103" pageNumber="104">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC3E431C0A9FF85C8C7B01EE" box="[395,542,1943,1967]" italics="true" pageId="103" pageNumber="104">Graecandrena</emphasis>
)
<emphasis id="DBF6EAF8AC3E431C0939F85C8CE201EE" box="[557,647,1943,1967]" italics="true" pageId="103" pageNumber="104">balticola</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC3E431C0999F8538C9101F1" bold="true" box="[653,756,1944,1968]" pageId="103" pageNumber="104">
<taxonomicNameLabel id="C0C55783AC3E431C0999F8538C9101F1" box="[653,756,1944,1968]" pageId="103" pageNumber="104" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC3143130BD3FE808C890458" blockId="104.[151,1437,151,2013]" pageId="104" pageNumber="105">
<emphasis id="DBF6EAF8AC3143130BD3FE808F200725" box="[199,325,331,356]" italics="true" pageId="104" pageNumber="105">Metasoma:</emphasis>
Tergal discs dark, T1 with apical rim narrowly lightened hyaline, T24 with marginal areas more broadly lightened hyaline-brown (
<figureCitation id="71B92A6FAC3143130904FEA48CF307C8" box="[528,662,366,393]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="104" pageNumber="105">Figure 65H</figureCitation>
). Tergal discs with regular granular microreticulation, weakly shining, surface impunctate. T24 apically with narrow hairbands, broadly interrupted on T2, slightly interrupted on T3, complete on T4, not greatly extending beyond apical rim of marginal areas, hairbands obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate golden-brown. Pygidial plate large, rounded triangular, surface with fine granular microreticulation, broadly shining.
</paragraph>
<paragraph id="E93D36EAAC3143130BD3FDE88FEF047C" blockId="104.[151,1437,151,2013]" box="[199,394,547,573]" pageId="104" pageNumber="105">
<emphasis id="DBF6EAF8AC3143130BD3FDE88F68047C" bold="true" box="[199,269,547,573]" pageId="104" pageNumber="105">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3143130BD3FD8D8C8301D4" pageId="104" pageNumber="105" type="diagnosis">
<paragraph id="E93D36EAAC3143130BD3FD8D8E85023C" blockId="104.[151,1437,151,2013]" pageId="104" pageNumber="105">
<emphasis id="DBF6EAF8AC3143130BD3FD8D8F270421" bold="true" box="[199,322,582,608]" pageId="104" pageNumber="105">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC3143130A5DFD8C8C700420" authorityName="WOOD" authorityYear="2024" box="[329,533,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="balticola">
<emphasis id="DBF6EAF8AC3143130A5DFD8C8C700420" box="[329,533,583,609]" italics="true" pageId="104" pageNumber="105">Andrena balticola</emphasis>
</taxonomicName>
presents an unusual combination of morphological characters. Due to the small body size, moderately developed facial foveae that are not strongly narrowed ventrally, hind tarsal claws with inner tooth, propodeal triangle that is not laterally defined by raised carinae (no carinae present) but which has the internal surface entirely covered with a dense network of irregular winding rugae, and strongly antefurcal nervulus, it does not neatly fit into the typical
<taxonomicName id="2E824D69AC3143130AC5FD1C8C5504B0" box="[465,560,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC3143130AC5FD1C8C5504B0" box="[465,560,727,753]" italics="true" pageId="104" pageNumber="105">Andrena</emphasis>
</taxonomicName>
subgenera. It cannot be an
<taxonomicName id="2E824D69AC314313084CFD1C8DBF04B0" authorityName="Warncke" authorityYear="1968" box="[856,986,727,753]" pageId="104" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC314313084CFD1C8DBF04B0" box="[856,986,727,753]" italics="true" pageId="104" pageNumber="105">Aciandrena</emphasis>
</taxonomicName>
, since the hind tarsal claws have an inner subapical tooth, and the propodeal triangle is clearly covered in rugae. It also does not fit the concept any groups within the subgenus
<taxonomicName id="2E824D69AC3143130A69FCD48C750578" authorityName="Ashmead" authorityYear="1899" box="[381,528,799,825]" pageId="104" pageNumber="105" rank="subGenus" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC3143130A69FCD48C750578" box="[381,528,799,825]" italics="true" pageId="104" pageNumber="105">Micrandrena</emphasis>
</taxonomicName>
, since the propodeal triangle lacks lateral carinae, the foveae are not strongly and linearly narrowed ventrally (
<emphasis id="DBF6EAF8AC3143130AC2FC888C32051C" box="[470,599,835,861]" italics="true" pageId="104" pageNumber="105">longibarbis</emphasis>
-species group), and the nervulus is strongly antefurcal. The best placement is therefore in the subgenus
<taxonomicName id="2E824D69AC3143130AC3FCAC8D4905C0" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[471,812,871,897]" pageId="104" pageNumber="105" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130AC3FCAC8C1C05C0" box="[471,633,871,897]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
Warncke, 1968
</taxonomicName>
due to these aforementioned characters. It differs from most members of the subgenus due to the more extensive network of rugae on the propodeal triangle, whereas most European
<taxonomicName id="2E824D69AC3143130A1FFC648FC80588" authorityName="Warncke" authorityYear="1968" box="[267,429,943,969]" pageId="104" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130A1FFC648FC80588" box="[267,429,943,969]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
</taxonomicName>
have only a small area with rugae basally (e.g.
<taxonomicName id="2E824D69AC31431308C4FC648B750588" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[976,1296,942,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="graecella">
<emphasis id="DBF6EAF8AC31431308C4FC648A3E0588" box="[976,1115,943,969]" italics="true" pageId="104" pageNumber="105">A. graecella</emphasis>
<bibRefCitation id="8D134B1BAC3143130F77FC648B750588" author="Warncke, K." box="[1123,1296,942,969]" pageId="104" pageNumber="105" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2E824D69AC3143130E09FC648F2005AC" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="hyemala">
<emphasis id="DBF6EAF8AC3143130E09FC648BF90588" box="[1309,1436,943,969]" italics="true" pageId="104" pageNumber="105">A. hyemala</emphasis>
<bibRefCitation id="8D134B1BAC3143130B83FC188F2005AC" author="Warncke, K." box="[151,325,979,1005]" pageId="104" pageNumber="105" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
). However, this character is variable as
<taxonomicName id="2E824D69AC314313081CFC188A7805AC" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[776,1053,978,1005]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="verticalis">
<emphasis id="DBF6EAF8AC314313081CFC188DF505AC" box="[776,912,979,1005]" italics="true" pageId="104" pageNumber="105">A. verticalis</emphasis>
Pérez, 1895
</taxonomicName>
(which can be confidently placed in the
<taxonomicName id="2E824D69AC3143130BF5FC3C8FE60250" authorityName="Warncke" authorityYear="1968" box="[225,387,1015,1041]" pageId="104" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130BF5FC3C8FE60250" box="[225,387,1015,1041]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
</taxonomicName>
based on genetic evidence;
<bibRefCitation id="8D134B1BAC31431309D3FC3C8DC90250" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[711,940,1015,1041]" pageId="104" pageNumber="105" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC3143130836FC3C8D300250" box="[802,853,1015,1041]" italics="true" pageId="104" pageNumber="105">et al</emphasis>
. 2022b
</bibRefCitation>
) lacks any rugae on the propodeal triangle. Ultimately the location of the male with a typical
<taxonomicName id="2E824D69AC31431309AFFBD08D380274" authorityName="Warncke" authorityYear="1968" box="[699,861,1051,1077]" pageId="104" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC31431309AFFBD08D380274" box="[699,861,1051,1077]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
</taxonomicName>
genital capsule or genetic work will be necessary to confirm or disprove this phylogenetic placement (as well as more precisely defining the boundaries of the subgenus itself).
</paragraph>
<paragraph id="E93D36EAAC3143130BD3FB4C8B6700D8" blockId="104.[151,1437,151,2013]" pageId="104" pageNumber="105">
The most morphologically similar species are
<taxonomicName id="2E824D69AC31431309CAFB4C8AAF02E0" authority="Wood, 2023" authorityName="Wood" authorityYear="2023" box="[734,1226,1159,1185]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC31431309CAFB4C8C9302E0" box="[734,758,1159,1185]" italics="true" pageId="104" pageNumber="105">A.</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8AC31431308A3FB4C8A5202E0" box="[951,1079,1159,1185]" italics="true" pageId="104" pageNumber="105">orichalcum</emphasis>
Wood, 2023
</taxonomicName>
(southern
<collectingCountry id="9195767AAC3143130E53FB4C8BFD02E0" box="[1351,1432,1159,1185]" name="Turkey" pageId="104" pageNumber="105">Turkey</collectingCountry>
, northern
<collectingCountry id="9195767AAC3143130BE8FB608F5E0284" box="[252,315,1195,1221]" name="Syria" pageId="104" pageNumber="105">Syria</collectingCountry>
*) and
<taxonomicName id="2E824D69AC3143130A90FB608D0B0284" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[388,878,1194,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130A90FB608FF90284" box="[388,412,1195,1221]" italics="true" pageId="104" pageNumber="105">A.</emphasis>
(
<emphasis id="DBF6EAF8AC3143130ABFFB608C280284" box="[427,589,1195,1221]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
)
<emphasis id="DBF6EAF8AC3143130949FB608CA40284" box="[605,705,1195,1221]" italics="true" pageId="104" pageNumber="105">schwarzi</emphasis>
Warncke, 1975
</taxonomicName>
(
<collectingCountry id="9195767AAC3143130868FB608DBE0284" box="[892,987,1195,1221]" name="Ukraine" pageId="104" pageNumber="105">Ukraine</collectingCountry>
, European
<collectingCountry id="9195767AAC3143130F43FB608AC00284" box="[1111,1189,1195,1221]" name="Russia" pageId="104" pageNumber="105">Russia</collectingCountry>
,
<collectingCountry id="9195767AAC3143130FBAFB608A9A0284" box="[1198,1279,1195,1221]" name="Turkey" pageId="104" pageNumber="105">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC3143130E1DFB608BF50284" box="[1289,1424,1195,1221]" name="Kazakhstan" pageId="104" pageNumber="105">Kazakhstan</collectingCountry>
), though it should be noted that
<taxonomicName id="2E824D69AC3143130AE2FB048C1E02A8" authorityName="Warncke" authorityYear="1975" box="[502,635,1231,1257]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC3143130AE2FB048C1E02A8" box="[502,635,1231,1257]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
has not been confirmed to fall within the
<taxonomicName id="2E824D69AC3143130F4AFB048B6502A8" authorityName="Warncke" authorityYear="1968" box="[1118,1280,1231,1257]" pageId="104" pageNumber="139" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130F4AFB048B6502A8" box="[1118,1280,1231,1257]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
</taxonomicName>
due to a lack of genetic data. Due to the scutum of
<taxonomicName id="2E824D69AC3143130940FB388CBF034C" authorityName="WOOD" authorityYear="2024" box="[596,730,1267,1293]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="balticola">
<emphasis id="DBF6EAF8AC3143130940FB388CBF034C" box="[596,730,1267,1293]" italics="true" pageId="104" pageNumber="105">A. balticola</emphasis>
</taxonomicName>
which has weak metallic greenish-copper reflections (
<figureCitation id="71B92A6FAC3143130E45FB388EAD0370" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="104" pageNumber="105">Figure 65E</figureCitation>
), it can be compared to
<taxonomicName id="2E824D69AC3143130ACAFADC8C1A0370" authorityName="Wood" authorityYear="2023" box="[478,639,1303,1329]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC3143130ACAFADC8C1A0370" box="[478,639,1303,1329]" italics="true" pageId="104" pageNumber="105">A. orichalcum</emphasis>
</taxonomicName>
which is tentatively but not definitively associated with the subgenus
<taxonomicName id="2E824D69AC3143130B83FAF08F5C0314" authorityName="Warncke" authorityYear="1968" box="[151,313,1339,1365]" pageId="104" pageNumber="105" rank="subGenus" subGenus="Graecandrena">
<emphasis id="DBF6EAF8AC3143130B83FAF08F5C0314" box="[151,313,1339,1365]" italics="true" pageId="104" pageNumber="105">Graecandrena</emphasis>
</taxonomicName>
.
<taxonomicName id="2E824D69AC3143130A53FAF08C750314" authorityName="WOOD" authorityYear="2024" box="[327,528,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="balticola">
<emphasis id="DBF6EAF8AC3143130A53FAF08C750314" box="[327,528,1339,1365]" italics="true" pageId="104" pageNumber="105">Andrena balticola</emphasis>
</taxonomicName>
can be separated by the sparse and obscure scutal punctation, punctures separated by 2 puncture diameters, individual punctures difficult to discern against background sculpture (in
<taxonomicName id="2E824D69AC3143130FE8FA948BF80338" authorityName="Wood" authorityYear="2023" box="[1276,1437,1375,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC3143130FE8FA948BF80338" box="[1276,1437,1375,1401]" italics="true" pageId="104" pageNumber="105">A. orichalcum</emphasis>
</taxonomicName>
scutum with punctures separated by 12 puncture diameters, individual punctures clear and easily visible against underlying surface), facial foveae relatively broad, occupying ½ space between compound eye and lateral ocellus (
<figureCitation id="71B92A6FAC3143130B8BFA008F4203A4" box="[159,295,1482,1509]" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="104" pageNumber="105">Figure 65D</figureCitation>
), not noticeably narrowed ventrally (in
<taxonomicName id="2E824D69AC31431309E2FA008DFC03A4" authorityName="Wood" authorityYear="2023" box="[758,921,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC31431309E2FA008DFC03A4" box="[758,921,1483,1509]" italics="true" pageId="104" pageNumber="105">A. orichalcum</emphasis>
</taxonomicName>
with foveae occupying ⅓ to ¼ of the space between compound eye and lateral ocellus, narrowed further ventrally), clypeus with sparse, shallow, and obscure punctation, punctures separated by 13 puncture diameters (in
<taxonomicName id="2E824D69AC314313085BF9D88D88006C" authorityName="Wood" authorityYear="2023" box="[847,1005,1555,1581]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC314313085BF9D88D88006C" box="[847,1005,1555,1581]" italics="true" pageId="104" pageNumber="105">A. orichalcum</emphasis>
</taxonomicName>
with clypeus more densely and clearly punctate, punctures separated by 0.52 puncture diameters), and terga with narrow apical hairbands on T24 (
<figureCitation id="71B92A6FAC3143130E45F9FC8EAE0034" captionStart="FIGURE 65" captionStartId="103.[151,250,1943,1967]" captionTargetBox="[215,1373,181,1919]" captionTargetId="figure-16@103.[215,1373,181,1919]" captionTargetPageId="103" captionText="FIGURE 65. Andrena (Graecandrena) balticola spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus detail, frontal view; D. Head, dorsal view; E. Scutum, dorsal view; F. Propodeum, dorsal view; G. Forewing, dorsal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209447" httpUri="https://zenodo.org/record/13209447/files/figure.png" pageId="104" pageNumber="105">Figure 65H</figureCitation>
), hairbands extending only slightly beyond the apical margin of the marginal areas (in
<taxonomicName id="2E824D69AC3143130F9AF9908B490034" authorityName="Wood" authorityYear="2023" box="[1166,1324,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC3143130F9AF9908B490034" box="[1166,1324,1627,1653]" italics="true" pageId="104" pageNumber="105">A. orichalcum</emphasis>
</taxonomicName>
with long hairbands on T24, hairbands clearly extending far beyond the apical margins of the marginal areas).
</paragraph>
<paragraph id="E93D36EAAC3143130BD3F9688C8301D4" blockId="104.[151,1437,151,2013]" pageId="104" pageNumber="105">
Separation from
<taxonomicName id="2E824D69AC3143130A9DF9688C6A00FC" authorityName="Warncke" authorityYear="1975" box="[393,527,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC3143130A9DF9688C6A00FC" box="[393,527,1699,1725]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
can be made using similar characters, specifically
<taxonomicName id="2E824D69AC3143130F74F9688A8300FC" authorityName="WOOD" authorityYear="2024" box="[1120,1254,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="balticola">
<emphasis id="DBF6EAF8AC3143130F74F9688A8300FC" box="[1120,1254,1699,1725]" italics="true" pageId="104" pageNumber="105">A. balticola</emphasis>
</taxonomicName>
has the scutum with weak metallic greenish-copper reflections (completely black in
<taxonomicName id="2E824D69AC314313088DF90C8A7900A0" authorityName="Warncke" authorityYear="1975" box="[921,1052,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC314313088DF90C8A7900A0" box="[921,1052,1735,1761]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
), weak scutal punctation blending into clear scutal sculpture, scutum weakly shining (in
<taxonomicName id="2E824D69AC31431309ECF9208D1E0144" authorityName="Warncke" authorityYear="1975" box="[760,891,1771,1797]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC31431309ECF9208D1E0144" box="[760,891,1771,1797]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
with the scutum polished and shining with deep distinct punctures), the clypeus is shagreened and weakly shining, with obscure shallow punctation (in
<taxonomicName id="2E824D69AC3143130E0EF8C48BF80168" authorityName="Warncke" authorityYear="1975" box="[1306,1437,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC3143130E0EF8C48BF80168" box="[1306,1437,1807,1833]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
with the clypeus polished and shining, with deep distinct punctures), and the tergal hairbands are short and barely surpass the apical margin of the marginal areas (in
<taxonomicName id="2E824D69AC31431309C5F89C8D300130" authorityName="Warncke" authorityYear="1975" box="[721,853,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="schwarzi">
<emphasis id="DBF6EAF8AC31431309C5F89C8D300130" box="[721,853,1879,1905]" italics="true" pageId="104" pageNumber="105">A. schwarzi</emphasis>
</taxonomicName>
with the apical hairbands of T24 clearly extending far beyond the apical margins of the marginal areas).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3143120BD3F8548E850790" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105" type="discussion">
<paragraph id="E93D36EAAC3143120BD3F8548E850790" blockId="104.[151,1437,151,2013]" lastBlockId="105.[151,1437,151,647]" lastPageId="105" lastPageNumber="106" pageId="104" pageNumber="105">
<emphasis id="DBF6EAF8AC3143130BD3F8548F5E01F8" bold="true" box="[199,315,1951,1977]" pageId="104" pageNumber="105">Remarks.</emphasis>
Based on the distribution, it could be expected that
<taxonomicName id="2E824D69AC31431308A7F8548A5901F8" authorityName="WOOD" authorityYear="2024" box="[947,1084,1951,1977]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="balticola">
<emphasis id="DBF6EAF8AC31431308A7F8548A5901F8" box="[947,1084,1951,1977]" italics="true" pageId="104" pageNumber="105">A. balticola</emphasis>
</taxonomicName>
might be conspecific with
<taxonomicName id="2E824D69AC3143130E90F8548C00019C" authority="Osytshnjuk, 1994" authorityName="Osytshnjuk" authorityYear="1994" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="volgensis" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC3143130E90F8548BF801F8" box="[1412,1437,1951,1977]" italics="true" pageId="104" pageNumber="105">A.</emphasis>
(
<emphasis id="DBF6EAF8AC3143130BB5F8088F46019C" box="[161,291,1987,2013]" italics="true" pageId="104" pageNumber="105">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC3143130A26F8088FFF019C" box="[306,410,1987,2013]" italics="true" pageId="104" pageNumber="105">volgensis</emphasis>
Osytshnjuk, 1994
</taxonomicName>
(southern European
<collectingCountry id="9195767AAC3143130859F8088DFD019C" box="[845,920,1987,2013]" name="Russia" pageId="104" pageNumber="105">Russia</collectingCountry>
and
<collectingCountry id="9195767AAC31431308D9F8088A37019C" box="[973,1106,1987,2013]" name="Kazakhstan" pageId="104" pageNumber="105">Kazakhstan</collectingCountry>
) or
<taxonomicName id="2E824D69AC3143120F69F8088F2706F0" authority="Warncke, 1972" authorityName="Warncke" authorityYear="1972" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" lastPageId="105" lastPageNumber="106" order="Hymenoptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="chersona" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC3143130F69F8088AF0019C" box="[1149,1173,1987,2013]" italics="true" pageId="104" pageNumber="105">A.</emphasis>
(
<emphasis id="DBF6EAF8AC3143130FB0F8088B43019C" box="[1188,1318,1987,2013]" italics="true" pageId="104" pageNumber="105">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC3143130E22F8088BF8019C" box="[1334,1437,1987,2013]" italics="true" pageId="104" pageNumber="105">chersona</emphasis>
Warncke, 1972
</taxonomicName>
(
<collectingCountry id="9195767AAC3043120A45FF5C8FC806F0" box="[337,429,151,177]" name="Ukraine" pageId="105" pageNumber="106">Ukraine</collectingCountry>
to
<collectingCountry id="9195767AAC3043120AC0FF5C8C4006F0" box="[468,549,151,177]" name="Turkey" pageId="105" pageNumber="106">Turkey</collectingCountry>
, the Caucasus, northern
<collectingCountry id="9195767AAC3043120829FF5C8D0806F0" box="[829,877,151,177]" name="Iran" pageId="105" pageNumber="106">Iran</collectingCountry>
, and
<collectingCountry id="9195767AAC30431208BEFF5C8A2206F0" box="[938,1095,151,177]" name="Turkmenistan" pageId="105" pageNumber="106">Turkmenistan</collectingCountry>
) as these species are found in dry open habitats in eastern Europe. However, these are clearly members of the subgenus
<taxonomicName id="2E824D69AC3043120F84FF708B770694" authorityName="Warncke" authorityYear="1968" box="[1168,1298,187,213]" pageId="105" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC3043120F84FF708B770694" box="[1168,1298,187,213]" italics="true" pageId="105" pageNumber="106">Aciandrena</emphasis>
</taxonomicName>
because the propodeal triangle is covered in the typical granular shagreen found in members of this subgenus, and the facial foveae are more clearly narrowed ventrally. It seems remarkable that this taxon has not previously been recognised from European
<collectingCountry id="9195767AAC3043120A5EFEEC8FFD0700" box="[330,408,295,321]" name="Russia" pageId="105" pageNumber="106">Russia</collectingCountry>
. If this was an older specimen, it could be suggested that it had been mislabelled, but this specimen was collected recently during a survey of known providence. New surveys at the
<emphasis id="DBF6EAF8AC3043120F8FFE808B480725" box="[1179,1325,331,357]" italics="true" pageId="105" pageNumber="106">locus typicus</emphasis>
(which is a small park on sandy soil along the Emajõgi river) have unfortunately not revealed any new specimens (V. Soon,
<emphasis id="DBF6EAF8AC3043120B83FE588EB807EC" box="[151,221,403,429]" italics="true" pageId="105" pageNumber="106">in litt.</emphasis>
), but other surveys in sandy habitats across the Baltic region may produce additional material, including males.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3043120BD3FE108C850458" pageId="105" pageNumber="106" type="etymology">
<paragraph id="E93D36EAAC3043120BD3FE108C850458" blockId="105.[151,1437,151,647]" pageId="105" pageNumber="106">
<emphasis id="DBF6EAF8AC3043120BD3FE108F2B07B4" bold="true" box="[199,334,475,501]" pageId="105" pageNumber="106">Etymology.</emphasis>
From the Latin
<emphasis id="DBF6EAF8AC3043120900FE108C1507B4" box="[532,624,475,501]" italics="true" pageId="105" pageNumber="106">Balticus</emphasis>
, the name for the geographic region comprising
<collectingCountry id="9195767AAC3043120FA0FE108B6E07B4" box="[1204,1291,475,501]" name="Estonia" pageId="105" pageNumber="106">Estonia</collectingCountry>
,
<collectingCountry id="9195767AAC3043120E0EFE108B0107B4" box="[1306,1380,475,501]" name="Latvia" pageId="105" pageNumber="106">Latvia</collectingCountry>
, and
<collectingCountry id="9195767AAC3043120B83FE348F670458" box="[151,258,511,537]" name="Lithuania" pageId="105" pageNumber="106">Lithuania</collectingCountry>
with the suffix
<emphasis id="DBF6EAF8AC3043120AA7FE348F880458" box="[435,493,511,537]" italics="true" pageId="105" pageNumber="106">-cola</emphasis>
, meaning “living in”.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC3043120BD3FDE98FD104C6" pageId="105" pageNumber="106" type="distribution">
<paragraph id="E93D36EAAC3043120BD3FDE98AB8047C" blockId="105.[151,1437,151,647]" box="[199,1245,546,573]" pageId="105" pageNumber="106">
<emphasis id="DBF6EAF8AC3043120BD3FDE98F04047D" bold="true" box="[199,353,546,572]" pageId="105" pageNumber="106">Distribution.</emphasis>
<collectingCountry id="9195767AAC3043120A73FDE88FDB047C" box="[359,446,547,573]" name="Estonia" pageId="105" pageNumber="106">Estonia</collectingCountry>
, probably present in at least neighbouring
<collectingCountry id="9195767AAC304312088EFDE88D87047C" box="[922,994,547,573]" name="Latvia" pageId="105" pageNumber="106">Latvia</collectingCountry>
and European
<collectingCountry id="9195767AAC3043120F9FFDE88ABC047C" box="[1163,1241,547,573]" name="Russia" pageId="105" pageNumber="106">Russia</collectingCountry>
.
</paragraph>
<paragraph id="E93D36EAAC3043120BD3FD8D8FD104C6" blockId="105.[151,1437,151,647]" pageId="105" pageNumber="106">
<emphasis id="DBF6EAF8AC3043120BD3FD8D8F9C0421" bold="true" box="[199,505,582,608]" pageId="105" pageNumber="106">Other material examined</emphasis>
(
<taxonomicName id="2E824D69AC304312091FFD8C8C910420" authorityName="Wood" authorityYear="2023" box="[523,756,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="105" pageNumber="106" phylum="Arthropoda" rank="species" species="orichalcum">
<emphasis id="DBF6EAF8AC304312091FFD8C8C910420" box="[523,756,583,609]" italics="true" pageId="105" pageNumber="106">Andrena orichalcum</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC3043120812FD8D8FD504C6" collectingDate="1992-04-19" collectorName="Jabbul-See, SE &amp; K. Warncke" country="Syria" elevation="300" location="Aleppo" pageId="105" pageNumber="106" specimenCount="16" specimenCount-female="16" stateProvince="Halab">
<emphasis id="DBF6EAF8AC3043120812FD8D8D030421" bold="true" box="[774,870,582,608]" pageId="105" pageNumber="106">
<collectingCountry id="9195767AAC3043120812FD8D8D040421" box="[774,865,582,608]" name="Syria" pageId="105" pageNumber="106">SYRIA</collectingCountry>
:
</emphasis>
<collectorName id="4477533CAC304312087BFD8C8A7E0420" box="[879,1051,583,609]" pageId="105" pageNumber="106">Jabbul-See, SE</collectorName>
<collectingRegion id="2B46F808AC3043120F36FD8C8A120420" box="[1058,1143,583,609]" country="Syria" name="Halab" pageId="105" pageNumber="106">Aleppo</collectingRegion>
,
<quantity id="2E7A9B0FAC3043120F91FD8C8AB50420" box="[1157,1232,583,609]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="105" pageNumber="106" unit="m" value="300.0">
<elevation id="62AFD1D9AC3043120F91FD8C8AB50420" box="[1157,1232,583,609]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="105" pageNumber="106" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="9D3C102AAC3043120FCAFD8C8B330420" box="[1246,1366,583,609]" pageId="105" pageNumber="106" value="1992-04-19">
<collectingDate id="8D78E9C2AC3043120FCAFD8C8B330420" box="[1246,1366,583,609]" pageId="105" pageNumber="106" value="1992-04-19">19.iv.1992</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC3043120E70FD8C8BF20420" box="[1380,1431,583,609]" count="16" pageId="105" pageNumber="106" type="female">16♀</specimenCount>
, leg.
<collectorName id="4477533CAC3043120BD3FDA08F2A04C4" box="[199,335,619,645]" pageId="105" pageNumber="106">K. Warncke</collectorName>
, OÖLM
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,417 @@
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<mods:title id="4BB79E626283B3363F349C2A25938732">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
</mods:titleInfo>
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<mods:identifier id="2BC655385DC6294A1CBD09C276D0843B" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="0A003C3D80B697E1390531E5521404A0" type="Zenodo-Dep">13209307</mods:identifier>
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</mods:mods>
<treatment id="612B87FCAC4743590B83F9B58C7007C8" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8" httpUri="http://treatment.plazi.org/id/612B87FCAC4743590B83F9B58C7007C8" lastPageId="34" lastPageNumber="35" pageId="30" pageNumber="31">
<subSubSection id="A1986561AC4743650B83F9B58CCF00D8" box="[151,682,1662,1689]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="E93D36EAAC4743650B83F9B58CCF00D8" blockId="30.[151,682,1662,1725]" box="[151,682,1662,1689]" pageId="30" pageNumber="31">
<heading id="B2758186AC4743650B83F9B58CCF00D8" bold="true" box="[151,682,1662,1689]" fontSize="11" level="2" pageId="30" pageNumber="31" reason="4">
<emphasis id="DBF6EAF8AC4743650B83F9B58CCF00D8" bold="true" box="[151,682,1662,1689]" pageId="30" pageNumber="31">
<taxonomicName id="2E824D69AC4743650B83F9B58C4900D8" authorityName="WOOD" authorityYear="2024" box="[151,556,1662,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="incustodita" status="sp. nov." subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC4743650B83F9B58F6400D8" bold="true" box="[151,257,1662,1689]" italics="true" pageId="30" pageNumber="31">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4743650A06F9B58FF100D8" bold="true" box="[274,404,1662,1689]" italics="true" pageId="30" pageNumber="31">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4743650AB2F9B58C4900D8" bold="true" box="[422,556,1662,1689]" italics="true" pageId="30" pageNumber="31">incustodita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC4743650927F9B48CCF00D8" box="[563,682,1663,1689]" pageId="30" pageNumber="31" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4743650B83F9688FC200FC" box="[151,423,1699,1725]" pageId="30" pageNumber="31" type="description">
<paragraph id="E93D36EAAC4743650B83F9688FC200FC" blockId="30.[151,682,1662,1725]" box="[151,423,1699,1725]" pageId="30" pageNumber="31">
<figureCitation id="71B92A6FAC4743650B83F9688F2D00FC" box="[151,328,1699,1725]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="30" pageNumber="31">Figures 17AD</figureCitation>
;
<figureCitation id="71B92A6FAC4743650A47F9688FC700FC" box="[339,418,1699,1725]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="30" pageNumber="31">18AF</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4743650B83F9218C81019E" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="E93D36EAAC4743650B83F9218F0B016A" blockId="30.[151,1437,1770,2015]" pageId="30" pageNumber="31">
<materialsCitation id="59EA3CB7AC4743650B83F9218F0B016A" collectingDate="1978-03-21" collectorName="Hohmann" country="Greece" location="Hagios Nicolaos" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" stateProvince="Kriti" typeStatus="holotype">
<emphasis id="DBF6EAF8AC4743650B83F9218FAD0145" bold="true" box="[151,456,1770,1796]" pageId="30" pageNumber="31">
<typeStatus id="36398848AC4743650B83F9218F5F0145" box="[151,314,1770,1796]" pageId="30" pageNumber="31" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC4743650A5CF9218FA60145" box="[328,451,1770,1796]" name="Greece" pageId="30" pageNumber="31">GREECE</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC4743650AC5F9208C770144" box="[465,530,1771,1797]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Kreta</collectingRegion>
,
<location id="EC5D6031AC474365090BF9208CB90144" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC474365090BF9208CB90144" box="[543,732,1771,1797]" country="Greece" name="Hagios Nicolaos" pageId="30" pageNumber="31" stateProvince="Kriti">Hagios Nicolaos</location>
[Agios Nikolaos; 35.1919
<superScript id="1EF79BA2AC4743650F1EF9208A4F0144" attach="right" box="[1034,1066,1771,1797]" fontSize="6" pageId="30" pageNumber="31">oN</superScript>
, 25.7067
<superScript id="1EF79BA2AC4743650F86F9208ACA0144" attach="right" box="[1170,1199,1771,1797]" fontSize="6" pageId="30" pageNumber="31">oE</superScript>
],
<date id="9D3C102AAC4743650FD1F9208B250144" box="[1221,1344,1771,1797]" pageId="30" pageNumber="31" value="1978-03-21">
<collectingDate id="8D78E9C2AC4743650FD1F9208B250144" box="[1221,1344,1771,1797]" pageId="30" pageNumber="31" value="1978-03-21">21.iii.1978</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650E5AF9208B000144" box="[1358,1381,1771,1797]" count="1" pageId="30" pageNumber="31" type="male"></specimenCount>
, leg.
<collectorName id="4477533CAC4743650B83F8C48F6D0168" box="[151,264,1807,1833]" pageId="30" pageNumber="31">Hohmann</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC4743650BD3F8F98C81019E" blockId="30.[151,1437,1770,2015]" pageId="30" pageNumber="31">
<materialsCitation id="59EA3CB7AC4743650BD3F8F98A55010C" box="[199,1072,1842,1869]" collectingDate="1996-04-06" collectionCode="TJWC" collectorName="M. Archer" country="Greece" location="Kissamos" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="DBF6EAF8AC4743650BD3F8F98F98010D" bold="true" box="[199,509,1842,1868]" pageId="30" pageNumber="31">
<typeStatus id="36398848AC4743650BD3F8F98F0A010D" box="[199,367,1842,1868]" pageId="30" pageNumber="31" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="9195767AAC4743650A69F8F98F9D010D" box="[381,504,1842,1868]" name="Greece" pageId="30" pageNumber="31">GREECE</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC4743650912F8F88C13010C" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC4743650912F8F88C13010C" box="[518,630,1843,1869]" country="Greece" name="Kissamos" pageId="30" pageNumber="31">Kissamos</location>
,
<date id="9D3C102AAC4743650997F8F88C88010C" box="[643,749,1843,1869]" pageId="30" pageNumber="31" value="1996-04-06">
<collectingDate id="8D78E9C2AC4743650997F8F88C88010C" box="[643,749,1843,1869]" pageId="30" pageNumber="31" value="1996-04-06">6.iv.1996</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC47436509EFF8F88D7A010C" box="[763,799,1843,1869]" count="1" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC4743650874F8F88DB2010C" box="[864,983,1843,1869]" pageId="30" pageNumber="31">M. Archer</collectorName>
,
<collectionCode id="8F93AE2FAC47436508F7F8F88A55010C" box="[995,1072,1843,1869]" pageId="30" pageNumber="31">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4743650F29F8F88CB80132" collectingDate="1978-03-21" collectionCode="OOLM" collectorName="Hohmann" country="Greece" location="Hagios Nicolaos" pageId="30" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Kriti" typeStatus="paratype">
<collectingRegion id="2B46F808AC4743650F29F8F88A1B010C" box="[1085,1150,1843,1869]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Kreta</collectingRegion>
,
<location id="EC5D6031AC4743650F9FF8F88B22010C" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC4743650F9FF8F88B22010C" box="[1163,1351,1843,1869]" country="Greece" name="Hagios Nicolaos" pageId="30" pageNumber="31" stateProvince="Kriti">Hagios Nicolaos</location>
[Agios Nikolaos],
<date id="9D3C102AAC4743650A03F89C8FF70130" box="[279,402,1879,1905]" pageId="30" pageNumber="31" value="1978-03-21">
<collectingDate id="8D78E9C2AC4743650A03F89C8FF70130" box="[279,402,1879,1905]" pageId="30" pageNumber="31" value="1978-03-21">21.iii.1978</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650AB5F89C8FA30130" box="[417,454,1879,1905]" count="2" pageId="30" pageNumber="31" type="male">2♂</specimenCount>
, leg.
<collectorName id="4477533CAC474365091CF89C8C1C0130" box="[520,633,1879,1905]" pageId="30" pageNumber="31">Hohmann</collectorName>
,
<collectionCode id="8F93AE2FAC4743650993F8938CB80132" box="[647,733,1880,1907]" pageId="30" pageNumber="31">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC47436509F8F89C8E8801D6" collectingDate="1973-04-25" collectionCode="OOLM" collectorName="H. Teunissen" country="Greece" location="Knossos" municipality="Iraklion" pageId="30" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Kriti" typeStatus="paratype">
<collectingRegion id="2B46F808AC47436509F8F89C8D490130" box="[748,812,1879,1905]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Creta</collectingRegion>
,
<collectingMunicipality id="0959AC90AC474365082DF89C8DF00130" box="[825,917,1879,1905]" pageId="30" pageNumber="31">Iraklion</collectingMunicipality>
,
<location id="EC5D6031AC47436508B7F89C8A600130" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC47436508B7F89C8A600130" box="[931,1029,1879,1905]" country="Greece" municipality="Iraklion" name="Knossos" pageId="30" pageNumber="31" stateProvince="Kriti">Knossos</location>
,
<date id="9D3C102AAC4743650F07F89D8AE90130" box="[1043,1164,1878,1905]" pageId="30" pageNumber="31" value="1973-04-25">
<collectingDate id="8D78E9C2AC4743650F07F89D8AE90130" box="[1043,1164,1878,1905]" pageId="30" pageNumber="31" value="1973-04-25">25.iv.1973</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650F8EF89C8ADA0130" box="[1178,1215,1879,1905]" count="2" pageId="30" pageNumber="31" type="male">2♂</specimenCount>
, leg.
<collectorName id="4477533CAC4743650E15F89C8BFC0130" box="[1281,1433,1879,1905]" pageId="30" pageNumber="31">H. Teunissen</collectorName>
,
<collectionCode id="8F93AE2FAC4743650B83F8B78E8801D6" box="[151,237,1916,1943]" pageId="30" pageNumber="31">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4743650BE2F8B08D3C01D6" box="[246,857,1915,1943]" collectingDate="1979-04-17" collectionCode="OOLM" collectorName="Hohmann" country="Greece" location="Mochlos" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="paratype">
<collectingRegion id="2B46F808AC4743650BE2F8B08F5201D4" box="[246,311,1915,1941]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Kreta</collectingRegion>
,
<location id="EC5D6031AC4743650A2BF8B08FC101D4" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC4743650A2BF8B08FC101D4" box="[319,420,1915,1941]" country="Greece" name="Mochlos" pageId="30" pageNumber="31" stateProvince="Kriti">Mochlos</location>
,
<date id="9D3C102AAC4743650AB8F8B08C4001D4" box="[428,549,1915,1941]" pageId="30" pageNumber="31" value="1979-04-17">
<collectingDate id="8D78E9C2AC4743650AB8F8B08C4001D4" box="[428,549,1915,1941]" pageId="30" pageNumber="31" value="1979-04-17">17.iv.1979</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC474365093AF8B08C3401D4" box="[558,593,1915,1941]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC474365099DF8B08C9F01D4" box="[649,762,1915,1941]" pageId="30" pageNumber="31">Hohmann</collectorName>
,
<collectionCode id="8F93AE2FAC4743650817F8B78D3C01D6" box="[771,857,1916,1943]" pageId="30" pageNumber="31">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4743650876F8B08FBE01FA" collectingDate="1976-06-14" collectionCode="OOLM" collectorName="K. Warncke" country="Greece" elevation="2100" location="Ida Geb." pageId="30" pageNumber="31" specimenCount="2" specimenCount-female="2" stateProvince="Kriti" typeStatus="paratype">
<collectingRegion id="2B46F808AC4743650876F8B08DC601D4" box="[866,931,1915,1941]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Kreta</collectingRegion>
,
<location id="EC5D6031AC47436508BFF8B08A6C01D4" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC47436508BFF8B08A6C01D4" box="[939,1033,1915,1941]" country="Greece" name="Ida Geb." pageId="30" pageNumber="31" stateProvince="Kriti">Ida Geb.</location>
[Ida Mountain],
<quantity id="2E7A9B0FAC4743650FD4F8B08B7101D4" box="[1216,1300,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="30" pageNumber="31" unit="m" value="2100.0">
<elevation id="62AFD1D9AC4743650FD4F8B08B7101D4" box="[1216,1300,1915,1941]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="30" pageNumber="31" unit="m" value="2100.0">2100 m</elevation>
</quantity>
,
<date id="9D3C102AAC4743650E09F8B08BF201D4" box="[1309,1431,1915,1941]" pageId="30" pageNumber="31" value="1976-06-14">
<collectingDate id="8D78E9C2AC4743650E09F8B08BF201D4" box="[1309,1431,1915,1941]" pageId="30" pageNumber="31" value="1976-06-14">14.vi.1976</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650B83F8548ED901F8" box="[151,188,1951,1977]" count="2" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC4743650BE2F8548F1901F8" box="[246,380,1951,1977]" pageId="30" pageNumber="31">K. Warncke</collectorName>
,
<collectionCode id="8F93AE2FAC4743650A91F86B8FBE01FA" box="[389,475,1952,1979]" pageId="30" pageNumber="31">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4743650AF1F8548B4701FA" box="[485,1314,1950,1979]" collectingDate="1963-05-15" collectionCode="OOLM" collectorName="M. Schwarz &amp; J. Gusenleitner" country="Greece" location="Knossos" pageId="30" pageNumber="31" specimenCount="2" specimenCount-male="2" stateProvince="Kriti" typeStatus="paratype">
<collectingRegion id="2B46F808AC4743650AF1F8548C4001F8" box="[485,549,1951,1977]" country="Greece" name="Kriti" pageId="30" pageNumber="31">Creta</collectingRegion>
,
<location id="EC5D6031AC474365093BF8548CF401F8" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC474365093BF8548CF401F8" box="[559,657,1951,1977]" country="Greece" name="Knossos" pageId="30" pageNumber="31" stateProvince="Kriti">Knossos</location>
,
<date id="9D3C102AAC474365098EF8558D6E01F8" box="[666,779,1950,1977]" pageId="30" pageNumber="31" value="1963-05-15">
<collectingDate id="8D78E9C2AC474365098EF8558D6E01F8" box="[666,779,1950,1977]" pageId="30" pageNumber="31" value="1963-05-15">15.v.1963</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650801F8548D5F01F8" box="[789,826,1951,1977]" count="2" pageId="30" pageNumber="31" type="male">2♂</specimenCount>
, leg.
<collectorName id="4477533CAC4743650867F8548D9C01F8" box="[883,1017,1951,1977]" pageId="30" pageNumber="31">M. Schwarz</collectorName>
&amp;
<collectorName id="4477533CAC4743650F0EF8548AA601F8" box="[1050,1219,1951,1977]" pageId="30" pageNumber="31">J. Gusenleitner</collectorName>
,
<collectionCode id="8F93AE2FAC4743650FD8F86B8B4701FA" box="[1228,1314,1952,1979]" pageId="30" pageNumber="31">OÖLM</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4743650E38F8548C1A019C" collectingDate="1963-05-22" collectorName="J. Gusenleitner" country="Greece" location="Heraklion - Strand" pageId="30" pageNumber="31" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<location id="EC5D6031AC4743650E38F8548E88019C" LSID="urn:lsid:plazi:treatment:612B87FCAC4743590B83F9B58C7007C8:EC5D6031AC4743650E38F8548E88019C" country="Greece" name="Heraklion - Strand" pageId="30" pageNumber="31">Heraklion -Strand</location>
,
<date id="9D3C102AAC4743650BECF8088F0C019C" box="[248,361,1987,2013]" pageId="30" pageNumber="31" value="1963-05-22">
<collectingDate id="8D78E9C2AC4743650BECF8088F0C019C" box="[248,361,1987,2013]" pageId="30" pageNumber="31" value="1963-05-22">22.v.1963</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4743650A60F8088FF2019C" box="[372,407,1987,2013]" count="1" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4743650AC1F8088C1A019C" box="[469,639,1987,2013]" pageId="30" pageNumber="31">J. Gusenleitner</collectorName>
</materialsCitation>
, OÖLM.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4643640BD3FF5D8F6201B2" pageId="31" pageNumber="32" type="description">
<paragraph id="E93D36EAAC4643640BD3FF5D8A43078E" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3FF5D8FA506F0" bold="true" box="[199,448,150,177]" pageId="31" pageNumber="32">Description: Female:</emphasis>
Body length:
<quantity id="2E7A9B0FAC464364094CFF5C8CD706F0" box="[600,690,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="31" pageNumber="32" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
(
<figureCitation id="71B92A6FAC46436409AAFF5C8D2706F0" box="[702,834,151,177]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="31" pageNumber="32">Figure 17A</figureCitation>
).
<emphasis id="DBF6EAF8AC4643640846FF5C8DFD06F0" box="[850,920,151,177]" italics="true" pageId="31" pageNumber="32">Head:</emphasis>
Dark, 1.35 times wider than long (
<figureCitation id="71B92A6FAC4643640E1AFF5C8BEA06F0" box="[1294,1423,151,177]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="31" pageNumber="32">Figure 17B</figureCitation>
). Clypeus domed, densely but obscurely punctate, punctures separated by 0.51 puncture diameters, disappearing into underlying surface which is shagreened to microreticulate, weakly shining to dull. Process of labrum rounded trapezoidal, 2 times wider than long, anterior margin widely but shallowly emarginate. Gena slightly exceeding width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Foveae moderately broad, occupying slightly &lt;½ space between compound eye and lateral ocellus, slightly constricted medially, ventrally extending to lower margin of antennal insertions; foveae filled with light brown hairs. Face, gena, vertex, and scape with moderately long whitish to light brownish hairs, none equalling length of scape. Antennae basally dark, A612 ventrally lightened by presence of dark brown-orange scales; A3 equalling A4+5.
</paragraph>
<paragraph id="E93D36EAAC4643640BD3FE128F6A053A" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3FE128F2D07B3" box="[199,328,473,498]" italics="true" pageId="31" pageNumber="32">Mesosoma:</emphasis>
Scutum and scutellum very densely punctate, punctures separated by &lt;0.5 puncture diameters, almost confluent, interspaces polished and shining (
<figureCitation id="71B92A6FAC46436409E0FE378D190457" box="[756,892,508,534]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="31" pageNumber="32">Figure 17C</figureCitation>
). Pronotum rounded. Mesepisternum with fine granular microreticulation, weakly shining. Dorsolateral parts of propodeum microreticulate, dull, overlain by weak network of raised rugosity; propodeal triangle laterally delineated by clear carinae, broadly triangular, internal surface with fine reticulation, weakly shining, covered with sparse raised rugae in basal half. Mesepisternum with pale hairs, medially and ventrally whitish, becoming light brown dorsally. Scutum and scutellum with short light brown hairs, almost sub-squamous. Propodeal corbicula incomplete, dorsal fringe composed of plumose light brown hairs, internal surface with sparse light brown simple hairs. Legs brown, pubescence light brown to orange-brown. Inner hind tibial spur broadened submedially. Flocculus moderately developed, composed of white plumose hairs, femoral and tibial scopae composed of pale predominantly simple hairs, some tibial hairs weakly plumose apically. Hind tarsal claws without inner tooth. Wings hyaline, stigma dark brown, venation orange-brown, nervulus slightly antefurcal.
</paragraph>
<paragraph id="E93D36EAAC4643640BD3FC4D8DFA0234" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3FC4D8F2005DE" box="[199,325,902,927]" italics="true" pageId="31" pageNumber="32">Metasoma:</emphasis>
Tergal discs dark, marginal areas broadly lightened hyaline white-yellow-brown (
<figureCitation id="71B92A6FAC4643640E12FC4E8BF505DE" box="[1286,1424,901,927]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="31" pageNumber="32">Figure 17D</figureCitation>
). Tergal discs regularly but weakly and obscurely punctate, punctures separated by 12 puncture diameters, punctation strongest on T1, weakest on T4; underlying surface shagreened, weakly shining, punctation somewhat disappearing into underlying sculpture. Tergal margins with punctation dense but weaker than on discs, obscure. Tergal discs with short scattered hairs, T24 with light brown apical hairbands, interrupted on T23, complete on T4, obscuring underlying surface in fresh individuals. Apical fringe of T5 and hairs flanking pygidial plate golden orange. Pygidial plate rounded triangular, surface domed, impunctate, weakly shining.
</paragraph>
<paragraph id="E93D36EAAC4643640BD3FBB48FA10331" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3FBB48F6802D8" bold="true" box="[199,269,1151,1177]" pageId="31" pageNumber="32">Male:</emphasis>
Body length:
<quantity id="2E7A9B0FAC4643640AA3FBB48C7D02DB" box="[439,536,1151,1178]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="31" pageNumber="32" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
(
<figureCitation id="71B92A6FAC464364093DFBB48CD602D8" box="[553,691,1151,1177]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="31" pageNumber="32">Figure 18A</figureCitation>
).
<emphasis id="DBF6EAF8AC46436409DEFBB48D7502D8" box="[714,784,1151,1177]" italics="true" pageId="31" pageNumber="32">Head:</emphasis>
Dark, 1.5 times wider than long (
<figureCitation id="71B92A6FAC4643640FB5FBB48B4D02D8" box="[1185,1320,1151,1177]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="31" pageNumber="32">Figure 18B</figureCitation>
). Clypeus weakly domed, densely punctate, punctures separated by ≤0.5 puncture diameters, interspaces polished and shining. Process of labrum rounded trapezoidal, 2 times wider than long, anterior margin widely but shallowly emarginate. Gena equalling width of compound eye; ocelloccipital distance 1.7 times diameter of lateral ocellus (
<figureCitation id="71B92A6FAC4643640E1FFB218BEA0344" box="[1291,1423,1258,1285]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="31" pageNumber="32">Figure 18C</figureCitation>
). Head with long pale hairs, ventrally white, becoming light brownish dorsally, hairs exceeding length of scape. Antennae basally dark, A513 ventrally lightened by presence of dark orange-brown scales; A3 exceeding A4, slightly shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAAC4643640BD3FAB18C6200CC" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3FAB18F2D03D2" box="[199,328,1402,1427]" italics="true" pageId="31" pageNumber="32">Mesosoma:</emphasis>
Scutum and scutellum densely but irregularly punctate, punctures separated by 0.51 puncture diameters, underlying surface predominantly polished and shining, laterally with weak shagreen. Pronotum rounded. Mesepisternum and dorsolateral parts of propodeum with dense granular microreticulation, overlain by weak network of raised rugosity, dull. Propodeal triangle laterally defined by fine carinae, internal surface with regular reticulation, basally with weakly raised rugae, surface weakly shining. Mesosoma with long pale hairs, ventrally whitish, becoming light brown dorsally, hairs exceeding length of scape. Legs black to dark brown, pubescence light brown. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation dark brown, nervulus interstitial to strongly antefurcal.
</paragraph>
<paragraph id="E93D36EAAC4643640BD3F9538F6201B2" blockId="31.[151,1437,150,2035]" pageId="31" pageNumber="32">
<emphasis id="DBF6EAF8AC4643640BD3F9538F2000F0" box="[199,325,1688,1713]" italics="true" pageId="31" pageNumber="32">Metasoma:</emphasis>
Tergal discs dark, marginal areas broadly lightened hyaline white-yellow-brown (
<figureCitation id="71B92A6FAC4643640E12F95C8BEA00F0" box="[1286,1423,1687,1713]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="31" pageNumber="32">Figure 18D</figureCitation>
). Tergal discs finely and obscurely punctate, punctures separated by 23 puncture diameters, underlying surface shagreened and weakly shining, sculpture strongest basally, becoming weakly shining on marginal areas. Tergal discs with scattered short pale hairs, not forming hairbands. T67 with long whitish hairs overlying pygidial plate of T7. S8 apically evenly broadened, ventrally with short light brown hairs, apical process truncate. Genital capsule complex, gonocoxae with inner margins widely separated, apically weakly diverging, produced into moderately long projecting apical teeth (
<figureCitation id="71B92A6FAC4643640AFCF8A58CFD01C9" box="[488,664,1902,1928]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="31" pageNumber="32">Figures 18EF</figureCitation>
). Gonostyli in dorsal view with outer margins curved, converging apically and produced into short spatulate flattened apexes, internal margin strongly raised, apexes with short pale hairs on dorsal surface. Penis valves conspicuously broadened, occupying entire space between gonocoxae and gonostyli.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC79435B0B83F9A68AA6008C" ID-DOI="http://doi.org/10.5281/zenodo.13209345" ID-Zenodo-Dep="13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1645,1669]" targetBox="[151,1436,181,1621]" targetPageId="32" targetType="figure">
<paragraph id="E93D36EAAC79435B0B83F9A68AA6008C" blockId="32.[151,1436,1645,1742]" pageId="32" pageNumber="33">
<emphasis id="DBF6EAF8AC79435B0B83F9A68F4400C4" bold="true" box="[151,289,1645,1669]" pageId="32" pageNumber="33">FIGURE 17.</emphasis>
<taxonomicName id="2E824D69AC79435B0A3CF9A68C1E00C4" authorityName="WOOD" authorityYear="2024" box="[296,635,1645,1670]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="incustodita" status="sp. nov." subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC79435B0A3CF9A68F1B00C4" box="[296,382,1645,1669]" italics="true" pageId="32" pageNumber="33">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC79435B0A9AF9A68F9E00C4" box="[398,507,1645,1669]" italics="true" pageId="32" pageNumber="33">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC79435B091EF9A68C1E00C4" box="[522,635,1645,1669]" italics="true" pageId="32" pageNumber="33">incustodita</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC79435B0996F9A58C8F00C7" bold="true" box="[642,746,1646,1670]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="C0C55783AC79435B0996F9A58C8F00C7" box="[642,746,1646,1670]" pageId="32" pageNumber="33" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view;
<taxonomicName id="2E824D69AC79435B0912F95A8DBE00EB" authority="Dours, 1873" authorityName="Dours" authorityYear="1873" box="[518,987,1681,1706]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" sensu="stricto" species="abbreviata" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC79435B0912F95A8C3900E8" box="[518,604,1681,1705]" italics="true" pageId="32" pageNumber="33">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC79435B0978F95A8CBC00E8" box="[620,729,1681,1705]" italics="true" pageId="32" pageNumber="33">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC79435B09FDF95A8D3300E8" box="[745,854,1681,1705]" italics="true" pageId="32" pageNumber="33">abbreviata</emphasis>
<bibRefCitation id="8D134B1BAC79435B0849F9598DBE00EB" author="Dours, J. A." box="[861,987,1682,1706]" pageId="32" pageNumber="33" pagination="274 - 325" refId="ref88485" refString="Dours, J. A. (1873) Hymenopteres du bassin mediterraneen Andrena (suite). Biareolina, Eucera. Revue et magasin de zoologie pure et appliquee, 1, 274 - 325." type="journal article" year="1873">Dours, 1873</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC79435B08F6F9598A0700E8" box="[994,1122,1682,1705]" italics="true" pageId="32" pageNumber="33">
<taxonomicNameLabel id="C0C55783AC79435B08F6F9598A0700E8" box="[994,1122,1682,1705]" pageId="32" pageNumber="33" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
female. E. Terga, dorsal view;
<taxonomicName id="2E824D69AC79435B0B83F97E8D72008C" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" box="[151,791,1717,1742]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="abbreviata" subGenus="Ulandrena" subSpecies="abnormis">
<emphasis id="DBF6EAF8AC79435B0B83F97E8E88008C" box="[151,237,1717,1741]" italics="true" pageId="32" pageNumber="33">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC79435B0BE8F97E8F0C008C" box="[252,361,1717,1741]" italics="true" pageId="32" pageNumber="33">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC79435B0A6CF97E8F80008C" box="[376,485,1717,1741]" italics="true" pageId="32" pageNumber="33">abbreviata</emphasis>
ssp.
<emphasis id="DBF6EAF8AC79435B090CF97E8C1D008C" box="[536,632,1717,1741]" italics="true" pageId="32" pageNumber="33">abnormis</emphasis>
<bibRefCitation id="8D134B1BAC79435B096AF97E8D72008C" author="Warncke, K." box="[638,791,1717,1741]" pageId="32" pageNumber="33" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
female (dark form). F. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC7943590BD3F9308C32075C" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="diagnosis">
<paragraph id="E93D36EAAC79435A0BD3F9308C0A06B8" blockId="32.[151,1437,1787,2030]" lastBlockId="33.[151,1437,151,249]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="DBF6EAF8AC79435B0BD3F9308F270154" bold="true" box="[199,322,1787,1813]" pageId="32" pageNumber="33">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC79435B0A53F9378C4C0157" authorityName="WOOD" authorityYear="2024" box="[327,553,1788,1814]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="incustodita">
<emphasis id="DBF6EAF8AC79435B0A53F9378C4C0157" box="[327,553,1788,1814]" italics="true" pageId="32" pageNumber="33">Andrena incustodita</emphasis>
</taxonomicName>
can be recognised as an
<taxonomicName id="2E824D69AC79435B082DF9378DD40157" authorityName="Warncke" authorityYear="1968" box="[825,945,1788,1814]" pageId="32" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC79435B082DF9378DD40157" box="[825,945,1788,1814]" italics="true" pageId="32" pageNumber="33">Ulandrena</emphasis>
</taxonomicName>
in the female sex due to the inner hind tibial spur which is broadened submedially (combined with the absence of a latitudinal carina on the posterior face of the hind femur) and in the male sex due to the complex genital capsule with conspicuously broadened penis valves and hyaline-lightened tergal margins. Females can be recognised as part of the
<taxonomicName id="2E824D69AC79435B08C4F8A38A2D01C3" authorityName="Dours" authorityYear="1873" box="[976,1096,1896,1922]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="DBF6EAF8AC79435B08C4F8A38A2D01C3" box="[976,1096,1896,1922]" italics="true" pageId="32" pageNumber="33">abbreviata</emphasis>
</taxonomicName>
-group due to its moderate size (
<quantity id="2E7A9B0FAC79435B0BB4F8478F6501E7" box="[160,256,1932,1958]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="32" pageNumber="33" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
) combined with the non-metallic integument and the extremely densely punctate scutum with punctures separated by &lt;0.5 puncture diameters, punctures almost confluent (
<figureCitation id="71B92A6FAC79435B0897F87B8A62018B" box="[899,1031,1968,1994]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="32" pageNumber="33">Figure 17C</figureCitation>
). It can be separated from other taxa within a broad interpretation of
<taxonomicName id="2E824D69AC79435B091CF81F8CC701AF" authorityName="Dours" authorityYear="1873" box="[520,674,2004,2030]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="DBF6EAF8AC79435B091CF81F8CC701AF" box="[520,674,2004,2030]" italics="true" pageId="32" pageNumber="33">A. abbreviata</emphasis>
</taxonomicName>
due to the tergal punctation which is very weak, tergal discs with fine shagreen, only weakly shining, punctures almost disappearing into the underlying shagreen (
<figureCitation id="71B92A6FAC78435A0FC2FF5C8B3906F0" box="[1238,1372,151,177]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="33" pageNumber="34">Figure 17D</figureCitation>
; in
<emphasis id="DBF6EAF8AC78435A0E90FF5C8FE90694" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="2E824D69AC78435A0E90FF5C8F6A0694" authorityName="Dours" authorityYear="1873" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="abbreviata">A. abbreviata</taxonomicName>
<taxonomicNameLabel id="C0C55783AC78435A0A0DFF708FE90694" box="[281,396,187,213]" pageId="33" pageNumber="34" sensu="lato">sensu lato</taxonomicNameLabel>
</emphasis>
with terga discs smooth and shining, deeply punctate, punctures clearly visible against the underlying polished disc,
<figureCitation id="71B92A6FAC78435A0AADFF148C0606B8" box="[441,611,223,249]" captionStart="FIGURE 17" captionStartId="32.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-28@32.[151,1436,181,1621]" captionTargetPageId="32" captionText="FIGURE 17. Andrena (Ulandrena) incustodita spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Terga, dorsal view; Andrena (Ulandrena) abbreviata Dours, 1873 sensu stricto female. E. Terga, dorsal view; Andrena (Ulandrena) abbreviata ssp. abnormis Warncke, 1967 female (dark form). F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209345" httpUri="https://zenodo.org/record/13209345/files/figure.png" pageId="33" pageNumber="34">Figures 17EF</figureCitation>
).
</paragraph>
<caption id="BDFD6662AC78435A0B83F9018AED0147" ID-DOI="http://doi.org/10.5281/zenodo.13209347" ID-Zenodo-Dep="13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,250,1738,1762]" targetBox="[151,1436,273,1715]" targetPageId="33" targetType="figure">
<paragraph id="E93D36EAAC78435A0B83F9018AED0147" blockId="33.[151,1437,1738,1798]" pageId="33" pageNumber="34">
<emphasis id="DBF6EAF8AC78435A0B83F9018F4700A3" bold="true" box="[151,290,1738,1762]" pageId="33" pageNumber="34">FIGURE 18.</emphasis>
<taxonomicName id="2E824D69AC78435A0A3FF9018CE700A3" authorityName="WOOD" authorityYear="2024" box="[299,642,1738,1762]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="incustodita" status="sp. nov." subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC78435A0A3FF9018FE400A3" box="[299,385,1738,1762]" italics="true" pageId="33" pageNumber="34">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC78435A0A87F9018C6500A3" box="[403,512,1738,1762]" italics="true" pageId="33" pageNumber="34">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC78435A0905F9018CE700A3" box="[529,642,1738,1762]" italics="true" pageId="33" pageNumber="34">incustodita</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC78435A099FF9018C9000A3" bold="true" box="[651,757,1738,1762]" pageId="33" pageNumber="34">
<taxonomicNameLabel id="C0C55783AC78435A099FF9018C9000A3" box="[651,757,1738,1762]" pageId="33" pageNumber="34" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC7843590BD3F8FF8C32075C" blockId="33.[151,1437,1844,2015]" lastBlockId="34.[151,1436,151,393]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Males are easier to diagnose. They can be recognised as part of the
<taxonomicName id="2E824D69AC78435A08CEF8FF8A37010F" authorityName="Dours" authorityYear="1873" box="[986,1106,1844,1870]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="DBF6EAF8AC78435A08CEF8FF8A37010F" box="[986,1106,1844,1870]" italics="true" pageId="33" pageNumber="34">abbreviata</emphasis>
</taxonomicName>
-group due to their moderate size (
<quantity id="2E7A9B0FAC78435A0BC6F8938F570133" box="[210,306,1880,1906]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="33" pageNumber="34" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
) combined with their genital capsule with clear gonocoxal teeth (
<figureCitation id="71B92A6FAC78435A0F05F8938AD80133" box="[1041,1213,1880,1906]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="33" pageNumber="34">Figures 18EF</figureCitation>
). However, the face and lower paraocular areas are black (
<figureCitation id="71B92A6FAC78435A0951F8B78CAC01D7" box="[581,713,1916,1942]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="33" pageNumber="34">Figure 18B</figureCitation>
; marked yellow-white in all other
<taxonomicName id="2E824D69AC78435A0F45F8B78AAC01D7" authorityName="Warncke" authorityYear="1968" box="[1105,1225,1916,1942]" pageId="33" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC78435A0F45F8B78AAC01D7" box="[1105,1225,1916,1942]" italics="true" pageId="33" pageNumber="34">Ulandrena</emphasis>
</taxonomicName>
species), the terga are weakly punctate in a manner comparable to the female sex (
<figureCitation id="71B92A6FAC78435A0861F86B8D9901FB" box="[885,1020,1952,1978]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="33" pageNumber="34">Figure 18D</figureCitation>
), the head is noticeably broader that comparison species at 1.5 times wider than long (usually &lt;1.3 times wider than long in other
<taxonomicName id="2E824D69AC78435A0FAFF80F8B56019F" authorityName="Warncke" authorityYear="1968" box="[1211,1331,1988,2014]" pageId="33" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC78435A0FAFF80F8B56019F" box="[1211,1331,1988,2014]" italics="true" pageId="33" pageNumber="34">Ulandrena</emphasis>
</taxonomicName>
species), and the ocelloccipital distance is relatively short, only 1.7 times the diameter of a lateral ocellus (
<figureCitation id="71B92A6FAC7B43590FE7FF5C8B1C06F0" box="[1267,1401,151,177]" captionStart="FIGURE 18" captionStartId="33.[151,250,1738,1762]" captionTargetBox="[151,1436,273,1715]" captionTargetId="figure-53@33.[151,1436,273,1715]" captionTargetPageId="33" captionText="FIGURE 18. Andrena (Ulandrena) incustodita spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209347" httpUri="https://zenodo.org/record/13209347/files/figure.png" pageId="34" pageNumber="35">Figure 18C</figureCitation>
; in
<emphasis id="DBF6EAF8AC7B43590B83FF708F120695" box="[151,375,187,213]" italics="true" pageId="34" pageNumber="35">
<taxonomicName id="2E824D69AC7B43590B83FF708F550694" authorityName="Dours" authorityYear="1873" box="[151,304,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="abbreviata">A. abbreviata</taxonomicName>
<taxonomicNameLabel id="C0C55783AC7B43590A2EFF708F120695" box="[314,375,187,212]" pageId="34" pageNumber="35" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
the ocelloccipital distance is 2.5 times the diameter of a lateral ocellus). Combined with the distribution that is restricted to
<collectingRegion id="2B46F808AC7B43590AEEFF148C5C06B8" box="[506,569,223,249]" country="Greece" name="Kriti" pageId="34" pageNumber="35">Crete</collectingRegion>
,
<taxonomicName id="2E824D69AC7B43590951FF148C8406B8" authorityName="WOOD" authorityYear="2024" box="[581,737,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="incustodita">
<emphasis id="DBF6EAF8AC7B43590951FF148C8406B8" box="[581,737,223,249]" italics="true" pageId="34" pageNumber="35">A. incustodita</emphasis>
</taxonomicName>
is therefore consistently morphologically diagnosable against other members of the
<taxonomicName id="2E824D69AC7B43590A99FEC88C60075C" authorityName="Dours" authorityYear="1873" box="[397,517,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="abbreviata">
<emphasis id="DBF6EAF8AC7B43590A99FEC88C60075C" box="[397,517,259,285]" italics="true" pageId="34" pageNumber="35">abbreviata</emphasis>
</taxonomicName>
-group.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7B43590BD3FEEC8D800724" pageId="34" pageNumber="35" type="etymology">
<paragraph id="E93D36EAAC7B43590BD3FEEC8D800724" blockId="34.[151,1436,151,393]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B43590BD3FEEC8F2B0700" bold="true" box="[199,334,295,321]" pageId="34" pageNumber="35">Etymology.</emphasis>
Feminine singular nominative form of the Latin adjective
<emphasis id="DBF6EAF8AC7B43590F1FFEEC8AE20700" box="[1035,1159,295,321]" italics="true" pageId="34" pageNumber="35">incustodito</emphasis>
meaning unwatched or unguarded, in reference to the lack of consistent recognition of this species.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7B43590BD3FEA58C7007C8" box="[199,533,366,393]" pageId="34" pageNumber="35" type="distribution">
<paragraph id="E93D36EAAC7B43590BD3FEA58C7007C8" blockId="34.[151,1436,151,393]" box="[199,533,366,393]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B43590BD3FEA58F0407C9" bold="true" box="[199,353,366,392]" pageId="34" pageNumber="35">Distribution.</emphasis>
<collectingCountry id="9195767AAC7B43590A73FEA48FD307C8" box="[359,438,367,393]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
(
<collectingRegion id="2B46F808AC7B43590AD2FEA48C6C07C8" box="[454,521,367,393]" country="Greece" name="Kriti" pageId="34" pageNumber="35">Crete</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="EF0391DD37086F9DF3720E2BA3B7BCA5">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:date id="721F5410072240F34172D261D92CD8CB">2024</mods:date>
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<mods:number id="45A66CAE36F32DDDF7BB5A060BC0B4DC">5483</mods:number>
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<subSubSection id="A1986561AC49436B0B83FE358CAF0458" box="[151,714,510,537]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="E93D36EAAC49436B0B83FE358CAF0458" blockId="16.[151,714,510,573]" box="[151,714,510,537]" pageId="16" pageNumber="17">
<heading id="B2758186AC49436B0B83FE358CAF0458" bold="true" box="[151,714,510,537]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<emphasis id="DBF6EAF8AC49436B0B83FE358CAF0458" bold="true" box="[151,714,510,537]" pageId="16" pageNumber="17">
<taxonomicName id="2E824D69AC49436B0B83FE358C290458" authorityName="WOOD" authorityYear="2024" box="[151,588,510,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="melacanoides" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC49436B0B83FE358F640458" bold="true" box="[151,257,510,537]" italics="true" pageId="16" pageNumber="17">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC49436B0A06FE358FF00458" bold="true" box="[274,405,510,537]" italics="true" pageId="16" pageNumber="17">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC49436B0AB3FE358C290458" bold="true" box="[423,588,510,537]" italics="true" pageId="16" pageNumber="17">melacanoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC49436B0947FE348CAF0458" box="[595,714,511,537]" pageId="16" pageNumber="17" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC49436B0B83FDE88FF1047C" box="[151,404,547,573]" pageId="16" pageNumber="17" type="description">
<paragraph id="E93D36EAAC49436B0B83FDE88FF1047C" blockId="16.[151,714,510,573]" box="[151,404,547,573]" pageId="16" pageNumber="17">
<figureCitation id="71B92A6FAC49436B0B83FDE88F53047C" box="[151,310,547,573]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figures 9AF</figureCitation>
;
<figureCitation id="71B92A6FAC49436B0A55FDE88FF5047C" box="[321,400,547,573]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="16" pageNumber="17">10AF</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC49436B0B83FDA18FC70578" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="E93D36EAAC49436B0B83FDA18AD104E8" blockId="16.[151,1437,618,2013]" pageId="16" pageNumber="17">
<materialsCitation id="59EA3CB7AC49436B0B83FDA18AD104E8" collectingDate="2023-03-18" collectorName="T. J. Wood" country="Spain" county="Finca Murtas" location="Cadiz" municipality="Las Algamitas" pageId="16" pageNumber="17" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC49436B0B83FDA18FC204C5" bold="true" box="[151,423,618,644]" pageId="16" pageNumber="17">
<typeStatus id="36398848AC49436B0B83FDA18F5F04C5" box="[151,314,618,644]" pageId="16" pageNumber="17" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC49436B0A58FDA18FC704C5" box="[332,418,618,644]" name="Spain" pageId="16" pageNumber="17">SPAIN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC49436B0AA0FDA08F9D04C4" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC49436B0AA0FDA08F9D04C4" box="[436,504,619,645]" country="Spain" county="Finca Murtas" municipality="Las Algamitas" name="Cadiz" pageId="16" pageNumber="17">Cádiz</location>
,
<location id="EC5D6031AC49436B091DFDA08DF504C4" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC49436B091DFDA08DF504C4" box="[521,912,619,645]" country="Spain" county="Finca Murtas" municipality="Las Algamitas" name="Parque Natural Los Alcornocales" pageId="16" pageNumber="17">Parque Natural Los Alcornocales</location>
,
<collectingMunicipality id="0959AC90AC49436B08B4FDA08A2C04C4" box="[928,1097,619,645]" pageId="16" pageNumber="17">Las Algamitas</collectingMunicipality>
,
<collectingCounty id="005C4E66AC49436B0F4EFDA08A9204C4" box="[1114,1271,619,645]" pageId="16" pageNumber="17">Finca Murtas</collectingCounty>
, 36.3273
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, - 5.5986
<superScript id="1EF79BA2AC49436B0BF0FD448F6204E8" attach="right" box="[228,263,655,681]" fontSize="6" pageId="16" pageNumber="17">oW</superScript>
,
<date id="9D3C102AAC49436B0A06FD448FEB04E8" box="[274,398,655,681]" pageId="16" pageNumber="17" value="2023-03-18">
<collectingDate id="8D78E9C2AC49436B0A06FD448FEB04E8" box="[274,398,655,681]" pageId="16" pageNumber="17" value="2023-03-18">18.iii.2023</collectingDate>
</date>
,
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, leg.
<collectorName id="4477533CAC49436B0AF8FD448C0504E8" box="[492,608,655,681]" pageId="16" pageNumber="17">T.J. Wood</collectorName>
, OÖLM (BOLD accession number WPATW986-23).
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAAC49436B0BD3FD798FC70578" blockId="16.[151,1437,618,2013]" pageId="16" pageNumber="17">
<materialsCitation id="59EA3CB7AC49436B0BD3FD798A3F048E" box="[199,1114,690,719]" collectionCode="TJWC" country="Spain" location="SPAIN" pageId="16" pageNumber="17" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<emphasis id="DBF6EAF8AC49436B0BD3FD798FBF048D" bold="true" box="[199,474,690,716]" pageId="16" pageNumber="17">
<typeStatus id="36398848AC49436B0BD3FD798F0A048D" box="[199,367,690,716]" pageId="16" pageNumber="17" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="9195767AAC49436B0A6BFD798FB0048D" box="[383,469,690,716]" name="Spain" pageId="16" pageNumber="17">SPAIN</collectingCountry>
:
</emphasis>
same information as
<typeStatus id="36398848AC49436B09CCFD788D58048C" box="[728,829,691,717]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
,
<specimenCount id="FF84FD63AC49436B085FFD788D14048C" box="[843,881,691,717]" count="2" pageId="16" pageNumber="17" type="male">2♂</specimenCount>
,
<specimenCount id="FF84FD63AC49436B0894FD788DC6048C" box="[896,931,691,717]" count="1" pageId="16" pageNumber="17" type="female">1♀</specimenCount>
, OÖLM/
<collectionCode id="8F93AE2FAC49436B0F04FD7F8A3F048E" box="[1040,1114,692,719]" pageId="16" pageNumber="17">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC49436B0F7DFD788AE904B0" collectingDate="2023-03-18" collectionCode="TJWC" collectorName="T. J. Wood" country="Spain" county="Las Algamitas" location="Cadiz" municipality="Parque Natural Los Alcornocales" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="EC5D6031AC49436B0F7DFD788AC8048C" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC49436B0F7DFD788AC8048C" box="[1129,1197,691,717]" country="Spain" county="Las Algamitas" municipality="Parque Natural Los Alcornocales" name="Cadiz" pageId="16" pageNumber="17">Cádiz</location>
,
<collectingMunicipality id="0959AC90AC49436B0FA8FD788F4B04B0" pageId="16" pageNumber="17">Parque Natural Los Alcornocales</collectingMunicipality>
,
<collectingCounty id="005C4E66AC49436B0A2FFD1C8F8404B0" box="[315,481,727,753]" pageId="16" pageNumber="17">Las Algamitas</collectingCounty>
, immediately north,
<date id="9D3C102AAC49436B09DBFD1C8D2E04B0" box="[719,843,727,753]" pageId="16" pageNumber="17" value="2023-03-18">
<collectingDate id="8D78E9C2AC49436B09DBFD1C8D2E04B0" box="[719,843,727,753]" pageId="16" pageNumber="17" value="2023-03-18">18.iii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC49436B084CFD1C8D1904B0" box="[856,892,727,753]" count="1" pageId="16" pageNumber="17" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC49436B08A9FD1C8A5604B0" box="[957,1075,727,753]" pageId="16" pageNumber="17">T.J. Wood</collectorName>
,
<collectionCode id="8F93AE2FAC49436B0F2BFD1C8AE904B0" box="[1087,1164,727,753]" pageId="16" pageNumber="17">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC49436B0F8DFD1C8D130556" collectingDate="2023-03-19" collectionCode="TJWC" collectorName="T. J. Wood" country="Spain" location="Cadiz" municipality="Tarifa" pageId="16" pageNumber="17" specimenCount="10" specimenCount-female="6" specimenCount-male="4" typeStatus="paratype">
<location id="EC5D6031AC49436B0F8DFD1C8AB804B0" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC49436B0F8DFD1C8AB804B0" box="[1177,1245,727,753]" country="Spain" municipality="Tarifa" name="Cadiz" pageId="16" pageNumber="17">Cádiz</location>
,
<collectingMunicipality id="0959AC90AC49436B0FFEFD1C8B4A04B0" box="[1258,1327,727,753]" pageId="16" pageNumber="17">Tarifa</collectingMunicipality>
,
<quantity id="2E7A9B0FAC49436B0E29FD1C8B1D04B0" box="[1341,1400,727,753]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="km" value="1.0">1 km</quantity>
N, grazing fields,
<date id="9D3C102AAC49436B0A29FD308FDC0554" box="[317,441,763,789]" pageId="16" pageNumber="17" value="2023-03-19">
<collectingDate id="8D78E9C2AC49436B0A29FD308FDC0554" box="[317,441,763,789]" pageId="16" pageNumber="17" value="2023-03-19">19.iii.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC49436B0AD7FD308F8D0554" box="[451,488,763,789]" count="4" pageId="16" pageNumber="17" type="male">4♂</specimenCount>
,
<specimenCount id="FF84FD63AC49436B0AE7FD308C720554" box="[499,535,763,789]" count="6" pageId="16" pageNumber="17" type="female">6♀</specimenCount>
, leg.
<collectorName id="4477533CAC49436B0946FD308CA00554" box="[594,709,763,789]" pageId="16" pageNumber="17">T.J. Wood</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FAC49436B0838FD378D130556" box="[812,886,764,791]" pageId="16" pageNumber="17">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC49436B0894FD308FFB0578" collectingDate="1989-05-09" collectionCode="TJWC" collectorName="A. Jimenez" country="Spain" county="Granada" location="Loja" municipality="Charco" pageId="16" pageNumber="17" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="EC5D6031AC49436B0894FD308DD30554" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC49436B0894FD308DD30554" box="[896,950,763,789]" country="Spain" county="Granada" municipality="Charco" name="Loja" pageId="16" pageNumber="17">Loja</location>
,
<collectingMunicipality id="0959AC90AC49436B08D5FD308A770554" box="[961,1042,763,789]" pageId="16" pageNumber="17">Charco</collectingMunicipality>
del negro (
<collectingCounty id="005C4E66AC49436B0F9AFD308A960554" box="[1166,1267,763,789]" pageId="16" pageNumber="17">Granada</collectingCounty>
),
<date id="9D3C102AAC49436B0E11FD308B0D0554" box="[1285,1384,763,789]" pageId="16" pageNumber="17" value="1989-05-09">
<collectingDate id="8D78E9C2AC49436B0E11FD308B0D0554" box="[1285,1384,763,789]" pageId="16" pageNumber="17" value="1989-05-09">9.v.1989</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC49436B0E66FD308BF20554" box="[1394,1431,763,789]" count="1" pageId="16" pageNumber="17" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC49436B0BD2FCD48F220578" box="[198,327,799,825]" pageId="16" pageNumber="17">A. Jiménez</collectorName>
,
<collectionCode id="8F93AE2FAC49436B0A45FCD48FFB0578" box="[337,414,799,825]" pageId="16" pageNumber="17">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4943680BD3FC898A2607C8" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17" type="description">
<paragraph id="E93D36EAAC49436B0BD3FC898CA00284" blockId="16.[151,1437,618,2013]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3FC898FD8051C" bold="true" box="[199,445,834,861]" pageId="16" pageNumber="17">Description. Female.</emphasis>
Body length: 7.5
<quantity id="2E7A9B0FAC49436B0999FC898CAE051C" box="[653,715,834,861]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="17" unit="mm" value="8.0">8 mm</quantity>
(
<figureCitation id="71B92A6FAC49436B09CCFC888D2B051C" box="[728,846,835,861]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9A</figureCitation>
).
<emphasis id="DBF6EAF8AC49436B0876FC888DC2051C" box="[866,935,835,861]" italics="true" pageId="16" pageNumber="17">Head:</emphasis>
Dark, 1.4 times wider than long (
<figureCitation id="71B92A6FAC49436B0E0EFC888BF5051C" box="[1306,1424,835,861]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9B</figureCitation>
). Clypeus very weakly domed, more or less flattened, densely and shallowly punctate, punctures separated by 0.51 puncture diameters, underlying surface shagreened and weakly shining. Process of labrum rounded rectangular, 34 times broader than long, anterior margin straight to weakly emarginate. Gena slightly exceeding width of compound eye; ocelloccipital distance equalling diameter of lateral ocellus. Foveae dorsally occupying ⅔ of space between compound eye and lateral ocellus, ventrally slightly narrowing, reaching upper level of antennal insertions; foveae filled with dark brown to black hairs. Face and scape medially covered with pale whitish plumose hairs, becoming intermixed with black hairs laterally, on frons, and along inner margins of compound eyes. Gena ventrally with predominantly pale hairs, becoming predominantly black dorsolaterally, vertex with long black to light brown hairs, hairs clearly exceeding length of scape. Antennae basally dark, A512 ventrally lightened by presence of grey scales; A3 exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="E93D36EAAC49436B0BD3FB048A2500FC" blockId="16.[151,1437,618,2013]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3FB048F2D02A9" box="[199,328,1231,1256]" italics="true" pageId="16" pageNumber="17">Mesosoma:</emphasis>
Scutum and scutellum finely shagreened, weakly shining, with very slight hints of metallic greasy purple reflections (
<figureCitation id="71B92A6FAC49436B0A73FB388FBE034C" box="[359,475,1267,1293]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9D</figureCitation>
) laterally; surface irregularly punctate, punctures separated by 13 puncture diameters. Pronotum rounded, surface with granular shagreen, weakly shining (
<figureCitation id="71B92A6FAC49436B08B0FADC8A790370" box="[932,1052,1303,1329]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9C</figureCitation>
). Mesepisternum and dorsolateral parts of propodeum with fine granular reticulation, weakly shining, surface almost smooth; propodeal triangle weakly differentiated, laterally delineated by weakly differentiated rim, internal surface very slightly depressed, with granular reticulation matching remaining parts of propodeum, hence blending together (
<figureCitation id="71B92A6FAC49436B0F7CFA488AB803DC" box="[1128,1245,1411,1437]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9E</figureCitation>
). Mesepisternum with pubescence variable, typically with long plumose pale whitish hairs, with intermixed black plumose hairs, black hairs typically representing &lt;30% of hairs but sometimes up to 60%, hairs exceeding scape in length. Scutum and scutellum with shorter intermixed plumose light brown and black hairs. Propodeal corbicula incomplete, corbicular fringe composed of long light brown plumose hairs with occasional intermixed black hairs, internal surface with abundant simple light brown hairs. Legs dark, apical tarsal segments lightened dark brown, pubescence light brown. Flocculus complete, strongly produced, composed of whitish hairs; femoral and tibial scopae composed of simple hairs, predominantly whitish, dorsal fringe of tibial scopae becoming darker brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark orange, nervulus weakly antefurcal.
</paragraph>
<paragraph id="E93D36EAAC49436B0BD3F90C8FB201D4" blockId="16.[151,1437,618,2013]" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3F90C8F2000A1" box="[199,325,1735,1760]" italics="true" pageId="16" pageNumber="17">Metasoma:</emphasis>
Tergal discs dark, apical rims narrowly lightened hyaline-brown (
<figureCitation id="71B92A6FAC49436B0F29F90C8AD700A0" box="[1085,1202,1735,1761]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="16" pageNumber="17">Figure 9F</figureCitation>
). Terga very weakly shagreened, shining; surface with scattered hair-bearing punctures, punctures separated by 34 puncture diameters. Tergal discs with long upstanding pale hairs, longest on T1, becoming shorter on subsequent terga, T24 with very weak apical hairbands, not obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate rounded triangular, lateral margin impunctate and dull, medially with dense granular reticulation, weakly shining.
</paragraph>
<paragraph id="E93D36EAAC49436A0BD3F8548FBF07C8" blockId="16.[151,1437,618,2013]" lastBlockId="17.[151,1437,151,393]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="DBF6EAF8AC49436B0BD3F8548F6E01F8" bold="true" box="[199,267,1951,1977]" pageId="16" pageNumber="17">Male.</emphasis>
Body length: 77.5 mm (
<figureCitation id="71B92A6FAC49436B0922F8548CDB01F8" box="[566,702,1951,1977]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="16" pageNumber="17">Figure 10A</figureCitation>
).
<emphasis id="DBF6EAF8AC49436B09C0F8548D7F01F8" box="[724,794,1951,1977]" italics="true" pageId="16" pageNumber="17">Head:</emphasis>
Dark, 1.4 times wider than long (
<figureCitation id="71B92A6FAC49436B0FB0F8548B4F01F8" box="[1188,1322,1951,1977]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="16" pageNumber="17">Figure 10B</figureCitation>
). Clypeus weakly domed, densely and shallowly punctate, punctures separated by 0.51 puncture diameters, underlying surface shagreened and dull to weakly shining. Process of labrum rounded trapezoidal, 2 times wider than long, anterior margin weakly emarginate. Gena exceeding width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Face and scape medially covered with long white plumose hairs, becoming intermixed with black hairs laterally, on frons, and along inner margins of compound eyes. Gena ventrally with predominantly white hairs, becoming predominantly black dorsolaterally, vertex with long black to white hairs, hairs clearly exceeding length of scape. Antennae basally dark, A513 ventrally lightened by presence of grey scales; A3 slightly exceeding A4+5, clearly shorter than A4+5+6.
</paragraph>
<caption id="BDFD6662AC48436A0B83F8918A7401D7" ID-DOI="http://doi.org/10.5281/zenodo.13209325" ID-Zenodo-Dep="13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1882,1906]" targetBox="[151,1436,417,1859]" targetPageId="17" targetType="figure">
<paragraph id="E93D36EAAC48436A0B83F8918A7401D7" blockId="17.[151,1437,1882,1942]" pageId="17" pageNumber="18">
<emphasis id="DBF6EAF8AC48436A0B83F8918F770133" bold="true" box="[151,274,1882,1906]" pageId="17" pageNumber="18">FIGURE 9.</emphasis>
<taxonomicName id="2E824D69AC48436A0A02F8918C1A0133" authorityName="WOOD" authorityYear="2024" box="[278,639,1882,1906]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="melacanoides" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC48436A0A02F8918F090133" box="[278,364,1882,1906]" italics="true" pageId="17" pageNumber="18">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC48436A0A6EF8918F820133" box="[378,487,1882,1906]" italics="true" pageId="17" pageNumber="18">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC48436A0AE0F8918C1A0133" box="[500,639,1882,1906]" italics="true" pageId="17" pageNumber="18">melacanoides</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC48436A0991F8918C8F0133" bold="true" box="[645,746,1882,1906]" pageId="17" pageNumber="18">
<taxonomicNameLabel id="C0C55783AC48436A0991F8918C8F0133" box="[645,746,1882,1906]" pageId="17" pageNumber="18" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC4B43690BD3FF5C8D6307C8" blockId="18.[151,1437,151,393]" pageId="18" pageNumber="19">
<emphasis id="DBF6EAF8AC4B43690BD3FF5C8F2D06F1" box="[199,328,151,176]" italics="true" pageId="18" pageNumber="19">Mesosoma:</emphasis>
Scutum and scutellum with dense granular shagreen, dull, becoming slightly weaker medially, here weakly shining; surface weakly and obscurely punctate, punctures separated by 13 puncture diameters (
<figureCitation id="71B92A6FAC4B43690E45FF708EAF06B8" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="18" pageNumber="19">Figure 10C</figureCitation>
). Pronotum rounded. Mesepisternum and propodeum structurally as in female (
<figureCitation id="71B92A6FAC4B43690F47FF148ABE06B8" box="[1107,1243,223,249]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="18" pageNumber="19">Figure 10D</figureCitation>
). Mesepisternum with very long white plumose hairs, with occasional scattered shorter black hairs. Scutum and scutellum with long white plumose hairs, with shorter black plumose hairs intermixed, particularly on scutellum. Legs dark, apical tarsal segments lightened dark brown, pubescence white. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark orange, nervulus interstitial to antefurcal.
</paragraph>
<caption id="BDFD6662AC4B43690B83F8928A3A01D4" ID-DOI="http://doi.org/10.5281/zenodo.13209327" ID-Zenodo-Dep="13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1881,1905]" targetBox="[151,1436,417,1857]" targetPageId="18" targetType="figure">
<paragraph id="E93D36EAAC4B43690B83F8928A3A01D4" blockId="18.[151,1436,1881,1941]" pageId="18" pageNumber="19">
<emphasis id="DBF6EAF8AC4B43690B83F8928F450130" bold="true" box="[151,288,1881,1905]" pageId="18" pageNumber="19">FIGURE 10.</emphasis>
<taxonomicName id="2E824D69AC4B43690A33F8928CF60130" authorityName="WOOD" authorityYear="2024" box="[295,659,1881,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="melacanoides" status="sp. nov." subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4B43690A33F8928F180130" box="[295,381,1881,1905]" italics="true" pageId="18" pageNumber="19">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4B43690A98F8928F9C0130" box="[396,505,1881,1905]" italics="true" pageId="18" pageNumber="19">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4B4369091CF8928CF60130" box="[520,659,1881,1905]" italics="true" pageId="18" pageNumber="19">melacanoides</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC4B4369098EF8928D670130" bold="true" box="[666,770,1881,1905]" pageId="18" pageNumber="19">
<taxonomicNameLabel id="C0C55783AC4B4369098EF8928D670130" box="[666,770,1881,1905]" pageId="18" pageNumber="19" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC4A43680BD3FF5C8A2607C8" blockId="19.[151,1436,151,395]" pageId="19" pageNumber="20">
<emphasis id="DBF6EAF8AC4A43680BD3FF5C8F2006F1" box="[199,325,151,176]" italics="true" pageId="19" pageNumber="20">Metasoma:</emphasis>
Terga structurally as in female, covered with sparse long whitish hairs, with weak hints of apical hairbands laterally on T24 (
<figureCitation id="71B92A6FAC4A43680AF4FF708C000694" box="[480,613,187,213]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="19" pageNumber="20">Figure 10E</figureCitation>
). T6 laterally with long white plumose hairs, medially and T7 with long dark brown hairs overlying pygidial plate of T7. S8 columnar, ventrally with dense fan of brown hairs, apical process slightly broadened and thickened. Genital capsule slightly elongate, gonocoxae apically produced into long narrow rounded teeth (
<figureCitation id="71B92A6FAC4A43680A51FEEC8FAF0700" box="[325,458,295,321]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="19" pageNumber="20">Figure 10F</figureCitation>
). Gonostyli strongly broadened and triangular apically, inner margin strongly raised, surface covered with light brown hairs. Penis valves basally slightly broadened, with slight hyaline extensions at level of gonocoxal teeth, occupying ½ space between gonostyli, narrowing apically.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC4A43680B83F8938EAB01F9" ID-DOI="http://doi.org/10.5281/zenodo.13209329" ID-Zenodo-Dep="13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1880,1904]" targetBox="[151,1436,417,1856]" targetPageId="19" targetType="figure">
<paragraph id="E93D36EAAC4A43680B83F8938EAB01F9" blockId="19.[151,1437,1880,1976]" pageId="19" pageNumber="20">
<emphasis id="DBF6EAF8AC4A43680B83F8938F440131" bold="true" box="[151,289,1880,1904]" pageId="19" pageNumber="20">FIGURE 11.</emphasis>
<taxonomicName id="2E824D69AC4A43680A3EF8938D670131" authority="Friese, 1922" authorityName="Friese" authorityYear="1922" box="[298,770,1880,1904]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melaleuca" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4A43680A3EF8938FE50131" box="[298,384,1880,1904]" italics="true" pageId="19" pageNumber="20">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4A43680A86F8938F9A0131" box="[402,511,1880,1904]" italics="true" pageId="19" pageNumber="20">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4A43680905F8938C1C0131" box="[529,633,1880,1904]" italics="true" pageId="19" pageNumber="20">melaleuca</emphasis>
<bibRefCitation id="8D134B1BAC4A43680996F8938D670131" author="Friese, H." box="[642,770,1880,1904]" pageId="19" pageNumber="20" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese, 1922</bibRefCitation>
</taxonomicName>
nec. Pérez =
<taxonomicName id="2E824D69AC4A43680883F8938AD80131" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" box="[919,1213,1880,1904]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4A43680883F8938A7D0131" box="[919,1048,1880,1904]" italics="true" pageId="19" pageNumber="20">A. melacana</emphasis>
<bibRefCitation id="8D134B1BAC4A43680F36F8938AD80131" author="Warncke, K." box="[1058,1213,1880,1904]" pageId="19" pageNumber="20" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC4A43680FD2F8938B420131" bold="true" box="[1222,1319,1880,1904]" pageId="19" pageNumber="20">lectotype</emphasis>
female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC4D436F0BD3FF5D8C8905E5" pageId="20" pageNumber="21" type="diagnosis">
<paragraph id="E93D36EAAC4D436F0BD3FF5D8DC107ED" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
<emphasis id="DBF6EAF8AC4D436F0BD3FF5D8F2706F1" bold="true" box="[199,322,150,176]" pageId="20" pageNumber="21">Diagnosis.</emphasis>
<taxonomicName id="2E824D69AC4D436F0A53FF5D8C2206F1" authorityName="WOOD" authorityYear="2024" box="[327,583,150,176]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0A53FF5D8C2206F1" box="[327,583,150,176]" italics="true" pageId="20" pageNumber="21">Andrena melacanoides</emphasis>
</taxonomicName>
can be recognised as an
<taxonomicName id="2E824D69AC4D436F0843FF5D8DAA06F1" authorityName="Warncke" authorityYear="1968" box="[855,975,150,176]" pageId="20" pageNumber="9" rank="subGenus" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4D436F0843FF5D8DAA06F1" box="[855,975,150,176]" italics="true" pageId="20" pageNumber="21">Avandrena</emphasis>
</taxonomicName>
due to its short and broad head with short and broad foveae; recognition of the spineless
<taxonomicName id="2E824D69AC4D436F098CFF718D750695" authorityName="Warncke" authorityYear="1968" box="[664,784,186,212]" pageId="20" pageNumber="9" rank="subGenus" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4D436F098CFF718D750695" box="[664,784,186,212]" italics="true" pageId="20" pageNumber="21">Avandrena</emphasis>
</taxonomicName>
is always more challenging, since the subgenus is partially defined on the presence of this character, and
<taxonomicName id="2E824D69AC4D436F09B6FF148D3806B9" authorityName="WOOD" authorityYear="2024" box="[674,861,222,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F09B6FF148D3806B9" box="[674,861,222,249]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
lacks spines on the hind face of the hind femorae. It is best diagnosed against the most morphologically similar species which is
<taxonomicName id="2E824D69AC4D436F0F1CFEC88B21075D" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" box="[1032,1348,258,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0F1CFEC88AF0075D" box="[1032,1173,258,285]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
<bibRefCitation id="8D134B1BAC4D436F0F8FFEC98B21075D" author="Warncke, K." box="[1179,1348,258,284]" pageId="20" pageNumber="21" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
as both species show a propodeum and propodeal triangle with smooth granular reticulation, without any raised reticulation or rugae (contrast
<taxonomicName id="2E824D69AC4D436F0A79FE808FB50725" authorityName="Wood" authorityYear="2023" box="[365,464,330,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="juliae">
<emphasis id="DBF6EAF8AC4D436F0A79FE808FB50725" box="[365,464,330,357]" italics="true" pageId="20" pageNumber="21">A. juliae</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC4D436F0918FE808CC10725" authorityName="Wood &amp; Ortiz-Sanchez" authorityYear="2022" box="[524,676,330,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="erodiorum">
<emphasis id="DBF6EAF8AC4D436F0918FE808CC10725" box="[524,676,330,357]" italics="true" pageId="20" pageNumber="21">A. erodiorum</emphasis>
</taxonomicName>
which have the propodeum with conspicuously raised sculpture), intermixed pale and black hairs on the face and mesepisternum, a broad process of the labrum, and facial foveae that occupy ⅔ of the space between the compound eye and lateral ocellus.
</paragraph>
<paragraph id="E93D36EAAC4D436F0BD3FE7D8C8905E5" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
<taxonomicName id="2E824D69AC4D436F0BD3FE7D8FAC0791" authorityName="WOOD" authorityYear="2024" box="[199,457,438,464]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0BD3FE7D8FAC0791" box="[199,457,438,464]" italics="true" pageId="20" pageNumber="21">Andrena melacanoides</emphasis>
</taxonomicName>
was previously reported from southern
<collectingCountry id="9195767AAC4D436F089BFE7D8DAA0791" box="[911,975,438,464]" name="Spain" pageId="20" pageNumber="21">Spain</collectingCountry>
as
<taxonomicName id="2E824D69AC4D436F08E2FE7C8AE60791" authorityName="Warncke" authorityYear="1967" box="[1014,1155,438,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F08E2FE7C8AE60791" box="[1014,1155,438,465]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC4D436F0F86FE7D8EB607B5" author="Wood, T. J. &amp; Ortiz-Sanchez, F. J." pageId="20" pageNumber="21" pagination="114 - 123" refId="ref92242" refString="Wood, T. J. &amp; Ortiz-Sanchez, F. J. (2022) Description of three new Andrena Fabricius, 1775 species from understudied parts of Iberia (Hymenoptera: Andrenidae). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 70, 114 - 123." type="book chapter" year="2022">Wood &amp; Ortiz-Sánchez 2022</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC4D436F0BF6FE118F1F07B5" author="Wood, T. J." box="[226,378,474,500]" pageId="20" pageNumber="21" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
), but examination of the type series in the ZMHB collection (
<figureCitation id="71B92A6FAC4D436F0F4DFE118AAB07B5" box="[1113,1230,474,500]" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
; see Remarks for
<typeStatus id="36398848AC4D436F0B83FE358E9B0459" box="[151,254,510,536]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
designation) shows that the two species cannot be conspecific. True female
<taxonomicName id="2E824D69AC4D436F0F2BFE348AAC0459" authorityName="Warncke" authorityYear="1967" box="[1087,1225,510,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0F2BFE348AAC0459" box="[1087,1225,510,537]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
have the pronotum laterally with a conspicuous longitudinal shining impression (
<figureCitation id="71B92A6FAC4D436F0852FDE98DAF047D" box="[838,970,546,572]" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="20" pageNumber="21">Figure 11D</figureCitation>
), whereas in
<taxonomicName id="2E824D69AC4D436F0F49FDE88B73047D" authorityName="WOOD" authorityYear="2024" box="[1117,1302,546,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0F49FDE88B73047D" box="[1117,1302,546,573]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
females the pronotum is evenly rounded and dull (
<figureCitation id="71B92A6FAC4D436F0959FD8D8CB60421" box="[589,723,582,608]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="20" pageNumber="21">Figure 10C</figureCitation>
). More subtly, the fore margin of the clypeus is comparatively more strongly upturned in
<taxonomicName id="2E824D69AC4D436F0AD1FDA08C3704C5" authorityName="Warncke" authorityYear="1967" box="[453,594,618,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0AD1FDA08C3704C5" box="[453,594,618,645]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
and only very weakly upturned in
<taxonomicName id="2E824D69AC4D436F08CAFDA08AFD04C5" authorityName="WOOD" authorityYear="2024" box="[990,1176,618,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F08CAFDA08AFD04C5" box="[990,1176,618,645]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
, and the shagreenation of the terga of
<taxonomicName id="2E824D69AC4D436F0A51FD448FB604E9" authorityName="Warncke" authorityYear="1967" box="[325,467,654,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0A51FD448FB604E9" box="[325,467,654,681]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
is slightly stronger compared to
<taxonomicName id="2E824D69AC4D436F0846FD448A6804E9" authorityName="WOOD" authorityYear="2024" box="[850,1037,654,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0846FD448A6804E9" box="[850,1037,654,681]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC4D436F0F0AFD458AFE04E9" box="[1054,1179,654,680]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="20" pageNumber="21">Figures 9F</figureCitation>
and
<figureCitation id="71B92A6FAC4D436F0FC2FD458B6604E9" box="[1238,1283,654,680]" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="20" pageNumber="21">11F</figureCitation>
; comparative material required). Based on examined specimens from
<collectingCountry id="9195767AAC4D436F081AFD798D2B048D" box="[782,846,690,716]" name="Spain" pageId="20" pageNumber="21">Spain</collectingCountry>
and North Africa, no intermediate individuals could be found, and the known distributions do not overlap as all Moroccan female specimens show the clear longitudinal ridge on the pronotum. Separation of males is more challenging, as the two species appear to be almost identical. In direct comparison, based on a limited number of specimens, the gonocoxal teeth of
<taxonomicName id="2E824D69AC4D436F0F6AFCD48B770579" authorityName="Friese" authorityYear="1922" box="[1150,1298,798,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaleuca">
<emphasis id="DBF6EAF8AC4D436F0F6AFCD48B770579" box="[1150,1298,798,825]" italics="true" pageId="20" pageNumber="21">A. melaleuca</emphasis>
</taxonomicName>
are broader compared to
<taxonomicName id="2E824D69AC4D436F0A33FC888FBA051D" authorityName="WOOD" authorityYear="2024" box="[295,479,834,861]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0A33FC888FBA051D" box="[295,479,834,861]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
, and the tergal discs of
<taxonomicName id="2E824D69AC4D436F09F6FC888D16051D" authorityName="Friese" authorityYear="1922" box="[738,883,834,861]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaleuca">
<emphasis id="DBF6EAF8AC4D436F09F6FC888D16051D" box="[738,883,834,861]" italics="true" pageId="20" pageNumber="21">A. melaleuca</emphasis>
</taxonomicName>
are comparatively more strongly shagreened than in
<taxonomicName id="2E824D69AC4D436F0BA2FCAC8F1705C1" authorityName="WOOD" authorityYear="2024" box="[182,370,870,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0BA2FCAC8F1705C1" box="[182,370,870,897]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC4D436F0A97FCAD8C6B05C1" box="[387,526,870,896]" captionStart="FIGURE 10" captionStartId="18.[151,250,1881,1905]" captionTargetBox="[151,1436,417,1857]" captionTargetId="figure-125@18.[151,1436,417,1857]" captionTargetPageId="18" captionText="FIGURE 10. Andrena (Avandrena) melacanoides spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209327" httpUri="https://zenodo.org/record/13209327/files/figure.png" pageId="20" pageNumber="21">Figures 10F</figureCitation>
and
<figureCitation id="71B92A6FAC4D436F095DFCAD8CFD05C1" box="[585,664,870,896]" captionStart="FIGURE 12" captionStartId="21.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@21.[151,1436,181,1624]" captionTargetPageId="21" captionText="FIGURE 12. Andrena (Avandrena) melaleuca Friese, 1922 nec. Provancher = A. melacana Warncke, 1967 male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, dorsal view alternate." figureDoi="http://doi.org/10.5281/zenodo.13209331" httpUri="https://zenodo.org/record/13209331/files/figure.png" pageId="20" pageNumber="21">12EF</figureCitation>
). Additional specimens are required to establish whether additional characters to differentiate these species can be found.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4D436F0BD3FC658FC30049" pageId="20" pageNumber="21" type="discussion">
<paragraph id="E93D36EAAC4D436F0BD3FC658D690339" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
<emphasis id="DBF6EAF8AC4D436F0BD3FC658F5E0589" bold="true" box="[199,315,942,968]" pageId="20" pageNumber="21">Remarks.</emphasis>
Resolving the name of the comparison species
<taxonomicName id="2E824D69AC4D436F0854FC648DAF0589" authorityName="Warncke" authorityYear="1967" box="[832,970,942,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0854FC648DAF0589" box="[832,970,942,969]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
is more challenging than it appears.
<bibRefCitation id="8D134B1BAC4D436F0E4CFC658F7E05AD" author="Friese, H." pageId="20" pageNumber="21" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese (1922: 216)</bibRefCitation>
described
<taxonomicName id="2E824D69AC4D436F0A87FC188C4005AD" authorityName="Friese" authorityYear="1922" box="[403,549,978,1005]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaleuca">
<emphasis id="DBF6EAF8AC4D436F0A87FC188C4005AD" box="[403,549,978,1005]" italics="true" pageId="20" pageNumber="21">A. melaleuca</emphasis>
</taxonomicName>
[nec. Pérez, 1895] based on “
<specimenCount id="FF84FD63AC4D436F0862FC198DEC05AD" box="[886,905,978,1004]" count="1" pageId="20" pageNumber="21" type="female"></specimenCount>
von
<collectingCountry id="9195767AAC4D436F08ABFC198A4405AD" box="[959,1057,978,1004]" name="Algeria" pageId="20" pageNumber="21">Algerien</collectingCountry>
und
<collectingCountry id="9195767AAC4D436F0F43FC198AFE05AD" box="[1111,1179,978,1004]" name="Tunisia" pageId="20" pageNumber="21">Tunis</collectingCountry>
”. Friese notes “olim
<taxonomicName id="2E824D69AC4D436F0E90FC188E940251" baseAuthorityName="Robertson" baseAuthorityYear="1902" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="DBF6EAF8AC4D436F0E90FC188E940251" italics="true" pageId="20" pageNumber="21">A. obscura</emphasis>
</taxonomicName>
1921”, a name he was using but which he never published.
<bibRefCitation id="8D134B1BAC4D436F0886FC3D8AE50251" author="Warncke, K." box="[914,1152,1014,1040]" pageId="20" pageNumber="21" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke (1967: 201)</bibRefCitation>
correctly recognised that the name
<taxonomicName id="2E824D69AC4D436F0A15FBD08FF70275" authorityName="Friese" authorityYear="1922" box="[257,402,1050,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melaleuca">
<emphasis id="DBF6EAF8AC4D436F0A15FBD08FF70275" box="[257,402,1050,1077]" italics="true" pageId="20" pageNumber="21">A. melaleuca</emphasis>
</taxonomicName>
was preoccupied, and created the replacement name
<taxonomicName id="2E824D69AC4D436F08C8FBD08A020275" authorityName="Warncke" authorityYear="1967" box="[988,1127,1050,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F08C8FBD08A020275" box="[988,1127,1050,1077]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
. Examination of material in the ZMHB and DEI collections produced three potentially syntypic specimens of
<taxonomicName id="2E824D69AC4D436F0F37FBF48ACA0219" authorityName="Warncke" authorityYear="1967" box="[1059,1199,1086,1113]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0F37FBF48ACA0219" box="[1059,1199,1086,1113]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
,
<specimenCount id="FF84FD63AC4D436F0FA8FBF48B260219" box="[1212,1347,1086,1113]" count="2" pageId="20" pageNumber="21" type="female">two females</specimenCount>
(
<figureCitation id="71B92A6FAC4D436F0E45FBF58ED7023D" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="20" pageNumber="21">Figure 11</figureCitation>
) and
<specimenCount id="FF84FD63AC4D436F0BE1FBA88F39023D" box="[245,348,1122,1149]" count="1" pageId="20" pageNumber="21" type="male">one male</specimenCount>
(
<figureCitation id="71B92A6FAC4D436F0A78FBA98FBA023D" box="[364,479,1122,1148]" captionStart="FIGURE 12" captionStartId="21.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@21.[151,1436,181,1624]" captionTargetPageId="21" captionText="FIGURE 12. Andrena (Avandrena) melaleuca Friese, 1922 nec. Provancher = A. melacana Warncke, 1967 male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, dorsal view alternate." figureDoi="http://doi.org/10.5281/zenodo.13209331" httpUri="https://zenodo.org/record/13209331/files/figure.png" pageId="20" pageNumber="21">Figure 12</figureCitation>
). A male and female specimen in the ZMHB collection are both marked with labels “
<taxonomicName id="2E824D69AC4D436F0BB0FB4D8F0002E1" baseAuthorityName="Robertson" baseAuthorityYear="1902" box="[164,357,1158,1184]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="obscura">
<emphasis id="DBF6EAF8AC4D436F0BB0FB4D8F0002E1" box="[164,357,1158,1184]" italics="true" pageId="20" pageNumber="21">Andrena obscura</emphasis>
</taxonomicName>
” in Frieses handwriting. However, only the male bears a type label (
<figureCitation id="71B92A6FAC4D436F0F6FFB4D8B6402E1" box="[1147,1281,1158,1184]" captionStart="FIGURE 12" captionStartId="21.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@21.[151,1436,181,1624]" captionTargetPageId="21" captionText="FIGURE 12. Andrena (Avandrena) melaleuca Friese, 1922 nec. Provancher = A. melacana Warncke, 1967 male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, dorsal view; F. Genital capsule, dorsal view alternate." figureDoi="http://doi.org/10.5281/zenodo.13209331" httpUri="https://zenodo.org/record/13209331/files/figure.png" pageId="20" pageNumber="21">Figure 12A</figureCitation>
), whereas the publication of
<bibRefCitation id="8D134B1BAC4D436F0A2EFB618FB50285" author="Friese, H." box="[314,464,1194,1220]" pageId="20" pageNumber="21" pagination="209 - 217" refId="ref88734" refString="Friese, H. (1922) Neue Formen der Bienengattung Andrena. (Hym.). Konowia, 1, 209 - 217." type="journal article" year="1922">Friese (1922)</bibRefCitation>
indicates that the species was described in the female sex only. This failure to mention the male specimen is likely a lapsus by Friese when drafting the paper. In any case, the female in the ZMHB collection is selected as a
<typeStatus id="36398848AC4D436F0AD0FB398C4B034D" box="[452,558,1266,1292]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
, by present designation. The
<emphasis id="DBF6EAF8AC4D436F086BFB398A76034D" box="[895,1043,1266,1292]" italics="true" pageId="20" pageNumber="21">locus typicus</emphasis>
is also ambiguous, because labels are marked with a printed “
<collectingCountry id="9195767AAC4D436F0ADBFADD8C750371" box="[463,528,1302,1328]" name="Tunisia" pageId="20" pageNumber="21">Tunis</collectingCountry>
merid.”, but Friese has written “Alg” [=Algers] on the side of the label (
<figureCitation id="71B92A6FAC4D436F0E45FADD8EAF0315" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="20" pageNumber="21">Figure 11A</figureCitation>
). It is therefore impossible to establish whether these specimens come from
<collectingCountry id="9195767AAC4D436F0F08FAF18A150315" box="[1052,1136,1338,1364]" name="Algeria" pageId="20" pageNumber="21">Algeria</collectingCountry>
or
<collectingCountry id="9195767AAC4D436F0F81FAF18A890315" box="[1173,1260,1338,1364]" name="Tunisia" pageId="20" pageNumber="21">Tunisia</collectingCountry>
. For the sake of convenience, it is assumed that they come from
<collectingCountry id="9195767AAC4D436F09A6FA958D6C0339" box="[690,777,1374,1400]" name="Tunisia" pageId="20" pageNumber="21">Tunisia</collectingCountry>
.
</paragraph>
<paragraph id="E93D36EAAC4D436F0BD3FA488FC30049" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
In this context, the specimens reported from southern
<collectingCountry id="9195767AAC4D436F0833FA498D0203DD" box="[807,871,1410,1436]" name="Spain" pageId="20" pageNumber="21">Spain</collectingCountry>
by
<bibRefCitation id="8D134B1BAC4D436F0885FA498A8F03DD" author="Wood, T. J. &amp; Ortiz-Sanchez, F. J." box="[913,1258,1410,1436]" pageId="20" pageNumber="21" pagination="114 - 123" refId="ref92242" refString="Wood, T. J. &amp; Ortiz-Sanchez, F. J. (2022) Description of three new Andrena Fabricius, 1775 species from understudied parts of Iberia (Hymenoptera: Andrenidae). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 70, 114 - 123." type="book chapter" year="2022">Wood &amp; Ortiz-Sánchez (2022)</bibRefCitation>
as
<taxonomicName id="2E824D69AC4D436F0E04FA488BF903DD" authorityName="Warncke" authorityYear="1967" box="[1296,1436,1410,1437]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0E04FA488BF903DD" box="[1296,1436,1410,1437]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
probably all belong to
<taxonomicName id="2E824D69AC4D436F0A8FFA6C8C330381" authorityName="WOOD" authorityYear="2024" box="[411,598,1446,1473]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0A8FFA6C8C330381" box="[411,598,1446,1473]" italics="true" pageId="20" pageNumber="21">A. melacanoides</emphasis>
</taxonomicName>
, though this must be confirmed through re-examination of all specimens.
<taxonomicName id="2E824D69AC4D436F0B83FA018FF203A5" authorityName="WOOD" authorityYear="2024" box="[151,407,1482,1508]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacanoides">
<emphasis id="DBF6EAF8AC4D436F0B83FA018FF203A5" box="[151,407,1482,1508]" italics="true" pageId="20" pageNumber="21">Andrena melacanoides</emphasis>
</taxonomicName>
is broadly oligolectic on
<taxonomicName id="2E824D69AC4D436F09A7FA018D2503A5" box="[691,832,1482,1508]" class="Magnoliopsida" family="Geraniaceae" kingdom="Plantae" order="Geraniales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="family">Geraniaceae</taxonomicName>
, and possibly narrowly oligolectic on
<taxonomicName id="2E824D69AC4D436F0FFDFA018B2F03A5" box="[1257,1354,1482,1508]" class="Magnoliopsida" family="Geraniaceae" genus="Erodium" kingdom="Plantae" order="Geraniales" pageId="20" pageNumber="21" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8AC4D436F0FFDFA018B2F03A5" box="[1257,1354,1482,1508]" italics="true" pageId="20" pageNumber="21">Erodium</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC4D436F0E4CFA018E850049" author="Wood, T. J." pageId="20" pageNumber="21" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
, as
<taxonomicName id="2E824D69AC4D436F0A1EFA248FF30049" authorityName="Warncke" authorityYear="1967" box="[266,406,1518,1545]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0A1EFA248FF30049" box="[266,406,1518,1545]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4D436F0BD3F9D98B72006D" box="[199,1303,1554,1581]" pageId="20" pageNumber="21" type="etymology">
<paragraph id="E93D36EAAC4D436F0BD3F9D98B72006D" blockId="20.[151,1437,150,1907]" box="[199,1303,1554,1581]" pageId="20" pageNumber="21">
<emphasis id="DBF6EAF8AC4D436F0BD3F9D98F2B006D" bold="true" box="[199,334,1554,1580]" pageId="20" pageNumber="21">Etymology.</emphasis>
From the name
<taxonomicName id="2E824D69AC4D436F0911F9D98C17006D" authorityName="Warncke" authorityYear="1967" box="[517,626,1554,1580]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0911F9D98C17006D" box="[517,626,1554,1580]" italics="true" pageId="20" pageNumber="21">melacana</emphasis>
</taxonomicName>
combined with the suffix
<emphasis id="DBF6EAF8AC4D436F088CF9D98DB8006D" box="[920,989,1554,1580]" italics="true" pageId="20" pageNumber="21">-oides</emphasis>
, indicating close proximity.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4D436F0BD3F9FD8E940132" pageId="20" pageNumber="21" type="distribution">
<paragraph id="E93D36EAAC4D436F0BD3F9FD8D830035" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
<emphasis id="DBF6EAF8AC4D436F0BD3F9FD8F040011" bold="true" box="[199,353,1590,1616]" pageId="20" pageNumber="21">Distribution.</emphasis>
Southern Spain (Cádiz and Granada, probably also Albacete and Málaga, see
<bibRefCitation id="8D134B1BAC4D436F0FFAF9FD8F230035" author="Wood, T. J. &amp; Ortiz-Sanchez, F. J." pageId="20" pageNumber="21" pagination="114 - 123" refId="ref92242" refString="Wood, T. J. &amp; Ortiz-Sanchez, F. J. (2022) Description of three new Andrena Fabricius, 1775 species from understudied parts of Iberia (Hymenoptera: Andrenidae). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 70, 114 - 123." type="book chapter" year="2022">Wood &amp; Ortiz-Sánchez (2022)</bibRefCitation>
for additional Spanish specimens reported as
<taxonomicName id="2E824D69AC4D436F085EF9908DB30035" authorityName="Warncke" authorityYear="1967" box="[842,982,1626,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F085EF9908DB30035" box="[842,982,1626,1653]" italics="true" pageId="20" pageNumber="21">A. melacana</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="E93D36EAAC4D436F0BD3F9B58E940132" blockId="20.[151,1437,150,1907]" pageId="20" pageNumber="21">
<materialsCitation id="59EA3CB7AC4D436F0BD3F9B58F3600FD" collectingDate="1910-04-01" collectingDateMax="1910-04-30" collectingDateMin="1910-04-01" collectorName="O. Schmiedeknecht" country="Algeria" location="Hammam Bou Hadjar" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2">
<emphasis id="DBF6EAF8AC4D436F0BD3F9B58F9100D9" bold="true" box="[199,500,1662,1688]" pageId="20" pageNumber="21">
<typeStatus id="36398848AC4D436F0BD3F9B58F9100D9" box="[199,500,1662,1688]" isOtherTypeStatus="true" pageId="20" pageNumber="21">Other material examined</typeStatus>
</emphasis>
(
<taxonomicName id="2E824D69AC4D436F0911F9B58CBD00D9" authorityName="Warncke" authorityYear="1967" box="[517,728,1662,1688]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4D436F0911F9B58CBD00D9" box="[517,728,1662,1688]" italics="true" pageId="20" pageNumber="21">Andrena melacana</emphasis>
</taxonomicName>
)
<emphasis id="DBF6EAF8AC4D436F09FCF9B58D1100D9" bold="true" box="[744,884,1662,1688]" pageId="20" pageNumber="21">
<collectingCountry id="9195767AAC4D436F09FCF9B58D0A00D9" box="[744,879,1662,1688]" name="Algeria" pageId="20" pageNumber="21">ALGERIA</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC4D436F086FF9B48A1000D9" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F086FF9B48A1000D9" box="[891,1141,1662,1689]" country="Algeria" name="Hammam Bou Hadjar" pageId="20" pageNumber="21">Hammam Bou Hadjar</location>
,
<date id="9D3C102AAC4D436F0F94F9B58B7100D9" box="[1152,1300,1662,1688]" pageId="20" pageNumber="21" value="1910-04-01" valueMax="1910-04-30" valueMin="1910-04-01">
<collectingDate id="8D78E9C2AC4D436F0F94F9B58B7100D9" box="[1152,1300,1662,1688]" pageId="20" pageNumber="21" value="1910-04-01" valueMax="1910-04-30" valueMin="1910-04-01">130.iv.1910</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F0E34F9B58B2100D9" box="[1312,1348,1662,1688]" count="2" pageId="20" pageNumber="21" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0E95F9B58F3600FD" pageId="20" pageNumber="21">O. Schmiedeknecht</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC4D436F0A48F9698AF700FD" box="[348,1170,1698,1725]" collectingDate="1910-04-08" collectorName="O. Schmiedeknecht" country="Algeria" location="Hammam Bou Hadjar" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
SMFD;
<location id="EC5D6031AC4D436F0AA2F9688CCE00FD" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F0AA2F9688CCE00FD" box="[438,683,1698,1725]" country="Algeria" name="Hammam Bou Hadjar" pageId="20" pageNumber="21">Hammam Bou Hadjar</location>
,
<date id="9D3C102AAC4D436F09A0F9698D7A00FD" box="[692,799,1698,1724]" pageId="20" pageNumber="21" value="1910-04-08">
<collectingDate id="8D78E9C2AC4D436F09A0F9698D7A00FD" box="[692,799,1698,1724]" pageId="20" pageNumber="21" value="1910-04-08">8.iv.1910</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F083DF9698D2800FD" box="[809,845,1698,1724]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC4D436F084CF9698D1E00FD" box="[856,891,1698,1724]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F08A1F9698AF700FD" box="[949,1170,1698,1724]" pageId="20" pageNumber="21">O. Schmiedeknecht</collectorName>
</materialsCitation>
, OÖLM;
<materialsCitation id="59EA3CB7AC4D436F0FEEF9698D1900A1" collectingDate="2017-04-14" collectionCode="AMC" collectorName="A. Muller" country="Morocco" elevation="1220" location="Tafraoute" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC4D436F0FEEF9698BF900FD" bold="true" box="[1274,1436,1698,1724]" pageId="20" pageNumber="21">
<collectingCountry id="9195767AAC4D436F0FEEF9698BF200FD" box="[1274,1431,1698,1724]" name="Morocco" pageId="20" pageNumber="21">MOROCCO</collectingCountry>
:
</emphasis>
<quantity id="2E7A9B0FAC4D436F0B83F90D8EBB00A1" box="[151,222,1734,1760]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="km" value="20.0">20 km</quantity>
N
<location id="EC5D6031AC4D436F0A14F90D8F0B00A1" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F0A14F90D8F0B00A1" box="[256,366,1734,1760]" country="Morocco" name="Tafraoute" pageId="20" pageNumber="21">Tafraoute</location>
,
<quantity id="2E7A9B0FAC4D436F0A6DF90D8FB500A0" box="[377,464,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="20" pageNumber="21" unit="m" value="1220.0">
<elevation id="62AFD1D9AC4D436F0A6DF90D8FB500A0" box="[377,464,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="1.22" pageId="20" pageNumber="21" unit="m" value="1220.0">1220 m</elevation>
</quantity>
,
<date id="9D3C102AAC4D436F0AC8F90D8C3100A1" box="[476,596,1734,1760]" pageId="20" pageNumber="21" value="2017-04-14">
<collectingDate id="8D78E9C2AC4D436F0AC8F90D8C3100A1" box="[476,596,1734,1760]" pageId="20" pageNumber="21" value="2017-04-14">14.iv.2017</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F0974F90D8CE600A1" box="[608,643,1734,1760]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F09ABF90D8D5500A1" box="[703,816,1734,1760]" pageId="20" pageNumber="21">A. Müller</collectorName>
,
<collectionCode id="8F93AE2FAC4D436F082DF90D8D1900A1" box="[825,892,1734,1760]" country="USA" httpUri="http://grbio.org/cool/2xyg-zxc2" name="Department of Biologics Research" pageId="20" pageNumber="21">AMC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4D436F089CF90D8BFD00A1" box="[904,1432,1734,1761]" collectingDate="1997-05-09" collectorName="J. Halada" country="Morocco" location="Ifrane" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
<quantity id="2E7A9B0FAC4D436F089CF90D8DAA00A1" box="[904,975,1734,1760]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="km" value="12.0">12 km</quantity>
E of
<location id="EC5D6031AC4D436F0F18F90D8A3700A1" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F0F18F90D8A3700A1" box="[1036,1106,1734,1760]" country="Morocco" name="Ifrane" pageId="20" pageNumber="21">Ifrane</location>
,
<date id="9D3C102AAC4D436F0F49F90D8ADA00A1" box="[1117,1215,1734,1760]" pageId="20" pageNumber="21" value="1997-05-09">
<collectingDate id="8D78E9C2AC4D436F0F49F90D8ADA00A1" box="[1117,1215,1734,1760]" pageId="20" pageNumber="21" value="1997-05-09">9.v.1997</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F0FD8F90D8A8B00A1" box="[1228,1262,1734,1760]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0E39F90C8BFD00A1" box="[1325,1432,1734,1761]" pageId="20" pageNumber="21">J. Halada</collectorName>
</materialsCitation>
, OÖLM;
<materialsCitation id="59EA3CB7AC4D436F0BEEF9218AA30145" box="[250,1222,1770,1797]" collectionCode="ZMHB" country="Tunisia" location="Tunis" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tunis">
<emphasis id="DBF6EAF8AC4D436F0BEEF9218F1D0145" bold="true" box="[250,376,1770,1796]" pageId="20" pageNumber="21">
<collectingCountry id="9195767AAC4D436F0BEEF9218F160145" box="[250,371,1770,1796]" name="Tunisia" pageId="20" pageNumber="21">TUNISIA</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC4D436F0A95F9218FA50145" box="[385,448,1770,1796]" country="Tunisia" name="Tunis" pageId="20" pageNumber="21">Tunis</collectingRegion>
merid.,
<specimenCount id="FF84FD63AC4D436F0935F9218C200145" box="[545,581,1770,1796]" count="1" pageId="20" pageNumber="21" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC4D436F0940F9218C120145" box="[596,631,1770,1796]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FAC4D436F0992F9208CB20144" box="[646,727,1771,1797]" pageId="20" pageNumber="21">ZMHB</collectionCode>
(female
<emphasis id="DBF6EAF8AC4D436F082AF9218DCC0145" bold="true" box="[830,937,1770,1796]" pageId="20" pageNumber="21">
<typeStatus id="36398848AC4D436F082AF9218DCC0145" box="[830,937,1770,1796]" pageId="20" pageNumber="21" type="lectotype">lectotype</typeStatus>
</emphasis>
, by present designation)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4D436F0FC7F9218F250169" collectionCode="DEI" country="Tunisia" location="Tunis" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Tunis">
<collectingRegion id="2B46F808AC4D436F0FC7F9218B770145" box="[1235,1298,1770,1796]" country="Tunisia" name="Tunis" pageId="20" pageNumber="21">Tunis</collectingRegion>
merid.,
<specimenCount id="FF84FD63AC4D436F0E66F9218BF00145" box="[1394,1429,1770,1796]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FAC4D436F0B83F8C48EA30168" box="[151,198,1807,1833]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="20" pageNumber="21" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<emphasis id="DBF6EAF8AC4D436F0BCDF8C58F500169" bold="true" box="[217,309,1806,1832]" pageId="20" pageNumber="21">
<typeStatus id="36398848AC4D436F0BCDF8C58F500169" box="[217,309,1806,1832]" pageId="20" pageNumber="21" type="syntype">syntype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4D436F0A5BF8C58C830169" box="[335,742,1806,1832]" collectorName="O. Schmiedeknecht" country="Tunisia" location="Tunis" pageId="20" pageNumber="21" specimenCount="2" specimenCount-female="2" stateProvince="Tunis">
<collectingRegion id="2B46F808AC4D436F0A5BF8C58FF40169" box="[335,401,1806,1832]" country="Tunisia" name="Tunis" pageId="20" pageNumber="21">Tunis</collectingRegion>
,
<specimenCount id="FF84FD63AC4D436F0A8BF8C58FA60169" box="[415,451,1806,1832]" count="2" pageId="20" pageNumber="21" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0911F8C58C830169" box="[517,742,1806,1832]" pageId="20" pageNumber="21">O. Schmiedeknecht</collectorName>
</materialsCitation>
, OÖLM;
<materialsCitation id="59EA3CB7AC4D436F084CF8C58A900168" box="[856,1269,1806,1833]" collectingDate="1927-04-06" collectorName="R. Meyer" country="Tunisia" location="Tunis" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Tunis">
<collectingRegion id="2B46F808AC4D436F084CF8C58DFF0169" box="[856,922,1806,1832]" country="Tunisia" name="Tunis" pageId="20" pageNumber="21">Tunis</collectingRegion>
,
<date id="9D3C102AAC4D436F08B3F8C58A770169" box="[935,1042,1806,1832]" pageId="20" pageNumber="21" value="1927-04-06">
<collectingDate id="8D78E9C2AC4D436F08B3F8C58A770169" box="[935,1042,1806,1832]" pageId="20" pageNumber="21" value="1927-04-06">6.iv.1927</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F0F34F8C58A260169" box="[1056,1091,1806,1832]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0F93F8C48A900168" box="[1159,1269,1807,1833]" pageId="20" pageNumber="21">R. Meyer</collectorName>
</materialsCitation>
, OÖLM; Gov.
<materialsCitation id="59EA3CB7AC4D436F0B83F8F98D5E010D" box="[151,827,1842,1869]" collectingDate="2018-04-01" collectionCode="TJWC" collectorName="H. Zettel" country="Tunisia" elevation="5" location="Skanes" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1" stateProvince="Monastir">
<collectingRegion id="2B46F808AC4D436F0B83F8F98E9B010D" box="[151,254,1842,1868]" country="Tunisia" name="Monastir" pageId="20" pageNumber="21">Monastir</collectingRegion>
,
<location id="EC5D6031AC4D436F0A1EF8F98F39010D" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F0A1EF8F98F39010D" box="[266,348,1842,1868]" country="Tunisia" name="Skanes" pageId="20" pageNumber="21" stateProvince="Monastir">Skanes</location>
,
<quantity id="2E7A9B0FAC4D436F0A7CF8F98FF3010C" box="[360,406,1842,1869]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="5.0">
<elevation id="62AFD1D9AC4D436F0A7CF8F98FF3010C" box="[360,406,1842,1869]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="21" unit="m" value="5.0">5 m</elevation>
</quantity>
,
<date id="9D3C102AAC4D436F0AB6F8F98C68010D" box="[418,525,1842,1868]" pageId="20" pageNumber="21" value="2018-04-01">
<collectingDate id="8D78E9C2AC4D436F0AB6F8F98C68010D" box="[418,525,1842,1868]" pageId="20" pageNumber="21" value="2018-04-01">1.iv.2018</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F090DF8F98C59010D" box="[537,572,1842,1868]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0968F8F88C86010D" box="[636,739,1842,1869]" pageId="20" pageNumber="21">H. Zettel</collectorName>
,
<collectionCode id="8F93AE2FAC4D436F09FAF8F98D5E010D" box="[750,827,1842,1868]" pageId="20" pageNumber="21">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC4D436F0853F8F98BFD010D" box="[839,1432,1842,1869]" collectingDate="1976-03-25" collectorName="P. Robinson" country="Tunisia" location="Testour" pageId="20" pageNumber="21" specimenCount="1" specimenCount-female="1">
<quantity id="2E7A9B0FAC4D436F0853F8F98DEA010D" box="[839,911,1842,1868]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="km" value="10.0">10 km</quantity>
NE
<location id="EC5D6031AC4D436F08D7F8F88A7D010C" LSID="urn:lsid:plazi:treatment:612B87FCAC49436F0B83FE358E940132:EC5D6031AC4D436F08D7F8F88A7D010C" box="[963,1048,1843,1869]" country="Tunisia" name="Testour" pageId="20" pageNumber="21">Testour</location>
,
<date id="9D3C102AAC4D436F0F30F8F98AC5010C" box="[1060,1184,1842,1869]" pageId="20" pageNumber="21" value="1976-03-25">
<collectingDate id="8D78E9C2AC4D436F0F30F8F98AC5010C" box="[1060,1184,1842,1869]" pageId="20" pageNumber="21" value="1976-03-25">25.iii.1976</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC4D436F0FB8F8F98AAA010D" box="[1196,1231,1842,1868]" count="1" pageId="20" pageNumber="21" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC4D436F0E1BF8F88BFD010D" box="[1295,1432,1842,1869]" pageId="20" pageNumber="21">P. Robinson</collectorName>
</materialsCitation>
, OÖLM.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,507 @@
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<mods:title id="0C1DA725D1EF6098A4C3CBEB5F25AD1E">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<treatment id="612B87FCAC4C43630B83F8E98BF904C2" LSID="urn:lsid:plazi:treatment:612B87FCAC4C43630B83F8E98BF904C2" httpUri="http://treatment.plazi.org/id/612B87FCAC4C43630B83F8E98BF904C2" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="A1986561AC4C436E0B83F8E98C3B017D" box="[151,606,1825,1852]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="E93D36EAAC4C436E0B83F8E98C3B017D" blockId="21.[151,1436,1825,1996]" box="[151,606,1825,1852]" pageId="21" pageNumber="22">
<emphasis id="DBF6EAF8AC4C436E0B83F8E98C3B017D" bold="true" box="[151,606,1825,1852]" pageId="21" pageNumber="22">
Key to
<taxonomicName id="2E824D69AC4C436E0BE5F8EA8F9C017A" authority="Warncke, 1968" box="[241,505,1825,1852]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subGenus" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4C436E0BE5F8EA8F3E017D" bold="true" box="[241,347,1825,1852]" italics="true" pageId="21" pageNumber="22">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4C436E0A78F8EA8F8A017D" bold="true" box="[364,495,1825,1852]" italics="true" pageId="21" pageNumber="22">Avandrena</emphasis>
)
</taxonomicName>
females
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC4C436E0B83F88D8CAB018D" pageId="21" pageNumber="22" type="description">
<paragraph id="E93D36EAAC4C436E0B83F88D8CAB018D" blockId="21.[151,1436,1825,1996]" pageId="21" pageNumber="22">
Note, the female of
<taxonomicName id="2E824D69AC4C436E0A6EF88D8FBD0121" ID-CoL="5FLJ2" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[378,472,1862,1888]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="gavia">
<emphasis id="DBF6EAF8AC4C436E0A6EF88D8FBD0121" box="[378,472,1862,1888]" italics="true" pageId="21" pageNumber="22">A. gavia</emphasis>
</taxonomicName>
is unknown, and is therefore not included in the key. Care should be taken in central
<collectingCountry id="9195767AAC4C436E0B83F8A18EB201C5" box="[151,215,1898,1924]" name="Spain" pageId="21" pageNumber="22">Spain</collectingCountry>
to ensure that material conforms to
<taxonomicName id="2E824D69AC4C436E0963F8A18C8E01C5" ID-CoL="5FLJ3" authorityName="Warncke" authorityYear="1975" box="[631,747,1898,1924]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="DBF6EAF8AC4C436E0963F8A18C8E01C5" box="[631,747,1898,1924]" italics="true" pageId="21" pageNumber="22">A. liturata</emphasis>
</taxonomicName>
; if the tergal margins are strongly depressed, it is likely that it belongs to
<taxonomicName id="2E824D69AC4C436E0A3EF8458FE201E9" ID-CoL="5FLJ2" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[298,391,1934,1960]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="gavia">
<emphasis id="DBF6EAF8AC4C436E0A3EF8458FE201E9" box="[298,391,1934,1960]" italics="true" pageId="21" pageNumber="22">A. gavia</emphasis>
</taxonomicName>
, but this should be established. For additional illustrations of
<taxonomicName id="2E824D69AC4C436E0F2BF8458AB101E9" authorityName="Wood &amp; Ortiz-Sanchez" authorityYear="2022" box="[1087,1236,1934,1960]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="erodiorum">
<emphasis id="DBF6EAF8AC4C436E0F2BF8458AB101E9" box="[1087,1236,1934,1960]" italics="true" pageId="21" pageNumber="22">A. erodiorum</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC4C436E0E1FF8458B0F01E9" authorityName="Wood" authorityYear="2023" box="[1291,1386,1934,1960]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="juliae">
<emphasis id="DBF6EAF8AC4C436E0E1FF8458B0F01E9" box="[1291,1386,1934,1960]" italics="true" pageId="21" pageNumber="22">A. juliae</emphasis>
</taxonomicName>
, see
<bibRefCitation id="8D134B1BAC4C436E0B83F8798F8A018D" author="Wood, T. J. &amp; Ortiz-Sanchez, F. J." box="[151,495,1970,1996]" pageId="21" pageNumber="22" pagination="114 - 123" refId="ref92242" refString="Wood, T. J. &amp; Ortiz-Sanchez, F. J. (2022) Description of three new Andrena Fabricius, 1775 species from understudied parts of Iberia (Hymenoptera: Andrenidae). Boletin de la Sociedad Entomologica Aragonesa (S. E. A.), 70, 114 - 123." type="book chapter" year="2022">Wood &amp; Ortiz-Sánchez (2022)</bibRefCitation>
and
<bibRefCitation id="8D134B1BAC4C436E0931F8798CAF018D" author="Wood, T. J." box="[549,714,1970,1996]" pageId="21" pageNumber="22" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood (2023b)</bibRefCitation>
.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC4F436D0B83F8BA8C5801B7" ID-DOI="http://doi.org/10.5281/zenodo.13209335" ID-Zenodo-Dep="13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1905,1929]" targetBox="[214,1373,151,1889]" targetPageId="22" targetType="figure">
<paragraph id="E93D36EAAC4F436D0B83F8BA8C5801B7" blockId="22.[151,1437,1905,2038]" pageId="22" pageNumber="23">
<emphasis id="DBF6EAF8AC4F436D0B83F8BA8F4601C8" bold="true" box="[151,291,1905,1929]" pageId="22" pageNumber="23">FIGURE 13.</emphasis>
<taxonomicName id="2E824D69AC4F436D0A39F8B98D6501CB" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[301,768,1906,1930]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="eureka" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4F436D0A39F8B98FE601CB" box="[301,387,1906,1930]" italics="true" pageId="22" pageNumber="23">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4F436D0A82F8B98C6601CB" box="[406,515,1906,1930]" italics="true" pageId="22" pageNumber="23">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4F436D0901F8B98C3C01CB" box="[533,601,1906,1930]" italics="true" pageId="22" pageNumber="23">eureka</emphasis>
<bibRefCitation id="8D134B1BAC4F436D0977F8B98D6501CB" author="Warncke, K." box="[611,768,1906,1930]" pageId="22" pageNumber="23" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
female. A. Head, lateral view; B. Terga, dorsal view;
<taxonomicName id="2E824D69AC4F436D0E52F8B98C6C01EF" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="caligata" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4F436D0E52F8B98BF901CB" box="[1350,1436,1906,1930]" italics="true" pageId="22" pageNumber="23">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4F436D0BB4F85D8F6801EF" box="[160,269,1942,1966]" italics="true" pageId="22" pageNumber="23">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4F436D0A0EF85D8F0801EF" box="[282,365,1942,1966]" italics="true" pageId="22" pageNumber="23">caligata</emphasis>
<bibRefCitation id="8D134B1BAC4F436D0A66F85D8C6C01EF" author="Warncke, K." box="[370,521,1942,1966]" pageId="22" pageNumber="23" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
female. C. Head, lateral view; D. Terga, dorsal view;
<taxonomicName id="2E824D69AC4F436D0F0BF85D8EAF0193" authority="Warncke, 1967" authorityName="Warncke" authorityYear="1967" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" sensu="stricto" species="avara" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4F436D0F0BF85D8A1001EF" box="[1055,1141,1942,1966]" italics="true" pageId="22" pageNumber="23">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4F436D0F97F85D8A9501EF" box="[1155,1264,1942,1966]" italics="true" pageId="22" pageNumber="23">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4F436D0FE9F85D8B5D01EC" box="[1277,1336,1942,1965]" italics="true" pageId="22" pageNumber="23">avara</emphasis>
<bibRefCitation id="8D134B1BAC4F436D0E29F85D8EAF0193" author="Warncke, K." pageId="22" pageNumber="23" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke, 1967</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC4F436D0BC4F8718F2A0190" box="[208,335,1978,2001]" italics="true" pageId="22" pageNumber="23">
<taxonomicNameLabel id="C0C55783AC4F436D0BC4F8718F2A0190" box="[208,335,1978,2001]" pageId="22" pageNumber="23" sensu="stricto">sensu stricto</taxonomicNameLabel>
</emphasis>
female. E. Head, lateral view; F. Terga, dorsal view;
<taxonomicName id="2E824D69AC4F436D0870F8718B480193" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[868,1325,1978,2002]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="liturata" subGenus="Avandrena">
<emphasis id="DBF6EAF8AC4F436D0870F8718DDF0193" box="[868,954,1978,2002]" italics="true" pageId="22" pageNumber="23">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC4F436D08DCF8718A500193" box="[968,1077,1978,2002]" italics="true" pageId="22" pageNumber="23">Avandrena</emphasis>
)
<emphasis id="DBF6EAF8AC4F436D0F57F8718AEA0193" box="[1091,1167,1978,2002]" italics="true" pageId="22" pageNumber="23">liturata</emphasis>
<bibRefCitation id="8D134B1BAC4F436D0F81F8718B480193" author="Warncke, K." box="[1173,1325,1978,2002]" pageId="22" pageNumber="23" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
female. G. Head, lateral view; H. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC4E43630B83FF538BF904C2" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="key">
<paragraph id="E93D36EAAC4E436C0B83FF538BF9068B" blockId="23.[151,1437,151,437]" pageId="23" pageNumber="24">1. Posterior face of the hind femur with a row of long apically pointed spines, clearly distinct against whatever hairs may be present............................................................................................. 2</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FF188BF906A6" blockId="23.[151,1437,151,437]" box="[151,1436,210,232]" pageId="23" pageNumber="24">- Posterior face of the hind femur without spines, with only scattered hairs........................................ 8</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FF3B8BF90744" blockId="23.[151,1437,151,437]" box="[151,1436,239,261]" pageId="23" pageNumber="24">2. Face with entirely black pubescence, mesosoma with abundant black hairs....................................... 3</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FEC68BF9077E" blockId="23.[151,1437,151,437]" pageId="23" pageNumber="24">
- Face never with entirely black pubescence, with at least some pale hairs medially (
<figureCitation id="71B92A6FAC4E436C0F16FEC68A150763" box="[1026,1136,269,291]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="23" pageNumber="24">Figures 1C</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0F68FEC68AFF0763" box="[1148,1178,269,290]" captionStart="FIGURE 4" captionStartId="9.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-79@9.[151,1436,309,1752]" captionTargetPageId="9" captionText="FIGURE 4. Andrena (Avandrena) baldocki spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Face, anterolateral view; D. Scutum, dorsal view; E. Propodeal triangle, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209315" httpUri="https://zenodo.org/record/13209315/files/figure.png" pageId="23" pageNumber="24">4C</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0FB1FEC68B400763" box="[1189,1317,269,290]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13A; C; E; F</figureCitation>
), mesosoma without abundant black hairs, mesepisternum predominantly pale-haired (
<figureCitation id="71B92A6FAC4E436C08B2FEE18A69077E" box="[934,1036,298,320]" captionStart="FIGURE 4" captionStartId="9.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-79@9.[151,1436,309,1752]" captionTargetPageId="9" captionText="FIGURE 4. Andrena (Avandrena) baldocki spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Face, anterolateral view; D. Scutum, dorsal view; E. Propodeal triangle, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209315" httpUri="https://zenodo.org/record/13209315/files/figure.png" pageId="23" pageNumber="24">Figure 4A</figureCitation>
;
<taxonomicName id="2E824D69AC4E436C0F02FEE08A2B0701" ID-CoL="DR3X" authorityName="Warncke" authorityYear="1967" box="[1046,1102,299,320]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="avara">
<emphasis id="DBF6EAF8AC4E436C0F02FEE08A2B0701" box="[1046,1102,299,320]" italics="true" pageId="23" pageNumber="24">avara</emphasis>
</taxonomicName>
-group).................... 4
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FE838BF9073B" blockId="23.[151,1437,151,437]" pageId="23" pageNumber="24">
3. Process of the labrum very narrow, triangular with a pointed apex. Posterior face of hind femur with 46 spines.......................................
<taxonomicName id="2E824D69AC4E436C0923FEAE8D72073A" ID-CoL="DSJP" authority="De Steffani" authorityName="De Steffani" box="[567,791,357,379]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="panurgina">
<emphasis id="DBF6EAF8AC4E436C0923FEAE8D72073A" bold="true" box="[567,791,357,379]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0923FEAE8CFA073B" bold="true" box="[567,671,357,378]" italics="true" pageId="23" pageNumber="24">panurgina</emphasis>
De Steffani
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0837FEAE8D1D073A" box="[803,888,357,379]" name="Portugal" pageId="23" pageNumber="24">Portugal</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0896FEAE8DDE073A" box="[898,955,357,379]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C08D1FEAE8A6C073B" box="[965,1033,357,378]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0F06FEAE8A0E073B" box="[1042,1131,357,378]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0F67FEAE8AD8073A" box="[1139,1213,357,379]" name="Algeria" pageId="23" pageNumber="24">Algeria</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0FD3FEAE8B74073A" box="[1223,1297,357,379]" name="Tunisia" pageId="23" pageNumber="24">Tunisia</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0E0EFEAE8B22073A" box="[1306,1351,357,379]" name="Italy" pageId="23" pageNumber="24">Italy</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0E44FEAE8BF3073B" box="[1360,1430,357,378]" name="Greece" pageId="23" pageNumber="24">Greece</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FE488BF807F4" blockId="23.[151,1437,151,437]" pageId="23" pageNumber="24">
- Process of the labrum broad, rounded rectangular, 4 times wider than long. Posterior face of hind femur with 912 spines.............................................................................
<taxonomicName id="2E824D69AC4E436C0F7CFE548B7307F4" ID-CoL="66MRX" authority="Perez" authorityName="Perez" authorityYear="1903" box="[1128,1302,415,437]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="heterodoxa">
<emphasis id="DBF6EAF8AC4E436C0F7CFE548B7307F4" bold="true" box="[1128,1302,415,437]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0F7CFE548AB207F4" bold="true" box="[1128,1239,415,437]" italics="true" pageId="23" pageNumber="24">heterodoxa</emphasis>
Pérez
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0E37FE548B3107F4" box="[1315,1364,415,437]" name="Italy" pageId="23" pageNumber="24">Italy</collectingCountry>
:
<collectingRegion id="2B46F808AC4E436C0E48FE548BF007F4" box="[1372,1429,415,437]" country="Italy" name="Sicilia" pageId="23" pageNumber="24">Sicily</collectingRegion>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FE168BFF045A" blockId="23.[151,1436,477,540]" pageId="23" pageNumber="24">
The condition of specimens is critical in identifying members of the
<taxonomicName id="2E824D69AC4E436C088EFE158DBE07B6" ID-CoL="DR3X" authorityName="Warncke" authorityYear="1967" box="[922,987,478,503]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="avara">
<emphasis id="DBF6EAF8AC4E436C088EFE158DBE07B6" box="[922,987,478,503]" italics="true" pageId="23" pageNumber="24">avara</emphasis>
</taxonomicName>
-group, since many characters are based on pubescence characters. Particular care should be taken with abraded specimens or specimens with matted hairs.
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FD8B8BF904CE" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
4. Terga with strongly developed apical hairbands, hairbands exceeding length of marginal areas, obscuring underlying surface (
<figureCitation id="71B92A6FAC4E436C0BFCFD968F390433" box="[232,348,605,627]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">Figure 13B</figureCitation>
); tergal discs with short pale hairs, intermixing with apical hairbands. Facial foveae relatively long, ventrally extending to below level of antennal insertions. Terminal fringe golden-orange..
<taxonomicName id="2E824D69AC4E436C08E8FDB18AC604CE" ID-CoL="5FLHZ" authority="Warncke" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[1020,1187,634,656]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="eureka">
<emphasis id="DBF6EAF8AC4E436C08E8FDB18AC604CE" bold="true" box="[1020,1187,634,656]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C08E8FDB18A2504D1" bold="true" box="[1020,1088,634,656]" italics="true" pageId="23" pageNumber="24">eureka</emphasis>
Warncke
</emphasis>
</taxonomicName>
(south-western
<collectingCountry id="9195767AAC4E436C0E2AFDB18BF304CE" box="[1342,1430,634,655]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FD538BF90544" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Terga with comparatively more weakly developed hairbands, hairbands either not exceeding length of marginal areas or weak and not obscuring underlying surface (beware abraded specimens;
<figureCitation id="71B92A6FAC4E436C084FFD7E8DA1048B" box="[859,964,693,715]" captionStart="FIGURE 4" captionStartId="9.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-79@9.[151,1436,309,1752]" captionTargetPageId="9" captionText="FIGURE 4. Andrena (Avandrena) baldocki spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Face, anterolateral view; D. Scutum, dorsal view; E. Propodeal triangle, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209315" httpUri="https://zenodo.org/record/13209315/files/figure.png" pageId="23" pageNumber="24">Figures 4F</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C08D8FD7E8A4B048B" box="[972,1070,693,714]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13D; F; H</figureCitation>
), tergal discs with sparse and scattered hairs, not intermixing with apical hairbands. Facial foveae shorter, terminating at or above level of antennal insertions. Terminal fringe orange-brown to brown........................................................................... 5
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FCC68BF9053A" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
5. Tergal margins strongly depressed, most abruptly so on T34, forming a clear step with tergal discs AND tergal discs with clearly visible punctures that contrast the underlying surface (
<figureCitation id="71B92A6FAC4E436C0851FCE18DCE057E" box="[837,939,810,832]" captionStart="FIGURE 4" captionStartId="9.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-79@9.[151,1436,309,1752]" captionTargetPageId="9" captionText="FIGURE 4. Andrena (Avandrena) baldocki spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Face, anterolateral view; D. Scutum, dorsal view; E. Propodeal triangle, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209315" httpUri="https://zenodo.org/record/13209315/files/figure.png" pageId="23" pageNumber="24">Figure 4F</figureCitation>
). Margins of T34 with loose hairband occupying entire width of margin, hairs not obscuring underlying surface, apical extent of hairbands slightly but distinctly exceeding apex of marginal areas..............................................
<emphasis id="DBF6EAF8AC4E436C08AFFCAF8A12053B" bold="true" box="[955,1143,868,890]" pageId="23" pageNumber="24">
<taxonomicName id="2E824D69AC4E436C08AFFCAF8A6A053B" authorityName="WOOD" authorityYear="2024" box="[955,1039,868,890]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="baldocki" status="sp. nov.">
<emphasis id="DBF6EAF8AC4E436C08AFFCAF8A6A053B" bold="true" box="[955,1039,868,890]" italics="true" pageId="23" pageNumber="24">baldocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC4E436C0F01FCAE8A12053B" box="[1045,1143,869,890]" pageId="23" pageNumber="24" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(southern
<collectingCountry id="9195767AAC4E436C0FCAFCAE8B70053A" box="[1246,1301,869,891]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
and
<collectingCountry id="9195767AAC4E436C0E57FCAE8BF3053A" box="[1347,1430,869,891]" name="Portugal" pageId="23" pageNumber="24">Portugal</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FC488BF90592" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Tergal margins either not strongly depressed (
<figureCitation id="71B92A6FAC4E436C09A4FC498D4605D6" box="[688,803,898,920]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="23" pageNumber="24">Figures 1D</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0826FC498D1905D6" box="[818,892,898,919]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13F; H</figureCitation>
), or if depressed then tergal discs obscurely punctate, punctures small, weak, and disappearing into underlying microsculpture (
<figureCitation id="71B92A6FAC4E436C08BAFC548A4705F4" box="[942,1058,927,949]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">Figure 13D</figureCitation>
). Margins of T34 with hairbands of a different form, either shorter (occupying a smaller proportion of the marginal area) or weaker........................ 6
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FC118BF9026A" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
6. Tergal margins comparatively strongly depressed, most visible on T34, forming a clear step with tergal discs (
<figureCitation id="71B92A6FAC4E436C0E09FC118BF405AE" box="[1309,1425,986,1008]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">Figure 13D</figureCitation>
). Margins of T24 with apical hairbands well-developed, short but dense and obscuring underlying surface.......................................................................
<taxonomicName id="2E824D69AC4E436C0895FBDF8A57026B" ID-CoL="5FLHY" authority="Warncke" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[897,1074,1044,1066]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="caligata">
<emphasis id="DBF6EAF8AC4E436C0895FBDF8A57026B" bold="true" box="[897,1074,1044,1066]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0895FBDF8DB5026B" bold="true" box="[897,976,1044,1066]" italics="true" pageId="23" pageNumber="24">caligata</emphasis>
Warncke
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0F2BFBDE8AEF026A" box="[1087,1162,1045,1067]" name="Tunisia" pageId="23" pageNumber="24">Tunisia</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0F80FBDE8AAB026A" box="[1172,1230,1045,1067]" name="Libya" pageId="23" pageNumber="24">Libya</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0FCCFBDE8B6D026A" box="[1240,1288,1045,1067]" name="Italy" pageId="23" pageNumber="24">Italy</collectingCountry>
:
<collectingRegion id="2B46F808AC4E436C0E05FBDE8B2F026A" box="[1297,1354,1045,1067]" country="Italy" name="Sicilia" pageId="23" pageNumber="24">Sicily</collectingRegion>
,
<collectingCountry id="9195767AAC4E436C0E47FBDE8BE9026A" box="[1363,1420,1045,1067]" name="Israel" pageId="23" pageNumber="24">Israel</collectingCountry>
*)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FBF88BF90224" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Tergal margins weakly depressed, not forming clear step with tergal discs (
<figureCitation id="71B92A6FAC4E436C08A1FBF98A400206" box="[949,1061,1074,1096]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="23" pageNumber="24">Figures 1D</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0F24FBF98A120206" box="[1072,1143,1074,1095]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13F; H</figureCitation>
). Margins of T24 with apical hairbands weakly developed, long or short but not obscuring underlying surface................................... 7
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FBA68BF9029B" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
7. Pubescence dark, face with abundant black and dark brown hairs, with pale hairs restricted to area around antennal insertions (
<figureCitation id="71B92A6FAC4E436C0BFCFB418F3002DE" box="[232,341,1162,1184]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="23" pageNumber="24">Figures 1C</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0A4BFB418FEC02DE" box="[351,393,1162,1183]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13E</figureCitation>
). Terga with light brown to brown pubescence, T24 with narrow apical hairbands, apical extent of hairbands not or only slightly exceeding apex of marginal areas, T24 with apical rims narrowly lightened hyaline-brown (
<figureCitation id="71B92A6FAC4E436C0E3CFB6C8BFD02FC" box="[1320,1432,1191,1213]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="23" pageNumber="24">Figures 1D</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C0BF6FB0E8F6D029B" box="[226,264,1221,1242]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">13F</figureCitation>
).....................................................
<taxonomicName id="2E824D69AC4E436C08B6FB0E8A58029B" ID-CoL="DR3X" authority="Warncke" authorityName="Warncke" authorityYear="1967" box="[930,1085,1221,1242]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="avara">
<emphasis id="DBF6EAF8AC4E436C08B6FB0E8A58029B" bold="true" box="[930,1085,1221,1242]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C08B6FB0E8DBF029B" bold="true" box="[930,986,1221,1242]" italics="true" pageId="23" pageNumber="24">avara</emphasis>
Warncke
</emphasis>
</taxonomicName>
(southern
<collectingCountry id="9195767AAC4E436C0FB0FB0E8AB8029A" box="[1188,1245,1221,1243]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
, northern
<collectingCountry id="9195767AAC4E436C0E2AFB0E8BF3029B" box="[1342,1430,1221,1242]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FB288BF9030E" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Pubescence bright, face with whitish pubescence medially, black hairs restricted to the inner margins of the compound eyes (
<figureCitation id="71B92A6FAC4E436C0BFCFB348F3E0354" box="[232,347,1279,1301]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">Figure 13G</figureCitation>
). Terga with pale pubescence, T24 with long apical hairbands, apical extent of hairbands clearly exceeding apex of marginal areas, T24 with apical rims more broadly lightened hyaline-yellow (
<figureCitation id="71B92A6FAC4E436C08FBFAD68A050373" box="[1007,1120,1309,1331]" captionStart="FIGURE 13" captionStartId="22.[151,250,1905,1929]" captionTargetBox="[214,1373,151,1889]" captionTargetId="figure-28@22.[214,1373,151,1889]" captionTargetPageId="22" captionText="FIGURE 13. Andrena (Avandrena) eureka Warncke, 1974 female. A. Head, lateral view; B. Terga, dorsal view; Andrena (Avandrena) caligata Warncke, 1974 female. C. Head, lateral view; D. Terga, dorsal view; Andrena (Avandrena) avara Warncke, 1967 sensu stricto female. E. Head, lateral view; F. Terga, dorsal view; Andrena (Avandrena) liturata Warncke, 1974 female. G. Head, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209335" httpUri="https://zenodo.org/record/13209335/files/figure.png" pageId="23" pageNumber="24">Figure 13H</figureCitation>
)...............................................................................................
<taxonomicName id="2E824D69AC4E436C0F27FAF18ABA030E" ID-CoL="5FLJ3" authority="Warncke" authorityName="Warncke" authorityYear="1974" baseAuthorityName="Warncke" baseAuthorityYear="1974" box="[1075,1247,1338,1360]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="liturata">
<emphasis id="DBF6EAF8AC4E436C0F27FAF18ABA030E" bold="true" box="[1075,1247,1338,1360]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0F27FAF18A190311" bold="true" box="[1075,1148,1338,1360]" italics="true" pageId="23" pageNumber="24">liturata</emphasis>
Warncke
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0FF8FAF18B410311" box="[1260,1316,1338,1360]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
and
<collectingCountry id="9195767AAC4E436C0E47FAF18BF3030E" box="[1363,1430,1338,1359]" name="France" pageId="23" pageNumber="24">France</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FA938BF903CB" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
8. Mesepisternum entirely black-haired (
<figureCitation id="71B92A6FAC4E436C0947FA9C8CA1032C" box="[595,708,1367,1389]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figures 6A</figureCitation>
;
<figureCitation id="71B92A6FAC4E436C09DAFA938D69032C" box="[718,780,1368,1389]" captionStart="FIGURE 8" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[213,1374,181,1919]" captionTargetId="figure-16@15.[213,1374,181,1919]" captionTargetPageId="15" captionText="FIGURE 8. Andrena (Avandrena) canohirta (Friese, 1923) holotype female.A. Label details; B. Habitus, lateral view; Andrena (Avandrena) balcanica Warncke, 1965 holotype female. C. Label details; D. Habitus, lateral view; E. Face, frontal view; F. Scutum, dorsal view; G. Mesepisternum, lateral view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209323" httpUri="https://zenodo.org/record/13209323/files/figure.png" pageId="23" pageNumber="24">8B; D</figureCitation>
). Process of the labrum narrow (quadrate, slightly wider than long) with deep emargination medially........................................................................ 9
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FA588BF90384" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Mesepisternum never entirely black-haired, usually with abundant pale hairs, at most with 6070% of hairs black (
<figureCitation id="71B92A6FAC4E436C0E3FFA598BF403E6" box="[1323,1425,1426,1448]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="23" pageNumber="24">Figure 9A</figureCitation>
). Process of the labrum variable, usually several times broader than long......................................... 10
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83FA068BF9005C" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
9. Clypeus with shallow but large, round, and clearly visible punctures (
<figureCitation id="71B92A6FAC4E436C0888FA068A6303A2" box="[924,1030,1485,1507]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figure 6A</figureCitation>
). Scutum uniformly shagreened and dull (
<figureCitation id="71B92A6FAC4E436C0BFCFA218F2E0041" box="[232,331,1514,1536]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figure 6F</figureCitation>
). Terminal fringe dark brown. Slightly larger,
<quantity id="2E7A9B0FAC4E436C0812FA218D300041" box="[774,853,1514,1536]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="23" pageNumber="24" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">78 mm</quantity>
.........................................................................................
<taxonomicName id="2E824D69AC4E436C09CEF9CC8DEA005C" ID-CoL="DR8Y" authority="(Friese)" baseAuthorityName="Friese" baseAuthorityYear="1923" box="[730,911,1543,1565]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="canohirta">
<emphasis id="DBF6EAF8AC4E436C09CEF9CC8DEA005C" bold="true" box="[730,911,1543,1565]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C09CEF9CC8D5F005C" bold="true" box="[730,826,1543,1565]" italics="true" pageId="23" pageNumber="24">canohirta</emphasis>
(Friese)
</emphasis>
</taxonomicName>
(North
<collectingCountry id="9195767AAC4E436C08CEF9CC8A22005C" box="[986,1095,1543,1565]" name="Macedonia" pageId="23" pageNumber="24">Macedonia</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0F45F9CC8AC3005C" box="[1105,1190,1543,1565]" name="Bulgaria" pageId="23" pageNumber="24">Bulgaria</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0FA4F9C38A93005C" box="[1200,1270,1544,1565]" name="Greece" pageId="23" pageNumber="24">Greece</collectingCountry>
, western
<collectingCountry id="9195767AAC4E436C0E44F9CC8BF3005C" box="[1360,1430,1543,1565]" name="Turkey" pageId="23" pageNumber="24">Turkey</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F9EE8BF80034" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Clypeus with obscure and latitudinally compressed punctures, disappearing into underlying sculpture of clypeus (
<figureCitation id="71B92A6FAC4E436C0E3FF9EE8BF4007A" box="[1323,1425,1573,1595]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figure 6C</figureCitation>
). Scutum laterally shagreened, becoming polished and shining medially (
<figureCitation id="71B92A6FAC4E436C0898F9898D8A0019" box="[908,1007,1602,1624]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figure 6E</figureCitation>
). Terminal fringe golden-brown (
<figureCitation id="71B92A6FAC4E436C0E38F9898BF40019" box="[1324,1425,1602,1624]" captionStart="FIGURE 6" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@12.[214,1373,181,1919]" captionTargetPageId="12" captionText="FIGURE 6. Andrena (Avandrena) collisa spec. nov. female. A. Habitus, lateral view; B. Terga, dorsal view; C. Face, frontal view; E. Scutum, dorsal view; G. Propodeum, dorsal view; Andrena (Avandrena) canohirta (Friese, 1923) female. D. Face, frontal view; F. Scutum, dorsal view; H. Propodeum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209319" httpUri="https://zenodo.org/record/13209319/files/figure.png" pageId="23" pageNumber="24">Figure 6B</figureCitation>
). Slightly smaller,
<quantity id="2E7A9B0FAC4E436C0A92F9AB8FD80034" box="[390,445,1632,1653]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="23" pageNumber="24" unit="mm" value="7.0">7 mm</quantity>
............................................................
<emphasis id="DBF6EAF8AC4E436C0F8DF9948B240034" bold="true" box="[1177,1345,1631,1653]" pageId="23" pageNumber="24">
<taxonomicName id="2E824D69AC4E436C0F8DF9948ABC0034" authorityName="WOOD" authorityYear="2024" box="[1177,1241,1631,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="collisa" status="sp. nov.">
<emphasis id="DBF6EAF8AC4E436C0F8DF9948ABC0034" bold="true" box="[1177,1241,1631,1653]" italics="true" pageId="23" pageNumber="24">collisa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC4E436C0FCBF9AB8B240034" box="[1247,1345,1632,1653]" pageId="23" pageNumber="24" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<collectingCountry id="9195767AAC4E436C0E5AF9AB8BF30034" box="[1358,1430,1632,1653]" name="Cyprus" pageId="23" pageNumber="24">Cyprus</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F9B68BF800F1" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
10. Small, 77.5 mm. Process of the labrum narrowly triangular. Associated with
<taxonomicName id="2E824D69AC4E436C08D9F9B68A5100D3" ID-CoL="8VRY5" box="[973,1076,1661,1682]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8AC4E436C08D9F9B68A5100D3" box="[973,1076,1661,1682]" italics="true" pageId="23" pageNumber="24">Euphorbia</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69AC4E436C0F54F9B68AB700D2" ID-CoL="9Y7" box="[1088,1234,1661,1683]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
)..................................................................................
<taxonomicName id="2E824D69AC4E436C0F04F9518AB700F1" authority="Pisanty" authorityName="Pisanty" authorityYear="2022" box="[1040,1234,1690,1712]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="euphorbiae">
<emphasis id="DBF6EAF8AC4E436C0F04F9518AB700F1" bold="true" box="[1040,1234,1690,1712]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0F04F9518AE500F1" bold="true" box="[1040,1152,1690,1712]" italics="true" pageId="23" pageNumber="24">euphorbiae</emphasis>
Pisanty
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0FCBF9518B7300F1" box="[1247,1302,1690,1712]" name="Israel" pageId="23" pageNumber="24">Israel</collectingCountry>
&amp;
<collectingCountry id="9195767AAC4E436C0E20F9518BE900F1" box="[1332,1420,1690,1712]" name="Lebanon" pageId="23" pageNumber="24">Lebanon</collectingCountry>
*)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F9738BF900AB" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Larger, usually clearly in excess of
<quantity id="2E7A9B0FAC4E436C0955F9738C18008C" box="[577,637,1720,1741]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="24" unit="mm" value="8.0">8 mm</quantity>
. Process of the labrum more or less rounded rectangular, with or without median indentation, never narrowly triangular. Not associated with
<taxonomicName id="2E824D69AC4E436C0809F91E8DE100AB" ID-CoL="8VRY5" box="[797,900,1749,1770]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8AC4E436C0809F91E8DE100AB" box="[797,900,1749,1770]" italics="true" pageId="23" pageNumber="24">Euphorbia</emphasis>
</taxonomicName>
.......................................... 11
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F9398BF90102" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
11. Mesepisternum predominantly dark-haired, at least 50% of hairs black. Pronotum laterally with deep longitudinal shining impression in ventral half (
<figureCitation id="71B92A6FAC4E436C0AF0F8C48C330164" box="[484,598,1807,1829]" captionStart="FIGURE 11" captionStartId="19.[151,250,1880,1904]" captionTargetBox="[151,1436,417,1856]" captionTargetId="figure-125@19.[151,1436,417,1856]" captionTargetPageId="19" captionText="FIGURE 11. Andrena (Avandrena) melaleuca Friese, 1922 nec. Pérez = A. melacana Warncke, 1967 lectotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Pronotum, lateral view; E. Scutum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209329" httpUri="https://zenodo.org/record/13209329/files/figure.png" pageId="23" pageNumber="24">Figure 11D</figureCitation>
). Fore margin of the clypeus upturned.............................................................................................
<taxonomicName id="2E824D69AC4E436C0897F8E78A230103" ID-CoL="66N3R" authority="Warncke" authorityName="Warncke" authorityYear="1967" box="[899,1094,1836,1858]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="melacana">
<emphasis id="DBF6EAF8AC4E436C0897F8E78A230103" bold="true" box="[899,1094,1836,1858]" pageId="23" pageNumber="24">
<emphasis id="DBF6EAF8AC4E436C0897F8E78D860103" bold="true" box="[899,995,1836,1858]" italics="true" pageId="23" pageNumber="24">melacana</emphasis>
Warncke
</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC4E436C0F47F8E68ACB0103" box="[1107,1198,1837,1858]" name="Morocco" pageId="23" pageNumber="24">Morocco</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0FA2F8E68B650102" box="[1206,1280,1837,1859]" name="Algeria" pageId="23" pageNumber="24">Algeria</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0E1DF8E68B360102" box="[1289,1363,1837,1859]" name="Tunisia" pageId="23" pageNumber="24">Tunisia</collectingCountry>
,
<collectingCountry id="9195767AAC4E436C0E48F8E68BF30102" box="[1372,1430,1837,1859]" name="Libya" pageId="23" pageNumber="24">Libya</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F8808BF901DB" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
- Mesepisternum predominantly pale-haired, usually with no more than 30% of hairs black. Pronotum with or without (
<figureCitation id="71B92A6FAC4E436C0E48F8818E9A013C" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="23" pageNumber="24">Figure 9C</figureCitation>
) a longitudinal shining impression; if mesepisternum with abundant black hairs, then never with a longitudinal impression on the pronotum. Fore margin of the clypeus not upturned................................................... 12
</paragraph>
<paragraph id="E93D36EAAC4E436C0B83F8698BF901B3" blockId="23.[151,1437,575,2035]" pageId="23" pageNumber="24">
12. Western species, present in
<collectingCountry id="9195767AAC4E436C0AF1F8698C7A01F9" box="[485,543,1954,1976]" name="Spain" pageId="23" pageNumber="24">Spain</collectingCountry>
. Pronotum rounded, without a humeral angle, at most with hints of an angulation dorsolaterally but this does not descend and form a longitudinal angle (
<figureCitation id="71B92A6FAC4E436C09E7F8748D320194" box="[755,855,1983,2005]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="23" pageNumber="24">Figure 9C</figureCitation>
). Dorsal surface of propodeum with weak to strong sculpture, in some species with distinctly raised network of raised rugosity.............................................. 13
</paragraph>
<paragraph id="E93D36EAAC4143630B83FF538BF9068A" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
- Eastern species, present in
<collectingCountry id="9195767AAC4143630AE2FF5C8C5506EC" box="[502,560,151,173]" name="Libya" pageId="24" pageNumber="25">Libya</collectingCountry>
,
<collectingCountry id="9195767AAC414363092AFF5C8CE106EC" box="[574,644,151,173]" name="Turkey" pageId="24" pageNumber="25">Turkey</collectingCountry>
, or
<collectingCountry id="9195767AAC41436309BBFF5C8C8206EC" box="[687,743,151,173]" name="Israel" pageId="24" pageNumber="25">Israel</collectingCountry>
. Pronotum laterally with a distinct humeral angle. Dorsal surface of propodeum entirely microreticulate, without strong or weak raised network of reticulation.......................... 15
</paragraph>
<paragraph id="E93D36EAAC4143630B83FF198BF80744" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
13. Dorsolateral parts of propodeum and propodeal triangle with fine granular reticulation, without any rugosity (
<figureCitation id="71B92A6FAC4143630FE9FF198B3A06A6" box="[1277,1375,210,232]" captionStart="FIGURE 9" captionStartId="17.[151,250,1882,1906]" captionTargetBox="[151,1436,417,1859]" captionTargetId="figure-117@17.[151,1436,417,1859]" captionTargetPageId="17" captionText="FIGURE 9. Andrena (Avandrena) melacanoides spec. nov. female.A. Habitus, lateral view; B. Face, frontal view; C. Pronotum, lateral view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209325" httpUri="https://zenodo.org/record/13209325/files/figure.png" pageId="24" pageNumber="25">Figure 9E</figureCitation>
)........................
<emphasis id="DBF6EAF8AC4143630AFBFF248CBB0744" bold="true" box="[495,734,239,261]" pageId="24" pageNumber="25">
<taxonomicName id="2E824D69AC4143630AFBFF248C130744" authorityName="WOOD" authorityYear="2024" box="[495,630,239,261]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="melacanoides" status="sp. nov.">
<emphasis id="DBF6EAF8AC4143630AFBFF248C130744" bold="true" box="[495,630,239,261]" italics="true" pageId="24" pageNumber="25">melacanoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC4143630968FF3B8CBB0744" box="[636,734,240,261]" pageId="24" pageNumber="25" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(southern
<collectingCountry id="9195767AAC4143630850FF248D1B0744" box="[836,894,239,261]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
: Cádiz and Granada, probably also Albacete &amp; Málaga).
</paragraph>
<paragraph id="E93D36EAAC4143630B83FEC68BF9077E" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">- Dorsolateral parts of propodeum and propodeal triangle with raised sculpture, either forming an interlinked network of raised rugae or more irregular pattern of raised rugosity........................................................... 14</paragraph>
<paragraph id="E93D36EAAC4143630B83FE838BF907D9" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
14. Propodeum (including propodeal triangle) and mesepisternum with strongly produced but fine interlinked network of raised rugae. Facial foveae occupying ¾ of space between the compound eye and lateral ocellus.Apical fringe of T5 and hairs flanking pygidial plate golden-brown......................
<taxonomicName id="2E824D69AC4143630819FE498A0C07D6" authority="Wood &amp; Ortiz-Sanchez" authorityName="Wood &amp; Ortiz-Sanchez" authorityYear="2022" box="[781,1129,385,408]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="erodiorum">
<emphasis id="DBF6EAF8AC4143630819FE498A0C07D6" bold="true" box="[781,1129,385,408]" pageId="24" pageNumber="25">
<emphasis id="DBF6EAF8AC4143630819FE498D1007D9" bold="true" box="[781,885,386,408]" italics="true" pageId="24" pageNumber="25">erodiorum</emphasis>
Wood &amp; Ortiz-Sánchez
</emphasis>
</taxonomicName>
(south-eastern
<collectingCountry id="9195767AAC4143630FE9FE498B5207D9" box="[1277,1335,386,408]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
: Albacete)
</paragraph>
<paragraph id="E93D36EAAC4143630B83FE6B8BF807B1" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
- Propodeum with fine granular reticulation, with weak network of raised rugosity, propodeal triangle slightly depressed, basal ⅔ with raised longitudinal rugae. Facial foveae occupying ½ space between the compound eye and a lateral ocellus. Apical fringe of T5 and hairs flanking pygidial plate dark brown.........................
<taxonomicName id="2E824D69AC4143630F11FE118A1B07AE" authority="Wood" authorityName="Wood" authorityYear="2023" box="[1029,1150,474,496]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="juliae">
<emphasis id="DBF6EAF8AC4143630F11FE118A1B07AE" bold="true" box="[1029,1150,474,496]" pageId="24" pageNumber="25">
<emphasis id="DBF6EAF8AC4143630F11FE118A5807B1" bold="true" box="[1029,1085,474,496]" italics="true" pageId="24" pageNumber="25">juliae</emphasis>
Wood
</emphasis>
</taxonomicName>
(south-western
<collectingCountry id="9195767AAC4143630E0DFE118B3607B1" box="[1305,1363,474,496]" name="Spain" pageId="24" pageNumber="25">Spain</collectingCountry>
: Cádiz)
</paragraph>
<paragraph id="E93D36EAAC4143630B83FE3C8BF80409" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
15. Scutum and scutellum (particularly scutellum) with weak shagreen, more or less smooth and shining between punctures. Tibial scopae composed of uniformly golden weakly plumose hairs. Clypeus with an impunctate longitudinal mid-line.......................................................................
<taxonomicName id="2E824D69AC41436308D0FDF98A130406" ID-CoL="DR9V" authority="Warncke" authorityName="Warncke" authorityYear="1965" box="[964,1142,562,584]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="caudata">
<emphasis id="DBF6EAF8AC41436308D0FDF98A130406" bold="true" box="[964,1142,562,584]" pageId="24" pageNumber="25">
<emphasis id="DBF6EAF8AC41436308D0FDF98A760409" bold="true" box="[964,1043,562,584]" italics="true" pageId="24" pageNumber="25">caudata</emphasis>
Warncke
</emphasis>
</taxonomicName>
(south-western
<collectingCountry id="9195767AAC4143630E04FDF98B330409" box="[1296,1366,562,584]" name="Turkey" pageId="24" pageNumber="25">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC4143630E4BFDF98BF30409" box="[1375,1430,562,584]" name="Israel" pageId="24" pageNumber="25">Israel</collectingCountry>
)
</paragraph>
<paragraph id="E93D36EAAC4143630B83FD9B8BF904C2" blockId="24.[151,1437,151,643]" pageId="24" pageNumber="25">
- Scutum and scutellum shagreened, at most weakly shining between punctures. Tibial scopae composed of simple hairs, dorsally dark brown, ventrally white. Clypeus uniformly punctate............
<taxonomicName id="2E824D69AC41436308B3FDA78A3E04C3" ID-CoL="DSG7" authority="Warncke" authorityName="Warncke" authorityYear="1974" box="[935,1115,620,642]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ochropa">
<emphasis id="DBF6EAF8AC41436308B3FDA78A3E04C3" bold="true" box="[935,1115,620,642]" pageId="24" pageNumber="25">
<emphasis id="DBF6EAF8AC41436308B3FDA78D9D04C3" bold="true" box="[935,1016,620,642]" italics="true" pageId="24" pageNumber="25">ochropa</emphasis>
Warncke
</emphasis>
</taxonomicName>
(north-eastern
<collectingCountry id="9195767AAC4143630FFAFDA68B4F04C2" box="[1262,1322,621,643]" name="Libya" pageId="24" pageNumber="25">Libya</collectingCountry>
: Cyrenaica)
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,375 @@
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<subSubSection id="A1986561AC6243400B83FAB38C9F03D3" box="[151,762,1400,1427]" pageId="59" pageNumber="60" type="nomenclature">
<paragraph id="E93D36EAAC6243400B83FAB38C9F03D3" blockId="59.[151,762,1400,1427]" box="[151,762,1400,1427]" pageId="59" pageNumber="60">
<heading id="B2758186AC6243400B83FAB38C9F03D3" bold="true" box="[151,762,1400,1427]" fontSize="11" level="2" pageId="59" pageNumber="60" reason="4">
<taxonomicName id="2E824D69AC6243400B83FAB38C9F03D3" ID-CoL="DSVB" authority="Stockhert, 1935" authorityName="Stockhert" authorityYear="1935" box="[151,762,1400,1427]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6243400B83FAB38C9F03D3" bold="true" box="[151,762,1400,1427]" pageId="59" pageNumber="60">
<emphasis id="DBF6EAF8AC6243400B83FAB38F6403D2" bold="true" box="[151,257,1400,1427]" italics="true" pageId="59" pageNumber="60">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6243400A06FAB38FD703D2" bold="true" box="[274,434,1400,1427]" italics="true" pageId="59" pageNumber="60">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6243400AD7FAB38C4A03D3" bold="true" box="[451,559,1400,1426]" italics="true" pageId="59" pageNumber="60">saxonica</emphasis>
<bibRefCitation id="8D134B1BAC6243400922FAB38C9F03D3" author="Stockhert, E." box="[566,762,1400,1426]" pageId="59" pageNumber="60" pagination="65 - 85" refId="ref91208" refString="Stockhert, E. (1935) Uber einige neue deutsche Arten der Andrena minutula - Gruppe (Hym. Apid.). Deutsche Entomologische Zeitschrift, 1935, 65 - 85. https: // doi. org / 10.1002 / mmnd. 48019350102" type="journal article" year="1935">Stöckhert, 1935</bibRefCitation>
</emphasis>
</taxonomicName>
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<subSubSection id="A1986561AC6243400B83FA768F7F0070" pageId="59" pageNumber="60" type="reference_group">
<paragraph id="E93D36EAAC6243400B83FA768DF60394" blockId="59.[151,1437,1469,1585]" box="[151,915,1469,1493]" pageId="59" pageNumber="60">
<treatmentCitationGroup id="C99211C4AC6243400B83FA768DF60394" box="[151,915,1469,1493]" pageId="59" pageNumber="60">
<treatmentCitation id="682310FBAC6243400B83FA768C7E0394" author="Stockhert, E." box="[151,539,1469,1493]" page="82" pageId="59" pageNumber="60" year="1935">
<taxonomicName id="2E824D69AC6243400B83FA768C7E0394" ID-CoL="DSVB" authority="Stockhert, 1935: 82" authorityName="Stockhert" authorityPageNumber="82" authorityYear="1935" box="[151,539,1469,1493]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6243400B83FA768F2B0394" box="[151,334,1469,1493]" italics="true" pageId="59" pageNumber="60">Andrena saxonica</emphasis>
<materialsCitation id="59EA3CB7AC6243400A40FA768C7E0394" box="[340,539,1469,1493]" collectionCode="ZSM" country="Germany" location="Germany" pageId="59" pageNumber="60" specimenCount="83" specimenCount-female="82" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC6243400A40FA768C7E0394" author="Stockhert, E." box="[340,539,1469,1493]" pageId="59" pageNumber="60" pagination="65 - 85" refId="ref91208" refString="Stockhert, E. (1935) Uber einige neue deutsche Arten der Andrena minutula - Gruppe (Hym. Apid.). Deutsche Entomologische Zeitschrift, 1935, 65 - 85. https: // doi. org / 10.1002 / mmnd. 48019350102" type="journal article" year="1935">
Stöckhert, 1935:
<specimenCount id="FF84FD63AC6243400916FA768C7E0394" box="[514,539,1469,1493]" count="82" pageId="59" pageNumber="60" type="female">82</specimenCount>
</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
<specimenCount id="FF84FD63AC6243400925FA768C220395" box="[561,583,1469,1492]" count="1" pageId="59" pageNumber="60" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6243400940FA768CD30394" box="[596,694,1469,1493]" name="Germany" pageId="59" pageNumber="60">Germany</collectingCountry>
, ZSM, not examined]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC6243400B83FA178BFE03B5" blockId="59.[151,1437,1469,1585]" box="[151,1435,1499,1526]" pageId="59" pageNumber="60">
<taxonomicName id="2E824D69AC6243400B83FA178C1903B2" ID-CoL="66N38" authority="Warncke, 1975 b" authorityName="Warncke" authorityYear="1975" box="[151,636,1499,1524]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6243400B83FA178E8803B5" box="[151,237,1500,1524]" italics="true" pageId="59" pageNumber="60">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6243400BE8FA178FE403B5" box="[252,385,1500,1524]" italics="true" pageId="59" pageNumber="60">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6243400A84FA178FB403B2" box="[400,465,1500,1523]" italics="true" pageId="59" pageNumber="60">crypta</emphasis>
<materialsCitation id="59EA3CB7AC6243400AC2FA178C1903B2" box="[470,636,1499,1524]" collectionCode="OOLM" country="Turkey" location="Turkey" pageId="59" pageNumber="60" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC6243400AC2FA178C1903B2" author="Warncke, K." box="[470,636,1499,1524]" pageId="59" pageNumber="60" pagination="29 - 102" refId="ref91833" refString="Warncke, K. (1975 b) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke, 1975b</bibRefCitation>
</materialsCitation>
</taxonomicName>
(nec.
<taxonomicName id="2E824D69AC62434009AEFA178A7C03B5" ID-CoL="4PTFS" authority="Viereck, 1904" authorityName="Viereck" authorityYear="1904" box="[698,1049,1500,1524]" class="Insecta" family="Andrenidae" genus="Pterandrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="DBF6EAF8AC62434009AEFA178DE403B2" box="[698,897,1500,1524]" italics="true" pageId="59" pageNumber="60">Pterandrena crypta</emphasis>
Viereck, 1904
</taxonomicName>
): 96,
<specimenCount id="FF84FD63AC6243400F44FA178A0503B2" box="[1104,1120,1500,1523]" count="1" pageId="59" pageNumber="60" type="female"></specimenCount>
<specimenCount id="FF84FD63AC6243400F74FA178A1303B2" box="[1120,1142,1500,1523]" count="1" pageId="59" pageNumber="60" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6243400F96FA178AAB03B5" box="[1154,1230,1500,1524]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
, OÖLM, examined]
</paragraph>
<paragraph id="E93D36EAAC6243400B83FA318F7F0070" blockId="59.[151,1437,1469,1585]" pageId="59" pageNumber="60">
<treatmentCitation id="682310FBAC6243400B83FA318D320053" author="Gusenleitner, F. &amp; Schwarz, M." box="[151,855,1530,1554]" page="112" pageId="59" pageNumber="60" year="2000">
<taxonomicName id="2E824D69AC6243400B83FA318D320053" ID-CoL="DTBB" authority="Gusenleitner &amp; Schwarz, 2000: 112" authorityName="Gusenleitner &amp; Schwarz" authorityPageNumber="112" authorityYear="2000" box="[151,855,1530,1554]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="warnckei" status="nom. nov." subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6243400B83FA318E880053" box="[151,237,1530,1554]" italics="true" pageId="59" pageNumber="60">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6243400BEFFA318FE50053" box="[251,384,1530,1554]" italics="true" pageId="59" pageNumber="60">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6243400A99FA318F8F0053" box="[397,490,1530,1554]" italics="true" pageId="59" pageNumber="60">warnckei</emphasis>
<bibRefCitation id="8D134B1BAC6243400AFBFA318D320053" author="Gusenleitner, F. &amp; Schwarz, M." box="[495,855,1530,1554]" pageId="59" pageNumber="60" pagination="105 - 116" refId="ref88896" refString="Gusenleitner, F. &amp; Schwarz, M. (2000) Nomenklatorische Aktualisierungen in der Bienengattung Andrena sowie Beschreibung einer neuen Art (Hymenoptera: Apidae: Andreninae). Entomofauna, 10, 105 - 116." type="journal article" year="2000">Gusenleitner &amp; Schwarz, 2000: 112</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="DBF6EAF8AC6243400875FA308DA70053" box="[865,962,1531,1554]" italics="true" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C0C55783AC6243400875FA308DA70053" box="[865,962,1531,1554]" pageId="59" pageNumber="60">nom. nov.</taxonomicNameLabel>
</emphasis>
for
<taxonomicName id="2E824D69AC62434008FDFA308A840053" ID-CoL="4PTFS" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[1001,1249,1530,1554]" class="Insecta" family="Andrenidae" genus="Pterandrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta" status="syn. nov.">
<emphasis id="DBF6EAF8AC62434008FDFA308A200053" box="[1001,1093,1531,1554]" italics="true" pageId="59" pageNumber="60">A. crypta</emphasis>
Warncke, 1975
</taxonomicName>
<emphasis id="DBF6EAF8AC6243400FF2FA318B270053" bold="true" box="[1254,1346,1530,1554]" pageId="59" pageNumber="60">
<taxonomicNameLabel id="C0C55783AC6243400FF2FA318B270053" box="[1254,1346,1530,1554]" pageId="59" pageNumber="60">syn. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="71B92A6FAC6243400E5BFA318F760070" captionStart="FIGURE 39" captionStartId="60.[151,250,1159,1183]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-28@60.[151,1436,181,1135]" captionTargetPageId="60" captionText="FIGURE 39. Andrena (Micrandrena) cryptaWarncke, 1975 nec. Viereck = A. warnckei Gusenleitner &amp; Schwarz, 2000 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209391" httpUri="https://zenodo.org/record/13209391/files/figure.png" pageId="59" pageNumber="60">Figures 39AD</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6243460B83F9948A760694" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" type="discussion">
<paragraph id="E93D36EAAC6243400B83F9948C4C0110" blockId="59.[151,1437,1631,1873]" pageId="59" pageNumber="60">
<emphasis id="DBF6EAF8AC6243400B83F9948F6E0038" bold="true" box="[151,267,1631,1657]" pageId="59" pageNumber="60">Remarks.</emphasis>
<bibRefCitation id="8D134B1BAC6243400A00F9948FBD003B" author="Warncke, K." box="[276,472,1631,1658]" pageId="59" pageNumber="60" pagination="29 - 102" refId="ref91833" refString="Warncke, K. (1975 b) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke (1975b)</bibRefCitation>
described
<taxonomicName id="2E824D69AC6243400941F9948CDB0038" authorityName="Warncke" authorityYear="1975" box="[597,702,1631,1657]" class="Insecta" family="Andrenidae" genus="Pterandrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="DBF6EAF8AC6243400941F9948CDB0038" box="[597,702,1631,1657]" italics="true" pageId="59" pageNumber="60">A. crypta</emphasis>
</taxonomicName>
from eastern
<collectingCountry id="9195767AAC6243400849F9948DCB0038" box="[861,942,1631,1657]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
, though the name was unavailable and was replaced by
<bibRefCitation id="8D134B1BAC6243400A0BF9488CE300DC" author="Gusenleitner, F. &amp; Schwarz, M." box="[287,646,1667,1693]" pageId="59" pageNumber="60" pagination="105 - 116" refId="ref88896" refString="Gusenleitner, F. &amp; Schwarz, M. (2000) Nomenklatorische Aktualisierungen in der Bienengattung Andrena sowie Beschreibung einer neuen Art (Hymenoptera: Apidae: Andreninae). Entomofauna, 10, 105 - 116." type="journal article" year="2000">Gusenleitner &amp; Schwarz (2000)</bibRefCitation>
. Warncke diagnosed
<taxonomicName id="2E824D69AC6243400865F9488DBD00DC" authorityName="Warncke" authorityYear="1975" box="[881,984,1667,1693]" class="Insecta" family="Andrenidae" genus="Pterandrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="DBF6EAF8AC6243400865F9488DBD00DC" box="[881,984,1667,1693]" italics="true" pageId="59" pageNumber="60">A. crypta</emphasis>
</taxonomicName>
against
<taxonomicName id="2E824D69AC6243400F20F9488AD200DC" authorityName="" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[1076,1207,1667,1693]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="DBF6EAF8AC6243400F20F9488AD200DC" box="[1076,1207,1667,1693]" italics="true" pageId="59" pageNumber="60">A. minutula</emphasis>
</taxonomicName>
and surprisingly did not mention
<taxonomicName id="2E824D69AC6243400A36F96C8FC10080" authorityName="Stockhert" authorityYear="1935" box="[290,420,1703,1729]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6243400A36F96C8FC10080" box="[290,420,1703,1729]" italics="true" pageId="59" pageNumber="60">A. saxonica</emphasis>
</taxonomicName>
; the two taxa are extremely similar morphologically, and the genital capsule of the
<typeStatus id="36398848AC6243400E2EF96C8BF90080" box="[1338,1436,1703,1729]" pageId="59" pageNumber="60" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC6243400BA0F9008F7F00A4" authorityName="Warncke" authorityYear="1975" box="[180,282,1739,1765]" class="Insecta" family="Andrenidae" genus="Pterandrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="crypta">
<emphasis id="DBF6EAF8AC6243400BA0F9008F7F00A4" box="[180,282,1739,1765]" italics="true" pageId="59" pageNumber="60">A. crypta</emphasis>
</taxonomicName>
is identical to that of
<taxonomicName id="2E824D69AC6243400913F9008CEC00A4" authorityName="Stockhert" authorityYear="1935" box="[519,649,1739,1765]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6243400913F9008CEC00A4" box="[519,649,1739,1765]" italics="true" pageId="59" pageNumber="60">A. saxonica</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC6243400982F9008D7E00A4" box="[662,795,1739,1765]" captionStart="FIGURE 39" captionStartId="60.[151,250,1159,1183]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-28@60.[151,1436,181,1135]" captionTargetPageId="60" captionText="FIGURE 39. Andrena (Micrandrena) cryptaWarncke, 1975 nec. Viereck = A. warnckei Gusenleitner &amp; Schwarz, 2000 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209391" httpUri="https://zenodo.org/record/13209391/files/figure.png" pageId="59" pageNumber="60">Figure 39D</figureCitation>
). The two taxa are therefore synonymised
<emphasis id="DBF6EAF8AC6243400FFAF9008B3000A4" bold="true" box="[1262,1365,1739,1765]" pageId="59" pageNumber="60">syn. nov.</emphasis>
<bibRefCitation id="8D134B1BAC6243400E4EF9008F110148" author="Wood, T. J. &amp; Monfared, A." pageId="59" pageNumber="60" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared (2022)</bibRefCitation>
reported
<taxonomicName id="2E824D69AC6243400ACAF9248C040148" authorityName="Stockhert" authorityYear="1935" box="[478,609,1775,1801]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6243400ACAF9248C040148" box="[478,609,1775,1801]" italics="true" pageId="59" pageNumber="60">A. saxonica</emphasis>
</taxonomicName>
from eastern
<collectingCountry id="9195767AAC62434009EDF9248D2F0148" box="[761,842,1775,1801]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
, this species thus occurring within the reported range of
<taxonomicName id="2E824D69AC6243400BA3F8D88F5B016C" authorityName="Gusenleitner &amp; Schwarz" authorityYear="2000" box="[183,318,1811,1837]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="warnckei">
<emphasis id="DBF6EAF8AC6243400BA3F8D88F5B016C" box="[183,318,1811,1837]" italics="true" pageId="59" pageNumber="60">A. warnckei</emphasis>
</taxonomicName>
, but
<taxonomicName id="2E824D69AC6243400A6DF8D88F98016C" authorityName="Stockhert" authorityYear="1935" box="[377,509,1811,1837]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6243400A6DF8D88F98016C" box="[377,509,1811,1837]" italics="true" pageId="59" pageNumber="60">A. saxonica</emphasis>
</taxonomicName>
is also widespread in mountainous areas throughout
<collectingCountry id="9195767AAC6243400F43F8D88ACD016C" box="[1111,1192,1811,1837]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
, including the Taurus mountains in south-western
<collectingCountry id="9195767AAC6243400AC7F8FC8C400110" box="[467,549,1847,1873]" name="Turkey" pageId="59" pageNumber="60">Turkey</collectingCountry>
.
</paragraph>
<caption id="BDFD6662AC6543470B83FB4C8AC30282" ID-DOI="http://doi.org/10.5281/zenodo.13209391" ID-Zenodo-Dep="13209391" httpUri="https://zenodo.org/record/13209391/files/figure.png" pageId="60" pageNumber="61" startId="60.[151,250,1159,1183]" targetBox="[151,1436,181,1135]" targetPageId="60" targetType="figure">
<paragraph id="E93D36EAAC6543470B83FB4C8AC30282" blockId="60.[151,1436,1159,1220]" pageId="60" pageNumber="61">
<emphasis id="DBF6EAF8AC6543470B83FB4C8F7B02DE" bold="true" box="[151,286,1159,1183]" pageId="60" pageNumber="61">FIGURE 39.</emphasis>
<taxonomicName id="2E824D69AC6543470A37FB4C8C9102DE" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[291,756,1159,1184]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="crypta" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6543470A37FB4C8F1C02DE" box="[291,377,1159,1183]" italics="true" pageId="60" pageNumber="61">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6543470A92FB4C8C6E02DE" box="[390,523,1159,1183]" italics="true" pageId="60" pageNumber="61">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC654347090CFB438C3C02DE" box="[536,601,1160,1183]" italics="true" pageId="60" pageNumber="61">crypta</emphasis>
Warncke, 1975
</taxonomicName>
nec. Viereck =
<taxonomicName id="2E824D69AC6543470898FB438B5E02E1" authority="Gusenleitner &amp; Schwarz, 2000" authorityName="Gusenleitner &amp; Schwarz" authorityYear="2000" box="[908,1339,1159,1184]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="warnckei">
<emphasis id="DBF6EAF8AC6543470898FB438A6602DE" box="[908,1027,1159,1183]" italics="true" pageId="60" pageNumber="61">A. warnckei</emphasis>
<bibRefCitation id="8D134B1BAC6543470F1CFB4C8B5E02E1" author="Gusenleitner, F. &amp; Schwarz, M." box="[1032,1339,1159,1184]" pageId="60" pageNumber="61" pagination="105 - 116" refId="ref88896" refString="Gusenleitner, F. &amp; Schwarz, M. (2000) Nomenklatorische Aktualisierungen in der Bienengattung Andrena sowie Beschreibung einer neuen Art (Hymenoptera: Apidae: Andreninae). Entomofauna, 10, 105 - 116." type="journal article" year="2000">Gusenleitner &amp; Schwarz, 2000</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC6543470E2BFB438BF902E1" bold="true" box="[1343,1436,1160,1184]" pageId="60" pageNumber="61">holotype</emphasis>
male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Genital capsule, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC6543470BD3FB398A480011" blockId="60.[151,1437,1265,2012]" pageId="60" pageNumber="61">
Furthermore,
<bibRefCitation id="8D134B1BAC6543470A4BFB398C2C034D" author="Warncke, K." box="[351,585,1265,1292]" pageId="60" pageNumber="61" pagination="29 - 102" refId="ref91833" refString="Warncke, K. (1975 b) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke (1975b: 53)</bibRefCitation>
described
<taxonomicName id="2E824D69AC65434709ABFB398DAA034D" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" box="[703,975,1265,1292]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC65434709ABFB398D46034D" box="[703,803,1266,1292]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
Warncke, 1975
</taxonomicName>
from
<collectingCountry id="9195767AAC6543470F04FB398A04034D" box="[1040,1121,1266,1292]" name="Turkey" pageId="60" pageNumber="61">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC6543470F7EFB398AD9034D" box="[1130,1212,1266,1292]" name="Greece" pageId="60" pageNumber="61">Greece</collectingCountry>
,
<collectingCountry id="9195767AAC6543470FD2FB398B45034D" box="[1222,1312,1266,1292]" name="Ukraine" pageId="60" pageNumber="61">Ukraine</collectingCountry>
(
<collectingRegion id="2B46F808AC6543470E39FB398BF5034D" box="[1325,1424,1266,1292]" country="Ukraine" name="Donets'k" pageId="60" pageNumber="61">Donetsk</collectingRegion>
), and southern European
<collectingCountry id="9195767AAC6543470AB2FADD8F940371" box="[422,497,1302,1328]" name="Russia" pageId="60" pageNumber="61">Russia</collectingCountry>
(
<collectingRegion id="2B46F808AC6543470915FADD8C100371" box="[513,629,1302,1328]" country="Russia" name="Smolensk" pageId="60" pageNumber="61">Smolensk</collectingRegion>
), and diagnosed it against
<taxonomicName id="2E824D69AC65434708B2FADD8A4E036E" authorityName="Stockhert" authorityYear="1935" box="[934,1067,1302,1328]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC65434708B2FADD8A4E036E" box="[934,1067,1302,1328]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
. The
<emphasis id="DBF6EAF8AC6543470F7AFADD8B64036E" box="[1134,1281,1302,1328]" italics="true" pageId="60" pageNumber="61">locus typicus</emphasis>
is the Taurus mountains in south-western
<collectingCountry id="9195767AAC6543470AC5FAF18C440315" box="[465,545,1338,1364]" name="Turkey" pageId="60" pageNumber="61">Turkey</collectingCountry>
(
<figureCitation id="71B92A6FAC654347093AFAF18CDB0315" box="[558,702,1338,1364]" captionStart="FIGURE 40" captionStartId="61.[151,250,1217,1241]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@61.[151,1436,237,1194]" captionTargetPageId="61" captionText="FIGURE 40. Andrena (Micrandrena) sillata Warncke, 1975 paratype male.A. Label details; B. Habitus, lateral view; C. Terga, dorsal view; D. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209393" httpUri="https://zenodo.org/record/13209393/files/figure.png" pageId="60" pageNumber="61">Figures 40A</figureCitation>
;
<figureCitation id="71B92A6FAC65434709DCFAF18C990315" box="[712,764,1338,1364]" captionStart="FIGURE 41" captionStartId="62.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@62.[214,1373,181,1919]" captionTargetPageId="62" captionText="FIGURE 41. Andrena (Micrandrena) sillata Warncke, 1975 holotype female. A. Label details; C. Habitus, lateral view; E. Face, frontal view; G. Terga, dorsal view; Andrena (Micrandrena) sillata Warncke, 1975 paratype from Ukraine. B. Label details; D. Habitus, lateral view; F. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209395" httpUri="https://zenodo.org/record/13209395/files/figure.png" pageId="60" pageNumber="61">41A</figureCitation>
), and hence the inclusion of material from Eastern Europe in the type series is perplexing. A single female
<typeStatus id="36398848AC654347098CFA958C9D0339" box="[664,760,1374,1400]" pageId="60" pageNumber="61" type="paratype">paratype</typeStatus>
from
<collectingCountry id="9195767AAC654347082AFA958DFD0339" box="[830,920,1374,1400]" name="Ukraine" pageId="60" pageNumber="61">Ukraine</collectingCountry>
is available in the OÖLM collection (
<figureCitation id="71B92A6FAC6543470E52FA958F4B03DD" captionStart="FIGURE 41" captionStartId="62.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@62.[214,1373,181,1919]" captionTargetPageId="62" captionText="FIGURE 41. Andrena (Micrandrena) sillata Warncke, 1975 holotype female. A. Label details; C. Habitus, lateral view; E. Face, frontal view; G. Terga, dorsal view; Andrena (Micrandrena) sillata Warncke, 1975 paratype from Ukraine. B. Label details; D. Habitus, lateral view; F. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209395" httpUri="https://zenodo.org/record/13209395/files/figure.png" pageId="60" pageNumber="61">Figures 41B; D; F; H</figureCitation>
), and in direct comparison is larger and presents terga that are more strongly shagreened than typical
<taxonomicName id="2E824D69AC6543470B83FA6D8E980381" authorityName="Warncke" authorityYear="1975" box="[151,253,1446,1472]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC6543470B83FA6D8E980381" box="[151,253,1446,1472]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
(compare
<figureCitation id="71B92A6FAC6543470A61FA6D8C4D0381" box="[373,552,1446,1472]" captionStart="FIGURE 41" captionStartId="62.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-28@62.[214,1373,181,1919]" captionTargetPageId="62" captionText="FIGURE 41. Andrena (Micrandrena) sillata Warncke, 1975 holotype female. A. Label details; C. Habitus, lateral view; E. Face, frontal view; G. Terga, dorsal view; Andrena (Micrandrena) sillata Warncke, 1975 paratype from Ukraine. B. Label details; D. Habitus, lateral view; F. Face, frontal view; H. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209395" httpUri="https://zenodo.org/record/13209395/files/figure.png" pageId="60" pageNumber="61">Figures 41GH</figureCitation>
). No males were reported by Warncke from the Eastern European populations. Comparison of the genital capsule of typical
<taxonomicName id="2E824D69AC65434709B7FA018D6E03A5" authorityName="Warncke" authorityYear="1975" box="[675,779,1482,1508]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC65434709B7FA018D6E03A5" box="[675,779,1482,1508]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC6543470808FA018DC703A5" box="[796,930,1482,1508]" captionStart="FIGURE 40" captionStartId="61.[151,250,1217,1241]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@61.[151,1436,237,1194]" captionTargetPageId="61" captionText="FIGURE 40. Andrena (Micrandrena) sillata Warncke, 1975 paratype male.A. Label details; B. Habitus, lateral view; C. Terga, dorsal view; D. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209393" httpUri="https://zenodo.org/record/13209393/files/figure.png" pageId="60" pageNumber="61">Figure 40D</figureCitation>
) and
<taxonomicName id="2E824D69AC65434708F3FA018A0803A2" authorityName="Stockhert" authorityYear="1935" box="[999,1133,1482,1508]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC65434708F3FA018A0803A2" box="[999,1133,1482,1508]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC6543470F6AFA018B6103A5" box="[1150,1284,1482,1508]" captionStart="FIGURE 39" captionStartId="60.[151,250,1159,1183]" captionTargetBox="[151,1436,181,1135]" captionTargetId="figure-28@60.[151,1436,181,1135]" captionTargetPageId="60" captionText="FIGURE 39. Andrena (Micrandrena) cryptaWarncke, 1975 nec. Viereck = A. warnckei Gusenleitner &amp; Schwarz, 2000 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209391" httpUri="https://zenodo.org/record/13209391/files/figure.png" pageId="60" pageNumber="61">Figure 39D</figureCitation>
) shows clear differences in the shape of the gonocoxal teeth (narrow and pointed in
<taxonomicName id="2E824D69AC65434708BFFA258A750049" authorityName="Warncke" authorityYear="1975" box="[939,1040,1518,1544]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC65434708BFFA258A750049" box="[939,1040,1518,1544]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
, broad and truncate in
<taxonomicName id="2E824D69AC6543470E05FA258BF60046" authorityName="Stockhert" authorityYear="1935" box="[1297,1427,1518,1544]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470E05FA258BF60046" box="[1297,1427,1518,1544]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
) and penis valves (relatively narrow in
<taxonomicName id="2E824D69AC6543470947F9D98CDE006D" authorityName="Warncke" authorityYear="1975" box="[595,699,1554,1580]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC6543470947F9D98CDE006D" box="[595,699,1554,1580]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
, strongly basally broadened and triangular in
<taxonomicName id="2E824D69AC6543470FDBF9D98B31006A" authorityName="Stockhert" authorityYear="1935" box="[1231,1364,1554,1580]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470FDBF9D98B31006A" box="[1231,1364,1554,1580]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
). One newly examined male specimen from
<collectingRegion id="2B46F808AC6543470956F9FD8CF10011" box="[578,660,1590,1616]" country="Ukraine" name="Crimea" pageId="60" pageNumber="61">Crimea</collectingRegion>
shows typical
<taxonomicName id="2E824D69AC6543470828F9FD8DDA000E" authorityName="Stockhert" authorityYear="1935" box="[828,959,1590,1616]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470828F9FD8DDA000E" box="[828,959,1590,1616]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
genitalia.
</paragraph>
<paragraph id="E93D36EAAC6543460BD3F9918A760694" blockId="60.[151,1437,1265,2012]" lastBlockId="61.[151,1436,151,213]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
It was possible to generate sequences (
<figureCitation id="71B92A6FAC654347096EF9918C8F0035" box="[634,746,1626,1652]" captionStart="FIGURE 37" captionStartId="57.[151,250,1819,1843]" captionTargetBox="[151,1433,311,1794]" captionTargetId="figure-78@57.[151,1435,309,1796]" captionTargetPageId="57" captionText="FIGURE 37. Phylogenetic tree (maximum likelihood) of Andrena subgenus Micrandrena Ashmead, 1899 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209387" httpUri="https://zenodo.org/record/13209387/files/figure.png" pageId="60" pageNumber="61">Figure 37</figureCitation>
) of
<taxonomicName id="2E824D69AC654347080CF9918D1B0035" authorityName="Warncke" authorityYear="1975" box="[792,894,1626,1652]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC654347080CF9918D1B0035" box="[792,894,1626,1652]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC6543470899F9918D960035" box="[909,1011,1626,1652]" name="Lebanon" pageId="60" pageNumber="61">Lebanon</collectingCountry>
) and
<taxonomicName id="2E824D69AC6543470F25F9918AD00032" authorityName="Stockhert" authorityYear="1935" box="[1073,1205,1626,1652]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470F25F9918AD00032" box="[1073,1205,1626,1652]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC6543470FD0F9918B600035" box="[1220,1285,1626,1652]" name="Spain" pageId="60" pageNumber="61">Spain</collectingCountry>
and
<collectingCountry id="9195767AAC6543470E29F9918BF40035" box="[1341,1425,1626,1652]" name="Greece" pageId="60" pageNumber="61">Greece</collectingCountry>
). The sequence of
<taxonomicName id="2E824D69AC6543470A4BF9B58FA200D9" authorityName="Warncke" authorityYear="1975" box="[351,455,1662,1688]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC6543470A4BF9B58FA200D9" box="[351,455,1662,1688]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
from
<collectingCountry id="9195767AAC6543470906F9B58C1100D9" box="[530,628,1662,1688]" name="Lebanon" pageId="60" pageNumber="61">Lebanon</collectingCountry>
was separated from Spanish and Greek
<taxonomicName id="2E824D69AC6543470F58F9B58AB700D6" authorityName="Stockhert" authorityYear="1935" box="[1100,1234,1662,1688]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470F58F9B58AB700D6" box="[1100,1234,1662,1688]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
by 6.67% (range 6.287.15%).
<taxonomicName id="2E824D69AC6543470A23F9698C6600FA" authorityName="Stockhert" authorityYear="1935" box="[311,515,1698,1724]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470A23F9698C6600FA" box="[311,515,1698,1724]" italics="true" pageId="60" pageNumber="61">Andrena saxonica</emphasis>
</taxonomicName>
itself showed average intraspecific variation of 2.49% (range 0.004.10%), with Spanish and Greek sequences forming clades with 89 and 98 bootstrap support, respectively. The division between the Spanish and Greek sequences is considered to represent separation by distance, since
<taxonomicName id="2E824D69AC6543470F7BF9218A940142" authorityName="Stockhert" authorityYear="1935" box="[1135,1265,1770,1796]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470F7BF9218A940142" box="[1135,1265,1770,1796]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
is found across most of Central Europe, and therefore Spanish and Greek specimens represent the east and west extremes of the European population. The same can be seen to a slightly lesser extent in
<taxonomicName id="2E824D69AC65434708D0F8F98B3A010D" authority="Noskiewicz, 1939" authorityName="Noskiewicz" authorityYear="1939" box="[964,1375,1842,1868]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="simontornyella">
<emphasis id="DBF6EAF8AC65434708D0F8F98AEE010D" box="[964,1163,1842,1868]" italics="true" pageId="60" pageNumber="61">A. simontornyella</emphasis>
<bibRefCitation id="8D134B1BAC6543470F86F8F98B3A010D" author="Noskiewicz, J." box="[1170,1375,1842,1868]" pageId="60" pageNumber="61" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz, 1939</bibRefCitation>
</taxonomicName>
, with average intraspecific variation of 1.90 (range 0.153.81%) between specimens from
<collectingCountry id="9195767AAC6543470F4AF89D8AC40131" box="[1118,1185,1878,1904]" name="Spain" pageId="60" pageNumber="61">Spain</collectingCountry>
,
<collectingCountry id="9195767AAC6543470FBAF89D8A980131" box="[1198,1277,1878,1904]" name="France" pageId="60" pageNumber="61">France</collectingCountry>
, and southern
<collectingCountry id="9195767AAC6543470B83F8B18E8F01D5" box="[151,234,1914,1940]" name="Greece" pageId="60" pageNumber="61">Greece</collectingCountry>
. The overall
<taxonomicName id="2E824D69AC6543470A6AF8B18C6701D2" authorityName="Stockhert" authorityYear="1935" box="[382,514,1914,1940]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470A6AF8B18C6701D2" box="[382,514,1914,1940]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
clade received bootstrap support of 90, and the inclusion of additional sequences from Central Europe (not currently available) is highly likely to reduce this average intraspecific variation further. In conclusion, Whilst
<taxonomicName id="2E824D69AC6543470A64F8098FB0019D" authorityName="Warncke" authorityYear="1975" box="[368,469,1986,2012]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC6543470A64F8098FB0019D" box="[368,469,1986,2012]" italics="true" pageId="60" pageNumber="61">A. sillata</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC6543470918F8098CEA019A" authorityName="Stockhert" authorityYear="1935" box="[524,655,1986,2012]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6543470918F8098CEA019A" box="[524,655,1986,2012]" italics="true" pageId="60" pageNumber="61">A. saxonica</emphasis>
</taxonomicName>
are clearly distinct species, material from Eastern Europe and eastern
<collectingCountry id="9195767AAC6443460B83FF5C8E8206F0" box="[151,231,151,177]" name="Turkey" pageId="61" pageNumber="62">Turkey</collectingCountry>
is considered to belong to
<taxonomicName id="2E824D69AC6443460903FF5C8CFE06F1" authorityName="Stockhert" authorityYear="1935" box="[535,667,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="saxonica">
<emphasis id="DBF6EAF8AC6443460903FF5C8CFE06F1" box="[535,667,151,177]" italics="true" pageId="61" pageNumber="62">A. saxonica</emphasis>
</taxonomicName>
.
<taxonomicName id="2E824D69AC64434609BDFF5C8D3006F0" authorityName="Warncke" authorityYear="1975" box="[681,853,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="sillata">
<emphasis id="DBF6EAF8AC64434609BDFF5C8D3006F0" box="[681,853,151,177]" italics="true" pageId="61" pageNumber="62">Andrena sillata</emphasis>
</taxonomicName>
is therefore considered to have a range restricted to
<collectingCountry id="9195767AAC6443460B83FF708E830694" box="[151,230,187,213]" name="Greece" pageId="61" pageNumber="62">Greece</collectingCountry>
(Rhodes), south-western
<collectingCountry id="9195767AAC6443460912FF708C320694" box="[518,599,187,213]" name="Turkey" pageId="61" pageNumber="62">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC6443460976FF708CAF0694" box="[610,714,187,213]" name="Lebanon" pageId="61" pageNumber="62">Lebanon</collectingCountry>
*,
<collectingCountry id="9195767AAC64434609F4FF708D450694" box="[736,800,187,213]" name="Syria" pageId="61" pageNumber="62">Syria</collectingCountry>
*,
<collectingCountry id="9195767AAC6443460823FF708DE20694" box="[823,903,187,213]" name="Jordan" pageId="61" pageNumber="62">Jordan</collectingCountry>
*, and
<collectingCountry id="9195767AAC64434608D9FF708A6A0694" box="[973,1039,187,213]" name="Israel" pageId="61" pageNumber="62">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:date id="11699C038481C070E262BEDC2F0E12A2">2024</mods:date>
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<mods:number id="D55EA1C2152771CF50D4D4A20AF0034A">2024-07-22</mods:number>
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<mods:identifier id="CC29576F7C3CC7984956D0D4A386939F" type="DOI">10.11646/zootaxa.5483.1.1</mods:identifier>
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<treatment id="612B87FCAC66433B0B83F9C98D8007EC" LSID="urn:lsid:plazi:treatment:612B87FCAC66433B0B83F9C98D8007EC" httpUri="http://treatment.plazi.org/id/612B87FCAC66433B0B83F9C98D8007EC" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<subSubSection id="A1986561AC6643440B83F9C98C83005D" box="[151,742,1538,1565]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="E93D36EAAC6643440B83F9C98C83005D" blockId="63.[151,1156,1538,1626]" box="[151,742,1538,1565]" pageId="63" pageNumber="64">
<heading id="B2758186AC6643440B83F9C98C83005D" bold="true" box="[151,742,1538,1565]" fontSize="11" level="2" pageId="63" pageNumber="64" reason="4">
<taxonomicName id="2E824D69AC6643440B83F9C98C83005D" ID-CoL="3ZKQH" authority="(Kirby, 1802)" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[151,742,1538,1565]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="afzeliella" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC6643440B83F9C98C83005D" bold="true" box="[151,742,1538,1565]" pageId="63" pageNumber="64">
<emphasis id="DBF6EAF8AC6643440B83F9C98F64005C" bold="true" box="[151,257,1538,1565]" italics="true" pageId="63" pageNumber="64">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6643440A06F9C98FDF005C" bold="true" box="[274,442,1538,1565]" italics="true" pageId="63" pageNumber="64">Taeniandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6643440AD8F9C98C5C005C" bold="true" box="[460,569,1538,1565]" italics="true" pageId="63" pageNumber="64">afzeliella</emphasis>
(Kirby, 1802)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6643440B83F9E88AE1001B" pageId="63" pageNumber="64" type="reference_group">
<paragraph id="E93D36EAAC6643440B83F9E88A4A007A" blockId="63.[151,1156,1538,1626]" box="[151,1071,1571,1595]" pageId="63" pageNumber="64">
<taxonomicName id="2E824D69AC6643440B83F9E88F9D007A" authority="Kirby, 1802: 169" authorityName="Kirby" authorityPageNumber="169" authorityYear="1802" box="[151,504,1571,1595]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="afzeliella">
<emphasis id="DBF6EAF8AC6643440B83F9E88F21007A" box="[151,324,1571,1595]" italics="true" pageId="63" pageNumber="64">Melitta afzeliella</emphasis>
<materialsCitation id="59EA3CB7AC6643440A5EF9E88F9D007A" box="[330,504,1571,1595]" collectionCode="NHMUK" country="United Kingdom" location="England" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1">Kirby, 1802: 169</materialsCitation>
</taxonomicName>
,
<specimenCount id="FF84FD63AC6643440917F9EF8C73007A" box="[515,534,1572,1595]" count="1" pageId="63" pageNumber="64" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC6643440936F9E88C1E007A" box="[546,635,1571,1595]" name="United Kingdom" pageId="63" pageNumber="64">England</collectingCountry>
, NHMUK, examined, see
<bibRefCitation id="8D134B1BAC664344089CF9EF8A4C007A" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[904,1065,1571,1595]" pageId="63" pageNumber="64" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC66434408AFF9EF8D8D007A" box="[955,1000,1571,1595]" italics="true" pageId="63" pageNumber="64">et al</emphasis>
. 2022
</bibRefCitation>
]
</paragraph>
<paragraph id="E93D36EAAC6643440B83F9898AE1001B" blockId="63.[151,1156,1538,1626]" box="[151,1156,1602,1626]" pageId="63" pageNumber="64">
<treatmentCitationGroup id="C99211C4AC6643440B83F9898AE1001B" box="[151,1156,1602,1626]" pageId="63" pageNumber="64">
<treatmentCitation id="682310FBAC6643440B83F9898C21001B" author="Cockerell, T. D. A." box="[151,580,1602,1626]" page="243" pageId="63" pageNumber="64" year="1922">
<taxonomicName id="2E824D69AC6643440B83F9898C21001B" ID-CoL="66N3X" authority="Cockerell, 1922: 243" authorityName="Cockerell" authorityPageNumber="243" authorityYear="1922" box="[151,580,1602,1626]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="heteropoda">
<emphasis id="DBF6EAF8AC6643440B83F9898F03001B" box="[151,358,1602,1626]" italics="true" pageId="63" pageNumber="64">Andrena heteropoda</emphasis>
<materialsCitation id="59EA3CB7AC6643440A79F9898C21001B" box="[365,580,1602,1626]" collectionCode="USNM" country="Pakistan" location="Pakistan" pageId="63" pageNumber="64" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC6643440A79F9898C21001B" author="Cockerell, T. D. A." box="[365,580,1602,1626]" pageId="63" pageNumber="64" pagination="49 - 54" refId="ref88273" refString="Cockerell, T. D. A. (1922) XXVI. - Descriptions and records of bees. - XCII. Annals and Magazine of Natural History, Series 9, 9, 49 - 54. https: // doi. org / 10.1080 / 00222932208632666" type="journal article" year="1922">Cockerell, 1922: 243</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC664344095AF9898C3B0018" box="[590,606,1602,1625]" count="1" pageId="63" pageNumber="64" type="female"></specimenCount>
<specimenCount id="FF84FD63AC664344094AF9898C110018" box="[606,628,1602,1625]" count="1" pageId="63" pageNumber="64" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6643440995F9898CB9001B" box="[641,732,1602,1626]" name="Pakistan" pageId="63" pageNumber="64">Pakistan</collectingCountry>
, USNM, photograph examined]
<emphasis id="DBF6EAF8AC6643440F33F9898AE1001B" bold="true" box="[1063,1156,1602,1626]" pageId="63" pageNumber="64">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC66433B0B83F9438FFE06B8" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64" type="discussion">
<paragraph id="E93D36EAAC66433B0B83F9438FFE06B8" blockId="63.[151,1437,1672,2023]" lastBlockId="64.[151,1436,151,429]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
<emphasis id="DBF6EAF8AC6643440B83F9438F6E00E3" bold="true" box="[151,267,1672,1698]" pageId="63" pageNumber="64">Remarks.</emphasis>
<bibRefCitation id="8D134B1BAC6643440A04F9438FA800E3" author="Cockerell, T. D. A." box="[272,461,1672,1698]" pageId="63" pageNumber="64" pagination="49 - 54" refId="ref88273" refString="Cockerell, T. D. A. (1922) XXVI. - Descriptions and records of bees. - XCII. Annals and Magazine of Natural History, Series 9, 9, 49 - 54. https: // doi. org / 10.1080 / 00222932208632666" type="journal article" year="1922">Cockerell (1922)</bibRefCitation>
described
<taxonomicName id="2E824D69AC6643440956F9438CBA00E3" authorityName="Cockerell" authorityYear="1922" box="[578,735,1672,1698]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="heteropoda">
<emphasis id="DBF6EAF8AC6643440956F9438CBA00E3" box="[578,735,1672,1698]" italics="true" pageId="63" pageNumber="64">A. heteropoda</emphasis>
</taxonomicName>
based on male and female specimens either taken from the C.G. Nurse collection (NHMUK) or received from him directly. The taxon clearly belongs to the subgenus
<taxonomicName id="2E824D69AC6643440E17F9678BF90087" authorityName="Hedicke" authorityYear="1933" box="[1283,1436,1708,1734]" pageId="63" pageNumber="64" rank="subGenus" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC6643440E17F9678BF90087" box="[1283,1436,1708,1734]" italics="true" pageId="63" pageNumber="64">Taeniandrena</emphasis>
</taxonomicName>
due to the flattened clypeus, and indeed Cockerell draws comparison with
<taxonomicName id="2E824D69AC66434408DBF91B8BFD00AB" authority="(Kirby, 1802)" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[975,1432,1744,1770]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="wilkella" subGenus="Taeniandrena">
<emphasis id="DBF6EAF8AC66434408DBF91B8D8D00AB" box="[975,1000,1744,1770]" italics="true" pageId="63" pageNumber="64">A.</emphasis>
(
<emphasis id="DBF6EAF8AC66434408E3F91B8AF500AB" box="[1015,1168,1744,1770]" italics="true" pageId="63" pageNumber="64">Taeniandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6643440F8BF91B8A9D00AB" box="[1183,1272,1744,1770]" italics="true" pageId="63" pageNumber="64">wilkella</emphasis>
(Kirby, 1802)
</taxonomicName>
. Based on the size (female
<quantity id="2E7A9B0FAC6643440AA9F93F8C69014E" box="[445,524,1780,1807]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="63" pageNumber="64" unit="mm" value="10.0">10 mm</quantity>
in length, male 8.5 mm in length), complete apical hairband on T3, light terminal fringe and light hairs flanking the basitibial plate, dull scutum, and moderately punctate terga,
<taxonomicName id="2E824D69AC6643440FCBF8D38BE50173" authorityName="Cockerell" authorityYear="1922" box="[1247,1408,1816,1842]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="heteropoda">
<emphasis id="DBF6EAF8AC6643440FCBF8D38BE50173" box="[1247,1408,1816,1842]" italics="true" pageId="63" pageNumber="64">A. heteropoda</emphasis>
</taxonomicName>
is clearly conspecific with
<taxonomicName id="2E824D69AC6643440ABAF8F78C530117" authorityName="Kirby" authorityYear="1802" box="[430,566,1852,1878]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="afzeliella">
<emphasis id="DBF6EAF8AC6643440ABAF8F78C530117" box="[430,566,1852,1878]" italics="true" pageId="63" pageNumber="64">A. afzeliella</emphasis>
</taxonomicName>
which was previously known as far east as
<collectingRegion id="2B46F808AC6643440F3BF8F68A960117" box="[1071,1267,1852,1879]" country="Iran" name="Kerman" pageId="63" pageNumber="64">Kerman province</collectingRegion>
in central
<collectingCountry id="9195767AAC6643440E7BF8F68BF90116" box="[1391,1436,1853,1879]" name="Iran" pageId="63" pageNumber="64">Iran</collectingCountry>
(
<bibRefCitation id="8D134B1BAC6643440B8BF8AB8F36013B" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[159,339,1888,1915]" pageId="63" pageNumber="64" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC6643440BCDF8AA8F6C013B" box="[217,265,1888,1914]" italics="true" pageId="63" pageNumber="64">et al</emphasis>
. 2022
</bibRefCitation>
). Cockerell mentions that one of the male specimens was dated
<date id="9D3C102AAC6643440F02F8AB8AE3013B" box="[1046,1158,1888,1914]" pageId="63" pageNumber="64" value="1903-07">July 1903</date>
, which is consistent with the known bivoltine behaviour of
<taxonomicName id="2E824D69AC6643440902F84F8CF901DF" authorityName="Kirby" authorityYear="1802" box="[534,668,1924,1950]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="afzeliella">
<emphasis id="DBF6EAF8AC6643440902F84F8CF901DF" box="[534,668,1924,1950]" italics="true" pageId="63" pageNumber="64">A. afzeliella</emphasis>
</taxonomicName>
with both spring (typically MayJune) and summer (typically July August) generations (
<bibRefCitation id="8D134B1BAC6643440A98F8638C2C0183" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[396,585,1960,1987]" pageId="63" pageNumber="64" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC6643440ADDF8628F990183" box="[457,508,1960,1986]" italics="true" pageId="63" pageNumber="64">et al</emphasis>
. 2022
</bibRefCitation>
). A series of previously undetermined
<taxonomicName id="2E824D69AC6643440F17F8638AE90183" authorityName="Kirby" authorityYear="1802" box="[1027,1164,1960,1986]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="afzeliella">
<emphasis id="DBF6EAF8AC6643440F17F8638AE90183" box="[1027,1164,1960,1986]" italics="true" pageId="63" pageNumber="64">A. afzeliella</emphasis>
</taxonomicName>
specimens from Quetta collected by Nurse between MarchJuly remain in the NHMUK collection, supporting this characterisation. The presence of
<taxonomicName id="2E824D69AC19433B0A0AFF5C8FC106F0" authorityName="Kirby" authorityYear="1802" box="[286,420,151,177]" class="Insecta" family="Melittidae" genus="Melitta" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="afzeliella">
<emphasis id="DBF6EAF8AC19433B0A0AFF5C8FC106F0" box="[286,420,151,177]" italics="true" pageId="64" pageNumber="65">A. afzeliella</emphasis>
</taxonomicName>
in western
<collectingCountry id="9195767AAC19433B0930FF5C8CE606F0" box="[548,643,151,177]" name="Pakistan" pageId="64" pageNumber="65">Pakistan</collectingCountry>
is consistent with recent finding of
<taxonomicName id="2E824D69AC19433B0F07FF5C8EAA0694" authority="Warncke, 1975" authorityName="Warncke" authorityYear="1975" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="saettana" subGenus="Troandrena">
<emphasis id="DBF6EAF8AC19433B0F07FF5C8A4E06F0" box="[1043,1067,151,177]" italics="true" pageId="64" pageNumber="65">A.</emphasis>
(
<emphasis id="DBF6EAF8AC19433B0F2FFF5C8AD806F0" box="[1083,1213,151,177]" italics="true" pageId="64" pageNumber="65">Troandrena</emphasis>
)
<emphasis id="DBF6EAF8AC19433B0FD9FF5C8B4906F1" box="[1229,1324,151,176]" italics="true" pageId="64" pageNumber="65">saettana</emphasis>
Warncke,
<quantity id="2E7A9B0FAC19433B0B83FF718EAA0694" box="[151,207,186,213]" metricMagnitude="1" metricUnit="m" metricValue="5.0165" pageId="64" pageNumber="65" unit="in" value="1975.0">1975</quantity>
</taxonomicName>
in Quetta, also collected by C.G. Nurse, when it was also thought to have an eastern range limit in Kerman province (
<bibRefCitation id="8D134B1BAC19433B0A13FF148FF506B8" author="Wood, T. J." box="[263,400,223,249]" pageId="64" pageNumber="65" pagination="167 - 188" refId="ref92492" refString="Wood, T. J. (2024) New Asian Andrena species, with notes on the subgenus Cnemidandrena (Hymenoptera: Andrenidae). Zootaxa, 5404 (1), 167 - 188. https: // doi. org / 10.11646 / zootaxa. 5404.1.11" type="journal article" year="2024">Wood 2024</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC19433B0BD3FEC98A4D07C8" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="E93D36EAAC19433B0BD3FEC98A4D07C8" blockId="64.[151,1436,151,429]" pageId="64" pageNumber="65">
<emphasis id="DBF6EAF8AC19433B0BD3FEC98FD4075D" bold="true" box="[199,433,258,284]" pageId="64" pageNumber="65">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC19433B0AA3FEC98CE90700" collectionCode="USNM" collectorName="C. G. Nurse" country="Pakistan" location="Quetta" pageId="64" pageNumber="65" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<emphasis id="DBF6EAF8AC19433B0AA3FEC98C2B075D" bold="true" box="[439,590,258,284]" pageId="64" pageNumber="65">
<collectingCountry id="9195767AAC19433B0AA3FEC98C2D075D" box="[439,584,258,284]" name="Pakistan" pageId="64" pageNumber="65">PAKISTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC19433B0940FEC88CC7075C" LSID="urn:lsid:plazi:treatment:612B87FCAC66433B0B83F9C98D8007EC:EC5D6031AC19433B0940FEC88CC7075C" box="[596,674,259,285]" country="Pakistan" name="Quetta" pageId="64" pageNumber="65">Quetta</location>
, [undated],
<specimenCount id="FF84FD63AC19433B0832FEC88D2C075C" box="[806,841,259,285]" count="1" pageId="64" pageNumber="65" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC19433B089CFEC88A6C075C" box="[904,1033,259,285]" pageId="64" pageNumber="65">C.G. Nurse</collectorName>
,
<collectionCode id="8F93AE2FAC19433B0F00FEC88A00075C" box="[1044,1125,259,285]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="64" pageNumber="65" type="Museum">USNM</collectionCode>
(
<emphasis id="DBF6EAF8AC19433B0F62FEC88AB7075C" bold="true" box="[1142,1234,259,285]" pageId="64" pageNumber="65">
<typeStatus id="36398848AC19433B0F62FEC88AB7075C" box="[1142,1234,259,285]" pageId="64" pageNumber="65" type="syntype">syntype</typeStatus>
</emphasis>
of
<taxonomicName id="2E824D69AC19433B0FE3FEC88BF1075C" authorityName="Cockerell" authorityYear="1922" box="[1271,1428,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="heteropoda">
<emphasis id="DBF6EAF8AC19433B0FE3FEC88BF1075C" box="[1271,1428,259,285]" italics="true" pageId="64" pageNumber="65">A. heteropoda</emphasis>
</taxonomicName>
; type no. 25053, examined by photograph)
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, leg.
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,
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,
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,
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,
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;
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<location id="EC5D6031AC19433B0AD9FEA48C7E07C8" LSID="urn:lsid:plazi:treatment:612B87FCAC66433B0B83F9C98D8007EC:EC5D6031AC19433B0AD9FEA48C7E07C8" box="[461,539,367,393]" country="Pakistan" name="Quetta" pageId="64" pageNumber="65">Quetta</location>
,
<date id="9D3C102AAC19433B0931FEA48CD807C8" box="[549,701,367,393]" pageId="64" pageNumber="65" value="1903-03-01" valueMax="1903-03-31" valueMin="1903-03-01">
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, leg.
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,
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</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC19433B0BD3FE598D8007EC" box="[199,997,402,429]" pageId="64" pageNumber="65" type="distribution">
<paragraph id="E93D36EAAC19433B0BD3FE598D8007EC" blockId="64.[151,1436,151,429]" box="[199,997,402,429]" pageId="64" pageNumber="65">
<emphasis id="DBF6EAF8AC19433B0BD3FE598F0407ED" bold="true" box="[199,353,402,428]" pageId="64" pageNumber="65">Distribution.</emphasis>
West and probably Central Palaearctic (
<bibRefCitation id="8D134B1BAC19433B0837FE588DBC07EC" author="Praz, C. &amp; Genoud, D. &amp; Vaucher, K. &amp; Benon, D. &amp; Monks, J. &amp; Wood, T. J." box="[803,985,403,429]" pageId="64" pageNumber="65" pagination="375 - 428" refId="ref90337" refString="Praz, C., Genoud, D., Vaucher, K., Benon, D., Monks, J. &amp; Wood, T. J. (2022) Unexpected levels of cryptic diversity in European bees of the genus Andrena subgenus Taeniandrena (Hymenoptera, Andrenidae): implications for conservation. Journal of Hymenoptera Research, 91, 375 - 428. https: // doi. org / 10.3897 / jhr. 91.82761" type="journal article" year="2022">
Praz
<emphasis id="DBF6EAF8AC19433B0849FE588DEB07EC" box="[861,910,403,429]" italics="true" pageId="64" pageNumber="65">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
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</treatment>
</document>

View file

@ -0,0 +1,265 @@
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<emphasis id="DBF6EAF8AC6643440B83FC898F64051C" bold="true" box="[151,257,834,861]" italics="true" pageId="63" pageNumber="64">Andrena</emphasis>
(
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)
<emphasis id="DBF6EAF8AC6643440AA0FC888C28051C" bold="true" box="[436,589,835,861]" italics="true" pageId="63" pageNumber="64">trimarginata</emphasis>
(
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<emphasis id="DBF6EAF8AC6643440B83FC4C8F1805DE" box="[151,381,903,927]" italics="true" pageId="63" pageNumber="64">Halictus trimarginatus</emphasis>
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,
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[
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, ZMHB, examined] (
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<paragraph id="E93D36EAAC6643440B83FC6D8A2D05FF" blockId="63.[151,1302,903,989]" box="[151,1096,934,960]" pageId="63" pageNumber="64">
<treatmentCitationGroup id="C99211C4AC6643440B83FC6D8A2D05FF" box="[151,1096,934,960]" pageId="63" pageNumber="64">
<treatmentCitation id="682310FBAC6643440B83FC6D8CC105FF" author="Warncke, K." box="[151,676,934,958]" page="99" pageId="63" pageNumber="64" year="1975">
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<emphasis id="DBF6EAF8AC6643440B83FC6D8E8805FF" box="[151,237,934,958]" italics="true" pageId="63" pageNumber="64">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6643440BE8FC6D8F1205FF" box="[252,375,934,958]" italics="true" pageId="63" pageNumber="64">Carandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6643440A92FC6C8FAA05FF" box="[390,463,935,958]" italics="true" pageId="63" pageNumber="64">zostera</emphasis>
<materialsCitation id="59EA3CB7AC6643440AC1FC6D8CC105FF" box="[469,676,934,958]" collectionCode="OOLM" country="Turkey" location="Turkey" pageId="63" pageNumber="64" specimenCount="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC6643440AC1FC6D8CC105FF" author="Warncke, K." box="[469,676,934,958]" pageId="63" pageNumber="64" pagination="29 - 102" refId="ref91833" refString="Warncke, K. (1975 b) Die Sandbienen der Turkei (Hymenoptera, Apoidea, Andrena), Teil B. Mitteilungen der Munchner Entomologischen Gesellschaft, 65, 29 - 102." type="journal article" year="1975">Warncke, 1975b: 99</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC66434409BAFC6D8CA405FC" box="[686,705,934,957]" count="1" pageId="63" pageNumber="64" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC66434409D9FC6D8D7C05FF" box="[717,793,934,958]" name="Turkey" pageId="63" pageNumber="64">Turkey</collectingCountry>
, OÖLM, examined]
<emphasis id="DBF6EAF8AC66434408FFFC6D8A2D05FF" bold="true" box="[1003,1096,934,958]" pageId="63" pageNumber="64">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC6643440B83FC0F8B73059C" blockId="63.[151,1302,903,989]" box="[151,1302,964,989]" pageId="63" pageNumber="64">
<treatmentCitationGroup id="C99211C4AC6643440B83FC0F8B73059C" box="[151,1302,964,989]" pageId="63" pageNumber="64">
<treatmentCitation id="682310FBAC6643440B83FC0F8D61059C" author="Osytshnjuk, A. Z." box="[151,772,964,989]" page="13" pageId="63" pageNumber="64" year="1984">
<taxonomicName id="2E824D69AC6643440B83FC0F8D61059C" ID-CoL="DT3T" authority="Osytshnjuk, 1984: 13" authorityName="Osytshnjuk" authorityPageNumber="13" authorityYear="1984" box="[151,772,964,989]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="subsmaragdina" subGenus="Carandrena">
<emphasis id="DBF6EAF8AC6643440B83FC0F8E88059D" box="[151,237,964,988]" italics="true" pageId="63" pageNumber="64">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6643440BE8FC0F8F12059D" box="[252,375,964,988]" italics="true" pageId="63" pageNumber="64">Carandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6643440A92FC0F8C47059D" box="[390,546,964,988]" italics="true" pageId="63" pageNumber="64">subsmaragdina</emphasis>
<materialsCitation id="59EA3CB7AC664344093DFC0F8D61059C" box="[553,772,964,989]" collectionCode="SIZK" country="Turkmenistan" location="Turkmenistan" pageId="63" pageNumber="64" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC664344093DFC0F8D61059C" author="Osytshnjuk, A. Z." box="[553,772,964,989]" pageId="63" pageNumber="64" pagination="3 - 15" refId="ref89639" refString="Osytshnjuk, A. Z. (1984) New and little known bees of the subgenus Carandrena War. (Hymenoptera, Andrenidae, Andrena F.) in the fauna of the USSR. Trudy Zoologicheskogo Instituta, Leningrad, 128, 3 - 15. [in Russian]" type="journal article" year="1984">Osytshnjuk, 1984: 13</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC664344081BFC0E8D7A059D" box="[783,799,965,988]" count="1" pageId="63" pageNumber="64" type="female"></specimenCount>
<specimenCount id="FF84FD63AC664344080AFC0E8D51059D" box="[798,820,965,988]" count="1" pageId="63" pageNumber="64" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6643440855FC0F8DB7059D" box="[833,978,964,988]" name="Turkmenistan" pageId="63" pageNumber="64">Turkmenistan</collectingCountry>
, SIZK, not examined]
<emphasis id="DBF6EAF8AC6643440FADFC0E8B73059C" bold="true" box="[1209,1302,965,989]" pageId="63" pageNumber="64">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6643440B83FBC08A3A02BC" pageId="63" pageNumber="64" type="discussion">
<paragraph id="E93D36EAAC6643440B83FBC08A3A02BC" blockId="63.[151,1437,1035,1457]" pageId="63" pageNumber="64">
<emphasis id="DBF6EAF8AC6643440B83FBC08F6E0264" bold="true" box="[151,267,1035,1061]" pageId="63" pageNumber="64">Remarks.</emphasis>
<bibRefCitation id="8D134B1BAC6643440A06FBC08FA20264" author="Warncke, K." box="[274,455,1035,1061]" pageId="63" pageNumber="64" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
and
<bibRefCitation id="8D134B1BAC6643440AEBFBC08D000264" author="Gusenleitner, F. &amp; Schwarz, M." box="[511,869,1035,1061]" pageId="63" pageNumber="64" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
indicated that the
<typeStatus id="36398848AC6643440F21FBC08A000264" box="[1077,1125,1035,1061]" pageId="63" pageNumber="64">type</typeStatus>
material of
<taxonomicName id="2E824D69AC6643440FFBFBC08BF90265" baseAuthorityName="Radoszkowski" baseAuthorityYear="1886" box="[1263,1436,1035,1061]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="trimarginata">
<emphasis id="DBF6EAF8AC6643440FFBFBC08BF90265" box="[1263,1436,1035,1061]" italics="true" pageId="63" pageNumber="64">A. trimarginata</emphasis>
</taxonomicName>
was present in the ZMHB, consisting of a head and mesosoma fragment only. This fragment was examined (
<figureCitation id="71B92A6FAC6643440E45FBE48ED3022C" captionStart="FIGURE 42" captionStartId="64.[151,250,1434,1458]" captionTargetBox="[151,1436,453,1411]" captionTargetId="figure-139@64.[151,1436,453,1411]" captionTargetPageId="64" captionText="FIGURE 42. Halictus trimarginatus Radoszkowski, 1886 holotype female.A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsolateral view." figureDoi="http://doi.org/10.5281/zenodo.13209397" httpUri="https://zenodo.org/record/13209397/files/figure.png" pageId="63" pageNumber="64">Figure 42</figureCitation>
), and confirmed its identity as a small
<emphasis id="DBF6EAF8AC6643440994FB988D60022C" box="[640,773,1107,1133]" italics="true" pageId="63" pageNumber="64">Notandrena</emphasis>
species with metallic green-purple reflections. Described from
<collectingRegion id="2B46F808AC6643440BC6FBBC8F5902D0" box="[210,316,1143,1169]" country="Turkmenistan" name="Ashgabat" pageId="63" pageNumber="64">Ashgabat</collectingRegion>
in
<collectingCountry id="9195767AAC6643440A49FBBC8F9E02D0" box="[349,507,1143,1169]" name="Turkmenistan" pageId="63" pageNumber="64">Turkmenistan</collectingCountry>
, this fragment is sufficient to allow recognition that this material is conspecific with
<taxonomicName id="2E824D69AC6643440B83FB508F6C02F5" authorityName="Warncke" authorityYear="1975" box="[151,265,1179,1205]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zostera">
<emphasis id="DBF6EAF8AC6643440B83FB508F6C02F5" box="[151,265,1179,1205]" italics="true" pageId="63" pageNumber="64">A. zostera</emphasis>
</taxonomicName>
. The revision of
<bibRefCitation id="8D134B1BAC6643440AD9FB508C9202F4" author="Wood, T. J. &amp; Monfared, A." box="[461,759,1179,1205]" pageId="63" pageNumber="64" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared (2022)</bibRefCitation>
demonstrated that
<taxonomicName id="2E824D69AC66434408C5FB508A2702F5" authorityName="Warncke" authorityYear="1975" box="[977,1090,1179,1205]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="zostera">
<emphasis id="DBF6EAF8AC66434408C5FB508A2702F5" box="[977,1090,1179,1205]" italics="true" pageId="63" pageNumber="64">A. zostera</emphasis>
</taxonomicName>
was the senior synonym of
<taxonomicName id="2E824D69AC6643440E90FB508C770298" authority="Osytshnjuk, 1984" authorityName="Osytshnjuk" authorityYear="1984" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="subsmaragdina">
<emphasis id="DBF6EAF8AC6643440E90FB508F210298" italics="true" pageId="63" pageNumber="64">A. subsmaragdina</emphasis>
<bibRefCitation id="8D134B1BAC6643440A5EFB748C770298" author="Osytshnjuk, A. Z." box="[330,530,1215,1241]" pageId="63" pageNumber="64" pagination="3 - 15" refId="ref89639" refString="Osytshnjuk, A. Z. (1984) New and little known bees of the subgenus Carandrena War. (Hymenoptera, Andrenidae, Andrena F.) in the fauna of the USSR. Trudy Zoologicheskogo Instituta, Leningrad, 128, 3 - 15. [in Russian]" type="journal article" year="1984">Osytshnjuk, 1984</bibRefCitation>
</taxonomicName>
, which was considered to be the only metallic green
<emphasis id="DBF6EAF8AC6643440F4FFB748A850298" box="[1115,1248,1215,1241]" italics="true" pageId="63" pageNumber="64">Notandrena</emphasis>
found in Central Asia.
<taxonomicName id="2E824D69AC6643440BCDFB288FA802BD" baseAuthorityName="Radoszkowski" baseAuthorityYear="1886" box="[217,461,1251,1277]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="trimarginata">
<emphasis id="DBF6EAF8AC6643440BCDFB288FA802BD" box="[217,461,1251,1277]" italics="true" pageId="63" pageNumber="64">Andrena trimarginata</emphasis>
</taxonomicName>
therefore becomes the senior name for both taxa
<emphasis id="DBF6EAF8AC66434408E3FB288A3A02BC" bold="true" box="[1015,1119,1251,1277]" pageId="63" pageNumber="64">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6643440BD3FACD8D840328" pageId="63" pageNumber="64" type="materials_examined">
<paragraph id="E93D36EAAC6643440BD3FACD8D840328" blockId="63.[151,1437,1035,1457]" pageId="63" pageNumber="64">
<emphasis id="DBF6EAF8AC6643440BD3FACD8FCB0361" bold="true" box="[199,430,1286,1312]" pageId="63" pageNumber="64">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC6643440AA6FACD8BF30360" box="[434,1430,1286,1313]" collectionCode="ZMHB" country="Turkmenistan" location="Ashabad" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1" stateProvince="Ashgabat" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6643440AA6FACD8CC40361" bold="true" box="[434,673,1286,1312]" pageId="63" pageNumber="64">
<collectingCountry id="9195767AAC6643440AA6FACD8CF90361" box="[434,668,1286,1312]" name="Turkmenistan" pageId="63" pageNumber="64">TURKMENISTAN</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC66434409B2FACC8D6D0360" box="[678,776,1287,1313]" country="Turkmenistan" name="Ashgabat" pageId="63" pageNumber="64">Ashabad</collectingRegion>
[
<collectingRegion id="2B46F808AC6643440807FACC8DE60360" box="[787,899,1287,1313]" country="Turkmenistan" name="Ashgabat" pageId="63" pageNumber="64">Ashgabat</collectingRegion>
; no further information],
<specimenCount id="FF84FD63AC6643440F83FACC8ADF0360" box="[1175,1210,1287,1313]" count="1" pageId="63" pageNumber="64" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FAC6643440FD2FACC8B720360" box="[1222,1303,1287,1313]" pageId="63" pageNumber="64">ZMHB</collectionCode>
(
<emphasis id="DBF6EAF8AC6643440E30FACC8BEE0360" bold="true" box="[1316,1419,1287,1313]" pageId="63" pageNumber="64">
<typeStatus id="36398848AC6643440E30FACC8BEE0360" box="[1316,1419,1287,1313]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6643440B83FAE18DFB0304" box="[151,926,1322,1349]" collectingDate="1902-04-01" collectingDateMax="1902-04-30" collectingDateMin="1902-04-01" collectionCode="NHMUK" collectorName="C. G. Nurse" country="Pakistan" location="Quetta" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6643440B83FAE18F480305" bold="true" box="[151,301,1322,1348]" pageId="63" pageNumber="64">
<collectingCountry id="9195767AAC6643440B83FAE18F4D0305" box="[151,296,1322,1348]" name="Pakistan" pageId="63" pageNumber="64">PAKISTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC6643440A23FAE08FE00304" LSID="urn:lsid:plazi:treatment:612B87FCAC6643440B83FC898CD003F0:EC5D6031AC6643440A23FAE08FE00304" box="[311,389,1323,1349]" country="Pakistan" name="Quetta" pageId="63" pageNumber="64">Quetta</location>
,
<date id="9D3C102AAC6643440A87FAE08C420304" box="[403,551,1323,1349]" pageId="63" pageNumber="64" value="1902-04-01" valueMax="1902-04-30" valueMin="1902-04-01">
<collectingDate id="8D78E9C2AC6643440A87FAE08C420304" box="[403,551,1323,1349]" pageId="63" pageNumber="64" value="1902-04-01" valueMax="1902-04-30" valueMin="1902-04-01">130.iv.1902</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6643440922FAE08C3C0304" box="[566,601,1323,1349]" count="1" pageId="63" pageNumber="64" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC664344098AFAE08D460304" box="[670,803,1323,1349]" pageId="63" pageNumber="64">C.G. Nurse</collectorName>
,
<collectionCode id="8F93AE2FAC6643440825FAE08DFB0304" box="[817,926,1323,1349]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="63" pageNumber="64" type="Museum">NHMUK</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC66434408B9FAE08F600328" collectingDate="1903-04-01" collectingDateMax="1903-04-30" collectingDateMin="1903-04-01" collectionCode="NHMUK" collectorName="C. G. Nurse" country="Pakistan" location="Quetta" pageId="63" pageNumber="64" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<location id="EC5D6031AC66434408B9FAE08D9E0304" LSID="urn:lsid:plazi:treatment:612B87FCAC6643440B83FC898CD003F0:EC5D6031AC66434408B9FAE08D9E0304" box="[941,1019,1323,1349]" country="Pakistan" name="Quetta" pageId="63" pageNumber="64">Quetta</location>
,
<date id="9D3C102AAC6643440F1DFAE08AF80304" box="[1033,1181,1323,1349]" pageId="63" pageNumber="64" value="1903-04-01" valueMax="1903-04-30" valueMin="1903-04-01">
<collectingDate id="8D78E9C2AC6643440F1DFAE08AF80304" box="[1033,1181,1323,1349]" pageId="63" pageNumber="64" value="1903-04-01" valueMax="1903-04-30" valueMin="1903-04-01">130.iv.1903</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6643440FB8FAE08AAA0304" box="[1196,1231,1323,1349]" count="1" pageId="63" pageNumber="64" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6643440E07FAE08BFD0304" box="[1299,1432,1323,1349]" pageId="63" pageNumber="64">C.G. Nurse</collectorName>
,
<collectionCode id="8F93AE2FAC6643440B83FA848F600328" box="[151,261,1359,1385]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="63" pageNumber="64" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
<materialsCitation id="59EA3CB7AC6643440A1BFA848DB90328" box="[271,988,1359,1385]" country="Pakistan" location="See" pageId="63" pageNumber="64" specimenCount="1" typeStatus="holotype">
<location id="EC5D6031AC6643440A1BFA848F520328" LSID="urn:lsid:plazi:treatment:612B87FCAC6643440B83FC898CD003F0:EC5D6031AC6643440A1BFA848F520328" box="[271,311,1359,1385]" country="Pakistan" name="See" pageId="63" pageNumber="64">See</location>
<bibRefCitation id="8D134B1BAC6643440A2AFA848C010328" author="Wood, T. J. &amp; Monfared, A." box="[318,612,1359,1385]" pageId="63" pageNumber="64" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared (2022)</bibRefCitation>
for additional examined material
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6643440BD3FAB98CD003F0" pageId="63" pageNumber="64" type="distribution">
<paragraph id="E93D36EAAC6643440BD3FAB98CD003F0" blockId="63.[151,1437,1035,1457]" pageId="63" pageNumber="64">
<emphasis id="DBF6EAF8AC6643440BD3FAB98F0403CD" bold="true" box="[199,353,1394,1420]" pageId="63" pageNumber="64">Distribution.</emphasis>
<collectingCountry id="9195767AAC6643440A79FAB88FCE03CC" box="[365,427,1395,1421]" name="Israel" pageId="63" pageNumber="64">Israel</collectingCountry>
and the
<collectingCountry id="9195767AAC664344090EFAB88CF903CC" box="[538,668,1395,1421]" name="West Bank" pageId="63" pageNumber="64">West Bank</collectingCountry>
,
<collectingCountry id="9195767AAC66434409B9FAB88C8E03CC" box="[685,747,1395,1421]" name="Syria" pageId="63" pageNumber="64">Syria</collectingCountry>
,
<collectingCountry id="9195767AAC66434409EFFAB88D2903CC" box="[763,844,1395,1421]" name="Turkey" pageId="63" pageNumber="64">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC664344084FFAB88DBC03CC" box="[859,985,1395,1421]" name="Azerbaijan" pageId="63" pageNumber="64">Azerbaijan</collectingCountry>
,
<collectingCountry id="9195767AAC66434408FDFAB88A7C03CC" box="[1001,1049,1395,1421]" name="Iraq" pageId="63" pageNumber="64">Iraq</collectingCountry>
,
<collectingCountry id="9195767AAC6643440F3DFAB88A3C03CC" box="[1065,1113,1395,1421]" name="Iran" pageId="63" pageNumber="64">Iran</collectingCountry>
,
<collectingCountry id="9195767AAC6643440F7DFAB88B6203CC" box="[1129,1287,1395,1421]" name="Turkmenistan" pageId="63" pageNumber="64">Turkmenistan</collectingCountry>
,
<collectingCountry id="9195767AAC6643440E03FAB88BFD03CC" box="[1303,1432,1395,1421]" name="Uzbekistan" pageId="63" pageNumber="64">Uzbekistan</collectingCountry>
,
<collectingCountry id="9195767AAC6643440B83FA5C8F6D03F0" box="[151,264,1431,1457]" name="Tajikistan" pageId="63" pageNumber="64">Tajikistan</collectingCountry>
,
<collectingCountry id="9195767AAC6643440A07FA5C8F1003F0" box="[275,373,1431,1457]" name="Pakistan" pageId="63" pageNumber="64">Pakistan</collectingCountry>
* (
<bibRefCitation id="8D134B1BAC6643440A9AFA5C8CCC03F0" author="Wood, T. J. &amp; Monfared, A." box="[398,681,1431,1457]" pageId="63" pageNumber="64" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="BE6A60B1E98E7D607D25228A5E63C1A9">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<subSubSection id="A1986561AC68434A0B83FBDB8D58026B" box="[151,829,1040,1067]" pageId="49" pageNumber="50" type="nomenclature">
<paragraph id="E93D36EAAC68434A0B83FBDB8D58026B" blockId="49.[151,829,1040,1067]" box="[151,829,1040,1067]" pageId="49" pageNumber="50">
<heading id="B2758186AC68434A0B83FBDB8D58026B" bold="true" box="[151,829,1040,1067]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
<taxonomicName id="2E824D69AC68434A0B83FBDB8D58026B" ID-CoL="DSSQ" authority="Osytshnjuk, 1995" authorityName="Osytshnjuk" authorityYear="1995" box="[151,829,1040,1067]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A0B83FBDB8D58026B" bold="true" box="[151,829,1040,1067]" pageId="49" pageNumber="50">
<emphasis id="DBF6EAF8AC68434A0B83FBDB8F64026A" bold="true" box="[151,257,1040,1067]" italics="true" pageId="49" pageNumber="50">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC68434A0A06FBDB8FD6026A" bold="true" box="[274,435,1040,1067]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
)
<emphasis id="DBF6EAF8AC68434A0AD1FBDB8C3E026A" bold="true" box="[453,603,1040,1067]" italics="true" pageId="49" pageNumber="50">romankovae</emphasis>
<bibRefCitation id="8D134B1BAC68434A0976FBDB8D58026B" author="Osytshnjuk, A. Z." box="[610,829,1040,1066]" pageId="49" pageNumber="50" pagination="489 - 527" refId="ref89691" refString="Osytshnjuk, A. Z. (1995) Family Andrenidae. In: Lehr, P. A. (Ed.), Keys to Insects of the Far East of Russia. Vol. 4. Megaloptera, Raphidioptera, Neuroptera, Mecoptera, Hymenoptera. Pt 1. Nauka, St. Petersburg, pp. 489 - 527. [in Russian]" type="book chapter" year="1995">Osytshnjuk, 1995</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC68434A0B83FB9E8A9802CD" pageId="49" pageNumber="50" type="reference_group">
<paragraph id="E93D36EAAC68434A0B83FB9E8A9802CD" blockId="49.[151,1376,1109,1164]" pageId="49" pageNumber="50">
<treatmentCitation id="682310FBAC68434A0B83FB9E8C99022C" author="Osytshnjuk, A. Z." box="[151,764,1109,1133]" page="501" pageId="49" pageNumber="50" year="1995">
<taxonomicName id="2E824D69AC68434A0B83FB9E8C99022C" ID-CoL="DSSQ" authority="Osytshnjuk, 1995: 501" authorityName="Osytshnjuk" authorityPageNumber="501" authorityYear="1995" box="[151,764,1109,1133]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A0B83FB9E8E88022C" box="[151,237,1109,1133]" italics="true" pageId="49" pageNumber="50">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC68434A0BE8FB9E8FE7022C" box="[252,386,1109,1133]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
)
<emphasis id="DBF6EAF8AC68434A0A85FB9E8C68022C" box="[401,525,1109,1133]" italics="true" pageId="49" pageNumber="50">romankovae</emphasis>
<bibRefCitation id="8D134B1BAC68434A0900FB9E8C99022C" author="Osytshnjuk, A. Z." box="[532,764,1109,1133]" pageId="49" pageNumber="50" pagination="489 - 527" refId="ref89691" refString="Osytshnjuk, A. Z. (1995) Family Andrenidae. In: Lehr, P. A. (Ed.), Keys to Insects of the Far East of Russia. Vol. 4. Megaloptera, Raphidioptera, Neuroptera, Mecoptera, Hymenoptera. Pt 1. Nauka, St. Petersburg, pp. 489 - 527. [in Russian]" type="book chapter" year="1995">Osytshnjuk, 1995: 501</bibRefCitation>
</taxonomicName>
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,
<specimenCount id="FF84FD63AC68434A0813FB9E8D7F022D" box="[775,794,1109,1132]" count="1" pageId="49" pageNumber="50" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC68434A0832FB9E8D15022C" box="[806,880,1109,1133]" name="Russia" pageId="49" pageNumber="50">Russia</collectingCountry>
: Far East, SIZK, not examined] (
<figureCitation id="71B92A6FAC68434A0FA9FB9E8B3C022C" box="[1213,1369,1109,1133]" captionStart="FIGURE 32" captionStartId="50.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-28@50.[151,1436,181,1618]" captionTargetPageId="50" captionText="FIGURE 32. Andrena (Hoplandrena) romankovae Osytshnjuk, 1995 paratype female.A. Label details; B. Face, frontal view; C. Process of labrum, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209377" httpUri="https://zenodo.org/record/13209377/files/figure.png" pageId="49" pageNumber="50">Figures 32AF</figureCitation>
)
<treatmentCitation id="682310FBAC68434A0B83FBBF8C9402CD" author="Xu, H. - L. &amp; Tadauchi, O." box="[151,753,1139,1164]" page="32" pageId="49" pageNumber="50" year="2005">
<taxonomicName id="2E824D69AC68434A0B83FBBF8C9402CD" ID-CoL="DRM3" authority="Xu &amp; Tadauchi, 2005: 32" authorityName="Xu &amp; Tadauchi" authorityPageNumber="32" authorityYear="2005" box="[151,753,1139,1164]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="fagopyri" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A0B83FBBF8E8802CD" box="[151,237,1140,1164]" italics="true" pageId="49" pageNumber="50">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC68434A0BE8FBBF8FE702CD" box="[252,386,1140,1164]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
)
<emphasis id="DBF6EAF8AC68434A0A85FBBF8F8202CD" box="[401,487,1140,1164]" italics="true" pageId="49" pageNumber="50">fagopyri</emphasis>
<bibRefCitation id="8D134B1BAC68434A0AF9FBBF8C9402CD" author="Xu, H. - L. &amp; Tadauchi, O." box="[493,753,1139,1164]" pageId="49" pageNumber="50" pagination="19 - 40" refId="ref92746" refString="Xu, H. - L. &amp; Tadauchi, O. (2005) A revision of the subgenus Hoplandrena of the genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 45, 19 - 40. https: // doi. org / 10.5109 / 2700" type="journal article" year="2005">Xu &amp; Tadauchi, 2005: 32</bibRefCitation>
</taxonomicName>
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,
<specimenCount id="FF84FD63AC68434A09EFFBBF8D6E02CA" box="[763,779,1140,1163]" count="1" pageId="49" pageNumber="50" type="female"></specimenCount>
<specimenCount id="FF84FD63AC68434A081FFBBF8D4402CA" box="[779,801,1140,1163]" count="1" pageId="49" pageNumber="50" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC68434A083AFBBF8D1402CD" box="[814,881,1140,1164]" name="China" pageId="49" pageNumber="50">China</collectingCountry>
:
<collectingRegion id="2B46F808AC68434A086EFBBF8DDC02CD" box="[890,953,1140,1164]" country="China" name="Hebei" pageId="49" pageNumber="50">Hebei</collectingRegion>
, IZAS, not examined]
<emphasis id="DBF6EAF8AC68434A0FB4FBBF8A9802CD" bold="true" box="[1184,1277,1140,1164]" pageId="49" pageNumber="50">syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC68434A0B83FB718A1E011D" pageId="49" pageNumber="50" type="discussion">
<paragraph id="E93D36EAAC68434A0B83FB718DAB0021" blockId="49.[151,1437,1210,1884]" pageId="49" pageNumber="50">
<emphasis id="DBF6EAF8AC68434A0B83FB718F6E0295" bold="true" box="[151,267,1210,1236]" pageId="49" pageNumber="50">Remarks.</emphasis>
In revising the
<taxonomicName id="2E824D69AC68434A0AA1FB718C2C0295" box="[437,585,1210,1236]" pageId="49" pageNumber="150" rank="subGenus" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A0AA1FB718C2C0295" box="[437,585,1210,1236]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
</taxonomicName>
of
<collectingCountry id="9195767AAC68434A0978FB718CB30295" box="[620,726,1210,1236]" name="China" pageId="49" pageNumber="50">East Asia</collectingCountry>
,
<bibRefCitation id="8D134B1BAC68434A09F4FB718DBF0294" author="Xu, H. - L. &amp; Tadauchi, O." box="[736,986,1210,1237]" pageId="49" pageNumber="50" pagination="19 - 40" refId="ref92746" refString="Xu, H. - L. &amp; Tadauchi, O. (2005) A revision of the subgenus Hoplandrena of the genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 45, 19 - 40. https: // doi. org / 10.5109 / 2700" type="journal article" year="2005">Xu &amp; Tadauchi (2005)</bibRefCitation>
examined all relevant type material with the exception of
<taxonomicName id="2E824D69AC68434A0A5BFB158F9302B9" authorityName="Osytshnjuk" authorityYear="1995" box="[335,502,1246,1272]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A0A5BFB158F9302B9" box="[335,502,1246,1272]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
which was described from the Russian Far East (
<bibRefCitation id="8D134B1BAC68434A0F07FB158AB302B8" author="Osytshnjuk, A. Z." box="[1043,1238,1246,1273]" pageId="49" pageNumber="50" pagination="489 - 527" refId="ref89691" refString="Osytshnjuk, A. Z. (1995) Family Andrenidae. In: Lehr, P. A. (Ed.), Keys to Insects of the Far East of Russia. Vol. 4. Megaloptera, Raphidioptera, Neuroptera, Mecoptera, Hymenoptera. Pt 1. Nauka, St. Petersburg, pp. 489 - 527. [in Russian]" type="book chapter" year="1995">Osytshnjuk 1995</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC68434A0FF4FB158EB6035D" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." pageId="49" pageNumber="50" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC68434A0E71FB148BF002B9" box="[1381,1429,1246,1272]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2008
</bibRefCitation>
), specifically Khasan Lake, Golubinyy Utes (42.4099
<superScript id="1EF79BA2AC68434A083DFAC98D2D035D" attach="right" box="[809,840,1282,1308]" fontSize="6" pageId="49" pageNumber="50">oN</superScript>
, 130.7536
<superScript id="1EF79BA2AC68434A08A8FAC98DB3035D" attach="right" box="[956,982,1282,1308]" fontSize="6" pageId="49" pageNumber="50">oE</superScript>
) which is approximately
<quantity id="2E7A9B0FAC68434A0FE5FAC98B52035D" box="[1265,1335,1282,1308]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="50" unit="km" value="20.0">20 km</quantity>
from the border with
<collectingCountry id="9195767AAC68434A0A09FAED8F010301" box="[285,356,1318,1344]" name="China" pageId="49" pageNumber="50">China</collectingCountry>
. Xu &amp; Tadauchi then described
<taxonomicName id="2E824D69AC68434A09D5FAED8D5B0301" authorityName="Xu &amp; Tadauchi" authorityYear="2005" box="[705,830,1318,1344]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="fagopyri">
<emphasis id="DBF6EAF8AC68434A09D5FAED8D5B0301" box="[705,830,1318,1344]" italics="true" pageId="49" pageNumber="50">A. fagopyri</emphasis>
</taxonomicName>
from north-eastern
<collectingCountry id="9195767AAC68434A0F0CFAED8A3B0301" box="[1048,1118,1318,1344]" name="China" pageId="49" pageNumber="50">China</collectingCountry>
, including from the province of
<collectingRegion id="2B46F808AC68434A0BACFA818F2F0325" box="[184,330,1354,1380]" country="China" name="Heilongjiang" pageId="49" pageNumber="50">Heilongjiang</collectingRegion>
which borders the Russian Far East. Careful reading of the descriptions as well as inspection of a
<typeStatus id="36398848AC68434A0BB9FAA58F6803C9" box="[173,269,1390,1416]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
of
<taxonomicName id="2E824D69AC68434A0A23FAA58F8703C9" authorityName="Osytshnjuk" authorityYear="1995" box="[311,482,1390,1416]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A0A23FAA58F8703C9" box="[311,482,1390,1416]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
from the
<emphasis id="DBF6EAF8AC68434A0942FAA58C8F03C9" box="[598,746,1390,1416]" italics="true" pageId="49" pageNumber="50">locus typicus</emphasis>
and non-type material from
<collectingCountry id="9195767AAC68434A0F23FAA58AC103C9" box="[1079,1188,1390,1416]" name="Mongolia" pageId="49" pageNumber="50">Mongolia</collectingCountry>
shows that these two taxa are conspecific due to their shared characters of weakly shagreened terga, T24 with clear complete apical hairbands composed of dense white hairs (
<figureCitation id="71B92A6FAC68434A0995FA7D8D610391" box="[641,772,1462,1488]" captionStart="FIGURE 32" captionStartId="50.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-28@50.[151,1436,181,1618]" captionTargetPageId="50" captionText="FIGURE 32. Andrena (Hoplandrena) romankovae Osytshnjuk, 1995 paratype female.A. Label details; B. Face, frontal view; C. Process of labrum, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209377" httpUri="https://zenodo.org/record/13209377/files/figure.png" pageId="49" pageNumber="50">Figure 32F</figureCitation>
), A3 exceeding the length of A4+5 (prompting comments by
<bibRefCitation id="8D134B1BAC68434A0BAFFA118C4403B5" author="Gusenleitner, F. &amp; Schwarz, M." box="[187,545,1498,1524]" pageId="49" pageNumber="50" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
who suggested a possible affiliation between
<taxonomicName id="2E824D69AC68434A0F33FA118AB503B5" authorityName="Osytshnjuk" authorityYear="1995" box="[1063,1232,1498,1524]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A0F33FA118AB503B5" box="[1063,1232,1498,1524]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
and the European species
<taxonomicName id="2E824D69AC68434A0BFAFA358F9E0059" authority="Perez, 1902" authorityName="Perez" authorityYear="1902" box="[238,507,1534,1560]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="nuptialis">
<emphasis id="DBF6EAF8AC68434A0BFAFA358F0B0059" box="[238,366,1534,1560]" italics="true" pageId="49" pageNumber="50">A. nuptialis</emphasis>
Pérez, 1902
</taxonomicName>
), and black facial hair including the facial foveae (
<figureCitation id="71B92A6FAC68434A0F33FA358ACF0059" box="[1063,1194,1534,1560]" captionStart="FIGURE 32" captionStartId="50.[151,250,1642,1666]" captionTargetBox="[151,1436,181,1618]" captionTargetId="figure-28@50.[151,1436,181,1618]" captionTargetPageId="50" captionText="FIGURE 32. Andrena (Hoplandrena) romankovae Osytshnjuk, 1995 paratype female.A. Label details; B. Face, frontal view; C. Process of labrum, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209377" httpUri="https://zenodo.org/record/13209377/files/figure.png" pageId="49" pageNumber="50">Figure 32B</figureCitation>
). The species is likely to be bivoltine (bivoltine behaviour found in many species of
<taxonomicName id="2E824D69AC68434A082AF9E98DB7007D" box="[830,978,1570,1596]" pageId="49" pageNumber="150" rank="subGenus" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A082AF9E98DB7007D" box="[830,978,1570,1596]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
</taxonomicName>
), with
<taxonomicName id="2E824D69AC68434A0F34F9E98AA2007D" authorityName="Osytshnjuk" authorityYear="1995" box="[1056,1223,1570,1596]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A0F34F9E98AA2007D" box="[1056,1223,1570,1596]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
described from the spring generation and
<taxonomicName id="2E824D69AC68434A0A85F98D8C6A0021" authorityName="Xu &amp; Tadauchi" authorityYear="2005" box="[401,527,1606,1632]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="fagopyri">
<emphasis id="DBF6EAF8AC68434A0A85F98D8C6A0021" box="[401,527,1606,1632]" italics="true" pageId="49" pageNumber="50">A. fagopyri</emphasis>
</taxonomicName>
described from the summer generation.
</paragraph>
<paragraph id="E93D36EAAC68434A0BD3F9A18A1E011D" blockId="49.[151,1437,1210,1884]" pageId="49" pageNumber="50">
Finally, examination of Radoszkowski material, a specimen under the unpublished name
<taxonomicName id="2E824D69AC68434A0FB8F9A18B5300C5" box="[1196,1334,1642,1668]" class="Insecta" family="Andrenidae" genus="Cilissa" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="seuli">
<emphasis id="DBF6EAF8AC68434A0FB8F9A18B5300C5" box="[1196,1334,1642,1668]" italics="true" pageId="49" pageNumber="50">Cilissa seuli</emphasis>
</taxonomicName>
was also found (
<figureCitation id="71B92A6FAC68434A0BF3F9458F3C00E9" box="[231,345,1678,1704]" captionStart="FIGURE 33" captionStartId="51.[151,250,1161,1185]" captionTargetBox="[151,1436,181,1138]" captionTargetId="figure-16@51.[151,1436,181,1138]" captionTargetPageId="51" captionText="FIGURE 33. Cilissa seuli Radoszkowski, nomen nudum female. A. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209379" httpUri="https://zenodo.org/record/13209379/files/figure.png" pageId="49" pageNumber="50">Figure 33</figureCitation>
).
<bibRefCitation id="8D134B1BAC68434A0A79F9458CEA00E8" author="Gusenleitner, F. &amp; Schwarz, M. &amp; Ascher, J. &amp; Scheuchl, E." box="[365,655,1678,1705]" pageId="49" pageNumber="50" pagination="437 - 472" refId="ref89016" refString="Gusenleitner, F., Schwarz, M., Ascher, J. &amp; Scheuchl, E. (2005) Korrekturen und Nachtrage zu Gusenleitner &amp; Schwarz (2002): &quot; Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andrenidae, Andrena). Entomofauna, 26, 437 - 472." type="journal article" year="2005">
Gusenleitner
<emphasis id="DBF6EAF8AC68434A0910F9448C5300E9" box="[516,566,1678,1704]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. (2005)
</bibRefCitation>
commented that the specimen belongs to the subgenus
<taxonomicName id="2E824D69AC68434A0E15F9458BF000E9" box="[1281,1429,1678,1704]" pageId="49" pageNumber="50" rank="subGenus" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC68434A0E15F9458BF000E9" box="[1281,1429,1678,1704]" italics="true" pageId="49" pageNumber="50">Hoplandrena</emphasis>
</taxonomicName>
, and I concur with this assessment; indeed, the specimen is conspecific with
<taxonomicName id="2E824D69AC68434A08F0F9798AE9008D" authorityName="Osytshnjuk" authorityYear="1995" box="[996,1164,1714,1740]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A08F0F9798AE9008D" box="[996,1164,1714,1740]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
. It is labelled as coming from “Koreja” (
<figureCitation id="71B92A6FAC68434A0A53F91D8FA800B1" box="[327,461,1750,1776]" captionStart="FIGURE 33" captionStartId="51.[151,250,1161,1185]" captionTargetBox="[151,1436,181,1138]" captionTargetId="figure-16@51.[151,1436,181,1138]" captionTargetPageId="51" captionText="FIGURE 33. Cilissa seuli Radoszkowski, nomen nudum female. A. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209379" httpUri="https://zenodo.org/record/13209379/files/figure.png" pageId="49" pageNumber="50">Figure 33A</figureCitation>
), and had it been published, it would be the senior synonym. As it was not, the seniority of
<taxonomicName id="2E824D69AC68434A0BA2F9318F3A0155" authorityName="Osytshnjuk" authorityYear="1995" box="[182,351,1786,1812]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A0BA2F9318F3A0155" box="[182,351,1786,1812]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
is maintained. The identity of the published species
<taxonomicName id="2E824D69AC68434A08A6F9318B000155" authority="Radoszkowski, 1891" authorityName="Radoszkowski" authorityYear="1891" box="[946,1381,1786,1812]" class="Insecta" family="Andrenidae" genus="Cilissa" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="DBF6EAF8AC68434A08A6F9318A160155" box="[946,1139,1786,1812]" italics="true" pageId="49" pageNumber="50">Cilissa thoracica</emphasis>
<bibRefCitation id="8D134B1BAC68434A0F6FF9318B000155" author="Radoszkowski, O." box="[1147,1381,1786,1812]" pageId="49" pageNumber="50" pagination="236 - 243" refId="ref90583" refString="Radoszkowski, O. (1891) Revision des armures copulatrices des males des genres Cilissa e Pseudocilissa. Horae Societatis Entomologicae Rossicae, 25, 236 - 243." type="journal article" year="1891">Radoszkowski, 1891</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="2E824D69AC68434A0E90F9318FA50179" authority="(Radoszkowski)" baseAuthorityName="Radoszkowski" class="Insecta" family="Andrenidae" genus="Cilissa" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="DBF6EAF8AC68434A0E90F9318F670179" italics="true" pageId="49" pageNumber="50">A. thoracica</emphasis>
(Radoszkowski)
</taxonomicName>
nec.
<taxonomicName id="2E824D69AC68434A0AEFF8D58D410179" authority="(Kirby, 1802)" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[507,804,1822,1848]" class="Insecta" family="Andrenidae" genus="Cilissa" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="DBF6EAF8AC68434A0AEFF8D58CE10179" box="[507,644,1822,1848]" italics="true" pageId="49" pageNumber="50">A. thoracica</emphasis>
(Kirby, 1802)
</taxonomicName>
=
<taxonomicName id="2E824D69AC68434A0855F8D58ACD0178" authority="Engel, 2005" authorityName="Engel" authorityYear="2005" box="[833,1192,1822,1849]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pseudothoracica">
<emphasis id="DBF6EAF8AC68434A0855F8D58A7E0179" box="[833,1051,1822,1848]" italics="true" pageId="49" pageNumber="50">A. pseudothoracica</emphasis>
<bibRefCitation id="8D134B1BAC68434A0F35F8D58ACD0178" author="Engel, M. S." box="[1057,1192,1822,1849]" pageId="49" pageNumber="50" pagination="179 - 180" refId="ref88585" refString="Engel, M. S. (2005) Three Replacement Names in the Bee Genus Andrena (Hymenoptera: Andrenidae). Journal of the Kansas Entomological Society, 78, 179 - 180. https: // doi. org / 10.2317 / 0022 - 8567 (2005) 78 [179: TRNITB] 2.0. CO; 2" type="journal article" year="2005">Engel, 2005</bibRefCitation>
</taxonomicName>
must be investigated; it was also described from Vladivostok, and may be a junior synonym of
<taxonomicName id="2E824D69AC68434A08D8F8898A11011D" authorityName="Osytshnjuk" authorityYear="1995" box="[972,1140,1858,1884]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="romankovae">
<emphasis id="DBF6EAF8AC68434A08D8F8898A11011D" box="[972,1140,1858,1884]" italics="true" pageId="49" pageNumber="50">A. romankovae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC6B43490B83F9A18A9300E7" ID-DOI="http://doi.org/10.5281/zenodo.13209377" ID-Zenodo-Dep="13209377" httpUri="https://zenodo.org/record/13209377/files/figure.png" pageId="50" pageNumber="51" startId="50.[151,250,1642,1666]" targetBox="[151,1436,181,1618]" targetPageId="50" targetType="figure">
<paragraph id="E93D36EAAC6B43490B83F9A18A9300E7" blockId="50.[151,1436,1642,1703]" pageId="50" pageNumber="51">
<emphasis id="DBF6EAF8AC6B43490B83F9A18F4500C3" bold="true" box="[151,288,1642,1666]" pageId="50" pageNumber="51">FIGURE 32.</emphasis>
<taxonomicName id="2E824D69AC6B43490A32F9A18D3000C3" authority="Osytshnjuk, 1995" authorityName="Osytshnjuk" authorityYear="1995" box="[294,853,1642,1667]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="romankovae" subGenus="Hoplandrena">
<emphasis id="DBF6EAF8AC6B43490A32F9A18F1900C3" box="[294,380,1642,1666]" italics="true" pageId="50" pageNumber="51">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6B43490A9FF9A18C7400C3" box="[395,529,1642,1666]" italics="true" pageId="50" pageNumber="51">Hoplandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6B43490934F9A18CF900C3" box="[544,668,1642,1666]" italics="true" pageId="50" pageNumber="51">romankovae</emphasis>
<bibRefCitation id="8D134B1BAC6B434909B6F9A18D3000C3" author="Osytshnjuk, A. Z." box="[674,853,1642,1666]" pageId="50" pageNumber="51" pagination="489 - 527" refId="ref89691" refString="Osytshnjuk, A. Z. (1995) Family Andrenidae. In: Lehr, P. A. (Ed.), Keys to Insects of the Far East of Russia. Vol. 4. Megaloptera, Raphidioptera, Neuroptera, Mecoptera, Hymenoptera. Pt 1. Nauka, St. Petersburg, pp. 489 - 527. [in Russian]" type="book chapter" year="1995">Osytshnjuk, 1995</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC6B4349084FF9A08DD900C2" bold="true" box="[859,956,1643,1667]" pageId="50" pageNumber="51">paratype</emphasis>
female. A. Label details; B. Face, frontal view; C. Process of labrum, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC6B43490BD3F91F8ACF013E" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="E93D36EAAC6B43490BD3F91F8ACF013E" blockId="50.[151,1437,1748,2027]" pageId="50" pageNumber="51">
<emphasis id="DBF6EAF8AC6B43490BD3F91F8C8800AF" bold="true" box="[199,749,1748,1774]" pageId="50" pageNumber="51">Material examined. KOREAN PENINSULA:</emphasis>
<materialsCitation id="59EA3CB7AC6B43490814F91E8B200178" collectingDate="2005-07-31" collectingDateMax="2005-08-02" collectingDateMin="2005-07-31" collectionCode="PASK" collectorName="J. Halada &amp; M. Kadlecova" country="Mongolia" elevation="1320" location="Mongol Els" pageId="50" pageNumber="51" specimenCount="4" specimenCount-female="4" stateProvince="Ulaanbaatar">
Koreja” [no further information],
<specimenCount id="FF84FD63AC6B43490F9CF91E8ACE00AE" box="[1160,1195,1749,1775]" count="1" pageId="50" pageNumber="51" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FAC6B43490FAFF91E8B6500AE" box="[1211,1280,1749,1775]" pageId="50" pageNumber="51">PASK</collectionCode>
(unpublished name
<taxonomicName id="2E824D69AC6B43490BC9F9328C7F0152" authority="Radoszkowski" authorityName="Radoszkowski" box="[221,538,1785,1811]" class="Insecta" family="Andrenidae" genus="Cilissa" kingdom="Animalia" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="seuli">
<emphasis id="DBF6EAF8AC6B43490BC9F9328F0D0152" box="[221,360,1785,1811]" italics="true" pageId="50" pageNumber="51">Cilissa seuli</emphasis>
Radoszkowski
</taxonomicName>
);
<emphasis id="DBF6EAF8AC6B4349093AF9338CBF0153" bold="true" box="[558,730,1784,1810]" pageId="50" pageNumber="51">
<collectingCountry id="9195767AAC6B4349093AF9338CB10153" box="[558,724,1784,1810]" name="Mongolia" pageId="50" pageNumber="51">MONGOLIA</collectingCountry>
:
</emphasis>
<quantity id="2E7A9B0FAC6B434909F6F9338D4E0152" box="[738,811,1784,1811]" metricMagnitude="4" metricUnit="m" metricValue="7.5" pageId="50" pageNumber="51" unit="km" value="75.0">75 km</quantity>
W
<collectingRegion id="2B46F808AC6B43490842F9328DBA0152" box="[854,991,1785,1811]" country="Mongolia" name="Ulaanbaatar" pageId="50" pageNumber="51">Ulaanbaatar</collectingRegion>
, dunes,
<date id="9D3C102AAC6B43490F29F9338ADC0152" box="[1085,1209,1784,1811]" pageId="50" pageNumber="51" value="2005-08-02">
<collectingDate id="8D78E9C2AC6B43490F29F9338ADC0152" box="[1085,1209,1784,1811]" pageId="50" pageNumber="51" value="2005-08-02">2.viii.2005</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6B43490FD2F9328A8C0152" box="[1222,1257,1785,1811]" count="1" pageId="50" pageNumber="51" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6B43490E3FF9328BFD0152" box="[1323,1432,1785,1811]" pageId="50" pageNumber="51">J. Halada</collectorName>
, OÖLM;
<location id="EC5D6031AC6B43490BE9F8D68FE10176" LSID="urn:lsid:plazi:treatment:612B87FCAC6843490B83FBDB8C5401AA:EC5D6031AC6B43490BE9F8D68FE10176" box="[253,388,1821,1847]" country="Mongolia" name="Mongol Els" pageId="50" pageNumber="51" stateProvince="Ulaanbaatar">Mongol Els</location>
n. res., dunes,
<quantity id="2E7A9B0FAC6B43490929F8D68CFD0176" box="[573,664,1821,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="50" pageNumber="51" unit="m" value="1320.0">
<elevation id="62AFD1D9AC6B43490929F8D68CFD0176" box="[573,664,1821,1847]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="50" pageNumber="51" unit="m" value="1320.0">1320 m</elevation>
</quantity>
,
<date id="9D3C102AAC6B434909BCF8D78D4F0176" box="[680,810,1820,1847]" pageId="50" pageNumber="51" value="2005-07-31">
<collectingDate id="8D78E9C2AC6B434909BCF8D78D4F0176" box="[680,810,1820,1847]" pageId="50" pageNumber="51" value="2005-07-31">31.vii.2005</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6B4349082EF8D68D3B0176" box="[826,862,1821,1847]" count="2" pageId="50" pageNumber="51" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC6B434908B0F8D68A220176" box="[932,1095,1821,1847]" pageId="50" pageNumber="51">M. Kadlecová</collectorName>
&amp;
<collectorName id="4477533CAC6B43490F67F8D68A870176" box="[1139,1250,1821,1847]" pageId="50" pageNumber="51">J. Halada</collectorName>
, OÖLM
</materialsCitation>
/
<materialsCitation id="59EA3CB7AC6B43490E5AF8D58AC3013E" collectingDate="1983-05-15" collectionCode="TJWC" collectorName="T. Romankova" country="Russia" location="Golubinyy Utes" municipality="Khasan Lake" pageId="50" pageNumber="51" specimenCount="1" specimenCount-female="1" stateProvince="Primorskiy" typeStatus="paratype">
<collectionCode id="8F93AE2FAC6B43490E5AF8D58BFD0178" box="[1358,1432,1822,1849]" pageId="50" pageNumber="51">TJWC</collectionCode>
;
<emphasis id="DBF6EAF8AC6B43490B83F88B8F62011B" bold="true" box="[151,263,1856,1882]" pageId="50" pageNumber="51">
<collectingCountry id="9195767AAC6B43490B83F88B8F67011B" box="[151,258,1856,1882]" name="Russia" pageId="50" pageNumber="51">RUSSIA</collectingCountry>
:
</emphasis>
Приморье, оЗ. Хасан Голубиный утес [
<collectingRegion id="2B46F808AC6B434909FFF88A8DC7011A" box="[747,930,1857,1883]" country="Russia" name="Primorskiy" pageId="50" pageNumber="51">Primorsky Krai</collectingRegion>
,
<collectingMunicipality id="0959AC90AC6B434908A4F88A8A2C011A" box="[944,1097,1857,1883]" pageId="50" pageNumber="51">Khasan Lake</collectingMunicipality>
,
<location id="EC5D6031AC6B43490F43F88A8B6A011A" LSID="urn:lsid:plazi:treatment:612B87FCAC6843490B83FBDB8C5401AA:EC5D6031AC6B43490F43F88A8B6A011A" box="[1111,1295,1857,1883]" country="Russia" municipality="Khasan Lake" name="Golubinyy Utes" pageId="50" pageNumber="51" stateProvince="Primorskiy">Golubinyy Utes</location>
, 42.4099
<superScript id="1EF79BA2AC6B43490E6DF88A8BFD011A" attach="right" box="[1401,1432,1857,1883]" fontSize="6" pageId="50" pageNumber="51">oN</superScript>
, 130.7536
<superScript id="1EF79BA2AC6B43490A14F8AE8F78013E" attach="right" box="[256,285,1893,1919]" fontSize="6" pageId="50" pageNumber="51">oE</superScript>
],
<date id="9D3C102AAC6B43490A25F8AF8FC7013E" box="[305,418,1892,1919]" pageId="50" pageNumber="51" value="1983-05-15">
<collectingDate id="8D78E9C2AC6B43490A25F8AF8FC7013E" box="[305,418,1892,1919]" pageId="50" pageNumber="51" value="1983-05-15">15.v.1983</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6B43490AB9F8AE8FB5013E" box="[429,464,1893,1919]" count="1" pageId="50" pageNumber="51" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6B4349091AF8AE8CD0013E" box="[526,693,1893,1919]" pageId="50" pageNumber="51">T. Romankova</collectorName>
, ZISP (
<emphasis id="DBF6EAF8AC6B4349081DF8AE8D11013E" bold="true" box="[777,884,1893,1919]" pageId="50" pageNumber="51">
<typeStatus id="36398848AC6B4349081DF8AE8D11013E" box="[777,884,1893,1919]" pageId="50" pageNumber="51" type="paratype">paratype</typeStatus>
</emphasis>
, examined by photograph)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6B43490BD3F8438C5401AA" pageId="50" pageNumber="51" type="distribution">
<paragraph id="E93D36EAAC6B43490BD3F8438C5401AA" blockId="50.[151,1437,1748,2027]" pageId="50" pageNumber="51">
<emphasis id="DBF6EAF8AC6B43490BD3F8438F0401E3" bold="true" box="[199,353,1928,1954]" pageId="50" pageNumber="51">Distribution.</emphasis>
<collectingCountry id="9195767AAC6B43490A65F8428FD901E2" box="[369,444,1929,1955]" name="Russia" pageId="50" pageNumber="51">Russia</collectingCountry>
(Eastern Siberia, Far East),
<collectingCountry id="9195767AAC6B43490832F8428DFC01E2" box="[806,921,1929,1955]" name="Mongolia" pageId="50" pageNumber="51">Mongolia</collectingCountry>
*, northern
<collectingCountry id="9195767AAC6B43490F33F8428A0F01E2" box="[1063,1130,1929,1955]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
(
<collectingRegion id="2B46F808AC6B43490F96F8428ABC01E2" box="[1154,1241,1929,1955]" country="China" name="Beijing" pageId="50" pageNumber="51">Beijing</collectingRegion>
,
<collectingRegion id="2B46F808AC6B43490FF9F8428B5601E2" box="[1261,1331,1929,1955]" country="China" name="Hebei" pageId="50" pageNumber="51">Hebei</collectingRegion>
,
<collectingRegion id="2B46F808AC6B43490E53F8428BFD01E2" box="[1351,1432,1929,1955]" country="China" name="Shanxi" pageId="50" pageNumber="51">Shanxi</collectingRegion>
,
<collectingRegion id="2B46F808AC6B43490B83F8668F4E0186" box="[151,299,1965,1991]" country="China" name="Heilongjiang" pageId="50" pageNumber="51">Heilongjiang</collectingRegion>
) and probably
<collectingCountry id="9195767AAC6B43490AC0F8668C050186" box="[468,608,1965,1991]" name="North Korea" pageId="50" pageNumber="51">North Korea</collectingCountry>
based on Radoszkowskis specimen (
<bibRefCitation id="8D134B1BAC6B43490F12F8668A930186" author="Xu, H. - L. &amp; Tadauchi, O." box="[1030,1270,1964,1991]" pageId="50" pageNumber="51" pagination="19 - 40" refId="ref92746" refString="Xu, H. - L. &amp; Tadauchi, O. (2005) A revision of the subgenus Hoplandrena of the genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 45, 19 - 40. https: // doi. org / 10.5109 / 2700" type="journal article" year="2005">Xu &amp; Tadauchi 2005</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6B43490E15F8668E9301AA" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." pageId="50" pageNumber="51" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC6B43490E9CF8668EC801AA" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. 2008
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6B43490A16F81A8C4001AA" author="Proshchalykin M. Y. &amp; Astafurova Y. V. &amp; Sidorov D. A." box="[258,549,2001,2027]" pageId="50" pageNumber="51" pagination="263 - 276" refId="ref90418" refString="Proshchalykin M. Y., Astafurova Y. V. &amp; Sidorov D. A. (2017) Family Andrenidae. In: Lelej, A. S., Proshchalykin, M. Y. &amp; Loktionov, V. M. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences, Supplement 6, 263 - 276." type="journal article" year="2017">
Proshchalykin
<emphasis id="DBF6EAF8AC6B43490ABDF81A8FBF01AA" box="[425,474,2001,2027]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,631 @@
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<treatment id="612B87FCAC6D434D0B83FA158AFA05C0" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0" httpUri="http://treatment.plazi.org/id/612B87FCAC6D434D0B83FA158AFA05C0" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
<subSubSection id="A1986561AC6D434F0B83FA158CA103B9" box="[151,708,1502,1529]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="E93D36EAAC6D434F0B83FA158CA103B9" blockId="52.[151,708,1502,1529]" box="[151,708,1502,1529]" pageId="52" pageNumber="53">
<heading id="B2758186AC6D434F0B83FA158CA103B9" bold="true" box="[151,708,1502,1529]" fontSize="11" level="2" pageId="52" pageNumber="53" reason="4">
<taxonomicName id="2E824D69AC6D434F0B83FA158CA103B9" ID-CoL="DS7N" authority="Nurse, 1904" authorityName="Nurse" authorityYear="1904" box="[151,708,1502,1529]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="marmora" subGenus="Melandrena">
<emphasis id="DBF6EAF8AC6D434F0B83FA158CA103B9" bold="true" box="[151,708,1502,1529]" pageId="52" pageNumber="53">
<emphasis id="DBF6EAF8AC6D434F0B83FA158F6403B8" bold="true" box="[151,257,1502,1529]" italics="true" pageId="52" pageNumber="53">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6D434F0A06FA158FC303B8" bold="true" box="[274,422,1502,1529]" italics="true" pageId="52" pageNumber="53">Melandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6D434F0AACFA148C4D03B8" bold="true" box="[440,552,1503,1529]" italics="true" pageId="52" pageNumber="53">marmora</emphasis>
<bibRefCitation id="8D134B1BAC6D434F093BFA148CA103B9" author="Nurse, C. G." box="[559,708,1502,1529]" pageId="52" pageNumber="53" pagination="557 - 585" refId="ref89529" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse, 1904</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6D434F0B83F9E88A430171" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="E93D36EAAC6D434F0B83F9E88F3C001B" blockId="52.[151,1436,1571,1841]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9E88C36007A" ID-CoL="DT6R" authority="Radoszkowski, 1893" authorityName="Radoszkowski" authorityYear="1893" box="[151,595,1571,1595]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="DBF6EAF8AC6D434F0B83F9E88F13007A" box="[151,374,1571,1595]" italics="true" pageId="52" pageNumber="53">Andrena transcaspica</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A6BF9E88C36007A" author="Radoszkowski, O." box="[383,595,1571,1595]" pageId="52" pageNumber="53" pagination="38 - 81" refId="ref90613" refString="Radoszkowski, O. (1893) Fauna Hymenopterologique Transcaspienne. Horae Societatis Entomologicae Rossicae, 27, 38 - 81." type="journal article" year="1893">Radoszkowski, 1893</bibRefCitation>
</taxonomicName>
(nec.
<taxonomicName id="2E824D69AC6D434F0981F9E88A30007A" ID-CoL="DT6R" authority="Radoszkowski, 1886" authorityName="Radoszkowski" authorityYear="1886" box="[661,1109,1571,1595]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="DBF6EAF8AC6D434F0981F9E88D11007A" box="[661,884,1571,1595]" italics="true" pageId="52" pageNumber="53">Andrena transcaspica</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0868F9E88A30007A" author="Radoszkowski, O." box="[892,1109,1571,1595]" pageId="52" pageNumber="53" pagination="3 - 56" refId="ref90561" refString="Radoszkowski, O. (1886) Fauna Hymenopterologique Transcaspienne. Horae Societatis Entomologicae Rossicae, 20, 3 - 56." type="journal article" year="1886">Radoszkowski, 1886</bibRefCitation>
</taxonomicName>
): 56,
<specimenCount id="FF84FD63AC6D434F0F9BF9EF8AC7007A" box="[1167,1186,1572,1595]" count="1" pageId="52" pageNumber="53" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC6D434F0FA4F9E88B24007A" box="[1200,1345,1571,1595]" name="Turkmenistan" pageId="52" pageNumber="53">Turkmenistan</collectingCountry>
, ZMHB, not examined]
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F9AB8FFE00D6" blockId="52.[151,1436,1571,1841]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9AB8C4B0039" ID-CoL="DSRD" authority="Dalla Torre, 1896" authorityName="Dalla Torre" authorityYear="1896" box="[151,558,1632,1656]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="radoszkowski">
<emphasis id="DBF6EAF8AC6D434F0B83F9AB8F1E0039" box="[151,379,1632,1656]" italics="true" pageId="52" pageNumber="53">Andrena radoszkowski</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A6BF9AB8C4B0039" author="Dalla Torre, K. W." box="[383,558,1632,1656]" pageId="52" pageNumber="53" refId="ref88449" refString="Dalla Torre, K. W. (1896) Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. X. Apidae (Anthophila). Engelmann, Lipsiae, VIII + 643 pp." type="book" year="1896">Dalla Torre, 1896</bibRefCitation>
</taxonomicName>
(nec.
<taxonomicName id="2E824D69AC6D434F097DF9AB8A5F0038" ID-CoL="DSRD" authority="Schmiedeknecht, 1883" authorityName="Schmiedeknecht" authorityYear="1883" box="[617,1082,1632,1657]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="radoszkowski">
<emphasis id="DBF6EAF8AC6D434F097DF9AB8D280039" box="[617,845,1632,1656]" italics="true" pageId="52" pageNumber="53">Andrena radoszkowski</emphasis>
Schmiedeknecht, 1883
</taxonomicName>
): 149,
<emphasis id="DBF6EAF8AC6D434F0F6EF9AA8ABE0039" box="[1146,1243,1633,1656]" italics="true" pageId="52" pageNumber="53">nom. nov.</emphasis>
for
<taxonomicName id="2E824D69AC6D434F0E14F9AA8BF90039" authority="Radoszkowski, 1893" box="[1280,1436,1633,1656]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="DBF6EAF8AC6D434F0E14F9AA8BF90039" box="[1280,1436,1633,1656]" italics="true" pageId="52" pageNumber="53">A. transcaspica</emphasis>
</taxonomicName>
<bibRefCitation id="8D134B1BAC6D434F0BD3F9B48FFE00D6" author="Radoszkowski, O." box="[199,411,1663,1687]" pageId="52" pageNumber="53" pagination="38 - 81" refId="ref90613" refString="Radoszkowski, O. (1893) Fauna Hymenopterologique Transcaspienne. Horae Societatis Entomologicae Rossicae, 27, 38 - 81." type="journal article" year="1893">Radoszkowski, 1893</bibRefCitation>
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F9558DE600F7" blockId="52.[151,1436,1571,1841]" box="[151,899,1694,1718]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9558C6F00F7" authority="Nurse, 1904: 562" authorityName="Nurse" authorityPageNumber="562" authorityYear="1904" box="[151,522,1694,1718]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6D434F0B83F9558F3700F7" box="[151,338,1694,1718]" italics="true" pageId="52" pageNumber="53">Andrena marmora</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A4CF9558C6F00F7" author="Nurse, C. G." box="[344,522,1694,1718]" pageId="52" pageNumber="53" pagination="557 - 585" refId="ref89529" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse, 1904: 562</bibRefCitation>
</taxonomicName>
,
<specimenCount id="FF84FD63AC6D434F0900F9558C4100F4" box="[532,548,1694,1717]" count="1" pageId="52" pageNumber="53" type="female"></specimenCount>
<specimenCount id="FF84FD63AC6D434F0930F9558C5F00F4" box="[548,570,1694,1717]" count="1" pageId="52" pageNumber="53" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6D434F0952F9558CC400F7" box="[582,673,1694,1718]" name="Pakistan" pageId="52" pageNumber="53">Pakistan</collectingCountry>
, NHMUK, examined]
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F9778A620094" blockId="52.[151,1436,1571,1841]" box="[151,1031,1724,1749]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9778F9E0095" authority="Cockerell, 1907: 50" authorityName="Cockerell" authorityPageNumber="50" authorityYear="1907" box="[151,507,1724,1749]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="ducis" status="nom. nov.">
<emphasis id="DBF6EAF8AC6D434F0B83F9778F4C0095" box="[151,297,1724,1748]" italics="true" pageId="52" pageNumber="53">Andrena ducis</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A24F9778F9E0095" author="Cockerell, T. D. A." box="[304,507,1724,1749]" pageId="52" pageNumber="53" pagination="49 - 51" refId="ref88197" refString="Cockerell, T. D. A. (1907) Notes on the nomenclature of some Hymenoptera. Entomologist, 40, 49 - 51." type="journal article" year="1907">Cockerell, 1907: 50</bibRefCitation>
</taxonomicName>
,
<emphasis id="DBF6EAF8AC6D434F0911F9768C020095" box="[517,615,1725,1748]" italics="true" pageId="52" pageNumber="53">
<taxonomicNameLabel id="C0C55783AC6D434F0911F9768C020095" box="[517,615,1725,1748]" pageId="52" pageNumber="53">nom. nov.</taxonomicNameLabel>
</emphasis>
for
<taxonomicName id="2E824D69AC6D434F0984F9768A620094" authority="Radoszkowski, 1893" authorityName="Radoszkowski" authorityYear="1893" box="[656,1031,1724,1749]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="DBF6EAF8AC6D434F0984F9768D480095" box="[656,813,1725,1748]" italics="true" pageId="52" pageNumber="53">A. transcaspica</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0820F9778A620094" author="Radoszkowski, O." box="[820,1031,1724,1749]" pageId="52" pageNumber="53" pagination="38 - 81" refId="ref90613" refString="Radoszkowski, O. (1893) Fauna Hymenopterologique Transcaspienne. Horae Societatis Entomologicae Rossicae, 27, 38 - 81." type="journal article" year="1893">Radoszkowski, 1893</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F9108AED00B2" blockId="52.[151,1436,1571,1841]" box="[151,1160,1755,1779]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9108C2D00B2" authority="Cockerell, 1917: 283" authorityName="Cockerell" authorityPageNumber="283" authorityYear="1917" box="[151,584,1755,1779]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="chionospila">
<emphasis id="DBF6EAF8AC6D434F0B83F9108F0F00B2" box="[151,362,1755,1779]" italics="true" pageId="52" pageNumber="53">Andrena chionospila</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A64F9108C2D00B2" author="Cockerell, T. D. A." box="[368,584,1755,1779]" pageId="52" pageNumber="53" pagination="282 - 290" refId="ref88224" refString="Cockerell, T. D. A. (1917) XXV. - Descriptions and records of bees. - LXXIV. Annals and Magazine of Natural History, Series 8, 19, 282 - 290. https: // doi. org / 10.1080 / 00222931709486938" type="journal article" year="1917">Cockerell, 1917: 283</bibRefCitation>
</taxonomicName>
,
<specimenCount id="FF84FD63AC6D434F0946F9178C0700B2" box="[594,610,1756,1779]" count="1" pageId="52" pageNumber="53" type="female"></specimenCount>
<specimenCount id="FF84FD63AC6D434F0976F9178C1D00B2" box="[610,632,1756,1779]" count="1" pageId="52" pageNumber="53" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6D434F0990F9108CBA00B2" box="[644,735,1755,1779]" name="Pakistan" pageId="52" pageNumber="53">Pakistan</collectingCountry>
, USNM, photograph examined]
<emphasis id="DBF6EAF8AC6D434F0F3EF9108AED00B2" bold="true" box="[1066,1160,1755,1779]" pageId="52" pageNumber="53">syn. nov.</emphasis>
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F9318AF30153" blockId="52.[151,1436,1571,1841]" box="[151,1174,1786,1810]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F9318C410153" authority="Alfken, 1931: 828" authorityName="Alfken" authorityPageNumber="828" authorityYear="1931" box="[151,548,1786,1810]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pullipennis">
<emphasis id="DBF6EAF8AC6D434F0B83F9318F000153" box="[151,357,1786,1810]" italics="true" pageId="52" pageNumber="53">Andrena pullipennis</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A7DF9318C410153" author="Alfken, J. D." box="[361,548,1786,1810]" pageId="52" pageNumber="53" pagination="823 - 844" refId="ref87056" refString="Alfken, J. D. (1931) Hymenoptera IV. Apidae. In: Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II.). Mitteilungen aus dem Zoologischen Museum in Berlin, 16, pp. 823 - 844." type="journal article" year="1931">Alfken, 1931: 828</bibRefCitation>
</taxonomicName>
,
<specimenCount id="FF84FD63AC6D434F093BF9318C270150" box="[559,578,1786,1809]" count="1" pageId="52" pageNumber="53" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC6D434F095AF9318CD20153" box="[590,695,1786,1810]" name="Tajikistan" pageId="52" pageNumber="53">Tajikistan</collectingCountry>
, ZMHB, examined]
<emphasis id="DBF6EAF8AC6D434F089CF9318D800153" bold="true" box="[904,997,1786,1810]" pageId="52" pageNumber="53">syn. nov.</emphasis>
(
<figureCitation id="71B92A6FAC6D434F08E7F9318AEB0153" box="[1011,1166,1786,1810]" captionStart="FIGURE 34" captionStartId="53.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-74@53.[151,1436,309,1752]" captionTargetPageId="53" captionText="FIGURE 34. Andrena (Melandrena) pullipennis Alfken, 1931 holotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Head, lateral view; E. Hind tibia, lateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209381" httpUri="https://zenodo.org/record/13209381/files/figure.png" pageId="52" pageNumber="53">Figures 34AF</figureCitation>
)
</paragraph>
<paragraph id="E93D36EAAC6D434F0B83F8D38A430171" blockId="52.[151,1436,1571,1841]" box="[151,1062,1816,1841]" pageId="52" pageNumber="53">
<taxonomicName id="2E824D69AC6D434F0B83F8D38C520171" authority="Lebedev, 1932: 65" authorityName="Lebedev" authorityPageNumber="65" authorityYear="1932" box="[151,567,1816,1841]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="DBF6EAF8AC6D434F0B83F8D38F110171" box="[151,372,1816,1840]" italics="true" pageId="52" pageNumber="53">Andrena gussakovskii</emphasis>
<bibRefCitation id="8D134B1BAC6D434F0A6EF8D38C520171" author="Lebedev, A. G." box="[378,567,1816,1841]" pageId="52" pageNumber="53" pagination="65 - 73" refId="ref89326" refString="Lebedev, A. G. (1932) Einige neue Andrena - Arten aus S. S. S. R. Konowia, 11, 65 - 73." type="journal article" year="1932">Lebedev, 1932: 65</bibRefCitation>
</taxonomicName>
,
<specimenCount id="FF84FD63AC6D434F0956F8D28C300171" box="[578,597,1817,1840]" count="1" pageId="52" pageNumber="53" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC6D434F0975F8D38CBD0171" box="[609,728,1816,1840]" name="Uzbekistan" pageId="52" pageNumber="53">Uzbekistan</collectingCountry>
, ZISP, examined by photograph]
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6D434D0B83F8948D4607EC" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="discussion">
<paragraph id="E93D36EAAC6D434E0B83F8948FBE075C" blockId="52.[151,1437,1887,2021]" lastBlockId="53.[151,1436,151,285]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="DBF6EAF8AC6D434F0B83F8948F6E0138" bold="true" box="[151,267,1887,1913]" pageId="52" pageNumber="53">Remarks.</emphasis>
<taxonomicName id="2E824D69AC6D434F0A04F8948FB80139" authorityName="Nurse" authorityYear="1904" box="[272,477,1887,1913]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6D434F0A04F8948FB80139" box="[272,477,1887,1913]" italics="true" pageId="52" pageNumber="53">Andrena marmora</emphasis>
</taxonomicName>
has been misinterpreted by some authors, but is a valid species found in dry parts of the Middle East and Central Asia (
<bibRefCitation id="8D134B1BAC6D434F0AF0F8488C9201DC" author="Wood, T. J. &amp; Monfared, A." box="[484,759,1923,1949]" pageId="52" pageNumber="53" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared 2022</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6D434F09EBF8488DF501DC" author="Wood, T. J." box="[767,912,1923,1949]" pageId="52" pageNumber="53" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood 2023a</bibRefCitation>
).
<bibRefCitation id="8D134B1BAC6D434F088AF8488AC001DC" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[926,1189,1923,1949]" pageId="52" pageNumber="53" pagination="411 - 447" refId="ref87593" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2024) The type specimens of bees (Hymenoptera: Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution VII. Family Andrenidae, genus Andrena Fabricius, 1775 m taxa described by A. Osytshnjuk and A. Lebedev. Zootaxa, 5453 (3), 411 - 447. https: // doi. org / 10.11646 / zootaxa. 5453.3.9" type="journal article" year="2024">
Astafurova
<emphasis id="DBF6EAF8AC6D434F0F0BF8488A2B01DC" box="[1055,1102,1923,1949]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. (2024)
</bibRefCitation>
recently synonymised
<taxonomicName id="2E824D69AC6D434F0B83F86C8F260180" authorityName="Lebedev" authorityYear="1932" box="[151,323,1959,1985]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gussakovskii">
<emphasis id="DBF6EAF8AC6D434F0B83F86C8F260180" box="[151,323,1959,1985]" italics="true" pageId="52" pageNumber="53">A. gussakovskii</emphasis>
</taxonomicName>
with
<taxonomicName id="2E824D69AC6D434F0A6BF86C8C600181" authorityName="Nurse" authorityYear="1904" box="[383,517,1959,1985]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6D434F0A6BF86C8C600181" box="[383,517,1959,1985]" italics="true" pageId="52" pageNumber="53">A. marmora</emphasis>
</taxonomicName>
following type illustration.
<bibRefCitation id="8D134B1BAC6D434F082DF86C8D900180" author="Cockerell, T. D. A." box="[825,1013,1959,1985]" pageId="52" pageNumber="53" pagination="282 - 290" refId="ref88224" refString="Cockerell, T. D. A. (1917) XXV. - Descriptions and records of bees. - LXXIV. Annals and Magazine of Natural History, Series 8, 19, 282 - 290. https: // doi. org / 10.1080 / 00222931709486938" type="journal article" year="1917">Cockerell (1917)</bibRefCitation>
described
<taxonomicName id="2E824D69AC6D434F0F78F86C8B690180" authorityName="Cockerell" authorityYear="1917" box="[1132,1292,1959,1985]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="chionospila">
<emphasis id="DBF6EAF8AC6D434F0F78F86C8B690180" box="[1132,1292,1959,1985]" italics="true" pageId="52" pageNumber="53">A. chionospila</emphasis>
</taxonomicName>
from what is now north-western
<collectingCountry id="9195767AAC6D434F0A7BF8008FB401A4" box="[367,465,1995,2021]" name="Pakistan" pageId="52" pageNumber="53">Pakistan</collectingCountry>
, comparing it to
<taxonomicName id="2E824D69AC6D434F099DF8008A1C01A4" authority="(Rossi, 1792)" baseAuthorityName="Rossi" baseAuthorityYear="1792" box="[649,1145,1995,2021]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="albopunctata" subGenus="Melandrena">
<emphasis id="DBF6EAF8AC6D434F099DF8008CC701A4" box="[649,674,1995,2021]" italics="true" pageId="52" pageNumber="53">A.</emphasis>
(
<emphasis id="DBF6EAF8AC6D434F09A4F8008D5D01A4" box="[688,824,1995,2021]" italics="true" pageId="52" pageNumber="53">Melandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6D434F0853F8008DBE01A4" box="[839,987,1995,2021]" italics="true" pageId="52" pageNumber="53">albopunctata</emphasis>
(Rossi, 1792)
</taxonomicName>
, but indicating the weaker and sparser metasomal punctation. No comparison is made to
<taxonomicName id="2E824D69AC6C434E0843FF5C8DBA06F1" authorityName="Nurse" authorityYear="1904" box="[855,991,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6C434E0843FF5C8DBA06F1" box="[855,991,151,177]" italics="true" pageId="53" pageNumber="54">A. marmora</emphasis>
</taxonomicName>
which was described from
<collectingCountry id="9195767AAC6C434E0E0DFF5C8B1D06F0" box="[1305,1400,151,177]" name="Pakistan" pageId="53" pageNumber="54">Pakistan</collectingCountry>
by
<bibRefCitation id="8D134B1BAC6C434E0B83FF708F480694" author="Nurse, C. G." box="[151,301,187,213]" pageId="53" pageNumber="54" pagination="557 - 585" refId="ref89529" refString="Nurse, C. G. (1904) New species of Indian Hymenoptera. Apidae. Journal of Bombay Natural History Society, 15, 557 - 585." type="journal article" year="1904">Nurse (1904)</bibRefCitation>
, and the two species are clearly conspecific (type material of
<taxonomicName id="2E824D69AC6C434E08CDFF708A050695" authorityName="Nurse" authorityYear="1904" box="[985,1120,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6C434E08CDFF708A050695" box="[985,1120,187,213]" italics="true" pageId="53" pageNumber="54">A. marmora</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="8D134B1BAC6C434E0FEBFF708BF50694" author="Wood, T. J." box="[1279,1424,187,213]" pageId="53" pageNumber="54" pagination="1 - 72" refId="ref92341" refString="Wood, T. J. (2023 a) New Old World Andrena species, with a focus on Turkey (Hymenoptera: Andrenidae). Zootaxa, 5266 (1), 1 - 72. https: // doi. org / 10.11646 / zootaxa. 5266.1.1" type="journal article" year="2023">Wood 2023a</bibRefCitation>
), and hence are synonymised here. The photographed specimen in the USNM is a
<typeStatus id="36398848AC6C434E0F21FF148AEA06B8" box="[1077,1167,223,249]" pageId="53" pageNumber="54" type="syntype">syntype</typeStatus>
, as Cockerell described both sexes of
<taxonomicName id="2E824D69AC6C434E0A26FEC88FB1075C" authorityName="Cockerell" authorityYear="1917" box="[306,468,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="chionospila">
<emphasis id="DBF6EAF8AC6C434E0A26FEC88FB1075C" box="[306,468,259,285]" italics="true" pageId="53" pageNumber="54">A. chionospila</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BDFD6662AC6C434E0B83F9248A48016A" ID-DOI="http://doi.org/10.5281/zenodo.13209381" ID-Zenodo-Dep="13209381" httpUri="https://zenodo.org/record/13209381/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,250,1775,1799]" targetBox="[151,1436,309,1752]" targetPageId="53" targetType="figure">
<paragraph id="E93D36EAAC6C434E0B83F9248A48016A" blockId="53.[151,1437,1775,1836]" pageId="53" pageNumber="54">
<emphasis id="DBF6EAF8AC6C434E0B83F9248F440146" bold="true" box="[151,289,1775,1799]" pageId="53" pageNumber="54">FIGURE 34.</emphasis>
<taxonomicName id="2E824D69AC6C434E0A3CF9248D730146" authority="Alfken, 1931" authorityName="Alfken" authorityYear="1931" box="[296,790,1775,1799]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pullipennis" subGenus="Melandrena">
<emphasis id="DBF6EAF8AC6C434E0A3CF9248F1B0146" box="[296,382,1775,1799]" italics="true" pageId="53" pageNumber="54">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6C434E0A9AF9248C6C0146" box="[398,521,1775,1799]" italics="true" pageId="53" pageNumber="54">Melandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6C434E090DF9248CEF0146" box="[537,650,1775,1799]" italics="true" pageId="53" pageNumber="54">pullipennis</emphasis>
<bibRefCitation id="8D134B1BAC6C434E099BF9248D730146" author="Alfken, J. D." box="[655,790,1775,1799]" pageId="53" pageNumber="54" pagination="823 - 844" refId="ref87056" refString="Alfken, J. D. (1931) Hymenoptera IV. Apidae. In: Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II.). Mitteilungen aus dem Zoologischen Museum in Berlin, 16, pp. 823 - 844." type="journal article" year="1931">Alfken, 1931</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC6C434E0809F9248D1F0146" bold="true" box="[797,890,1775,1799]" pageId="53" pageNumber="54">holotype</emphasis>
female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Head, lateral view; E. Hind tibia, lateral view; F. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC6C434D0BD3F8918BE10694" blockId="53.[151,1437,1881,2016]" lastBlockId="54.[151,1437,151,897]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
In a similar manner,
<bibRefCitation id="8D134B1BAC6C434E0AB3F8928C200132" author="Alfken, J. D." box="[423,581,1881,1907]" pageId="53" pageNumber="54" pagination="823 - 844" refId="ref87056" refString="Alfken, J. D. (1931) Hymenoptera IV. Apidae. In: Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II.). Mitteilungen aus dem Zoologischen Museum in Berlin, 16, pp. 823 - 844." type="journal article" year="1931">Alfken (1931)</bibRefCitation>
described
<taxonomicName id="2E824D69AC6C434E09AFF8918D300132" authorityName="Alfken" authorityYear="1931" box="[699,853,1881,1908]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="pullipennis">
<emphasis id="DBF6EAF8AC6C434E09AFF8918D300132" box="[699,853,1881,1908]" italics="true" pageId="53" pageNumber="54">A. pullipennis</emphasis>
</taxonomicName>
from a single specimen from the Pamir mountains in
<collectingCountry id="9195767AAC6C434E0B83F8B68F6001D6" box="[151,261,1917,1943]" name="Tajikistan" pageId="53" pageNumber="54">Tajikistan</collectingCountry>
(
<figureCitation id="71B92A6FAC6C434E0A07F8B68FE001D6" box="[275,389,1917,1943]" captionStart="FIGURE 34" captionStartId="53.[151,250,1775,1799]" captionTargetBox="[151,1436,309,1752]" captionTargetId="figure-74@53.[151,1436,309,1752]" captionTargetPageId="53" captionText="FIGURE 34. Andrena (Melandrena) pullipennis Alfken, 1931 holotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Head, lateral view; E. Hind tibia, lateral view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209381" httpUri="https://zenodo.org/record/13209381/files/figure.png" pageId="53" pageNumber="54">Figure 34</figureCitation>
). He indicated that the species can be recognised due to its white hairs of the face, mesosoma, and terminal fringe. He noted that he could not diagnose the systematic position of the species without seeing the male, possibly contributing to the lack of confident subgeneric placement by
<bibRefCitation id="8D134B1BAC6C434E0F30F80E8BFC019E" author="Gusenleitner, F. &amp; Schwarz, M." box="[1060,1433,1989,2015]" pageId="53" pageNumber="54" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
. Inspection of the
<typeStatus id="36398848AC6F434D0A4AFF5C8FEB06F0" box="[350,398,151,177]" pageId="54" pageNumber="55">type</typeStatus>
shows that
<taxonomicName id="2E824D69AC6F434D090CFF5C8CD106F0" authorityName="Alfken" authorityYear="1931" box="[536,692,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pullipennis">
<emphasis id="DBF6EAF8AC6F434D090CFF5C8CD106F0" box="[536,692,151,177]" italics="true" pageId="54" pageNumber="55">A. pullipennis</emphasis>
</taxonomicName>
is a
<taxonomicName id="2E824D69AC6F434D09F9FF5C8D1006F0" authorityName="Perez" authorityYear="1890" box="[749,885,151,177]" pageId="54" pageNumber="150" rank="subGenus" subGenus="Melandrena">
<emphasis id="DBF6EAF8AC6F434D09F9FF5C8D1006F0" box="[749,885,151,177]" italics="true" pageId="54" pageNumber="55">Melandrena</emphasis>
</taxonomicName>
, and that it is conspecific with
<taxonomicName id="2E824D69AC6F434D0FC8FF5C8B0106F1" authorityName="Nurse" authorityYear="1904" box="[1244,1380,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="marmora" status="syn. nov.">
<emphasis id="DBF6EAF8AC6F434D0FC8FF5C8B0106F1" box="[1244,1380,151,177]" italics="true" pageId="54" pageNumber="55">A. marmora</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC6F434D0E79FF5C8EAD0694" bold="true" pageId="54" pageNumber="55">
<taxonomicNameLabel id="C0C55783AC6F434D0E79FF5C8EAD0694" pageId="54" pageNumber="55">syn. nov.</taxonomicNameLabel>
</emphasis>
; although the lateral white hair patches on the terga are weak to absent, it is otherwise structurally identical.
</paragraph>
<paragraph id="E93D36EAAC6F434D0BD3FF148D4607EC" blockId="54.[151,1437,151,897]" pageId="54" pageNumber="55">
Finally, it was not possible to locate the
<typeStatus id="36398848AC6F434D0984FF148CA506B8" box="[656,704,223,249]" pageId="54" pageNumber="55">type</typeStatus>
of
<taxonomicName id="2E824D69AC6F434D09FCFF148AEC06B8" authority="Radoszkowski, 1893" authorityName="Radoszkowski" authorityYear="1893" box="[744,1161,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="transcaspica">
<emphasis id="DBF6EAF8AC6F434D09FCFF148DF206B9" box="[744,919,223,249]" italics="true" pageId="54" pageNumber="55">A. transcaspica</emphasis>
<bibRefCitation id="8D134B1BAC6F434D088BFF148AEC06B8" author="Radoszkowski, O." box="[927,1161,223,249]" pageId="54" pageNumber="55" pagination="38 - 81" refId="ref90613" refString="Radoszkowski, O. (1893) Fauna Hymenopterologique Transcaspienne. Horae Societatis Entomologicae Rossicae, 27, 38 - 81." type="journal article" year="1893">
Radoszkowski,
<quantity id="2E7A9B0FAC6F434D0F45FF148AEC06B8" box="[1105,1161,223,249]" metricMagnitude="1" metricUnit="m" metricValue="4.80822" pageId="54" pageNumber="55" unit="in" value="1893.0">1893</quantity>
</bibRefCitation>
</taxonomicName>
in Berlin, the suggested
<typeStatus id="36398848AC6F434D0B83FEC88EA2075C" box="[151,199,259,285]" pageId="54" pageNumber="55">type</typeStatus>
repository (
<bibRefCitation id="8D134B1BAC6F434D0A44FEC88CC9075C" author="Gusenleitner, F. &amp; Schwarz, M." box="[336,684,259,285]" pageId="54" pageNumber="55" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
).
<bibRefCitation id="8D134B1BAC6F434D09ABFEC88D10075C" author="Warncke, K." box="[703,885,259,285]" pageId="54" pageNumber="55" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke (1967)</bibRefCitation>
did not examine the
<typeStatus id="36398848AC6F434D0F71FEC88AFD075C" box="[1125,1176,259,285]" pageId="54" pageNumber="55">type</typeStatus>
, but mentioned an oral communication from Grünwaldt who said that the two taxa were conspecific. It is not clear if Grünwaldt examined the
<typeStatus id="36398848AC6F434D0BD7FE808E960724" box="[195,243,331,357]" pageId="54" pageNumber="55">type</typeStatus>
material, but the synonymy is plausible based on the description and the examination of material from
<collectingCountry id="9195767AAC6F434D0B83FEA48F5007C8" box="[151,309,367,393]" name="Turkmenistan" pageId="54" pageNumber="55">Turkmenistan</collectingCountry>
; in any case,
<taxonomicName id="2E824D69AC6F434D0ACCFEA48C5307C8" authorityName="Cockerell" authorityYear="1907" box="[472,566,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="ducis">
<emphasis id="DBF6EAF8AC6F434D0ACCFEA48C5307C8" box="[472,566,367,393]" italics="true" pageId="54" pageNumber="55">A. ducis</emphasis>
</taxonomicName>
(the replacement name later proposed by
<bibRefCitation id="8D134B1BAC6F434D0F30FEA48A8A07C8" author="Cockerell, T. D. A." box="[1060,1263,367,393]" pageId="54" pageNumber="55" pagination="49 - 51" refId="ref88197" refString="Cockerell, T. D. A. (1907) Notes on the nomenclature of some Hymenoptera. Entomologist, 40, 49 - 51." type="journal article" year="1907">Cockerell (1907))</bibRefCitation>
is junior to
<taxonomicName id="2E824D69AC6F434D0E90FEA48E9A07ED" authorityName="Nurse" authorityYear="1904" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="marmora">
<emphasis id="DBF6EAF8AC6F434D0E90FEA48E9A07ED" italics="true" pageId="54" pageNumber="55">A. marmora</emphasis>
</taxonomicName>
. Searches must be made in the PASK collection.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6F434D0BD3FE7D8B040578" pageId="54" pageNumber="55" type="materials_examined">
<paragraph id="E93D36EAAC6F434D0BD3FE7D8B040578" blockId="54.[151,1437,151,897]" pageId="54" pageNumber="55">
<emphasis id="DBF6EAF8AC6F434D0BD3FE7D8FD30791" bold="true" box="[199,438,438,464]" pageId="54" pageNumber="55">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC6F434D0AD7FE7D8BFD0790" box="[451,1432,438,465]" collectingDate="1988-03-31" collectionCode="RMNH" collectorName="R. Leys" country="Israel" location="Negev" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC6F434D0AD7FE7D8C510791" bold="true" box="[451,564,438,464]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D0AD7FE7D8C550791" box="[451,560,438,464]" name="Israel" pageId="54" pageNumber="55">ISRAEL</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC6F434D0955FE7C8CEE0790" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0955FE7C8CEE0790" box="[577,651,439,465]" country="Israel" name="Negev" pageId="54" pageNumber="55">Negev</location>
,
<locationDeviation id="F88F2FC8AC6F434D098FFE7D8DA30790" box="[667,966,438,465]" location="Be'er Sheva" pageId="54" pageNumber="55">
<quantity id="2E7A9B0FAC6F434D098FFE7D8C8D0790" box="[667,744,438,465]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="km" value="15.0">15 km</quantity>
S of Beer Sheva
</locationDeviation>
,
<date id="9D3C102AAC6F434D08C3FE7C8A370790" box="[983,1106,439,465]" pageId="54" pageNumber="55" value="1988-03-31">
<collectingDate id="8D78E9C2AC6F434D08C3FE7C8A370790" box="[983,1106,439,465]" pageId="54" pageNumber="55" value="1988-03-31">31.iii.1988</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0F70FE7C8AE20790" box="[1124,1159,439,465]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0FC4FE7C8B4A0790" box="[1232,1327,439,465]" pageId="54" pageNumber="55">R. Leys</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0E54FE7C8BFD0790" box="[1344,1432,439,465]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="54" pageNumber="55" type="Museum">RMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D0B83FE118DA607B4" box="[151,963,474,501]" collectingDate="1978-05-05" collectorName="J. Visa" country="Kyrgyzstan" location="Kadamzoj" pageId="54" pageNumber="55" specimenCount="2" specimenCount-female="2">
<emphasis id="DBF6EAF8AC6F434D0B83FE118F0107B5" bold="true" box="[151,356,474,500]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D0B83FE118F3A07B5" box="[151,351,474,500]" name="Kyrgyzstan" pageId="54" pageNumber="55">KYRGYZSTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC6F434D0A65FE108F8107B4" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0A65FE108F8107B4" box="[369,484,475,501]" country="Kyrgyzstan" name="Kadamzoj" pageId="54" pageNumber="55">Kadamžoj</location>
[Kadamjay],
<date id="9D3C102AAC6F434D0998FE118C8B07B4" box="[652,750,474,501]" pageId="54" pageNumber="55" value="1978-05-05">
<collectingDate id="8D78E9C2AC6F434D0998FE118C8B07B4" box="[652,750,474,501]" pageId="54" pageNumber="55" value="1978-05-05">5.v.1978</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0814FE108D4007B4" box="[768,805,475,501]" count="2" pageId="54" pageNumber="55" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D087AFE108DA607B4" box="[878,963,475,501]" pageId="54" pageNumber="55">J. Víša</collectorName>
</materialsCitation>
, OÖLM;
<materialsCitation id="59EA3CB7AC6F434D0F28FE118F41047C" collectingDate="1906-03" collectionCode="USNM" collectorName="Menserah, N. W &amp; F. Benton" country="Pakistan" location="India" municipality="Provinces" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<emphasis id="DBF6EAF8AC6F434D0F28FE118AB707B5" bold="true" box="[1084,1234,474,500]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D0F28FE118AA807B5" box="[1084,1229,474,500]" name="Pakistan" pageId="54" pageNumber="55">PAKISTAN</collectingCountry>
:
</emphasis>
<collectorName id="4477533CAC6F434D0FCBFE108BFD07B4" box="[1247,1432,475,501]" pageId="54" pageNumber="55">Menserah, N.W</collectorName>
.
<collectingMunicipality id="0959AC90AC6F434D0B83FE348F6D0458" box="[151,264,511,537]" pageId="54" pageNumber="55">Provinces</collectingMunicipality>
,
<location id="EC5D6031AC6F434D0A07FE348F2A0458" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0A07FE348F2A0458" box="[275,335,511,537]" country="Pakistan" municipality="Provinces" name="India" pageId="54" pageNumber="55">India</location>
,
<date id="9D3C102AAC6F434D0A4EFE348F800458" box="[346,485,511,537]" pageId="54" pageNumber="55" value="1906-03">
<collectingDate id="8D78E9C2AC6F434D0A4EFE348F800458" box="[346,485,511,537]" pageId="54" pageNumber="55" value="1906-03">March 1906</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0AFBFE348C770458" box="[495,530,511,537]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D095BFE348CDB0458" box="[591,702,511,537]" pageId="54" pageNumber="55">F. Benton</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D09DDFE348D7F0458" box="[713,794,511,537]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="54" pageNumber="55" type="Museum">USNM</collectionCode>
(
<emphasis id="DBF6EAF8AC6F434D083DFE348DE00458" bold="true" box="[809,901,511,537]" pageId="54" pageNumber="55">
<typeStatus id="36398848AC6F434D083DFE348DE00458" box="[809,901,511,537]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
</emphasis>
of
<taxonomicName id="2E824D69AC6F434D08BEFE348A2E0458" authorityName="Cockerell" authorityYear="1917" box="[938,1099,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="chionospila">
<emphasis id="DBF6EAF8AC6F434D08BEFE348A2E0458" box="[938,1099,511,537]" italics="true" pageId="54" pageNumber="55">A. chionospila</emphasis>
</taxonomicName>
; type no. 23137, examined by photograph)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D0A25FDE88DA2047C" box="[305,967,547,573]" collectingDate="1903-04" collectionCode="NHMUK" collectorName="C. G. Nurse" country="Pakistan" location="Peshin" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
<location id="EC5D6031AC6F434D0A25FDE88F1A047C" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0A25FDE88F1A047C" box="[305,383,547,573]" country="Pakistan" name="Peshin" pageId="54" pageNumber="55">Peshin</location>
,
<date id="9D3C102AAC6F434D0A9FFDE88F84047C" box="[395,481,547,573]" pageId="54" pageNumber="55" value="1903-04">
<collectingDate id="8D78E9C2AC6F434D0A9FFDE88F84047C" box="[395,481,547,573]" pageId="54" pageNumber="55" value="1903-04">iv.1903</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0AFAFDE88C74047C" box="[494,529,547,573]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0940FDE88CBD047C" box="[596,728,547,573]" pageId="54" pageNumber="55">C.G. Nurse</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D09F0FDE88D2B047C" box="[740,846,547,573]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="54" pageNumber="55" type="Museum">NHMUK</collectionCode>
(
<emphasis id="DBF6EAF8AC6F434D0875FDE88DD8047C" bold="true" box="[865,957,547,573]" pageId="54" pageNumber="55">
<typeStatus id="36398848AC6F434D0875FDE88DD8047C" box="[865,957,547,573]" pageId="54" pageNumber="55" type="syntype">syntype</typeStatus>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D08C2FDE98C8E04C4" collectingDate="1928-08-15" collectionCode="ZMHB" collectorName="W. F. Reinig" country="Tajikistan" elevation="3580" location="Maz" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6F434D08C2FDE98AE2047D" bold="true" box="[982,1159,546,572]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D08C2FDE98AE7047D" box="[982,1154,546,572]" name="Tajikistan" pageId="54" pageNumber="55">TAJIKISTAN</collectingCountry>
:
</emphasis>
W-Pamir,
<quantity id="2E7A9B0FAC6F434D0E10FDE98B38047C" box="[1284,1373,546,573]" metricMagnitude="3" metricUnit="m" metricValue="3.5799999999999996" pageId="54" pageNumber="55" unit="m" value="3580.0">
<elevation id="62AFD1D9AC6F434D0E10FDE98B38047C" box="[1284,1373,546,573]" metricMagnitude="3" metricUnit="m" metricValue="3.5799999999999996" pageId="54" pageNumber="55" unit="m" value="3580.0">3580 m</elevation>
</quantity>
,
<location id="EC5D6031AC6F434D0E7EFDE88BF9047C" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0E7EFDE88BF9047C" box="[1386,1436,547,573]" country="Tajikistan" name="Maz" pageId="54" pageNumber="55">Maz</location>
[
<bibRefCitation id="8D134B1BAC6F434D0B89FD8C8F5A0420" author="Alfken, J. D." box="[157,319,583,609]" pageId="54" pageNumber="55" pagination="823 - 844" refId="ref87056" refString="Alfken, J. D. (1931) Hymenoptera IV. Apidae. In: Entomologische Ergebnisse der Deutsch-Russischen Alai-Pamir-Expedition 1928 (II.). Mitteilungen aus dem Zoologischen Museum in Berlin, 16, pp. 823 - 844." type="journal article" year="1931">Alfken (1931)</bibRefCitation>
adds the note “westl. Täler” which is inferred to be Talbar, 38.5947
<superScript id="1EF79BA2AC6F434D0F3EFD8C8A2C0420" attach="right" box="[1066,1097,583,609]" fontSize="6" pageId="54" pageNumber="55">oN</superScript>
, 70.3644
<superScript id="1EF79BA2AC6F434D0FBAFD8C8AAE0420" attach="right" box="[1198,1227,583,609]" fontSize="6" pageId="54" pageNumber="55">oE</superScript>
],
<date id="9D3C102AAC6F434D0FCAFD8D8B0D0420" box="[1246,1384,582,609]" pageId="54" pageNumber="55" value="1928-08-15">
<collectingDate id="8D78E9C2AC6F434D0FCAFD8D8B0D0420" box="[1246,1384,582,609]" pageId="54" pageNumber="55" value="1928-08-15">15.viii.1928</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0E66FD8C8BF00420" box="[1394,1429,583,609]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0BD3FDA08F2B04C4" box="[199,334,619,645]" pageId="54" pageNumber="55">W.F. Reinig</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0A4DFDA08FCF04C4" box="[345,426,619,645]" pageId="54" pageNumber="55">ZMHB</collectionCode>
(
<emphasis id="DBF6EAF8AC6F434D0AADFDA08C4504C4" bold="true" box="[441,544,619,645]" pageId="54" pageNumber="55">
<typeStatus id="36398848AC6F434D0AADFDA08C4504C4" box="[441,544,619,645]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
</emphasis>
of
<taxonomicName id="2E824D69AC6F434D0951FDA08CBA04C4" authorityName="Alfken" authorityYear="1931" box="[581,735,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="pullipennis">
<emphasis id="DBF6EAF8AC6F434D0951FDA08CBA04C4" box="[581,735,619,645]" italics="true" pageId="54" pageNumber="55">A. pullipennis</emphasis>
</taxonomicName>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D09ECFDA18F4904E8" collectingDate="1977-04-26" collectorName="Dr. A. Svozil" country="Turkmenistan" location="Kara-kum" municipality="Annau" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6F434D09ECFDA18D8204C5" bold="true" box="[760,999,618,644]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D09ECFDA18D8704C5" box="[760,994,618,644]" name="Turkmenistan" pageId="54" pageNumber="55">TURKMENISTAN</collectingCountry>
:
</emphasis>
<collectingMunicipality id="0959AC90AC6F434D08F9FDA08A5D04C4" box="[1005,1080,619,645]" pageId="54" pageNumber="55">Annau</collectingMunicipality>
/
<location id="EC5D6031AC6F434D0F55FDA08AD004C4" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0F55FDA08AD004C4" box="[1089,1205,619,645]" country="Turkmenistan" municipality="Annau" name="Kara-kum" pageId="54" pageNumber="55">Kara-kum</location>
,
<date id="9D3C102AAC6F434D0FABFDA08B5204C4" box="[1215,1335,619,645]" pageId="54" pageNumber="55" value="1977-04-26">
<collectingDate id="8D78E9C2AC6F434D0FABFDA08B5204C4" box="[1215,1335,619,645]" pageId="54" pageNumber="55" value="1977-04-26">26.iv.1977</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0E56FDA08B0004C4" box="[1346,1381,619,645]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0B83FD448F4904E8" box="[151,300,655,681]" pageId="54" pageNumber="55">Dr. A. Svozil</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC6F434D0A2CFD5B8D5404E8" box="[312,817,655,683]" collectingDate="1977-04" collectorName="J. Strejcek" country="Turkmenistan" location="Firjuza" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
OÖLM;
<location id="EC5D6031AC6F434D0A8DFD448F8E04E8" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0A8DFD448F8E04E8" box="[409,491,655,681]" country="Turkmenistan" name="Firjuza" pageId="54" pageNumber="55">Firjuza</location>
,
<date id="9D3C102AAC6F434D0AE3FD448C2904E8" box="[503,588,655,681]" pageId="54" pageNumber="55" value="1977-04">
<collectingDate id="8D78E9C2AC6F434D0AE3FD448C2904E8" box="[503,588,655,681]" pageId="54" pageNumber="55" value="1977-04">iv.1977</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D094CFD448C1E04E8" box="[600,635,655,681]" count="1" pageId="54" pageNumber="55" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D09ADFD448D5404E8" box="[697,817,655,681]" pageId="54" pageNumber="55">J. Strejček</collectorName>
</materialsCitation>
, OÖLM;
<materialsCitation id="59EA3CB7AC6F434D088AFD458F94048C" collectingDate="1991-04-30" collectorName="J. Halada" country="Uzbekistan" location="Qarshi" pageId="54" pageNumber="55" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6F434D088AFD458A3804E9" bold="true" box="[926,1117,654,680]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D088AFD458A3D04E9" box="[926,1112,654,680]" name="Uzbekistan" pageId="54" pageNumber="55">UZBEKISTAN</collectingCountry>
:
</emphasis>
<locationDeviation id="F88F2FC8AC6F434D0F70FD448B5404E8" box="[1124,1329,655,681]" location="Karschi" pageId="54" pageNumber="55">
<quantity id="2E7A9B0FAC6F434D0F70FD448ACE04E8" box="[1124,1195,655,681]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="54" pageNumber="55" unit="km" value="80.0">80 km</quantity>
SE Karschi
</locationDeviation>
[
<location id="EC5D6031AC6F434D0E2BFD448BF404E8" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0E2BFD448BF404E8" box="[1343,1425,655,681]" country="Uzbekistan" name="Qarshi" pageId="54" pageNumber="55">Qarshi</location>
],
<date id="9D3C102AAC6F434D0B83FD788F75048C" box="[151,272,691,717]" pageId="54" pageNumber="55" value="1991-04-30">
<collectingDate id="8D78E9C2AC6F434D0B83FD788F75048C" box="[151,272,691,717]" pageId="54" pageNumber="55" value="1991-04-30">30.iv.1991</collectingDate>
</date>
,
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, leg.
<collectorName id="4477533CAC6F434D0A90FD788F94048C" box="[388,497,691,717]" pageId="54" pageNumber="55">J. Halada</collectorName>
</materialsCitation>
,
<materialsCitation id="59EA3CB7AC6F434D0AEBFD7F8F2704B0" collectingDate="2022-04-09" collectionCode="NMPC" collectorName="D. Benda" country="Uzbekistan" elevation="973" location="Surxondaryo" municipality="Daganadzham" pageId="54" pageNumber="55" specimenCount="4" specimenCount-male="4" typeStatus="holotype">
OÖLM;
<collectingMunicipality id="0959AC90AC6F434D0977FD788D66048C" box="[611,771,691,717]" pageId="54" pageNumber="55">Daganadzham</collectingMunicipality>
env., II, near
<location id="EC5D6031AC6F434D08B3FD788A5E048C" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D08B3FD788A5E048C" box="[935,1083,691,717]" country="Uzbekistan" municipality="Daganadzham" name="Surxondaryo" pageId="54" pageNumber="55">Surxondaryo</location>
,
<quantity id="2E7A9B0FAC6F434D0F5CFD788AF1048C" box="[1096,1172,691,717]" metricMagnitude="2" metricUnit="m" metricValue="9.73" pageId="54" pageNumber="55" unit="m" value="973.0">
<elevation id="62AFD1D9AC6F434D0F5CFD788AF1048C" box="[1096,1172,691,717]" metricMagnitude="2" metricUnit="m" metricValue="9.73" pageId="54" pageNumber="55" unit="m" value="973.0">973 m</elevation>
</quantity>
,
<date id="9D3C102AAC6F434D0FB6FD788B69048C" box="[1186,1292,691,717]" pageId="54" pageNumber="55" value="2022-04-09">
<collectingDate id="8D78E9C2AC6F434D0FB6FD788B69048C" box="[1186,1292,691,717]" pageId="54" pageNumber="55" value="2022-04-09">9.iv.2022</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0E0FFD788B25048C" box="[1307,1344,691,717]" count="4" pageId="54" pageNumber="55" type="male">4♂</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0E95FD788E8404B0" pageId="54" pageNumber="55">D. Benda</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0BFBFD1C8F2704B0" box="[239,322,727,753]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="54" pageNumber="55">NMPC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D0A5BFD1C8B7604B0" box="[335,1299,726,753]" collectingDate="2022-04-10" collectionCode="NMPC" collectorName="D. Benda" country="Uzbekistan" elevation="752" location="Kashkadarya" municipality="Kul' - Kishlak" pageId="54" pageNumber="55" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<collectingMunicipality id="0959AC90AC6F434D0A5BFD1C8F8604B0" box="[335,483,727,753]" pageId="54" pageNumber="55">Kul-Kishlak</collectingMunicipality>
env.,
<location id="EC5D6031AC6F434D093EFD1C8CD904B0" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D093EFD1C8CD904B0" box="[554,700,727,753]" country="Uzbekistan" municipality="Kul' - Kishlak" name="Kashkadarya" pageId="54" pageNumber="55">Kashkadarya</location>
reg.,
<quantity id="2E7A9B0FAC6F434D0814FD1D8D2E04B0" box="[768,843,726,753]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="54" pageNumber="55" unit="m" value="752.0">
<elevation id="62AFD1D9AC6F434D0814FD1D8D2E04B0" box="[768,843,726,753]" metricMagnitude="2" metricUnit="m" metricValue="7.52" pageId="54" pageNumber="55" unit="m" value="752.0">752 m</elevation>
</quantity>
,
<date id="9D3C102AAC6F434D084DFD1C8DB404B0" box="[857,977,727,753]" pageId="54" pageNumber="55" value="2022-04-10">
<collectingDate id="8D78E9C2AC6F434D084DFD1C8DB404B0" box="[857,977,727,753]" pageId="54" pageNumber="55" value="2022-04-10">10.iv.2022</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D08CBFD1C8A6004B0" box="[991,1029,727,753]" count="2" pageId="54" pageNumber="55" type="male">2♂</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0F50FD1C8AD604B0" box="[1092,1203,727,753]" pageId="54" pageNumber="55">D. Benda</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0FD4FD1C8B7604B0" box="[1216,1299,727,753]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="54" pageNumber="55">NMPC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D0E34FD1C8DDF0554" collectingDate="2022-04-13" collectionCode="NMPC" collectorName="D. Benda" country="Uzbekistan" elevation="722" location="Surxondaryo" municipality="Tangimush" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingMunicipality id="0959AC90AC6F434D0E34FD1C8BF904B0" box="[1312,1436,727,753]" pageId="54" pageNumber="55">Tangimush</collectingMunicipality>
env.,
<location id="EC5D6031AC6F434D0BC1FD308F030554" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0BC1FD308F030554" box="[213,358,763,789]" country="Uzbekistan" municipality="Tangimush" name="Surxondaryo" pageId="54" pageNumber="55">Surxondaryo</location>
reg.,
<quantity id="2E7A9B0FAC6F434D0ABDFD308F960554" box="[425,499,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.22" pageId="54" pageNumber="55" unit="m" value="722.0">
<elevation id="62AFD1D9AC6F434D0ABDFD308F960554" box="[425,499,763,789]" metricMagnitude="2" metricUnit="m" metricValue="7.22" pageId="54" pageNumber="55" unit="m" value="722.0">722 m</elevation>
</quantity>
,
<date id="9D3C102AAC6F434D0915FD308C1C0554" box="[513,633,763,789]" pageId="54" pageNumber="55" value="2022-04-13">
<collectingDate id="8D78E9C2AC6F434D0915FD308C1C0554" box="[513,633,763,789]" pageId="54" pageNumber="55" value="2022-04-13">13.iv.2022</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0993FD308CCE0554" box="[647,683,763,789]" count="1" pageId="54" pageNumber="55" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D09FFFD308D3C0554" box="[747,857,763,789]" pageId="54" pageNumber="55">D. Benda</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0873FD308DDF0554" box="[871,954,763,789]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="54" pageNumber="55">NMPC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D08D3FD308C220578" collectingDate="2022-04-13" collectionCode="NMPC" collectorName="D. Benda" country="Uzbekistan" elevation="1178" location="Surxondaryo" municipality="Charbaq" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingMunicipality id="0959AC90AC6F434D08D3FD308A430554" box="[967,1062,763,789]" pageId="54" pageNumber="55">Charbaq</collectingMunicipality>
env.,
<location id="EC5D6031AC6F434D0F78FD308A980554" LSID="urn:lsid:plazi:treatment:612B87FCAC6D434D0B83FA158AFA05C0:EC5D6031AC6F434D0F78FD308A980554" box="[1132,1277,763,789]" country="Uzbekistan" municipality="Charbaq" name="Surxondaryo" pageId="54" pageNumber="55">Surxondaryo</location>
reg.,
<quantity id="2E7A9B0FAC6F434D0E54FD308BFD0554" box="[1344,1432,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="54" pageNumber="55" unit="m" value="1178.0">
<elevation id="62AFD1D9AC6F434D0E54FD308BFD0554" box="[1344,1432,763,789]" metricMagnitude="3" metricUnit="m" metricValue="1.178" pageId="54" pageNumber="55" unit="m" value="1178.0">1178 m</elevation>
</quantity>
,
<date id="9D3C102AAC6F434D0B83FCD48F750578" box="[151,272,799,825]" pageId="54" pageNumber="55" value="2022-04-13">
<collectingDate id="8D78E9C2AC6F434D0B83FCD48F750578" box="[151,272,799,825]" pageId="54" pageNumber="55" value="2022-04-13">13.iv.2022</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0A0FFCD48F250578" box="[283,320,799,825]" count="1" pageId="54" pageNumber="55" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0A68FCD48F8C0578" box="[380,489,799,825]" pageId="54" pageNumber="55">D. Benda</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0AE0FCD48C220578" box="[500,583,799,825]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="54" pageNumber="55">NMPC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6F434D0946FCD48B380578" box="[594,1373,799,825]" collectingDate="1982-04-16" collectionCode="NHMUK" collectorName="I. L. Hamer" country="Yemen" location="Sanaa" pageId="54" pageNumber="55" specimenCount="1" specimenCount-male="1" stateProvince="San'a'" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6F434D0946FCD48CA10578" bold="true" box="[594,708,799,825]" pageId="54" pageNumber="55">
<collectingCountry id="9195767AAC6F434D0946FCD48CDA0578" box="[594,703,799,825]" name="Yemen" pageId="54" pageNumber="55">YEMEN</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808AC6F434D09DFFCD48D6B0578" box="[715,782,799,825]" country="Yemen" name="San'a'" pageId="54" pageNumber="55">Sanaa</collectingRegion>
[
<collectingRegion id="2B46F808AC6F434D0808FCD48D0A0578" box="[796,879,799,825]" country="Yemen" name="San'a'" pageId="54" pageNumber="55">Sanaa</collectingRegion>
],
<date id="9D3C102AAC6F434D0895FCD48D9F0578" box="[897,1018,799,825]" pageId="54" pageNumber="55" value="1982-04-16">
<collectingDate id="8D78E9C2AC6F434D0895FCD48D9F0578" box="[897,1018,799,825]" pageId="54" pageNumber="55" value="1982-04-16">16.iv.1982</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6F434D0F11FCD48A4F0578" box="[1029,1066,799,825]" count="1" pageId="54" pageNumber="55" type="male">1♂</specimenCount>
, leg.
<collectorName id="4477533CAC6F434D0F73FCD48A810578" box="[1127,1252,799,825]" pageId="54" pageNumber="55">I.L. Hamer</collectorName>
,
<collectionCode id="8F93AE2FAC6F434D0FFBFCD48B380578" box="[1263,1373,799,825]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="54" pageNumber="55" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6F434D0BD3FC898AFA05C0" pageId="54" pageNumber="55" type="distribution">
<paragraph id="E93D36EAAC6F434D0BD3FC898AFA05C0" blockId="54.[151,1437,151,897]" pageId="54" pageNumber="55">
<emphasis id="DBF6EAF8AC6F434D0BD3FC898F04051D" bold="true" box="[199,353,834,860]" pageId="54" pageNumber="55">Distribution.</emphasis>
<collectingCountry id="9195767AAC6F434D0A7CFC888FCC051C" box="[360,425,835,861]" name="Israel" pageId="54" pageNumber="55">Israel</collectingCountry>
,
<collectingCountry id="9195767AAC6F434D0AA1FC888C6C051C" box="[437,521,835,861]" name="Yemen" pageId="54" pageNumber="55">Yemen</collectingCountry>
*,
<collectingCountry id="9195767AAC6F434D0934FC888C0E051C" box="[544,619,835,861]" name="Russia" pageId="54" pageNumber="55">Russia</collectingCountry>
(Western Siberia),
<collectingCountry id="9195767AAC6F434D0852FC888D81051C" box="[838,996,835,861]" name="Turkmenistan" pageId="54" pageNumber="55">Turkmenistan</collectingCountry>
,
<collectingCountry id="9195767AAC6F434D08FBFC888A15051C" box="[1007,1136,835,861]" name="Uzbekistan" pageId="54" pageNumber="55">Uzbekistan</collectingCountry>
,
<collectingCountry id="9195767AAC6F434D0F68FC888A95051C" box="[1148,1264,835,861]" name="Tajikistan" pageId="54" pageNumber="55">Tajikistan</collectingCountry>
*,
<collectingCountry id="9195767AAC6F434D0E13FC888BE8051C" box="[1287,1421,835,861]" name="Kyrgyzstan" pageId="54" pageNumber="55">Kyrgyzstan</collectingCountry>
*,
<collectingCountry id="9195767AAC6F434D0B83FCAC8EA205C0" box="[151,199,871,897]" name="Iran" pageId="54" pageNumber="55">Iran</collectingCountry>
,
<collectingCountry id="9195767AAC6F434D0BC6FCAC8F5405C0" box="[210,305,871,897]" name="Pakistan" pageId="54" pageNumber="55">Pakistan</collectingCountry>
(
<bibRefCitation id="8D134B1BAC6F434D0A2BFCAC8C2705C0" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[319,578,871,897]" pageId="54" pageNumber="55" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC6F434D0ADCFCAC8F9C05C0" box="[456,505,871,897]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2008
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6F434D095AFCAC8D0A05C0" author="Proshchalykin M. Y. &amp; Astafurova Y. V. &amp; Sidorov D. A." box="[590,879,871,897]" pageId="54" pageNumber="55" pagination="263 - 276" refId="ref90418" refString="Proshchalykin M. Y., Astafurova Y. V. &amp; Sidorov D. A. (2017) Family Andrenidae. In: Lelej, A. S., Proshchalykin, M. Y. &amp; Loktionov, V. M. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences, Supplement 6, 263 - 276." type="journal article" year="2017">
Proshchalykin
<emphasis id="DBF6EAF8AC6F434D09E1FCAC8D4305C0" box="[757,806,871,897]" italics="true" pageId="54" pageNumber="55">et al</emphasis>
. 2017
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6F434D086EFCAC8AF605C0" author="Wood, T. J. &amp; Monfared, A." box="[890,1171,871,897]" pageId="54" pageNumber="55" pagination="1 - 136" refId="ref92183" refString="Wood, T. J. &amp; Monfared, A. (2022) A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species. European Journal of Taxonomy, 843, 1 - 136. https: // doi. org / 10.5852 / ejt. 2022.843.1947" type="journal article" year="2022">Wood &amp; Monfared 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,434 @@
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<heading id="B2758186AC6E434C0B83FAD28C880375" bold="true" box="[151,749,1305,1332]" fontSize="11" level="2" pageId="55" pageNumber="56" reason="4">
<taxonomicName id="2E824D69AC6E434C0B83FAD28C880375" ID-CoL="DQXM" authority="Perkins, 1914" authorityName="Perkins" authorityYear="1914" box="[151,749,1305,1332]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6E434C0B83FAD28C880375" bold="true" box="[151,749,1305,1332]" pageId="55" pageNumber="56">
<emphasis id="DBF6EAF8AC6E434C0B83FAD28F640375" bold="true" box="[151,257,1305,1332]" italics="true" pageId="55" pageNumber="56">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6E434C0A06FAD28FD70375" bold="true" box="[274,434,1305,1332]" italics="true" pageId="55" pageNumber="56">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6E434C0AD7FAD28C590375" bold="true" box="[451,572,1305,1332]" italics="true" pageId="55" pageNumber="56">alfkenella</emphasis>
<bibRefCitation id="8D134B1BAC6E434C0957FAD18C880375" author="Perkins, R. C. L." box="[579,749,1306,1332]" pageId="55" pageNumber="56" pagination="71 - 75" refId="ref89893" refString="Perkins, R. C. L. (1914) Synopsis of the British Forms of the Andrena minutula Group. Entomologist's Monthly Magazine, 25, 71 - 75 + 112 - 115." type="journal article" year="1914">Perkins, 1914</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6E434C0B83FA958AEB03D4" pageId="55" pageNumber="56" type="reference_group">
<paragraph id="E93D36EAAC6E434C0B83FA958DD80337" blockId="55.[151,1166,1374,1429]" box="[151,957,1374,1398]" pageId="55" pageNumber="56">
<treatmentCitationGroup id="C99211C4AC6E434C0B83FA958DD80337" box="[151,957,1374,1398]" pageId="55" pageNumber="56">
<treatmentCitation id="682310FBAC6E434C0B83FA958C7B0337" author="Perkins, R. C. L." box="[151,542,1374,1398]" page="112" pageId="55" pageNumber="56" year="1914">
<taxonomicName id="2E824D69AC6E434C0B83FA958C7B0337" ID-CoL="DQXM" authority="Perkins, 1914: 112" authorityName="Perkins" authorityPageNumber="112" authorityYear="1914" box="[151,542,1374,1398]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C0B83FA958F3D0337" box="[151,344,1374,1398]" italics="true" pageId="55" pageNumber="56">Andrena alfkenella</emphasis>
<materialsCitation id="59EA3CB7AC6E434C0A4AFA958C7B0337" box="[350,542,1374,1398]" collectionCode="OUMNH" country="United Kingdom" location="England" pageId="55" pageNumber="56" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<bibRefCitation id="8D134B1BAC6E434C0A4AFA958C7B0337" author="Perkins, R. C. L." box="[350,542,1374,1398]" pageId="55" pageNumber="56" pagination="71 - 75" refId="ref89893" refString="Perkins, R. C. L. (1914) Synopsis of the British Forms of the Andrena minutula Group. Entomologist's Monthly Magazine, 25, 71 - 75 + 112 - 115." type="journal article" year="1914">Perkins, 1914: 112</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC6E434C093CFA948C5D0337" box="[552,568,1375,1398]" count="1" pageId="55" pageNumber="56" type="female"></specimenCount>
<specimenCount id="FF84FD63AC6E434C092CFA948C2B0337" box="[568,590,1375,1398]" count="1" pageId="55" pageNumber="56" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC6E434C094FFA958CD10337" box="[603,692,1374,1398]" name="United Kingdom" pageId="55" pageNumber="56">England</collectingCountry>
, OUMNH, not examined]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC6E434C0B83FAB68AEB03D4" blockId="55.[151,1166,1374,1429]" box="[151,1166,1405,1429]" pageId="55" pageNumber="56">
<treatmentCitationGroup id="C99211C4AC6E434C0B83FAB68AEB03D4" box="[151,1166,1405,1429]" pageId="55" pageNumber="56">
<treatmentCitation id="682310FBAC6E434C0B83FAB68C7603D4" author="Strand, E." box="[151,531,1405,1429]" page="153" pageId="55" pageNumber="56" year="1915">
<taxonomicName id="2E824D69AC6E434C0B83FAB68C7603D4" ID-CoL="DR8W" authority="Strand, 1915 a: 153" authorityName="Strand" authorityPageNumber="153" authorityYear="1915" box="[151,531,1405,1429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6E434C0B83FAB68F2E03D4" box="[151,331,1405,1429]" italics="true" pageId="55" pageNumber="56">Andrena caneibia</emphasis>
<materialsCitation id="59EA3CB7AC6E434C0A45FAB68C7603D4" box="[337,531,1405,1429]" collectionCode="DEI" country="Greece" location="Crete" pageId="55" pageNumber="56" specimenCount="1" specimenCount-female="1" stateProvince="Kriti">
<bibRefCitation id="8D134B1BAC6E434C0A45FAB68C7603D4" author="Strand, E." box="[337,531,1405,1429]" pageId="55" pageNumber="56" pagination="145 - 168" refId="ref91314" refString="Strand, E. (1915 a) Apidae von Creta. Archiv fur Naturgeschichte, 81 A, 145 - 168." type="journal article" year="1915">Strand, 1915a: 153</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
,
<specimenCount id="FF84FD63AC6E434C090AFAB68C5403D5" box="[542,561,1405,1428]" count="1" pageId="55" pageNumber="56" type="female"></specimenCount>
[
<collectingCountry id="9195767AAC6E434C0929FAB68CEE03D4" box="[573,651,1405,1429]" name="Greece" pageId="55" pageNumber="56">Greece</collectingCountry>
:
<collectingRegion id="2B46F808AC6E434C0981FAB68CAB03D4" box="[661,718,1405,1429]" country="Greece" name="Kriti" pageId="55" pageNumber="56">Crete</collectingRegion>
, DEI, examined]
<emphasis id="DBF6EAF8AC6E434C0895FAB68DBB03D4" bold="true" box="[897,990,1405,1429]" pageId="55" pageNumber="56">syn. nov.</emphasis>
(
<figureCitation id="71B92A6FAC6E434C08FFFAB68AE203D4" box="[1003,1159,1405,1429]" captionStart="FIGURE 36" captionStartId="56.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@56.[151,1436,181,1623]" captionTargetPageId="56" captionText="FIGURE 36. Andrena (Micrandrena) caneibia Strand, 1915 holotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209385" httpUri="https://zenodo.org/record/13209385/files/figure.png" pageId="55" pageNumber="56">Figures 36AF</figureCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6E43420B83FA088D6F075C" lastPageId="57" lastPageNumber="58" pageId="55" pageNumber="56" type="discussion">
<paragraph id="E93D36EAAC6E434C0B83FA088B7401CC" blockId="55.[151,1437,1475,1934]" pageId="55" pageNumber="56">
<emphasis id="DBF6EAF8AC6E434C0B83FA088F6E039C" bold="true" box="[151,267,1475,1501]" pageId="55" pageNumber="56">Remarks.</emphasis>
The identity of
<taxonomicName id="2E824D69AC6E434C0AD6FA0F8C27039C" authorityName="Strand" authorityYear="1915" box="[450,578,1475,1502]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6E434C0AD6FA0F8C27039C" box="[450,578,1475,1502]" italics="true" pageId="55" pageNumber="56">A. caneibia</emphasis>
</taxonomicName>
has long remained a mystery. Described from a single specimen from
<collectingRegion id="2B46F808AC6E434C0E74FA088BF8039C" box="[1376,1437,1475,1501]" country="Greece" name="Kriti" pageId="55" pageNumber="56">Crete</collectingRegion>
(
<figureCitation id="71B92A6FAC6E434C0B8BFA2C8F400040" box="[159,293,1511,1537]" captionStart="FIGURE 36" captionStartId="56.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@56.[151,1436,181,1623]" captionTargetPageId="56" captionText="FIGURE 36. Andrena (Micrandrena) caneibia Strand, 1915 holotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209385" httpUri="https://zenodo.org/record/13209385/files/figure.png" pageId="55" pageNumber="56">Figure 36A</figureCitation>
),
<bibRefCitation id="8D134B1BAC6E434C0A2EFA2C8F8C0043" author="Warncke, K." box="[314,489,1511,1538]" pageId="55" pageNumber="56" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke (1965</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6E434C0AE2FA2C8C4A0040" author="Warncke, K." box="[502,559,1511,1537]" pageId="55" pageNumber="56" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">1967</bibRefCitation>
) did not treat the species further other than listing it as valid.
<bibRefCitation id="8D134B1BAC6E434C0FFBFA2C8F2F0064" author="Gusenleitner, F. &amp; Schwarz, M." pageId="55" pageNumber="56" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
uncertainly placed the species in the subgenus
<taxonomicName id="2E824D69AC6E434C0877F9C08AD50064" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[867,1200,1547,1573]" pageId="55" pageNumber="56" rank="subGenus" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6E434C0877F9C08D930064" box="[867,1014,1547,1573]" italics="true" pageId="55" pageNumber="56">Micrandrena</emphasis>
Ashmead, 1899
</taxonomicName>
, commenting that its taxonomic status was unclear. Examination of the
<typeStatus id="36398848AC6E434C09C3F9E48D590008" box="[727,828,1583,1609]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
, along with a long series of new material from
<collectingRegion id="2B46F808AC6E434C0E74F9E48BF80008" box="[1376,1437,1583,1609]" country="Greece" name="Kriti" pageId="55" pageNumber="56">Crete</collectingRegion>
and new genetic analysis has allowed a confident link with the widespread West Palaearctic species
<taxonomicName id="2E824D69AC6E434C0FE7F99F8B1A002C" authorityName="Perkins" authorityYear="1914" box="[1267,1407,1619,1646]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C0FE7F99F8B1A002C" box="[1267,1407,1619,1646]" italics="true" pageId="55" pageNumber="56">A. alfkenella</emphasis>
</taxonomicName>
to be made. The
<typeStatus id="36398848AC6E434C0A21F9BC8FF200D0" box="[309,407,1655,1681]" pageId="55" pageNumber="56" type="holotype">holotype</typeStatus>
female closely resembles
<taxonomicName id="2E824D69AC6E434C09A9F9B38D2F00D0" authorityName="Perkins" authorityYear="1914" box="[701,842,1655,1682]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C09A9F9B38D2F00D0" box="[701,842,1655,1682]" italics="true" pageId="55" pageNumber="56">A. alfkenella</emphasis>
</taxonomicName>
, but the punctation of the tergal discs (one of the key characters allowing separation of
<taxonomicName id="2E824D69AC6E434C0906F9578CFA00F4" authorityName="Perkins" authorityYear="1914" box="[530,671,1691,1718]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C0906F9578CFA00F4" box="[530,671,1691,1718]" italics="true" pageId="55" pageNumber="56">A. alfkenella</emphasis>
</taxonomicName>
females from those of
<taxonomicName id="2E824D69AC6E434C08B7F9578AAA00F4" authority="(Kirby, 1802))" authorityName="" baseAuthorityName="Kirby" baseAuthorityYear="1802" box="[931,1231,1691,1718]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="minutula">
<emphasis id="DBF6EAF8AC6E434C08B7F9578A4200F4" box="[931,1063,1691,1718]" italics="true" pageId="55" pageNumber="56">A. minutula</emphasis>
(Kirby, 1802))
</taxonomicName>
is slightly weaker than in typical
<taxonomicName id="2E824D69AC6E434C0A55F90B8FAB0098" authorityName="Perkins" authorityYear="1914" box="[321,462,1727,1754]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C0A55F90B8FAB0098" box="[321,462,1727,1754]" italics="true" pageId="55" pageNumber="56">A. alfkenella</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC6E434C0ACAF9748C050098" box="[478,608,1727,1753]" captionStart="FIGURE 36" captionStartId="56.[151,250,1646,1670]" captionTargetBox="[151,1436,181,1623]" captionTargetId="figure-28@56.[151,1436,181,1623]" captionTargetPageId="56" captionText="FIGURE 36. Andrena (Micrandrena) caneibia Strand, 1915 holotype female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209385" httpUri="https://zenodo.org/record/13209385/files/figure.png" pageId="55" pageNumber="56">Figure 36F</figureCitation>
). Strands description is unhelpful, as he only compares
<taxonomicName id="2E824D69AC6E434C0FCBF90B8B3B0098" authorityName="Strand" authorityYear="1915" box="[1247,1374,1727,1754]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6E434C0FCBF90B8B3B0098" box="[1247,1374,1727,1754]" italics="true" pageId="55" pageNumber="56">A. caneibia</emphasis>
</taxonomicName>
to
<taxonomicName id="2E824D69AC6E434C0E90F90B8FE700BF" authority="Strand, 1915" authorityName="Strand" authorityYear="1915" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="nanana">
<emphasis id="DBF6EAF8AC6E434C0E90F90B8E8E00BC" italics="true" pageId="55" pageNumber="56">A. nanana</emphasis>
Strand, 1915
</taxonomicName>
(described from
<collectingRegion id="2B46F808AC6E434C0950F9288CE600BC" box="[580,643,1763,1789]" country="Greece" name="Kriti" pageId="55" pageNumber="56">Crete</collectingRegion>
, but a synonym of
<taxonomicName id="2E824D69AC6E434C084FF92F8AA300BC" authority="Morawitz, 1873" authorityName="Morawitz" authorityYear="1873" box="[859,1222,1763,1790]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="colletiformis">
<emphasis id="DBF6EAF8AC6E434C084FF92F8A6D00BC" box="[859,1032,1763,1790]" italics="true" pageId="55" pageNumber="56">A. colletiformis</emphasis>
Morawitz, 1873
</taxonomicName>
), this latter species falling in a completely different subgenus (
<taxonomicName id="2E824D69AC6E434C099CF8CC8DB80160" authority="Pittioni, 1948" authorityName="Pittioni" authorityYear="1948" box="[648,989,1799,1825]" class="Insecta" family="Andrenidae" genus="Brachyandrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC6E434C099CF8CC8D500160" box="[648,821,1799,1825]" italics="true" pageId="55" pageNumber="56">Brachyandrena</emphasis>
Pittioni, 1948
</taxonomicName>
), and consequently there are numerous morphological differences. Given the weak difference from
<taxonomicName id="2E824D69AC6E434C0858F8E78DB90104" authorityName="Perkins" authorityYear="1914" box="[844,988,1835,1862]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6E434C0858F8E78DB90104" box="[844,988,1835,1862]" italics="true" pageId="55" pageNumber="56">A. alfkenella</emphasis>
</taxonomicName>
and the shortage of material, previous authors have not drawn strong conclusions as to the status of
<taxonomicName id="2E824D69AC6E434C084AF89B8DBA0128" authorityName="Strand" authorityYear="1915" box="[862,991,1871,1898]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6E434C084AF89B8DBA0128" box="[862,991,1871,1898]" italics="true" pageId="55" pageNumber="56">A. caneibia</emphasis>
</taxonomicName>
, as opposed to many of Strands other
<taxonomicName id="2E824D69AC6E434C0B83F8B88E9301CC" baseAuthorityName="Popov" baseAuthorityYear="1960" box="[151,246,1907,1933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC6E434C0B83F8B88E9301CC" box="[151,246,1907,1933]" italics="true" pageId="55" pageNumber="56">Andrena</emphasis>
</taxonomicName>
taxa described from
<collectingRegion id="2B46F808AC6E434C0AF7F8B88C4501CC" box="[483,544,1907,1933]" country="Greece" name="Kriti" pageId="55" pageNumber="56">Crete</collectingRegion>
which have been synonymised with other species (
<bibRefCitation id="8D134B1BAC6E434C0F49F8B88B6001CC" author="Warncke, K." box="[1117,1285,1907,1933]" pageId="55" pageNumber="56" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke 1967</bibRefCitation>
).
</paragraph>
<caption id="BDFD6662AC6143430B83F9A58A3400EB" ID-DOI="http://doi.org/10.5281/zenodo.13209385" ID-Zenodo-Dep="13209385" httpUri="https://zenodo.org/record/13209385/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1646,1670]" targetBox="[151,1436,181,1623]" targetPageId="56" targetType="figure">
<paragraph id="E93D36EAAC6143430B83F9A58A3400EB" blockId="56.[151,1437,1646,1707]" pageId="56" pageNumber="57">
<emphasis id="DBF6EAF8AC6143430B83F9A58F4700C7" bold="true" box="[151,290,1646,1670]" pageId="56" pageNumber="57">FIGURE 36.</emphasis>
<taxonomicName id="2E824D69AC6143430A3EF9A58D6E00C7" authority="Strand, 1915" authorityName="Strand" authorityYear="1915" box="[298,779,1646,1671]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="caneibia" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC6143430A3EF9A58FE500C7" box="[298,384,1646,1670]" italics="true" pageId="56" pageNumber="57">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC6143430A85F9A58C7300C7" box="[401,534,1646,1670]" italics="true" pageId="56" pageNumber="57">Micrandrena</emphasis>
)
<emphasis id="DBF6EAF8AC6143430933F9A58C1B00C7" box="[551,638,1646,1670]" italics="true" pageId="56" pageNumber="57">caneibia</emphasis>
Strand, 1915
</taxonomicName>
<emphasis id="DBF6EAF8AC6143430807F9A48D1500C6" bold="true" box="[787,880,1647,1671]" pageId="56" pageNumber="57">holotype</emphasis>
female. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC6143420BD3F9128D6F075C" blockId="56.[151,1437,1753,2031]" lastBlockId="57.[151,1436,151,285]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
New collections have produced long series of
<taxonomicName id="2E824D69AC61434309CDF9128D3F00B2" authorityName="Strand" authorityYear="1915" box="[729,858,1753,1779]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC61434309CDF9128D3F00B2" box="[729,858,1753,1779]" italics="true" pageId="56" pageNumber="57">A. caneibia</emphasis>
</taxonomicName>
suitable for genetic analysis. Two sequences of
<taxonomicName id="2E824D69AC6143430E90F9128E920156" authorityName="Strand" authorityYear="1915" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6143430E90F9128E920156" italics="true" pageId="56" pageNumber="57">A. caneibia</emphasis>
</taxonomicName>
were generated from specimens from Crete, and they were compared to
<taxonomicName id="2E824D69AC6143430F58F9368AB80156" authorityName="Perkins" authorityYear="1914" box="[1100,1245,1789,1815]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6143430F58F9368AB80156" box="[1100,1245,1789,1815]" italics="true" pageId="56" pageNumber="57">A. alfkenella</emphasis>
</taxonomicName>
specimens from
<collectingCountry id="9195767AAC6143430B83F8EA8EBF017A" box="[151,218,1825,1851]" name="Spain" pageId="56" pageNumber="57">Spain</collectingCountry>
,
<collectingCountry id="9195767AAC6143430BF2F8EA8F2D017A" box="[230,328,1825,1851]" name="Portugal" pageId="56" pageNumber="57">Portugal</collectingCountry>
,
<collectingCountry id="9195767AAC6143430A47F8EA8FD9017A" box="[339,444,1825,1851]" name="Germany" pageId="56" pageNumber="57">Germany</collectingCountry>
, and
<collectingCountry id="9195767AAC6143430AECF8EA8C38017A" box="[504,605,1825,1851]" name="Romania" pageId="56" pageNumber="57">Romania</collectingCountry>
(
<figureCitation id="71B92A6FAC6143430979F8EA8CBB017A" box="[621,734,1825,1851]" captionStart="FIGURE 37" captionStartId="57.[151,250,1819,1843]" captionTargetBox="[151,1433,311,1794]" captionTargetId="figure-78@57.[151,1435,309,1796]" captionTargetPageId="57" captionText="FIGURE 37. Phylogenetic tree (maximum likelihood) of Andrena subgenus Micrandrena Ashmead, 1899 based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted)." figureDoi="http://doi.org/10.5281/zenodo.13209387" httpUri="https://zenodo.org/record/13209387/files/figure.png" pageId="56" pageNumber="57">Figure 37</figureCitation>
; not including
<taxonomicName id="2E824D69AC6143430893F8EA8AAC017A" authority="Noskiewicz, 1939" authorityName="Noskiewicz" authorityYear="1939" box="[903,1225,1825,1851]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC6143430893F8EA8D93017A" box="[903,1014,1825,1851]" italics="true" pageId="56" pageNumber="57">A. pillichi</emphasis>
<bibRefCitation id="8D134B1BAC61434308EAF8EA8AAC017A" author="Noskiewicz, J." box="[1022,1225,1825,1851]" pageId="56" pageNumber="57" pagination="240 - 265" refId="ref89503" refString="Noskiewicz, J. (1939) Beitrage zur Kenntnis der Bienenfauna Ungarns. Polskie pismo entomologiczne, 16 / 17, 240 - 265." type="journal article" year="1939">Noskiewicz, 1939</bibRefCitation>
</taxonomicName>
which is currently lumped under
<taxonomicName id="2E824D69AC6143430A29F88E8FA9011E" authorityName="Perkins" authorityYear="1914" box="[317,460,1861,1887]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6143430A29F88E8FA9011E" box="[317,460,1861,1887]" italics="true" pageId="56" pageNumber="57">A. alfkenella</emphasis>
</taxonomicName>
). The sequences from Crete were separated from continental European sequences by an average genetic distance of 1.77% (range 0.792.29%). The greatest distance was seen between specimens from north-western
<collectingCountry id="9195767AAC6143430A68F8468FD901E6" box="[380,444,1933,1959]" name="Spain" pageId="56" pageNumber="57">Spain</collectingCountry>
and Crete, but the Cretan specimens were separated from the Romanian specimen by only 0.791.07%. Collectively, all sequences formed a clade with bootstrap support of 91, and were consistently separated from a clade formed by
<taxonomicName id="2E824D69AC6143430930F81E8CF001AE" authorityName="Noskiewicz" authorityYear="1939" box="[548,661,2005,2031]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC6143430930F81E8CF001AE" box="[548,661,2005,2031]" italics="true" pageId="56" pageNumber="57">A. pillichi</emphasis>
</taxonomicName>
that had bootstrap support of 83.
<taxonomicName id="2E824D69AC6143430F34F81E8A9201AE" authorityName="Perkins" authorityYear="1914" box="[1056,1271,2005,2031]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6143430F34F81E8A9201AE" box="[1056,1271,2005,2031]" italics="true" pageId="56" pageNumber="57">Andrena alfkenella</emphasis>
</taxonomicName>
+
<taxonomicName id="2E824D69AC6143430E0EF81E8BF901AE" authorityName="Strand" authorityYear="1915" box="[1306,1436,2005,2031]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6143430E0EF81E8BF901AE" box="[1306,1436,2005,2031]" italics="true" pageId="56" pageNumber="57">A. caneibia</emphasis>
</taxonomicName>
sequences were separated from
<taxonomicName id="2E824D69AC6043420913FF5C8C9506F0" box="[519,752,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC6043420913FF5C8C9506F0" box="[519,752,151,177]" italics="true" pageId="57" pageNumber="58">A. alfkenella pillichi</emphasis>
</taxonomicName>
sequences by an average genetic distance of 4.08% (range 3.654.56%). In this context, it is best to consider
<taxonomicName id="2E824D69AC60434209C9FF708D3B0694" authorityName="Strand" authorityYear="1915" box="[733,862,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC60434209C9FF708D3B0694" box="[733,862,187,213]" italics="true" pageId="57" pageNumber="58">A. caneibia</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="2E824D69AC6043420F6EFF708B6C0694" authorityName="Perkins" authorityYear="1914" box="[1146,1289,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="alfkenella" status="syn. nov.">
<emphasis id="DBF6EAF8AC6043420F6EFF708B6C0694" box="[1146,1289,187,213]" italics="true" pageId="57" pageNumber="58">A. alfkenella</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC6043420E06FF708B190694" bold="true" box="[1298,1404,187,213]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="C0C55783AC6043420E06FF708B190694" box="[1298,1404,187,213]" pageId="57" pageNumber="58">syn. nov.</taxonomicNameLabel>
</emphasis>
as the morphological and genetic differences are slight, especially when compared to those observed for
<taxonomicName id="2E824D69AC6043420E1BFF148E83075C" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="subSpecies" species="alfkenella" subSpecies="pillichi">
<emphasis id="DBF6EAF8AC6043420E1BFF148E83075C" italics="true" pageId="57" pageNumber="58">A. alfkenella pillichi</emphasis>
</taxonomicName>
which is formally raised to species status below.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC6043420B83F8D08AB30116" ID-DOI="http://doi.org/10.5281/zenodo.13209387" ID-Zenodo-Dep="13209387" httpUri="https://zenodo.org/record/13209387/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1819,1843]" targetBox="[151,1433,311,1794]" targetPageId="57" targetType="figure">
<paragraph id="E93D36EAAC6043420B83F8D08AB30116" blockId="57.[151,1436,1819,1880]" pageId="57" pageNumber="58">
<emphasis id="DBF6EAF8AC6043420B83F8D08F400172" bold="true" box="[151,293,1819,1843]" pageId="57" pageNumber="58">FIGURE 37.</emphasis>
Phylogenetic tree (maximum likelihood) of
<taxonomicName id="2E824D69AC604342081CF8D08D3B0172" baseAuthorityName="Popov" baseAuthorityYear="1960" box="[776,862,1819,1843]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC604342081CF8D08D3B0172" box="[776,862,1819,1843]" italics="true" pageId="57" pageNumber="58">Andrena</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="2E824D69AC60434208C0F8D08B6D0175" authority="Ashmead, 1899" authorityName="Ashmead" authorityYear="1899" box="[980,1288,1819,1844]" pageId="57" pageNumber="58" rank="subGenus" subGenus="Micrandrena">
<emphasis id="DBF6EAF8AC60434208C0F8D08A3C0172" box="[980,1113,1819,1843]" italics="true" pageId="57" pageNumber="58">Micrandrena</emphasis>
Ashmead, 1899
</taxonomicName>
based on the mitochondrial COI gene. Numbers above branches represent bootstrap support (values of &lt;75 are omitted).
</paragraph>
</caption>
<subSubSection id="A1986561AC6043410BD3F84E8E8F0694" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="E93D36EAAC6043410BD3F84E8E8F0694" blockId="57.[151,1436,1925,2024]" lastBlockId="58.[151,1437,150,357]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
<emphasis id="DBF6EAF8AC6043420BD3F84E8FCA01DE" bold="true" box="[199,431,1925,1951]" pageId="57" pageNumber="58">Material examined.</emphasis>
<materialsCitation id="59EA3CB7AC6043420AA1F84E8F470185" collectingDate="1914-03-01" collectingDateMax="1914-06-30" collectingDateMin="1914-03-01" collectionCode="DEI" collectorName="Paganetti" country="Greece" location="Canea" pageId="57" pageNumber="58" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="holotype">
<emphasis id="DBF6EAF8AC6043420AA1F84E8C5101DE" bold="true" box="[437,564,1925,1951]" pageId="57" pageNumber="58">
<collectingCountry id="9195767AAC6043420AA1F84E8C5501DE" box="[437,560,1925,1951]" name="Greece" pageId="57" pageNumber="58">GREECE</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC604342092EF84D8CE501E1" LSID="urn:lsid:plazi:treatment:612B87FCAC6E43410B83FAD28F0E0724:EC5D6031AC604342092EF84D8CE501E1" box="[570,640,1926,1952]" country="Greece" name="Canea" pageId="57" pageNumber="58" stateProvince="Kriti">Canea</location>
[Chania] (
<collectingRegion id="2B46F808AC60434209E1F84D8D5201E1" box="[757,823,1926,1952]" country="Greece" name="Kriti" pageId="57" pageNumber="58">Creta</collectingRegion>
),
<date id="9D3C102AAC604342085DF84D8D9801E1" box="[841,1021,1926,1952]" pageId="57" pageNumber="58" value="1914-03-01" valueMax="1914-06-30" valueMin="1914-03-01">
<collectingDate id="8D78E9C2AC604342085DF84D8D9801E1" box="[841,1021,1926,1952]" pageId="57" pageNumber="58" value="1914-03-01" valueMax="1914-06-30" valueMin="1914-03-01">1.iii30.vi.1914</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6043420F1CF84D8A4E01E1" box="[1032,1067,1926,1952]" count="1" pageId="57" pageNumber="58" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC6043420F72F84D8AB401E1" box="[1126,1233,1926,1952]" pageId="57" pageNumber="58">Paganetti</collectorName>
,
<collectionCode id="8F93AE2FAC6043420FCFF84D8B6F01E1" box="[1243,1290,1926,1952]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="57" pageNumber="58" type="Other type of research institution/biorepository">DEI</collectionCode>
(
<emphasis id="DBF6EAF8AC6043420E0DF84D8BE501E1" bold="true" box="[1305,1408,1926,1952]" pageId="57" pageNumber="58">
<typeStatus id="36398848AC6043420E0DF84D8BE501E1" box="[1305,1408,1926,1952]" pageId="57" pageNumber="58" type="holotype">holotype</typeStatus>
</emphasis>
of
<taxonomicName id="2E824D69AC6043420B83F8618F720185" authorityName="Strand" authorityYear="1915" box="[151,279,1962,1988]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="caneibia">
<emphasis id="DBF6EAF8AC6043420B83F8618F720185" box="[151,279,1962,1988]" italics="true" pageId="57" pageNumber="58">A. caneibia</emphasis>
</taxonomicName>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6043420A3BF8618B280185" box="[303,1357,1961,1988]" collectingDate="2023-06-06" collectorName="R. Le Divelec" country="Greece" elevation="450" location="Foinikilas" pageId="57" pageNumber="58" specimenCount="16" specimenCount-female="15" specimenCount-male="1" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC6043420A3BF8618F0B0185" box="[303,366,1962,1988]" country="Greece" name="Kriti" pageId="57" pageNumber="58">Crete</collectingRegion>
,
<location id="EC5D6031AC6043420A6EF8618F8F0185" LSID="urn:lsid:plazi:treatment:612B87FCAC6E43410B83FAD28F0E0724:EC5D6031AC6043420A6EF8618F8F0185" box="[378,490,1962,1988]" country="Greece" name="Foinikilas" pageId="57" pageNumber="58" stateProvince="Kriti">Foinikilas</location>
[
<locationDeviation id="F88F2FC8AC6043420AEDF8618CB60185" box="[505,723,1962,1988]" location="Agios Ioannis" pageId="57" pageNumber="58">E of Agios Ioannis</locationDeviation>
],
<quantity id="2E7A9B0FAC60434209F1F8628D4B0185" box="[741,814,1961,1988]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="57" pageNumber="58" unit="m" value="450.0">
<elevation id="62AFD1D9AC60434209F1F8628D4B0185" box="[741,814,1961,1988]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="57" pageNumber="58" unit="m" value="450.0">450 m</elevation>
</quantity>
,
<date id="9D3C102AAC604342082EF8618DC20185" box="[826,935,1962,1988]" pageId="57" pageNumber="58" value="2023-06-06">
<collectingDate id="8D78E9C2AC604342082EF8618DC20185" box="[826,935,1962,1988]" pageId="57" pageNumber="58" value="2023-06-06">6.vi.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC60434208A6F8618DB20185" box="[946,983,1962,1988]" count="1" pageId="57" pageNumber="58" type="male">1♂</specimenCount>
,
<specimenCount id="FF84FD63AC60434208F0F8628A730185" box="[996,1046,1961,1988]" count="15" pageId="57" pageNumber="58" type="female">15♀</specimenCount>
, leg.
<collectorName id="4477533CAC6043420F47F8618A960185" box="[1107,1267,1962,1988]" pageId="57" pageNumber="58">R. Le Divelec</collectorName>
, RLDC
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6043420E4CF8618DC301A9" collectingDate="2023-06-10" collectionCode="TJWC" collectorName="R. Le Divelec" country="Greece" elevation="500" location="Mesavlia" pageId="57" pageNumber="58" specimenCount="6" specimenCount-female="6" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC6043420E4CF8618BF20185" box="[1368,1431,1962,1988]" country="Greece" name="Kriti" pageId="57" pageNumber="58">Crete</collectingRegion>
,
<location id="EC5D6031AC6043420B83F8058F6401A9" LSID="urn:lsid:plazi:treatment:612B87FCAC6E43410B83FAD28F0E0724:EC5D6031AC6043420B83F8058F6401A9" box="[151,257,1998,2024]" country="Greece" name="Mesavlia" pageId="57" pageNumber="58" stateProvince="Kriti">Mesavlia</location>
,
<quantity id="2E7A9B0FAC6043420A1AF8068F3C01A9" box="[270,345,1997,2024]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="57" pageNumber="58" unit="m" value="500.0">
<elevation id="62AFD1D9AC6043420A1AF8068F3C01A9" box="[270,345,1997,2024]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="57" pageNumber="58" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<date id="9D3C102AAC6043420A73F8058F8701A9" box="[359,482,1998,2024]" pageId="57" pageNumber="58" value="2023-06-10">
<collectingDate id="8D78E9C2AC6043420A73F8058F8701A9" box="[359,482,1998,2024]" pageId="57" pageNumber="58" value="2023-06-10">10.vi.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6043420AFBF8058C7101A9" box="[495,532,1998,2024]" count="6" pageId="57" pageNumber="58" type="female">6♀</specimenCount>
, leg.
<collectorName id="4477533CAC6043420941F8058C9C01A9" box="[597,761,1998,2024]" pageId="57" pageNumber="58">R. Le Divelec</collectorName>
, RLDC/
<collectionCode id="8F93AE2FAC604342084FF8058DC301A9" box="[859,934,1998,2024]" pageId="57" pageNumber="58">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC60434108A7F8058C1506F0" collectingDate="2023-06-08" collectorName="R. Le Divelec" country="Greece" lastPageId="58" lastPageNumber="59" location="Skafidaras" pageId="57" pageNumber="58" specimenCount="3" specimenCount-female="3" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC60434208A7F8058D9701A9" box="[947,1010,1998,2024]" country="Greece" name="Kriti" pageId="57" pageNumber="58">Crete</collectingRegion>
,
<location id="EC5D6031AC6043420F14F8058A1301A9" LSID="urn:lsid:plazi:treatment:612B87FCAC6E43410B83FAD28F0E0724:EC5D6031AC6043420F14F8058A1301A9" box="[1024,1142,1998,2024]" country="Greece" name="Skafidaras" pageId="57" pageNumber="58" stateProvince="Kriti">Skafidaras</location>
[
<locationDeviation id="F88F2FC8AC6043420F9DF8058BF701A9" box="[1161,1426,1998,2024]" location="Ag. Dimitrios" pageId="57" pageNumber="58">
<quantity id="2E7A9B0FAC6043420F9DF8058AA101A9" box="[1161,1220,1998,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="57" pageNumber="58" unit="km" value="1.0">1 km</quantity>
W Ag. Dimitrios
</locationDeviation>
],
<date id="9D3C102AAC6343410B83FF5C8F6106F0" box="[151,260,151,177]" pageId="58" pageNumber="59" value="2023-06-08">
<collectingDate id="8D78E9C2AC6343410B83FF5C8F6106F0" box="[151,260,151,177]" pageId="58" pageNumber="59" value="2023-06-08">8.vi.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6343410A04FF5C8F5106F0" box="[272,308,151,177]" count="3" pageId="58" pageNumber="59" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CAC6343410A60FF5C8C7006F0" box="[372,533,151,177]" pageId="58" pageNumber="59">R. Le Divelec</collectorName>
, RLDC
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC6343410968FF5C8E800694" collectingDate="2023-06-08" collectorName="R. Le Divelec" country="Greece" location="Vamos Chania" pageId="58" pageNumber="59" specimenCount="5" specimenCount-female="5" stateProvince="Kriti" typeStatus="holotype">
<collectingRegion id="2B46F808AC6343410968FF5C8CDE06F0" box="[636,699,151,177]" country="Greece" name="Kriti" pageId="58" pageNumber="59">Crete</collectingRegion>
,
<location id="EC5D6031AC63434109DCFF5C8D0E06F0" LSID="urn:lsid:plazi:treatment:612B87FCAC6E43410B83FAD28F0E0724:EC5D6031AC63434109DCFF5C8D0E06F0" box="[712,875,151,177]" country="Greece" name="Vamos Chania" pageId="58" pageNumber="59" stateProvince="Kriti">Vamos Chania</location>
[
<locationDeviation id="F88F2FC8AC634341086FFF5C8A6206F0" box="[891,1031,151,177]" location="Vryses" pageId="58" pageNumber="59">E of Vryses</locationDeviation>
],
<date id="9D3C102AAC6343410F0EFF5C8AE306F0" box="[1050,1158,151,177]" pageId="58" pageNumber="59" value="2023-06-08">
<collectingDate id="8D78E9C2AC6343410F0EFF5C8AE306F0" box="[1050,1158,151,177]" pageId="58" pageNumber="59" value="2023-06-08">8.vi.2023</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC6343410F87FF5D8AD206F0" box="[1171,1207,150,177]" count="5" pageId="58" pageNumber="59" type="female">5♀</specimenCount>
, leg.
<collectorName id="4477533CAC6343410FE3FF5C8BFD06F0" box="[1271,1432,151,177]" pageId="58" pageNumber="59">R. Le Divelec</collectorName>
, RLDC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC6343410BD3FF158F0E0724" pageId="58" pageNumber="59" type="distribution">
<paragraph id="E93D36EAAC6343410BD3FF158F0E0724" blockId="58.[151,1437,150,357]" pageId="58" pageNumber="59">
<emphasis id="DBF6EAF8AC6343410BD3FF158F0406B9" bold="true" box="[199,353,222,248]" pageId="58" pageNumber="59">Distribution.</emphasis>
West Palaearctic, from
<collectingCountry id="9195767AAC6343410965FF148CB306B8" box="[625,726,223,249]" name="Morocco" pageId="58" pageNumber="59">Morocco</collectingCountry>
(Atlas Mountains) across Europe to
<collectingCountry id="9195767AAC6343410F6EFF148AAE06B8" box="[1146,1227,223,249]" name="Turkey" pageId="58" pageNumber="59">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC6343410FCCFF148B5806B8" box="[1240,1341,223,249]" name="Lebanon" pageId="58" pageNumber="59">Lebanon</collectingCountry>
, and the Caucasus (
<bibRefCitation id="8D134B1BAC6343410A05FEC88C15075C" author="Gusenleitner, F. &amp; Schwarz, M." box="[273,624,259,285]" pageId="58" pageNumber="59" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC6343410969FEC88D30075C" author="Wood, T. J. &amp; Boustani, M. &amp; Rasmont, P." box="[637,853,259,285]" pageId="58" pageNumber="59" pagination="279 - 312" refId="ref91916" refString="Wood, T. J., Boustani, M. &amp; Rasmont, P. (2020 a) A revision of the Andrena (Hymenoptera: Andrenidae) of Lebanon with the description of six new species. Annales de la Societe entomologique de France, New Series, 56, 279 - 312. https: // doi. org / 10.1080 / 00379271.2020.1794960" type="journal article" year="2020">
Wood
<emphasis id="DBF6EAF8AC63434109DCFEC88C9E075C" box="[712,763,259,285]" italics="true" pageId="58" pageNumber="59">et al</emphasis>
. 2020a
</bibRefCitation>
). Not found in the remaining parts of North Africa or the Middle East. See notes below on the status of
<taxonomicName id="2E824D69AC63434109F9FEEC8D390700" authorityName="Noskiewicz" authorityYear="1939" box="[749,860,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="pillichi">
<emphasis id="DBF6EAF8AC63434109F9FEEC8D390700" box="[749,860,295,321]" italics="true" pageId="58" pageNumber="59">A. pillichi</emphasis>
</taxonomicName>
which has previously been lumped into a broad
<taxonomicName id="2E824D69AC6343410E90FEEC8F600724" authorityName="Perkins" authorityYear="1914" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="58" pageNumber="59" phylum="Arthropoda" rank="species" species="alfkenella">
<emphasis id="DBF6EAF8AC6343410E90FEEC8F600724" italics="true" pageId="58" pageNumber="59">A. alfkenella</emphasis>
</taxonomicName>
concept.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,510 @@
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<paragraph id="E93D36EAAC7743550B83FA948CBA0338" blockId="46.[151,735,1375,1402]" box="[151,735,1375,1402]" pageId="46" pageNumber="47">
<heading id="B2758186AC7743550B83FA948CBA0338" bold="true" box="[151,735,1375,1402]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
<taxonomicName id="2E824D69AC7743550B83FA948CBA0338" ID-CoL="DRPZ" authority="(Christ, 1791)" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[151,735,1375,1402]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550B83FA948CBA0338" bold="true" box="[151,735,1375,1402]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550B83FA948F64033B" bold="true" box="[151,257,1375,1402]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7743550A06FA948FDA033B" bold="true" box="[274,447,1375,1402]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7743550AC4FA948C4E033B" bold="true" box="[464,555,1375,1402]" italics="true" pageId="46" pageNumber="47">fulvago</emphasis>
(
<bibRefCitation id="8D134B1BAC774355092FFA948CB20338" author="Christ, J. L." box="[571,727,1375,1401]" pageId="46" pageNumber="47" refId="ref88151" refString="Christ, J. L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten vom Bienen, Wespen und Ameisengeschlecht. Herrmann, Frankfurt, Main, 535 pp. https: // doi. org / 10.5962 / bhl. title. 87724" type="book" year="1791">Christ, 1791</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7743550B83FA6E8A0F03BB" pageId="46" pageNumber="47" type="reference_group">
<paragraph id="E93D36EAAC7743550B83FA6E8CC003FD" blockId="46.[151,1130,1444,1530]" box="[151,677,1444,1469]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA6E8CC003FD" box="[151,677,1444,1469]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA6E8FAB03FD" author="Christ, J. L." box="[151,462,1444,1468]" page="189" pageId="46" pageNumber="47" year="1791">
<taxonomicName id="2E824D69AC7743550B83FA6E8FAB03FD" authority="Christ, 1791: 189" authorityName="Christ" authorityPageNumber="189" authorityYear="1791" box="[151,462,1444,1468]" class="Insecta" family="Apidae" genus="Apis" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7743550B83FA6E8F7203FD" box="[151,279,1444,1468]" italics="true" pageId="46" pageNumber="47">Apis fulvago</emphasis>
<materialsCitation id="59EA3CB7AC7743550A09FA6F8FAB03FD" box="[285,462,1444,1468]" country="Germany" location="Germany" pageId="46" pageNumber="47" specimenCount="1">
<bibRefCitation id="8D134B1BAC7743550A09FA6F8FAB03FD" author="Christ, J. L." box="[285,462,1444,1468]" pageId="46" pageNumber="47" refId="ref88151" refString="Christ, J. L. (1791) Naturgeschichte, Klassification und Nomenclatur der Insekten vom Bienen, Wespen und Ameisengeschlecht. Herrmann, Frankfurt, Main, 535 pp. https: // doi. org / 10.5962 / bhl. title. 87724" type="book" year="1791">Christ, 1791: 189</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
[
<collectingCountry id="9195767AAC7743550ACEFA6F8C5903FD" box="[474,572,1444,1468]" name="Germany" pageId="46" pageNumber="47">Germany</collectingCountry>
,
<typeStatus id="36398848AC7743550952FA6E8C1703FC" box="[582,626,1445,1469]" pageId="46" pageNumber="47">type</typeStatus>
lost]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC7743550B83FA088DFB039A" blockId="46.[151,1130,1444,1530]" box="[151,926,1475,1499]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA088DFB039A" box="[151,926,1475,1499]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA088C6A039A" author="Smith, F." box="[151,527,1475,1499]" page="59" pageId="46" pageNumber="47" year="1849">
<taxonomicName id="2E824D69AC7743550B83FA088C6A039A" authority="Smith, 1849: 59" authorityName="Smith" authorityPageNumber="59" authorityYear="1849" box="[151,527,1475,1499]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="DBF6EAF8AC7743550B83FA088F06039A" box="[151,355,1475,1499]" italics="true" pageId="46" pageNumber="47">Andrena constrictus</emphasis>
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</materialsCitation>
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,
<specimenCount id="FF84FD63AC774355090DFA088C49039B" box="[537,556,1475,1498]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC774355092CFA088CF3039A" box="[568,662,1475,1499]" name="United Kingdom" pageId="46" pageNumber="47">Scotland</collectingCountry>
, OUMNH, not examined]
</treatmentCitationGroup>
</paragraph>
<paragraph id="E93D36EAAC7743550B83FA2A8A0F03BB" blockId="46.[151,1130,1444,1530]" box="[151,1130,1505,1530]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="C99211C4AC7743550B83FA2A8A0F03BB" box="[151,1130,1505,1530]" pageId="46" pageNumber="47">
<treatmentCitation id="682310FBAC7743550B83FA2A8C4D03B8" author="Morawitz, F." box="[151,552,1505,1529]" page="68" pageId="46" pageNumber="47" year="1886">
<taxonomicName id="2E824D69AC7743550B83FA2A8C4D03B8" ID-CoL="DRCJ" authority="Morawitz, 1886: 68" authorityName="Morawitz" authorityPageNumber="68" authorityYear="1886" box="[151,552,1505,1529]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550B83FA2A8F3D03B8" box="[151,344,1505,1529]" italics="true" pageId="46" pageNumber="47">Andrena colonialis</emphasis>
<materialsCitation id="59EA3CB7AC7743550A4AFA2A8C4D03B8" box="[350,552,1505,1529]" collectionCode="ZISP" country="Azerbaijan" location="Azerbaijan" pageId="46" pageNumber="47" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
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</materialsCitation>
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,
<specimenCount id="FF84FD63AC7743550927FA298C2603B8" box="[563,579,1506,1529]" count="1" pageId="46" pageNumber="47" type="female"></specimenCount>
<specimenCount id="FF84FD63AC7743550957FA298C3C03B8" box="[579,601,1506,1529]" count="1" pageId="46" pageNumber="47" type="male"></specimenCount>
[
<collectingCountry id="9195767AAC7743550971FA2A8CBF03B8" box="[613,730,1505,1529]" name="Azerbaijan" pageId="46" pageNumber="47">Azerbaijan</collectingCountry>
, ZISP, photograph examined]
<emphasis id="DBF6EAF8AC7743550F19FA298A0F03BB" bold="true" box="[1037,1130,1506,1530]" pageId="46" pageNumber="47">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
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<subSubSection id="A1986561AC77434B0B83F9E38B17048C" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="discussion">
<paragraph id="E93D36EAAC7743550B83F9E38C6E015B" blockId="46.[151,1437,1576,2035]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550B83F9E38F6E0003" bold="true" box="[151,267,1576,1602]" pageId="46" pageNumber="47">Remarks.</emphasis>
<taxonomicName id="2E824D69AC7743550A00F9E38FB50003" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[276,464,1576,1602]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7743550A00F9E38FB50003" box="[276,464,1576,1602]" italics="true" pageId="46" pageNumber="47">Andrena fulvago</emphasis>
</taxonomicName>
is one of the oldest described
<taxonomicName id="2E824D69AC7743550825F9E38D8B0003" box="[817,1006,1576,1602]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550825F9E38DF50003" box="[817,912,1576,1602]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
species
</taxonomicName>
, and the concept has remained robust for more than two centuries despite the lack of
<typeStatus id="36398848AC77435509A5F9878C840027" box="[689,737,1612,1638]" pageId="46" pageNumber="47">type</typeStatus>
material. It is the
<typeStatus id="36398848AC77435508A6F9878D870027" box="[946,994,1612,1638]" pageId="46" pageNumber="47">type</typeStatus>
species of the subgenus
<taxonomicName id="2E824D69AC7743550FE9F9878F5C00CB" authority="Hedicke, 1933" authorityName="Hedicke" authorityYear="1933" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550FE9F9878BF90027" box="[1277,1436,1612,1638]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
Hedicke, 1933
</taxonomicName>
that is found across the Palaearctic and which was confirmed as distinct from the Palaearctic subgenus
<taxonomicName id="2E824D69AC7743550B83F95F8FA500EF" authority="Hedicke, 1933" authorityName="Hedicke" authorityYear="1933" box="[151,448,1684,1710]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC7743550B83F95F8F7700EF" box="[151,274,1684,1710]" italics="true" pageId="46" pageNumber="47">Euandrena</emphasis>
Hedicke, 1933
</taxonomicName>
by
<bibRefCitation id="8D134B1BAC7743550AF9F95F8C8300EF" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[493,742,1684,1710]" pageId="46" pageNumber="47" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC774355095CF95E8C1F00EF" box="[584,634,1684,1710]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (2022b)
</bibRefCitation>
. The two subgenera are very similar, but can be most clearly separated in the female sex by the presence of plumose scopal hairs in female
<taxonomicName id="2E824D69AC77435508E1F9738AF10093" authorityName="Hedicke" authorityYear="1933" box="[1013,1172,1720,1746]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC77435508E1F9738AF10093" box="[1013,1172,1720,1746]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
as an adaptation for the collection of
<taxonomicName id="2E824D69AC7743550A38F9178FAC00B7" box="[300,457,1756,1782]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="subFamily" subFamily="Cichorioideae">Cichorioideae</taxonomicName>
(
<taxonomicName id="2E824D69AC7743550ACCF9178C3200B7" box="[472,599,1756,1782]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="46" pageNumber="47" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) pollen, all known species of
<taxonomicName id="2E824D69AC77435508B3F9178A2300B7" authorityName="Hedicke" authorityYear="1933" box="[935,1094,1756,1782]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC77435508B3F9178A2300B7" box="[935,1094,1756,1782]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
specialising on this subfamily (
<bibRefCitation id="8D134B1BAC7743550B8AF8CB8F20015B" author="Westrich, P." box="[158,325,1792,1818]" pageId="46" pageNumber="47" refId="ref91897" refString="Westrich, P. (1989) Die wildbienen Baden-Wurttembergs. Eugen Ulmer, Stuttgart, 972 pp." type="book" year="1989">Westrich 1989</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7743550A45F8CB8F86015B" author="Wood, T. J." box="[337,483,1792,1818]" pageId="46" pageNumber="47" pagination="241 - 484" refId="ref92391" refString="Wood, T. J. (2023 b) The genus Andrena Fabricius, 1775 in the Iberian Peninsula (Hymenoptera: Andrenidae). Journal of Hymenoptera Research, 96, 241 - 484. https: // doi. org / 10.3897 / jhr. 96.101873" type="journal article" year="2023">Wood 2023b</bibRefCitation>
; c).
</paragraph>
<paragraph id="E93D36EAAC77434B0BD3F8EF8F7F06B8" blockId="46.[151,1437,1576,2035]" lastBlockId="48.[151,1437,151,933]" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47">
The majority of the important
<taxonomicName id="2E824D69AC7743550902F8EF8CA9017F" box="[534,716,1828,1854]" pageId="46" pageNumber="47">
<emphasis id="DBF6EAF8AC7743550902F8EF8C10017F" box="[534,629,1828,1854]" italics="true" pageId="46" pageNumber="47">Andrena</emphasis>
species
</taxonomicName>
described by Ferdinand Morawitz have recently been illustrated, including the designation of
<typeStatus id="36398848AC7743550AC9F8838C2A0123" box="[477,591,1864,1890]" pageId="46" pageNumber="47" type="lectotype">lectotypes</typeStatus>
where appropriate (
<bibRefCitation id="8D134B1BAC7743550827F8838A530123" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[819,1078,1864,1890]" pageId="46" pageNumber="47" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC77435508AEF8828D890123" box="[954,1004,1864,1890]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2021
</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7743550F57F8838AEB0123" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A." box="[1091,1166,1864,1890]" pageId="46" pageNumber="47" pagination="105 - 176" refId="ref87433" refString="Astafurova, Y. A., Proshchalykin, M. Y. &amp; Sidorov, D. A. (2022 b) The bees of the genus Andrena Fabricius, 1775 (Hymenoptera, Andrenidae) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko. Zookeys, 1120, 105 - 176. https: // doi. org / 10.3897 / zookeys. 1120.90206" type="journal article" year="2022">2022b</bibRefCitation>
). This includes the type material for
<taxonomicName id="2E824D69AC7743550A3DF8A78FDF01C7" authorityName="Morawitz" authorityYear="1886" box="[297,442,1900,1926]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550A3DF8A78FDF01C7" box="[297,442,1900,1926]" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
which was described from the Caucasus (
<bibRefCitation id="8D134B1BAC77435508BEF8A78AD001C7" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[938,1205,1900,1926]" pageId="46" pageNumber="47" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC7743550F27F8A68A0D01C7" box="[1075,1128,1900,1926]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2021
</bibRefCitation>
). The identity of
<taxonomicName id="2E824D69AC7743550E90F8A78F6001EB" authorityName="Morawitz" authorityYear="1886" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550E90F8A78F6001EB" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
has been obscure due to its differential treatment in the literature.
<bibRefCitation id="8D134B1BAC7743550F12F85B8B1401EB" author="Gusenleitner, F. &amp; Schwarz, M." box="[1030,1393,1936,1962]" pageId="46" pageNumber="47" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
list
<taxonomicName id="2E824D69AC7743550B83F87F8F40018F" authorityName="Morawitz" authorityYear="1886" box="[151,293,1972,1998]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7743550B83F87F8F40018F" box="[151,293,1972,1998]" italics="true" pageId="46" pageNumber="47">A. colonialis</emphasis>
</taxonomicName>
as a
<taxonomicName id="2E824D69AC7743550A76F87F8C64018F" authorityName="Hedicke" authorityYear="1933" box="[354,513,1972,1998]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550A76F87F8C64018F" box="[354,513,1972,1998]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
, but
<bibRefCitation id="8D134B1BAC7743550928F87F8D35018F" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[572,848,1972,1998]" pageId="46" pageNumber="47" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC77435509D0F87E8C93018F" box="[708,758,1972,1998]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (2008)
</bibRefCitation>
list it as a
<taxonomicName id="2E824D69AC77435508DAF87F8A2C018F" authorityName="Hedicke" authorityYear="1933" box="[974,1097,1972,1998]" pageId="46" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC77435508DAF87F8A2C018F" box="[974,1097,1972,1998]" italics="true" pageId="46" pageNumber="47">Euandrena</emphasis>
</taxonomicName>
, and do not include it in their revision of
<taxonomicName id="2E824D69AC7743550A0CF8138FDC01B3" authorityName="Hedicke" authorityYear="1933" box="[280,441,2008,2034]" pageId="46" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7743550A0CF8138FDC01B3" box="[280,441,2008,2034]" italics="true" pageId="46" pageNumber="47">Chrysandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC7743550AD3F8138CAB01B2" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Cierzniak, T." box="[455,718,2008,2035]" pageId="46" pageNumber="47" refId="ref89759" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Cierzniak, T. (2005) Andreninae of the Central and Eastern Palaearctic. Part 1. Polish Entomological Monographs II. Polish Entomological Society, Poznan, 235 pp." type="book" year="2005">
Osytshnjuk
<emphasis id="DBF6EAF8AC7743550945F8128CE601B3" box="[593,643,2008,2034]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. 2005
</bibRefCitation>
). This is surprising, because the female
<typeStatus id="36398848AC7743550F85F8138A9C01B3" box="[1169,1273,2008,2034]" pageId="46" pageNumber="47" type="lectotype">lectotype</typeStatus>
designated by
<bibRefCitation id="8D134B1BAC7643540B89F9358FCC0159" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[157,425,1790,1816]" pageId="47" pageNumber="48" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC7643540A36F9348F370159" box="[290,338,1790,1816]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2008)
</bibRefCitation>
and illustrated by
<bibRefCitation id="8D134B1BAC7643540966F9358DC10158" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[626,932,1790,1817]" pageId="47" pageNumber="48" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC76435409E0F9348D460159" box="[756,803,1790,1816]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2021: 15)
</bibRefCitation>
is clearly a
<taxonomicName id="2E824D69AC7643540F31F9358AA10159" authorityName="Hedicke" authorityYear="1933" box="[1061,1220,1790,1816]" pageId="47" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7643540F31F9358AA10159" box="[1061,1220,1790,1816]" italics="true" pageId="47" pageNumber="48">Chrysandrena</emphasis>
</taxonomicName>
, as it has obviously plumose tibial scopal hairs. Structurally, there is no difference between
<taxonomicName id="2E824D69AC76435408C3F8E98A02017D" authorityName="Morawitz" authorityYear="1886" box="[983,1127,1826,1852]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC76435408C3F8E98A02017D" box="[983,1127,1826,1852]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7643540FB0F8E98B7F017D" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1188,1306,1826,1852]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7643540FB0F8E98B7F017D" box="[1188,1306,1826,1852]" italics="true" pageId="47" pageNumber="48">A. fulvago</emphasis>
</taxonomicName>
; both have narrow facial foveae that narrow further ventrally, plumose tibial scopae, hind tibiae that are lightened orange, pale pubescence over the remaining parts of the body, a light terminal fringe, and the clypeus, scutum, and terga display a polished integument with distinct isolated punctures. The only major difference is that the colouration of the tergal margins is lighter and more extensive in
<taxonomicName id="2E824D69AC7643540976F8798C95018D" authorityName="Morawitz" authorityYear="1886" box="[610,752,1970,1996]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7643540976F8798C95018D" box="[610,752,1970,1996]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
. However, this is variable within
<taxonomicName id="2E824D69AC7643540F79F8798A85018D" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1133,1248,1970,1996]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC7643540F79F8798A85018D" box="[1133,1248,1970,1996]" italics="true" pageId="47" pageNumber="48">A. fulvago</emphasis>
</taxonomicName>
across its range, and is considered to represent only variation. Furthermore,
<bibRefCitation id="8D134B1BAC7643540838F81D8A5E01B1" author="Osytshnjuk, A. Z. &amp; Romasenko, L. &amp; Banaszak, J. &amp; Motyka, E." box="[812,1083,2006,2032]" pageId="47" pageNumber="48" refId="ref89808" refString="Osytshnjuk, A. Z., Romasenko, L., Banaszak, J. &amp; Motyka, E. (2008) Andreninae of the Central and Eastern Palearctic. Part 2. Polish Entomological Monographs V. Polish Entomological Society, Poznan, 234 pp." type="book" year="2008">
Osytshnjuk
<emphasis id="DBF6EAF8AC76435408A6F81C8D8601B1" box="[946,995,2006,2032]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2008)
</bibRefCitation>
report that
<taxonomicName id="2E824D69AC7643540FA8F81D8B2C01B1" authorityName="Morawitz" authorityYear="1886" box="[1212,1353,2006,2032]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="colonialis">
<emphasis id="DBF6EAF8AC7643540FA8F81D8B2C01B1" box="[1212,1353,2006,2032]" italics="true" pageId="47" pageNumber="48">A. colonialis</emphasis>
</taxonomicName>
flies in July and August and is a specialist of
<taxonomicName id="2E824D69AC69434B092AFF5C8CD906F0" box="[574,700,151,177]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
, foraging predominantly on
<taxonomicName id="2E824D69AC69434B08EFFF5C8AAE06F0" box="[1019,1227,151,177]" class="Magnoliopsida" family="Asteraceae" genus="Cichorium" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="intybus">
<emphasis id="DBF6EAF8AC69434B08EFFF5C8AAE06F0" box="[1019,1227,151,177]" italics="true" pageId="48" pageNumber="49">Cichorium intybus</emphasis>
</taxonomicName>
. This is consistent with the flight period and dietary niche of
<taxonomicName id="2E824D69AC69434B0997FF708C9C0694" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[643,761,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0997FF708C9C0694" box="[643,761,187,213]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
; oligolecty on
<taxonomicName id="2E824D69AC69434B08B3FF708A470694" box="[935,1058,187,213]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
is currently unknown in the true
<taxonomicName id="2E824D69AC69434B0B83FF148F7706B8" authorityName="Hedicke" authorityYear="1933" box="[151,274,223,249]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0B83FF148F7706B8" box="[151,274,223,249]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BDFD6662AC7643540B83F9A48C280091" ID-DOI="http://doi.org/10.5281/zenodo.13209373" ID-Zenodo-Dep="13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1647,1671]" targetBox="[151,1436,181,1624]" targetPageId="47" targetType="figure">
<paragraph id="E93D36EAAC7643540B83F9A48C280091" blockId="47.[151,1437,1647,1744]" pageId="47" pageNumber="48">
<emphasis id="DBF6EAF8AC7643540B83F9A48F4500C6" bold="true" box="[151,288,1647,1671]" pageId="47" pageNumber="48">FIGURE 30.</emphasis>
<taxonomicName id="2E824D69AC7643540A31F9BB8DBE00C6" authority="Tadauchi, Miyanaga &amp; Dawut, 2005" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[293,987,1647,1672]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="almas" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC7643540A31F9BB8F1E00C9" box="[293,379,1648,1672]" italics="true" pageId="47" pageNumber="48">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7643540A9EF9BB8C7F00C9" box="[394,538,1648,1672]" italics="true" pageId="47" pageNumber="48">Chrysandrena</emphasis>
)
<emphasis id="DBF6EAF8AC764354093CF9BB8C0000C9" box="[552,613,1648,1672]" italics="true" pageId="47" pageNumber="48">almas</emphasis>
<bibRefCitation id="8D134B1BAC764354097EF9BB8DBE00C6" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[618,987,1647,1672]" pageId="47" pageNumber="48" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">Tadauchi, Miyanaga &amp; Dawut, 2005</bibRefCitation>
</taxonomicName>
female from Kazakhstan. A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC69434B0BD3FEC88A250420" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
This association between
<taxonomicName id="2E824D69AC69434B0AFBFEC88CEB075C" authorityName="Hedicke" authorityYear="1933" box="[495,654,259,285]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC69434B0AFBFEC88CEB075C" box="[495,654,259,285]" italics="true" pageId="48" pageNumber="49">Chrysandrena</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC69434B09DCFEC88D26075C" box="[712,835,259,285]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
can be reinforced by the new transfer made here of
<taxonomicName id="2E824D69AC69434B0B83FEEC8CBA0700" authority="Tadauchi, Miyanaga &amp; Dawut, 2005" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[151,735,294,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B0B83FEEC8F250700" box="[151,320,295,321]" italics="true" pageId="48" pageNumber="49">Andrena almas</emphasis>
<bibRefCitation id="8D134B1BAC69434B0A53FEEC8CBA0700" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[327,735,294,321]" pageId="48" pageNumber="49" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">Tadauchi, Miyanaga &amp; Dawut, 2005</bibRefCitation>
</taxonomicName>
from
<taxonomicName id="2E824D69AC69434B0837FEEC8DFB0700" authorityName="Hedicke" authorityYear="1933" box="[803,926,295,321]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0837FEEC8DFB0700" box="[803,926,295,321]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
to
<taxonomicName id="2E824D69AC69434B08D6FEEC8A040700" authorityName="Hedicke" authorityYear="1933" box="[962,1121,295,321]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">
<emphasis id="DBF6EAF8AC69434B08D6FEEC8A040700" box="[962,1121,295,321]" italics="true" pageId="48" pageNumber="49">Chrysandrena</emphasis>
</taxonomicName>
. This species was described from
<collectingRegion id="2B46F808AC69434B0BC0FE808F530724" box="[212,310,331,357]" country="China" name="Xinjiang Uygur" pageId="48" pageNumber="49">Xinjiang</collectingRegion>
in western
<collectingCountry id="9195767AAC69434B0AA3FE808F9F0724" box="[439,506,331,357]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
where it flies in August and is associated with
<taxonomicName id="2E824D69AC69434B0F12FE808B6C0724" box="[1030,1289,331,357]" class="Magnoliopsida" family="Asteraceae" genus="Chondrilla" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="species" species="brevirostris">
<emphasis id="DBF6EAF8AC69434B0F12FE808B6C0724" box="[1030,1289,331,357]" italics="true" pageId="48" pageNumber="49">Chondrilla brevirostris</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69AC69434B0E02FE808BF00724" box="[1302,1429,331,357]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
) (
<bibRefCitation id="8D134B1BAC69434B0B8AFEA48FF107C8" author="Tadauchi, O. &amp; Miyanaga, R. &amp; Dawut, A." box="[158,404,366,393]" pageId="48" pageNumber="49" pagination="9 - 17" refId="ref91507" refString="Tadauchi, O., Miyanaga, R. &amp; Dawut, A. (2005) A new species belonging to the subgenus Euandrena of the genus Andrena from Xinjiang Uygur, China with notes on nest structure (Hymenoptera, Andrenidae). Esakia, 45, 9 - 17. https: // doi. org / 10.5109 / 2699" type="journal article" year="2005">
Tadauchi
<emphasis id="DBF6EAF8AC69434B0A05FEA48F2307C8" box="[273,326,367,393]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2005
</bibRefCitation>
). In the description, the authors clearly mention the plumose scopal hairs, but concluded on a placement in the
<taxonomicName id="2E824D69AC69434B0A8CFE588C7607EC" authorityName="Hedicke" authorityYear="1933" box="[408,531,403,429]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0A8CFE588C7607EC" box="[408,531,403,429]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
. Confusingly,
<taxonomicName id="2E824D69AC69434B09A8FE588D4507EC" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[700,800,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B09A8FE588D4507EC" box="[700,800,403,429]" italics="true" pageId="48" pageNumber="49">A. almas</emphasis>
</taxonomicName>
was later not included in a subsequent revision of East Asian
<taxonomicName id="2E824D69AC69434B0BF7FE7C8F3B0790" authorityName="Hedicke" authorityYear="1933" box="[227,350,439,465]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0BF7FE7C8F3B0790" box="[227,350,439,465]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
(
<bibRefCitation id="8D134B1BAC69434B0A64FE7C8C0D0790" author="Xu, H. - L. &amp; Tadauchi, O." box="[368,616,439,465]" pageId="48" pageNumber="49" pagination="77 - 90" refId="ref92855" refString="Xu, H. - L. &amp; Tadauchi, O. (2012) A revision of the subgenus Euandrena of the genus Andrena of Eastern Asia (Hymenoptera: Apoidea: Andrenidae). Esakia, 52, 77 - 90. https: // doi. org / 10.5109 / 25400" type="journal article" year="2012">Xu &amp; Tadauchi 2012</bibRefCitation>
) for unspecified reasons even though its description was mentioned in the introduction of this work. The combination of plumose scopal hairs, summer flight period, and association with
<taxonomicName id="2E824D69AC69434B0BC5FE348F290458" box="[209,332,511,537]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="48" pageNumber="49" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
supports movement from
<taxonomicName id="2E824D69AC69434B0968FE348C920458" authorityName="Hedicke" authorityYear="1933" box="[636,759,511,537]" pageId="48" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8AC69434B0968FE348C920458" box="[636,759,511,537]" italics="true" pageId="48" pageNumber="49">Euandrena</emphasis>
</taxonomicName>
to
<emphasis id="DBF6EAF8AC69434B080BFE348DA30458" box="[799,966,511,537]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="2E824D69AC69434B080BFE348DA70458" authorityName="Hedicke" authorityYear="1933" box="[799,962,511,537]" pageId="48" pageNumber="47" rank="subGenus" subGenus="Chrysandrena">Chrysandrena</taxonomicName>
.
</emphasis>
Examination of additional material from
<collectingCountry id="9195767AAC69434B0B83FDE88F7F047C" box="[151,282,547,573]" name="Kazakhstan" pageId="48" pageNumber="49">Kazakhstan</collectingCountry>
(
<figureCitation id="71B92A6FAC69434B0A24FDE88F8E047C" box="[304,491,547,573]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30AD</figureCitation>
),
<collectingCountry id="9195767AAC69434B0911FDE88CEB047C" box="[517,654,547,573]" name="Afghanistan" pageId="48" pageNumber="49">Afghanistan</collectingCountry>
(
<figureCitation id="71B92A6FAC69434B09B0FDE88D32047C" box="[676,855,547,573]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30EF</figureCitation>
), and
<collectingCountry id="9195767AAC69434B08BDFDE88D91047C" box="[937,1012,547,573]" name="Russia" pageId="48" pageNumber="49">Russia</collectingCountry>
(Siberia) substantially expands the geographic range, and the species may be present in other dry parts of Central Asia.
</paragraph>
<paragraph id="E93D36EAAC69434B0BD3FDA08B17048C" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
Separately,
<bibRefCitation id="8D134B1BAC69434B0A58FDA08F8704C4" author="Smith, F." box="[332,482,619,645]" pageId="48" pageNumber="49" refId="ref91052" refString="Smith, F. (1849) Descriptions of New Species of British Bees. Zoologist, 7, LVII - LX." type="journal volume" year="1849">Smith (1849)</bibRefCitation>
described
<taxonomicName id="2E824D69AC69434B0974FDA08C9804C5" authorityName="Smith" authorityYear="1849" box="[608,765,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="DBF6EAF8AC69434B0974FDA08C9804C5" box="[608,765,619,645]" italics="true" pageId="48" pageNumber="49">A. constrictus</emphasis>
</taxonomicName>
[recte
<taxonomicName id="2E824D69AC69434B085BFDA08D8504C5" authorityName="Smith" authorityYear="1849" box="[847,992,619,645]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constricta">
<emphasis id="DBF6EAF8AC69434B085BFDA08D8504C5" box="[847,992,619,645]" italics="true" pageId="48" pageNumber="49">A. constricta</emphasis>
</taxonomicName>
] from
<collectingCountry id="9195767AAC69434B0F26FDA08AFD04C4" box="[1074,1176,619,645]" name="United Kingdom" pageId="48" pageNumber="49">Scotland</collectingCountry>
. This species has been listed as a synonym of
<taxonomicName id="2E824D69AC69434B0A80FD448C6204E8" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[404,519,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0A80FD448C6204E8" box="[404,519,655,681]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
for a long time (e.g.
<bibRefCitation id="8D134B1BAC69434B09F9FD448DF704E8" author="Warncke, K." box="[749,914,655,681]" pageId="48" pageNumber="49" pagination="171 - 318" refId="ref91690" refString="Warncke, K. (1967) Beitrag zur Klarung palaarktischer Andrena - Arten. Eos, 43, 171 - 318." type="journal article" year="1967">Warncke 1967</bibRefCitation>
). However,
<taxonomicName id="2E824D69AC69434B0F03FD448AEC04E8" baseAuthorityName="Christ" baseAuthorityYear="1791" box="[1047,1161,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="fulvago">
<emphasis id="DBF6EAF8AC69434B0F03FD448AEC04E8" box="[1047,1161,655,681]" italics="true" pageId="48" pageNumber="49">A. fulvago</emphasis>
</taxonomicName>
has never been recorded from
<collectingCountry id="9195767AAC69434B0BC1FD788F52048C" box="[213,311,691,717]" name="United Kingdom" pageId="48" pageNumber="49">Scotland</collectingCountry>
(
<bibRefCitation id="8D134B1BAC69434B0A51FD788C24048C" author="Else, G. R. &amp; Edwards, M." box="[325,577,691,717]" pageId="48" pageNumber="49" refId="ref88553" refString="Else, G. R. &amp; Edwards, M. (2018) Handbook of the bees of the British Isles. The Ray Society, London, 775 pp." type="book" year="2018">Else &amp; Edwards 2018</bibRefCitation>
).
<typeStatus id="36398848AC69434B0947FD788CEE048C" box="[595,651,691,717]" pageId="48" pageNumber="49">Type</typeStatus>
inspection is necessary to clarify the true identity of
<taxonomicName id="2E824D69AC69434B0FC8FD788B0E048D" authorityName="Smith" authorityYear="1849" box="[1244,1387,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="constricta">
<emphasis id="DBF6EAF8AC69434B0FC8FD788B0E048D" box="[1244,1387,691,717]" italics="true" pageId="48" pageNumber="49">A. constricta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC69434B0BD3FD1D8AAD05E4" pageId="48" pageNumber="49" type="distribution">
<paragraph id="E93D36EAAC69434B0BD3FD1D8D540554" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
<emphasis id="DBF6EAF8AC69434B0BD3FD1D8F0404B1" bold="true" box="[199,353,726,752]" pageId="48" pageNumber="49">Distribution.</emphasis>
Palaearctic, from
<collectingCountry id="9195767AAC69434B093BFD1C8C1504B0" box="[559,624,727,753]" name="Georgia" pageId="48" pageNumber="49">Iberia</collectingCountry>
across Europe and northern
<collectingCountry id="9195767AAC69434B08A7FD1C8A6604B0" box="[947,1027,727,753]" name="Turkey" pageId="48" pageNumber="49">Turkey</collectingCountry>
to the Caucasus and Eastern Siberia (
<bibRefCitation id="8D134B1BAC69434B0B8BFD308F920554" author="Gusenleitner, F. &amp; Schwarz, M." box="[159,503,763,789]" pageId="48" pageNumber="49" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC69434B0916FD308D400554" author="Proshchalykin M. Y. &amp; Astafurova Y. V. &amp; Sidorov D. A." box="[514,805,763,789]" pageId="48" pageNumber="49" pagination="263 - 276" refId="ref90418" refString="Proshchalykin M. Y., Astafurova Y. V. &amp; Sidorov D. A. (2017) Family Andrenidae. In: Lelej, A. S., Proshchalykin, M. Y. &amp; Loktionov, V. M. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute Russian Academy of Sciences, Supplement 6, 263 - 276." type="journal article" year="2017">
Proshchalykin
<emphasis id="DBF6EAF8AC69434B09BDFD308CBF0554" box="[681,730,763,789]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
. 2017
</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAAC69434B0BD3FCD58AAD05E4" blockId="48.[151,1437,151,933]" pageId="48" pageNumber="49">
<emphasis id="DBF6EAF8AC69434B0BD3FCD58F990579" bold="true" box="[199,508,798,824]" pageId="48" pageNumber="49">Other material examined</emphasis>
(
<taxonomicName id="2E824D69AC69434B0904FCD48CD80578" authorityName="Tadauchi, Miyanaga &amp; Dawut" authorityYear="2005" box="[528,701,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="almas">
<emphasis id="DBF6EAF8AC69434B0904FCD48CD80578" box="[528,701,799,825]" italics="true" pageId="48" pageNumber="49">Andrena almas</emphasis>
</taxonomicName>
)
<materialsCitation id="59EA3CB7AC69434B09C5FCD58BFD051C" collectingDate="1962-08-07" collectorName="Baloutchah, W &amp; K. Lindberg &amp; No A &amp; Entomology Collection &amp; Lund Museum" country="Afghanistan" location="AFGHANISTAN" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1">
<emphasis id="DBF6EAF8AC69434B09C5FCD58DCD0579" bold="true" box="[721,936,798,824]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B09C5FCD58DC70579" box="[721,930,798,824]" name="Afghanistan" pageId="48" pageNumber="49">AFGHANISTAN</collectingCountry>
:
</emphasis>
<collectorName id="4477533CAC69434B08A6FCD48A380578" box="[946,1117,799,825]" pageId="48" pageNumber="49">Baloutchah, W</collectorName>
Tagab Echnan,
<date id="9D3C102AAC69434B0E08FCD48BFD0578" box="[1308,1432,799,825]" pageId="48" pageNumber="49" value="1962-08-07">
<collectingDate id="8D78E9C2AC69434B0E08FCD48BFD0578" box="[1308,1432,799,825]" pageId="48" pageNumber="49" value="1962-08-07">7.viii.1962</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B0B83FC888EDF051C" box="[151,186,835,861]" count="1" pageId="48" pageNumber="49" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B0A15FC888FF5051C" box="[257,400,835,861]" pageId="48" pageNumber="49">K. Lindberg</collectorName>
, OÖLM (
<collectorName id="4477533CAC69434B0910FC888C20051C" box="[516,581,835,861]" pageId="48" pageNumber="49">No A</collectorName>
1075, ex. coll.
<collectorName id="4477533CAC69434B0816FC888A75051C" box="[770,1040,835,861]" pageId="48" pageNumber="49">Entomology Collection</collectorName>
,
<collectorName id="4477533CAC69434B0F0AFC888AAE051C" box="[1054,1227,835,861]" pageId="48" pageNumber="49">Lund Museum</collectorName>
;
<figureCitation id="71B92A6FAC69434B0FCEFC888BF5051C" box="[1242,1424,835,861]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30AD</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC69434B0B83FCAD8AAC05C0" box="[151,1225,870,899]" collectingDate="1991-08-07" collectorName="M. Snizek" country="Kazakhstan" location="Alma-Ata" pageId="48" pageNumber="49" specimenCount="1" specimenCount-female="1" stateProvince="Almaty Oblysy">
<emphasis id="DBF6EAF8AC69434B0B83FCAD8F0305C1" bold="true" box="[151,358,870,896]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B0B83FCAD8F0405C1" box="[151,353,870,896]" name="Kazakhstan" pageId="48" pageNumber="49">KAZAKHSTAN</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC69434B0A7FFCAC8FBE05C0" LSID="urn:lsid:plazi:treatment:612B87FCAC77434B0B83FA948AAD05E4:EC5D6031AC69434B0A7FFCAC8FBE05C0" box="[363,475,871,897]" country="Kazakhstan" name="Alma-Ata" pageId="48" pageNumber="49" stateProvince="Almaty Oblysy">Alma-Ata</location>
[
<collectingRegion id="2B46F808AC69434B0AF3FCAC8C2605C0" box="[487,579,871,897]" country="Kazakhstan" name="Almaty Oblysy" pageId="48" pageNumber="49">Almaty</collectingRegion>
],
<date id="9D3C102AAC69434B0940FCAC8CB505C0" box="[596,720,871,897]" pageId="48" pageNumber="49" value="1991-08-07">
<collectingDate id="8D78E9C2AC69434B0940FCAC8CB505C0" box="[596,720,871,897]" pageId="48" pageNumber="49" value="1991-08-07">7.viii.1991</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B09CEFCAC8C9805C0" box="[730,765,871,897]" count="1" pageId="48" pageNumber="49" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B082CFCAC8DC805C0" box="[824,941,871,897]" pageId="48" pageNumber="49">M. Snižek</collectorName>
, OÖLM (
<figureCitation id="71B92A6FAC69434B0F02FCAC8AA405C0" box="[1046,1217,871,897]" captionStart="FIGURE 30" captionStartId="47.[151,250,1647,1671]" captionTargetBox="[151,1436,181,1624]" captionTargetId="figure-16@47.[151,1436,181,1624]" captionTargetPageId="47" captionText="FIGURE 30. Andrena (Chrysandrena) almas Tadauchi, Miyanaga &amp; Dawut, 2005 female from Kazakhstan.A. Habitus, lateral view; B. Scutum and head, dorsal view; C. Terga, dorsal view; D. Tibial scopae, lateral view; Female from Afghanistan. E. Habitus, lateral view; F. Face, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209373" httpUri="https://zenodo.org/record/13209373/files/figure.png" pageId="48" pageNumber="49">Figures 30EF</figureCitation>
)
</materialsCitation>
;
<materialsCitation id="59EA3CB7AC69434B0FC7FCAD8AA105E4" collectingDate="2010-08-03" collectionCode="NHMUK" collectorName="Michez &amp; DeMeule" country="Russia" elevation="282" location="Cherepanovo" municipality="Eastern Siberia" pageId="48" pageNumber="49" specimenCount="5" specimenCount-female="5">
<emphasis id="DBF6EAF8AC69434B0FC7FCAD8B2605C1" bold="true" box="[1235,1347,870,896]" pageId="48" pageNumber="49">
<collectingCountry id="9195767AAC69434B0FC7FCAD8B5B05C1" box="[1235,1342,870,896]" name="Russia" pageId="48" pageNumber="49">RUSSIA</collectingCountry>
:
</emphasis>
<collectingMunicipality id="0959AC90AC69434B0E5CFCAC8E8C05E4" pageId="48" pageNumber="49">Eastern Siberia</collectingMunicipality>
,
<location id="EC5D6031AC69434B0BE1FC408FE805E4" LSID="urn:lsid:plazi:treatment:612B87FCAC77434B0B83FA948AAD05E4:EC5D6031AC69434B0BE1FC408FE805E4" box="[245,397,907,933]" country="Russia" municipality="Eastern Siberia" name="Cherepanovo" pageId="48" pageNumber="49">Cherepanovo</location>
, train station,
<quantity id="2E7A9B0FAC69434B093FFC408C1105E4" box="[555,628,907,933]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="48" pageNumber="49" unit="m" value="282.0">
<elevation id="62AFD1D9AC69434B093FFC408C1105E4" box="[555,628,907,933]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="48" pageNumber="49" unit="m" value="282.0">282 m</elevation>
</quantity>
,
<date id="9D3C102AAC69434B0994FC408C9E05E4" box="[640,763,907,933]" pageId="48" pageNumber="49" value="2010-08-03">
<collectingDate id="8D78E9C2AC69434B0994FC408C9E05E4" box="[640,763,907,933]" pageId="48" pageNumber="49" value="2010-08-03">3.viii.2010</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC69434B0813FC418D4E05E4" box="[775,811,906,933]" count="5" pageId="48" pageNumber="49" type="female">5♀</specimenCount>
, leg.
<collectorName id="4477533CAC69434B087CFC408DD905E4" box="[872,956,907,933]" pageId="48" pageNumber="49">Michez</collectorName>
&amp;
<collectorName id="4477533CAC69434B08F4FC408A2E05E4" box="[992,1099,907,933]" pageId="48" pageNumber="49">DeMeule</collectorName>
,
<collectionCode id="8F93AE2FAC69434B0F42FC408AA105E4" box="[1110,1220,907,933]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="48" pageNumber="49" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,554 @@
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<subSubSection id="A1986561AC7B43590B83FE118A0A07B5" box="[151,1135,474,501]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="E93D36EAAC7B43590B83FE118A0A07B5" blockId="34.[151,1135,474,501]" box="[151,1135,474,501]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B43590B83FE118A0A07B5" bold="true" box="[151,1135,474,501]" pageId="34" pageNumber="35">
3) Revision of the
<emphasis id="DBF6EAF8AC7B43590A6CFE108F8D07B4" bold="true" box="[376,488,475,501]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
-group (within subgenus
<taxonomicName id="2E824D69AC7B43590834FE118A0207B5" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[800,1127,474,501]" class="Insecta" family="Andrenidae" genus="Ulandrena" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC7B43590834FE118DC707B4" bold="true" box="[800,930,474,501]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
Warncke, 1968
</taxonomicName>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7B435D0B83FDE88B640420" lastPageId="38" lastPageNumber="39" pageId="34" pageNumber="35" type="discussion">
<paragraph id="E93D36EAAC7B43590B83FDE88FED051C" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
The
<taxonomicName id="2E824D69AC7B43590BD3FDE88FF7047D" authorityName="Smith" authorityYear="1853" box="[199,402,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590BD3FDE88FF7047D" box="[199,402,547,573]" italics="true" pageId="34" pageNumber="35">Andrena concinna</emphasis>
</taxonomicName>
-group consists of a small number of very large-sized
<taxonomicName id="2E824D69AC7B435908CDFDE88A5D047C" baseAuthorityName="Popov" baseAuthorityYear="1960" box="[985,1080,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8AC7B435908CDFDE88A5D047C" box="[985,1080,547,573]" italics="true" pageId="34" pageNumber="35">Andrena</emphasis>
</taxonomicName>
within the subgenus
<taxonomicName id="2E824D69AC7B43590E09FDE88BF0047C" authorityName="Warncke" authorityYear="1968" box="[1309,1429,547,573]" pageId="34" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7B43590E09FDE88BF0047C" box="[1309,1429,547,573]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
</taxonomicName>
. These bees are instantly recognised within the subgenus by their huge body size, with specimens usually exceeding
<quantity id="2E7A9B0FAC7B43590B83FDA18E8F04C4" box="[151,234,618,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="mm" value="15.0">15 mm</quantity>
in length which makes them some of the largest
<taxonomicName id="2E824D69AC7B4359083BFDA08D8F04C4" box="[815,1002,619,645]" pageId="34" pageNumber="35">
<emphasis id="DBF6EAF8AC7B4359083BFDA08DEB04C4" box="[815,910,619,645]" italics="true" pageId="34" pageNumber="35">Andrena</emphasis>
species
</taxonomicName>
globally. The
<taxonomicName id="2E824D69AC7B43590F81FDA08A9904C5" authorityName="Smith" authorityYear="1853" box="[1173,1276,619,644]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F81FDA08A9904C5" box="[1173,1276,619,644]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
-group can be recognised further because the hind tarsal claws of the females are bidentate, presenting an inner subapical tooth. This separates them from the other large-bodied
<taxonomicName id="2E824D69AC7B435909D9FD788D20048C" authorityName="Warncke" authorityYear="1968" box="[717,837,691,717]" pageId="34" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7B435909D9FD788D20048C" box="[717,837,691,717]" italics="true" pageId="34" pageNumber="35">Ulandrena</emphasis>
</taxonomicName>
, specifically
<taxonomicName id="2E824D69AC7B435908F5FD788B72048C" authority="Popov, 1940" authorityName="Popov" authorityYear="1940" box="[993,1303,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="armeniaca">
<emphasis id="DBF6EAF8AC7B435908F5FD788A1E048D" box="[993,1147,691,717]" italics="true" pageId="34" pageNumber="35">A. armeniaca</emphasis>
Popov, 1940
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590E31FD788F5404B0" authority="Giraud, 1863" authorityName="Giraud" authorityYear="1863" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="DBF6EAF8AC7B43590E31FD788BF8048C" box="[1317,1437,691,717]" italics="true" pageId="34" pageNumber="35">A. elegans</emphasis>
Giraud, 1863
</taxonomicName>
, and
<taxonomicName id="2E824D69AC7B43590A66FD1C8C8504B0" authority="Morawitz, 1876" authorityName="Morawitz" authorityYear="1876" box="[370,736,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="DBF6EAF8AC7B43590A66FD1C8C4504B0" box="[370,544,727,753]" italics="true" pageId="34" pageNumber="35">A. fedtschenkoi</emphasis>
Morawitz, 1876
</taxonomicName>
which have the hind tarsal claws unidentate. The
<taxonomicName id="2E824D69AC7B43590E38FD1C8BF604B1" authorityName="Smith" authorityYear="1853" box="[1324,1427,727,752]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E38FD1C8BF604B1" box="[1324,1427,727,752]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
- group therefore nominally contains seven species:
<taxonomicName id="2E824D69AC7B435909D9FD308A620554" authority="Morawitz, 1877" authorityName="Morawitz" authorityYear="1877" box="[717,1031,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="DBF6EAF8AC7B435909D9FD308D2E0555" box="[717,843,763,789]" italics="true" pageId="34" pageNumber="35">A. carinata</emphasis>
<bibRefCitation id="8D134B1BAC7B43590845FD308A620554" author="Morawitz, F." box="[849,1031,763,789]" pageId="34" pageNumber="35" pagination="3 - 112" refId="ref89413" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14, 3 - 112." type="journal article" year="1877">Morawitz, 1877</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590F05FD308B4C0554" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[1041,1321,762,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F05FD308AF20555" box="[1041,1175,763,789]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F8AFD308B4C0554" author="Smith, F." box="[1182,1321,762,789]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590E20FD308F210578" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590E20FD308BF80555" box="[1332,1437,763,789]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
<bibRefCitation id="8D134B1BAC7B43590B83FCD48F210578" author="Warncke, K." box="[151,324,798,825]" pageId="34" pageNumber="35" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B43590A44FCD48CFA0578" authority="Grunwaldt, 2005" authorityName="Grunwaldt" authorityYear="2005" box="[336,671,798,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="heinrichi">
<emphasis id="DBF6EAF8AC7B43590A44FCD48FB00578" box="[336,469,799,825]" italics="true" pageId="34" pageNumber="35">A. heinrichi</emphasis>
<bibRefCitation id="8D134B1BAC7B43590ACAFCD48CFA0578" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[478,671,798,825]" pageId="34" pageNumber="35" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">Grünwaldt, 2005</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B435909BFFCD48DA80578" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" box="[683,973,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7B435909BFFCD48D520578" box="[683,823,799,825]" italics="true" pageId="34" pageNumber="35">A. paradoxa</emphasis>
Friese, 1921
</taxonomicName>
,
<taxonomicName id="2E824D69AC7B435908CEFCD48B6C0578" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[986,1289,799,825]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B435908CEFCD48A300578" box="[986,1109,799,825]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F48FCD48B6C0578" author="Warncke, K." box="[1116,1289,799,825]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="2E824D69AC7B43590E52FCD48FE1051C" authority="Radoszkowski, 1876" authorityName="Radoszkowski" authorityYear="1876" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7B43590E52FCD48BF90579" box="[1350,1436,799,825]" italics="true" pageId="34" pageNumber="35">A. tecta</emphasis>
Radoszkowski, 1876
</taxonomicName>
.
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FCAC8F7202A8" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
The oldest species name within this group is
<taxonomicName id="2E824D69AC7B435909C2FCAC8D3A05C1" authorityName="Smith" authorityYear="1853" box="[726,863,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B435909C2FCAC8D3A05C1" box="[726,863,871,897]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
which was described from
<collectingCountry id="9195767AAC7B43590FB4FCAC8A9905C0" box="[1184,1276,871,897]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
, and which is currently known only from Mediterranean
<collectingCountry id="9195767AAC7B4359096DFC408CB705E4" box="[633,722,907,933]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
and
<collectingCountry id="9195767AAC7B4359081EFC408D3805E4" box="[778,861,907,933]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
. The type series of
<taxonomicName id="2E824D69AC7B43590F2DFC408AA505E5" authorityName="Smith" authorityYear="1853" box="[1081,1216,907,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F2DFC408AA505E5" box="[1081,1216,907,933]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
could not be found in NHMUK which is listed as the type depository by
<bibRefCitation id="8D134B1BAC7B435909E6FC648A390588" author="Gusenleitner, F. &amp; Schwarz, M." box="[754,1116,943,969]" pageId="34" pageNumber="35" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
.
<bibRefCitation id="8D134B1BAC7B43590F73FC648B530588" author="Smith, F." box="[1127,1334,942,969]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith (1853: 106)</bibRefCitation>
lists that the type material comes from the collection of W.W. Saunders, but there is no entry in the type catalogue of the NHMUK. In contrast,
<bibRefCitation id="8D134B1BAC7B43590A83FC3C8C030250" author="Smith, F." box="[407,614,1014,1041]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith (1853: 107)</bibRefCitation>
does not list a collection for
<taxonomicName id="2E824D69AC7B435908A7FC3C8AB10250" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[947,1236,1014,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="DBF6EAF8AC7B435908A7FC3C8A5B0250" box="[947,1086,1015,1041]" italics="true" pageId="34" pageNumber="35">A. variabilis</emphasis>
<bibRefCitation id="8D134B1BAC7B43590F53FC3C8AB10250" author="Smith, F." box="[1095,1236,1014,1041]" pageId="34" pageNumber="35" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
, and this material can be found in the NHMUK type collection. This suggests that since Smith indicated a private collection for
<taxonomicName id="2E824D69AC7B43590E90FBD08E9B0219" authorityName="Smith" authorityYear="1853" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E90FBD08E9B0219" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, the material did not come from the NHMUK general collection and hence may be lost. However, it is not considered necessary to designate a
<typeStatus id="36398848AC7B4359097CFBA88CA4023C" box="[616,705,1123,1149]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
at this point, as the identity of this taxon is clear based on the description and also based on biogeography, since only one member of the
<taxonomicName id="2E824D69AC7B435908E6FB4C8A3C02E1" authorityName="Smith" authorityYear="1853" box="[1010,1113,1159,1184]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B435908E6FB4C8A3C02E1" box="[1010,1113,1159,1184]" italics="true" pageId="34" pageNumber="35">concinna</emphasis>
</taxonomicName>
-group is present in
<collectingCountry id="9195767AAC7B43590E29FB4C8BFC02E0" box="[1341,1433,1159,1185]" name="Albania" pageId="34" pageNumber="35">Albania</collectingCountry>
, i.e.
<taxonomicName id="2E824D69AC7B43590BD5FB608F220285" authorityName="Smith" authorityYear="1853" box="[193,327,1195,1221]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590BD5FB608F220285" box="[193,327,1195,1221]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
. Additional searches should be made in the NHMUK and OUMNH collections before a
<typeStatus id="36398848AC7B43590E3EFB608BE60284" box="[1322,1411,1195,1221]" pageId="34" pageNumber="35" type="neotype">neotype</typeStatus>
is designated.
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FB388CF50380" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
Warncke took a very broad approach to this species complex. As females are almost morphologically inseparable, differing most obviously in the strength of the tergal hairbands, he treated taxa as subspecies of
<taxonomicName id="2E824D69AC7B43590E90FADC8E9B0315" authorityName="Smith" authorityYear="1853" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590E90FADC8E9B0315" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, describing
<taxonomicName id="2E824D69AC7B43590A91FAF08C3F0315" authorityName="Warncke" authorityYear="1965" box="[389,602,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
<emphasis id="DBF6EAF8AC7B43590A91FAF08C3F0315" box="[389,602,1339,1365]" italics="true" pageId="34" pageNumber="35">A. concinna crecca</emphasis>
</taxonomicName>
from western
<collectingCountry id="9195767AAC7B435909ECFAF08D2D0314" box="[760,840,1339,1365]" name="Turkey" pageId="34" pageNumber="35">Turkey</collectingCountry>
(Brussa =
<collectingRegion id="2B46F808AC7B435908ABFAF08A610314" box="[959,1028,1339,1365]" country="Turkey" name="Bursa" pageId="34" pageNumber="35">Bursa</collectingRegion>
;
<figureCitation id="71B92A6FAC7B43590F1AFAF08A1F0314" box="[1038,1146,1339,1365]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="34" pageNumber="35">Figure 19</figureCitation>
) and
<taxonomicName id="2E824D69AC7B43590FA3FAF08BF90314" authorityName="Warncke" authorityYear="1973" box="[1207,1436,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="resoluta">
<emphasis id="DBF6EAF8AC7B43590FA3FAF08BF90314" box="[1207,1436,1339,1365]" italics="true" pageId="34" pageNumber="35">A. concinna resoluta</emphasis>
</taxonomicName>
from North
<collectingCountry id="9195767AAC7B43590A37FA948FFA0338" box="[291,415,1375,1401]" name="Macedonia" pageId="34" pageNumber="35">Macedonia</collectingCountry>
(Skopje) (
<bibRefCitation id="8D134B1BAC7B4359090FFA948CA10338" author="Warncke, K." box="[539,708,1374,1401]" pageId="34" pageNumber="35" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke 1965</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7B435909C7FA948D6A0338" author="Warncke, K." box="[723,783,1375,1401]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">1973</bibRefCitation>
). However, within this group, there are clear differences in the genital capsule, and
<bibRefCitation id="8D134B1BAC7B43590AD3FA488D4A03DC" author="Gusenleitner, F. &amp; Schwarz, M." box="[455,815,1411,1437]" pageId="34" pageNumber="35" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz (2002)</bibRefCitation>
recognised six of these taxa as species, with a seventh described soon after (
<bibRefCitation id="8D134B1BAC7B43590A93FA6C8CE10380" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[391,644,1446,1473]" pageId="34" pageNumber="35" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">
Grünwaldt
<emphasis id="DBF6EAF8AC7B4359091CFA6C8C5C0380" box="[520,569,1447,1473]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2005
</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAAC7B43590BD3FA008CB50130" blockId="34.[151,1437,547,1905]" pageId="34" pageNumber="35">
Both
<taxonomicName id="2E824D69AC7B43590A12FA008FE703A4" authorityName="Warncke" authorityYear="1973" box="[262,386,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B43590A12FA008FE703A4" box="[262,386,1483,1509]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7B43590AAFFA008C4303A5" authorityName="Warncke" authorityYear="1965" box="[443,550,1483,1509]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590AAFFA008C4303A5" box="[443,550,1483,1509]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
have been recognised as distinct species, but the differences between the two are obscure. Moreover, the distributional ranges are unclear;
<taxonomicName id="2E824D69AC7B4359085FFA248DA30048" authorityName="Warncke" authorityYear="1973" box="[843,966,1519,1545]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B4359085FFA248DA30048" box="[843,966,1519,1545]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
was considered a species of the southern Balkans around North
<collectingCountry id="9195767AAC7B43590AB4F9D88C79006C" box="[416,540,1555,1581]" name="Macedonia" pageId="34" pageNumber="35">Macedonia</collectingCountry>
and south-western
<collectingCountry id="9195767AAC7B43590816F9D88D07006C" box="[770,866,1555,1581]" name="Bulgaria" pageId="34" pageNumber="35">Bulgaria</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7B43590860F9D88A7A006C" author="Warncke, K." box="[884,1055,1555,1581]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke 1973</bibRefCitation>
), but specimens can be found in south-eastern
<collectingCountry id="9195767AAC7B43590A22F9FC8FF30010" box="[310,406,1591,1617]" name="Bulgaria" pageId="34" pageNumber="35">Bulgaria</collectingCountry>
close to the Turkish border (see below) and hence not far from Bursa, and specimens of
<taxonomicName id="2E824D69AC7B43590B83F9908F640035" authorityName="Warncke" authorityYear="1965" box="[151,257,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590B83F9908F640035" box="[151,257,1627,1653]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
have been reported from the East
<collectingRegion id="2B46F808AC7B4359099CF9908CBB0034" box="[648,734,1627,1653]" country="Greece" name="Voreio Aigaio" pageId="34" pageNumber="35">Aegean</collectingRegion>
Islands of
<collectingCountry id="9195767AAC7B43590849F9908DC90034" box="[861,940,1627,1653]" name="Greece" pageId="34" pageNumber="35">Greece</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7B435908A8F9908ACD0034" author="Rasmont, P. &amp; Devalez, J. &amp; Pauly, A. &amp; Michez, D. &amp; Radchenko, V." box="[956,1192,1627,1653]" pageId="34" pageNumber="35" pagination="17 - 32" refId="ref90635" refString="Rasmont, P., Devalez, J., Pauly, A., Michez, D. &amp; Radchenko, V. (2017) Addition to the checklist of IUCN European wild bees (Hymenoptera: Apoidea). Annales de la Societe entomologique de France, New Series, 53, 17 - 32. https: // doi. org / 10.1080 / 00379271.2017.1307696" type="journal article" year="2017">
Rasmont
<emphasis id="DBF6EAF8AC7B43590F3DF9908A390034" box="[1065,1116,1627,1653]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2017
</bibRefCitation>
).
<bibRefCitation id="8D134B1BAC7B43590FAFF9908BF90034" author="Warncke, K." box="[1211,1436,1627,1653]" pageId="34" pageNumber="35" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke (1973: 28)</bibRefCitation>
gives small differences between
<taxonomicName id="2E824D69AC7B4359091EF9B48CE000D8" authorityName="Warncke" authorityYear="1973" box="[522,645,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B4359091EF9B48CE000D8" box="[522,645,1663,1689]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
and what can be assumed is the nominate
<taxonomicName id="2E824D69AC7B43590F7AF9B48A9000D9" authorityName="Smith" authorityYear="1853" box="[1134,1269,1663,1689]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7B43590F7AF9B48A9000D9" box="[1134,1269,1663,1689]" italics="true" pageId="34" pageNumber="35">A. concinna</emphasis>
</taxonomicName>
, since he does not specify the comparison taxon until mentioning
<taxonomicName id="2E824D69AC7B435909CEF9688D3200FD" authorityName="Warncke" authorityYear="1965" box="[730,855,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
ssp.
<emphasis id="DBF6EAF8AC7B43590819F9688D3200FD" box="[781,855,1699,1724]" italics="true" pageId="34" pageNumber="35">crecca</emphasis>
</taxonomicName>
, where he argues that the gonocoxae of
<taxonomicName id="2E824D69AC7B43590E35F9688BF900FC" authorityName="Warncke" authorityYear="1973" box="[1313,1436,1699,1725]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7B43590E35F9688BF900FC" box="[1313,1436,1699,1725]" italics="true" pageId="34" pageNumber="35">A. resoluta</emphasis>
</taxonomicName>
have a wide and sharply developed longitudinal ridge on their inner margin, whereas in
<taxonomicName id="2E824D69AC7B43590F75F90C8AAC00A1" authorityName="Warncke" authorityYear="1965" box="[1121,1225,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7B43590F75F90C8AAC00A1" box="[1121,1225,1735,1761]" italics="true" pageId="34" pageNumber="35">A. crecca</emphasis>
</taxonomicName>
the transition from the gonocoxae to the gonostyli is smooth [“
<emphasis id="DBF6EAF8AC7B43590986F9208D140169" italics="true" pageId="34" pageNumber="35">Gonocoxiten besitzen auf der Innenseite je eine scharf ausgebildete und breite, längsverlaufende Leiste (bei der östlichen Unterart</emphasis>
<taxonomicName id="2E824D69AC7B4359086FF8C48A000168" authority="Warncke" authorityName="Warncke" authorityYear="1965" box="[891,1125,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subSpecies" species="concinna" subSpecies="crecca">
<emphasis id="DBF6EAF8AC7B4359086FF8C48DC20169" box="[891,935,1807,1832]" italics="true" pageId="34" pageNumber="35">ssp.</emphasis>
crecca
<emphasis id="DBF6EAF8AC7B43590F10F8C48A000168" box="[1028,1125,1807,1833]" italics="true" pageId="34" pageNumber="35">Warncke</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC7B43590F7BF8C48CD70130" italics="true" pageId="34" pageNumber="35">ist der Übergang von den Gonocoxiten zu den Gonostylen gleichmässig, und der etwas höhere Penisaufsatz ist nach hinten zu noch starker verlängert, die Penisspitze nach unten gebogen)</emphasis>
”].
</paragraph>
<caption id="BDFD6662AC7A43580B83F85C8CCD01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209349" ID-Zenodo-Dep="13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1943,1967]" targetBox="[214,1374,181,1919]" targetPageId="35" targetType="figure">
<paragraph id="E93D36EAAC7A43580B83F85C8CCD01B6" blockId="35.[151,1437,1943,2040]" pageId="35" pageNumber="36">
<emphasis id="DBF6EAF8AC7A43580B83F85C8F4401EE" bold="true" box="[151,289,1943,1967]" pageId="35" pageNumber="36">FIGURE 19.</emphasis>
<taxonomicName id="2E824D69AC7A43580A3CF85C8C8801EE" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[296,749,1943,1968]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="crecca" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7A43580A3CF85C8F1B01EE" box="[296,382,1943,1967]" italics="true" pageId="35" pageNumber="36">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7A43580A9AF85C8F9E01EE" box="[398,507,1943,1967]" italics="true" pageId="35" pageNumber="36">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7A4358091EF8538C2801EE" box="[522,589,1944,1967]" italics="true" pageId="35" pageNumber="36">crecca</emphasis>
<bibRefCitation id="8D134B1BAC7A43580940F85C8C8801EE" author="Warncke, K." box="[596,749,1943,1967]" pageId="35" pageNumber="36" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC7A435809E0F8538D3401F1" bold="true" box="[756,849,1944,1968]" pageId="35" pageNumber="36">holotype</emphasis>
male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail.
</paragraph>
</caption>
<paragraph id="E93D36EAAC7D435F0BD3FF5C8A5307B4" blockId="36.[151,1437,151,501]" pageId="36" pageNumber="37">
It is not clear what Warncke meant by this, because examination of the respective type material shows that the
<typeStatus id="36398848AC7D435F0B83FF708E9C0694" box="[151,249,187,213]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC7D435F0A35FF708FEE0695" authorityName="Warncke" authorityYear="1965" box="[289,395,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F0A35FF708FEE0695" box="[289,395,187,213]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
does have a developed longitudinal ridge on the inner margins of each gonocoxite (
<figureCitation id="71B92A6FAC7D435F0E52FF708E8E06B8" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figures 19F; H</figureCitation>
). In the
<typeStatus id="36398848AC7D435F0A5EFF148FC906B8" box="[330,428,223,249]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
itself, it is not so strongly developed as in type material of
<taxonomicName id="2E824D69AC7D435F0F42FF148AB406B8" authorityName="Warncke" authorityYear="1973" box="[1110,1233,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F0F42FF148AB406B8" box="[1110,1233,223,249]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
(
<figureCitation id="71B92A6FAC7D435F0FF5FF148B0606B8" box="[1249,1379,223,249]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37">Figure 20F</figureCitation>
), but examination of material from across
<collectingCountry id="9195767AAC7D435F0923FEC88CE2075C" box="[567,647,259,285]" name="Turkey" pageId="36" pageNumber="37">Turkey</collectingCountry>
shows that this is a little variable, and many individuals can be found that have the internal ridges as strongly produced as
<taxonomicName id="2E824D69AC7D435F09F3FEEC8D040700" authorityName="Warncke" authorityYear="1973" box="[743,865,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F09F3FEEC8D040700" box="[743,865,295,321]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
. Unfortunately, the penis valves of the
<typeStatus id="36398848AC7D435F0E08FEEC8B1B0700" box="[1308,1406,295,321]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
of
<taxonomicName id="2E824D69AC7D435F0B83FE808F650725" authorityName="Warncke" authorityYear="1965" box="[151,256,331,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F0B83FE808F650725" box="[151,256,331,357]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
are not in a normal position (
<figureCitation id="71B92A6FAC7D435F095CFE808C920724" box="[584,759,331,357]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figures 19FG</figureCitation>
), but the apex of the penis valves can be seen where they are straight (
<figureCitation id="71B92A6FAC7D435F0BE8FEA48FE607C8" box="[252,387,367,393]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="36" pageNumber="37">Figure 19G</figureCitation>
), whereas in
<taxonomicName id="2E824D69AC7D435F0908FEA48CFD07C8" authorityName="Warncke" authorityYear="1973" box="[540,664,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta">
<emphasis id="DBF6EAF8AC7D435F0908FEA48CFD07C8" box="[540,664,367,393]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
the apex is slightly upturned (
<figureCitation id="71B92A6FAC7D435F08E2FEA48A1E07C8" box="[1014,1147,367,393]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37">Figure 20E</figureCitation>
). However, intermediates can be found between these two states, for example from Malko Tarnovo in south-eastern
<collectingCountry id="9195767AAC7D435F0F86FE588A9307EC" box="[1170,1270,403,429]" name="Bulgaria" pageId="36" pageNumber="37">Bulgaria</collectingCountry>
. In the context of examination of material from the southern Balkans to the Caucasus, these differences are considered to be both small and variable, and hence
<taxonomicName id="2E824D69AC7D435F0AFEFE108C0107B4" authorityName="Warncke" authorityYear="1973" box="[490,612,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta" status="syn. nov.">
<emphasis id="DBF6EAF8AC7D435F0AFEFE108C0107B4" box="[490,612,475,501]" italics="true" pageId="36" pageNumber="37">A. resoluta</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC7D435F097FFE108CB607B4" bold="true" box="[619,723,475,501]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="C0C55783AC7D435F097FFE108CB607B4" box="[619,723,475,501]" pageId="36" pageNumber="37">syn. nov.</taxonomicNameLabel>
</emphasis>
is synonymised with
<taxonomicName id="2E824D69AC7D435F08D3FE108A5507B5" authorityName="Warncke" authorityYear="1965" box="[967,1072,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7D435F08D3FE108A5507B5" box="[967,1072,475,501]" italics="true" pageId="36" pageNumber="37">A. crecca</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="BDFD6662AC7D435F0B83F87C8AEB01B2" ID-DOI="http://doi.org/10.5281/zenodo.13209351" ID-Zenodo-Dep="13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1975,1999]" targetBox="[151,1436,510,1960]" targetPageId="36" targetType="figure">
<paragraph id="E93D36EAAC7D435F0B83F87C8AEB01B2" blockId="36.[151,1436,1975,2036]" pageId="36" pageNumber="37">
<emphasis id="DBF6EAF8AC7D435F0B83F87C8F45018E" bold="true" box="[151,288,1975,1999]" pageId="36" pageNumber="37">FIGURE 20.</emphasis>
<taxonomicName id="2E824D69AC7D435F0A31F87C8C930191" authority="Warncke, 1973" authorityName="Warncke" authorityYear="1973" box="[293,758,1975,2000]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="resoluta" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7D435F0A31F87C8F1E018E" box="[293,379,1975,1999]" italics="true" pageId="36" pageNumber="37">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7D435F0A9EF87C8F92018E" box="[394,503,1975,1999]" italics="true" pageId="36" pageNumber="37">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7D435F0912F87C8C3D018E" box="[518,600,1975,1999]" italics="true" pageId="36" pageNumber="37">resoluta</emphasis>
<bibRefCitation id="8D134B1BAC7D435F0949F87C8C930191" author="Warncke, K." box="[605,758,1975,2000]" pageId="36" pageNumber="37" pagination="13 - 36" refId="ref91713" refString="Warncke, K. (1973) Beitrag zur Bienenfauna Mazedoniens (Colletidae, Andrenidae und Melittidae / Apoidea). Mitteilungen aus dem Zoologischen Museum in Berlin, 49, 13 - 36. https: // doi. org / 10.1002 / mmnz. 4830490103" type="journal article" year="1973">Warncke, 1973</bibRefCitation>
</taxonomicName>
<emphasis id="DBF6EAF8AC7D435F09E8F8738D380191" bold="true" box="[764,861,1976,2000]" pageId="36" pageNumber="37">paratype</emphasis>
male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<caption id="BDFD6662AC7C435E0B83F8BF8B7201B9" ID-DOI="http://doi.org/10.5281/zenodo.13209355" ID-Zenodo-Dep="13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,243,1908,1932]" targetBox="[214,1373,161,1792]" targetPageId="37" targetType="figure">
<paragraph id="E93D36EAAC7C435E0B83F8BF8B7201B9" blockId="37.[151,1437,1908,2040]" pageId="37" pageNumber="38">
<emphasis id="DBF6EAF8AC7C435E0B83F8BF8F7001CD" bold="true" box="[151,277,1908,1932]" pageId="37" pageNumber="38">FIGURE 21.</emphasis>
<taxonomicName id="2E824D69AC7C435E0A0DF8BF8CD001CD" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[281,693,1908,1932]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="concinna" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0A0DF8BF8F0C01CD" box="[281,361,1908,1932]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0A62F8BF8FBE01CD" box="[374,475,1908,1932]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0AFCF8BE8C5B01CD" box="[488,574,1909,1932]" italics="true" pageId="37" pageNumber="38">concinna</emphasis>
<bibRefCitation id="8D134B1BAC7C435E0957F8BF8CD001CD" author="Smith, F." box="[579,693,1908,1932]" pageId="37" pageNumber="38" refId="ref91075" refString="Smith, F. (1853) Catalogue of Hymenopterous insects in the collection of the British Museum. Andrenidae and Apidae. London, 197 pp." type="book" year="1853">Smith, 1853</bibRefCitation>
</taxonomicName>
male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E0E58F8BF8F8701F1" authority="Friese, 1921" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="paradoxa" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0E58F8BF8BF901CD" box="[1356,1436,1908,1932]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0B8BF8538F6101F1" box="[159,260,1944,1968]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0A05F8538F0E01F1" box="[273,363,1944,1968]" italics="true" pageId="37" pageNumber="38">paradoxa</emphasis>
Friese, 1921
</taxonomicName>
male. B. Genital capsule, lateral view; D. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E0F6DF8538F520195" authority="Grunwaldt, 2005" authorityName="Grunwaldt" authorityYear="2005" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="heinrichi" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0F6DF8538AAC01F1" box="[1145,1225,1944,1968]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0FC2F8538B5E01F1" box="[1238,1339,1944,1968]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0E53F8538BF901F1" box="[1351,1436,1944,1968]" italics="true" pageId="37" pageNumber="38">heinrichi</emphasis>
<bibRefCitation id="8D134B1BAC7C435E0B83F8778F520195" author="Grunwaldt, W. &amp; Osytshnjuk, A. Z. &amp; Scheuchl, E." box="[151,311,1980,2004]" pageId="37" pageNumber="38" pagination="349 - 368" refId="ref88854" refString="Grunwaldt, W., Osytshnjuk, A. Z. &amp; Scheuchl, E. (2005) Neue Andrena - Arten aus der Palaarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26, 349 - 368." type="journal article" year="2005">Grünwaldt, 2005</bibRefCitation>
</taxonomicName>
male. E. Genital capsule, dorsal view;
<taxonomicName id="2E824D69AC7C435E09B3F8778A0A0195" authority="Radoszkowski, 1876" authorityName="Radoszkowski" authorityYear="1876" box="[679,1135,1980,2004]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="tecta" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E09B3F8778C920195" box="[679,759,1980,2004]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0811F8778D0F0195" box="[773,874,1980,2004]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E0863F8768DC00195" box="[887,933,1981,2004]" italics="true" pageId="37" pageNumber="38">tecta</emphasis>
Radoszkowski, 1876
</taxonomicName>
male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view;
<taxonomicName id="2E824D69AC7C435E0913F82B8DCC01B9" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[519,937,2016,2040]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="crecca" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7C435E0913F82B8C3201B9" box="[519,599,2016,2040]" italics="true" pageId="37" pageNumber="38">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC7C435E0971F82B8CAF01B9" box="[613,714,2016,2040]" italics="true" pageId="37" pageNumber="38">Ulandrena</emphasis>
)
<emphasis id="DBF6EAF8AC7C435E09CCF82A8D7301B9" box="[728,790,2017,2040]" italics="true" pageId="37" pageNumber="38">crecca</emphasis>
<bibRefCitation id="8D134B1BAC7C435E080FF82B8DCC01B9" author="Warncke, K." box="[795,937,2016,2040]" pageId="37" pageNumber="38" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
male. H. Genital capsule, lateral view.
</paragraph>
</caption>
<paragraph id="E93D36EAAC7F435D0BD3FF5C8CB507B4" blockId="38.[151,1437,151,609]" pageId="38" pageNumber="39">
Searches in the ZMHB were unable to find material of
<taxonomicName id="2E824D69AC7F435D0845FF5C8DCC06F1" authorityName="Radoszkowski" authorityYear="1876" box="[849,937,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0845FF5C8DCC06F1" box="[849,937,151,177]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
, the listed
<typeStatus id="36398848AC7F435D0F3AFF5C8A3B06F0" box="[1070,1118,151,177]" pageId="38" pageNumber="39">type</typeStatus>
repository (
<bibRefCitation id="8D134B1BAC7F435D0FFFFF5C8F580694" author="Gusenleitner, F. &amp; Schwarz, M." pageId="38" pageNumber="39" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
). In the relevant box, there is a pinned note indicating that
<typeStatus id="36398848AC7F435D08CBFF708A6A0694" box="[991,1039,187,213]" pageId="38" pageNumber="39">type</typeStatus>
material from both
<taxonomicName id="2E824D69AC7F435D0FE0FF708B2E0695" authorityName="Radoszkowski" authorityYear="1876" box="[1268,1355,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0FE0FF708B2E0695" box="[1268,1355,187,213]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC7F435D0E90FF708F6706B8" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7F435D0E90FF708F6706B8" italics="true" pageId="38" pageNumber="39">A. paradoxa</emphasis>
</taxonomicName>
have been on loan to Robert Wilhelm Grünwaldt (1909†2003) since 1977. Since Grünwaldts collection was donated to the ZSM before his death, the material is likely to be there. No taxonomic action is therefore taken regarding
<taxonomicName id="2E824D69AC7F435D0A1FFEEC8F070701" authorityName="Radoszkowski" authorityYear="1876" box="[267,354,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0A1FFEEC8F070701" box="[267,354,295,321]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
. Finally,
<taxonomicName id="2E824D69AC7F435D0ADBFEEC8C2B0701" authorityName="Morawitz" authorityYear="1877" box="[463,590,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="DBF6EAF8AC7F435D0ADBFEEC8C2B0701" box="[463,590,295,321]" italics="true" pageId="38" pageNumber="39">A. carinata</emphasis>
</taxonomicName>
was described in the male sex only from modern day
<collectingCountry id="9195767AAC7F435D0FAFFEEC8B780700" box="[1211,1309,295,321]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
(
<bibRefCitation id="8D134B1BAC7F435D0E39FEEC8EB70724" author="Morawitz, F." pageId="38" pageNumber="39" pagination="3 - 112" refId="ref89413" refString="Morawitz, F. (1877) Nachtrag zur Bienenfauna Caucasiens. Horae Societatis Entomologicae Rossicae, 14, 3 - 112." type="journal article" year="1877">Morawitz 1877</bibRefCitation>
;
<bibRefCitation id="8D134B1BAC7F435D0BCAFE808FFA0724" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[222,415,331,357]" pageId="38" pageNumber="39" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">Astafurova 2021</bibRefCitation>
). Although it can confidently be placed into the group of large-bodied
<taxonomicName id="2E824D69AC7F435D0FDEFE808B270724" authorityName="Warncke" authorityYear="1968" box="[1226,1346,331,357]" pageId="38" pageNumber="35" rank="subGenus" subGenus="Ulandrena">
<emphasis id="DBF6EAF8AC7F435D0FDEFE808B270724" box="[1226,1346,331,357]" italics="true" pageId="38" pageNumber="39">Ulandrena</emphasis>
</taxonomicName>
species based on its declared size (16.5 mm) and declared morphological similarity to
<taxonomicName id="2E824D69AC7F435D0F0BFEA48AA907C8" authorityName="Morawitz" authorityYear="1876" box="[1055,1228,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="fedtschenkoi">
<emphasis id="DBF6EAF8AC7F435D0F0BFEA48AA907C8" box="[1055,1228,367,393]" italics="true" pageId="38" pageNumber="39">A. fedtschenkoi</emphasis>
</taxonomicName>
, it lacks a precise description or illustration of the genital capsule. As such, it could be a junior synonym of either
<taxonomicName id="2E824D69AC7F435D0FDDFE588B7A07ED" authorityName="Radoszkowski" authorityYear="1876" box="[1225,1311,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0FDDFE588B7A07ED" box="[1225,1311,403,429]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
or a senior synonym of
<taxonomicName id="2E824D69AC7F435D0A35FE7C8FEF0791" authorityName="Warncke" authorityYear="1965" box="[289,394,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7F435D0A35FE7C8FEF0791" box="[289,394,439,465]" italics="true" pageId="38" pageNumber="39">A. crecca</emphasis>
</taxonomicName>
since both taxa occur in the Caucasus, but as the
<typeStatus id="36398848AC7F435D08BFFE7C8DBE0790" box="[939,987,439,465]" pageId="38" pageNumber="39">type</typeStatus>
material is lost (
<bibRefCitation id="8D134B1BAC7F435D0F86FE7C8BF50790" author="Astafurova, Y. A. &amp; Proshchalykin, M. Y. &amp; Sidorov, D. A. &amp; Osytshnjuk, A. Z." box="[1170,1424,439,465]" pageId="38" pageNumber="39" pagination="1 - 78" refId="ref87243" refString="Astafurova, Y. A., Proshchalykin, M. Y., Sidorov, D. A. &amp; Osytshnjuk, A. Z. (2021) The type specimens of bees (Hymenoptera, Apoidea) deposited in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg. Contribution IV. Family Andrenidae, genus Andrena Fabricius, 1775, species described by F. Morawitz. Zootaxa, 5037 (1), 1 - 78. https: // doi. org / 10.11646 / zootaxa. 5037.1.1" type="journal article" year="2021">
Astafurova
<emphasis id="DBF6EAF8AC7F435D0E03FE7C8B220790" box="[1303,1351,439,465]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2021
</bibRefCitation>
), it is better to officially declare it a
<emphasis id="DBF6EAF8AC7F435D090EFE108CB507B4" bold="true" box="[538,720,474,501]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D090EFE108CAC07B4" bold="true" box="[538,713,474,501]" italics="true" pageId="38" pageNumber="39">nomen dubium</emphasis>
.
</emphasis>
</paragraph>
<paragraph id="E93D36EAAC7F435D0BD3FE348B640420" blockId="38.[151,1437,151,609]" pageId="38" pageNumber="39">
The
<taxonomicName id="2E824D69AC7F435D0BE8FE348F060459" authorityName="Smith" authorityYear="1853" box="[252,355,511,536]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D0BE8FE348F060459" box="[252,355,511,536]" italics="true" pageId="38" pageNumber="39">concinna</emphasis>
</taxonomicName>
-group is therefore reduced to five species,
<taxonomicName id="2E824D69AC7F435D084EFE348D860459" authorityName="Smith" authorityYear="1853" box="[858,995,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D084EFE348D860459" box="[858,995,511,537]" italics="true" pageId="38" pageNumber="39">A. concinna</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D08E0FE348A340458" box="[1012,1105,511,537]" name="Albania" pageId="38" pageNumber="39">Albania</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0F4BFE348AD60458" box="[1119,1203,511,537]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D0FDCFE348B560459" authorityName="Warncke" authorityYear="1965" box="[1224,1331,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7F435D0FDCFE348B560459" box="[1224,1331,511,537]" italics="true" pageId="38" pageNumber="39">A. crecca</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D0E50FE348BFD0458" box="[1348,1432,511,537]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0B83FDE88E81047C" box="[151,228,547,573]" name="Serbia" pageId="38" pageNumber="39">Serbia</collectingCountry>
*, North
<collectingCountry id="9195767AAC7F435D0A58FDE88FAE047C" box="[332,459,547,573]" name="Macedonia" pageId="38" pageNumber="39">Macedonia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0ACFFDE88C5B047C" box="[475,574,547,573]" name="Bulgaria" pageId="38" pageNumber="39">Bulgaria</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D095AFDE88CFA047C" box="[590,671,547,573]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D09B9FDE88D71047C" box="[685,788,547,573]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D083FFDE88DD1047C" authorityName="Grunwaldt" authorityYear="2005" box="[811,948,547,573]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="heinrichi">
<emphasis id="DBF6EAF8AC7F435D083FFDE88DD1047C" box="[811,948,547,573]" italics="true" pageId="38" pageNumber="39">A. heinrichi</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D08D3FDE88A79047C" box="[967,1052,547,573]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
: Samos and western
<collectingCountry id="9195767AAC7F435D0E0CFDE88B09047C" box="[1304,1388,547,573]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
),
<taxonomicName id="2E824D69AC7F435D0E90FDE88F670420" authorityName="Friese" authorityYear="1921" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7F435D0E90FDE88F670420" italics="true" pageId="38" pageNumber="39">A. paradoxa</emphasis>
</taxonomicName>
(southern
<collectingCountry id="9195767AAC7F435D0A6EFD8C8FAE0420" box="[378,459,583,609]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0AC2FD8C8C710420" box="[470,532,583,609]" name="Syria" pageId="38" pageNumber="39">Syria</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D090BFD8C8C040420" box="[543,609,583,609]" name="Israel" pageId="38" pageNumber="39">Israel</collectingCountry>
), and
<taxonomicName id="2E824D69AC7F435D09B0FD8C8C9C0421" authorityName="Radoszkowski" authorityYear="1876" box="[676,761,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D09B0FD8C8C9C0421" box="[676,761,583,609]" italics="true" pageId="38" pageNumber="39">A. tecta</emphasis>
</taxonomicName>
(
<collectingCountry id="9195767AAC7F435D081CFD8C8D3E0420" box="[776,859,583,609]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0872FD8C8DA60420" box="[870,963,583,609]" name="Georgia" pageId="38" pageNumber="39">Georgia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D08D8FD8C8A540420" box="[972,1073,583,609]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0F2EFD8C8ADD0420" box="[1082,1208,583,609]" name="Azerbaijan" pageId="38" pageNumber="39">Azerbaijan</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0FD7FD8C8A900420" box="[1219,1269,583,609]" name="Iran" pageId="38" pageNumber="39">Iran</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,196 @@
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<subSubSection id="A1986561AC7F435D0B83FD788C60048C" box="[151,517,691,717]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="E93D36EAAC7F435D0B83FD788C60048C" blockId="38.[151,517,691,717]" box="[151,517,691,717]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0B83FD788C60048C" bold="true" box="[151,517,691,717]" pageId="38" pageNumber="39">
Key to
<taxonomicName id="2E824D69AC7F435D0BE5FD788F04048C" ID-CoL="DRD7" authority="Smith, 1853" authorityName="Smith" authorityYear="1853" box="[241,353,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D0BE5FD788F04048C" bold="true" box="[241,353,691,717]" italics="true" pageId="38" pageNumber="39">concinna</emphasis>
</taxonomicName>
-group males
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC7F435D0B83FD338BF8035C" pageId="38" pageNumber="39" type="key">
<key id="9D1361DBAC7F435D0B83FD338BF8035C" pageId="38" pageNumber="39">
<keyStep id="52762E4FAC7F435D0B83FD338BF90524" pageId="38" pageNumber="39">
<paragraph id="E93D36EAAC7F435D0B83FD338BF9056B" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FD338BF9056B" pageId="38" pageNumber="39">
1. Genital capsule with gonocoxae presenting a strong longitudinal ridge along their inner margins, the apex of this ridge forming a rounded 60
<superScript id="1EF79BA2AC7F435D0A4BFCDF8F030561" attach="none" box="[351,358,788,800]" fontSize="5" pageId="38" pageNumber="39">o</superScript>
80
<superScript id="1EF79BA2AC7F435D0A9EFCDF8FF40561" attach="left" box="[394,401,788,800]" fontSize="5" pageId="38" pageNumber="39">o</superScript>
angle (
<figureCitation id="71B92A6FAC7F435D0ACCFCDE8C08056B" box="[472,621,789,811]" captionStart="FIGURE 19" captionStartId="35.[151,250,1943,1967]" captionTargetBox="[214,1374,181,1919]" captionTargetId="figure-16@35.[214,1374,181,1919]" captionTargetPageId="35" captionText="FIGURE 19. Andrena (Ulandrena) crecca Warncke, 1965 holotype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Scutum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view; G. Genital capsule, lateral view; H. Genital capsule, inner margins of gonocoxae, detail." figureDoi="http://doi.org/10.5281/zenodo.13209349" httpUri="https://zenodo.org/record/13209349/files/figure.png" pageId="38" pageNumber="39">Figures 19F; H</figureCitation>
;
<figureCitation id="71B92A6FAC7F435D0963FCDE8CDC056B" box="[631,697,789,810]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="38" pageNumber="39">20EF</figureCitation>
)........................................................... 2
</keyLead>
</paragraph>
<paragraph id="E93D36EAAC7F435D0B83FCF88BF90524" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FCF88BF90524" pageId="38" pageNumber="39">
- Genital capsule with gonocoxae presenting a smooth internal margin, without any sign of a longitudinal ridge or carina (
<figureCitation id="71B92A6FAC7F435D0E47FCF98F4D0524" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figures 21CD</figureCitation>
) ............................................................................................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="52762E4FAC7F435D0B83FCA68BF905B6" pageId="38" pageNumber="39">
<paragraph id="E93D36EAAC7F435D0B83FCA68BF905FC" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FCA68BF905FC" pageId="38" pageNumber="39">
2. Penis valves bulbous, grossly expanded and filling almost the entire space between the gonostyli (
<figureCitation id="71B92A6FAC7F435D0F8AFCA68B6A05C3" box="[1182,1295,877,899]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21E</figureCitation>
). Restricted to
<collectingCountry id="9195767AAC7F435D0BF6FC418F4305DE" box="[226,294,906,927]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
(island of Samos) and a small part of western
<collectingCountry id="9195767AAC7F435D09F0FC418D4D05E1" box="[740,808,906,928]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
on the Aegean coast only (around Kuşadası)..................................................................................................
<taxonomicName id="2E824D69AC7F435D0FDEFC6C8BF905FC" ID-CoL="DRTX" authority="Grunwaldt" authorityName="Grunwaldt" authorityYear="2005" box="[1226,1436,935,957]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="heinrichi">
<emphasis id="DBF6EAF8AC7F435D0FDEFC6C8BF905FC" bold="true" box="[1226,1436,935,957]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0FDEFC6C8B4005FC" bold="true" box="[1226,1317,935,957]" italics="true" pageId="38" pageNumber="39">heinrichi</emphasis>
Grünwaldt
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E93D36EAAC7F435D0B83FC0E8BF905B6" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FC0E8BF905B6" pageId="38" pageNumber="39">
- Penis valves not greatly expanded laterally, not occupying almost the entire space between the gonostyli (
<figureCitation id="71B92A6FAC7F435D0FF1FC0E8B3B059B" box="[1253,1374,965,987]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="38" pageNumber="39">Figures 20F</figureCitation>
;
<figureCitation id="71B92A6FAC7F435D0E7DFC0E8BF7059B" box="[1385,1426,965,986]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">21F</figureCitation>
). Species more widespread.............................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="52762E4FAC7F435D0B83FBCB8BF802E9" pageId="38" pageNumber="39">
<paragraph id="E93D36EAAC7F435D0B83FBCB8BF9020E" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FBCB8BF9020E" pageId="38" pageNumber="39">
3. In profile view, penis valves relatively long, two times longer than broad (
<figureCitation id="71B92A6FAC7F435D08B9FBCB8A420254" box="[941,1063,1024,1046]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="38" pageNumber="39">Figures 20E</figureCitation>
;
<figureCitation id="71B92A6FAC7F435D0F25FBCB8A3B0254" box="[1073,1118,1024,1045]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">21H</figureCitation>
). In dorsal view, surface of penis valves flattened and somewhat “leathery”, with pattern of latitudinal striations, surface dull (
<figureCitation id="71B92A6FAC7F435D0F7AFBD68ABA0273" box="[1134,1247,1053,1075]" captionStart="FIGURE 20" captionStartId="36.[151,250,1975,1999]" captionTargetBox="[151,1436,510,1960]" captionTargetId="figure-216@36.[151,1436,510,1960]" captionTargetPageId="36" captionText="FIGURE 20. Andrena (Ulandrena) resoluta Warncke, 1973 paratype male. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view; E. Genital capsule, lateral view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209351" httpUri="https://zenodo.org/record/13209351/files/figure.png" pageId="38" pageNumber="39">Figure 20F</figureCitation>
). Southern Balkans across
<collectingCountry id="9195767AAC7F435D0A37FBF18F020211" box="[291,359,1082,1104]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
to
<collectingCountry id="9195767AAC7F435D0A91FBF18FBC0211" box="[389,473,1082,1104]" name="Armenia" pageId="38" pageNumber="39">Armenia</collectingCountry>
.................................................................
<taxonomicName id="2E824D69AC7F435D0FEDFBF08BF9020E" ID-CoL="66MPK" authority="Warncke" authorityName="Warncke" authorityYear="1965" box="[1273,1436,1082,1104]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="crecca">
<emphasis id="DBF6EAF8AC7F435D0FEDFBF08BF9020E" bold="true" box="[1273,1436,1082,1104]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0FEDFBF08B5C0211" bold="true" box="[1273,1337,1083,1104]" italics="true" pageId="38" pageNumber="39">crecca</emphasis>
Warncke
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E93D36EAAC7F435D0B83FB938BF802E9" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FB938BF802E9" pageId="38" pageNumber="39">
- In profile view, penis valves relatively short, clearly less than two times longer than wide (
<figureCitation id="71B92A6FAC7F435D0F55FB938AD1022C" box="[1089,1204,1112,1134]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21G</figureCitation>
). In dorsal view, surface of penis valves more or less domed, shining with only narrow longitudinal dull strip (
<figureCitation id="71B92A6FAC7F435D0F23FBBE8ACC02CB" box="[1079,1193,1141,1163]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21F</figureCitation>
).
<collectingCountry id="9195767AAC7F435D0FAFFBBE8A9A02CA" box="[1211,1279,1141,1163]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
(predominantly eastern
<collectingCountry id="9195767AAC7F435D0A3FFB598F1602E9" box="[299,371,1170,1192]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
), the Caucasus, and northern
<collectingCountry id="9195767AAC7F435D099AFB588CD002E9" box="[654,693,1171,1192]" name="Iran" pageId="38" pageNumber="39">Iran</collectingCountry>
............................................
<taxonomicName id="2E824D69AC7F435D0FC1FB588BF802E9" ID-CoL="DT55" authority="Radoszkowski" authorityName="Radoszkowski" authorityYear="1876" box="[1237,1437,1170,1192]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="tecta">
<emphasis id="DBF6EAF8AC7F435D0FC1FB588BF802E9" bold="true" box="[1237,1437,1170,1192]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0FC1FB588B6102E9" bold="true" box="[1237,1284,1171,1192]" italics="true" pageId="38" pageNumber="39">tecta</emphasis>
Radoszkowski
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="52762E4FAC7F435D0B83FB7B8BF8035C" pageId="38" pageNumber="39">
<paragraph id="E93D36EAAC7F435D0B83FB7B8BF902A2" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FB7B8BF902A2" pageId="38" pageNumber="39">
4. In profile view, penis valves relatively small (
<figureCitation id="71B92A6FAC7F435D0982FB7B8D6D0284" box="[662,776,1200,1222]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21A</figureCitation>
). In dorsal view, penis valves apically with the two halves presenting a deep “V-shaped” emargination (
<figureCitation id="71B92A6FAC7F435D0931FB068CF202A3" box="[549,663,1229,1251]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21C</figureCitation>
). Mediterranean
<collectingCountry id="9195767AAC7F435D082CFB068DE102A2" box="[824,900,1229,1251]" name="Albania" pageId="38" pageNumber="39">Albania</collectingCountry>
and
<collectingCountry id="9195767AAC7F435D08A7FB068D9202A3" box="[947,1015,1229,1250]" name="Greece" pageId="38" pageNumber="39">Greece</collectingCountry>
....................
<taxonomicName id="2E824D69AC7F435D0FE9FB068BF902A2" ID-CoL="DRD7" authority="Smith" authorityName="Smith" authorityYear="1853" box="[1277,1436,1229,1251]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="concinna">
<emphasis id="DBF6EAF8AC7F435D0FE9FB068BF902A2" bold="true" box="[1277,1436,1229,1251]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0FE9FB068B3C02A3" bold="true" box="[1277,1369,1229,1250]" italics="true" pageId="38" pageNumber="39">concinna</emphasis>
Smith
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="E93D36EAAC7F435D0B83FB208BF8035C" blockId="38.[151,1437,760,1310]" pageId="38" pageNumber="39">
<keyLead id="527395DFAC7F435D0B83FB208BF8035C" pageId="38" pageNumber="39">
- In profile view, penis valves relatively large (
<figureCitation id="71B92A6FAC7F435D0987FB218D6102BE" box="[659,772,1258,1280]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21B</figureCitation>
). In dorsal view, penis valves apically with the two halves presenting a relatively small and rounded emargination (
<figureCitation id="71B92A6FAC7F435D0981FAC38D62035C" box="[661,775,1288,1310]" captionStart="FIGURE 21" captionStartId="37.[151,243,1908,1932]" captionTargetBox="[214,1373,161,1792]" captionTargetId="figure-16@37.[214,1373,161,1899]" captionTargetPageId="37" captionText="FIGURE 21. Andrena (Ulandrena) concinna Smith, 1853 male.A. Genital capsule, lateral view; C. Genital capsule, dorsal view; Andrena (Ulandrena) paradoxa Friese, 1921 male.B. Genital capsule, lateral view; D.Genital capsule, dorsal view; Andrena (Ulandrena) heinrichi Grünwaldt, 2005 male. E. Genital capsule, dorsal view; Andrena (Ulandrena) tecta Radoszkowski, 1876 male. F. Genital capsule, dorsal view; G. Genital capsule, lateral view; Andrena (Ulandrena) crecca Warncke, 1965 male. H. Genital capsule, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13209355" httpUri="https://zenodo.org/record/13209355/files/figure.png" pageId="38" pageNumber="39">Figure 21D</figureCitation>
). Southern
<collectingCountry id="9195767AAC7F435D0861FAC38DDE035F" box="[885,955,1288,1310]" name="Turkey" pageId="38" pageNumber="39">Turkey</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D08D0FAC38D9C035F" box="[964,1017,1288,1310]" name="Syria" pageId="38" pageNumber="39">Syria</collectingCountry>
,
<collectingCountry id="9195767AAC7F435D0F16FAC38A52035F" box="[1026,1079,1288,1310]" name="Israel" pageId="38" pageNumber="39">Israel</collectingCountry>
...............
<taxonomicName id="2E824D69AC7F435D0FEEFACC8BF8035C" ID-CoL="66N5L" authority="Friese" authorityName="Friese" authorityYear="1921" box="[1274,1437,1287,1309]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="DBF6EAF8AC7F435D0FEEFACC8BF8035C" bold="true" box="[1274,1437,1287,1309]" pageId="38" pageNumber="39">
<emphasis id="DBF6EAF8AC7F435D0FEEFACC8B32035C" bold="true" box="[1274,1367,1287,1309]" italics="true" pageId="38" pageNumber="39">paradoxa</emphasis>
Friese
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,493 @@
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<emphasis id="DBF6EAF8ACD443F60B83FE818F640724" bold="true" box="[151,257,330,357]" italics="true" pageId="141" pageNumber="142">Andrena</emphasis>
(
<emphasis id="DBF6EAF8ACD443F60A06FE808FD30724" bold="true" box="[274,438,330,357]" italics="true" pageId="141" pageNumber="142">incertae sedis</emphasis>
)
<emphasis id="DBF6EAF8ACD443F60AD3FE808C3D0724" bold="true" box="[455,600,331,357]" italics="true" pageId="141" pageNumber="142">tenasserima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783ACD443F6094BFE808CB20724" box="[607,727,331,357]" pageId="141" pageNumber="142" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD443F60B83FEA48FC707C8" box="[151,418,366,393]" pageId="141" pageNumber="142" type="description">
<paragraph id="E93D36EAACD443F60B83FEA48FC707C8" blockId="141.[151,727,330,393]" box="[151,418,366,393]" pageId="141" pageNumber="142">
<figureCitation id="71B92A6FACD443F60B83FEA48F2107C8" box="[151,324,366,393]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figures 85AF</figureCitation>
;
<figureCitation id="71B92A6FACD443F60A5BFEA48FFB07C8" box="[335,414,367,393]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="141" pageNumber="142">86AF</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD443F60B83FE7D8A4D047C" pageId="141" pageNumber="142" type="materials_examined">
<paragraph id="E93D36EAACD443F60B83FE7D8D7D07B4" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<materialsCitation id="59EA3CB7ACD443F60B83FE7D8D7D07B4" collectingDate="1898-04-01" collectingDateMax="1898-04-30" collectingDateMin="1898-04-01" collectionCode="ZMHB" collectorName="C. T. Bingham" country="Myanmar" elevation="1219" location="Tenasserim" municipality="Thanduang" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8ACD443F60B83FE7D8F880791" bold="true" box="[151,493,438,464]" pageId="141" pageNumber="142">
<typeStatus id="36398848ACD443F60B83FE7D8F5F0791" box="[151,314,438,464]" pageId="141" pageNumber="142" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AACD443F60A58FE7D8F8D0791" box="[332,488,438,464]" name="Myanmar" pageId="141" pageNumber="142">MYANMAR</collectingCountry>
:
</emphasis>
<location id="EC5D6031ACD443F60AEDFE7C8C1E0790" LSID="urn:lsid:plazi:treatment:612B87FCACD443EB0B83FE818D1401D4:EC5D6031ACD443F60AEDFE7C8C1E0790" box="[505,635,439,465]" country="Myanmar" municipality="Thanduang" name="Tenasserim" pageId="141" pageNumber="142">Tenasserim</location>
,
<collectingMunicipality id="0959AC90ACD443F6099FFE7C8D6C0790" box="[651,777,439,465]" pageId="141" pageNumber="142">Thanduang</collectingMunicipality>
[Than Daung Gyi; 19.0769
<superScript id="1EF79BA2ACD443F60F42FE7C8A100790" attach="right" box="[1110,1141,439,465]" fontSize="6" pageId="141" pageNumber="142">oN</superScript>
, 96.6803
<superScript id="1EF79BA2ACD443F60FF5FE7C8A9B0790" attach="right" box="[1249,1278,439,465]" fontSize="6" pageId="141" pageNumber="142">oE</superScript>
],
<quantity id="2E7A9B0FACD443F60E03FE7C8B150790" box="[1303,1392,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="141" pageNumber="142" unit="ft" value="4000.0">
<elevation id="62AFD1D9ACD443F60E03FE7C8B150790" box="[1303,1392,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.2192" pageId="141" pageNumber="142" unit="ft" value="4000.0">4000 ft</elevation>
</quantity>
,
<date id="9D3C102AACD443F60E94FE7C8F6A07B4" pageId="141" pageNumber="142" value="1898-04-01" valueMax="1898-04-30" valueMin="1898-04-01">
<collectingDate id="8D78E9C2ACD443F60E94FE7C8F6A07B4" pageId="141" pageNumber="142" value="1898-04-01" valueMax="1898-04-30" valueMin="1898-04-01">1 30.iv.1898</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACD443F60A0FFE108F5507B4" box="[283,304,475,501]" count="1" pageId="141" pageNumber="142" type="female"></specimenCount>
, leg.
<collectorName id="4477533CACD443F60A7AFE108C6A07B4" box="[366,527,475,501]" pageId="141" pageNumber="142">C.T. Bingham</collectorName>
,
<collectionCode id="8F93AE2FACD443F6090EFE108C0E07B4" box="[538,619,475,501]" pageId="141" pageNumber="142">ZMHB</collectionCode>
(code 9edc81).
</materialsCitation>
</paragraph>
<paragraph id="E93D36EAACD443F60BD3FE358A4D047C" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<materialsCitation id="59EA3CB7ACD443F60BD3FE358E8F047C" collectingDate="2015-04-09" collectionCode="NMPC" collectorName="M. Mikat" country="Thailand" location="Doi Suthep-Pui National Park" pageId="141" pageNumber="142" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Chiang Mai" typeStatus="paratype">
<emphasis id="DBF6EAF8ACD443F60BD3FE358C790459" bold="true" box="[199,540,510,536]" pageId="141" pageNumber="142">
<typeStatus id="36398848ACD443F60BD3FE358F0A0459" box="[199,367,510,536]" pageId="141" pageNumber="142" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="9195767AACD443F60A68FE358C720459" box="[380,535,510,536]" name="Thailand" pageId="141" pageNumber="142">THAILAND</collectingCountry>
:
</emphasis>
<collectingRegion id="2B46F808ACD443F60931FE348CCB0458" box="[549,686,511,537]" country="Thailand" name="Chiang Mai" pageId="141" pageNumber="142">Chiang Mai</collectingRegion>
,
<location id="EC5D6031ACD443F609AEFE348A750458" LSID="urn:lsid:plazi:treatment:612B87FCACD443EB0B83FE818D1401D4:EC5D6031ACD443F609AEFE348A750458" box="[698,1040,511,537]" country="Thailand" name="Doi Suthep-Pui National Park" pageId="141" pageNumber="142" stateProvince="Chiang Mai">Doi Suthep-Pui National Park</location>
,
<date id="9D3C102AACD443F60F09FE358AE20458" box="[1053,1159,510,537]" pageId="141" pageNumber="142" value="2015-04-09">
<collectingDate id="8D78E9C2ACD443F60F09FE358AE20458" box="[1053,1159,510,537]" pageId="141" pageNumber="142" value="2015-04-09">9.iv.2015</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACD443F60F80FE348ADC0458" box="[1172,1209,511,537]" count="3" pageId="141" pageNumber="142" type="male">3♂</specimenCount>
,
<specimenCount id="FF84FD63ACD443F60FD2FE348A8E0458" box="[1222,1259,511,537]" count="2" pageId="141" pageNumber="142" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CACD443F60E3EFE348BFD0458" box="[1322,1432,511,537]" pageId="141" pageNumber="142">M. Mikát</collectorName>
,
<collectionCode id="8F93AE2FACD443F60B83FDE88E8F047C" box="[151,234,547,573]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="141" pageNumber="142">NMPC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7ACD443F60BE1FDE98A41047C" box="[245,1060,546,573]" collectionCode="ZMHB" country="Myanmar" location="MYANMAR" pageId="141" pageNumber="142" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="DBF6EAF8ACD443F60BE1FDE98FF3047D" bold="true" box="[245,406,546,572]" pageId="141" pageNumber="142">
<collectingCountry id="9195767AACD443F60BE1FDE98FF4047D" box="[245,401,546,572]" name="Myanmar" pageId="141" pageNumber="142">MYANMAR</collectingCountry>
:
</emphasis>
same information as
<typeStatus id="36398848ACD443F60993FDE88C89047C" box="[647,748,547,573]" pageId="141" pageNumber="142" type="holotype">holotype</typeStatus>
,
<specimenCount id="FF84FD63ACD443F609E3FDE88D7F047C" box="[759,794,547,573]" count="1" pageId="141" pageNumber="142" type="female">1♀</specimenCount>
,
<collectionCode id="8F93AE2FACD443F6083CFDE88D1C047C" box="[808,889,547,573]" pageId="141" pageNumber="142">ZMHB</collectionCode>
(code dcd15b)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD443F40BD3FD8D8DD906B8" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142" type="description">
<paragraph id="E93D36EAACD443F60BD3FD8D8BFC05E4" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<emphasis id="DBF6EAF8ACD443F60BD3FD8D8FA40420" bold="true" box="[199,449,582,609]" pageId="141" pageNumber="142">Description. Female.</emphasis>
Body length:
<quantity id="2E7A9B0FACD443F6097EFD8C8CAF0420" box="[618,714,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="141" pageNumber="142" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
(
<figureCitation id="71B92A6FACD443F609C8FD8C8D000420" box="[732,869,582,609]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85A</figureCitation>
).
<emphasis id="DBF6EAF8ACD443F60868FD8C8DA40420" box="[892,961,583,609]" italics="true" pageId="141" pageNumber="142">Head:</emphasis>
Dark, 1.4 times wider than long (
<figureCitation id="71B92A6FACD443F60E45FD8C8EAF04C4" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85B</figureCitation>
). Clypeus strongly domed and flattened medially, thus weakly three-faced, irregularly punctate, punctures separated by 0.52 puncture diameters, interspaces shining. Process of labrum trapezoidal, 2 times broader than long, apical margin truncate. Gena exceeding width of compound eye; ocelloccipital distance 2.5 times diameter of lateral ocellus (
<figureCitation id="71B92A6FACD443F60A73FD1C8F8B04B0" box="[359,494,726,753]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85C</figureCitation>
). Vertex with surface posterior to ocellar triangle slightly but distinctly depressed, forming slight latitudinal furrow. Foveae dorsally occupying slightly more than ½ space between compound eye and lateral ocellus, separated from lateral ocellus by distance greater than diameter of lateral ocellus; foveae narrowing slightly ventrally, extending slightly beyond lower margin of antennal insertions, dorsally very slightly impressed, filled with brown hairs. Head covered with short and sparse whitish hairs, becoming brownish on vertex. Antennae basally dark, A512 ventrally lightened by presence of orange-brown scales; A3 exceeding A4, shorter than A4+5.
</paragraph>
<paragraph id="E93D36EAACD443F60BD3FC648B190370" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<emphasis id="DBF6EAF8ACD443F60BD3FC648F2D0589" box="[199,328,943,968]" italics="true" pageId="141" pageNumber="142">Mesosoma:</emphasis>
Scutum and scutellum very weakly sculptured, more or less polished and shining, densely punctate, punctures separated by ≤0.5 puncture diameters (
<figureCitation id="71B92A6FACD443F60838FC188DD605AC" box="[812,947,978,1005]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85D</figureCitation>
). Pronotum rounded. Mesepisternum finely microreticulate, weakly shining. Dorsolateral parts of propodeum finely reticulate, forming slightly raised network. Propodeal triangle broad, broader than long, laterally defined by narrow slightly raised carinae, internal surface with fine granular reticulation, basally and medially with weakly raised network of rugosity (
<figureCitation id="71B92A6FACD443F60F48FBF48ABB0218" box="[1116,1246,1086,1113]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85E</figureCitation>
). Mesepisternum laterally with whitish hairs, scutum with short and densely plumose whitish and brownish hairs, almost subsquamous; scutellum with dense tuft of longer dark brown hairs, these less strongly plumose. Propodeal corbicula incomplete, dorsal fringe composed of whitish plumose hairs, internal surface with scattered long pale hairs. Legs dark, apically becoming reddish-brown, pubescence brownish. Flocculus complete, composed of white plumose hairs; femoral scopae composed of pale simple hairs, tibial scopae composed of simple hairs, ventrally pale, dorsally dark brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma and venation dark orange-brown, nervulus antefurcal.
</paragraph>
<paragraph id="E93D36EAACD443F60BD3FAF08A2B03A4" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<emphasis id="DBF6EAF8ACD443F60BD3FAF08F200315" box="[199,325,1339,1364]" italics="true" pageId="141" pageNumber="142">Metasoma:</emphasis>
Tergal discs dark, marginal areas lightened hyaline-brown (
<figureCitation id="71B92A6FACD443F608E2FAF08A1F0314" box="[1014,1146,1338,1365]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="141" pageNumber="142">Figure 85F</figureCitation>
). Terga densely punctate, punctures separated by 0.51 puncture diameters, becoming slightly sparser on marginal areas, underlying surface polished and shining. Tergal discs with very short brown hairs, laterally with scattered pale hairs, not forming hairbands or apical fringes. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate large, rounded triangular, medially with large swollen raised area, surface impunctate, dull.
</paragraph>
<paragraph id="E93D36EAACD443F60BD3FA248FD400A0" blockId="141.[151,1437,438,2013]" pageId="141" pageNumber="142">
<emphasis id="DBF6EAF8ACD443F60BD3FA248F6E0048" bold="true" box="[199,267,1519,1545]" pageId="141" pageNumber="142">Male.</emphasis>
Body length: 6.5
<quantity id="2E7A9B0FACD443F60AF5FA258C470048" box="[481,546,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="141" pageNumber="142" unit="mm" value="7.0">7 mm</quantity>
(
<figureCitation id="71B92A6FACD443F60926FA248CDF0048" box="[562,698,1519,1545]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="141" pageNumber="142">Figure 86A</figureCitation>
).
<emphasis id="DBF6EAF8ACD443F609DAFA248D710048" box="[718,788,1519,1545]" italics="true" pageId="141" pageNumber="142">Head:</emphasis>
Dark, 1.35 times wider than long (
<figureCitation id="71B92A6FACD443F60FB2FA248B490048" box="[1190,1324,1519,1545]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="141" pageNumber="142">Figure 86B</figureCitation>
). Clypeus dark, domed, irregularly punctate, punctures separated by 0.52 puncture diameters, underlying surface polished and shining over majority of area, laterally shagreened and dull. Process of labrum small, trapezoidal, slightly broader than long, apical margin truncate. Gena equalling width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Gena, vertex, clypeus, and scape covered with short light brown hairs, paraocular areas laterally and frons with longer black hairs, weakly intermixed. Antennae basally dark, A413 ventrally lightened orange; A3 equalling A4.
</paragraph>
<paragraph id="E93D36EAACD443F50BD3F9208D550694" blockId="141.[151,1437,438,2013]" lastBlockId="142.[151,1436,151,213]" lastPageId="142" lastPageNumber="143" pageId="141" pageNumber="142">
<emphasis id="DBF6EAF8ACD443F60BD3F9208F2D0145" box="[199,328,1771,1796]" italics="true" pageId="141" pageNumber="142">Mesosoma:</emphasis>
Scutum and scutellum with fine granular shagreen, medially weak and almost shining, laterally and anteriorly stronger, almost dull; surface punctate, punctures separated by 0.51 puncture diameters (
<figureCitation id="71B92A6FACD443F60E45F8C48EAF010C" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="141" pageNumber="142">Figure 86C</figureCitation>
). Pronotum rounded with faint hint of humeral angle. Mesepisternum microreticulate, dull to weakly shining ventrally. Dorsolateral parts of propodeum finely reticulate, forming slightly raised network of wrinkles. Propodeal triangle broad, broader than long, laterally defined by narrow slightly raised carinae, internal surface with fine granular reticulation, basally and medially with weakly raised network of rugosity (
<figureCitation id="71B92A6FACD443F60F46F8548ABF01F8" box="[1106,1242,1951,1977]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="141" pageNumber="142">Figure 86D</figureCitation>
). Mesepisternum laterally with light brown plumose hair, becoming slightly shorter on scutum and scutellum. Legs dark, apical tarsal segments lightened orange, pubescence whitish to light brown. Hind tarsal claws with inner tooth. Wings hyaline, stigma brown, venation orange-brown, nervulus antefurcal.
</paragraph>
<caption id="BDFD6662ACD743F50B83F96C8A6A00A2" ID-DOI="http://doi.org/10.5281/zenodo.13209499" ID-Zenodo-Dep="13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="142" pageNumber="143" startId="142.[151,250,1703,1727]" targetBox="[151,1436,237,1680]" targetPageId="142" targetType="figure">
<paragraph id="E93D36EAACD743F50B83F96C8A6A00A2" blockId="142.[151,1437,1703,1763]" pageId="142" pageNumber="143">
<emphasis id="DBF6EAF8ACD743F50B83F96C8F7A00FE" bold="true" box="[151,287,1703,1727]" pageId="142" pageNumber="143">FIGURE 85.</emphasis>
<taxonomicName id="2E824D69ACD743F50A31F96C8CFF00FE" authorityName="WOOD" authorityYear="2024" box="[293,666,1703,1727]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="tenasserima" status="sp. nov.">
<emphasis id="DBF6EAF8ACD743F50A31F96C8F1E00FE" box="[293,379,1703,1727]" italics="true" pageId="142" pageNumber="143">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8ACD743F50934F9638CFF00FE" box="[544,666,1704,1727]" italics="true" pageId="142" pageNumber="143">tenasserima</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8ACD743F509B4F96C8D6300FE" bold="true" box="[672,774,1703,1727]" pageId="142" pageNumber="143">
<taxonomicNameLabel id="C0C55783ACD743F509B4F96C8D6300FE" box="[672,774,1703,1727]" pageId="142" pageNumber="143" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<paragraph id="E93D36EAACD743F40BD3F8D98DD906B8" blockId="142.[151,1437,1809,2016]" lastBlockId="143.[151,1437,151,251]" lastPageId="143" lastPageNumber="144" pageId="142" pageNumber="143">
<emphasis id="DBF6EAF8ACD743F50BD3F8D98F20016A" box="[199,325,1810,1835]" italics="true" pageId="142" pageNumber="143">Metasoma:</emphasis>
Tergal discs dark, marginal areas with apical rims narrowly lightened hyaline-brown (
<figureCitation id="71B92A6FACD743F50E19F8DA8BF5016A" box="[1293,1424,1809,1835]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="142" pageNumber="143">Figure 86E</figureCitation>
). Terga densely punctate, punctures separated by 0.51 puncture diameters, becoming slightly sparser on marginal areas, underlying surface polished and shining. Tergal discs with very short brown hairs, laterally with scattered pale hairs, not forming hairbands or apical fringes. T67 with long brown hairs overlying pygidial plate of T7. S8 narrow, columnar, ventrally with strong laterally-projecting fringes of orange hairs, these not projecting apically, hence giving impression of fishtail-like apex to S8. Genital capsule slightly elongate, gonocoxae produced into pointed and apically diverging teeth (
<figureCitation id="71B92A6FACD643F40958FF5C8CB606F0" box="[588,723,151,177]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="143" pageNumber="144">Figure 86F</figureCitation>
). Gonostyli flattened apically, apexes roughly triangular with strongly raised internal margin, outer surface weakly punctate. Penis valves slightly inflated medially, occupying slightly less than ½ space between gonostyli, gently narrowing apically.
</paragraph>
</subSubSection>
<caption id="BDFD6662ACD643F40B83F9028A3A0144" ID-DOI="http://doi.org/10.5281/zenodo.13209503" ID-Zenodo-Dep="13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="143" pageNumber="144" startId="143.[151,250,1737,1761]" targetBox="[151,1436,273,1713]" targetPageId="143" targetType="figure">
<paragraph id="E93D36EAACD643F40B83F9028A3A0144" blockId="143.[151,1436,1737,1797]" pageId="143" pageNumber="144">
<emphasis id="DBF6EAF8ACD643F40B83F9028F4500A0" bold="true" box="[151,288,1737,1761]" pageId="143" pageNumber="144">FIGURE 86.</emphasis>
<taxonomicName id="2E824D69ACD643F40A32F9028CF900A0" authorityName="WOOD" authorityYear="2024" box="[294,668,1737,1761]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="tenasserima" status="sp. nov.">
<emphasis id="DBF6EAF8ACD643F40A32F9028F1900A0" box="[294,380,1737,1761]" italics="true" pageId="143" pageNumber="144">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8ACD643F40936F9018CF900A0" box="[546,668,1738,1761]" italics="true" pageId="143" pageNumber="144">tenasserima</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8ACD643F409B6F9028D6C00A0" bold="true" box="[674,777,1737,1761]" pageId="143" pageNumber="144">
<taxonomicNameLabel id="C0C55783ACD643F409B6F9028D6C00A0" box="[674,777,1737,1761]" pageId="143" pageNumber="144" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561ACD643EB0BD3F8F88CF40284" lastPageId="144" lastPageNumber="145" pageId="143" pageNumber="144" type="diagnosis">
<paragraph id="E93D36EAACD643EB0BD3F8F88F010724" blockId="143.[151,1436,1843,2014]" lastBlockId="144.[151,1437,150,1941]" lastPageId="144" lastPageNumber="145" pageId="143" pageNumber="144">
<emphasis id="DBF6EAF8ACD643F40BD3F8F88F27010C" bold="true" box="[199,322,1843,1869]" pageId="143" pageNumber="144">Diagnosis.</emphasis>
<taxonomicName id="2E824D69ACD643F40A5CF8F88C50010C" authorityName="WOOD" authorityYear="2024" box="[328,565,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACD643F40A5CF8F88C50010C" box="[328,565,1843,1869]" italics="true" pageId="143" pageNumber="144">Andrena tenasserima</emphasis>
</taxonomicName>
presents a combination of characters that do not currently allow for confident subgeneric placement. Morphologically, due to the head with relatively long ocelloccipital distance (2.5 times the diameter of a lateral ocellus,
<figureCitation id="71B92A6FACD643F40AC8F8B08C3901D7" box="[476,604,1915,1942]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="143" pageNumber="144">Figure 85C</figureCitation>
) and vertex which is slightly but distinctly depressed, forming a slight but visible furrow along the posterior margins of the hind ocelli, facial foveae that dorsally extend to the anterior margins of the lateral ocelli, gena broader than the width of the compound eye, broad and relatively unsculptured propodeal triangle (
<figureCitation id="71B92A6FACC943EB0BE8FF5C8F1B06F0" box="[252,382,150,177]" captionStart="FIGURE 85" captionStartId="142.[151,250,1703,1727]" captionTargetBox="[151,1436,237,1680]" captionTargetId="figure-50@142.[151,1436,237,1680]" captionTargetPageId="142" captionText="FIGURE 85. Andrena (incertae sedis) tenasserima spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Vertex, dorsal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209499" httpUri="https://zenodo.org/record/13209499/files/figure.png" pageId="144" pageNumber="145">Figure 85E</figureCitation>
), and lack of any other distinctive characters, it can be placed close to the group of subgenera
<taxonomicName id="2E824D69ACC943EB0B83FF708C0A0694" authority="Hirashima &amp; LaBerge, 1963" authorityName="Hirashima &amp; LaBerge" authorityYear="1963" box="[151,623,187,213]" class="Insecta" genus="Calomelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB0B83FF708F440694" box="[151,289,187,213]" italics="true" pageId="144" pageNumber="145">Calomelissa</emphasis>
Hirashima &amp; LaBerge, 1963
</taxonomicName>
,
<taxonomicName id="2E824D69ACC943EB096FFF708A3F0694" authority="Hirashima &amp; Tadauchi, 1975" authorityName="Hirashima &amp; Tadauchi" authorityYear="1975" box="[635,1114,186,213]" class="Insecta" family="Andrenidae" genus="Oreomelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB096FFF708D6D0694" box="[635,776,187,213]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
Hirashima &amp; Tadauchi, 1975
</taxonomicName>
, and
<taxonomicName id="2E824D69ACC943EB0F83FF708B410694" authorityName="Osytshnjuk" authorityYear="1984" box="[1175,1316,187,213]" pageId="144" pageNumber="150" rank="subGenus" subGenus="Tarsandrena">
<emphasis id="DBF6EAF8ACC943EB0F83FF708B410694" box="[1175,1316,187,213]" italics="true" pageId="144" pageNumber="145">Tarsandrena</emphasis>
</taxonomicName>
. Based on limited taxon sampling, these three subgenera form a monophyletic and relatively basal clade within
<taxonomicName id="2E824D69ACC943EB0E29FF148FEB075C" authority="(Pisanty et al. 2022 b)" baseAuthorityName="Pisanty" baseAuthorityYear="2022" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB0E29FF148BF906B8" box="[1341,1436,223,249]" italics="true" pageId="144" pageNumber="145">Andrena</emphasis>
(
<bibRefCitation id="8D134B1BACC943EB0B8BFEC88FE0075C" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[159,389,259,285]" pageId="144" pageNumber="145" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8ACC943EB0BEFFEC88F49075C" box="[251,300,259,285]" italics="true" pageId="144" pageNumber="145">et al</emphasis>
. 2022b
</bibRefCitation>
)
</taxonomicName>
. The greatest diversity of these three subgenera is clearly found in Asia, predominantly
<collectingCountry id="9195767AACC943EB0E78FEC88EA80700" name="China" pageId="144" pageNumber="145">East Asia</collectingCountry>
, with only three species entering the West Palaearctic (
<bibRefCitation id="8D134B1BACC943EB083CFEEC8A710700" author="Xu, H. - L. &amp; Tadauchi, O." box="[808,1044,294,321]" pageId="144" pageNumber="145" pagination="621 - 631" refId="ref92541" refString="Xu, H. - L. &amp; Tadauchi, O. (1995) Revision of the Subgenus Calomelissa of the Genus Andrena (Hymenoptera, Andrenidae) of Eastern Asia. Japanese Journal of Entomology, 63, 621 - 631." type="journal article" year="1995">Xu &amp; Tadauchi 1995</bibRefCitation>
; 1999;
<bibRefCitation id="8D134B1BACC943EB0F77FEEC8B610700" author="Xu, H. - L. &amp; Tadauchi, O. &amp; Wu, Y. - R." box="[1123,1284,295,321]" pageId="144" pageNumber="145" pagination="41 - 61" refId="ref92909" refString="Xu, H. - L., Tadauchi, O. &amp; Wu, Y. - R. (2000) A Revision of the Subgenus Oreomelissa of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 40, 41 - 61. https: // doi. org / 10.5109 / 2641" type="journal article" year="2000">
Xu
<emphasis id="DBF6EAF8ACC943EB0F9FFEEC8ADE0700" box="[1163,1211,295,321]" italics="true" pageId="144" pageNumber="145">et al</emphasis>
. 2000
</bibRefCitation>
;
<bibRefCitation id="8D134B1BACC943EB0E19FEEC8F3D0724" author="Gusenleitner, F. &amp; Schwarz, M." pageId="144" pageNumber="145" pagination="1 - 1280" refId="ref88973" refString="Gusenleitner, F. &amp; Schwarz, M. (2002) Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Erganzungen zu palaarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna, Supplement 10, 1 - 1280." type="journal article" year="2002">Gusenleitner &amp; Schwarz 2002</bibRefCitation>
).
</paragraph>
<paragraph id="E93D36EAACC943EB0BD3FEA48F61051C" blockId="144.[151,1437,150,1941]" pageId="144" pageNumber="145">
These subgenera are not strongly defined morphologically, with many of the highlighted characters variable (e.g. head length, humeral angle, tergal sculpture, foveae width, forewing venation;
<bibRefCitation id="8D134B1BACC943EB0F5EFE588A9407EC" author="Xu, H. - L. &amp; Tadauchi, O. &amp; Wu, Y. - R." box="[1098,1265,403,429]" pageId="144" pageNumber="145" pagination="41 - 61" refId="ref92909" refString="Xu, H. - L., Tadauchi, O. &amp; Wu, Y. - R. (2000) A Revision of the Subgenus Oreomelissa of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 40, 41 - 61. https: // doi. org / 10.5109 / 2641" type="journal article" year="2000">
Xu
<emphasis id="DBF6EAF8ACC943EB0F60FE588AC307EC" box="[1140,1190,403,429]" italics="true" pageId="144" pageNumber="145">et al</emphasis>
. 2000
</bibRefCitation>
). Further taxon sampling is needed to establish whether retention of the three subgenera is merited. In this context, placement is closest to the subgenus
<taxonomicName id="2E824D69ACC943EB0AB7FE108C5507B4" authorityName="Hirashima &amp; Tadauchi" authorityYear="1975" box="[419,560,475,501]" class="Insecta" family="Andrenidae" genus="Oreomelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB0AB7FE108C5507B4" box="[419,560,475,501]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
</taxonomicName>
due to the shape of the vertex, which is long and clearly impressed along the posterior margins of the hind ocelli. However, there are some differences; in most
<taxonomicName id="2E824D69ACC943EB0F2BFE348B4D0458" box="[1087,1320,511,537]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0F2BFE348AA90458" box="[1087,1228,511,537]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
species
</taxonomicName>
, the facial foveae extend further dorsally, reaching to the posterior margins of the lateral ocelli, and are noticeably impressed, whereas in
<taxonomicName id="2E824D69ACC943EB0A06FD8C8FD30421" authorityName="WOOD" authorityYear="2024" box="[274,438,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0A06FD8C8FD30421" box="[274,438,583,609]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
the facial foveae are only very slightly impressed dorsally. Furthermore, most
<taxonomicName id="2E824D69ACC943EB0E04FD8C8E8D04C4" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0E04FD8C8BF80420" box="[1296,1437,583,609]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
species
</taxonomicName>
have the dorsal (horizontal) section of the propodeum relatively elongate, so in dorsal view the propodeal triangle is about as long as broad. In contrast, in
<taxonomicName id="2E824D69ACC943EB09D2FD448D0804E9" authorityName="WOOD" authorityYear="2024" box="[710,877,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB09D2FD448D0804E9" box="[710,877,655,681]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
the dorsal section of the propodeum is relatively short and so the propodeal triangle is clearly broader than long. The female pronotum also usually presents a weak humeral angle (though this is variable), but this is lacking in
<taxonomicName id="2E824D69ACC943EB0829FD1C8D8704B1" authorityName="WOOD" authorityYear="2024" box="[829,994,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0829FD1C8D8704B1" box="[829,994,727,753]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
. Moreover, the male clypeus and lower paraocular areas are black (
<figureCitation id="71B92A6FACC943EB0AC0FD308C3F0554" box="[468,602,763,789]" captionStart="FIGURE 86" captionStartId="143.[151,250,1737,1761]" captionTargetBox="[151,1436,273,1713]" captionTargetId="figure-54@143.[151,1436,273,1713]" captionTargetPageId="143" captionText="FIGURE 86. Andrena (incertae sedis) tenasserima spec. nov. male. A. Habitus, lateral view; B. Face, frontal view; C. Scutum, dorsal view; D. Propodeum, dorsal view; E. Terga, dorsal view; F. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209503" httpUri="https://zenodo.org/record/13209503/files/figure.png" pageId="144" pageNumber="145">Figure 86B</figureCitation>
; yellow or white-marked in
<taxonomicName id="2E824D69ACC943EB08B0FD308A540554" authorityName="Hirashima &amp; Tadauchi" authorityYear="1975" box="[932,1073,763,789]" class="Insecta" family="Andrenidae" genus="Oreomelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB08B0FD308A540554" box="[932,1073,763,789]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
</taxonomicName>
). Consequently,
<taxonomicName id="2E824D69ACC943EB0FE7FD308BF90555" authorityName="WOOD" authorityYear="2024" box="[1267,1436,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0FE7FD308BF90555" box="[1267,1436,763,789]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
is cautiously associated with the subgenus
<taxonomicName id="2E824D69ACC943EB0993FCD48D710578" authorityName="Hirashima &amp; Tadauchi" authorityYear="1975" box="[647,788,799,825]" class="Insecta" family="Andrenidae" genus="Oreomelissa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB0993FCD48D710578" box="[647,788,799,825]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
</taxonomicName>
, but is classified as
<emphasis id="DBF6EAF8ACC943EB0F17FCD48AC50578" box="[1027,1184,799,825]" italics="true" pageId="144" pageNumber="145">incertae sedis</emphasis>
until genetic data are available.
</paragraph>
<paragraph id="E93D36EAACC943EB0BD3FCAC8CF40284" blockId="144.[151,1437,150,1941]" pageId="144" pageNumber="145">
Diagnosis is provided against Chinese
<taxonomicName id="2E824D69ACC943EB0994FCAC8D0305C0" box="[640,870,871,897]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0994FCAC8D6805C0" box="[640,781,871,897]" italics="true" pageId="144" pageNumber="145">Oreomelissa</emphasis>
species
</taxonomicName>
which are currently known to extend south to the northern part of
<collectingRegion id="2B46F808ACC943EB0A47FC408C7605E4" box="[339,531,907,933]" country="China" name="Yunnan" pageId="144" pageNumber="145">Yunnan province</collectingRegion>
in southern
<collectingCountry id="9195767AACC943EB09B0FC408C8205E4" box="[676,743,907,933]" name="China" pageId="144" pageNumber="145">China</collectingCountry>
(
<bibRefCitation id="8D134B1BACC943EB09ECFC408DC105E4" author="Xu, H. - L. &amp; Tadauchi, O. &amp; Wu, Y. - R." box="[760,932,907,933]" pageId="144" pageNumber="145" pagination="41 - 61" refId="ref92909" refString="Xu, H. - L., Tadauchi, O. &amp; Wu, Y. - R. (2000) A Revision of the Subgenus Oreomelissa of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 40, 41 - 61. https: // doi. org / 10.5109 / 2641" type="journal article" year="2000">
Xu
<emphasis id="DBF6EAF8ACC943EB0831FC408D3D05E4" box="[805,856,907,933]" italics="true" pageId="144" pageNumber="145">et al</emphasis>
. 2000
</bibRefCitation>
), approximately
<quantity id="2E7A9B0FACC943EB0F70FC408AAC05E4" box="[1124,1225,907,933]" metricMagnitude="6" metricUnit="m" metricValue="1.0" pageId="144" pageNumber="145" unit="km" value="1000.0">1000 km</quantity>
north of the
<emphasis id="DBF6EAF8ACC943EB0E75FC408E830589" italics="true" pageId="144" pageNumber="145">locus typicus</emphasis>
of
<taxonomicName id="2E824D69ACC943EB0A19FC648FD60589" authorityName="WOOD" authorityYear="2024" box="[269,435,943,969]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0A19FC648FD60589" box="[269,435,943,969]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
, and which are consequently likely to be present in northern
<collectingCountry id="9195767AACC943EB0F72FC648AB00588" box="[1126,1237,943,969]" name="Myanmar" pageId="144" pageNumber="145">Myanmar</collectingCountry>
. Due to the broad gena (broader than the width of the compound eye), lack of spines along the posterior face of the hind femur (with spines in
<taxonomicName id="2E824D69ACC943EB0A53FC3C8C0E0250" authority="Wu, 1982" authorityName="Wu" authorityYear="1982" box="[327,619,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="submontana">
<emphasis id="DBF6EAF8ACC943EB0A53FC3C8F940250" box="[327,497,1015,1041]" italics="true" pageId="144" pageNumber="145">A. submontana</emphasis>
Wu, 1982
</taxonomicName>
,
<taxonomicName id="2E824D69ACC943EB0963FC3C8A3A0250" authority="Gusenleitner &amp; Schwarz, 2000" authorityName="Gusenleitner &amp; Schwarz" authorityYear="2000" box="[631,1119,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="malickyi">
<emphasis id="DBF6EAF8ACC943EB0963FC3C8C930250" box="[631,758,1015,1041]" italics="true" pageId="144" pageNumber="145">A. malickyi</emphasis>
<bibRefCitation id="8D134B1BACC943EB09EBFC3C8A3A0250" author="Gusenleitner, F. &amp; Schwarz, M." box="[767,1119,1015,1041]" pageId="144" pageNumber="145" pagination="105 - 116" refId="ref88896" refString="Gusenleitner, F. &amp; Schwarz, M. (2000) Nomenklatorische Aktualisierungen in der Bienengattung Andrena sowie Beschreibung einer neuen Art (Hymenoptera: Apidae: Andreninae). Entomofauna, 10, 105 - 116." type="journal article" year="2000">Gusenleitner &amp; Schwarz, 2000</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="2E824D69ACC943EB0F88FC3C8EB60274" authority="Wu, 2000" authorityName="Wu" authorityYear="2000" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="setosifemoralis">
<emphasis id="DBF6EAF8ACC943EB0F88FC3C8B030250" box="[1180,1382,1015,1041]" italics="true" pageId="144" pageNumber="145">A. setosifemoralis</emphasis>
Wu, 2000
</taxonomicName>
), polished and densely punctate terga (punctures separated by 0.51 puncture diameters; in
<taxonomicName id="2E824D69ACC943EB0E15FBD08F200218" authority="Xu &amp; Tadauchi, 2000" authorityName="Xu &amp; Tadauchi" authorityYear="2000" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="fani">
<emphasis id="DBF6EAF8ACC943EB0E15FBD08B350274" box="[1281,1360,1051,1077]" italics="true" pageId="144" pageNumber="145">A. fani</emphasis>
<bibRefCitation id="8D134B1BACC943EB0E4EFBD08F200218" author="Xu, H. - L. &amp; Tadauchi, O. &amp; Wu, Y. - R." pageId="144" pageNumber="145" pagination="41 - 61" refId="ref92909" refString="Xu, H. - L., Tadauchi, O. &amp; Wu, Y. - R. (2000) A Revision of the Subgenus Oreomelissa of the Genus Andrena of Eastern Asia (Hymenoptera, Andrenidae). Esakia, 40, 41 - 61. https: // doi. org / 10.5109 / 2641" type="journal article" year="2000">Xu &amp; Tadauchi, 2000</bibRefCitation>
</taxonomicName>
with terga shagreened with obscure and scattered punctures, punctures separated by 13 puncture diameters), and dark male face and genital capsule with relatively narrow penis valves, separation is straightforward. Additionally,
<taxonomicName id="2E824D69ACC943EB0A25FB4C8FB202E1" authorityName="WOOD" authorityYear="2024" box="[305,471,1159,1185]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0A25FB4C8FB202E1" box="[305,471,1159,1185]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
</taxonomicName>
can be separated from all of these species by the only weakly impressed facial foveae and the broader than long propodeal triangle.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACC943EB0BD3FB048AEC0144" pageId="144" pageNumber="145" type="discussion">
<paragraph id="E93D36EAACC943EB0BD3FB048D87006C" blockId="144.[151,1437,150,1941]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0BD3FB048F5D02A8" bold="true" box="[199,312,1231,1257]" pageId="144" pageNumber="145">Remarks.</emphasis>
Charles Thomas Bingham (1848†1908) was an Irish military officer and naturalist. He was posted in what is now
<collectingCountry id="9195767AACC943EB0A08FB388FE3034C" box="[284,390,1267,1293]" name="Myanmar" pageId="144" pageNumber="145">Myanmar</collectingCountry>
in 1877, and developed an interest in entomology. He later worked in the NHMUK, and edited the first two
<taxonomicName id="2E824D69ACC943EB0BE3FADC8FEE0370" box="[247,395,1303,1329]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
volumes of the Fauna of
<collectingCountry id="9195767AACC943EB098EFADC8D470370" box="[666,802,1303,1329]" name="India" pageId="144" pageNumber="145">British India</collectingCountry>
(
<bibRefCitation id="8D134B1BACC943EB083BFADC8DB10370" author="Bingham, C. T." box="[815,980,1303,1329]" pageId="144" pageNumber="145" refId="ref87688" refString="Bingham, C. T. (1897) Hymenoptera. Vol. I. Wasps and Bees. British Fauna of India. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1897">Bingham 1897</bibRefCitation>
;
<bibRefCitation id="8D134B1BACC943EB08C9FADC8A700370" author="Bingham, C. T." box="[989,1045,1303,1329]" pageId="144" pageNumber="145" refId="ref87735" refString="Bingham, C. T. (1903) Hymenoptera. Vol. II. Ants and Cuckoo-wasps. British Fauna of India. Taylor and Francis, London, 506 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1903">1903</bibRefCitation>
) and described two
<taxonomicName id="2E824D69ACC943EB0FF8FADC8BF90370" box="[1260,1436,1303,1329]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0FF8FADC8B2C0370" box="[1260,1353,1303,1329]" italics="true" pageId="144" pageNumber="145">Andrena</emphasis>
species
</taxonomicName>
from
<collectingCountry id="9195767AACC943EB0BC6FAF08F6F0314" box="[210,266,1339,1365]" name="India" pageId="144" pageNumber="145">India</collectingCountry>
(
<bibRefCitation id="8D134B1BACC943EB0A0CFAF08FDB0314" author="Bingham, C. T." box="[280,446,1339,1365]" pageId="144" pageNumber="145" pagination="347 - 368" refId="ref87782" refString="Bingham, C. T. (1908) Notes of aculeate Hymenoptera in the Indian Museum. Records of the Indian Museum, 2, 347 - 368. https: // doi. org / 10.26515 / rzsi / v 2 / i 4 / 1908 / 163353" type="journal article" year="1908">Bingham 1908</bibRefCitation>
). For
<taxonomicName id="2E824D69ACC943EB0AE9FAF08D6A0314" authority=", Bingham" authorityName="Bingham" box="[509,783,1339,1365]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="tenasserima">
<emphasis id="DBF6EAF8ACC943EB0AE9FAF08CF80315" box="[509,669,1339,1365]" italics="true" pageId="144" pageNumber="145">A. tenasserima</emphasis>
, Bingham
</taxonomicName>
indicated “Tenasserim” on the labels, but some geographers separate the Tenasserim Range from the more northerly Dawna Range, and others consider them collectively as “Dawna Tenasserim”. The collecting locality of Than Daung Gyi falls within the Dawna Range. The village itself was used by the British during the colonial period as a summer retreat from the heat of the lowlands, as it is found above
<quantity id="2E7A9B0FACC943EB0E3AFA6C8BE50380" box="[1326,1408,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="144" pageNumber="145" unit="m" value="1000.0">1000 m</quantity>
in elevation, and hence has a cooler climate more suited to
<taxonomicName id="2E824D69ACC943EB09E0FA008DCC03A4" box="[756,937,1483,1509]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB09E0FA008D3403A4" box="[756,849,1483,1509]" italics="true" pageId="144" pageNumber="145">Andrena</emphasis>
species
</taxonomicName>
; this is the first
<taxonomicName id="2E824D69ACC943EB0F44FA008B6403A4" box="[1104,1281,1483,1509]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0F44FA008AC803A4" box="[1104,1197,1483,1509]" italics="true" pageId="144" pageNumber="145">Andrena</emphasis>
species
</taxonomicName>
reported from this predominantly tropical country (
<bibRefCitation id="8D134B1BACC943EB0935FA248D5D0048" author="Ascher, J. S. &amp; Pickering, J." box="[545,824,1519,1545]" pageId="144" pageNumber="145" refId="ref87181" refString="Ascher, J. S. &amp; Pickering, J. (2023) Discover Life bee species guide and world checklist (Hymenoptera: Apidae: Anthophila). Available from: https: // www. discoverlife. org / mp / 20 q? guide = Apoidea _ species &amp; flags = HAS (accessed 28 May 2024)" type="url" year="2023">Ascher &amp; Pickering 2023</bibRefCitation>
), with only
<taxonomicName id="2E824D69ACC943EB08A2FA248BF90048" authority="Baker, 1995" authorityName="Baker" authorityYear="1995" box="[950,1436,1518,1545]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="chrysochersonus" subGenus="Malayapis">
<emphasis id="DBF6EAF8ACC943EB08A2FA248DAB0048" box="[950,974,1519,1545]" italics="true" pageId="144" pageNumber="145">A.</emphasis>
(
<emphasis id="DBF6EAF8ACC943EB08C8FA248A2A0048" box="[988,1103,1519,1545]" italics="true" pageId="144" pageNumber="145">Malayapis</emphasis>
)
<emphasis id="DBF6EAF8ACC943EB0F49FA248B710048" box="[1117,1300,1519,1545]" italics="true" pageId="144" pageNumber="145">chrysochersonus</emphasis>
Baker, 1995
</taxonomicName>
known from further south in south-east Asia (Cameron Highlands,
<collectingCountry id="9195767AACC943EB087AF9D88DB2006C" box="[878,983,1555,1581]" name="Malaysia" pageId="144" pageNumber="145">Malaysia</collectingCountry>
).
</paragraph>
<paragraph id="E93D36EAACC943EB0BD3F9FC8AEC0144" blockId="144.[151,1437,150,1941]" pageId="144" pageNumber="145">
Bingham collected extensively in
<collectingCountry id="9195767AACC943EB095FF9FC8CF20010" box="[587,663,1591,1617]" name="Myanmar" pageId="144" pageNumber="145">Burma</collectingCountry>
(now
<collectingCountry id="9195767AACC943EB09F6F9FC8D360010" box="[738,851,1591,1617]" name="Myanmar" pageId="144" pageNumber="145">Myanmar</collectingCountry>
), writing in 1897 that the majority of his collecting in
<collectingCountry id="9195767AACC943EB0BA3F9908F2D0034" box="[183,328,1627,1653]" name="India" pageId="144" pageNumber="145">British India</collectingCountry>
was carried out during the preceding 12 years in
<collectingCountry id="9195767AACC943EB0885F9908DB80034" box="[913,989,1627,1653]" name="Myanmar" pageId="144" pageNumber="145">Burma</collectingCountry>
and Tenasserim (
<bibRefCitation id="8D134B1BACC943EB0FBEF9908B3C0034" author="Bingham, C. T." box="[1194,1369,1627,1653]" pageId="144" pageNumber="145" refId="ref87688" refString="Bingham, C. T. (1897) Hymenoptera. Vol. I. Wasps and Bees. British Fauna of India. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1897">Bingham 1897</bibRefCitation>
: xiii). He described other bee species from the Tenasserim region (e.g.
<taxonomicName id="2E824D69ACC943EB0862F9B48B7C00D8" authority="Bingham, 1897" authorityName="Bingham" authorityYear="1897" box="[886,1305,1663,1689]" class="Insecta" family="Apidae" genus="Anthophora" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="fraterna">
<emphasis id="DBF6EAF8ACC943EB0862F9B48A0500D8" box="[886,1120,1663,1689]" italics="true" pageId="144" pageNumber="145">Anthophora fraterna</emphasis>
<bibRefCitation id="8D134B1BACC943EB0F7CF9B48B7C00D8" author="Bingham, C. T." box="[1128,1305,1663,1689]" pageId="144" pageNumber="145" refId="ref87688" refString="Bingham, C. T. (1897) Hymenoptera. Vol. I. Wasps and Bees. British Fauna of India. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1897">Bingham, 1897</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2E824D69ACC943EB0E32F9B48FB100FC" authority="Bingham, 1897" authorityName="Bingham" authorityYear="1897" class="Insecta" family="Megachilidae" genus="Megachile" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="elizabethae">
<emphasis id="DBF6EAF8ACC943EB0E32F9B48F7200FC" italics="true" pageId="144" pageNumber="145">Megachile elizabethae</emphasis>
<bibRefCitation id="8D134B1BACC943EB0A35F9688FB100FC" author="Bingham, C. T." box="[289,468,1699,1725]" pageId="144" pageNumber="145" refId="ref87688" refString="Bingham, C. T. (1897) Hymenoptera. Vol. I. Wasps and Bees. British Fauna of India. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1897">Bingham, 1897</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="2E824D69ACC943EB0AF7F9688DD100FC" authority="(Bingham, 1897))" authorityName="" baseAuthorityName="Bingham" baseAuthorityYear="1897" box="[483,948,1699,1725]" class="Insecta" family="Megachilidae" genus="Protosmia" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="burmanica">
<emphasis id="DBF6EAF8ACC943EB0AF7F9688CBF00FC" box="[483,730,1699,1725]" italics="true" pageId="144" pageNumber="145">Protosmia burmanica</emphasis>
(
<bibRefCitation id="8D134B1BACC943EB09F9F9688DC500FC" author="Bingham, C. T." box="[749,928,1699,1725]" pageId="144" pageNumber="145" refId="ref87688" refString="Bingham, C. T. (1897) Hymenoptera. Vol. I. Wasps and Bees. British Fauna of India. Taylor and Francis, London, 579 pp. https: // doi. org / 10.5962 / bhl. title. 10364" type="book" year="1897">Bingham, 1897</bibRefCitation>
))
</taxonomicName>
, and mentioned numerous species records from this region for other bees, but none for
<taxonomicName id="2E824D69ACC943EB096BF90C8CBB00A0" box="[639,734,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="144" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="DBF6EAF8ACC943EB096BF90C8CBB00A0" box="[639,734,1735,1761]" italics="true" pageId="144" pageNumber="145">Andrena</emphasis>
</taxonomicName>
. His collection was split between the NHMUK and the ZMHB, with the latter containing the specimens described here in a box of undetermined material.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACC943EB0BD3F8C48F0A0130" pageId="144" pageNumber="145" type="etymology">
<paragraph id="E93D36EAACC943EB0BD3F8C48F0A0130" blockId="144.[151,1437,150,1941]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0BD3F8C48F2B0168" bold="true" box="[199,334,1807,1833]" pageId="144" pageNumber="145">Etymology.</emphasis>
From the Tenasserim Hills or Tenasserim Range that broadly run along the border between
<collectingCountry id="9195767AACC943EB0B83F8F88F61010C" box="[151,260,1843,1869]" name="Myanmar" pageId="144" pageNumber="145">Myanmar</collectingCountry>
and
<collectingCountry id="9195767AACC943EB0A29F8F88FC4010C" box="[317,417,1843,1869]" name="Thailand" pageId="144" pageNumber="145">Thailand</collectingCountry>
(the name originally deriving from the Malay
<emphasis id="DBF6EAF8ACC943EB08ADF8F88A51010C" box="[953,1076,1843,1869]" italics="true" pageId="144" pageNumber="145">Tanah Seri</emphasis>
, meaning glowing land). It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACC943EB0BD3F8B18D1401D4" box="[199,881,1914,1941]" pageId="144" pageNumber="145" type="distribution">
<paragraph id="E93D36EAACC943EB0BD3F8B18D1401D4" blockId="144.[151,1437,150,1941]" box="[199,881,1914,1941]" pageId="144" pageNumber="145">
<emphasis id="DBF6EAF8ACC943EB0BD3F8B18F0401D5" bold="true" box="[199,353,1914,1940]" pageId="144" pageNumber="145">Distribution.</emphasis>
Central
<collectingCountry id="9195767AACC943EB0AD5F8B08C4B01D4" box="[449,558,1915,1941]" name="Myanmar" pageId="144" pageNumber="145">Myanmar</collectingCountry>
and north-western
<collectingCountry id="9195767AACC943EB0812F8B08D0801D4" box="[774,877,1915,1941]" name="Thailand" pageId="144" pageNumber="145">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:location id="824F8E227CDEF5EF0A45FDC561EC67B2">
<mods:url id="A06E6E0A69BBA0569416A249C6FD45A4">http://dx.doi.org/10.11646/zootaxa.5483.1.1</mods:url>
</mods:location>
<mods:classification id="37E07C0D50907021854434567AA922B8">journal article</mods:classification>
<mods:identifier id="34E10CEC24A30C93B8A2115910EEE1F4" type="DOI">10.11646/zootaxa.5483.1.1</mods:identifier>
<mods:identifier id="5CC131657B9937C18D23ACD82D0657A9" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="0691EA654D90CAB39C2525F3CACE13D3" type="Zenodo-Dep">13209307</mods:identifier>
<mods:identifier id="4A1525262B361DF6FD7558CF88A87762" type="ZooBank">AF0272DB-5588-411D-9EAE-DED4785BF170</mods:identifier>
</mods:mods>
<treatment id="612B87FCACD943F80B83F8F98FD703A4" LSID="urn:lsid:plazi:treatment:612B87FCACD943F80B83F8F98FD703A4" httpUri="http://treatment.plazi.org/id/612B87FCACD943F80B83F8F98FD703A4" lastPageId="131" lastPageNumber="132" pageId="128" pageNumber="129">
<subSubSection id="A1986561ACD943FB0B83F8F98CE2010C" box="[151,647,1842,1869]" pageId="128" pageNumber="129" type="nomenclature">
<paragraph id="E93D36EAACD943FB0B83F8F98CE2010C" blockId="128.[151,647,1842,1905]" box="[151,647,1842,1869]" pageId="128" pageNumber="129">
<heading id="B2758186ACD943FB0B83F8F98CE2010C" bold="true" box="[151,647,1842,1869]" fontSize="11" level="2" pageId="128" pageNumber="129" reason="4">
<emphasis id="DBF6EAF8ACD943FB0B83F8F98CE2010C" bold="true" box="[151,647,1842,1869]" pageId="128" pageNumber="129">
<taxonomicName id="2E824D69ACD943FB0B83F8F98C6D010C" authorityName="WOOD" authorityYear="2024" box="[151,520,1842,1869]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="dacia" status="sp. nov.">
<emphasis id="DBF6EAF8ACD943FB0B83F8F98F64010C" bold="true" box="[151,257,1842,1869]" italics="true" pageId="128" pageNumber="129">Andrena</emphasis>
(
<emphasis id="DBF6EAF8ACD943FB0A06F8F88FD3010C" bold="true" box="[274,438,1842,1869]" italics="true" pageId="128" pageNumber="129">incertae sedis</emphasis>
)
<emphasis id="DBF6EAF8ACD943FB0AD3F8F98C6D010C" bold="true" box="[455,520,1842,1869]" italics="true" pageId="128" pageNumber="129">dacia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783ACD943FB0904F8F88CE2010C" box="[528,647,1843,1869]" pageId="128" pageNumber="129" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD943FB0B83F89C8FD50130" box="[151,432,1879,1905]" pageId="128" pageNumber="129" type="description">
<paragraph id="E93D36EAACD943FB0B83F89C8FD50130" blockId="128.[151,647,1842,1905]" box="[151,432,1879,1905]" pageId="128" pageNumber="129">
<figureCitation id="71B92A6FACD943FB0B83F89C8FCE0130" box="[151,427,1879,1905]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="128" pageNumber="129">Figures 79AC; E; GH</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACD943F90B83F8558B68075E" lastPageId="130" lastPageNumber="131" pageId="128" pageNumber="129" type="materials_examined">
<paragraph id="E93D36EAACD943FB0B83F8558A7F019E" blockId="128.[151,1437,1950,2015]" pageId="128" pageNumber="129">
<materialsCitation id="59EA3CB7ACD943FB0B83F8558A7F019E" collectingDate="1954-04-06" collectingDateMax="1996-05-01" collectingDateMin="1954-04-06" collectionCode="TJWC" collectorName="X. Scobiola &amp; L. Rakosy" country="Romania" location="Stefanesti" pageId="128" pageNumber="129" specimenCount="4" specimenCount-female="4" stateProvince="Ilfov" typeStatus="holotype">
<emphasis id="DBF6EAF8ACD943FB0B83F8558FB801F9" bold="true" box="[151,477,1950,1976]" pageId="128" pageNumber="129">
<typeStatus id="36398848ACD943FB0B83F8558F5F01F9" box="[151,314,1950,1976]" pageId="128" pageNumber="129" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AACD943FB0A5CF8558FBD01F9" box="[328,472,1950,1976]" name="Romania" pageId="128" pageNumber="129">ROMANIA</collectingCountry>
:
</emphasis>
<location id="EC5D6031ACD943FB0AF2F86B8C3101FA" LSID="urn:lsid:plazi:treatment:612B87FCACD943F80B83F8F98FD703A4:EC5D6031ACD943FB0AF2F86B8C3101FA" box="[486,596,1952,1979]" country="Romania" name="Stefanesti" pageId="128" pageNumber="129" stateProvince="Ilfov">Ștefănești</location>
jud.
<collectingRegion id="2B46F808ACD943FB0984F8548CAD01F8" box="[656,712,1951,1977]" country="Romania" name="Ilfov" pageId="128" pageNumber="129">Ilfov</collectingRegion>
, 44.5356
<superScript id="1EF79BA2ACD943FB083BF8548D2A01F8" attach="right" box="[815,847,1951,1977]" fontSize="6" pageId="128" pageNumber="129">oN</superScript>
, 26.1968
<superScript id="1EF79BA2ACD943FB08A2F8548DB701F8" attach="right" box="[950,978,1951,1977]" fontSize="6" pageId="128" pageNumber="129">oE</superScript>
,
<date id="9D3C102AACD943FB08CBF8558A2F01F8" box="[991,1098,1950,1977]" pageId="128" pageNumber="129" value="1954-04-06">
<collectingDate id="8D78E9C2ACD943FB08CBF8558A2F01F8" box="[991,1098,1950,1977]" pageId="128" pageNumber="129" value="1954-04-06">6.iv.1954</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACD943FB0F43F8548A0901F8" box="[1111,1132,1951,1977]" count="1" pageId="128" pageNumber="129" type="female"></specimenCount>
, leg.
<collectorName id="4477533CACD943FB0FB9F8548B5301F8" box="[1197,1334,1951,1977]" pageId="128" pageNumber="129">X. Scobiola</collectorName>
, OÖLM. Schwarzmeer-Kuste, Vama Veche,
<date id="9D3C102AACD943FB0934F8088CE7019C" box="[544,642,1987,2013]" pageId="128" pageNumber="129" value="1996-05-01">
<collectingDate id="8D78E9C2ACD943FB0934F8088CE7019C" box="[544,642,1987,2013]" pageId="128" pageNumber="129" value="1996-05-01">1.v.1996</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACD943FB099AF8088CD7019C" box="[654,690,1987,2013]" count="3" pageId="128" pageNumber="129" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CACD943FB09FBF8088D06019C" box="[751,867,1987,2013]" pageId="128" pageNumber="129">L. Rákosy</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FACD943FB08DFF80F8A73019E" box="[971,1046,1988,2015]" pageId="128" pageNumber="129">TJWC</collectionCode>
.
</materialsCitation>
</paragraph>
<caption id="BDFD6662ACD843FA0B83F85C8DEB01B6" ID-DOI="http://doi.org/10.5281/zenodo.13209483" ID-Zenodo-Dep="13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="129" pageNumber="130" startId="129.[151,250,1943,1967]" targetBox="[214,1373,181,1919]" targetPageId="129" targetType="figure">
<paragraph id="E93D36EAACD843FA0B83F85C8DEB01B6" blockId="129.[151,1436,1943,2040]" pageId="129" pageNumber="130">
<emphasis id="DBF6EAF8ACD843FA0B83F85C8F7A01EE" bold="true" box="[151,287,1943,1967]" pageId="129" pageNumber="130">FIGURE 79.</emphasis>
<taxonomicName id="2E824D69ACD843FA0A31F85C8C3D01EE" authorityName="WOOD" authorityYear="2024" box="[293,600,1943,1967]" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="dacia" status="sp. nov.">
<emphasis id="DBF6EAF8ACD843FA0A31F85C8F1E01EE" box="[293,379,1943,1967]" italics="true" pageId="129" pageNumber="130">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8ACD843FA0934F85C8C3D01EE" box="[544,600,1943,1967]" italics="true" pageId="129" pageNumber="130">dacia</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8ACD843FA094AF8538CA001F1" bold="true" box="[606,709,1944,1968]" pageId="129" pageNumber="130">
<taxonomicNameLabel id="C0C55783ACD843FA094AF8538CA001F1" box="[606,709,1944,1968]" pageId="129" pageNumber="130" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view.
<taxonomicName id="2E824D69ACD843FA0F1BF8708F1401B9" authority="Scheuchl &amp; Gusenleitner, 2009" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" class="Insecta" family="Andrenidae" genus="Andrena" isHigherUncertain="true" kingdom="Animalia" order="Hymenoptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACD843FA0F1BF8708A000192" box="[1039,1125,1979,2003]" italics="true" pageId="129" pageNumber="130">Andrena</emphasis>
(incertae sedis)
<emphasis id="DBF6EAF8ACD843FA0E1FF8708B5C0192" box="[1291,1337,1979,2003]" italics="true" pageId="129" pageNumber="130">elsei</emphasis>
<bibRefCitation id="8D134B1BACD843FA0E2BF8708F1401B9" author="Scheuchl, E. &amp; Gusenleitner, F." pageId="129" pageNumber="130" pagination="947 - 962" refId="ref90737" refString="Scheuchl, E. &amp; Gusenleitner, F. (2009) Andrena (Margandrena) elsei nov. sp., eine neue turkische Sandbienenart (Hymenoptera, Apidae, Andreninae). Linzer biologische Beitrage, 41, 947 - 962." type="journal article" year="2009">Scheuchl &amp; Gusenleitner, 2009</bibRefCitation>
</taxonomicName>
female. D. Face, frontal view; F. Galea, frontal view.
</paragraph>
</caption>
<paragraph id="E93D36EAACDB43F90BD3FF5D8B68075E" blockId="130.[151,1437,150,2013]" pageId="130" pageNumber="131">
<materialsCitation id="59EA3CB7ACDB43F90BD3FF5D8A0806F2" box="[199,1133,150,179]" collectionCode="TJWC" country="Romania" location="ROMANIA" pageId="130" pageNumber="131" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="DBF6EAF8ACDB43F90BD3FF5D8C7306F1" bold="true" box="[199,534,150,176]" pageId="130" pageNumber="131">
<typeStatus id="36398848ACDB43F90BD3FF5D8F0A06F1" box="[199,367,150,176]" pageId="130" pageNumber="131" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="9195767AACDB43F90A94FF5D8C7506F1" box="[384,528,150,176]" name="Romania" pageId="130" pageNumber="131">ROMANIA</collectingCountry>
:
</emphasis>
same information as
<typeStatus id="36398848ACDB43F9080FFF5C8DE506F0" box="[795,896,151,177]" pageId="130" pageNumber="131" type="holotype">holotype</typeStatus>
,
<specimenCount id="FF84FD63ACDB43F90884FF5C8DD106F0" box="[912,948,151,177]" count="3" pageId="130" pageNumber="131" type="female">3♀</specimenCount>
, OÖLM/
<collectionCode id="8F93AE2FACDB43F90F37FF538A0806F2" box="[1059,1133,152,179]" pageId="130" pageNumber="131">TJWC</collectionCode>
</materialsCitation>
;
<materialsCitation id="59EA3CB7ACDB43F90F6AFF5C8EAA0694" collectorName="Mogosoaia" country="Romania" location="Ilfov" pageId="130" pageNumber="131" specimenCount="1" stateProvince="Ilfov" typeStatus="paratype">
<collectorName id="4477533CACDB43F90F6AFF5C8A9F06F0" box="[1150,1274,151,177]" pageId="130" pageNumber="131">Mogosoaia</collectorName>
[
<collectorName id="4477533CACDB43F90E19FF538BF706F2" box="[1293,1426,152,179]" pageId="130" pageNumber="131">Mogoșoaia</collectorName>
],
<collectingRegion id="2B46F808ACDB43F90B83FF708EAA0694" box="[151,207,187,213]" country="Romania" name="Ilfov" pageId="130" pageNumber="131">Ilfov</collectingRegion>
</materialsCitation>
,
<materialsCitation id="59EA3CB7ACDB43F90BC8FF718AB90694" box="[220,1244,186,215]" collectingDate="1954-04-15" collectorName="X. Scobiola &amp; Caldarusani J." country="Romania" location="Gradistea" municipality="Caldarusani" pageId="130" pageNumber="131" specimenCount="2" specimenCount-female="2" stateProvince="Ilfov" typeStatus="paratype">
<date id="9D3C102AACDB43F90BC8FF718F310694" box="[220,340,186,213]" pageId="130" pageNumber="131" value="1954-04-15">
<collectingDate id="8D78E9C2ACDB43F90BC8FF718F310694" box="[220,340,186,213]" pageId="130" pageNumber="131" value="1954-04-15">15.iv.1954</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACDB43F90A75FF708FE30694" box="[353,390,187,213]" count="2" pageId="130" pageNumber="131" type="female">2♀</specimenCount>
, leg.
<collectorName id="4477533CACDB43F90AD2FF708C2A0694" box="[454,591,187,213]" pageId="130" pageNumber="131">X. Scobiola</collectorName>
, OÖLM;
<collectorName id="4477533CACDB43F909ABFF708D050694" box="[703,864,187,213]" pageId="130" pageNumber="131">Caldarusani J.</collectorName>
<collectingRegion id="2B46F808ACDB43F9087DFF708DFA0694" box="[873,927,187,213]" country="Romania" name="Ilfov" pageId="130" pageNumber="131">Ilfov</collectingRegion>
[
<collectingMunicipality id="0959AC90ACDB43F908BAFF778A520696" box="[942,1079,188,215]" pageId="130" pageNumber="131">Căldărușani</collectingMunicipality>
by
<location id="EC5D6031ACDB43F90F71FF708AB30694" LSID="urn:lsid:plazi:treatment:612B87FCACD943F80B83F8F98FD703A4:EC5D6031ACDB43F90F71FF708AB30694" box="[1125,1238,187,213]" country="Romania" municipality="Caldarusani" name="Gradistea" pageId="130" pageNumber="131" stateProvince="Ilfov">Grădiştea</location>
]
</materialsCitation>
,
<materialsCitation id="59EA3CB7ACDB43F90FFDFF718AD706B8" collectingDate="1957-03-24" collectorName="X. Scobiola &amp; Decembrie" country="Romania" location="Reg. Buc." pageId="130" pageNumber="131" specimenCount="1" specimenCount-female="1" stateProvince="Bucuresti" typeStatus="paratype">
<date id="9D3C102AACDB43F90FFDFF718B000694" box="[1257,1381,186,213]" pageId="130" pageNumber="131" value="1957-03-24">
<collectingDate id="8D78E9C2ACDB43F90FFDFF718B000694" box="[1257,1381,186,213]" pageId="130" pageNumber="131" value="1957-03-24">24.iii.1957</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACDB43F90E66FF708BF00694" box="[1394,1429,187,213]" count="1" pageId="130" pageNumber="131" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CACDB43F90BD2FF148F2906B8" box="[198,332,223,249]" pageId="130" pageNumber="131">X. Scobiola</collectorName>
, OÖLM; 30
<collectorName id="4477533CACDB43F90ACAFF148C3906B8" box="[478,604,223,249]" pageId="130" pageNumber="131">Decembrie</collectorName>
,
<location id="EC5D6031ACDB43F90972FF148CB106B8" LSID="urn:lsid:plazi:treatment:612B87FCACD943F80B83F8F98FD703A4:EC5D6031ACDB43F90972FF148CB106B8" box="[614,724,223,249]" country="Romania" name="Reg. Buc." pageId="130" pageNumber="131" stateProvince="Bucuresti">Reg. Buc.</location>
[around
<collectingRegion id="2B46F808ACDB43F90823FF148DCF06B8" box="[823,938,223,249]" country="Romania" name="Bucuresti" pageId="130" pageNumber="131">Bucharest</collectingRegion>
, exact location unclear]
</materialsCitation>
,
<materialsCitation id="59EA3CB7ACDB43F90FA9FF158D6D075C" collectingDate="1954-03-15" collectorName="X. Scobiola" country="Romania" location="Vama Veche" municipality="Schwarzmeer-Kuste" pageId="130" pageNumber="131" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<date id="9D3C102AACDB43F90FA9FF158B5D06B8" box="[1213,1336,222,249]" pageId="130" pageNumber="131" value="1954-03-15">
<collectingDate id="8D78E9C2ACDB43F90FA9FF158B5D06B8" box="[1213,1336,222,249]" pageId="130" pageNumber="131" value="1954-03-15">15.iii.1954</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACDB43F90E57FF148B0306B8" box="[1347,1382,223,249]" count="1" pageId="130" pageNumber="131" type="female">1♀</specimenCount>
, leg.
<collectorName id="4477533CACDB43F90B83FEC88F7B075C" box="[151,286,259,285]" pageId="130" pageNumber="131">X. Scobiola</collectorName>
, OÖLM;
<collectingMunicipality id="0959AC90ACDB43F90A9FFEC88C15075C" box="[395,624,259,285]" pageId="130" pageNumber="131">Schwarzmeer-Kuste</collectingMunicipality>
,
<location id="EC5D6031ACDB43F9096FFEC88D6D075C" LSID="urn:lsid:plazi:treatment:612B87FCACD943F80B83F8F98FD703A4:EC5D6031ACDB43F9096FFEC88D6D075C" box="[635,776,259,285]" country="Romania" municipality="Schwarzmeer-Kuste" name="Vama Veche" pageId="130" pageNumber="131">Vama Veche</location>
</materialsCitation>
,
<materialsCitation id="59EA3CB7ACDB43F90807FEC88B6C075E" box="[787,1289,259,287]" collectingDate="1996-05-01" collectionCode="TJWC" collectorName="L. Rakosy" country="Romania" pageId="130" pageNumber="131" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<date id="9D3C102AACDB43F90807FEC88D13075C" box="[787,886,259,285]" pageId="130" pageNumber="131" value="1996-05-01">
<collectingDate id="8D78E9C2ACDB43F90807FEC88D13075C" box="[787,886,259,285]" pageId="130" pageNumber="131" value="1996-05-01">1.v.1996</collectingDate>
</date>
,
<specimenCount id="FF84FD63ACDB43F90895FEC88DC0075C" box="[897,933,259,285]" count="3" pageId="130" pageNumber="131" type="female">3♀</specimenCount>
, leg.
<collectorName id="4477533CACDB43F908F7FEC88A32075C" box="[995,1111,259,285]" pageId="130" pageNumber="131">L. Rákosy</collectorName>
, OÖLM/
<collectionCode id="8F93AE2FACDB43F90FAAFECF8B6C075E" box="[1214,1289,260,287]" pageId="130" pageNumber="131">TJWC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDB43F90BD3FEED8FEF0314" pageId="130" pageNumber="131" type="description">
<paragraph id="E93D36EAACDB43F90BD3FEED8BE704E8" blockId="130.[151,1437,150,2013]" pageId="130" pageNumber="131">
<emphasis id="DBF6EAF8ACDB43F90BD3FEED8FA40700" bold="true" box="[199,449,294,321]" pageId="130" pageNumber="131">Description: Female:</emphasis>
Body length:
<quantity id="2E7A9B0FACDB43F9094FFEEC8CD20700" box="[603,695,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="130" pageNumber="131" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
(
<figureCitation id="71B92A6FACDB43F909D7FEEC8D2D0700" box="[707,840,295,321]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79A</figureCitation>
).
<emphasis id="DBF6EAF8ACDB43F9084DFEEC8DFA0700" box="[857,927,295,321]" italics="true" pageId="130" pageNumber="131">Head:</emphasis>
Dark, 1.2 times wider than long (
<figureCitation id="71B92A6FACDB43F90E19FEEC8BF50700" box="[1293,1424,295,321]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79C</figureCitation>
). Clypeus weakly domed, flattened over majority of its area, densely punctate laterally, punctures separated by 0.51 puncture diameters, medially with a broad impunctate longitudinal mid-line, as much as 78 puncture diameters wide; underlying surface with strong granular shagreen, weakly shining. Process of labrum trapezoidal, 3 times wider than long, surface with irregular latitudinal striations. Galea with outer surface covered with strong granular microreticulation, dull (
<figureCitation id="71B92A6FACDB43F90AB7FE108C4D07B4" box="[419,552,475,501]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79E</figureCitation>
). Gena slightly exceeding width of compound eye; ocelloccipital distance 1.5 times diameter of lateral ocellus. Foveae dorsally narrow, dorsally occupying ¼ of space between compound eye and lateral ocellus, slightly narrower than width of flagellum, uniformly wide, ventrally extending to slightly below level of antennal insertions; foveae filled with dark brown hairs. Face, gena, and scape covered with black hairs, vertex with mixture of black and light brown hairs (fading to whitish); hairs not equalling length of scape. Antennae dark basally, A512 ventrally lightened by presence of silvery scales; A3 exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="E93D36EAACDB43F90BD3FD788DAC0274" blockId="130.[151,1437,150,2013]" pageId="130" pageNumber="131">
<emphasis id="DBF6EAF8ACDB43F90BD3FD788F2D048D" box="[199,328,691,716]" italics="true" pageId="130" pageNumber="131">Mesosoma:</emphasis>
Scutum and scutellum finely shagreened, weakly shining, sparsely and irregularly punctate, punctures separated by 0.54 puncture diameters, punctures sparsest medially (
<figureCitation id="71B92A6FACDB43F90F1BFD1C8AF104B0" box="[1039,1172,727,753]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79B</figureCitation>
). Pronotum with strong humeral angle, surface with weak longitudinal striations. Mesepisternum and dorsolateral parts of propodeum with fine granular microreticulation, weakly shining; propodeal triangle very weakly indicated by obscure fine carinae laterally, internal surface almost structurally identical to surrounding propodeum, with weak, short, and obscure rugae basally (
<figureCitation id="71B92A6FACDB43F90A29FCAC8FA105C0" box="[317,452,871,897]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79G</figureCitation>
). Mesepisternum with long black hairs, longest exceeding length of scape, scutum and scutellum with shorter light brown hair (fading to whitish). Propodeal corbicula incomplete, dorsal fringe composed of black plumose hairs, internal surface with short brownish simple hairs. Legs dark.Flocculus incomplete, composed of short whitish hairs; femoral and tibial scopae composed of simple whitish hairs, tibial scopae basally with brown hairs adjacent to basitibial plate. Remaining leg pubescence brownish. Hind tarsal claws with inner tooth. Wings hyaline, stigma dark brown, venation brown-orange, nervulus postfurcal.
</paragraph>
<paragraph id="E93D36EAACDB43F90BD3FBF48E940370" blockId="130.[151,1437,150,2013]" pageId="130" pageNumber="131">
<emphasis id="DBF6EAF8ACDB43F90BD3FBF48F200219" box="[199,325,1087,1112]" italics="true" pageId="130" pageNumber="131">Metasoma:</emphasis>
Tergal discs dark, marginal areas lightened brownish over majority of area (
<figureCitation id="71B92A6FACDB43F90FACFBF48B240218" box="[1208,1345,1087,1113]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79H</figureCitation>
). Tergal discs finely shagreened, shining, sculpture strongest basally and laterally; surface with irregular obscure punctures, punctures separated by 0.52 puncture diameters. T14 discs laterally with short and erect light brownish hairs, on T24 coalescing to form weak, widely separated apical hair-fringes, not obscuring underlying surface. Apical fringe of T5 and hairs flanking pygidial plate dark brown. Pygidial plate broad, apically rounded, lateral margin slightly thickened, internal surface densely punctate, punctures separated by 0.5 puncture diameters, interspaces raised and shining.
</paragraph>
<paragraph id="E93D36EAACDB43F90BD3FAF08FEF0314" blockId="130.[151,1437,150,2013]" box="[199,394,1339,1365]" pageId="130" pageNumber="131">
<emphasis id="DBF6EAF8ACDB43F90BD3FAF08F680314" bold="true" box="[199,269,1339,1365]" pageId="130" pageNumber="131">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDB43F80BD3FA958D5A048C" lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131" type="diagnosis">
<paragraph id="E93D36EAACDB43F90BD3FA958DAC0034" blockId="130.[151,1437,150,2013]" pageId="130" pageNumber="131">
<emphasis id="DBF6EAF8ACDB43F90BD3FA958F270339" bold="true" box="[199,322,1374,1400]" pageId="130" pageNumber="131">Diagnosis.</emphasis>
<taxonomicName id="2E824D69ACDB43F90A5DFA948F8A0338" authorityName="WOOD" authorityYear="2024" box="[329,495,1375,1401]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDB43F90A5DFA948F8A0338" box="[329,495,1375,1401]" italics="true" pageId="130" pageNumber="131">Andrena dacia</emphasis>
</taxonomicName>
can be recognised as part of the currently undescribed subgenus best referred to as the
<emphasis id="DBF6EAF8ACDB43F90BD5FA488F5003DC" box="[193,309,1411,1437]" italics="true" pageId="130" pageNumber="131">crocusella</emphasis>
-group (
<bibRefCitation id="8D134B1BACDB43F90A9AFA488C1703DC" author="Pisanty, G. &amp; Scheuchl, E. &amp; Martin, T. &amp; Cardinal, S. &amp; Wood, T. J." box="[398,626,1411,1437]" pageId="130" pageNumber="131" pagination="1 - 109" refId="ref90052" refString="Pisanty, G., Scheuchl, E., Martin, T., Cardinal, S. &amp; Wood, T. J. (2022 a) Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant. Zootaxa, 5185 (1), 1 - 109. https: // doi. org / 10.11646 / zootaxa. 5185.1.1" type="journal article" year="2022">
Pisanty
<emphasis id="DBF6EAF8ACDB43F90AFEFA488C7903DC" box="[490,540,1411,1437]" italics="true" pageId="130" pageNumber="131">et al</emphasis>
. 2022a
</bibRefCitation>
; b) due to the pronotum with clear humeral angle (in the closely related subgenus
<taxonomicName id="2E824D69ACDB43F90A1CFA6C8FE60380" authorityName="Hedicke" authorityYear="1933" box="[264,387,1447,1473]" pageId="130" pageNumber="131" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8ACDB43F90A1CFA6C8FE60380" box="[264,387,1447,1473]" italics="true" pageId="130" pageNumber="131">Euandrena</emphasis>
</taxonomicName>
with the pronotum rounded), the barely indicated propodeal triangle (
<figureCitation id="71B92A6FACDB43F90FB1FA6C8B480380" box="[1189,1325,1447,1473]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79G</figureCitation>
), oviform metasoma (
<figureCitation id="71B92A6FACDB43F90A02FA008FFF03A4" box="[278,410,1483,1509]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79H</figureCitation>
), and long and narrow foveae that remain a consistent width along their length (in
<taxonomicName id="2E824D69ACDB43F90E35FA008BF903A4" authorityName="Hedicke" authorityYear="1933" box="[1313,1436,1483,1509]" pageId="130" pageNumber="150" rank="subGenus" subGenus="Euandrena">
<emphasis id="DBF6EAF8ACDB43F90E35FA008BF903A4" box="[1313,1436,1483,1509]" italics="true" pageId="130" pageNumber="131">Euandrena</emphasis>
</taxonomicName>
typically narrowing further ventrally).
<taxonomicName id="2E824D69ACDB43F9095BFA248C900048" authorityName="WOOD" authorityYear="2024" box="[591,757,1519,1545]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDB43F9095BFA248C900048" box="[591,757,1519,1545]" italics="true" pageId="130" pageNumber="131">Andrena dacia</emphasis>
</taxonomicName>
has dark-marked terga, the head is relatively short, and the mandibular and labial palpi have a regular number of segments (six and four, respectively) that are not noticeably elongate. This places it close to
<taxonomicName id="2E824D69ACDB43F90910F9FC8DA40010" authority="Scheuchl &amp; Gusenleitner, 2009" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[516,961,1591,1617]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F90910F9FC8C320010" box="[516,599,1591,1617]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
<bibRefCitation id="8D134B1BACDB43F90974F9FC8DA40010" author="Scheuchl, E. &amp; Gusenleitner, F." box="[608,961,1591,1617]" pageId="130" pageNumber="131" pagination="947 - 962" refId="ref90737" refString="Scheuchl, E. &amp; Gusenleitner, F. (2009) Andrena (Margandrena) elsei nov. sp., eine neue turkische Sandbienenart (Hymenoptera, Apidae, Andreninae). Linzer biologische Beitrage, 41, 947 - 962." type="journal article" year="2009">Scheuchl &amp; Gusenleitner, 2009</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="9195767AACDB43F908C5F9FC8A420010" box="[977,1063,1591,1617]" name="Greece" pageId="130" pageNumber="131">Greece</collectingCountry>
*, North
<collectingCountry id="9195767AACDB43F90F9DF9FC8B6E0010" box="[1161,1291,1591,1617]" name="Macedonia" pageId="130" pageNumber="131">Macedonia</collectingCountry>
*,
<collectingCountry id="9195767AACDB43F90E36F9FC8BE80010" box="[1314,1421,1591,1617]" name="Romania" pageId="130" pageNumber="131">Romania</collectingCountry>
*, and
<collectingCountry id="9195767AACDB43F90BD2F9908F7D0034" box="[198,280,1627,1653]" name="Turkey" pageId="130" pageNumber="131">Turkey</collectingCountry>
) and
<taxonomicName id="2E824D69ACDB43F90A42F9908C9D0034" authority="Pisanty &amp; Wood, 2022" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[342,760,1627,1653]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDB43F90A42F9908F960034" box="[342,499,1627,1653]" italics="true" pageId="130" pageNumber="131">A. ornithogali</emphasis>
<bibRefCitation id="8D134B1BACDB43F90AEEF9908C9D0034" author="Pisanty, G. &amp; Scheuchl, E. &amp; Martin, T. &amp; Cardinal, S. &amp; Wood, T. J." box="[506,760,1627,1653]" pageId="130" pageNumber="131" pagination="1 - 109" refId="ref90052" refString="Pisanty, G., Scheuchl, E., Martin, T., Cardinal, S. &amp; Wood, T. J. (2022 a) Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant. Zootaxa, 5185 (1), 1 - 109. https: // doi. org / 10.11646 / zootaxa. 5185.1.1" type="journal article" year="2022">Pisanty &amp; Wood, 2022</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="9195767AACDB43F90812F9908D230034" box="[774,838,1627,1653]" name="Israel" pageId="130" pageNumber="131">Israel</collectingCountry>
and
<collectingCountry id="9195767AACDB43F90869F9908DD80034" box="[893,957,1627,1653]" name="Syria" pageId="130" pageNumber="131">Syria</collectingCountry>
).
</paragraph>
<paragraph id="E93D36EAACDB43F80BD3F9B48CB3075C" blockId="130.[151,1437,150,2013]" lastBlockId="131.[151,1437,151,1511]" lastPageId="131" lastPageNumber="132" pageId="130" pageNumber="131">
Diagnosis must also take into account biogeographical context. Only two of the 10 species within this group (
<bibRefCitation id="8D134B1BACDB43F90BFDF9688FB500FC" author="Pisanty, G. &amp; Scheuchl, E. &amp; Martin, T. &amp; Cardinal, S. &amp; Wood, T. J." box="[233,464,1699,1725]" pageId="130" pageNumber="131" pagination="1 - 109" refId="ref90052" refString="Pisanty, G., Scheuchl, E., Martin, T., Cardinal, S. &amp; Wood, T. J. (2022 a) Twenty-five new species of mining bees (Hymenoptera: Andrenidae: Andrena) from Israel and the Levant. Zootaxa, 5185 (1), 1 - 109. https: // doi. org / 10.11646 / zootaxa. 5185.1.1" type="journal article" year="2022">
Pisanty
<emphasis id="DBF6EAF8ACDB43F90A53F9688F1F00FC" box="[327,378,1699,1725]" italics="true" pageId="130" pageNumber="131">et al</emphasis>
. 2022a
</bibRefCitation>
) have previously been recorded from continental Europe (i.e. not including
<collectingCountry id="9195767AACDB43F90E2FF9688BF500FC" box="[1339,1424,1699,1725]" name="Cyprus" pageId="130" pageNumber="131">Cyprus</collectingCountry>
), specifically
<taxonomicName id="2E824D69ACDB43F90A37F90C8C1D00A0" authority="Grunwaldt, 1976" authorityName="Grunwaldt" authorityYear="1976" box="[291,632,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="grossella">
<emphasis id="DBF6EAF8ACDB43F90A37F90C8FC900A0" box="[291,428,1735,1761]" italics="true" pageId="130" pageNumber="131">A. grossella</emphasis>
Grünwaldt, 1976
</taxonomicName>
and
<taxonomicName id="2E824D69ACDB43F909A2F90C8D9D00A0" authority="Warncke, 1965" authorityName="Warncke" authorityYear="1965" box="[694,1016,1734,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="muscaria">
<emphasis id="DBF6EAF8ACDB43F909A2F90C8D2400A1" box="[694,833,1735,1761]" italics="true" pageId="130" pageNumber="131">A. muscaria</emphasis>
<bibRefCitation id="8D134B1BACDB43F9085FF90C8D9D00A0" author="Warncke, K." box="[843,1016,1734,1761]" pageId="130" pageNumber="131" pagination="27 - 76" refId="ref91663" refString="Warncke, K. (1965) Beitrag zur Kenntnis der Bienengattung Andrena Fabricius in Griechenland. Beitrage zur Entomologie, 15, 27 - 76." type="journal article" year="1965">Warncke, 1965</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="9195767AACDB43F90F50F90C8AF200A0" box="[1092,1175,1735,1761]" name="Greece" pageId="130" pageNumber="131">Greece</collectingCountry>
. Surprisingly,
<taxonomicName id="2E824D69ACDB43F90E2BF90C8BF000A0" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[1343,1429,1735,1761]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F90E2BF90C8BF000A0" box="[1343,1429,1735,1761]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
</taxonomicName>
, a species previously known only from the Taurus mountains of
<collectingCountry id="9195767AACDB43F90861F9208DA30144" box="[885,966,1771,1797]" name="Turkey" pageId="130" pageNumber="131">Turkey</collectingCountry>
, can now be reported from south-eastern Europe including southern
<collectingCountry id="9195767AACDB43F90ADEF8C48C4A0168" box="[458,559,1807,1833]" name="Romania" pageId="130" pageNumber="131">Romania</collectingCountry>
(see below).
<taxonomicName id="2E824D69ACDB43F909DCF8C48D0B0168" authorityName="WOOD" authorityYear="2024" box="[712,878,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDB43F909DCF8C48D0B0168" box="[712,878,1807,1833]" italics="true" pageId="130" pageNumber="131">Andrena dacia</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="2E824D69ACDB43F90F63F8C48AAF0168" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[1143,1226,1807,1833]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F90F63F8C48AAF0168" box="[1143,1226,1807,1833]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
</taxonomicName>
due to the smaller body length of
<quantity id="2E7A9B0FACDB43F90A52F8F88FC1010C" box="[326,420,1843,1869]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="130" pageNumber="131" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
(
<taxonomicName id="2E824D69ACDB43F90AA2F8F88C6C010C" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[438,521,1843,1869]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F90AA2F8F88C6C010C" box="[438,521,1843,1869]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
</taxonomicName>
with body length of
<quantity id="2E7A9B0FACDB43F909EFF8F88D1D010C" box="[763,888,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="130" pageNumber="131" unit="mm" value="10.5" valueMax="11.0" valueMin="10.0">1011 mm</quantity>
), the shorter ocelloccipital distance which is 1.5 times the diameter of a lateral ocellus (in
<taxonomicName id="2E824D69ACDB43F9097FF89C8CD80130" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[619,701,1879,1905]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F9097FF89C8CD80130" box="[619,701,1879,1905]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
</taxonomicName>
with the ocelloccipital distance equalling 2 times the diameter of a lateral ocellus), the surface of the galea which is covered with strong granular microreticulation and dull (
<figureCitation id="71B92A6FACDB43F90E45F8B08EAD01F8" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79E</figureCitation>
; in
<taxonomicName id="2E824D69ACDB43F90BE4F8548F2701F8" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[240,322,1951,1977]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="130" pageNumber="131" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDB43F90BE4F8548F2701F8" box="[240,322,1951,1977]" italics="true" pageId="130" pageNumber="131">A. elsei</emphasis>
</taxonomicName>
with the surface of the galea finely shagreened and shining,
<figureCitation id="71B92A6FACDB43F908FEF8548A0C01F8" box="[1002,1129,1951,1977]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="130" pageNumber="131">Figure 79F</figureCitation>
), and with the clypeus with strong granular shagreen, weakly shining, covered with dense large punctures, laterally separated by 0.51 puncture diameters, medially with a broad impunctate longitudinal mid-line, as much as 78 puncture diameters wide (
<figureCitation id="71B92A6FACDA43F80E45FF5C8EAF0694" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="131" pageNumber="132">Figure 79C</figureCitation>
; in
<taxonomicName id="2E824D69ACDA43F80BE4FF708F270694" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[240,322,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDA43F80BE4FF708F270694" box="[240,322,187,213]" italics="true" pageId="131" pageNumber="132">A. elsei</emphasis>
</taxonomicName>
with the clypeus more finely shagreened, more strongly shining, more densely and evenly punctate, punctures separated by ≤0.5 puncture diameters with exception of narrow impunctate longitudinal mid-line, no more than 3 puncture diameters wide,
<figureCitation id="71B92A6FACDA43F80951FEC88CAF075C" box="[581,714,259,285]" captionStart="FIGURE 79" captionStartId="129.[151,250,1943,1967]" captionTargetBox="[214,1373,181,1919]" captionTargetId="figure-16@129.[214,1373,181,1919]" captionTargetPageId="129" captionText="FIGURE 79. Andrena (incertae sedis) dacia spec. nov. female. A. Habitus, lateral view; B. Scutum, dorsolateral view; C. Face, frontal view; E. Galea, frontal view; G. Propodeum, dorsal view; H. Terga, dorsal view. Andrena (incertae sedis) elsei Scheuchl &amp; Gusenleitner, 2009 female. D. Face, frontal view; F. Galea, frontal view." figureDoi="http://doi.org/10.5281/zenodo.13209483" httpUri="https://zenodo.org/record/13209483/files/figure.png" pageId="131" pageNumber="132">Figure 79D</figureCitation>
).
</paragraph>
<paragraph id="E93D36EAACDA43F80BD3FEEC8D5A048C" blockId="131.[151,1437,151,1511]" pageId="131" pageNumber="132">
Differences from
<taxonomicName id="2E824D69ACDA43F80A98FEEC8C4D0700" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[396,552,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80A98FEEC8C4D0700" box="[396,552,295,321]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
are smaller. The male of
<taxonomicName id="2E824D69ACDA43F80854FEEC8DB80700" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[832,989,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80854FEEC8DB80700" box="[832,989,295,321]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
displays a unique character which is the presence of a short distinct tooth on the gena at the base of the mandible, but unfortunately the male of
<taxonomicName id="2E824D69ACDA43F80E30FE808BE70724" authorityName="WOOD" authorityYear="2024" box="[1316,1410,331,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDA43F80E30FE808BE70724" box="[1316,1410,331,357]" italics="true" pageId="131" pageNumber="132">A. dacia</emphasis>
</taxonomicName>
is not available for comparison. However, separation of
<taxonomicName id="2E824D69ACDA43F809E2FEA48D3607C8" authorityName="WOOD" authorityYear="2024" box="[758,851,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDA43F809E2FEA48D3607C8" box="[758,851,367,393]" italics="true" pageId="131" pageNumber="132">A. dacia</emphasis>
</taxonomicName>
in the female sex is still possible due to the slightly larger body size of
<quantity id="2E7A9B0FACDA43F80A7BFE588FAE07EC" box="[367,459,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="131" pageNumber="132" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
(77.5 mm in
<taxonomicName id="2E824D69ACDA43F80964FE588D6807EC" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[624,781,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80964FE588D6807EC" box="[624,781,403,429]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
), the comparatively narrower process of the labrum that is trapezoidal, 3 times wider than long, surface with irregular latitudinal striations (in
<taxonomicName id="2E824D69ACDA43F80F55FE7C8ABB0790" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[1089,1246,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80F55FE7C8ABB0790" box="[1089,1246,439,465]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
with the process of the labrum rounded rectangular, 4 times wider than long, surface polished and shining), and by the same clypeus structural character, with strong granular shagreen, weakly shining, covered with dense large punctures, laterally separated by 0.51 puncture diameters, medially with a broad impunctate longitudinal mid-line, as much as 78 puncture diameters wide (in
<taxonomicName id="2E824D69ACDA43F80ACBFD8C8C1B0420" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[479,638,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80ACBFD8C8C1B0420" box="[479,638,583,609]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
with the clypeus shagreened and dull, but more evenly punctate over its surface, punctures shallower and separated by 0.51 puncture diameter, with narrower impunctate longitudinal mid-line, no more than 3 puncture diameters wide). The strongly disjunct distribution (southern
<collectingCountry id="9195767AACDA43F80FD6FD448B4204E8" box="[1218,1319,655,681]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
compared to the central Levant in
<taxonomicName id="2E824D69ACDA43F80AB6FD788C5A048C" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[418,575,691,717]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80AB6FD788C5A048C" box="[418,575,691,717]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
) also aids recognition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDA43F80BD3FD1C8F22023C" pageId="131" pageNumber="132" type="discussion">
<paragraph id="E93D36EAACDA43F80BD3FD1C8F22023C" blockId="131.[151,1437,151,1511]" pageId="131" pageNumber="132">
<emphasis id="DBF6EAF8ACDA43F80BD3FD1C8F5E04B0" bold="true" box="[199,315,727,753]" pageId="131" pageNumber="132">Remarks.</emphasis>
Inspection of museum material unexpectedly produced many records of
<taxonomicName id="2E824D69ACDA43F80F67FD1C8AA004B0" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[1139,1221,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDA43F80F67FD1C8AA004B0" box="[1139,1221,727,753]" italics="true" pageId="131" pageNumber="132">A. elsei</emphasis>
</taxonomicName>
from Europe, with the first country records for
<collectingCountry id="9195767AACDA43F80AF4FD308C570554" box="[480,562,763,789]" name="Greece" pageId="131" pageNumber="132">Greece</collectingCountry>
, North
<collectingCountry id="9195767AACDA43F8099FFD308D6F0554" box="[651,778,763,789]" name="Macedonia" pageId="131" pageNumber="132">Macedonia</collectingCountry>
, and
<collectingCountry id="9195767AACDA43F8085EFD308DD60554" box="[842,947,763,789]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
. The Romanian specimens are particularly intriguing, as they were present in undetermined material in the Warncke collection, but they were clearly taken from the Paris museum; they are labelled as part of the Vachal collection. They were collected from the same part of southern
<collectingCountry id="9195767AACDA43F80BEAFCAC8F0605C0" box="[254,355,871,897]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
as
<taxonomicName id="2E824D69ACDA43F80A93FCAC8F8605C0" authorityName="WOOD" authorityYear="2024" box="[391,483,871,897]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDA43F80A93FCAC8F8605C0" box="[391,483,871,897]" italics="true" pageId="131" pageNumber="132">A. dacia</emphasis>
</taxonomicName>
, presumably prior to 1911 as this is the printed date corresponding to the accession of the Vachal collection to the MNHN, since the specimens themselves are undated.
<taxonomicName id="2E824D69ACDA43F80F27FC408AA805E4" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[1075,1229,907,933]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDA43F80F27FC408AA805E4" box="[1075,1229,907,933]" italics="true" pageId="131" pageNumber="132">Andrena elsei</emphasis>
</taxonomicName>
is associated with
<taxonomicName id="2E824D69ACDA43F80B83FC648E820588" box="[151,231,943,969]" class="Liliopsida" family="Iridaceae" genus="Crocus" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8ACDA43F80B83FC648E820588" box="[151,231,943,969]" italics="true" pageId="131" pageNumber="132">Crocus</emphasis>
</taxonomicName>
flowers (
<taxonomicName id="2E824D69ACDA43F80A5BFC648FDE0588" box="[335,443,943,969]" class="Liliopsida" family="Iridaceae" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="family">Iridaceae</taxonomicName>
;
<bibRefCitation id="8D134B1BACDA43F80AD2FC648D7A0588" author="Scheuchl, E. &amp; Gusenleitner, F." box="[454,799,943,969]" pageId="131" pageNumber="132" pagination="947 - 962" refId="ref90737" refString="Scheuchl, E. &amp; Gusenleitner, F. (2009) Andrena (Margandrena) elsei nov. sp., eine neue turkische Sandbienenart (Hymenoptera, Apidae, Andreninae). Linzer biologische Beitrage, 41, 947 - 962." type="journal article" year="2009">Scheuchl &amp; Gusenleitner 2009</bibRefCitation>
; label details from
<collectingCountry id="9195767AACDA43F808E0FC648A2D0588" box="[1012,1096,943,969]" name="Greece" pageId="131" pageNumber="132">Greece</collectingCountry>
), suggesting that new searches in southern
<collectingCountry id="9195767AACDA43F80A09FC188FE705AC" box="[285,386,979,1005]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
must be made in March and April on spring-flowering
<taxonomicName id="2E824D69ACDA43F808E2FC188A2305AC" box="[1014,1094,979,1005]" class="Liliopsida" family="Iridaceae" genus="Crocus" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8ACDA43F808E2FC188A2305AC" box="[1014,1094,979,1005]" italics="true" pageId="131" pageNumber="132">Crocus</emphasis>
</taxonomicName>
and potentially
<taxonomicName id="2E824D69ACDA43F80FEAFC188BF805AC" box="[1278,1437,979,1005]" class="Liliopsida" family="Asparagaceae" genus="Ornithogalum" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8ACDA43F80FEAFC188BF805AC" box="[1278,1437,979,1005]" italics="true" pageId="131" pageNumber="132">Ornithogalum</emphasis>
</taxonomicName>
(
<taxonomicName id="2E824D69ACDA43F80B8BFC3C8F5A0250" authorityName="de Jussieu" authorityYear="1789" box="[159,319,1015,1041]" class="Liliopsida" family="Asparagaceae" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="family">Asparagaceae</taxonomicName>
, the host plant of
<taxonomicName id="2E824D69ACDA43F80914FC3C8CF90250" authorityName="Pisanty &amp; Wood" authorityYear="2022" box="[512,668,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="ornithogali">
<emphasis id="DBF6EAF8ACDA43F80914FC3C8CF90250" box="[512,668,1015,1041]" italics="true" pageId="131" pageNumber="132">A. ornithogali</emphasis>
</taxonomicName>
) in order to determine whether
<taxonomicName id="2E824D69ACDA43F808ECFC3C8A310250" authorityName="WOOD" authorityYear="2024" box="[1016,1108,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDA43F808ECFC3C8A310250" box="[1016,1108,1015,1041]" italics="true" pageId="131" pageNumber="132">A. dacia</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69ACDA43F80F9CFC3C8ABC0250" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[1160,1241,1015,1041]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDA43F80F9CFC3C8ABC0250" box="[1160,1241,1015,1041]" italics="true" pageId="131" pageNumber="132">A. elsei</emphasis>
</taxonomicName>
still persist there. Visual inspection of pollen grains in the scopae of
<taxonomicName id="2E824D69ACDA43F809CEFBD08D5C0274" authorityName="WOOD" authorityYear="2024" box="[730,825,1051,1077]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="dacia">
<emphasis id="DBF6EAF8ACDA43F809CEFBD08D5C0274" box="[730,825,1051,1077]" italics="true" pageId="131" pageNumber="132">A. dacia</emphasis>
</taxonomicName>
indicate that they conform to
<taxonomicName id="2E824D69ACDA43F80F87FBD08A9F0274" authorityName="de Jussieu" authorityYear="1789" box="[1171,1274,1051,1077]" class="Liliopsida" family="Liliaceae" kingdom="Plantae" order="Liliales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="family">Liliaceae</taxonomicName>
<emphasis id="DBF6EAF8ACDA43F80E16FBD08B470274" box="[1282,1314,1051,1077]" italics="true" pageId="131" pageNumber="132">s.l.</emphasis>
(large and elongate oval shaped,
<taxonomicName id="2E824D69ACDA43F80A84FBF48F850218" box="[400,480,1087,1113]" class="Liliopsida" family="Iridaceae" genus="Crocus" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8ACDA43F80A84FBF48F850218" box="[400,480,1087,1113]" italics="true" pageId="131" pageNumber="132">Crocus</emphasis>
</taxonomicName>
pollen is spherical), and hence monocotyledon plants likely represent the pollen host for this species.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDA43F80BD3FB4C8C230284" pageId="131" pageNumber="132" type="etymology">
<paragraph id="E93D36EAACDA43F80BD3FB4C8C230284" blockId="131.[151,1437,151,1511]" pageId="131" pageNumber="132">
<emphasis id="DBF6EAF8ACDA43F80BD3FB4C8F2B02E0" bold="true" box="[199,334,1159,1185]" pageId="131" pageNumber="132">Etymology.</emphasis>
From the Roman province of “Dacia Traiana” which roughly corresponds to the modern-day state of
<collectingCountry id="9195767AACDA43F80BA2FB608F7A0284" box="[182,287,1195,1221]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
. It is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A1986561ACDA43F80BD3FB058FD703A4" pageId="131" pageNumber="132" type="distribution">
<paragraph id="E93D36EAACDA43F80BD3FB058D0A034C" blockId="131.[151,1437,151,1511]" pageId="131" pageNumber="132">
<emphasis id="DBF6EAF8ACDA43F80BD3FB058F0402A9" bold="true" box="[199,353,1230,1256]" pageId="131" pageNumber="132">Distribution.</emphasis>
Southern
<collectingCountry id="9195767AACDA43F80AC4FB048C5002A8" box="[464,565,1231,1257]" name="Romania" pageId="131" pageNumber="132">Romania</collectingCountry>
around
<collectingRegion id="2B46F808ACDA43F80999FB048D6402A8" box="[653,769,1231,1257]" country="Romania" name="Bucuresti" pageId="131" pageNumber="132">Bucharest</collectingRegion>
. Probably present also in northern
<collectingCountry id="9195767AACDA43F80F6EFB048ABF02A8" box="[1146,1242,1231,1257]" name="Bulgaria" pageId="131" pageNumber="132">Bulgaria</collectingCountry>
given the records from Vame Veche which is only
<quantity id="2E7A9B0FACDA43F80910FB388C58034C" box="[516,573,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="131" pageNumber="132" unit="km" value="1.0">1 km</quantity>
from the Bulgarian border.
</paragraph>
<paragraph id="E93D36EAACDA43F80BD3FADD8FD703A4" blockId="131.[151,1437,151,1511]" pageId="131" pageNumber="132">
<emphasis id="DBF6EAF8ACDA43F80BD3FADD8F900371" bold="true" box="[199,501,1302,1328]" pageId="131" pageNumber="132">Other material examined</emphasis>
(
<taxonomicName id="2E824D69ACDA43F80912FADC8CFA0370" authorityName="Scheuchl &amp; Gusenleitner" authorityYear="2009" box="[518,671,1303,1329]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="131" pageNumber="132" phylum="Arthropoda" rank="species" species="elsei">
<emphasis id="DBF6EAF8ACDA43F80912FADC8CFA0370" box="[518,671,1303,1329]" italics="true" pageId="131" pageNumber="132">Andrena elsei</emphasis>
</taxonomicName>
)
<emphasis id="DBF6EAF8ACDA43F809A4FADD8D550371" bold="true" box="[688,816,1302,1328]" pageId="131" pageNumber="132">
<collectingCountry id="9195767AACDA43F809A4FADD8D4E0371" box="[688,811,1302,1328]" name="Greece" pageId="131" pageNumber="132">GREECE</collectingCountry>
:
</emphasis>
Melano [Menalon?] Mountains,
<quantity id="2E7A9B0FACDA43F80FB0FADC8A990370" box="[1188,1276,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="131" pageNumber="132" unit="m" value="1600.0">1600 m</quantity>
,
<collectingRegion id="2B46F808ACDA43F80E1CFADC8BFD0370" box="[1288,1432,1303,1329]" country="Greece" name="Peloponnisos" pageId="131" pageNumber="132">Peloponnese</collectingRegion>
,
<date id="9D3C102AACDA43F80B83FAF08F570314" box="[151,306,1339,1365]" pageId="131" pageNumber="132" value="1987-05-13" valueMax="1987-05-19" valueMin="1987-05-13">1319.v.1987</date>
,
<specimenCount id="FF84FD63ACDA43F80A29FAF08F040314" box="[317,353,1339,1365]" count="6" pageId="131" pageNumber="132" type="female">6♀</specimenCount>
, leg. H. Teunissen, RMNH, on
<taxonomicName id="2E824D69ACDA43F809D7FAF08D760314" box="[707,787,1339,1365]" class="Liliopsida" family="Iridaceae" genus="Crocus" kingdom="Plantae" order="Asparagales" pageId="131" pageNumber="132" phylum="Tracheophyta" rank="genus">
<emphasis id="DBF6EAF8ACDA43F809D7FAF08D760314" box="[707,787,1339,1365]" italics="true" pageId="131" pageNumber="132">Crocus</emphasis>
</taxonomicName>
;
<emphasis id="DBF6EAF8ACDA43F80835FAF18A2F0315" bold="true" box="[801,1098,1338,1364]" pageId="131" pageNumber="132">
NORTH
<collectingRegion id="2B46F808ACDA43F8089AFAF18A200315" box="[910,1093,1338,1364]" country="Greece" name="Dytiki Makedonia" pageId="131" pageNumber="132">MACEDONIA</collectingRegion>
:
</emphasis>
Vardar reg., Papeliste, steppe,
<quantity id="2E7A9B0FACDA43F80B83FA948E840338" box="[151,225,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="131" pageNumber="132" unit="m" value="130.0">130 m</quantity>
,
<date id="9D3C102AACDA43F80BFAFA948F030338" box="[238,358,1375,1401]" pageId="131" pageNumber="132" value="2021-04-16">16.iv.2021</date>
,
<specimenCount id="FF84FD63ACDA43F80A67FA948FF30338" box="[371,406,1375,1401]" count="1" pageId="131" pageNumber="132" type="female">1♀</specimenCount>
, leg. V. Bureš, TJWC;
<emphasis id="DBF6EAF8ACDA43F8098AFA958D560339" bold="true" box="[670,819,1374,1400]" pageId="131" pageNumber="132">
<collectingCountry id="9195767AACDA43F8098AFA958D4B0339" box="[670,814,1374,1400]" name="Romania" pageId="131" pageNumber="132">ROMANIA</collectingCountry>
:
</emphasis>
Comana, Vlasca [Vlașca],
<specimenCount id="FF84FD63ACDA43F80F7DFA948AE90338" box="[1129,1164,1375,1401]" count="1" pageId="131" pageNumber="132" type="female">1♀</specimenCount>
, leg. A.L. Mountandon, labelled “Museum of Paris, coll. J. Vachal 1911”, OÖLM; Valachie [Wallachia], Comana,
<specimenCount id="FF84FD63ACDA43F80F66FA488AF003DC" box="[1138,1173,1411,1437]" count="1" pageId="131" pageNumber="132" type="female">1♀</specimenCount>
, leg.A.L. Mountandon, labelled “Museum of Paris, coll. J. Vachal 1911”, OÖLM;
<emphasis id="DBF6EAF8ACDA43F80838FA6C8DCE0380" bold="true" box="[812,939,1447,1473]" pageId="131" pageNumber="132">
<collectingCountry id="9195767AACDA43F80838FA6C8DC30380" box="[812,934,1447,1473]" name="Turkey" pageId="131" pageNumber="132">TURKEY</collectingCountry>
:
</emphasis>
Taurus, Eskibag,
<date id="9D3C102AACDA43F80F61FA6C8A880380" box="[1141,1261,1447,1473]" pageId="131" pageNumber="132" value="1977-04-18">18.iv.1977</date>
,
<specimenCount id="FF84FD63ACDA43F80FEDFA6C8B790380" box="[1273,1308,1447,1473]" count="1" pageId="131" pageNumber="132" type="female">1♀</specimenCount>
, leg. P.A.J. Ball, OÖLM (
<emphasis id="DBF6EAF8ACDA43F80A23FA008FC703A4" bold="true" box="[311,418,1483,1509]" pageId="131" pageNumber="132">
<typeStatus id="36398848ACDA43F80A23FA008FC703A4" box="[311,418,1483,1509]" pageId="131" pageNumber="132" type="paratype">paratype</typeStatus>
</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>