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<mods:title id="B5B4C7E5008AB308C1D6F5B36417D314">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="85F60E99E35FB00CE83ED758203518BF">Bistroem, Olof</mods:namePart>
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<mods:namePart id="337B9E1118F9CF97705B37FBD06AA509">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="BB9421996668F62967951002807BED9D">Bergsten, Johannes</mods:namePart>
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<paragraph id="A35DBB7C3CAF8B195305CF6EB69A1FBE" pageId="32" pageNumber="33">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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<subSubSection id="9FB78B81FE60F8DBB06BC9DA0C80F11F" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="D6084A9ACCD5F60A175B2DEB49C87172" pageId="32" pageNumber="33">
<taxonomicName id="6F70CA7B3FBE69B538D39DBC44EA36EF" ID-CoL="3RM4W" authority="Regimbart, 1900" authorityName="Regimbart" authorityYear="1900" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="alluaudi">
Laccophilus alluaudi
<normalizedToken id="149923BB5863515731897FCDEF412970" originalValue="Régimbart">Regimbart</normalizedToken>
, 1900
</taxonomicName>
Figs 35-36, 233-234, 392, 531
</paragraph>
</subSubSection>
<subSubSection id="E15B4E3E628D971CCB6228D2A2677D11" pageId="33" pageNumber="34" type="reference_group">
<paragraph id="B5366FACCC7E94AA1C6693B3196AAD0A" pageId="33" pageNumber="34">
<taxonomicName id="004560B4D9B0948D35AB512D7F0C3AC3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">
<pageBreakToken id="8471F5814A5B349DB8F6F0F06A9F3C75" pageId="33" pageNumber="34" start="start">Laccophilus</pageBreakToken>
alluaudi
</taxonomicName>
<bibRefCitation id="DF2961C09970984BA4703B75B240BD24" author="Regimbart, M" journalOrPublisher="Bulletin de la Societe Entomologique de France" pageId="262" pageNumber="263" pagination="371 - 374" title="Diagnoses d'especes nouvelles de Dytiscidae de la Region malgache (Col.)." volume="1899" year="1900">
<normalizedToken id="B8923181C1FD9FF772AB31CCCC90676D" originalValue="Régimbart">Regimbart</normalizedToken>
1900
</bibRefCitation>
: 373 (original description, faunistics);
<bibRefCitation id="309DC8A4C56F062E6E4686134DBA8A30" author="Zimmermann, A" journalOrPublisher="Junk, Berlin" pageId="263" pageNumber="264" title="Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: Schenkling S (Ed.) Coleopterorum Catalogus. Vol. 4 pars 71. W." year="1920 a">Zimmermann 1920a</bibRefCitation>
:16 (catalogue);
<bibRefCitation id="76EB22B62E861F402655D6354684159B" author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="257" pageNumber="258" pagination="63 - 70" title="Dytiscides et gyrinides nouveaux de Madagascar (Coleopteres)." volume="7" year="1955 d">Guignot 1955d</bibRefCitation>
: 67 (discussion);
<bibRefCitation id="12D7811FA3ECB4695A4CAE1ACBC77A57" author="Guignot, F" journalOrPublisher="Le Naturaliste Malgache" pageId="257" pageNumber="258" pagination="139 - 144" title="Nouveaux dytiscides de Madagascar (Coleopteres)." volume="7" year="1955 f">Guignot 1955f</bibRefCitation>
: 141 (faunistics, discussion);
<bibRefCitation id="30A56A9896C6B73C026474DA1F8FCF1D" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
: 544, 548 (description, faunistics);
<bibRefCitation id="4A696C491F9F5A50EFD3887BE10B619A" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Scientifique de Madagascar" pageId="258" pageNumber="259" pagination="75 - 79" title="Dytiscides et gyrinides des Comores." volume="10" year="1959 c">Guignot 1959c</bibRefCitation>
: 76, 77, 78, 79 (discussion, faunistics);
<bibRefCitation id="1DECDA589CC1287C0D5E423FB62F530E" author="Guignot, F" journalOrPublisher="Memoires de l'Institut Scientifique de Madagascar (E)" pageId="258" pageNumber="259" pagination="71 - 73" title="Dytiscides et gyrinides de l'Ile Moheli." volume="10" year="1959 e">Guignot 1959e</bibRefCitation>
: 72 (faunistics);
<bibRefCitation id="2323152CC652B8191EF9E221D19FDB5D" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="653 - 995" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 3." volume="90" year="1961 a">Guignot 1961a</bibRefCitation>
: 931 (faunistics);
<bibRefCitation id="D03BE86A5550DF26ED7D968311F4B79B" author="Betrand, H" journalOrPublisher="Cahiers Office de la Recherche Scientifique et Technique Outre-Mer, Paris" pageId="253" pageNumber="254" pagination="241 - 249" title="Hydrocanthares (excl. Gyrinidae) recueillis a Madagascar (Mission Bertrand, 1960)" year="1971">Bertrand and Legros 1971</bibRefCitation>
: 244 (faunistics, biology);
<bibRefCitation id="3838D89D9A5F8D740B72151DB7508903" author="Wewalka, G" journalOrPublisher="Annalen des Naturhistorischen Museum in Wien" pageId="263" pageNumber="264" pagination="723 - 732" title="Results of the Austrian Hydrobiological Mission, 1974, to the Seychelles-, Comores- and Mascarene Archipelagos. Part VII: Dytiscidae, Gyrinidae (Coleoptera)." volume="83" year="1980">Wewalka 1980</bibRefCitation>
: 724, 726 (faunistics, discussion);
<bibRefCitation id="D12A7496A167B1B29FEE7A0DEC4B2969" author="Rocchi, S" journalOrPublisher="Frustula Entomologica (n. s.)" pageId="262" pageNumber="263" pagination="71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
: 79, 86 (faunistics);
<bibRefCitation id="6EF3FF85BF0DCE7CE484622E9F333A1B" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
: 240 (catalogue, faunistics);
<bibRefCitation id="BFA708BAA4F72456434D7597689B723B" author="Pederzani, F" journalOrPublisher="Atti Dell'Accademia Roveretana degli Agiati, Contributi della Classe di Scienze Matematiche, Fisiche e Naturali" pageId="261" pageNumber="262" pagination="81 - 100" title="On some Madagascan Hydradephaga of the Natural History Museum, London, with the description of five new species of Dytiscidae (ColeopteraHaliplidae, Dytiscidae, Noteridae)." volume="8 B" year="2009">Pederzani and Rocchi 2009</bibRefCitation>
: 95 (faunistics, list);
<bibRefCitation id="D675C62B59EA959D41FFD74F7C437187" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
: 208 (catalogue, faunistics).
</paragraph>
</subSubSection>
<subSubSection id="9231469AB2BFA96EA71D2F3BE1571C9D" pageId="33" pageNumber="34" type="type locality">
<paragraph id="EC6BD0802921527E8638433B6EF8C7A8" pageId="33" pageNumber="34">Type locality.</paragraph>
<paragraph id="7E1502A42D2D5EE62045547A4A22B48F" pageId="33" pageNumber="34">Madagascar: Diego Suarez.</paragraph>
</subSubSection>
<subSubSection id="C6CC8B674ED8B5EAC504B5FAE1E1A987" pageId="33" pageNumber="34" type="type material">
<paragraph id="03A48BC2DC36C8A722845147A6E0A52D" pageId="33" pageNumber="34">Type material, studied</paragraph>
<paragraph id="A292FAFED188DAC0AFAE0E6CFB41F4FF" pageId="33" pageNumber="34">(5 exs). Lectotype (by present designation): male: &quot;Madagascar Diego-Suarez Ch. Alluaud 1893 / male symbol / Cotype / Museum Paris col. Guignot&quot; (MNHN). - Paralectotypes: Same data as in lectotype, but two of the specimens with female symbol (4 exx. MNHN; habitus in Fig. 392).</paragraph>
</subSubSection>
<subSubSection id="9F6CA01B382D07FC1EF2541117D7335C" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="577275EB51582BD505F714D3E3A7F1E7" pageId="33" pageNumber="34">Additional material studied</paragraph>
<paragraph id="A0788EE529A3100027751ECDFAC51570" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
(189 exs.): Madagascar:
<normalizedToken id="8C7180344753A7E90B4691C1EA55B4F2" originalValue="“Forét">&quot;Foret</normalizedToken>
<normalizedToken id="4EF2841704848068A1714DC6B8A9A167" originalValue="dAmbre">d'Ambre</normalizedToken>
Lat -12.4754 Lon 49.2173 coll. Balke &amp; Monaghan, BMNH(E)670568_MSL007: B10, 19.11.04 /
<taxonomicName id="A3830729BE3C5706D73E24FFA0FB27DA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
<normalizedToken id="9B3A22A14AE79491D1ADC779D959FD44" originalValue="Régb">Regb</normalizedToken>
. det. Bergsten&quot; (1 ex. NHRS); same data but &quot;BMNH(E)670572_MSL007: CO2&quot; (1 ex. NHRS); &quot;Antsiranana II Mt
<normalizedToken id="CBE410D0C17AD5806028628E3358C39A" originalValue="dAmbre">d'Ambre</normalizedToken>
, Grande cascade stream parallel to GC in deep very steep gorge, mostly isolated pools 17.11. 2004, N: -
<geoCoordinate id="A45CB95A4BBDE9033FA469EA61102758" direction="east" orientation="longitude" precision="1" value="12.4992">12,49920 E</geoCoordinate>
: 49,17600, 800 m Balke et al&quot; (29 exs. NHRS, 4 exx. MZH); &quot;Antsiranana II Foret
<normalizedToken id="67E494AD55306AEE04E99C503435E56E" originalValue="dAmbre">d'Ambre</normalizedToken>
, small water hole in dry streambed, gardenland at edge of dry forest 19.11. 2004
<geoCoordinate id="A887CA163E62451361C7F0D173A946C5" direction="north" orientation="latitude" precision="1" value="12.4754">N-12,47540</geoCoordinate>
,
<geoCoordinate id="0D52603F9891CFBD20451696CB87E76B" direction="east" orientation="longitude" precision="1" value="49.2173">E49,21730</geoCoordinate>
, 545m leg. Balke&quot; (2 exs. NHRS); &quot;Antsiranana 1 Mtd French Streampool 12.11. 2004
<geoCoordinate id="C9B81671130ABF46CDB6EB7AFC33408B" direction="north" orientation="latitude" precision="1" value="12.3336">N-12,33360</geoCoordinate>
,
<geoCoordinate id="42B6BF539277EC90B5624A287616858F" direction="east" orientation="longitude" precision="1" value="49.3535">E49,35350</geoCoordinate>
, 171 m, leg Balke et al&quot; (12 exs. NHRS); &quot;Antsiranana 1 Mt.
<normalizedToken id="943B247B82C0D8694430241E2240DFE7" originalValue="dAmbre">d'Ambre</normalizedToken>
16.11. 2004,
<geoCoordinate id="33A9495E67B5091225442E361915C066" direction="north" orientation="latitude" precision="1" value="12.5283">N-12,52830</geoCoordinate>
,
<geoCoordinate id="A53D656073F18ECF4524EFDED16E304C" direction="east" orientation="longitude" precision="1" value="49.17253">E49,17253</geoCoordinate>
, 1020 m Balke et al.&quot; (1 ex. NHRS); &quot;Toli, NW Ft Dauphin, forest, watersource, P54E: N: E:: m 19.5. 2006 Bergsten et al. /
<taxonomicName id="3531886A4C5505B2E34F516634F90212" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten/BMNH(E): &lt;?94192&gt; DNA voucher /
<taxonomicName id="7E2142CE328F566CDB240B64E45B9D6A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Toli, NW Ft. Dauphin, creek with gravel, stones and sand in rainforest along the creek, in small water holes and (
<taxonomicName id="2559F54966B149A765C7B2956CB756E1" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Madaglymbus</taxonomicName>
) in waterpool on a large rock with wood and leaves 19.5. 2006 N-24°45.583, E46°51.821, 300 m Bergsten et al.&quot; (2 exs. NHRS); &quot;Ambilobe, Anjiabe Ambony Antsabe: Galoko Mts, hygropetric cascade; alt. 50 m, P2506M N -
<geoCoordinate id="BFF004A6CD9ACF3727C819E57BFA3209" direction="east" orientation="longitude" precision="5" value="13.6093">
13.6093
<geoCoordinate id="872C03DFC322C606C335CEBB8EBBF6A8" direction="east" orientation="longitude" precision="5" value="48.7212">E 48.7212</geoCoordinate>
</geoCoordinate>
, 23.11. 2004 leg. Monaghan, Andriamparany, Balke / BMNH(E) &lt;794160&gt; DNA voucher /
<taxonomicName id="BE43D828F5441FC36B45D1D827C175EA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Anjiabe Ambony, Ambilobe Antsabe stairways-like cascade with vertical steps, exposed, extremely hot day 23.11. 2004
<geoCoordinate id="C9EF97BD589B9509BD00F1CD42BA8D9A" direction="north" orientation="latitude" precision="1" value="13.6093">N-13,60930</geoCoordinate>
,
<geoCoordinate id="D61050389D24CDE837BE8A0D3C440752" direction="east" orientation="longitude" precision="1" value="48.7212">E48,72120</geoCoordinate>
, 303 m Balke et al.&quot; (17 exx. NHRS); &quot;Ambilobe, Anjiabe Ambony Antsabe: waterhole in streambed on clearing: 50 m, P25MD11 N -
<geoCoordinate id="BB0DC7937524EED50E8012A8EE71B55A" direction="east" orientation="longitude" precision="55" value="13.648">
13.648
<geoCoordinate id="2977520FB667B7382B63AEFA3702CCCC" direction="east" orientation="longitude" precision="55" value="48.721">E 48.721</geoCoordinate>
</geoCoordinate>
, 21.11. 2004, leg. Balke et al/BMNH(E) &lt;794189&gt; DNA voucher /
<taxonomicName id="1F654DEFCB793B902DF81F6A6B921C92" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Toli, Taolanaro; Isaky Ivondro, Foret Manangotry, running water, P67B: N: -24.859: E: 46.862: 310 m, 9.4. 2007 leg. Ranarilalatiana et al / DNA voucher BMNH &lt;830767&gt; MSL399:H4 /
<taxonomicName id="2D5B1C08FD4970430F7913133D95EC00" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Ants, Sambava, Marojejy, Marojejy NP: Forest stream P57BI01: N: -14.437: E: 49.773: 464 m, 6.12. 2006 leg. Isambert et al. / DNA voucher BMNH &lt;830690&gt; MSL399:A11 /
<taxonomicName id="10670196869B1C4159C15D7C4E5DCD8B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Toam, Ambatondrazaka, Zahamena; Zahamena NP: Stream P60BI08 N: -17.505: E: 48.724: 1054 m, 30.12.2006 leg. Isambert et al. / DNA voucher BMNH &lt;830696&gt; MSL399:B5 /
<taxonomicName id="D4BD54E0913A0971640B4D988CE34836" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); same data but &quot;P60BI06 N: -17.508: E: 48.724: 1068 m&quot; and &quot;BMNH &lt;830701&gt; MSL399:B10&quot; (1 ex. NHRS); same data but &quot;Zahamena NP, Analamaintsoa 1st stream before Camp 1 within a few m from the GPS point P60BI06, 30.12. 2006
<geoCoordinate id="3BEE4356B300394DE63876ABEF99F053" direction="north" orientation="latitude" precision="1" value="17.50617">N-17,50617</geoCoordinate>
,
<geoCoordinate id="B6AE9E919008CDB3983A4F0971396692" direction="east" orientation="longitude" precision="1" value="48.724">E48,72400</geoCoordinate>
, 1068 m&quot; (4 exs. NHRS); &quot;TOAM Ambatondrazaka Zahamena NP on the way to camp 2 to Fenerive Est &quot;Route des contrebandiers&quot; P60BI29 02.1. 2007, N: -
<geoCoordinate id="A97600B9622C3AF7DFC4A7AB8E12FF48" direction="east" orientation="longitude" precision="1" value="17.54167">17,54167 E</geoCoordinate>
: 48,72183, 1322m leg. Isambert et al&quot; (37 exs. NHRS); same data but &quot;Analamaintsoa, 2nd stream btw Camp 1 and 2, stream of pools 31.12. 2006
<geoCoordinate id="7630E662F6D7B47E8B8637A2BCA05EFC" direction="north" orientation="latitude" precision="1" value="17.5185">N-17,51850</geoCoordinate>
,
<geoCoordinate id="61C10E9D3B57667445EB56C7D2397FE2" direction="east" orientation="longitude" precision="1" value="48.72217">E48,72217</geoCoordinate>
, 1075 m&quot; (4 exs. NHRS); same data but &quot;Analamaintsoa, 3rd stream before Camp 1, stream of pools, almost dry P60BI08, 30.12. 2006,
<geoCoordinate id="7E22906A54F5D14D26EF76C7293AFE9D" direction="north" orientation="latitude" precision="1" value="17.505">N-17,50500</geoCoordinate>
,
<geoCoordinate id="166C2BB9D4CD924D70F99C4269721FCE" direction="east" orientation="longitude" precision="1" value="48.7245">E48,72450</geoCoordinate>
, 1054 m Isambert et al&quot; (10 exx. NHRS); &quot;Montagne des Francais, Lat. -12.3336 Lon. 49.3535 leg. Balke &amp; Otke, BMNH(E)_671210 MSL014; 1/1/1904 / DNA voucher BMNH &lt;671210&gt; MSL014: H02/
<taxonomicName id="3D04430E92354CB1DA1EC2D63F3CA2CE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Saratanana, leg. Lees &amp; Ranaivosolo MNH(E)_672835_MSL028; 1/1/1904 /DNA voucher BMNH &lt;672835&gt; MSL028: B05 /
<taxonomicName id="3B9C4AF4BBFCC144BC5C3B8C847E0B7D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Antsabe Lat -13.6093 Lon 49.7212, Balke leg. BMNH(E) _670700_MSL008 23.9. 2004/1904/DNA voucher BMNH &lt;670700&gt; MSL008: E10/
<taxonomicName id="0CA34E1048B5206C2087433CB0E47E0F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;Ants, Nosy Be: Lokobe R.N.1, 50 m, 2004/DNA voucher BMNH(E) &lt;794171&gt; /
<taxonomicName id="9E1A2D6D17923536652522EC4DCD62D8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
det. Bergsten&quot; (1 ex. NHRS); &quot;R.N.I. Lokobe Nosy Be, 50 m, 15.12. 2004 Ravo&quot; (1 ex. NHRS); &quot;Mahajanga: Boeny Mahavavy Kinkony RS.
<geoCoordinate id="5CF678369BE3354C6E93288C8135521D" direction="south" orientation="latitude" precision="1" value="-16.05648">S16.05648</geoCoordinate>
,
<geoCoordinate id="1B6987C802850FCE86E5C618FAA0A617" direction="east" orientation="longitude" precision="1" value="45.76371">E045.76371</geoCoordinate>
, 55 m.a.o., 5.12. 2009 water net, field, Bergsten et al. (19 exs. NHRS); same data, add: &quot;000000465 NHRS-JLKB&quot; (1 ex. NHRS); &quot;Mahajanga: Melaky: Tsingy de Bemaraha NP,
<geoCoordinate id="16412D36706191C1BDB2F825B8498FB1" direction="south" orientation="latitude" precision="1" value="-19.03572">S19.03572</geoCoordinate>
,
<geoCoordinate id="4CE92FB2945CC12193247F6A341F1F7B" direction="east" orientation="longitude" precision="1" value="44.77507">E044.77507</geoCoordinate>
, 66m m.a.o., 15.12. 2009, water net, field, Bergsten et al.&quot; (12 exs. NHRS); same data but &quot;
<geoCoordinate id="4041E61C97D60976DD9FBC2C635D7E60" direction="south" orientation="latitude" precision="1" value="-18.75643">S18.75643</geoCoordinate>
,
<geoCoordinate id="16AEC9449E288FCACE7F0A2D375AACF0" direction="east" orientation="longitude" precision="1" value="44.71398">E044.71398</geoCoordinate>
, 119 m.a.o., 17.12. 2009&quot; (2 exs. NHRS); same data but &quot;
<geoCoordinate id="2CC356DA65F7AE93E1EC4FE626A2F527" direction="south" orientation="latitude" precision="1" value="-19.03419">S19.03419</geoCoordinate>
,
<geoCoordinate id="5F7FDA1F29617E8BA6BAF5DE7B6C93A2" direction="east" orientation="longitude" precision="1" value="44.77499">E044.77499</geoCoordinate>
, 41 m.a.o., 15.12. 2009&quot; (2 exs. NHRS); same data but
<pageBreakToken id="435987C202F1386D9B16F7880E964C27" pageId="34" pageNumber="35" start="start">&quot;</pageBreakToken>
<geoCoordinate id="9D1AEF3719F691898B6CBB4A64635D4C" direction="south" orientation="latitude" precision="1" value="-18.75724">S18.75724</geoCoordinate>
,
<geoCoordinate id="896DD642158944421D381CCF04BC667D" direction="east" orientation="longitude" precision="1" value="44.71239">E044.71239</geoCoordinate>
, 72 m.a.o., 17.12. 2009&quot; (3 exx. NHRS); same data, add: &quot;000000468 NHRS-JLKB&quot; (1 ex. NHRS); same data but &quot;
<geoCoordinate id="42E5EB1A70DA4C006631428800A6F08D" direction="south" orientation="latitude" precision="1" value="-19.14114">S19.14114</geoCoordinate>
,
<geoCoordinate id="9510D6D1F50765555776AA33431022D9" direction="east" orientation="longitude" precision="1" value="44.81245">E044.81245</geoCoordinate>
, 45 m.a.o., 14.12. 2009/000000466 NHRS-JLKB&quot; (1 ex.NHRS); &quot;Prov. Antsirarana, P.N. Montagne
<normalizedToken id="7DC40D6139DAB84ED12283485D28F91A" originalValue="dAmbre">d'Ambre</normalizedToken>
, elev. 960 m 26-29.1. 2001/ N -12*30'52&quot; E 49*10'53&quot; leg. Irwin &amp; al, malaise trap&quot; (1 ex. CAS); &quot;Prov. Antsirarana, P.N. Montagne
<normalizedToken id="4EAE26803905C7331422F0D9E91BEE58" originalValue="dAmbre">d'Ambre</normalizedToken>
, elev. 1125 m 30.5.-6.6. 2001/ N -
<geoCoordinate id="25BF4FD66CDF48B86C820298F9BC5445" direction="east" orientation="longitude" precision="15" value="12.520278">
12°31'13&quot;
<geoCoordinate id="69B97DDE5EFF4A1EF485ED2604ACEA24" direction="east" orientation="longitude" precision="15" value="49.17917">E 49°10'45&quot;</geoCoordinate>
</geoCoordinate>
leg. Irwin &amp; Hala, malaise trap&quot; (1 ex. CAS); &quot;Prov. Antsirarana, P.N. Montagne
<normalizedToken id="3A15A5FE6F9BB79277CBFE3E919DE282" originalValue="dAmbre">d'Ambre</normalizedToken>
, elev. 1125 m 21-26.4. 2001/ N -
<geoCoordinate id="6E005671D1CA487FAED89ABE55E2119E" direction="east" orientation="longitude" precision="15" value="12.520278">
12°31'13&quot;
<geoCoordinate id="C2B187363630D255A78478B9FE6ED889" direction="east" orientation="longitude" precision="15" value="49.17917">E 49°10'45&quot;</geoCoordinate>
</geoCoordinate>
leg. Irwin &amp; Hala, malaise trap&quot; (1 ex. CAS); &quot;Prov. Antsirarana, Sakalava beach, dwarf littoral forest 10 m, 13-20.8. 2001/ N -
<geoCoordinate id="1473C2EFD649A41080D76319E5DA6AB2" direction="east" orientation="longitude" precision="15" value="12.262777">
12°15'46&quot;
<geoCoordinate id="4176862964C9E577971910FA68E2789F" direction="east" orientation="longitude" precision="15" value="49.397503">E 49°23'51&quot;</geoCoordinate>
</geoCoordinate>
Irwin &amp; Hala leg., malaise trap - across sandy trail&quot; (1 ex. CAS); &quot;Andjamangirana (Majunga) 19.10. 2001/stream in dry forest, upstream. Rice field area (road to Tsaratanana) 200 m a.s.l., 30,8 °C, 0.008 mS/cm/Gerecke &amp; Goldschmidt leg.&quot; (1 ex. BMNH, 1 ex. MZH); &quot;Anjozorobe (Antananarivo) Ravoandrina Riv. Ampanakamonty 21.7.2001 / 1280 m asl, 12,8 °C, 0,078 mS/cm/Gerecke &amp; Goldschmidt leg.&quot; (1 ex. BMNH); &quot;SE-Mad.
<normalizedToken id="83C484A3F86AF9B8483378C1ADC94178" originalValue="Rés">Res</normalizedToken>
. Nat. Integr. de Andohahela (NW Ft. Dauphin) Parcelle 1 (versante E) - 300 m foresta pluviale 26.5. 1991/Bartolozzi, Taiti, Raharimina leg. /
<taxonomicName id="AB712D1D8C9A4C8F006ED89607AE87E2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
<normalizedToken id="212CDA8663794AC0E8F75CC84F8C2E7A" originalValue="Rég">Reg</normalizedToken>
. det.
<bibRefCitation id="FF1589FF3C251E057094EA2B0854F946" author="Rocchi, S" journalOrPublisher="Frustula Entomologica (n. s.)" pageId="262" pageNumber="263" pagination="71 - 89" title="Contributo alla conoscenza degli Haliplidae e dei Dytiscidae del Madagascar con descrizione di due nuove specie (Coleoptera)." volume="14" year="1991">Rocchi 1991</bibRefCitation>
&quot;(2 exs. CSR); &quot;E-Mad. Ampamoho nr Andilamena 1200-1300 m asl, 18-20.1. 1995 Dunay &amp; Janak&quot; (1 ex. MZH); &quot;Fiananrantsoa, Mania River S Ilaka, 900 m 27 rd km NNW Ambositra 23.10. 2001 Schuh leg.&quot; (1 ex. NMW); &quot;Foret de Fito, ex. coll. Dr. Breuning&quot; (1 ex. MRAC); &quot;Prov. de Tamatave, Foret de Perinet 17.7. 1970 Pederzani /
<taxonomicName id="BFFFC0509DF622206B2144623A7AE934" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus lateralis" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lateralis">Laccophilus lateralis</taxonomicName>
Sharp det. Pederzani&quot; (1 ex. CSR; determination uncertain).
</paragraph>
</subSubSection>
<subSubSection id="FBF57B4FF7915BE7E20C2198E4BC474E" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="D2FD9E661CB43560A1D365186C2B3B44" pageId="34" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="880CEDDE82A18FCA24912A9988F42714" pageId="34" pageNumber="35">
<taxonomicName id="C1767D9071F922CFA56126511B75C5EF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
forms together with
<taxonomicName id="09A279B46044CD8BAF6D03CED3401B8F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
,
<taxonomicName id="37070C24E0807B30A5BB9C10ADD4327D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tigrinus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tigrinus">Laccophilus tigrinus</taxonomicName>
,
<taxonomicName id="36CD511B07993453A426EFF7CFB054E8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pseustes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pseustes">Laccophilus pseustes</taxonomicName>
and
<taxonomicName id="BB66734FC9F379F95BDE0470D4F72D44" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
a distinct group of species characterized by quite similar colour pattern of body and male genitalia exhibiting same ground plan. Penis of
<taxonomicName id="1F9FDA261FC55963F56245BC447C91FB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
,
<taxonomicName id="C133DE8124145BA364652FCE20065379" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
and
<taxonomicName id="E1CD2D212D75E8111CBA42B370CB8386" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
is not evenly curved in lateral view as in
<taxonomicName id="5D813D951F73F089571E8A17EF17ECFC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus tigrinus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="tigrinus">Laccophilus tigrinus</taxonomicName>
, but angled. Longitudinal markings of
<taxonomicName id="C5AF84C573B7FE8455053EDC2430B2C5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
reach humeral region while in
<taxonomicName id="AF6A4A0EB9D9F805525849C2A5654D3A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
and
<taxonomicName id="444445F08CEB33531422AE2447565CCC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
corresponding markings fade away before reaching humeral region. Finally, penis (dorsal aspect) in
<taxonomicName id="649971563DB7B091433723CF68740906" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
is strongly twisted, while almost straight in
<taxonomicName id="EA46AC68B0AE902001CA66E0B1ABE2E5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus furthi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="furthi">Laccophilus furthi</taxonomicName>
. Furthermore
<taxonomicName id="DADB23FC2E931CE06AFB04C6EC49817C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
penis is in lateral view medially slightly depressed and immediately after angle towards apex there is a minute, sharp knob which is lacking in
<taxonomicName id="98389C9426CC013AEB59AD49CC124FF2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="5EAA09F222787E03A1534EB3CC9043D9" pageId="34" pageNumber="35" type="description">
<paragraph id="60EF2722F63E6904E068B035AD70FD0F" pageId="34" pageNumber="35">Description.</paragraph>
<paragraph id="0B81B8E2541E7F1AF37BE1558E0A4487" pageId="34" pageNumber="35">
(See description of
<taxonomicName id="9F711A3F672ABA3734A383FF808F72C8" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
; only diagnostically important differences noted): Body length 3.4-3.9 mm, width 1.9-2.1 mm. Dorsal, colour pattern (Fig. 392); slightly less pronounced in comparison with
<taxonomicName id="06BF9BC90862827FD3A81C27F2D0BE5F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
(Fig. 390).
</paragraph>
<paragraph id="3C2BE45266B60F9545BBC181F6F40D8C" pageId="34" pageNumber="35">
Elytra: Longitudinal markings brownish to ferrugineous; less pronounced in comparison with material of
<taxonomicName id="118D660C912609E9ABFADE2BE67AFBF3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
; especially humeral region with reduced dark markings (Fig. 392).
</paragraph>
<paragraph id="74D9BB05F109780B85E9ABC66A004ACB" pageId="34" pageNumber="35">Ventral aspect: Apical ventrite as in Fig. 35.</paragraph>
<paragraph id="EFB1560F66F28AD1DF6587ACD38872B2" pageId="34" pageNumber="35">
Male genitalia: Penis in lateral aspect from base region, almost straight to extreme apex, which is slightly bent upwards; in dorsal aspect, penis bent at right but less so than in
<taxonomicName id="EC94B5959157455875C66AB29E939A14" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus comes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="comes">Laccophilus comes</taxonomicName>
(Figs 233-234).
</paragraph>
<paragraph id="821712FDC48660ACF56BFB66FC9E8A2E" pageId="34" pageNumber="35">Female apical ventrite as in Fig. 34.</paragraph>
</subSubSection>
<subSubSection id="4CE05056DBC8BE7D2D4D4D98E571E746" pageId="35" pageNumber="36" type="distribution">
<paragraph id="969D3337E3F8B69E41909FBB69B46811" pageId="35" pageNumber="36">
<pageBreakToken id="24D8AA6BF0D6D2A5F2C5D25004E72342" pageId="35" pageNumber="36" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph id="8E0FE60AA21D5A28095CBFC29FBF915C" pageId="35" pageNumber="36">
Madagascar (Fig. 531). Also reported from the Comoros (e.g.
<bibRefCitation id="C2940FC08C4F2538902D776DDB3D1385" author="Guignot, F" journalOrPublisher="Bulletin de l'Institut Royal des Sciences Naturelles de Belgique" pageId="258" pageNumber="259" pagination="1 - 8" title="Haliplidae et Dytiscidae nouveaux du Parc National de la Garamba. Deuxieme serie (1)." volume="34" year="1958">Guignot 1958</bibRefCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="C3F7E5D0A55048CFFC107DCF860F1ED3" pageId="35" pageNumber="36" type="collecting circumstances">
<paragraph id="580F4DBB51D4023BAD8BC6F5794F110B" pageId="35" pageNumber="36">Collecting circumstances.</paragraph>
<paragraph id="C48E33AEF0AA86727995601A5992F3D6" pageId="35" pageNumber="36">
Label data indicate that
<taxonomicName id="E02F4B9200D09A632667FD76F945B365" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus alluaudi" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="alluaudi">Laccophilus alluaudi</taxonomicName>
occurs in both standing and running waters. Collected in a creek with gravel, stones and sand in rainforest along the creek, in small water holes and together with
<taxonomicName id="096F34EC151A426ED6BF0DF6C89BAF18" class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Madaglymbus</taxonomicName>
in water pool on a large rock with wood and leaves.
</paragraph>
</subSubSection>
</treatment>
</document>